A Checklist of the Beetles of Cuba with Data on Distributions and ...

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10 Arthropods <strong>of</strong> Florida and Neighboring Land Areas, Vol. 18 <strong>of</strong> a multiplicity <strong>of</strong> users; from agronomists, to ecologists, land-use managers, taxonomists, quarantine entomologists, ecologists and biogeographers. Users can insert <strong>the</strong>ir own new data as <strong>the</strong>y come upon it. Organization The core <strong>of</strong> <strong>the</strong> work is a list <strong>of</strong> valid species names, synonyms, and <strong>the</strong> more recent references about <strong>the</strong>se species in <strong>the</strong> scientific literature. The following explains <strong>the</strong> selection and organization <strong>of</strong> <strong>the</strong> included data. Higher classification. The checklist generally follows <strong>the</strong> sequence and phylogenetic arrangement <strong>of</strong> <strong>the</strong> world list <strong>of</strong> beetle families, subfamilies, and tribes <strong>of</strong> Lawrence and Newton (1995 and at http:www. coleopsoc.org/). Arnett and Thomas (2000), and Arnett et al. (2002). <strong>Data</strong> on distribution and habits <strong>of</strong> beetles outside <strong>of</strong> <strong>Cuba</strong> are generally from Lawrence (1982). The families are numbered so that finding <strong>the</strong>m in sequence is easier. Numbering from <strong>the</strong> world list is used because several families not now known from <strong>Cuba</strong> may be expected to be found in <strong>the</strong> future. Included species. The list contains all species <strong>of</strong> beetles which are known or reported to occur in <strong>Cuba</strong>. If <strong>the</strong> known distribtion <strong>of</strong> a species suggests that it very probably occurs in <strong>Cuba</strong>, but <strong>the</strong>re are seemingly no records for it, its name and references are enclosed in paren<strong>the</strong>ses ( ). Some lists or catalogues only vaguely give distributions as “West Indies” or “Antilles.” If <strong>the</strong> total distribution suggests that this includes <strong>Cuba</strong> I have conservatively included <strong>the</strong> species. The function <strong>of</strong> such additions is to alert future workers to species which are named but not yet explicitly recorded from <strong>Cuba</strong>. Such citations could undoubtedly be far more numerous than those I have included. If <strong>the</strong> species has been recorded in <strong>the</strong> past from <strong>Cuba</strong>, but <strong>the</strong> application <strong>of</strong> <strong>the</strong> name is now known to be an error, <strong>the</strong> name and references are enclosed in brackets [ ]. I ask that users <strong>of</strong> <strong>the</strong> catalogue bring errors <strong>of</strong> omission or inclusion to my attention. Citations. The author, year <strong>of</strong> publication, and page <strong>of</strong> publication are given (when known) for each valid species. These data will help to locate <strong>the</strong> original references. When a species was originally placed in a different genus than now, I give this original genus name when it is known to me. Synonomies. I list synonym names below <strong>the</strong> valid name <strong>of</strong> <strong>the</strong> species. I do not give complete lists <strong>of</strong> synonyms for <strong>the</strong> species in this checklist. My intention is to give synonomies only for names which have been applied to <strong>Cuba</strong>n records. Catalogs. I do not cite catalogs or lists <strong>of</strong> names if no new taxonomic, distributional, or bionomic information is given in <strong>the</strong>m. I do not give citations to <strong>the</strong> older primary literature which is included in Blackwelder (1944, 1957; reprinted in 1982). The literature cited in Blackwelder is from 1944 and earlier. To include all <strong>the</strong>se older references here would greatly increase <strong>the</strong> length <strong>of</strong> this listing. References. It is my intention to give references for “younger” literature only; i.e., that published since Blackwelder (1944, 1957) which has contributed data on <strong>Cuba</strong>n species. I have also tried to include references to recent faunal summaries or revisions which may not contain direct information on <strong>Cuba</strong>n species but which may contain keys, descriptions, or illustrations or o<strong>the</strong>r information on species in <strong>Cuba</strong> or which may be expected in <strong>Cuba</strong>. Primary older literature not cited in Blackwelder can be found in appropriate issues <strong>of</strong> “Zoological Record” or in <strong>the</strong> several catalogues cited. The relevant family chapters in Arnett and Thomas (2000) and Arnett et al. (2002) are valuable references for all beetle families in <strong>Cuba</strong>, and are usually not cited for each family. Types. Type status and depository are given (if known). Older species usually do not have designated types. Co-types or syntypes from which lectotypes may be designated may be in collections in Havana (<strong>the</strong> Gundlach collection), or in Paris (Chevrolat and Jaquelin du Val), or elsewhere (see table 3). Jacquelin du Val published new species names from specimens in both <strong>the</strong> Chevrolat and Guérin-Méneville collections, so those will probably have appropriate type material. Horn and Kahle (1935-1937) may be <strong>of</strong> help in locating historically important collections and specimens. Type localities. I list published type localities (t.l.) only for endemic species. In many older works, no type localities were specified. In <strong>the</strong>se cases “<strong>Cuba</strong>” can be seen to serve as a type area. If this is <strong>the</strong> case, I usually do not cite “<strong>Cuba</strong>.” Distributions. Distributions <strong>of</strong> <strong>the</strong> species are presented in two ways. (A). They are first summarized as <strong>Cuba</strong> and <strong>the</strong>n as a list <strong>of</strong> <strong>the</strong> known major islands or countries outside <strong>of</strong> <strong>Cuba</strong> from where <strong>the</strong> species is known. Distribution <strong>with</strong>in <strong>the</strong> USA is by abbreviations <strong>of</strong> states at <strong>the</strong> edges <strong>of</strong> <strong>the</strong> range in <strong>the</strong> USA. “Endemic” is used as a term to indicate that <strong>the</strong> species is naturally limited to <strong>Cuba</strong>. “Intro-
A <strong>Checklist</strong> <strong>of</strong> <strong>the</strong> <strong>Beetles</strong> <strong>of</strong> <strong>Cuba</strong> 11 Table 3. Locations, available codens, addresses, and annotations for major collections known to have identified <strong>Cuba</strong>n beetles. Codens for collections follow usage <strong>of</strong> Arnett et al. (1997). Amsterdam. ZMAN. Instituut voor Taxonomische Zoologie, Zoologisch Museum, Universiteit van Amsterdam, Plantage Middenlaan 64, 1018 DH, Amsterdam, The Ne<strong>the</strong>rlands. Aylmer. CMNC. Entomology Collections, Canadian Museum <strong>of</strong> Nature, Aylmer, Quebec, (P.O. Box 3443, Station D, Ottawa ON K1P 6P4), Canada. Basel. NHMB. Entomology Department, Naturhistorisches Museum, Augustinergasse 2, 4001, Basel, Switzerland. Especially containing <strong>the</strong> large collection <strong>of</strong> G. Frey and W. Wittmer. Berkeley. EMEC. Essig Museum <strong>of</strong> Entomology, Department <strong>of</strong> Integrative Biology, University <strong>of</strong> California, Berkeley, California, 94720. USA. Berlin. ZMHB. Museum für Naturkunde der Humbolt Universitat zu Berlin, Bereich Zoologisches Museum, Invalidenstrasse 43, 1040, Berlin, Germany. Bozeman. MTEC. Montana State University Entomology Collection, Entomology Research Laboratory, Department <strong>of</strong> Biology, Montana State University, Bozeman, Montana, 59715, USA. Containing <strong>the</strong> important collection <strong>of</strong> West Indian insects <strong>of</strong> M. A. Ivie. Bruner. S.C. Bruner collection, from <strong>the</strong> Santiago de las Vegas Agriculture experiment station (EEA), supposedly now in Havana at <strong>the</strong> Institute <strong>of</strong> Ecology and Systematics (IES). Brno. MMBC. Department <strong>of</strong> Entomology, Moravské Museum, Preslova ul. 659 37, Brno, Czech Republic. Brussels. ISNB. Institut Royal des Sciences Naturelles de Belgique, 29 rue Vautier, B1040, Brussels, Belgium. Bucarest. Institute <strong>of</strong> Speologie “Emile Racovitza”, Str. Mikail Moxa 9, Bucarest 12, Romania. This may contain material from <strong>the</strong> <strong>Cuba</strong>n-Roumanian cave biology expeditions. Cambridge. MCZC. Entomology Department, Museum <strong>of</strong> Comparative Zoology, Harvard University, 22 Oxford St., Cambridge, Massachusetts, 02138, USA. A collection strong in <strong>Cuba</strong>n material from field work <strong>of</strong> P. J. Darlington, Jr, and o<strong>the</strong>rs. Champaign. INHS. Illinois Natural History Survey Insect Collection, 607 E. Peabody Dr., Champaign, Illinois 61820, USA. With <strong>Cuba</strong> material collected by M. W. Sanderson. Chicago. FMNH. Insect Collection, Division <strong>of</strong> Insects, Field Museum <strong>of</strong> Natural History, Roosevelt Road and Lake Shore Drive, Chicago, Illinois, 60605, USA. Containing <strong>the</strong> important Bierig collection <strong>of</strong> Staphylinoidea. Columbus. OSUC. Ohio State University Collection <strong>of</strong> Insects and Spiders, 1735 Neil Avenue, Columbus, Ohio, 43210, USA. Eberswalde. DEIC. Deutsche Entomologische Institut, Eberswalde Finow 1, 1300, Germany. Franz. H. Franz collection, Mödling, Austria. Now placed in <strong>the</strong> Vienna Museum, Austria. Geneva. MHNG. Department des Insectes, Museum d’Histoire Naturelle, CP 434, CH-1211, Geneva 6, Switzerland. Gundlach. J. Gundlach collection. Now a part <strong>of</strong> <strong>the</strong> collections <strong>of</strong> <strong>the</strong> Institute <strong>of</strong> Ecology and Systematics (IES) in Havana. The Gundlach code number relates <strong>the</strong> specimen to <strong>the</strong> name and locality as published in volume 3 <strong>of</strong> this “Contribucion a la Entomologia <strong>Cuba</strong>na.” Garrido. Orlando Garrido, Havana, <strong>Cuba</strong>. A private collection specializing in Tenebrionidae. Gainesville. FSCA. Florida State Collection <strong>of</strong> Arthropods, Division <strong>of</strong> Plant Industry, 1911 34 th St. SW, PO Box 147100, Gainesville, Florida, 32614, USA. The Strohecker collection <strong>of</strong> Endomychidae and o<strong>the</strong>r large collections from throughout <strong>the</strong> Caribbean are here. This institution also houses <strong>the</strong> important and large private collection <strong>of</strong> New World scarabs and o<strong>the</strong>r families <strong>of</strong> beetles <strong>of</strong> R. E. Woodruff Havana. IZAC. Collections <strong>of</strong> <strong>the</strong> Institute <strong>of</strong> Ecology and Systematics (IES) <strong>of</strong> <strong>the</strong> Academia de Ciencias de <strong>Cuba</strong>, at Carretera de Varona km 3 ½, Capdevila, Boyeros, La Habana, <strong>Cuba</strong>; containing <strong>the</strong> historically important Gundlach collection, and <strong>the</strong> former collection <strong>of</strong> <strong>the</strong> Instituto de Zoologia de la Academia de Ciencias de <strong>Cuba</strong> (IZAC), Havana. It may also contain some or all <strong>of</strong> (1) <strong>the</strong> former collections <strong>of</strong> <strong>the</strong> Estacion Experimental de Agronomia (EEA), once located at Santiago de las Vegas, near Havana, (2) <strong>the</strong> reference collections <strong>of</strong> <strong>the</strong> <strong>Cuba</strong>n Sugar Club (CSC), (3) <strong>of</strong> <strong>the</strong> Department <strong>of</strong> Speleology, Academy <strong>of</strong> Sciences, <strong>with</strong> some <strong>of</strong> <strong>the</strong> specimens from <strong>the</strong> <strong>Cuba</strong>n-Roumanian cave-biology expeditions, and (4) <strong>the</strong> tenebrionid collection <strong>of</strong> O. Garrido. Part or all <strong>of</strong> <strong>the</strong>se collections may move to <strong>the</strong> National Museum <strong>of</strong> Natural History, located in downtown Old Havana (Habana Vieja). Halle. MLUH. Wissenschaftsbereich Zoologie, Sektion Biowissenschaften Martin-Lu<strong>the</strong>r Universitat Halle, WB Zoologie, Domplatz 4, 4020 Halle (Salle), Germany. Possibly contains <strong>Cuba</strong>n species in <strong>the</strong> large and important collections <strong>of</strong> Burmeister, Germar, and especially that <strong>of</strong> Suffrian. Helsinki. MZHF. Zoological Museum, Finnish Museum <strong>of</strong> Natural History, Universitets Helsinki, P. Rautienkatu 13, SF-00100, Helsinki 10, Finland . Howden. HAHC. Henry and Anne Howden Collection, Ottawa, Ontario, Canada. Placed in <strong>the</strong> Entomology Division, Canadian Museum <strong>of</strong> Nature, Aylmer, Quebec, (PO Box 3443, Station D, Ottawa, ON K1P 6P4), Canada.
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