Zootaxa, Taxonomy of Serpulidae (Annelida ... - Magnolia Press

Schröder 1971), France (Fauvel 1927, Zibrowius 1968a), west coast of Africa (Zibrowius 1973b), north coast
of Tunis (Zibrowius 1979a). The species was confused with Rhodopsis (by Straughan 1967a fig. 5i). Ben-
Eliahu & Payiatas (1999) provided SEMs of Josephella chaetae. Dew (1959) mentioned a single specimen
with two opercula equal in size.
Josephella marenzelleri Caullery & Mesnil, 1896, circum(sub)tropical, temperate.
19. Laminatubus ten Hove & Zibrowius, 1986
(Fig. 26)
Type-species: Laminatubus alvini ten Hove & Zibrowius, 1986
Number of species: 1
Tube white, more or less triangular in cross-section, with large undulating longitudinal keel; consisting of two
layers: an inner opaque layer and an outer (not granular) hyaline one. Operculum globular, with bulbous
proximal ampulla and more or less flattened distal part with thickened cuticle. Peduncle cylindrical, gradually
merging into opercular ampulla, constriction absent; inserted to left side, proximal from first and second
normal radiole. Pseudoperculum absent. Radioles not connected by inter-radiolar membrane, arranged into
slightly ascending spiral of up to two whorls. Up to 33 radioles per lobe. Stylodes and branchial eyes absent.
Mouth palps not observed. 6 thoracic chaetigerous segments. Collar with medio-ventral and two latero-dorsal
lobes, continuous with thoracic membranes, forming apron. Tonguelets absent. Collar chaetae Spirobranchustype
(Fig. 26A) and limbate. Apomatus chaetae absent. All uncini saw-shaped with 5–7 teeth, anterior fang
simple, pointed (Fig. 26D, C). Thoracic tori converging posteriorly, forming triangular depression. Abdominal
chaetae long, with hollow trumpet-shaped tip, smoothly bent (Fig. 26D). Posterior chaetae become longer, but
posterior capillary chaetae absent. Achaetous anterior abdominal zone absent. Posterior glandular pad absent.
Remarks. The species is a common element of the bathyal hydrothermal vent communities found in the
Galapagos rift and the East Pacific Rise.
Laminatubus alvini ten Hove & Zibrowius, 1986, East Pacific, 1–21º N; bathyal.
20. Marifugia Absolon & Hrabĕ, 1930
(Fig. 27)
Type-species: Marifugia cavatica Absolon & Hrabĕ, 1930
Number of species: 1
Tube white, opaque, circular in cross-section; irregular longitudinal keel and collar like rings may be present.
Thin hyaline granular overlay of the tube present. Operculum fig-shaped to inverse conical, with (or without)
chitinous endplate. Peduncle flattened cylindrical, smooth, without distal wings, gradually merging into
opercular ampulla; inserted just below and between first and second dorsal radiole on left side (in large
specimen almost covering base of branchial lobe). Pseudoperculum absent. Radioles arranged in semi-circles,
up to 6 per lobe. Inter-radiolar membrane, branchial eyes and stylodes absent. Mouth palps not found. 6
thoracic chaetigerous segments. Collar non-lobed but with low medio-ventral projection. Thoracic
membranes narrow but forming apron. Tonguelets absent. Collar chaetae absent. Thoracic chaetae limbate,
Apomatus chaetae absent (Fig. 27A). Thoracic uncini saw-to-rasp-shaped, with about 8 teeth in profile, up to
4 in a row above blunt almost square shallowly gouged anterior peg (dental formula P:4:3:2:1:1:1:1:1, Fig.
27B). Triangular depression absent. Abdominal chaetae trumpet-shaped, long, smoothly bent, with hollow tip
bordered with pointed teeth (Fig. 27D). Posterior abdominal capillaries not observed. Uncini saw-to-rasp-
TAXONOMY OF SERPULIDS: STATE OF AFFAIRS
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