state and is gathering data hitherto unavailable.Even as the information is collected, large tracts arebeing threatened by major development.Occurrences <strong>of</strong> <strong>cedar</strong> in the state may begrouped in three broad classes (1) pure forest standswith little other canopy vegetation (the most common<strong>cedar</strong> community <strong>of</strong> the mainland), (2) mixed stands,with <strong>cedar</strong> occurring among other wetland trees,primarily red maple, and (3) in kettles with an openbody <strong>of</strong> water surrounded by a succession <strong>of</strong> zonesin which <strong>cedar</strong> is one <strong>of</strong> the concentric rings <strong>of</strong>vegetation.An example <strong>of</strong> the vegetation sequence surroundinga kettle pond would be: a band <strong>of</strong> emergentswamp loosestrife (Decodon verticillatus) rimmedby a Sphagnum-based mat, on which there is a succession<strong>of</strong> narrow shrub zones starting with perhapssome dwarf huckleberry (Gaylussacia dumosa),leatherleaf (Chamaedaphne calyculata), blueberry(Vaccinium spp.), and swamD azalea(~hododendron~viscosurn), which sharply gradeinto <strong>Atlantic</strong> <strong>white</strong> <strong>cedar</strong>, and finally <strong>white</strong> pine, hemlock,and upland species. Somet~~ical plants <strong>of</strong>thewestern sections <strong>of</strong> Kent and Providence Counties(D. Lowry and F. Golet, pers. comm.) <strong>The</strong>re is very little<strong>cedar</strong> on the east side <strong>of</strong> the Bay, although placenames such as "Cedar Swamp" suggest that thespecies was more common there in the past.<strong>The</strong> largest stands <strong>of</strong> <strong>cedar</strong> occur within thestate's three largest <strong>wetlands</strong>, all <strong>of</strong> which aresituated on broad expanses <strong>of</strong> stratified drift less than30 m above sea level. Cedar forest covers 240 ha <strong>of</strong>the 870-ha Chapman Swamp in Westerly. <strong>The</strong>remainder <strong>of</strong> this highly diverse wetland includesdeciduous forest, shrub swamp, bog, marsh, andopen water. Two-thirds <strong>of</strong> the 390-halndian <strong>cedar</strong>Swamp in Charlestown supports <strong>cedar</strong>, but redmaple (Acerrubrum) is the dominant species in most<strong>of</strong> the stands in which <strong>cedar</strong> occurs. In the GreatSwamp, which occupies 1200 ha in SouthKingstown, Richmond, and Charlestown, <strong>cedar</strong>covers some 90 ha; the great majority <strong>of</strong> this wetlandconsists <strong>of</strong> deciduous forest and shrub swamp.Smaller stands <strong>of</strong> <strong>cedar</strong> are commonlyfound in glacial kettles (ice-block basins) whichformed in stratified drift or in thick deposits <strong>of</strong>Open Sphagnum =One would be pitcher plant (Sar- morainal material. A highly unusual stand <strong>of</strong> <strong>Atlantic</strong>racenia ~ur~urea), s~rldew (Drosera intermedia), <strong>white</strong> <strong>cedar</strong> occupies a kettle situated in omash atand occasional orchids such as rose ~ogonia the edge <strong>of</strong> Factory Pond, 9 m above sea level in(Pogonia o~hioglossoides) or grass pink South Kingstown. <strong>The</strong> trees in this 5-ha "forest" are(Calopogon pulchellus) .80 years old, but only 1-1.5 m tall. Bordered by theA variation <strong>of</strong> this vegetation type is found on pond On One side, the stand is separated from the ad-Cape Cod, where <strong>cedar</strong>s may occupy relatively flat- jacent upland by a moat <strong>of</strong> open water and a quakingsurfaced kettles rimmed by a moat slightly deeper mat low shrubs- <strong>The</strong> surface Of this dwarf <strong>cedar</strong>than the body <strong>of</strong> the wetland. <strong>The</strong> <strong>cedar</strong>s, <strong>of</strong>ten the bog is carpeted throughout with Sphagnum moss.sole canopytree, cluster on small hummocksthatare <strong>The</strong> Water table Stays within a few centimeters <strong>of</strong> thespotted over the entire basin. <strong>The</strong> concentricvegeta- surface all Year, and the pH <strong>of</strong> the soil water drops astion pattern is condensed on each hummock, with IOW as 3.1. <strong>The</strong> soil is a p00dy decomposed, fibricericaceous shrubs, sweet pepperbush (Clethra a/- Peat. Growing in association with the <strong>cedar</strong>s arenifolia), and ferns in tight array rising from a sphag- leatherleaf, cranberries (Vaccinium macrocarpon, V.nous carpet that continues into the water <strong>of</strong> the OXYCOCCOS)I cOtfongraSS (Erio~horum SP.)~ andhollows.pitcher plant. At its deepest point, this kettle contains9 m <strong>of</strong> peat.Species otherwise rare in southern NewEngland are found in Chamaecyparis <strong>wetlands</strong>, e.g.,dwarf mistletoe (Arceuthobium pusillum), a tinyflowering parasite that causes deformation anddeath <strong>of</strong> at least the branches <strong>of</strong> the black spruce onwhich it grows; and heartleaf twayblade (Listera tordata),a northern species at its southern limit in CapeCod (the only known extant location in the state).<strong>The</strong> northern parula warbler (Parula americana) inMassachusetts now breeds primarily in a few <strong>cedar</strong><strong>wetlands</strong>, as the hanging lichen Usnea, its favorednesting material, is fast disappearing outside the<strong>cedar</strong> swamps.Rhode. In Rhode Island, <strong>Atlantic</strong><strong>white</strong> <strong>cedar</strong> is most abundant west <strong>of</strong> NarragansettBay, particularly in Washington County and in the14Cedar <strong>wetlands</strong> along the Connecticut borderin western Rhode Island generally lie at elevationsranging from 90 to 180 m. Most <strong>of</strong> these havedeveloped over valley train deposits <strong>of</strong> stratified driftor in association with ice contact deposits. A verysmall percentage <strong>of</strong> these swamps lie directly onbedrock or on unstratified drift (more commonlyknown as glacial till). Most wetland basins in till orbedrock tend to be small, and peat deposits seldomexceed 2-3 m in thickness.Red maple and black gum (Nyssa sylvatica)are the two tree species most commonly associatedwith <strong>Atlantic</strong> <strong>white</strong> <strong>cedar</strong> throughout Rhode Island,but eastern hemlock (Tsuga canadensis) is animportant associate in many <strong>of</strong> the swamps lying
above 90 m. In a small number <strong>of</strong> <strong>wetlands</strong> in northwesternRhode Island, <strong>cedar</strong> grows in associationwith two boreal species, black spruce (Piceamariana) and larch (Larix larina) (R. Enser, pers.comm.).Great laurel (Rhododendron maximum), abroad-leaved evergreen shrub which is common inupland areas <strong>of</strong> the southern Appalachians (Fernald1950), is locally common as an understory species inboth deciduous and evergreen wetland forests insouthern Rhode Island and nearby Connecticut.This shrub grows to a height <strong>of</strong> 2.5 to 4.5 m and <strong>of</strong>tenforms such dense tangles that travel through theswamps is exceedingly difficult. As a result <strong>of</strong> thedeep shade created by a dense canopy <strong>of</strong> <strong>cedar</strong> anda thick understory <strong>of</strong> great laurel, herbs are scarce tononexistent in these swamps (Lowry 1984).A striking example <strong>of</strong> the <strong>Atlantic</strong> <strong>white</strong><strong>cedar</strong>-great laurel association can be seen in the EllPond-Long Pond Natural Areas Complex near theConnecticut line in Hopkinton. <strong>The</strong>re a dense, 90-year old <strong>cedar</strong> forest containing hemlock as well asgreat laurel borders the northern and western shores<strong>of</strong> Ell Pond, which lies in a deep fracture in the localbedrock. <strong>The</strong> surrounding relief is rugged andbedrock outcrops are numerous. Between the forestand the water's edge is a narrow bog mat dominatedby leatherleaf. Peat thickness ranges from 4 m in theforest interior to 8-9 m at the water's edge. <strong>The</strong> EllPond stand, which averages 13 m in height, is 98 mabove sea level. Ell Pond and its associated <strong>wetlands</strong>represent Rhode Island's only <strong>National</strong> NaturalLandmark. For further description <strong>of</strong> Rhode Islandsites, see Lowry (1 984) and Golet and Lowry (1987).Connecticut. Thirty-nine <strong>cedar</strong> <strong>wetlands</strong>, allbut six <strong>of</strong> them east <strong>of</strong> the Connecticut River, areknown to contain living <strong>cedar</strong> in Connecticut atpresent (K. Metzler, pers. comm.). Some sitesare reportedto be in near-pristine condition, some aretrampled and debris-strewn, and some are still beinglogged for <strong>cedar</strong>. A few are in public ownership, butmost have no active conservation manaaement. -Two <strong>cedar</strong> <strong>wetlands</strong> were designated as <strong>National</strong>Natural Landmarks in 1973: Chester CedarSwamp, and Pachaug Great Meadow in Voluntown.A <strong>cedar</strong> log walkway and marked trail traverse a section<strong>of</strong> the Pachaug preserve containing over 200 ha<strong>of</strong> <strong>cedar</strong> in an approximately 350 ha swamp-bogsedgemeadow complex (K. Metzler, pers. comm.)drained by the Pachaug River. Pachaug and at leasttwo other stands are known to contain sizable, vigorous,dense great laurel populations (LedyardCedar Swamp, and Bell Cedar Swamp in NorthStonington) (K. Metzler, pers. comm.). Creepingsnowberry (Gaultheria hispidula) is reputed to growin one privately-owned swamp. North WindhamPeat Bog contains a dense 30-ha <strong>white</strong> <strong>cedar</strong> swampwith black spruce, unusual in Connecticut. It is acombination not seen south <strong>of</strong> this point except in themontane Sterling Forest, New York and High Point,New Jersey forests (Laderman, unpubl.).Monographs by Nichols (1913) and Taylor(1915), and a master's thesis by Noyes (1939) constitutethe major sources <strong>of</strong> historical botanical dataabout Chamaecyparis in the state. <strong>The</strong> papers containlists <strong>of</strong> associated species, brief site descriptions,and maps, indicating that <strong>of</strong> 86 <strong>cedar</strong> standsknown at the time, 85% were east <strong>of</strong> the ConnecticutRiver.New York State. Before the agricultural andsuburban development <strong>of</strong> Long Island, <strong>cedar</strong>swamps were believed to form an almost continuouschain from Brooklyn to Montauk Point (Nichols1913), clustered along the southern edge <strong>of</strong> the terminalmoraine that forms the island's spine. As civilizationspread, <strong>cedar</strong> <strong>wetlands</strong> declined drastically(Torrey 1843; Harper 1907; Bicknell 1908; Taylor1916).<strong>The</strong> primary cause <strong>of</strong> <strong>cedar</strong> loss in NassauCounty was lowering <strong>of</strong> the water table when streamswere dammed to create reservoirs for the rapidly expandingpopulace. Nassau County today holds fewmature <strong>cedar</strong>s, with no evidence <strong>of</strong> regeneration (J.Turner, pers. comm.).In Suffolk County, earlier in this century,many <strong>wetlands</strong>were lumbered, drained, and clearedfor farming. Those remaining are being rapidlyredaced bv summer resorts and second homes.he county now contains only 11 known <strong>cedar</strong>stands, most <strong>of</strong> them quite small. SouthamptonTownship harbors the greatest abundance <strong>of</strong> <strong>cedar</strong>sin Long Island. <strong>The</strong> largest New York wetland complexcontaining Chamaecyparis is in a 40-ha area <strong>of</strong>Southampton's Cranberry Bog County Park, alongthe southern reaches <strong>of</strong> the Peconic River (J. Turner,pers. comm.).Outside Long Island, the only <strong>cedar</strong> standsremaining in the state are two small bogs in SterlingForest, each less than 0.5 ha (Lynn 1984; Lynn andKarlin 1985).New Jersey. Glaciated New Jersey has onlyseven known <strong>cedar</strong> stands, but it bears the distinction<strong>of</strong> harboring an <strong>Atlantic</strong> <strong>white</strong> <strong>cedar</strong> swamp inHigh Point at the greatest altitude recorded for thespecies. Its elevation <strong>of</strong> 457 m exceeds that <strong>of</strong> thenext highest stand (in New Hampshire) by 69 m.Only three northern New Jersey sites contain morethan a few trees at present: High Point andWawayanda in Sussex County in the far northwest
- Page 1: Biological Report 85(7.21)July 1989
- Page 5 and 6: PREFACEThis monograph on the ecolog
- Page 7 and 8: CONTENTSea9tl.PREFACE .............
- Page 9 and 10: NumberTABLESEarliest records of Atl
- Page 11 and 12: - CHAPTER 1 -INTRODUCTION1 .I GENER
- Page 13 and 14: TEMPORARLYSEASONALLY1I SATURATED1Fi
- Page 15 and 16: Stagnant,,- melting Icel Block d~ag
- Page 17: Table I. Earliest records of Atlant
- Page 20 and 21: CHAPTER 2 -REGIONAL OVERVIEW2.1 INT
- Page 22 and 23: stands are scattered north and west
- Page 26 and 27: corner of the state, and Uttertown
- Page 28 and 29: hardwood stands, or as isolated tre
- Page 30 and 31: cedar have comprised 40%-60% of the
- Page 32 and 33: tablished. Drainage from 224 krn of
- Page 34 and 35: perennial streams (Figure 18) and i
- Page 36 and 37: - CHAPTER 3 -CHAMAECYPARIS THYOIDES
- Page 38 and 39: Ms&gua As early as 1923, Akermandes
- Page 40 and 41: - CHAPTER 4 -STRUCTURE AND FUNCTION
- Page 42 and 43: ottomland-isolatedtill & bedrockupl
- Page 44 and 45: Table 5. Mean August tissue nutrien
- Page 46 and 47: - CHAPTER 5 -BIOLOGICAL COMPONENTS
- Page 48 and 49: Figure 24a. Companions: plants freq
- Page 50 and 51: Figure 24c. Companions: Plants freq
- Page 52 and 53: ?able 7. Comparison of bird species
- Page 54 and 55: Meanley (1979) emphasized the impor
- Page 56 and 57: - CHAPTER 6 -MANAGEMENT AND HARVEST
- Page 58 and 59: FireBurnedtreesPeatGlacialrubble-Tr
- Page 60 and 61: Mixedhardwood& shrubsPeatGlacialrub
- Page 62 and 63: acidity is reduced, and ammonia, ph
- Page 64 and 65: and condition of the soil surface a
- Page 66 and 67: sprouts and shade-tolerant shrubs g
- Page 68 and 69: Figure 32. Atlantic white cedar reg
- Page 70 and 71: sedhflings, conversely, canopy open
- Page 72 and 73: FORESTMANAGEMENTSCHEMAT I CLEGEND-
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-.- Rsfupe BoundaryState BoundaryRo
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NATIONAL WILDLIFE RERX;EVirginia an
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Figure 38. Alligator River (North C
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Mainland Dare County is located on
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The cedar swamp forests along the A
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IPure Atlanticwhite cedar standsy-J
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Generally, the mixed swamp forest s
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species of mammals are recorded by
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Braun-Blanquet, J. [I 9321 1983. Pl
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Ferguson, R.H., and C.E. Meyer. 197
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Littte, S. 1953. Prescribed burning
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Porter, D.M. 1979. Rare and endange
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Gov. Print. Off., Washington, DC. (
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APPENDIX A. Flora Associated with C
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APPENDIX A. Flora: Trees (6ontin~ed
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APPENDIX A. Flora: Shrubs (Continue
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APPENDIX A. Fiora: Herbs (Continud)
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APPENDIX A. Flora: Herbs (Continued
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APPENDIX A. flora: Herbs (Continued
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APPENDIX A. Flora: Herbs (Continued
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APPENDIX A. Flora: Herbs (Continued
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APPENDIX B. FAUNA OF ATLANTIC WHITE
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APPENDIX C. Hydric Soilsis a soil t
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Frost, CecilFuller, ManleyFunk, Dav
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Ward, Daniel B.Whigham, DennisWidof