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IntensIve sIlvIculture - Forest Science Labs - Research Network ...

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136<br />

Keywords: nursery operations<br />

nursery fertilization<br />

tree physiology<br />

Abstract: Seedlings of Douglas fir (Pseudotsuga menziesii) and ponderosa pine (Pinus<br />

ponderosa) were grown in a controlled environment and given nutrient solutions containing 0 (-<br />

N) or 150 ppm nitrogen (+N). Seedling growth, and the concentrations of N and carbohydrates in<br />

their tissues were affected by the treatments. Metabolically active tissues, such as roots, incubated<br />

with a limited supply of O2 became hypoxic faster and synthesized more ethanol than less active<br />

tissues, such as needles. All tissues that were incubated for 4 h in N2 synthesized ethanol.<br />

Needles incubated in N2 and light had much lower quantities of ethanol than needles in N2 and<br />

dark, suggesting that O2 from photosynthesis limited internal anoxia. Most tissues from +N<br />

seedlings synthesized greater quantities of ethanol in N2 anoxia than tissues from -N seedlings,<br />

probably because they were able to produce more enzymes with a greater availability of N.<br />

However, this increase in ethanol synthesis between N treatments was most pronounced in the<br />

phloem. Ethanol and soluble sugar concentrations were negatively related in needles and<br />

positively related in roots of +N seedlings, but not -N seedlings. Starch concentrations had no<br />

effect on the amount of ethanol produced by any tissue. Regardless of N treatments, all tissues<br />

from ponderosa pine produced more N2-induced ethanol than Douglas-fir, in part because its<br />

tissues contained different concentrations of soluble sugars and N as a consequence of<br />

phenological differences between the species. However, ponderosa pine tissues may also<br />

maintain greater quantities of anaerobic enzymes, or their isoenzymes than Douglas fir.<br />

309. Kennedy, R.W. 1995. Coniferous wood quality in the future: concerns and strategies. Wood<br />

<strong>Science</strong> and Technology 29:321-338.<br />

Keywords: planting operations<br />

wood quality<br />

Abstract: As the raw material base for forest products manufacturing shifts from oldgrowth<br />

to short-rotation plantation stock, the wood from these younger trees will contain larger<br />

proportions of juvenile wood. This in turn will influence the quality of forest products obtained.<br />

The pattern of specific gravity variation in these trees, which varies among the five most<br />

important Pacific Northwest species groups, is reviewed, and the nature of their differences is<br />

related to growth habit. It is speculated that the shade intolerance of some species manifests itself<br />

in an early culmination of annual height increment, after which specific gravity increases rapidly<br />

to a maximum. This is contrasted to shade-tolerant species, in which specific gravity may take<br />

several decades to attain a minimum value, followed by only moderate increases thereafter. In<br />

addition, faster growth rates in widely spaced plantation trees tend to depress specific gravity and<br />

advance the age at which these trees reach their minimum value, thereby compounding the<br />

overall wood density deficit of short-rotation trees. Lower specific gravity, compounded with<br />

reduced lignin content in juvenile wood, negatively influences kraft pulp yield, but not pulp<br />

quality parameters such as sheet density, burst and tensile strength. Reduced wood density,<br />

coupled with larger fibril angles in juvenile wood, reduces average strength and stiffness of<br />

lumber from younger plantation trees. Mechanical stress rating needs to be adopted to segregate<br />

the strong, stiff material for engineered construction uses, because a large proportion of visually<br />

graded lumber from juvenile wood zones will not meet currently assigned stress values.<br />

Mechanical stress rating can ensure a continued stream of appropriate engineering grades from<br />

future tree supplies. These conclusions are drawn from data in the literature on Pseudotsuga<br />

menziesii, Pinus contorta, Tsuga heterophylla, Picea spp., and Abies spp.

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