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Common Trees of Puerto Rico and the Virgin Islands

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SOAPBERRY FAMILY (SAPINDACEAE)<br />

Key to <strong>the</strong> 6 species illustrated (Nos. 138-143)<br />

A. Leaves simple, elliptic, <strong>the</strong> edges flnely too<strong>the</strong>d-142. I?loui?tfa pmtwtcaeie.<br />

AA. Leaves compound.<br />

B. Leaflets 3, elliptic or obovate, broadest beyond middle, shorbpointed at both ends, too<strong>the</strong>d on edges-143.<br />

Thouinia striata.<br />

BB. Leaflets pinnate, 4 or more (sometimes only 2).<br />

C. Leaflets 4-8, not paired, with wavy-too<strong>the</strong>d edges, elliptic, those toward apex largest, rounded or notched<br />

at apex-138. Cupanfa americana.<br />

CC. Leaflets mostly mired, not too<strong>the</strong>d.<br />

D. Leaflets 2-8, rounded or blunt-pointed at apex, on slender reddish or dark brown axis; fruit a<br />

flattened seed capsul+l3& dlatal/ba domingensls.<br />

DD. Leaflets long- or short-pointed at both ends, on axis <strong>of</strong>ten winged; fruit round, fleshy.<br />

E. Lenflets 4, <strong>the</strong> pair at apex larger, fruit Y'-l% inches in diameter, edible-140. llielicoccue<br />

bijftgatu8.*<br />

BE. Leaflet8 usually 8-12, sometimes only 1 at end; fruit %-I inch in diameter, yellow, inediblc+141.<br />

f3apindus saponaria.<br />

138. Guara Cupania americana L.<br />

This tree is characterized by : (1) twi4s <strong>and</strong> leaf<br />

axes brown hairy ; (2) pinnate leaves with 4-8 alternate<br />

elliptic or obovate leaflets, those toward<br />

apex largest, rounded or notched at apex, shortpointed<br />

at base, with aavy too<strong>the</strong>d ed es, <strong>the</strong> up-<br />

per surface shiny grem <strong>and</strong> hairy on f<br />

y on veins,<br />

<strong>and</strong> <strong>the</strong> lower surface aler <strong>and</strong> densely s<strong>of</strong>t<br />

hairy; (3) numerous sma P 1 whitish 5-parted flowers<br />

y8 inch across, in most1 terminal branched<br />

clusters; <strong>and</strong> (4) <strong>the</strong> rounde I seed capsules 1/2-S/4<br />

inch long, bluntly 3-lobed, velvety-brown or rustybrown<br />

hairy, splitting into 3 parts <strong>and</strong> exposing 3<br />

rounded shiny blackish seeds 5/ls inch long, each<br />

in an omnge cup.<br />

A small to medium-sized evergreen tree attaining<br />

20-50 feet in height <strong>and</strong> 10 inches in trunk<br />

diameter with a broadly spreading rounded crown.<br />

The gray bark is smoothish or becoming rough <strong>and</strong><br />

fissured into plates. Inner bark is light brown <strong>and</strong><br />

sli htly bitter. The twigs are stout.<br />

h e leaves are alternate, 5-10 inches long, with<br />

stout axes. Leaflets have short hairy stalks l,--1/4<br />

inch long <strong>and</strong> blades 1y2-6 inches long <strong>and</strong> 3/4-2%<br />

inches broad <strong>and</strong> slightly thickened.<br />

The flower clusters (panicles) 4-8 inches long<br />

have brown hairy branches. Flowers are male,<br />

female, <strong>and</strong> bisexual (pol gamous). There are 5<br />

hairy sepals nearly l/s inc i: long; 5 hairy, narroxv<br />

stalked petals about as long as sepals, each with 2<br />

scales on <strong>the</strong> outer edges; 8 stamens on a disk; <strong>and</strong><br />

pistil composed <strong>of</strong> hairy 3-celled ovary with short<br />

style <strong>and</strong> 3 stigmas.<br />

Seed capsules common1 are many <strong>and</strong> crowded<br />

in terminal branched c ?' usters, short-stalked at<br />

base, opening widely into 3 parts, retaining <strong>the</strong> 3<br />

seeds attached for some time. Flowering in winter<br />

<strong>and</strong> early spring (December to March) <strong>and</strong> maturing<br />

fruits in spring <strong>and</strong> summer.<br />

The mood is Iight brown <strong>and</strong> hard, <strong>of</strong> medium<br />

weight (specific gravity 0.4). Very susceptible to<br />

attack b dry-mood termites. Used in <strong>Puerto</strong> <strong>Rico</strong><br />

P<br />

chiefly or posts ,<strong>and</strong> oles <strong>and</strong> elsewhere for con-<br />

struction <strong>and</strong> shipbui f' ding.<br />

This s ecies has been su gested as an ornamental<br />

<strong>and</strong> sha i e tree. The see 2 s <strong>and</strong> leaves sometimes<br />

serve for medicinal Also a honey plant.<br />

Forests, especially<br />

coast, moist limestone,<br />

<strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong>.<br />

PUBLIC FORESTS..-Cambalache, Carite, Guajataca,<br />

Luquillo, Maricao, Rio Abajo, Susfia, Vega.<br />

RANGE.-Greater Antilles <strong>and</strong> in Lesser Antilles<br />

on1 in Dominica, Martinique, <strong>and</strong> Barbados, <strong>and</strong><br />

in grinidad <strong>and</strong> Tobago. Also in Colombia <strong>and</strong><br />

Venezuela.<br />

OTHER COJ~MON ~~~rn.--guara blanca (<strong>Puerto</strong><br />

<strong>Rico</strong>) ; guiirana, guara (Dominican Republic) ;<br />

guara, guara comGn, phrano, gudrana macho,<br />

gukrana hembra (Cuba) ; guacharaco (Colombia)<br />

; par&, patillo, zapatero, cabimo, guamo<br />

guarB, guamo matias (Venezuela) ; c<strong>and</strong>lewoodtree<br />

(Barbados) ; maraquil (Trinidad) ; bois de<br />

satanier (Haiti).<br />

A closely related s ecies <strong>of</strong> para (Cupania<br />

tvipuetm A. Rich.) in f?uerto <strong>Rico</strong> differs in hm ing twi~ <strong>and</strong> seed capsules with shorter yellowbrown<br />

hairs <strong>and</strong> <strong>the</strong> seed capsules sharply 3-angled<br />

<strong>and</strong> on longer stalks.

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