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Common Trees of Puerto Rico and the Virgin Islands

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Within <strong>the</strong> area described are four distinct forest<br />

regions or ecological provinces, each giving rise<br />

to a distinct type <strong>of</strong> forest. These re~ions or provinces,<br />

designated as moist coast, moist limestone,<br />

dry coast, <strong>and</strong> dry limestone, are shown in figure 2.<br />

?he more common or characteristic species <strong>of</strong><br />

<strong>the</strong> moist coastal forest included <strong>the</strong> following:<br />

Acrocomia media Psidium guajasa<br />

Nect<strong>and</strong>ra coriacea Nanilkara btdentata<br />

Hernundia sonora Siderox lon<br />

Hymnuea courbaril /oe t idstimum<br />

Andira inemia C%t harexylum<br />

Pterocarpw <strong>of</strong>ficinal& fmtico&um<br />

Zanthoxylum<br />

Petitia domingensis<br />

martinicense<br />

Tabebuia heterophylla<br />

Calophyllum<br />

Genipa americam<br />

brasilieme<br />

Guettarda scabra<br />

illammea americana R<strong>and</strong>ia acuteata<br />

The moist limestone forest was similar to that<br />

along <strong>the</strong> coast <strong>and</strong> had many <strong>of</strong> <strong>the</strong> same species.<br />

The chief differences appear to be due to <strong>the</strong> drier<br />

soils on <strong>the</strong> well-drained limestone hills <strong>and</strong> <strong>the</strong><br />

ater humidity in <strong>the</strong> protected areas between<br />

T'<br />

t e hills, especially in <strong>the</strong> sout.hern part <strong>of</strong> this<br />

area which is close to <strong>the</strong> central mountains. .The<br />

trm species <strong>of</strong> <strong>the</strong> moist limestone forest include :<br />

Aiphanes<br />

aeanthophylla<br />

Baussia attenuata<br />

Coccoloba diversifolia<br />

Coccoloba pubescens<br />

Licaria salicifolia<br />

Zant hox y Zum<br />

nuzrtinicense<br />

Bursera simarmba<br />

Cedrela oderata<br />

Hyeronima cluswides<br />

Sapium lauroceras.tu<br />

Thouinia striata<br />

Montesuma<br />

speciosissima<br />

Ochroma pyramidale<br />

Clu8ia. rosea<br />

Bucida bucem<br />

Tetrw gia ekagnaides<br />

I<br />

Dipho is salicifolia<br />

Siderox lon<br />

oeti&si-m<br />

Guettarda scabra<br />

Terebraria resinosa<br />

R<strong>and</strong>ia aculeata<br />

On <strong>the</strong> sou<strong>the</strong>rn, dry side <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>the</strong><br />

more adverse moisture conditions excluded many<br />

<strong>of</strong> <strong>the</strong> tree species common on <strong>the</strong> north side. In<br />

<strong>the</strong>ir places grew a few o<strong>the</strong>r species especially<br />

adapted to such conditions. The tree species <strong>of</strong><br />

<strong>the</strong> dry coastal forest include :<br />

Coccoloba venosa Pot gala cowellii<br />

Cap aris cynopha2b- Cei fa pent<strong>and</strong>m<br />

p I ora<br />

Guazuma dmif oZia<br />

Stahlia monospemza CanelZu wintemna<br />

Lonchocarpus dontin- Bucida buceras<br />

Prn?tSh<br />

Rauvolfia nitida<br />

Pictetm aculeata Cordia nitida<br />

Erythroxyhn areola- Citharez ylum<br />

turn<br />

fruticosum<br />

Guaiocum <strong>of</strong>ficim.de<br />

In <strong>the</strong> limestone region <strong>of</strong> <strong>the</strong> south coast, as on<br />

<strong>the</strong> north coast, excessive soil drainage accentuates<br />

<strong>the</strong> dryness <strong>of</strong> <strong>the</strong> environment to a point that some<br />

species <strong>of</strong> trees cannot subsist. O<strong>the</strong>rs which are<br />

more hardy replace <strong>the</strong>se. The trees <strong>of</strong> <strong>the</strong> dry<br />

limestone forest include :<br />

Pbonia dbida<br />

Guaiacum, <strong>of</strong>ficinale<br />

Guaiacum sanctum<br />

A?nyris elemif era<br />

Bursera &mamba<br />

G ymnnn<strong>the</strong>s lucida<br />

Thouinia portorlcmia<br />

Colubrina n~*borescens<br />

Su)~cmphaZua reticulalus<br />

I~cphnlocereus royed<br />

0 y u~ttia rubescens<br />

Btrcidcl bucerua<br />

Dipholb sdicifolia<br />

Plumeria dba<br />

The constal forests <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong><br />

Isl<strong>and</strong>s, unlike those <strong>of</strong> <strong>the</strong> o<strong>the</strong>r Greater<br />

Antilles, con tnined no mahogany (Swietenia mahngoni).<br />

Widespread use <strong>of</strong> <strong>the</strong> wood <strong>and</strong> early<br />

introduction <strong>of</strong> <strong>the</strong> tree to <strong>the</strong>se isl<strong>and</strong>s has led to<br />

n generi11 impression that this species is native.<br />

However, whereas <strong>the</strong> young trees develop abundantly<br />

in <strong>Puerto</strong> <strong>Rico</strong> beneath or near planted<br />

trees <strong>of</strong> this species, <strong>the</strong>y are never encountered in<br />

native forest at any distance from such trees. Had<br />

<strong>the</strong> species been native, <strong>the</strong>re would still be young<br />

trees throughout <strong>the</strong> coastal forests <strong>of</strong> <strong>the</strong> isl<strong>and</strong><br />

without relation to <strong>the</strong> location <strong>of</strong> planted trees.<br />

It is extremely unlikely that it could ever have<br />

been exterminated, since aceitillo (Zanthoaylum,<br />

flavton), a tree in greater dem<strong>and</strong> <strong>and</strong> with much<br />

wealter reproductive capacity than mahogany, is<br />

still to be found in remote forests.<br />

Typical mountain forests are confined to <strong>Puerto</strong><br />

<strong>Rico</strong>, although :L small patch <strong>of</strong> similar but unique<br />

forest is found on <strong>the</strong> top <strong>of</strong> Sage Mountain, Tortola.<br />

Between about 500 to 2,000 feet elevation in<br />

<strong>the</strong> enstern mountains, known as Luquillo Mountains,<br />

<strong>and</strong> to 3,000 feet in <strong>the</strong> central mountains or<br />

Central Cordillera, slightly higher on <strong>the</strong> south<br />

slope than on <strong>the</strong> north, was probably <strong>the</strong> most<br />

magnificent forest <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong>. bluch <strong>of</strong> <strong>the</strong><br />

original vegetation <strong>of</strong> this area is described as<br />

tropical moist forest; in <strong>the</strong> wettest areas it is tropical<br />

rain forest. At its maximum development this<br />

forest reached 110 feet in height, with trees to 8<br />

feet. in diameter. Three forests <strong>of</strong> distinct size <strong>and</strong><br />

composition grew toge<strong>the</strong>r here, each forming a<br />

separate story <strong>of</strong> vegetation. Throughout <strong>the</strong><br />

range <strong>of</strong> this type <strong>of</strong> forest <strong>the</strong>re mere probably<br />

about 170 tree species.<br />

Within <strong>the</strong> mountain area are two forest regions<br />

or ecological provinces <strong>and</strong> corresponding distinct<br />

forest types. These provinces are desi ated as<br />

lower Cordillera <strong>and</strong> lower Luquillo. f" t is seen<br />

in figure 2 that <strong>the</strong> lower Cordillera province includes<br />

both <strong>the</strong> north <strong>and</strong> south lower slopes <strong>of</strong><br />

<strong>the</strong> central mountains <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong><br />

Sierra de Cnyey <strong>and</strong> also <strong>the</strong> upper slopes <strong>of</strong> <strong>the</strong><br />

disconnected Sierra de Atalaya in <strong>the</strong> northwest.<br />

The Lu uillo Mountains are separate both geographica<br />

'I ly <strong>and</strong> ecologically from <strong>the</strong> Central<br />

Cordillera.<br />

The trees <strong>of</strong> <strong>the</strong> lower Cordillera forest include<br />

<strong>the</strong> following :

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