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December 1, 2012 EvolDir 43<br />

cesses that determine the assembly of wood decomposer<br />

communities in freshwater-estuarine river systems<br />

in Coiba Island National Park on the Pacific<br />

coast of Panama. This project will provide unique,<br />

cross-disciplinary training in tropical ecology, metagenomics,<br />

bioinformatics, microbiology, and fungal taxonomy.<br />

Students will also have the opportunity to apply<br />

to a recently-funded NSF IGERT (graduate training<br />

grant: http://sib.illinois.edu/igert/index.php), which<br />

is focused on integrating genomic data with organismal<br />

biology. Interested applicants should contact<br />

one of the PIs: Astrid Ferrer (aferrer@life.illinois.edu),<br />

Jim Dalling (dallingj@life.illinois.edu), or Katy Heath<br />

(kheath@life.illinois.edu) for more information.<br />

kheath@life.illinois.edu<br />

UJyvaskyla Finland SocialEvolution<br />

PhD opportunity in Social Evolution Theory at the<br />

University of Jyväskylä, Finland<br />

Eusocial organisms such as bees and ants pose a challenge<br />

for evolutionary theory because they exhibit altruistic<br />

traits that contradict the Darwinian principle<br />

of individual fitness maximization. Although a large<br />

body of theory exists on the evolution of altruistic<br />

traits, few studies have explicitly modelled the transition<br />

from a solitary lifestyle to eusociality, including<br />

feedbacks between gene frequencies, colony dynamics<br />

and population ecology that are a necessary part<br />

of this process. We aim to fill this gap by building<br />

models based on recent theoretical advances. Applicants<br />

should have a degree in a relevant area (e.g. behavioural<br />

ecology, evolutionary biology, economics with<br />

focus on game theory, physics, mathematics, computer<br />

science), with some mathematical and computing skills<br />

and a keen interest in evolutionary biology. The project<br />

will be supervised by Lutz Fromhage at the University<br />

of Jyväskylä, in collaboration with Hanna Kokko at<br />

Australian National University, Canberra. Funding is<br />

provided by the Department of Biological and Environmental<br />

Science to fill positions in two out of four<br />

competing projects, of which the above project is one.<br />

The department seeks to recruit new students into its<br />

PhD programme, starting 1 January 2013. The goal<br />

of the department is that the student completes the<br />

PhD degree in four years. Initially, the successful candidates<br />

will receive a one year employment contract,<br />

which can be extended up to four years after positive<br />

progress evaluation. The application deadline is<br />

on 23 November 2012, 4:15 pm. Applications should<br />

be submitted exclusively through www.jyu.fi/science/en/applicationformphdstudents,<br />

and should include a<br />

CV, and a research and study plan. Before submitting<br />

the application, the applicant is advised to contact Dr.<br />

Lutz Fromhage: lutz.fromhage@jyu.fi<br />

Dr. Lutz Fromhage University of Jyväskylä PO Box<br />

35, 40014 Finland tel: +358 404834256<br />

Lutz Fromhage <br />

UKonstanz<br />

ParallelEvolutionCichlids<br />

The Meyer lab at the University of Konstanz, Germany,<br />

member of the International Max Planck Research<br />

School (IMPRS) for Organismal Biology, is offering<br />

two fully-funded three-year PhD positions<br />

Ecology and genomics of parallel speciation of cichlid<br />

fishes in Nicaraguan crater lakes<br />

Project abstract The crater lakes in Nicaragua each<br />

contain small adaptive radiations of endemic cichlid<br />

fish. The species in these lakes often look astonishingly<br />

similar (Elmer and Meyer 2011). These cases of<br />

parallel evolution are the focus of research in the Meyer<br />

lab and we are studying this phenomenon at different<br />

levels of biological organization. We are looking for<br />

two PhD students to join our team who are interested<br />

in this from a (1) genetic (e.g., QTL-analyses) and genomic<br />

(e.g., comparative transcriptomics) perspective,<br />

and (2) an ecological/experimental viewpoint. The ecological<br />

genomics work aims to discover the genetic basis<br />

of (parallel) adaptations and next the functional<br />

characterization of candidate genes. This approach involves<br />

nextgeneration DNA sequencing technologies (Illumina<br />

and Roche FLX platforms) in the Genomics<br />

Center Konstanz and evo-devo work using transgenic<br />

model fish (e.g., zebrafish and medaka). We also plan<br />

to conduct mark-recapture studies and other ecological<br />

work in crater lakes in Nicaragua and are looking for<br />

a Spanish-speaking student who would work for longer<br />

periods of time in Nicaragua.<br />

The successful applicants will participate in the International<br />

Max Planck Research School (IMPRS) for Organismal<br />

Biology, the PhD program of the Max Planck<br />

Institute for Ornithology in Seewiesen and Radolfzell<br />

and the University of Konstanz. All IMPRS students<br />

are supported by stipends or contracts. The pro-

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