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mens from this genus appear to have been determined,<br />

the odds must be considerable that the ages of older animals—maybe<br />

vastly older animals—await discovery.<br />

FIRE IN THE HOLE<br />

There is really no way to age the calcareous skeletons<br />

of hydrocorals, and that presents particular problems in<br />

determining their ages. They form small colonies, but it<br />

certainly appears that some of them, particularly some<br />

of the colder water stylasterines, may live quite long.<br />

The few ecological studies on them indicate that once<br />

the colony gets about a year old, it may last decades, but<br />

there are no definitive data on how long they last. No<br />

ecological studies have been undertaken that give any<br />

idea of the parameters of individual growth rates, mortality<br />

rates, or causes of death.<br />

On the other hand, the milleporines, or fire corals,<br />

of tropical reefs that form the other end of the coral age<br />

scale from the deep-water “oldies” are reasonably well<br />

studied. Fire corals are rapidly growing, highly adventitious,<br />

and temporally transitory animals. In other words,<br />

they are the coral equivalent of weeds and possess an<br />

ecological growth strategy that can be summed up with<br />

the phrase: “Live Fast, Die Young, and Leave a Butt-Ugly<br />

Corpse.” Varying from reef to reef and species to species,<br />

a reasonable average for the maximum age of most fire<br />

coral colonies is about a decade. Maximal growth rates<br />

appear to be around an inch (2.5 cm) a year.<br />

SO…HOW OLD CAN A CORAL GET?<br />

Obviously, it depends on the coral—both the coral type<br />

and the specific coral species within any given type.<br />

What it doesn’t seem to depend upon is geographical<br />

region; old corals of one sort or another may be found<br />

anywhere and everywhere. In Table 1, I have given a few<br />

representative data about the extreme life spans for various<br />

types of coral living in various habitats. Perhaps a<br />

general rule of coral longevity may be developed, but if<br />

so that rule—and the work to support it—are still largely<br />

in the future. In the interim, it is probably best to deal<br />

with those few coral species whose longevities have been<br />

determined on a case-by-case basis.<br />

Given the paucity of examples, it is prudent to avoid<br />

drawing too many conclusions about coral ages, but<br />

gingerly stepping out on the limb of the fragile branching<br />

Acropora, I will make some. First, it appears that on<br />

average, deep-water corals probably live longer than do<br />

shallow-water ones. Second, it appears that long-lived<br />

corals are slow-growing corals. Finally, there is the wild<br />

card of asexual reproduction. To date, there has been<br />

no study that gives us a handle on the longevity of the<br />

genomes of highly fragmenting corals, such as various<br />

84 CORAL

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