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Continental trace fossils and museum exhibits - Geological Curators ...

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‘forgotten’. The Natural History Museum in London(BMNH) has a good collection of <strong>trace</strong> <strong>fossils</strong> <strong>and</strong> yetfor years these specimens were in a state which wasless than user-friendly. Only when resources wereavailable, in the form of unpaid volunteers supervisedby hard-pressed <strong>museum</strong> staff, did the curatoriallevel of the collection rise <strong>and</strong> improve to a moreusable state. The collection is small relative to manyother taxa in the department, only occupying about1.6 % of the storage area of one floor, not includinglarger <strong>trace</strong>s like dinosaur tracks/trackways whichhave to be stored elsewhere within the Museumcomplex, nor other smaller <strong>trace</strong>s which are keptalongside their known <strong>and</strong> recognised formingorganisms.A further consequence of ‘Cinderella’ groups like<strong>trace</strong> <strong>fossils</strong> is that a good collection made by anenthusiast is unlikely to be received with muchenthusiasm by a <strong>museum</strong> whose storage space islimited. This may lead to the collection being lost toscience.However, <strong>trace</strong> <strong>fossils</strong>, especially when placed intheir sedimentological <strong>and</strong> palaeobiological context,are important environmental <strong>and</strong> ecologicalindicators. They are useful for trying to determinesuch things as the mode of life, even if the organismitself was either not preserved or unknown. Forexample, the carpoid Rhenocystis from the Devonianof Bundenbach in Germany belongs to a group thathave been the subject of debate concerning theirfunctional morphology <strong>and</strong> mode of life; even suchbasic characteristics as life orientation are contended.Jefferies (1984) provided detailed descriptions of hisinterpretation of the movement of the organism thatis now supported by the discovery of severalspecimens of Rhenocystis at the end of their trails(Sutcliffe et al. 2000). Even when the ichnotaxa arewell documented <strong>and</strong> their relationships are known,e.g., the Diplichnites-Cruziana-Rusophycussequence, the evidence of the actual producers isscanty. Although the producing organism may befound at the end of the trail (for example, a trilobite),there are other organisms in entirely separatetaxonomic groups that produce very similar <strong>trace</strong><strong>fossils</strong> (see discussion in Whittington 1992, p. 39).Correlating the two is very difficult, but the interest<strong>and</strong> benefits of doing so are great. So why is thereonly limited interest in <strong>trace</strong> <strong>fossils</strong>? The reasonsmight include any or all of the following.• Collections failure - people do not recognisethem or they cannot be extracted from theirlocalities (Donovan et al. 2006).• Too big to collect <strong>and</strong>/or store.• Not aesthetic - <strong>trace</strong> <strong>fossils</strong> are generallyconsidered unattractive, although ichnotaxa suchas Zoophycos <strong>and</strong> Paleodictyon, for example, arenonetheless pleasing to the eye.• Not identifiable - they require a good deal ofexpertise to appreciate their importance (althoughthis may be said for many fossil groups).Suggestions for organising <strong>trace</strong> fossilcollectionsTrace <strong>fossils</strong> are morphological <strong>and</strong> functional entitieswith the overlay of a Linnean style of classification,distinguished from true biological taxonomy by theprefix ‘ichno’ - ichnogenus or ichnospecies. They arenot body <strong>fossils</strong> in the biological sense, but are the<strong>trace</strong>s produced by the organisms in the course ofliving, that is, sedimentary structures. In order that acollection can be useful some form of organisationneeds to be imposed upon the specimens. Variousalternatives are possible.• Alphabetical (commonly by ichnogenus) – Thesimplest organisation <strong>and</strong> one that enables aconstant layout to be made of ichnogenera.Revision of the interpretation of the <strong>trace</strong>s wouldnot lead to major problems of reorganisation forcurators, unless an ichnogenus is ‘split’ orsynonymised. This is currently the way in whichspecimens are organised in the BMNH. It has theadvantage of not requiring constant reorganisationaccording to differing interpretations,<strong>and</strong> is usable by expert <strong>and</strong> novice alike.• Stratigraphical – Trace <strong>fossils</strong> can be restrictedto certain stratigraphic levels, so that astratigraphical organisation will reflect theirdifferent morphologies <strong>and</strong> likely producers, ormay occur at many horizons (some taxa occurthroughout the Phanerozoic).• Geographical – Organisation according tolocation will show similarities <strong>and</strong> groupingsover large areas, may help to correlate stratigraphichorizons <strong>and</strong> will support ichnofaunal studies.• Ethological – Organisation follows a functionalinterpretation of ichnotaxa. Similar modes ofproduction are grouped together, such as feeding<strong>trace</strong>s, dwelling <strong>trace</strong>s, locomotion <strong>trace</strong>s, etc.Problems may arise when interpretations ofbehaviour change, which will probably give acurator problems of reorganisation.• Combination – An assembly of these can also beused, though this will inevitably increase the-256-

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