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Common Trees of Puerto Rico and the Virgin Islands

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DOGBANE FAMILY (APOCYNACEAE)<br />

219. Palo amargo, bitter-ash Rauvoliia nitida Jacq.<br />

This small tree or shrub with white latex is distinguished<br />

by its shiny yellow-pn lance-shaped<br />

or narrowly elliptic leaves 2%-6 inches Ion <strong>and</strong><br />

a/4-ls/4 inches wide, long-pointed at apex an I f base,<br />

slightly turned under at edges, attached to <strong>the</strong><br />

oups <strong>of</strong> 4 <strong>of</strong> unequal size (whorled).<br />

%fs,"afwhitish flowers are about lh inch across<br />

<strong>and</strong> <strong>the</strong> reddish to blackish rounded fruits about<br />

% inch long <strong>and</strong> inch brond.<br />

<strong>Common</strong>ly a shrub or small tree to 20-40 feet<br />

high <strong>and</strong> to 1% feet in trunk diameter, evergreen<br />

with open rounded crown. The bark is smoothish,<br />

slightly fissured <strong>and</strong> warty, very light brown, <strong>and</strong><br />

thin. Inner bark is light green <strong>and</strong> bitter. The<br />

twigs are bright green with rnised brown dots<br />

(lenticels), enlarged at <strong>the</strong> nodes, <strong>and</strong> becoming<br />

brownish.<br />

The slightly thickened lea<strong>the</strong>ry leaves have<br />

petioles I/a-s/4 inch long <strong>and</strong> blades green or yel-<br />

low green on upper surface <strong>and</strong> lighter yellow<br />

green beneath.<br />

The small flower clusters (cymes) are terminal<br />

or lateral, branched, 1-11h inches long <strong>and</strong> brond,<br />

shorter than <strong>the</strong> leaves, with several to many flow-<br />

ers. The white <strong>and</strong> greenish flowers are tubular,<br />

about % inch long, with 5 spreading lobes 4/4 inch<br />

across; <strong>the</strong> green 5-lobed calyx is less than 4/8 inch<br />

long; corolla has a narrow greenish tube j/4 inch<br />

long <strong>and</strong> 5 rounded white lobes; 5 minute stamens<br />

are inserted near mouth <strong>of</strong> corolla tube; <strong>and</strong> <strong>the</strong><br />

pistil is composed <strong>of</strong> n 2-lobed ovary with slender<br />

style <strong>and</strong> enlarged stigma.<br />

The fleshy fruits, with milky juice, turn from<br />

green to reddish <strong>and</strong> blackish <strong>and</strong> contain 1 or 2<br />

brownish seeds s/16 inch long. Flowering <strong>and</strong><br />

fruiting nearly throughout tho year.<br />

The sapwood is light brown, <strong>and</strong> <strong>the</strong> heartwood<br />

clenr yellow. The hard, lightweight wood is emoyed<br />

for posts in <strong>Puerto</strong> <strong>Rico</strong>.<br />

PI or furniture <strong>and</strong> musical instruments E1sewhe%ase has een<br />

suggested.<br />

In <strong>the</strong> coastal <strong>and</strong> limestone forests <strong>and</strong> thickets<br />

<strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong>, growing in <strong>the</strong> o en <strong>and</strong> robably<br />

requiring light. Also in Mona, 6eques, &. Croir,<br />

St. Thomas, St. John, <strong>and</strong> Tortola<br />

P~LIC FORE~TS.-Cnmbalache, au6nica.<br />

RANGE.-Bahamas, Cuba, Jamaica, Hispaniola,<br />

<strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>Virgin</strong> Isl<strong>and</strong>s, St. Kitts, Guadeloupe,<br />

<strong>and</strong> Martinique. Also recorded long ago<br />

from St. Barth6lemy.<br />

OTHER CONMON NAMES.- a10 de muiieco, ca-<br />

i<br />

chimbo (<strong>Puerto</strong> <strong>Rico</strong>) ; bitter ush, milkbush (Virfin<br />

,Isl<strong>and</strong>s) ; palo de leche (Dominican Repubic)<br />

, huevo de gallo (Cuba) ; smooth rauvolfia<br />

(Bahamas) ; bois lnit femelle (Haiti).<br />

The name <strong>of</strong> a low shrub (Ra.uvuljia tetraphylla<br />

L. has been misapplied to this s ecies.<br />

?l!he genus Rauvol* (also spe f' led Rawwol*)<br />

has attracted much attention in recent years becnuse<br />

<strong>the</strong> root <strong>of</strong> a shrubby species in India has<br />

yielded a drug for <strong>the</strong> treatment <strong>of</strong> high blood<br />

pressure <strong>and</strong> certain mental illnesses. Related species<br />

are also under investigation.<br />

Ano<strong>the</strong>r native species is a shrub 3-10 feet high<br />

known as bitterbush (Rauvol$a viridis Roem. &<br />

Schult. ; synonym R. Zumarclcii A. DC.). It has<br />

slightly unequal leaves in groups <strong>of</strong> 4 (whorled),<br />

ovate to elliptic, most1 11h-4 inches long, thin,<br />

dull green instead <strong>of</strong> diny, <strong>and</strong> smaller blackish<br />

fruits about 4/4 inch broad. It grows in coastal<br />

thickets <strong>of</strong> eastern <strong>Puerto</strong> <strong>Rico</strong> Mona, Icacos,<br />

Culebrn, Vieques, St. Croix, St. ~Aomas, St. John,<br />

nnd <strong>Virgin</strong> Gorda <strong>and</strong> o<strong>the</strong>r isl<strong>and</strong>s to Hispaniola<br />

<strong>and</strong> nor<strong>the</strong>rn South America.

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