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around 1,476 feet (450 m), where the water temperature<br />

is between 46 and 54°F (8–12°C), or roughly the<br />

same as that found in Puget Sound, Washington; the age<br />

of one of these was determined to be 2,742 years. The<br />

specimens were not very large, less than 6.5 feet (2 m)<br />

in height, and the radial growth rates of the branches,<br />

or how fast a branch got thicker, were extremely low at<br />

0.013–0.001 inch (4–35 μm) per year. Savalia individuals<br />

of such ancient age are not restricted to Hawai’ian<br />

waters; a Savalia skeleton that had lived 1,800 years before<br />

it died was recently recovered near the Bahamas.<br />

Large stand of staghorn Acropora.<br />

Because they reproduce by fragmenting,<br />

a particular genome could theoretically<br />

be millions of years old.<br />

THINGS ARE LOOKING BLACK<br />

The Antipatharians, or black corals, are corals that have<br />

wholly proteinaceous skeletons and small polyps with<br />

only six tentacles. Typically found in deeper waters, a few<br />

species are common at diving depths, and occasionally a<br />

specimen is collected for the aquarium hobby. They may<br />

be quite large, particularly the whip-like wire or whip<br />

corals, Cirripathes species, which may reach over 16 feet<br />

(5 m) in length as they extend from deep reef walls, but<br />

most branching black corals look rather like gorgonians<br />

and are found in the same size range, up to 65 feet (a<br />

couple of meters) high. The common name, black coral,<br />

reflects the color of the skeleton, not the color of the living<br />

tissue, which typically is in hues of orange or yellow<br />

for shallower forms.<br />

Some of the deep-water species are stark white, as are<br />

individuals of a number of other deep-water species, particularly<br />

those of the stony coral Lophelia. Because the<br />

skeletons of black corals are proteinaceous, their ages are<br />

typically determined radiometrically, primarily using the<br />

abundance of the radioactive 14 C isotope of carbon incorporated<br />

into them as it was deposited. This isotope decays<br />

into the stable 12 C at a known and highly calibrated rate.<br />

By measuring the proportional abundances of both carbon<br />

isotopes, the age of the sample can be determined.<br />

To measure the ages of these corals, very small samples<br />

of the skeleton are vaporized and the isotope abundances<br />

are determined; then, using some reference data, it is<br />

relatively easy to determine the age of the sample. Examination<br />

of a few specimens of Antipathes dichotoma<br />

from shallow, 164-foot-deep (50-m) Hawai’ian waters<br />

showed ages from 12 to 32 years. These colonies were<br />

also growing quite rapidly, adding as much as 0.04 inch<br />

(1.1 mm) in branch diameter per year. On the other<br />

hand, samples from several individual corals in what are<br />

probably several different species of Leiopathes are the<br />

“All-Coral” winners of the “olde age lottery.” One specimen<br />

each of an indeterminate species and Leiopathes glaberrima<br />

from near Hawai’i were 4,265 and 2,377 years<br />

old, respectively. The radial growth rates of these colonies<br />

were less than 5μm per year. Leiopathes species are ubiquitous<br />

in the deep sea and have been collected from the<br />

Northwest Pacific, near Antarctica, and many areas in<br />

between. Given that the ages of fewer than a dozen speci-<br />

CBPIX/SHUTTERSTOCK<br />

82 CORAL

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