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phylogenetic relationships and classification of didelphid marsupials ...

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2009 VOSS AND JANSA: DIDELPHID MARSUPIALS 171<br />

00111 00101 00010 01100 00000 00011 00000 00000<br />

11000 10021 00000 00000 00000 01001 0000<br />

Marmosops incanus: 00000 10010 00100 00000 02010<br />

0-110 02020 1010? 00100 01010 00-00 00(01)11<br />

00101 00010 01100 00000 00011 00000 00000 11000<br />

10021 00000 0001? 00000 01001 0000<br />

Marmosops noctivagus: 00000 10010 00100 00000<br />

02010 0-010 02020 10101 00000 01010 00-00<br />

00(01)11 00101 00010 01100 00000 00011 00000<br />

00000 11000 10021 00000 00000 00000 01001 0000<br />

Marmosops parvidens: 00000 10000 00100 00000<br />

02010 0-010 02020 10101 00000 01010 00-00<br />

00111 00100 00010 01100 00000 00011 00000<br />

00020 11000 10020 00000 00000 00000 01001 ????<br />

Marmosops pinheiroi: 00000 10000 00100 00000 02010<br />

0-010 02020 10101 00000 01010 00-00 00111 00100<br />

00010 01100 00000 00011 00000 00020 11000 10021<br />

00000 00000 00000 01001 ????<br />

Metachirus nudicaudatus: 01000 11110 00000 00000<br />

02010 0-010 02010 00000 00000 01010 00-00 10011<br />

00100 00010 01100 00000 00121 00000 00000 11000<br />

10021 00100 00000 00000 01001 0000<br />

Monodelphis brevicaudata: 01000 0-014 00000 00000<br />

00010 0-010 00000 00000 00001 01010 10-00 01011<br />

(01)0100 0(01)010 01000 00000 00011 00000 00000<br />

21000 10021 00100 00010 00001 11001 0101<br />

Monodelphis emiliae: 01000 0-015 00000 00000 00010<br />

0-0?0 00000 00000 00001 01010 10-00 01011<br />

(01)0100 00010 01000 10000 (01)0011 00000<br />

00000 21000 10021 00?00 00020 00001 11001 0101<br />

Monodelphis peruviana: 01000 0-010 00000 00000<br />

00010 0-0?0 02000 00000 00001 01010 10-00<br />

00011 10100 00010 01000 00000 10011 00000<br />

00000 21?00 10021 00000 0001? 00001 11001 ????<br />

Monodelphis theresa: 01000 0-003 00000 00000 00010<br />

????0 02000 00000 00001 01010 10-00 00011 00100<br />

00010 01200 00000 10011 00000 00000 21?00 10021<br />

00?00 0001? 00001 11001 ????<br />

Phil<strong>and</strong>er frenatus: 01000 1(01)100 00000 00000 02010<br />

11010 01120 11000 10000 010(01)0 01010 02011<br />

10101 00010 01200 00100 00121 00001 00000 21?00<br />

10011 10?10 0000? 00000 01001 1111<br />

Phil<strong>and</strong>er mcilhennyi: 01000 11100 00000 00000 02010<br />

11010 01120 11000 10000 01010 01010 02011 10101<br />

00010 01200 00100 00121 00001 00000 21000 10011<br />

10210 00000 00000 01001 1111<br />

Phil<strong>and</strong>er opossum: 01000 11100 00000 00000 02010<br />

110?0 01120 11000 10000 01010 01010 02011 10101<br />

00010 01200 00100 00121 00001 00000 21000 10011<br />

10210 00000 00000 01001 1111<br />

Thylamys macrurus: 00000 10016 00000 00001 02011<br />

0-1?0 02000 10010 00100 01010 00-00 00111 00101<br />

10110 01100 10000 00011 00000 00000 21?00 10021<br />

00000 0001? 00000 01001 0000<br />

Thylamys pallidior: 01000 10016 00000 00001 02011<br />

????0 02000 10010 00100 01010 00-00 00111 00101<br />

00110 01100 10000 00011 00000 00000 21000 10021<br />

00000 00011 00000 01001 0000<br />

Thylamys pusillus: 00000 10016 00000 00001 02011 0-<br />

110 02000 10010 00100 01010 00-00 00111 00101<br />

10110 01100 10000 00011 00000 00000 21000 10021<br />

00000 0001(01) 00000 01001 0000<br />

Thylamys venustus: 00000 10016 00000 00001 02011<br />

????0 02000 10010 00100 01010 00-00 00111 00101<br />

10110 01100 10000 00011 00000 00000 21000 10021<br />

00000 00011 00000 01001 0000<br />

Tlacuatzin canescens: 00000 10000 00100 01000 02010<br />

0-010 02000 10000 00000 01010 01000 00011 00100<br />

10010 01000 00000 00011 00000 00000 11000 10020<br />

00000 00001 00000 01001 1111<br />

Caenolestes fuliginosus: 1-110 0-000 00001 10010<br />

11000 0-000 02000 00001 0001? 00000 00-11 000??<br />

?0100 00001 01000 00000 10011 01102 1-0(03)0<br />

21?11 -1--- 01211 ???22 01110 01101 0000<br />

Rhyncholestes raphanurus: 1-110 0-000 00001 10010<br />

11000 0-010 02000 0?010 0001? 00000 00-11 000??<br />

?0110 00001 01000 00000 10011 00(01)02 1-0(02)0<br />

21?11 -1--- 01211 ???2? 01110 01101 0000<br />

Murexia longicaudata: 01000 0-010 00000 00001<br />

11000 0-000 02000 0?000 20000 01000 00-0(01)<br />

01010 00100 01001 01010 21000 10011 00000 1-100<br />

21200 10020 00?00 01122 00001 01000 0000<br />

Sminthopsis crassicaudata: 01000 10010 00000 00021<br />

11001 13000 02000 0?010 20000 01000 00-00 1000-<br />

00101 01001 01010 21000 10011 00000 1-100 21100<br />

10021 00000 01112 00001 00-00 0000<br />

Dromiciops gliroides: 01000 10001 00000 01000 02010<br />

10001 00000 11011 00000 01011 00-00 10010 00100<br />

00011 11010 20011 00000 00000 00100 21000 10000<br />

00200 10020 00000 01001 0000<br />

Echymipera kalubu: 1-000 11000 02002 201-1 12220<br />

12000 02000 0?000 11101 10000 00-01 1000- 00100<br />

00001 01200 00000 11011 00013 1-130 21200 10130<br />

01010 01122 10000 01001 0000<br />

Perameles gunnii: 1-000 0-007 02002 201-1 12220<br />

12000 02000 0?001 01101 10100 00-01 1000- 01111<br />

00001 01211 01000 11012 00013 01100 21200 1113-<br />

01?10 01112 10000 00-01 0000<br />

APPENDIX 5<br />

Nonmolecular Apomorphy List<br />

Here we list all <strong>of</strong> the unambiguous<br />

changes (those identically located on the<br />

tree under both ACCTRAN <strong>and</strong> DEL-<br />

TRAN optimizations; Sw<strong>of</strong>ford <strong>and</strong> Maddison,<br />

1987) reconstructed by parsimony<br />

on our best-supported ingroup topology<br />

(without Chacodelphys, fig. 35). In effect,<br />

these morphological <strong>and</strong> karyotypic transformations<br />

summarize phenotypic character<br />

support for our <strong>classification</strong>, although<br />

it should be noted that we do not formally<br />

recognize all <strong>of</strong> the nodes in this tree as<br />

taxa.<br />

Internal Branches<br />

Caluromyinae + Glironia<br />

Character 32. Body pelage on tail: noneRlonger<br />

dorsally than ventrally.<br />

Character 35. Base <strong>of</strong> tail: not prehensileRprehensile.<br />

Character 52. Postorbital process 1: absentRpresent.<br />

Caluromyinae

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