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SPIROGYRA 217<br />

252. Spirogyra inconstans Collins 1912. Tufts College Studies.<br />

3, p. 73; Phycoth. Bor.-Amer., No. 1768. Wittrock and<br />

Nordstedt Algae Exsiccatae, No. 958. 1889.<br />

Vegetative cells 28-40 /a x /-i, 70-350 with replicate end walls; (2-)<br />

to 4 chromatophores, nearly straight, or making .5 to 1.5 turns in the<br />

cell; conjugation scalar iform or rarely lateral; tubes formed by both<br />

gametangia; at the time of tube formation both gametangia become<br />

reflexed and usually the conjugation between the same pair of filaments<br />

is limited to single, or to 2 adjoining pairs of cells; cross conjugation<br />

is not infrequent; receptive gametangia shortened and inflated up<br />

to 50-75 m; zygospores ellipsoid to broadly ellipsoid, 45-70 jU, x 75-140 /x;<br />

median wall reticulate-aculeate, brown. (PI. XXXVIII, Figs. 18-20.)<br />

United States: Illinois; Michigan; Indiana; Ohio; Massachusetts; Wash-<br />

ington (Bodenberg Coll.).<br />

Europe, from Sweden to Rumania.<br />

At the time of tube formation an outer pectic ring is formed at the<br />

junction, and at maturity there is usually an outer and an inner tube wall.<br />

The history of the specific name of this species is a striking example of<br />

a scientific "Comedy of Errors." In 1889 Nordstedt described and distrib-<br />

uted specimens of this alga, as S. insignis (Hassall) Kiitzing forma. In 1899<br />

Wille attached the name S. jallax to it, based on a variety described by<br />

Hansgirg as S. insignis var. jallax which has smooth-walled spores, alleging<br />

that Hansgirg had overlooked the ornamented median spore wall. In 1907<br />

Teodoresco found an alga similar to Nordstedt's forma, and named it<br />

S. insignis var. Nordstedtii. In 19 12 Collins described the same alga as<br />

S. inconstans. In 1913 Borge perpetuated Wille's error. In 1915 Transeau<br />

called attention to the discrepancy between Hansgirg's figure and descrip-<br />

tion and those of Wille, but, following Borge's key, decided in favor of<br />

S. jallax. In 191 8 Collins changed the name in his key to S. jallax. In 1928<br />

Skuja renamed the species S. willei. Czurda in 1932 relegated Collins'<br />

S. inconstans to the discard, and used the name S. willei. Collins, however,<br />

first described the alga as a species and his specific name is therefore valid.<br />

253. Spirogyra borysthenica Kasanofsky & Smirnoff 1913.<br />

Oesterr. Bot. Zeitschr. 63, p. 137, PL 3.<br />

Vegetative cells 30-40 /x x 1 80-450 /x, with replicate end walls; 2 to 4<br />

chromatophores, straight, or making up to 2.5 turns in the cell; con-<br />

jugation scalariform; tubes formed by both gametangia; fertile cells<br />

inflated up to 70 /a; zygospores ellipsoid, 52-62^1 x 110-160/J.; median<br />

spore wall with spinelike or mammaeform papillae, yellow-brown.<br />

(PI. XXXIX, Figs. 1-3.)<br />

United States: Michigan (Ackley Coll.); Iowa (Tiffany Coll.).<br />

U.S.S.R., Bukovina.

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