amount of space needed to accommodate thecollection if it is not to be cluttered, cramped orconfusing. Thus, the following organisation couldbe followed throughout the collection, each drawerbearing a label showing:1. Geographical location - e.g., Isle of Wight,British Isles, North America.2. Stratigraphical level - how old is it?3. Ichnotaxon - its classification within the system.4. Alphabetical - arranged within theichnotaxonomic organisation.Such a scheme of organisation is used by the NationaalNatuurhistorisch Museum, Leiden (NNHM), aprimary stratigraphic organisation being secondarilygrouped into geographic areas, e.g., Devonian –Germany, Devonian – France; Upper Cretaceous –The Netherl<strong>and</strong>s, Upper Cretaceous – Germany.Ichno<strong>fossils</strong> should not be classified according totheir perceived producing organism. Any givenichnofossil morphology may potentially have beenproduced by more than one biological species, whichmay belong to different phyla (Pickerill 1994). Onlyin cases where the producing organism is preservedin intimate juxtaposition with the <strong>trace</strong> can such anidentity be established with confidence.Whichever style the institution or curator chooses tofollow is not so important as long as specimens can belocated. There may also be a necessity to divide thecollection into large <strong>and</strong> small components, wherebylarge slabs requiring shelf space have to be separatedoff from small ‘h<strong>and</strong> specimens’ which can be storedin drawers. If this has to be done, the st<strong>and</strong>ard layoutshould be followed as far as possible.Two national collectionsThe evolution of a collection of items in a <strong>museum</strong>can tell a lot about the development of the <strong>museum</strong> asan institution of scientific excellence. Some of theolder <strong>museum</strong>s throughout the world may have startedlife as simple displays of curios without any particularregard to order. As scientific experience grew, bettermethods of display developed. Storage methods soonfollowed suit, for the collections began to grow asinterest was fired amongst the population, <strong>and</strong> noteverything could be put on show. Some <strong>museum</strong>swere set up around a core collection, such as that ofHans Sloane for the British Museum, whose naturalhistory collection was the core which eventually ledto the founding of the daughter institution, the BritishMuseum (Natural History), as the Natural HistoryMuseum (BMNH) in London was known untilrecently. The BMNH is now completely independentof the parent <strong>museum</strong>.The Natural History Museum, LondonThe <strong>trace</strong> fossil collection in the Department ofPalaeontology at the BMNH is modest in size <strong>and</strong>represents collections made from the nineteenthcentury onwards. In the past it contained pretty muchanything that was not readily identifiable, withartifacts (including lumps of concrete with theimpression of the sacking that had once contained it),curiously shaped stones, wind-polished rocks fromdeserts <strong>and</strong> lumps of rock with strange markings,including mineral dendrites resembling plants. Inamongst these were to be found true <strong>trace</strong> <strong>fossils</strong> - theremains of the lifestyles of organisms. It alsocontained, <strong>and</strong> indeed still does, the Problematica,that is, body <strong>fossils</strong> of unknown identity.Eventually, the whole mixed collection was sortedout <strong>and</strong>, with some exceptions, the non-<strong>trace</strong> fossilcomponent was removed from the collection <strong>and</strong>transferred to the correct locations elsewhere in the<strong>museum</strong>. The exceptions include plaster models ofBeringer’s iconoliths (Taylor 2004) <strong>and</strong> curiouslyshaped flints that have a fanciful resemblance tovarious animals (see Lewis 2000). Other nonbiologicalspecimens are also retained for the purposeof illustrating to interested parties the pitfalls for theunwary. One fine example of these, amongst many, isan object which resembles an eviscerated stomach<strong>and</strong> which is actually a ceramic bottle that collapsedduring firing.The remaining true <strong>trace</strong> <strong>fossils</strong> are now stored intheir own part of the storage system of the Departmentof Palaeontology. They occupy 125 drawers in three<strong>and</strong> a half cabinets, six shelves in two cabinets <strong>and</strong> afew slots in the roller storage set aside for large slabs(for a description of the storage units see Owen et al.1982). These do not include the <strong>trace</strong> <strong>fossils</strong> whoseproducers are well known <strong>and</strong> which are stored withthe relevant taxa in other parts of the collections,including the dinosaur tracks stored with fossilreptiles, most of the Gastrochaenolites specimensstored with the molluscs <strong>and</strong> Gnathichnus which arekept with the fossil echinoids.Material from the Ediacara Formation of Australia isalso kept together rather than being distributedalphabetically, with the rock specimens beingsupplemented by plaster casts of the originals held inAustralian <strong>museum</strong>s. These <strong>fossils</strong> also come into thecategory of ‘problematica’, a mixture of <strong>trace</strong> <strong>and</strong>body <strong>fossils</strong> preserved as natural moulds <strong>and</strong> casts.Even though some of these remains may eventually-257-
prove to be nothing more that sedimentary structures<strong>and</strong> not biologically produced entities, they shouldbe kept together for contextual <strong>and</strong> historical reasons.This may seem to be returning to the earlier days ofkeeping together everything that was not identifiable,but now there is the difference of experience <strong>and</strong> newknowledge rather than ignorance <strong>and</strong> not knowingwhat to do with them. Research into another part ofthe problematica collections, the so-called ‘Muschia’from the Devonian Gogo Formation of Australia,reveal that parts can be re-distributed to fossil fishes<strong>and</strong> crustaceans. The name ‘Muschia’, an unofficialone, indicates the general state of the preservation, ageneralised mush.The arrangement of the collection in a user-friendlystate was non-existent for many years, with everythingmixed in together. This made finding a specimenextremely difficult <strong>and</strong> time consuming, such that thewhole collection had to be trawled through in order tolocate something required. In the early 1990s a startwas made to re-organise the <strong>trace</strong> <strong>fossils</strong> collectionsinto something more user-friendly. Volunteers,including several from the Museum front-of-housestaff (as part of their training to see how the ScienceDepartments worked), carried out an initial sortthrough the material, firstly to re-box <strong>and</strong> re-labelspecimens, <strong>and</strong> then to sort roughly into similaritems. Later on, another volunteer, a non-member ofthe staff who had an interest in <strong>trace</strong> <strong>fossils</strong>, againsorted through the collections, identifying as best aspossible the ichnotaxa present. It was then very mucheasier to re-organise the collection, doing soalphabetically for the most part, but with certaindiscrete parts kept together, usually because theywere cited in various scientific papers.When the re-arrangement was complete, the drawerswere re-labelled <strong>and</strong> location indexes wereconstructed so that specimens could be extractedeasily <strong>and</strong> quickly. Furthermore, both the drawerlabels <strong>and</strong> index can be updated easily.The next phase of the operation will be to sort out <strong>and</strong>re-organise the material on the shelf units. This willbe rather more tricky as some of the specimens arelarge <strong>and</strong> heavy, so that placing them in their correctalphabetic location may not be possible withoutendangering those who may want to look at them <strong>and</strong>remove them from the collections. Heavy specimenson a high shelf can be a problem without specialisth<strong>and</strong>ling equipment.Currently, the <strong>trace</strong> fossil collections are used onlyoccasionally by external visitors (e.g., Donovan 2002)<strong>and</strong>, more often, by internal staff. With pressure onthe storage capacity of the Department ofPalaeontology there is a possibility that the wholecollection will be removed from the South Kensingtonsite to the outstation at W<strong>and</strong>sworth, where storage ismuch less cramped <strong>and</strong> environmental conditions arethe equal of the main <strong>museum</strong>. However, access isless immediate <strong>and</strong> transport to the site is moredifficult.Nationaal Natuurhistorisch Museum, LeidenThe <strong>trace</strong> fossil collection of the NNHM is small,reflecting a previous lack of interest in ichnology inThe Netherl<strong>and</strong>s <strong>and</strong> the <strong>museum</strong>. There are currentlyabout 60 drawers of ichno<strong>fossils</strong>, ranging throughoutthe Phanerozoic. The collection is richest in specimensfrom northwest Europe <strong>and</strong> Spain, particularly theDevonian <strong>and</strong> Mesozoic. Many specimens areawaiting identification or re-identification. Type <strong>and</strong>figured specimens are few. However, S.K.D. <strong>and</strong> coworkersare currently actively researching theichnology of the Upper Cretaceous of northern Europe(e.g., Donovan <strong>and</strong> Jagt 2005) <strong>and</strong> the Cenozoic ofthe Antilles (e.g., Pickerill et al. 2003), resulting in acurrent steady influx of new, correctly identifiedspecimens. Contributions concerning significantdonated material are being encouraged to the<strong>museum</strong>’s journal, Scripta Geologica (e.g., Blissett<strong>and</strong> Pickerill 2004).AcknowledgementsWe thank our referees, Drs J.W.M. Jagt(Natuurhistorisch Museum Maastricht) <strong>and</strong> J.T.Hannibal (Clevel<strong>and</strong> Museum of Natural History),for their constructive reviews. This is a contributionto S.K.D.’s NNHM project “Trace fossil studies”.ReferencesBLISSETT, D.J. <strong>and</strong> PICKERILL, R.K. 2004. Softsedimentichnotaxa from the Cenozoic WhiteLimestone Group, Jamaica, West Indies. ScriptaGeologica 127, 341-378.BRIGGS, D.E.G. <strong>and</strong> ROLFE, W.D.I. 1983. A giantarthropod trackway from the Lower Mississippian ofPennsylvania. Journal of Paleontology 57, 377-390.DONOVAN, S.K. 2002. A new ichnospecies ofGastrochaenolites Leymerie from the PleistocenePort Morant Formation of southeast Jamaica <strong>and</strong> thetaphonomy of calcareous linings in clavate borings.Ichnos 9, 61-66.DONOVAN, S.K. <strong>and</strong> JAGT, J.W.M. 2005. Anadditional record of Oichnus excavatus Donovan &Jagt from the Maastrichtian (Upper Cretaceous) ofsouthern Limburg, The Netherl<strong>and</strong>s. ScriptaGeologica 129, 147-150.-258-
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