able physiognonies in this species mightbe considered analogous to <strong>the</strong> heightforins <strong>of</strong> smooth cordgrass whi ch del ineatezones in salt narshes.Where zonation, as an organizationalexpression <strong>of</strong> species distributions, actuallyexists in <strong>the</strong> tidal freshwater marshhabitat, it is probably controlled by acombination <strong>of</strong> physical variables and eco-1ogi cal processes. Prel irninary evidencesugyests that <strong>the</strong>re may be a certain degree<strong>of</strong> consistency in <strong>the</strong> zonation foundin tidal freshwater oarshes. However, <strong>the</strong>extent <strong>of</strong> this patterning with respect tovarious community types, as we1 1 as itsregularity frorn location to location, isunknown.species within a general structural frameworkfor tidal freshwater wetlands. As afirst approximation to community structurewithin this habitat, most <strong>of</strong> <strong>the</strong> comnon1.yoccurring vegetation fa1 1 s into one <strong>of</strong> <strong>the</strong>fol lowing categories: (1) submerged orfloating-leaved plants, ( ) emergentplants with basal leaves and/or leaflessstems, (3) emergent or damp soil herbswith stems bearing alternate or opposite1 eaves ,. (4) grass1 ike or rush1 i ke ol ants,and (5) broad-1 eaved shrubs and trees(Magee 1381). By recognizing <strong>the</strong> aporoximatemodal distributions <strong>of</strong> common plantspecies or communi ty-type indicatorspecies within <strong>the</strong>se structural subgroups,a typical marsh pr<strong>of</strong>ile can be visual izedand described.<strong>The</strong> Marsh Pr<strong>of</strong>ile<strong>The</strong> marsh pr<strong>of</strong>ile de~icted in Fiqure13 is most characteristic <strong>of</strong> mid-AtlanticExcept for <strong>the</strong> most obvious community tidal marshes. Seds <strong>of</strong> submerqed, rootedtypes, it is difficult to place a given aquatic plants (see Section 2.6) make upBALD CYPRESS - BLACK GUMWAX-MYRTLEWILD RICEGlA NT CUTGRASS4 CATTAILSEDGES - RUSHESBIG CORDGRASSIIIIIROSE-MALLOWJEWELWEED3 BURMA RlGOL DTEARTHUMBSMARTWEED1ARROW-ARUMPICKERELWEEDI IIIIII I I I IHIGH MARSHWOODED SWAMP1FORESTED UPLANDFigure 13.Characteristic pr<strong>of</strong>ile <strong>of</strong> mid-At1 antic tidal freshwater marsh.25
an invisible, suspended mat <strong>of</strong> vegetationat <strong>the</strong> foot <strong>of</strong> <strong>the</strong> marsh where inundationis constant. Verging with this subtidal1 ayer and extending variable distances uponto <strong>the</strong> muck surface <strong>of</strong> <strong>the</strong> marsh are ahost <strong>of</strong> fleshy-leaved, emergent macrophytes:spatterdock, arrow-arum, pickerelweed,and arrowhead. <strong>The</strong>se species, pluswild rice, big cordgrass, and numeroussedges and rushes, comprise <strong>the</strong> bulk <strong>of</strong>1 ow marsh vegetation. <strong>The</strong> transition fromlow to high marsh is generally marked byan increase in species number, presumahlydue to reduced periods <strong>of</strong> inundation. <strong>The</strong>predominant components <strong>of</strong> <strong>the</strong> high marshzone include low swards <strong>of</strong> tangled grass(rice cutgrass), erect or sprawl ing herbaceousthickets (burmarigold, tearthumb,jewelweed, smartweed), tall grasses orgrass1 ike plants (giant cutgrass, wildrice, cattail, sweetflag), and shrub1 ikethickets (rose-ma1 low, swamp rose, loosestrife). <strong>The</strong> most 1 andward extent <strong>of</strong> <strong>the</strong>marsh usually coincides with <strong>the</strong> mean highwater mark and is indicated structurallyby a dense wall <strong>of</strong> shrubbery (wax-myrtle)and associated overstory (bald cypress,black gum, red maple) and understory (jewelweed,Asiatic spiderwort) species.Variations on this scheme are numerous,and are <strong>of</strong>ten associated with <strong>the</strong>physiographic characteristics <strong>of</strong> <strong>the</strong> marshpr<strong>of</strong> i 1 e. One physiographic feature consistentlyfound in <strong>the</strong> tidal freshwaterhabitat is an elevated levee forming <strong>the</strong>crest <strong>of</strong> <strong>the</strong> channel bank. This featurecreates a niche for facultative hydrophytesor less water-tolerant specieswithin <strong>the</strong> low marsh zone. Plant speciescommonly taking advantage <strong>of</strong> <strong>the</strong> leveenichein Virginia are water-hemp, commonthreesquare, squarestem spi ke-rus h, rosemallow,giant ragweed, and o<strong>the</strong>r highmarsh herbs. Simi 1 arly, subsidence areaswithin <strong>the</strong> high marsh zone can create aniche for obl igate hydrophytes. Thisphenomenon can usually be attributed togeologic maturation <strong>of</strong> riverine and estuarinewetlands (see Section 1.6).Metzler and Rosza (1982) describe amarsh pr<strong>of</strong>i le for nor<strong>the</strong>astern At1 anticcoast tidal freshwater wet1 ands. For comparison,it is presented in Figure 14.<strong>The</strong> definition and extent <strong>of</strong> zones isquite similar to that described for mid-At1 antic marshes, a1 though <strong>the</strong>re aresignificant differences in species composition.fMHWI I I1 I II I 1I I 1 C__IIIID I EIZONE DESCRIPTION SPECIES CHARACTERISTICSA SUBTIDAL PONDWEEDS, WATERWEED, HORNWORTB LOWER INTERTIDAL ARROWHEADS, SEEDBOX, BULRUSHESC MID-TIDAL MARSH BULRUSHES, WATER HEMP, WATER PARSNIP,BORDERSNEEZEWEED, SMOOTH BURMARIGOLD, WILDRICE, PICKERELWEED, ARROWHEADSD HIGH MARSH SWEETFLAG, CATTAIL, SWAMP ROSE, REEDBENTGRASSE UPLAND RED MAPLE, WATER WILLOW, ARROW-WOODFigure 14. Nor<strong>the</strong>astern marsh pr<strong>of</strong>ile. Modified from bletzl er and Rosza (1982).
- Page 2 and 3: LibraryNational Wetlands Restarch C
- Page 4 and 5: PREFACEThis report is part of a ser
- Page 6 and 7: Page.......6.1 Species Composition
- Page 8: TABLESNumber12345678910111213141516
- Page 13 and 14: (DURINGLowFLOWCONMTIONS)Figure 1. T
- Page 15 and 16: MERRIMACK RlVERNORTH RlVERCONNECTIC
- Page 17 and 18: Figure 3,Tidal freshwater marsh on
- Page 19 and 20: In general the river systems of the
- Page 21 and 22: and Powers (1978) measured the cati
- Page 24 and 25: CHAPTER 2, COMMUNITY COMPONENTS: PL
- Page 26 and 27: Table 4.Common species of vascular
- Page 28 and 29: Nyssa sylvanticaN~s;;~;;uaticaMediu
- Page 30 and 31: progresses, some pl ants wi 11 be o
- Page 32 and 33: many of the tidal freshwater marshe
- Page 34 and 35: Rose-Mallow \i':JewelweedFigure 10.
- Page 38 and 39: 2.4 FACTORS CONT3OLLIVG PLANT DEMOG
- Page 40 and 41: Table 5. Species composition of fiv
- Page 42 and 43: of sorne form (e.g . , major commun
- Page 44 and 45: naiads, wild celery, and dwarf arro
- Page 46 and 47: govern the seasonal blooms and succ
- Page 48 and 49: CHAPTER 3. ECOSYSTEM PROCESSES3.1 P
- Page 50 and 51: Table 7. Continued.Peak standing cr
- Page 52 and 53: ing the sampling effort to just one
- Page 54 and 55: and completely decompose in 4 to 5
- Page 56 and 57: SOURCES(LARGELY INORGANIC, OXIDIZED
- Page 58 and 59: is an approximate 100% turnover of
- Page 60 and 61: solved carbon as particulate carbon
- Page 62 and 63: polychaete, Scolecol epides viridus
- Page 64 and 65: 4.3 MARSH PLANT INSECT COMMUVITYPub
- Page 66 and 67: minnows occupy midwater and benthic
- Page 68 and 69: Tab1 e 11.Characteristics of anadro
- Page 70 and 71: Figure 24. Striped bass, the most i
- Page 72 and 73: Table 13.Numerically dominant fishe
- Page 74 and 75: - Table 15. Fishes using tidal fres
- Page 76 and 77: to the ocean. The freshwater reside
- Page 78 and 79: Lizards and 1 izard-1 ike reptiles
- Page 80: Tab1 e 17. Efficiency of secondary
- Page 83 and 84: The birds of tidal freshwater wetla
- Page 85 and 86: Table 18. Distribution of waterfowl
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Table 29. Breadth of diet of select
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other rails) gather to feed on the
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the effects of seabirds on nutrient
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the various soecies of mammals in t
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h?ve not been directly studied. Stu
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CHAPTER 9- VALUES, ALTERATIONS, AND
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estuary to the rlext dependinq upon
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Section 3.3). Simpson et al. (1981)
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Tab1 e 24. Hypothetical comparisons
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Table 24.Concluded.Characteristics
- Page 107 and 108:
REFERENCESAdams, D.A. 1963. Factors
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J.E. Clark eds. Wetland functions a
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Sci. Rep. 40.4. Nat. Resour. Inst.U
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University Press, Ithaca, N.Y.345 p
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fish fauna of Tivoli Bays. Bard Col
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Patrick, eds. Two studies of Tinicu
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Sci . 16:77-78.Penney, J.T. 1950, D
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Smith, B.A. 1971. The fishes of fou
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R.L. Simpson, eds. Freshwater wet1
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APPENDIX APlants of the Tidal Fresh
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PontederiaceaePontederia cordataZos
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Ona graceaeJossiaeare~en~LudwuriaDa
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APPENDIX BFISH OF TIDAL FRESHWATERS
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Amiidae -bowfins&Labrvf inElopidae
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Geographic Salinity Relativerange r
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NameCatostomidae -suckersGeographic
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Name GeographicrangeSalinityrangeRe
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NameGeographicrangeSalinity Relativ
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Centrarchidae -sunfishesName Geogra
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Percidae -perchesEtheostomafusiform
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Name Geographic Salinity Relativera
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Reference Numbers Key1. Adams 19702
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l m L o .Arl- >4J 3o m uc, mEU mmm
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Family / SpeciesRegionStatusFood ha
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- fIn-f W--7.40 -> 0171- Lw m mUtu
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APPENDIX D:Avifauna of tidal freshw
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Hooded merganser(LoDbodvtes-)Common
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FamilySpeciThreskiornithidae - ibis
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Family / SpeciesRegionSeasonStatusH
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Cathartidae - vulturesRegionDIURNAL
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CULLS, TERNS, KINGFISHERS, AND CROW
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ARBOREAL BIRDSFamily / Species Regi
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Family / Species Region Season Stat
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Family / Species Region Season Stat
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Family / Species Region Seasonnorth
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Family / SpeciesMarsh wren(Long-bil
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drQFamily / SpeciesFringilidae - fi
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APPENDIX E:Mammals of tidal freshwa
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Family / SpeciesRegionStatusFood ha
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50272 -to1 1REPORT DOCUMENTATION2.3