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not forget. So far we have only mentioned immigration into Ireland, by Anglo-Normans at first and<br />

then through the plantations. But these are numerically dwarfed by the departures. Religious<br />

intolerance and persecution from the sixteenth century onwards, closely coupled to land seizure,<br />

drove many Catholic landowners abroad, mainly to France and Spain. Though doubtless traumatic<br />

for them, these exiles did not really affect ordinary Irish agricultural workers, for whom life<br />

continued much as before, though the land was under new ownership. However, in the nineteenth<br />

century, Irish emigration on a large scale began in earnest.<br />

In the first decades of the century, agricultural prices fell, estate rentals declined, investment in<br />

the land was reduced to a trickle, and the rural population grew. Whatever the ultimate causes of<br />

this cycle of economic decline, the effects on the rural poor were catastrophic. Reduced to almost<br />

complete dependence on the potato as the staple crop, the countryside was decimated when the<br />

crop was infested with the potato blight and rotted in the ground. During the Great Famine of the<br />

mid 1840s, thousands died of starvation or of the infectious diseases which swept through the<br />

malnourished population. Though many thousands died, thousands also made their escape. Ireland’s<br />

mid-nineteenth-century population of 8 million began a steady decline that has only very recently<br />

stabilized at 4.1 million in the Republic and 1.7 million in Ulster. The desparate diaspora of the<br />

Irish saw massive immigration both to Britain and to the New World, especially the United States.<br />

Today, there are far more ‘Irish’ genes abroad than there are in Ireland itself.<br />

Though Ireland is not yet united into a single political state, the poverty and suffering which<br />

suffuse all accounts of the history of the last centuries cannot be equated with Ireland today. The<br />

economy is transformed. The bars and cafés of Dublin are as lively and as sophisticated as<br />

anywhere in Europe. There is a tangible feeling of optimism in the air wherever you go. Though we<br />

will have to wait to see how much of the turmoil of past centuries is remembered by the genes, I<br />

suspect the main effect will be of emigration and the dispersal of Irish genes around the globe.<br />

Now that the future of Ireland as an independent country is looking so good, this is the time to move<br />

the sad centuries to one side and examine Ireland before the day when Henry II arrived to begin the<br />

English occupation. That is where we must seek to interpret the patterns of the genes. What do we<br />

know of these earlier times?<br />

The appeal that Dermot MacMurrough made to Richard de Clare to come to his aid, the appeal<br />

de Clare used as an excuse to invade, is a clear indication of the state of affairs in medieval Ireland<br />

– the struggle for dominance of one minor king against another. It is so very typical of the middle<br />

stage of evolution of any modern society and one that is only too visible in other parts of the world.<br />

Except that in those places, like Afghanistan or unstable African countries, these men are not<br />

dignified with the title ‘king’ but denigrated as ‘warlords’. In Ireland during the first millennium AD<br />

there was a constant struggle for dominance between different minor kings. According to one<br />

source, there may have been 150 of them at any one time, lending some credibility to the common<br />

Irish boast that they are all descended from lines of Irish kings. This may be something we can test<br />

as it could be visible in the Y-chromosome gene pool by what has come to be known as the<br />

‘Genghis Khan effect’.<br />

A few years ago, researchers from Oxford found a Y-chromosome that was very widespread<br />

throughout Asia, more or less within the geographical limits of the Mongol Empire. Finding a<br />

particular Y-chromosome with a specific fingerprint across such a wide area is highly unusual. Y-<br />

chromosomes are generally much more localized. The explanation, which I think is the correct one,

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