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eogrāfiski raksti folia geographica xii - Ģeogrāfijas un Zemes zinātņu ...

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27<br />

NATURE RESEARCH<br />

3.5. Vegetation period<br />

Characteristic of Latvia’s climate is a half-year long vegetation period, when, <strong>un</strong>like the<br />

other half-year (dormant period), the plants are growing, blooming, producing fruits and<br />

ripening them. In terms of phenology it might be the time from the <strong>un</strong>folding of leaves to their<br />

falling [Schaefer, Tischler 1983; Leser 1997], yet more often this stage is restricted to climatic<br />

or agro-climatic interpretation – in the mid-latitudes it is associated with the period when the<br />

mean daily temperature exceeds 5ºC [Schultz 1995] or with the number of days when the mean<br />

daily temperature exceeds 10ºC [Ozenda 1994; Hendl <strong>un</strong>d Liedtke (Hrsg.) 1997].<br />

In accordance with the phenological observations [Latvijas dabas <strong>un</strong> vēstures kalendārs<br />

1967-2004], in most of Latvia the first signs of vegetation renewal (bud burst of several tree and<br />

shrub species, blooming of the willow, and the occasional decoration of bare spots by the first<br />

spring ephemeroids) are already seen in early spring, when the daily temperature rises above<br />

0ºC, and even earlier. The remnants of summer greenness and some blooms may be even seen<br />

<strong>un</strong>til the first winter frost in November. Yet, in general, the vegetation matches the features<br />

referred to in the definition of the vegetation period, when the daily temperature exceeds 5ºC. In<br />

most cases it sets in about 2-3 weeks after vernal equinox (mid-April) and comes to an end a<br />

month after autumn equinox (end of October).<br />

In the beginning of the described period the landscape outside coniferous forests is still<br />

dominated by gray color. Yet, the vegetation processes are already getting more intensive.<br />

Commonly the circulation of birch sap has started, and the shrub species widespread in Latvia,<br />

such as alder and hazel-trees, are blooming and the grass germinates. Soon after that leaves start<br />

to <strong>un</strong>fold, first the shrubs, then the trees. In the herb layer, the spring ephemeriods of nemoral<br />

zone (mainly Hepatica nobilis, Corydalis Halleri, Anemone nemorosa and A.ran<strong>un</strong>coloides) far<br />

and wide form blue, white and sometimes yellow flower carpets, which appear as bright<br />

colourful clusters in the generally gray landscape. In a short time the mean daily temperature<br />

rises by up to 10ºC (it usually occurs by the end of the first decade of May), all trees and shrubs<br />

have burst into leaf, the grasslands have become green.<br />

Thus, the beginning of vegetation period, a season less than one month in duration, which<br />

is typical of the change from a gray (except coniferous forests) early spring landscape with melt<br />

waters in the bare spots and isolated snow patches to a green and blooming landscape, might<br />

most appropriately be named Full spring. Afterwards, when daily air temperature rises above<br />

10ºC, the main phase of vegetation period sets in and lasts <strong>un</strong>til the end of the culminant phase<br />

of net radiation (end of July), encompassing the three following stages of summer (Figure 5).<br />

The first stage of the main phase of the vegetation period is pre-summer, a season of the<br />

renewal of highly active life processes, which, despite the still possible “return of cold”,<br />

distinguishes itself with a rapid increase of the number of blooming species and high biomass<br />

production. It follows from the phenological studies carried out in Estonia that during the next<br />

stage – early summer, the number of blooming species in the forest biogeocenosis of boreonemoral<br />

type (the dominant tree species is spruce, but in gro<strong>un</strong>d cover and shrub layer and tree<br />

stand, too, the species of deciduous broad-leaved forest hold a significant position) reach the<br />

maximum (shortly before/after summer solstice). Apparently, early summer and the following<br />

high summer are the main seasons of the annual course of biomass production, decomposition<br />

of dead biomass and biosphere-atmosphere gas exchange [Schultz 1981].<br />

In high summer the state of vegetation is much the same. Yet, as regards zonal forests in<br />

July, the phenospectra of the herb layer generated at the Vooremaa Forest Ecology Station in<br />

the growing season of 1973 showed hardly any not-yet blooming species, while the number of<br />

blooming species was gradually decreasing. Some time later, almost half of herb-layer species<br />

will have already come to the phenophase of developing and ripening of fruits or even seed<br />

dispersal, and a quarter will have reached pre-dormancy quiescence [Kannukene 1979]. In other<br />

words, during high summer the processes which ensure reproduction and dissemination of<br />

species and prepare the plants for survival during the period of winter dormancy and for<br />

resuming activity during the next vegetation period, gradually come to the forefront. Moreover,<br />

the biomass production decreases essentially and green mass dies off more intensively. Later on,<br />

in the second half of vegetation period, these processes are the main ones.

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