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Taxonomy and Ecology Of Inland Sand Dune ... - Mycorrhizae

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11wall thickening or spore inner wall.Width <strong>and</strong> length measurement of thesubtending hyphae are used in speciesidentification.c. SPOROCARP MORPHOLOGYSporocarps are formed in peridial <strong>and</strong>possibly in glebal hyphae of Sclerocystis<strong>and</strong> in some species of Glomus(Gerdemann <strong>and</strong> Trappe, 1974).Sporocarps are not known inEntrophospora, Gigaspora, <strong>and</strong>Scutellospora (Ames <strong>and</strong> Schneider,1979; Berch, 1985; Walker <strong>and</strong> S<strong>and</strong>ers,1986). Sporocarps typically are absentin Acaulospora. The exceptions are A.myriocarpa Spain, Sieverding <strong>and</strong>Schenck, in which spores are in a cluster(Schenck et al., 1986), <strong>and</strong> A.sporocarpa Berch which has anaggregation of spores in a network ofhyphae (Berch, 1985). Externalsporocarp colorranges from white to brown. However,internal sporocarp color ranges fromwhite to black in Glomus <strong>and</strong> brown inSclerocystis species. Sporocarp shape isirregular in Glomus, but it is globose,subglobose or ellipsoid in Sclerocystisspecies (Morton, 1988). Two terms areused to describe hyphal arrangement insporocarps: the term peridium is usedwhen the hyphae form a loosely ortightly interwoven network on thesurface of the sporocarp, <strong>and</strong> the termglebal is used when the hyphae form amatrix in the sporocarp. Epigeoussporocarps can be large <strong>and</strong> brightlycolored. Hypogeous sporocarps aresmall <strong>and</strong> can be blend with soil texture<strong>and</strong> color. Sporocarps of diameters lessthan 1 mm can be separated by wetsieving <strong>and</strong> centrifugation (Powell <strong>and</strong>Bagyaraj, 1984).2. SPORE DISPERSALAll VAM fungi depend on passivemeans of spore dispersal. Wind <strong>and</strong>animals are good vectors for sporedispersal (Friese, 1984; MacMahon <strong>and</strong>Warner, 1984). In arid ecosystems,wind might be the most importantdispersal agent for VAM inoculum(Trappe, 1981; MacMahon <strong>and</strong> Warner,1984; Warner et al., 1987). However,animals are the major vectors for VAMinoculum in mesic habitats (Marx, 1975;

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