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the north – substantial in terms of numbers, but really only denting the Celtic substructure.<br />
Lastly, I have found a tiny number of very unusual clans in the southern part of England. Two of<br />
these are from sub-Saharan Africa, three from Syria or Jordan. These exotic sequences are found<br />
only in England, with one exception, and among people with no knowledge of, or family<br />
connections with, those distant parts of the world. I think they might be the descendants of Roman<br />
slaves, whose lines have kept going through unbroken generations of women. If this was the genetic<br />
legacy of the Romans, they have left only the slightest traces on the female side. I have not found<br />
any in Wales, or in Ireland and only one in Scotland. This is an African sequence from Stornoway<br />
in the Western Isles, for which I have absolutely no explanation. These exotic dustings, and the<br />
more substantial layers of Viking maternal lines, are the exception. Everything else in the Isles, on<br />
the maternal side, is both Celtic and ancient. But what about the men?<br />
Here again, the strongest signal is a Celtic one, in the form of the clan of Oisin, which<br />
dominates the scene all over the Isles. The predominance in every part of the Isles of the Atlantis<br />
chromosome (the most frequent in the Oisin clan), with its strong affinities to Iberia, along with<br />
other matches and the evidence from the maternal side convinces me that it is from this direction<br />
that we must look for the origin of Oisin and the great majority of our Y-chromosomes. The sea<br />
routes of the Atlantic fringe conveyed both men and women to the Isles. I can find no evidence at<br />
all of a large-scale arrival from the heartland of the Celts of central Europe among the paternal<br />
genetic ancestry of the Isles, just as there is none on the maternal side.<br />
The pockets of ancient Wodans in mid-Wales and the ‘Pictland’ regions of Grampian and<br />
Tayside are, I believe, the echoes of the very first Mesolithic settlers who arrived from continental<br />
Europe, perhaps even travelling by foot while there was still a land connection. They look old to<br />
me, and for an apparently contradictory reason. That is because they are all very similar. The same<br />
applies to the Oisins. And yet the customs of genetics state that the longer a gene has been in a<br />
place, the more diversity should have accumulated. That was how I was able to fix the homelands<br />
of the seven European clan matriarchs. Using that rule I placed them at the locations where the<br />
present-day diversity was highest, and thus where they had had longest to accumulate mutations<br />
away from the original.<br />
But this rule does not seem to work with the paternal lines delineated by the Y-chromosome.<br />
The very striking thing about the clan of Oisin throughout the Isles is how very similar they all are.<br />
Or at least, how there are very large clusters of very similar chromosomes in one location, and not<br />
in others. For instance, the Ui Neill chromosome reaches a very high frequency in north-west<br />
Ireland but is rare elsewhere, and the Somerled chromosome is common in the Highlands and the<br />
Hebrides, but virtually unknown elsewhere – unless carried by a member of Clan Donald or Clan<br />
Dugall. This dramatically reduces the genetic diversity, and leads to very recent settlement dates,<br />
sometimes obviously incorrect. This has been noticed before with the Y-chromosome but has been<br />
attributed to what is called ‘patrilocality’. This is the practice of men staying put, while the women<br />
move to marry. However, I don’t think<br />
this works well enough to explain the amazing similarity in the Oisin chromosomes. The<br />
explanation is less cosy.<br />
This is the ‘Genghis effect’ and it is not confined to the Mongol Empire. In the Isles very large<br />
numbers of men, perhaps all of them in the clan of Oisin, are descended from only a few<br />
genetically successful ancestors. All the conditions are here in the Isles. From the Iron Age