Keys to the macrolichens and checklist of the lichens and ...

19a. Medulla C+ red; lecanoric acid present ............................................. H. woitapensis
19b. Medulla C-; lecanoric acid absent.......................................................................... 20
20a. Medulla P+ orange or red; fumarprotocetraric, protocetraric and/or quaesitic acids
present.................................................................................................................... 21
20b. Medulla P-; fumarprotocetraric, protocetraric and quaesitic acids absent............. 24
21a. Medulla K+ yellow then dark red; quaesitic acid present .......................H. quaesita
21b. Medulla K- or K+ yellow-brown; quaesitic acid absent........................................ 22
22a. Medulla K-, KC+ orange ; fumarprotocetraric and barbatic acids present.
...............................................................................................................H. malesiana
22b. Medulla K+ yellow-brown; KC+ pink; protocetraric acid present........................ 23
23a. Medulla yellow in part; pigmentosin A present.................................... H. consimilis
23b. Medulla white thoughout; pigmentosin A absent ..................................H. koyaensis
24a. Medulla KC+ orange; barbatic acid present .......................................................... 25
24b. Medulla KC-; fatty acids present ........................................................................... 28
25a. Isidia inflated; apices becoming erumpent ................................................. H. toiana
25b. Isidia cylindrical; apices intact............................................................................... 26
26a. Echinocarpic acid present ........................................................................... H. addita
26b. Echinocarpic acid absent........................................................................................ 27
27a. Upper surface distinctly maculate; obtusatic and norobtusatic acids (submajor)
present.................................................................................................H. imbricatula
27b. Upper surface emaculate; obtusatic and norobtusatic acids present in trace
quantities or absent ................................................................................ H. orientalis
28a. Thallus loosely adnate, saxicolous or terricolous; lobes broad, 4-12 mm wide.
................................................................................................................. H. terricola
28b. Thallus adnate, corticolous; lobes narrower, 1-6 mm wide................................... 29
29a. Lobes narrow, 1-3 mm wide; ascospores large, 20-23 x 12-15 µm; lichesterinic
and protolichesterinic acids present.................................................H. lichesterinica
29b. Lobes broader, 2-6 mm wide; ascospores small, 8-10 x 4-6 µm; constipatic,
protoconstipatic and protopraesorediosic acids present................... H. costaricensis
30a. Thallus pustulate-sorediate .................................................................................... 31
30b. Thallus sorediate; pustules absent.......................................................................... 35
31a. Upper cortex UV+ yellow; lichexanthone present................................ H. osseoalba
31b. Upper cortex UV-; lichexanthone absent............................................................... 32
32a. Thallus laciniate; medulla C+ rose-red; gyrophoric acid present ......... H. monticola
32b. Thallus elaciniate; medulla C-; gyrophoric acid absent......................................... 33