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http://jurassic.ru/

АКАДЕМИЯ НАУК СССР<br />

МИНИСТЕРСТВО ГЕОЛОГИИ<br />

И ОХРАНЫ НЕДР СССР<br />

МИНИСТЕРСТВО ВЫСШЕГО<br />

ОБРАЗОВАНИЯ СССР<br />

http://jurassic.ru/

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<br />

ПАЛЕОНТОЛОГИИ<br />

С ПРАВОЧНИК<br />

ДЛЯ ПАЛЕОНТОЛОГОВ И ГЕОЛОГОВ СССР<br />

ЧЛЕНИСТОНОГИЕ<br />

ТРАХЕЙНЫЕ И ХЕЛИЦЕРОВЫЕ<br />

От<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енный ред<strong>а</strong><strong>к</strong>т<strong>о</strong>р т<strong>о</strong>м<strong>а</strong><br />

Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф<br />

ИЗДАТЕЛЬСТВО АКАДЕМИИ НАУК СССР<br />

*<strong>Л1</strong> <strong>о</strong> <strong>с</strong> <strong>к</strong> <strong>в</strong> <strong>а</strong><br />

\ 9 6 2<br />

http://jurassic.ru/

УЧРЕЖДЕНИЯ, ПРИНИМАВШИЕ УЧАСТИЕ<br />

В СОСТАВЛЕНИИ ТОМА:<br />

ПАЛЕОНТОЛОГИЧЕСКИЙ ИНСТИТУТ АКАДЕМИИ НАУК СССР<br />

ЗООЛОГИЧЕСКИЙ ИНСТИТУТ АКАДЕМИИ НАУК СССР<br />

МОСКОВСКИЙ ГОСУДАРСТВЕННЫЙ УНИВЕРСИТЕТ «м. М. В. ЛОМОНОСОВА<br />

http://jurassic.ru/

(••г<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ь<br />

ПАЛЕОНТОЛОГИИ<br />

С П РАБ ОЧНИК<br />

ДЛЯ ПАЛЕОНТОЛОГОВ И ГЕОЛОГОВ СССР<br />

В ПЯТНАДЦАТИ ТОМАХ<br />

Гл<strong>а</strong><strong>в</strong>ный ред<strong>а</strong><strong>к</strong>т<strong>о</strong>р Ю. А. Орл<strong>о</strong><strong>в</strong><br />

З<strong>а</strong>м. гл<strong>а</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> ред<strong>а</strong><strong>к</strong>т<strong>о</strong>р<strong>а</strong>: Б. П. М<strong>а</strong>р<strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий, В. Е. Руженце<strong>в</strong>,<br />

Б. С. С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><br />

Ученые <strong>с</strong>е<strong>к</strong>рет<strong>а</strong>ри: Л. Д. Кип<strong>а</strong>ри<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, В. Н. Шим<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

Члены гл<strong>а</strong><strong>в</strong>н<strong>о</strong>й ред<strong>а</strong><strong>к</strong>ции: В. А. В<strong>а</strong>хр<strong>а</strong>мее<strong>в</strong>, Р. Ф. Ге<strong>к</strong><strong>к</strong>ер,<br />

В. И. Гр<strong>о</strong>м<strong>о</strong><strong>в</strong><strong>а</strong>, Л. Ш. Д<strong>а</strong><strong>в</strong>ит<strong>а</strong>ш<strong>в</strong>или, Г. Я. Крымг<strong>о</strong>льц,<br />

Н. П. Лупп<strong>о</strong><strong>в</strong>, Д. В. Обруче<strong>в</strong>, Н. К. О<strong>в</strong>еч<strong>к</strong>ин\,<br />

И. М. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я, В. Ф. Пчелинце<strong>в</strong>, Г. П. Р<strong>а</strong>дчен<strong>к</strong><strong>о</strong>,<br />

Д. М. Р<strong>а</strong>узер-Ч<strong>в</strong>рн<strong>о</strong>у<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф,<br />

А. К. Р<strong>о</strong>жде<strong>с</strong>т<strong>в</strong>ен<strong>с</strong><strong>к</strong>ий, Т. Г. С<strong>а</strong>рыче<strong>в</strong><strong>а</strong>, Н. Н. Субб<strong>о</strong>тин<strong>а</strong>,<br />

А. Л. Т<strong>а</strong>хт<strong>а</strong>дж<strong>а</strong>н, К. К. Флер<strong>о</strong><strong>в</strong>, А. В. Фур<strong>с</strong>ен<strong>к</strong><strong>о</strong>,<br />

А. В. Х<strong>а</strong>б<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>, Н. Е. Черныше<strong>в</strong><strong>а</strong>, А. Г. Эберзин<br />

ИЗДАТЕЛЬСТВО АКАДЕМИИ НАУК СССР<br />

jH <strong>о</strong> <strong>с</strong> 1<strong>с</strong> <strong>в</strong> <strong>а</strong><br />

\ 9 6 2<br />

http://jurassic.ru/

СОДЕРЖАНИЕ ТОМОВ<br />

Общ<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть. Пр<strong>о</strong><strong>с</strong>тейшие. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Д. М. Р<strong>а</strong>узер-Черн<strong>о</strong>у<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й и А. В. Фур<strong>с</strong>ен<strong>к</strong><strong>о</strong>.<br />

Губ<strong>к</strong>и, <strong>а</strong>рхе<strong>о</strong>ци<strong>а</strong>ты, <strong>к</strong>ишечн<strong>о</strong>п<strong>о</strong>л<strong>о</strong><strong>с</strong>тные. Прил<strong>о</strong>жение — чер<strong>в</strong>и. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией<br />

Б. С. С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>.<br />

М<strong>о</strong>ллю<strong>с</strong><strong>к</strong>и — п<strong>а</strong>нцирные,д<strong>в</strong>у<strong>с</strong>т<strong>в</strong><strong>о</strong>рч<strong>а</strong>тые, л<strong>о</strong>п<strong>а</strong>т<strong>о</strong>н<strong>о</strong>гие. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией А. Г.Эберзин<strong>а</strong>.<br />

М<strong>о</strong>ллю<strong>с</strong><strong>к</strong>и — брюх<strong>о</strong>н<strong>о</strong>гие. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией В. Ф. Пчелинце<strong>в</strong><strong>а</strong> и Я. Л. К<strong>о</strong>р<strong>о</strong>б<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong><br />

М<strong>о</strong>ллю<strong>с</strong><strong>к</strong>и — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>н<strong>о</strong>гие. I: н<strong>а</strong>утил<strong>о</strong>идеи, б<strong>а</strong><strong>к</strong>трит<strong>о</strong>идеи, <strong>а</strong>мм<strong>о</strong>н<strong>о</strong>идеи (<strong>а</strong>г<strong>о</strong>ни<strong>а</strong>-<br />

титы, г<strong>о</strong>ни<strong>а</strong>титы, <strong>к</strong>лимении). П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией В. Е. Руженце<strong>в</strong><strong>а</strong>.<br />

М<strong>о</strong>ллю<strong>с</strong><strong>к</strong>и — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>н<strong>о</strong>гие. II: <strong>а</strong>мм<strong>о</strong>н<strong>о</strong>идеи (цер<strong>а</strong>титы, <strong>а</strong>мм<strong>о</strong>ниты), <strong>в</strong>нутреннер<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>инные.<br />

Прил<strong>о</strong>жение — <strong>к</strong><strong>о</strong>ни<strong>к</strong><strong>о</strong>нхии. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Я. Я. Лупп<strong>о</strong><strong>в</strong><strong>а</strong> и<br />

В. В. Друщиц<strong>а</strong>.<br />

Мш<strong>а</strong>н<strong>к</strong>и, бр<strong>а</strong>хи<strong>о</strong>п<strong>о</strong>ды. Прил<strong>о</strong>жение — ф<strong>о</strong>р<strong>о</strong>ниды. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Т. Г. С<strong>а</strong>рыче<strong>в</strong><strong>о</strong>й.<br />

Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие — трил<strong>о</strong>бит<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные и р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Я. Е. Черныше<strong>в</strong><strong>о</strong>й.<br />

Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие — тр<strong>а</strong>хейные и хелицер<strong>о</strong><strong>в</strong>ые. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф<strong>а</strong>.<br />

Игл<strong>о</strong><strong>к</strong><strong>о</strong>жие, п<strong>о</strong>лух<strong>о</strong>рд<strong>о</strong><strong>в</strong>ые. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Р. Ф. Ге<strong>к</strong><strong>к</strong>ер<strong>а</strong>.<br />

Бе<strong>с</strong>челю<strong>с</strong>тные, рыбы. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Д. В. Обруче<strong>в</strong><strong>а</strong>.<br />

Земн<strong>о</strong><strong>в</strong><strong>о</strong>дные, пре<strong>с</strong>мы<strong>к</strong><strong>а</strong>ющие<strong>с</strong>я, птицы. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией А. К. Р<strong>о</strong>жде<strong>с</strong>т<strong>в</strong>ен<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>.<br />

Мле<strong>к</strong><strong>о</strong>пит<strong>а</strong>ющие. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией В. И. Гр<strong>о</strong>м<strong>о</strong><strong>в</strong><strong>о</strong>й.<br />

В<strong>о</strong>д<strong>о</strong>р<strong>о</strong><strong>с</strong>ли, мхи, п<strong>с</strong>ил<strong>о</strong>фиты, пл<strong>а</strong>ун<strong>о</strong><strong>в</strong>ые, члени<strong>с</strong>т<strong>о</strong><strong>с</strong>тебельные, п<strong>а</strong>п<strong>о</strong>р<strong>о</strong>тни<strong>к</strong>и. П<strong>о</strong>д<br />

ред<strong>а</strong><strong>к</strong>цией В. А. В<strong>а</strong>хр<strong>а</strong>мее<strong>в</strong><strong>а</strong>, Г. П. Р<strong>а</strong>дчен<strong>к</strong><strong>о</strong>, А. Л. Т<strong>а</strong>хт<strong>а</strong>дж<strong>а</strong>н<strong>а</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>с</strong>еменные, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong><strong>с</strong>еменные. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией В. А. В<strong>а</strong>хр<strong>а</strong>мее<strong>в</strong><strong>а</strong>, Г. П. Р<strong>а</strong>дчен<strong>к</strong><strong>о</strong>,<br />

А. Л. Т<strong>а</strong>хт<strong>а</strong>дж<strong>а</strong>н<strong>а</strong>.<br />

http://jurassic.ru/

ОГЛАВЛЕНИЕ<br />

Стр.<br />

Гл<strong>а</strong><strong>в</strong>нейшие <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>ие п<strong>о</strong>др<strong>а</strong>зделения, принятые <strong>в</strong> изд<strong>а</strong>нии «О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии» 13<br />

Преди<strong>с</strong>л<strong>о</strong><strong>в</strong>ие 15<br />

П<strong>о</strong>дтип Mandibulata. Ж<strong>в</strong><strong>а</strong>л<strong>о</strong>н<strong>о</strong><strong>с</strong>ные члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 17<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Tracheata. Тр<strong>а</strong>хейные 20<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Symphyla. Симфилы (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 21<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Pauropoda. П<strong>а</strong>ур<strong>о</strong>п<strong>о</strong>ды (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 21<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Diplopoda. Д<strong>в</strong>уп<strong>а</strong>рн<strong>о</strong>н<strong>о</strong>гие (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 22<br />

Н<strong>а</strong>д<strong>о</strong>тряд Pselaphognatha 22<br />

Н<strong>а</strong>д<strong>о</strong>тряд Chilognatha 23<br />

Отряд Palaeocoxopleura 23<br />

Отряд Eurysterna 23<br />

Отряд Palaeomorpha 23<br />

Отряд Oniscomorpha 24<br />

Отряд Polydesmodea 24<br />

Отряд Nematophora 24<br />

Отряд Juliformia 24<br />

Отряд Colobognatha 24<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Chilopoda. Губ<strong>о</strong>н<strong>о</strong>гие (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 25<br />

Отряд Epimorpha 25<br />

П<strong>о</strong>д<strong>о</strong>тряд Geophilomorpha 25<br />

П<strong>о</strong>д<strong>о</strong>тряд Scolopendromorpha. С<strong>к</strong><strong>о</strong>л<strong>о</strong>пендры 25<br />

Отряд Anamorpha 26<br />

П<strong>о</strong>д<strong>о</strong>тряд Lithobiomorpha. К<strong>о</strong><strong>с</strong>тян<strong>к</strong>и 26<br />

П<strong>о</strong>д<strong>о</strong>тряд Scutigeromorpha. Мух<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>к</strong>и 27<br />

Литер<strong>а</strong>тур<strong>а</strong>.... 28<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Insecta. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые 29<br />

Общ<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 29<br />

Си<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 45<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Apterygota. Пер<strong>в</strong>ичн<strong>о</strong>бе<strong>с</strong><strong>к</strong>рылые (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 45<br />

Отряд Collembola. Н<strong>о</strong>г<strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>к</strong>и 45<br />

П<strong>о</strong>д<strong>о</strong>тряд Arthropleona. Члени<strong>с</strong>т<strong>о</strong>брюхие 45<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Poduromorpha 46<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protentomobryomorpha 46<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Entomobryomorpha 46<br />

П<strong>о</strong>д<strong>о</strong>тряд Symphypleona. Цельн<strong>о</strong>брюхие 47<br />

Отряд Diplura. Д<strong>в</strong>ух<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>к</strong>и 47<br />

Отряд Thysanura. Щетин<strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>к</strong>и 48<br />

Отряд Monura. Одн<strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>к</strong>и 48<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Pterygota. Крыл<strong>а</strong>тые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 49<br />

Инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Palaeoptera. Дре<strong>в</strong>не<strong>к</strong>рылые 49<br />

Н<strong>а</strong>д<strong>о</strong>тряд Palaeodictyopteroidea 50<br />

Отряд Palaeodictyoptera. П<strong>а</strong>ле<strong>о</strong>ди<strong>к</strong>ти<strong>о</strong>птеры 50<br />

П<strong>о</strong>д<strong>о</strong>тряд Eupalaeodictyoptera 50<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Ljthomanteidea 50<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dictyoneuridea 50<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Breyeriidea 50<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Homoiopteridea 52<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Polycreagridea 52<br />

П<strong>о</strong>д<strong>о</strong>тряд Parapalaeodictyoptera 52<br />

П<strong>о</strong>д<strong>о</strong>тряд Metapalaeodictyoptera 53<br />

Отряд Eubleptodea 54<br />

Отряд Archodonata 55<br />

Отряд Ephemeroptera. П<strong>о</strong>ден<strong>к</strong>и (О. А. Черн<strong>о</strong><strong>в</strong><strong>а</strong>) 55<br />

http://jurassic.ru/<br />

6

П<strong>о</strong>д<strong>о</strong>тряд Protephemeroptera , 55<br />

П<strong>о</strong>д<strong>о</strong>тряд Plectoptera 56<br />

Инфр<strong>а</strong><strong>о</strong>тряд Permoplectoptera 56<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protereismatidea 56<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesephemeridea 56<br />

Инфр<strong>а</strong><strong>о</strong>тряд Euplectoptera 57<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Paedephemeridea 58<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Ephemeridea 59<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Siphlonuridea ' 60<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Heptageniidea 64<br />

Н<strong>а</strong>д<strong>о</strong>тряд Megasecopteroidea (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 64<br />

Отряд Megasecoptera 64<br />

П<strong>о</strong>д<strong>о</strong>тряд Eumegasecoptera 64<br />

П<strong>о</strong>д<strong>о</strong>тряд Protohymenoptera . : 66<br />

Отряд Diaphanopterodea 69<br />

П<strong>о</strong>д<strong>о</strong>тряд Protodiaphanopterodea 69<br />

П<strong>о</strong>д<strong>о</strong>тряд Eudiaphanopterodea 69<br />

П<strong>о</strong>д<strong>о</strong>тряд Metadiaphanopterodea 71<br />

Отряд Campylopterodea 72<br />

Н<strong>а</strong>д<strong>о</strong>тряд Odonatoidea. Стре<strong>к</strong><strong>о</strong>з<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные 73<br />

Отряд Odonata. Стре<strong>к</strong><strong>о</strong>зы 73<br />

П<strong>о</strong>д<strong>о</strong>тряд Eomeganisoptera 73<br />

П<strong>о</strong>д<strong>о</strong>тряд Meganisoptera. Гиг<strong>а</strong>нт<strong>с</strong><strong>к</strong>ие <strong>с</strong>тре<strong>к</strong><strong>о</strong>зы 73<br />

П<strong>о</strong>д<strong>о</strong>тряд Protanisoptera. Пр<strong>о</strong>т<strong>а</strong>низ<strong>о</strong>птеры 74<br />

П<strong>о</strong>д<strong>о</strong>тряд Zygoptera. Р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые <strong>с</strong>тре<strong>к</strong><strong>о</strong>зы 76<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Kennedyidea 76<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Permagrionidea 77<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Permepallagidea 78<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protomyrmeleontidea . . 78<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hemiphlebiidea 78<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Coenagrionidea . . , 79<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Agrionidea ..... 80<br />

П<strong>о</strong>д<strong>о</strong>тряд Anisozygoptera . . . • 81<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Heterophlebiidea 81<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tarsophlebiidea 81<br />

П<strong>о</strong>д<strong>о</strong>тряд Anisoptera. Нер<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые <strong>с</strong>тре<strong>к</strong><strong>о</strong>зы 83<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Aeshnidea 83<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Libellulidea 85<br />

Инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Neoptera. Н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылые (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 85<br />

К<strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Archaeoneoptera . 86<br />

Отряд Archaeoptera • . . . 86<br />

К<strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Pclyneoptera. Мн<strong>о</strong>г<strong>о</strong>жил<strong>к</strong><strong>о</strong><strong>в</strong>ые 88<br />

Н<strong>а</strong>д<strong>о</strong>тряд Blattopteroidea. Т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные 88<br />

Отряд Blattodea. Т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ые (Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>) . . . . 88<br />

Отряд Manteodea. Б<strong>о</strong>г<strong>о</strong>м<strong>о</strong>лы (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 111<br />

Отряд Isoptera. Термиты (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 112<br />

Отряд Dermaptera. К<strong>о</strong>жи<strong>с</strong>т<strong>о</strong><strong>к</strong>рылые (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 113<br />

П<strong>о</strong>д<strong>о</strong>тряд Archidermaptera 114<br />

П<strong>о</strong>д<strong>о</strong>тряд Eudermaptera 114<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protodermaptera 114<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Forficulidea 115<br />

Отряд Protelytroptera (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 115<br />

Отряд Protoblattodea (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 116<br />

Н<strong>а</strong>д<strong>о</strong>тряд Plecopteroidea. Ве<strong>с</strong>нян<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) . . . 118<br />

Отряд Paraplecoptera (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 119<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Ideliidea 119<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cacurgidea 122<br />

http://jurassic.ru/<br />

7

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Geraridea 122<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Strephocladidea 124<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridea 124<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Megakhosaridea 132<br />

Отряд Plecoptera. Ве<strong>с</strong>нян<strong>к</strong>и (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 134<br />

Отряд Embioptera. Эмбии (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 139<br />

П<strong>о</strong>д<strong>о</strong>тряд Sheimioidea 139<br />

П<strong>о</strong>д<strong>о</strong>тряд Euembiaria 139<br />

Отряд Miomoptera (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 140<br />

Отряд Caloneurodea (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 142<br />

Н<strong>а</strong>д<strong>о</strong>тряд Orthopteroidea. Прям<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) . . 144<br />

Отряд Protorthoptera (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 145<br />

Отряд Orthoptera. Прям<strong>о</strong><strong>к</strong>рылые (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 146<br />

П<strong>о</strong>д<strong>о</strong>тряд Ensifera 146<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Oedischiidea 146<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Gryllidea 150<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tettigoniidea. Кузнечи<strong>к</strong><strong>о</strong><strong>в</strong>ые 154<br />

П<strong>о</strong>д<strong>о</strong>тряд Caelifera 155<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Locustopseidea 155<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Acrididea. С<strong>а</strong>р<strong>а</strong>нч<strong>о</strong><strong>в</strong>ые 156<br />

Отряд Glosselytrodea (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 157<br />

Отряд Phasmatodea. П<strong>а</strong>л<strong>о</strong>чни<strong>к</strong>и (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 159<br />

П<strong>о</strong>д<strong>о</strong>тряд Chresmododea 160<br />

П<strong>о</strong>д<strong>о</strong>тряд Euphasmatodea 160<br />

К<strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Paraneoptera. П<strong>а</strong>р<strong>а</strong>не<strong>о</strong>птеры (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 160<br />

Н<strong>а</strong>д<strong>о</strong>тряд Rhynchota. Х<strong>о</strong>б<strong>о</strong>тные (Е. Э: Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>) 161<br />

Отряд Homoptera. Р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые (Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>) . . . . 162<br />

П<strong>о</strong>д<strong>о</strong>тряд Auchenorrhyncha 163<br />

Инфр<strong>а</strong><strong>о</strong>тряд Blattoprosbolomorpha 163<br />

Инфр<strong>а</strong><strong>о</strong>тряд Cicadomorpha 164<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Palaeontinidea 164<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cicadidea 167<br />

Инфр<strong>а</strong><strong>о</strong>тряд Cicadellomorpha 172<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cicadellidea 173<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cercopidea 179<br />

Инфр<strong>а</strong><strong>о</strong>тряд Fulgoromorpha 181<br />

П<strong>о</strong>д<strong>о</strong>тряд Sternorrhyncha 190<br />

Инфр<strong>а</strong><strong>о</strong>тряд Archescytinomorpha 190<br />

Инфр<strong>а</strong><strong>о</strong>тряд Aphidomorpha (Е. Е. Айзенберг, Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

199<br />

Инфр<strong>а</strong><strong>о</strong>тряд Coccomorpha (Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>) . . . . 199<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protopsyllidiidea 200<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Coecidea . . 200<br />

Инфр<strong>а</strong><strong>о</strong>тряд Psyllomorpha 203<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Aleurodidea 203<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cicadopsyllidea 204<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Psyllidea 206<br />

П<strong>о</strong>д<strong>о</strong>тряд Coleorrhyncha 207<br />

Отряд Heteroptera. Р<strong>а</strong>зн<strong>о</strong><strong>к</strong>рылые, или <strong>к</strong>л<strong>о</strong>пы (Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

208<br />

Н<strong>а</strong>д<strong>о</strong>тряд Psocopteroidea. Сен<strong>о</strong>ед<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) . . . 226<br />

Отряд Psocoptera. Сен<strong>о</strong>еды (Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, В. Н. Вишня<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

226<br />

П<strong>о</strong>д<strong>о</strong>тряд Permopsocida 227<br />

П<strong>о</strong>д<strong>о</strong>тряд Parapsocida 228<br />

П<strong>о</strong>д<strong>о</strong>тряд Eupsocida 234<br />

Отряд Perielytrodea (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 23<br />

8<br />

http://jurassic.ru/

Н<strong>а</strong>д<strong>о</strong>тряд Thysanopteroidea. Трип<strong>с</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) . . 236<br />

Отряд Thysanoptera. Пузырен<strong>о</strong>гие или трип<strong>с</strong>ы (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 236<br />

П<strong>о</strong>д<strong>о</strong>тряд Terebrantia. Яйце<strong>к</strong>л<strong>а</strong>дные трип<strong>с</strong>ы 237<br />

П<strong>о</strong>д<strong>о</strong>тряд Tubulifera. Труб<strong>к</strong><strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>тые трип<strong>с</strong>ы<br />

К<strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Holometabola (Oligoneoptera). Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением<br />

237<br />

или м<strong>а</strong>л<strong>о</strong>жил<strong>к</strong><strong>о</strong><strong>в</strong>ые н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылые (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 239<br />

Н<strong>а</strong>д<strong>о</strong>тряд Coleopteroidea. Же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 240<br />

Отряд Coleoptera. Же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые, или жу<strong>к</strong>и (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф,<br />

А. Г. П<strong>о</strong>н<strong>о</strong>м<strong>а</strong>рен<strong>к</strong><strong>о</strong>) 241<br />

Отряд Strepsiptera. Веер<strong>о</strong><strong>к</strong>рылые (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 267<br />

Н<strong>а</strong>д<strong>о</strong>тряд Neuropteroidea. Сетч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 269<br />

Отряд Megaloptera. Ви<strong>с</strong>л<strong>о</strong><strong>к</strong>рылые (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 269<br />

Отряд Raphidioptera. Верблюд<strong>к</strong>и (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 270<br />

Отряд Neuroptera. Сетч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 272<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Myrmeleontidea<br />

1<br />

273<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Polystoechotidea 274<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hemerobiidea 277<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Coniopterygidea 282<br />

Н<strong>а</strong>д<strong>о</strong>тряд Mecopteroidea (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 283<br />

Отряд Mecoptera. С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>нницы (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 283<br />

П<strong>о</strong>д<strong>о</strong>тряд Protomecoptera 284<br />

П<strong>о</strong>д<strong>о</strong>тряд Eumecoptera 286<br />

П<strong>о</strong>д<strong>о</strong>тряд Paratrichoptera 292<br />

Отряд Trichoptera. Ручейни<strong>к</strong>и (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 294<br />

П<strong>о</strong>д<strong>о</strong>тряд Permotrichoptera. Дре<strong>в</strong>нещупи<strong>к</strong><strong>о</strong><strong>в</strong>ые 295<br />

П<strong>о</strong>д<strong>о</strong>тряд Annulipalpia. К<strong>о</strong>льч<strong>а</strong>т<strong>о</strong>щупи<strong>к</strong><strong>о</strong><strong>в</strong>ые 296<br />

П<strong>о</strong>д<strong>о</strong>тряд Integripalipia. Цельн<strong>о</strong>щупи<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

Отряд Lepidoptera. Б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong>и или чешуе<strong>к</strong>рылые (А. С. Д<strong>а</strong>ниле<strong>в</strong><strong>с</strong><strong>к</strong>ий<br />

299<br />

и О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) 303<br />

П<strong>о</strong>д<strong>о</strong>тряд Jugata. Низшие р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong>и 304<br />

П<strong>о</strong>д<strong>о</strong>тряд Frenata. Р<strong>а</strong>зи<strong>о</strong><strong>к</strong>рылые б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong>и 304<br />

Инфр<strong>а</strong><strong>о</strong>тряд Microfrenata 304<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tineidea. М<strong>о</strong>ле<strong>о</strong>бр<strong>а</strong>зные 304<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tortricidea. Ли<strong>с</strong>т<strong>о</strong><strong>в</strong>ерт<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные . . . . 307<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Gelechiidea 307<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pyralididea. Огне<strong>в</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные 307<br />

Инфр<strong>а</strong><strong>о</strong>тряд Macrofrenata 307<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Noctuidea. С<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные 307<br />

Отряд Diptera. Д<strong>в</strong>у<strong>к</strong>рылые (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф) 307<br />

П<strong>о</strong>д<strong>о</strong>тряд Archidiptera * 308<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dictyodipteridea 308<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hyperpolyneuridea 309<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dyspolyneuridea 310<br />

П<strong>о</strong>д<strong>о</strong>тряд Eudiptera 310<br />

Инфр<strong>а</strong><strong>о</strong>тряд Diplopolyneuromorpha 310<br />

Инфр<strong>а</strong><strong>о</strong>тряд Tipulomorpha 310<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tipulodictyidea 310<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eopolyneuridea 311<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tipulidea 312<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eoptychopteridea 314<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tanyderophryneidea 314<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dixidea 314<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Culicidea 314<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Psychodidea 314<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Chironomidea 316<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhaetomyiidea 318<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesophantasmatidea 318<br />

9<br />

http://jurassic.ru/

Инфр<strong>а</strong><strong>о</strong>тряд Bibionomorpha 319<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pleciodictyidea 319<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protoligoneuridea .' 319<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Fungivoridea 319<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bibionidea 328<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bolitophilidea 330<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Scatopsidea 330<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhyphidea 331<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phragmoligoneridea 332<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cecidomyiidea 332<br />

Инфр<strong>а</strong><strong>о</strong>тряд Asilomorpha 334<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Stratiomyiidea 334<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tabanidea 336<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Asilidea 337<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bombyliidea 339<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Empididea 339<br />

Инфр<strong>а</strong><strong>о</strong>тряд Phoromorpha 339<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phoridea 339<br />

Инфр<strong>а</strong><strong>о</strong>тряд Musidoromorpha 340<br />

Инфр<strong>а</strong><strong>о</strong>тряд Myiomropha 340<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Platypezidea 341<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Syrphidea 341<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Conopidea 342<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Muscidea 343<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hippoboscidea 344<br />

Отряд Aphaniptera. Бл<strong>о</strong>хи (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 345<br />

Н<strong>а</strong>д<strong>о</strong>тряд Hymenopteroidea. Переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

345<br />

Отряд Hymenoptera. Переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые (О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>) . . . . 345<br />

П<strong>о</strong>д<strong>о</strong>тряд Symphyta. Сидячебрюхие переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые 346<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Megalodontidea 346<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Siricidea 347<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Orussidea 348<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cephidea 348<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tenthredinidea 348<br />

П<strong>о</strong>д<strong>о</strong>тряд Apocrita 349<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Ichneumonidea 349<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cynipidea 352<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Chalcididea. Х<strong>а</strong>льциды 352<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Proctotrupidea 353<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bethylidea 354<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cretavidea 356<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Scoliidea 356<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Formicidea. Мур<strong>а</strong><strong>в</strong>ьи 357<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pompilidea. Д<strong>о</strong>р<strong>о</strong>жные <strong>о</strong><strong>с</strong>ы 357<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Apidea. Пчелиные 359<br />

Литер<strong>а</strong>тур<strong>а</strong> 360<br />

П<strong>о</strong>дтип Chelicerophora. Хелицер<strong>о</strong>н<strong>о</strong><strong>с</strong>ные члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие (|В. Б. Дубинин)) 375<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Chelicerata. Хелицер<strong>о</strong><strong>в</strong>ые 378<br />

Общ<strong>а</strong>я х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong><strong>а</strong> хелицер<strong>о</strong><strong>в</strong>ых 378<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Merostomata. Мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты (Не<strong>с</strong>т<strong>о</strong>р Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>) 386<br />

Отряд Aglaspidida 390<br />

Отряд Chasmataspidida 394<br />

Отряд Synziphosura 394<br />

Отряд Limulida. Н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты 397<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Belinuridea 398<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomesaspididea 399<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Limulidea 400<br />

10<br />

http://jurassic.ru/

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Scorpionomorpha. С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>м<strong>о</strong>рфы (| В. Б. Дубинин |) 402<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Holactinochitinosi. С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные 403<br />

Отряд Eurypterida (Не<strong>с</strong>т<strong>о</strong>р Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>) 404<br />

П<strong>о</strong>д<strong>о</strong>тряд Eurypterina 408<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridea • • • 408<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pterygotidea 409<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mixopteridea 412<br />

П<strong>о</strong>д<strong>о</strong>тряд Stylonurina 415<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Stylonuridea 416<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, Mycteropidea 420<br />

Отряд Scorpionida. С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны ( В. Б. Дубинин ) 423<br />

П<strong>о</strong>д<strong>о</strong>тряд Apoxypodes. З<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>н<strong>о</strong>гие <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны 425<br />

П<strong>о</strong>д<strong>о</strong>тряд Dionychopodes. Д<strong>в</strong>у<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны . . . . 426<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mazoniidea 428<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Euscorpiidea. Н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны . . . . 428<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Pedipalpides. Жгут<strong>о</strong>н<strong>о</strong>гие (| В. Б. Дубинин ) - 433<br />

Отряд Uropygi. Жгут<strong>о</strong>н<strong>о</strong>гие <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны 434<br />

П<strong>о</strong>д<strong>о</strong>тряд Schizopeltidia 436<br />

П<strong>о</strong>д<strong>о</strong>тряд Holopeltidia 436<br />

Отряд Amblypygi. Жгут<strong>о</strong>н<strong>о</strong>гие п<strong>а</strong>у<strong>к</strong>и 436<br />

Отряд Kustarachnida 437<br />

Отряд Palpigradi. П<strong>а</strong>льпигр<strong>а</strong>ды 437<br />

Отряд Pseudoscorpionodea. Л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны 439<br />

П<strong>о</strong>д<strong>о</strong>тряд Chthoniinea 441<br />

П<strong>о</strong>д<strong>о</strong>тряд Neobisinea 441<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Neobisidea 441<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Garypidea 441<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Feaellidea 441<br />

П<strong>о</strong>д<strong>о</strong>тряд Cheliferinea 442<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cheliferidea 442<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cheiridiidea 443<br />

Отряд Ricinulei 445<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha. С<strong>о</strong>льпуг<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бные (|В. Б. Дубинин|) 446<br />

Отряд Solifugae. Ф<strong>а</strong>л<strong>а</strong>нги, или <strong>с</strong><strong>о</strong>льпуги 447<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Acaromorpha. Клещи (|В. Б. Дубинин |) 450<br />

Отряд AcarHormes. Н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие <strong>к</strong>лещи 451<br />

П<strong>о</strong>д<strong>о</strong>тряд Acaridiae. С<strong>а</strong>р<strong>к</strong><strong>о</strong>пт<strong>о</strong>идные 452<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Acaridea. Хлебные (<strong>а</strong>мб<strong>а</strong>рные) <strong>к</strong>лещи . . . 452<br />

П<strong>о</strong>д<strong>о</strong>тряд Oribatei. П<strong>а</strong>нцирные <strong>к</strong>лещи 454<br />

Инфр<strong>а</strong><strong>о</strong>тряд Aptyctima 454<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hypochthoniidea 454<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Camisiidea 454<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hermannidea 454<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Damaeidea 456<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Carabodidea 456<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hermanniellidea 457<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Notospididea 457<br />

Инфр<strong>а</strong><strong>о</strong>тряд Ptyetima 461<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phthiracaridea 461<br />

П<strong>о</strong>д<strong>о</strong>тряд Trombidiformes. Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>тел<strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>к</strong>лещи 461<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pachygnathidea 462<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhaphignathidea 465<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tydeidea • 465<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Anystidea 466<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bdellidea 466<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cheyletidea. Хищные <strong>к</strong>лещи 467<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Smaridiidea 468<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Trombidiidea. Клещи-<strong>к</strong>р<strong>а</strong><strong>с</strong>н<strong>о</strong>тел<strong>к</strong>и . . . . 468<br />

11<br />

http://jurassic.ru/

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tetranychidea. П<strong>а</strong>утинные, или тетр<strong>а</strong>них<strong>о</strong><strong>в</strong>ые<br />

<strong>к</strong>лещи 468<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Limnocharidea. Лимн<strong>о</strong>х<strong>а</strong>риды 469<br />

П<strong>о</strong>д<strong>о</strong>тряд Tetrapodili. Четырехн<strong>о</strong>гие <strong>к</strong>лещи 471<br />

Отряд Parasitiformes. Г<strong>а</strong>м<strong>а</strong>з<strong>о</strong>идные <strong>к</strong>лещи 471<br />

П<strong>о</strong>д<strong>о</strong>тряд Mesostigmata. Г<strong>а</strong>м<strong>а</strong>з<strong>о</strong><strong>в</strong>ые <strong>к</strong>лещи 472<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Gamasidea 472<br />

П<strong>о</strong>д<strong>о</strong>тряд Ixodides. И<strong>к</strong><strong>с</strong><strong>о</strong>д<strong>о</strong><strong>в</strong>ые <strong>к</strong>лещи 473<br />

Отряд Opilioacarina. Клещи <strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цы 473<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida. П<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные ( В. Б. Дубинин|) 474<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Opiliomorphae. Сен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цеп<strong>о</strong>д<strong>о</strong>бные, или л<strong>о</strong>жн<strong>о</strong>п<strong>а</strong>у<strong>к</strong>и 476<br />

Отряд Haptopoda 478<br />

Отряд Phalangiotarbi 477<br />

Отряд Opiliones. П<strong>а</strong>у<strong>к</strong>и-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цы 480<br />

П<strong>о</strong>д<strong>о</strong>тряд Cyphophthalmi 482<br />

П<strong>о</strong>д<strong>о</strong>тряд Laniatores 483<br />

П<strong>о</strong>д<strong>о</strong>тряд Palpatores 483<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Trogulidea 483<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phalangiidea 483<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Soluta. П<strong>а</strong>нцирные п<strong>а</strong>у<strong>к</strong>и 485<br />

Отряд Trigonotarbi 486<br />

Отряд Anthracomarti 490<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Аг<strong>а</strong>пе<strong>а</strong>е. П<strong>а</strong>у<strong>к</strong>и 492<br />

Отряд Liphistiomorphae. Члени<strong>с</strong>т<strong>о</strong>брюхие п<strong>а</strong>у<strong>к</strong>и 494<br />

Отряд Arachnomorphae 497<br />

Отряд Mygalomorphae. П<strong>а</strong>у<strong>к</strong>и-птицеяды 497<br />

П<strong>о</strong>д<strong>о</strong>тряд Nelipoda 497<br />

П<strong>о</strong>д<strong>о</strong>тряд Hypodemata 498<br />

Отряд Araneomorphae. Н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие п<strong>а</strong>у<strong>к</strong>и 498<br />

П<strong>о</strong>д<strong>о</strong>тряд Ecribellatae 498<br />

Инфр<strong>а</strong><strong>о</strong>тряд Haplogynae 498<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dysderidea 498<br />

Инфр<strong>а</strong><strong>о</strong>тряд Entelogynae 499<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pholcidea 499<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Zodariidea 500<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hersiliidea 501<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Araneidea 501<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Lycosidea 505<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Drassodidea 507<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Clubionidea 507<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Thomisidea 509<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Salticidea 509<br />

П<strong>о</strong>д<strong>о</strong>тряд Cribellatae 515<br />

Инфр<strong>а</strong><strong>о</strong>тряд Palaeocribellatae 515<br />

Инфр<strong>а</strong><strong>о</strong>тряд Neocribellatae 515<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Zoropseidea 515<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eresidea 515<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dictynidea 515<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Amaurobiidea 517<br />

Литер<strong>а</strong>тур<strong>а</strong> 518<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Pycnogonides. М<strong>о</strong>р<strong>с</strong><strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и ( В. Б. Дубинин ) 525<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Protopantopoda. Пр<strong>о</strong>т<strong>о</strong>п<strong>а</strong>нт<strong>о</strong>п<strong>о</strong>ды, или члени<strong>с</strong>т<strong>о</strong>брюхие м<strong>о</strong>р<strong>с</strong><strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и . . 527<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Pantopoda • • 529<br />

Отряд Paleopantopoda 529<br />

Отряд Eupantopoda 529<br />

Литер<strong>а</strong>тур<strong>а</strong> 530<br />

Тип Malacopoda. Мяг<strong>к</strong><strong>о</strong>н<strong>о</strong>гие, или пер<strong>в</strong>ичн<strong>о</strong>тр<strong>а</strong>хейные (А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>) 531<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Onychophora 533<br />

Отряд Protonychophora 533<br />

Литер<strong>а</strong>тур<strong>а</strong> 535<br />

Т<strong>а</strong>блицы (I — XXII) и <strong>о</strong>бъя<strong>с</strong>нения <strong>к</strong> ним . . - 537—539<br />

Алф<strong>а</strong><strong>в</strong>итный у<strong>к</strong><strong>а</strong>з<strong>а</strong>тель 539<br />

http://jurassic.ru/

ГЛАВНЕЙШИЕ СТРАТИГРАФИЧЕСКИЕ ПОДРАЗДЕЛЕНИЯ,<br />

ПРИНЯТЫЕ В ИЗДАНИИ «ОСНОВЫ ПАЛЕОНТОЛОГИИ»<br />

Схем<strong>а</strong> ут<strong>в</strong>ержден<strong>а</strong> для «О<strong>с</strong>н<strong>о</strong><strong>в</strong> п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии» Меж<strong>в</strong>ед<strong>о</strong>м<strong>с</strong>т<strong>в</strong>енным <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>им<br />

<strong>к</strong><strong>о</strong>митет<strong>о</strong>м СССР 30 июня 1955 г.<br />

Группы Си<strong>с</strong>темы Отделы Яру<strong>с</strong>ы (и др. п<strong>о</strong>др<strong>а</strong>зд.)<br />

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Группы Си<strong>с</strong>темы Отделы Яру<strong>с</strong>ы (и др. п<strong>о</strong>др<strong>а</strong>зд.)<br />

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Верхний Т<strong>о</strong><strong>а</strong>р<strong>с</strong><strong>к</strong>ий<br />

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С<strong>а</strong><strong>к</strong>м<strong>а</strong>р<strong>с</strong><strong>к</strong>ий<br />

Жигуле<strong>в</strong><strong>с</strong><strong>к</strong>ий<br />

О<strong>к</strong><strong>о</strong>нч<strong>а</strong>ние<br />

Д<strong>о</strong>мер<strong>с</strong><strong>к</strong>ий<br />

Плин<strong>с</strong>б<strong>а</strong>х<strong>с</strong><strong>к</strong>ий<br />

Л<strong>о</strong>т<strong>а</strong>ринг<strong>с</strong><strong>к</strong>ий<br />

Синемюр<strong>с</strong><strong>к</strong>ий<br />

Гетт<strong>а</strong>нг<strong>с</strong><strong>к</strong>ий<br />

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Н<strong>о</strong>рий<strong>с</strong><strong>к</strong>ий<br />

К<strong>а</strong>рний<strong>с</strong><strong>к</strong>ий<br />

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К<strong>а</strong>мпиль<strong>с</strong><strong>к</strong>ий 1<br />

Сей<strong>с</strong><strong>с</strong><strong>к</strong>ий 1<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong>ий<br />

К<strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

Кунгур<strong>с</strong><strong>к</strong>ий<br />

Артин<strong>с</strong><strong>к</strong>ий<br />

С<strong>а</strong><strong>к</strong>м<strong>а</strong>р<strong>с</strong><strong>к</strong>ий<br />

А<strong>с</strong><strong>с</strong>ель<strong>с</strong><strong>к</strong>ий<br />

Оренбург<strong>с</strong><strong>к</strong>ий<br />

Гжель<strong>с</strong><strong>к</strong>ий<br />

К<strong>а</strong><strong>с</strong>им<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий<br />

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М<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий<br />

Б<strong>а</strong>ш<strong>к</strong>ир<strong>с</strong><strong>к</strong>ий, или <strong>к</strong><strong>а</strong>яль<strong>с</strong><strong>к</strong>ий<br />

м<br />

Н<strong>а</strong>мюр<strong>с</strong><strong>к</strong>ий<br />

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Визей<strong>с</strong><strong>к</strong>ий<br />

Турней<strong>с</strong><strong>к</strong>ий<br />

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Ф<strong>а</strong>мен<strong>с</strong><strong>к</strong>ий<br />

Фр<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

Средний<br />

Жи<strong>в</strong>ет<strong>с</strong><strong>к</strong>ий<br />

Эйфель<strong>с</strong><strong>к</strong>ий<br />

Нижний<br />

К<strong>о</strong>бленц<strong>с</strong><strong>к</strong>ий<br />

Жедин<strong>с</strong><strong>к</strong>ий<br />

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Венл<strong>о</strong><strong>к</strong><strong>с</strong><strong>к</strong>ий<br />

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Ашгиль<strong>с</strong><strong>к</strong>ий<br />

К<strong>а</strong>р<strong>а</strong>д<strong>о</strong><strong>к</strong><strong>с</strong><strong>к</strong>ий<br />

Лл<strong>а</strong>ндейль<strong>с</strong><strong>к</strong>ий<br />

Лл<strong>а</strong>ндейль<strong>с</strong><strong>к</strong>ий<br />

<strong>о</strong>.<br />

О<br />

Нижний<br />

Арениг<strong>с</strong><strong>к</strong>ий<br />

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Лен<strong>с</strong><strong>к</strong>ий<br />

Алд<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

С<strong>о</strong>гл<strong>а</strong><strong>с</strong>н<strong>о</strong> решению меж<strong>в</strong>ед<strong>о</strong>м<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>митет<strong>а</strong> <strong>о</strong>т 10 м<strong>а</strong>я 1956 г., ре<strong>к</strong><strong>о</strong>мендует<strong>с</strong>я деление<br />

нижнег<strong>о</strong> три<strong>а</strong><strong>с</strong><strong>а</strong> н<strong>а</strong> инд<strong>с</strong><strong>к</strong>ий и <strong>о</strong>лене<strong>к</strong><strong>с</strong><strong>к</strong>ий яру<strong>с</strong>ы.<br />

http://jurassic.ru/

ПРЕДИСЛОВИЕ<br />

Н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящий т<strong>о</strong>м я<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>лжением т<strong>о</strong>м<strong>а</strong> «Трил<strong>о</strong>бит<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные<br />

и р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные» и <strong>с</strong><strong>о</strong>держит <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> тр<strong>а</strong>хейных, хелицер<strong>о</strong><strong>в</strong>ых<br />

и м<strong>о</strong>р<strong>с</strong><strong>к</strong>их п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> (пи<strong>к</strong>н<strong>о</strong>г<strong>о</strong>н<strong>о</strong><strong>в</strong>). Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong> эт<strong>о</strong>м т<strong>о</strong>ме при<strong>в</strong><strong>о</strong>дит<strong>с</strong>я<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong><strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> мяг<strong>к</strong><strong>о</strong>н<strong>о</strong>гих (Mala,copoda),<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии п<strong>о</strong>чти не пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>о</strong>г<strong>о</strong>.<br />

Пер<strong>в</strong>ый, н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бширный р<strong>а</strong>здел, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляющий б<strong>о</strong>лее д<strong>в</strong>ух третей<br />

<strong>о</strong>бъем<strong>а</strong> т<strong>о</strong>м<strong>а</strong>, <strong>в</strong><strong>к</strong>люч<strong>а</strong>ет <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние четырех <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong> и <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых; эт<strong>о</strong>т р<strong>а</strong>здел тр<strong>а</strong>хейных <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен Е. Е. Айзенберг<strong>о</strong>м,<br />

Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й, В. Н. Вишня<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й, А. С. Д<strong>а</strong>ниле<strong>в</strong><strong>с</strong><strong>к</strong>им,<br />

О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й, Н. И. Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>ым, А. Г. П<strong>о</strong>н<strong>о</strong>м<strong>а</strong>рен<strong>к</strong><strong>о</strong>, Ю. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>ым,<br />

Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф<strong>о</strong>м, О. А. Черн<strong>о</strong><strong>в</strong><strong>о</strong>й и А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>ым. Ред<strong>а</strong><strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

пр<strong>о</strong><strong>в</strong>еден<strong>о</strong> Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф<strong>о</strong>м.<br />

Вт<strong>о</strong>р<strong>о</strong>й р<strong>а</strong>здел <strong>в</strong><strong>к</strong>люч<strong>а</strong>ет <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> хелицер<strong>о</strong><strong>в</strong>ых и м<strong>о</strong>р<strong>с</strong><strong>к</strong>их<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>а</strong> третий — тип<strong>а</strong> мяг<strong>к</strong><strong>о</strong>н<strong>о</strong>гих; эти р<strong>а</strong>зделы <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

|В Б. Дубининым|, Н. И. Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>ым и А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>ым. Ред<strong>а</strong><strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>­<br />

ние пр<strong>о</strong><strong>в</strong>еден<strong>о</strong>|<strong>в</strong>. Б. Дубининым! -<br />

, п<strong>о</strong><strong>с</strong>ле ег<strong>о</strong> <strong>к</strong><strong>о</strong>нчины не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые д<strong>о</strong>п<strong>о</strong>лне­<br />

ния <strong>в</strong>не<strong>с</strong> и ред<strong>а</strong><strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>ние пр<strong>о</strong><strong>в</strong>ел Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф.<br />

Общий пл<strong>а</strong>н изл<strong>о</strong>жения и <strong>о</strong>х<strong>в</strong><strong>а</strong>т м<strong>а</strong>тери<strong>а</strong>л<strong>а</strong> <strong>в</strong> цел<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>уют<br />

принят<strong>о</strong>му <strong>в</strong> эт<strong>о</strong>м изд<strong>а</strong>нии. О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>е <strong>о</strong>т<strong>с</strong>тупление <strong>о</strong>т пл<strong>а</strong>н<strong>а</strong> з<strong>а</strong><strong>к</strong>люч<strong>а</strong>л<strong>о</strong><strong>с</strong>ь<br />

<strong>в</strong> б<strong>о</strong>лее <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенн<strong>о</strong>м <strong>о</strong>бз<strong>о</strong>ре <strong>к</strong><strong>а</strong>йн<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уны. Были <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>и лишь <strong>в</strong>ы<strong>с</strong>ших <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их единиц третичных тр<strong>а</strong>хейных<br />

и хелицер<strong>о</strong><strong>в</strong>ых, д<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> и п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong><strong>к</strong>лючительн<strong>о</strong>, р<strong>о</strong>ды<br />

же, <strong>о</strong>бн<strong>а</strong>руженные <strong>в</strong> третичных <strong>о</strong>тл<strong>о</strong>жениях, т<strong>о</strong>ль<strong>к</strong><strong>о</strong> перечи<strong>с</strong>ляли<strong>с</strong>ь (хелицер<strong>о</strong><strong>в</strong>ые)<br />

или у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>л<strong>о</strong><strong>с</strong>ь их <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>, н<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние их не при<strong>в</strong><strong>о</strong>дил<strong>о</strong><strong>с</strong>ь.<br />

Т<strong>а</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щение <strong>о</strong>бъем<strong>а</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>лен<strong>о</strong> <strong>о</strong>чень<br />

б<strong>о</strong>льшим <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных <strong>в</strong>ид<strong>о</strong><strong>в</strong> из третичных <strong>о</strong>тл<strong>о</strong>жений,<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м прин<strong>а</strong>длеж<strong>а</strong>щих р<strong>о</strong>д<strong>а</strong>м, р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненным <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>чень низ<strong>к</strong>ий ур<strong>о</strong><strong>в</strong>ень изученн<strong>о</strong><strong>с</strong>ти третичных <strong>в</strong>ид<strong>о</strong><strong>в</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, не п<strong>о</strong>з<strong>в</strong><strong>о</strong>лил <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ить п<strong>о</strong>лные х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>и<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> п<strong>а</strong>ле<strong>о</strong>ген<strong>о</strong><strong>в</strong>ых и не<strong>о</strong>ген<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и хелицер<strong>о</strong><strong>в</strong>ых.<br />

Ряд <strong>с</strong>пеци<strong>а</strong>ли<strong>с</strong>т<strong>о</strong><strong>в</strong> п<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым <strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ли <strong>а</strong><strong>в</strong>т<strong>о</strong>р<strong>а</strong>м<br />

<strong>к</strong><strong>о</strong>н<strong>с</strong>ульт<strong>а</strong>ти<strong>в</strong>ную п<strong>о</strong>м<strong>о</strong>щь п<strong>о</strong> <strong>с</strong>и<strong>с</strong>теме <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных групп. В ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти,<br />

т<strong>а</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> <strong>к</strong><strong>о</strong>н<strong>с</strong>ульт<strong>а</strong>ции были п<strong>о</strong>лучены <strong>о</strong>т Г. Я. Бей-Биен<strong>к</strong><strong>о</strong> п<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременным прям<strong>о</strong><strong>к</strong>рылым, <strong>о</strong>т А. Н. Киричен<strong>к</strong><strong>о</strong> п<strong>о</strong> п<strong>о</strong>луже<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылым,<br />

15<br />

http://jurassic.ru/

<strong>о</strong>т С. Г. Лепне<strong>в</strong><strong>о</strong>й п<strong>о</strong> ручейни<strong>к</strong><strong>а</strong>м, <strong>о</strong>т М. Н. Ни<strong>к</strong><strong>о</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>й, В. В. П<strong>о</strong>п<strong>о</strong><strong>в</strong><strong>а</strong><br />

и А. П. Р<strong>а</strong><strong>с</strong>ницын<strong>а</strong> п<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылым. А<strong>в</strong>т<strong>о</strong>ры <strong>в</strong>ыр<strong>а</strong>ж<strong>а</strong>ют бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>рн<strong>о</strong><strong>с</strong>ть<br />

<strong>в</strong><strong>с</strong>ем перечи<strong>с</strong>ленным лиц<strong>а</strong>м.<br />

В<strong>с</strong>ег<strong>о</strong> <strong>в</strong> т<strong>о</strong>ме при<strong>в</strong><strong>о</strong>дит<strong>с</strong>я <strong>с</strong><strong>в</strong>ыше 1900 <strong>о</strong>пи<strong>с</strong><strong>а</strong>ний <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

единиц, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле 660 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Зн<strong>а</strong>чительн<strong>а</strong> иллю<strong>с</strong>тр<strong>а</strong>ти<strong>в</strong>н<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть; ее з<strong>а</strong>д<strong>а</strong>чей был<strong>о</strong> д<strong>а</strong>ть из<strong>о</strong>бр<strong>а</strong>жения<br />

<strong>в</strong><strong>с</strong>ех <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их единиц, х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>и <strong>к</strong><strong>о</strong>т<strong>о</strong>рых были<br />

п<strong>о</strong>мещены ,<strong>в</strong> <strong>к</strong>ниге; т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>ие<br />

из<strong>о</strong>бр<strong>а</strong>жения. Меньш<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть иллю<strong>с</strong>тр<strong>а</strong>ций — ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых.<br />

Вып<strong>о</strong>лнение иллю<strong>с</strong>тр<strong>а</strong>ций был<strong>о</strong> <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>лен<strong>о</strong> <strong>к</strong><strong>о</strong>лле<strong>к</strong>ти<strong>в</strong><strong>о</strong>м худ<strong>о</strong>жни<strong>к</strong><strong>о</strong><strong>в</strong><br />

К<strong>о</strong>мбин<strong>а</strong>т<strong>а</strong> гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>их р<strong>а</strong>б<strong>о</strong>т (Т. Ф. Бел<strong>о</strong>ц<strong>в</strong>ет<strong>о</strong><strong>в</strong><strong>о</strong>й, И. П. Вели<strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

В. И. Д<strong>о</strong>р<strong>о</strong>фее<strong>в</strong>ым, Н. Л. К<strong>о</strong>рч<strong>а</strong>жин<strong>с</strong><strong>к</strong><strong>о</strong>й, Г. М. Лях<strong>о</strong><strong>в</strong>ым, К. П. Меш<strong>к</strong><strong>о</strong><strong>в</strong>ым,<br />

В. Н. П<strong>о</strong>п<strong>о</strong><strong>в</strong><strong>о</strong>й, А. Н. П<strong>о</strong>т<strong>а</strong>п<strong>о</strong><strong>в</strong><strong>о</strong>й и Т. Л. С<strong>а</strong><strong>в</strong>р<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>й) и ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>ф<strong>о</strong><strong>в</strong><br />

(А. В. С<strong>к</strong>индер<strong>о</strong>м, В. А. Пре<strong>с</strong>ня<strong>к</strong><strong>о</strong><strong>в</strong>ым, Н. П. Фин<strong>о</strong>ген<strong>о</strong><strong>в</strong>ым<br />

и Ю. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>ым). Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые ри<strong>с</strong>ун<strong>к</strong>и <strong>к</strong> <strong>с</strong><strong>в</strong><strong>о</strong>им р<strong>а</strong>здел<strong>а</strong>м были <strong>с</strong>дел<strong>а</strong>ны<br />

1<br />

<strong>с</strong><strong>а</strong>мими <strong>а</strong><strong>в</strong>т<strong>о</strong>р<strong>а</strong>ми — О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й, Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф<strong>о</strong>м,<br />

О. А. Черн<strong>о</strong><strong>в</strong><strong>о</strong>й, Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й и А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>ым.<br />

П<strong>о</strong>дг<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>к</strong><strong>а</strong> т<strong>о</strong>м<strong>а</strong> <strong>к</strong> печ<strong>а</strong>ти и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ление т<strong>а</strong>блиц иллю<strong>с</strong>тр<strong>а</strong>ций пр<strong>о</strong><strong>в</strong>едены<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й, Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

В. А. И<strong>в</strong><strong>а</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>й, Н. Л. Гр<strong>о</strong>мбче<strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й, А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>ым, Н. П. Меш<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

И. Д. Су<strong>к</strong><strong>а</strong>че<strong>в</strong><strong>о</strong>й, В. Н. Вишня<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й и Е. М. С<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й.<br />

http://jurassic.ru/<br />

Ред<strong>а</strong><strong>к</strong>т<strong>о</strong>р т<strong>о</strong>м<strong>а</strong> Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф

Z О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

ПОДТИП MANDIBULATA.<br />

ЖВАЛОНОСНЫЕ<br />

ЧЛЕНИСТОНОГИЕ<br />

http://jurassic.ru/

http://jurassic.ru/

ПОДТИП MANDIBULATA.<br />

ЖВАЛОНОСНЫЕ ЧЛЕНИСТОНОГИЕ<br />

Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующие<strong>с</strong>я н<strong>а</strong>личием<br />

д<strong>в</strong>ух или <strong>о</strong>дн<strong>о</strong>й п<strong>а</strong>ры передних <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

(<strong>а</strong>нтеннул и <strong>а</strong>нтенн), пре<strong>в</strong>р<strong>а</strong>щением<br />

третьей п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>в</strong> ж<strong>в</strong><strong>а</strong>льг (м<strong>а</strong>ндибулы)<br />

и <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нием <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>о</strong>тдел<strong>а</strong>,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> из пер<strong>в</strong>ичн<strong>о</strong>г<strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> (<strong>а</strong><strong>к</strong>р<strong>о</strong>н<strong>а</strong>) и четырех п<strong>о</strong><strong>с</strong>ледующих<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Передний <strong>к</strong><strong>о</strong>нец <strong>о</strong><strong>с</strong>и тел<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<br />

<strong>с</strong><strong>в</strong><strong>о</strong>е пер<strong>в</strong>ичн<strong>о</strong>е прям<strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение и не<br />

з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я <strong>к</strong><strong>в</strong>ерху. Х<strong>о</strong>дильные н<strong>о</strong>ги р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> тул<strong>о</strong><strong>в</strong>ище и лишены гн<strong>а</strong>т<strong>о</strong>б<strong>а</strong>з.<br />

К эт<strong>о</strong>му п<strong>о</strong>дтипу члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я<br />

д<strong>в</strong><strong>а</strong> <strong>о</strong>бширных н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: Crustaceomorpha,<br />

или р<strong>а</strong><strong>к</strong><strong>о</strong>п<strong>о</strong>д<strong>о</strong>бные, и Tracheata, или тр<strong>а</strong>хейные.<br />

Пер<strong>в</strong>ый н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> <strong>о</strong>пи<strong>с</strong>ы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> друг<strong>о</strong>м<br />

т<strong>о</strong>ме «О<strong>с</strong>н<strong>о</strong><strong>в</strong> п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии»; н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящий<br />

т<strong>о</strong>м з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ет <strong>о</strong>бз<strong>о</strong>р тр<strong>а</strong>хейных члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих.<br />

http://jurassic.ru/

НАДКЛАСС TRACHEATA. ТРАХЕЙНЫЕ<br />

Н<strong>а</strong>земные члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие, дыш<strong>а</strong>щие <strong>а</strong>тм<strong>о</strong><strong>с</strong>ферным<br />

<strong>в</strong><strong>о</strong>здух<strong>о</strong>м п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ленн<strong>о</strong>й<br />

<strong>с</strong>и<strong>с</strong>темы хитин<strong>о</strong><strong>в</strong>ых <strong>с</strong><strong>о</strong><strong>с</strong>уд<strong>о</strong><strong>в</strong>: тр<strong>а</strong>хей, <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><br />

н<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong> тел<strong>а</strong>, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я<br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>с</strong>егмент<strong>а</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями—<br />

дых<strong>а</strong>льц<strong>а</strong>ми, или <strong>с</strong>тигм<strong>а</strong>ми, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

п<strong>а</strong>рными и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

пле<strong>в</strong>р, или (ред<strong>к</strong><strong>о</strong>) неп<strong>а</strong>рными и <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я<br />

н<strong>а</strong> тергите. В ред<strong>к</strong>их <strong>с</strong>луч<strong>а</strong>ях тр<strong>а</strong>хеи<br />

нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>иты или з<strong>а</strong>м<strong>к</strong>нуты и не <strong>с</strong><strong>о</strong><strong>о</strong>бщ<strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong> н<strong>а</strong>ружн<strong>о</strong>й <strong>с</strong>ред<strong>о</strong>й (у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых личин<strong>о</strong><strong>к</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

жи<strong>в</strong>ущих <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х). Имеет<strong>с</strong>я <strong>о</strong>дн<strong>а</strong><br />

п<strong>а</strong>р<strong>а</strong> <strong>а</strong>нтенн и три п<strong>а</strong>ры челю<strong>с</strong>тей; <strong>о</strong>чень ред<strong>к</strong><strong>о</strong><br />

<strong>а</strong>нтенны и третья п<strong>а</strong>р<strong>а</strong> челю<strong>с</strong>тей <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

В<strong>с</strong>егд<strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> передняя т<strong>а</strong>гм<strong>а</strong> тел<strong>а</strong> —<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, не<strong>с</strong>ущ<strong>а</strong>я <strong>а</strong>нтенны, челю<strong>с</strong>ти и гл<strong>а</strong>з<strong>а</strong>.<br />

Тел<strong>о</strong> длинн<strong>о</strong>е, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящее из мн<strong>о</strong>гих <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> и <strong>с</strong>редняя т<strong>а</strong>гм<strong>а</strong> тел<strong>а</strong> — грудь —<br />

лишь <strong>в</strong> <strong>с</strong><strong>а</strong>мых и<strong>с</strong><strong>к</strong>лючительных <strong>с</strong>луч<strong>а</strong>ях <strong>о</strong>бъединяют<strong>с</strong>я<br />

<strong>в</strong>ме<strong>с</strong>те (у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из<br />

<strong>о</strong>тряд<strong>а</strong> Heteroptera), <strong>о</strong>бычн<strong>о</strong> же чет<strong>к</strong><strong>о</strong> р<strong>а</strong>згр<strong>а</strong>ничены.<br />

Кишечни<strong>к</strong> <strong>с</strong> немн<strong>о</strong>гими <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми<br />

<strong>с</strong>лепыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми, ди<strong>в</strong>ерти<strong>к</strong>ул<strong>а</strong>ми. Силур<br />

— ныне. Пять <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>: Pauropoda, Diplopoda,<br />

Chilopoda, Symphyla и Insecta.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения <strong>в</strong>нутри н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

тр<strong>а</strong>хейных <strong>о</strong><strong>с</strong><strong>в</strong>ещены еще <strong>о</strong>чень нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>;<br />

н<strong>а</strong>меч<strong>а</strong>ют<strong>с</strong>я я<strong>с</strong>ные <strong>с</strong><strong>в</strong>язи д<strong>в</strong>ух п<strong>о</strong><strong>с</strong>ледних<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> и <strong>в</strong> т<strong>о</strong> же <strong>в</strong>ремя <strong>о</strong>че<strong>в</strong>идн<strong>а</strong><br />

б<strong>о</strong>льш<strong>а</strong>я <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong><strong>с</strong>ть пер<strong>в</strong>ых трех <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>.<br />

От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерных п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong> з<strong>а</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>б<strong>о</strong>бщениях лишь н<strong>а</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>т­<br />

20<br />

http://jurassic.ru/<br />

рении <strong>с</strong>тр<strong>о</strong>ения и <strong>о</strong>нт<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм.<br />

Пер<strong>в</strong>ые четыре <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> тр<strong>а</strong>хейных члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

<strong>о</strong>бычн<strong>о</strong> <strong>о</strong>бъединяют п<strong>о</strong>д <strong>о</strong>бщим п<strong>о</strong>нятием<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong> (Myriopoda). Одн<strong>а</strong><strong>к</strong><strong>о</strong> эти <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы<br />

между <strong>с</strong><strong>о</strong>б<strong>о</strong>й <strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>ют <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> лишь<br />

<strong>в</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ии <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бления з<strong>а</strong>дней т<strong>а</strong>гмы тел<strong>а</strong>,<br />

брюш<strong>к</strong><strong>а</strong>; ре<strong>а</strong>льные р<strong>а</strong>зличия групп мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong><br />

<strong>о</strong>чень <strong>в</strong>ели<strong>к</strong>и, безу<strong>с</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong> <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>уя<br />

<strong>о</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>их <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их р<strong>а</strong>нг<strong>а</strong>х, <strong>а</strong> именн<strong>о</strong><br />

<strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>х.<br />

Пер<strong>в</strong><strong>о</strong>е н<strong>а</strong>учн<strong>о</strong>е <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong> д<strong>а</strong>л С<strong>к</strong>еддер (Scudder,<br />

1882). Кл<strong>а</strong><strong>с</strong><strong>с</strong>иче<strong>с</strong><strong>к</strong><strong>о</strong>е и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ние Фрич<strong>а</strong><br />

(Fritsch, 1889) п<strong>о</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емым мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong>м<br />

перми Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии и <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<br />

<strong>с</strong><strong>в</strong><strong>о</strong>е зн<strong>а</strong>чение, я<strong>в</strong>ляя<strong>с</strong>ь <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong><strong>в</strong><strong>о</strong>д<strong>к</strong><strong>о</strong>й<br />

п<strong>о</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уне. Опи<strong>с</strong><strong>а</strong>ние б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong><br />

чи<strong>с</strong>л<strong>а</strong> третичных мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря д<strong>а</strong>л Б<strong>а</strong>х<strong>о</strong>фен Эхт (Bachofen von Echt,<br />

1942).<br />

* П<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющее б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong> <strong>о</strong>бит<strong>а</strong>ет <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жных би<strong>о</strong>т<strong>о</strong>п<strong>а</strong>х —<br />

п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями, <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, ле<strong>с</strong>н<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е, гниющих<br />

р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>х и пр., '<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

чег<strong>о</strong> <strong>к</strong>р<strong>а</strong>йне ред<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии; не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые из из<strong>в</strong>е<strong>с</strong>тных и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong>, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong>ели <strong>а</strong>мфиби<strong>о</strong>тиче<strong>с</strong><strong>к</strong>ий<br />

<strong>о</strong>бр<strong>а</strong>з жизни. С<strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги, фит<strong>о</strong>ф<strong>а</strong>ги и<br />

хищные. С д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерн<strong>о</strong><strong>с</strong>тью из<strong>в</strong>е<strong>с</strong>тны из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>,<br />

н<strong>о</strong>, <strong>в</strong>идим<strong>о</strong>, уже <strong>о</strong>бит<strong>а</strong>ли <strong>в</strong> <strong>с</strong>илуре, <strong>о</strong>т<strong>к</strong>уд<strong>а</strong><br />

из<strong>в</strong>е<strong>с</strong>тны пр<strong>о</strong>блем<strong>а</strong>тичные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и (Peach,<br />

1882, 1898).

КЛАСС SYMPHYLA. СИМФИЛЫ<br />

Мел<strong>к</strong>ие (менее 8 мм) ф<strong>о</strong>рмы <strong>с</strong> т<strong>о</strong>н<strong>к</strong>ими,<br />

непигментир<strong>о</strong><strong>в</strong><strong>а</strong>нными п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми, пр<strong>о</strong>гн<strong>а</strong>тн<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, лишенн<strong>о</strong>й гл<strong>а</strong>з, длинными мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми и ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>р<strong>о</strong><strong>с</strong>шими<strong>с</strong>я<br />

<strong>в</strong>т<strong>о</strong>рыми м<strong>а</strong><strong>к</strong><strong>с</strong>илл<strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими нижнюю<br />

губу; 12 п<strong>а</strong>р н<strong>о</strong>г. Тул<strong>о</strong><strong>в</strong>ище у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из 14 <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong>лит<strong>а</strong> <strong>с</strong> п<strong>о</strong><strong>с</strong>ледним <strong>с</strong>егмент<strong>о</strong>м. Обычн<strong>о</strong><br />

XIII <strong>с</strong>егмент <strong>с</strong>лит <strong>с</strong> XIV и не<strong>с</strong>ет нечлени<strong>с</strong>тые<br />

цер<strong>к</strong>и. Н<strong>а</strong> третьем и <strong>с</strong>ледующих не<strong>с</strong>ущих н<strong>о</strong>ги<br />

<strong>с</strong>егмент<strong>а</strong>х <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>о</strong>г е<strong>с</strong>ть <strong>с</strong>тили и <strong>в</strong>ыпячи<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я<br />

меш<strong>о</strong>ч<strong>к</strong>и (ри<strong>с</strong>. 1). С<strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги<br />

и фит<strong>о</strong>ф<strong>а</strong>ги. Третичные — ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Scolopendrellidae, Geophilellidae и Scutigerellidae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

не н<strong>а</strong>йдены.<br />

СЕМЕЙСТВО SCOLOPENDRELLIDAE BAGNAL, 1913.<br />

СКОЛОПЕНДРЕЛЛЫ<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

Тергит<strong>о</strong><strong>в</strong> 15. Н<strong>о</strong>ги пер<strong>в</strong><strong>о</strong>й п<strong>а</strong>ры <strong>о</strong>бычн<strong>о</strong><br />

меньше или р<strong>а</strong><strong>в</strong>ны длине н<strong>о</strong>г <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й п<strong>а</strong>ры.<br />

Стили нея<strong>с</strong>ные. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три р<strong>о</strong>д<strong>а</strong>,<br />

из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы 1<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Ри<strong>с</strong>. 1—2. Кл<strong>а</strong><strong>с</strong><strong>с</strong>ы Symphyla, Pauropoda<br />

1. Scutigerella immaculate. Newport; X20.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Snodgrass,<br />

1952). 2. Pauropus silvaticus Latzel; x50,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Tiegs, 1947)<br />

КЛАСС PAUROPODA. ПАУРОПОДЫ<br />

Мел<strong>к</strong>ие (менее 1,5 мм) <strong>с</strong>л<strong>а</strong>б<strong>о</strong> хитинизир<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и <strong>с</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

лишенн<strong>о</strong>й гл<strong>а</strong>з, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми (у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых),<br />

р<strong>а</strong>зд<strong>в</strong><strong>о</strong>енными н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>а</strong>нтенн<strong>а</strong>ми, пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ющими<strong>с</strong>я<br />

<strong>в</strong> мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые жгуты;<br />

<strong>в</strong>т<strong>о</strong>рые м<strong>а</strong><strong>к</strong><strong>с</strong>иллы 1<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; 11 —12 <strong>с</strong>егмен­<br />

т<strong>о</strong><strong>в</strong> тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong> у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых и <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть;<br />

пер<strong>в</strong>ый <strong>с</strong>егмент (шейный) лишен н<strong>о</strong>г. У <strong>в</strong>зр<strong>о</strong><strong>с</strong>­<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

лых <strong>о</strong>бычн<strong>о</strong> де<strong>в</strong>ять, реже де<strong>с</strong>ять п<strong>а</strong>р н<strong>о</strong>г. Тергиты<br />

третьег<strong>о</strong>, пят<strong>о</strong>г<strong>о</strong>, <strong>с</strong>едьм<strong>о</strong>г<strong>о</strong> и де<strong>в</strong>ят<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м <strong>в</strong><strong>о</strong><strong>о</strong>ружены трих<strong>о</strong>б<strong>о</strong>триями;<br />

тергиты пер<strong>в</strong><strong>о</strong>г<strong>о</strong>, чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong>, <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> и<br />

де<strong>с</strong>ят<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны (ри<strong>с</strong>. 2).<br />

С<strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тны.<br />

http://jurassic.ru/

КЛАСС DIPLOPODA. ДВУПАРНОНОГИЕ<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

Средней <strong>в</strong>еличины и <strong>к</strong>рупные (0,7—5 <strong>с</strong>м)<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и. К<strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>а</strong>нтеяны, <strong>в</strong><strong>о</strong><strong>с</strong>ьми-де<strong>с</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые;<br />

групп<strong>а</strong> пр<strong>о</strong><strong>с</strong>тьгх гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> или<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; <strong>с</strong>егмент <strong>в</strong>т<strong>о</strong>рых<br />

м<strong>а</strong><strong>к</strong>еилл редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н, <strong>в</strong> п<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льный<br />

пери<strong>о</strong>д <strong>о</strong>т нег<strong>о</strong> не <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я ни<strong>к</strong><strong>а</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>лед<strong>а</strong>.<br />

Тергиты <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые; .пер<strong>в</strong>ый <strong>с</strong>егмент<br />

пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, лишен <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, три <strong>с</strong>ледующие<br />

з<strong>а</strong> ним т<strong>а</strong><strong>к</strong>же пр<strong>о</strong><strong>с</strong>тые, не<strong>с</strong>ут п<strong>о</strong> п<strong>а</strong>ре <strong>к</strong><strong>о</strong>неч­<br />

н<strong>о</strong><strong>с</strong>тей. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>с</strong>ближены<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие реду<strong>к</strong>ции <strong>с</strong>тернит<strong>о</strong><strong>в</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия<br />

п<strong>а</strong>рные, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> третьем тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>м<br />

<strong>с</strong>егменте. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленье н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бильн<strong>о</strong>, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляя<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 80% <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Н<strong>а</strong>д<strong>о</strong>тряды: Pselaphognatha, Chilognatha.<br />

НАДОТРЯД P S E L A P H O G N A T H A<br />

Стен<strong>к</strong>и тел<strong>а</strong> мяг<strong>к</strong>ие, без из<strong>в</strong>е<strong>с</strong>ти <strong>в</strong> хитине.<br />

В<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> теле <strong>о</strong>чень <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения<br />

(зубч<strong>а</strong>тые, пильч<strong>а</strong>тые, чешуйч<strong>а</strong>тые и др.),<br />

<strong>с</strong><strong>о</strong>бр<strong>а</strong>нные <strong>в</strong> пуч<strong>к</strong>и. Н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>е при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

трих<strong>о</strong>б<strong>о</strong>трии. Г<strong>о</strong>н<strong>о</strong>п<strong>о</strong>ды и тел<strong>о</strong>п<strong>о</strong>ды <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Третичные — ныне. Один <strong>о</strong>тряд <strong>с</strong> <strong>о</strong>дним<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>м — Polyxenidae.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong><br />

3. Polyxenus largus Linnaeus; х 25,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Attems, 1926).<br />

4. Acantherpestes vicinus Fritsch; x 5,0, н. Пермь, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Laurentiaux,<br />

1953; no Fritsch, 1900). 5. Euphoberia llarenai Melendez;<br />

<strong>с</strong>егмент тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong>, x 3,0, н. Пермь, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Laurentiaux,<br />

1953; п<strong>о</strong> Melendez, 1948). 6. Tomiulus angulatus Martynov; A —<br />

з<strong>а</strong>дний <strong>к</strong><strong>о</strong>нец жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong>; x 5,0; Б — нижняя ч<strong>а</strong><strong>с</strong>ть трех тул<strong>о</strong><strong>в</strong>ищных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong>н<strong>а</strong>ружи, х 10,0; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

{М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1939). 7. Pleurojulus levis Fritsch; х 7,0, н. Пермь,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Laurentiaux, 1953; п<strong>о</strong> Fritsch, 1900). 8. Archiscud-<br />

22<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО POLYXENIDAE LATREILLE,<br />

1825<br />

Тул<strong>о</strong><strong>в</strong>ище из 10—12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти. Пле<strong>в</strong>ры р<strong>а</strong>з<strong>в</strong>иты (ри<strong>с</strong>. 3). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Де<strong>с</strong>ять <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

д<strong>в</strong><strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Mplopoda<br />

deria tapeta Fritsch; x 2,0. н. пермь, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Laurentiaux,<br />

1953,no Fritsch, 1900). 9. Glomeris marginata Villers; x2,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Attems, 1926), 10. Polydesmus sp.; x 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Attems, 1926),<br />

11. Julus ligulifer Latzel; x 2,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Attems, 1926). 12. Arthropleura<br />

armata Jordan et Meyer; A — <strong>а</strong>бд<strong>о</strong>мен (x 0,05); Б —<br />

передний тергит; В — ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция <strong>о</strong>дн<strong>о</strong>й п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей;<br />

st — <strong>с</strong>тернит, t— <strong>в</strong>ентр<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть, tg —тергит, х—р<strong>о</strong>зет<strong>к</strong><strong>о</strong><strong>в</strong>идный<br />

<strong>о</strong>рг<strong>а</strong>н, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> и К<strong>а</strong>р<strong>а</strong>г<strong>а</strong>ндин<strong>с</strong><strong>к</strong>ий ;б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(St0rmer, 1944)

НАДОТРЯД CHILOGNATHA<br />

Хитин п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong> тел<strong>а</strong> пр<strong>о</strong>пит<strong>а</strong>н из<strong>в</strong>е<strong>с</strong>тью.<br />

В<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и, е<strong>с</strong>ли <strong>о</strong>ни <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я, пр<strong>о</strong><strong>с</strong>тые и не<br />

<strong>с</strong><strong>о</strong>бр<strong>а</strong>ны <strong>в</strong> пуч<strong>к</strong>и. Трих<strong>о</strong>б<strong>о</strong>трии <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

С<strong>а</strong>мцы <strong>с</strong> г<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>а</strong>ми н<strong>а</strong> <strong>с</strong>едьм<strong>о</strong>м <strong>с</strong>егменте или<br />

тел<strong>о</strong>п<strong>о</strong>д<strong>а</strong>ми н<strong>а</strong> <strong>к</strong><strong>о</strong>нце тел<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Средней <strong>в</strong>еличины и <strong>к</strong>рупные (д<strong>о</strong> 50 <strong>с</strong>м).<br />

Тел<strong>о</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>е; н<strong>а</strong> <strong>о</strong>тдельных <strong>с</strong>егмент<strong>а</strong>х<br />

имеют<strong>с</strong>я длинные шипы 1<br />

р<strong>а</strong>зличн<strong>о</strong>й ф<strong>о</strong>рмы; не<br />

<strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>ют<strong>с</strong>я ни <strong>в</strong> ш<strong>а</strong>р, ни <strong>в</strong> <strong>с</strong>пир<strong>а</strong>ль. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

б<strong>о</strong>льш<strong>а</strong>я, <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я; <strong>а</strong>нтенны ше<strong>с</strong>ти^<strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые;<br />

гл<strong>а</strong>з<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> чи<strong>с</strong>л<strong>а</strong><br />

те<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>о</strong>м<strong>к</strong>нутых пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong>. Тер гиты<br />

<strong>о</strong>бр<strong>а</strong>зуют лишь п<strong>о</strong>лу<strong>к</strong><strong>о</strong>льц<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>. Пле<strong>в</strong>ры<br />

и <strong>с</strong>терниты <strong>с</strong>п<strong>а</strong>яны (<strong>о</strong>т<strong>с</strong>юд<strong>а</strong> и н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние<br />

<strong>о</strong>тряд<strong>а</strong>) и <strong>о</strong>бр<strong>а</strong>зуют нижнюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину <strong>к</strong><strong>о</strong>льц<strong>а</strong><br />

ОТРЯД PALAEOCOXOPLEURA<br />

ОТРЯД EURYSTERNA<br />

Крупные мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и <strong>с</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

не<strong>с</strong>ущей, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong>,<br />

<strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong>им тел<strong>о</strong>м и б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

(д<strong>о</strong> 60 и б<strong>о</strong>лее) <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; лишены шип<strong>о</strong><strong>в</strong> и<br />

<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бны <strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong> <strong>с</strong>пир<strong>а</strong>ль. Желези<strong>с</strong>тые<br />

п<strong>о</strong>ры имеют<strong>с</strong>я или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. К<strong>а</strong>ждый<br />

тергит п<strong>о</strong>др<strong>а</strong>зделен н<strong>а</strong> пр<strong>о</strong>тергит и мет<strong>а</strong>тергит.<br />

Пле<strong>в</strong>ры <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>о</strong>членены <strong>с</strong> тел<strong>о</strong>м, чет<strong>к</strong><strong>о</strong><br />

<strong>о</strong>тделены' <strong>о</strong>т тергит<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>к</strong><strong>а</strong>ймлены <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й<br />

линией р<strong>а</strong>зличн<strong>о</strong>й ширины. К<strong>а</strong>рб<strong>о</strong>н — три<strong>а</strong><strong>с</strong>.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Archijulidae.<br />

СЕМЕЙСТВО ARCHIJULIDAE SCUDDER,<br />

1873<br />

(Projulidae Fritsch, 1899)<br />

Тергиты т<strong>о</strong>н<strong>к</strong><strong>о</strong> пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>нные или п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>а</strong>тые.<br />

Стерниты <strong>к</strong>рупные, шир<strong>о</strong><strong>к</strong>ие, ин<strong>о</strong>гд<strong>а</strong><br />

Средней <strong>в</strong>еличины мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, не<strong>с</strong>ущей <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>, и <strong>с</strong> неб<strong>о</strong>льшим<br />

чи<strong>с</strong>л<strong>о</strong>м тергит<strong>о</strong><strong>в</strong> (16—17). С<strong>в</strong>ерты<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременные Oniscomorpha, н<strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не<br />

з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>л<strong>а</strong><strong>с</strong>ь <strong>а</strong>н<strong>а</strong>льны'м щит<strong>о</strong>м, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый был<br />

<strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н. Тергиты <strong>о</strong><strong>к</strong><strong>а</strong>ймлены ши­<br />

Де<strong>в</strong>ять <strong>о</strong>тряд<strong>о</strong><strong>в</strong>: Palaeocoxopleura, Eurysterna,<br />

Palaeomorpha, Oniscomorpha, Polydesmodea,<br />

Nematophora, Juliformia, Colobognatha, Limacomorpha;<br />

из них т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong><strong>с</strong>ледний не н<strong>а</strong>йден<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

<strong>с</strong>егмент<strong>а</strong>. Н<strong>а</strong> <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong>-<strong>с</strong>терн<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>х н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я<br />

щел<strong>а</strong><strong>в</strong>идные <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые я<strong>в</strong>ляют<strong>с</strong>я<br />

<strong>с</strong>тигм<strong>а</strong>ми или желези<strong>с</strong>тыми п<strong>о</strong>р<strong>а</strong>ми, <strong>а</strong> т<strong>а</strong><strong>к</strong>же<br />

<strong>о</strong><strong>к</strong><strong>а</strong>ймленные <strong>в</strong><strong>а</strong>ли<strong>к</strong><strong>а</strong>ми <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия, зн<strong>а</strong>чение<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых нея<strong>с</strong>н<strong>о</strong>. У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>с</strong>тигмы и <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия н<strong>а</strong>х<strong>о</strong>дят<strong>с</strong>я н<strong>а</strong> тергит<strong>а</strong>х.<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; т<strong>а</strong>зи<strong>к</strong>и<br />

<strong>с</strong>ближены (ри<strong>с</strong>. 4—5). С<strong>в</strong>язи <strong>с</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременными<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong>ми нея<strong>с</strong>ны. К<strong>а</strong>рб<strong>о</strong>н—пермь.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Euphoberidae — <strong>в</strong>не СССР.<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> единый <strong>с</strong><strong>к</strong>лерит. К<strong>а</strong>рб<strong>о</strong>н — три<strong>а</strong><strong>с</strong>.<br />

Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Tomiulus М <strong>а</strong> г t у п <strong>о</strong> v, 1936. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Т. angulatus Martynov, 1936; три<strong>а</strong><strong>с</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (м<strong>а</strong>льце<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Б<strong>а</strong>бий К<strong>а</strong>мень).<br />

Пле<strong>в</strong>риты' <strong>о</strong>чень уз<strong>к</strong>ие, не<strong>о</strong>тгр<strong>а</strong>ниченные; нижние<br />

<strong>к</strong>р<strong>а</strong>я их <strong>в</strong>ыпу<strong>к</strong>лые, ут<strong>о</strong>лщенные, <strong>в</strong>ыд<strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong>ружу. Пр<strong>о</strong>д<strong>о</strong>льные п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> тергит<strong>а</strong>х я<strong>с</strong>ные,<br />

з<strong>а</strong>х<strong>о</strong>дят и н<strong>а</strong> пр<strong>о</strong>тергиты. Вы<strong>с</strong><strong>о</strong>т<strong>а</strong> <strong>с</strong>егмент<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4,5 мм (ри<strong>с</strong>. % А, Б). Один <strong>в</strong>ид.<br />

Три<strong>а</strong><strong>с</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Archijulus Scudder, 1873; Pylojulus<br />

Fritsch, 1899 (Xyloblus Dawson, 1859);<br />

Isojutus Fritsch, 1899; Pteurojulus Fritsch, 1899<br />

(ри<strong>с</strong>. 7); Anthracojulus Fritsch, 1899; Purkynia<br />

Fritsch, 1899.<br />

ОТРЯД PALAEOMORPHA<br />

23<br />

http://jurassic.ru/<br />

п<strong>а</strong>ми или гл<strong>а</strong>д<strong>к</strong>ие, ин<strong>о</strong>гд<strong>а</strong> р<strong>а</strong>зделены н<strong>а</strong> пр<strong>о</strong>и<br />

мет<strong>а</strong>тергиты, <strong>с</strong> п<strong>а</strong>р<strong>а</strong>терпит<strong>а</strong>ми, п<strong>о</strong>х<strong>о</strong>жими н<strong>а</strong><br />

пле<strong>в</strong>ры (ри<strong>с</strong>. 8). К<strong>а</strong>рб<strong>о</strong>н. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: А<strong>с</strong>roglomeridae,<br />

Proglomeridae, Sphaerherpestidae<br />

— <strong>в</strong>не СССР.

Средней <strong>в</strong>еличины мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными гл<strong>а</strong>з<strong>а</strong>ми и немн<strong>о</strong>гими<br />

тергит<strong>а</strong>ми (14—16). Н<strong>о</strong>г <strong>о</strong>т 17 д<strong>о</strong><br />

21 п<strong>а</strong>ры. Шипы и п<strong>а</strong>р<strong>а</strong>тергиты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Пре<strong>а</strong>н<strong>а</strong>льный члени<strong>к</strong> б<strong>о</strong>льш<strong>о</strong>й, при <strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>нии<br />

жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет г<strong>о</strong>л<strong>о</strong><strong>в</strong>у.<br />

Мел — ныне. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Glomeridae.<br />

Тел<strong>о</strong> упл<strong>о</strong>щенн<strong>о</strong>е. К<strong>о</strong>лец тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong> <strong>о</strong>т 19<br />

д<strong>о</strong> 22. Прядильные железы н<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>м <strong>с</strong>егменте<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. С<strong>в</strong>ерты<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>с</strong>пир<strong>а</strong>ль<br />

К<strong>о</strong>лец тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong> 26 и б<strong>о</strong>лее. Н<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>м<br />

<strong>с</strong>егменте д<strong>в</strong>е —три п<strong>а</strong>ры прядильных желез,<br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>й — трех п<strong>а</strong>р<strong>а</strong>х прядильных<br />

<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>. Третичные — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

30 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Craspedosomatidae.<br />

Мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и <strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong>им тел<strong>о</strong>м,<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, <strong>с</strong>н<strong>а</strong>бженн<strong>о</strong>й гл<strong>а</strong>з<strong>а</strong>ми<br />

из те<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>о</strong>м<strong>к</strong>нутых пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Чи<strong>с</strong>л<strong>о</strong> <strong>к</strong><strong>о</strong>лец тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong> б<strong>о</strong>лее 40. Прядильные<br />

железы <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Мез<strong>о</strong>з<strong>о</strong>й — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 15 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<br />

них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Julidae.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я, треуг<strong>о</strong>льн<strong>а</strong>я, <strong>с</strong> м<strong>а</strong><strong>с</strong><strong>с</strong>и<strong>в</strong>ными<br />

<strong>а</strong>нтенн<strong>а</strong>ми. Верхняя губ<strong>а</strong> без зубц<strong>о</strong><strong>в</strong>.<br />

Тул<strong>о</strong><strong>в</strong>ище или <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е, или<br />

длинн<strong>о</strong>е и уз<strong>к</strong><strong>о</strong>е. Третичные—ныне. Пять<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Polyzoniidae.<br />

ОТРЯД ONISCOMORPHA<br />

ОТРЯД POLYDESMODEA<br />

ОТРЯД NEMATOPHORA<br />

ОТРЯД JULIFORM1A<br />

ОТРЯД COLOBOGNATHA<br />

24<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО GLOMERIDAE BRANDT,<br />

1833<br />

Префемур 19-й п<strong>а</strong>ры н<strong>о</strong>г у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>с</strong> <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м,<br />

не<strong>с</strong>ущим щетин<strong>к</strong>у, или т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong> щетин<strong>к</strong><strong>о</strong>й.<br />

Бедр<strong>о</strong> и г<strong>о</strong>лень <strong>с</strong> <strong>в</strong>нутренними л<strong>о</strong>п<strong>а</strong><strong>с</strong>те<strong>в</strong>идными<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Сп<strong>о</strong><strong>с</strong><strong>о</strong>бны <strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>ть<strong>с</strong>я<br />

<strong>в</strong> ш<strong>а</strong>р (ри<strong>с</strong>. 9). Мел — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>дин из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

(ри<strong>с</strong>. 10). Третичные — ныне. Из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тны <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и,<br />

не <strong>о</strong>пределимые б<strong>о</strong>лее т<strong>о</strong>чн<strong>о</strong>.<br />

СЕМЕЙСТВО CRASPEDOSOMATIDAE<br />

VERHOEFF, 1889<br />

С<strong>а</strong>м<strong>о</strong>е б<strong>о</strong>льш<strong>о</strong>е 30, ч<strong>а</strong>ще 28 тул<strong>о</strong><strong>в</strong>ищных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Ментум п<strong>о</strong>др<strong>а</strong>зделен н<strong>а</strong> <strong>в</strong>т<strong>о</strong>ричный<br />

ментум и пр<strong>о</strong>ментум. Вт<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong> н<strong>о</strong>г <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин из них <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО JULIDAE VERHOEFF, Ш1<br />

Обе п<strong>а</strong>ры 1<br />

г<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д <strong>с</strong>прят<strong>а</strong>ны <strong>в</strong> углубления.<br />

Пени<strong>с</strong> шир<strong>о</strong><strong>к</strong>ий, <strong>с</strong> д<strong>в</strong>умя <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями и д<strong>в</strong>умя<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>о</strong><strong>с</strong>триями (ри<strong>с</strong>. 11). Мел — ныне.<br />

Б<strong>о</strong>лее 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин из<br />

них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

и <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из мел<strong>а</strong> Гренл<strong>а</strong>ндии.<br />

СЕМЕЙСТВО POLYZONIIDAE GERVAIS, 1844<br />

Стерниты 1<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные; gnathochilarium <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из <strong>о</strong>дн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и; гл<strong>а</strong>з<strong>а</strong> е<strong>с</strong>ть; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, не п<strong>о</strong><strong>к</strong>рыт<strong>а</strong>я тергит<strong>а</strong>ми <strong>с</strong>ледующих<br />

з<strong>а</strong> нею <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин из<br />

них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).

СЕМЕЙСТВО ARTHROPLEURIDAE<br />

ZITTEL, 1848<br />

(Не<strong>с</strong>т<strong>о</strong>р Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>)<br />

Крупные члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong><strong>в</strong>шие 1,5 м<br />

длины. Б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты 1<br />

тергит<strong>о</strong><strong>в</strong> шир<strong>о</strong><strong>к</strong>ие,<br />

<strong>о</strong>тделены <strong>о</strong>т <strong>с</strong>рединн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>, н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>я <strong>с</strong><strong>о</strong>члененные<br />

пле<strong>в</strong>ры у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong><br />

Eurysterna. Сегменты п<strong>о</strong>чти <strong>в</strong><strong>с</strong>е <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы 1<br />

и<br />

лишь три-четыре з<strong>а</strong>дних немн<strong>о</strong>г<strong>о</strong> меньше<br />

п<strong>о</strong> <strong>в</strong>еличине и -ин<strong>о</strong>й ф<strong>о</strong>рмы. Чи<strong>с</strong>л<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти при<strong>к</strong>реплены <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

<strong>с</strong>егмент<strong>а</strong>, имеющему <strong>в</strong>ыпу<strong>к</strong>лую <strong>в</strong>ентр<strong>а</strong>льную<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong><br />

ж<strong>а</strong>бр<strong>а</strong>, и четыре <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие р<strong>а</strong>ди<strong>а</strong>льные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и,<br />

<strong>о</strong>бр<strong>а</strong>зующие <strong>о</strong><strong>с</strong><strong>о</strong>бый <strong>о</strong>рг<strong>а</strong>н — «р<strong>о</strong>зет<strong>к</strong>у».<br />

К<strong>а</strong>жд<strong>а</strong>я <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>в</strong>яти<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их члени<strong>к</strong><strong>о</strong><strong>в</strong>, п<strong>о</strong><strong>с</strong>ледний из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых —<br />

прет<strong>а</strong>рз. Дет<strong>а</strong>ли <strong>с</strong>тр<strong>о</strong>ения <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей еще не<br />

я<strong>с</strong>ны; В<strong>а</strong>терл<strong>о</strong> (Waterlot, 1934) <strong>с</strong>чит<strong>а</strong>л их д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>ближенными <strong>в</strong>ет<strong>в</strong>ями; Ст<strong>о</strong>рмер<br />

(Stormer, 1944) п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет, чт<strong>о</strong> предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>емые<br />

д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и м<strong>о</strong>гут быть ч<strong>а</strong><strong>с</strong>тями <strong>с</strong>пинн<strong>о</strong>й<br />

и брюшн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тей <strong>о</strong>дн<strong>о</strong>й и т<strong>о</strong>й же <strong>в</strong>ет<strong>в</strong>и.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> и тель<strong>с</strong><strong>о</strong>н неиз<strong>в</strong>е<strong>с</strong>тны.<br />

ORDO INCERTAE SEDIS<br />

От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие д<strong>а</strong>нных п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы и<br />

д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерных <strong>с</strong><strong>в</strong>едений <strong>о</strong> <strong>с</strong>тр<strong>о</strong>ении <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> з<strong>а</strong>трудняют у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ление <strong>с</strong><strong>в</strong>язей этих<br />

члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих <strong>с</strong> другими групп<strong>а</strong>ми. Шульце<br />

(Schulze, 1939) <strong>о</strong>тн<strong>о</strong><strong>с</strong>ил <strong>а</strong>ртр<strong>о</strong>пле<strong>в</strong>рид <strong>к</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зны'м;<br />

Ст<strong>о</strong>рмер (1944) <strong>с</strong>чит<strong>а</strong>л <strong>в</strong><strong>о</strong>зм<strong>о</strong>жным<br />

<strong>в</strong>ыделить <strong>а</strong>ртр<strong>о</strong>пле<strong>в</strong>рид <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бый <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>. Н<strong>о</strong><strong>в</strong>ые<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния К<strong>о</strong>м<strong>а</strong>ре<strong>к</strong><strong>а</strong> (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1960)<br />

у<strong>с</strong>т<strong>а</strong>н<strong>а</strong><strong>в</strong>ли<strong>в</strong><strong>а</strong>ют близ<strong>о</strong><strong>с</strong>ть этих члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

<strong>к</strong> д<strong>в</strong>уп<strong>а</strong>рн<strong>о</strong>н<strong>о</strong>гим мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong>м (именн<strong>о</strong> <strong>к</strong> Р<strong>о</strong>lydesmodea).<br />

К<strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д.<br />

Arthropleura Jordan et Meyer, 1854. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. artnata Jordan et Meyer, 1854; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С<strong>а</strong><strong>а</strong>р<strong>с</strong><strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн. Пле<strong>в</strong>ры <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими<br />

шип<strong>а</strong>ми п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й <strong>в</strong>еличины 1<br />

и <strong>с</strong> <strong>к</strong>илем,,<br />

пр<strong>о</strong>х<strong>о</strong>дящим ближе <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю пле<strong>в</strong>р<br />

и <strong>о</strong>бр<strong>а</strong>зующим пл<strong>а</strong><strong>в</strong>ный из_гиб, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый п<strong>о</strong><strong>в</strong>т<strong>о</strong>ряет<br />

изгиб переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Передний <strong>к</strong>р<strong>а</strong>й<br />

пле<strong>в</strong>р <strong>с</strong>илын<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый, з<strong>а</strong>дний — <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

<strong>в</strong>ыем<strong>к</strong><strong>о</strong>й. Н<strong>а</strong> <strong>с</strong><strong>о</strong>б<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> тергит<strong>а</strong>х п<strong>о</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong><br />

буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>, не<strong>с</strong>ущих шипы (ри<strong>с</strong>. 12 А—В). Семь<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. Ср. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н К<strong>а</strong>р<strong>а</strong>г<strong>а</strong>ндин<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> (еще не <strong>о</strong>пи<strong>с</strong><strong>а</strong>нные<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и).<br />

КЛАСС CHILOPODA. ГУБОНОГИЕ<br />

Крупные мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>тделен<strong>а</strong><br />

<strong>о</strong>т тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong>. Антенны <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong><br />

чи<strong>с</strong>л<strong>а</strong> (14 и б<strong>о</strong>лее) г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных члени<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Вт<strong>о</strong>рые м<strong>а</strong><strong>к</strong><strong>с</strong>иллы х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итые, <strong>с</strong>р<strong>о</strong><strong>с</strong>шие<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong><strong>о</strong> <strong>с</strong>тернит<strong>о</strong>м. Тул<strong>о</strong><strong>в</strong>ище<br />

ОТРЯД ЕР<br />

Тергиты г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, и <strong>к</strong><strong>а</strong>ждый из них<br />

<strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует п<strong>а</strong>ре <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие эпим<strong>о</strong>рфн<strong>о</strong>е: м<strong>о</strong>л<strong>о</strong>дые<br />

мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и п<strong>о</strong><strong>к</strong>ид<strong>а</strong>ют яйц<strong>о</strong> <strong>с</strong> п<strong>о</strong>лным чи<strong>с</strong>л<strong>о</strong>м<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Третичные —<br />

ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Geophilomorpha, Scolopendromorpha.<br />

ПОДОТРЯД<br />

GEOPHILOMORPHA<br />

Тел<strong>о</strong> <strong>о</strong>чень длинн<strong>о</strong>е и т<strong>о</strong>н<strong>к</strong><strong>о</strong>е; чи<strong>с</strong>л<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong>т 35 д<strong>о</strong> 177; члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенны 14;<br />

гл<strong>а</strong>з нет. Мез<strong>о</strong>з<strong>о</strong>й — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Geophilidae.<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

упл<strong>о</strong>щенн<strong>о</strong> д<strong>о</strong>р<strong>с</strong><strong>о</strong><strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пер<strong>в</strong><strong>о</strong>г<strong>о</strong><br />

тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> н<strong>о</strong>г<strong>о</strong>челю<strong>с</strong>ти.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне. В <strong>к</strong><strong>а</strong>рб<strong>о</strong>не 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

н<strong>а</strong>йдены лишь пр<strong>о</strong>блем<strong>а</strong>тичные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

Chilopoda. Отряды: Epimorpha, Anamorpha.<br />

10RPHA<br />

СЕМЕЙСТВО GEOPHILIDAE NEWPORT, 1844<br />

Верхняя губ<strong>а</strong> из трех <strong>с</strong><strong>к</strong>лерит<strong>о</strong><strong>в</strong>. Н<strong>а</strong> <strong>к</strong>липеу<strong>с</strong>е<br />

<strong>о</strong>дн<strong>о</strong>-д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>в</strong>етлых пятн<strong>а</strong>. П<strong>а</strong>р<strong>а</strong>тергитьг<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>о</strong>ги ше<strong>с</strong>ти-<strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

(ри<strong>с</strong>. 13). Мел — ныне. Б<strong>о</strong>лее 60 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и <strong>о</strong>дин <strong>в</strong>ымерший<br />

р<strong>о</strong>д из мел<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ПОДОТРЯД<br />

SCOLOPENDROMORPHA.<br />

СКОЛОПЕНДРЫ<br />

Тел<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> 25—27, н<strong>о</strong>г 21—23 п<strong>а</strong>ры. Третичные<br />

— ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Scolopendridae,<br />

Cryptopidae.<br />

25<br />

http://jurassic.ru/

СЕМЕЙСТВО SCOLOPENDRIDAE<br />

NEWPORT, 1844<br />

Гл<strong>а</strong>з<strong>а</strong> имеют<strong>с</strong>я; л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

(ри<strong>с</strong>. 14). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>лее 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

ОТРЯД ANAMORPHA<br />

Тергиты или гетер<strong>о</strong>н<strong>о</strong>мные, либ<strong>о</strong> <strong>о</strong>дн<strong>о</strong>му<br />

тергиту <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>уют д<strong>в</strong>е п<strong>а</strong>ры н<strong>о</strong>г. Р<strong>а</strong>з<strong>в</strong>итие<br />

<strong>а</strong>н<strong>а</strong>м<strong>о</strong>рфн<strong>о</strong>е: из яйц<strong>а</strong> <strong>в</strong>ы'х<strong>о</strong>дит мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong><br />

<strong>с</strong> неп<strong>о</strong>лным чи<strong>с</strong>л<strong>о</strong>м <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong> п<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льный пери<strong>о</strong>д.<br />

Сегмент<strong>о</strong><strong>в</strong> тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong> 19, п<strong>а</strong>р <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

15. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Lithobiomorpha,<br />

Scutigeromorpha.<br />

26<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО CRYPTOPIDAE<br />

NEWPORT, 1844<br />

Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, л<strong>а</strong>п<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>лее де<strong>с</strong>яти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>о</strong>дин<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДОТРЯД LITHOBIOMORPHA.<br />

<strong>к</strong><strong>о</strong><strong>с</strong>тян<strong>к</strong>и<br />

Сегменты <strong>с</strong> х<strong>о</strong>дильными <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями имеют<br />

15 чередующих<strong>с</strong>я б<strong>о</strong>льших и м<strong>а</strong>лых тергит<strong>о</strong><strong>в</strong>.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 30 и б<strong>о</strong>лее. Пр<strong>о</strong><strong>с</strong>тые<br />

гл<strong>а</strong>з<strong>к</strong>и. Третичные — ныне. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Lithobiidae.

СЕМЕЙСТВО LITHOBIIDAE<br />

NEWPORT, 1844<br />

Гл<strong>а</strong>з<strong>а</strong> н<strong>а</strong> <strong>к</strong>р<strong>а</strong>ю г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> щит<strong>к</strong><strong>а</strong>. Н<strong>о</strong>г<strong>а</strong> <strong>о</strong>т<br />

пер<strong>в</strong><strong>о</strong>й д<strong>о</strong> пят<strong>о</strong>й п<strong>а</strong>ры <strong>с</strong> <strong>к</strong><strong>о</strong><strong>к</strong>е<strong>а</strong>льными желез<strong>а</strong>ми.<br />

Г<strong>о</strong>н<strong>о</strong>л<strong>о</strong>ды <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>о</strong>дн<strong>о</strong>- или д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

(ри<strong>с</strong>. 15). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДОТРЯД SCUTIGEROMORPHА.<br />

МУХОЛОВКИ<br />

Н<strong>а</strong> не<strong>с</strong>ущие н<strong>о</strong>ги <strong>с</strong>егменты <strong>о</strong>т 2-г<strong>о</strong> д<strong>о</strong> 16-г<strong>о</strong><br />

прих<strong>о</strong>дит<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емь я<strong>с</strong>ных тергит<strong>о</strong><strong>в</strong>.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн д<strong>о</strong> 400. Н<strong>о</strong>ги не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong><br />

длинные и т<strong>о</strong>н<strong>к</strong>ие. Л<strong>а</strong>п<strong>к</strong><strong>а</strong> жгути<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>а</strong>я, п<strong>о</strong>др<strong>а</strong>зделен<strong>а</strong><br />

н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> мел<strong>к</strong>их <strong>в</strong>т<strong>о</strong>ричных<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong>. Гл<strong>а</strong>з<strong>а</strong> <strong>к</strong>рупные, ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные.<br />

Третичные —ныне. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Scutigeridae.<br />

СЕМЕЙСТВО SCUTIGERIDAE GERVAIS,<br />

1847<br />

Тергит н<strong>о</strong>г<strong>о</strong>челю<strong>с</strong>тей м<strong>а</strong>лень<strong>к</strong>ий. Стигмы<br />

неп<strong>а</strong>рные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е<br />

тергит<strong>о</strong><strong>в</strong>, з<strong>а</strong> и<strong>с</strong><strong>к</strong>лючением п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> (ри<strong>с</strong>. 16).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

http://jurassic.ru/

ЛИТЕРАТУРА<br />

М<strong>а</strong>ртын<strong>о</strong><strong>в</strong> А. В. 1936. О не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>о</strong><strong>в</strong>ых м<strong>а</strong>тери<strong>а</strong>л<strong>а</strong>х<br />

члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих жи<strong>в</strong><strong>о</strong>тных из Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> 'б<strong>а</strong><strong>с</strong><strong>с</strong>ейне<br />

Из<strong>в</strong>. А<strong>к</strong><strong>а</strong>д. н<strong>а</strong>у<strong>к</strong> СССР, би<strong>о</strong>л. <strong>с</strong>ер., № 6, <strong>с</strong>тр. 1258—<br />

1260.<br />

Ш<strong>а</strong>р<strong>о</strong><strong>в</strong> А. Г. 1960. Ю. К<strong>о</strong>м<strong>а</strong>ре<strong>к</strong> <strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>о</strong><strong>к</strong><strong>о</strong>м<br />

п<strong>о</strong>л<strong>о</strong>жении Arthropleura. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. журн., № 3,<br />

<strong>с</strong>тр. 140.<br />

Andrée К. 1909. Neue Funde von Arthropleura armata<br />

Jordan. Zbl. Miner. usw., S. 753— .1910. Zur Kenntnis<br />

der Crustaceen-Gattung Arthropleura und deren systematische<br />

Stellung. Palaeontographica, Bd. 57, S. 67.<br />

A 11 e m s С. 1926. Myriopoda. In: Kükental «Handbuch<br />

der Zoologie», iBd. 4, H. il, S. 1—402.<br />

Bachofen von Echt A. 1942. Über die Myriapoden<br />

des Bernstein. Palaeobiologica, Bd. 7, H. 5—6, S.<br />

394—403. Baldwin W. 1911. Fossil Myriapoda from<br />

the middle coal measures of sparth bottoms Rochdale,<br />

Lancashire. Geol. Mag., v. 8, ip. 74—80. В a g n a 1 A. S.<br />

1913. On the classification of the order Symphyla. Journ.<br />

Linn. Soc. London, v. 32.<br />

F r i t s <strong>с</strong> h A. .1899. Fauna der Gaskohle und der Kalk- -<br />

steine der Permformation Böhmens, Bd. 4, H. 1—2, S. 1 —<br />

64.<br />

Gervais P. 1844. Sur la ponte et le développement<br />

des Glomerides. Procès verbaux soc. philomat. Paris.—<br />

1847. Myriapodes, in Walkenaer «Histoire naturelle des<br />

insectes aptères», Bd. 4. Guthör 1 P. 1934. Die Arthropoden<br />

aus dem Karbon und Perm des Saar-Nahe-Pfalz-<br />

Gebietes. Abh. Geol. Landesanst., N. F., H. 164.— 1936.<br />

Arthropleura, der Riesengliederfüssler des Oberkarbons,<br />

und seine Verbreitung in den europäischen Steinkohlenbecken.<br />

Glückauf, Nr. 39, S. 966—975.<br />

Jordan H. und Meyer H. 1854. Uber die Crustaccen<br />

der Steinkohlenformation von Saarbrücken. Palaeontographica,<br />

Bd. 4, S. 13—15.<br />

L a t r e i 11 e P. A. 1825. Familles naturelles du règne<br />

animal. Laurentiaux D. 1953. Classe des Myriapodes.<br />

In: Piveteau «Traité de paléontologie», t. III, Paris.<br />

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Melendez B. 1948. Un Miriapodo fosil en ei Estefaniense<br />

de Llombera (Léon). Madrid, Bol. R. Soc. espanola<br />

Hist. Nat., t. XIV.<br />

Newport G. 1844. A list of the species of Myriapoda,<br />

order Chilopoda, contained in the cabinets of the<br />

British Museum. Ann. and Mag. iNat. Hist., v. 13.<br />

Peach B. N. 1882. On some fossil myriapods from<br />

the lower old Red Sandstone of Forfarshire. Proc. Roy.<br />

Phys. Soc, Edinburgh, v. 7, .p. 177—188. 1898. One some<br />

new myriapods from the Palaeozoic Rocks of Scottland.<br />

Proc. Roy. Phys. Soc, Edinburgh, v. 14, p. 113—126. P г uv<br />

<strong>о</strong> s t P. 1912. Sur la présence du genre Arthropleura<br />

dans le terrain houiller du Nord et du Pas-de-Calais. Ann.<br />

Soc. Géol. Nord, t. 41, p. 59.—1930. La faune continentale<br />

du terrain houiller de la Belgique. Mém. Mus. Roy.<br />

Hist. Nat. Belg., t. 44, p. 171.<br />

Schulze P. 1939. Bemerkenswerte palaeozoische<br />

Arthropoden, die wahrscheinlich in die Spinnentierreiche<br />

gehören. Ztschr. Morph, u. Ökol. d. Tiere, Bd. 35. S <strong>с</strong> u dder<br />

S. H. 1882. Archipolypoda, a subordinal type of s pinned<br />

myriapods from the Carboniferous formation. Mém.<br />

Bost. Soc. Nat. Hist., t. III. Snodgrass R. E. 1952.<br />

A texbook of Arthropod anatomy, p. 1—363, N. Y. S t ö rm<br />

e r L. 1944. On the relationships and phylogeny of<br />

fossil and recent Arachnomorpha. Skrifter Utgitt av Det<br />

Ncrske Videnskaps-Akademi i Oslo. I. Mat.-Naturv.<br />

Klasse, Nr. 5, S. 1—.152.<br />

Tiegs О. W. 1947. The development and affinities<br />

on the Pauropoda, based on a study of Pauropus silvaticus.<br />

Quart. Journ. Micr. Sei., v. 88, pt. 2, ip. 165—336.<br />

Verhoeff K. W. 1911. Zur Systematik, Phylogenie<br />

und vergleichenden Morphologie der Juliden und über<br />

einige andere Diplopoden. Archiv f. Naturgeschichte, Bd.<br />

I. —1911. Zur Kenntni's des Mentum der Juliden und über<br />

Protojuliden. Zool. Anz., Bd. 38, Nr. 24.<br />

Waterlot G. 1934. La faune continentale du terrain<br />

houiller Sarro-Lorrain. Études de gîtes minéraux de<br />

la France. Lille, p. 52—110.

КЛАСС INSECTA. НАСЕКОМЫЕ<br />

И<strong>с</strong>т<strong>о</strong>рия изучения<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые — н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бширный и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих, зн<strong>а</strong>чительн<strong>о</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ющий п<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ему мн<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зию любые<br />

другие группы жи<strong>в</strong><strong>о</strong>тных. Эт<strong>о</strong> <strong>к</strong>р<strong>а</strong>йнее<br />

<strong>о</strong>билие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>пределяет <strong>о</strong>чень б<strong>о</strong>льшую<br />

нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><strong>с</strong>ть изученн<strong>о</strong><strong>с</strong>ти <strong>о</strong>тдельных групп<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: н<strong>а</strong>ряду <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> изученными <strong>о</strong>тряд<strong>а</strong>ми<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>уют еще м<strong>а</strong>л<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тные, прит<strong>о</strong>м<br />

<strong>о</strong>бширные группы, и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я<br />

н<strong>а</strong> пер<strong>в</strong>ичн<strong>о</strong>й, ф<strong>а</strong>униети<strong>к</strong><strong>о</strong>-реги<strong>с</strong>тр<strong>а</strong>ци<strong>о</strong>нн<strong>о</strong>й<br />

<strong>с</strong>т<strong>а</strong>дии. Одн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>в</strong> цел<strong>о</strong>м эт<strong>о</strong>т <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих <strong>в</strong> з<strong>о</strong><strong>о</strong>л<strong>о</strong>гии я<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>дним из<br />

н<strong>а</strong>иб<strong>о</strong>лее д<strong>а</strong><strong>в</strong>н<strong>о</strong> и п<strong>о</strong>лн<strong>о</strong> и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>нных.<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> изучены<br />

г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> меньше, чем <strong>с</strong><strong>о</strong><strong>в</strong>ременные. Причины<br />

эт<strong>о</strong>г<strong>о</strong> з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> б<strong>о</strong>льших трудн<strong>о</strong><strong>с</strong>тях<br />

<strong>о</strong>пределения и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> неп<strong>о</strong>лных, прит<strong>о</strong>м немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

и трудн<strong>о</strong> н<strong>а</strong>х<strong>о</strong>димых.<br />

Опи<strong>с</strong><strong>а</strong>ние н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> был<strong>о</strong> н<strong>а</strong>ч<strong>а</strong>т<strong>о</strong><br />

<strong>о</strong>чень д<strong>а</strong><strong>в</strong>н<strong>о</strong>. Т<strong>а</strong><strong>к</strong>, еще <strong>в</strong> пер<strong>в</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

XVIII <strong>в</strong>., <strong>в</strong> д<strong>о</strong>линее<strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>е <strong>в</strong>ремя, Зенделем<br />

(Sendelius, 1742) был<strong>о</strong> из<strong>о</strong>бр<strong>а</strong>жен<strong>о</strong> и <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<strong>о</strong><br />

б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.<br />

Н<strong>о</strong> б<strong>о</strong>лее пл<strong>а</strong>н<strong>о</strong>мерн<strong>о</strong>е <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong><strong>с</strong>ь лишь <strong>с</strong><br />

20-х г<strong>о</strong>д<strong>о</strong><strong>в</strong> пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> <strong>в</strong>е<strong>к</strong><strong>а</strong>, причем пер<strong>в</strong>ые р<strong>а</strong>б<strong>о</strong>ты<br />

были п<strong>о</strong><strong>с</strong><strong>в</strong>ящены третичным и мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>им<br />

ф<strong>а</strong>ун<strong>а</strong>м. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы <strong>о</strong>пи<strong>с</strong><strong>а</strong>ния е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong>их<br />

третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых Ту<strong>с</strong><strong>с</strong>эн<strong>а</strong> де Серре (Toussaint<br />

de Serres, 1829), д<strong>а</strong>лее Геер<strong>а</strong> (Heer,<br />

1847—1860), Г<strong>а</strong>ген<strong>а</strong> (Hagen, 1854, 1862),<br />

К. Г<strong>а</strong>йден<strong>а</strong> и Л. Г<strong>а</strong>йден (С. Heyden, L. Неу-<br />

ОБЩАЯ ЧАСТЬ<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

den, 1856—1870), н<strong>а</strong><strong>к</strong><strong>о</strong>нец, У<strong>с</strong>т<strong>а</strong> л э (Oustalet,<br />

1870, 1873, 1874). Примерн<strong>о</strong> <strong>с</strong> эт<strong>о</strong>г<strong>о</strong> же <strong>в</strong>ремени<br />

п<strong>о</strong> е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong>им мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>им н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым<br />

п<strong>о</strong>я<strong>в</strong>или<strong>с</strong>ь р<strong>а</strong>б<strong>о</strong>ты Герм<strong>а</strong>р<strong>а</strong> (Germar, 1837,<br />

1839, 1842), Бр<strong>о</strong>ди (Brodie, 1845), Гибеля<br />

(Giebel, 1856) и д<strong>а</strong>лее Вейенберг<strong>а</strong> (Weyenbergh,<br />

1869, 1874) и Онпенгейм<strong>а</strong> (Oppenheim,<br />

1885, 1888).<br />

Пер<strong>в</strong>ые п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong>т<strong>а</strong>ли изуч<strong>а</strong>ть<strong>с</strong>я<br />

лишь <strong>с</strong> <strong>с</strong>ередины XIX <strong>в</strong>. Г<strong>о</strong>льденберг<strong>о</strong>м<br />

(Goldenberg, 1854, 1873, 1877). Б<strong>о</strong>лее интен<strong>с</strong>и<strong>в</strong>н<strong>о</strong><br />

дре<strong>в</strong>ние н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>о</strong>пи<strong>с</strong>ы<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь <strong>к</strong><br />

<strong>к</strong><strong>о</strong>нцу <strong>в</strong>е<strong>к</strong><strong>а</strong> С<strong>к</strong>еддер<strong>о</strong>м — из С. Амери<strong>к</strong>и (Scudder,<br />

1878, 1884, 1890), Вуд<strong>в</strong><strong>а</strong>рд<strong>о</strong>м (Woodward,<br />

1887) и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> Бр<strong>о</strong>ньяр<strong>о</strong>м — из 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

(Brongniart, 1885, 1893).<br />

Ит<strong>о</strong>ги р<strong>а</strong>б<strong>о</strong>т пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> <strong>в</strong>е<strong>к</strong><strong>а</strong> были п<strong>о</strong>д<strong>в</strong>едены<br />

<strong>о</strong>чень <strong>в</strong><strong>а</strong>жными и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниями Г<strong>а</strong>ндлирш<strong>а</strong><br />

(Handlirsch, 1904, 1906, 1908, 1919,<br />

1925); <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>а</strong>я им м<strong>о</strong>н<strong>о</strong>гр<strong>а</strong>фия н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>в</strong><strong>с</strong>ех ф<strong>а</strong>ун пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> Земли д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р<br />

я<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong><strong>а</strong>жнейшей <strong>с</strong><strong>в</strong><strong>о</strong>д<strong>к</strong><strong>о</strong>й, з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющей<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>ния и из<strong>о</strong>бр<strong>а</strong>жения п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их и мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и п<strong>о</strong>лный <strong>к</strong><strong>а</strong>т<strong>а</strong>л<strong>о</strong>г <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных<br />

<strong>к</strong> т<strong>о</strong>му <strong>в</strong>ремени <strong>к</strong><strong>а</strong>йн<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

И<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния Г<strong>а</strong>ндлирш<strong>а</strong> <strong>с</strong>тимулир<strong>о</strong><strong>в</strong><strong>а</strong>ли б<strong>о</strong>лее<br />

интен<strong>с</strong>и<strong>в</strong>н<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний <strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе, н<strong>а</strong>пример,<br />

р<strong>а</strong>б<strong>о</strong>ты* Б<strong>о</strong>лт<strong>о</strong>н<strong>а</strong> п<strong>о</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>им н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым<br />

(Botton, 1911, 1925) или Менье — п<strong>о</strong> третичным<br />

(Meunier, 1899—1910) и. <strong>в</strong> С. Амери<strong>к</strong>е,<br />

н<strong>а</strong>пример, Селл<strong>а</strong>рд<strong>с</strong><strong>а</strong> — п<strong>о</strong> перм<strong>с</strong><strong>к</strong>им (Sellards,<br />

1909) или К<strong>о</strong><strong>к</strong>ерелл<strong>а</strong> и других и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей<br />

— п<strong>о</strong> третичным ф<strong>а</strong>ун<strong>а</strong>м (Cockerell,<br />

1906—1915).<br />

29<br />

http://jurassic.ru/

Д<strong>а</strong>льнейшее р<strong>а</strong>з<strong>в</strong>итие п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гии <strong>о</strong>пределил<strong>о</strong><strong>с</strong>ь<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниями Тильярд<strong>а</strong> (Tillyard,<br />

1916—1948), изуч<strong>а</strong><strong>в</strong>шег<strong>о</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии, Е<strong>в</strong>р<strong>о</strong>пы и <strong>в</strong> д<strong>а</strong>льнейшем<br />

С. Амери<strong>к</strong>и, прит<strong>о</strong>м <strong>с</strong><strong>а</strong>мых р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

<strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>, н<strong>а</strong>чин<strong>а</strong>я <strong>с</strong> третичн<strong>о</strong>г<strong>о</strong>, д<strong>а</strong>лее<br />

три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> и, 'н<strong>а</strong><strong>к</strong><strong>о</strong>нец, п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>, преиму­<br />

ще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> перм<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>. Р<strong>а</strong>б<strong>о</strong>ты 1<br />

Тильярд<strong>а</strong>, безу­<br />

<strong>с</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>, м<strong>о</strong>гут быть <strong>о</strong>х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ны <strong>к</strong><strong>а</strong><strong>к</strong><br />

зн<strong>а</strong>чительн<strong>о</strong>е углубление и <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ция<br />

п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний <strong>в</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лении<br />

р<strong>а</strong>зр<strong>а</strong>б<strong>о</strong>т<strong>к</strong>и и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития <strong>о</strong>тдельных<br />

групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Изучение ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>л<strong>о</strong><strong>с</strong>ь ф<strong>о</strong>н<strong>о</strong>м, м<strong>а</strong>тери<strong>а</strong>л<strong>о</strong>м,<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м при<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ли<strong>с</strong>ь<br />

<strong>о</strong>тдельные группы и <strong>о</strong><strong>с</strong><strong>в</strong>ещ<strong>а</strong>ли<strong>с</strong>ь их фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие<br />

<strong>о</strong>тн<strong>о</strong>шения.<br />

Н<strong>о</strong> н<strong>а</strong>иб<strong>о</strong>лее <strong>в</strong><strong>а</strong>жные <strong>в</strong>ы<strong>в</strong><strong>о</strong>ды и <strong>о</strong>б<strong>о</strong>бщения<br />

п<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> были п<strong>о</strong>лучены г<strong>а</strong> р<strong>а</strong>б<strong>о</strong>т<strong>а</strong>х<br />

А. В. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong> (1924—1940), <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

п<strong>о</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>л фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ление <strong>в</strong> п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гии и <strong>в</strong><strong>с</strong>емерн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширял <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>ие и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

п<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мы'м для б<strong>о</strong>лее<br />

глуб<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong><strong>в</strong>ещения <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>в</strong>ел <strong>о</strong>бширные<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния п<strong>о</strong> юр<strong>с</strong><strong>к</strong>им, перм<strong>с</strong><strong>к</strong>им и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым<br />

другим ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>им <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>м из<br />

р<strong>а</strong>зличных ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений Ср. Азии, Ур<strong>а</strong>л<strong>а</strong>,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и мн<strong>о</strong>гих других р<strong>а</strong>й<strong>о</strong>н<strong>о</strong><strong>в</strong>.<br />

П<strong>о</strong>чти <strong>в</strong><strong>с</strong>е эти ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения были <strong>о</strong>т<strong>к</strong>рыты<br />

<strong>в</strong>пер<strong>в</strong>ые. Р<strong>а</strong>б<strong>о</strong>ты М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong><strong>а</strong>жны<br />

тем, чт<strong>о</strong> <strong>о</strong>ни п<strong>о</strong>л<strong>о</strong>жили н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> изучению<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> террит<strong>о</strong>рии<br />

СССР. Д<strong>о</strong> ег<strong>о</strong> р<strong>а</strong>б<strong>о</strong>т п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гии у н<strong>а</strong><strong>с</strong><br />

не <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>л<strong>о</strong> <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е, <strong>в</strong><strong>с</strong>е немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых изуч<strong>а</strong>ли<strong>с</strong>ь<br />

ин<strong>о</strong><strong>с</strong>тр<strong>а</strong>нными учеными, т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и<br />

<strong>о</strong>бширные и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>знейшие м<strong>а</strong>тери<strong>а</strong>лы п<strong>о</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ил<br />

<strong>о</strong>бз<strong>о</strong>р <strong>в</strong><strong>с</strong>ех ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

50), <strong>о</strong>пи<strong>с</strong><strong>а</strong>л ряд ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

и предл<strong>о</strong>жил <strong>а</strong>ргументир<strong>о</strong><strong>в</strong><strong>а</strong>нную фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ую<br />

<strong>с</strong>хему <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (1938).<br />

В <strong>к</strong><strong>о</strong>нце 20-х г<strong>о</strong>д<strong>о</strong><strong>в</strong> н<strong>а</strong>шег<strong>о</strong> <strong>в</strong>е<strong>к</strong><strong>а</strong> <strong>а</strong>мери<strong>к</strong><strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>г Ф. М. К<strong>а</strong>рпентер (Carpenter,<br />

1926—1959), <strong>с</strong>пеци<strong>а</strong>ли<strong>с</strong>т п<strong>о</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым групп<strong>а</strong>м<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й <strong>с</strong>е<strong>в</strong>ер<strong>о</strong><strong>а</strong>мери<strong>к</strong><strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ф<strong>а</strong>уны, н<strong>а</strong>ч<strong>а</strong>л р<strong>а</strong>б<strong>о</strong>т<strong>а</strong>ть п<strong>о</strong> п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гии;<br />

эти и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>о</strong>н пр<strong>о</strong><strong>в</strong><strong>о</strong>дил <strong>в</strong> н<strong>а</strong>учн<strong>о</strong>м<br />

<strong>к</strong><strong>о</strong>нт<strong>а</strong><strong>к</strong>те <strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ими <strong>с</strong><strong>о</strong><strong>в</strong>ременни<strong>к</strong><strong>а</strong>ми, <strong>с</strong>т<strong>а</strong>ршими<br />

<strong>к</strong><strong>о</strong>ллег<strong>а</strong>ми Тильярд<strong>о</strong>м и М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>ым.<br />

В <strong>с</strong><strong>в</strong><strong>о</strong>их р<strong>а</strong>б<strong>о</strong>т<strong>а</strong>х К<strong>а</strong>рпентер <strong>о</strong>чень п<strong>о</strong>лн<strong>о</strong> <strong>о</strong>пи<strong>с</strong>ы<strong>в</strong><strong>а</strong>л<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лял мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ции <strong>о</strong>бщег<strong>о</strong> <strong>в</strong>ид<strong>а</strong> мн<strong>о</strong>гих<br />

п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и <strong>к</strong>ритиче<strong>с</strong><strong>к</strong>и р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>л<br />

<strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>ы <strong>с</strong>и<strong>с</strong>темы р<strong>а</strong>зных групп.<br />

П<strong>о</strong>мим<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>ний н<strong>о</strong><strong>в</strong>ых ф<strong>о</strong>рм н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

30<br />

http://jurassic.ru/<br />

К<strong>а</strong>рпентер <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лял <strong>с</strong><strong>в</strong><strong>о</strong>дные <strong>о</strong>чер<strong>к</strong>и п<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым<br />

ф<strong>а</strong>уны пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> Земли. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти,<br />

р<strong>а</strong>здел п<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>е <strong>в</strong>ымерших н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>в</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>й <strong>к</strong>ниге Бру<strong>с</strong><strong>а</strong>, Мел<strong>а</strong>ндер<strong>а</strong><br />

и К<strong>а</strong>рпентер<strong>а</strong> (Brues, Melander, Carpenter,.<br />

1952). В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя К<strong>а</strong>рпентер я<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong>дним из н<strong>а</strong>иб<strong>о</strong>лее <strong>а</strong><strong>в</strong>т<strong>о</strong>ритетных п<strong>а</strong>лэнт<strong>о</strong>м<strong>с</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong>.<br />

П<strong>о</strong><strong>с</strong>ле <strong>в</strong>ых<strong>о</strong>д<strong>а</strong> м<strong>о</strong>н<strong>о</strong>гр<strong>а</strong>фии Г<strong>а</strong>ндлирш<strong>а</strong> <strong>в</strong><strong>а</strong>жнейшим<br />

перел<strong>о</strong>мным м<strong>о</strong>мент<strong>о</strong>м <strong>в</strong> р<strong>а</strong>з<strong>в</strong>итии<br />

п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их зн<strong>а</strong>ний, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ли<strong>с</strong>ь<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>. Специ<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е<br />

изучение <strong>о</strong>пределенных групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

их фил<strong>о</strong>генез<strong>о</strong><strong>в</strong> и м<strong>о</strong>рф<strong>о</strong>-фун<strong>к</strong>ци<strong>о</strong>н<strong>а</strong>льных<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей рез<strong>к</strong><strong>о</strong> п<strong>о</strong><strong>в</strong>ы<strong>с</strong>ил<strong>о</strong> д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерн<strong>о</strong><strong>с</strong>ть<br />

<strong>в</strong>ы<strong>в</strong><strong>о</strong>д<strong>о</strong><strong>в</strong>. Эти и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>с</strong>тимулир<strong>о</strong><strong>в</strong><strong>а</strong>ли<br />

р<strong>а</strong><strong>с</strong>ширение <strong>в</strong> н<strong>а</strong>шей <strong>с</strong>тр<strong>а</strong>не р<strong>а</strong>б<strong>о</strong>т п<strong>о</strong><br />

п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гии, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong>т<strong>а</strong>ли пр<strong>о</strong><strong>в</strong><strong>о</strong>дить<strong>с</strong>я<br />

ряд<strong>о</strong>м и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы 1<br />

р<strong>а</strong>б<strong>о</strong>ты п<strong>о</strong><br />

перм<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уне Ур<strong>а</strong>л<strong>а</strong> Ю. М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

(1931—1956), р<strong>а</strong>б<strong>о</strong>т<strong>а</strong> п<strong>о</strong> чешуе<strong>к</strong>рылым б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря Н. Я. Кузнец<strong>о</strong><strong>в</strong><strong>а</strong> (1941), и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф<strong>а</strong> (1938, 1946,<br />

1949, 1951) п<strong>о</strong> и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>му р<strong>а</strong>з<strong>в</strong>итию д<strong>в</strong>у<strong>к</strong>рылых<br />

и э<strong>в</strong><strong>о</strong>люции лет<strong>а</strong>тельных <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong><strong>в</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

(1940—1948, 1949, 1952) п<strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гии<br />

и э<strong>в</strong><strong>о</strong>люции р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых, О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

(1942, 1948, 1949, 1952) п<strong>о</strong> э<strong>в</strong><strong>о</strong>люции<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>няиц и <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых. Изучение реги<strong>о</strong>н<strong>а</strong>льных<br />

ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> при<strong>о</strong>брел<strong>о</strong><br />

ин<strong>о</strong>й х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер и <strong>с</strong>т<strong>а</strong>л<strong>о</strong> пр<strong>о</strong><strong>в</strong><strong>о</strong>дить<strong>с</strong>я<br />

<strong>к</strong><strong>о</strong>лле<strong>к</strong>ти<strong>в</strong>н<strong>о</strong> мн<strong>о</strong>гими <strong>с</strong>пеци<strong>а</strong>ли<strong>с</strong>т<strong>а</strong>ми; т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы,<br />

н<strong>а</strong>пример, р<strong>а</strong>б<strong>о</strong>ты п<strong>о</strong> изучению п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ф<strong>а</strong>уны Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>, п<strong>о</strong>з<strong>в</strong><strong>о</strong>ли<strong>в</strong>шие <strong>с</strong>удить<br />

<strong>о</strong> <strong>с</strong>мен<strong>а</strong>х ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> и<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ть <strong>о</strong>пределенные г<strong>о</strong>риз<strong>о</strong>нты и<br />

<strong>с</strong>л<strong>о</strong>и <strong>о</strong>тл<strong>о</strong>жений <strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й т<strong>о</strong>чн<strong>о</strong><strong>с</strong>тью.<br />

Общ<strong>а</strong>я х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong><strong>а</strong><br />

и м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые — члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие жи<strong>в</strong><strong>о</strong>тные <strong>с</strong>тел<strong>о</strong>й,<br />

рез<strong>к</strong><strong>о</strong> р<strong>а</strong>зделенным н<strong>а</strong> три гл<strong>а</strong><strong>в</strong>ных <strong>о</strong>тдел<strong>а</strong>:<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>у, грудь и брюш<strong>к</strong><strong>о</strong>; эт<strong>о</strong> деление не<br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у ф<strong>о</strong>рм, <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> п<strong>о</strong>д<strong>в</strong>ергших<strong>с</strong>я<br />

реду<strong>к</strong>ции, или у личин<strong>о</strong>чных ф<strong>а</strong>з. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из ше<strong>с</strong>ти эмбри<strong>о</strong>н<strong>а</strong>льных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й п<strong>а</strong>р<strong>о</strong>й <strong>а</strong>нтенн н<strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>м<br />

ее <strong>с</strong>егменте, без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> пер<strong>в</strong><strong>о</strong>м и<br />

третьем <strong>с</strong>егмент<strong>а</strong>х и <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми челю<strong>с</strong>тей<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льных трех, причем р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые прид<strong>а</strong>т<strong>к</strong>и,<br />

челю<strong>с</strong>ти, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>в</strong>ид<strong>о</strong>изменены.<br />

Грудь <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из трех <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, у им<strong>а</strong>г<strong>о</strong> <strong>о</strong>н<strong>а</strong><br />

н<strong>о</strong>рм<strong>а</strong>льн<strong>о</strong> не<strong>с</strong>ет три п<strong>а</strong>ры н<strong>о</strong>г н<strong>а</strong> <strong>с</strong>терн<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти и д<strong>в</strong>е п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong><br />

— н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й. И<strong>с</strong>х<strong>о</strong>дн<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> брюшных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> 11; <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти или прид<strong>а</strong>т<strong>к</strong>и

<strong>о</strong>бычн<strong>о</strong> имеют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> п<strong>о</strong><strong>с</strong>ледних трех<br />

или четырех. Дых<strong>а</strong>ние, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>в</strong><strong>о</strong>здушн<strong>о</strong>е,<br />

<strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляем<strong>о</strong>е тр<strong>а</strong>хейн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>тем<strong>о</strong>й.<br />

Тр<strong>а</strong>хеи <strong>о</strong>бычн<strong>о</strong> ,<strong>в</strong> <strong>в</strong>иде <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>й мет<strong>а</strong>мерн<strong>о</strong>й<br />

<strong>с</strong>и<strong>с</strong>темы, ж<strong>а</strong>берн<strong>о</strong>е дых<strong>а</strong>ние н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я у<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых личин<strong>о</strong><strong>к</strong> и, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong><strong>о</strong>зни<strong>к</strong>л<strong>о</strong> <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>.<br />

Нер<strong>в</strong>н<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> <strong>в</strong> <strong>в</strong>иде брюшн<strong>о</strong>й цеп<strong>о</strong>ч<strong>к</strong>и;<br />

имеет<strong>с</strong>я д<strong>в</strong><strong>а</strong> <strong>с</strong>л<strong>о</strong>жных г<strong>а</strong>нглия <strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>е,<br />

три <strong>в</strong> груди и д<strong>о</strong> 11 — <strong>в</strong> брюш<strong>к</strong>е. Брюшные<br />

г<strong>а</strong>нглии ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>литы <strong>в</strong> <strong>с</strong>л<strong>о</strong>жные <strong>с</strong>инг<strong>а</strong>нглии.<br />

Пище<strong>в</strong><strong>а</strong>рительный тр<strong>а</strong><strong>к</strong>т <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из трех<br />

<strong>о</strong>тдел<strong>о</strong><strong>в</strong>: передней, <strong>с</strong>редней и з<strong>а</strong>дней <strong>к</strong>иш<strong>о</strong><strong>к</strong>;<br />

<strong>о</strong>бычн<strong>о</strong> имеет<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>а</strong>р <strong>с</strong>люнных же­<br />

лез. Выделительные <strong>о</strong>рг<strong>а</strong>ны 1<br />

<strong>в</strong> <strong>в</strong>иде м<strong>а</strong>льпигие-<br />

<strong>в</strong>ых <strong>с</strong><strong>о</strong><strong>с</strong>уд<strong>о</strong><strong>в</strong>. Жен<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие лежит<br />

<strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong>, муж<strong>с</strong><strong>к</strong><strong>о</strong>е — <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

де<strong>в</strong>ят<strong>о</strong>г<strong>о</strong> брюшных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

р<strong>а</strong>здельн<strong>о</strong>п<strong>о</strong>лы; их п<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие<br />

<strong>в</strong> п<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющем б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> <strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>ет<strong>с</strong>я<br />

мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>о</strong>м, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным<br />

<strong>к</strong>р<strong>а</strong>йне рез<strong>к</strong><strong>о</strong>. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong>ни<br />

я<strong>в</strong>ляют<strong>с</strong>я <strong>с</strong><strong>а</strong>м<strong>о</strong>й б<strong>о</strong>льш<strong>о</strong>й п<strong>о</strong> чи<strong>с</strong>лу <strong>в</strong>ид<strong>о</strong><strong>в</strong> и п<strong>о</strong><br />

б<strong>о</strong>г<strong>а</strong>т<strong>с</strong>т<strong>в</strong>у ф<strong>о</strong>рм групп<strong>о</strong>й н<strong>а</strong>земных жи<strong>в</strong><strong>о</strong>тных.<br />

Лишь немн<strong>о</strong>гие ф<strong>о</strong>рмы, прит<strong>о</strong>м преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

личин<strong>о</strong>чные ф<strong>а</strong>зы, жи<strong>в</strong>ут <strong>в</strong> пре<strong>с</strong>н<strong>о</strong>й<br />

<strong>в</strong><strong>о</strong>де; <strong>в</strong> м<strong>о</strong>рях и <strong>о</strong><strong>к</strong>е<strong>а</strong>н<strong>а</strong>х н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

<strong>о</strong>чень немн<strong>о</strong>гими <strong>в</strong>ид<strong>а</strong>ми. В <strong>с</strong><strong>о</strong>л<strong>о</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>тых<br />

<strong>в</strong><strong>о</strong>д<strong>а</strong>х <strong>в</strong>нутренних м<strong>о</strong>рей личин<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> б<strong>о</strong>льш<strong>о</strong>м <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых я<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>с</strong><strong>а</strong>мым <strong>о</strong>бширным и, п<strong>о</strong>чти н<strong>а</strong><strong>в</strong>ерн<strong>о</strong>е,<br />

пре<strong>в</strong><strong>о</strong><strong>с</strong>х<strong>о</strong>дит п<strong>о</strong> чи<strong>с</strong>лу <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong><strong>с</strong>ех других жи<strong>в</strong><strong>о</strong>тных<br />

<strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых. Чи<strong>с</strong>л<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных п<strong>о</strong><br />

н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя ф<strong>о</strong>рм н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых приближ<strong>а</strong>ет<strong>с</strong>я<br />

<strong>к</strong> милли<strong>о</strong>ну <strong>в</strong>ид<strong>о</strong><strong>в</strong>, причем д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р, п<strong>о</strong><br />

мере изучения ф<strong>а</strong>ун не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>а</strong>й<strong>о</strong>н<strong>о</strong><strong>в</strong> Земли,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н, чи<strong>с</strong>л<strong>о</strong> <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>емых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ет <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ть.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет и<strong>с</strong><strong>к</strong>лючительный<br />

интере<strong>с</strong> не т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong> би<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>им<br />

<strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong><strong>а</strong>м этих жи<strong>в</strong><strong>о</strong>тных, н<strong>о</strong> и п<strong>о</strong> п<strong>о</strong>льзе или<br />

<strong>в</strong>реду, прин<strong>о</strong><strong>с</strong>имым ими чел<strong>о</strong><strong>в</strong>е<strong>к</strong>у и ег<strong>о</strong> х<strong>о</strong>зяй<strong>с</strong>т<strong>в</strong>у.<br />

В <strong>с</strong>реде н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых е<strong>с</strong>ть мн<strong>о</strong>же<strong>с</strong>т<strong>в</strong><strong>о</strong> ф<strong>о</strong>рм<br />

<strong>с</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й нер<strong>в</strong>н<strong>о</strong>й деятельн<strong>о</strong><strong>с</strong>тью, <strong>в</strong>ыр<strong>а</strong>ж<strong>а</strong>ющей<strong>с</strong>я<br />

<strong>в</strong> <strong>с</strong>л<strong>о</strong>жных пр<strong>о</strong>я<strong>в</strong>лениях з<strong>а</strong>б<strong>о</strong>т<br />

<strong>о</strong> п<strong>о</strong>т<strong>о</strong>м<strong>с</strong>т<strong>в</strong>е и <strong>в</strong> н<strong>а</strong>личии <strong>о</strong>бще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бр<strong>а</strong>з<strong>а</strong><br />

жизни. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>к</strong> <strong>о</strong><strong>к</strong>руж<strong>а</strong>ющей <strong>с</strong>реде.<br />

Величин<strong>а</strong> тел<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> <strong>о</strong>чень изменчи<strong>в</strong><strong>а</strong>:<br />

<strong>о</strong>н<strong>а</strong> <strong>к</strong><strong>о</strong>леблет<strong>с</strong>я <strong>о</strong>т 200—250 мм (у р<strong>а</strong>зличных<br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их п<strong>а</strong>л<strong>о</strong>чни<strong>к</strong><strong>о</strong><strong>в</strong>, Phasmatodea)<br />

д<strong>о</strong> 0,25 мм (у жуч<strong>к</strong><strong>о</strong><strong>в</strong> из <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Ptiliidae)<br />

и д<strong>а</strong>же д<strong>о</strong> еще меньших р<strong>а</strong>змер<strong>о</strong><strong>в</strong> (у переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых<br />

яйцеед<strong>о</strong><strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Mymaridae).<br />

Р<strong>а</strong>зличные <strong>о</strong>тряды н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

цруг <strong>о</strong>т друг<strong>а</strong> п<strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гии, инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>му<br />

р<strong>а</strong>з<strong>в</strong>итию и э<strong>к</strong><strong>о</strong>л<strong>о</strong>гии. Орг<strong>а</strong>низ<strong>а</strong>ция и<br />

би<strong>о</strong>л<strong>о</strong>гия <strong>с</strong><strong>о</strong><strong>в</strong>ременных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых лег<strong>к</strong><strong>о</strong> м<strong>о</strong>гут<br />

быть изучены. С<strong>о</strong><strong>в</strong><strong>с</strong>ем ин<strong>о</strong>е н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я <strong>в</strong><br />

пр<strong>а</strong><strong>к</strong>ти<strong>к</strong>е п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний:<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong><strong>с</strong>егд<strong>а</strong> фр<strong>а</strong>гмент<strong>а</strong>рны,<br />

<strong>а</strong> е<strong>с</strong>ли и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> целых<br />

тел н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, т<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

н<strong>а</strong><strong>с</strong>т<strong>о</strong>ль<strong>к</strong><strong>о</strong> не<strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong>, чт<strong>о</strong> не д<strong>а</strong>ют <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ть гл<strong>а</strong><strong>в</strong>нейшие <strong>с</strong>и<strong>с</strong>темы <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>,<br />

<strong>с</strong>тр<strong>о</strong>ение <strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong>иб<strong>о</strong>лее <strong>в</strong><strong>а</strong>жн<strong>о</strong> для<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>зн<strong>а</strong><strong>в</strong><strong>а</strong>ния их <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений.<br />

П<strong>о</strong>эт<strong>о</strong>му <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> неизбежн<strong>о</strong> неп<strong>о</strong>лны и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>нии и изучении лишь немн<strong>о</strong>гих<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>ших<strong>с</strong>я <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> тел<strong>а</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

именн<strong>о</strong> тех, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые у <strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм м<strong>а</strong>л<strong>о</strong><br />

и<strong>с</strong>п<strong>о</strong>льзуют<strong>с</strong>я <strong>в</strong> целях <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и и п<strong>о</strong>эт<strong>о</strong>му<br />

нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тны. Не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong> <strong>с</strong>ч<strong>а</strong><strong>с</strong>тли<strong>в</strong>ым<br />

<strong>о</strong>б<strong>с</strong>т<strong>о</strong>ятель<strong>с</strong>т<strong>в</strong><strong>о</strong>м я<strong>в</strong>ляет<strong>с</strong>я т<strong>о</strong>, чт<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>о</strong>чных<br />

п<strong>о</strong>р<strong>о</strong>д<strong>а</strong>х <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и <strong>к</strong>рылье<strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, т. е. тех <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>,<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е <strong>к</strong><strong>о</strong>т<strong>о</strong>рых мы р<strong>а</strong>злич<strong>а</strong>ем <strong>к</strong><strong>а</strong>тег<strong>о</strong>рии<br />

<strong>с</strong>и<strong>с</strong>темы <strong>с</strong><strong>о</strong><strong>в</strong>ременных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Неп<strong>о</strong>лн<strong>о</strong>т<strong>а</strong><br />

п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> м<strong>а</strong>тери<strong>а</strong>л<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong><br />

и<strong>с</strong><strong>к</strong>уп<strong>а</strong>ет<strong>с</strong>я тем, чт<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я именн<strong>о</strong><br />

<strong>к</strong>рылья — <strong>о</strong>дни из н<strong>а</strong>иб<strong>о</strong>лее <strong>в</strong><strong>а</strong>жных для<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> тел<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Вме<strong>с</strong>те<br />

<strong>с</strong> тем эт<strong>о</strong> <strong>о</strong>пределяет <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зие изучения и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е <strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong>й<br />

мере <strong>с</strong><strong>в</strong><strong>о</strong>дит<strong>с</strong>я <strong>к</strong> изучению <strong>о</strong>дних<br />

<strong>к</strong>рылье<strong>в</strong>, <strong>к</strong> дет<strong>а</strong>льн<strong>о</strong>му <strong>о</strong><strong>с</strong><strong>в</strong>ещению р<strong>а</strong>зличных<br />

ч<strong>а</strong><strong>с</strong>тей этих <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. П<strong>о</strong>эт<strong>о</strong>му п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>г<br />

д<strong>о</strong>лжен уметь р<strong>а</strong>злич<strong>а</strong>ть группы н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не п<strong>о</strong> <strong>с</strong><strong>а</strong>мым х<strong>а</strong>р<strong>а</strong><strong>к</strong>терным для них призн<strong>а</strong><strong>к</strong><strong>а</strong>м,<br />

н<strong>а</strong>пример, п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong><br />

или пре<strong>в</strong>р<strong>а</strong>щению, <strong>а</strong> <strong>о</strong>бяз<strong>а</strong>тельн<strong>о</strong> п<strong>о</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

<strong>к</strong>рылье<strong>в</strong>, их ф<strong>о</strong>рме и р<strong>а</strong>змер<strong>а</strong>м.<br />

Крылья н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ены д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

<strong>с</strong>л<strong>о</strong>жн<strong>о</strong> и,— чт<strong>о</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong><strong>а</strong>жн<strong>о</strong>,— <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зличн<strong>о</strong><br />

у р<strong>а</strong>зных групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Не т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>о</strong>тряды н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых лег<strong>к</strong><strong>о</strong> р<strong>а</strong>злич<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ению<br />

<strong>к</strong>рылье<strong>в</strong>, н<strong>о</strong> и б<strong>о</strong>лее низ<strong>к</strong>ие <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие<br />

<strong>к</strong><strong>а</strong>тег<strong>о</strong>рии <strong>в</strong> гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>м б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е <strong>с</strong>луч<strong>а</strong>е<strong>в</strong><br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я <strong>с</strong>тр<strong>о</strong>ением <strong>к</strong>рылье<strong>в</strong>.<br />

Эт<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>о</strong>д<strong>а</strong> уни<strong>в</strong>ер<strong>с</strong><strong>а</strong>льн<strong>о</strong>е зн<strong>а</strong>чение <strong>к</strong>рылье<strong>в</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых для р<strong>а</strong>зличения <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>а</strong>тег<strong>о</strong>рий з<strong>а</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>о</strong>бр<strong>а</strong>тить <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>е<br />

<strong>в</strong>ним<strong>а</strong>ние при зн<strong>а</strong><strong>к</strong><strong>о</strong>м<strong>с</strong>т<strong>в</strong>е <strong>с</strong> гл<strong>а</strong><strong>в</strong>нейшими и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емыми<br />

групп<strong>а</strong>ми н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ение<br />

их <strong>к</strong>рылье<strong>в</strong>.<br />

Крылья н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых я<strong>в</strong>ляют<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>быми<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми, <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>с</strong>пин<strong>к</strong>и грудных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong><strong>о</strong>единены<br />

<strong>с</strong> тел<strong>о</strong>м п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong>членения,<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляющег<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ерш<strong>а</strong>ть <strong>о</strong>пределенные д<strong>в</strong>ижения—<br />

<strong>в</strong>зм<strong>а</strong>хи (ри<strong>с</strong>. 17—19). Крылья р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

лишь н<strong>а</strong> д<strong>в</strong>ух <strong>с</strong>егмент<strong>а</strong>х грудн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>тдел<strong>а</strong>: н<strong>а</strong> <strong>с</strong>реднегруди и н<strong>а</strong> з<strong>а</strong>днегруди. Из<strong>в</strong>е<strong>с</strong>тны<br />

п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

31<br />

http://jurassic.ru/

CuP<br />

17. Р<strong>а</strong>п<strong>в</strong>рр<strong>а</strong> sp.; <strong>с</strong>тр<strong>о</strong>ение грудн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тдел<strong>а</strong>, <strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у; <strong>с</strong>ле<strong>в</strong><strong>а</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong><strong>о</strong>:<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, Гпереднегрудь ^<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями переднегрудных<br />

н<strong>о</strong>г; <strong>с</strong>реднегрудь? <strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями' <strong>с</strong>реднегрудных н<strong>о</strong>г и передних<br />

<strong>к</strong>рылье<strong>в</strong>; з<strong>а</strong>днегрудь <strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями з<strong>а</strong>днегрудных н<strong>о</strong>г и з<strong>а</strong>дних<br />

<strong>к</strong>рылье<strong>в</strong>; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние брюш<strong>к</strong><strong>а</strong> (Ш<strong>в</strong><strong>а</strong>н<strong>в</strong>ич, 1949). 18. Perla sp.;<br />

р<strong>а</strong><strong>с</strong>членение тел<strong>а</strong>; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>; переднегрудь'<strong>с</strong> переднегруднымн н<strong>о</strong>г<strong>а</strong>­<br />

Palaeodictyoptera и Paraplecoptera), у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

неб<strong>о</strong>льшие <strong>к</strong>рыл<strong>о</strong><strong>в</strong>идные прид<strong>а</strong>т<strong>к</strong>и при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли<br />

и н<strong>а</strong> переднегрудн<strong>о</strong>м <strong>о</strong>тделе; эти<br />

прид<strong>а</strong>т<strong>к</strong>и не были н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящими <strong>к</strong>рыльями, т<strong>а</strong><strong>к</strong><br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>ни не были <strong>с</strong><strong>о</strong>единены <strong>с</strong> <strong>с</strong>егмент<strong>о</strong>м <strong>о</strong><strong>с</strong><strong>о</strong>бым<br />

<strong>с</strong><strong>о</strong>членением и не <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ли п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><strong>с</strong>тью.<br />

Крылья, <strong>к</strong>р<strong>о</strong>ме <strong>с</strong><strong>в</strong><strong>о</strong>ей <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й фун<strong>к</strong>ции<br />

^<strong>о</strong>рг<strong>а</strong>н<strong>а</strong> <strong>в</strong>зм<strong>а</strong>х<strong>а</strong>, <strong>в</strong>ып<strong>о</strong>лняют т<strong>а</strong><strong>к</strong>же и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

другие, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле и фун<strong>к</strong>цию п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

.<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>. К<strong>а</strong><strong>к</strong> у<strong>в</strong>идим д<strong>а</strong>льше, у<strong>к</strong><strong>а</strong>з<strong>а</strong>нн<strong>о</strong>е <strong>о</strong>б<strong>с</strong>т<strong>о</strong>ятель<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

имеет <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>е зн<strong>а</strong>чение при<br />

.<strong>о</strong>пен<strong>к</strong>е р<strong>о</strong>ли этих <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> для целей <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и:<br />

изменчи<strong>в</strong><strong>о</strong><strong>с</strong>ть жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рылье<strong>в</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я,<br />

з<strong>а</strong><strong>в</strong>и<strong>с</strong>ит <strong>о</strong>т т<strong>о</strong>г<strong>о</strong>, я<strong>в</strong>ляют<strong>с</strong>я ли <strong>к</strong>ры-<br />

,лья <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми <strong>в</strong>зм<strong>а</strong>х<strong>а</strong> или п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ными <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми.<br />

Ф<strong>о</strong>рм<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> р<strong>а</strong>зличн<strong>а</strong> у р<strong>а</strong>зных групп<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Ин<strong>о</strong>гд<strong>а</strong> <strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> удлинены'<br />

и пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> уз<strong>к</strong>ие пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong>, н<strong>а</strong>пример,<br />

у <strong>с</strong>тре<strong>к</strong><strong>о</strong>з и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых других дре<strong>в</strong>не<strong>к</strong>рылых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (Megasecoptera), у пе<strong>к</strong><strong>о</strong>-<br />

-т<strong>о</strong>рых <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых, <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц, д<strong>в</strong>у<strong>к</strong>ры­<br />

Ри<strong>с</strong>. 17—19. Стр<strong>о</strong>ение тел<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong><br />

32<br />

http://jurassic.ru/<br />

ми; <strong>с</strong>реднегрудь <strong>с</strong><strong>о</strong> <strong>с</strong>реднегрудными н<strong>о</strong>г<strong>а</strong>ми и передними <strong>к</strong>рыльями;<br />

з<strong>а</strong>днегрудь <strong>с</strong> з<strong>а</strong>днегрудными н<strong>о</strong>г<strong>а</strong>ми и з<strong>а</strong>дними <strong>к</strong>рыльями;<br />

члени<strong>с</strong>т<strong>о</strong>е брюш<strong>к</strong><strong>о</strong> <strong>с</strong> цер<strong>к</strong><strong>а</strong>ми (Handlirsch, 1925). 19. Стр<strong>о</strong>ение<br />

<strong>к</strong>рыл<strong>а</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> груди н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> (<strong>с</strong>хем<strong>а</strong>); <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чения <strong>с</strong>м. <strong>в</strong><br />

те<strong>к</strong><strong>с</strong>те (Ш<strong>в</strong><strong>а</strong>н<strong>в</strong>ич, 1949).<br />

лых. Н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бычн<strong>а</strong>я для <strong>к</strong>рылье<strong>в</strong> ф<strong>о</strong>рм<strong>а</strong><br />

непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>г<strong>о</strong> треуг<strong>о</strong>льни<strong>к</strong><strong>а</strong> или п<strong>а</strong>р<strong>а</strong>ллел<strong>о</strong>гр<strong>а</strong>мм<strong>а</strong><br />

<strong>в</strong><strong>с</strong>треч<strong>а</strong>ет<strong>с</strong>я у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Ф<strong>о</strong>рм<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong>ими<br />

п<strong>о</strong>требн<strong>о</strong><strong>с</strong>тями <strong>а</strong>эр<strong>о</strong>дин<strong>а</strong>ми<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong><br />

при ег<strong>о</strong> <strong>в</strong>зм<strong>а</strong>хе, т<strong>а</strong><strong>к</strong> и при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блением<br />

<strong>с</strong>тр<strong>о</strong>ения <strong>к</strong>рыл<strong>а</strong> <strong>к</strong> <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>нию при п<strong>о</strong><strong>к</strong><strong>о</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>.<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным (<strong>с</strong>м. ниже); перед <strong>в</strong>ершин<strong>о</strong>й <strong>к</strong>рыл<strong>а</strong><br />

эт<strong>о</strong>т <strong>к</strong>р<strong>а</strong>й з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бляя <strong>в</strong>ершину,<br />

или <strong>а</strong>пе<strong>к</strong><strong>с</strong> (apex). З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>лее <strong>в</strong>ыпу<strong>к</strong>лый, чем передний,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделен н<strong>а</strong> термин<strong>а</strong>льный<br />

(termen) и <strong>а</strong>н<strong>а</strong>льный (analis) <strong>к</strong>р<strong>а</strong>я, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

р<strong>а</strong>зделяют<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>бым угл<strong>о</strong>м, или т<strong>о</strong>рну<strong>с</strong><strong>о</strong>м (tornus).<br />

При пре<strong>в</strong>р<strong>а</strong>щении <strong>к</strong>рылье<strong>в</strong> <strong>в</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ные<br />

<strong>о</strong>рг<strong>а</strong>ны, <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>я<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> п<strong>о</strong><strong>к</strong>рыш<strong>к</strong>и или элитры,<br />

ф<strong>о</strong>рм<strong>а</strong> их рез<strong>к</strong><strong>о</strong> изменяет<strong>с</strong>я и <strong>о</strong>ни приним<strong>а</strong>ют<br />

<strong>в</strong>ид <strong>в</strong>ыпу<strong>к</strong>лых чешуе<strong>к</strong> или .п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><strong>к</strong>р<strong>а</strong>йних<br />

пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>.

Н<strong>а</strong>иб<strong>о</strong>лее <strong>в</strong><strong>а</strong>жн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть —их <strong>о</strong>п<strong>о</strong>рный<br />

<strong>с</strong><strong>к</strong>елет — т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>о</strong>е жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние: <strong>с</strong>и<strong>с</strong>тем<strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я труб<strong>о</strong><strong>к</strong>, н<strong>а</strong>п<strong>о</strong>лненных у жи<strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

гем<strong>о</strong>лимф<strong>о</strong>й. Эти труб<strong>к</strong>и-жил<strong>к</strong>и р<strong>а</strong>зличн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е <strong>к</strong>рыл<strong>а</strong>; их<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение <strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>лен<strong>о</strong> не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>тью<br />

мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> у<strong>к</strong>репления р<strong>а</strong>зличных уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>к</strong>рыл<strong>о</strong><strong>в</strong><strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых при<br />

<strong>в</strong>зм<strong>а</strong>хе <strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ют <strong>с</strong>ильные н<strong>а</strong>пряжения. Для<br />

н<strong>а</strong>ших целей не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong> лишь <strong>о</strong>тметить н<strong>а</strong>блюд<strong>а</strong>ющую<strong>с</strong>я<br />

<strong>а</strong><strong>с</strong>имметрию <strong>в</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жении<br />

жил<strong>о</strong><strong>к</strong> н<strong>а</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е <strong>к</strong>рыл<strong>а</strong>. Жил<strong>к</strong>и н<strong>а</strong> передней<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> близ ег<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я,<br />

леж<strong>а</strong>т <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем ин<strong>а</strong>че, чем н<strong>а</strong> з<strong>а</strong>дней. У передней<br />

<strong>к</strong>р<strong>о</strong>м<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong> жил<strong>к</strong>и <strong>о</strong>чень ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я,<br />

ин<strong>о</strong>гд<strong>а</strong> ут<strong>о</strong>лщ<strong>а</strong>ют<strong>с</strong>я, чт<strong>о</strong> <strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ли<strong>в</strong><strong>а</strong>ет<br />

<strong>с</strong>ильн<strong>о</strong>е у<strong>к</strong>репление переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нению<br />

<strong>с</strong> з<strong>а</strong>дним. Эт<strong>о</strong>т пр<strong>о</strong>це<strong>с</strong><strong>с</strong> н<strong>о</strong><strong>с</strong>ит н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции, и ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итие <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ет<br />

б<strong>о</strong>льшую или меньшую <strong>с</strong>тепень при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления<br />

<strong>к</strong>рыл<strong>а</strong> <strong>к</strong> <strong>с</strong>ильн<strong>о</strong>му и бы<strong>с</strong>тр<strong>о</strong>му п<strong>о</strong>лету.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции р<strong>а</strong>зличен у р<strong>а</strong>зных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Отдельные жил<strong>к</strong>и <strong>о</strong>бъединяют<strong>с</strong>я <strong>в</strong> <strong>о</strong>пределенные<br />

<strong>с</strong>и<strong>с</strong>темы жил<strong>о</strong><strong>к</strong>, н<strong>о</strong><strong>с</strong>ящие <strong>о</strong><strong>с</strong><strong>о</strong>бые н<strong>а</strong>з<strong>в</strong><strong>а</strong>ния<br />

(ри<strong>с</strong>. 19). Н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>, или <strong>к</strong> <strong>о</strong><strong>с</strong>т <strong>а</strong> (С). В гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>м б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е<br />

<strong>с</strong>луч<strong>а</strong>е<strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> имеет<strong>с</strong>я <strong>в</strong><br />

един<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>м чи<strong>с</strong>ле; п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>н<strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>о</strong>бх<strong>о</strong>дит <strong>в</strong><strong>с</strong>е <strong>к</strong>рыл<strong>о</strong> <strong>к</strong>руг<strong>о</strong>м, д<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ег<strong>о</strong><br />

ч<strong>а</strong><strong>с</strong>ти. Ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти передней<br />

<strong>к</strong>р<strong>о</strong>м<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong>переди <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong>рыл<strong>а</strong>, н<strong>о</strong><strong>с</strong>ящее н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние<br />

пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и не<strong>с</strong>ущее <strong>о</strong>дну<br />

или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льных жил<strong>о</strong><strong>к</strong><br />

(PC). Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льные жил<strong>к</strong>и р<strong>а</strong>з<strong>в</strong>иты у <strong>о</strong>чень<br />

немн<strong>о</strong>гих групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — у дре<strong>в</strong>них <strong>с</strong>тре<strong>к</strong><strong>о</strong>з,<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых прям<strong>о</strong><strong>к</strong>рылых. Сз<strong>а</strong>ди <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>, или <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong> (SC), <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

у примити<strong>в</strong>ных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>, р<strong>а</strong>зличных<br />

прям<strong>о</strong><strong>к</strong>рылых, <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц и др.) и,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, я<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, не<strong>в</strong>ет<strong>в</strong>ящей<strong>с</strong>я<br />

у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> других групп. Следующ<strong>а</strong>я групп<strong>а</strong><br />

жил<strong>о</strong><strong>к</strong> н<strong>о</strong><strong>с</strong>ит н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние р<strong>а</strong>ди<strong>а</strong>льных (R).<br />

Эти жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>бширную <strong>с</strong>и<strong>с</strong>тему мн<strong>о</strong>г<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>в</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong>ним<strong>а</strong>я<br />

б<strong>о</strong>льшую ч<strong>а</strong><strong>с</strong>ть передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>в</strong><strong>с</strong>ей пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и<br />

<strong>к</strong>рыл<strong>а</strong>. Ч<strong>а</strong><strong>с</strong>ть р<strong>а</strong>ди<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

<strong>с</strong>з<strong>а</strong>ди <strong>о</strong>т их передней <strong>в</strong>ет<strong>в</strong>и и <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ющих<br />

<strong>о</strong>бщим <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м, ин<strong>о</strong>гд<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленным,<br />

н<strong>о</strong><strong>с</strong>ит н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>е<strong>к</strong>т<strong>о</strong>р<strong>а</strong> (RS). Следующей <strong>с</strong>и<strong>с</strong>тем<strong>о</strong>й жил<strong>о</strong><strong>к</strong><br />

я<strong>в</strong>ляют<strong>с</strong>я меди<strong>а</strong>льньге жил<strong>к</strong>и, ин<strong>о</strong>гд<strong>а</strong><br />

н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емые <strong>в</strong> цел<strong>о</strong>м меди<strong>а</strong>н<strong>о</strong>й (М). Обычн<strong>о</strong><br />

эти жил<strong>к</strong>и п<strong>о</strong>др<strong>а</strong>зделяют<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

— передний и з<strong>а</strong>дний, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>б<strong>о</strong>-<br />

3 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

зн<strong>а</strong>ч<strong>а</strong>емые <strong>к</strong><strong>а</strong><strong>к</strong> передняя (МА) и з<strong>а</strong>дняя (MP)<br />

меди<strong>а</strong>льные жил<strong>к</strong>и. Эти жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

примерн<strong>о</strong> <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> и делят<strong>с</strong>я н<strong>а</strong><br />

мн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей. З<strong>а</strong>дние <strong>с</strong>и<strong>с</strong>темы жил<strong>о</strong><strong>к</strong> н<strong>о</strong><strong>с</strong>ят н<strong>а</strong>з<strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>убит<strong>а</strong>льных (Си) и <strong>а</strong>н<strong>а</strong>льных<br />

(А) жил<strong>о</strong><strong>к</strong>. У<strong>к</strong><strong>а</strong>з<strong>а</strong>нные жил<strong>к</strong>и, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые т<strong>а</strong><strong>к</strong>же<br />

м<strong>о</strong>гут быть мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>ют<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> CuA, CuP, Аь А2, A3<br />

и т. д. П<strong>о</strong>з<strong>а</strong>ди <strong>в</strong><strong>с</strong>ех <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных <strong>с</strong>и<strong>с</strong>тем жил<strong>о</strong><strong>к</strong> н<strong>а</strong><br />

<strong>к</strong>рыльях н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылы'х н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>бычн<strong>о</strong> при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

еще <strong>о</strong><strong>с</strong><strong>о</strong>бые юг<strong>а</strong>льные жил<strong>к</strong>и<br />

(Ju).<br />

Пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>а</strong> переп<strong>о</strong>н<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong>, лишенные<br />

жил<strong>о</strong><strong>к</strong>, н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я ячей<strong>к</strong><strong>а</strong>ми, <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>емыми,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, п<strong>о</strong> жил<strong>к</strong>е, <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ющей<br />

ячей<strong>к</strong>у <strong>с</strong>переди, реже — п<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жению.<br />

Между <strong>о</strong>пи<strong>с</strong><strong>а</strong>нными <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ми жил<strong>о</strong><strong>к</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

еще т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е имеют <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем ин<strong>о</strong>е<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение, шляя<strong>с</strong>ь пр<strong>о</strong>из<strong>в</strong><strong>о</strong>дным <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>етч<strong>а</strong>т<strong>о</strong>г<strong>о</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния пер<strong>в</strong>ичных <strong>к</strong>рылье<strong>в</strong><br />

дре<strong>в</strong>них н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>о</strong>г<strong>о</strong> <strong>а</strong> р х ед<br />

и <strong>к</strong> т и я (ри<strong>с</strong>. 41).<br />

Археди<strong>к</strong>тий <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ем пер<strong>в</strong>ичн<strong>о</strong>м <strong>в</strong>иде м<strong>о</strong>жн<strong>о</strong><br />

н<strong>а</strong>блюд<strong>а</strong>ть <strong>в</strong> <strong>к</strong>рыльях дре<strong>в</strong>них <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных<br />

п<strong>а</strong>ле<strong>о</strong>ди<strong>к</strong>ти<strong>о</strong>птер, прям<strong>о</strong><strong>к</strong>рылых и<br />

т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>; <strong>в</strong> д<strong>а</strong>льнейшей э<strong>в</strong><strong>о</strong>люции жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>рылье<strong>в</strong> <strong>а</strong>рхеди<strong>к</strong>тий пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> п<strong>о</strong>перечные<br />

жил<strong>к</strong>и.<br />

О<strong>с</strong><strong>о</strong>б<strong>о</strong>е зн<strong>а</strong>чение имеет п<strong>о</strong>л<strong>о</strong>жение <strong>к</strong>рылье<strong>в</strong><br />

<strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> ег<strong>о</strong> тел<strong>а</strong>.<br />

Эт<strong>а</strong> черт<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ет при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бление<br />

<strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong>му п<strong>о</strong>лету, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

и при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бление <strong>к</strong> <strong>о</strong>бит<strong>а</strong>нию <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>я<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong><br />

у<strong>к</strong>рытых, те<strong>с</strong>ных ме<strong>с</strong>т<strong>а</strong>х, <strong>с</strong> друг<strong>о</strong>й. У дре<strong>в</strong>не<strong>к</strong>рылых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> их и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>м р<strong>а</strong>з<strong>в</strong>итии<br />

<strong>о</strong>чень р<strong>а</strong>н<strong>о</strong> <strong>с</strong>т<strong>а</strong>ли <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ть<strong>с</strong>я лет<strong>а</strong>тельные<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления: <strong>к</strong>рылья их у<strong>в</strong>еличили<strong>с</strong>ь,<br />

и <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ть <strong>к</strong> бы<strong>с</strong>тр<strong>о</strong>му <strong>в</strong>зм<strong>а</strong>ху <strong>с</strong>т<strong>а</strong>л<strong>а</strong><br />

лег<strong>к</strong><strong>о</strong> <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>им<strong>о</strong>й <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие п<strong>о</strong><strong>с</strong>т<strong>о</strong>янн<strong>о</strong>й<br />

мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>й г<strong>о</strong>т<strong>о</strong><strong>в</strong>н<strong>о</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> <strong>к</strong> д<strong>в</strong>ижению.<br />

Крылья у дре<strong>в</strong>не<strong>к</strong>рылых <strong>в</strong><strong>с</strong>егд<strong>а</strong> н<strong>а</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>терт<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, будучи <strong>о</strong>т<strong>в</strong>еденными<br />

<strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>о</strong>т тел<strong>а</strong>. Ин<strong>о</strong>е н<strong>а</strong>блюд<strong>а</strong>л<strong>о</strong><strong>с</strong>ь у<br />

пер<strong>в</strong>ичных н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти<br />

у т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>; у них <strong>о</strong>чень р<strong>а</strong>н<strong>о</strong> <strong>с</strong>т<strong>а</strong>ли р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ть<strong>с</strong>я<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления <strong>к</strong> жизни <strong>в</strong> у<strong>к</strong>рытых,<br />

те<strong>с</strong>ньгх ме<strong>с</strong>т<strong>о</strong><strong>о</strong>бит<strong>а</strong>ниях, их <strong>к</strong>рылья при<strong>о</strong>брели<br />

<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ть <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong>д<strong>о</strong>ль тел<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>или<strong>с</strong>ь<br />

<strong>с</strong>л<strong>о</strong>жные <strong>с</strong><strong>о</strong>членения <strong>к</strong>рылье<strong>в</strong> <strong>с</strong> грудным<br />

<strong>о</strong>тдел<strong>о</strong>м, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енный <strong>с</strong><strong>к</strong>елетный и<br />

мышечный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т; н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е п<strong>о</strong>лучил<strong>о</strong> <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ть<br />

у<strong>к</strong>ры<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>я<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> те<strong>с</strong>ных<br />

пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>а</strong>х (щелях п<strong>о</strong>ч<strong>в</strong>ы, п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями,<br />

между р<strong>а</strong><strong>с</strong>тениями).<br />

З<strong>а</strong><strong>к</strong>люч<strong>а</strong>я р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трение <strong>к</strong>рылье<strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>с</strong>ледует еще у<strong>к</strong><strong>а</strong>з<strong>а</strong>ть н<strong>а</strong> их р<strong>а</strong>змеры и<br />

п<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> тел<strong>а</strong>. У дре<strong>в</strong>-<br />

33<br />

http://jurassic.ru/

них прям<strong>о</strong><strong>к</strong>рылых <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>, у т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

примити<strong>в</strong>ных дре<strong>в</strong>не<strong>к</strong>рылых длин<strong>а</strong><br />

<strong>к</strong>рылье<strong>в</strong> примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> длине тел<strong>а</strong>. Вме<strong>с</strong>те<br />

<strong>с</strong> тем у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> дре<strong>в</strong>не<strong>к</strong>рылых, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

прям<strong>о</strong><strong>к</strong>рылых и др. р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>л<strong>а</strong><strong>с</strong>ь длинн<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ть:<br />

их <strong>к</strong>рылья <strong>с</strong>т<strong>а</strong>ли пре<strong>в</strong>ыш<strong>а</strong>ть длину<br />

тел<strong>а</strong>. Ин<strong>о</strong>е н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ление э<strong>в</strong><strong>о</strong>люции <strong>к</strong>рылье<strong>в</strong><br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>л<strong>о</strong><strong>с</strong>ь .<strong>в</strong> уменьшении их р<strong>а</strong>змер<strong>о</strong><strong>в</strong> и <strong>в</strong><br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong>м ит<strong>о</strong>ге — <strong>в</strong> п<strong>о</strong>лн<strong>о</strong>й их реду<strong>к</strong>ции, <strong>в</strong><br />

р<strong>а</strong>з<strong>в</strong>итии бе<strong>с</strong><strong>к</strong>ры'л<strong>о</strong><strong>с</strong>ти. В эт<strong>о</strong>м <strong>с</strong>луч<strong>а</strong>е <strong>с</strong>ледует<br />

р<strong>а</strong>злич<strong>а</strong>ть <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щение р<strong>а</strong>змер<strong>о</strong><strong>в</strong> <strong>к</strong>рылье<strong>в</strong>, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нн<strong>о</strong>е<br />

<strong>с</strong> реду<strong>к</strong>цией лет<strong>а</strong>тельн<strong>о</strong>й фун<strong>к</strong>ции и<br />

р<strong>а</strong>з<strong>в</strong>итием бе<strong>с</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ти, <strong>о</strong>т <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щения длины<br />

<strong>к</strong>рылье<strong>в</strong> <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие у<strong>с</strong>иления <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции, <strong>с</strong>илы<br />

и <strong>с</strong><strong>к</strong><strong>о</strong>р<strong>о</strong><strong>с</strong>ти <strong>в</strong>зм<strong>а</strong>х<strong>а</strong>; п<strong>о</strong><strong>с</strong>леднее н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

у н<strong>а</strong>иб<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нных, м<strong>о</strong>л<strong>о</strong>дьгх ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

групп — у д<strong>в</strong>у<strong>к</strong>рылых и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у<br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых.<br />

Р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трение и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емы'х <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>ших<strong>с</strong>я цели<strong>к</strong><strong>о</strong>м, п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет <strong>о</strong>пределить х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер лет<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong><br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong>, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти п<strong>о</strong>л<strong>о</strong>жение <strong>к</strong>рылье<strong>в</strong>, чт<strong>о</strong><br />

д<strong>а</strong>ет <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ть у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ить прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ть<br />

<strong>к</strong> т<strong>о</strong>му или друг<strong>о</strong>му инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>у.<br />

Стр<strong>о</strong>ение б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> тел<strong>а</strong><br />

(ри<strong>с</strong>. 17—19) н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых имеет для п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong><strong>с</strong>тепенн<strong>о</strong>е<br />

зн<strong>а</strong>чение.<br />

Ф<strong>о</strong>рм<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мы'х р<strong>а</strong>зличн<strong>а</strong>, н<strong>о</strong> <strong>в</strong><br />

<strong>о</strong>бщем приближ<strong>а</strong>ет<strong>с</strong>я <strong>к</strong> неп<strong>о</strong>лн<strong>о</strong>му ш<strong>а</strong>ру или<br />

п<strong>о</strong>луш<strong>а</strong>рию. Антенны, гл<strong>а</strong>з<strong>а</strong> и р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны<br />

п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ены <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. Их <strong>с</strong>тр<strong>о</strong>ение<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> для <strong>о</strong>тдельных <strong>о</strong>тряд<strong>о</strong><strong>в</strong> и групп.<br />

Грудн<strong>о</strong>й <strong>о</strong>тдел, <strong>к</strong>р<strong>о</strong>ме <strong>к</strong>рылье<strong>в</strong>, не<strong>с</strong>ет н<strong>о</strong>ги,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из м<strong>а</strong><strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>о</strong>г<strong>о</strong>, <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> — т<strong>а</strong>зи<strong>к</strong><strong>а</strong> (<strong>с</strong><strong>о</strong>х<strong>а</strong>), п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть при<strong>к</strong>репляет<strong>с</strong>я<br />

<strong>к</strong> пле<strong>в</strong>р<strong>а</strong>м грудн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>. К т<strong>а</strong>зи<strong>к</strong>у причленяет<strong>с</strong>я<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий <strong>в</strong>ертлуг, или тр<strong>о</strong>х<strong>а</strong>нтер (trochanter),<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из д<strong>в</strong>ух члени<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Он имеет <strong>в</strong><strong>а</strong>жн<strong>о</strong>е мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>е зн<strong>а</strong>чение, я<strong>в</strong>ляя<strong>с</strong>ь<br />

ме<strong>с</strong>т<strong>о</strong>м при<strong>к</strong>репления не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых мышц,<br />

упр<strong>а</strong><strong>в</strong>ляющих д<strong>в</strong>ижениями <strong>в</strong><strong>с</strong>ей н<strong>о</strong>ги <strong>в</strong> цел<strong>о</strong>м.<br />

Следующий, <strong>с</strong><strong>а</strong>мый м<strong>а</strong><strong>с</strong><strong>с</strong>и<strong>в</strong>ный и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> длинный<br />

<strong>с</strong>егмент н<strong>о</strong>ги — бедр<strong>о</strong> (femur), з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющий<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные мышцы, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляют<br />

д<strong>в</strong>ижения диет<strong>а</strong>льны'х <strong>о</strong>тдел<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти: г<strong>о</strong>лени<br />

(tibia) и л<strong>а</strong>п<strong>к</strong>и (tarsus). Г<strong>о</strong>лень примерн<strong>о</strong><br />

р<strong>а</strong><strong>в</strong>ных <strong>с</strong> бедр<strong>о</strong>м р<strong>а</strong>змер<strong>о</strong><strong>в</strong>, лишь п<strong>о</strong>чти<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> т<strong>о</strong>ньше ег<strong>о</strong>; <strong>о</strong>н<strong>а</strong> <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>а</strong> те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближ<strong>а</strong>ть<strong>с</strong>я<br />

<strong>с</strong> бедр<strong>о</strong>м, <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляя, н<strong>а</strong>ряду <strong>с</strong> д<strong>в</strong>ижениями<br />

<strong>в</strong><strong>с</strong>ей н<strong>о</strong>ги <strong>в</strong> цел<strong>о</strong>м, пр<strong>о</strong>це<strong>с</strong><strong>с</strong> ш<strong>а</strong>г<strong>а</strong>.<br />

Л<strong>а</strong>п<strong>к</strong><strong>а</strong> у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я;<br />

чи<strong>с</strong>л<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> ее <strong>к</strong><strong>о</strong>леблет<strong>с</strong>я <strong>в</strong> р<strong>а</strong>зных<br />

групп<strong>а</strong>х <strong>о</strong>т <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> д<strong>о</strong> пяти. П<strong>о</strong><strong>с</strong>ледний —<br />

ди<strong>с</strong>т<strong>а</strong>льный — члени<strong>к</strong> л<strong>а</strong>п<strong>к</strong>и не<strong>с</strong>ет <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и и<br />

р<strong>а</strong>зличные прид<strong>а</strong>т<strong>к</strong>и, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные н<strong>а</strong> <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>м<br />

<strong>с</strong><strong>к</strong>лерите, н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>о</strong>м прет<strong>а</strong>рз<strong>о</strong>м. К<strong>а</strong>ж-<br />

25 Б<br />

Ри<strong>с</strong>. 20—25. Личин<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

20. Личин<strong>к</strong>и <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых: А — Sisyra sp.; Б —'Raphldia<br />

sp. (Handlirsch, 1925). 21. Личин<strong>к</strong>и же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых: А — Catops<br />

sp.; Б — Silpha sp..(Handlirsch, 1925). 22. Личин<strong>к</strong><strong>а</strong> чешуе<strong>к</strong>рыл<strong>о</strong>г<strong>о</strong>:<br />

гу<strong>с</strong>ениц<strong>а</strong> бр<strong>а</strong>жни<strong>к</strong><strong>а</strong> Sphingidae (Handlirsch, 1925). 23. Личин<strong>к</strong><strong>а</strong><br />

ж<strong>а</strong>лящег<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong>г<strong>о</strong>: д<strong>о</strong>м<strong>а</strong>шняя пчел<strong>а</strong> (Handlirsch,<br />

1925). 24. Личин<strong>к</strong><strong>а</strong> длинн<strong>о</strong>у<strong>с</strong><strong>о</strong>г<strong>о</strong> д<strong>в</strong>у<strong>к</strong>рыл<strong>о</strong>г<strong>о</strong> из <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Bibionidae (Handlirsch, 1925). 25. Личин<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>у<strong>с</strong>ых д<strong>в</strong>у<strong>к</strong>рылых:<br />

А —личин<strong>к</strong><strong>а</strong> журч<strong>а</strong>л<strong>к</strong>и из р<strong>о</strong>д<strong>а</strong> Microdon, жи<strong>в</strong>ущ<strong>а</strong>я<br />

<strong>в</strong> мур<strong>а</strong><strong>в</strong>ейни<strong>к</strong><strong>а</strong>х; Б —личин<strong>к</strong><strong>а</strong> Muscidae (Handlirsch, 1925)<br />

34<br />

http://jurassic.ru/<br />

дый <strong>с</strong>егмент груди <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из тергит<strong>а</strong> (tergum<br />

или scutum), <strong>с</strong>тернит<strong>а</strong> (sternum) и пле<strong>в</strong>р (pleura).<br />

Тергит ин<strong>о</strong>гд<strong>а</strong> р<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> <strong>о</strong>тдельные<br />

уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и <strong>с</strong>пин<strong>к</strong>и (notum, postnotum) и щит<strong>к</strong>и<br />

(praescutum, scutum, scutellum); <strong>в</strong> <strong>с</strong>луч<strong>а</strong>е<br />

б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития <strong>о</strong>тдельных <strong>с</strong>егмент<strong>о</strong><strong>в</strong> груди<br />

эти <strong>с</strong>егменты (<strong>с</strong>реднегрудь у д<strong>в</strong>у<strong>к</strong>рылых<br />

или переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых, з<strong>а</strong>днегрудь у же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых)<br />

<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют н<strong>а</strong>иб<strong>о</strong>лее <strong>с</strong>л<strong>о</strong>жным <strong>с</strong>тр<strong>о</strong>ением<br />

и т<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из мн<strong>о</strong>гих <strong>о</strong>тдельных<br />

пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>. Пле<strong>в</strong>ры грудн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> р<strong>а</strong>зделяют<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong>е гл<strong>а</strong><strong>в</strong>ные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и: переднюю<br />

— эпи<strong>с</strong>тернит и з<strong>а</strong>днюю — эпимерит. <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> имеют и <strong>в</strong>т<strong>о</strong>ричные п<strong>о</strong>др<strong>а</strong>зделения.<br />

Грудные <strong>с</strong>терниты т<strong>а</strong><strong>к</strong>же <strong>о</strong>бычн<strong>о</strong> делят<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>тдельные уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енных п<strong>о</strong> <strong>о</strong>бщему пл<strong>а</strong>ну; ч<strong>а</strong><strong>с</strong>ти

брюшн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> <strong>о</strong>чень р<strong>а</strong>зличны п<strong>о</strong> р<strong>а</strong>змер<strong>а</strong>м<br />

и п<strong>о</strong>л<strong>о</strong>жению. Обычн<strong>о</strong> н<strong>а</strong>иб<strong>о</strong>лее <strong>в</strong>ели<strong>к</strong>и<br />

тергиты <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, ин<strong>о</strong>гд<strong>а</strong> з<strong>а</strong>х<strong>о</strong>дящие д<strong>а</strong>ле<strong>к</strong><strong>о</strong><br />

н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong> или д<strong>а</strong>же н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну (у<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых и д<strong>в</strong>у<strong>к</strong>рылых).<br />

В других <strong>с</strong>луч<strong>а</strong>ях б<strong>о</strong>льш<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие п<strong>о</strong>луч<strong>а</strong>ют<br />

<strong>с</strong>терниты 1<br />

(у же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых). Н<strong>а</strong> <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены р<strong>а</strong>зличные п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые прид<strong>а</strong>т<strong>к</strong>и,<br />

<strong>с</strong>тр<strong>о</strong>ение <strong>к</strong><strong>о</strong>т<strong>о</strong>рых ч<strong>а</strong><strong>с</strong>т<strong>о</strong> имеет б<strong>о</strong>льш<strong>о</strong>е зн<strong>а</strong>чение<br />

для р<strong>а</strong>зличения <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>, реже — р<strong>о</strong>д<strong>о</strong><strong>в</strong> мн<strong>о</strong>гих н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>егменте брюш<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong><br />

п<strong>а</strong>р<strong>а</strong> цер<strong>о</strong><strong>к</strong>. У дре<strong>в</strong>них групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

цер<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>чень<br />

длинные (у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> дре<strong>в</strong>не<strong>к</strong>рылых, мн<strong>о</strong>гих<br />

<strong>о</strong>рт<strong>о</strong>птер<strong>о</strong>ид<strong>о</strong><strong>в</strong>); у других н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых цер<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, нечлени<strong>с</strong>тые; ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>ни нер<strong>а</strong>зличимы<br />

1<br />

.<br />

П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>о</strong>чень <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> и р<strong>а</strong>зличн<strong>о</strong> <strong>в</strong> р<strong>а</strong>зных групп<strong>а</strong>х;<br />

<strong>в</strong> цел<strong>о</strong>м гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>е б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я н<strong>а</strong>личием пре<strong>в</strong>р<strong>а</strong>щения,<br />

или мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>, з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющег<strong>о</strong><strong>с</strong>я <strong>в</strong> т<strong>о</strong>м, чт<strong>о</strong><br />

<strong>в</strong>ых<strong>о</strong>дящее из яйц<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я<br />

л<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции <strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>й, п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрел<strong>о</strong>й<br />

<strong>с</strong>т<strong>а</strong>дии (ри<strong>с</strong>. 20—25). Лишь не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

пер<strong>в</strong>ичн<strong>о</strong>бе<strong>с</strong><strong>к</strong>рылые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>в</strong>ых<strong>о</strong>дят<br />

из яиц <strong>о</strong>чень <strong>с</strong>х<strong>о</strong>дными п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению <strong>с</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лыми,<br />

<strong>о</strong>тлич<strong>а</strong>я<strong>с</strong>ь меньшими р<strong>а</strong>змер<strong>а</strong>ми и нер<strong>а</strong>з<strong>в</strong>итыми<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ыми <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми: у <strong>в</strong><strong>с</strong>ех <strong>к</strong>рыл<strong>а</strong>тых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong>ылупляющие<strong>с</strong>я из яйц<strong>а</strong> личин<strong>к</strong>и<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong> т<strong>о</strong>й или ин<strong>о</strong>й мере рез<strong>к</strong><strong>о</strong> <strong>о</strong>тличны<br />

<strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых.<br />

Р<strong>а</strong>злич<strong>а</strong>ют д<strong>в</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные ф<strong>о</strong>рмы пре<strong>в</strong>р<strong>а</strong>щения<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых; неп<strong>о</strong>лный мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з, или<br />

гемимет<strong>а</strong>б<strong>о</strong>лию, и п<strong>о</strong>лный мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з, или<br />

г<strong>о</strong>л<strong>о</strong>мет<strong>а</strong>б<strong>о</strong>лию. При гемимет<strong>а</strong>б<strong>о</strong>лии пер<strong>в</strong>ые<br />

<strong>с</strong>т<strong>а</strong>дии р<strong>а</strong>з<strong>в</strong>ития н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> имеют мн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бщег<strong>о</strong><br />

<strong>с</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лым н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым: <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>,<br />

три п<strong>а</strong>ры 1<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Р<strong>а</strong>з<strong>в</strong>итие при гемимет<strong>а</strong>б<strong>о</strong>лии<br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> у<strong>в</strong>еличении р<strong>а</strong>змер<strong>о</strong><strong>в</strong>,<br />

п<strong>о</strong>я<strong>в</strong>лении и р<strong>а</strong>з<strong>в</strong>итии з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong>рылье<strong>в</strong> и п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых<br />

прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>; <strong>с</strong> п<strong>о</strong><strong>с</strong>ледней линь<strong>к</strong><strong>о</strong>й личин<strong>к</strong><strong>а</strong>,<br />

т<strong>о</strong>чнее н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>а</strong>я нимф<strong>о</strong>й, пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>е — <strong>к</strong>рыл<strong>а</strong>т<strong>о</strong>е и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрел<strong>о</strong>е —<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е. Ф<strong>а</strong>зы п<strong>о</strong><strong>к</strong><strong>о</strong>я (<strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>и) при эт<strong>о</strong>м<br />

пре<strong>в</strong>р<strong>а</strong>щении не н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я, т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong><br />

<strong>о</strong>бычн<strong>о</strong> не н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я и р<strong>а</strong>з<strong>в</strong>ития <strong>к</strong><strong>а</strong><strong>к</strong>их-либ<strong>о</strong><br />

<strong>о</strong><strong>с</strong><strong>о</strong>бых личин<strong>о</strong>чных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> и при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений:<br />

пит<strong>а</strong>ние и э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия п<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льных<br />

<strong>с</strong>т<strong>а</strong>дий р<strong>а</strong>з<strong>в</strong>ития у н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> неп<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением<br />

не <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых.<br />

Гемимет<strong>а</strong>б<strong>о</strong>лия <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong>енн<strong>а</strong>, н<strong>а</strong>пример,<br />

прям<strong>о</strong><strong>к</strong>рылым и т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>а</strong>м.<br />

При п<strong>о</strong>лн<strong>о</strong>м пре<strong>в</strong>р<strong>а</strong>щении из яиц <strong>в</strong>ых<strong>о</strong>дят<br />

личин<strong>к</strong>и, рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ющие<strong>с</strong>я п<strong>о</strong> <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции<br />

<strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Т<strong>а</strong><strong>к</strong>ие личин<strong>к</strong>и <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют<br />

пр<strong>о</strong><strong>с</strong>тыми гл<strong>а</strong>з<strong>а</strong>ми, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> имеют д<strong>о</strong>п<strong>о</strong>л­<br />

нительные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти н<strong>а</strong> брюш<strong>к</strong>е или п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

лишены их. У личин<strong>о</strong><strong>к</strong> и им<strong>а</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мы'х<br />

<strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти р<strong>а</strong>зличны. Личин<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong><br />

п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют <strong>с</strong>пеци<strong>а</strong>льными<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блениями <strong>к</strong> <strong>о</strong>бит<strong>а</strong>нию <strong>в</strong> <strong>о</strong>пределенных<br />

у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях. При р<strong>а</strong>з<strong>в</strong>итии <strong>о</strong>ни пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бую, не приним<strong>а</strong>ющую пищи, <strong>с</strong>т<strong>а</strong>дию<br />

— <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>у, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>в</strong> д<strong>а</strong>льнейшем, п<strong>о</strong><strong>с</strong>ле<br />

пери<strong>о</strong>д<strong>а</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>я, пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>к</strong>рыл<strong>а</strong>т<strong>о</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е.<br />

П<strong>о</strong>лн<strong>о</strong>е пре<strong>в</strong>р<strong>а</strong>щение (г<strong>о</strong>л<strong>о</strong>мет<strong>а</strong>б<strong>о</strong>лия)<br />

<strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылым, д<strong>в</strong>у<strong>к</strong>рылым,<br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылым, <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылым и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым<br />

другим н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым.<br />

Эти гл<strong>а</strong><strong>в</strong>ные д<strong>в</strong><strong>а</strong> тип<strong>а</strong> пре<strong>в</strong>р<strong>а</strong>щения н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

имеют р<strong>а</strong>зличные ф<strong>о</strong>рмы и п<strong>о</strong>др<strong>а</strong>зделяют<strong>с</strong>я<br />

н<strong>а</strong> мн<strong>о</strong>гие <strong>в</strong>т<strong>о</strong>ричные типы, р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong> д<strong>а</strong>нн<strong>о</strong>м ру<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>д<strong>с</strong>т<strong>в</strong>е ед<strong>в</strong><strong>а</strong> ли<br />

не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong>. В<strong>а</strong>жн<strong>о</strong> <strong>о</strong>тметить <strong>с</strong>л<strong>о</strong>жн<strong>о</strong><strong>с</strong>ть п<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ития н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и глуб<strong>о</strong><strong>к</strong>ие<br />

р<strong>а</strong>зличия их неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых <strong>с</strong>т<strong>а</strong>дий, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>о</strong>чень <strong>о</strong>тлич<strong>а</strong>ющих<strong>с</strong>я не т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

ф<strong>о</strong>рм, н<strong>о</strong> д<strong>а</strong>же и <strong>о</strong>т <strong>о</strong>бщег<strong>о</strong> пл<strong>а</strong>н<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ения<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> <strong>в</strong><strong>о</strong><strong>о</strong>бще (н<strong>а</strong>пример, безн<strong>о</strong>гие личин<strong>к</strong>и<br />

жу<strong>к</strong><strong>о</strong><strong>в</strong>, переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых и д<strong>в</strong>у<strong>к</strong>рылых<br />

или безн<strong>о</strong>гие и безг<strong>о</strong>л<strong>о</strong><strong>в</strong>ые личин<strong>к</strong>и <strong>в</strong>ы<strong>с</strong>ших<br />

д<strong>в</strong>у<strong>к</strong>рылых).<br />

Принципы <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и<br />

Крупные п<strong>о</strong>др<strong>а</strong>зделения <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

(п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы, инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы, <strong>к</strong><strong>о</strong>г<strong>о</strong>рты, н<strong>а</strong>д<strong>о</strong>тряды<br />

и <strong>о</strong>тряды) х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я <strong>о</strong>пределенными<br />

черт<strong>а</strong>ми <strong>о</strong>нт<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития, м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гии<br />

и э<strong>к</strong><strong>о</strong>л<strong>о</strong>гии личин<strong>о</strong>чных и <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм.<br />

Для у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ления <strong>к</strong>рупных <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

п<strong>о</strong>др<strong>а</strong>зделений <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>е зн<strong>а</strong>чение имеют <strong>о</strong>бщий<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер пре<strong>в</strong>р<strong>а</strong>щения и э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых<br />

<strong>с</strong>т<strong>а</strong>дий, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>с</strong>тр<strong>о</strong>ения р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> и лет<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

ф<strong>о</strong>рм. П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

Pterygota, <strong>в</strong><strong>к</strong>люч<strong>а</strong>ет д<strong>в</strong><strong>а</strong> инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: Palaeoptera<br />

(дре<strong>в</strong>не<strong>к</strong>рылые) и Neoptera (н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылые);<br />

п<strong>о</strong><strong>с</strong>ледний р<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> четыре<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>рты 1<br />

: Archaeoneoptera, Polyneoptera, Р<strong>а</strong>-<br />

raneoptera и Holometabola (или Oligoneoptera).<br />

Эт<strong>о</strong> деление, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е н<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ении<br />

<strong>к</strong>рылье<strong>в</strong>, п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я т<strong>а</strong><strong>к</strong>же <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тями<br />

пре<strong>в</strong>р<strong>а</strong>щения и <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м жизни неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых<br />

<strong>с</strong>т<strong>а</strong>дий. Перечи<strong>с</strong>ленные инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы<br />

и <strong>к</strong><strong>о</strong>г<strong>о</strong>рты <strong>к</strong>рыл<strong>а</strong>тьгх н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ют<br />

р<strong>а</strong>зличные <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong>,<br />

и <strong>о</strong>тряд<strong>о</strong><strong>в</strong>; х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>и этих группир<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong><br />

<strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong>й мере д<strong>о</strong>п<strong>о</strong>лняют<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тями<br />

<strong>с</strong>тр<strong>о</strong>ения р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. Чт<strong>о</strong> <strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong> б<strong>о</strong>лее низ<strong>к</strong>их единиц<strong>с</strong>и<strong>с</strong>темы<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> п<<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>), т<strong>о</strong> не<strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong> <strong>с</strong> д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерн<strong>о</strong><strong>с</strong>тью) <strong>о</strong><strong>с</strong>юб-<br />

35 3*<br />

http://jurassic.ru/

щить х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер их р<strong>а</strong>зличии: <strong>в</strong> р<strong>а</strong>зных <strong>о</strong>тряд<strong>а</strong>х<br />

эти <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие единицы х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я<br />

<strong>о</strong>чень р<strong>а</strong>зличн<strong>о</strong>, и их ди<strong>а</strong>гн<strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тях <strong>с</strong><strong>а</strong>мых р<strong>а</strong>зных <strong>с</strong>и<strong>с</strong>тем <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>.<br />

Об <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>х <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и р<strong>а</strong>зных <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

и групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых г<strong>о</strong><strong>в</strong><strong>о</strong>рит<strong>с</strong>я <strong>в</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных<br />

<strong>о</strong>чер<strong>к</strong><strong>а</strong>х, зде<strong>с</strong>ь же п<strong>о</strong>дчер<strong>к</strong>нем, чт<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ение<br />

<strong>к</strong>рылье<strong>в</strong> п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> и<strong>с</strong>п<strong>о</strong>льзует<strong>с</strong>я<br />

для р<strong>а</strong>зличения <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их единиц<br />

п<strong>о</strong>чти люб<strong>о</strong>г<strong>о</strong> р<strong>а</strong>нг<strong>а</strong>, <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>их<br />

<strong>к</strong><strong>а</strong>тег<strong>о</strong>рий.<br />

И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>с</strong> другими члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гими д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р не м<strong>о</strong>гут<br />

<strong>с</strong>чит<strong>а</strong>ть<strong>с</strong>я <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>тельн<strong>о</strong> <strong>в</strong>ыя<strong>с</strong>ненными. П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

н<strong>а</strong>иб<strong>о</strong>лее д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерн<strong>о</strong> <strong>с</strong>ближение н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>с</strong> мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong>ми, именн<strong>о</strong> <strong>с</strong> групп<strong>о</strong>й<br />

Symphyla. Другие т<strong>о</strong>ч<strong>к</strong>и зрения, н<strong>а</strong>пример, п<strong>о</strong>и<strong>с</strong><strong>к</strong>и<br />

пред<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong>реди трил<strong>о</strong>бит<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

ф<strong>о</strong>рм или р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, не з<strong>а</strong><strong>с</strong>лужи<strong>в</strong><strong>а</strong>ют<br />

<strong>с</strong>ерьезн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ним<strong>а</strong>ния, будучи м<strong>а</strong>л<strong>о</strong><strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нными.<br />

В<strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>л<strong>о</strong><strong>с</strong>ь<br />

прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> яр<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным<br />

пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>о</strong>м т<strong>а</strong>гм<strong>а</strong>тиз<strong>а</strong>ции — <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блением <strong>с</strong>реднег<strong>о</strong><br />

<strong>о</strong>тдел<strong>а</strong> тел<strong>а</strong>, груди, н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рый легл<strong>а</strong><br />

фун<strong>к</strong>ция л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>ции; т<strong>о</strong>ль<strong>к</strong><strong>о</strong> три <strong>с</strong>егмент<strong>а</strong> груди<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нили д<strong>в</strong>иг<strong>а</strong>тельные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти — х<strong>о</strong>дильные<br />

н<strong>о</strong>ги. Эт<strong>о</strong> <strong>о</strong>пределил<strong>о</strong> <strong>с</strong>т<strong>о</strong>ль типичн<strong>о</strong>е<br />

п<strong>о</strong>др<strong>а</strong>зделение тел<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> н<strong>а</strong> три т<strong>а</strong>гмы:<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>у, грудь и брюш<strong>к</strong><strong>о</strong>.<br />

Д<strong>а</strong>льнейший эт<strong>а</strong>п и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>а</strong> именн<strong>о</strong>, <strong>в</strong>ыделение н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными р<strong>о</strong>т<strong>о</strong><strong>в</strong>ыми <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми,<br />

тр<strong>а</strong>хейн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>тем<strong>о</strong>й, 11-члени<strong>к</strong><strong>о</strong><strong>в</strong>ым брюш<strong>к</strong><strong>о</strong>м,<br />

длинными <strong>а</strong>нтенн<strong>а</strong>ми и цер<strong>к</strong><strong>а</strong>ми, эмбри<strong>о</strong>н<strong>а</strong>льным<br />

р<strong>а</strong>з<strong>в</strong>итием <strong>с</strong> интен<strong>с</strong>и<strong>в</strong>ным бл<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong>инез<strong>о</strong>м<br />

и <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нием з<strong>а</strong>р<strong>о</strong>дыше<strong>в</strong>ых <strong>о</strong>б<strong>о</strong>л<strong>о</strong>че<strong>к</strong>,<br />

д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р еще <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тен. Т<strong>а</strong><strong>к</strong> же<br />

м<strong>а</strong>л<strong>о</strong> изучен<strong>о</strong> и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е 'р<strong>а</strong>з<strong>в</strong>итие пер<strong>в</strong>ичных<br />

<strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, их ди<strong>в</strong>ергенция<br />

<strong>с</strong> групп<strong>о</strong>й щетин<strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> (Thysanura), <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение<br />

<strong>к</strong>рылье<strong>в</strong>. Эти пр<strong>о</strong>це<strong>с</strong><strong>с</strong>ы д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р<br />

<strong>о</strong>бъя<strong>с</strong>няют<strong>с</strong>я и р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ют<strong>с</strong>я лишь н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е<br />

чи<strong>с</strong>т<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>их д<strong>а</strong>нных<br />

и <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных, <strong>в</strong> т<strong>о</strong>й или ин<strong>о</strong>й мере д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерных<br />

предп<strong>о</strong>л<strong>о</strong>жений <strong>о</strong>б у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях, <strong>о</strong>предели<strong>в</strong>ших<br />

т<strong>а</strong><strong>к</strong>ие н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ления фил<strong>о</strong>генез<strong>а</strong>; <strong>к</strong><strong>а</strong><strong>к</strong>ие-<br />

Ри<strong>с</strong>. 26. Схем<strong>а</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>, инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>, <strong>к</strong><strong>о</strong>г<strong>о</strong>рт и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

Об<strong>о</strong>зн<strong>а</strong>чения: С - <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Cr - мел, D - де<strong>в</strong><strong>о</strong>н, J - юр<strong>а</strong>. Pal - Palaeoptera, Par - Paraneoptera, Pip - Plecopteroidea,<br />

N _не 0 Г ен p - пермь, Pg - п<strong>а</strong>ле<strong>о</strong>ген, S - <strong>с</strong>илур, T - три- Pol - Polyneoptera, Pt - Pterygota; <strong>с</strong>хем<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong> 1958 г.<br />

<strong>а</strong><strong>с</strong> АР - Apterygota, Blp - Blattopteroidea, Cf - Coleopte- и <strong>к</strong><strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Archaeoneoptera не п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>а</strong> (Р<strong>о</strong>денд<strong>о</strong>рф, Бе<strong>к</strong><strong>к</strong>ерrol'dea,<br />

Hm - Hymenopteroidea, М<strong>с</strong> - Mecopteroldea, Nr - Ne- Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)<br />

uropteroidea, Ollg — Oligoneoptefa, Orth — Orthopteroidea,<br />

36<br />

http://jurassic.ru/

Ри<strong>с</strong>. 27. Схем<strong>а</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений <strong>о</strong>тряд<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Apterygota, инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Palaeoptera и <strong>к</strong><strong>о</strong>г<strong>о</strong>рты<br />

Polyneoptera.<br />

Об<strong>о</strong>зн<strong>а</strong>чения; Ар — Apterygota, Bl — Blattodea, С — Col- Pal — Palaeoptera, Par — Paraneoptera, Ph — Phasmatodea,<br />

lembola, CG — Caloneurodea и Glosselytrodea, D — Diplura, PI — Plecoptera, Ppl — Paraplecoptera, Pr — Protelytroptera,<br />

Der — Dermaptera, Em — Embioptera, Eph — Ephemeroptera, Prb — Protoblattodea, Sal — Saltatoria, Thy — Thysanura<br />

Is — Isoptera, M —Monura, Man —Manteodea, Meg —Megase- (Р<strong>о</strong>денд<strong>о</strong>рф и др., 1961).<br />

coptera, Od — Odonata, Olig — Oligoneoptera, P — Protura,<br />

либ<strong>о</strong> т<strong>о</strong>чные д<strong>о</strong><strong>к</strong>умент<strong>а</strong>льные п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong><strong>а</strong> д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют и<br />

не <strong>о</strong><strong>с</strong><strong>в</strong>ещ<strong>а</strong>ют эти <strong>в</strong><strong>а</strong>жные м<strong>о</strong>менты э<strong>в</strong><strong>о</strong>люции.<br />

Г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> б<strong>о</strong>лее д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерны <strong>с</strong><strong>в</strong>едения <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong><br />

<strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ения д<strong>в</strong>ух и<strong>в</strong>фр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

<strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых: н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых и дре<strong>в</strong>не<strong>к</strong>рылых.<br />

П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, н<strong>а</strong>иб<strong>о</strong>лее пр<strong>о</strong><strong>с</strong>т<strong>о</strong> п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енными<br />

и близ<strong>к</strong>ими <strong>к</strong> пер<strong>в</strong>ичным и<strong>с</strong>х<strong>о</strong>дным<br />

ф<strong>о</strong>рм<strong>а</strong>м <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых были пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Neoptera, <strong>а</strong> именн<strong>о</strong>,<br />

нед<strong>а</strong><strong>в</strong>н<strong>о</strong> <strong>о</strong>бн<strong>а</strong>руженный <strong>в</strong> де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уне<br />

<strong>о</strong>тряд Archaeoptera, <strong>о</strong>бр<strong>а</strong>зующий <strong>о</strong><strong>с</strong><strong>о</strong>бую <strong>к</strong><strong>о</strong>г<strong>о</strong>рту<br />

н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых — Archaeoneoptera. Б<strong>о</strong>льш<strong>а</strong>я<br />

пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>ть <strong>к</strong>рылье<strong>в</strong> этих н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>й <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шую<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ть их<br />

и<strong>с</strong>х<strong>о</strong>дных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>, <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> <strong>с</strong>пин<strong>к</strong>и грудн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> — п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>лий. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние у них<br />

р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong> еще <strong>с</strong>л<strong>а</strong>б<strong>о</strong>, х<strong>о</strong>тя ф<strong>о</strong>рм<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> уже<br />

при<strong>о</strong>брел<strong>а</strong> <strong>а</strong>эр<strong>о</strong>дин<strong>а</strong>миче<strong>с</strong><strong>к</strong>ие при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления.<br />

Эти де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые уже были «н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылыми»,<br />

т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> м<strong>о</strong>гли <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ть <strong>к</strong>рылья<br />

<strong>в</strong>д<strong>о</strong>ль тел<strong>а</strong>; п<strong>о</strong>эт<strong>о</strong>му <strong>с</strong>ледует предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ть, чт<strong>о</strong><br />

пер<strong>в</strong>ичные дре<strong>в</strong>не<strong>к</strong>рылые <strong>в</strong><strong>о</strong>зни<strong>к</strong>ли именн<strong>о</strong> н<strong>а</strong><br />

их <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е, путем <strong>о</strong>чень р<strong>а</strong>нней <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>к</strong>и<br />

б<strong>о</strong>льших переп<strong>о</strong>нч<strong>а</strong>тых <strong>к</strong>рылье<strong>в</strong>, п<strong>о</strong>з<strong>в</strong><strong>о</strong>ли<strong>в</strong>ших<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым, <strong>о</strong>бит<strong>а</strong>телям <strong>о</strong>т<strong>к</strong>рытых пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong>,<br />

<strong>о</strong>чень р<strong>а</strong>н<strong>о</strong> д<strong>о</strong><strong>с</strong>тичь <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

лет<strong>а</strong>тельных при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений.<br />

Друг<strong>о</strong>й путь — д<strong>а</strong>льнейшее <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рыл<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — <strong>о</strong>бит<strong>а</strong>телей<br />

<strong>с</strong><strong>к</strong>рытых пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong> — з<strong>а</strong><strong>к</strong>люч<strong>а</strong>л<strong>с</strong>я <strong>в</strong><br />

р<strong>а</strong>з<strong>в</strong>итии жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния и при<strong>о</strong>бретении передними<br />

<strong>к</strong>рыльями х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong><br />

<strong>с</strong> р<strong>а</strong>з<strong>в</strong>итием гетер<strong>о</strong>н<strong>о</strong>мии <strong>к</strong>рылье<strong>в</strong>.<br />

В эт<strong>о</strong>м <strong>с</strong>луч<strong>а</strong>е т<strong>о</strong>ль<strong>к</strong><strong>о</strong> з<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong>т<strong>а</strong>ли<br />

лег<strong>к</strong>ими и эл<strong>а</strong><strong>с</strong>тичными лет<strong>а</strong>тельными <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми,<br />

<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бными <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ть<strong>с</strong>я и <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>лять<br />

бы<strong>с</strong>трые <strong>в</strong>зм<strong>а</strong>хи. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы <strong>к</strong><strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Polyneoptera<br />

и прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong>тряд т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ых (Blattodea).<br />

Д<strong>а</strong>льнейш<strong>а</strong>я и<strong>с</strong>т<strong>о</strong>рия этих инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong>чень р<strong>а</strong>зличн<strong>а</strong>. Дре<strong>в</strong>не<strong>к</strong>рылые <strong>о</strong>чень <strong>с</strong><strong>к</strong><strong>о</strong>р<strong>о</strong>,<br />

еще <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>е, дифференцир<strong>о</strong><strong>в</strong><strong>а</strong>ли<strong>с</strong>ь н<strong>а</strong> ряд<br />

<strong>в</strong>т<strong>о</strong>ричных группир<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>: не менее чем н<strong>а</strong> три<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>, мн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>о</strong><strong>в</strong> и п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong>. Н<strong>о</strong><br />

еще б<strong>о</strong>лее <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>х<strong>о</strong>дил<strong>о</strong> <strong>в</strong><br />

37<br />

http://jurassic.ru/

инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые р<strong>а</strong>зделили<strong>с</strong>ь<br />

н<strong>а</strong> четыре <strong>о</strong>бширные <strong>к</strong><strong>о</strong>г<strong>о</strong>рты <strong>с</strong> мн<strong>о</strong>же<strong>с</strong>т<strong>в</strong><strong>о</strong>м<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> и <strong>о</strong>тряд<strong>о</strong><strong>в</strong>.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных<br />

групп <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых при<strong>в</strong>едены н<strong>а</strong> прил<strong>а</strong>г<strong>а</strong>емых<br />

<strong>с</strong>хем<strong>а</strong>х (ри<strong>с</strong>. 26 и 27), <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти<br />

и<strong>с</strong>т<strong>о</strong>рии инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> и <strong>к</strong><strong>о</strong>г<strong>о</strong>рт р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных <strong>о</strong>чер<strong>к</strong><strong>а</strong>х. Н<strong>а</strong> прил<strong>а</strong>г<strong>а</strong>емых<br />

<strong>с</strong>хем<strong>а</strong>х п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>о</strong> <strong>в</strong>ремя ди<strong>в</strong>ергенции<br />

гл<strong>а</strong><strong>в</strong>ных групп. Эти ди<strong>в</strong>ергенции пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дили<br />

<strong>в</strong> течение де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong><br />

пери<strong>о</strong>д<strong>о</strong><strong>в</strong>. Не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong> з<strong>а</strong>метить, чт<strong>о</strong> ре<strong>а</strong>льные<br />

п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong><strong>а</strong> р<strong>а</strong>нней<br />

и<strong>с</strong>т<strong>о</strong>рии <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, и <strong>о</strong> них мы <strong>с</strong>удим лишь н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>в</strong>енных (<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их и<br />

иных) д<strong>а</strong>нных.<br />

С<strong>а</strong>мые дре<strong>в</strong>ние п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие д<strong>о</strong><strong>к</strong>ументы<br />

п<strong>о</strong> и<strong>с</strong>т<strong>о</strong>рии Pterygota д<strong>а</strong>тируют<strong>с</strong>я <strong>в</strong>ерхним<br />

де<strong>в</strong><strong>о</strong>н<strong>о</strong>м. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong> уже у<strong>к</strong><strong>а</strong>з<strong>а</strong>нн<strong>а</strong>я н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong><br />

<strong>в</strong> Се<strong>в</strong>ерн<strong>о</strong>м Приур<strong>а</strong>лье <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>т<strong>о</strong>г<strong>о</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> Eopterum devonicum Rohdendorf<br />

(<strong>с</strong>тр. 86), прин<strong>а</strong>длеж<strong>а</strong>щег<strong>о</strong> <strong>к</strong> <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>му <strong>о</strong>тряду<br />

Archaeoptera, н<strong>а</strong>иб<strong>о</strong>лее близ<strong>к</strong><strong>о</strong>му <strong>к</strong> т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ым<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). Эт<strong>а</strong> ф<strong>о</strong>рм<strong>а</strong> <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>л<strong>а</strong><br />

упл<strong>о</strong>тненными передними <strong>к</strong>рыльями <strong>с</strong> немн<strong>о</strong>гими<br />

пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми. Не<strong>с</strong>м<strong>о</strong>тря<br />

н<strong>а</strong> б<strong>о</strong>льшую дре<strong>в</strong>н<strong>о</strong><strong>с</strong>ть эт<strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>, <strong>о</strong>н<strong>о</strong><br />

н<strong>о</strong><strong>с</strong>ил<strong>о</strong> уже не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые черты <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ции.<br />

Р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>в</strong>пер<strong>в</strong>ые п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

лет<strong>о</strong>пи<strong>с</strong>и <strong>в</strong> <strong>в</strong>ерх<strong>а</strong>х нижнег<strong>о</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> —<br />

н<strong>а</strong>мюре. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых дре<strong>в</strong>не<strong>к</strong>рылых,<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых, прям<strong>о</strong><strong>к</strong>рылых, дре<strong>в</strong>нейших<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц <strong>в</strong> Е<strong>в</strong>р<strong>о</strong>пе и С. Амери<strong>к</strong>е.<br />

В <strong>в</strong>е<strong>с</strong>тф<strong>а</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>м яру<strong>с</strong>е <strong>с</strong>реднег<strong>о</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>в</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных <strong>о</strong>тл<strong>о</strong>жениях<br />

С. Амери<strong>к</strong>и <strong>о</strong>бн<strong>а</strong>ружены уже б<strong>о</strong>г<strong>а</strong>тые ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы из пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

<strong>о</strong>тряд<strong>о</strong><strong>в</strong>. П<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>у, п<strong>о</strong>я<strong>в</strong>ление<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных<br />

<strong>о</strong>тл<strong>о</strong>жениях был<strong>о</strong> <strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong>й<br />

мере <strong>в</strong>нез<strong>а</strong>пным и <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует <strong>о</strong> <strong>к</strong>р<strong>а</strong>йней<br />

нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><strong>с</strong>ти п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их д<strong>а</strong>нных.<br />

В<strong>с</strong>е <strong>о</strong>бн<strong>а</strong>руженные ф<strong>о</strong>рмы прин<strong>а</strong>длеж<strong>а</strong>т <strong>к</strong> <strong>о</strong>тряд<strong>а</strong>м,<br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненным <strong>в</strong> 'б<strong>о</strong>лее п<strong>о</strong>здние<br />

ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие пери<strong>о</strong>ды, и лишь <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>й м<strong>а</strong>л<strong>о</strong>й<br />

мере п<strong>о</strong>м<strong>о</strong>г<strong>а</strong>ют <strong>о</strong><strong>с</strong><strong>в</strong>етить м<strong>о</strong>менты <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ления<br />

и ди<strong>в</strong>ергенции <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных групп <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>.<br />

О<strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong><strong>а</strong>жны н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и <strong>в</strong> этих дре<strong>в</strong>нейших<br />

ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>о</strong>тряд<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, прин<strong>а</strong>длеж<strong>а</strong>щих <strong>к</strong><br />

групп<strong>а</strong>м, н<strong>а</strong>иб<strong>о</strong>льший р<strong>а</strong><strong>с</strong>ц<strong>в</strong>ет <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<br />

зн<strong>а</strong>чительн<strong>о</strong> п<strong>о</strong>зднее. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и<br />

дре<strong>в</strong>ней <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нницы (Metropator pusillus<br />

Handlirsch) <strong>в</strong> <strong>в</strong>ерх<strong>а</strong>х нижнег<strong>о</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> (п<strong>о</strong>т<strong>с</strong><strong>в</strong>ил)<br />

С. Амери<strong>к</strong>и, дре<strong>в</strong>нейших р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых<br />

(Protoprosbole straeleni Laurentiaux) из н<strong>а</strong>мюр<strong>а</strong><br />

38<br />

http://jurassic.ru/<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong><strong>в</strong>иты <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

Ю. Сибири (Blattoprosbole tomiensis Becker-<br />

Migdisova). Эти ф<strong>а</strong><strong>к</strong>ты з<strong>а</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>о</strong>т<strong>о</strong>д<strong>в</strong>иг<strong>а</strong>ть <strong>в</strong> пр<strong>о</strong>шл<strong>о</strong>е м<strong>о</strong>менты <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ения<br />

инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> и <strong>к</strong><strong>о</strong>г<strong>о</strong>рт <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> н<strong>а</strong>иб<strong>о</strong>лее пр<strong>о</strong>гре<strong>с</strong><strong>с</strong>и<strong>в</strong>ных групп<br />

Paraneoptera и Holometabola.<br />

Нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><strong>с</strong>ть п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их д<strong>а</strong>нных<br />

у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет н<strong>а</strong> т<strong>о</strong>, чт<strong>о</strong> решение <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития <strong>к</strong>рупных групп п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

Pterygota м<strong>о</strong>жет <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ть<strong>с</strong>я преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниях, изучении инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ития и э<strong>к</strong><strong>о</strong>л<strong>о</strong>гии; ф<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong>ие п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>о</strong>чень<br />

нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чными и не д<strong>а</strong>ют <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти <strong>в</strong>ыя<strong>с</strong>нить<br />

<strong>к</strong><strong>о</strong>н<strong>к</strong>ретные пути фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития.<br />

Обр<strong>а</strong>з жизни, э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и т<strong>а</strong>ф<strong>о</strong>н<strong>о</strong>мия<br />

Р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong> Земле <strong>в</strong> <strong>о</strong>бщих<br />

черт<strong>а</strong>х м<strong>о</strong>жет быть <strong>к</strong>р<strong>а</strong>т<strong>к</strong><strong>о</strong> <strong>о</strong>х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong><br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющее <strong>с</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нением<br />

н<strong>а</strong>земн<strong>о</strong>й фл<strong>о</strong>ры. Лишь <strong>к</strong>р<strong>а</strong>йние п<strong>о</strong>лярные з<strong>о</strong>ны<br />

и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>рья <strong>с</strong> <strong>в</strong>ечными ледни<strong>к</strong><strong>а</strong>ми лишены<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые т<strong>а</strong><strong>к</strong>же чужды м<strong>о</strong>рю,<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м <strong>о</strong>бит<strong>а</strong>ют лишь <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

и <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зные ф<strong>о</strong>рмы: <strong>о</strong><strong>с</strong><strong>о</strong>бые пл<strong>а</strong><strong>в</strong><strong>а</strong>ющие<br />

<strong>к</strong>л<strong>о</strong>пы, жи<strong>в</strong>ущие н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

м<strong>о</strong>рей и личин<strong>к</strong>и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых д<strong>в</strong>у<strong>к</strong>рылых,<br />

<strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong> прибрежных, приб<strong>о</strong>йных з<strong>о</strong>н<strong>а</strong>х.<br />

Отн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong> личин<strong>о</strong><strong>к</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong><br />

<strong>о</strong>пре<strong>с</strong>ненных м<strong>о</strong>р<strong>с</strong><strong>к</strong>их <strong>о</strong><strong>к</strong>р<strong>а</strong>инных <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х,<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых жи<strong>в</strong>ут мн<strong>о</strong>гие д<strong>в</strong>у<strong>к</strong>рылые и ручейни<strong>к</strong>и.<br />

Гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>е б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — <strong>о</strong>бит<strong>а</strong>тели<br />

<strong>с</strong>уши и <strong>к</strong><strong>о</strong>нтинент<strong>а</strong>льных <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong> (ре<strong>к</strong><br />

и пре<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>дных <strong>о</strong>зер).<br />

Г<strong>о</strong><strong>в</strong><strong>о</strong>ря <strong>о</strong>б <strong>о</strong>бщем р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нении н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong> прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>в</strong><strong>с</strong>п<strong>о</strong>мнить <strong>о</strong><br />

не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong>м р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зии и <strong>о</strong>билии этих жи<strong>в</strong><strong>о</strong>тных<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>с</strong> любыми другими <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>ми<br />

и тип<strong>а</strong>ми <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong>. Причин<strong>а</strong>ми эт<strong>о</strong>г<strong>о</strong><br />

я<strong>в</strong>ляют<strong>с</strong>я прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> мел<strong>к</strong>ие р<strong>а</strong>змеры и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong>тепень <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, чт<strong>о</strong><br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>лил<strong>о</strong> им <strong>в</strong> пр<strong>о</strong>це<strong>с</strong><strong>с</strong>е э<strong>в</strong><strong>о</strong>люции <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>а</strong>ть<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ершенные при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления <strong>к</strong> <strong>с</strong><strong>а</strong>мым<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным у<strong>с</strong>л<strong>о</strong><strong>в</strong>иям <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния, з<strong>а</strong>х<strong>в</strong><strong>а</strong>тить<br />

и з<strong>а</strong><strong>с</strong>елить бу<strong>к</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong> <strong>в</strong><strong>с</strong>е пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

н<strong>а</strong> Земле, где т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>а</strong> <strong>о</strong>рг<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

жизнь.<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые р<strong>а</strong>зличных <strong>к</strong>лим<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их п<strong>о</strong>я<strong>с</strong><strong>о</strong><strong>в</strong>,<br />

т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>о</strong>пределенных з<strong>о</strong><strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>их<br />

р<strong>а</strong>й<strong>о</strong>н<strong>о</strong><strong>в</strong>, <strong>о</strong>чень р<strong>а</strong>зличны: м<strong>о</strong>жн<strong>о</strong> <strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть,<br />

чт<strong>о</strong> ф<strong>а</strong>уны н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых д<strong>а</strong>ют н<strong>а</strong>иб<strong>о</strong>лее<br />

убедительный п<strong>о</strong> п<strong>о</strong>лн<strong>о</strong>те м<strong>а</strong>тери<strong>а</strong>л для з<strong>о</strong><strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>й<strong>о</strong>нир<strong>о</strong><strong>в</strong><strong>а</strong>ния и х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>и<br />

люб<strong>о</strong>й <strong>с</strong>тр<strong>а</strong>ны и люб<strong>о</strong>г<strong>о</strong> м<strong>а</strong>тери<strong>к</strong><strong>а</strong>. О<strong>с</strong><strong>о</strong>-

бенн<strong>о</strong> р<strong>а</strong>зительны, <strong>к</strong><strong>о</strong>нечн<strong>о</strong>, <strong>о</strong>тличия тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

и умеренных п<strong>о</strong>я<strong>с</strong><strong>о</strong><strong>в</strong>; <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> тем х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>зличимы между <strong>с</strong><strong>о</strong>б<strong>о</strong>й и р<strong>а</strong>зличные тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие<br />

или умеренные ф<strong>а</strong>уны, н<strong>а</strong>пример,<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Ю. Амери<strong>к</strong>и, Афри<strong>к</strong>и и Ю. Азии<br />

или ф<strong>а</strong>уны С. Амери<strong>к</strong>и и Е<strong>в</strong>р<strong>а</strong>зии.<br />

В<strong>с</strong>ех н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых м<strong>о</strong>жн<strong>о</strong> груб<strong>о</strong> р<strong>а</strong>збить н<strong>а</strong><br />

д<strong>в</strong>е нер<strong>а</strong><strong>в</strong>ные э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие группы. Меньш<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ит группу <strong>в</strong><strong>о</strong>дных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, т. е.<br />

т<strong>а</strong><strong>к</strong>их, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые или <strong>в</strong> течение <strong>в</strong><strong>с</strong>ей жизни жи<strong>в</strong>ут<br />

<strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые жу<strong>к</strong>и и <strong>к</strong>л<strong>о</strong>пы), или<br />

т<strong>а</strong><strong>к</strong>их, личин<strong>к</strong>и <strong>к</strong><strong>о</strong>т<strong>о</strong>рых жи<strong>в</strong>ут <strong>в</strong> <strong>в</strong><strong>о</strong>де, <strong>а</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лые<br />

ф<strong>о</strong>рмы <strong>в</strong>едут <strong>о</strong>бычный, <strong>в</strong><strong>о</strong>здушный, <strong>о</strong>бр<strong>а</strong>з<br />

жизни. К п<strong>о</strong><strong>с</strong>ледней <strong>к</strong><strong>а</strong>тег<strong>о</strong>рии прин<strong>а</strong>длежит<br />

гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>е б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong><strong>о</strong>дных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

(<strong>с</strong>тре<strong>к</strong><strong>о</strong>зы, п<strong>о</strong>ден<strong>к</strong>и, <strong>в</strong>е<strong>с</strong>нян<strong>к</strong>и, ручейни<strong>к</strong>и, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые, д<strong>в</strong>у<strong>к</strong>рылые и другие).<br />

К<strong>о</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й группе н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — чи<strong>с</strong>т<strong>о</strong> н<strong>а</strong>земных,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляющих пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ющее б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong>,—<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ует п<strong>о</strong><strong>с</strong>тепенный перех<strong>о</strong>д <strong>о</strong>т <strong>в</strong><strong>о</strong>дных<br />

ф<strong>о</strong>рм через н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, жи<strong>в</strong>ущих <strong>в</strong> <strong>в</strong><strong>о</strong>дн<strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>ти,<br />

н<strong>а</strong> <strong>с</strong>ырых луг<strong>а</strong>х, <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й п<strong>о</strong>ч<strong>в</strong>е,<br />

<strong>к</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым — <strong>о</strong>бит<strong>а</strong>телям ле<strong>с</strong><strong>а</strong>, п<strong>о</strong>лей и<br />

<strong>с</strong>тепей. Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, пу<strong>с</strong>тыни <strong>в</strong><strong>с</strong>ех <strong>с</strong>тр<strong>а</strong>н <strong>с</strong><strong>в</strong>ет<strong>а</strong> н<strong>а</strong><strong>с</strong>елены<br />

<strong>о</strong>чень <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й и б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>й пу<strong>с</strong>тынн<strong>о</strong>й<br />

ф<strong>а</strong>ун<strong>о</strong>й. Р<strong>а</strong>зличные э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти<br />

<strong>о</strong>бит<strong>а</strong>ния н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют чет<strong>к</strong>ий<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong> н<strong>а</strong> <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>цию жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong>. Не<br />

г<strong>о</strong><strong>в</strong><strong>о</strong>ря уже <strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — <strong>в</strong><strong>о</strong>дных <strong>о</strong>бит<strong>а</strong>телях,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong> их <strong>о</strong>бр<strong>а</strong>зу жизни<br />

<strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> яр<strong>к</strong>и и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метны, т<strong>а</strong><strong>к</strong>же и иные<br />

э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие у<strong>с</strong>л<strong>о</strong><strong>в</strong>ия <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> у<strong>с</strong>тр<strong>о</strong>й<strong>с</strong>т<strong>в</strong>е<br />

тел<strong>а</strong>. П<strong>о</strong>эт<strong>о</strong>му мы <strong>в</strong><strong>с</strong>егд<strong>а</strong> м<strong>о</strong>жем <strong>с</strong> т<strong>о</strong>й или<br />

ин<strong>о</strong>й <strong>с</strong>тепенью <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти <strong>с</strong>удить <strong>о</strong>б <strong>о</strong>бр<strong>а</strong>зе<br />

жизни н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> п<strong>о</strong> ег<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению.<br />

О<strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти пит<strong>а</strong>ния, р<strong>а</strong>змн<strong>о</strong>жения и р<strong>а</strong>з<strong>в</strong>ития<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны.<br />

М<strong>о</strong>жн<strong>о</strong> <strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть, чт<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>в</strong> цел<strong>о</strong>м<br />

я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong> п<strong>о</strong>лн<strong>о</strong>м <strong>с</strong>мы<strong>с</strong>ле <strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong><strong>с</strong>еядными<br />

жи<strong>в</strong><strong>о</strong>тными: нет, п<strong>о</strong>ж<strong>а</strong>луй, ни <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> <strong>о</strong>рг<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е<br />

не <strong>с</strong>лужил<strong>о</strong> бы пищей <strong>к</strong><strong>а</strong><strong>к</strong>им-либ<strong>о</strong> <strong>в</strong>ид<strong>а</strong>м<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, и<strong>с</strong><strong>к</strong>люч<strong>а</strong>я р<strong>а</strong>з<strong>в</strong>е лишь не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

пр<strong>о</strong>ду<strong>к</strong>ты р<strong>а</strong><strong>с</strong>п<strong>а</strong>д<strong>а</strong> <strong>а</strong>з<strong>о</strong>ти<strong>с</strong>тых <strong>в</strong>еще<strong>с</strong>т<strong>в</strong>. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пит<strong>а</strong>ет<strong>с</strong>я р<strong>а</strong><strong>с</strong>тительными<br />

<strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong>ми (фит<strong>о</strong>ф<strong>а</strong>ги), меньш<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть — хищни<strong>к</strong>и,<br />

н<strong>а</strong>п<strong>а</strong>д<strong>а</strong>ющие н<strong>а</strong> других н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

иных жи<strong>в</strong><strong>о</strong>тных. Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, мн<strong>о</strong>гие<br />

группы я<strong>в</strong>ляют<strong>с</strong>я п<strong>а</strong>р<strong>а</strong>зит<strong>а</strong>ми <strong>с</strong><strong>а</strong>мых р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

жи<strong>в</strong><strong>о</strong>тных—<strong>о</strong>чень мн<strong>о</strong>гих п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных,<br />

м<strong>о</strong>ллю<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>, члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих (<strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле<br />

других н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong><br />

и д<strong>а</strong>же не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных), <strong>к</strong><strong>о</strong>льч<strong>а</strong>тых<br />

чер<strong>в</strong>ей. О<strong>с</strong><strong>о</strong>бую би<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ую группу<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют п<strong>о</strong>требители гниющих<br />

р<strong>а</strong><strong>с</strong>тительных и жи<strong>в</strong><strong>о</strong>тных <strong>в</strong>еще<strong>с</strong>т<strong>в</strong>, труп<strong>о</strong><strong>в</strong> и<br />

э<strong>к</strong><strong>с</strong><strong>к</strong>ремент<strong>о</strong><strong>в</strong> жи<strong>в</strong><strong>о</strong>тных (<strong>с</strong>хиз<strong>о</strong>ф<strong>а</strong>ги). В<strong>с</strong>еядн<strong>о</strong><strong>с</strong>ть<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых не<strong>о</strong>быч<strong>а</strong>йн<strong>а</strong>: <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>уют н<strong>а</strong>­<br />

39<br />

http://jurassic.ru/<br />

<strong>с</strong>е<strong>к</strong><strong>о</strong>мые, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые м<strong>о</strong>гут жить <strong>в</strong> <strong>с</strong>ыр<strong>о</strong>й нефти;<br />

т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>, н<strong>а</strong>пример, из<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>я нефтян<strong>а</strong>я муш<strong>к</strong><strong>а</strong>,<br />

личин<strong>к</strong>и <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й жи<strong>в</strong>ут <strong>в</strong> луж<strong>а</strong>х, пит<strong>а</strong>я<strong>с</strong>ь ми<strong>к</strong>р<strong>о</strong><strong>о</strong>рг<strong>а</strong>низм<strong>а</strong>ми<br />

нефти, <strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные термиты—<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бые тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>в</strong>редные,— пит<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>чти чи<strong>с</strong>т<strong>о</strong>й целлюл<strong>о</strong>з<strong>о</strong>й,<br />

п<strong>о</strong>жир<strong>а</strong>я <strong>с</strong>ух<strong>о</strong>е дере<strong>в</strong><strong>о</strong> <strong>в</strong> п<strong>о</strong>дел<strong>к</strong><strong>а</strong>х.<br />

Пище<strong>в</strong><strong>а</strong>я <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ция н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>чень <strong>о</strong><strong>с</strong>л<strong>о</strong>жняет<strong>с</strong>я<br />

н<strong>а</strong>личием у них пре<strong>в</strong>р<strong>а</strong>щения: <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, личин<strong>к</strong>и пит<strong>а</strong>ют<strong>с</strong>я ин<strong>о</strong>й пищей,<br />

чем <strong>в</strong>зр<strong>о</strong><strong>с</strong>лые ф<strong>о</strong>рмы. При эт<strong>о</strong>м ин<strong>о</strong>гд<strong>а</strong> н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я<br />

и<strong>с</strong><strong>к</strong>лючительные примеры <strong>а</strong>ф<strong>а</strong>гии<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем лиш<strong>а</strong>ют<strong>с</strong>я <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ти приним<strong>а</strong>ть<br />

пищу и жи<strong>в</strong>ут з<strong>а</strong> <strong>с</strong>чет н<strong>а</strong><strong>к</strong><strong>о</strong>пленных жир<strong>о</strong><strong>в</strong> <strong>в</strong><strong>о</strong><br />

<strong>в</strong>ремя их личин<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития. К т<strong>а</strong><strong>к</strong>им н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я мн<strong>о</strong>гие д<strong>в</strong>у<strong>к</strong>рылые (<strong>о</strong><strong>в</strong><strong>о</strong>ды,<br />

<strong>к</strong><strong>о</strong>м<strong>а</strong>ры-з<strong>в</strong><strong>о</strong>нцы и др.), чешуе<strong>к</strong>рылые (мн<strong>о</strong>гие<br />

шел<strong>к</strong><strong>о</strong>пряды), н<strong>а</strong><strong>к</strong><strong>о</strong>нец, п<strong>о</strong>ден<strong>к</strong>и, <strong>с</strong><strong>а</strong>мцы <strong>в</strong>еер<strong>о</strong><strong>к</strong>рылых<br />

и <strong>к</strong><strong>о</strong><strong>к</strong>цид. Эт<strong>о</strong> я<strong>в</strong>ление шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нен<strong>о</strong><br />

<strong>с</strong>реди н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>уя<br />

<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>м н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лении и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ития — пед<strong>о</strong>м<strong>о</strong>рф<strong>о</strong>зе, при <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м <strong>с</strong>т<strong>а</strong>дия<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>г<strong>о</strong> жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ит<strong>с</strong>я <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>к</strong>р<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й,<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующей зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>лее<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е <strong>в</strong>ремя п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нению <strong>с</strong> неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелыми<br />

<strong>с</strong>т<strong>а</strong>диями р<strong>а</strong>з<strong>в</strong>ития. Т<strong>а</strong><strong>к</strong>ие пед<strong>о</strong>м<strong>о</strong>рфные<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>в</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ып<strong>о</strong>лняют фун<strong>к</strong>цию р<strong>а</strong>змн<strong>о</strong>жения: <strong>в</strong><strong>с</strong>е другие<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны их жизнедеятельн<strong>о</strong><strong>с</strong>ти <strong>о</strong>чень <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щ<strong>а</strong>ют<strong>с</strong>я.<br />

Би<strong>о</strong>л<strong>о</strong>гия р<strong>а</strong>змн<strong>о</strong>жения т<strong>а</strong><strong>к</strong>же <strong>о</strong>чень р<strong>а</strong>зличн<strong>а</strong>.<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые р<strong>а</strong>здельн<strong>о</strong>п<strong>о</strong>лы. С<strong>а</strong>м<strong>к</strong>и <strong>в</strong><br />

гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>м б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<br />

яйц<strong>а</strong>, реже р<strong>о</strong>жд<strong>а</strong>ют жи<strong>в</strong>ых личин<strong>о</strong><strong>к</strong>, ин<strong>о</strong>гд<strong>а</strong><br />

д<strong>а</strong>же <strong>в</strong>п<strong>о</strong>лне зрелых, р<strong>а</strong>з<strong>в</strong>и<strong>в</strong>ших<strong>с</strong>я и пит<strong>а</strong><strong>в</strong>ших<strong>с</strong>я<br />

<strong>в</strong> теле <strong>с</strong><strong>а</strong>м<strong>к</strong>и. Пл<strong>о</strong>д<strong>о</strong><strong>в</strong>ит<strong>о</strong><strong>с</strong>ть н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>к</strong><strong>о</strong>леблет<strong>с</strong>я <strong>в</strong> р<strong>а</strong>зличных групп<strong>а</strong>х <strong>в</strong> <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>их<br />

предел<strong>а</strong>х: е<strong>с</strong>ть <strong>в</strong>иды, <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ющие з<strong>а</strong><br />

<strong>в</strong><strong>с</strong>ю <strong>с</strong><strong>в</strong><strong>о</strong>ю жизнь <strong>в</strong><strong>с</strong>ег<strong>о</strong> меньше де<strong>с</strong>ят<strong>к</strong><strong>а</strong> яиц;<br />

<strong>с</strong> друг<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong>иды, х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные<br />

не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong>й пл<strong>о</strong>д<strong>о</strong><strong>в</strong>ит<strong>о</strong><strong>с</strong>тью, чи<strong>с</strong>л<strong>о</strong> яиц у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

м<strong>о</strong>жет д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ть 20 ООО.<br />

Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бны <strong>к</strong> м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>му<br />

р<strong>а</strong>змн<strong>о</strong>жению. Эт<strong>о</strong> я<strong>в</strong>ление х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>:<br />

д<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> <strong>в</strong><strong>с</strong>п<strong>о</strong>мнить м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>е п<strong>о</strong>я<strong>в</strong>ление<br />

<strong>с</strong><strong>а</strong>р<strong>а</strong>нчи — прям<strong>о</strong><strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

и тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н, р<strong>а</strong>зличных <strong>в</strong>редных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> чешуе<strong>к</strong>рылых, же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых и других<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>ые лёты п<strong>о</strong>ден<strong>о</strong><strong>к</strong>, п<strong>о</strong>я<strong>в</strong>ление<br />

<strong>в</strong> гр<strong>о</strong>м<strong>а</strong>дных <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>а</strong>х не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>с</strong>тре<strong>к</strong><strong>о</strong>з, чешуе<strong>к</strong>рылых и мн<strong>о</strong>гих других н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

не имеющих неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> х<strong>о</strong>зяй<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong><br />

зн<strong>а</strong>чения. Я<strong>в</strong>ления м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>змн<strong>о</strong>жения<br />

<strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong>л<strong>о</strong>жным <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong>м<br />

у<strong>с</strong>л<strong>о</strong><strong>в</strong>ий <strong>о</strong>бит<strong>а</strong>ния <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующих <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых: <strong>к</strong>лим<strong>а</strong>т<strong>о</strong>м, у<strong>с</strong>л<strong>о</strong><strong>в</strong>иями пит<strong>а</strong>ния,<br />

<strong>с</strong>нижением <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>а</strong> е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енных <strong>в</strong>р<strong>а</strong>г<strong>о</strong><strong>в</strong>.

Следует <strong>о</strong>тметить <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех этих <strong>с</strong>луч<strong>а</strong>ях б<strong>о</strong>льшую<br />

<strong>с</strong>л<strong>о</strong>жн<strong>о</strong><strong>с</strong>ть и,—чт<strong>о</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong><strong>а</strong>жн<strong>о</strong>,—<br />

<strong>к</strong>р<strong>а</strong>йнюю <strong>с</strong>пецифичн<strong>о</strong><strong>с</strong>ть, <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зие причин<br />

м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>змн<strong>о</strong>жения р<strong>а</strong>зных <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Б<strong>о</strong>лее<br />

глуб<strong>о</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трение этих я<strong>в</strong>лений не пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<br />

неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> интере<strong>с</strong><strong>а</strong> для п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии,—<br />

<strong>в</strong><strong>а</strong>жен лишь <strong>с</strong><strong>а</strong>м ф<strong>а</strong><strong>к</strong>т шир<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нения м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых лёт<strong>о</strong><strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, чт<strong>о</strong> имеет неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>е <strong>о</strong>тн<strong>о</strong>шение<br />

<strong>к</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>с</strong><strong>к</strong><strong>о</strong>плений и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>я<strong>с</strong>ь <strong>о</strong>дним из <strong>в</strong><strong>а</strong>жнейших у<strong>с</strong>л<strong>о</strong><strong>в</strong>ий<br />

для эт<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>а</strong>. В<strong>о</strong> <strong>в</strong>ремя м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых<br />

лёт<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дит н<strong>а</strong><strong>к</strong><strong>а</strong>пли<strong>в</strong><strong>а</strong>ние их<br />

труп<strong>о</strong><strong>в</strong> или <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>тдельных <strong>с</strong><strong>к</strong>елетных<br />

ч<strong>а</strong><strong>с</strong>тей, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong> -б<strong>о</strong>льших <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>а</strong>х, причем<br />

<strong>в</strong> у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях, <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>с</strong>т<strong>в</strong>ующих их з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нению:<br />

<strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, где <strong>с</strong><strong>к</strong><strong>а</strong>пли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>к</strong>и и <strong>с</strong><strong>о</strong>зд<strong>а</strong>ют<strong>с</strong>я<br />

у<strong>с</strong>л<strong>о</strong><strong>в</strong>ия для их ф<strong>о</strong><strong>с</strong><strong>с</strong>илиз<strong>а</strong>ции (<strong>в</strong> <strong>о</strong>зер<strong>а</strong>х,<br />

л<strong>а</strong>гун<strong>а</strong>х, лим<strong>а</strong>н<strong>а</strong>х, з<strong>а</strong><strong>в</strong><strong>о</strong>дях ре<strong>к</strong>"и их дельт<strong>а</strong>х).<br />

Эт<strong>о</strong>т путь <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ения ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> результ<strong>а</strong>те<br />

м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых лёт<strong>о</strong><strong>в</strong> <strong>о</strong>тдельных <strong>в</strong>ид<strong>о</strong><strong>в</strong>, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>,<br />

игр<strong>а</strong>л <strong>в</strong><strong>а</strong>жную р<strong>о</strong>ль. Т<strong>а</strong><strong>к</strong>им путем <strong>в</strong><strong>о</strong>зни<strong>к</strong>ли<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющие<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и <strong>о</strong>пределенных, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

немн<strong>о</strong>гих, <strong>в</strong>ид<strong>о</strong><strong>в</strong>. К эт<strong>о</strong>й <strong>к</strong><strong>а</strong>тег<strong>о</strong>рии <strong>с</strong>ледует <strong>о</strong>тне<strong>с</strong>ти,<br />

н<strong>а</strong>пример, ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждение <strong>в</strong>ерхнеперм<strong>с</strong><strong>к</strong>их<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. п<strong>о</strong> р. С<strong>о</strong>яне,<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения "мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

В. Сибири — тургин<strong>с</strong><strong>к</strong><strong>о</strong>й и у<strong>с</strong>ть-б<strong>а</strong>лей<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ф<strong>а</strong>уны, н<strong>а</strong><strong>к</strong><strong>о</strong>нец, третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> ми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых<br />

<strong>о</strong>тл<strong>о</strong>жениях Ст<strong>а</strong><strong>в</strong>р<strong>о</strong>п<strong>о</strong>лья н<strong>а</strong> С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>зе<br />

и, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, мн<strong>о</strong>гие другие. В<strong>о</strong> <strong>в</strong><strong>с</strong>ех этих<br />

ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных э<strong>к</strong>земпляр<strong>о</strong><strong>в</strong><br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong>, чт<strong>о</strong> <strong>с</strong> <strong>о</strong>че<strong>в</strong>идн<strong>о</strong><strong>с</strong>тью <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует<br />

<strong>о</strong>б их м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong><strong>с</strong>ти.<br />

Н<strong>а</strong>ряду <strong>с</strong> <strong>с</strong>ерийными <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми <strong>о</strong>пределенных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> этих же ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждениях мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и и <strong>о</strong>дин<strong>о</strong>чных э<strong>к</strong>земпляр<strong>о</strong><strong>в</strong><br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Суще<strong>с</strong>т<strong>в</strong>уют т<strong>а</strong><strong>к</strong>же целые<br />

ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящие из уни<strong>к</strong><strong>а</strong>льных э<strong>к</strong>земпляр<strong>о</strong><strong>в</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В<strong>о</strong> <strong>в</strong><strong>с</strong>ех этих <strong>с</strong>луч<strong>а</strong>ях<br />

нет <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ть н<strong>а</strong>личия<br />

м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых лёт<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>с</strong>чит<strong>а</strong>ть<br />

и<strong>с</strong>т<strong>о</strong>чни<strong>к</strong><strong>а</strong>ми и у<strong>с</strong>л<strong>о</strong><strong>в</strong>иями <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> не м<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>е<br />

р<strong>а</strong>змн<strong>о</strong>жение н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>а</strong> <strong>с</strong>луч<strong>а</strong>йный, <strong>в</strong>ыб<strong>о</strong>р<strong>о</strong>чный<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер п<strong>о</strong>п<strong>а</strong>д<strong>а</strong>ния н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> у<strong>с</strong>л<strong>о</strong><strong>в</strong>ия<br />

з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нения. Т<strong>а</strong><strong>к</strong><strong>о</strong>й путь ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых я<strong>в</strong>ляет<strong>с</strong>я <strong>с</strong><strong>а</strong>мым<br />

<strong>в</strong><strong>а</strong>жным и н<strong>а</strong>иб<strong>о</strong>лее шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненным.<br />

Случ<strong>а</strong>йный х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нения н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лн<strong>о</strong> <strong>с</strong>удить <strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>е<br />

ре<strong>а</strong>льн<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong><strong>в</strong>шей ф<strong>а</strong>уны. Пример<strong>о</strong>м<br />

т<strong>а</strong><strong>к</strong>их <strong>в</strong>ыб<strong>о</strong>р<strong>о</strong>чных, н<strong>а</strong>иб<strong>о</strong>лее б<strong>о</strong>г<strong>а</strong>тых<br />

<strong>в</strong>ид<strong>а</strong>ми ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> я<strong>в</strong>ляют<strong>с</strong>я<br />

40<br />

http://jurassic.ru/<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения перм<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых Ур<strong>а</strong>л<strong>а</strong><br />

(н<strong>а</strong>пример, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>), мн<strong>о</strong>гих третичных ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

(н<strong>а</strong>пример, <strong>о</strong>лиг<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

из Р<strong>о</strong>тт<strong>а</strong> <strong>в</strong> Герм<strong>а</strong>нии), н<strong>о</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тные<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения юр<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>не (К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у), з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющие <strong>о</strong>чень<br />

б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> (измеряемых <strong>с</strong><strong>о</strong>тнями),<br />

прит<strong>о</strong>м пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленных лишь единичными <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми.<br />

М<strong>о</strong>жн<strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> <strong>о</strong>тн<strong>о</strong>шении р<strong>а</strong>змер<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ня<strong>в</strong>ших<strong>с</strong>я<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> и их п<strong>о</strong>лн<strong>о</strong>ты, т. е. т<strong>о</strong>й или<br />

друг<strong>о</strong>й <strong>с</strong>тепени их фр<strong>а</strong>гмент<strong>а</strong>рн<strong>о</strong><strong>с</strong>ти. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong>р<strong>а</strong>йне фр<strong>а</strong>гмент<strong>а</strong>рн<strong>о</strong>:<br />

<strong>в</strong> лучшем <strong>с</strong>луч<strong>а</strong>е <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я целые<br />

<strong>к</strong>рылья, <strong>о</strong>бычн<strong>о</strong> же п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>ги имеют дел<strong>о</strong><br />

<strong>с</strong> <strong>о</strong>бры<strong>в</strong><strong>к</strong><strong>а</strong>ми <strong>к</strong>рылье<strong>в</strong>, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми ч<strong>а</strong><strong>с</strong>тями<br />

тел<strong>а</strong>, <strong>о</strong>тдельными <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями или <strong>с</strong><strong>к</strong>лерит<strong>а</strong>ми<br />

(н<strong>а</strong>пример, <strong>с</strong><strong>о</strong> <strong>с</strong>тернит<strong>а</strong>ми груди жу<strong>к</strong><strong>о</strong><strong>в</strong>).<br />

Г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> реже <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я целые или п<strong>о</strong>чти<br />

целые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Суще<strong>с</strong>т<strong>в</strong>уют ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные п<strong>о</strong>чти<br />

и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> из н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых мел<strong>к</strong>их р<strong>а</strong>змер<strong>о</strong><strong>в</strong>,<br />

н<strong>а</strong>пример, ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения <strong>в</strong>ерхнеперм<strong>с</strong><strong>к</strong>их<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> Кузнец<strong>к</strong><strong>о</strong>м б<strong>а</strong><strong>с</strong><strong>с</strong>ейне (С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II) или три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> Средней Азии.<br />

В этих <strong>с</strong>луч<strong>а</strong>ях з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нение <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>,<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<strong>о</strong> п<strong>о</strong><strong>с</strong>ле перен<strong>о</strong><strong>с</strong><strong>а</strong> п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>а</strong>ми<br />

и мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>й их <strong>о</strong>т<strong>с</strong><strong>о</strong>ртир<strong>о</strong><strong>в</strong><strong>к</strong>и <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енными<br />

мел<strong>к</strong><strong>о</strong>зерни<strong>с</strong>тыми <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>к</strong><strong>а</strong>ми. С<strong>о</strong>хр<strong>а</strong>нение<br />

целых тел н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, чт<strong>о</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждениях тип<strong>а</strong> К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у, н<strong>а</strong><strong>о</strong>б<strong>о</strong>р<strong>о</strong>т,<br />

п<strong>о</strong>чти н<strong>а</strong><strong>в</strong>ерня<strong>к</strong><strong>а</strong> <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ил<strong>о</strong><strong>с</strong>ь <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем<br />

без перен<strong>о</strong><strong>с</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых перед пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>о</strong>м их<br />

ф<strong>о</strong><strong>с</strong><strong>с</strong>илиз<strong>а</strong>ции. Об эт<strong>о</strong>м мы м<strong>о</strong>жем <strong>с</strong>удить, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>я<strong>с</strong>ь<br />

н<strong>а</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>тере <strong>с</strong><strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ти н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых;<br />

<strong>в</strong> ряде <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ть н<strong>а</strong><strong>с</strong>т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>а</strong>,<br />

чт<strong>о</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью и<strong>с</strong><strong>к</strong>люч<strong>а</strong>ет<strong>с</strong>я их перен<strong>о</strong><strong>с</strong><br />

<strong>в</strong><strong>о</strong>дным п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>о</strong>м.<br />

Случ<strong>а</strong>йный х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нения н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

ин<strong>о</strong>гд<strong>а</strong> при<strong>в</strong><strong>о</strong>дит <strong>к</strong> т<strong>о</strong>му, чт<strong>о</strong> их <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

мы н<strong>а</strong>х<strong>о</strong>дим <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>е <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем не<strong>о</strong>жид<strong>а</strong>нных и<br />

не<strong>о</strong>бычных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong>, н<strong>а</strong>пример,<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и з<strong>а</strong>дних н<strong>о</strong>г прыг<strong>а</strong>ющих прям<strong>о</strong><strong>к</strong>рылых<br />

<strong>с</strong>реди <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> третичных глуб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>дных<br />

рыб <strong>в</strong> 'мел<strong>к</strong><strong>о</strong>зерни<strong>с</strong>тых п<strong>о</strong>р<strong>о</strong>д<strong>а</strong>х я<strong>в</strong>н<strong>о</strong><br />

м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>, не<strong>к</strong><strong>о</strong>нтинент<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>а</strong> (<strong>с</strong><strong>а</strong>рм<strong>а</strong>т<strong>с</strong><strong>к</strong>ие<br />

<strong>о</strong>тл<strong>о</strong>жения С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>), <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичные<br />

н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> н<strong>а</strong>земных же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых<br />

<strong>с</strong>реди м<strong>о</strong>р<strong>с</strong><strong>к</strong>их <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong>. В<strong>о</strong> <strong>в</strong><strong>с</strong>ех этих <strong>с</strong>луч<strong>а</strong>ях<br />

<strong>о</strong>чень б<strong>о</strong>льш<strong>а</strong>я мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong><strong>а</strong>я пр<strong>о</strong>чн<strong>о</strong><strong>с</strong>ть и<br />

<strong>а</strong>б<strong>с</strong><strong>о</strong>лютн<strong>а</strong>я лег<strong>к</strong><strong>о</strong><strong>с</strong>ть <strong>с</strong><strong>к</strong>елетных ч<strong>а</strong><strong>с</strong>тей н<strong>а</strong>з<strong>в</strong><strong>а</strong>нных<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых п<strong>о</strong>з<strong>в</strong><strong>о</strong>лили им перене<strong>с</strong>ти длительную<br />

тр<strong>а</strong>н<strong>с</strong>п<strong>о</strong>ртир<strong>о</strong><strong>в</strong><strong>к</strong>у (<strong>в</strong><strong>о</strong>здушными и <strong>в</strong><strong>о</strong>дными<br />

п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>а</strong>ми) и п<strong>о</strong>п<strong>а</strong><strong>с</strong>ть <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем не<strong>о</strong>бычные<br />

для <strong>о</strong>бит<strong>а</strong>телей <strong>с</strong>уши у<strong>с</strong>л<strong>о</strong><strong>в</strong>ия з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нения.<br />

О<strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти труп<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых —их м<strong>а</strong>лые<br />

р<strong>а</strong>змеры, пр<strong>о</strong>чн<strong>о</strong><strong>с</strong>ть <strong>с</strong><strong>к</strong>елет<strong>а</strong> и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые

другие мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong><strong>а</strong> —<strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ли<strong>в</strong><strong>а</strong>ют<br />

н<strong>а</strong>х<strong>о</strong>ждение <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емыми <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> ли<strong>с</strong>т<strong>о</strong>нргих (Conchostraca)<br />

и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>а</strong><strong>к</strong>уш<strong>к</strong><strong>о</strong><strong>в</strong>ых (Ostracoda)<br />

р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>т<strong>о</strong>ж<strong>а</strong>берных м<strong>о</strong>ллю<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых плечен<strong>о</strong>гих (Lingulidae).<br />

Эт<strong>о</strong> <strong>о</strong>пределяет<strong>с</strong>я прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> р<strong>а</strong>змер<strong>а</strong>ми<br />

н<strong>а</strong>з<strong>в</strong><strong>а</strong>нных р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, м<strong>о</strong>ллю<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong> и<br />

плечен<strong>о</strong>гих и <strong>о</strong>бит<strong>а</strong>нием их <strong>в</strong> тех же <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х,<br />

где пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<strong>о</strong> з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нение <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Среди <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> других жи<strong>в</strong><strong>о</strong>тных, <strong>в</strong><strong>с</strong>треч<strong>а</strong>ющих<strong>с</strong>я<br />

реже, прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>с</strong>ледует уп<strong>о</strong>мянуть<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и рыб, <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ти<strong>с</strong>тых (н<strong>а</strong>пример, Clupeidae). О<strong>с</strong>т<strong>а</strong>т<strong>к</strong>и рыб<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> чешуей, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

<strong>в</strong> <strong>о</strong>тдельных ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждениях не<strong>о</strong>ген<strong>о</strong><strong>в</strong>ых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>чень <strong>о</strong>бычн<strong>а</strong> и <strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>ет<br />

п<strong>о</strong>чти <strong>в</strong><strong>с</strong>е н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и <strong>в</strong> <strong>о</strong>пределенных г<strong>о</strong>риз<strong>о</strong>нт<strong>а</strong>х<br />

(н<strong>а</strong>пример, <strong>в</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>г<strong>а</strong>не ми<strong>о</strong>цен<strong>а</strong> Ст<strong>а</strong><strong>в</strong>р<strong>о</strong>п<strong>о</strong>лья).<br />

Ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ме<strong>с</strong>тн<strong>о</strong> <strong>с</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мыми <strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шие<strong>с</strong>я, п<strong>о</strong>чти целые <strong>с</strong><strong>к</strong>елеты<br />

рыб; т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы из<strong>в</strong>е<strong>с</strong>тные юр<strong>с</strong><strong>к</strong>ие «рыбные»<br />

или «бум<strong>а</strong>жные <strong>с</strong>л<strong>а</strong>нцы» К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у, мн<strong>о</strong>гие мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения М<strong>о</strong>нг<strong>о</strong>лии и З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лья.<br />

Пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение этих ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

ин<strong>о</strong>е, и <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>о</strong>н<strong>о</strong> <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong> <strong>с</strong> пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>а</strong>ми<br />

пери<strong>о</strong>диче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>в</strong>ы<strong>с</strong>ых<strong>а</strong>ния и п<strong>о</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>тл<strong>о</strong>жения мел<strong>к</strong><strong>о</strong>зерни<strong>с</strong>т<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х<br />

<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong>: рыбы гибли <strong>о</strong>т удушья<br />

и пере<strong>с</strong>ых<strong>а</strong>ния <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>, н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые з<strong>а</strong>н<strong>о</strong><strong>с</strong>или<strong>с</strong>ь<br />

<strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем <strong>в</strong>етр<strong>о</strong>м и бы<strong>с</strong>тр<strong>о</strong> з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>няли<strong>с</strong>ь,<br />

п<strong>о</strong>чти без п<strong>о</strong><strong>с</strong>ледующег<strong>о</strong> перен<strong>о</strong><strong>с</strong><strong>а</strong> п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>а</strong>ми.<br />

Кр<strong>о</strong>ме <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> жи<strong>в</strong><strong>о</strong>тных, и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>ют<strong>с</strong>я <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми<br />

р<strong>а</strong><strong>с</strong>тений. Н<strong>а</strong>иб<strong>о</strong>лее ч<strong>а</strong><strong>с</strong>ты н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и ч<strong>а</strong><strong>с</strong>тиц<br />

дре<strong>в</strong>е<strong>с</strong>ины, <strong>о</strong>бры<strong>в</strong><strong>к</strong><strong>о</strong><strong>в</strong> ли<strong>с</strong>тье<strong>в</strong> и целых<br />

неб<strong>о</strong>льших 'пл<strong>о</strong>д<strong>о</strong><strong>в</strong> или <strong>с</strong>емян р<strong>а</strong><strong>с</strong>тений. С<strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ть<br />

р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> з<strong>а</strong><strong>в</strong>и<strong>с</strong>ит <strong>о</strong>т<br />

их мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong>их <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong> и <strong>о</strong>т <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти<br />

перен<strong>о</strong><strong>с</strong><strong>а</strong> их <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Этим <strong>о</strong>бъя<strong>с</strong>няет<strong>с</strong>я <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие или ред<strong>к</strong><strong>о</strong><strong>с</strong>ть<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> <strong>с</strong>л<strong>о</strong>ях, б<strong>о</strong>г<strong>а</strong>тых <strong>к</strong>рупными р<strong>а</strong><strong>с</strong>тительными<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми: ли<strong>с</strong>тьями, <strong>в</strong>ет<strong>в</strong>ями, <strong>с</strong>теблями<br />

и <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми р<strong>а</strong><strong>с</strong>тений; зде<strong>с</strong>ь н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

п<strong>о</strong>п<strong>а</strong>д<strong>а</strong>ют<strong>с</strong>я ред<strong>к</strong><strong>о</strong>, уд<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong>х<strong>о</strong>дить лишь<br />

<strong>к</strong>рупные и тяжелые ф<strong>о</strong>рмы или <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> б<strong>о</strong>льшие<br />

<strong>к</strong>рылья. З<strong>а</strong>т<strong>о</strong> н<strong>а</strong>личие <strong>в</strong> <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>о</strong>чн<strong>о</strong>й п<strong>о</strong>р<strong>о</strong>де<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных <strong>с</strong>емян р<strong>а</strong><strong>с</strong>тений <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>ет<strong>с</strong>я и н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong>ми <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

В з<strong>а</strong><strong>к</strong>лючение <strong>с</strong>ледует уп<strong>о</strong>мянуть <strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>знейшем<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нении и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>в</strong> <strong>о</strong>т<strong>в</strong>ерде<strong>в</strong>шей <strong>с</strong>м<strong>о</strong>ле х<strong>в</strong><strong>о</strong>йных р<strong>а</strong><strong>с</strong>тений —<br />

<strong>в</strong> янт<strong>а</strong>ре и <strong>к</strong><strong>о</strong>п<strong>а</strong>ле. См<strong>о</strong>л<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых х<strong>в</strong><strong>о</strong>йных<br />

<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ет <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>тью не пере<strong>к</strong>ри<strong>с</strong>т<strong>а</strong>ллиз<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ть<strong>с</strong>я<br />

при <strong>в</strong>ы<strong>с</strong>ых<strong>а</strong>нии и при упл<strong>о</strong>тнении <strong>с</strong><strong>о</strong>хр<strong>а</strong>нять<br />

<strong>с</strong>те<strong>к</strong>л<strong>о</strong><strong>в</strong>идную, пр<strong>о</strong>зр<strong>а</strong>чную <strong>с</strong>тру<strong>к</strong>туру.<br />

Эт<strong>о</strong> п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет без з<strong>а</strong>труднений р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть<br />

<strong>в</strong><strong>с</strong>е <strong>в</strong><strong>к</strong>лючения (ин<strong>к</strong>люзы), <strong>к</strong><strong>о</strong>т<strong>о</strong>рые п<strong>о</strong>п<strong>а</strong>ли<br />

<strong>в</strong> <strong>с</strong>м<strong>о</strong>лу р<strong>а</strong><strong>с</strong>тения при ее <strong>в</strong>ыте<strong>к</strong><strong>а</strong>нии и е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>м<br />

з<strong>а</strong>т<strong>в</strong>ерде<strong>в</strong><strong>а</strong>нии, пре<strong>в</strong>р<strong>а</strong>щении <strong>в</strong> пл<strong>о</strong>тн<strong>о</strong>е,<br />

т<strong>в</strong>ерд<strong>о</strong>е тел<strong>о</strong> — янт<strong>а</strong>рь или <strong>к</strong><strong>о</strong>п<strong>а</strong>л. В результ<strong>а</strong>те<br />

эт<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>а</strong> <strong>с</strong><strong>о</strong>зд<strong>а</strong>ли<strong>с</strong>ь, <strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>ж<strong>а</strong>я<strong>с</strong>ь, е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енные «п<strong>о</strong><strong>с</strong>т<strong>о</strong>янные преп<strong>а</strong>р<strong>а</strong>ты»<br />

из <strong>с</strong>м<strong>о</strong>лы дре<strong>в</strong>них р<strong>а</strong><strong>с</strong>тений. Пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong><strong>с</strong>ть<br />

янт<strong>а</strong>ря или <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бы<strong>в</strong><strong>а</strong>ет <strong>о</strong>чень<br />

<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>й, т<strong>а</strong><strong>к</strong> чт<strong>о</strong> мн<strong>о</strong>гие <strong>в</strong><strong>к</strong>лючения <strong>в</strong>идны<br />

<strong>с</strong> и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong>й п<strong>о</strong>лн<strong>о</strong>т<strong>о</strong>й и я<strong>с</strong>н<strong>о</strong><strong>с</strong>тью. Приним<strong>а</strong>я<br />

<strong>в</strong><strong>о</strong> <strong>в</strong>ним<strong>а</strong>ние б<strong>о</strong>льшую дре<strong>в</strong>н<strong>о</strong><strong>с</strong>ть янт<strong>а</strong>ря<br />

(д<strong>о</strong> <strong>в</strong>ерхнег<strong>о</strong> мел<strong>а</strong>!), <strong>о</strong>че<strong>в</strong>идн<strong>а</strong> гр<strong>о</strong>м<strong>а</strong>дн<strong>а</strong>я ценн<strong>о</strong><strong>с</strong>ть<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния янт<strong>а</strong>рных <strong>в</strong><strong>к</strong>лючений для<br />

п<strong>о</strong>зн<strong>а</strong>ния ф<strong>а</strong>уны пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong>. Зн<strong>а</strong>чение и ценн<strong>о</strong><strong>с</strong>ть<br />

этих м<strong>а</strong>тери<strong>а</strong>л<strong>о</strong><strong>в</strong> п<strong>о</strong> ф<strong>а</strong>уне янт<strong>а</strong>ря <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

их <strong>о</strong>дн<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>о</strong>нним <strong>о</strong>тб<strong>о</strong>р<strong>о</strong>м: <strong>о</strong>ни<br />

<strong>о</strong><strong>с</strong><strong>в</strong>ещ<strong>а</strong>ют лишь <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong><br />

ле<strong>с</strong><strong>а</strong>, прит<strong>о</strong>м з<strong>а</strong><strong>к</strong>люч<strong>а</strong>я преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

неб<strong>о</strong>льших и мельч<strong>а</strong>йших н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>о</strong>бит<strong>а</strong><strong>в</strong>ших<br />

н<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>х, <strong>к</strong><strong>о</strong>ре дере<strong>в</strong>ье<strong>в</strong>, реже н<strong>а</strong><br />

п<strong>о</strong>ч<strong>в</strong>е или н<strong>а</strong> ли<strong>с</strong>тьях. Крупных или <strong>с</strong><strong>в</strong>яз<strong>а</strong>нных<br />

<strong>с</strong> <strong>о</strong>т<strong>к</strong>рытыми л<strong>а</strong>ндш<strong>а</strong>фт<strong>а</strong>ми н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых мы <strong>в</strong><br />

янт<strong>а</strong>ре не н<strong>а</strong>х<strong>о</strong>дим. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> янт<strong>а</strong>ре <strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>ют<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и р<strong>а</strong><strong>с</strong>тений<br />

(д<strong>о</strong> пре<strong>к</strong>р<strong>а</strong><strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няющих<strong>с</strong>я ц<strong>в</strong>ет<strong>к</strong><strong>о</strong><strong>в</strong>!) и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

жи<strong>в</strong><strong>о</strong>тных —п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong>.<br />

Ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>с</strong>м<strong>о</strong>л —<br />

янт<strong>а</strong>ря и <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong>— д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> ред<strong>к</strong>и: н<strong>а</strong>ши зн<strong>а</strong>ния<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых этих ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я из<strong>в</strong>е<strong>с</strong>тным<br />

б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>им янт<strong>а</strong>рем.<br />

Зн<strong>а</strong>чение н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

Зн<strong>а</strong>чение н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, при их <strong>о</strong>билии, р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зии<br />

и шир<strong>о</strong>ч<strong>а</strong>йшем р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нении, <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong><br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> и <strong>о</strong>чень <strong>в</strong><strong>а</strong>жн<strong>о</strong> для чел<strong>о</strong><strong>в</strong>е<strong>к</strong><strong>а</strong>.<br />

Вредн<strong>а</strong>я деятельн<strong>о</strong><strong>с</strong>ть н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти и<strong>с</strong><strong>к</strong>уп<strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й п<strong>о</strong>льз<strong>о</strong>й; д<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

<strong>в</strong><strong>с</strong>п<strong>о</strong>мнить прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> шел<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong>, пчел<strong>о</strong><strong>в</strong><strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong>,<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — <strong>о</strong>пылителей р<strong>а</strong><strong>с</strong>тений,<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых-п<strong>а</strong>р<strong>а</strong>зит<strong>о</strong><strong>в</strong>, <strong>в</strong>редителей и, н<strong>а</strong><strong>к</strong><strong>о</strong>нец,<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых—пр<strong>о</strong>из<strong>в</strong><strong>о</strong>дителей л<strong>а</strong><strong>к</strong><strong>а</strong>, <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>к</strong>р<strong>а</strong><strong>с</strong>ителей и т. д.<br />

С<strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>е зн<strong>а</strong>чение имеют н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Я<strong>в</strong>ляя<strong>с</strong>ь <strong>о</strong>бит<strong>а</strong>телями <strong>с</strong>уши,<br />

прит<strong>о</strong>м <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>я и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> шир<strong>о</strong><strong>к</strong>им р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нением<br />

<strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тях, н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляют лег<strong>к</strong><strong>о</strong><br />

и т<strong>о</strong>чн<strong>о</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ть р<strong>а</strong>зличные р<strong>а</strong>й<strong>о</strong>ны<br />

и <strong>с</strong>т<strong>а</strong>ции. Их <strong>о</strong>билие и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зие, н<strong>а</strong><strong>к</strong><strong>о</strong>нец,<br />

мел<strong>к</strong>ие <strong>а</strong>б<strong>с</strong><strong>о</strong>лютные р<strong>а</strong>змеры тел<strong>а</strong> дел<strong>а</strong>ют их<br />

<strong>о</strong>чень уд<strong>о</strong>бным и <strong>в</strong><strong>а</strong>жным 'м<strong>а</strong>тери<strong>а</strong>л<strong>о</strong>м при<br />

э<strong>к</strong><strong>о</strong>л<strong>о</strong>г<strong>о</strong>-ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>й х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>е люб<strong>о</strong>г<strong>о</strong><br />

уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong> <strong>с</strong>уши. Этим <strong>о</strong>пределяет<strong>с</strong>я и их<br />

зн<strong>а</strong>чение для ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й би<strong>о</strong><strong>с</strong>тр<strong>а</strong>типр<strong>а</strong>фии;<br />

ред<strong>к</strong><strong>о</strong><strong>с</strong>ть <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>к</strong><strong>о</strong>нтинент<strong>а</strong>льных<br />

<strong>о</strong>тл<strong>о</strong>жениях <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> п<strong>о</strong>дчер<strong>к</strong>и<strong>в</strong><strong>а</strong>ет п<strong>о</strong>тен-<br />

41<br />

http://jurassic.ru/

ци<strong>а</strong>льную ценн<strong>о</strong><strong>с</strong>ть и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Е<strong>с</strong>ли зн<strong>а</strong>чение и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

для <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фии не м<strong>о</strong>жет <strong>в</strong>ыз<strong>в</strong><strong>а</strong>ть<br />

принципи<strong>а</strong>льных <strong>в</strong><strong>о</strong>зр<strong>а</strong>жений, т<strong>о</strong> ф<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

применение результ<strong>а</strong>т<strong>о</strong><strong>в</strong> л<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний и <strong>в</strong><strong>о</strong><strong>о</strong>бще и<strong>с</strong>п<strong>о</strong>льз<strong>о</strong><strong>в</strong><strong>а</strong>ние и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых при ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

р<strong>а</strong>б<strong>о</strong>т<strong>а</strong>х м<strong>о</strong>жет быть не <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем я<strong>с</strong>ным. Причины<br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><strong>с</strong>ти <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующих<br />

д<strong>а</strong>нных п<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong>: д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> из<strong>в</strong>е<strong>с</strong>тным пр<strong>а</strong><strong>в</strong><strong>о</strong>м<br />

<strong>с</strong>чит<strong>а</strong>ют<strong>с</strong>я ред<strong>к</strong>ими и <strong>с</strong><strong>а</strong>ми н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

чуть ли не единичными и и<strong>с</strong><strong>к</strong>лючительными.<br />

Дей<strong>с</strong>т<strong>в</strong>ительн<strong>о</strong>, д<strong>о</strong> 20-х г<strong>о</strong>д<strong>о</strong><strong>в</strong> н<strong>а</strong>шег<strong>о</strong> <strong>в</strong>е<strong>к</strong><strong>а</strong> ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong> террит<strong>о</strong>рии<br />

С<strong>о</strong><strong>в</strong>ет<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> С<strong>о</strong>юз<strong>а</strong> н<strong>а</strong><strong>с</strong>читы<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь<br />

единиц<strong>а</strong>ми, <strong>а</strong> <strong>с</strong><strong>а</strong>ми <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — немн<strong>о</strong>гими<br />

де<strong>с</strong>ят<strong>к</strong><strong>а</strong>ми. Одн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя<br />

чи<strong>с</strong>л<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тных т<strong>о</strong>че<strong>к</strong>, где были <strong>о</strong>бн<strong>а</strong>ружены<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, приближ<strong>а</strong>ет<strong>с</strong>я <strong>к</strong><br />

300, <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> же <strong>с</strong><strong>о</strong>бр<strong>а</strong>нных и изученных<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет уже мн<strong>о</strong>гих<br />

де<strong>с</strong>ят<strong>к</strong><strong>о</strong><strong>в</strong> ты<strong>с</strong>яч. Эти <strong>о</strong>бширные м<strong>а</strong>тери<strong>а</strong>лы<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>лили д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> п<strong>о</strong>лн<strong>о</strong> <strong>о</strong>х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ть<br />

ф<strong>а</strong>уны н<strong>а</strong>шей террит<strong>о</strong>рии <strong>в</strong> р<strong>а</strong>зличные м<strong>о</strong>менты<br />

ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени, н<strong>а</strong>чин<strong>а</strong>я <strong>с</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>.<br />

З<strong>а</strong> <strong>в</strong><strong>с</strong>е <strong>в</strong>ремя <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>о</strong>чень меняли<strong>с</strong>ь: ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы р<strong>а</strong>зличных пери<strong>о</strong>д<strong>о</strong><strong>в</strong> и эп<strong>о</strong>х<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я не т<strong>о</strong>ль<strong>к</strong><strong>о</strong> друг <strong>о</strong>т друг<strong>а</strong>, н<strong>о</strong><br />

и <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ют чи<strong>с</strong>т<strong>о</strong> ге<strong>о</strong>гр<strong>а</strong>ф<strong>о</strong>-<strong>к</strong>лим<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти р<strong>а</strong>зных <strong>о</strong>бл<strong>а</strong><strong>с</strong>тей.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong><strong>а</strong> изученн<strong>о</strong><strong>с</strong>ти<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

р<strong>а</strong>зличных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

К<strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> я<strong>в</strong>ляет<strong>с</strong>я <strong>с</strong><strong>а</strong>м<strong>о</strong>й<br />

дре<strong>в</strong>ней, е<strong>с</strong>ли не <strong>с</strong>чит<strong>а</strong>ть единичных н<strong>а</strong>х<strong>о</strong>д<strong>о</strong><strong>к</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> де<strong>в</strong><strong>о</strong>не. Лучше <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong>н<strong>а</strong> изучен<strong>а</strong><br />

<strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе и С. Амери<strong>к</strong>е, н<strong>а</strong> террит<strong>о</strong>рии же<br />

СССР <strong>о</strong>н<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>а</strong> еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>. Лишь<br />

<strong>в</strong> п<strong>о</strong><strong>с</strong>ледние г<strong>о</strong>ды <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> п<strong>о</strong>лн<strong>о</strong> изучены<br />

д<strong>а</strong><strong>в</strong>н<strong>о</strong> <strong>о</strong>т<strong>к</strong>рытые ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения Сибири<br />

<strong>в</strong> <strong>о</strong>тл<strong>о</strong>жениях нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong><strong>в</strong>иты Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. В цел<strong>о</strong>м <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льные<br />

ф<strong>а</strong>уны х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я <strong>о</strong>билием п<strong>а</strong>ле<strong>о</strong>ди<strong>к</strong>ти<strong>о</strong>птер<strong>о</strong>ид<strong>о</strong><strong>в</strong>,<br />

т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ых и пр<strong>о</strong>т<strong>о</strong>бл<strong>а</strong>тт<strong>о</strong>идей;<br />

<strong>к</strong> ним при<strong>с</strong><strong>о</strong>единяют<strong>с</strong>я не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые группы прям<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных,<br />

дре<strong>в</strong>ние п<strong>о</strong>д<strong>о</strong>тряды <strong>с</strong>тре<strong>к</strong><strong>о</strong>з<br />

(Meganisoptera) и мег<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>птер<strong>о</strong>идеи. Н<strong>а</strong><br />

эт<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м ф<strong>о</strong>не п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong><br />

немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители мн<strong>о</strong>гих других<br />

групп, д<strong>о</strong><strong>с</strong>тигших р<strong>а</strong><strong>с</strong>ц<strong>в</strong>ет<strong>а</strong> п<strong>о</strong>зднее (дре<strong>в</strong>ние<br />

Homoptera, Mecoptera, Ephemeroptera). Д<strong>о</strong><br />

<strong>с</strong>их п<strong>о</strong>р <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т<strong>к</strong>рытым <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong> <strong>о</strong> <strong>к</strong>лим<strong>а</strong>ти­<br />

42<br />

http://jurassic.ru/<br />

че<strong>с</strong><strong>к</strong><strong>о</strong>м р<strong>а</strong>й<strong>о</strong>нир<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных ф<strong>а</strong>ун:<br />

р<strong>а</strong>зрешение ег<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong> изучением <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> Анг<strong>а</strong>риды и Г<strong>о</strong>нд<strong>в</strong><strong>а</strong>ны.<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> изучен<strong>а</strong> г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> п<strong>о</strong>лнее<br />

<strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>й. Он<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>а</strong> п<strong>о</strong> ряду ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

<strong>в</strong> Е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong><strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти СССР<br />

(Приур<strong>а</strong>лье и Ур<strong>а</strong>л, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.), <strong>в</strong><br />

Ю. Сибири (Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн). Перм<strong>с</strong><strong>к</strong>ие<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения из<strong>в</strong>е<strong>с</strong>тны и <strong>в</strong> С. Амери<strong>к</strong>е,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и 3. Е<strong>в</strong>р<strong>о</strong>пе, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лные<br />

м<strong>а</strong>тери<strong>а</strong>лы <strong>с</strong><strong>о</strong>бр<strong>а</strong>ны п<strong>о</strong> перм<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уне<br />

именн<strong>о</strong> н<strong>а</strong> н<strong>а</strong>шей террит<strong>о</strong>рии. Н<strong>а</strong>иб<strong>о</strong>лее <strong>в</strong><strong>а</strong>жн<strong>о</strong>й<br />

черт<strong>о</strong>й перм<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уны я<strong>в</strong>ляет<strong>с</strong>я бурн<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц, Mecoptera (ф<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong>и,<br />

е<strong>с</strong>ли не <strong>с</strong>чит<strong>а</strong>ть един<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>й<br />

н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и Metropatridae, <strong>в</strong>пер<strong>в</strong>ые п<strong>о</strong>я<strong>в</strong>и<strong>в</strong>ших<strong>с</strong>я<br />

<strong>в</strong> перм<strong>с</strong><strong>к</strong>ий пери<strong>о</strong>д) и р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых<br />

(Homoptera), <strong>а</strong> т<strong>а</strong><strong>к</strong>же пер<strong>в</strong><strong>о</strong>е п<strong>о</strong>я<strong>в</strong>ление же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых<br />

(Coleoptera).<br />

Очень х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> для перми <strong>о</strong>билие р<strong>а</strong>зличных<br />

дре<strong>в</strong>не<strong>к</strong>рылых из н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong> мег<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>птер<strong>о</strong>идеи<br />

(Megasecopteroidea), <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти<br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Protohymenoptera, и ряд<strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

<strong>с</strong>тре<strong>к</strong><strong>о</strong>з (Protanisoptera, Permanisoptera). Т<strong>а</strong><strong>к</strong>же<br />

чи<strong>с</strong>т<strong>о</strong> перм<strong>с</strong><strong>к</strong>им я<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>тряд Protelytroptera.<br />

Три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> изучен<strong>а</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong>: ее<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения <strong>к</strong>р<strong>а</strong>йне немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны и<br />

из<strong>в</strong>е<strong>с</strong>тны лишь из Ср. Азии, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

и немн<strong>о</strong>гих пун<strong>к</strong>т<strong>о</strong><strong>в</strong> <strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ть три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ую ф<strong>а</strong>уну з<strong>а</strong>труднительн<strong>о</strong>;<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>ть ее м<strong>о</strong>жн<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>а</strong>мых<br />

<strong>о</strong>бщих черт<strong>а</strong>х. Следует <strong>о</strong>тметить б<strong>о</strong>льш<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых, п<strong>о</strong>я<strong>в</strong>ление пер<strong>в</strong>ых<br />

д<strong>в</strong>у<strong>к</strong>рылых, р<strong>а</strong>зн<strong>о</strong><strong>к</strong>рылых (Heteroptera), <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

немн<strong>о</strong>гих рели<strong>к</strong>т<strong>о</strong><strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

прям<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных перми (Glosselytrodea),<br />

п<strong>о</strong>лн<strong>о</strong>е и<strong>с</strong>чезн<strong>о</strong><strong>в</strong>ение п<strong>а</strong>ле<strong>о</strong>ди<strong>к</strong>ти<strong>о</strong>птер.<br />

Юр<strong>с</strong><strong>к</strong><strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> изучен<strong>а</strong> лучше <strong>в</strong><strong>с</strong>ех других<br />

мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их. О<strong>с</strong><strong>о</strong>бенн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong> д<strong>а</strong>нных был<strong>о</strong><br />

п<strong>о</strong>лучен<strong>о</strong> при изучении нижнеюр<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>в</strong> Ср. Азии (ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения Шур<strong>а</strong>б,<br />

Кизил-Кия и др.) и <strong>в</strong>ерхнеюр<strong>с</strong><strong>к</strong>их <strong>в</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>не<br />

(из<strong>в</strong>е<strong>с</strong>тные ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения <strong>в</strong> г<strong>о</strong>р<strong>а</strong>х К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

З<strong>а</strong> рубеж<strong>о</strong>м юр<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из<strong>в</strong>е<strong>с</strong>тны<br />

лишь из немн<strong>о</strong>гих т<strong>о</strong>че<strong>к</strong> <strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе (Герм<strong>а</strong>нии<br />

и Англии), Кит<strong>а</strong>е, М<strong>о</strong>нг<strong>о</strong>лии и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Одн<strong>а</strong><strong>к</strong><strong>о</strong> гл<strong>а</strong><strong>в</strong>ные и н<strong>а</strong>иб<strong>о</strong>лее б<strong>о</strong>г<strong>а</strong>тые ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

юр<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уны н<strong>а</strong>х<strong>о</strong>дят<strong>с</strong>я н<strong>а</strong> террит<strong>о</strong>рии<br />

СССР. Кр<strong>о</strong>ме <strong>о</strong>тмеченных <strong>в</strong>ыше <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных<br />

юр<strong>с</strong><strong>к</strong>их ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений, <strong>с</strong>ледует у<strong>к</strong><strong>а</strong>з<strong>а</strong>ть<br />

еще н<strong>а</strong> н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых эт<strong>о</strong>г<strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong><br />

<strong>в</strong> 3. Сибири (ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения <strong>в</strong> Кузнец<strong>к</strong><strong>о</strong>м<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейне и Челябин<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.), В. Сибири<br />

(ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения У<strong>с</strong>ть-Б<strong>а</strong>лей и др.), К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>не<br />

(Тург<strong>а</strong>й<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>в</strong>п<strong>а</strong>дин<strong>а</strong>), н<strong>а</strong> К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>зе и <strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

других р<strong>а</strong>й<strong>о</strong>н<strong>а</strong>х. Х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ть юр<strong>с</strong><strong>к</strong>ую<br />

ф<strong>а</strong>уну <strong>с</strong>ледует б<strong>о</strong>г<strong>а</strong>тым р<strong>а</strong>з<strong>в</strong>итием же<strong>с</strong>т<strong>к</strong><strong>о</strong>-

<strong>к</strong>рылых (Coleoptera) и д<strong>в</strong>у<strong>к</strong>рылых (Diptera).<br />

Среди этих <strong>о</strong>тряд<strong>о</strong><strong>в</strong> уже н<strong>а</strong>чин<strong>а</strong>ют п<strong>о</strong>я<strong>в</strong>лять<strong>с</strong>я<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>, д<strong>о</strong>жи<strong>в</strong>шие и д<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й эп<strong>о</strong>хи,<br />

<strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>о</strong>бщий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> <strong>в</strong><strong>с</strong>ей ф<strong>а</strong>уны <strong>о</strong><strong>с</strong><strong>о</strong>бый: <strong>в</strong>ымерших<br />

юр<strong>с</strong><strong>к</strong>их <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> б<strong>о</strong>льше <strong>в</strong><strong>с</strong>ег<strong>о</strong>. Впер<strong>в</strong>ые<br />

<strong>в</strong> юре п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые (Нутеnoptera).<br />

Ве<strong>с</strong>ьм<strong>а</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> для юр<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уны<br />

р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зных <strong>к</strong>рупных р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых<br />

Palaeontinidae, <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е <strong>в</strong>ремя <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong><br />

приним<strong>а</strong><strong>в</strong>ших<strong>с</strong>я з<strong>а</strong> б<strong>а</strong>б<strong>о</strong>че<strong>к</strong>. Эти гр<strong>о</strong>м<strong>а</strong>дные<br />

ци<strong>к</strong><strong>а</strong>д<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые я<strong>в</strong>ляют<strong>с</strong>я я<strong>в</strong>н<strong>о</strong><br />

«ру<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>дящими» и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емыми для <strong>в</strong><strong>с</strong>ех юр<strong>с</strong><strong>к</strong>их<br />

ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong>. Среди дре<strong>в</strong>не<strong>к</strong>рылых<br />

<strong>с</strong>ледует <strong>о</strong>тметить п<strong>о</strong>я<strong>в</strong>ление <strong>в</strong> юре<br />

н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих п<strong>о</strong>ден<strong>о</strong><strong>к</strong>, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующих<strong>с</strong>я гетер<strong>о</strong>н<strong>о</strong>мией<br />

<strong>к</strong>рылье<strong>в</strong> (нер<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>тью), б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> <strong>с</strong>тре<strong>к</strong><strong>о</strong>з Anisozygoptera<br />

(ныне <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шег<strong>о</strong><strong>с</strong>я лишь <strong>в</strong> чи<strong>с</strong>ле <strong>о</strong>дн<strong>о</strong>г<strong>о</strong><br />

рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> Азии) и п<strong>о</strong>я<strong>в</strong>ление н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong>,<br />

ныне шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong><br />

Anisoptera.<br />

Мел<strong>о</strong><strong>в</strong><strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> изучен<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> меньше<br />

юр<strong>с</strong><strong>к</strong><strong>о</strong>й и д<strong>а</strong>же три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й: <strong>в</strong><strong>с</strong>ег<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>ят<strong>к</strong><strong>а</strong> ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений <strong>в</strong> 3. Сибири,<br />

П<strong>о</strong><strong>в</strong><strong>о</strong>лжье и 3. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>не (р. Эмб<strong>а</strong>), где<br />

н<strong>а</strong>йдены единичные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и мел<strong>о</strong><strong>в</strong><strong>о</strong>й ф<strong>а</strong>уны.<br />

З<strong>а</strong>рубежные ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения мел<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе, С. Амери<strong>к</strong>е и Кит<strong>а</strong>е т<strong>а</strong><strong>к</strong>же<br />

д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р изучены <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong> и не м<strong>о</strong>гут <strong>с</strong><br />

д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерн<strong>о</strong><strong>с</strong>тью и миним<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>лн<strong>о</strong>т<strong>о</strong>й <strong>о</strong><strong>с</strong><strong>в</strong>етить<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти ф<strong>а</strong>уны. Одн<strong>о</strong> из ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых эт<strong>о</strong>г<strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>, <strong>о</strong>т<strong>к</strong>рыт<strong>о</strong>е <strong>в</strong><br />

п<strong>о</strong><strong>с</strong>ледние де<strong>с</strong>ятилетия, п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет н<strong>а</strong>деять<strong>с</strong>я<br />

н<strong>а</strong> п<strong>о</strong>лучение интере<strong>с</strong>ных <strong>с</strong><strong>в</strong>едений: т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> <strong>к</strong><strong>а</strong>н<strong>а</strong>д<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждение мел<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря<br />

Сидэрлэй<strong>к</strong> (Cedar Lake); <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т эт<strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует <strong>с</strong><strong>а</strong>мым <strong>в</strong>ерх<strong>а</strong>м мел<strong>а</strong>, гр<strong>а</strong>нич<strong>а</strong>щим<br />

<strong>с</strong> п<strong>а</strong>ле<strong>о</strong>цен<strong>о</strong>м, и, т<strong>а</strong><strong>к</strong>им <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я<br />

т<strong>о</strong>лщ<strong>а</strong> <strong>в</strong><strong>с</strong>ей <strong>с</strong>и<strong>с</strong>темы <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>чень<br />

м<strong>а</strong>л<strong>о</strong>изученн<strong>о</strong>й. Ф<strong>а</strong>ун<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых мел<strong>а</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong>т юр<strong>с</strong><strong>к</strong>их и х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я н<strong>а</strong>личием<br />

мн<strong>о</strong>гих н<strong>о</strong><strong>в</strong>ых групп, уже д<strong>о</strong>жи<strong>в</strong><strong>а</strong>ющих д<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й эп<strong>о</strong>хи. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

д<strong>в</strong>у<strong>к</strong>рылых (Ceratopogonidae, Chironomidae),<br />

<strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых (Psychopseidae), <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>,<br />

мн<strong>о</strong>гих же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых. П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>в</strong> мелу<br />

чи<strong>с</strong>т<strong>о</strong> юр<strong>с</strong><strong>к</strong>ие группы р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых уже <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Третичн<strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лн<strong>о</strong><br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>к</strong><strong>о</strong>лле<strong>к</strong>циями <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> их <strong>в</strong><br />

музеях мир<strong>а</strong>. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>бн<strong>а</strong>ружены н<strong>а</strong> <strong>в</strong><strong>с</strong>ех<br />

м<strong>а</strong>тери<strong>к</strong><strong>а</strong>х, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле и н<strong>а</strong> террит<strong>о</strong>рии СССР.<br />

У<strong>к</strong><strong>а</strong>жем прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> н<strong>а</strong> ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые из<strong>в</strong>е<strong>с</strong>тны н<strong>а</strong> берег<strong>а</strong>х<br />

Б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> м<strong>о</strong>ря <strong>в</strong> Лит<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й ССР и <strong>в</strong><br />

К<strong>а</strong>линингр<strong>а</strong>д<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл., д<strong>а</strong>лее н<strong>а</strong> б<strong>о</strong>г<strong>а</strong>тые ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

ми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong><br />

С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>зе (<strong>в</strong> Ст<strong>а</strong><strong>в</strong>р<strong>о</strong>п<strong>о</strong>лье), н<strong>а</strong> Тянь-Ш<strong>а</strong>не<br />

(<strong>в</strong> Ч<strong>о</strong>н-Тузе), <strong>в</strong> М<strong>о</strong>лд<strong>а</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й ССР, 3. У<strong>к</strong>р<strong>а</strong>ине<br />

и др. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения третичных<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе, н.<strong>а</strong>пример, <strong>о</strong>лиг<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> Герм<strong>а</strong>нии (Р<strong>о</strong>тт),<br />

Фр<strong>а</strong>нции, Ш<strong>в</strong>ейц<strong>а</strong>рии, Ит<strong>а</strong>лии. Очень б<strong>о</strong>г<strong>а</strong>ты<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждение ми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых н<strong>а</strong><br />

з<strong>а</strong>п<strong>а</strong>де С. Амери<strong>к</strong>и (Фл<strong>о</strong>ри<strong>с</strong><strong>с</strong><strong>а</strong>нт) и мн<strong>о</strong>гие<br />

другие.<br />

Не<strong>с</strong>м<strong>о</strong>тря н<strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зие и шир<strong>о</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение<br />

ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

третичн<strong>о</strong>й ф<strong>а</strong>уны, <strong>с</strong>тепень их изученн<strong>о</strong><strong>с</strong>ти еще<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>о</strong>чень не<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>. Причин<strong>а</strong> м<strong>а</strong>л<strong>о</strong>й<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong><strong>с</strong>ти третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> б<strong>о</strong>льш<strong>о</strong>й близ<strong>о</strong><strong>с</strong>ти их <strong>к</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне: эт<strong>о</strong> <strong>о</strong>б<strong>с</strong>т<strong>о</strong>ятель<strong>с</strong>т<strong>в</strong><strong>о</strong> требует<br />

<strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> п<strong>о</strong>лных зн<strong>а</strong>ний ныне жи<strong>в</strong>ущих н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

и тем <strong>с</strong><strong>а</strong>мым <strong>о</strong>пределяет <strong>о</strong>бяз<strong>а</strong>тельн<strong>о</strong><strong>с</strong>ть<br />

уз<strong>к</strong><strong>о</strong>й <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ции и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong>,<br />

изуч<strong>а</strong>ющих третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р эт<strong>о</strong> треб<strong>о</strong><strong>в</strong><strong>а</strong>ние п<strong>о</strong>чти не <strong>в</strong>ып<strong>о</strong>лнял<strong>о</strong><strong>с</strong>ь,<br />

и третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых изуч<strong>а</strong>ли и <strong>о</strong>пи<strong>с</strong>ы<strong>в</strong><strong>а</strong>ли<br />

не<strong>с</strong>пеци<strong>а</strong>ли<strong>с</strong>ты. П<strong>о</strong>эт<strong>о</strong>му <strong>в</strong> гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>м<br />

б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е р<strong>а</strong>б<strong>о</strong>ты п<strong>о</strong> 'н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым третичн<strong>о</strong>й<br />

ф<strong>а</strong>уны нет<strong>о</strong>чны и требуют п<strong>о</strong><strong>в</strong>т<strong>о</strong>рн<strong>о</strong>г<strong>о</strong> <strong>к</strong>ритиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

пере<strong>с</strong>м<strong>о</strong>тр<strong>а</strong>. М<strong>о</strong>жн<strong>о</strong> <strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть, чт<strong>о</strong> эт<strong>о</strong>т<br />

р<strong>а</strong>здел п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гии н<strong>а</strong>иб<strong>о</strong>лее труд<strong>о</strong>ем<strong>о</strong><strong>к</strong><br />

и еще <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong> р<strong>а</strong>зр<strong>а</strong>б<strong>о</strong>т<strong>а</strong>н. П<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью х<strong>а</strong>р<strong>а</strong><strong>к</strong>териз<strong>о</strong><strong>в</strong><strong>а</strong>ть<br />

третичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых з<strong>а</strong>труднительн<strong>о</strong>:<br />

<strong>о</strong>ни <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>о</strong>чень близ<strong>к</strong>и <strong>к</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременным. М<strong>о</strong>жн<strong>о</strong> лишь <strong>в</strong> грубых черт<strong>а</strong>х<br />

у<strong>к</strong><strong>а</strong>з<strong>а</strong>ть, чт<strong>о</strong> <strong>в</strong> третичн<strong>о</strong>й ф<strong>а</strong>уне при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

п<strong>о</strong>чти <strong>в</strong><strong>с</strong>е <strong>с</strong><strong>о</strong><strong>в</strong>ременные <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>о</strong>тряд<strong>о</strong><strong>в</strong>;<br />

<strong>в</strong>ымершие <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>о</strong>чень немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

и для мн<strong>о</strong>гих групп неиз<strong>в</strong>е<strong>с</strong>тны <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е. Ср<strong>а</strong><strong>в</strong>ни<strong>в</strong><strong>а</strong>я<br />

ф<strong>а</strong>уны п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> <strong>с</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й, м<strong>о</strong>жн<strong>о</strong><br />

<strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть, чт<strong>о</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>в</strong>ымершие п<strong>а</strong>ле<strong>о</strong>ген<strong>о</strong><strong>в</strong>ые<br />

р<strong>о</strong>ды н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>о</strong>бычны. Ин<strong>о</strong>е н<strong>а</strong>блю-<br />

,д<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> не<strong>о</strong>ген<strong>о</strong><strong>в</strong>ых ф<strong>а</strong>ун<strong>а</strong>х: <strong>в</strong>ымершие р<strong>о</strong>ды<br />

п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>в</strong> лучше <strong>в</strong><strong>с</strong>ег<strong>о</strong><br />

из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>м ми<strong>о</strong>цене, и <strong>в</strong><strong>с</strong>е <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зие этих<br />

ф<strong>а</strong>ун п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нению <strong>с</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременными <strong>с</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

лишь <strong>в</strong> ин<strong>о</strong>м <strong>в</strong>ид<strong>о</strong><strong>в</strong><strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>е. Впр<strong>о</strong>чем, <strong>в</strong><br />

ми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых ф<strong>а</strong>ун<strong>а</strong>х <strong>о</strong>тнюдь не ред<strong>к</strong>и и <strong>с</strong><strong>о</strong><strong>в</strong>ременные<br />

<strong>в</strong>иды. Эт<strong>а</strong> <strong>о</strong>бщ<strong>а</strong>я х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong><strong>а</strong> третичных<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>к</strong><strong>о</strong>нечн<strong>о</strong>, <strong>о</strong>чень <strong>с</strong>хем<strong>а</strong>тичн<strong>а</strong><br />

и нужд<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>о</strong>н<strong>к</strong>ретиз<strong>а</strong>ции и ут<strong>о</strong>чнении н<strong>а</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трения <strong>о</strong>пределенных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong>.<br />

Чет<strong>в</strong>ертичные ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие '<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы<br />

изучены еще <strong>о</strong>чень нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>; п<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>у,<br />

их и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ние нер<strong>а</strong>зры<strong>в</strong>н<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong> р<strong>а</strong>б<strong>о</strong>т<strong>о</strong>й<br />

з<strong>о</strong><strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong>-ф<strong>а</strong>уни<strong>с</strong>т<strong>о</strong><strong>в</strong>, з<strong>а</strong>ним<strong>а</strong>ющих<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>в</strong>ещением<br />

и<strong>с</strong>т<strong>о</strong>рии и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны.<br />

Чет<strong>в</strong>ертичные ф<strong>а</strong>уны <strong>о</strong>чень <strong>в</strong><strong>а</strong>жны для з<strong>о</strong><strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>их<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний, <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>я <strong>с</strong><strong>а</strong>мую нед<strong>а</strong><strong>в</strong>нюю<br />

и<strong>с</strong>т<strong>о</strong>рию и изменения ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нения <strong>о</strong>пределенных <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

http://jurassic.ru/

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. Специ<strong>а</strong>лиз<strong>а</strong>ция и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей <strong>в</strong> <strong>с</strong>луч<strong>а</strong>е<br />

изучения чет<strong>в</strong>ертичных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong>т<strong>о</strong>ль<br />

же <strong>о</strong>бяз<strong>а</strong>тельн<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>в</strong> р<strong>а</strong>б<strong>о</strong>т<strong>а</strong>х п<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне.<br />

В з<strong>а</strong><strong>к</strong>лючение <strong>с</strong>ледует <strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>с</strong>л<strong>о</strong><strong>в</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующег<strong>о</strong> мнения <strong>о</strong><br />

б<strong>о</strong>льш<strong>о</strong>й ред<strong>к</strong><strong>о</strong><strong>с</strong>ти и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>о</strong> трудн<strong>о</strong><strong>с</strong>тях их <strong>с</strong>б<strong>о</strong>р<strong>о</strong><strong>в</strong>, д<strong>а</strong>же п<strong>о</strong>дч<strong>а</strong><strong>с</strong> <strong>о</strong><br />

не<strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти пл<strong>а</strong>н<strong>о</strong>мерн<strong>о</strong>г<strong>о</strong> <strong>с</strong>б<strong>о</strong>р<strong>а</strong> их <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

и п<strong>о</strong>и<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Т<strong>а</strong><strong>к</strong>ие пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ления <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ны<br />

н<strong>а</strong> нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><strong>с</strong>ти и <strong>с</strong>луч<strong>а</strong>йн<strong>о</strong><strong>с</strong>ти д<strong>а</strong>нных, <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>л<strong>а</strong> п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гия <strong>в</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

нед<strong>а</strong><strong>в</strong>нее <strong>в</strong>ремя. Пер<strong>в</strong>ые н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых дел<strong>а</strong>ли<strong>с</strong>ь <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

п<strong>о</strong>путн<strong>о</strong> и были <strong>с</strong>луч<strong>а</strong>йными при ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>с</strong>ъем<strong>к</strong>е и <strong>с</strong>б<strong>о</strong>р<strong>а</strong>х <strong>к</strong><strong>а</strong><strong>к</strong>их-либ<strong>о</strong> других, «<strong>а</strong>рхи<strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>их»<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее<br />

<strong>в</strong>ремя т<strong>а</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> мнение <strong>с</strong>пр<strong>а</strong><strong>в</strong>едли<strong>в</strong><strong>о</strong> лишь<br />

<strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>й м<strong>а</strong>л<strong>о</strong>й мере. К<strong>а</strong><strong>к</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>л х<strong>о</strong>д р<strong>а</strong>з<strong>в</strong>ития<br />

п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>'гиче<strong>с</strong><strong>к</strong>их зн<strong>а</strong>ний <strong>в</strong> СССР, <strong>в</strong><br />

ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти р<strong>а</strong>б<strong>о</strong>ты А. В. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>тнюдь не я<strong>в</strong>ляют<strong>с</strong>я<br />

и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> ред<strong>к</strong>ими и трудн<strong>о</strong> <strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>емыми.<br />

При п<strong>о</strong><strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong><strong>к</strong>е <strong>с</strong>пеци<strong>а</strong>льных<br />

п<strong>о</strong>и<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых р<strong>а</strong>б<strong>о</strong>т, <strong>в</strong>ним<strong>а</strong>тельн<strong>о</strong>м и тщ<strong>а</strong>тельн<strong>о</strong>м<br />

изучении <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>о</strong>чных п<strong>о</strong>р<strong>о</strong>д<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong><strong>с</strong>егд<strong>а</strong>.<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>е у<strong>с</strong>л<strong>о</strong><strong>в</strong>ие н<strong>а</strong>х<strong>о</strong>ждения <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых — н<strong>а</strong>личие т<strong>о</strong>н<strong>к</strong><strong>о</strong>зерни<strong>с</strong>тых <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>о</strong>чных<br />

п<strong>о</strong>р<strong>о</strong>д, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong><strong>о</strong>нтинент<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

http://jurassic.ru/<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждения. Вним<strong>а</strong>тельный п<strong>о</strong><strong>с</strong>л<strong>о</strong>йный пр<strong>о</strong><strong>с</strong>м<strong>о</strong>тр<br />

р<strong>а</strong><strong>с</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong> п<strong>о</strong>р<strong>о</strong>ды при п<strong>о</strong>м<strong>о</strong>щи лупы, д<strong>а</strong>ющей<br />

у<strong>в</strong>еличение <strong>в</strong> 10 р<strong>а</strong>з, п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет <strong>о</strong>бн<strong>а</strong>ружить<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е не<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> м<strong>о</strong>гут быть <strong>к</strong>рупными и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метными.<br />

Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>ч<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>а</strong>жущие<strong>с</strong>я бе<strong>с</strong>пл<strong>о</strong>дными<br />

п<strong>о</strong>и<strong>с</strong><strong>к</strong>и п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляют <strong>в</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong><strong>о</strong>нц<strong>о</strong><strong>в</strong><br />

<strong>о</strong>бн<strong>а</strong>ружить н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>н<strong>о</strong><strong>с</strong>ный г<strong>о</strong>риз<strong>о</strong>нт, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

<strong>в</strong> д<strong>а</strong>льнейшем д<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет б<strong>о</strong>г<strong>а</strong>тый и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

м<strong>а</strong>тери<strong>а</strong>л.<br />

Изучение п<strong>о</strong>лученн<strong>о</strong>г<strong>о</strong> м<strong>а</strong>тери<strong>а</strong>л<strong>а</strong> п<strong>о</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емым<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых я<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong>иб<strong>о</strong>лее<br />

<strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енным м<strong>о</strong>мент<strong>о</strong>м <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>я<strong>к</strong><strong>о</strong>м п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>нии: и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong>е<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> любых ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х, н<strong>а</strong>чин<strong>а</strong>я <strong>с</strong> <strong>в</strong>ерх<strong>о</strong><strong>в</strong> нижнег<strong>о</strong><br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>, <strong>о</strong>чень з<strong>а</strong>трудняет т<strong>о</strong>чн<strong>о</strong>е <strong>о</strong>пределение<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>й прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ти <strong>о</strong>бн<strong>а</strong>руженных<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong>. Ф<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong>и<br />

т<strong>о</strong>чн<strong>о</strong>е <strong>о</strong>пределение <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>-либ<strong>о</strong> <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я нед<strong>о</strong><strong>с</strong>тупным<br />

<strong>о</strong>дн<strong>о</strong>му и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телю. П<strong>о</strong>эт<strong>о</strong>му п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>ги<br />

принуждены <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>ть<strong>с</strong>я н<strong>а</strong><br />

р<strong>а</strong>зличных групп<strong>а</strong>х н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичн<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующему<br />

п<strong>о</strong>л<strong>о</strong>жению при и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниях<br />

п<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>е <strong>с</strong><strong>о</strong><strong>в</strong>ременных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

при эт<strong>о</strong>м у<strong>с</strong>л<strong>о</strong><strong>в</strong>ии уд<strong>а</strong>ет<strong>с</strong>я избеж<strong>а</strong>ть нет<strong>о</strong>чных<br />

или не<strong>в</strong>ерных <strong>о</strong>пределений и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>бычных <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех <strong>с</strong>луч<strong>а</strong>ях и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний<br />

целых ф<strong>а</strong>ун <strong>о</strong>дним п<strong>а</strong>лэнт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>г<strong>о</strong>м (<strong>с</strong>м. у Р<strong>о</strong>денд<strong>о</strong>рф<strong>а</strong>,<br />

1957; Р<strong>о</strong>денд<strong>о</strong>рф<strong>а</strong>, Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й и Ш<strong>а</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>, 1961).

СИСТЕМАТИЧЕСКАЯ ЧАСТЬ<br />

ПОДКЛАСС APTERYGOTA. ПЕРВИЧНОБЕСКРЫЛЫЕ'<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые: 0,2—15 мм <strong>в</strong> длину,<br />

ред<strong>к</strong><strong>о</strong> 40 мм. Крылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> три и меньше. Н<strong>а</strong> пер<strong>в</strong>ых<br />

<strong>с</strong>егмент<strong>а</strong>х брюш<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> п<strong>а</strong>рные или неп<strong>а</strong>рные<br />

(<strong>с</strong>р<strong>о</strong><strong>с</strong>шие<strong>с</strong>я) прид<strong>а</strong>т<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде меш<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>с</strong>тержень<strong>к</strong><strong>о</strong><strong>в</strong> или рудимент<strong>а</strong>рных н<strong>о</strong>же<strong>к</strong>.<br />

Мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з неп<strong>о</strong>лный или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Жи<strong>в</strong>ут <strong>с</strong>реди р<strong>а</strong><strong>с</strong>тений (<strong>в</strong> тр<strong>а</strong><strong>в</strong>е, <strong>в</strong><strong>о</strong> мху и т. д.);<br />

<strong>в</strong> земле, п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями, п<strong>о</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>й гнилых пней,<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые ф<strong>о</strong>рмы — <strong>в</strong> <strong>о</strong>т<strong>с</strong>ыре<strong>в</strong>ших ме<strong>с</strong>т<strong>а</strong>х д<strong>о</strong>м<strong>о</strong><strong>в</strong>,<br />

мур<strong>а</strong><strong>в</strong>ейни<strong>к</strong><strong>а</strong>х, н<strong>а</strong> <strong>с</strong><strong>к</strong><strong>а</strong>л<strong>а</strong>х. Пит<strong>а</strong>ют<strong>с</strong>я р<strong>а</strong><strong>с</strong>тительными<br />

<strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong>ми (пле<strong>с</strong>енью, грибным<br />

мицелием, лиш<strong>а</strong>йни<strong>к</strong><strong>а</strong>ми, ин<strong>о</strong>гд<strong>а</strong> т<strong>а</strong><strong>к</strong>же т<strong>к</strong><strong>а</strong>нями<br />

<strong>с</strong><strong>о</strong><strong>с</strong>уди<strong>с</strong>тых р<strong>а</strong><strong>с</strong>тений, гниющими р<strong>а</strong><strong>с</strong>тительными<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми).<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>чень ред<strong>к</strong>и. Лучше<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong>ни из<strong>в</strong>е<strong>с</strong>тны из янт<strong>а</strong>ря Приб<strong>а</strong>лти<strong>к</strong>и,<br />

<strong>о</strong>т<strong>к</strong>уд<strong>а</strong> был<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<strong>о</strong> неб<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> ф<strong>о</strong>рм.<br />

Отряды Protura, Collembola и Diplura близ<strong>к</strong>и<br />

<strong>к</strong> мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong>м, и лишь Thysanura и Monura<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны <strong>к</strong>рыл<strong>а</strong>тым н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым. Не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>,<br />

чт<strong>о</strong> тиз<strong>а</strong>нур<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бные ф<strong>о</strong>рмы д<strong>а</strong>ли н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong><br />

<strong>в</strong><strong>с</strong>ем <strong>к</strong>рыл<strong>а</strong>тым н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong> п<strong>о</strong>лным, т<strong>а</strong><strong>к</strong><br />

и <strong>с</strong> неп<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением. Кр<strong>о</strong>ме Protura,<br />

<strong>в</strong><strong>с</strong>е другие <strong>о</strong>тряды пер<strong>в</strong>ичн<strong>о</strong>бе<strong>с</strong><strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии. Де<strong>в</strong><strong>о</strong>н<br />

— ныне. Пять <strong>о</strong>тряд<strong>о</strong><strong>в</strong>: Collembola, Protura,<br />

Diplura, Thysanura, Monura.<br />

ОТРЯД COLLEMBOLA. НОГОХВОСТКИ<br />

Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны <strong>в</strong>тянуты <strong>в</strong>нутрь г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн четыре-пять; гл<strong>а</strong>з<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> <strong>о</strong>дним <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м.<br />

Брюш<strong>к</strong><strong>о</strong> из ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; н<strong>а</strong> пер<strong>в</strong><strong>о</strong>м<br />

<strong>с</strong>егменте п<strong>а</strong>рный <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т или <strong>с</strong>р<strong>о</strong><strong>с</strong>ший<strong>с</strong>я, <strong>в</strong>тяги<strong>в</strong><strong>а</strong>ющий<strong>с</strong>я<br />

меш<strong>о</strong>че<strong>к</strong>; н<strong>а</strong> пят<strong>о</strong>м <strong>с</strong>егменте у<br />

прыг<strong>а</strong>ющих н<strong>о</strong>г<strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> прыг<strong>а</strong>тельн<strong>а</strong>я <strong>в</strong>ил<strong>к</strong><strong>а</strong>;<br />

цер<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong> тел<strong>а</strong> 0,2—10 мм.<br />

Де<strong>в</strong><strong>о</strong>н — ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Arthropleona,<br />

Symphypleona.<br />

1<br />

В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя ряд з<strong>а</strong>рубежных и <strong>с</strong><strong>о</strong><strong>в</strong>ет<strong>с</strong><strong>к</strong>их<br />

энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong> предл<strong>а</strong>г<strong>а</strong>ют р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть Protura, Collembola<br />

и Diplura, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельные <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы. А<strong>в</strong>т<strong>о</strong>р<br />

45<br />

ПОДОТРЯД ARTHROPLEONA.<br />

ЧЛЕНИСТОБРЮХИЕ<br />

Тел<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е, цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>е; грудные<br />

и брюшные <strong>с</strong>егменты <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные, не<strong>с</strong>р<strong>о</strong><strong>с</strong>шие<strong>с</strong>я.<br />

Де<strong>в</strong><strong>о</strong>н — ныне. Три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Poduromorpha,<br />

Protentomobryomorpha, Entomobryomorpha.<br />

при<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>к</strong> эт<strong>о</strong>й т<strong>о</strong><strong>а</strong><strong>к</strong>е зрения, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> р<strong>а</strong>ди уд<strong>о</strong>б<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

изл<strong>о</strong>жения и ди<strong>с</strong><strong>к</strong>у<strong>с</strong><strong>с</strong>и<strong>о</strong>нн<strong>о</strong>е <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong><strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<br />

<strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящей р<strong>а</strong>б<strong>о</strong>те <strong>с</strong>т<strong>а</strong>рую <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию Apterygota.<br />

http://jurassic.ru/

НАДСЕМЕЙСТВО PODUROMORPHA<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы <strong>с</strong> м<strong>а</strong>л<strong>о</strong>зерни<strong>с</strong>т<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тью.<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; п<strong>о</strong><strong>с</strong>т<strong>а</strong>нтенн<strong>а</strong>льный <strong>о</strong>рг<strong>а</strong>н<br />

из <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> центр<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>с</strong>идящими<br />

н<strong>а</strong> нем периферийными буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми. Среднеи<br />

з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че, не при<strong>к</strong>рыт<strong>а</strong> передним <strong>к</strong>р<strong>а</strong>ем<br />

<strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>и. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Hypogastruridae, Poduridae, Onychiuridae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йдены.<br />

СЕМЕЙСТВО HYPOGASTRURIDAE BORNER, 1906<br />

[п<strong>о</strong>т. transl. Вбгпег, 1913<br />

(ex Hypogastrurinae Вбгпег, 1906) 1<br />

Прыг<strong>а</strong>тельн<strong>а</strong>я <strong>в</strong>ил<strong>к</strong><strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>а</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

н<strong>о</strong> <strong>в</strong>ыд<strong>а</strong>ет<strong>с</strong>я <strong>в</strong>перед з<strong>а</strong> брюшную труб<strong>к</strong>у.<br />

Длин<strong>а</strong> тел<strong>а</strong> 0,8—1 мм (ри<strong>с</strong>. 28). П<strong>а</strong>ле<strong>о</strong>ген<br />

Ри<strong>с</strong>. 28—32. П<strong>о</strong>д<strong>о</strong>тряд Arthropleona<br />

28. Hypogastrura protoviatica Handschin; <strong>о</strong>бщий<br />

<strong>в</strong>ид, х 23,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Handschin, 1926). 29. Rhyniella<br />

praecursor Hirst et Maulik; передняя ч<strong>а</strong><strong>с</strong>ть тел<strong>а</strong>,<br />

x 57,0, де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия (Scourfield,<br />

1940). 30. Laotoma saltans Nicolet; <strong>о</strong>бщий <strong>в</strong>ид,<br />

x 16,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handschin, 1929). 31.<br />

Lepidocyrtus ambricus Handschin; <strong>о</strong>бщий <strong>в</strong>ид,<br />

x 45,0. п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Handschin, 1926). 32. Tomocerus longicornis<br />

Lubbock-, <strong>о</strong>бщий <strong>в</strong>ид, х 12,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handschin, 1929)<br />

46<br />

http://jurassic.ru/<br />

ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<br />

них лишь <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь), <strong>о</strong><strong>с</strong>т<strong>а</strong>льные пять <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йдены.<br />

НАДСЕМЕЙСТВО<br />

PROTENTOMOBRYOMORPHA<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>че или <strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> и при<strong>к</strong>рыт<strong>а</strong><br />

п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью передним <strong>к</strong>р<strong>а</strong>ем <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>и;<br />

п<strong>о</strong><strong>с</strong>ледняя р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong>е. Де<strong>в</strong><strong>о</strong>н<br />

— мел. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protentomobryidae.<br />

СЕМЕЙСТВО PROTENTOMOBRYIDAE<br />

FOLSOM, 1938<br />

Гл<strong>а</strong>зных ф<strong>а</strong><strong>с</strong>ет<strong>о</strong><strong>к</strong> ше<strong>с</strong>ть-<strong>с</strong>емь. Пер<strong>в</strong>ый<br />

<strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>о</strong><strong>с</strong>т<strong>а</strong>льных;<br />

прыг<strong>а</strong>тельн<strong>а</strong>я <strong>в</strong>ил<strong>к</strong><strong>а</strong> из д<strong>в</strong>ух пр<strong>о</strong><strong>с</strong>тых <strong>в</strong>ет<strong>в</strong>ей.<br />

Длин<strong>а</strong> тел<strong>а</strong> 0,65—5 мм (ри<strong>с</strong>. 29). Де<strong>в</strong><strong>о</strong>н —<br />

мел. Три р<strong>о</strong>д<strong>а</strong>.<br />

Вне СССР: Rhyniella Hirst et Maulik, 1926;<br />

Rhyniognatha Tillyard, 1928; Protentomobrya<br />

Folsom, 1937.<br />

НАДСЕМЕЙСТВО ENTOMOBRYOMORPHA<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> гл<strong>а</strong>д<strong>к</strong>ие, без зерни<strong>с</strong>т<strong>о</strong><strong>с</strong>ти;<br />

ин<strong>о</strong>гд<strong>а</strong> е<strong>с</strong>ть чешуй<strong>к</strong>и. Антенны длиннее<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; п<strong>о</strong><strong>с</strong>т<strong>а</strong>нтенн<strong>а</strong>льный <strong>о</strong>рг<strong>а</strong>н <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

или <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

м<strong>а</strong>л<strong>а</strong>, п<strong>о</strong>чти не р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью при<strong>к</strong>рыт<strong>а</strong><br />

передним <strong>к</strong>р<strong>а</strong>ем <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>и. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 600<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>в</strong>ид<strong>о</strong><strong>в</strong>, б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>а</strong>ле<strong>а</strong>р<strong>к</strong>тиче<strong>с</strong><strong>к</strong>ие.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Isotomidae, Entomobryidae, Tomoceridae, Aetaletidae,<br />

Cyphoderidae, из них д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йдены.<br />

СЕМЕЙСТВО ISOTOMIDAE CARL, 1899<br />

[nom. transl. Вбгпег, 1913<br />

(ex Isotominae Carl, 1899)1<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> четырьмя п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мными<br />

члени<strong>к</strong><strong>а</strong>ми. Тр<strong>о</strong>х<strong>а</strong>нтерный <strong>о</strong>рг<strong>а</strong>н н<strong>а</strong><br />

з<strong>а</strong>дней п<strong>а</strong>ре н<strong>о</strong>г <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Чет<strong>в</strong>ертый <strong>с</strong>егмент<br />

брюш<strong>к</strong><strong>а</strong> немн<strong>о</strong>г<strong>о</strong> длиннее третьег<strong>о</strong>. Длин<strong>а</strong><br />

тел<strong>а</strong> 0,6—3 мм (ри<strong>с</strong>. 30). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ENTOMOBRYIDAE BORNER, 1913<br />

Антенны р<strong>а</strong>зличн<strong>о</strong>й длины, из четырех —<br />

ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>; чет<strong>в</strong>ертый члени<strong>к</strong> <strong>с</strong><strong>а</strong>мый<br />

длинный. Тр<strong>о</strong>х<strong>а</strong>нтерный <strong>о</strong>рг<strong>а</strong>н з<strong>а</strong>дней н<strong>о</strong>ги <strong>с</strong><br />

групп<strong>о</strong>й т<strong>о</strong>рч<strong>а</strong>щих шип<strong>о</strong><strong>в</strong>. Чет<strong>в</strong>ертый <strong>с</strong>епмент<br />

брюш<strong>к</strong><strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> длиннее третьег<strong>о</strong>. Длин<strong>а</strong><br />

тел<strong>а</strong> 1,5—3,5 мм (ри<strong>с</strong>. 31). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, три <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).

СЕМЕЙСТВО TOMOCER1DAE<br />

SCHAFFER, 1896<br />

[nom. transl. Borner, 1913<br />

(ex Tomocerinae Schaffer, 1896)]<br />

Антенны длинные, из четырех члени<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

третий члени<strong>к</strong> <strong>с</strong><strong>а</strong>мый длинный, <strong>к</strong><strong>о</strong>льч<strong>а</strong>тый.<br />

Чет<strong>в</strong>ертый <strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че третьег<strong>о</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 1,5—6 мм (ри<strong>с</strong>. 32). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Один р<strong>о</strong>д. П<strong>а</strong>ле<strong>о</strong>ген Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

ПОДОТРЯД SYMPHYPLEONA<br />

ЦЕЛЬНОБРЮХИЕ<br />

Тел<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>о</strong>е; пер<strong>в</strong>ые четыре <strong>с</strong>егмент<strong>а</strong><br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong>р<strong>о</strong><strong>с</strong>ли<strong>с</strong>ь <strong>с</strong> т<strong>о</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>льными <strong>с</strong>егмент<strong>а</strong>ми;<br />

пятый и ше<strong>с</strong>т<strong>о</strong>й <strong>с</strong>егменты <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Sminthuridae, Neelidae,<br />

Dicyrtomidae.<br />

СЕМЕЙСТВО SMINTHURIDAE<br />

NICOLET, 1841<br />

[nom. transl. Lubbock, 1870<br />

(ex Sminthurelles Nicolet, 1841)]<br />

Антенны р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; чет<strong>в</strong>ертый члени<strong>к</strong> длиннее третьег<strong>о</strong>.<br />

Гр<strong>а</strong>ницы <strong>с</strong>р<strong>о</strong><strong>с</strong>ших<strong>с</strong>я <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> <strong>с</strong>терты;<br />

<strong>а</strong>н<strong>а</strong>льный и генит<strong>а</strong>льный <strong>с</strong>егменты рез<strong>к</strong><strong>о</strong><br />

<strong>о</strong>тделены. Длин<strong>а</strong> тел<strong>а</strong> 0,2—1 мм (ри<strong>с</strong>. 33).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

200 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>в</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> п<strong>а</strong>ле<strong>а</strong>р<strong>к</strong>тиче<strong>с</strong><strong>к</strong>их,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> пяти р<strong>о</strong>д<strong>а</strong>м; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые ч<strong>а</strong><strong>с</strong>ти <strong>в</strong>тянуты <strong>в</strong>нутрь г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; гл<strong>а</strong>з<br />

нет; <strong>а</strong>нтенны длинные, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые.<br />

Один или д<strong>в</strong><strong>а</strong> члени<strong>к</strong><strong>а</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Брюш<strong>к</strong><strong>о</strong> из де<strong>с</strong>яти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х<br />

грифель<strong>к</strong>и; п<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> <strong>с</strong><br />

длинными члени<strong>с</strong>тыми или цельными <strong>к</strong>леще<strong>в</strong>идными<br />

цер<strong>к</strong><strong>а</strong>ми. Жи<strong>в</strong>ут <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями,<br />

п<strong>о</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>й гнилых пней. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядны.<br />

Р<strong>а</strong>з<strong>в</strong>итие без мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>бычн<strong>о</strong><br />

2—5 мм, ред<strong>к</strong><strong>о</strong> 40 мм. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Campodidae, Japygidae, Projapygidae;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не<br />

н<strong>а</strong>йден<strong>о</strong>.<br />

Ри<strong>с</strong>. 33—37. П<strong>о</strong>д<strong>о</strong>тряд Symphypleona, <strong>о</strong>тряды Diplura,<br />

Thysanura<br />

33. Sminthurus minnesotensis Guthrie; <strong>о</strong>бщий <strong>в</strong>ид, x 27,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

Амери<strong>к</strong><strong>а</strong> (Guthrie, 1903). 34. Campodea sp., <strong>о</strong>бщий <strong>в</strong>ид, x 4,5,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925). 35. Triassomachilis urethritis<br />

Sharov; x 6,0; A — <strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; Б — <strong>в</strong>ид <strong>с</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; три<strong>а</strong><strong>с</strong>, Приур<strong>а</strong>лье (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1948). 36. Lepisma<br />

sacharinum Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, х 3,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1925). 37. Machilis sp., <strong>о</strong>бщий <strong>в</strong>ид, х 4,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Snodgrass, 1952)<br />

ОТРЯД DIPLURA. ДВУХВОСТКИ<br />

СЕМЕЙСТВО CAMPODEIDAE LUBBOCK, 1873<br />

[nom. transl. Handlirsch, 1906<br />

(ex Campodeidea Lubbock, 1873)<br />

Антенны <strong>о</strong>бычн<strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы и груди<br />

<strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых. Цер<strong>к</strong>и ните<strong>в</strong>идные, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong> брюш<strong>к</strong><strong>о</strong>м (ри<strong>с</strong>. 34).<br />

П<strong>а</strong>ле<strong>о</strong>ген—ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong>;<br />

<strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь), <strong>о</strong>дин из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО JAPYGIDAE LUBBOCK, 1873<br />

Антенны немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы и груди<br />

<strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых. Цер<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

щипце<strong>в</strong>идные. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них д<strong>в</strong><strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

47<br />

http://jurassic.ru/

ОТРЯД THYSANURA.<br />

Тел<strong>о</strong> удлиненн<strong>о</strong>е. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные;<br />

<strong>о</strong>т пяти д<strong>о</strong> ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong> м<strong>а</strong><strong>к</strong><strong>с</strong>илярных<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong>; <strong>а</strong>нтенны длинные, члени<strong>с</strong>тые; гл<strong>а</strong>з<strong>а</strong><br />

<strong>с</strong>л<strong>о</strong>жные. Л<strong>а</strong>п<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>члененн<strong>а</strong>я. Брюш<strong>к</strong><strong>о</strong> из 11<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>; н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х грифель<strong>к</strong>и и <strong>в</strong>ыпячи<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я<br />

меш<strong>о</strong>ч<strong>к</strong>и; длинные члени<strong>с</strong>тые цер<strong>к</strong>и<br />

и <strong>с</strong>редняя неп<strong>а</strong>рн<strong>а</strong>я нить. Н<strong>а</strong> теле чешуй<strong>к</strong>и.<br />

Очень п<strong>о</strong>д<strong>в</strong>ижны, жи<strong>в</strong>ут <strong>в</strong> у<strong>к</strong>рытых ме<strong>с</strong>т<strong>а</strong>х<br />

(п<strong>о</strong>ч<strong>в</strong>е, трещин<strong>а</strong>х <strong>с</strong><strong>к</strong><strong>а</strong>л и т. д.). Три<strong>а</strong><strong>с</strong> — ныне.<br />

Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Triassomachilidae, Lepismatidae,<br />

Machilidae, Meinertellidae, Nicoletidae; д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йдены.<br />

СЕМЕЙСТВО TRIASSOMACHLIDAE<br />

SHAROV, 1948<br />

Гл<strong>а</strong>з<strong>а</strong> из неб<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> чи<strong>с</strong>л<strong>а</strong> ф<strong>а</strong><strong>с</strong>ет<strong>о</strong><strong>к</strong>, не <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я.<br />

Пле<strong>в</strong>р<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть брюш<strong>к</strong><strong>а</strong><br />

<strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; грифель<strong>к</strong>и н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых ч<strong>а</strong><strong>с</strong>тях<br />

пер<strong>в</strong>ых де<strong>с</strong>яти <strong>с</strong>тернит<strong>о</strong><strong>в</strong>. Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Triassomachilis S h <strong>а</strong> г <strong>о</strong> v, 1948. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Т. uralensis Sharov, 1948; <strong>в</strong> три<strong>а</strong><strong>с</strong>, Б<strong>а</strong>ш<strong>к</strong>ир<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (р. Н<strong>а</strong><strong>к</strong>ыз). Грифель<strong>к</strong>и члени<strong>с</strong>тые. Длин<strong>а</strong><br />

тел<strong>а</strong> 6 мм (ри<strong>с</strong>. 35). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong><br />

Приур<strong>а</strong>лья.<br />

ОТРЯД MONURA,<br />

Тел<strong>о</strong> удлиненн<strong>о</strong>е. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные;<br />

ше<strong>с</strong>ть члени<strong>к</strong><strong>о</strong><strong>в</strong> м<strong>а</strong><strong>к</strong><strong>с</strong>илярных щупи<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты; <strong>а</strong>нтенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, члени<strong>с</strong>тые;<br />

гл<strong>а</strong>з<strong>а</strong> мн<strong>о</strong>г<strong>о</strong>ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные; <strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>к</strong><strong>а</strong>п<strong>с</strong>уле<br />

имеют<strong>с</strong>я рудименты тергит<strong>о</strong><strong>в</strong> м<strong>а</strong>ндибулярн<strong>о</strong>г<strong>о</strong>,<br />

м<strong>а</strong><strong>к</strong><strong>с</strong>илярн<strong>о</strong>г<strong>о</strong> и нижнегубн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, б<strong>о</strong>лее<br />

р<strong>а</strong>з<strong>в</strong>итые, чем у Thysanura. Л<strong>а</strong>п<strong>к</strong><strong>а</strong> нер<strong>а</strong><strong>с</strong>члененн<strong>а</strong>я,<br />

<strong>с</strong> един<strong>с</strong>т<strong>в</strong>енным <strong>к</strong>рупным прет<strong>а</strong>рз<strong>о</strong>м.<br />

Тул<strong>о</strong><strong>в</strong>ище г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мн<strong>о</strong> р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>е, без<br />

я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>зделения н<strong>а</strong> грудь и<br />

брюш<strong>к</strong><strong>о</strong>; <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из 14 <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

неп<strong>а</strong>рн<strong>о</strong>й члени<strong>с</strong>т<strong>о</strong>й нитью, приблизительн<strong>о</strong><br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong> длине тул<strong>о</strong><strong>в</strong>ищу; цер<strong>о</strong>,<strong>к</strong> нет. Чешуе<strong>к</strong><br />

н<strong>а</strong> теле нет. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — н. пермь. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Dasyleptidae.<br />

СЕМЕЙСТВО DASYLEPTIDAE SHAROV, 1951<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, м<strong>а</strong>л<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> д<strong>в</strong>умя<br />

б<strong>о</strong>льшими б<strong>а</strong>з<strong>а</strong>льными члени<strong>к</strong><strong>а</strong>ми. Яйце-<br />

ЩЕТИНОХВОСТКИ<br />

СЕМЕЙСТВО LEP1SMATIDAE LEACH, 1815.<br />

ЧЕШУИНИЦЫ<br />

[nom transl. Escherich, 1904<br />

(ex Lepismida Leach, 1815)]<br />

• Ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> м<strong>а</strong>лень<strong>к</strong>ие, р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Грифель<strong>к</strong>и<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> д<strong>в</strong>ух п<strong>о</strong><strong>с</strong>ледних <strong>с</strong>егмент<strong>а</strong>х брюш<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 1—12 мм (ри<strong>с</strong>. 36). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

<strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь), <strong>о</strong>дин из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО MACHILIDAE<br />

GRASSI, 1888<br />

Ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льшие, <strong>с</strong>ближены; три<br />

гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. Грифель<strong>к</strong>и н<strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>с</strong>егмент<strong>а</strong>х брюш<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 7—17 мм (ри<strong>с</strong>. 37). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

<strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> ми<strong>о</strong>цене С. Амери<strong>к</strong>и.<br />

<strong>о</strong>дн<strong>о</strong>х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>к</strong>и<br />

<strong>к</strong>л<strong>а</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, нечлени<strong>с</strong>тый; тергит трин<strong>а</strong>дц<strong>а</strong>т<strong>о</strong>г<strong>о</strong><br />

тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> б<strong>о</strong>льше тергит<strong>а</strong><br />

д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — н. пермь.<br />

Один р<strong>о</strong>д.<br />

Dasyleptus Brongniart, 1885. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. lucasi Brongniart, 1855; <strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Фр<strong>а</strong>нция. Гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>лулунные, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

нижним и прямым или <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong><strong>о</strong>гнутым <strong>в</strong>ерхним<br />

<strong>к</strong>р<strong>а</strong>ем. Тергит пер<strong>в</strong><strong>о</strong>г<strong>о</strong> тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> уже <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>; н<strong>а</strong> <strong>с</strong><strong>в</strong>еши<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я<br />

<strong>в</strong>низ б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>к</strong>р<strong>а</strong>ях, ближе <strong>к</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю и п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> ему,— углубление<br />

<strong>в</strong> <strong>в</strong>иде ш<strong>в</strong><strong>а</strong>; тергит трин<strong>а</strong>дц<strong>а</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> шире четырн<strong>а</strong>дц<strong>а</strong>т<strong>о</strong>г<strong>о</strong> (ри<strong>с</strong>. 38). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Фр<strong>а</strong>нции; н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ри<strong>с</strong>. 38. Отряд Monura<br />

Dasyleptus brongniarti Sharov; ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция, x 5,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1957)<br />

http://jurassic.ru/

Н<strong>а</strong> <strong>с</strong>реднегрудн<strong>о</strong>м и з<strong>а</strong>днегрудн<strong>о</strong>м <strong>с</strong>егмент<strong>а</strong>х<br />

имеет<strong>с</strong>я п<strong>о</strong> п<strong>а</strong>ре <strong>к</strong>рылье<strong>в</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ующих<br />

у неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых <strong>с</strong>т<strong>а</strong>дий р<strong>а</strong>з<strong>в</strong>ития и у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

ф<strong>о</strong>рм, утр<strong>а</strong>ти<strong>в</strong>ших их <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны не <strong>в</strong>тянуты <strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ную <strong>к</strong><strong>а</strong>п<strong>с</strong>улу,<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные, не <strong>с</strong>чит<strong>а</strong>я ред<strong>к</strong>их <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> их п<strong>о</strong>лн<strong>о</strong>й<br />

реду<strong>к</strong>ции при <strong>а</strong>ф<strong>а</strong>гии. Р<strong>а</strong>з<strong>в</strong>итие <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><br />

пре<strong>в</strong>р<strong>а</strong>щением. Де<strong>в</strong><strong>о</strong>н — ныне. Д<strong>в</strong><strong>а</strong> инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>:<br />

Palaeoptera, или дре<strong>в</strong>не<strong>к</strong>рылые, и<br />

ПОДКЛАСС PTERYGOTA<br />

КРЫЛАТЫЕ НАСЕКОМЫЕ<br />

(В. 'Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

Neoptera, или н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылые. Об<strong>о</strong><strong>с</strong><strong>о</strong>бление и<br />

ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ние инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> пр<strong>о</strong>х<strong>о</strong>дил<strong>о</strong>, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>,<br />

<strong>в</strong> де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>м пери<strong>о</strong>де: <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующие<br />

п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>о</strong><strong>с</strong><strong>в</strong>ещ<strong>а</strong>ют<br />

лишь б<strong>о</strong>лее п<strong>о</strong>здние эт<strong>а</strong>пы и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

<strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

<strong>о</strong>б<strong>о</strong>их инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> <strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>е<br />

ф<strong>а</strong>уны <strong>в</strong>ерх<strong>о</strong><strong>в</strong> нижнег<strong>о</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> (н<strong>а</strong>мюр<strong>а</strong>).<br />

ИНФРАКЛАСС PALAEOPTERA. ДРЕВНЕКРЫЛЫЕ<br />

Крылья удлиненные, лишенные р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й<br />

юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, не <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бные, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>о</strong>т<strong>в</strong><strong>о</strong>дить<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д и <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong>д<strong>о</strong>ль тел<strong>а</strong>,<br />

п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>я брюш<strong>к</strong><strong>о</strong>. Крылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>терты<br />

<strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны. Лишь <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>а</strong> из мег<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>птер<strong>о</strong>ид<strong>о</strong><strong>в</strong><br />

(Diaphanopterodea) <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>а</strong>ли <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ть<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ть <strong>к</strong>рылья <strong>в</strong>д<strong>о</strong>ль тел<strong>а</strong>. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны грызущие или <strong>в</strong> <strong>в</strong>иде длинн<strong>о</strong>г<strong>о</strong> х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong>,<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong> <strong>в</strong><strong>с</strong>еми р<strong>о</strong>т<strong>о</strong><strong>в</strong>ыми ч<strong>а</strong><strong>с</strong>тями.<br />

Р<strong>а</strong>з<strong>в</strong>итие из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> лишь для д<strong>в</strong>ух <strong>о</strong>тряд<strong>о</strong><strong>в</strong>,<br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>н<strong>о</strong><br />

пр<strong>о</strong>х<strong>о</strong>дит <strong>в</strong> <strong>в</strong><strong>о</strong>дн<strong>о</strong>й <strong>с</strong>реде, причем пре<strong>в</strong>р<strong>а</strong>щение<br />

неп<strong>о</strong>лн<strong>о</strong>е, без п<strong>о</strong><strong>к</strong><strong>о</strong>ящей<strong>с</strong>я <strong>с</strong>т<strong>а</strong>дии (<strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>и).<br />

Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне; н<strong>а</strong>иб<strong>о</strong>лее р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>е. Три н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>: Palaeodictyopteroidea,<br />

Megasecopteroidea, Odonatoidea. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие<br />

<strong>о</strong>тн<strong>о</strong>шения этих н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> изуче­<br />

4 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

49<br />

http://jurassic.ru/<br />

ны м<strong>а</strong>л<strong>о</strong>, Palaeodictyopteroidea — н<strong>а</strong>иб<strong>о</strong>лее<br />

дре<strong>в</strong>ний и и<strong>с</strong>х<strong>о</strong>дный для д<strong>в</strong>ух других. В<strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение<br />

д<strong>в</strong>ух других н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ет лет<strong>а</strong>тельную<br />

<strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>цию и еще не <strong>в</strong>ыя<strong>с</strong>ненные<br />

изменения <strong>о</strong>бр<strong>а</strong>з<strong>а</strong> жизни. Б<strong>о</strong>лее <strong>в</strong><strong>с</strong>ег<strong>о</strong> эт<strong>о</strong><br />

я<strong>с</strong>н<strong>о</strong> <strong>в</strong> <strong>о</strong>тн<strong>о</strong>шении <strong>с</strong>тре<strong>к</strong><strong>о</strong>з<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я рез<strong>к</strong><strong>о</strong> удлиненными <strong>к</strong>рыльями,<br />

р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ющими <strong>с</strong>ильные тяг<strong>о</strong><strong>в</strong>ые у<strong>с</strong>илия и<br />

у<strong>в</strong>еличи<strong>в</strong><strong>а</strong>ющими т<strong>а</strong><strong>к</strong>им <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м <strong>с</strong><strong>к</strong><strong>о</strong>р<strong>о</strong><strong>с</strong>ть п<strong>о</strong>лет<strong>а</strong>.<br />

Эт<strong>о</strong> <strong>о</strong>тр<strong>а</strong>зил<strong>о</strong><strong>с</strong>ь н<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ении грудн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тдел<strong>а</strong>,<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м <strong>о</strong>чень б<strong>о</strong>льш<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие при<strong>о</strong>брели<br />

пле<strong>в</strong>р<strong>а</strong>льные мышцы, <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляющие<br />

д<strong>в</strong>ижения <strong>к</strong>рылье<strong>в</strong> неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>, <strong>а</strong> не <strong>к</strong><strong>о</strong><strong>с</strong><strong>в</strong>енн<strong>о</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> у <strong>в</strong><strong>с</strong>ех других н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. В<strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение<br />

б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> и <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong><br />

Megasecopteroidea, ныне п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>в</strong>ымершег<strong>о</strong>,<br />

нея<strong>с</strong>н<strong>о</strong>, х<strong>о</strong>тя <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зие м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей ег<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>чень <strong>в</strong>ели<strong>к</strong><strong>о</strong>.

НАДОТРЯД PALAEOD I CTYOPTEROIDEA<br />

Крылья <strong>с</strong> прямым или умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

передним и <strong>в</strong>ыпу<strong>к</strong>лым з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ями (ни<strong>к</strong><strong>о</strong>гд<strong>а</strong><br />

не бы<strong>в</strong><strong>а</strong>ют <strong>с</strong>тебельч<strong>а</strong>тыми); <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля имеют<strong>с</strong>я, <strong>в</strong><strong>с</strong>егд<strong>а</strong> уз<strong>к</strong>ие. П<strong>о</strong>перечные<br />

жил<strong>к</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т <strong>к</strong>реп<strong>к</strong>их<br />

пр<strong>о</strong>д<strong>о</strong>льных; <strong>о</strong>ни ч<strong>а</strong><strong>с</strong>т<strong>о</strong> непр<strong>а</strong><strong>в</strong>ильные, <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зуя <strong>с</strong>еть, ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют,<br />

или же имеет<strong>с</strong>я н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящий <strong>а</strong>рхеди<strong>к</strong>тий. П<strong>о</strong>пе­<br />

речные жил<strong>к</strong>и ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> чет<strong>к</strong>ие<br />

п<strong>о</strong>перечные ряды. В<strong>с</strong>егд<strong>а</strong> имеют<strong>с</strong>я длинные<br />

цер<strong>к</strong>и, ин<strong>о</strong>гд<strong>а</strong> и неп<strong>а</strong>рный п<strong>а</strong>р<strong>а</strong>цер<strong>к</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне. Четыре <strong>о</strong>тряд<strong>а</strong>, из<strong>в</strong>е<strong>с</strong>тных<br />

<strong>с</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>я: Palaeodictyoptera, Eubleptodea,<br />

Archodonata, Ephemeroptera; п<strong>о</strong><strong>с</strong>ледний пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<br />

<strong>в</strong> мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их и <strong>к</strong><strong>а</strong>йн<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>а</strong>ун<strong>а</strong>х.<br />

ОТРЯД PALAEODICTYOPTERA. ПАЛЕОДИКТИОПТЕРЫ<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong>лющими р<strong>о</strong>т<strong>о</strong><strong>в</strong>ыми<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>к</strong>рыл<strong>о</strong><strong>в</strong>идные<br />

прид<strong>а</strong>т<strong>к</strong>и н<strong>а</strong> передне<strong>с</strong>пин<strong>к</strong>е. Обычн<strong>о</strong> имеет<strong>с</strong>я<br />

<strong>а</strong>рхеди<strong>к</strong>тий или непр<strong>а</strong><strong>в</strong>ильн<strong>а</strong>я гу<strong>с</strong>т<strong>а</strong>я <strong>с</strong>еть п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>ни т<strong>о</strong>н<strong>к</strong>и и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны,<br />

реже <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Неп<strong>а</strong>рный п<strong>а</strong>р<strong>а</strong>цер<strong>к</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. К<strong>а</strong>рб<strong>о</strong>н — пермь. Три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>:<br />

Eupalaeodictyoptera, Parapalaeodictyoptera,<br />

Metapalaeodictyoptera.<br />

ПОДОТРЯД<br />

EUPALAEODICTYOPTERA<br />

З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лее переднег<strong>о</strong>. RS, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, гребенч<strong>а</strong>т<strong>а</strong>я;<br />

SC д<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>в</strong>ершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> умеренн<strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>а</strong>я, <strong>о</strong>бычн<strong>о</strong><br />

притуплённ<strong>а</strong>я; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен <strong>а</strong>рхеди<strong>к</strong>тий.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Пять н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Lithomanteidea, Dictyoneuridea, Breyeriidea,<br />

Homoiopteridea, Polycreagridea.<br />

НАДСЕМЕЙСТВО<br />

LITHOMANTEIDEA<br />

Крылья п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, з<strong>а</strong>дние <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширенные; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные <strong>о</strong>тделы гл<strong>а</strong><strong>в</strong>ных жил<strong>о</strong><strong>к</strong><br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я; б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> <strong>с</strong>л<strong>а</strong>бых и непр<strong>а</strong><strong>в</strong>ильных<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>, <strong>о</strong>бр<strong>а</strong>зующих <strong>с</strong>еть;<br />

ин<strong>о</strong>гд<strong>а</strong> ме<strong>с</strong>т<strong>а</strong>ми н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящий <strong>а</strong>рхеди<strong>к</strong>тий. К<strong>а</strong>рб<strong>о</strong>н<br />

— пермь. Ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: три из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы (Lithomanteidae, Lycocercidae и<br />

Lusiellidae), д<strong>в</strong><strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и [Syntonopteridae<br />

(ри<strong>с</strong>. 39) и Thesoneuridae] и <strong>о</strong>дн<strong>о</strong><br />

из перми 3. Е<strong>в</strong>р<strong>о</strong>пы (Eugereonidae).<br />

СЕМЕЙСТВО EUGEREONIDAE<br />

HANDLIRSCH, 1906<br />

Передние и з<strong>а</strong>дние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие; гу<strong>с</strong>т<strong>о</strong>й<br />

<strong>а</strong>рхеди<strong>к</strong>тий <strong>в</strong> з<strong>а</strong>дних ч<strong>а</strong><strong>с</strong>тях <strong>к</strong>рылье<strong>в</strong>, ч<strong>а</strong><strong>с</strong>тые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>с</strong>переди (ри<strong>с</strong>. 40, А, Б). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я; т<strong>о</strong>н<strong>к</strong>ие ните<strong>в</strong>идные <strong>а</strong>нтенны;<br />

<strong>в</strong>ыпу<strong>к</strong>лые гл<strong>а</strong>з<strong>а</strong>; члени<strong>с</strong>тый <strong>к</strong><strong>о</strong>лющий х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>,<br />

50<br />

http://jurassic.ru/<br />

<strong>в</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>а</strong>з длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие,<br />

л<strong>а</strong>п<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Пермь. Один р<strong>о</strong>д из н. перми<br />

Герм<strong>а</strong>нии.<br />

НАДСЕМЕЙСТВО DICTYONEURIDEA<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные или з<strong>а</strong>дние <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong><strong>с</strong>ширены<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>а</strong>рхеди<strong>к</strong>тий х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен,<br />

реже имеют<strong>с</strong>я мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные, <strong>о</strong>бычн<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ящие<strong>с</strong>я т<strong>о</strong>н<strong>к</strong>ие п<strong>о</strong>перечные жил<strong>к</strong>и. К<strong>а</strong>рб<strong>о</strong>н.<br />

Ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: <strong>о</strong>дн<strong>о</strong> из <strong>с</strong>р. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы,<br />

Азии и С. Амери<strong>к</strong>и (Dictyoneuridae), четыре<br />

из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы (Peromapteridae,<br />

Cockerelliellidae, Mecynopteridae и Protagriidae)<br />

и <strong>о</strong>дн<strong>о</strong> из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и (Нурегmegethidae).<br />

СЕМЕЙСТВО DICTYONEURIDAE HANDLIRSCH, 1906<br />

(Stenodictyopteridae Brongniart, 1893, nom.<br />

invalid., Megaptilidae Handlirsch, 1906;<br />

Fouqueidae Handlirsch, 1906; Pteronidiidae<br />

Bolton, 1912; Saarlandiidae Gutnorl, 1930;<br />

Rochlingiidae guthorl, 1934. Syn. nov.<br />

Крылья р<strong>а</strong><strong>в</strong>н<strong>о</strong>й длины, умеренн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенные<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; типичный <strong>а</strong>рхеди<strong>к</strong>тий, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий<br />

из мел<strong>к</strong>их ячее<strong>к</strong>, лишь ме<strong>с</strong>т<strong>а</strong>ми <strong>о</strong>бр<strong>а</strong>зующий<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные нежные п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

К<strong>а</strong>рб<strong>о</strong>н. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе и<br />

С. Амери<strong>к</strong>е; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> СССР.<br />

Asiodictya Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. rosslca Rohdendorf, 1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, З<strong>а</strong><strong>в</strong>ьял<strong>о</strong><strong>в</strong><strong>о</strong>). SC и R рез<strong>к</strong><strong>о</strong><br />

<strong>с</strong>ближены <strong>с</strong> С; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong>ыми <strong>в</strong>ет<strong>в</strong>ями<br />

RS; другие ячей<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> типичным <strong>а</strong>рхеди<strong>к</strong>тием.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 65 мм, ширин<strong>а</strong><br />

18 мм (ри<strong>с</strong>. 41, А, Б). Один <strong>в</strong>ид. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

НАДСЕМЕЙСТВО BREYERIIDEA<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong> рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширенным <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> т<strong>о</strong>н<strong>к</strong>их, из<strong>о</strong>гнутых п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>, ме<strong>с</strong>т<strong>а</strong>ми ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуюших

•40В<br />

89. Liihoneura lameerl Carpenter; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 1,7;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,7; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1933).<br />

40. Eugereon boeckingi Dohrn; A —г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, si; £ —переднее<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,1; В—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1; Пермь, Герм<strong>а</strong>ния (Handlirsch,<br />

1906). 41. Asiodictya rossica Rohdendorf; A— <strong>к</strong>рыл<strong>о</strong>, x 1,9;<br />

Б — уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> <strong>к</strong>рыл<strong>а</strong>, x 12,0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>ден­<br />

\<br />

Ри<strong>с</strong>. 39—44. П<strong>о</strong>д<strong>о</strong>тряд Eupalaeodictyoptera<br />

д<strong>о</strong>рф, 1961). 42. Breyeria borinensts Borre; <strong>к</strong>рыл<strong>о</strong>, х 0,9, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1906). 43. Homoioptera woodwardi<br />

Brongniart; <strong>к</strong>рыл<strong>о</strong>, x 0,8, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1906); 44. Polycreagra elegans Handlirsch; <strong>к</strong>рыл<strong>о</strong>, x 0,9, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1906).<br />

51 4*<br />

http://jurassic.ru/

<strong>к</strong>рупн<strong>о</strong>ячеи<strong>с</strong>тую <strong>с</strong>еть; <strong>а</strong>рхеди<strong>к</strong>тия нет. К<strong>а</strong>рб<strong>о</strong>н.<br />

Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> из <strong>с</strong>р. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы:<br />

Breyeriidae (ри<strong>с</strong>. 42) и Cryptoveniidae.<br />

НАДСЕМЕЙСТВО HOMOIOPTERIDEA<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, умеренн<strong>о</strong> удлиненные,<br />

<strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чными <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми <strong>в</strong><strong>о</strong><br />

<strong>в</strong><strong>с</strong>ех п<strong>о</strong>лях; мн<strong>о</strong>г<strong>о</strong> т<strong>о</strong>н<strong>к</strong>их п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>;<br />

<strong>а</strong>рхеди<strong>к</strong>тия нет. К<strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> из<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы: Homoiopteridae (ри<strong>с</strong>. 43)<br />

и Rhabdoptilidae.<br />

НАДСЕМЕЙСТВО POLYCREAGRIDEA<br />

Крыл<strong>о</strong> рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>в</strong>ершин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>с</strong>тр<strong>а</strong>я; RS и М <strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

п<strong>а</strong>р<strong>а</strong>ллельных д<strong>о</strong>п<strong>о</strong>лнительных <strong>в</strong>ет<strong>в</strong>ей (д<strong>о</strong> 30);<br />

SC д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong> не д<strong>о</strong>х<strong>о</strong>дит; <strong>а</strong>рхеди<strong>к</strong>тий<br />

не р<strong>а</strong>з<strong>в</strong>ит; б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> т<strong>о</strong>н<strong>к</strong>их и непр<strong>а</strong><strong>в</strong>ильных<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>, не <strong>о</strong>бр<strong>а</strong>зующих<br />

<strong>с</strong>ети (ри<strong>с</strong>. 44). Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Polycreagridae<br />

<strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ПОДОТРЯД<br />

PAR АР AL A EODICTYOP Т ERA<br />

Крыл<strong>о</strong> умеренн<strong>о</strong> удлиненн<strong>о</strong>е^ <strong>с</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лыми передним и з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ями; <strong>в</strong>ерши­<br />

н<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>с</strong>тр<strong>а</strong>я или уз<strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>а</strong>я; п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> немн<strong>о</strong>г<strong>о</strong>; <strong>о</strong>ни непр<strong>а</strong><strong>в</strong>ильные и<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зуют <strong>с</strong>ети или <strong>к</strong>рупные ячей<strong>к</strong>и; <strong>а</strong>рхеди<strong>к</strong>тия<br />

нет; RS <strong>о</strong>бычн<strong>о</strong> дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и <strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я,<br />

ин<strong>о</strong>гд<strong>а</strong> непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>-гребенч<strong>а</strong>т<strong>а</strong>я; SC<br />

д<strong>о</strong> <strong>в</strong>ершины не д<strong>о</strong>х<strong>о</strong>дит, ин<strong>о</strong>гд<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>я. К<strong>а</strong>рб<strong>о</strong>н<br />

— пермь. В<strong>о</strong><strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них четьь<br />

ре из н. и <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы (Macropteridae,<br />

Synarmogidae, Jongmansiidae, Orthocostidae),<br />

<strong>о</strong>дн<strong>о</strong> из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и<br />

(Heolidae), три из перми СССР и С. Амери<strong>к</strong>и<br />

(Eohymenidae, Bardapteridae и Calvertiellidae).<br />

П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, не дре<strong>в</strong>не<strong>к</strong>рылые!<br />

СЕМЕЙСТВО EOHYMENIDAE MARTY NOV, 1937<br />

Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е: SC нея<strong>с</strong>н<strong>а</strong>; М и Си лишь из<br />

д<strong>в</strong>ух пр<strong>о</strong><strong>с</strong>тых <strong>в</strong>ет<strong>в</strong>ей <strong>к</strong><strong>а</strong>жд<strong>а</strong>я; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

т<strong>о</strong>н<strong>к</strong>ие, <strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>о</strong>лях <strong>о</strong>бр<strong>а</strong>зуют д<strong>в</strong>урядные<br />

ячей<strong>к</strong>и. Пермь. Один р<strong>о</strong>д.<br />

Eohymen М <strong>а</strong> г t у п <strong>о</strong> v, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Е. maculipennis Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong><br />

прямыми п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; д<strong>в</strong><strong>а</strong> п<strong>о</strong>ля <strong>с</strong><br />

д<strong>в</strong><strong>о</strong>йными ряд<strong>а</strong>ми ячее<strong>к</strong>: между <strong>в</strong>ет<strong>в</strong>ями RS и<br />

между М и Си. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 38,8 мм, ширин<strong>а</strong><br />

11,8 мм (ри<strong>с</strong>. 45). Один <strong>в</strong>ид. Пермь Ю. Приур<strong>а</strong>лья.<br />

Ри<strong>с</strong>. 45—50. П<strong>о</strong>д<strong>о</strong>тряды Parapalaeodictyoptera, Metapalaeodictyoptera<br />

45. Eohymen maculipennis Martynov; <strong>к</strong>рыл<strong>о</strong>, x 1,9, пермь, При- x 1,4, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий'б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1946). 49. Ретур<strong>а</strong>лье<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,-»1937). 46. Bardapteron ovale G. Zalessky; miakovia quinquefasciata Martynov; <strong>о</strong>бщий <strong>в</strong>ид, x 1,6, пермь,<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,2, пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1944). 47. Neuburgia Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1941). 50. Doropteron mirum G. Zalessky;<br />

altaica Martynov;?<strong>к</strong>рыл<strong>о</strong>, x 2,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн x 1,5, пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1946).<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931).f48. Abaptilon sibiricum G. Zalessky; <strong>к</strong>рыл<strong>о</strong>,<br />

52<br />

http://jurassic.ru/

СЕМЕЙСТВО BARDAPTERIDAE<br />

G. ZALESSKY, 1944<br />

Крыл<strong>о</strong> умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, н<strong>о</strong><br />

з<strong>а</strong>метн<strong>а</strong>я; М из трех, Си из д<strong>в</strong>ух <strong>в</strong>ет<strong>в</strong>ей; з<strong>а</strong>дние<br />

<strong>в</strong>ет<strong>в</strong>и М и <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и Си <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я у<br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и непр<strong>а</strong><strong>в</strong>ильные,<br />

п<strong>о</strong>чти <strong>в</strong>езде <strong>о</strong>бр<strong>а</strong>зуют д<strong>в</strong>урядные ячей<strong>к</strong>и.<br />

Пермь. Один р<strong>о</strong>д.<br />

Bardapteron G. Z ales sky, 1944. Тип<br />

р<strong>о</strong>д<strong>а</strong> — В. ovate G. Zalessky, 1944; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Б<strong>а</strong>рд<strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными <strong>к</strong><strong>о</strong><strong>с</strong>ыми п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; <strong>в</strong>ет<strong>в</strong>ей RS пять. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>в</strong>ыше 50 мм, ширин<strong>а</strong> 18 мм (ри<strong>с</strong>. 46)-<br />

Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

ПОДОТРЯД<br />

METAPALAEODICTYOPTERA<br />

Крыл<strong>о</strong> удлиненн<strong>о</strong>е, <strong>с</strong> прямым или <strong>с</strong>лег<strong>к</strong><strong>а</strong><br />

<strong>в</strong><strong>о</strong>гнутым <strong>в</strong> <strong>с</strong>ередине передним <strong>к</strong>р<strong>а</strong>ем и <strong>с</strong>ильн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лым з<strong>а</strong>дним; <strong>в</strong>ершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

<strong>о</strong><strong>с</strong>тр<strong>а</strong>я; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля уз<strong>к</strong>ие,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, без п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; <strong>а</strong>рхеди<strong>к</strong>тий<br />

или <strong>с</strong>еть из п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и прямые, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong><br />

чет<strong>к</strong>ие, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>дн<strong>а</strong> <strong>о</strong>т друг<strong>о</strong>й,<br />

ин<strong>о</strong>гд<strong>а</strong> п<strong>о</strong>перечные жил<strong>к</strong>и п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

SC д<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>, из них<br />

<strong>о</strong>дн<strong>о</strong> из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и (Homothetidae),<br />

<strong>о</strong>дн<strong>о</strong> из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. (Lamproptilidae),<br />

<strong>о</strong>дн<strong>о</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

(Neuburgiidae) и <strong>о</strong>дн<strong>о</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы,<br />

Амери<strong>к</strong>и и перми Ю. Приур<strong>а</strong>лья (Spilapteridae).<br />

СЕМЕЙСТВО NEUBURGIIDAE ROHDENDORF,<br />

1961<br />

Крыл<strong>о</strong> удлиненн<strong>о</strong>е; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й умеренн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R п<strong>о</strong>чти <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>, негребенч<strong>а</strong>т<strong>а</strong>я, дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я. К<strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д.<br />

Neuburgia М <strong>а</strong> г t у п <strong>о</strong> v, 1931. Тип р<strong>о</strong>д<strong>а</strong>—<br />

N. alfaica Martynov, 1931; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Верх<strong>о</strong>т<strong>о</strong>м<strong>с</strong><strong>к</strong><strong>о</strong>е). Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> немн<strong>о</strong>гими п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>; н<strong>а</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong>ых<strong>о</strong>дят де<strong>в</strong>ять <strong>в</strong>ет<strong>в</strong>ей RS и <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>в</strong>ет<strong>в</strong>ей М;<br />

SC и R <strong>о</strong>чень <strong>с</strong>ближены. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

30 мм (ри<strong>с</strong>. 47). Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО SPILAPTERIDAE<br />

BRONGNIART, 1893<br />

Крыл<strong>о</strong> умеренн<strong>о</strong> удлиненн<strong>о</strong>е, <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

з<strong>а</strong>дним и я<strong>с</strong>н<strong>о</strong> <strong>в</strong><strong>о</strong>гнутым передним<br />

<strong>к</strong>р<strong>а</strong>ями; RS <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R <strong>в</strong> <strong>о</strong>тд<strong>а</strong>лении<br />

<strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong><strong>с</strong>егд<strong>а</strong> гребенч<strong>а</strong>т<strong>а</strong>я;<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> мн<strong>о</strong>г<strong>о</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>ни нежные<br />

и нея<strong>с</strong>ные. К<strong>а</strong>рб<strong>о</strong>н — пермь. 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Abaptilon G. Zalessky, 1946. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. sibiricum G. Zalessky, 1946; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>в</strong>ерхнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>.<br />

Пр<strong>о</strong><strong>к</strong><strong>о</strong>пье<strong>в</strong><strong>с</strong><strong>к</strong>). RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong> и не<strong>с</strong>ет <strong>с</strong>емь<br />

<strong>в</strong>ет<strong>в</strong>ей; М и СиА <strong>с</strong> Ше<strong>с</strong>тью <strong>в</strong>ет<strong>в</strong>ями <strong>к</strong><strong>а</strong>жд<strong>а</strong>я;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм, ширин<strong>а</strong> 15 мм<br />

(ри<strong>с</strong>. 48). Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permiakovia Martynov, 1941. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. quinquefasciata Martynov, 1941; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>,<br />

не<strong>с</strong>ет гребень из <strong>с</strong>еми <strong>в</strong>ет<strong>в</strong>ей, д<strong>в</strong>е из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>бр<strong>а</strong>зуют р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и; н<strong>а</strong> М пять, н<strong>а</strong> СиА ше<strong>с</strong>ть<br />

<strong>в</strong>ет<strong>в</strong>ей; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> я<strong>с</strong>ными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 34 мм, ширин<strong>а</strong><br />

12,5 мм (ри<strong>с</strong>. 49). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Doropteron G. Zalessky, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. mirum G. Zalessky, 1946; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти<br />

<strong>к</strong>рыл<strong>а</strong> и не<strong>с</strong>ет <strong>в</strong> <strong>в</strong>иде пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>г<strong>о</strong> гребня<br />

де<strong>с</strong>ять <strong>в</strong>ет<strong>в</strong>ей; н<strong>а</strong> М четыре <strong>в</strong>ет<strong>в</strong>и; СиА <strong>с</strong> пятью<br />

<strong>в</strong>ет<strong>в</strong>ями н<strong>а</strong> передних <strong>к</strong>рыльях и <strong>с</strong> ше<strong>с</strong>тью н<strong>а</strong><br />

з<strong>а</strong>дних; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong><br />

<strong>к</strong>рылье<strong>в</strong> 36—37 мм, ширин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

10 мм, з<strong>а</strong>днег<strong>о</strong> — 13 мм (ри<strong>с</strong>. 50). Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Вне СССР: <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы — Severinula<br />

Pruvost, 1930; Severinopsis Kukalova, 1958;<br />

Bojoptera Kukalova, 1958; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и—<br />

McLuckiepteron Richardson, 1956; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Е<strong>в</strong>р<strong>о</strong>пы— Palaeoptillus Brongniart, 1893;<br />

Spilaptera Brongniart, 1885; Epitethe Handlirsch,<br />

1906; Compsoneura Brongniart, 1885;<br />

Becquerelia Brongniart, 1893; Homaloneurina<br />

Handlirsch, 1906; Homaloneurites Handlirsch,<br />

1906; Homaloneura Brongniart, 1885.<br />

53<br />

http://jurassic.ru/

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, умеренн<strong>о</strong> удлиненные;<br />

передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый; С, SC<br />

и R <strong>с</strong>ближены; RS и М <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и<br />

и три<strong>а</strong>д не <strong>о</strong>бр<strong>а</strong>зуют; Си и А пр<strong>о</strong><strong>с</strong>тые, дуг<strong>о</strong><strong>о</strong>б-<br />

ОТРЯД EUBLEPTODEA<br />

Ри<strong>с</strong>; 51—54. Отряды Eubleptodea, Archodonata<br />

ные <strong>с</strong>егменты п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й длины; переднегрудь<br />

без п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>. Сегмент<strong>о</strong><strong>в</strong><br />

брюш<strong>к</strong><strong>а</strong> де<strong>с</strong>ять; п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> нет;<br />

п<strong>а</strong>р<strong>а</strong> длинных ните<strong>в</strong>идных цер<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

1. Eubleptus danietsl Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид, x 4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, <strong>к</strong>рыл<strong>о</strong>, х 3,6, пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1932). 54<br />

С. Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1906). 52. Permothemidla caudata Roh- Ideliella decora G. Zalessky; <strong>к</strong>рыл<strong>о</strong>, x 3,2, Пермь, риур<strong>а</strong>ль.<br />

dendorf; <strong>о</strong>бщий <strong>в</strong>ид, x 6,0 (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция), пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1937)<br />

Р<strong>о</strong>денд<strong>о</strong>рф, 1949). 53. Permothemis libelluloides (Martynov);<br />

р<strong>а</strong>зные; <strong>а</strong>рхеди<strong>к</strong>тия нет; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

прямые, ред<strong>к</strong>ие. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими р<strong>о</strong>т<strong>о</strong><strong>в</strong>ыми<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми; <strong>о</strong>чень <strong>к</strong>рупные, <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющие<br />

гл<strong>а</strong>з<strong>а</strong>; <strong>а</strong>нтенны, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Груд-<br />

54<br />

http://jurassic.ru/<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 14 мм (ри<strong>с</strong>. 51). К<strong>а</strong>рб<strong>о</strong>н. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eubleptidae<br />

<strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.

Одн<strong>а</strong> п<strong>а</strong>р<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong>; з<strong>а</strong>дние<br />

<strong>к</strong>рылья и п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты н<strong>а</strong> переднегруди<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й<br />

или <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>ильн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый; <strong>в</strong><strong>с</strong>е пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и р<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

т<strong>о</strong>лщины и три<strong>а</strong>д не <strong>о</strong>бр<strong>а</strong>зуют; RS <strong>о</strong>бычн<strong>о</strong> гребенч<strong>а</strong>тый;<br />

М и Си р<strong>а</strong>з<strong>в</strong>иты <strong>с</strong>л<strong>а</strong>б<strong>о</strong>; <strong>с</strong>л<strong>о</strong>жн<strong>а</strong>я,<br />

мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> А. Сегмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong><br />

де<strong>в</strong>ять; п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> нет; п<strong>а</strong>р<strong>а</strong><br />

длинных цер<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 10—17 мм<br />

(ри<strong>с</strong>. 52). К<strong>а</strong>рб<strong>о</strong>н — пермь. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Permothemistidae, Rectineuridae, Doteridae,<br />

Permoneuridae, из них д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних из н. Перми<br />

С. Амери<strong>к</strong>и, <strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>е из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PERMOTHEMISTIDAE MARTYNOV, 1934<br />

Вет<strong>в</strong>и RS нея<strong>с</strong>н<strong>о</strong>-гребенч<strong>а</strong>тые или дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>ие;<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>в</strong><strong>с</strong>ех жил<strong>к</strong><strong>а</strong>х <strong>к</strong><strong>о</strong>нце<strong>в</strong>ые р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные,<br />

н<strong>о</strong> я<strong>с</strong>ные п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

Пермь. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Permothemidia Rohdendorf, 1940 (Uralothemis<br />

G. Zalessky, 1951 syn. nov.). Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. caudata Rohdendorf, 1940: н. пермь,<br />

ОТРЯД ARCHODONATA<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й; RS <strong>с</strong> б<strong>о</strong>лее или<br />

менее гребенч<strong>а</strong>тым р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жением <strong>в</strong>ет<strong>в</strong>ей;<br />

з<strong>а</strong>дний <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>с</strong> д<strong>в</strong>умя — четырьмя <strong>в</strong>ет<strong>в</strong>ями.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 10,5—12 мм, ширин<strong>а</strong> 3,5—4 мм.<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Ю. Ур<strong>а</strong>л<strong>а</strong>.<br />

Permothemis Martynov, 1934 (Palaeothemis<br />

Martynov, 1932). Тип р<strong>о</strong>д<strong>а</strong> — Palaeothemis<br />

libelluloides Martynov, 1932; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й; RS <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и;<br />

з<strong>а</strong>дний <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>с</strong> пятью <strong>в</strong>ет<strong>в</strong>ями.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 17 мм, ширин<strong>а</strong> 5,75 мм (ри<strong>с</strong>. 53).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Ideliella G. Zalessky, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

/. decora G. Zalessky, 1937; <strong>в</strong> пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры). Передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>, н<strong>о</strong> з<strong>а</strong>метн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый;<br />

RS <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и; з<strong>а</strong>дний <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>с</strong><br />

тремя <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 12,3 мм, ширин<strong>а</strong><br />

4,5 мм (ри<strong>с</strong>. 54). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Вне СССР: Kansasia Tillyard, 1937<br />

(н. пермь С. Амери<strong>к</strong>и).<br />

ОТРЯД EPHEMEROPTERA. ПОДЕНКИ<br />

Неб<strong>о</strong>льш<strong>а</strong>я г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, у <strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм <strong>с</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными р<strong>о</strong>т<strong>о</strong><strong>в</strong>ыми <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>а</strong>нтенн<strong>а</strong>ми. Переднегрудь без п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>. Крылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> прямым<br />

передним <strong>к</strong>р<strong>а</strong>ем, <strong>в</strong>д<strong>о</strong>ль <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

длинные и прямые SC и R. Пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я, <strong>о</strong>бр<strong>а</strong>зуя п<strong>о</strong> <strong>к</strong>р<strong>а</strong>йней<br />

мере <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме RS т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емые три<strong>а</strong>ды<br />

жил<strong>о</strong><strong>к</strong> — х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные <strong>а</strong><strong>с</strong>имметричные дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>ие<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления. RS негребенч<strong>а</strong>тый. Археди<strong>к</strong>тий<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, п<strong>о</strong>перечные жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные,<br />

т<strong>о</strong>н<strong>к</strong>ие и прямые, ред<strong>к</strong><strong>о</strong> п<strong>о</strong>чти<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Имеют<strong>с</strong>я длинные цер<strong>к</strong>и и п<strong>а</strong>р<strong>а</strong>цер<strong>к</strong>;<br />

п<strong>о</strong><strong>с</strong>ледний ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щ<strong>а</strong>ет<strong>с</strong>я<br />

д<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>г<strong>о</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong><strong>а</strong>. Личин<strong>к</strong>и жи<strong>в</strong>ут <strong>в</strong> <strong>в</strong><strong>о</strong>де.<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. П<strong>о</strong>д<strong>о</strong>тряды: Protephemeroptera,<br />

Plectoptera.<br />

ПОДОТРЯД<br />

PROTEPH EMEROP T ERA<br />

(О. А. Черн<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Крылья п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми<br />

три<strong>а</strong>д<strong>а</strong>ми <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме RS. RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

R н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния ,<br />

<strong>к</strong>рыл<strong>а</strong> и не <strong>с</strong><strong>в</strong>яз<strong>а</strong>н <strong>с</strong> М. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й дуж<strong>к</strong>и <strong>в</strong> <strong>о</strong><strong>с</strong>-<br />

55<br />

http://jurassic.ru/<br />

Р и <strong>с</strong><br />

- 5 5<br />

Tnplosob a<br />

! :<br />

B ^^%%'^% )<br />

^Х е<br />

?^Г У<br />

- n<br />

°fl°W Protephemeroptera<br />

' * ^

н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> нет. Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е,<br />

<strong>с</strong> п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> тел<strong>а</strong><br />

24 мм, длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 21 мм( ри<strong>с</strong>. 55). В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Фр<strong>а</strong>нции. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Triplosobidae.<br />

ПОДОТРЯД PLECTOPTERA<br />

RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R при <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я дуж<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> е<strong>с</strong>ть.<br />

Пермь — ныне. Инфр<strong>а</strong><strong>о</strong>тряды: Permoplectopteг<strong>а</strong>,<br />

Euplectoptera.<br />

И Н ФРАОТРЯД<br />

PERMOPLECTOPTERA<br />

З<strong>а</strong>днегрудь <strong>к</strong>рупн<strong>а</strong>я, рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т<br />

<strong>с</strong>реднегруди. Крылья п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные. У <strong>с</strong><strong>а</strong>м<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я дуж<strong>к</strong><strong>а</strong> <strong>в</strong><br />

<strong>в</strong>иде уд<strong>а</strong>ленн<strong>о</strong>й <strong>о</strong>т переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>и <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

жил<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующей петлю, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R. Пермь — юр<strong>а</strong>. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Рг<strong>о</strong>tereismatidea,<br />

Mesephemeridea.<br />

НАДСЕМЕЙСТВО PROTEREISMATIDEA<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>ные пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии друг <strong>о</strong>т друг<strong>а</strong>, не <strong>с</strong>ближ<strong>а</strong>я<strong>с</strong>ь<br />

п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>. Пермь. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Protereismatidae,<br />

Misthodotidae, Eudoteridae.<br />

СЕМЕЙСТВО PROTEREISMATIDAE SELLARDS,<br />

1907<br />

[nom. corr. Tillyard, 1932 (ex Protereismephemeridae<br />

Sellards, 1907)]<br />

CuA <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й три<strong>а</strong>д<strong>о</strong>й; <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

примерн<strong>о</strong> н<strong>а</strong> <strong>с</strong>ередине <strong>с</strong><strong>в</strong><strong>о</strong>ей длины. В <strong>с</strong>и<strong>с</strong>теме<br />

М и Си п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> мн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>межут<strong>о</strong>ч­<br />

http://jurassic.ru/<br />

ных жил<strong>о</strong><strong>к</strong>. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные п<strong>о</strong>перечные жил<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены гу<strong>с</strong>т<strong>о</strong>. Пермь. Один р<strong>о</strong>д.<br />

Protereisma Sellards, 1907 (Loxophlebia<br />

Martynov, 1928). Тип р<strong>о</strong>д<strong>а</strong> — P. permiana Sellards,<br />

1907; н. пермь, С. Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

дуж<strong>к</strong><strong>а</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, н<strong>а</strong> переднем<br />

<strong>к</strong>рыле б<strong>о</strong>лее <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, чем н<strong>а</strong> з<strong>а</strong>днем (ри<strong>с</strong>. 56).<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 15—30 мм, ширин<strong>а</strong> 4,5—9,5 мм.<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> С. Амери<strong>к</strong>е; три <strong>в</strong>ид<strong>а</strong> из<br />

н. перми Ур<strong>а</strong>л<strong>а</strong> и <strong>в</strong>. перми Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО MISTHODOTIDAE TILLYARD, 1932<br />

CuA без трех длинных <strong>в</strong>ет<strong>в</strong>ей, лишь <strong>с</strong> <strong>к</strong><strong>о</strong>нечными<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми. Пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> имеют<strong>с</strong>я. П<strong>о</strong>перечные<br />

жил<strong>к</strong>и ред<strong>к</strong>ие (ри<strong>с</strong>. 57). Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 9—•<br />

15 мм, ширин<strong>а</strong> 3,5—5 мм. Один р<strong>о</strong>д. Н. пермь<br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО EUDOTERIDAE DEMOULIN, 1954<br />

П<strong>о</strong>перечные жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. CuA<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>с</strong> нея<strong>с</strong>ными <strong>в</strong>ет<strong>в</strong>ями. Пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Р<strong>а</strong>змеры<br />

мел<strong>к</strong>ие, длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 6 мм (ри<strong>с</strong>. 58). Н. пермь<br />

С. Амери<strong>к</strong>и. Один р<strong>о</strong>д.<br />

НАДСЕМЕЙСТВО<br />

MESEPHEMERIDEA<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и (RS 3 и RS 4; RS 5 и MAj<br />

и др.) <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong> <strong>к</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. П<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>бы и з<strong>а</strong>метны не <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех

Ри<strong>с</strong>. 59—62. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesephemeridae; Permoplectoptera incertae sedis<br />

59. Pallngeniopsis praecox Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, x 1,7,5 B<br />

- Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид, x 1,5, н. пермь, Приур<strong>а</strong>лье (Handlirsch,<br />

Пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 60. Mesephemera 1906). 62. Mesoplectopteron longipcs Handlirsch; Р <strong>о</strong>бщий^<strong>в</strong>ид,<br />

prisca (Germar); A — переднее <strong>к</strong>рыл<strong>о</strong>, x 3,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,5, три<strong>а</strong><strong>с</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1918)<br />

х 3,0, <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Demoulin, 1955). 61. Phthartus rossicus<br />

п<strong>о</strong>лях. Мн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>межут<strong>о</strong>чных жил<strong>о</strong><strong>к</strong> <strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными<br />

б<strong>а</strong>з<strong>а</strong>льными <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми. Пермь — юр<strong>а</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Mesephemeridae.<br />

СЕМЕЙСТВО MESEPHEMERIDAE CARPENTER, 1932<br />

(Palingeniopsidae Martynov, 1938, Syn nov.)<br />

П<strong>о</strong>перечные жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итые, нея<strong>с</strong>ные.<br />

СиА и Ai <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми. Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле неб<strong>о</strong>льш<strong>о</strong>е. Пермь — юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Palingeniopsis Martynov, 1932. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. praecox Martynov, 1932; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong><strong>о</strong>гнутый; СиР<br />

<strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 32 мм,<br />

ширин<strong>а</strong> 11,5 мм (ри<strong>с</strong>. 59). Один <strong>в</strong>ид. В. пермь<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Mesephemera Handlirsch, 1906,<br />

<strong>в</strong>. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы (ри<strong>с</strong>. 60).<br />

PERMOPLECTOPTERA<br />

INCERTAE SEDIS<br />

Phthartus Handlirsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Ph. rossicus Handlirsch, 1904; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Личин­<br />

<strong>к</strong><strong>а</strong>. З<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и <strong>к</strong>рылье<strong>в</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные. Н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

Де<strong>в</strong>ять п<strong>а</strong>р п<strong>а</strong>л<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идных ж<strong>а</strong>берных<br />

ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Кр<strong>а</strong>я <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> р<strong>о</strong><strong>в</strong>ные. Х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ые<br />

нити <strong>о</strong>пушены <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; л<strong>а</strong>р<strong>а</strong>цер<strong>к</strong> немн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че цер<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 61). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Пермь<br />

Приур<strong>а</strong>лья.<br />

Mesoplectopteron Handlirsch, 1918. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. longipes Handlirsch, 1918; три<strong>а</strong><strong>с</strong>,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (В<strong>о</strong>гезы). Личин<strong>к</strong><strong>а</strong>. З<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и <strong>к</strong>рылье<strong>в</strong><br />

п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и <strong>о</strong>чень<br />

длинные. В<strong>о</strong><strong>с</strong>емь п<strong>а</strong>р ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти брюш<strong>к</strong><strong>а</strong>.<br />

Сегменты брюш<strong>к</strong><strong>а</strong> р<strong>о</strong><strong>в</strong>ные. Х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ые нити<br />

<strong>о</strong>пушены <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; п<strong>а</strong>р<strong>а</strong>цер<strong>к</strong> длиннее цер<strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 62). Один <strong>в</strong>ид. Три<strong>а</strong><strong>с</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ИНФРАОТРЯД EUPLECTOPTERA<br />

З<strong>а</strong>днегрудь меньше <strong>с</strong>реднегруди и те<strong>с</strong>н<strong>о</strong> <strong>с</strong><br />

нею <strong>с</strong>лит<strong>а</strong>. Крылья рез<strong>к</strong><strong>о</strong> гетер<strong>о</strong>н<strong>о</strong>мные, з<strong>а</strong>дние<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

С <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> н<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>..Юр<strong>а</strong> — ныне. Ше<strong>с</strong>ть н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Paedephemeridea, Ephemeridea, Siphlonuridea,<br />

Heptageniidea, Neoephemeridea, Caenidea; д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong><strong>с</strong>ледних т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

http://jurassic.ru/

НАДСЕМЕЙСТВО PAEDEPHEMERIDEA<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны и ин<strong>о</strong>гд<strong>а</strong><br />

п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong> <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я: MPi и CuAj <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельны. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> зн<strong>а</strong>чительн<strong>о</strong><br />

б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong>. Верхняя<br />

юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Paedephemeridae и Нех<strong>а</strong>genitidae.<br />

63. Paedephemera multlnervosa (Oppenheim): А — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

x 2,9; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,9; <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1906). 64. Ephemeropsis trisetalis Eichwald (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция),<br />

x 1,4, <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (Черн<strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 65. Ephemeropsis<br />

martynovae Tshernova; <strong>к</strong>рылья, х^2,4, <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье<br />

СЕМЕЙСТВО PAEDEPHEMERIDAE LAMEERE, 1917<br />

(Stenodicranidae Demoulin, 1954, syn, nov.)<br />

Ри<strong>с</strong>. 63—67. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Paedephemeridae, Hexagenitidae<br />

58<br />

http://jurassic.ru/<br />

З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> з<strong>а</strong><strong>к</strong>ругленный,<br />

т<strong>о</strong>рну<strong>с</strong> не <strong>в</strong>ыр<strong>а</strong>жен; <strong>с</strong>ближение жил<strong>о</strong><strong>к</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

CuAi переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> не имеет<br />

я<strong>с</strong>н<strong>о</strong>г<strong>о</strong> р<strong>а</strong>зд<strong>в</strong><strong>о</strong>ения. Личин<strong>к</strong>и неиз<strong>в</strong>е<strong>с</strong>тны<br />

(ри<strong>с</strong>. 63). Один р<strong>о</strong>д. В. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

64<br />

(Черн<strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 66. Ephemeropsis trisetalis Eichwald;<br />

личин<strong>к</strong><strong>а</strong>, х 3,0, <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (Н. П. Меш<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961).<br />

67. Hexagenites weyenberghl Scudder; переднее <strong>к</strong>рыл<strong>о</strong>, х 3.2.<br />

<strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter, 1932)

СЕМЕЙСТВО HEXAGENITIDAE LAMEERE, 1917<br />

[nom. transl. Demoulin, 1954<br />

(ex Hexagenitinae Lameere, 1917)]<br />

(Ephemeropsidae Cockerell, 1924, syn. nov.)<br />

Переднее <strong>к</strong>рыл<strong>о</strong> треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы, т<strong>о</strong>рну<strong>с</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен; <strong>с</strong>ближение жил<strong>о</strong><strong>к</strong> <strong>о</strong>чень я<strong>с</strong>н<strong>о</strong>е;<br />

CuA н<strong>а</strong> переднем <strong>к</strong>рыле р<strong>а</strong>зд<strong>в</strong><strong>о</strong>ен<strong>а</strong>, причем<br />

из р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> <strong>о</strong>тх<strong>о</strong>дит <strong>в</strong>ет<strong>в</strong>ь, д<strong>а</strong>ющ<strong>а</strong>я ряд<br />

пр<strong>а</strong><strong>в</strong>ильных петле<strong>о</strong>бр<strong>а</strong>зных р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>. Личин<strong>к</strong><strong>а</strong><br />

<strong>с</strong> <strong>с</strong>емью п<strong>а</strong>р<strong>а</strong>ми <strong>о</strong>дин<strong>о</strong>чных ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идных ж<strong>а</strong>бр;<br />

х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ые нити гу<strong>с</strong>т<strong>о</strong> <strong>о</strong>пушены <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

Юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Ephemeropsis Eichwald, 1864 (Phacelobranchus<br />

Handlirsch, 1906). Тип р<strong>о</strong>д<strong>а</strong> — E. trisetalis<br />

Eichewald, 1864; <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (тургин<strong>о</strong>-<strong>в</strong>итим<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Т<strong>о</strong><strong>в</strong>ег<strong>а</strong>). Очень <strong>к</strong>рупные<br />

п<strong>о</strong>ден<strong>к</strong>и. Ан<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> чуть<br />

длиннее термин<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; CuAi рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я,<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>чень мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны; п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>е<br />

<strong>с</strong>ближение пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>о</strong> у мн<strong>о</strong>гих жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> тел<strong>а</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>г<strong>о</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> (без х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ых нитей) 50 мм,<br />

длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 35—42 мм, з<strong>а</strong>днег<strong>о</strong><br />

17—24 мм. Личин<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>к</strong>рупными гл<strong>а</strong>з<strong>а</strong>ми; <strong>с</strong>егменты<br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими з<strong>а</strong>дними угл<strong>а</strong>ми;<br />

длин<strong>а</strong> тел<strong>а</strong> 45—55 м, х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ых нитей 15—<br />

25 мм (ри<strong>с</strong>. 64—66; т<strong>а</strong>бл. I, фиг. 1—3). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лья, М<strong>о</strong>нг<strong>о</strong>лии, С. Кит<strong>а</strong>я.<br />

Hexagenites S <strong>с</strong> u d d е г, 1880. Тип р<strong>о</strong>д<strong>а</strong> —<br />

N. weyenberghi Scudder, 1880; <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(м<strong>а</strong>льм, З<strong>о</strong>ленг<strong>о</strong>фен). Средней <strong>в</strong>еличины<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Термин<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

длиннее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; CuAi п<strong>о</strong>чти прям<strong>а</strong>я; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны. П<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>е<br />

<strong>с</strong>ближение пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>о</strong> лишь<br />

у <strong>в</strong>ет<strong>в</strong>ей RS. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 15—19 мм<br />

(ри<strong>с</strong>. 67). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО EPHEMERIDEA<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>; MPi и CuAi <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я. З<strong>а</strong>дние <strong>к</strong>рылья <strong>к</strong>рупные, немн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины передних <strong>к</strong>рылье<strong>в</strong>.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Palingeniidae,<br />

Ephemeridae, Potamanthidae, Behningiidae,<br />

Polymitarcidae; п<strong>о</strong><strong>с</strong>ледние д<strong>в</strong><strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PALINGENIIDAE KLAPALEK, 1909<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>;<br />

CuA <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; пр<strong>о</strong>межут<strong>о</strong>чные жил<strong>к</strong>и<br />

<strong>в</strong> <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле прямые. Личин<strong>к</strong>и з<strong>а</strong><strong>к</strong><strong>а</strong>пы<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я,<br />

<strong>с</strong> шир<strong>о</strong><strong>к</strong>ими передними г<strong>о</strong>ленями<br />

и пл<strong>о</strong><strong>с</strong><strong>к</strong>ими <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми <strong>в</strong>ерхних челю<strong>с</strong>тей, не<strong>с</strong>ущих<br />

зубцы п<strong>о</strong> <strong>в</strong>нешнему <strong>к</strong>р<strong>а</strong>ю. Крупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Palingenia Burmeister, 1839. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ephemera longicauda Olivier, 1791; <strong>с</strong><strong>о</strong><strong>в</strong>ременный,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. Пр<strong>о</strong>д<strong>о</strong>льные и п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>в</strong> передних и з<strong>а</strong>дних <strong>к</strong>рыльях мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны.<br />

МА переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong> (ри<strong>с</strong>. 68). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне три <strong>в</strong>ид<strong>а</strong>,<br />

<strong>о</strong>дин <strong>в</strong>ид из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО EPHEMERIDAE KLAPALEK, 1909<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong><br />

не <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я; CuA без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и <strong>в</strong> <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>о</strong>тх<strong>о</strong>дят<br />

<strong>о</strong>т CuA и <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>; CuP <strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; между Ai и <strong>к</strong>р<strong>а</strong>ем<br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ерия п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Личин<strong>к</strong>и з<strong>а</strong><strong>к</strong><strong>а</strong>пы<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я,<br />

<strong>с</strong> т<strong>о</strong>н<strong>к</strong>ими, гл<strong>а</strong>д<strong>к</strong>ими, длинными<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми <strong>в</strong>ерхних челю<strong>с</strong>тей, <strong>к</strong><strong>о</strong>нцы <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

з<strong>а</strong>гнуты н<strong>а</strong>ружу, и <strong>с</strong> д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми пери<strong>с</strong>тыми<br />

ж<strong>а</strong>бр<strong>а</strong>ми, з<strong>а</strong>гнутыми н<strong>а</strong> <strong>с</strong>пинную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну<br />

брюш<strong>к</strong><strong>а</strong>. Крупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые (ри<strong>с</strong>. 69). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из них Hexagenia Walsh, 1863 из не<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>пи<strong>с</strong><strong>а</strong>ны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> изученный р<strong>о</strong>д Р<strong>а</strong>г<strong>а</strong>baetis<br />

Haupt, 1956, <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> нея<strong>с</strong>ны (Demoulin, 1957), и ряд нет<strong>о</strong>чн<strong>о</strong><br />

<strong>о</strong>пределенных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>: ?Ephemera Linnaeus,<br />

1746 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря), не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. Амери<strong>к</strong>и<br />

и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

59<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО POTAMANTHIDAE KLAPALEK, 1909<br />

Н<strong>а</strong> переднем <strong>к</strong>рыле CuP <strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>;<br />

между Ai и <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong> нет <strong>с</strong>ерии п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Личин<strong>к</strong>и жи<strong>в</strong>ут <strong>с</strong>реди р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>ти,<br />

д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые пери<strong>с</strong>тые ж<strong>а</strong>бры р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м брюш<strong>к</strong><strong>а</strong>. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong>редней <strong>в</strong>еличины<br />

(ри<strong>с</strong>. 70). Не<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них р<strong>о</strong>д Potamanthus<br />

Pictet, 1845 из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО POLYMITARCIDAE KLAPALEK, 1909<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> не<br />

<strong>с</strong>ближены п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>; CuA <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, <strong>в</strong>нутри<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> н<strong>а</strong>х<strong>о</strong>дят<strong>с</strong>я прямые пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и. Личин<strong>к</strong>и з<strong>а</strong><strong>к</strong><strong>а</strong>пы<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я, <strong>с</strong> длинными<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми челю<strong>с</strong>тей, н<strong>а</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>х з<strong>а</strong>гиб<strong>а</strong>ющих<strong>с</strong>я<br />

<strong>в</strong>нутрь. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>к</strong>рупные и <strong>с</strong>редней <strong>в</strong>еличины<br />

(ри<strong>с</strong>. 71). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них Ephoron Williamson,<br />

1802 <strong>о</strong>бн<strong>а</strong>ружен <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).

Ри<strong>с</strong>. 68—71. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Palingeniidae, Ephemeridae, Potamanthidae, Polymitarcidae<br />

<strong>к</strong> 8<br />

' P<br />

2lL n<br />

/JI l<br />

i a<br />

Jf' n<br />

J li<br />

i a<br />

'i d<br />

l i v i e r f '° ): A<br />

~ переднее <strong>к</strong>рыл<strong>о</strong>, x 1,7;<br />

•£„7и ?25т<br />

реднее <strong>к</strong>рыл<strong>о</strong>, x 5,5; Б - з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 4,5; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Schoene-<br />

Р<br />

Ри<strong>с</strong>. 72—76. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Siphlonuridae, Isonychiidae, Baefidae<br />

72. Stphlonurus aestivalis Eaton; переднее <strong>к</strong>рыл<strong>о</strong>, x 3,5 и з<strong>а</strong>днее (Demoulin, 1955). 75. Baetis pumilus (Bueister); ^"етм"<br />

<strong>к</strong>рыл<strong>о</strong>, хЗ,5; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Schoenemund, 1930). 73. IsonycMa ignota <strong>к</strong>рыл<strong>о</strong>, x 10,0 и з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, x 10,0; j<strong>с</strong><strong>о</strong><strong>в</strong>рем. (S=hoenemunQ<br />

Walker; переднее <strong>к</strong>рыл<strong>о</strong>, x 4,2 и з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 4,2; <strong>с</strong><strong>о</strong><strong>в</strong>рем. 1930>. 76. Mesotefis s.b.nca ^<br />

(Schoenemund, 1930). 74. Cronicus anomalus (Plctet); <strong>о</strong>бщий <strong>в</strong>ид, er; <strong>о</strong>бщий <strong>в</strong>ид, x 3,8, н юр<strong>а</strong>, Сибирь (Brauer, Redtenbacner,<br />

х 3,0 и ф<strong>о</strong>рцеп<strong>с</strong>, х 1,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), uangmauer, iee»j<br />

СЕМЕЙСТВО SIPHLONURIDAE KLAPALEK, 1909<br />

Н<strong>а</strong> переднем <strong>к</strong>рыле <strong>о</strong>т CuAi <strong>к</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> идут мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные из<strong>о</strong>гнутые<br />

пр<strong>о</strong>межут<strong>о</strong>чные жил<strong>к</strong>и, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

м<strong>о</strong>гут <strong>в</strong>ет<strong>в</strong>ить<strong>с</strong>я. З<strong>а</strong>дние <strong>к</strong>рылья х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Siphlonurinae;<br />

Isonychiinae.<br />

ПОДСЕМЕЙСТВО SIPHLONURINAE<br />

Klapalek, 1909 [nom. transl.<br />

Edmunds et Traver., 1954<br />

(ex Siphlonuridae Klapalek, 1909)]<br />

CuAi <strong>в</strong>ых<strong>о</strong>дит <strong>в</strong> т<strong>о</strong>ч<strong>к</strong>е т<strong>о</strong>рну<strong>с</strong><strong>а</strong> (з<strong>а</strong>дний уг<strong>о</strong>л<br />

<strong>к</strong>рыл<strong>а</strong>); пр<strong>о</strong>межут<strong>о</strong>чные жил<strong>к</strong>и, идущие <strong>о</strong>т<br />

CuA <strong>к</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, из<strong>о</strong>гнутые. Личин<strong>к</strong>и<br />

<strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong>им брюш<strong>к</strong><strong>о</strong>м и <strong>с</strong> <strong>с</strong>емью<br />

п<strong>а</strong>р<strong>а</strong>ми ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

тергиты брюш<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>о</strong><strong>с</strong>трыми б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными н<strong>а</strong>з<strong>а</strong>д. Х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ые нити<br />

гу<strong>с</strong>т<strong>о</strong> <strong>о</strong>пушены длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми (цер<strong>к</strong>и<br />

<strong>о</strong>пушены лишь <strong>с</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны)<br />

(ри<strong>с</strong>. 72). Крупные и <strong>с</strong>редней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

Не<strong>о</strong>ген — ныне. 19 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д Siphlurites Cockerel!,<br />

1923 из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

61<br />

http://jurassic.ru/<br />

ПОДСЕМЕЙСТВО ISONYCHIINAE EDMUNDS ET<br />

TRAVER, 1954<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>к</strong>рупные; CuAi <strong>в</strong>ых<strong>о</strong>дит зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>в</strong>переди <strong>о</strong>т т<strong>о</strong>ч<strong>к</strong>и т<strong>о</strong>рну<strong>с</strong><strong>а</strong>, т. е. н<strong>а</strong> термин<strong>а</strong>льн<strong>о</strong>м<br />

<strong>к</strong>р<strong>а</strong>е; пр<strong>о</strong>межут<strong>о</strong>чные жил<strong>к</strong>и, идущие<br />

<strong>о</strong>т CuAi, <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я (ри<strong>с</strong>. 73). Личин<strong>к</strong>и,<br />

<strong>к</strong>р<strong>о</strong>ме <strong>с</strong>еми п<strong>а</strong>р ж<strong>а</strong>бер, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

брюш<strong>к</strong><strong>а</strong>, имеют еще р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые и <strong>с</strong>терн<strong>о</strong>-т<strong>о</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>льные<br />

ж<strong>а</strong>бры. Средней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне и <strong>о</strong>дин <strong>в</strong>ымерший из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Cronicus Eaton, 1871. Тип р<strong>о</strong>д<strong>а</strong> — Baetis<br />

anomalus Pictet, 1854; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь). В <strong>о</strong>тличие <strong>о</strong>т <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, третий члени<strong>к</strong> ф<strong>о</strong>рцеп<strong>с</strong><strong>о</strong><strong>в</strong> длинный, п<strong>о</strong>чти<br />

р<strong>а</strong><strong>в</strong>ен длине <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> (ри<strong>с</strong>. 74). Один <strong>в</strong>ид.<br />

П<strong>а</strong>ле<strong>о</strong>ген Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО OLIGONEURI1DAE ULMER, 1920<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong><br />

<strong>в</strong><strong>с</strong>ем пр<strong>о</strong>тяжении п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong> <strong>с</strong>ближены; МА 2,<br />

<strong>с</strong>ближен<strong>а</strong> <strong>с</strong> МР Ь МР 2 <strong>с</strong> СиАь п<strong>о</strong>перечных

жил<strong>о</strong><strong>к</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong>. Личин<strong>к</strong>и лишены <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong><strong>в</strong><br />

н<strong>а</strong> <strong>в</strong>ерхних челю<strong>с</strong>тях; <strong>с</strong>емь п<strong>а</strong>р ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из неб<strong>о</strong>льш<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и, п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющей<br />

пуч<strong>о</strong><strong>к</strong> нитей. Средней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

Не<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 11<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д Protoligoneuria Demoulin,<br />

1955 из не<strong>о</strong>ген<strong>а</strong> Бр<strong>а</strong>зилии.<br />

СЕМЕЙСТВО BAETIDAE<br />

KLAPALEK, 1909<br />

Передние <strong>к</strong>рылья <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы; МА не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> м<strong>а</strong>л<strong>о</strong>. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>о</strong>чень м<strong>а</strong>лы, <strong>с</strong> д<strong>в</strong>умя-тремя пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми и немн<strong>о</strong>гими п<strong>о</strong>перечными;<br />

ин<strong>о</strong>гд<strong>а</strong> з<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Верхний<br />

<strong>о</strong>тдел <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> гл<strong>а</strong>з<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>о</strong>чень <strong>в</strong>ели<strong>к</strong>, гриб<strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

или тюрб<strong>а</strong>нн<strong>о</strong>й ф<strong>о</strong>рмы, <strong>с</strong><strong>в</strong>етл<strong>о</strong> <strong>о</strong><strong>к</strong>р<strong>а</strong>шенный.<br />

Личин<strong>к</strong>и пл<strong>а</strong><strong>в</strong><strong>а</strong>ющие, <strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong>им<br />

брюш<strong>к</strong><strong>о</strong>м, без <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>, <strong>с</strong> <strong>с</strong>емью п<strong>а</strong>р<strong>а</strong>ми<br />

пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тых ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м<br />

брюш<strong>к</strong><strong>а</strong> и <strong>с</strong> гу<strong>с</strong>т<strong>о</strong> <strong>о</strong>пушенными длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми<br />

х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ыми нитями (цер<strong>к</strong>и <strong>о</strong>пушены<br />

лишь <strong>с</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны). Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

16 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них д<strong>в</strong><strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

Baetis Leach, 1815. Тип р<strong>о</strong>д<strong>а</strong> — Ephemera<br />

bioculata Linnaeus, 1746; <strong>с</strong><strong>о</strong><strong>в</strong>ременный, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

Б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> и <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

без п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> яйце<strong>в</strong>идн<strong>о</strong>й<br />

ф<strong>о</strong>рмы <strong>с</strong> д<strong>в</strong>умя или тремя пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 75). Личин<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>с</strong>емью п<strong>а</strong>р<strong>а</strong>ми<br />

<strong>о</strong>дин<strong>о</strong>чных ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 150 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, три <strong>в</strong>ид<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин <strong>в</strong>ид <strong>в</strong> н. ми<strong>о</strong>цене<br />

С. Амери<strong>к</strong>и.<br />

Cloeon Leach, 1815. Тип р<strong>о</strong>д<strong>а</strong> — Ephemera<br />

diptera Linnaeus, 1746; <strong>с</strong><strong>о</strong><strong>в</strong>ременный, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Пер<strong>в</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> между R и RS р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> ближе<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>, чем п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

<strong>с</strong>ледующег<strong>о</strong> п<strong>о</strong>ля. У личин<strong>к</strong>и пер<strong>в</strong>ые ше<strong>с</strong>ть п<strong>а</strong>р<br />

ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> д<strong>в</strong><strong>о</strong>йные. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 70 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>дин <strong>в</strong>ид из<strong>в</strong>е<strong>с</strong>тен из<br />

<strong>в</strong>. ми<strong>о</strong>цен<strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО LEPTOPHLEBIIDAE ,<br />

WESTWOOD, 1840<br />

Крылья <strong>с</strong><strong>о</strong> мн<strong>о</strong>гими п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

з<strong>а</strong>дние <strong>к</strong>рылья имеют<strong>с</strong>я, <strong>о</strong>чень ред<strong>к</strong><strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Пр<strong>о</strong>межут<strong>о</strong>чные жил<strong>к</strong>и между MPi и<br />

МР 2 и между МР 2 и CuAi переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Ф<strong>о</strong>рцеп<strong>с</strong>ы <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong> д<strong>в</strong>умя или тремя<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>к</strong><strong>о</strong>нечными члени<strong>к</strong><strong>а</strong>ми. Личин<strong>к</strong>и <strong>с</strong><br />

<strong>с</strong>емью п<strong>а</strong>р<strong>а</strong>ми д<strong>в</strong>ур<strong>а</strong>здельных ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идных или<br />

ните<strong>в</strong>идных г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных ж<strong>а</strong>бер, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен­<br />

'62<br />

http://jurassic.ru/<br />

ных п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м брюш<strong>к</strong><strong>а</strong>. Средней <strong>в</strong>еличины<br />

и мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Н. юр<strong>а</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 45 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них д<strong>в</strong><strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>дин пл<strong>о</strong>х<strong>о</strong> изученный<br />

<strong>в</strong>ымерший р<strong>о</strong>д из н. юры.<br />

Mesobaetis Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — М. sibirica<br />

Brauer, Redtenbacher, Ganglbauer, 1889; н. юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-Б<strong>а</strong>лей).<br />

Личин<strong>к</strong><strong>а</strong> <strong>с</strong> т<strong>о</strong>н<strong>к</strong>им брюш<strong>к</strong><strong>о</strong>м, тремя х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ыми<br />

нитями и т<strong>о</strong>н<strong>к</strong>ими, нея<strong>с</strong>ными ж<strong>а</strong>бр<strong>а</strong>ми<br />

(ри<strong>с</strong>. 76). Юр<strong>а</strong> — не<strong>о</strong>ген. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из Сибири<br />

и С. Амери<strong>к</strong>и.<br />

Leptophlebia Westwood, 1840. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ephemera vespertine Linnaeus, 1746,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременный, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и <strong>в</strong> <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельны;<br />

их б<strong>а</strong>з<strong>а</strong>льные <strong>к</strong><strong>о</strong>нцы <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; передний <strong>к</strong>р<strong>а</strong>й з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>й, н<strong>о</strong> з<strong>а</strong>метн<strong>о</strong>й <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й (ри<strong>с</strong>. 77). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 <strong>в</strong>ид<strong>о</strong><strong>в</strong>;<br />

<strong>о</strong>дин <strong>в</strong>ид <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

Atatophlebia Eaton, 1881. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Ephemera australis Walker, 1850; <strong>с</strong><strong>о</strong><strong>в</strong>ременный,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия. В<strong>с</strong>е <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле, <strong>в</strong><strong>к</strong>люч<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигму,<br />

<strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми, не<strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; <strong>в</strong> <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и <strong>с</strong><strong>о</strong> <strong>с</strong>литыми, <strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я<br />

б<strong>а</strong>з<strong>а</strong>льными <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми; передний <strong>к</strong>р<strong>а</strong>й з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> без <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>в</strong>ыем<strong>к</strong>и. Не<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии,<br />

Ю. Азии, Афри<strong>к</strong>е и Ю. Амери<strong>к</strong>е; <strong>о</strong>дин <strong>в</strong>ид<br />

из ми<strong>о</strong>цен<strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО EPHEMERELHDAE<br />

KLAPALEK, 1909<br />

Крылья <strong>с</strong><strong>о</strong> мн<strong>о</strong>гими п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми,<br />

з<strong>а</strong>дние <strong>к</strong>рылья имеют<strong>с</strong>я; д<strong>в</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и <strong>в</strong> переднем <strong>к</strong>рыле между<br />

MP! и МР 2 и между МР 2 и CuAi (ри<strong>с</strong>. 78).<br />

Ф<strong>о</strong>рцеп<strong>с</strong>ы <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong> <strong>о</strong>дним <strong>к</strong><strong>о</strong>нечным члени<strong>к</strong><strong>о</strong>м.<br />

Личин<strong>к</strong><strong>а</strong> <strong>с</strong> пятью п<strong>а</strong>р<strong>а</strong>ми ж<strong>а</strong>берных ли<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не третьег<strong>о</strong> —<br />

<strong>с</strong>едьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong>; <strong>в</strong>ерхняя ч<strong>а</strong><strong>с</strong>ть<br />

<strong>к</strong><strong>а</strong>жд<strong>о</strong>й ж<strong>а</strong>бры <strong>в</strong> <strong>в</strong>иде п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>н<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и.<br />

Средней <strong>в</strong>еличины или мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

Юр<strong>а</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>дин из не<strong>о</strong>ген<strong>а</strong>; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong> из мез<strong>о</strong>з<strong>о</strong>я.<br />

Mesoneta Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — М. antiqua<br />

Brauer, Redtenbacher, Ganglbauer, 1889; н. юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-

Ри<strong>с</strong>. 77—83. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Leptophlebiidae, Ephemi<br />

77. Leptophlebia marginata (Linnaues); A —переднее <strong>к</strong>рыл<strong>о</strong>,<br />

x 5,7; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,7; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Ulmer, 1929), 78. Ephemerella<br />

ignita Poda; A — переднее <strong>к</strong>рыл<strong>о</strong>, x 5.2; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, х 5,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Schoenemund, 1930). 79. Mesoneta antiqua<br />

Brauer, Redtenbacher, Ganglbauer; <strong>о</strong>бщий <strong>в</strong>ид, x 5,0, н. юр<strong>а</strong>.<br />

В. Сибирь (<strong>о</strong>риг. ри<strong>с</strong>.). 80. Turfanella tlngi (Ping.); брюш<strong>к</strong><strong>о</strong><br />

личин<strong>к</strong>и, х 9,0, <strong>в</strong>. юр<strong>а</strong>, Ц. Азия (Demoulin, 1954). 81. Heptage-<br />

llidae, Siphlonuridae incertae sedis, Heptageniidae<br />

nia flava Rostock; A — переднее <strong>к</strong>рыл<strong>о</strong>, x 4,6; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

х 4,6; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Schoenemund, 1930). 82. Electrogenia dewalschei.<br />

Demoulin; A — <strong>о</strong>бщий <strong>в</strong>ид, x 6,0; Б — <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ние брюш<strong>к</strong><strong>а</strong>,<br />

х 40;0; п<strong>а</strong>ле<strong>о</strong>ген Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Demoulin, 1956).<br />

83. Мiocoenogenla gorbunovi ETsbernova; <strong>о</strong>бщий <strong>в</strong>ид, х 5,1;<br />

не<strong>о</strong>ген, 3. Сибирь (Черн<strong>о</strong><strong>в</strong><strong>а</strong>, 1962)<br />

http://jurassic.ru/

Б<strong>а</strong>лей). Личин<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>к</strong>рупными гл<strong>а</strong>з<strong>а</strong>ми, шир<strong>о</strong><strong>к</strong>им<br />

брюш<strong>к</strong><strong>о</strong>м, не<strong>с</strong>ущим <strong>о</strong><strong>с</strong>трые <strong>в</strong>ы<strong>с</strong>тупы н<strong>а</strong><br />

б<strong>о</strong><strong>к</strong><strong>а</strong>х <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, <strong>с</strong> <strong>о</strong>дин<strong>о</strong>чными ж<strong>а</strong>берными<br />

ли<strong>с</strong>т<strong>к</strong><strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

и <strong>с</strong> тремя х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ыми нитями (ри<strong>с</strong>. 79).<br />

Один <strong>в</strong>ид Н. юр<strong>а</strong> В. Сибири.<br />

Turfanella Demoulin, 1954. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Ephemeropsis tingi Ping, 1935; <strong>в</strong>. юр<strong>а</strong>, Кит<strong>а</strong>й<br />

Синьцзян (Турф<strong>а</strong>н). Личин<strong>к</strong><strong>а</strong>- <strong>с</strong> шир<strong>о</strong><strong>к</strong>им<br />

брюш<strong>к</strong><strong>о</strong>м, не<strong>с</strong>ущим <strong>о</strong><strong>с</strong>трые, <strong>о</strong>ттянутые <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты<br />

н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и три т<strong>о</strong>л<strong>с</strong>тые х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ые<br />

нити (ри<strong>с</strong>. 80). В. юр<strong>а</strong>. Один <strong>в</strong>ид.<br />

Вне СССР, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

р<strong>о</strong>д Ephemerella Walsh, 1862 п<strong>о</strong> нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

изученн<strong>о</strong>му м<strong>а</strong>тери<strong>а</strong>лу из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО HEPTAGENIIDEA<br />

Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> не<br />

<strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>; MPi и CuAi <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельны; <strong>в</strong> <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>о</strong>дн<strong>а</strong><br />

или д<strong>в</strong>е п<strong>а</strong>ры прямых пр<strong>о</strong>межут<strong>о</strong>чных жил<strong>о</strong><strong>к</strong>.<br />

З<strong>а</strong>дние <strong>к</strong>рылья х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины передних. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Heptageniidae и Ametropodidae;<br />

п<strong>о</strong><strong>с</strong>леднее лишь <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО HEPTAGENIIDAE ULMER, 1920<br />

В <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле д<strong>в</strong>е п<strong>а</strong>ры пр<strong>о</strong>межут<strong>о</strong>чных<br />

жил<strong>о</strong><strong>к</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong>иб<strong>о</strong>лее длинн<strong>а</strong>я н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я<br />

<strong>в</strong>близи СиА 2. Личин<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> бы<strong>с</strong>тр<strong>о</strong>те<strong>к</strong>ущих<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х или бы<strong>с</strong>трин<strong>а</strong>х, жи<strong>в</strong>ут н<strong>а</strong><br />

<strong>к</strong><strong>а</strong>мнях; пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>телые, <strong>с</strong> <strong>с</strong>емью п<strong>а</strong>р<strong>а</strong>ми ж<strong>а</strong>бер,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м брюш<strong>к</strong><strong>а</strong>; <strong>в</strong>ерхняя<br />

ч<strong>а</strong><strong>с</strong>ть ж<strong>а</strong>бры ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, нижняя — <strong>в</strong> <strong>в</strong>иде<br />

пуч<strong>к</strong><strong>а</strong> нитей (ри<strong>с</strong>. 81). Средней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 25 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong>.<br />

Electrogenia Demoulin, 1955. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Е. dewalschei Demoulin, 1955; п<strong>а</strong>ле<strong>о</strong>ген,<br />

р<strong>о</strong>д<strong>а</strong> — М. gorbunovi Tschernova, 1961; не<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь). В птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>лей<br />

переднег<strong>о</strong> и з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рылье<strong>в</strong> п<strong>о</strong>перечные жил<strong>к</strong>и<br />

пр<strong>о</strong><strong>с</strong>тые, не<strong>в</strong>ет<strong>в</strong>ящие<strong>с</strong>я (ри<strong>с</strong>. 82). Один <strong>в</strong>ид.<br />

П<strong>а</strong>ле<strong>о</strong>ген Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Miocoenogenia Tshernova, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. gorbunovi Tshernova, 1961; не<strong>о</strong>ген,<br />

3. Сибирь (<strong>с</strong>р. ми<strong>о</strong>цен, р. Тым). Личин<strong>к</strong><strong>а</strong> <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong>перед; з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и <strong>к</strong>рылье<strong>в</strong> <strong>к</strong>рупные;<br />

н<strong>о</strong>ги <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими бедр<strong>а</strong>ми; н<strong>а</strong> брюш<strong>к</strong>е и<br />

н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>в</strong>иде пятен; три х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ые<br />

нити; длин<strong>а</strong> тел<strong>а</strong> 13,5 мм (ри<strong>с</strong>. 83). Один <strong>в</strong>ид.<br />

Не<strong>о</strong>ген 3. Сибири.<br />

НАДОТРЯД MEGASECOPTEROIDEA<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

Крылья <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>о</strong>бычн<strong>о</strong> <strong>с</strong>уженные,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зующие н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящий <strong>с</strong>тебеле<strong>к</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>чень<br />

уз<strong>к</strong>ие, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие рез<strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции<br />

<strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и <strong>к</strong>реп<strong>к</strong>ие,<br />

Крылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>терты <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, не<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>я<strong>с</strong>ь н<strong>а</strong>з<strong>а</strong>д; передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

р<strong>о</strong><strong>в</strong>ный, без <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong><strong>в</strong>; х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен <strong>с</strong>тебеле<strong>к</strong><br />

— <strong>с</strong>уженн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong>;<br />

SC <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинн<strong>а</strong>я и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, неу<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>а</strong>я,<br />

ред<strong>к</strong><strong>о</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине нея<strong>с</strong>н<strong>а</strong>я;<br />

ин<strong>о</strong>гд<strong>а</strong> эт<strong>а</strong> жил<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> R; п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

пр<strong>а</strong><strong>в</strong>ильными ряд<strong>а</strong>ми. К<strong>а</strong>рб<strong>о</strong>н — пермь. Д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Eumegasecoptera, Protohymenoptera.<br />

немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>в</strong>иде ряд<strong>о</strong><strong>в</strong>;<br />

<strong>с</strong>еть, или <strong>а</strong>рхеди<strong>к</strong>тий, <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong> длинные цер<strong>к</strong>и; п<strong>а</strong>р<strong>а</strong>цер<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

нет. К<strong>а</strong>рб<strong>о</strong>н — пермь. Три <strong>о</strong>тряд<strong>а</strong>: Megasecoptera,<br />

Diaphanopterodea, Campylopterodea.<br />

ОТРЯД MEGASECOPTERA<br />

64<br />

http://jurassic.ru/<br />

ПОДОТРЯД EUMEGASECOPTERA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не<strong>с</strong>ущее п<strong>о</strong>перечные<br />

жил<strong>к</strong>и и я<strong>с</strong>ную SC; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong><br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из R и <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й RS.<br />

К<strong>а</strong>рб<strong>о</strong>н. В<strong>о</strong><strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Brodiidae, Parabrodiidae,<br />

Carbonopteridae, Foririidae, Corydaloididae,<br />

Raphidiopseidae, Mischopteridae, Vorkutiidae.

84. Bruata pnscoclnda Scudder; <strong>к</strong>рыл<strong>о</strong>, х 1,2, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1906). 85. Parabrodia sp.; х 1,9, 9р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1933). 86. Carbonoptera furcaradii Guthorl;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, x 2,5, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Guthorl,<br />

1940).^87. Foriria maculata Meunier; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 1,7;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,7, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter, 1951).<br />

88. Corydaloides sc'udderi Brongniart; A — переднее <strong>к</strong>рыл<strong>о</strong>, x 1,3;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,3; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter,<br />

1951). 89. Rhaphidiopsis diversipenna Scudder: A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,4; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,4, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

Ри<strong>с</strong>. 84—94. П<strong>о</strong>д<strong>о</strong>тряд Eumegasecoptera<br />

5 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 65<br />

http://jurassic.ru/<br />

90 А<br />

91Б<br />

90 Б<br />

91 А<br />

(Handlirsch, 1906). 90. Mischoptera nlg-a Brongniart: A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>; x 0,9; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 0,9, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Carpenter, 1951). 91. Sphecoptera gracilis Brongniart; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,2; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1.2; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Carpenter, 1951). 92. 1 schnoptilus elegans Brongniart; <strong>к</strong>рыл<strong>о</strong>,<br />

хЗ,1, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter, 1951). 93. Sibirionymen<br />

asiaticus Rohdendorf; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, x 5,4, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 94. Vorkutia tshernovi Rohdendorf;<br />

ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, х 3,7, <strong>в</strong>. пермь, В<strong>о</strong>р<strong>к</strong>ут<strong>с</strong><strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>дьнд<strong>о</strong>рф,<br />

1947).

СЕМЕЙСТВО BROD1IDAE HANDLIRSCH, 1906<br />

Вершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> притуплен<strong>а</strong>; <strong>с</strong>тебеле<strong>к</strong> <strong>к</strong>рыл<strong>а</strong><br />

длинный; SC <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля нея<strong>с</strong>н<strong>а</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>в</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong>м бе<strong>с</strong>п<strong>о</strong>ряд<strong>к</strong>е, не <strong>о</strong>бр<strong>а</strong>зуя<br />

п<strong>о</strong>перечных ряд<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 84). Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 50—60 мм. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д из<br />

<strong>в</strong>е<strong>с</strong>тф<strong>а</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong> 3." Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PARABRODIIDAE CARPENTER, 1933<br />

Стр<strong>о</strong>ение <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>;<br />

SC р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> <strong>в</strong> <strong>о</strong>тд<strong>а</strong>лении <strong>о</strong>т С и д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

<strong>в</strong>ершинн<strong>о</strong>й трети <strong>к</strong>рыл<strong>а</strong>; R <strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>м <strong>о</strong>тделе<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>тд<strong>а</strong>лен<strong>а</strong> <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные р<strong>а</strong>зделы<br />

МА и RS <strong>с</strong>литы; п<strong>о</strong><strong>с</strong>ледняя <strong>с</strong> д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями,<br />

т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и MP. Длин<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

24 мм, ширин<strong>а</strong> 7 мм (ри<strong>с</strong>. 85). К<strong>а</strong>рб<strong>о</strong>н. Один<br />

р<strong>о</strong>д из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО CARBONOPTERIDAE GUTHORL, 1940<br />

"Крыл<strong>о</strong> н<strong>а</strong> <strong>в</strong>ершине притуплённ<strong>о</strong>е; SC п<strong>о</strong>чти<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong> и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С; имеют<strong>с</strong>я<br />

передние <strong>в</strong>ет<strong>в</strong>и RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющие <strong>в</strong> R, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

н<strong>а</strong> <strong>в</strong>ершине не<strong>с</strong>ет неб<strong>о</strong>льш<strong>о</strong>й уз<strong>к</strong>ий р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>;<br />

RS <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями; М н<strong>а</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS и д<strong>а</strong>ет три п<strong>а</strong>р<strong>а</strong>ллельные<br />

<strong>в</strong>ет<strong>в</strong>и; имеют<strong>с</strong>я немн<strong>о</strong>гие <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>в</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> между СиА и MP, MP<br />

и MA, МА и RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 33 мм<br />

(ри<strong>с</strong>. 86). Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д из <strong>в</strong>е<strong>с</strong>тф<strong>а</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

яру<strong>с</strong><strong>а</strong> Герм<strong>а</strong>нии.<br />

СЕМЕЙСТВО FOR1R1IDAE HANDLIRSCH, 1919<br />

З<strong>а</strong>дние <strong>к</strong>рылья не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних,<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> притуплённые; МА + MP и СиА + СиР<br />

<strong>в</strong> <strong>в</strong>иде <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных пр<strong>о</strong><strong>с</strong>тых <strong>в</strong>ил<strong>о</strong><strong>к</strong>, не <strong>с</strong>литы<br />

друг <strong>с</strong> друг<strong>о</strong>м и <strong>с</strong> RS. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рылье<strong>в</strong> умеренн<strong>о</strong><br />

<strong>с</strong>уженные; SC д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>в</strong>ершинн<strong>о</strong>й чет<strong>в</strong>ерти<br />

<strong>к</strong>рыл<strong>а</strong> и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>в</strong>иде д<strong>в</strong>ух чет<strong>к</strong>их ряд<strong>о</strong><strong>в</strong>, з<strong>а</strong>метн<strong>о</strong><br />

<strong>в</strong><strong>о</strong>лни<strong>с</strong>тых. Длин<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> 30—35 мм<br />

(ри<strong>с</strong>. 87). В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д из <strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

яру<strong>с</strong><strong>а</strong> Фр<strong>а</strong>нции.<br />

СЕМЕЙСТВО CORYDALOIDIDAE HANDLIRSCH, 1906<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, умеренн<strong>о</strong> <strong>с</strong>уженные <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; МА ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS, <strong>а</strong> СиА —<br />

<strong>с</strong> <strong>о</strong>бщим <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м М; SC д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>в</strong>ершинн<strong>о</strong>й<br />

чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>, н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я; три —<br />

четыре <strong>в</strong>ет<strong>в</strong>и RS и MP; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

прямые, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong> <strong>в</strong>иде нея<strong>с</strong>ных трех<br />

ряд<strong>о</strong><strong>в</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> и <strong>с</strong>егменты брюш<strong>к</strong><strong>а</strong> <strong>с</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными <strong>о</strong><strong>с</strong>трыми п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льными<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 48 мм (ри<strong>с</strong>. 88).<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д из <strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong><br />

Фр<strong>а</strong>нции.<br />

66<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО RHAPHIDIOPSEIDAE HANDLIRSCH<br />

1906<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че и шире передних, <strong>с</strong><br />

б<strong>о</strong>лее уз<strong>к</strong>им <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным <strong>к</strong>р<strong>а</strong>ем; М <strong>с</strong>лит<strong>а</strong><br />

<strong>с</strong> RS. Длин<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 25—30 мм<br />

(ри<strong>с</strong>. 89).. К<strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО MISCHOPTERIDAE HANDLIRSCH, 1906;<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>уженные <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>в</strong> <strong>в</strong>иде <strong>с</strong>тебель<strong>к</strong><strong>а</strong>; длин<strong>а</strong> SC р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

д<strong>в</strong>ум третям <strong>к</strong>рыл<strong>а</strong>; МА <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м RS; СиА <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong> ИЛИ ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> MP; п<strong>о</strong>перечные жил<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>к</strong>реп<strong>к</strong>ие, <strong>в</strong><strong>о</strong>лни<strong>с</strong>тые, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong> <strong>в</strong>иде <strong>о</strong>дн<strong>о</strong>г<strong>о</strong><br />

— трех п<strong>о</strong>перечных ряд<strong>о</strong><strong>в</strong>. Длин<strong>а</strong> <strong>к</strong>рылье<strong>в</strong><br />

<strong>о</strong>т 30 д<strong>о</strong> 75 мм. К<strong>а</strong>рб<strong>о</strong>н. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

<strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, р<strong>а</strong><strong>с</strong>пределяющих<strong>с</strong>я<br />

<strong>с</strong>реди четырех п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Mischopterinae<br />

(ри<strong>с</strong>. 90), Sphecopterinae (ри<strong>с</strong>. 91),<br />

Cyclocelidinae и Ischnoptilinae (ри<strong>с</strong>. 92).<br />

СЕМЕЙСТВО VORKUTI1DAE ROHDENDORF, 1947<br />

SC т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я<br />

и я<strong>с</strong>н<strong>а</strong>я; п<strong>о</strong><strong>с</strong>ле <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления <strong>о</strong>т RS рез<strong>к</strong><strong>о</strong><br />

<strong>с</strong>ближ<strong>а</strong>ет<strong>с</strong>я <strong>с</strong> С, <strong>с</strong>уж<strong>а</strong>я <strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми<br />

<strong>о</strong>дин или д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

<strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>, ближе <strong>к</strong><br />

<strong>к</strong>р<strong>а</strong>ю. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 17—30 мм. К<strong>а</strong>рб<strong>о</strong>н —<br />

пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Sibiriohymen Rahdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. asiaticus Rohdendorf, 1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

(нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Верх<strong>о</strong>т<strong>о</strong>м<strong>с</strong><strong>к</strong><strong>о</strong>е). Один ряд п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> между <strong>в</strong>ет<strong>в</strong>ями RS и М; между<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м RS и МА п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 17 мм (ри<strong>с</strong>. 93). Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Vorkutia Rohdendorf, 1947. Тип р<strong>о</strong>д<strong>а</strong> —<br />

V. tshernovi Rohdendorf, 1947; н. пермь,<br />

В<strong>о</strong>р<strong>к</strong>ут<strong>с</strong><strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн. Д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> между RS и Си; между <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м RS и<br />

МА п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> нет. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

30 мм (ри<strong>с</strong>. 94). Один <strong>в</strong>ид. Н. пермь В<strong>о</strong>р<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

ПОДОТРЯД PROTOHYMENOPTERA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

те<strong>с</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>ближения С, SC и R; RS ин<strong>о</strong>гд<strong>а</strong> ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R. К<strong>а</strong>рб<strong>о</strong>н — пермь. Пять

Ри<strong>с</strong>.3.95—105. П<strong>о</strong>д<strong>о</strong>!<br />

95. Aspidothorax triangularis Brongniart; A — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

x 2,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter,<br />

1951). 96. Bardohymen magnipennifer G. Zalessky; <strong>к</strong>рыл<strong>о</strong>,<br />

x 1,5, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1937), 97. Sylvohymen г<strong>о</strong>bustus<br />

Martynov, <strong>к</strong>рыл<strong>о</strong>, x 2,1, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1941).<br />

98. Calohymen permianus Carpenter; <strong>к</strong>рыл<strong>о</strong>, х 2,8, н. пермь<br />

С Амери<strong>к</strong><strong>а</strong> (Carpenter, 1947). 99. Pseudohymen angustipennis<br />

Martynov (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x,l,8, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1931). 100."Ivahymen constrictus Martynov; <strong>к</strong>рыл<strong>о</strong>, х 3,3. <strong>в</strong>. пермь,<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Aspidothoracidae, Bardohymenidae,<br />

Protohymenidae, Scytohymenidae, Aspidohymenidae.<br />

СЕМЕЙСТВО ASPIDOTHORACIDAE HANDLIRSCH,<br />

1919<br />

MA чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т RS; п<strong>о</strong><strong>с</strong>ледняя не<strong>с</strong>ет<br />

четыре <strong>в</strong>ет<strong>в</strong>и <strong>в</strong> <strong>в</strong>иде гребня; R н<strong>а</strong> <strong>в</strong><strong>с</strong>ем пр<strong>о</strong>тяжении<br />

не <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

30—40 мм (ри<strong>с</strong>. 95). К<strong>а</strong>рб<strong>о</strong>н- Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

яру<strong>с</strong>е Фр<strong>а</strong>нции.<br />

[д Protohymenoptera<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 101. Protohymen permianus<br />

Tillyard; <strong>к</strong>рыл<strong>о</strong>, х 6,0, н. Пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter,<br />

1930). 102. Permohymen schucherti Tillyard; <strong>к</strong>рыл<strong>о</strong>, x 5,0, н.<br />

пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1930). 103. Tshekardohymen martynovi<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x 1,3, н. пермь, Ур<strong>а</strong>л (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1940). 104. Scytohymen extremusMartynov; <strong>к</strong>рыл<strong>о</strong>, xl,5; н. пермь,<br />

Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 105. Aspidohymen extensus Martynov;<br />

<strong>к</strong>рыл<strong>о</strong>, х 1,8, <strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1930)<br />

СЕМЕЙСТВО BARDOHYMENIDAE G. ZALESSKY,<br />

1937<br />

MA <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> или <strong>с</strong>ближен<strong>а</strong> н<strong>а</strong> незн<strong>а</strong>чительн<strong>о</strong>м<br />

пр<strong>о</strong>тяжении <strong>с</strong> RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я не<strong>с</strong>ет <strong>о</strong>т<br />

трех д<strong>о</strong> Пяти <strong>в</strong>ет<strong>в</strong>ей <strong>в</strong> <strong>в</strong>иде гребня. Н<strong>а</strong> <strong>в</strong>ершине<br />

<strong>к</strong>рыл<strong>а</strong> R <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я и <strong>с</strong>н<strong>а</strong>бжен<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ями<br />

или п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

<strong>в</strong> <strong>в</strong>иде <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> или д<strong>в</strong>ух ряд<strong>о</strong><strong>в</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

25—45 мм. Пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Bardohymen G. Zalessky, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — В. magnipennifer G. Zalessky, 1937;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

67 Б*<br />

http://jurassic.ru/

Б<strong>а</strong>рд<strong>а</strong>). Между <strong>к</strong><strong>о</strong>нц<strong>о</strong>м R и <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong> четыре<br />

п<strong>о</strong>перечные жил<strong>к</strong>и; R н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м;<br />

RS <strong>с</strong> пятью <strong>в</strong>ет<strong>в</strong>ями; МА <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

RS п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 43 мм (ри<strong>с</strong>. 96). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Sylvohymen Martynov, 1940. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. robustus Martynov, 1940; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). К<strong>о</strong>нец R<br />

не<strong>с</strong>ет р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде <strong>о</strong>дн<strong>о</strong>й — трех <strong>к</strong><strong>о</strong><strong>с</strong>ых п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; RS <strong>с</strong> четырьмя <strong>в</strong>ет<strong>в</strong>ями; МА<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

40 мм (ри<strong>с</strong>. 97). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong><br />

и С. Амери<strong>к</strong>и.<br />

Calohymen Carpenter, 1947. Тип р<strong>о</strong>д<strong>а</strong> —<br />

С. permianus Carpenter, 1947; н. пермь,<br />

С. Амери<strong>к</strong><strong>а</strong>. К<strong>о</strong>нец R пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, <strong>с</strong><strong>о</strong>единенный<br />

<strong>с</strong> <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong> тремя прямыми п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; RS <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 25 мм (ри<strong>с</strong>. 98). Один <strong>в</strong>ид. Н. пермь<br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО PROTOHYMENIDAE<br />

TILLYARD, 1924<br />

МА <strong>с</strong>лит<strong>а</strong> н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении <strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тью RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из д<strong>в</strong>ух<br />

<strong>в</strong>ет<strong>в</strong>ей — <strong>в</strong> <strong>в</strong>иде <strong>в</strong>ил<strong>к</strong>и; н<strong>а</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong> R<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зуя <strong>о</strong><strong>в</strong><strong>а</strong>льную<br />

ячей<strong>к</strong>у; п<strong>о</strong>перечные жил<strong>к</strong>и <strong>в</strong><strong>с</strong>егд<strong>а</strong> имеют<strong>с</strong>я<br />

между MP и СиА и СиР. Пермь. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Pseudohymen Martynov, 1931. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ps. angustipennis Martynov, 1931;<br />

н. пермь. Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и между R и<br />

передней <strong>в</strong>ет<strong>в</strong>ью RS; <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, з<strong>а</strong>дний ег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>й м<strong>а</strong>л<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый; длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> примерн<strong>о</strong><br />

<strong>в</strong> 5 р<strong>а</strong>з б<strong>о</strong>льше ег<strong>о</strong> ширины. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 19—27,5 мм, ширин<strong>а</strong> ег<strong>о</strong> 3,5—5,2 мм,<br />

длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 24,5 мм (ри<strong>с</strong>. 99). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong> и С. Амери<strong>к</strong>и.<br />

Ivahymen Martynov, 1931. Тип р<strong>о</strong>д<strong>а</strong> —<br />

/. constrlctus Martynov, 1931; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Д<strong>в</strong>е<br />

п<strong>о</strong>перечные жил<strong>к</strong>и между R и передней <strong>в</strong>ет<strong>в</strong>ью<br />

RS; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е, <strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>е; длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> не б<strong>о</strong>лее чем <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ег<strong>о</strong><br />

ширину. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 18 мм (ри<strong>с</strong>. 100). Один<br />

<strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Protohymen Т i 11 у <strong>а</strong> г d, 1924. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. permianus Tillyard, 1924; н. пермь, С. Амери<strong>к</strong><strong>а</strong>.<br />

Одн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> между п<strong>о</strong><strong>с</strong>ледним<br />

<strong>о</strong>трез<strong>к</strong><strong>о</strong>м R и передней <strong>в</strong>ет<strong>в</strong>ью RS; между<br />

МА и MP д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и (не <strong>с</strong>чит<strong>а</strong>я<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния МА); з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый;<br />

длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong> 3,2—4,4 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ег<strong>о</strong> ширины и р<strong>а</strong><strong>в</strong>н<strong>а</strong> 10—20 мм (ри<strong>с</strong>. 101). Пять<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь С. Амери<strong>к</strong>и.<br />

Permohymen Tillyard, 1924. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. schucherti Tillyard, 1924; н. пермь, С. Амери<strong>к</strong><strong>а</strong>.<br />

Одн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> между R и передней<br />

<strong>в</strong>ет<strong>в</strong>ью RS; между МА и MP <strong>о</strong>дн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> (не <strong>с</strong>чит<strong>а</strong>я <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния МА).<br />

З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, длин<strong>а</strong> ег<strong>о</strong><br />

<strong>в</strong> 3—3,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины и р<strong>а</strong><strong>в</strong>н<strong>а</strong> 12—<br />

13 мм (ри<strong>с</strong>. 102). Один <strong>в</strong>ид. Н. пермь С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО SCYTOHYMEN1DAE<br />

MARTYNOV, 1937<br />

МА <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я примерн<strong>о</strong> <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е<br />

<strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из д<strong>в</strong>ух <strong>в</strong>ет<strong>в</strong>ей<br />

— <strong>в</strong> <strong>в</strong>иде <strong>в</strong>ил<strong>к</strong>и; <strong>в</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong> R п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong><br />

<strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я, не <strong>о</strong>бр<strong>а</strong>зуя <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong>и; п<strong>о</strong>перечные жил<strong>к</strong>и между MP, СиА и<br />

CuP <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Tshekardohymen Rohdendorf, 1940. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. martynovi Rohdendorf, 1940;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Ряд п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> между С, R,<br />

RSi, RS 2, МА и MP, <strong>в</strong><strong>с</strong>ег<strong>о</strong> пять п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>,<br />

п<strong>о</strong> <strong>о</strong>днЪй <strong>в</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м п<strong>о</strong>ле. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

35 мм (ри<strong>с</strong>. 103). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Scytohymen Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — 5. extremus Martynov, 1937; <strong>в</strong>. пермь,<br />

Приур<strong>а</strong>лье (С<strong>а</strong><strong>в</strong>ин<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий л<strong>о</strong>г). В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле лишь д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и между R и<br />

передней <strong>в</strong>ет<strong>в</strong>ью RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 41 мм<br />

(ри<strong>с</strong>. 104). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

68<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ASPIDOHYMENIDAE<br />

MARTYNOV, 1930<br />

R н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> SC,<br />

примерн<strong>о</strong> н<strong>а</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й "трети <strong>о</strong>т<br />

<strong>в</strong>ершины <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я и идет п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

С и RS; МА <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т RS, <strong>с</strong><strong>о</strong>единяя<strong>с</strong>ь <strong>с</strong><br />

нею лишь п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из<br />

д<strong>в</strong>ух длинных <strong>в</strong>ет<strong>в</strong>ей; п<strong>о</strong>перечные жил<strong>к</strong>и имеют<strong>с</strong>я<br />

между <strong>в</strong><strong>с</strong>еми пр<strong>о</strong>д<strong>о</strong>льными <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми.<br />

Пермь. Один р<strong>о</strong>д.<br />

Aspidohymen Martynov, 1930. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. extensus Martynov, 1930; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Между R и <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я,<br />

т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>о</strong>трез<strong>о</strong><strong>к</strong> R, у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RS<br />

<strong>с</strong> рез<strong>к</strong>им темным <strong>о</strong><strong>к</strong><strong>а</strong>ймлением. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

примерн<strong>о</strong> 27—35 мм (ри<strong>с</strong>. 105). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

B. пермь Приур<strong>а</strong>лья.

Крылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>о</strong>т<strong>о</strong>д<strong>в</strong>иг<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д и <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong>д<strong>о</strong>ль тел<strong>а</strong> п<strong>о</strong>д неб<strong>о</strong>льшим угл<strong>о</strong>м <strong>к</strong><br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> ни<strong>к</strong><strong>о</strong>гд<strong>а</strong><br />

не <strong>о</strong>бр<strong>а</strong>зует <strong>с</strong>тебель<strong>к</strong><strong>а</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

реже <strong>с</strong>уженн<strong>о</strong>е; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>че д<strong>в</strong>ух третей<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем нея<strong>с</strong>н<strong>а</strong>я,<br />

<strong>о</strong>чень ред<strong>к</strong><strong>о</strong> длинн<strong>а</strong>я. Передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми, реже прям<strong>о</strong>й и <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нный.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>:<br />

Protodiaphanopterodea, Eudiaphanopterodea,<br />

Metadiaphanopterodea.<br />

ПОДОТРЯД<br />

PROTODIAPHANOPTERODEA<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, умеренн<strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренные;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; RS имеет гребенч<strong>а</strong>т<strong>о</strong>е<br />

<strong>с</strong>тр<strong>о</strong>ение и не<strong>с</strong>ет д<strong>о</strong> пяти <strong>в</strong>ет<strong>в</strong>ей, <strong>в</strong>ых<strong>о</strong>дящих<br />

н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 12 мм<br />

(ри<strong>с</strong>. 106 А, Б). К<strong>а</strong>рб<strong>о</strong>н. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Sypharopteridae <strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

ПОДОТРЯД<br />

EUDIAPHANOPTERODEA<br />

Крылья <strong>о</strong>бычн<strong>о</strong> притуплённые; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ция<br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> — передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый<br />

<strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й SC; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле нер<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е и без <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong> <strong>к</strong>реп<strong>к</strong>их п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны,<br />

ин<strong>о</strong>гд<strong>а</strong> их зн<strong>а</strong>чительн<strong>о</strong> меньше, н<strong>о</strong> <strong>о</strong>ни<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>в</strong> <strong>в</strong>иде<br />

<strong>в</strong>ет<strong>в</strong>ей SC и R (п<strong>о</strong> <strong>к</strong>р<strong>а</strong>йней мере <strong>о</strong>дн<strong>а</strong> <strong>к</strong>реп<strong>к</strong><strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>). К<strong>а</strong>рб<strong>о</strong>н — пермь. Семь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Рг<strong>о</strong>choropteridae,<br />

Diaphanopteritidae, Diaphanopteridae,<br />

Elmoidae, Kaltanelmoidae, Parelmoidae,<br />

Martynoviidae.<br />

СЕМЕЙСТВО PROCHOROPTERIDAE HANDLIRSCH,<br />

1911<br />

SC з<strong>а</strong>метн<strong>о</strong> длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; <strong>о</strong><strong>к</strong>руглые пятн<strong>а</strong> н<strong>а</strong> <strong>к</strong>рыльях <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящее из<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong><br />

<strong>к</strong>рылье<strong>в</strong> 20—32 мм (ри<strong>с</strong>. 107). К<strong>а</strong>рб.<strong>о</strong>н. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong> из <strong>с</strong>р. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО DIAPHANOPTERITIDAE HANDLIRSCH,<br />

1911<br />

SC <strong>о</strong>чень длинн<strong>а</strong>я. Н<strong>а</strong> <strong>к</strong>рыльях <strong>о</strong><strong>к</strong>руглые<br />

темные пятн<strong>а</strong> <strong>с</strong>реди жил<strong>о</strong><strong>к</strong> <strong>с</strong>и<strong>с</strong>тем RS и М. М<strong>а</strong>л<strong>о</strong>из<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>я<br />

групп<strong>а</strong> из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Один р<strong>о</strong>д.<br />

ОТРЯД DIAPHANOPTERODEA<br />

СЕМЕЙСТВО DIAPHANOPTERIDAE<br />

HANDLIRSCH, 1906<br />

SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> примерн<strong>о</strong> д<strong>в</strong>ум третям переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>. Н<strong>а</strong> <strong>к</strong>рыльях чет<strong>к</strong>ие темные <strong>о</strong><strong>к</strong>руглые<br />

пятн<strong>а</strong> <strong>в</strong> р<strong>а</strong>зличных п<strong>о</strong>лях <strong>к</strong>рыл<strong>а</strong>. Ан<strong>а</strong>льные<br />

жил<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных <strong>в</strong>ет<strong>в</strong>ей, <strong>о</strong>тх<strong>о</strong>дящих<br />

<strong>о</strong>т <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н; Три р<strong>о</strong>д<strong>а</strong>.<br />

Diaphanoptera Brongniart, 1893. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. munieri Brongniart, 1893; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Фр<strong>а</strong>нция (<strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, К<strong>о</strong>м<strong>а</strong>нтри). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми, умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е; <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я;<br />

<strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, б<strong>о</strong>лее чем <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

ширины. Длин<strong>а</strong> 35—37 мм (ри<strong>с</strong>: 108). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>» Фр<strong>а</strong>нции.<br />

Philiasptilon М. Zalessky, 1931. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ph. maculosum М. Zalessky, 1931; <strong>с</strong>р.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Черемич<strong>к</strong>ин<strong>о</strong>).<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC <strong>с</strong><br />

един<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> R; н<strong>а</strong> б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>ля п<strong>о</strong>перечные<br />

жил<strong>к</strong>и нея<strong>с</strong>ны; RS <strong>с</strong> ше<strong>с</strong>тью <strong>в</strong>ет<strong>в</strong>ями <strong>в</strong> <strong>в</strong>иде<br />

гребня. Пятн<strong>а</strong> з<strong>а</strong>метны лишь н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>.<br />

Крыл<strong>о</strong> менее чем <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

ширины — ег<strong>о</strong> длин<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 45 мм (ри<strong>с</strong>. 109).<br />

Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tchirkovaea М. Zalessky, 1931. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. guttata М. Zalessky, 1931; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Черемич<strong>к</strong>ин<strong>о</strong>).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. SC длиннее<br />

д<strong>в</strong>ух третей длины п<strong>о</strong>перечнег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

<strong>о</strong>чень <strong>в</strong>ыпу<strong>к</strong>лый; три <strong>в</strong>ет<strong>в</strong>и SC; пятн<strong>а</strong> р<strong>а</strong>з<strong>в</strong>иты<br />

п<strong>о</strong>чти <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех п<strong>о</strong>лях <strong>к</strong>рыл<strong>а</strong>. Крыл<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, длин<strong>а</strong> ег<strong>о</strong> лишь <strong>в</strong> 2—2,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ширины и р<strong>а</strong><strong>в</strong>н<strong>а</strong> 56 мм (ри<strong>с</strong>. ПО). Один <strong>в</strong>ид.<br />

К<strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО ELMOIDAE TILLYARD, 1937<br />

Крылья р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>с</strong>уж<strong>а</strong>ют<strong>с</strong>я <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию.<br />

SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R и п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong> длине переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>г<strong>о</strong> и не<strong>с</strong>ущег<strong>о</strong><br />

<strong>о</strong>дин<strong>о</strong>чные п<strong>о</strong>перечные жил<strong>к</strong>и; МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

MP <strong>с</strong> длинным р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuA и CuP пр<strong>о</strong><strong>с</strong>тые;<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong> чи<strong>с</strong>ле <strong>о</strong>дн<strong>о</strong>й — д<strong>в</strong>ух <strong>в</strong>ет<strong>в</strong>ей.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 12 мм, ширин<strong>а</strong> б<strong>о</strong>лее 3 мм<br />

(ри<strong>с</strong>. 111). Пермь. Один р<strong>о</strong>д из н. перми<br />

С. Амери<strong>к</strong>и.<br />

69<br />

http://jurassic.ru/

Ри<strong>с</strong>, 106—114. П<strong>о</strong>д<strong>о</strong>тряды Protodii<br />

106. Sypharoptera pneuma Handlirsch; A — <strong>о</strong>бщий <strong>в</strong>ид и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>г<strong>о</strong>,<br />

x 2,8; Б — <strong>к</strong>рыл<strong>о</strong>, х 4.7, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Handlirsch,<br />

1911). 107. Prochoropttra calopteryx Handlirsch; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, x 3,1; £ — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,1, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

(Handlirsch, 1911). 108. Diaphanoptera munieri Brongniart;<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,4, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1906). 109. Phlliasptilon<br />

maculosum M. Zalessky, <strong>к</strong>рыл<strong>о</strong>, x 1,2, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (M. З<strong>а</strong> л <strong>в</strong>е<strong>с</strong><strong>к</strong>ий, 1931). 110. Tchirkovaea guttata<br />

М. Zalessky; <strong>к</strong>рыл<strong>о</strong>, х 0,94, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М. 3<strong>а</strong>-<br />

hanopterodea, Eudiaphanopterodea<br />

ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1931). 111. Elmoa trlsecta Tillyard; А— переднее <strong>к</strong>рыл<strong>о</strong>,<br />

x 5,6,; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,6; н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter,<br />

1943). 112. Parelmoa revelata Carpenter; <strong>к</strong>рыл<strong>о</strong>, x 4,7, н.<br />

пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1947). 113. Kaltanelmoa sibirlca<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x 5,7, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 114. Martynovia instgnis Tillyard; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 3,75; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 75, н. пе<strong>о</strong>мь, С. Амери<strong>к</strong><strong>а</strong><br />

(Carpenter, 1943).<br />

70<br />

http://jurassic.ru/

СЕМЕЙСТВО PARELMOIDAE ROHDENDORF,<br />

FAM. NOV.<br />

SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С и зн<strong>а</strong>чительн<strong>о</strong> длиннее<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

<strong>о</strong>бычн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый; MP <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; три <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рыл<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 15—<br />

22 мм (ри<strong>с</strong>. 112). Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Pseudelmoa<br />

Carpenter, 1947 и Parelmoa Carpenter,<br />

1947 из н. перми С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО KALTANELMOIDAE ROHDENDORF,<br />

1961<br />

SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С и R, длин<strong>а</strong> ее п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>г<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я;<br />

MP <strong>с</strong> длинным р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuA н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong><br />

<strong>к</strong>реп<strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я<br />

<strong>о</strong>т М, <strong>с</strong><strong>о</strong>единяющ<strong>а</strong>я<strong>с</strong>я <strong>с</strong> нею п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong> чи<strong>с</strong>ле трех <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных<br />

<strong>в</strong>ет<strong>в</strong>ей. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> умеренн<strong>о</strong><br />

<strong>с</strong>уженн<strong>о</strong>е. Пермь. Один р<strong>о</strong>д.<br />

т<strong>а</strong>н). Между С и SC четыре п<strong>о</strong>перечные жил<strong>к</strong>и;<br />

R <strong>с</strong> лег<strong>к</strong>им изгиб<strong>о</strong>м <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и прям<strong>а</strong>я<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м пр<strong>о</strong>тяжении, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>с</strong>х<strong>о</strong>дящ<strong>а</strong>я<strong>с</strong>я<br />

<strong>с</strong> С; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л М з<strong>а</strong>метн<strong>о</strong> длиннее<br />

<strong>о</strong>бщег<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 12 мм,<br />

ширин<strong>а</strong> 4,2 мм (ри<strong>с</strong>. 113). Один <strong>в</strong>ид. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО MARTYNOVIIDAE TILLYARD, 1932<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, не длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я, рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>г<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й трети<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>к</strong><strong>о</strong>нец SC <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дный, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С или<br />

<strong>в</strong> R; между SC и С изменчи<strong>в</strong><strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; MA, MP, CuA и CuP пр<strong>о</strong><strong>с</strong>тые, без<br />

<strong>в</strong>ет<strong>в</strong>ей. Длин<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> 7—22 мм (ри<strong>с</strong>. 114).<br />

Пермь. Три р<strong>о</strong>д<strong>а</strong> из н. перми С. Амери<strong>к</strong>и.<br />

ПОДОТРЯД<br />

METADIAPHANOPTERODEA<br />

Kaltanelmoa Rohdendorf, 1957. Тип. Вершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> притуплённ<strong>а</strong>я; <strong>к</strong>рыл<strong>о</strong> рез<strong>к</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>а</strong> — К. sibirica Rohdendorf, 1957; н. пермь, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>, без п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>к</strong><strong>о</strong>­<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>л<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле. SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

Ри<strong>с</strong>. 115—122. П<strong>о</strong>д<strong>о</strong>тряд Metadiaphanopterodea и <strong>о</strong>тряд Campylopterodea<br />

115. Blarmohymen bardense G. Zalessky; <strong>к</strong>рыл<strong>о</strong>, x 2,0, н. пермь,<br />

Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1937). 116. Asthenohymen dunbarl Tillyard<br />

(ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); х 3,2, н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Laurentiaux,<br />

1953). 11/. Asthenohymen uraltcus G. Zalessky; <strong>к</strong>рыл<strong>о</strong>, x 6,6, н.<br />

пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1939). 118. Aenigmatidia kaltanica<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, х 15,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 119. Kulo/a expansa Martynov, <strong>к</strong>рыл<strong>о</strong>, х 2,0,<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). 120. Parakuloja<br />

paurovenosa (Martynov); <strong>к</strong>рыл<strong>о</strong>, х 2.5, <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>. 1931). 121. Eukulola cubttalts (Martynov); <strong>к</strong>рыл<strong>о</strong>,<br />

х 1,9, <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>. 1931). 122.<br />

Campyloptera eatonl Brongniart; <strong>к</strong>рыл<strong>о</strong>, х 0,8, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter, 1943)<br />

71<br />

http://jurassic.ru/

<strong>к</strong>р<strong>а</strong>йней <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> м<strong>а</strong>л<strong>о</strong>, ин<strong>о</strong>гд<strong>а</strong> их нет<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем. Пермь. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Biarmohymenidae,<br />

Asthenohymenidae, Aenigmatidiidae,<br />

Kulojidae.<br />

СЕМЕЙСТВО BIARMOHYMENIDAE G. ZALESSKY, 1937<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>, <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь д<strong>а</strong>лее <strong>с</strong> С и R<br />

и уч<strong>а</strong><strong>с</strong>т<strong>в</strong>уя <strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нии длинн<strong>о</strong>й птер<strong>о</strong><strong>с</strong>тигмы;<br />

МА и MP пр<strong>о</strong><strong>с</strong>тые, без <strong>в</strong>ет<strong>в</strong>ей, пер<strong>в</strong><strong>а</strong>я из<br />

них н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

RS; CuA, CuP и А н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми.<br />

Пермь. Один р<strong>о</strong>д.<br />

Biarmohymen G. Zalessky, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — В. bardense G. Zalessky, 1937; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Б<strong>а</strong>рд<strong>а</strong>). Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; RS дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я, деля<strong>с</strong>ь н<strong>а</strong> четыре <strong>в</strong>ет<strong>в</strong>и;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>трез<strong>о</strong><strong>к</strong><br />

МА <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 24 мм, ширин<strong>а</strong> 7 мм (ри<strong>с</strong>. 115). Один<br />

<strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

СЕМЕЙСТВО ASTHENOHYMENIDAE TILLYARD, 1924<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>уженн<strong>о</strong>е; <strong>к</strong>рыл<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

<strong>с</strong> пл<strong>о</strong>х<strong>о</strong> р<strong>а</strong>зличим<strong>о</strong>й <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

SC; RS д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; МА и MP без <strong>в</strong>ет<strong>в</strong>ей,<br />

пер<strong>в</strong><strong>а</strong>я <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; CuA и CuP пр<strong>о</strong><strong>с</strong>тые; немн<strong>о</strong>гие чет<strong>к</strong>ие<br />

п<strong>о</strong>перечные жил<strong>к</strong>и между <strong>в</strong><strong>с</strong>еми пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми; А <strong>в</strong> чи<strong>с</strong>ле <strong>о</strong>дн<strong>о</strong>й <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й<br />

жил<strong>к</strong>и. Пермь. Один р<strong>о</strong>д.<br />

Asthenohymen Tillyard, 1924. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. dunbari Tillyard, 1924; н. пермь,<br />

С. Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). Вершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> <strong>к</strong><strong>о</strong>нце<br />

передней <strong>в</strong>ет<strong>в</strong>и RS <strong>с</strong> з<strong>а</strong>метным <strong>с</strong>л<strong>а</strong>бым з<strong>а</strong><strong>о</strong><strong>с</strong>трением.<br />

Между R и RS <strong>о</strong>дн<strong>а</strong> чет<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>; <strong>в</strong>ершин<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т 4,3 д<strong>о</strong> 10 мм, ширин<strong>а</strong> <strong>о</strong>т 1,5<br />

д<strong>о</strong> 3 мм (ри<strong>с</strong>. 116, 117). Н. пермь. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong><br />

<strong>с</strong> Ур<strong>а</strong>л<strong>а</strong>, <strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong> из С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО AENIGMATIDIIDAE ROHDENDORF, 1961<br />

SC нея<strong>с</strong>н<strong>а</strong>я, редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я;R <strong>о</strong>тд<strong>а</strong>лен<strong>а</strong> <strong>о</strong>т<br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong>реп<strong>к</strong><strong>а</strong>я; RS <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями; МА<br />

Крыл<strong>о</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным <strong>с</strong>тебель<strong>к</strong><strong>о</strong>м,<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>с</strong> прямым передним <strong>к</strong>р<strong>а</strong>ем<br />

и <strong>в</strong>ыпу<strong>к</strong>лым з<strong>а</strong>дним, <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е — не менее<br />

чем <strong>в</strong> 6 р<strong>а</strong>з длиннее ширины; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е;<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я — не б<strong>о</strong>льше трети <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С и <strong>о</strong>бр<strong>а</strong>зует узел<strong>о</strong><strong>к</strong>; RS <strong>с</strong> пятью<br />

<strong>в</strong>ет<strong>в</strong>ями, мн<strong>о</strong>гие из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых интер<strong>к</strong><strong>а</strong>лярные;<br />

ОТРЯД CAMPYLOPTERODEA<br />

72<br />

http://jurassic.ru/<br />

<strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т RS и <strong>о</strong>бр<strong>а</strong>зует <strong>с</strong> <strong>в</strong>ильч<strong>а</strong>т<strong>о</strong>й MP <strong>о</strong>бщую<br />

<strong>с</strong>и<strong>с</strong>тему; CuA, CuP и А пр<strong>о</strong><strong>с</strong>тые. Пермь.<br />

Один р<strong>о</strong>д.<br />

Aenigmatidia Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. kaltanica Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Крупн<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и между<br />

<strong>в</strong>ет<strong>в</strong>ями RS, МА и между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями М<br />

и Си. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,75 мм, ширин<strong>а</strong> 1,3 мм<br />

(ри<strong>с</strong>. 118). Один <strong>в</strong>ид. Н, пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО KULOJIDAE MARTYNOV, 1928<br />

SC н<strong>а</strong>цел<strong>о</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> С; R р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>о</strong>тд<strong>а</strong>лен<strong>а</strong><br />

<strong>о</strong>т <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong>реп<strong>к</strong><strong>а</strong>я; RS <strong>с</strong> д<strong>в</strong>умя пр<strong>о</strong><strong>с</strong>тыми<br />

<strong>в</strong>ет<strong>в</strong>ями; М <strong>в</strong> <strong>в</strong>иде <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong>й, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

длинн<strong>о</strong>й пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й <strong>в</strong>ил<strong>к</strong>и (МА + MP);<br />

Си <strong>в</strong> <strong>в</strong>иде пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й или <strong>в</strong>ильч<strong>а</strong>т<strong>о</strong>й жил<strong>к</strong>и; А <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. Пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Kuloja Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong> — К. ехpansa<br />

Martynov, 1928; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). К<strong>о</strong>нец R,<br />

передняя <strong>в</strong>ет<strong>в</strong>ь RS и <strong>в</strong>ет<strong>в</strong>и М п<strong>а</strong>р<strong>а</strong>ллельные,<br />

нер<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я; <strong>к</strong><strong>о</strong>нец R з<strong>а</strong>метн<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; з<strong>а</strong>метны <strong>с</strong>леды т<strong>о</strong>н<strong>к</strong>их п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм, ширин<strong>а</strong><br />

11 мм (ри<strong>с</strong>. 119). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Parakutoja Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong>—Kuloja paurovenosa Martynov, 1931; <strong>в</strong>.<br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). К<strong>о</strong>нец R, передняя <strong>в</strong>ет<strong>в</strong>ь RS, т<strong>а</strong><strong>к</strong> же<br />

<strong>к</strong><strong>а</strong><strong>к</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и RS и М, з<strong>а</strong>метн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я;<br />

<strong>к</strong><strong>о</strong>нец R п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> <strong>с</strong>ближ<strong>а</strong>ет<strong>с</strong>я <strong>с</strong> С; Си<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, без <strong>в</strong>ет<strong>в</strong>и; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> нет.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 25 мм, ширин<strong>а</strong> 8,4 мм<br />

(ри<strong>с</strong>. 120). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Eukuloja Rohdendorf, gen. nov. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Kuloja cubitatis Martynov, 1931; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и RS и М рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я; <strong>к</strong><strong>о</strong>нец R<br />

з<strong>а</strong>метн<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; Си <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из<br />

д<strong>в</strong>ух чет<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей — CuA и CuP; п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> нет. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 23—28 мм, ширин<strong>а</strong><br />

7,5—9 мм (ри<strong>с</strong>. 121). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

МА, MP, CuA и CuP пр<strong>о</strong><strong>с</strong>тые; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные, <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех п<strong>о</strong>лях <strong>к</strong>реп<strong>к</strong>ие;<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены бе<strong>с</strong>п<strong>о</strong>ряд<strong>о</strong>чн<strong>о</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

64 мм, ширин<strong>а</strong> 11 мм (ри<strong>с</strong>. 122). К<strong>а</strong>рб<strong>о</strong>н. Одн<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Campylopteridae <strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м из<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Фр<strong>а</strong>нции.

НАДОТРЯД ODONATOIDEA.<br />

Крылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> удлиненные, <strong>в</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>а</strong>з длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины, <strong>с</strong> прямым или <strong>в</strong><strong>о</strong>гнутым<br />

передним <strong>к</strong>р<strong>а</strong>ем. MP и СиА <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>иты 1<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; п<strong>о</strong>перечные жил­<br />

СТРЕКОЗООБРАЗНЫЕ<br />

ОТРЯД ODONATA. СТРЕКОЗЫ<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне. Ше<strong>с</strong>ть п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong>: Б<strong>о</strong>теganisoptera,<br />

Meganisoptera, Protanisoptera, Zygoptera,<br />

Anisozygoptera, Anisoptera.<br />

ПОДОТРЯД EOMEGANISOPTERA<br />

CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я; R<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т <strong>о</strong>бщег<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> МА + RS; МА <strong>с</strong><br />

немн<strong>о</strong>гими <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; узел<strong>о</strong><strong>к</strong> и п<strong>о</strong>дузел<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

жил<strong>к</strong>и, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, не р<strong>а</strong>з<strong>в</strong>иты;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и ред<strong>к</strong>ие и длинные. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60 мм (ри<strong>с</strong>. 123). Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Одн<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Erasipteridae <strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м из<br />

н<strong>а</strong>мюр<strong>а</strong> Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

Ри<strong>с</strong>. 123—130. П<strong>о</strong>д<strong>о</strong>тряды Eomeganisoptera, Meganisoptera<br />

123. Eraslpteron larlschl Pruvost; <strong>к</strong>рыл<strong>о</strong>, x 1,0, я. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия<br />

(Laurentiaux, 1953). 124. Oltgotypus tillyardl Carpenter;<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,9. н. пермь, С Амери<strong>к</strong><strong>а</strong> (Carpenter, 1947). 125.<br />

Meganeura monyl Brongniart, <strong>к</strong>рыл<strong>о</strong>, x 0,25, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1906). 126. Arctotypus slnuatus Martynov; <strong>к</strong>рыл<strong>о</strong>,<br />

x 0,75, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 127.<br />

<strong>к</strong>и <strong>к</strong>реп<strong>к</strong>ие; <strong>а</strong>рхеди<strong>к</strong>тия <strong>в</strong><strong>с</strong>егд<strong>а</strong> нет. Тел<strong>о</strong>><br />

удлиненн<strong>о</strong>е, длинные цер<strong>к</strong>и и п<strong>а</strong>р<strong>а</strong>цер<strong>к</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. К<strong>а</strong>рб<strong>о</strong>н — ныне. Один <strong>о</strong>тряд<br />

Odonata.<br />

ПОДОТРЯД<br />

MEGANISOPTERA.<br />

ГИГАНТСКИЕ СТРЕКОЗЫ<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й, без <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ти<br />

или изл<strong>о</strong>м<strong>а</strong>: узел<strong>о</strong><strong>к</strong> не р<strong>а</strong>з<strong>в</strong>ит. CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

<strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я; п<strong>о</strong>перечныежил<strong>к</strong>и<br />

<strong>о</strong>чень мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны; <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бые ячей<strong>к</strong>и — треуг<strong>о</strong>льни<strong>к</strong> и четырехуг<strong>о</strong>льни<strong>к</strong><br />

— <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Р<strong>а</strong>змеры' <strong>к</strong>рупные.<br />

К<strong>а</strong>рб<strong>о</strong>н — мез<strong>о</strong>з<strong>о</strong>й. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Paralogidae, Meganeuridae, Triadotypidae, Liadotypidae.<br />

Kargalotypus kargaltnsis (Martynov); <strong>к</strong>рыл<strong>о</strong>, х 1,1, Пермь, Ю.Приур<strong>а</strong>лье<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). '28. Petrotypus multivenosus Q. Zalessky;<br />

<strong>к</strong>рыл<strong>о</strong>, x 0,75, <strong>в</strong>. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1950). 129.<br />

Trladotypus guillaumei Laurentiaux et Grauvogel; <strong>к</strong>рыл<strong>о</strong>, x 0,75,<br />

н. три<strong>а</strong><strong>с</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Laurentiaux, 1953). 130. Ltadotypus reltctu&<br />

Martynov; <strong>к</strong>рыл<strong>о</strong>, x 1,1, <strong>в</strong>. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937).<br />

73<br />

http://jurassic.ru/

СЕМЕЙСТВО PARALOGIDAE HANDLIRSCH, 1906<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, не пре<strong>в</strong>ыш<strong>а</strong>ющ<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; RSi + RS 2HRS 3 + RS 4<br />

рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я; рудимент CuA <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 40—60 мм (ри<strong>с</strong>. 124).<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО MEGANEURIDAE HANDLIRSCH, 1907<br />

SC длинн<strong>а</strong>я — не <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я, <strong>о</strong>бычн<strong>о</strong> длиннее; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>тх<strong>о</strong>дят<br />

<strong>о</strong>дн<strong>а</strong> <strong>о</strong>т друг<strong>о</strong>й п<strong>о</strong>д неб<strong>о</strong>льшими угл<strong>а</strong>ми.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Meganeurinae,<br />

Typinae.<br />

ПОДСЕМЕЙСТВО MEGANEURINAE HANDLIRSCH,<br />

1919<br />

Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е, п<strong>о</strong>чти д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющее<br />

<strong>с</strong>ередины переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; з<strong>а</strong>дняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

RS <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>, нед<strong>а</strong>ле<strong>к</strong><strong>о</strong><br />

<strong>о</strong>т <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления МА. Р<strong>а</strong>змеры <strong>о</strong>чень <strong>к</strong>рупные.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 260—310 мм (ри<strong>с</strong>. 125). К<strong>а</strong>рб<strong>о</strong>н<br />

— пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и н. перми С. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО TYPINAE HANDLIRSCH, 1919<br />

Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е — зн<strong>а</strong>чительн<strong>о</strong><br />

меньше трети переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь<br />

RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RSi, причем,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления<br />

МА. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> не б<strong>о</strong>лее 250 мм.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Arctotypus Martynov, 1931. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. sinuatus Martynov, 1931; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е; SC зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>ей МА<br />

<strong>с</strong>емь-<strong>в</strong><strong>о</strong><strong>с</strong>емь, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле и <strong>с</strong><strong>а</strong>мых т<strong>о</strong>н<strong>к</strong>их;<br />

между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ными ч<strong>а</strong><strong>с</strong>тями CuP и А <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>, з<strong>а</strong>метн<strong>о</strong> б<strong>о</strong>лее пл<strong>о</strong>тн<strong>а</strong>я,<br />

чем другие. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 90—125 мм<br />

(ри<strong>с</strong>. 126). Пермь. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из Приур<strong>а</strong>лья,<br />

<strong>о</strong>дин <strong>в</strong>ид из Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.; из<strong>в</strong>е<strong>с</strong>тен т<strong>а</strong><strong>к</strong>же<br />

<strong>о</strong>дин м<strong>а</strong>л<strong>о</strong>изученный <strong>в</strong>ид из <strong>к</strong>унгур<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

яру<strong>с</strong><strong>а</strong> Ур<strong>а</strong>л<strong>а</strong>.<br />

Kargalotypus Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> Megatypus kargalensis Martynov,<br />

1932; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>).<br />

Стр<strong>о</strong>ение пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>, т<strong>а</strong><strong>к</strong><br />

же <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>с</strong>тр<strong>о</strong>ение <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>. SC<br />

не длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины' переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я;<br />

М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>в</strong>ет<strong>в</strong>ления<br />

RS, <strong>о</strong>бр<strong>а</strong>зуя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, н<strong>о</strong> шир<strong>о</strong><strong>к</strong>ий <strong>в</strong>еер из<br />

14 т<strong>о</strong>н<strong>к</strong>их жил<strong>о</strong><strong>к</strong>; между <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми М и CuP,<br />

CuP и А, МА и RS д<strong>в</strong>урядные ячей<strong>к</strong>и, р<strong>а</strong>зделенные<br />

зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 135 мм (ри<strong>с</strong>. 127).<br />

Один <strong>в</strong>ид. Пермь Ю. Приур<strong>а</strong>лья.<br />

Petrotypus G. Zalessky, 1950. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. multivenosus G. Zalessky, 1950; <strong>в</strong>.<br />

пермь, Приур<strong>а</strong>лье (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Вишер<strong>а</strong>).<br />

Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>; SC зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я;<br />

МА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> RS и <strong>о</strong>бр<strong>а</strong>зует<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные, те<strong>с</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>в</strong>ет<strong>в</strong>и (<strong>с</strong><strong>в</strong>ыше де<strong>в</strong>яти). CuP <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> приближен<strong>а</strong> <strong>к</strong> <strong>о</strong>бщему <strong>с</strong>т<strong>в</strong><strong>о</strong>лу<br />

R + М; CuP и А рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнутые; д<strong>в</strong>урядны'х<br />

ячее<strong>к</strong> нет. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 120 мм<br />

(ри<strong>с</strong>. 128). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Вне СССР: Meganeurula Handlirsch, 1906;<br />

Typus Sellards, 1907; Megatypus Tillyard, 1925.<br />

СЕМЕЙСТВО TRIADOTYPIDAE GRAUVOGEL<br />

ET LAURENTIAUX, 1952<br />

Передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й. SC меньше трети<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; МА и CuP пр<strong>о</strong><strong>с</strong>тые, у ме<strong>с</strong>т<strong>а</strong><br />

<strong>в</strong>п<strong>а</strong>дения <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ближенные; п<strong>о</strong><strong>с</strong>ледняя<br />

жил<strong>к</strong><strong>а</strong> умеренн<strong>о</strong> <strong>в</strong><strong>о</strong>лни<strong>с</strong>т<strong>а</strong>я; <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> CuP <strong>в</strong>ет<strong>о</strong>ч<strong>к</strong><strong>а</strong> CuA р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> <strong>в</strong><br />

<strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и А мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

и <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>ты. П<strong>о</strong>перечные<br />

жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны и непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

(ри<strong>с</strong>. 129).Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 125мм.<br />

Мез<strong>о</strong>з<strong>о</strong>й. Один р<strong>о</strong>д из н. три<strong>а</strong><strong>с</strong><strong>а</strong> Фр<strong>а</strong>нции.<br />

74<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО LIADOTYPIDAE MARTYNOV, 1937<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й, <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый. SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я и д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>, будучи <strong>о</strong>чень <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> С; МА <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; CuP <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т А. Мез<strong>о</strong>з<strong>о</strong>й. Один р<strong>о</strong>д из н.<br />

юры Ср. Азии.<br />

Liadotypus Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

L. relictus Martynov, 1937; н. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). От ме<strong>с</strong>т<strong>а</strong> <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления<br />

RS <strong>о</strong>т М <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

<strong>о</strong>дну линию д<strong>в</strong>е <strong>к</strong><strong>о</strong><strong>с</strong>ые п<strong>о</strong>перечные жил<strong>к</strong>и, з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти RS; между<br />

CuP и А <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>ля р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

мн<strong>о</strong>г<strong>о</strong>рядные ячей<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующие нея<strong>с</strong>ные<br />

пр<strong>о</strong>д<strong>о</strong>льные <strong>в</strong>ет<strong>в</strong>и. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

67 мм (ри<strong>с</strong>. 130). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

ПОДОТРЯД PROTANISOPTERA.<br />

ПРОТАНИЗОПТЕРЫ<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>гнут. RS<br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не или ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

SC; CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рыл<strong>а</strong> умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е или <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е,<br />

не<strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>е; четырехуг<strong>о</strong>льни<strong>к</strong> и треуг<strong>о</strong>льни<strong>к</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Пермь. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Kaltanoneuridae,<br />

Ditaxineuridae, Polytaxineuridae,<br />

Callimokaltaniidae, Permaeschnidae.

Ри<strong>с</strong>. 131—139. П<strong>о</strong>д<br />

131. Kaltanoneura bartenevi Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, х 5,6, н. пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 132. Dltaxineurella stlgmalis<br />

Martynov; <strong>к</strong>рыл<strong>о</strong>, x 2,6, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1941).<br />

133. Dltaxineurella uralensls (G. Zalessky); <strong>к</strong>рыл<strong>о</strong>, x 2,0, н. пермь,<br />

Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1955). 134. Dltaxineura anomalostlgma Tillyard;<br />

<strong>к</strong>рыл<strong>о</strong>, х 3,3, н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1931). 135.<br />

Pholidoptllon sp.; <strong>к</strong>рыл<strong>о</strong>, х 1,7, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

СЕМЕЙСТВО KALTANONEURIDAE ROHDENDORF,<br />

1961<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong>дуг<strong>о</strong><strong>в</strong>идн<strong>о</strong>-<strong>в</strong><strong>о</strong>гнутый;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> немн<strong>о</strong>гими п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> длинн<strong>а</strong>я, <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я;<br />

М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, RS <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями. Пермь. Один<br />

р<strong>о</strong>д.<br />

Kaltanoneura Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К- bartenevi Rohdendorf, 1961; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> удлиненн<strong>а</strong>я и <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле зн<strong>а</strong>чительн<strong>о</strong> шире<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>, п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

ше<strong>с</strong>ть; между RS5 и МА пять, между МА<br />

и CuP ше<strong>с</strong>ть п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле б<strong>о</strong>лее чем <strong>с</strong> <strong>с</strong>емью п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 мм (ри<strong>с</strong>. 131).<br />

Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

ряд Protanisoptera<br />

(Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1931). 136. Pholidoptllon camense G. Zalessky;<br />

<strong>к</strong>рыл<strong>о</strong>, x 1,7, <strong>в</strong>. пермь, Приур<strong>а</strong>лье (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1931). 137.<br />

Polytaxineura stanleyl Tillyard; <strong>к</strong>рыл<strong>о</strong>, х 1,2, <strong>в</strong>. Пермь. А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

(Tillyard, 1935). 138. Calllmokattanla martynovl G. Zalessky;<br />

<strong>к</strong>рыл<strong>о</strong>, x 2.4, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1955). 139. Permaeschna dolloi Martynov; <strong>к</strong>рыл<strong>о</strong>, х 1,3, <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931)<br />

СЕМЕЙСТВО DITAXINEURIDAE TILLYARD, 1925<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>гнут; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> немн<strong>о</strong>гими п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>в</strong>ели<strong>к</strong><strong>а</strong>; пять <strong>в</strong>ет<strong>в</strong>ей RS и<br />

пять-ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей М; меди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле не<br />

б<strong>о</strong>лее чем <strong>с</strong> ше<strong>с</strong>тью жил<strong>к</strong><strong>а</strong>ми. Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>,<br />

Ditaxineurella Martynov, 1940 (Hetntzygopteron<br />

G. Zalessky, 1955, syn. nov.).<br />

Тип р<strong>о</strong>д<strong>а</strong> — D. stigmalis Martynov, 1940; н.<br />

пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC<br />

<strong>с</strong> ше<strong>с</strong>тью п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>о</strong>чень <strong>в</strong>ели<strong>к</strong><strong>а</strong>; между <strong>в</strong>ет<strong>в</strong>ями RS не менее четырех<br />

п<strong>о</strong>перечных <strong>в</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м интер<strong>в</strong><strong>а</strong>ле. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 37 мм (ри<strong>с</strong>. 132 и 133). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Ditaxineura Tillyard, 1925. Тип р<strong>о</strong>д<strong>а</strong> —<br />

D. anomalostigma Tillyard, 1925; н. пермь, С.<br />

75<br />

http://jurassic.ru/

Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> SC <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й или <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е без п<strong>о</strong>перечные<br />

жил<strong>о</strong><strong>к</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> умеренн<strong>о</strong>й <strong>в</strong>еличины;<br />

между <strong>в</strong>ет<strong>в</strong>ями RS д<strong>в</strong>е-три п<strong>о</strong>перечные жил<strong>к</strong>и<br />

<strong>в</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м интер<strong>в</strong><strong>а</strong>ле. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 25 мм<br />

(ри<strong>с</strong>. 134). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО POLYTAXINEURIDAE TILLYARD, 1935<br />

Передний <strong>к</strong>р<strong>а</strong>й рез<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>гнут; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> умеренн<strong>о</strong>й <strong>в</strong>еличины; меди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> 10—13 жил<strong>к</strong><strong>а</strong>ми, <strong>в</strong>ых<strong>о</strong>дящими<br />

н<strong>а</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Pholidoptilon G. Zalessky, 1931. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ph. kamense G. Zalessky, 1931; <strong>в</strong>.<br />

пермь. Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие<br />

Г<strong>о</strong>ры). Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> тремя <strong>в</strong>ыем<strong>к</strong><strong>а</strong>ми;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

лишен<strong>о</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

60 мм (ри<strong>с</strong>. 135 и 136). Один <strong>в</strong>ид. В. пермь<br />

Приур<strong>а</strong>лья.<br />

Polytaxineura Tillyard, 1935. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. Stanleyi Tillyard, 1935; <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> един<strong>с</strong>т<strong>в</strong>енным<br />

изл<strong>о</strong>м<strong>о</strong>м н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и шир<strong>о</strong><strong>к</strong><strong>а</strong>я; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong><br />

мн<strong>о</strong>гими п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

50 мм (ри<strong>с</strong>. 137). Один <strong>в</strong>ид. В. пермь А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО CALLIMOKALTANIIDAE G. ZALESSKY,<br />

1955<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong><strong>о</strong>гнутый; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми.<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупн<strong>а</strong>я, <strong>к</strong>ругл<strong>а</strong>я. Меди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью жил<strong>к</strong><strong>а</strong>ми, <strong>в</strong>ых<strong>о</strong>дящими<br />

н<strong>а</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. Пермь. Один р<strong>о</strong>д.<br />

Callimokaltania G. Zalessky, 1955. Тип<br />

р<strong>о</strong>д<strong>а</strong> — С. martynovi G. Zalessky, 1955; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Узел<strong>к</strong><strong>о</strong><strong>в</strong>ые и п<strong>о</strong>дузел<strong>к</strong><strong>о</strong><strong>в</strong>ые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены; длинн<strong>а</strong>я з<strong>а</strong>дняя<br />

<strong>в</strong>ет<strong>в</strong>ь МА; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> д<strong>в</strong>умя ряд<strong>а</strong>ми ячее<strong>к</strong>.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 28 мм (ри<strong>с</strong>. 138). Один <strong>в</strong>ид.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.'<br />

СЕМЕЙСТВО PERMAESCHNIDAE MARTYNOV, 1931<br />

Передний <strong>к</strong>р<strong>а</strong>й рез<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>гнут; <strong>в</strong>ершинный<br />

<strong>о</strong>тдел <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong> МА; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я; меди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> <strong>о</strong>чень мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

жил<strong>к</strong><strong>а</strong>ми; н<strong>а</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ых<strong>о</strong>дит не<br />

менее 20 жил<strong>о</strong><strong>к</strong>. Пермь. Один р<strong>о</strong>д.<br />

Permaeschna Martynov, 1931. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. dolloi Martynov, 1931; <strong>в</strong>. пермь, Арх<strong>а</strong>н­<br />

гель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). П<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>в</strong> б<strong>о</strong>льш<strong>о</strong>м чи<strong>с</strong>ле <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех п<strong>о</strong>лях<br />

<strong>к</strong>рыл<strong>а</strong>; н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>ры 1<br />

-<br />

л<strong>а</strong> 13—15 ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 55—<br />

60 мм (ри<strong>с</strong>. 139). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

ПОДОТРЯД ZYGOPTERA.<br />

РАВНОКРЫЛЫЕ СТРЕКОЗЫ<br />

(PROTOZYQOPTERA)<br />

Крылья рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженные <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>в</strong> <strong>в</strong>иде <strong>с</strong>тебель<strong>к</strong><strong>а</strong>; <strong>о</strong>бе их п<strong>а</strong>ры п<strong>о</strong>чти <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

ф<strong>о</strong>рмы и р<strong>а</strong>змер<strong>о</strong><strong>в</strong>, з<strong>а</strong>дние п<strong>о</strong>чти не р<strong>а</strong><strong>с</strong>­<br />

ширены 1<br />

; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> меди­<br />

<strong>а</strong>льн<strong>о</strong>й, ч<strong>а</strong>ще <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не р<strong>а</strong>зделяя<strong>с</strong>ь<br />

н<strong>а</strong> треуг<strong>о</strong>льни<strong>к</strong>и. Тел<strong>о</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>е; брюш<strong>к</strong><strong>о</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>е;<br />

гл<strong>а</strong>з<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные.<br />

Личин<strong>к</strong>и н<strong>а</strong> <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong> не<strong>с</strong>ут три <strong>к</strong>рупные<br />

ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идные ж<strong>а</strong>бры. Пермь — ныне. Семь<br />

н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Kennedyidea, Permagrionidea,<br />

Permepallagidea, Protomyrmeleontidea, Hemiphlebiidea,<br />

Coenagrionidea, Agrionidea.<br />

76<br />

http://jurassic.ru/<br />

НАДСЕМЕЙСТВО KENNEDYIDEA<br />

Узел<strong>о</strong><strong>к</strong> не <strong>в</strong>ыр<strong>а</strong>жен; узел<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я и п<strong>о</strong>дузел<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я<br />

жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong><br />

ред<strong>к</strong>ими п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>й.<br />

Пермь. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО KENNEDYIDAE TILLYARD, 1923<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й, з<strong>а</strong>дний <strong>в</strong>ыпу<strong>к</strong>лый.<br />

В<strong>с</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>лях прямые; <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льных<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> не б<strong>о</strong>лее д<strong>в</strong>ух, п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льньгх<br />

три — ше<strong>с</strong>ть; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>-<strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>с</strong> <strong>о</strong>дним ряд<strong>о</strong>м ячее<strong>к</strong>. Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Kennedya Tillyard, 1925. Тип р<strong>о</strong>д<strong>а</strong>—<br />

К. mirabilis Tillyard, 1925; н. пермь, С. Амери<strong>к</strong><strong>а</strong><br />

(К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не<strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, неб<strong>о</strong>льш<strong>а</strong>я;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле р<strong>а</strong><strong>в</strong>н<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му<br />

или незн<strong>а</strong>чительн<strong>о</strong> шире нег<strong>о</strong>; п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> четыре-пять; между <strong>в</strong>ет<strong>в</strong>ями<br />

RSs и МА <strong>с</strong>емь-<strong>в</strong><strong>о</strong><strong>с</strong>емь п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>,<br />

между МА и CuP их <strong>в</strong><strong>о</strong><strong>с</strong>емь-де<strong>в</strong>ять; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> 12—18 п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 34—45 мм (ри<strong>с</strong>. 140). Пермь.<br />

Четыре <strong>в</strong>ид<strong>а</strong>. Н. пермь С. Амери<strong>к</strong>и.<br />

Progoneura Carpenter, 1931. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. minuta Carpenter, 1931; н. пермь,<br />

С. Амери<strong>к</strong><strong>а</strong>. Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, умеренн<strong>о</strong><br />

удлиненн<strong>а</strong>я; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле зн<strong>а</strong>чительн<strong>о</strong> шире<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных

Ри<strong>с</strong>. 140—148. П<strong>о</strong>д<strong>о</strong>тряд Zygoptera, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>!<br />

140. Kennedya mirabilis Tillyard; <strong>к</strong>рыл<strong>о</strong>, x 1,4, н. пермь, С. Амери<strong>к</strong><strong>а</strong><br />

(Tillyard, 1928). 141. Progoneura nobills Carpenter; <strong>к</strong>рыл<strong>о</strong>,<br />

x 4,6, и. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1947). 142. sPermagrlon<br />

falklandicum Tillyard; <strong>к</strong>рыл<strong>о</strong>, x2,3, <strong>в</strong>. пермь, Ф<strong>а</strong>л<strong>к</strong>л<strong>а</strong>нд<strong>с</strong><strong>к</strong>ие <strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>в</strong><strong>а</strong><br />

(Tillyard, 1928). 143. Permolestes gracilis Martynov; <strong>к</strong>рыл<strong>о</strong>,<br />

x 2, 0,<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 144. Sushkinia<br />

parvula Martynov; <strong>к</strong>рыл<strong>о</strong>, x 4,7, <strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>р-<br />

жил<strong>о</strong><strong>к</strong> три-четыре; между RS5 и МА — три<br />

п<strong>о</strong>перечные жил<strong>к</strong>и, между МА и CuP — д<strong>в</strong>е.<br />

Кубит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> д<strong>в</strong>умя-тремя п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 13—17 мм<br />

(ри<strong>с</strong>. 141). Один <strong>в</strong>ид из <strong>в</strong>. перми Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл., д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из н. перми С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО PERMAGRIONIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> д<strong>в</strong>умя-тремя <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льными<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; узел<strong>о</strong><strong>к</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чный;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> без изл<strong>о</strong>м<strong>а</strong>;<br />

узел<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я и п<strong>о</strong>дузел<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я п<strong>о</strong>перечные жил<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют пр<strong>о</strong>д<strong>о</strong>лжение<br />

<strong>о</strong>дн<strong>а</strong> друг<strong>о</strong>й; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>тделен<strong>а</strong><br />

<strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>й. Пермь. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Регmagrionidae,<br />

Permolestidae, Solikamptilonidae.<br />

СЕМЕЙСТВО PERMAGRIONIDAE TILLYARD, 1928<br />

В<strong>о</strong><strong>с</strong>емь п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>,<br />

т<strong>о</strong>чн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющих <strong>с</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле; <strong>о</strong><strong>с</strong>-<br />

77<br />

http://jurassic.ru/<br />

! Kennedyidea, Permagrionidea, Permepallagidea<br />

тын<strong>о</strong><strong>в</strong>, 1930). 145. Scytolestes stigmalis Martynov; <strong>к</strong>рыл<strong>о</strong>, x 3,0,<br />

<strong>в</strong>. пермь, Ю. Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 146. Epilestes kargalensis<br />

Martynov, <strong>к</strong>рыл<strong>о</strong>, x 2,4, <strong>в</strong>. пермь, Ю. Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1937). 147. Solikamptilon remullforme Q. Zalessky; <strong>к</strong>рыл<strong>о</strong>,<br />

X 1,7, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1948). 148. Permepallage<br />

angustisslma Martynov; <strong>к</strong>рыл<strong>о</strong>, х 1,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938)<br />

н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел Си <strong>в</strong> <strong>с</strong>тебель<strong>к</strong>е <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> я<strong>с</strong>н<strong>о</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> <strong>о</strong>дним ряд<strong>о</strong>м ячее<strong>к</strong>. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 34 мм (ри<strong>с</strong>. 142). Пермь. Один р<strong>о</strong>д из<br />

<strong>в</strong>. перми 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PERMOLESTIDAE MARTYNOV, 1931<br />

Пять-<strong>с</strong>емь п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>,<br />

не р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел Си<br />

<strong>о</strong>бычн<strong>о</strong> я<strong>с</strong>ен; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> д<strong>в</strong>умя или тремя<br />

ряд<strong>а</strong>ми мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных мел<strong>к</strong>их ячее<strong>к</strong>. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 25—50 мм. Пермь. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Permolestes Martynov, 1931. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. gracilis Martynov, 1931; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля <strong>в</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>й ширины; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>я;<br />

п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные жил<strong>к</strong>и <strong>к</strong>реп<strong>к</strong>ие, чет<strong>к</strong>ие; МА и<br />

CuP не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>я<strong>с</strong>ь <strong>в</strong><br />

т<strong>о</strong>н<strong>к</strong>ие п<strong>о</strong><strong>о</strong>межут<strong>о</strong>чные <strong>в</strong>ет<strong>в</strong>и между мн<strong>о</strong>г<strong>о</strong>чи-

<strong>с</strong>ленными мел<strong>к</strong>ими ячей<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

48 мм (ри<strong>с</strong>. 143). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Sushkinia Martynov, 1930. Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. parvula Martynov, 1930; <strong>о</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> длинн<strong>а</strong>я и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я;<br />

п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong>зличн<strong>о</strong>й<br />

т<strong>о</strong>лщины 1<br />

, <strong>к</strong>реп<strong>к</strong>ие и <strong>с</strong>л<strong>а</strong>бые; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле з<strong>а</strong>метн<strong>о</strong> шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. МА и<br />

CuP д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>к</strong>р<strong>а</strong>я. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 28—33 мм<br />

(ри<strong>с</strong>. 144). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Приур<strong>а</strong>лья.<br />

Scytolestes Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. stigmalis Martynov, 1937; <strong>в</strong>. пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, н<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я;<br />

п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные жил<strong>к</strong>и чет<strong>к</strong>ие; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле з<strong>а</strong>метн<strong>о</strong> шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; МА<br />

и CuP д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>к</strong>р<strong>а</strong>я. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 25 мм<br />

(ри<strong>с</strong>. 145). Один <strong>в</strong>ид. В. пермь Ю. Приур<strong>а</strong>лья.<br />

Epilestes Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong><br />

— Е. kargalensis Martynov, 1937; <strong>в</strong>. пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й<br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> длинн<strong>а</strong>я;<br />

п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>бые;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й ширины <strong>с</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным; МА и CuP д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>к</strong>р<strong>а</strong>я. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 31 мм (ри<strong>с</strong>. 146). Один <strong>в</strong>ид. В.<br />

пермь Ю. Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО SOLIKAMPT1LON1DAE G. ZALESSKY,<br />

1948<br />

Крыл<strong>о</strong> <strong>к</strong> <strong>в</strong>ершине ут<strong>о</strong>няет<strong>с</strong>я; д<strong>в</strong>е п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные<br />

п<strong>о</strong>перечные жил<strong>к</strong>и п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют<br />

<strong>с</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енными п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми <strong>в</strong><br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>к</strong>реп<strong>к</strong><strong>а</strong>я,<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> <strong>о</strong>дним<br />

ряд<strong>о</strong>м ячее<strong>к</strong>. Пермь. Один р<strong>о</strong>д.<br />

Solikamptilon G. Zalessky, 1948. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. remuliforme G. Zalessky, 1948; и.<br />

пермь, Ур<strong>а</strong>л (С<strong>о</strong>ли<strong>к</strong><strong>а</strong>м<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>о</strong>ли, Н. М<strong>о</strong>ше<strong>в</strong><strong>о</strong>).<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и RS, М, Си и<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л А п<strong>а</strong>р<strong>а</strong>ллельны; <strong>в</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле б<strong>о</strong>лее 25 <strong>в</strong>ет<strong>в</strong>ей. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

45 мм (ри<strong>с</strong>. 147). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

НАДСЕМЕЙСТВО PERMEPALLAGIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми — б<strong>о</strong>лее 15 <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льных<br />

и 25 п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных; узел<strong>о</strong><strong>к</strong> не <strong>в</strong>ыр<strong>а</strong>жен; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>й;<br />

<strong>в</strong> <strong>с</strong>тебель<strong>к</strong>е <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть Aj. Пермь. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PERMEPALLAGIDAE<br />

MARTYNOV, 1938<br />

Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е и длинн<strong>о</strong>е, <strong>с</strong> прямыми передним<br />

и з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ями; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong> <strong>к</strong>реп<strong>к</strong><strong>о</strong>й и длинн<strong>о</strong>й Ai; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>о</strong>чень мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех п<strong>о</strong>лях.<br />

Пермь. Один р<strong>о</strong>д.<br />

Permepallage М <strong>а</strong> г t у п <strong>о</strong>V, 1938. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. angustissima Martynov, 1938; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Пр<strong>о</strong>межут<strong>о</strong>чные д<strong>о</strong>п<strong>о</strong>лнительные пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и между <strong>в</strong>ет<strong>в</strong>ями RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

90 мм (ри<strong>с</strong>. 148). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

НАДСЕМЕЙСТВО<br />

PROTOMYRMELEONTIDEA<br />

(ARCHIZIGOPTERA)<br />

Узел<strong>о</strong><strong>к</strong> не <strong>в</strong>ыр<strong>а</strong>жен, <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны; SC <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>й;<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченных жил<strong>о</strong><strong>к</strong><br />

нет. Мез<strong>о</strong>з<strong>о</strong>й. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО PROTOMYRMELEONTIDAE<br />

HANDLIRSCH, 1906<br />

Крыл<strong>о</strong> <strong>с</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им <strong>с</strong>тебель<strong>к</strong><strong>о</strong>м; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

чет<strong>к</strong><strong>а</strong>я; р<strong>а</strong>ди<strong>а</strong>льные жил<strong>к</strong>и з<strong>а</strong>ним<strong>а</strong>ют<br />

б<strong>о</strong>льшую ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>; МА и CuP пр<strong>о</strong><strong>с</strong>тые;<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Мез<strong>о</strong>з<strong>о</strong>й. Три<br />

р<strong>о</strong>д<strong>а</strong>.<br />

Protomyrmeleon G е i n i t z, 1887. Тип р<strong>о</strong>д<strong>а</strong>—P.<br />

brunonis Geinitz, 1887; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(<strong>в</strong>. лей<strong>а</strong><strong>с</strong>). Четыре я<strong>с</strong>ные <strong>в</strong>ет<strong>в</strong>и RS; RS 4<br />

не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; между RS 2 и RS3 н<strong>а</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong>ых<strong>о</strong>дит б<strong>о</strong>лее де<strong>с</strong>ят<strong>к</strong><strong>а</strong> жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

22—24 мм (ри<strong>с</strong>. 149). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из <strong>в</strong>. юры<br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> (К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у) и <strong>о</strong>дин из н. юры 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Вне СССР: Triassagrion Tillyard, 1922 (<strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии) и Tillyardagrion Martynov,<br />

1927 (н. юр<strong>а</strong> Англии).<br />

НАДСЕМЕЙСТВО HEMIPHLEBIIDEA<br />

Узел<strong>о</strong><strong>к</strong> и ег<strong>о</strong> п<strong>о</strong>перечные жил<strong>к</strong>и н<strong>а</strong> <strong>к</strong>рыле<br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты; д<strong>в</strong>е чет<strong>к</strong>ие <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льные<br />

п<strong>о</strong>перечные жил<strong>к</strong>и; п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong> пять, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены' н<strong>а</strong> ин<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, чем<br />

п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>; р<strong>а</strong>змеры м<strong>а</strong>лы.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 10—12 мм (ри<strong>с</strong>. 150). Одн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hemiphlebiidae <strong>с</strong> <strong>о</strong>дним<br />

р<strong>о</strong>д<strong>о</strong>м.

Ри<strong>с</strong>. 149—158. П<strong>о</strong>д<strong>о</strong>тряд Zygoptera, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Protomyrmeleontidea, Hemiphlebiidea, jCoenagrionidea. Agrionidea<br />

149. Protomyrmeleon brunonis Geinitz; <strong>к</strong>рыл<strong>о</strong>, x 3,7, н. юр<strong>а</strong>, 3.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1906). 150. Hemiphlebla mlrabilist Selys;<br />

<strong>к</strong>рыл<strong>о</strong>, x 6,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Tillyard, 1938). 151. Argiolestes icteromelas<br />

Selys; <strong>к</strong>рыл<strong>о</strong>, x 2,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Tillyard, 1926). 152. Austrolestes<br />

cingulatus Burmeister; <strong>к</strong>рыл<strong>о</strong>, x 2,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Tillyard, |1926).<br />

153. Pseudagrion australasiae Selys; <strong>к</strong>рыл<strong>о</strong>, x 2,1, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Tillyard,<br />

1926). 154. Platycnemis pennipes Pall; <strong>к</strong>рыл<strong>о</strong>, x 3,1, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

НАДСЕМЕЙСТВО COENAGRIONIDEA<br />

Узел<strong>о</strong><strong>к</strong> и ег<strong>о</strong> п<strong>о</strong>перечные жил<strong>к</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>иты; д<strong>в</strong>е чет<strong>к</strong>ие <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные<br />

жил<strong>к</strong>и; п<strong>о</strong><strong>с</strong>тн<strong>о</strong>д<strong>а</strong>льные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

т<strong>о</strong>чн<strong>о</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енных п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>й. Не<strong>о</strong>ген<br />

— ныне. Де<strong>в</strong>ять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> .<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; из них четыре н<strong>а</strong>йдены 1<br />

и <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>е­<br />

м<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии: Megapodagrionidae, Lestidae,<br />

Coenagrionidae, Platycnemididae.<br />

СЕМЕЙСТВО MEGAPODAGRIONIDAE<br />

TILLYARD, 1917<br />

Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные д<strong>о</strong>п<strong>о</strong>лнительные пр<strong>о</strong>межут<strong>о</strong>чные<br />

<strong>в</strong>ет<strong>в</strong>и RS и М, <strong>в</strong>ых<strong>о</strong>дящие н<strong>а</strong> <strong>к</strong>р<strong>а</strong>й;<br />

лишь <strong>о</strong>дн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>ль-<br />

(Munz, 1919), 155. Steleopteron deichmuelleri Handlirsch; <strong>к</strong>рыл<strong>о</strong><br />

x 1,9, <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925). 156. Phenacolestes<br />

mirandus Cockerell; <strong>к</strong>рыл<strong>о</strong>, x 2,5, ми<strong>о</strong>цен, С. Амери<strong>к</strong><strong>а</strong> (Handlirsch,<br />

1925). 157. Agrion virgo Linnaeus; <strong>к</strong>рыл<strong>о</strong>, x 2,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем-<br />

(Munz, 1919). 158. Epaltage fatima Charpenter;J <strong>к</strong>рыл<strong>о</strong>, x 2,0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Munz, 1919)<br />

79<br />

нее ур<strong>о</strong><strong>в</strong>ня <strong>с</strong>убн<strong>о</strong>д<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенным ди<strong>с</strong>т<strong>а</strong>льным<br />

<strong>к</strong><strong>о</strong>нц<strong>о</strong>м (ри<strong>с</strong>. 151). Не<strong>о</strong>ген — ныне.<br />

13 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, три р<strong>о</strong>д<strong>а</strong> из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и и<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО LESTIDAE CALVERT, 1901.<br />

' ЛЮТКИ<br />

Д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>тх<strong>о</strong>дят <strong>о</strong>т гл<strong>а</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong> <strong>о</strong>т ур<strong>о</strong><strong>в</strong>ня <strong>с</strong>убн<strong>о</strong>д<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и; д<strong>о</strong>п<strong>о</strong>лнительные пр<strong>о</strong>межут<strong>о</strong>чные<br />

<strong>в</strong>ет<strong>в</strong>и RS е<strong>с</strong>ть; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>с</strong><br />

уз<strong>к</strong>им ди<strong>с</strong>т<strong>а</strong>льным <strong>к</strong><strong>о</strong>нц<strong>о</strong>м (ри<strong>с</strong>. 152). Не<strong>о</strong>ген<br />

— ныне. Де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне^<br />

из них <strong>о</strong>дин (testes Leach) <strong>в</strong> не<strong>о</strong>гене;<br />

<strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong> <strong>в</strong> не<strong>о</strong>гене<br />

С. Амери<strong>к</strong>и.<br />

http://jurassic.ru/

СЕМЕЙСТВО COENAGR10N1DAE TILLYARD, 1918<br />

Д<strong>о</strong>п<strong>о</strong>лнительные <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>о</strong><strong>с</strong>трым ди<strong>с</strong>т<strong>а</strong>льным<br />

<strong>к</strong><strong>о</strong>нц<strong>о</strong>м; RS 3, МА и Ai н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong><strong>в</strong> <strong>в</strong>иде рез<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женных зигз<strong>а</strong>г<strong>о</strong><strong>в</strong>; CuP и Ai е<strong>с</strong>ть<br />

(ри<strong>с</strong>. 153). Не<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>в</strong> ми<strong>о</strong>цене<br />

С. Амери<strong>к</strong>и и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых три <strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> не<strong>о</strong>гене.<br />

СЕМЕЙСТВО PLATYCNEMID1DAE JACOBSON ЕТ<br />

BIANCHI, 1905<br />

Д<strong>о</strong>п<strong>о</strong>лнительные <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

жил<strong>к</strong>и CuP и А] имеют<strong>с</strong>я; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>с</strong> тупым ди<strong>с</strong>т<strong>а</strong>льным <strong>к</strong><strong>о</strong>нц<strong>о</strong>м; <strong>в</strong> <strong>в</strong>иде<br />

зигз<strong>а</strong>г<strong>а</strong> из<strong>о</strong>гнуты лишь <strong>к</strong><strong>о</strong>нцы жил<strong>о</strong><strong>к</strong> МА,<br />

CuP и Ai (ри<strong>с</strong>. 154). Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, д<strong>в</strong><strong>а</strong> из них<br />

<strong>в</strong> ми<strong>о</strong>цене С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО AGRIONIDEA<br />

Крылья без <strong>с</strong>тебель<strong>к</strong><strong>а</strong>, реже <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

<strong>с</strong>тебель<strong>к</strong><strong>о</strong>м; узел<strong>о</strong><strong>к</strong> и ег<strong>о</strong> п<strong>о</strong>перечные жил<strong>к</strong>и<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты 1<br />

; пять и б<strong>о</strong>лее <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льных<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Юр<strong>а</strong> —ныне. Семь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>,<br />

из них Zacallitidae, Amphipterygidae,<br />

Agrionidae, Epallagidae н<strong>а</strong>йдены <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО ZACALLITIDAE FRASER, 1943<br />

Антен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> де<strong>с</strong>ять;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>я. Один р<strong>о</strong>д<br />

из э<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО AMPHIPTERYGIDAE TILLYARD, 1917<br />

Крыл<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>тебель<strong>к</strong><strong>о</strong>м; <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> де<strong>в</strong>ять; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л RSi <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии не з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я и не <strong>с</strong>ближен <strong>с</strong> R;<br />

Ри<strong>с</strong>. 159—164. П<strong>о</strong>д<strong>о</strong>тряд Anisozygcptera, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Heterophlebiidea<br />

159. Eplophlebla superstes^MacLachlan; <strong>о</strong>бщий <strong>в</strong>ид, х 1,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 163. Llassophlebia magnified Tillyard;<br />

(Handlirsch, 1925). 160. ProgonopMebia woodwardi Tillyard; <strong>к</strong>ры- <strong>к</strong>рыл<strong>о</strong>, x 0,«5, н. юр<strong>а</strong>. 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Tillvard, 1925). 164. Heteroл<strong>о</strong><br />

x 2,0, н юр<strong>а</strong>, .3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Tillyard, 1925). 161. Selenothemls phlebia dlslocata Brodie et Westwood; <strong>к</strong>рылья, x 0,68, и. юр<strong>а</strong>,<br />

<strong>а</strong> п <strong>а</strong> 1 1 Г<br />

.<strong>о</strong><strong>о</strong>п» ,? <strong>с</strong> 0 <strong>с</strong> ^ Ь <strong>к</strong> р ы л <strong>о</strong><br />

<strong>к</strong> * х 2<br />

-, 8<br />

- "• юр<strong>а</strong>. 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925)<br />

1925).: 162. Sogdothemis modesta Martynov; <strong>к</strong>рыл<strong>о</strong>, x 4,4, н. юр<strong>а</strong>.<br />

80<br />

http://jurassic.ru/<br />

160<br />

164

п<strong>о</strong>перечные жил<strong>к</strong>и н<strong>а</strong> <strong>к</strong>рыле немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> те<strong>с</strong>н<strong>о</strong>. Мез<strong>о</strong>з<strong>о</strong>й<br />

— ныне. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

и пять <strong>в</strong>ымерших р<strong>о</strong>д<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 155 и 156):<br />

Steleopteron Handlirsch, 1908 из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и четыре р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не'<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО AGRIONIDAE K1RBY, 1890<br />

Крыл<strong>о</strong> без <strong>с</strong>тебель<strong>к</strong><strong>а</strong>; <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> б<strong>о</strong>льше 12; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л<br />

RS ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ближен <strong>с</strong> R; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

<strong>о</strong>чень мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны 1<br />

, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

(ри<strong>с</strong>. 157). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО EPALLAGIDAE NEEDHAM, 1903<br />

Крыл<strong>о</strong> без <strong>с</strong>тебель<strong>к</strong><strong>а</strong>; <strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> 11 и б<strong>о</strong>лее; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong>ближен <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> R; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> без <strong>в</strong><strong>о</strong>з<strong>в</strong>р<strong>а</strong>тн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны (ри<strong>с</strong>. 158). В. юр<strong>а</strong> — ныне.<br />

В<strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

д<strong>в</strong><strong>а</strong> из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

ПОДОТРЯД ANISOZYGOPTERA.<br />

АНИЗОЗИГОПТЕРЫ<br />

Крылья без <strong>с</strong>тебель<strong>к</strong><strong>а</strong>, умеренн<strong>о</strong> <strong>с</strong>уженные<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>о</strong>бе п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong> п<strong>о</strong>чти <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

ф<strong>о</strong>рмы; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong> переднем<br />

<strong>к</strong>рыле не р<strong>а</strong>зделен<strong>а</strong> н<strong>а</strong> треуг<strong>о</strong>льни<strong>к</strong>и; <strong>в</strong><br />

з<strong>а</strong>днем <strong>к</strong>рыле ф<strong>о</strong>рм<strong>а</strong> эт<strong>о</strong>й ячей<strong>к</strong>и ин<strong>а</strong>я. Тел<strong>о</strong><br />

т<strong>о</strong>н<strong>к</strong><strong>о</strong>е; брюш<strong>к</strong><strong>о</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>е, <strong>к</strong> <strong>к</strong><strong>о</strong>нцу<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong>здут<strong>о</strong>е. Гл<strong>а</strong>з<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены. Юр<strong>а</strong> — ныне. Д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Heterophlebiidea и Tarsophlebiidea.<br />

НАДСЕМЕЙСТВО HETEROPHLEBIIDEA<br />

Ф<strong>о</strong>рм<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> и переднег<strong>о</strong> <strong>к</strong>рылье<strong>в</strong> з<strong>а</strong>метн<strong>о</strong><br />

р<strong>а</strong>зн<strong>а</strong>я; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> шире ячей<strong>к</strong>и переднег<strong>о</strong>; ее ди<strong>с</strong>т<strong>а</strong>льный<br />

передний уг<strong>о</strong>л, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong><strong>с</strong>трый. Юр<strong>а</strong> —<br />

ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Progonophlebiidae, Archithemistidae,<br />

Liassophlebiidae, Heterophlebiidae,<br />

Epiophlebiidae (ри<strong>с</strong>. 159); п<strong>о</strong><strong>с</strong>леднее т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PROGONOPHLEBIIDAE TILLYARD, 1925<br />

Субн<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я и н<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечные<br />

жил<strong>к</strong>и не р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>о</strong>дну линию; <strong>в</strong> <strong>о</strong>б<strong>о</strong>их<br />

<strong>к</strong>рыльях ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

6 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 81<br />

http://jurassic.ru/<br />

нер<strong>а</strong>зделенн<strong>а</strong>я; А не р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, и между ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong><strong>о</strong>й и з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong><br />

б<strong>о</strong>льш<strong>о</strong>е п<strong>о</strong>ле. Длин<strong>а</strong> 35—440 мм (ри<strong>с</strong>. 160).<br />

Н. юр<strong>а</strong>. Один р<strong>о</strong>д из лей<strong>а</strong><strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ARCHITHEMISTIDAE TILLYARD, 1917<br />

(Selenothemidae Handlirsch, 1939)<br />

Антен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> мн<strong>о</strong>г<strong>о</strong>;<br />

<strong>с</strong>убн<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я и н<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечные жил<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>о</strong>дну линию н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>ры-'<br />

л<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях нер<strong>а</strong>зделенн<strong>а</strong>я;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> е<strong>с</strong>ть (ри<strong>с</strong>.161).<br />

Юр<strong>а</strong>. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Sogdothemis М <strong>а</strong> г t у п <strong>о</strong> v, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

3. modesta Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б). Крыл<strong>о</strong> удлиненн<strong>о</strong>е<br />

<strong>с</strong> м<strong>а</strong>л<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>: МА з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я д<strong>о</strong> ур<strong>о</strong><strong>в</strong>ня<br />

птер<strong>о</strong><strong>с</strong>тигмы'; между птер<strong>о</strong><strong>с</strong>тигм<strong>о</strong>й и н<strong>о</strong>д<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й не менее 20 п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 37 мм<br />

(ри<strong>с</strong>. 162). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: Diastatommites Handlirsch,<br />

1906; Archithemis Handlirsch, 1906; Selenothemis<br />

Handlirsch, 1920; Heterothemis Handlirsch,<br />

1906; Liadothemis Handlirsch, 1906; Oryctothemis<br />

Handlirsch, 1906.<br />

СЕМЕЙСТВО LIASSOPHLEBIIDAE TILLYARD, 1925<br />

Антен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>етри;<br />

<strong>с</strong>убн<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я и н<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечные<br />

жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>о</strong>дну линию; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях нер<strong>а</strong>зделенн<strong>а</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 60—70 мм (ри<strong>с</strong>. 163). Н.<br />

юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из лей<strong>а</strong><strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы 1<br />

.<br />

СЕМЕЙСТВО HETEROPHLEBIIDAE HANDLIRSCH,<br />

1906<br />

Антен<strong>о</strong>д<strong>а</strong>льных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> пять:<br />

д<strong>в</strong>е <strong>к</strong>реп<strong>к</strong>ие, три <strong>с</strong>л<strong>а</strong>бые; <strong>с</strong>убн<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я и н<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я<br />

п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong><br />

<strong>о</strong>дн<strong>о</strong>й линии; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> р<strong>а</strong>зделен<strong>а</strong> н<strong>а</strong> треуг<strong>о</strong>льни<strong>к</strong> и н<strong>а</strong>дтреуг<strong>о</strong>льни<strong>к</strong>.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 мм (ри<strong>с</strong>. 164).<br />

Н. юр<strong>а</strong>. Четыре р<strong>о</strong>д<strong>а</strong> из лей<strong>а</strong><strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО TARSOPHLEBIIDEA<br />

Передние и з<strong>а</strong>дние <strong>к</strong>рылья п<strong>о</strong>чти <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы<br />

п<strong>о</strong> ф<strong>о</strong>рме; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льные ячей<strong>к</strong>и <strong>с</strong>х<strong>о</strong>дные,<br />

<strong>с</strong> тупыми ди<strong>с</strong>т<strong>а</strong>льными угл<strong>а</strong>ми. Брюш<strong>к</strong><strong>о</strong> длинн<strong>о</strong>е<br />

и т<strong>о</strong>н<strong>к</strong><strong>о</strong>е. Юр<strong>а</strong> — п<strong>а</strong>ле<strong>о</strong>ген. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Tarsophlebiidae, Karatawiidae, Stenophlebiidae,<br />

Isophlebiidae, Sieblosiidae.

Ри<strong>с</strong>. 165—171. П<strong>о</strong>д<strong>о</strong>тряд Anisozygopfera, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tarsophlebiidea и Anisozygoptera incertae sedis<br />

165. Tarsophlebia eximia (Hagen); <strong>к</strong>рылья, x 2,1, ». юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1925). 166. Karatawia turantca Martynov; <strong>к</strong>рыл<strong>о</strong>,<br />

x 1,6, ». юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 167. Stenophtebia<br />

latreilll German <strong>к</strong>рылья, x 1,3, <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1925). 168. Antsophlebla helle Hagen; <strong>к</strong>рыл<strong>о</strong>, x 1,2, <strong>в</strong>. юр<strong>а</strong>,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlircsh, 1925). 169. SUblosia \iucunda Hagen,<br />

СЕМЕЙСТВО TARSOPHLEBIIDAE HANDLIRSCH, 1906<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> меди<strong>а</strong>льн<strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>зуя <strong>с</strong><br />

п<strong>о</strong><strong>с</strong>ледней х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>е из<strong>о</strong>гнут<strong>о</strong>е и з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>е<br />

п<strong>о</strong>ле, лишенн<strong>о</strong>е п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет, п<strong>о</strong>чти <strong>с</strong>ередины з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я и <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими<br />

ряд<strong>а</strong>ми ячее<strong>к</strong>. Тел<strong>о</strong> <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong><strong>о</strong>е. В. юр<strong>а</strong>.<br />

Один р<strong>о</strong>д.<br />

Tarsophlebia Hagen, 1866. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Heterophlebia eximia Hagen, 1862; <strong>в</strong>. юр<strong>а</strong>, Гер­<br />

82<br />

http://jurassic.ru/<br />

<strong>к</strong>рылья, x 1,7, <strong>о</strong>диг<strong>о</strong>цен, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925). 170. Somarura<br />

glgantea Brauer, Redtenbacher, Ganglbauer; прид<strong>а</strong>т<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> брюш<strong>к</strong><strong>а</strong> личин<strong>к</strong>и, х 6,8, н. юр<strong>а</strong>, В. Сибирь (<strong>о</strong>риг. ри<strong>с</strong>.).<br />

171. Samarura minor Brauer, Redtenbacher, Ganglbauer; прид<strong>а</strong>т<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> брюш<strong>к</strong><strong>а</strong> личин<strong>к</strong>и, х 9,5, н. юр<strong>а</strong>. Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я |^бл.<br />

(<strong>о</strong>риг. ри<strong>с</strong>.)<br />

м<strong>а</strong>ния (м<strong>а</strong>льм). Между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ными <strong>в</strong>ет<strong>в</strong>ями<br />

RS <strong>в</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> <strong>к</strong>рупные ячей<strong>к</strong>и. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 35—60 мм (ри<strong>с</strong>. 165). Юр<strong>а</strong>. Один <strong>в</strong>ид<br />

из н. юры 3. Е<strong>в</strong>р<strong>о</strong>пы', три <strong>в</strong>ид<strong>а</strong> из <strong>в</strong>. юры 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин <strong>в</strong>ид <strong>в</strong>. юры К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО KARATAWIIDAE MARTYNOV, 1925<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>й; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й А <strong>о</strong>чень

м<strong>а</strong>л<strong>о</strong> и уз<strong>к</strong><strong>о</strong>, з<strong>а</strong>ним<strong>а</strong>я менее чет<strong>в</strong>ерти длины<br />

<strong>к</strong>рыл<strong>а</strong>. Один р<strong>о</strong>д из <strong>в</strong>. юры.<br />

Karatawia Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong>—<br />

К. turanica Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). МА н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong> пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> т<strong>о</strong>н<strong>к</strong>ую зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зную<br />

жил<strong>к</strong>у и не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 48 мм (ри<strong>с</strong>. 166). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО STENOPHLEBIIDAE<br />

HANDLIRSCH, 1906<br />

Ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>,<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>трен<strong>а</strong> и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д, р<strong>а</strong>зделен<strong>а</strong> н<strong>а</strong><br />

<strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях д<strong>в</strong>умя п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле б<strong>о</strong>льш<strong>о</strong>е, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>е<br />

<strong>к</strong>реп<strong>к</strong><strong>о</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>зующей <strong>с</strong>ильный<br />

изгиб. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 30—80 мм (ри<strong>с</strong>. 167).<br />

Один р<strong>о</strong>д из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ISOPHLEBIIDAE HANDLIRSCH, 1906<br />

Крылья <strong>о</strong>чень уз<strong>к</strong>ие, <strong>с</strong> <strong>о</strong><strong>с</strong>трыми <strong>в</strong>ершин<strong>а</strong>ми;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>в</strong>иде тр<strong>а</strong>пеции,<br />

неиз<strong>о</strong>гнут<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>е,<br />

<strong>о</strong>гр<strong>а</strong>ниченн<strong>о</strong>е рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>о</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 70—100 мм (ри<strong>с</strong>. 168).<br />

Три р<strong>о</strong>д<strong>а</strong> из <strong>с</strong>р. и <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО SIEBLOSIIDAE HANDLIRSCH, 1906<br />

Крылья м<strong>а</strong>л<strong>о</strong> удлиненные, <strong>с</strong> тупыми <strong>в</strong>ершин<strong>а</strong>ми;<br />

<strong>а</strong>нтен<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные жил<strong>к</strong>и<br />

нея<strong>с</strong>ны; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>в</strong>иде <strong>к</strong>рупн<strong>о</strong>й<br />

тр<strong>а</strong>пеции. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм<br />

(ри<strong>с</strong>. 169). П<strong>а</strong>ле<strong>о</strong>ген. Один р<strong>о</strong>д из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ANISOZYGOPTERA INCERTAE<br />

SEDIS<br />

К эт<strong>о</strong>му п<strong>о</strong>д<strong>о</strong>тряду <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

р<strong>о</strong>ды, еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> изученные, <strong>о</strong>тн<strong>о</strong>шения<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых нея<strong>с</strong>ны'.<br />

Samarura Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — 5. gigantea<br />

Brauer, Redtenbacher, Ganglbauer, 1889; н.<br />

юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

У<strong>с</strong>ть-Б<strong>а</strong>лей). Личин<strong>к</strong><strong>а</strong>. З<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и <strong>к</strong>рылье<strong>в</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я третьег<strong>о</strong> тергит<strong>а</strong><br />

брюш<strong>к</strong><strong>а</strong>. Брюш<strong>к</strong><strong>о</strong> удлиненн<strong>о</strong>е, гиб<strong>к</strong><strong>о</strong>е; три<br />

ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идные ж<strong>а</strong>бры н<strong>а</strong> <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong>. Длин<strong>а</strong><br />

ж<strong>а</strong>берных ли<strong>с</strong>т<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong> 2,7—5,5 мм (ри<strong>с</strong>. 170 и<br />

171). Н. юр<strong>а</strong>. Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong> из Сибири.<br />

83<br />

http://jurassic.ru/<br />

Вне СССР: из три<strong>а</strong><strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы — Pirontetia<br />

Meunier, 1907; из н. юры 3. Е<strong>в</strong>р<strong>о</strong>пы —<br />

Campterophlebia Bode, 1907; Petrothemis Handlirsch,<br />

1906; Systellothemis Handlirsch, 1939;<br />

Rhabdothemis Handlirsch, 1939; Temnostigma<br />

Handlirsch, 1939; Pycnothemis Handlirsch, 1939;<br />

Anomothemis Handlirsch, 1906; Parelthothemis<br />

Handlirsch, 1906; Petrophlebia Tillyard, 1925; из<br />

три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии — Mesophlebia Tillyard, 1916;<br />

Periassophlebia Tillyard, 1918; Triassophlebia<br />

Tillyard, 1922; Triassolestes Tillyard, 1918.<br />

ПОДОТРЯД ANISOPTERA.<br />

НЕРАВНОКРЫЛЫЕ СТРЕКОЗЫ<br />

Крылья без <strong>с</strong>тебель<strong>к</strong><strong>а</strong>, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

з<strong>а</strong>дние <strong>к</strong>рылья зн<strong>а</strong>чительн<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т передних, б<strong>о</strong>лее шир<strong>о</strong><strong>к</strong>ие; н<strong>о</strong>д<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> н<strong>а</strong> <strong>с</strong>ередине переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> н<strong>а</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях<br />

пре<strong>в</strong>р<strong>а</strong>щен<strong>а</strong> <strong>в</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные треуг<strong>о</strong>льни<strong>к</strong>и.<br />

Тел<strong>о</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> м<strong>а</strong><strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>о</strong>е, брюш<strong>к</strong><strong>о</strong> нет<strong>о</strong>н<strong>к</strong><strong>о</strong>е,<br />

ин<strong>о</strong>гд<strong>а</strong> упл<strong>о</strong>щенн<strong>о</strong>е или р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е <strong>к</strong><br />

<strong>к</strong><strong>о</strong>нцу. Личин<strong>к</strong>и без ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идных ж<strong>а</strong>бер.<br />

Юр<strong>а</strong> — ныне. Д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Aeshnidea и<br />

Libellulidea.<br />

НАДСЕМЕЙСТВО AESHNIDEA<br />

Антен<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные жил<strong>к</strong>и <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>лях р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>, не н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; треуг<strong>о</strong>льни<strong>к</strong><br />

<strong>в</strong> з<strong>а</strong>днем <strong>к</strong>рыле п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ен т<strong>а</strong><strong>к</strong> же, <strong>к</strong><strong>а</strong><strong>к</strong><br />

<strong>в</strong> переднем, и уд<strong>а</strong>лен <strong>о</strong>т меди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля.<br />

Юр<strong>а</strong> — ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Liassogomphidae,<br />

Petaluridae, Gomphidae, Aeshnidae, Aeschnidiidae.<br />

СЕМЕЙСТВО LIASSOGOMPHIDAE TILLYARD, 1935<br />

Треуг<strong>о</strong>льни<strong>к</strong> р<strong>а</strong>зделен <strong>о</strong>дн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>й<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> р<strong>а</strong>зделен<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ями <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и н<strong>а</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличимые, р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я<br />

<strong>о</strong>трез<strong>к</strong>и — <strong>с</strong>е<strong>к</strong>т<strong>о</strong>ры. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

37 мм (ри<strong>с</strong>. 172). Юр<strong>а</strong>. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> из н. юры<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PETALURIDAE NEEDHAM, 1903<br />

Треуг<strong>о</strong>льни<strong>к</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>ий или<br />

шир<strong>о</strong><strong>к</strong>ий, п<strong>о</strong>др<strong>а</strong>зделенный непр<strong>а</strong><strong>в</strong>ильными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми н<strong>а</strong> д<strong>в</strong>е — ше<strong>с</strong>ть ячее<strong>к</strong>;<br />

треуг<strong>о</strong>льни<strong>к</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong><br />

и имеет <strong>о</strong>дну п<strong>о</strong>перечную жил<strong>к</strong>у;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> без <strong>о</strong>бр<strong>а</strong>зующих<br />

петли жил<strong>о</strong><strong>к</strong>; <strong>в</strong>ыд<strong>а</strong>ющий<strong>с</strong>я яйце<strong>к</strong>л<strong>а</strong>д (ри<strong>с</strong>. 173).<br />

В. юр<strong>а</strong> — ныне. Три р<strong>о</strong>д<strong>а</strong> из <strong>в</strong>. юры 1<br />

Е<strong>в</strong>р<strong>о</strong>пы<br />

б*

Ри<strong>с</strong>. 172—179. П<strong>о</strong>д<strong>о</strong>тряд Anisoptera<br />

172. Llassogomphus brodiei Buckland; <strong>к</strong>рыл<strong>о</strong>, x 1,5, н. юр<strong>а</strong>, 3.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Tillyard, 1925). 173. Protolindenia wittei Qiebel; A —<br />

переднее <strong>к</strong>рыл<strong>о</strong>, x 1,3; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,3; <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1925). 174. Lanthus parvulus Selys; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, x 0,8; Б —з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 0,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch,<br />

1906). 185. Ап<strong>а</strong>х Junius Day; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 1,5; Б —<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Comstock, 1918). 176. Aeschnldium<br />

84<br />

http://jurassic.ru/<br />

Г?? А<br />

densum Hagen; <strong>к</strong>рылья, x 1,7. <strong>в</strong>. юр<strong>а</strong>. 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1925). 177. Aeschnldiopsis flindersiensis Woodward; <strong>к</strong>рыл<strong>о</strong>,<br />

x 3,0, мел, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1917). 178. Aeschnldlella kaba.<br />

novt G. Zalessky; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, x 5,2, мел, П<strong>о</strong><strong>в</strong><strong>о</strong>лжье (Ю. З<strong>а</strong>ле<strong>с</strong>.<br />

<strong>с</strong><strong>к</strong>ий, 1955). 179. Scapanea sp.; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 2,0;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch, 1925)

и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>дин из<strong>в</strong>е<strong>с</strong>тен из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО GOMPHIDAE BRAUER, 1856<br />

Ан<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без петель. Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и шир<strong>о</strong><strong>к</strong><strong>а</strong>я; треуг<strong>о</strong>льни<strong>к</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>о</strong>нний, н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>рыле пр<strong>о</strong>д<strong>о</strong>льный,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>б<strong>а</strong> лишены п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>;<br />

яйце<strong>к</strong>л<strong>а</strong>д редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нный (ри<strong>с</strong>. 174).<br />

В. юр<strong>а</strong> — ныне. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых д<strong>в</strong><strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы 1<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и из <strong>в</strong>. <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО AESHNIDAE LEACH, 1815<br />

(Aeschnidae auctorum)<br />

Треуг<strong>о</strong>льни<strong>к</strong>и <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рылье<strong>в</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong> и р<strong>а</strong>зделены п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женными петлями. Гл<strong>а</strong>з<strong>а</strong> <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> темени. Яйце<strong>к</strong>л<strong>а</strong>д не р<strong>а</strong>з<strong>в</strong>ит (ри<strong>с</strong>. 175). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Три р<strong>о</strong>д<strong>а</strong> из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> В. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong><br />

и из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и; мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

пять из<strong>в</strong>е<strong>с</strong>тны из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> В. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong><br />

и ми<strong>о</strong>цен<strong>а</strong> К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong> и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО AESCHNIDIIDAE<br />

HANDLIRSCH, 1906<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> д<strong>в</strong><strong>о</strong>йны'м ряд<strong>о</strong>м<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> умеренн<strong>о</strong><br />

удлиненн<strong>а</strong>я; треуг<strong>о</strong>льни<strong>к</strong>и н<strong>а</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях<br />

рез<strong>к</strong><strong>о</strong> п<strong>о</strong>перечные и п<strong>о</strong>др<strong>а</strong>зделенные н<strong>а</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

ячей<strong>к</strong>и; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>днег<strong>о</strong><br />

• <strong>к</strong>рыл<strong>а</strong> без петле<strong>о</strong>бр<strong>а</strong>зных жил<strong>о</strong><strong>к</strong>, <strong>с</strong> ряд<strong>а</strong>ми<br />

ячее<strong>к</strong>; яйце<strong>к</strong>л<strong>а</strong>д <strong>с</strong>ильн<strong>о</strong> удлиненный (ри<strong>с</strong>. 176).<br />

Юр<strong>а</strong> — мел. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Aeschnidiella G. Zalessky, 1953. Тип<br />

р<strong>о</strong>д<strong>а</strong> —Л. kabanovi G. Zalessky, 1953; мел, П<strong>о</strong><strong>в</strong><strong>о</strong>лжье<br />

(<strong>а</strong>пт). Треуг<strong>о</strong>льни<strong>к</strong> <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти<br />

<strong>с</strong> ячей<strong>к</strong><strong>а</strong>ми, <strong>о</strong> уз<strong>к</strong>им, <strong>о</strong><strong>с</strong>трым з<strong>а</strong>дним <strong>к</strong><strong>о</strong>нц<strong>о</strong>м;<br />

<strong>а</strong>р<strong>к</strong>улю<strong>с</strong> неп<strong>о</strong>лный; п<strong>о</strong>дтреуг<strong>о</strong>льни<strong>к</strong> м<strong>а</strong>лень<strong>к</strong>ий,<br />

н<strong>о</strong> я<strong>с</strong>ный. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм<br />

(ри<strong>с</strong>. 178). Один <strong>в</strong>ид. Мел П<strong>о</strong><strong>в</strong><strong>о</strong>лжья.<br />

Вне СССР: из юры 3. Е<strong>в</strong>р<strong>о</strong>пы — Aeschnidium<br />

Westwood, 1854; Urogompus Handlirsch,<br />

1906; из мел<strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии — Aeschnidiopsis Tillyard,<br />

1917 (ри<strong>с</strong>. 177).<br />

НАДСЕМЕЙСТВО LIBELLULIDEA<br />

Антен<strong>о</strong>д<strong>а</strong>льные п<strong>о</strong>перечные жил<strong>к</strong>и <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>лях р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> <strong>о</strong>дн<strong>а</strong> пр<strong>о</strong>ти<strong>в</strong> друг<strong>о</strong>й; треуг<strong>о</strong>льни<strong>к</strong>и<br />

н<strong>а</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ены 1<br />

р<strong>а</strong>зличн<strong>о</strong>: н<strong>а</strong><br />

переднем треуг<strong>о</strong>льни<strong>к</strong> рез<strong>к</strong><strong>о</strong> п<strong>о</strong>перечный, н<strong>а</strong><br />

з<strong>а</strong>днем <strong>о</strong>н р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>о</strong>нний или пр<strong>о</strong>д<strong>о</strong>льный<br />

(ри<strong>с</strong>. 179). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне: Libellulidae и Corduliidae;<br />

<strong>о</strong>дин р<strong>о</strong>д п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> из<strong>в</strong>е<strong>с</strong>тен<br />

из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и; д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>- <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

эт<strong>о</strong>г<strong>о</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>, т<strong>о</strong>чн<strong>о</strong> не <strong>о</strong>пределенных,<br />

у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я из мн<strong>о</strong>гих ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

<strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> и ми<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

С. Амери<strong>к</strong>и.<br />

ANISOPTERA INCERTAE SEDIS<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> эт<strong>о</strong>му п<strong>о</strong>д<strong>о</strong>тряду прин<strong>а</strong>длеж<strong>а</strong>т<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые р<strong>о</strong>ды <strong>в</strong>ерхнеюр<strong>с</strong><strong>к</strong>их <strong>с</strong>тре<strong>к</strong><strong>о</strong>з из<br />

3. Е<strong>в</strong>р<strong>о</strong>пы, б<strong>о</strong>лее т<strong>о</strong>чные <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых нея<strong>с</strong>ны 1<br />

: Cymatophlebiopsis<br />

Handlirsch, 1939; Morbaeschna Needham^ 1907;<br />

Mesogomphus Handlirsch, 1939; Aescfinopsis<br />

Handlirsch, 1939.<br />

ИНФРАКЛАСС NEOPTERA. НОВОКРЫЛЫЕ<br />

Крылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong>д<strong>о</strong>ль тел<strong>а</strong><br />

н<strong>а</strong>з<strong>а</strong>д, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>я<strong>с</strong>ь пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong> <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й<br />

пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>с</strong>ти <strong>с</strong><strong>в</strong>ерху брюш<strong>к</strong><strong>а</strong> и п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>я ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

друг друг<strong>а</strong>. Лишь у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых групп н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых<br />

<strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я п<strong>а</strong>ле<strong>о</strong>птерия,<br />

т. е. утр<strong>а</strong>чи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ть <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ть<br />

<strong>к</strong>рылья и <strong>о</strong>т<strong>в</strong><strong>о</strong>дить их н<strong>а</strong>з<strong>а</strong>д. У б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>а</strong>я з<strong>а</strong>дняя,<br />

юг<strong>а</strong>льн<strong>а</strong>я, <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> жил<strong>о</strong><strong>к</strong> <strong>к</strong>рыл<strong>а</strong> (Ju). Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения. Инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>х<strong>о</strong>дит <strong>в</strong> р<strong>а</strong>зличных у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях,<br />

<strong>в</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> <strong>в</strong> н<strong>а</strong>земных<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

85<br />

http://jurassic.ru/<br />

<strong>с</strong>т<strong>а</strong>циях, реже <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х. В. де<strong>в</strong><strong>о</strong>н —• ныне.<br />

Четыре <strong>к</strong><strong>о</strong>г<strong>о</strong>рты: Archaeoneoptera, Polyneoptera,<br />

Paraneoptera, Holometabola (=01igoneoptera).<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения <strong>к</strong><strong>о</strong>г<strong>о</strong>рт еще<br />

нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тны; решение этих <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> з<strong>а</strong>труднен<strong>о</strong> м<strong>а</strong>лым зн<strong>а</strong>нием <strong>с</strong>тр<strong>о</strong>ения<br />

дре<strong>в</strong>нейших де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. В<strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение<br />

н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong><br />

<strong>с</strong> п<strong>о</strong>я<strong>в</strong>лением пер<strong>в</strong>ых <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые п<strong>о</strong>чти н<strong>а</strong><strong>в</strong>ерн<strong>о</strong>е прин<strong>а</strong>длеж<strong>а</strong>ли<br />

<strong>к</strong> эт<strong>о</strong>му инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>у Pterygota. В эт<strong>о</strong>м пр<strong>о</strong>-

це<strong>с</strong><strong>с</strong>е, н<strong>а</strong>ряду <strong>с</strong> <strong>о</strong>тчленением п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong><strong>в</strong>, пре<strong>в</strong>р<strong>а</strong>щением их <strong>в</strong> <strong>к</strong>рылья, б<strong>о</strong>льш<strong>о</strong>е<br />

зн<strong>а</strong>чение имел<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нение <strong>к</strong>рыльями пер<strong>в</strong>ичн<strong>о</strong>й<br />

пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>ти п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>лий, <strong>в</strong> д<strong>а</strong>льнейшем<br />

<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong><strong>в</strong>шей<strong>с</strong>я <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>й для р<strong>а</strong>з<strong>в</strong>ития п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ных<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений лет<strong>а</strong>тельными <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми. Т<strong>а</strong><strong>к</strong>им<br />

путем <strong>в</strong><strong>о</strong>зни<strong>к</strong>л<strong>а</strong> дре<strong>в</strong>нейш<strong>а</strong>я <strong>к</strong><strong>о</strong>г<strong>о</strong>рт<strong>а</strong> н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых<br />

Archaeoneoptera, п<strong>о</strong><strong>с</strong>лужи<strong>в</strong>ш<strong>а</strong>я и<strong>с</strong>т<strong>о</strong>чни<strong>к</strong><strong>о</strong>м<br />

р<strong>а</strong>з<strong>в</strong>ития <strong>к</strong><strong>о</strong>г<strong>о</strong>рты Polyneoptera н<strong>а</strong>ряду<br />

<strong>с</strong> инфр<strong>а</strong><strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong>м дре<strong>в</strong>не<strong>к</strong>рылых. Мн<strong>о</strong>г<strong>о</strong>жил<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылые (Polyneoptera), бы<strong>в</strong>шие<br />

и<strong>с</strong>х<strong>о</strong>дными для д<strong>в</strong>ух других <strong>к</strong><strong>о</strong>г<strong>о</strong>рт, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я<br />

примити<strong>в</strong>ным <strong>с</strong>тр<strong>о</strong>ением тел<strong>а</strong> ( <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы) и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием п<strong>о</strong><strong>к</strong><strong>о</strong>ящей<strong>с</strong>я<br />

Крыл<strong>о</strong> <strong>о</strong>чень пл<strong>о</strong>тн<strong>о</strong>е и бле<strong>с</strong>тящее: п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

<strong>с</strong><strong>о</strong> <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й <strong>в</strong> <strong>в</strong>иде непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong><strong>с</strong>ти и мел<strong>к</strong><strong>о</strong>й ямч<strong>а</strong>т<strong>о</strong><strong>с</strong>ти,<br />

н<strong>а</strong>иб<strong>о</strong>лее я<strong>с</strong>н<strong>о</strong>й <strong>в</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>. Ан<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть лишен<strong>а</strong> жил<strong>о</strong><strong>к</strong> и д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> уз<strong>к</strong><strong>а</strong>я. Ф<strong>о</strong>рм<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> треуг<strong>о</strong>льн<strong>а</strong>я <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным<br />

термин<strong>а</strong>льным <strong>к</strong>р<strong>а</strong>ем, лишенным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

и <strong>с</strong>н<strong>а</strong>бженным <strong>с</strong><strong>к</strong>л<strong>а</strong>дч<strong>а</strong>т<strong>о</strong><strong>с</strong>тью, <strong>в</strong>ых<strong>о</strong>дящей н<strong>а</strong><br />

<strong>к</strong>р<strong>а</strong>й и з<strong>а</strong>меняющей жил<strong>к</strong>и. Стр<strong>о</strong>ение тел<strong>а</strong><br />

неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>. В. де<strong>в</strong><strong>о</strong>н. Один <strong>о</strong>тряд Archaeoptera.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

из<strong>в</strong>е<strong>с</strong>тны. Б<strong>о</strong>льш<strong>а</strong>я пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>ть <strong>к</strong>рылье<strong>в</strong> и бедн<strong>о</strong><strong>с</strong>ть<br />

их жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют эту <strong>к</strong><strong>о</strong>г<strong>о</strong>рту<br />

<strong>о</strong>т других н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых. Н<strong>а</strong>иб<strong>о</strong>лее близ<strong>к</strong><strong>а</strong><br />

<strong>о</strong>н<strong>а</strong> <strong>к</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым Paraneoptera п<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>му р<strong>а</strong>з<strong>в</strong>итию<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я т<strong>а</strong><strong>к</strong>же<br />

КОГОРТА ARCHAEONEOPTERA<br />

ОТРЯД ARCHAEOPTERA<br />

Крыл<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ы'пу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем,<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> туп<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й, <strong>в</strong>ыпу<strong>к</strong>лым<br />

термин<strong>а</strong>льным и прямым <strong>а</strong>н<strong>а</strong>льным<br />

<strong>к</strong>р<strong>а</strong>ями, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong>тделены 1<br />

друг <strong>о</strong>т друг<strong>а</strong> т<strong>о</strong>р-<br />

ну<strong>с</strong><strong>о</strong>м. Имеют<strong>с</strong>я <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я С и пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

SC, <strong>о</strong>бр<strong>а</strong>зующие уз<strong>к</strong><strong>о</strong>е <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле.<br />

Н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> немн<strong>о</strong>гие пр<strong>о</strong><strong>с</strong>тые пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и. В з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>а</strong>я <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я<br />

дуг<strong>о</strong><strong>в</strong>идн<strong>а</strong>я Си, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бляющ<strong>а</strong>я уз<strong>к</strong>ую<br />

<strong>а</strong>н<strong>а</strong>льную <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть. Срединные жил<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>ие, <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лые <strong>с</strong>низу (ри<strong>с</strong>. 180).<br />

Де<strong>в</strong><strong>о</strong>н. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eopteridae.<br />

СЕМЕЙСТВО EOPTERIDAE ROHDENDORF, 1961<br />

Жил<strong>к</strong><strong>а</strong> С <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я н<strong>а</strong> переднем <strong>с</strong><strong>в</strong><strong>о</strong>ем <strong>к</strong>р<strong>а</strong>е<br />

и пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я н<strong>а</strong> з<strong>а</strong>днем, п<strong>о</strong><strong>к</strong><strong>а</strong>т<strong>а</strong>я <strong>к</strong> мембр<strong>а</strong>не. Н<strong>а</strong><br />

<strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые жил<strong>к</strong>и, R и М,<br />

идущие <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>дн<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

друг<strong>о</strong>й: R п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> <strong>с</strong>ближ<strong>а</strong>ет<strong>с</strong>я <strong>с</strong> пе­<br />

86<br />

http://jurassic.ru/<br />

<strong>с</strong>т<strong>а</strong>дии (<strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>и) <strong>в</strong> инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>м р<strong>а</strong>з<strong>в</strong>итии.<br />

К<strong>о</strong>г<strong>о</strong>рт<strong>а</strong> Paraneoptera <strong>в</strong><strong>о</strong>зни<strong>к</strong>л<strong>а</strong> <strong>в</strong> результ<strong>а</strong>те<br />

р<strong>а</strong>з<strong>в</strong>ития при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нных <strong>с</strong> пит<strong>а</strong>нием<br />

<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>ми р<strong>а</strong><strong>с</strong>тений, прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>к</strong><strong>о</strong>лющег<strong>о</strong><br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong>, <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>ни<strong>к</strong><strong>а</strong>ть <strong>в</strong><br />

т<strong>к</strong><strong>а</strong>ни жи<strong>в</strong>ых р<strong>а</strong><strong>с</strong>тений. П<strong>о</strong><strong>с</strong>ледняя <strong>к</strong><strong>о</strong>г<strong>о</strong>рт<strong>а</strong> —<br />

Holometabola, или Oligoneoptera,— <strong>в</strong><strong>о</strong>зни<strong>к</strong>л<strong>а</strong> <strong>в</strong><br />

результ<strong>а</strong>те <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ития, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е <strong>с</strong>т<strong>а</strong>л<strong>о</strong> пр<strong>о</strong>х<strong>о</strong>дить <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем<br />

иных у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях <strong>с</strong>реды, чт<strong>о</strong> <strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ил<strong>о</strong> <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение<br />

<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>й п<strong>о</strong><strong>к</strong><strong>о</strong>ящей<strong>с</strong>я <strong>с</strong>т<strong>а</strong>дии — <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>и;<br />

<strong>в</strong><strong>о</strong> <strong>в</strong>ремя <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й шл<strong>о</strong> ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>о</strong>рг<strong>а</strong>низм<strong>а</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>, рез<strong>к</strong><strong>о</strong> <strong>о</strong>тличн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>т м<strong>о</strong>л<strong>о</strong>дых <strong>с</strong>т<strong>а</strong>дий р<strong>а</strong>з<strong>в</strong>ития, личин<strong>о</strong><strong>к</strong>.<br />

<strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> т<strong>а</strong><strong>к</strong>ими мн<strong>о</strong>г<strong>о</strong>жил<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

(Polyneoptera), <strong>к</strong><strong>а</strong><strong>к</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ые<br />

(Blattodea). Одн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>г<strong>о</strong><br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ет <strong>а</strong>рхе<strong>о</strong>не<strong>о</strong>птер<br />

<strong>о</strong>т других н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых. Не<strong>с</strong><strong>о</strong>мненн<strong>о</strong> де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>ие<br />

Archaeoneoptera иллю<strong>с</strong>трируют <strong>о</strong>дин из пер<strong>в</strong>ичных<br />

эт<strong>а</strong>п<strong>о</strong><strong>в</strong> <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ения <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых,<br />

<strong>а</strong> именн<strong>о</strong> — п<strong>о</strong>я<strong>в</strong>ление <strong>о</strong>тчлененных<br />

<strong>к</strong>рылье<strong>в</strong> н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong><strong>в</strong><br />

<strong>с</strong>пин<strong>к</strong>и грудн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>. Т<strong>а</strong><strong>к</strong>ие пер<strong>в</strong>ичные<br />

<strong>к</strong>рылья при<strong>о</strong>брели <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ть <strong>с</strong><strong>о</strong><strong>в</strong>ерш<strong>а</strong>ть<br />

<strong>в</strong>зм<strong>а</strong>хи, <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>а</strong>ли при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленную для п<strong>о</strong>лет<strong>а</strong><br />

ф<strong>о</strong>рму; <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>ть этих <strong>к</strong>рылье<strong>в</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>л<strong>а</strong><strong>с</strong>ь еще б<strong>о</strong>льш<strong>о</strong>ю, и жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние еще не<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>с</strong>ь <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е <strong>о</strong>п<strong>о</strong>рн<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>к</strong>елет<strong>а</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и<br />

<strong>к</strong>рыл<strong>а</strong>.<br />

редним <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong> и перед ег<strong>о</strong> <strong>в</strong>ершин<strong>о</strong>й <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я<br />

<strong>с</strong> SC и С п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м нея<strong>с</strong>н<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и. М прям<strong>а</strong>я, <strong>с</strong>пер<strong>в</strong><strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

<strong>с</strong> R, п<strong>о</strong>т<strong>о</strong>м р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящ<strong>а</strong>я<strong>с</strong>я <strong>с</strong> нею и <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>я. Б<strong>о</strong>льш<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льнее Си и <strong>с</strong>з<strong>а</strong>ди М<br />

лишен<strong>а</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния. Г<strong>о</strong>фр п<strong>о</strong> термин<strong>а</strong>льн<strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю чет<strong>к</strong>ий. Один р<strong>о</strong>д. В. де<strong>в</strong><strong>о</strong>н.<br />

Eopterum Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. devonicum Rohdendorf, 1961; <strong>в</strong>. де<strong>в</strong><strong>о</strong>н,<br />

К<strong>о</strong>ми АССР (фр<strong>а</strong>н<strong>с</strong><strong>к</strong>ий — ф<strong>а</strong>мен<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>ы,<br />

В<strong>о</strong>й-В<strong>о</strong>ж). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле и <strong>с</strong>ередин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong>р<strong>а</strong>шенные, <strong>к</strong><strong>о</strong>ричне<strong>в</strong>ые, <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е <strong>к</strong>рыл<strong>о</strong><br />

желт<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>-<strong>к</strong><strong>о</strong>ричне<strong>в</strong><strong>о</strong>е; г<strong>о</strong>фр н<strong>а</strong> термин<strong>а</strong>льн<strong>о</strong>м<br />

<strong>к</strong>р<strong>а</strong>е <strong>о</strong>бр<strong>а</strong>зует 23 чет<strong>к</strong>ие <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и. Интер<strong>в</strong><strong>а</strong>л<br />

между R и М примерн<strong>о</strong> <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> шире ширины<br />

М. Жил<strong>к</strong><strong>а</strong> Си рез<strong>к</strong><strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющ<strong>а</strong>я, темн<strong>о</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>шенн<strong>а</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 11 мм, ширин<strong>а</strong> 5 мм<br />

(ри<strong>с</strong>. 180). Один <strong>в</strong>ид. В. де<strong>в</strong><strong>о</strong>н К<strong>о</strong>ми АССР.

Ри<strong>с</strong>. 180—182. Отряды Archaeopfera, Blattodea<br />

180. Eopterum devonlcum Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид, x 8,0, <strong>в</strong>. де<strong>в</strong><strong>о</strong>н, 180Б. Panestia spadicea Shlraki; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, x 1,3; <strong>с</strong><strong>о</strong><strong>в</strong>-<br />

C. Приур<strong>а</strong>лье (Р<strong>о</strong>денд<strong>о</strong>рф, 19616). 181А. Cryptocercus rellctus рем.; <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Panestiidae (Бей-Биен<strong>к</strong><strong>о</strong>, 1950). 182. Blabera<br />

Bey-Bienko; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х 3,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Panesti- atropos Stoll; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х 1,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Blatdae,<br />

<strong>с</strong><strong>в</strong>яз<strong>а</strong>н^ рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й третичн<strong>о</strong>й фл<strong>о</strong>р<strong>о</strong>й (Бей^Биен<strong>к</strong><strong>о</strong>, 1950); tldae (Brunner, 1865)<br />

http://jurassic.ru/

КОГОРТА POLYNEOPTERA. МНОГОЖИЛКОВЫЕ<br />

Оп<strong>о</strong>рный <strong>с</strong><strong>к</strong>елет пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных пр<strong>о</strong>д<strong>о</strong>льных и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

т<strong>о</strong>же мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

или х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>г<strong>о</strong> <strong>а</strong>рхеди<strong>к</strong>тия. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны <strong>в</strong><strong>с</strong>егд<strong>а</strong> грызущег<strong>о</strong> тип<strong>а</strong>. Имеет<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

юг<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>. Инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие без п<strong>о</strong><strong>к</strong><strong>о</strong>ящей<strong>с</strong>я <strong>с</strong>т<strong>а</strong>дии: неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелые,<br />

м<strong>о</strong>л<strong>о</strong>дые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, жи<strong>в</strong>ут<br />

<strong>в</strong> т<strong>о</strong>й же <strong>с</strong>реде, чт<strong>о</strong> и <strong>в</strong>зр<strong>о</strong><strong>с</strong>лые. Ин<strong>о</strong>гд<strong>а</strong><br />

р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>х<strong>о</strong>дит <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х (Plecopteroidea).<br />

Ст<strong>а</strong>дии инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития <strong>о</strong>бычн<strong>о</strong><br />

п<strong>о</strong>чти лишены <strong>с</strong>пеци<strong>а</strong>льных при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений<br />

<strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении, <strong>о</strong>тлич<strong>а</strong>я<strong>с</strong>ь <strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

лишь нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>итыми п<strong>о</strong>л<strong>о</strong><strong>в</strong>ыми <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми<br />

и <strong>к</strong>рыльями, и н<strong>о</strong><strong>с</strong>ят н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние нимф.<br />

Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. Три н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>: Blattopteroidea,<br />

Plecopteroidea, Orthopteroidea.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

из<strong>в</strong>е<strong>с</strong>тны лишь <strong>в</strong> <strong>о</strong>бщих черт<strong>а</strong>х. Blattopteroidea<br />

и Plecopteroidea — н<strong>а</strong>иб<strong>о</strong>лее дре<strong>в</strong>ние н<strong>а</strong>д<strong>о</strong>тряды,<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующие<strong>с</strong>я р<strong>а</strong>зличными н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лениями<br />

и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития: <strong>с</strong>ильным<br />

р<strong>а</strong>з<strong>в</strong>итием п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ных при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений <strong>в</strong><br />

<strong>к</strong>рыльях, <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ития, п<strong>о</strong>ч<strong>в</strong>енным или н<strong>а</strong>земным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м<br />

жизни (Blattopteroidea), или <strong>с</strong><strong>о</strong>хр<strong>а</strong>нением<br />

пер<strong>в</strong>ичн<strong>о</strong>й г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мн<strong>о</strong><strong>с</strong>ти <strong>о</strong>беих п<strong>а</strong>р<br />

<strong>к</strong>рылье<strong>в</strong>, или м<strong>а</strong>л<strong>о</strong>й <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>а</strong>цией передних<br />

<strong>к</strong>рылье<strong>в</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием <strong>к</strong><strong>а</strong><strong>к</strong>их-либ<strong>о</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бых при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений<br />

<strong>в</strong> инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>м р<strong>а</strong>з<strong>в</strong>итии и<br />

<strong>в</strong><strong>о</strong>дным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м жизни неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых <strong>с</strong>т<strong>а</strong>дий<br />

(Plecopteroidea). П<strong>о</strong><strong>с</strong>ледний н<strong>а</strong>д<strong>о</strong>тряд, п<strong>о</strong><strong>в</strong>идим<strong>о</strong>му,<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я р<strong>а</strong>з<strong>в</strong>итием те<strong>с</strong>ных<br />

<strong>с</strong><strong>в</strong>язей <strong>с</strong> жи<strong>в</strong><strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>тью и <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>а</strong> перед<strong>в</strong>ижения п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м прыг<strong>а</strong>тельных<br />

з<strong>а</strong>дних н<strong>о</strong>г (Orthopteroidea).<br />

НАДОТРЯД BLATTOPTEROIDEA.<br />

ТАРАКАНООБРАЗНЫЕ<br />

Обе п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong> гетер<strong>о</strong>н<strong>о</strong>мны — передние<br />

пл<strong>о</strong>тные, з<strong>а</strong>дние р<strong>а</strong><strong>с</strong>ширенные, переп<strong>о</strong>нч<strong>а</strong>тые.<br />

Ан<strong>о</strong>юг<strong>а</strong>льный <strong>в</strong>еер з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> юг<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>тем<strong>о</strong>й<br />

жил<strong>о</strong><strong>к</strong>: <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>с</strong>мещены <strong>в</strong>перед.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних <strong>к</strong>рылье<strong>в</strong> не <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

р<strong>а</strong>ди<strong>а</strong>льные жил<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> ,не имеют<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й передней <strong>в</strong>ет<strong>в</strong>и, <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>в</strong>ели<strong>к</strong><strong>а</strong>, мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>. Тел<strong>о</strong> упл<strong>о</strong>щенн<strong>о</strong>е,<br />

н<strong>о</strong>ги <strong>с</strong>ильные, бег<strong>а</strong>тельные. Р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong><br />

<strong>с</strong> п<strong>о</strong>ч<strong>в</strong><strong>о</strong>й, <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>о</strong>бр<strong>а</strong>з жизни <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. Ше<strong>с</strong>ть <strong>о</strong>тряд<strong>о</strong><strong>в</strong>:<br />

Blattodea, Manteodea, Isoptera, Dermaptera,<br />

Protelytroptera, Protoblattodea.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения. Эт<strong>о</strong>т дре<strong>в</strong>нейший<br />

н<strong>а</strong>д<strong>о</strong>тряд, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>, я<strong>в</strong>ляет<strong>с</strong>я и<strong>с</strong>х<strong>о</strong>дн<strong>о</strong>й<br />

групп<strong>о</strong>й для других н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> и, <strong>в</strong>е­<br />

ОТРЯД BLATTODEA. ТАРАКАНОВЫЕ<br />

Передние <strong>к</strong>рылья <strong>в</strong> <strong>в</strong>иде пл<strong>о</strong>тны'х н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем,<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й и <strong>с</strong> <strong>о</strong>чень гу<strong>с</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

из мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных те<strong>с</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>с</strong> <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й, реже пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й SC; R, М и<br />

CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые; CuP дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я и<br />

р<strong>о</strong>ятн<strong>о</strong>, других <strong>к</strong><strong>о</strong>г<strong>о</strong>рт. И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие<br />

<strong>в</strong> эт<strong>о</strong>м н<strong>а</strong>д<strong>о</strong>тряде пр<strong>о</strong>х<strong>о</strong>дил<strong>о</strong> р<strong>а</strong>зными путями:<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>о</strong>бщей черт<strong>о</strong>й был<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

р<strong>а</strong>змн<strong>о</strong>жения путем ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>в</strong><strong>с</strong>я<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> з<strong>а</strong>щитных при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений для<br />

яиц (<strong>о</strong><strong>о</strong>те<strong>к</strong>и т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong> и б<strong>о</strong>г<strong>о</strong>м<strong>о</strong>л<strong>о</strong><strong>в</strong>, п<strong>о</strong>дземные<br />

гнезд<strong>а</strong> ух<strong>о</strong><strong>в</strong>ерт<strong>о</strong><strong>к</strong>) или д<strong>а</strong>же <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>к</strong>и<br />

<strong>о</strong>бще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бр<strong>а</strong>з<strong>а</strong> жизни (термиты). Кр<strong>о</strong>ме<br />

т<strong>о</strong>г<strong>о</strong>, пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>бмен<strong>а</strong><br />

<strong>в</strong>еще<strong>с</strong>т<strong>в</strong> и пит<strong>а</strong>ния, причем п<strong>о</strong>лучили<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение у мн<strong>о</strong>гих т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

<strong>с</strong>имби<strong>о</strong>тиче<strong>с</strong><strong>к</strong>ие ми<strong>к</strong>р<strong>о</strong><strong>о</strong>рг<strong>а</strong>низмы <strong>в</strong><br />

<strong>к</strong>ишечни<strong>к</strong>е или п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти тел<strong>а</strong>, <strong>о</strong>бе<strong>с</strong>печи<strong>в</strong><strong>а</strong>ющие<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong>е у<strong>с</strong><strong>в</strong><strong>о</strong>ение р<strong>а</strong><strong>с</strong>тительных <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><br />

(<strong>к</strong>летч<strong>а</strong>т<strong>к</strong>и и <strong>к</strong><strong>с</strong>ил<strong>о</strong>зы) и не<strong>о</strong>бх<strong>о</strong>димые <strong>в</strong> пит<strong>а</strong>нии<br />

<strong>в</strong>ит<strong>а</strong>мины.<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

88<br />

http://jurassic.ru/<br />

<strong>с</strong>лит<strong>а</strong>я <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью Ai, <strong>о</strong>бр<strong>а</strong>зует чет<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женную <strong>а</strong>н<strong>а</strong>льную б<strong>о</strong>р<strong>о</strong>зду, рез<strong>к</strong><strong>о</strong> <strong>о</strong>тделяющую<br />

<strong>а</strong>н<strong>а</strong>льную <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья. Обычн<strong>о</strong><br />

имеют<strong>с</strong>я <strong>а</strong>рхеди<strong>к</strong>тий или мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные т<strong>о</strong>н<strong>к</strong>ие<br />

п<strong>о</strong>перечные жил<strong>к</strong>и. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е, <strong>с</strong> прямым передним<br />

<strong>к</strong>р<strong>а</strong>ем, б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием и п<strong>о</strong>дгиб<strong>а</strong>ющей-

<strong>с</strong>я или <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ющей<strong>с</strong>я <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью.<br />

Тел<strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>- или <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я,<br />

<strong>о</strong>бычн<strong>о</strong> з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я <strong>с</strong><strong>в</strong>ерху передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й.<br />

Антенны длинные, мн<strong>о</strong>г<strong>о</strong>члени<strong>с</strong>тые. Н<strong>о</strong>ги<br />

бег<strong>а</strong>тельны'е, <strong>с</strong> <strong>к</strong>рупными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми; б<strong>о</strong>льшие<br />

шипы н<strong>а</strong> г<strong>о</strong>ленях и <strong>о</strong>бычн<strong>о</strong> н<strong>а</strong> бедр<strong>а</strong>х. Цер<strong>к</strong>и,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, члени<strong>с</strong>тые, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Ин<strong>о</strong>гд<strong>а</strong><br />

имеет<strong>с</strong>я длинный яйце<strong>к</strong>л<strong>а</strong>д; ч<strong>а</strong>ще <strong>о</strong>н <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

и яйц<strong>а</strong> п<strong>о</strong>мещ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>к</strong><strong>а</strong>п<strong>с</strong>улы<br />

— <strong>о</strong><strong>о</strong>те<strong>к</strong>и. Тепл<strong>о</strong>люби<strong>в</strong>ые ф<strong>о</strong>рмы 1<br />

, жи<strong>в</strong>ущие<br />

<strong>в</strong> у<strong>к</strong>рытых <strong>с</strong>т<strong>а</strong>циях: п<strong>о</strong>ч<strong>в</strong>е, р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>х<br />

или з<strong>а</strong>р<strong>о</strong><strong>с</strong>лях р<strong>а</strong><strong>с</strong>тений (ри<strong>с</strong>. 181 и<br />

182). К<strong>а</strong>рб<strong>о</strong>н — ныне. Де<strong>в</strong>ять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Archimylacridae,<br />

Mylacridae, Neorthroblattinidae, Р<strong>о</strong>roblattinidae,<br />

Blattidae, Corydiidae, Panestiidae,<br />

Diplopteridae, Oulopterigidae; три п<strong>о</strong><strong>с</strong>ледних<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО ARCHIMYLACRIDAE<br />

HANDLIRSCH, 1906<br />

(Spiloblattinidae Handlirsch, 1906)<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>а</strong>рхеди<strong>к</strong>тием; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние ег<strong>о</strong><br />

без р<strong>а</strong><strong>с</strong>ширения, ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления <strong>к</strong> груди<br />

ближе <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; SC длинн<strong>а</strong>я, <strong>с</strong> ряд<strong>о</strong>м<br />

<strong>в</strong>ет<strong>в</strong>ей; R и RS ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены; <strong>в</strong>ет<strong>в</strong>и<br />

R, RS и М н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>перед; <strong>с</strong>и<strong>с</strong>темы R и М<br />

р<strong>а</strong>з<strong>в</strong>иты <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>; СиА гребенч<strong>а</strong>т<strong>а</strong>я, <strong>о</strong>т чет<strong>в</strong>ерти<br />

д<strong>о</strong> трех чет<strong>в</strong>ертей длины <strong>к</strong>рыл<strong>а</strong>. В з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле п<strong>о</strong>дгиб<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>д <strong>к</strong>рыл<strong>о</strong>, не <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет <strong>в</strong>переди передне<strong>с</strong>пин<strong>к</strong>и.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я. Цер<strong>к</strong>и члени<strong>с</strong>тые.<br />

Длинный яйце<strong>к</strong>л<strong>а</strong>д, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из<br />

трех п<strong>а</strong>р <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>о</strong><strong>к</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Archimylacrinae и Spiloblattininae.<br />

ПОДСЕМЕЙСТВО ARCHIMYLACRINAE<br />

HANDLIRSCH, 1906<br />

[nom. transl. Cockerell, 1927<br />

(ex Archimylacridae Handlirsch, 1906)]<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> <strong>в</strong> 2—2,8 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше их<br />

ширины; SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> 0,50—0,75 длины н<strong>а</strong>д<strong>к</strong>рылья,<br />

<strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м делящих<strong>с</strong>я и пр<strong>о</strong><strong>с</strong>тых<br />

<strong>в</strong>ет<strong>в</strong>ей; R х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; СиА не д<strong>о</strong>х<strong>о</strong>дит<br />

д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я. Археди<strong>к</strong>тий я<strong>с</strong>ный. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н —<br />

юр<strong>а</strong>. С<strong>а</strong>м<strong>а</strong>я дре<strong>в</strong>няя групп<strong>а</strong> т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ых. О<strong>к</strong><strong>о</strong>д<strong>о</strong><br />

70 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Caenoblatta М. Zalessky, 1931. Тип р<strong>о</strong>д<strong>а</strong>—С.<br />

angaridensis М. Zalessky, 1931; <strong>с</strong>р!<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Черемич<strong>к</strong>ин<strong>о</strong>).<br />

Н<strong>а</strong>д<strong>к</strong>рылье длинн<strong>о</strong>е: ег<strong>о</strong> длин<strong>а</strong> <strong>в</strong> 2,8 р<strong>а</strong>з<strong>а</strong><br />

б<strong>о</strong>льше ширины; <strong>в</strong>ершин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ытянут<strong>а</strong>;<br />

передний и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>я <strong>с</strong>имметричн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лы; SC <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут<strong>а</strong>, <strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; R делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тых <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; М гребенч<strong>а</strong>т<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены<br />

<strong>в</strong>перед; СиА <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й<br />

б<strong>а</strong>з<strong>а</strong>льнее, чем R — <strong>в</strong> передний; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля уже п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины ширины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

27,5 мм (ри<strong>с</strong>. 183). Один <strong>в</strong>ид. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Phylobtatta Handlirsch, 1906 (Tomioblatta,<br />

Martynov, 1931). Тип р<strong>о</strong>д<strong>а</strong> — Blattina<br />

schroteri Giebel, 1857; IB. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Герм<strong>а</strong>ния.<br />

Н<strong>а</strong>д<strong>к</strong>рылье пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> эллиптиче<strong>с</strong><strong>к</strong><strong>о</strong>е; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле лент<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е; SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> 0,50—0,66<br />

длины н<strong>а</strong>д<strong>к</strong>рылья; R без я<strong>с</strong>н<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

RS,c ряд<strong>о</strong>м делящих<strong>с</strong>я <strong>в</strong>ет<strong>в</strong>ей, <strong>в</strong><strong>с</strong>егд<strong>а</strong> з<strong>а</strong>ним<strong>а</strong>ет<br />

переднюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину <strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь R <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не <strong>в</strong>п<strong>а</strong>дения ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и СиА <strong>в</strong> ег<strong>о</strong><br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее; СиА<br />

п<strong>о</strong>л<strong>о</strong>г<strong>а</strong>я, <strong>с</strong> ше<strong>с</strong>тью-д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>тью <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й близ <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти уз<strong>к</strong><strong>а</strong>я;<br />

длин<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля р<strong>а</strong><strong>в</strong>н<strong>а</strong> трети длины<br />

н<strong>а</strong>д<strong>к</strong>рылья; ряд не<strong>с</strong>ли<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 11—35 мм<br />

(pHc v<br />

184; т<strong>а</strong>бл. III, фиг. 1, т<strong>а</strong>бл. IV, фиг. 1 —<br />

2). О<strong>к</strong><strong>о</strong>л<strong>о</strong> 190 <strong>в</strong>ид<strong>о</strong><strong>в</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> и перми 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы, С. и Ю. Амери<strong>к</strong>и; три <strong>в</strong>ид<strong>а</strong> из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; три <strong>в</strong>ид<strong>а</strong> из <strong>в</strong>.Перми<br />

Приур<strong>а</strong>лья. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong>. пермь.<br />

Aissobldtta Handlirsch, 1904. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. rossica Handlirsch, 1904; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье эллип<strong>с</strong><strong>о</strong>идн<strong>о</strong><br />

<strong>в</strong>ытянут<strong>о</strong>, ег<strong>о</strong> длин<strong>а</strong> <strong>в</strong> 2,25 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ширины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong> <strong>в</strong>ершине <strong>с</strong>лег<strong>к</strong><strong>а</strong><br />

р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> 0,7 длины н<strong>а</strong>д<strong>к</strong>рылья,<br />

имеет мн<strong>о</strong>г<strong>о</strong> п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; М и R делят<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, гребенч<strong>а</strong>тые, их<br />

<strong>в</strong>ет<strong>в</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>перед; М <strong>с</strong>игм<strong>о</strong>ид<strong>а</strong>льн<strong>о</strong><br />

из<strong>о</strong>гнут<strong>а</strong>; СиА при<strong>с</strong><strong>о</strong>единяет ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей з<strong>а</strong>днег<strong>о</strong><br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М <strong>в</strong> <strong>в</strong>иде <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

СиА; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле р<strong>а</strong><strong>в</strong>н<strong>о</strong> 0,36 длины н<strong>а</strong>д<strong>к</strong>рылья<br />

и 0,6 ширины ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 30—36 мм (ри<strong>с</strong>. 185). Три <strong>в</strong>ид<strong>а</strong>.<br />

Пермь Приур<strong>а</strong>лья.<br />

89<br />

http://jurassic.ru/<br />

Kunguroblattina Martynov, 1930. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К. arcuata Martynov, 1930; н. пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Б<strong>а</strong>рд<strong>а</strong>).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем; R и<br />

М делят<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я<br />

дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е: шире п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья;<br />

передняя <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

з<strong>а</strong>дняя — гребенч<strong>а</strong>т<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 53—<br />

55 мм (ри<strong>с</strong>. 186, 187 А, Б). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.

188Б<br />

Ри<strong>с</strong>. 183—189. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacrinae<br />

183. Caenoblatta angaridensis M. Zalessky; н<strong>а</strong>д<strong>к</strong>рылье, x 2,7,<br />

cp <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (M. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1931). 184. Рпуloblatta<br />

distincta Martynov; н<strong>а</strong>д<strong>к</strong>рылье, х 2,5, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933). 185. Aissoblatta rosslca Hand-<br />

i<br />

1<br />

, ?f Д^рылье, х 1,5, пермь, Ю. Приур<strong>а</strong>лье (Handlirsch,<br />

1904). 18<strong>о</strong>. Kunguroblattina arcuata Martynov; н<strong>а</strong>д<strong>к</strong>рылье, x 1,15<br />

н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1930). 187. Kunguroblattina microuictya<br />

Becker-Migdisova et Vishnjkova; A — <strong>о</strong>бщий <strong>в</strong>ид, x 2,7-<br />

90<br />

http://jurassic.ru/<br />

Ant — <strong>а</strong>нтенны, Cx — т<strong>а</strong>зи<strong>к</strong>, Fm — бедр<strong>о</strong>, Hd — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, Tb —<br />

г<strong>о</strong>лень, Tr —<strong>в</strong>ертлуг, Ts —л<strong>а</strong>п<strong>к</strong><strong>а</strong>, О — яйце<strong>к</strong>л<strong>а</strong>д; Б — яйце<strong>к</strong>л<strong>а</strong>д,<br />

х 4,0; н. пермь, Ур<strong>а</strong>л (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1957). 188A Uraloblatta<br />

mslgnis Q. Zalessky; <strong>о</strong>бщий <strong>в</strong>ид, x 2,2, н. пермь, Ур<strong>а</strong>л<br />

(Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1939). 188Б. Patrodoron auspicatum Q. ZalessKy;<br />

н<strong>а</strong>д<strong>к</strong>рылье, xl, н. Пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1955). 189<br />

Kisylblatta unifasciata Martynov; н<strong>а</strong>д<strong>к</strong>рылье, х 9,0, н. юр<strong>а</strong>.<br />

Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)

Patrodoron G. Zalessky, 1955. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. auspicatum G. Zalessky, 1955; н. пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ая-К<strong>о</strong>шеле<strong>в</strong><strong>к</strong><strong>а</strong>).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем,<br />

длин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> 2,8 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; SG<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> 0,7 длины н<strong>а</strong>д<strong>к</strong>рылья; R и СиА делят<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, т. е. з<strong>а</strong>дний <strong>с</strong>т<strong>в</strong><strong>о</strong>л М при<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я<br />

<strong>к</strong> СиА; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я;<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, шир<strong>о</strong><strong>к</strong><strong>о</strong>е:<br />

шире п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья,<br />

длин<strong>а</strong> ег<strong>о</strong> менее трети длины н<strong>а</strong>д<strong>к</strong>рылья; <strong>о</strong>бе<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и гребенч<strong>а</strong>тые. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

50—55 мм (ри<strong>с</strong>. 188 Б). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н.<br />

пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Uraloblatta G. Zalessky, 1939. Тип р<strong>о</strong>д<strong>а</strong><br />

— U. insignis G. Zalessky, 1939; н. пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Б<strong>а</strong>рд<strong>а</strong>).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; SC р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

0,6 длины <strong>к</strong>рыл<strong>а</strong>, <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>в</strong>ет<strong>в</strong>ями; R и RS мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые, <strong>в</strong> <strong>с</strong>ередине<br />

н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong> изгиб<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д; п<strong>о</strong>ля между<br />

SC и R и между М и СиА <strong>с</strong>ужены. Яйце<strong>к</strong>л<strong>а</strong>д<br />

длинный. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 15 мм (ри<strong>с</strong>.<br />

188А). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Kisylblatta Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. unifasciata Martynov, 1937; н. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е;<br />

ег<strong>о</strong> длин<strong>а</strong> <strong>в</strong> 2,2 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; SC<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья, четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

<strong>в</strong>ет<strong>в</strong>и R пр<strong>о</strong><strong>с</strong>тые, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>перед;<br />

М делит<strong>с</strong>я* немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее, чем R, дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

из<strong>о</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д, гребенч<strong>а</strong>т<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены<br />

<strong>в</strong>перед; СиА <strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д,<br />

<strong>в</strong>ет<strong>в</strong>и длинные, мн<strong>о</strong>г<strong>с</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые, <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> 0,27 длины н<strong>а</strong>д<strong>к</strong>рылья и 0,6 ширины<br />

ег<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. Дли<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

6,6 мм (ри<strong>с</strong>. 189). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Balachonoblatta Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — В. zheltojarica Becker-Megdisova,<br />

1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, ег<strong>о</strong> длин<strong>а</strong> <strong>в</strong> 1,7 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ширины; ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>-притуплённ<strong>о</strong>е,<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е, передний и з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>я п<strong>о</strong>чти прямые; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC длинн<strong>а</strong>я, д<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти д<strong>о</strong><br />

<strong>в</strong>ершины; R и М р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты; М рез<strong>к</strong><strong>о</strong><br />

з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю, гребенч<strong>а</strong>т<strong>а</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>перед; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е; СиА <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, б<strong>о</strong>г<strong>а</strong>т<strong>а</strong> изред<strong>к</strong><strong>а</strong> делящими<strong>с</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ями; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я:<br />

менее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины,длины н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 25—30 мм (ри<strong>с</strong>. 190; т<strong>а</strong>бл. III,<br />

фиг. 2). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Hemimylacretta Handlirsch, 1922. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Hemimylacris ramificata Handlirsch,<br />

1922; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong><br />

шир<strong>о</strong><strong>к</strong>им и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем; SC <strong>с</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я: р<strong>а</strong><strong>в</strong>н<strong>а</strong> или<br />

менее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины н<strong>а</strong>д<strong>к</strong>рылья, <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я<br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; М <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>в</strong>ершину <strong>к</strong>рыл<strong>а</strong>;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я и <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти шир<strong>о</strong><strong>к</strong>ие;<br />

СиА б<strong>о</strong>г<strong>а</strong>т<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ями; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я,<br />

<strong>с</strong> трехр<strong>а</strong>здельн<strong>о</strong>й А. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

21—39 мм (ри<strong>с</strong>. 191). Четыре <strong>в</strong>ид<strong>а</strong>, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле<br />

<strong>о</strong>дин из Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

Sibiroblatta Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — S. flexuosa Becker-Megdisova, 1961;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр). Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>в</strong> 2,7 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

длинн<strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong>е 0,8 длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

R и М <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>с</strong>имметричн<strong>о</strong>: R з<strong>а</strong>ним<strong>а</strong>ет передний, <strong>а</strong> М —<br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й у <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; R <strong>с</strong> д<strong>в</strong>умя<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми и <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями; М делит<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong> длинными р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, и<br />

четырьмя п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями; <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть треуг<strong>о</strong>льн<strong>а</strong>я; СиА п<strong>о</strong>л<strong>о</strong>г<strong>а</strong>я, <strong>с</strong> ч<strong>а</strong><strong>с</strong>тыми<br />

<strong>в</strong>ет<strong>в</strong>ями, <strong>о</strong>дн<strong>а</strong> из них н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong>перед.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 25—28 мм (ри<strong>с</strong>. 192). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

Grypoblattites Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — G. flabellatus Becker-Migdisova,<br />

1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр). Н<strong>а</strong>д<strong>к</strong>рылье<br />

эллиптиче<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним<br />

<strong>к</strong>р<strong>а</strong>ем; длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ширины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е: б<strong>о</strong>лее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины н<strong>а</strong>д<strong>к</strong>рылья, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лег<strong>к</strong><strong>а</strong><br />

р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; SC <strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>к</strong><strong>о</strong><strong>с</strong>ых, нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>ильч<strong>а</strong>тых <strong>в</strong>ет<strong>в</strong>ей;<br />

Ri <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, лишь <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>д<strong>в</strong>инут<strong>а</strong><br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию, <strong>с</strong> длинными <strong>в</strong>ет<strong>в</strong>ями; М <strong>в</strong><br />

<strong>с</strong>ередине дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я, <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я <strong>к</strong><br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; СиА дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я, з<strong>а</strong>ним<strong>а</strong>ет<br />

0,3 или менее длины н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong> пятью —<br />

<strong>с</strong>емью <strong>с</strong>ближенными <strong>в</strong>ет<strong>в</strong>ями; А] <strong>о</strong>бычн<strong>о</strong> трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 25—30 мм<br />

(ри<strong>с</strong>. 193; т<strong>а</strong>бл. III, фиг. 3). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Ср.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Archaeotiphites Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — A. petroblattinoides Becker-<br />

Migdisova, 1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем и<br />

прямым з<strong>а</strong>дним; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>ним<strong>а</strong>ет 0,6<br />

длины н<strong>а</strong>д<strong>к</strong>рылья, SC <strong>о</strong>бр<strong>а</strong>зует б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong><br />

91<br />

http://jurassic.ru/

196<br />

Ри<strong>с</strong>. 190—197. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacrinae'<br />

190. Balachonoblatta zheltojarica Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

x 1,8 и <strong>а</strong>рхеди<strong>к</strong>тий, х 15,0, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 191. Hemimylacrella teptophlebtopsis<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 1,6, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 192. Sibiroblatta flexuosa Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 2,46, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 193. Grypoblattites jlabellatus<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 3,45, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 194. Archaeotiphites petro-<br />

http://jurassic.ru/<br />

btattinoldes Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 1,93, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 195. Miaroblatta<br />

balachonensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 2,53, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 196. Ignaroblatta<br />

panda Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 2,16, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 197. Aphthoroblattina<br />

jascigera (Scudder); н<strong>а</strong>д<strong>к</strong>рылье, x 1,4, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Handlircsh, 1906)

Ри<strong>с</strong>. 198—202. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacrinae и ф<strong>о</strong>рмы incertae sedis<br />

198. Tomacblatta bashklrlca M. Zalessky; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,3,<br />

<strong>в</strong>. пермь, Приур<strong>а</strong>лье (М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1929). 199. Sardycblatta<br />

tlkhvinskii М. Zalessky; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,7, <strong>в</strong>. пермь, Приур<strong>а</strong>лье<br />

(М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1928). 200. Ungoneurites pauctnervls<br />

Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,3, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

п<strong>о</strong>л<strong>о</strong>гих, нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных <strong>в</strong>ет<strong>о</strong>че<strong>к</strong>;<br />

R з<strong>а</strong>ним<strong>а</strong>ет ч<strong>а</strong><strong>с</strong>ть передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины н<strong>а</strong>д<strong>к</strong>рылья,<br />

гребенч<strong>а</strong>т<strong>а</strong>я или делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

и <strong>о</strong>бр<strong>а</strong>зует ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей; М з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю<br />

<strong>в</strong>ершину и треть з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я, делит<strong>с</strong>я н<strong>а</strong><br />

д<strong>в</strong><strong>а</strong> дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и <strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; СиА<br />

делит<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> R, рез<strong>к</strong><strong>о</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зует четрые-пять <strong>с</strong>ближенных<br />

<strong>в</strong>ет<strong>в</strong>ей; CuP дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я; ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей А <strong>с</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 27—30 мм<br />

(ри<strong>а</strong> 194; т<strong>а</strong>бл. III, фиг. 4). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Ср.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Miaroblatta Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong><br />

— М. elata Handlirsch, 1906; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>и,<br />

Фр<strong>а</strong>нция. Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> гребенч<strong>а</strong>т<strong>о</strong>й SC и<br />

длинным <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем, р<strong>а</strong><strong>в</strong>ным 0,66—<br />

0,77 длины н<strong>а</strong>д<strong>к</strong>рылья; R <strong>с</strong> четырьмя — ше<strong>с</strong>тью<br />

п<strong>о</strong>л<strong>о</strong>гими <strong>в</strong>ет<strong>в</strong>ями, д<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины<br />

н<strong>а</strong>д<strong>к</strong>рылья; М з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong>ершинную ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья,<br />

<strong>с</strong> прямыми <strong>в</strong>ет<strong>в</strong>ями, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными<br />

<strong>в</strong>перед; СиА <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и <strong>с</strong>ближены 1<br />

, п<strong>о</strong>чти<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>й длины, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь при<strong>с</strong><strong>о</strong>еди­<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 201. Olratia vallda Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

х 6,2, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 202.<br />

Parapanorpa ungensis М. Zalessky; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,3, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1931)<br />

няет<strong>с</strong>я <strong>к</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>лу <strong>с</strong>переди, у <strong>в</strong>ершины <strong>о</strong>н<strong>а</strong> п<strong>о</strong>л<strong>о</strong>г<strong>а</strong>я;<br />

<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле б<strong>о</strong>льш<strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong>е 0,4 длины н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong><br />

11 п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

35—40 мм (ри<strong>с</strong>. 195). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле<br />

<strong>о</strong>дин из Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Ignaroblatta Becker-Migdisova, 196L<br />

Тип р<strong>о</strong>д<strong>а</strong> —A panda Becker-Migdisova, 1961;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр). Передний <strong>к</strong>р<strong>а</strong>й<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>е 0,55—0,60 длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

SC <strong>с</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными, делящими<strong>с</strong>я <strong>в</strong>ет<strong>в</strong>ями;<br />

R б<strong>о</strong>г<strong>а</strong>т<strong>а</strong>я, з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю <strong>в</strong>ершину н<strong>а</strong>д<strong>к</strong>рылья,<br />

делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, или же пер<strong>в</strong>ые<br />

<strong>в</strong>ет<strong>в</strong>и <strong>с</strong>ближены <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; М делит<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, з<strong>а</strong>ним<strong>а</strong>ет ч<strong>а</strong><strong>с</strong>ть з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

у <strong>в</strong>ершины; СиА п<strong>о</strong>л<strong>о</strong>г<strong>о</strong>-прям<strong>а</strong>я ИЛИ <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я<br />

<strong>с</strong> ше<strong>с</strong>тью-<strong>с</strong>емью <strong>в</strong>ет<strong>в</strong>ями, ч<strong>а</strong><strong>с</strong>ть из<br />

них делит<strong>с</strong>я; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые.<br />

93<br />

http://jurassic.ru/

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 25—30 мм (ри<strong>с</strong>. 196). Три<br />

<strong>в</strong>ид<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Aphthoroblattina Handlirsch, 1906. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Blattina fascigera Scudder, 1876; <strong>с</strong>р.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> 2,5<br />

р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е<br />

и длинн<strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong>е 0,6 длины н<strong>а</strong>д<strong>к</strong>рылья; <strong>в</strong>ет<strong>в</strong>и<br />

SC делят<strong>с</strong>я пуч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>; Ri <strong>с</strong> пятью <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ями; RS делит<strong>с</strong>я<br />

н<strong>а</strong> четыре — ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей; М делит<strong>с</strong>я у<br />

<strong>с</strong>ередины 1<br />

н<strong>а</strong>д<strong>к</strong>рылья н<strong>а</strong> четыре <strong>в</strong>ет<strong>в</strong>и; CuA <strong>с</strong><br />

<strong>с</strong>емью — <strong>в</strong><strong>о</strong><strong>с</strong>емью <strong>в</strong>ет<strong>в</strong>ями, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>к</strong><br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 24—38 мм (ри<strong>с</strong>. 197). Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

С. Амери<strong>к</strong>и.<br />

Tomacblatta М. Z <strong>а</strong> 1 е s s <strong>к</strong> у, 1929. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. baschkirica М. Zalessky, 1929; <strong>в</strong>. пермь,<br />

Приур<strong>а</strong>лье (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, р. Уяз<strong>а</strong>). З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> идущей близ <strong>к</strong>р<strong>а</strong>я и <strong>в</strong>д<strong>о</strong>ль нег<strong>о</strong> SC;<br />

R з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>е пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>о</strong> д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>,<br />

делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>: м<strong>а</strong>л<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тую Ri<br />

или PR и гребенч<strong>а</strong>тую RS; М м<strong>а</strong>л<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

14 мм (ри<strong>с</strong>. 198). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Sardycblatta М. Zalessky, 1928. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. tikhvinskii М. Zalessky, 1928; <strong>в</strong>. пермь,<br />

Приур<strong>а</strong>лье (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, р. С<strong>а</strong>рды<strong>к</strong>). З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем; SC <strong>с</strong>лег<strong>к</strong><strong>а</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я, <strong>с</strong> длинными<br />

<strong>в</strong>ет<strong>в</strong>ями; R гребенч<strong>а</strong>т<strong>а</strong>я; Ri п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ям<br />

RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й р<strong>а</strong>нее <strong>в</strong>ершины;<br />

М х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тых <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

13 мм (ри<strong>с</strong>. 199). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Вне СССР: Acoblatta Cockerell, 1927; Acosmoblatta<br />

Handlirsch, 1906; Actinoblatta Pruvost,<br />

1912; Adetobtatta Handlirsch, 1906; Amblyblatta<br />

Handlirsch, 1906; Amoeboblatta Handlirsch,<br />

1906; Amorphoblatta Handlirsch, 1906; Anomobtatta<br />

Handlirsch, 1906; Anthracoblattina Scudder,<br />

1879; Apempherus Handlirsch, 1906;<br />

Aphthoroblattina Handlirsch, 1906; Apotypoma<br />

Handlirsch, 1906; Archaeotiphe Pruvost,<br />

1919; Archimylacris Scudder, 1868; Archoblattina<br />

Sellards, 1903; Asemoblatta Handlirsch,<br />

1906; Atimoblatta Handlirsch, 1906;<br />

Auxanoblatta Handlirsch, 1906; Barroisiblatta<br />

Pruvost, 1919; Bertrandiblatta Laurentiaux,<br />

1950; Boltonopruvostia Strand, 1929; Bradyblatta<br />

Handlirsch, 1906; Cebennibtatta Laurentiaux,<br />

1950; Cobalobtatta Cockerell, 1919; Compsoblattula<br />

Strand, 1929; Coraloblatta Cockerell,<br />

1918; Crokoblattina Guthorl, 1933; Dictyoblatta<br />

Handlirsch, 1906; Dimoblatta Cockerell, 1927;<br />

94<br />

http://jurassic.ru/<br />

Discobtatta Handlirsch, 1906; Distatoblatta Handlirsch,<br />

1906; Dromoblatta Handlirsch, 1906;<br />

Dysmenes Handlirsch, 1906; Elaphroblatta<br />

Handlirsch, 1906; Eneriblatta Laurentiaux,<br />

1949; Etoblattina Scudder, 1879; E amorphoblatta<br />

Handlirsch, 1906; Exochoblatta Handlirsch,<br />

1906; Flabellites Fritsch, 1895; Friciella<br />

Kukalova, 1955; Gondwanoblatta Handlirsch,<br />

1906; Gongylobtatta Handlirsch, 1906;<br />

Grypoblattina Pruvost, 1919; Gyroblatta<br />

Handlirsch, 1906; Hesperoblatta Handlirsch,<br />

1906; Kafar Handlirsch, 1920; Kinklidoblatta<br />

Handlirsch, 1906; Kinklidoptera Handlirsch,<br />

1906; Leptoblattina Handlirsch, 1906; Liparoblatta<br />

Handlirsch, 1906; Livetiblatta Laurentiaux,<br />

1950; Loxoblatta Cockerell, 1927; Manoblatta<br />

Pruvost, 1919; Mesitoblatta Handlirsch,<br />

1906; Metachorus Handlirsch, 1906;<br />

Metaphyloblatta Kukalova, 1955; Metapoblatta<br />

Cockerell, 1927; Metaxys Handlirsch, 1906; Pachyblatta<br />

Cockerell, 1927; Parelthoblatta Handlirsch,<br />

1906; Penetoblatta Handlirsch, 1906; Petrablattina<br />

Scudder, 1885; Phautoblatta Handlirsch,<br />

1906; Phoberoblatta Handlirsch, 1906;<br />

Phytoblatta Handlirsch, 1906; Plagioblatta Handlirsch<br />

1906; Platyblatta Handlirsch, 1906; Platyblattina<br />

Laurentiaux, 1950; Polyctoblatta Handlirsch<br />

1906; Prantlites Kukalova, 1955; Procoptoblatta<br />

Handlirsch, 1920; Progonoblattina Scudder,<br />

1879; Schizoblatta Handlirsch, 1906; Schizoblattina<br />

Sellards, 1904; Scuderutla Handlirsch,<br />

1920; Sellardsuta Handlirsch, 1920; Sooblatella<br />

Handlirsch, 1920; Sphaleroblattina Schlechtendal,<br />

1879; Stephanoblatta Handlirsch, 1906; Sterzelia<br />

Handlirsch, 1906; Stigetoblatta Handlirsch,<br />

1906; Sulcoboltonia Strand, 1929; Symphyoblatta<br />

Handlirsch, 1906; Syncoptoblatta Handlirsch,<br />

1906; Xenoblatta Handlirsch, 1906.<br />

ARCHIMYLACRINAE INCERTAE SEDIS<br />

З<strong>а</strong>дние <strong>к</strong>рылья неп<strong>о</strong>лн<strong>о</strong>й <strong>с</strong><strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ти. Р<strong>о</strong>ды:<br />

Ungoneurites Martynov, 1931 (ри<strong>с</strong>. 200),<br />

Oiratia Martynov, 1931 (ри<strong>с</strong>. 201), Parapanorpa<br />

M. Zalessky, 1931 (ри<strong>с</strong>. 202) из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

ПОДСЕМЕЙСТВО SPILOBLATTININAE<br />

HANDLIRSCH, 1906<br />

[nom. transl. Becker-Migdisova, 1961<br />

(ex Spiloblattinidae Handlirsch, 1906)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными передним и<br />

з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ями; длин<strong>а</strong> их <strong>в</strong> 2,5—3,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ширины; п<strong>о</strong>ля между SC и R, R и М, М и<br />

CuA <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны и р<strong>а</strong><strong>с</strong>ширены <strong>в</strong> <strong>с</strong>ередине; SC<br />

длинн<strong>а</strong>я, <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных пр<strong>о</strong><strong>с</strong>тых <strong>в</strong>ет-

<strong>в</strong>ей; СиА длинн<strong>а</strong>я—д<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины<br />

<strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями; <strong>к</strong>убительн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

уз<strong>к</strong><strong>а</strong>я; <strong>а</strong>рхеди<strong>к</strong>тий нежный, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong>метен<br />

лишь <strong>в</strong>д<strong>о</strong>ль жил<strong>о</strong><strong>к</strong>. К<strong>а</strong>рб<strong>о</strong>н — н. пермь.<br />

B. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; н. пермь Е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти СССР; е. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н и н. пермь<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и. 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Sysciophlebia Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong><br />

— Blattina euglyptica Germar, 1851; <strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Герм<strong>а</strong>ния. Н<strong>а</strong>д<strong>к</strong>рылье эллиптиче<strong>с</strong><strong>к</strong>и <strong>в</strong>ытянут<strong>о</strong>е,<br />

ред<strong>к</strong><strong>о</strong> п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>о</strong><strong>к</strong>ругленн<strong>о</strong>е, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем и прямым з<strong>а</strong>дним,<br />

длин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> 2,5—3,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е и длинн<strong>о</strong>е, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

0,50—0,75 длины н<strong>а</strong>д<strong>к</strong>рылья; SC п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю, <strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей;<br />

Ri не<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я или пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

или <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; R и М гребенч<strong>а</strong>тые,<br />

<strong>в</strong>ет<strong>в</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>перед; М з<strong>а</strong>ним<strong>а</strong>ет<br />

<strong>в</strong><strong>с</strong>ю <strong>с</strong>ередину н<strong>а</strong>д<strong>к</strong>рылья; СиА длинн<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и<br />

з<strong>а</strong>ним<strong>а</strong>ют п<strong>о</strong>чти <strong>в</strong>е<strong>с</strong>ь з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й; СиА и<br />

CuP рез<strong>к</strong><strong>о</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 11—32 мм (ри<strong>с</strong>. 203). Один <strong>в</strong>ид из<br />

н. перми Ур<strong>а</strong>л<strong>а</strong>, <strong>о</strong>дин <strong>в</strong>ид из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> (З<strong>а</strong><strong>в</strong>ьял<strong>о</strong><strong>в</strong><strong>о</strong>) и б<strong>о</strong>лее 50 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> и н. перми 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

C. Амери<strong>к</strong>и. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — н. пермь.<br />

Olethroblatta Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong>—<br />

О. americana Handlirsch, 1906; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong>. Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, передний<br />

и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>я <strong>в</strong>ыпу<strong>к</strong>лые; длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>в</strong> 2—2,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле лент<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е, умеренн<strong>о</strong>й ширины,<br />

длин<strong>а</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет 0,5—0,6 длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е, р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е и меди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширены; R и М делят<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

н<strong>а</strong>д<strong>к</strong>рылья или М — диет<strong>а</strong>льг<strong>а</strong>ее <strong>с</strong>ередины; <strong>в</strong>ет<strong>в</strong>и<br />

R и М <strong>с</strong>ближены, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены пуч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>;<br />

СиА <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле з<strong>а</strong>ним<strong>а</strong>ет 0,4 длины <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>рхеди<strong>к</strong>тий<br />

<strong>о</strong>чень нежный, <strong>в</strong> <strong>в</strong>иде из<strong>в</strong>или<strong>с</strong>тых п<strong>о</strong>перечных.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 17—25 мм (ри<strong>с</strong>.<br />

204). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>дин из <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Верхи <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и;<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н и н. пермь 3. Е<strong>в</strong>р<strong>о</strong>пы. К<strong>а</strong>рб<strong>о</strong>н—н.<br />

пермь.<br />

Zavjaloblatta Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong>—Z. rotundata Becker-Migdisova,<br />

1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр) • Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле лент<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е; SC<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong><br />

<strong>в</strong>п<strong>а</strong>дением СиА <strong>в</strong> з<strong>а</strong>дний '<strong>к</strong>р<strong>а</strong>й; R <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных,<br />

прямых, неделящих<strong>с</strong>я <strong>в</strong>ет<strong>в</strong>ей; М<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong><br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>в</strong>ершины, <strong>с</strong> ряд<strong>о</strong>м<br />

РИС. 203—205. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Spiloblattininae<br />

203- Sysciophlebia zavjalovensts Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

x 2,9, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 204. Olethroblatta tomiensis Becker-Migdisova; А —н<strong>а</strong>д<strong>к</strong>рылье<br />

х 2,9; Б и В — дет<strong>а</strong>ли <strong>а</strong>рхеди<strong>к</strong>тий, х 15,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 205. Zavjaloblatta<br />

rotundata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 2,8, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

п<strong>а</strong>р<strong>а</strong>ллельных дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных<br />

<strong>в</strong>перед и <strong>о</strong>бр<strong>а</strong>щенных <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>тью <strong>к</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю; <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й<br />

ширины; СиА <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю, ее <strong>в</strong>ет<strong>в</strong>и р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>; <strong>а</strong>рхеди<strong>к</strong>тий <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ий и мел<strong>к</strong><strong>о</strong><strong>с</strong>етч<strong>а</strong>тый<br />

п<strong>о</strong> <strong>в</strong><strong>с</strong>ей п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии СиА<br />

между пер<strong>в</strong><strong>о</strong>й и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ями п<strong>о</strong> ше<strong>с</strong>ть-<strong>с</strong>емь<br />

ячее<strong>к</strong> <strong>в</strong> п<strong>о</strong>перечни<strong>к</strong>е п<strong>о</strong>ля. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

20 мм (ри<strong>с</strong>. 205; т<strong>а</strong>бл. IV, фиг. 3). Один <strong>в</strong>ид.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Ametrobtatta Handlirsch, 1906;<br />

Atactoblatta Handlirsch, 1906; Doryblatta Handlirsch,<br />

1906; Syscioblatta Handlirsch, 1906: Dicladoblatta<br />

Handlirsch, 1906; Pareinobtatta Handlirsch,<br />

1906; Arrhythmoblatta Handlirsch, 1906;<br />

Permoblattina Tillyard, 1937; Spiloblattina<br />

Scudder, 1885.<br />

95 http://jurassic.ru/<br />

СиА<br />

Э05

CuP<br />

Ри<strong>с</strong>. 206—212. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mylacridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Jiemimylacrinae, Idiomylacrinae, Mylacrinae, Pteridomylacrinae<br />

206. Hemlmylacrts ctintoniana Scudder; н<strong>а</strong>д<strong>к</strong>рылье, x3,l,"cp<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Scudder, 1895). 207. Metaxybtatta gromotucnensis<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье.'х 2,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 208. Idiomylacrls gracilis<br />

Handlirsch; н<strong>а</strong>д<strong>к</strong>рылье, х 3,5, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Handlirsch,<br />

1906). 209. • Mylacris anthracophila Scudder; н<strong>а</strong>д<strong>к</strong>рылье,<br />

СЕМЕЙСТВО MYLACRIDAE SCUDDER, 1886<br />

(Hemimylacridae Waterlot, 1934)<br />

Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е; ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления<br />

<strong>к</strong> груди р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> п<strong>о</strong>чти т<strong>о</strong>чн<strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле треуг<strong>о</strong>льн<strong>о</strong>е,<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е, п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>е<br />

п<strong>о</strong> длине <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти; SC прям<strong>а</strong>я, ее<br />

<strong>в</strong>ет<strong>в</strong>и <strong>в</strong>ых<strong>о</strong>дят из <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния, реже <strong>о</strong>н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я или гребенч<strong>а</strong>т<strong>а</strong>я, н<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

96<br />

http://jurassic.ru/<br />

x 1,8, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С Амери<strong>к</strong><strong>а</strong> (Scudder, 1895). 210. Pseudomylacrls<br />

wettlnensis Schlechtendal; н<strong>а</strong>д<strong>к</strong>рылье, x 8,25, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Герм<strong>а</strong>ния (Handlirsch. 1906). 211. Neomylacrls major Handlirsch<br />

н<strong>а</strong>д<strong>к</strong>рылье, x 2,8, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Handlirsch,<br />

1906). 212. Pterldomylacrls paradoxa Handlirsch; н<strong>а</strong>д<strong>к</strong>рылье;<br />

x 3,0, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1906)<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельных пр<strong>о</strong><strong>с</strong>тых<br />

<strong>в</strong>ет<strong>в</strong>ей; R и CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>ты; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

б<strong>о</strong>льш<strong>а</strong>я, <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong><br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья, реже — <strong>в</strong> <strong>а</strong>н<strong>а</strong>льную<br />

б<strong>о</strong>р<strong>о</strong>зду. Ан<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

не <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, <strong>а</strong> п<strong>о</strong>дгиб<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>д <strong>к</strong>рыл<strong>о</strong>. Передне<strong>о</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>а</strong>я,<br />

п<strong>о</strong>перечн<strong>а</strong>я, <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Тел<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е. Ст<strong>в</strong><strong>о</strong>р<strong>к</strong>и яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong>. пермь. Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Hemimylacrinae, Mylacrinae, Idiomylacrinae,<br />

Pteridomylacrinae.

ПОДСЕМЕЙСТВО HEMIMYLACRINAE PRUVOST, 1919<br />

[nom. transl. Becker-Migdisova, 1961<br />

(ex Hemimylacrides Pruvost, 1919)]<br />

[Hemimylacridae Waterlot, 1934;<br />

Archimylacridae Handlirsch, 1906 (pars)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии умеренн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, треуг<strong>о</strong>льн<strong>о</strong>е; SC<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> гребенч<strong>а</strong>т<strong>а</strong>я <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными<br />

пр<strong>о</strong><strong>с</strong>тыми <strong>в</strong>ет<strong>в</strong>ями <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; М <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й, ди<strong>с</strong>т<strong>а</strong>льные <strong>к</strong><strong>о</strong>нцы их не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 19—40 мм (ри<strong>с</strong>. 206). Ср. и <strong>в</strong>.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Metaxybtatta Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. hadroptera Handlirsch, 1906; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong>. Н<strong>а</strong>д<strong>к</strong>рылье немн<strong>о</strong>г<strong>о</strong> б<strong>о</strong>лее чем <strong>в</strong><br />

2 р<strong>а</strong>з<strong>а</strong> длиннее ширины, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем и д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

SC; <strong>о</strong>дин <strong>с</strong>т<strong>в</strong><strong>о</strong>л гребенч<strong>а</strong>тый, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong><br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>в</strong>ет<strong>в</strong>ей пр<strong>о</strong>х<strong>о</strong>дит п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю н<strong>а</strong>д<strong>к</strong>рылья; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я; R прям<strong>а</strong>я, <strong>с</strong> четырьмя — <strong>с</strong>емью делящими<strong>с</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ями; М з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю <strong>в</strong>ершину,<br />

из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю, гребенч<strong>а</strong>т<strong>а</strong>я, <strong>с</strong><br />

ше<strong>с</strong>тью — <strong>в</strong><strong>о</strong><strong>с</strong>емью делящими<strong>с</strong>я <strong>в</strong>ет<strong>в</strong>ями, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными<br />

<strong>в</strong>перед; <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong> <strong>с</strong>емью-<strong>в</strong><strong>о</strong><strong>с</strong>емью р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

<strong>в</strong>ет<strong>в</strong>ями; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е,<br />

<strong>о</strong>тделен<strong>о</strong> п<strong>о</strong>л<strong>о</strong>г<strong>о</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 23—24 мм (ри<strong>с</strong>. 207). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С.<br />

tendal, 1879.<br />

Вне СССР: Hemimylacris Handlirsch, 1906;<br />

Cyphomylacris Cockerell, 1927; Discomylacris<br />

Handlirsch, 1920; Drybrookia Bolton, 1923; Loxoblatta<br />

Cockerell, 1927; Lusitanomytacris Teixeira,<br />

1939; Metachorus Handlirsch t 1906; Oxynoblatta<br />

Handlirsch, 1906; Phylomylacris Pruvost,<br />

1914; Sooblatta Handlirsch, 1906; Soomylacris<br />

Handlirsch, 1906; Cardioblatta Handlirsch, 1906,<br />

Stephanomylacris Teixeira, 1949; Trilophomylacris<br />

Pruvost, 1919.<br />

ПОДСЕМЕЙСТВО IDIOMYLACRINAE HANDLIRSCH,<br />

1906<br />

, [nom. transl. Becker-Migdisova, hie<br />

(ex Idiomylacridae Handlirsch, 1906)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; SC делит<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тых <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong><br />

я<strong>с</strong>н<strong>а</strong>я, <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые, их<br />

<strong>в</strong>ет<strong>в</strong>и п<strong>о</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong> <strong>в</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>о</strong>дн<strong>а</strong> <strong>в</strong> другую. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 15 мм (ри<strong>с</strong>.<br />

208). Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и. Один р<strong>о</strong>д —<br />

Idiomylacris Handlirsch, 1906.<br />

7 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 97<br />

http://jurassic.ru/<br />

ПОДСЕМЕЙСТВО MYLACRINAE SELLARDS, 1904<br />

[nom. transl. Meunier, 1912<br />

(ex Mylacrinariae Sellards, 1904)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е,<br />

<strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>е, треуг<strong>о</strong>льн<strong>о</strong>е или <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля <strong>с</strong>имметричны и<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине; <strong>в</strong>ет<strong>в</strong>и SC <strong>в</strong>ых<strong>о</strong>дят из <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

луче<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, передние ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м;<br />

М или СиА <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> ' я<strong>с</strong>н<strong>а</strong>я, дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

14—40 мм. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong> пермь. Три<br />

трибы: Mylacrini, Pseudomylacrini, Neomylacrini.<br />

ТРИБА MYLACRINI PRUVOST, 1919<br />

[nom. transl. Becker-Migdisova, hie<br />

(ex Mylacrides Pruvost, 1919)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>-треуг<strong>о</strong>льн<strong>о</strong>е, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>а</strong><strong>с</strong>имметричн<strong>о</strong>е, <strong>с</strong> т<strong>о</strong>н<strong>к</strong>им <strong>а</strong>рхеди<strong>к</strong>тием; R мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

М и СиА р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, <strong>с</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; <strong>в</strong>ет<strong>в</strong>и СиА з<strong>а</strong>ним<strong>а</strong>ют<br />

0,3 длины з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

14—40 мм (ри<strong>с</strong>. 209). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong> <strong>с</strong>рез<strong>а</strong>нн<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong> п<strong>а</strong>рэн<strong>о</strong>т<strong>а</strong>льными р<strong>а</strong><strong>с</strong>ширениями.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я, <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющ<strong>а</strong>я.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong>. пермь: <strong>с</strong>р. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и; <strong>в</strong>. пермь Ю. Р<strong>о</strong>дезии. 17 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Вне СССР: Mylacris Scudder, 1886; Actinomylacris<br />

Handlirsch, 1906; Exochomylacris<br />

Handlirsch, 1906; Symplicius Handlirsch, 1920;<br />

Anomomylacris Handlirsch, 1906; Chalepomylacris<br />

Handlirsch, 1906; Amblymylacris Handlirsch,<br />

1906; Platymylacris Handlirsch, 1911;<br />

Ptilomylacris Cockerell, 1918; Aphelomylacris<br />

Handlirsch, 1906; Paromylacris Scudder, 1885;<br />

Promylacris Scudder, 1885; Orthomylacris<br />

Handlirsch, 1906; Stenomylacris Handlirsch.<br />

1906; Lithomylacris Scudder, 1879; Brachymylacris<br />

Handlirsch, 1906: Rhodesiomylacris<br />

Zeuner, 1955; Goniomylacris Handlirsch,<br />

1906; Necymylacris Scudder, 1879; Necymylacrodes<br />

Strand, 1942; Neosimplicius Carpenter,<br />

1934; Phthinomylacris Handlirsch, 1906.<br />

ТРИБА PSEUDOMYLACRINI<br />

HANDLIRSCH, 1906<br />

[nom. transl. Becker-Migdisova, hie<br />

(ex Pseudomylacridae Handlirsch, 1906)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>-<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>с</strong> ред<strong>к</strong>ими<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; М х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

СиА <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями, з<strong>а</strong>ним<strong>а</strong>ющими<br />

0,25 длины з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>а</strong>н<strong>а</strong>льные

жил<strong>к</strong>и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 6,6 мм (ри<strong>с</strong>. 210). В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Pseudomylacris Schlechtendal,<br />

1879.<br />

ТРИБА NEOMYLACRINI HANDLIRSCH,<br />

1906<br />

[nom. transl. Becker-Migdisova, hie<br />

(ex Neomylacridae Handlirsch, 1906)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>-<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; М <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>:<br />

<strong>в</strong>ет<strong>в</strong>и CuA з<strong>а</strong>ним<strong>а</strong>ют <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 0,3 длины з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; ч<strong>а</strong><strong>с</strong>ть <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зную <strong>а</strong>н<strong>а</strong>льную б<strong>о</strong>р<strong>о</strong>зду. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

16—22 мм (ри<strong>с</strong>. 211). Ср. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н.<br />

B. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и.<br />

Один р<strong>о</strong>д — Neomylacrls Handlirsch, 1906.<br />

ПОДСЕМЕЙСТВО PTERIDOMYLACRINAE<br />

HANDLIRSCH, 1906<br />

[nom. transl. Becker-Migdisova, hie<br />

(ex Pteridomylacridae Handlirsch, 1906)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, «<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е; <strong>в</strong>ет<strong>в</strong>и SC прямые и<br />

длинные, <strong>в</strong>ых<strong>о</strong>дят из <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния; R прям<strong>а</strong>я,<br />

гребенч<strong>а</strong>т<strong>а</strong>я; М прям<strong>а</strong>я, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, лишь <strong>с</strong> неб<strong>о</strong>льшим<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; <strong>в</strong>ет<strong>в</strong>и CuA<br />

длинные, <strong>в</strong>ых<strong>о</strong>дят из <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> не <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>, CuP прям<strong>а</strong>я, п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ям CuA и <strong>а</strong>н<strong>а</strong>льным жил<strong>к</strong><strong>а</strong>м, п<strong>о</strong><strong>с</strong>ледние<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 18 мм (ри<strong>с</strong>. 212). Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

C. Амери<strong>к</strong>и. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Pteridomylacrls Handlirsch,<br />

1906 и Sphenomylacris Handlirsch, 1906.<br />

СЕМЕЙСТВО NEORTHROBLATTINIDAE<br />

HANDLIRSCH, 1906<br />

(Dictyomylacridae Handlirsch, 1906)<br />

Н<strong>а</strong>д<strong>к</strong>рылье и з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> ред<strong>к</strong>ими, я<strong>с</strong>ными<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Н<strong>а</strong>д<strong>к</strong>рылье без<br />

рез<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>ширения <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления<br />

<strong>к</strong> груди р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> ближе <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

длинн<strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC гребенч<strong>а</strong>т<strong>а</strong>я; М х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; CuA <strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м длинных<br />

<strong>в</strong>ет<strong>в</strong>ей; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> я<strong>с</strong>н<strong>а</strong>я, дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я;<br />

<strong>в</strong>ет<strong>в</strong>и Ai <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> <strong>а</strong>н<strong>а</strong>льную б<strong>о</strong>р<strong>о</strong>зду, <strong>в</strong>ет<strong>в</strong>и<br />

А 2 — <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья. В з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть не<strong>в</strong>ели<strong>к</strong><strong>а</strong>, не<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, <strong>а</strong> п<strong>о</strong>дгиб<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>д<br />

<strong>к</strong>рыл<strong>о</strong>. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — н. пермь. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Neorthroblattininae и Dictyomylacrinae.<br />

98<br />

http://jurassic.ru/<br />

ПОДСЕМЕЙСТВО NEORTHROBLATTININAE<br />

HANDLIRSCH, 1906<br />

[nom. transl. Becker-Migdisova, hie<br />

(ex Neorthroblattinidae Handlirsch, 1906)]<br />

В н<strong>а</strong>д<strong>к</strong>рыльях п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

ред<strong>к</strong><strong>о</strong>, р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>яния между ними ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

б<strong>о</strong>льше длины п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле умеренн<strong>о</strong>й ширины; R и М рез<strong>к</strong><strong>о</strong> <strong>о</strong>тделены;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е, прям<strong>о</strong>е,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> <strong>в</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 8—9 мм (ри<strong>с</strong>. 213 и 214). В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

— н. пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Mylacridium Schlechtendal,<br />

1879 и Neorthroblattina Scudder, 1885.<br />

ПОДСЕМЕЙСТВО DICTYOMYLACRINAE<br />

HANDLIRSCH, 1906<br />

[nom. transl. Becker-Migdisova hie. (ex Dictyomylacridae<br />

Handlirsch, 1906)]<br />

В н<strong>а</strong>д<strong>к</strong>рыльях <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближены,<br />

п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены ч<strong>а</strong><strong>с</strong>т<strong>о</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии шир<strong>о</strong><strong>к</strong><strong>о</strong>е. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 16—20 мм (ри<strong>с</strong>. 215). В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и. Один р<strong>о</strong>д — Dictyomylacris<br />

Brongniart, 1893.<br />

NEORTHROBLATTINIDAE INCERTAE SEDIS<br />

Limmatoblatta Handlirsch, 1904. Тип<br />

р<strong>о</strong>д<strong>а</strong> — L. permensis Handlirsch, 1904; <strong>в</strong>. пермь,<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Фр<strong>а</strong>гмент н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> я<strong>с</strong>ными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми, длин<strong>а</strong> фр<strong>а</strong>гмент<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 25 мм<br />

(ри<strong>с</strong>. 216). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Ideloblatta М. Zalessky, 1929. Тип р<strong>о</strong>д<strong>а</strong>—<br />

/. rossica М. Zalessky, 1929; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Фр<strong>а</strong>гмент б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины н<strong>а</strong>д<strong>к</strong>рылья.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> м<strong>а</strong>л<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong> CuA (три)<br />

и п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти CuA и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля. Длин<strong>а</strong> фр<strong>а</strong>гмент<strong>а</strong> 12 мм (ри<strong>с</strong>. 217).<br />

Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО POROBLATTINIDAE<br />

HANDLIRSCH, 1906<br />

Ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong> груди<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> ближе <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, з<strong>а</strong>ним<strong>а</strong>ет не б<strong>о</strong>лее<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины н<strong>а</strong>д<strong>к</strong>рылья; SC <strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

пр<strong>о</strong><strong>с</strong>тых п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> р<strong>а</strong>ди<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong> <strong>о</strong>бширнее <strong>с</strong>и<strong>с</strong>темы меди<strong>а</strong>льных,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong>х<strong>о</strong>дит <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины н<strong>а</strong>д<strong>к</strong>рылья,<br />

<strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; М х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует <strong>с</strong>рединный треуг<strong>о</strong>льни<strong>к</strong>;<br />

CuA <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, з<strong>а</strong>ним<strong>а</strong>ет лишь 7з—'/<strong>в</strong><br />

ч<strong>а</strong><strong>с</strong>ть з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я, <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями,<br />

перпенди<strong>к</strong>улярными ему; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>

123. Neorthroblattina albolineata Scudder; н<strong>а</strong>д<strong>к</strong>рыье, x 6,8, н.<br />

Пермь, С. Амери<strong>к</strong><strong>а</strong> (Scudder, 1895). 214. Mylacridium germari<br />

Schlechtendal; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, x 6,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

Handlirsch, 1906). 215. Dictyomylacris poraulti Brongniart;<br />

<strong>в</strong>нешний <strong>в</strong>ид, x 2,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Фр<strong>а</strong>нция (Brongniart, 1893).<br />

я<strong>с</strong>н<strong>а</strong>я, <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong>. три<strong>а</strong><strong>с</strong>. В<strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Ishanablattina Becker-Migdisova,<br />

gen. nov. Тип р<strong>о</strong>д<strong>а</strong> — Drepanoblattina fodinensis<br />

Becker-Migdisova, 1961; <strong>в</strong>. <strong>к</strong><strong>а</strong>р бен, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>в</strong>ерхнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, П<strong>о</strong>ры<strong>в</strong><strong>а</strong>й<strong>с</strong><strong>к</strong>ий<br />

рудни<strong>к</strong>). Передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong><br />

<strong>в</strong>ершине рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут, ближе <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й; SC <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я<br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья,<br />

<strong>с</strong> ше<strong>с</strong>тью-<strong>с</strong>емью <strong>к</strong><strong>о</strong><strong>с</strong>ыми п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>в</strong>ет<strong>в</strong>ями; R з<strong>а</strong>х<strong>о</strong>дит немн<strong>о</strong>г<strong>о</strong> д<strong>а</strong>льше <strong>с</strong>ередины<br />

н<strong>а</strong>д<strong>к</strong>рылья и з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю ди<strong>с</strong>т<strong>а</strong>льную<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я д<strong>о</strong> <strong>в</strong>ершины, гребенч<strong>а</strong>т<strong>а</strong>я;<br />

М з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю ди<strong>с</strong>т<strong>а</strong>льную п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину<br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я, делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; СиА <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я,<br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong><strong>о</strong> <strong>с</strong>ближенными <strong>в</strong>ет<strong>в</strong>ями;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>е; <strong>а</strong>рхеди<strong>к</strong>тий<br />

<strong>о</strong>чень <strong>с</strong>л<strong>а</strong>бый, б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> — <strong>с</strong>т<strong>о</strong>лбч<strong>а</strong>тый,<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> — <strong>в</strong> <strong>в</strong>иде ч<strong>а</strong><strong>с</strong>тых п<strong>о</strong>перечных. Дли­<br />

Ри<strong>с</strong>. 213—217. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Neorthroblattinidae<br />

216. Limmatoblatta permensis Handlirsch; фр<strong>а</strong>гмент н<strong>а</strong>д<strong>к</strong>рылья,<br />

х 3,0, <strong>в</strong>. пермь, Приур<strong>а</strong>лье (Handlirsch, 1904). 217. Ideloblatta<br />

rossica М. Zalessky; н<strong>а</strong>д<strong>к</strong>рылье, х 2,6, <strong>в</strong>. пермь, Приур<strong>а</strong>лье<br />

(М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1929)<br />

н<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 24 мм (ри<strong>с</strong>. 218; т<strong>а</strong>бл.<br />

IV, фиг. 4). Один <strong>в</strong>ид. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Amekeoblatta G. Zalessky, 1933. Тип р<strong>о</strong>д<strong>а</strong><br />

— A. elegans G. Zalessky, 1933; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е; SC длинн<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья, гребенч<strong>а</strong>т<strong>а</strong>я,<br />

<strong>с</strong> ред<strong>к</strong>ими, нея<strong>с</strong>ными <strong>в</strong>ет<strong>в</strong>ями; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле р<strong>а</strong><strong>с</strong>ширен<strong>о</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти;<br />

R <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти н<strong>а</strong>д<strong>к</strong>рылья делит<strong>с</strong>я н<strong>а</strong><br />

д<strong>в</strong><strong>а</strong> длинных <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья<br />

рез<strong>к</strong><strong>о</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> изгиб<strong>а</strong>ют<strong>с</strong>я и з<strong>а</strong>х<strong>о</strong>дят<br />

<strong>в</strong> з<strong>а</strong>днюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину н<strong>а</strong>д<strong>к</strong>рылья; М и СиА<br />

<strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я c'R; СиА бедн<strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>ями, не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

8,5 мм (ри<strong>с</strong>. 219). Один <strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

09 7*<br />

http://jurassic.ru/

R \<br />

Ри<strong>с</strong>. 218—221. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Poroblattinidae, Blattidae,<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Protereminae, Diechoblattininae<br />

218 Ishanoblattina fodinensis (Becker-Migdisova!; н<strong>а</strong>д<strong>к</strong>рылье,<br />

x 2 5 и <strong>а</strong>рхеди<strong>к</strong>тий, x 15,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 219. Amekeoblatta elegans G. Zalessky;<br />

н<strong>а</strong>д<strong>к</strong>рылье, x 7,6, н. пермь, Ю. Приур<strong>а</strong>лье (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1933) 220. Proterema rarinervis Goppert; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 3,5<br />

н. пермь, Герм<strong>а</strong>ния (Goppert, 1865). 221. Kokandoblattina analis<br />

Martynov, н<strong>а</strong>д<strong>к</strong>рылье, x 7,5, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

Вне СССР: Poroblattina Scudder, 1885;<br />

Autoblattina Schlechtendal, 1879; Premnobtatta<br />

Pruvost, 1919; Kebaona Handlirsch, 1906:<br />

Drepanoblattina Schlechtendal, 1879.<br />

СЕМЕЙСТВО BLATTIDAE STEPHENS, 1829<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые или п<strong>о</strong>чти переп<strong>о</strong>нч<strong>а</strong>тые<br />

<strong>с</strong> я<strong>в</strong><strong>с</strong>т<strong>в</strong>енным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong> груди р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong><br />

ближе <strong>к</strong> переднему «р<strong>а</strong>ю; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е<br />

— меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

SC <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, реже длинн<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями; R прям<strong>а</strong>я, <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

з<strong>а</strong>ним<strong>а</strong>ет переднюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину н<strong>а</strong>д<strong>к</strong>рылья и д<strong>а</strong>ет<br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю ряд п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей;<br />

М <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние ее бы<strong>в</strong><strong>а</strong>ет<br />

<strong>с</strong>литым <strong>с</strong> R или CuA. Ан<strong>а</strong>льные жил<strong>к</strong>и<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> <strong>а</strong>н<strong>а</strong>льную б<strong>о</strong>р<strong>о</strong>зду. З<strong>а</strong>дние <strong>к</strong>рылья<br />

<strong>с</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью, <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ющей<strong>с</strong>я<br />

<strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. Бедр<strong>а</strong> <strong>в</strong><strong>о</strong><strong>о</strong>ружены шип<strong>а</strong>ми<br />

или л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми между <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Н<strong>а</strong>лични<strong>к</strong> без я<strong>в</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделения н<strong>а</strong><br />

переднюю и з<strong>а</strong>днюю ч<strong>а</strong><strong>с</strong>ти. Ст<strong>в</strong><strong>о</strong>р<strong>к</strong>и яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong><br />

<strong>с</strong><strong>к</strong>рыты и у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены. Цер<strong>к</strong>и я<strong>с</strong>н<strong>о</strong> <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нные.<br />

Н. пермь — ныне. 12 п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>,<br />

из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии де<strong>в</strong>ять: Protereminae,<br />

Diechoblattininae, Mesoblattininae,<br />

Blattinae, Pseudomopinae, Epilamprinae, Ectobiinae,<br />

Anaplectinae, Nyctiborinae.<br />

ПОДСЕМЕЙСТВО PROTEREMINAE<br />

HANDLIRSCH, 1906<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>а</strong>рхеди<strong>к</strong>тием и <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им,<br />

треуг<strong>о</strong>льным <strong>а</strong>пи<strong>к</strong><strong>а</strong>льным п<strong>о</strong>лем; SC прям<strong>а</strong>я,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; R <strong>с</strong> д<strong>в</strong>умя длинными <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми,<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями у <strong>в</strong>ершины; М <strong>с</strong> д<strong>в</strong>умя длинными<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми; CuA <strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их,<br />

неп<strong>о</strong>лных <strong>в</strong>ет<strong>в</strong>ей; <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть не <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь,<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>л<strong>а</strong><strong>с</strong>ь <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>.<br />

Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 15 мм (ри<strong>с</strong>.<br />

220). Н. пермь 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Proterema<br />

Handlirsch, 1906.<br />

ПОДСЕМЕЙСТВО DIECHOBLATTININAE<br />

HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, <strong>о</strong><strong>к</strong>руглые или <strong>с</strong>лег<strong>к</strong><strong>а</strong><br />

<strong>с</strong>уженные у <strong>в</strong>ершины; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е—менее<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины н<strong>а</strong>д<strong>к</strong>рылья; SC<br />

Ри<strong>с</strong>. 222—235. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Blattidae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesoblattininae<br />

222. Sogdobtatta robusta Martynov; н<strong>а</strong>д<strong>к</strong>рылье,- x 3,4, н. юр<strong>а</strong>,<br />

Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 223. Sogdob/atta sp.; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

<strong>к</strong> 3,7, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 224. Taublatta klsylkiensis<br />

Martynov; н<strong>а</strong>д<strong>к</strong>рылье, x 3,8, н. юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1937). 225. Taublatta sp.; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,5. н. юр<strong>а</strong>, Ср. Азия<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 226. Rhipidoblattlna angustata Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 5,0, н. юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 227. Mesoblattlna<br />

multivenosa Martynov, н<strong>а</strong>д<strong>к</strong>рылье, x 5,1, н. юр<strong>а</strong>. Ср. Азия<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 228. Samarob/attula subacuta Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 5,1, н. юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 229. Santaroblattula<br />

sp., з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,0, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

100<br />

http://jurassic.ru/<br />

1937) 230. Mesoblattula shurabica (Martynov, 1937); н<strong>а</strong>д<strong>к</strong>рылье,<br />

х5 6 н юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 231. Euryblattula<br />

soarsa Martynov; н<strong>а</strong>д<strong>к</strong>рылье, х 1,8, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><br />

1937). 232. Samaroblatta tlllyardl Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х34 и юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 233. Blattula turamca<br />

Martynov; н<strong>а</strong>д<strong>к</strong>рылье х 8,5, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1937). 234. Blattula sp.; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> х 6,9, н. юр<strong>а</strong>, Ср. Азия<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 235. Ophismoblatta sibirica Brauer, Redtenbacher,<br />

Ganglbauer; н<strong>а</strong>д<strong>к</strong>рылье, x 3,0. юр<strong>а</strong>. Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(Brauer, Redtenbacher, Ganglbauer, 1889).

http://jurassic.ru/

<strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>в</strong>ет<strong>в</strong>ями; R мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>в</strong>ет<strong>в</strong>ями, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong>х<strong>о</strong>дит <strong>в</strong> з<strong>а</strong>днюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину<br />

н<strong>а</strong>д<strong>к</strong>рылья; М <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> CuA, п<strong>о</strong><strong>с</strong>ледняя б<strong>о</strong>г<strong>а</strong>т<strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>ями, з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю з<strong>а</strong>днюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину<br />

н<strong>а</strong>д<strong>к</strong>рылья, ин<strong>о</strong>гд<strong>а</strong> М <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет переднюю<br />

<strong>в</strong>ет<strong>в</strong>ь, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющую<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> R; ч<strong>а</strong><strong>с</strong>ть<br />

<strong>в</strong>ет<strong>в</strong>ей <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>а</strong>н<strong>а</strong>льную<br />

б<strong>о</strong>р<strong>о</strong>зду. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>. Н. пермь —<br />

мел. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Kokandoblattina Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К- analis Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле р<strong>а</strong><strong>в</strong>н<strong>о</strong> п<strong>о</strong> длине <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>му; SC д<strong>а</strong>ет <strong>о</strong>переди<br />

<strong>о</strong>т четырех д<strong>о</strong> пяти <strong>в</strong>ет<strong>в</strong>ей; R пр<strong>о</strong>х<strong>о</strong>дит <strong>в</strong><br />

передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине н<strong>а</strong>д<strong>к</strong>рылья, прям<strong>а</strong>я, <strong>о</strong>бр<strong>а</strong>зует<br />

де<strong>в</strong>ять <strong>в</strong>ет<strong>в</strong>ей, ч<strong>а</strong><strong>с</strong>тью делящих<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и;<br />

М <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й жил<strong>к</strong>и, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющей<strong>с</strong>я<br />

<strong>с</strong> R; CuA прям<strong>а</strong>я, пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong><strong>с</strong>редине<br />

н<strong>а</strong>д<strong>к</strong>рылья, з<strong>а</strong>ним<strong>а</strong>я ег<strong>о</strong> з<strong>а</strong>днюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину,<br />

б<strong>о</strong>г<strong>а</strong>т<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>я, Ai и ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей А 2 <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют<br />

<strong>в</strong> рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнутую <strong>а</strong>н<strong>а</strong>льную б<strong>о</strong>р<strong>о</strong>зду. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 8 мм (ри<strong>с</strong>. 221). Один <strong>в</strong>ид. Н. юр<strong>а</strong><br />

Ср. Азии.<br />

Вне СССР: Brephoblatta Handlirsch, 1906;<br />

Nepioblatta Handlirsch, 1906; Diechoblattina<br />

Scudder, 1886; Dipluroblattina Scudder, 1886.<br />

ПОДСЕМЕЙСТВО MESOBLATTININAE<br />

HANDLIRSCH, 1908<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ытянуты <strong>в</strong> длину; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е — менее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

SC <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>т д<strong>в</strong>ух д<strong>о</strong> четырех<br />

<strong>в</strong>ет<strong>в</strong>ей, <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бр<strong>а</strong>зует <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е<br />

<strong>о</strong>т жил<strong>о</strong><strong>к</strong> плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле; R <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

з<strong>а</strong>ним<strong>а</strong>ет переднюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину н<strong>а</strong>д<strong>к</strong>рылья<br />

и д<strong>а</strong>ет <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю мн<strong>о</strong>г<strong>о</strong> п<strong>а</strong>р<strong>а</strong>ллельных<br />

жил<strong>о</strong><strong>к</strong>; М и CuA не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е н<strong>о</strong>ле <strong>о</strong>тделен<strong>о</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й,<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья.<br />

Ан<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong><br />

неиз<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>. Бедр<strong>а</strong> <strong>с</strong>низу <strong>в</strong><strong>о</strong><strong>о</strong>ружены шип<strong>а</strong>ми.<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong>. юр<strong>а</strong>. 49 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Sogdoblatta Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. robusta Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Н<strong>а</strong>д<strong>к</strong>рылье удлиненн<strong>о</strong>е,<br />

<strong>с</strong> иятер<strong>к</strong><strong>а</strong>лярными жил<strong>к</strong><strong>а</strong>ми и п<strong>о</strong>перечными<br />

м<strong>о</strong>рщин<strong>к</strong><strong>а</strong>ми; SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> или немн<strong>о</strong>г<strong>о</strong><br />

длиннее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, д<strong>а</strong>ет <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти три-четыре <strong>в</strong>ет<strong>в</strong>и; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле неб<strong>о</strong>льш<strong>о</strong>е;<br />

R <strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; CuA б<strong>о</strong>г<strong>а</strong>че, чем<br />

М, и делит<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ее; <strong>в</strong>ет<strong>в</strong>и М и CuA<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>б­<br />

102<br />

http://jurassic.ru/<br />

л<strong>а</strong><strong>с</strong>ть б<strong>о</strong>льш<strong>а</strong>я, <strong>в</strong>ытянут<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

20—26 мм (ри<strong>с</strong>. 222). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> длинн<strong>о</strong>й<br />

SC; PR и R делят<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и; R и М делят<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; М <strong>с</strong> д<strong>в</strong>умя <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми;<br />

CuA делит<strong>с</strong>я р<strong>а</strong>ньше, мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong> (ри<strong>с</strong>. 223).<br />

Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. юр<strong>а</strong> Ср. Азии.<br />

Taublatta Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Т. curvata Martynov, 1937; н. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong><br />

интер<strong>к</strong><strong>а</strong>лярными и п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

SC длиннее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, д<strong>а</strong>ет д<strong>в</strong>е-три<br />

ди<strong>с</strong>т<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле р<strong>а</strong><strong>в</strong>н<strong>о</strong> или<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти; R ди<strong>с</strong>т<strong>а</strong>льнее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, <strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д и н<strong>а</strong>ружу;<br />

CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>, делит<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее М и<br />

<strong>в</strong>ершины <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

15—20 мм (ри<strong>с</strong>. 224). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й<br />

SC и гребенч<strong>а</strong>тым ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> PR; М <strong>с</strong> д<strong>в</strong>умя<br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми; R, М и CuA делят<strong>с</strong>я<br />

н<strong>а</strong> р<strong>а</strong>зных ур<strong>о</strong><strong>в</strong>нях, CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong> (ри<strong>с</strong>.<br />

225). Три <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ср. Азии.<br />

Rhipidoblattina Handlirsch, 1908. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Rh. geinitzi Scudder, 1886; н. юр<strong>а</strong>, Англия.<br />

Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е, <strong>с</strong> я<strong>с</strong>ными интер<strong>к</strong><strong>а</strong>лярными<br />

и п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; SC<br />

<strong>с</strong> <strong>о</strong>дн<strong>о</strong>й-д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями или пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; плече<strong>в</strong><strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong> или б<strong>о</strong>льше длины <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; R <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>, ед<strong>в</strong><strong>а</strong><br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>с</strong>ередины ширины н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong><br />

б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>к</strong><strong>о</strong><strong>с</strong>ых <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных<br />

<strong>в</strong>перед; передний <strong>с</strong>т<strong>в</strong><strong>о</strong>л М пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й; CuA делит<strong>с</strong>я<br />

б<strong>а</strong>з<strong>а</strong>льнее, чем М, д<strong>а</strong>ет д<strong>в</strong><strong>а</strong> <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тых <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>;<br />

<strong>в</strong>ет<strong>в</strong>и М и CuA идут <strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я и<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют у <strong>в</strong>ершины; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть удлиненн<strong>а</strong>я.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 8,5—13 мм (ри<strong>с</strong><br />

226). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Юр<strong>а</strong> Ср. Азии и Англии.<br />

Mesoblattina Geinitz, 1880. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Blattina protypa Geinitz, 1880; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(<strong>в</strong>. лей<strong>а</strong><strong>с</strong>, Герм<strong>а</strong>ния). Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е, <strong>с</strong><br />

п<strong>а</strong>р<strong>а</strong>ллельными <strong>к</strong>р<strong>а</strong>ями; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; плече<strong>в</strong><strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong> или длиннее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; R прям<strong>а</strong>я или <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>бым изгиб<strong>о</strong>м, не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

<strong>с</strong>ередины ширины; передний <strong>с</strong>т<strong>в</strong><strong>о</strong>л М<br />

пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, з<strong>а</strong>дний — гребенч<strong>а</strong>тый, без р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>лений<br />

или <strong>с</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong><strong>к</strong>руглен<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

6,0—11,5 мм (ри<strong>с</strong>. 227). О<strong>к</strong><strong>о</strong>л<strong>о</strong> 30 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Юр<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и Ср. Азии.<br />

Samaroblattula Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. subacuta Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>с</strong>редней <strong>в</strong>еличины, переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е, <strong>с</strong> интер<strong>к</strong><strong>а</strong>лярными<br />

жил<strong>к</strong><strong>а</strong>ми и п<strong>о</strong>перечными м<strong>о</strong>рщин<strong>к</strong><strong>а</strong>ми<br />

н<strong>а</strong> гребнях; SC длинн<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями;<br />

R д<strong>а</strong>ет пять пр<strong>о</strong><strong>с</strong>тых <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных <strong>в</strong>ет<strong>в</strong>ей;

CuA делит<strong>с</strong>я р<strong>а</strong>н<strong>о</strong>, б<strong>а</strong>з<strong>а</strong>льнее М, и <strong>о</strong>т<strong>с</strong>ыл<strong>а</strong>ет<br />

ше<strong>с</strong>ть-<strong>с</strong>емь <strong>в</strong>ет<strong>в</strong>ей, из<strong>о</strong>гнутых <strong>к</strong>н<strong>а</strong>ружи; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть не<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>я, удлиненн<strong>а</strong>я. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 10—12 мм (ри<strong>с</strong>. 228). В з<strong>а</strong>днем<br />

<strong>к</strong>рыле SC дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>, <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м,<br />

д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; R, М и<br />

CuA делят<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; PR <strong>с</strong>ильн<strong>а</strong>я,<br />

гребенч<strong>а</strong>т<strong>а</strong>я, <strong>с</strong> п<strong>о</strong>чти <strong>в</strong>ерти<strong>к</strong><strong>а</strong>льными <strong>в</strong>ет<strong>в</strong>ями;<br />

М <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong> (ри<strong>с</strong>.<br />

229). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ср. Азии.<br />

Mesoblattula Н a n d 1 i s <strong>с</strong> h, 1908. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Mesoblattina dobbertinensis Geinitz, 1887;<br />

н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (<strong>в</strong>. лей<strong>а</strong><strong>с</strong>, Герм<strong>а</strong>ния). Н<strong>а</strong>д<strong>к</strong>рылье<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>к</strong> <strong>к</strong><strong>о</strong>нцу <strong>с</strong>леп<strong>к</strong><strong>а</strong><br />

<strong>с</strong>уженн<strong>о</strong>е, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> нея<strong>с</strong>ными интер<strong>к</strong><strong>а</strong>лярными<br />

жил<strong>к</strong><strong>а</strong>ми и п<strong>о</strong>перечными м<strong>о</strong>рщин<strong>а</strong>ми; SC прям<strong>а</strong>я;<br />

плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е; R <strong>с</strong><br />

рез<strong>к</strong>им S-<strong>о</strong>бр<strong>а</strong>зным изгиб<strong>о</strong>м, б<strong>о</strong>г<strong>а</strong>т<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ями;<br />

М и CuA <strong>с</strong> д<strong>в</strong>умя <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми и шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными<br />

<strong>в</strong>ет<strong>в</strong>ями; CuA бедн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ями, <strong>с</strong> рез<strong>к</strong>им<br />

изгиб<strong>о</strong>м, п<strong>а</strong>р<strong>а</strong>ллельным CuP <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле б<strong>о</strong>льш<strong>о</strong>е, шир<strong>о</strong><strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

6—14 мм (ри<strong>с</strong>. 230). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н.<br />

юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и Ср. Азии.<br />

Euryblattula Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. sparsa Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

SC д<strong>а</strong>ет т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дну <strong>в</strong>ет<strong>в</strong>ь; плече<strong>в</strong><strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й; R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, из<strong>о</strong>гнут<strong>а</strong>я,<br />

д<strong>а</strong>ет ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей; CuA делит<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее М, <strong>в</strong>ет<strong>в</strong>ление <strong>о</strong>беих бедн<strong>о</strong>е, <strong>в</strong>ет<strong>в</strong>и<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены шир<strong>о</strong><strong>к</strong><strong>о</strong>; CuA <strong>с</strong> рез<strong>к</strong>им изгиб<strong>о</strong>м,<br />

п<strong>а</strong>р<strong>а</strong>ллельным CuP <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 17 мм<br />

(ри<strong>с</strong>. 231). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Samaroblatta Tillyard, 1919. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. reticulata Tillyard, 1919; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> интер<strong>к</strong><strong>а</strong>лярными и гу<strong>с</strong>т<strong>о</strong>й<br />

<strong>с</strong>етью п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> длине<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля, <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями;<br />

плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; R д<strong>а</strong>ет S-<strong>о</strong>бр<strong>а</strong>зный<br />

изгиб н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля, д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong><br />

<strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья или немн<strong>о</strong>г<strong>о</strong> д<strong>а</strong>лее; д<strong>в</strong><strong>а</strong><br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М делят<strong>с</strong>я мн<strong>о</strong>г<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong>, <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены<br />

<strong>в</strong>перед; CuA делит<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее, чем М,<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине рез<strong>к</strong><strong>о</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> ч<strong>а</strong><strong>с</strong>тыми<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

18 мм (ри<strong>с</strong>. 232). В. три<strong>а</strong><strong>с</strong> — н. юр<strong>а</strong>. Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

В. три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и н. юр<strong>а</strong> Ср. Азии.<br />

Blattula Handlirsch, 1908. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Mesoblattina dobbertinensis Geinitz, 1884; н.<br />

юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (<strong>в</strong>. лей<strong>а</strong><strong>с</strong>, Герм<strong>а</strong>ния). Н<strong>а</strong>д<strong>к</strong>рылье<br />

эллиптиче<strong>с</strong><strong>к</strong>и <strong>в</strong>ытянут<strong>о</strong>, <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>к</strong>р<strong>а</strong>ями; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, прям<strong>а</strong>я, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>в</strong>перед; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

R п<strong>о</strong>чти прям<strong>а</strong>я, пр<strong>о</strong>х<strong>о</strong>дит через <strong>с</strong>ередину,<br />

<strong>в</strong>ет<strong>в</strong>и R ред<strong>к</strong>ие, п<strong>а</strong>р<strong>а</strong>ллельные; М <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, передняя ее <strong>в</strong>ет<strong>в</strong>ь <strong>в</strong><strong>с</strong>егд<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями; CuA<br />

прям<strong>а</strong>я, гребенч<strong>а</strong>т<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле неб<strong>о</strong>льш<strong>о</strong>е,<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,0—7,0 мм<br />

(ри<strong>с</strong>. 233). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> прямыми SC, PR и<br />

М, <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я <strong>о</strong>чень ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>. CuA прям<strong>а</strong>я,<br />

гребенч<strong>а</strong>т<strong>а</strong>я (ри<strong>с</strong>. 234). О<strong>к</strong><strong>о</strong>л<strong>о</strong> 18 <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Юр<strong>а</strong> Ср. Азии и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Ophismoblatta Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong><br />

— О. sibirica Brauer, Redtenbacher, Ganglbauer,<br />

1889; н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-Б<strong>а</strong>лей). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> шир<strong>о</strong><strong>к</strong>им<br />

плече<strong>в</strong>ым п<strong>о</strong>лем; SC <strong>с</strong> четырьмя-пятью<br />

<strong>в</strong>ет<strong>в</strong>ями <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти, р<strong>а</strong><strong>в</strong>н<strong>а</strong> длине<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; R <strong>с</strong>л<strong>а</strong>б<strong>о</strong>дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>, из<strong>о</strong>гнут<strong>а</strong><br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и ее <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми;<br />

М <strong>с</strong> д<strong>в</strong>умя <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми и четырьмя <strong>в</strong>ет<strong>в</strong>ями;<br />

CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>, з<strong>а</strong>дний <strong>с</strong>т<strong>в</strong><strong>о</strong>л рез<strong>к</strong><strong>о</strong> изгиб<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong>д<strong>о</strong>ль CuP, передний <strong>с</strong>т<strong>в</strong><strong>о</strong>л <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными<br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>в</strong>ет<strong>в</strong>ями, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющими<br />

<strong>в</strong> <strong>в</strong>ершину; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле б<strong>о</strong>льш<strong>о</strong>е, н<strong>о</strong> не<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

жил<strong>к</strong>и <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

17—20 мм (ри<strong>с</strong>. 235). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Юр<strong>а</strong><br />

В. Сибири.<br />

Вне СССР: Acmeoblatta Handlirsch, 1906;<br />

Dichronoblatta Handlirsch, 1906; Nearob'atta<br />

Handlirsch, 1906; Haeberleoblatta Guthorl, 1934;<br />

Epheboblatta Handlirsch, 1906; Scutinoblattina<br />

Scudder, 1885; Austroblattula, Tillyard, 1919;<br />

Austromylacrites Tillyard, 1916; Notoblattites<br />

Tillyard, 1916; Triassoblatta Tillyard 1919; Rhaetiblattina<br />

Handlirsch, 1908; Hongaya Handlirsch,<br />

1908; Pedinoblatta Handlirsch, 1910; Actinoblattula<br />

Handlirsch, 1908; Liadoblattina<br />

Handlirsch, 1908; Mesoblattopsis Handlirsch,<br />

1908; Chiloblattula Handlirsch, 1939; Caloblattina<br />

Handlirsch, 1908; Ectinobtattula Handlirsch,<br />

1939; Eublattula Handlirsch, 1939; Metablattula<br />

Handlirsch, 1939; Pachyneuroblattina Handlirsch,<br />

1908; Parablattula Handlirsch, 1939;<br />

Paleoblattula Handlirsch, 1939; Schambeloblattina<br />

Handlirsch, 1908; Artitocoblatta<br />

Handlirsch, 1908; Blattidium Westwood, 1854;<br />

Ctenoblattina Scudder, 1886; Durdelstonia<br />

Handlirsch, 1908; Elisama Giebel, 1856;<br />

Malmoblattina Handlirsch, 1908; Nannoblattina<br />

Scudder, 1886; Rithma Giebel, 1856; Apistoblattuta<br />

Bode, 1953; Macroblattina Bode, 1953;<br />

Kulmbachiellon Kuhn, 1938; Thiiringoblatta<br />

Kuhn, 1939; Trirhabdoblattina Bode, 1953;<br />

Lithoblatta Handlirsch, 1908; Billia Handlirsch,<br />

1939; Palmoblattina Bode, 1953; Euglyptoblatta<br />

Bode, 1953; Strebloblattula Bode, 1953; Striatotegmen<br />

Haughton, 1924; Potyphleboblatta Bode,<br />

1953; Plyctoblattina Bode, 1953.<br />

103<br />

http://jurassic.ru/

Ри<strong>с</strong>. 236-237. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Blattidae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Blattinae, Pseudomopinae<br />

ПОДСЕМЕЙСТВО BLATTINAE STEPHENS, 1829<br />

[nom. transl. Handlirsch, 1925<br />

(ex Blattidae Stephens, 1829)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье пл<strong>о</strong>тн<strong>о</strong>е, п<strong>о</strong>лу<strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>е; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле умеренн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>;<br />

SC ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не имеет з<strong>а</strong>дних <strong>в</strong>ет<strong>в</strong>ей; <strong>в</strong>ет<strong>в</strong>и<br />

R <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>-р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ящие<strong>с</strong>я; М мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> СиА; СиА <strong>о</strong>бычн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>и ее <strong>к</strong><strong>о</strong><strong>с</strong>ые или п<strong>а</strong>р<strong>а</strong>ллельные з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю н<strong>а</strong>д<strong>к</strong>рылья; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющими <strong>в</strong> <strong>а</strong>н<strong>а</strong>льную б<strong>о</strong>р<strong>о</strong>зду.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> без д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля, <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я М и СиА. Темя г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет <strong>в</strong>переди передне<strong>с</strong>пин<strong>к</strong>и. Бедр<strong>а</strong> <strong>с</strong><br />

<strong>с</strong>ильными мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными шип<strong>а</strong>ми н<strong>а</strong> нижней<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не (ри<strong>с</strong>. 236, 237). Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>й<br />

ф<strong>а</strong>уне — Stantoniella Handlirsch, 1908<br />

из <strong>в</strong>. мел<strong>а</strong> С. Амери<strong>к</strong>и и <strong>о</strong>дин р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и из чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong> Афри<strong>к</strong>и. Мел — ныне.<br />

ПОДСЕМЕЙСТВО PSEUDOMOPINAE<br />

BURR, 1913<br />

[nom. transl. Bey-Bienko, 1950<br />

(ex Pseudomopidae Burr, 1913)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье и з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>а</strong>д<strong>к</strong>рылье нежн<strong>о</strong>е,<br />

104<br />

http://jurassic.ru/<br />

п<strong>о</strong>лу<strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>е, реже р<strong>о</strong>г<strong>о</strong><strong>в</strong><strong>о</strong>е; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, реже<br />

<strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; <strong>в</strong>ет<strong>в</strong>и R п<strong>а</strong>р<strong>а</strong>ллельные,<br />

пр<strong>о</strong><strong>с</strong>тые или <strong>о</strong>дн<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящие<strong>с</strong>я. М<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

СиА; е<strong>с</strong>ли СиА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М, т<strong>о</strong> <strong>в</strong>ет<strong>в</strong>и ее<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ые или дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, е<strong>с</strong>ли СиА <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я,<br />

т<strong>о</strong> <strong>в</strong>ет<strong>в</strong>и ее п<strong>а</strong>р<strong>а</strong>ллельны з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю. З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>тем<strong>о</strong>й М; СиА<br />

без <strong>к</strong><strong>о</strong><strong>с</strong>ых <strong>в</strong>ет<strong>в</strong>ей <strong>с</strong>з<strong>а</strong>ди или <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими п<strong>о</strong>лными<br />

и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими неп<strong>о</strong>лными <strong>в</strong>ет<strong>в</strong>ями; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть уз<strong>к</strong><strong>а</strong>я — меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины ширины<br />

г<strong>а</strong>ред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> нез<strong>а</strong><strong>к</strong>рытым<br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й теменем. Бедр<strong>а</strong> <strong>с</strong> <strong>с</strong>ильными<br />

немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными шип<strong>а</strong>ми н<strong>а</strong> нижней<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не или <strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идными<br />

шипи<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 238 и 239). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Одинн<strong>а</strong>дц<strong>а</strong>ть триб <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из них <strong>с</strong> третичн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени Symplocini, Supellini,<br />

Ischnopterini, Blattellini.<br />

ТРИБА SYMPLICINI REHN, 1951<br />

Н<strong>а</strong>д<strong>к</strong>рылье длинн<strong>о</strong>е и <strong>с</strong>леп<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е;<br />

SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, прям<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля;<br />

RS <strong>с</strong> ряд<strong>о</strong>м пр<strong>о</strong><strong>с</strong>тых н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ей;<br />

М <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, имеет три <strong>в</strong>ет<strong>в</strong>и; СиА делит<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и М и СиА п<strong>а</strong>р<strong>а</strong>ллельны<br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю н<strong>а</strong>д<strong>к</strong>рылья. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

б<strong>о</strong>льш<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,

Ри<strong>с</strong>. 238—242. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Blattidae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pseudomopinae<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>ем 1гитяТ'|°<strong>в</strong> е<br />

рй 5<br />

Ки^п W<br />

e r :<br />

^ n ° б и й В И <strong>с</strong> <strong>а</strong> м<br />

Ри<strong>с</strong>. 243—248. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Blattidae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Epilampririae<br />

243. Phlebonotus pallens Serville; <strong>с</strong><strong>о</strong><strong>в</strong>рем., М<strong>а</strong>л<strong>а</strong>й<strong>с</strong><strong>к</strong>ий <strong>а</strong>рхипел<strong>а</strong>г;<br />

А —з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>; Б — у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>о</strong>е з<strong>а</strong>! нее <strong>к</strong>"ыл<strong>о</strong> <strong>с</strong><strong>а</strong>мги,<br />

х 4,0. 244. Notolampra gibba Thunberg; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,5,<br />

Бр<strong>а</strong>зилия (Shelford, 1907). 245. Epilampra goliath SheHord;<br />

•<strong>о</strong>бщий <strong>в</strong>ид, х 0,75, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Б<strong>о</strong>рне<strong>о</strong> (Shelford, 1907). 246. Punch-<br />

ТРИБА ISCHNOPTERINI REHN, 1951<br />

Н<strong>а</strong>д<strong>к</strong>рылья удлиненные; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими<br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льными <strong>в</strong>ет<strong>в</strong>ями; RS мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>;<br />

М <strong>с</strong> четырьмя длинными пр<strong>о</strong><strong>с</strong>тыми<br />

<strong>в</strong>ет<strong>в</strong>ями; CuA <strong>с</strong> ше<strong>с</strong>тью <strong>в</strong>ет<strong>в</strong>ями, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>к</strong> з<strong>а</strong>дне<strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>му углу. В з<strong>а</strong>днем<br />

<strong>к</strong>рыле <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> умеренн<strong>о</strong>,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че пред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й; SC <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> Rr,<br />

RS п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> Ri, делит<strong>с</strong>я у <strong>в</strong>ершины; CuA<br />

<strong>с</strong> четырьмя пр<strong>о</strong><strong>с</strong>тыми <strong>а</strong>пи<strong>к</strong><strong>а</strong>льными <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющими <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ТРИБА BLATTELLINI REHN, 1951<br />

Н<strong>а</strong>д<strong>к</strong>рылья длинные; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, п<strong>о</strong>чти<br />

прям<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; RS <strong>с</strong> пр<strong>о</strong><strong>с</strong>ты­<br />

106<br />

http://jurassic.ru/<br />

248<br />

lora cubensis Saussure; н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x 3,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Центр.<br />

Амери<strong>к</strong><strong>а</strong> (Hebard, 1917). 247. Pycnoscelus surinamensis Linnaeus;<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и, хЗ,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем., тр<strong>о</strong>пи<strong>к</strong>и Ст<strong>а</strong>р<strong>о</strong>г<strong>о</strong> и Н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

С<strong>в</strong>ет<strong>а</strong> (Бей-Биен<strong>к</strong><strong>о</strong>, 1950). 248. Perispherini sp.; личин<strong>к</strong><strong>а</strong>, х 7,2.<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Shelford, 1910)<br />

ми, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ями; М и CuA<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я, д<strong>а</strong>ют ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей; ин<strong>о</strong>гд<strong>а</strong> CuA<br />

редуцирует<strong>с</strong>я д<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и. З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> уз<strong>к</strong><strong>о</strong>й пред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й и<br />

умеренн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тями;<br />

SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; Ri я<strong>с</strong>н<strong>а</strong>я, <strong>с</strong> <strong>в</strong>ершинными <strong>в</strong>ет<strong>в</strong>ями.<br />

RS д<strong>а</strong>ет <strong>в</strong>ет<strong>в</strong>и <strong>в</strong>перед и <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>; CuA п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

М, <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>а</strong>пи<strong>к</strong><strong>а</strong>льными<br />

<strong>в</strong>ет<strong>в</strong>ями. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

PSEUDOMOPINAE INCERTAE SEDIS<br />

Mesoblattina sinica Ping, 1928 из <strong>в</strong>. мел<strong>а</strong> Кит<strong>а</strong>я<br />

и Pseudophyllodromia Shelford, 1911 из <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong><br />

Афри<strong>к</strong>и.

ПОДСЕМЕЙСТВО EPILAMPRINAE SAUSSURE, 1864<br />

[nom. transl. Saussure, 1893<br />

(ex Epilampriens Saussure, 1864)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья и <strong>к</strong>рылья р<strong>а</strong>з<strong>в</strong>итые, у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые или<br />

р<strong>о</strong>г<strong>о</strong><strong>в</strong>ые, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> <strong>о</strong><strong>к</strong>ругленн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й; SC<br />

прям<strong>а</strong>я, <strong>о</strong>бычн<strong>о</strong> длиннее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями; <strong>в</strong>ет<strong>в</strong>и R <strong>с</strong>переди ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> SC, <strong>а</strong> у <strong>в</strong>ершины—<strong>с</strong> М. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми SC и М; <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

шире п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины ширины пред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й;<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных, неп<strong>о</strong>лных<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ых <strong>в</strong>ет<strong>в</strong>ей СиА <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> СиР. Бедр<strong>а</strong> <strong>с</strong>низу<br />

<strong>в</strong><strong>о</strong><strong>о</strong>ружены шип<strong>а</strong>ми или <strong>о</strong>дним <strong>в</strong>ершинным<br />

шип<strong>о</strong>м (ри<strong>с</strong>. 243—248). П<strong>а</strong>ле<strong>о</strong>ген — ныне. 12<br />

триб <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> третичных <strong>о</strong>тл<strong>о</strong>жениях<br />

Epilamprini, Panchlorini, Perisphaerini.<br />

ТРИБА EPILAMPRINI SAUSSURE, 1864<br />

[nom. transl. Rehn, 1951<br />

(ex Epilampriens Saussure, 1864)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>; SC <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>с</strong> <strong>в</strong>ершинными <strong>в</strong>ет<strong>в</strong>ями; R <strong>с</strong> длинными, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными<br />

<strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ями; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ближен<strong>а</strong><br />

<strong>с</strong> R; М и СиА мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>ты, их <strong>в</strong>ет<strong>в</strong>и<br />

п<strong>а</strong>р<strong>а</strong>ллельны з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью, р<strong>а</strong><strong>в</strong>н<strong>о</strong>й пред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й;<br />

SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й з<strong>а</strong> <strong>с</strong>ередин<strong>о</strong>й <strong>к</strong>рыл<strong>а</strong>;<br />

Ri <strong>с</strong> тремя-четырьмя <strong>в</strong>ершинными <strong>в</strong>ет<strong>в</strong>ями;<br />

RS <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ершинными<br />

<strong>в</strong>ет<strong>в</strong>ями; менее чем треть <strong>в</strong>ет<strong>в</strong>ей СиА н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong><br />

<strong>к</strong> <strong>в</strong>ершинн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю. Бедр<strong>а</strong> <strong>с</strong>низу <strong>в</strong><strong>о</strong><strong>о</strong>ружены<br />

шип<strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> п<strong>о</strong>душеч<strong>к</strong><strong>о</strong>й и при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>й.<br />

П<strong>о</strong>чти и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие ф<strong>о</strong>рмы, р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные<br />

<strong>в</strong><strong>с</strong>е<strong>с</strong><strong>в</strong>етн<strong>о</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Один <strong>в</strong>ымерший р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ТРИБА PANCHLORINI SAUSSURE, 1864<br />

[nom. transl. Rehn, 1951<br />

(ex Panchloriens Saussure, 1864)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ытянут<strong>о</strong>е, <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>к</strong>р<strong>а</strong>ями; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле не р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>; SC ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><br />

ряд<strong>о</strong>м з<strong>а</strong>дних <strong>в</strong>ет<strong>в</strong>ей; передние <strong>в</strong>ет<strong>в</strong>и R <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми;<br />

М. <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>ближен<strong>а</strong> <strong>с</strong> R; <strong>в</strong>ет<strong>в</strong>и СиА п<strong>а</strong>р<strong>а</strong>ллельны<br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю. В з<strong>а</strong>дних <strong>к</strong>рыльях пред<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> б<strong>о</strong>льше <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й;<br />

SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й р<strong>а</strong>нее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>; СиА имеет <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 12 <strong>в</strong>ет<strong>в</strong>ей,<br />

треть их <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й; <strong>в</strong>ершинный<br />

треуг<strong>о</strong>льни<strong>к</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует или <strong>в</strong>ыр<strong>а</strong>жен<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>. Бедр<strong>а</strong> <strong>с</strong>низу <strong>с</strong> <strong>о</strong>дним <strong>в</strong>ершинным шип<strong>о</strong>м,<br />

передний нижний <strong>к</strong>р<strong>а</strong>й передних бедер<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> имеет ряд <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>­<br />

ми. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Один <strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ТРИБА PERISPHAERINI SAUSSURE<br />

1870<br />

[nom. transl. Rehn, 1951<br />

(ex Perispherites Saussure, 1870)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е;<br />

плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, без<br />

з<strong>а</strong>дних <strong>в</strong>ет<strong>в</strong>ей; R <strong>с</strong> ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> делящими<strong>с</strong>я передними<br />

<strong>в</strong>ет<strong>в</strong>ями; М ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

СиА, <strong>с</strong> четырьмя <strong>в</strong>ет<strong>в</strong>ями; СиА мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>,<br />

делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, <strong>в</strong>ет<strong>в</strong>и из<strong>о</strong>гнуты. З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> я<strong>с</strong>н<strong>о</strong>й Ri, п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>й SC и <strong>с</strong>ильн<strong>о</strong>й<br />

RS, <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных жил<strong>о</strong><strong>к</strong>; СиА мнюг<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>,<br />

б<strong>о</strong>лее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины ее <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>о</strong><br />

<strong>к</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю. Бедр<strong>а</strong> <strong>с</strong>низу без<br />

шип<strong>о</strong><strong>в</strong>; <strong>в</strong>ершинный и <strong>к</strong><strong>о</strong>ленный шипы ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми между <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Цер<strong>к</strong>и ч<strong>а</strong>ете <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>с</strong> нея<strong>с</strong>н<strong>о</strong>й<br />

члени<strong>с</strong>т<strong>о</strong><strong>с</strong>тью. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Личин<strong>к</strong><strong>а</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО ECTOBIINAE SAUSSURE, 1864<br />

[nom. transl. Kirby, 1904<br />

(ex Ectobiens Saussure, 1864)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье умеренн<strong>о</strong> <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>е, <strong>с</strong>уженн<strong>о</strong>е<br />

<strong>к</strong> <strong>в</strong>ершине, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> пери<strong>с</strong>тым р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жением<br />

жил<strong>о</strong><strong>к</strong>; SC <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>му п<strong>о</strong>лю; R <strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ет<strong>с</strong>я д<strong>о</strong><br />

<strong>в</strong>ершины н<strong>а</strong>д<strong>к</strong>рылья; М и СиА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии,<br />

<strong>о</strong>бр<strong>а</strong>зуют <strong>с</strong>ближенный <strong>с</strong> R <strong>с</strong>т<strong>в</strong><strong>о</strong>л или<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> ним; е<strong>с</strong>ли М и СиА р<strong>а</strong>зделимы, т<strong>о</strong><br />

М из<strong>о</strong>гнут<strong>а</strong> и имеет три <strong>в</strong>ет<strong>в</strong>и и б<strong>о</strong>лее, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленные<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; не б<strong>о</strong>лее четырех жил<strong>о</strong><strong>к</strong> <strong>в</strong><br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> б<strong>о</strong>льшим <strong>в</strong>ершинным<br />

треуг<strong>о</strong>льни<strong>к</strong><strong>о</strong>м, п<strong>о</strong>дгиб<strong>а</strong>ющим<strong>с</strong>я <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

или <strong>с</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong><strong>о</strong>м ег<strong>о</strong>; R р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, пр<strong>о</strong>тяги<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

д<strong>о</strong> <strong>в</strong>ершинн<strong>о</strong>г<strong>о</strong> треуг<strong>о</strong>льни<strong>к</strong><strong>а</strong>, з<strong>а</strong>дние <strong>в</strong>ет<strong>в</strong>и<br />

ее <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; М прям<strong>а</strong>я или <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут<strong>а</strong>я;<br />

п<strong>о</strong>ле между RS и М р<strong>а</strong>зделен<strong>о</strong> п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми н<strong>а</strong> ряд ячее<strong>к</strong>; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я.<br />

З<strong>а</strong>дние бедр<strong>а</strong> <strong>с</strong> немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными шип<strong>а</strong>ми<br />

или без них. Р<strong>а</strong>змеры мел<strong>к</strong>ие или <strong>с</strong>редние<br />

(ри<strong>с</strong>. 249). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три трибы:<br />

Ectobiini, Plectopterini, Chorisoneurini; п<strong>о</strong><strong>с</strong>ледняя<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йден<strong>а</strong>.<br />

ТРИБА ECTOBI INI REDTENBACHER,<br />

1900<br />

Н<strong>а</strong>д<strong>к</strong>рылье л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идн<strong>о</strong>е; SC <strong>с</strong>игм<strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>;<br />

плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле умеренн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>; R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

ут<strong>о</strong>лщен<strong>а</strong>; М и СиА <strong>с</strong>ипм<strong>о</strong>ид<strong>а</strong>льны; М<br />

четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

107<br />

http://jurassic.ru/

Ри<strong>с</strong>. 249—251. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Blattidae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Ectobiinae, Anaplectinae, Nyctiborinae<br />

249. Ectobius lapponicus (Linnaeus); <strong>о</strong>бщий <strong>в</strong>ид, x6,6, <strong>с</strong><strong>о</strong><strong>в</strong>рем., x 9,6, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Shelford, 1907). 251. Megaloblatta longipennis.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Бей-Биен<strong>к</strong><strong>о</strong>, 1950). 250. Anaplecta sp.; А, Б — инди<strong>в</strong>и- Walker, x 3/4, <strong>с</strong><strong>о</strong><strong>в</strong>рем.. Центр. Амери<strong>к</strong><strong>а</strong><br />

ду<strong>а</strong>льн<strong>а</strong>я изменчи<strong>в</strong><strong>о</strong><strong>с</strong>ть ф<strong>о</strong>рмы и жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>с</strong> не<strong>в</strong>ытянутым <strong>в</strong>ершинным треуг<strong>о</strong>льни<strong>к</strong><strong>о</strong>м, ег<strong>о</strong><br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й шир<strong>о</strong><strong>к</strong><strong>о</strong> <strong>о</strong><strong>к</strong>руглен, рез<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я; Ri не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; R, М и CuA у <strong>в</strong>ершины<br />

при<strong>с</strong><strong>о</strong>единяют<strong>с</strong>я друг <strong>к</strong> другу; между М<br />

и R ряд непр<strong>а</strong><strong>в</strong>ильных ячее<strong>к</strong>, <strong>а</strong> между М и<br />

CuA ряд пр<strong>а</strong><strong>в</strong>ильных, прям<strong>о</strong>уг<strong>о</strong>льных ячее<strong>к</strong>.<br />

Средние и з<strong>а</strong>дние бедр<strong>а</strong> <strong>с</strong>низу <strong>в</strong><strong>о</strong><strong>о</strong>ружены <strong>с</strong>л<strong>а</strong>быми<br />

шип<strong>а</strong>ми, у <strong>в</strong>ершины ряд <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идных<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их шипи<strong>к</strong><strong>о</strong><strong>в</strong>. Л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>с</strong> не<strong>с</strong>имметричными<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 249). Немн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один <strong>с</strong><strong>о</strong><strong>в</strong>ременный<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

Ри<strong>с</strong>. 252—258. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Corydiidae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Corydiinae, Holocompsinae и Euthyrrhaphinae; Blattodea<br />

incertae sedis (<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cainoblattinidae)<br />

252. Corydia nuptialis Gerstatter, x 1,6, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Индия (Brunner,<br />

1865). 253. Polyphaga saussurei Dohrn; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>,<br />

x 2,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х 6,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Ср. Азия (Бей-Биен<strong>к</strong><strong>о</strong>,<br />

1950). 254. Polyphaga fossilis Shelford; личин<strong>к</strong><strong>а</strong>, х 15,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Shelford, 1910). 255. Нурегcompsa<br />

fieberi Brunner; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 10,0; Б — з<strong>а</strong>д­<br />

108<br />

http://jurassic.ru/<br />

нее <strong>к</strong>рыл<strong>о</strong>, х 10,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. Амери<strong>к</strong><strong>а</strong> (Brunner, 1865). 256. Н<strong>о</strong>locompsa<br />

fossils Shelford; <strong>с</strong><strong>а</strong>мец, х 6,3, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Shelford, 1910). 257. Еuthyrrhapa paclfica Coquebert;<br />

переднее и з<strong>а</strong>днее <strong>к</strong>рылья, х 10,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., тр<strong>о</strong>пи<strong>к</strong>и (Brunner,<br />

1865). 258. Cainoblattinopsis fushunensis Ping; <strong>о</strong>бщий <strong>в</strong>ид, х 3,0,<br />

<strong>о</strong>лиг<strong>о</strong>цен, Кит<strong>а</strong>й (янт<strong>а</strong>рь Фушун<strong>а</strong>,) (Ping, 1931)

109<br />

http://jurassic.ru/

ТРИБА^ PLECTOPTERINI SAUSSURE,<br />

1893<br />

[nom. transl. Rehn, 1951<br />

(ex Plectopterinae Saussure, 1893)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье п<strong>о</strong> длине р<strong>а</strong><strong>в</strong>н<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>рылу,<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>о</strong>н<strong>о</strong> 'р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е, треуг<strong>о</strong>льн<strong>о</strong>е, <strong>с</strong><br />

нея<strong>с</strong>ным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле неб<strong>о</strong>льш<strong>о</strong>е;<br />

R прям<strong>а</strong>я, не д<strong>а</strong>ет з<strong>а</strong>дних <strong>в</strong>ершинных <strong>в</strong>ет<strong>в</strong>ей;<br />

М и СиА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении<br />

<strong>в</strong> <strong>о</strong>бщий <strong>с</strong>тебель; у <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья<br />

р<strong>а</strong>зделяют<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong><br />

прям<strong>а</strong>я. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> б<strong>о</strong>льшим <strong>в</strong>ершинным<br />

треуг<strong>о</strong>льни<strong>к</strong><strong>о</strong>м, р<strong>а</strong><strong>в</strong>ным <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong><br />

и <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бным <strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>ть<strong>с</strong>я; Ri <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

между М и R ряд непр<strong>а</strong><strong>в</strong>ильных ячее<strong>к</strong>. Тел<strong>о</strong><br />

<strong>с</strong><strong>в</strong>ерху <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е. Бедр<strong>а</strong> <strong>с</strong>низу без шип<strong>о</strong><strong>в</strong>. Р<strong>а</strong>змеры<br />

мел<strong>к</strong>ие. Не<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле <strong>о</strong>дин из <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong><br />

Афри<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО ANAPLECTINAE WALKER, 1868<br />

[nom. transl. Hebard, 1929<br />

(ex Anaplectidae Walker, 1868)1<br />

Ф<strong>о</strong>рмы мел<strong>к</strong>ие, <strong>в</strong>нешне <strong>с</strong>х<strong>о</strong>дные <strong>с</strong> Coleoptera.<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>-<strong>в</strong>ытянут<strong>о</strong>е; плече<strong>в</strong><strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong>ытянут<strong>о</strong>; SC я<strong>с</strong>н<strong>а</strong>я, <strong>с</strong> ряд<strong>о</strong>м <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей; М <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ллельными з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>в</strong>ет<strong>в</strong>ями; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; R пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong><strong>с</strong>редине<br />

д<strong>о</strong> <strong>в</strong>ершины н<strong>а</strong>д<strong>к</strong>рылья без з<strong>а</strong>дних<br />

<strong>в</strong>ершинных <strong>в</strong>ет<strong>в</strong>ей. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итым<br />

<strong>в</strong>ершинным треуг<strong>о</strong>льни<strong>к</strong><strong>о</strong>м, <strong>с</strong>мещенным<br />

<strong>к</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong> и з<strong>а</strong>ним<strong>а</strong>ющим <strong>в</strong><strong>с</strong>ю ег<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льную<br />

ч<strong>а</strong><strong>с</strong>ть; жил<strong>к</strong>и R и М н<strong>а</strong> гр<strong>а</strong>нице п<strong>о</strong><strong>с</strong>ледней<br />

<strong>о</strong>бры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, длинн<strong>а</strong>я;<br />

R <strong>с</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й Ri и <strong>с</strong> ч<strong>а</strong><strong>с</strong>тью <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных<br />

н<strong>а</strong>з<strong>а</strong>д. В<strong>о</strong><strong>о</strong>ружение бедер <strong>с</strong>л<strong>а</strong>б<strong>о</strong>е (ри<strong>с</strong>.<br />

250). Не<strong>о</strong>ген — ныне. Пять <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

<strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле <strong>о</strong>дин из <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong> Афри<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО NYCTIBORINAE SAUSSURE, 1893<br />

(Nyctiboridae Brunner, 1893;<br />

Nyctiborides Sharp, 1895)<br />

Н<strong>а</strong>д<strong>к</strong>рылья и передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> п<strong>о</strong><strong>к</strong>рыты шел<strong>к</strong><strong>о</strong><strong>в</strong>и<strong>с</strong>тым<br />

пуш<strong>к</strong><strong>о</strong>м. Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле без жил<strong>о</strong><strong>к</strong> или ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми<br />

SC; R <strong>с</strong> мн<strong>о</strong>гими непр<strong>а</strong><strong>в</strong>ильными передними<br />

<strong>в</strong>ет<strong>в</strong>ями; М и СиА мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>ты, нер<strong>а</strong>зделимы,<br />

<strong>в</strong>ет<strong>в</strong>и <strong>к</strong><strong>о</strong><strong>с</strong>ые, п<strong>а</strong>р<strong>а</strong>ллельные з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние М <strong>с</strong>ближен<strong>о</strong> <strong>с</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым R и<br />

СиА. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й пред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью, я<strong>с</strong>н<strong>о</strong>й Ri; R <strong>с</strong> мн<strong>о</strong>гими передними<br />

и <strong>в</strong>ершинными <strong>в</strong>ет<strong>в</strong>ями и <strong>с</strong> делящей<strong>с</strong>я з<strong>а</strong>дней<br />

<strong>в</strong>ершинн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью; М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>ближен<strong>а</strong><br />

<strong>с</strong> RS; СиА <strong>с</strong><strong>о</strong> МНОГИМИ непр<strong>а</strong><strong>в</strong>ильными<br />

<strong>в</strong>ет<strong>в</strong>ями, п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> или д<strong>в</strong>е трети их <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й, <strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льные — <strong>в</strong> CuP; <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong><br />

http://jurassic.ru/<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет <strong>в</strong>переди передне<strong>с</strong>пин<strong>к</strong>и. Цер<strong>к</strong>и<br />

длинные. Бедр<strong>а</strong> <strong>с</strong> шип<strong>а</strong>ми. Р<strong>а</strong>змеры <strong>к</strong>рупные<br />

(ри<strong>с</strong>. 251). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один <strong>с</strong><strong>о</strong><strong>в</strong>ременный<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

BLATTIDAE 1NCERTAE SEDIS<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> Blattidae <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я м<strong>а</strong>л<strong>о</strong>изученные<br />

р<strong>о</strong>ды из н. мел<strong>а</strong> Кит<strong>а</strong>я: Sinoblatta Grabau,<br />

1923 и Layangia Grabau, 1923.<br />

СЕМЕЙСТВО CORYDIIDAE SAUSSURE, 1864<br />

[nom, transl. Handlirsch, 1925<br />

(ex Corydiens Saussure, 1864)1<br />

[Polyphagidae Walker, 1868;<br />

Melander-Brues, 1932;<br />

Archimylacridae Karny, 1921 (pars)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья и <strong>к</strong>рылья, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, неред<strong>к</strong><strong>о</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, и т<strong>о</strong>гд<strong>а</strong> тел<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е.<br />

Ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong> груди<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> ближе <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние или <strong>с</strong>х<strong>о</strong>дн<strong>о</strong> <strong>с</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым Archimylacridae<br />

или же <strong>с</strong> рез<strong>к</strong><strong>о</strong>й <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>цией: п<strong>о</strong>лн<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>а</strong>цией или <strong>с</strong> <strong>с</strong>ильн<strong>о</strong>й реду<strong>к</strong>цией жил<strong>о</strong><strong>к</strong><br />

и р<strong>а</strong>з<strong>в</strong>итием птер<strong>о</strong><strong>с</strong>тигмы; SC ч<strong>а</strong><strong>с</strong>т<strong>о</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

<strong>с</strong>ередины <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong>р<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong>ыми <strong>в</strong>ет<strong>в</strong>ями<br />

и л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния; R р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, без<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й Ri. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й<br />

пред<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью; <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть,<br />

неб<strong>о</strong>льш<strong>а</strong>я, не <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, <strong>а</strong><br />

лишь п<strong>о</strong>дгиб<strong>а</strong>ет<strong>с</strong>я. Н<strong>а</strong>лични<strong>к</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>тгр<strong>а</strong>ничен<br />

<strong>о</strong>т<strong>о</strong> лб<strong>а</strong>, я<strong>с</strong>н<strong>о</strong> р<strong>а</strong>зделен н<strong>а</strong> переднюю и з<strong>а</strong>днюю<br />

ч<strong>а</strong><strong>с</strong>ти. Бедр<strong>а</strong> <strong>с</strong>низу без шип<strong>о</strong><strong>в</strong> или <strong>с</strong> <strong>о</strong>днимд<strong>в</strong>умя<br />

шип<strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Пять п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне: Corydiinae, Н<strong>о</strong>locompsinae,<br />

Euthyrrhaphinae, Latindiinae, Tiviinae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не<br />

н<strong>а</strong>йдены.<br />

ПОДСЕМЕЙСТВО CORYDIINAE SAUSSURE, 1870<br />

[nom. transl. Saussure, 1893<br />

(ex Corydites Saussure, 1870)1<br />

(Polyphaginae Kirby, 1904;<br />

Karny, 1921; Rehn, 1951)<br />

Н<strong>а</strong>д<strong>к</strong>рылье и з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

плече<strong>в</strong><strong>о</strong>е п<strong>о</strong>ле умеренн<strong>о</strong>е; SC <strong>в</strong> н<strong>а</strong>д<strong>к</strong>рылье<br />

длинн<strong>а</strong>я, <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>в</strong>ет<strong>в</strong>ями; Ri мн<strong>о</strong>г<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong> делит<strong>с</strong>я; М имеет<br />

три <strong>в</strong>ет<strong>в</strong>и и б<strong>о</strong>лее; СиА х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>е, <strong>с</strong> п<strong>о</strong>лным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; SC и Ri<br />

пр<strong>о</strong><strong>с</strong>тые, реже делят<strong>с</strong>я; RS <strong>с</strong> непр<strong>а</strong><strong>в</strong>ильными<br />

передними и <strong>а</strong>пи<strong>к</strong><strong>а</strong>льными или <strong>с</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льными<br />

и з<strong>а</strong>дними <strong>в</strong>ет<strong>в</strong>ями; М <strong>с</strong> тремя или б<strong>о</strong>лее <strong>в</strong>ет<strong>в</strong>ями;<br />

СиА <strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей. Передние<br />

бедр<strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>о</strong>дним <strong>в</strong>ершинным шип<strong>о</strong>м. Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

<strong>к</strong>рупные ф<strong>о</strong>рмы (ри<strong>с</strong>. 252—254). П<strong>а</strong>ле- .

<strong>о</strong>ген — ныне. Д<strong>в</strong>е трибы <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне:<br />

Corydiini и Polyphagini, из них <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong>» п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО HOLOCOMPSINAE<br />

HANDLIRSCH, 1925<br />

[nom. transl. Rehn, 1951<br />

(ex Holocompsini Handlirsch, 1925)1<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong>зр<strong>а</strong>чны,<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> упл<strong>о</strong>тнены; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

неп<strong>о</strong>лн<strong>о</strong>е, ди<strong>с</strong>т<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и <strong>в</strong> <strong>с</strong>ередине<br />

<strong>о</strong>бры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я; SC <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

R <strong>с</strong> <strong>с</strong>ерией <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти; RS неп<strong>о</strong>лн<strong>а</strong>я; М и СиА б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> т<strong>о</strong>л<strong>с</strong>тый, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий <strong>с</strong>тебеле<strong>к</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> <strong>в</strong>ерти<strong>к</strong><strong>а</strong>льн<strong>а</strong>я, <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е т<strong>о</strong>ле без жил<strong>о</strong><strong>к</strong><br />

или <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми жил<strong>о</strong><strong>к</strong>. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й SC; <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я Ri и р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я<br />

RS <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> М <strong>в</strong> п<strong>с</strong>е<strong>в</strong>д<strong>о</strong><strong>с</strong>тигму; СиА<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> я<strong>с</strong>н<strong>а</strong>я, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>о</strong>бры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тым<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м (ри<strong>с</strong>. 255 и 256). П<strong>а</strong>ле<strong>о</strong>ген —<br />

Вне СССР м<strong>а</strong>л<strong>о</strong>изученные р<strong>о</strong>ды Cainoblattinopsis<br />

Ping, 1931 (Cainoblattinidae Ping,<br />

1931; ри<strong>с</strong>. 258) —из н. мел<strong>а</strong> Кит<strong>а</strong>я, Paralellophora<br />

Haupt, 1956 (Paralellophoridae Haupt,<br />

Средней <strong>в</strong>еличины или <strong>к</strong>рупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными <strong>к</strong>рыльями. Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и нет;<br />

SC длинн<strong>а</strong>я, <strong>в</strong>ыте<strong>с</strong>ни<strong>в</strong>ш<strong>а</strong>я передние <strong>в</strong>ет<strong>в</strong>и R,<br />

<strong>к</strong>р<strong>о</strong>ме <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я т<strong>а</strong><strong>к</strong>же длинн<strong>а</strong>я, п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

SC и R; <strong>в</strong>ет<strong>в</strong>ей М м<strong>а</strong>л<strong>о</strong>, <strong>о</strong>ни н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> н<strong>а</strong>з<strong>а</strong>д и н<strong>а</strong>ружу; Си мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>о</strong>бычн<strong>о</strong> меньше, чем<br />

у т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>; мн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>, ин<strong>о</strong>гд<strong>а</strong><br />

измененный <strong>а</strong>рхеди<strong>к</strong>тий. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> б<strong>о</strong>льшим<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льным <strong>в</strong>еер<strong>о</strong>м.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я, <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я; р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызущег<strong>о</strong> тип<strong>а</strong>; <strong>а</strong>нтенны не<br />

длиннее тел<strong>а</strong>, ните<strong>в</strong>идные; ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong>,<br />

три гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. Переднегрудь <strong>о</strong>бычн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>,<br />

длиннее д<strong>в</strong>ух других <strong>с</strong>егмент<strong>о</strong><strong>в</strong> груди,<br />

не при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы ; передние н<strong>о</strong>ги х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные;<br />

т<strong>а</strong>зи<strong>к</strong> <strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличен, п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ен<br />

п<strong>о</strong> длине шир<strong>о</strong><strong>к</strong><strong>о</strong>му бедру, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е не<strong>с</strong>ет ряд<br />

шип<strong>о</strong><strong>в</strong> п<strong>о</strong> нижнему <strong>к</strong>р<strong>а</strong>ю; г<strong>о</strong>лени <strong>к</strong><strong>о</strong>р<strong>о</strong>че, т<strong>а</strong><strong>к</strong>же<br />

<strong>с</strong> шип<strong>а</strong>ми п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю; <strong>с</strong>редние и з<strong>а</strong>дние н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие,<br />

длинные, х<strong>о</strong>дильные; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> пять.<br />

К<strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие члени<strong>с</strong>тые цер<strong>к</strong>и. Яйц<strong>а</strong> <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<br />

<strong>в</strong> пл<strong>о</strong>тных <strong>к</strong><strong>о</strong><strong>к</strong><strong>о</strong>н<strong>а</strong>х. Хищни<strong>к</strong>и тр<strong>о</strong>пи<strong>к</strong><strong>о</strong><strong>в</strong> и <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 259). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

BLATTODEA INCERTAE SED1S<br />

ОТРЯД MANTEODEA. БОГОМОЛЫ<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

111<br />

http://jurassic.ru/<br />

ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из них <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО EUTHYRRHAPHINAE<br />

HANDLIRSCH, 1925<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>о</strong><br />

лишь <strong>с</strong>лед SC и R, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>,<br />

у <strong>в</strong>ершины шир<strong>о</strong><strong>к</strong><strong>о</strong> <strong>о</strong><strong>к</strong>ругленн<strong>о</strong>е; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> <strong>в</strong> <strong>в</strong>иде <strong>с</strong>л<strong>а</strong>б<strong>о</strong>й из<strong>о</strong>гнут<strong>о</strong>й линии.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> длиннее н<strong>а</strong>д<strong>к</strong>рылья; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R, Ri <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>в</strong>ет<strong>в</strong>ей R упл<strong>о</strong>тненн<strong>а</strong>я п<strong>с</strong>е<strong>в</strong>д<strong>о</strong><strong>с</strong>тигм<strong>а</strong>,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>в</strong>ых<strong>о</strong>дит ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть четырех <strong>в</strong>ет<strong>в</strong>ей R; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>; СиА<br />

<strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью н<strong>а</strong><strong>к</strong>л<strong>о</strong>нными <strong>в</strong>ет<strong>в</strong>ями. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> ш<strong>а</strong>р<strong>о</strong><strong>в</strong>идн<strong>а</strong>я,<br />

без я<strong>с</strong>н<strong>о</strong>г<strong>о</strong> <strong>в</strong>здутия <strong>в</strong> нижней ч<strong>а</strong><strong>с</strong>ти, <strong>с</strong><br />

б<strong>о</strong>льшим postclypeus. Бедр<strong>а</strong> <strong>с</strong> длинными <strong>в</strong>ершинными<br />

шип<strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы (ри<strong>с</strong>. 257). Не<strong>о</strong>ген — ныне.<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

<strong>о</strong>дин р<strong>о</strong>д из <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong> Афри<strong>к</strong>и.<br />

1956)—из третичных <strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

Lygobius Mitchell, 1908 (Lygobiidae Mitchell,<br />

1908) — из IH. мел<strong>а</strong> С. Амери<strong>к</strong>и.<br />

Ри<strong>с</strong>. 259. Отряд Manteodea<br />

Paratenodera sinensis Saussure; <strong>о</strong>бщий <strong>в</strong>ид, х 1,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Essijj, 1947)

1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне. Д<strong>в</strong><strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong><br />

нея<strong>с</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жения из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

ОТРЯД ISOPTERA. ТЕРМИТЫ<br />

Мел<strong>к</strong>ие и <strong>с</strong>редней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые,<br />

жи<strong>в</strong>ущие б<strong>о</strong>льшими <strong>к</strong><strong>о</strong>л<strong>о</strong>ниями. Рез<strong>к</strong><strong>о</strong> п<strong>о</strong>лим<strong>о</strong>рфны:<br />

н<strong>о</strong>рм<strong>а</strong>льные п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>о</strong><strong>с</strong><strong>о</strong>би, <strong>к</strong>рыл<strong>а</strong>тые<br />

<strong>с</strong><strong>а</strong>мцы («ц<strong>а</strong>ри»), <strong>с</strong><strong>а</strong>м<strong>к</strong>и («ц<strong>а</strong>рицы>) и<br />

бе<strong>с</strong><strong>к</strong>рылые, нер<strong>а</strong>змн<strong>о</strong>ж<strong>а</strong>ющие<strong>с</strong>я р<strong>а</strong>б<strong>о</strong>чие и <strong>с</strong><strong>о</strong>л­<br />

д<strong>а</strong>ты 1<br />

р<strong>а</strong>зличных ф<strong>о</strong>рм. Д<strong>в</strong>е п<strong>а</strong>ры длинных<br />

г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong>; ин<strong>о</strong>гд<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>е р<strong>а</strong><strong>с</strong>ширение<br />

н<strong>а</strong> з<strong>а</strong>дних <strong>к</strong>рыльях (Mastotermitidae).<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е, н<strong>о</strong> нежн<strong>о</strong>е, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящее из<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных <strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong>;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и п<strong>о</strong>чти не <strong>в</strong>ыр<strong>а</strong>жены 1<br />

,<br />

реже <strong>о</strong>ни <strong>с</strong>л<strong>а</strong>бые и т<strong>о</strong>н<strong>к</strong>ие; <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я<br />

SC; пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> б<strong>о</strong>льшим <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных<br />

<strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ей R; М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или<br />

<strong>с</strong> тремя — ше<strong>с</strong>тью <strong>в</strong>ет<strong>в</strong>ями, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными н<strong>а</strong>з<strong>а</strong>д;<br />

Си х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> и не<strong>с</strong>ет мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленные н<strong>а</strong>з<strong>а</strong>д <strong>в</strong>ет<strong>в</strong>и; <strong>о</strong>бычн<strong>о</strong><br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть. О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>й чешуи,<br />

п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>к</strong>рылья <strong>о</strong>бл<strong>а</strong>мы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я при<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Ри<strong>с</strong>. 260—266. Отряд Isoptera<br />

260. Diatermes slblricus Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, x 2,5, <strong>о</strong>лиг<strong>о</strong>цен,<br />

В. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1929). 261. Mastotermes angllcanus<br />

Rosen; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,0, <strong>о</strong>лиг<strong>о</strong>цен, Англия (Rosen,<br />

1913). 262. Eotermes grandaeva Statz; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,2,<br />

<strong>о</strong>иг<strong>о</strong>цен, Герм<strong>а</strong>ния (Statz, 1939). 163. Termopsls bremii (Heer);<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

112<br />

http://jurassic.ru/<br />

<strong>о</strong>дин <strong>в</strong>ид из ми<strong>о</strong>цен<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы, три <strong>в</strong>ид<strong>а</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящие<strong>с</strong>я<br />

<strong>к</strong> д<strong>в</strong>ум р<strong>о</strong>д<strong>а</strong>м, из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

перех<strong>о</strong>де <strong>к</strong>рыл<strong>а</strong>т<strong>о</strong>й <strong>о</strong><strong>с</strong><strong>о</strong>би <strong>к</strong> п<strong>о</strong>ч<strong>в</strong>енн<strong>о</strong>му <strong>о</strong>бр<strong>а</strong>зу<br />

жизни. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

грызущег<strong>о</strong> тип<strong>а</strong>. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> 10—<br />

30 члени<strong>к</strong><strong>а</strong>ми. С<strong>а</strong>мцы и <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong> ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чными<br />

гл<strong>а</strong>з<strong>а</strong>ми и д<strong>в</strong>умя пр<strong>о</strong><strong>с</strong>тыми гл<strong>а</strong>з<strong>к</strong><strong>а</strong>ми. Грудные<br />

<strong>с</strong>егменты п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные; н<strong>о</strong>ги бег<strong>а</strong>тельные,<br />

т<strong>о</strong>н<strong>к</strong>ие; л<strong>а</strong>п<strong>к</strong>и четырех-, реже<br />

пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Цер<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>о</strong>дн<strong>о</strong>- или<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, ред<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Жи<strong>в</strong>ут<br />

<strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, дре<strong>в</strong>е<strong>с</strong>ине или <strong>в</strong> <strong>с</strong>пеци<strong>а</strong>льных<br />

п<strong>о</strong><strong>с</strong>тр<strong>о</strong>й<strong>к</strong><strong>а</strong>х — термитни<strong>к</strong><strong>а</strong>х. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные;<br />

<strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н<strong>а</strong>х прин<strong>о</strong><strong>с</strong>ят б<strong>о</strong>льш<strong>о</strong>й<br />

<strong>в</strong>ред, р<strong>а</strong>зруш<strong>а</strong>я дере<strong>в</strong>янные п<strong>о</strong><strong>с</strong>тр<strong>о</strong>й<strong>к</strong>и, мебель<br />

и другие п<strong>о</strong>дел<strong>к</strong>и. Р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены<br />

<strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти <strong>в</strong> <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х. Из<strong>в</strong>е<strong>с</strong>тны<br />

третичные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и дре<strong>в</strong>е<strong>с</strong>ины <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми<br />

п<strong>о</strong><strong>в</strong>реждений термит<strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Mastotermitidae, Calotermitidae,<br />

Hodotermitidae, Rhinotermitidae, Termitidae.<br />

(Rosen, 1913). 264. Ulmerielta cockerelli Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 2,8, <strong>о</strong>лиг<strong>о</strong>цен, В. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1929). 265.<br />

Parastylotermes washingtonensis (Snyder); переднее <strong>к</strong>рыл<strong>о</strong>, х 5,0,<br />

не<strong>о</strong>ген, С. Амери<strong>к</strong><strong>а</strong> (Snvder, 1931). 266. Constrictotermes tenuirostrls<br />

Desneux; <strong>о</strong>бщий <strong>в</strong>ид, x 11,0. <strong>с</strong><strong>о</strong><strong>в</strong>рем., Ме<strong>к</strong><strong>с</strong>и<strong>к</strong><strong>а</strong> (Handlirsch,<br />

1925)

СЕМЕЙСТВО MASTOTERMITIDAE DESNEUX, 1904<br />

[nom. transl. Silvestri, 1909<br />

(ex Mastotermitinae Desneux, 1904)]<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й С; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины' длины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зуя<br />

длинные <strong>в</strong>ет<strong>в</strong>и; R п<strong>о</strong>чти прям<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и<br />

R н<strong>а</strong>чин<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>чти <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>, длинные,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong>перед; М <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> R и<br />

не<strong>с</strong>ет три-четыре <strong>в</strong>ет<strong>в</strong>и; меди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

шире <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; Си <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>в</strong>ершине<br />

<strong>к</strong>рыл<strong>а</strong>, <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> ее <strong>в</strong>ет<strong>в</strong>ей з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю<br />

з<strong>а</strong>днюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину <strong>к</strong>рыл<strong>а</strong>. В з<strong>а</strong>днем <strong>к</strong>рыле б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

ячеи нет, <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>.<br />

Л<strong>а</strong>п<strong>к</strong>и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Стр<strong>о</strong>ят гнезд<strong>а</strong><br />

гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м п<strong>о</strong>д землей, р<strong>а</strong>зруш<strong>а</strong>ют<br />

<strong>к</strong><strong>о</strong>рни и <strong>с</strong>т<strong>в</strong><strong>о</strong>лы дере<strong>в</strong>ье<strong>в</strong>. Из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии (ри<strong>с</strong>. 260 и 261).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы, Ср. Азии и С. Амери<strong>к</strong>и, из них <strong>о</strong>дин<br />

(Mastoterm.es) — <strong>с</strong><strong>о</strong><strong>в</strong>ременный.<br />

СЕМЕЙСТВО CALOTERMITIDAE ENDERLE1N, 1909<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й п<strong>о</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> н<strong>а</strong>ч<strong>а</strong>ле <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>, без<br />

<strong>в</strong>ет<strong>в</strong>ей; R пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> длинными <strong>в</strong>ет<strong>в</strong>ями,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong>перед; М <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> R, <strong>в</strong>ет<strong>в</strong>и<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> нея<strong>с</strong>ные; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е и меди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ля р<strong>а</strong><strong>в</strong>ны между <strong>с</strong><strong>о</strong>б<strong>о</strong>й п<strong>о</strong> ширине; <strong>с</strong>и<strong>с</strong>тем<strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>ей Си з<strong>а</strong>ним<strong>а</strong>ет б<strong>о</strong>льшую ч<strong>а</strong><strong>с</strong>ть ширины 1<br />

<strong>к</strong>рыл<strong>а</strong>. Б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячея <strong>в</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях.<br />

Гл<strong>а</strong>з<strong>к</strong>и е<strong>с</strong>ть. Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> четыре. Цер<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, ин<strong>о</strong>гд<strong>а</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

пять. Жи<strong>в</strong>ут <strong>в</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии <strong>в</strong> пнях и <strong>с</strong>ух<strong>о</strong><strong>с</strong>т<strong>о</strong>е,<br />

р<strong>а</strong>б<strong>о</strong>чих нет (ри<strong>с</strong>. 262). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>с</strong> <strong>с</strong>емью р<strong>о</strong>д<strong>а</strong>ми из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы, ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и и плей<strong>с</strong>т<strong>о</strong>цен<strong>а</strong><br />

В. Афри<strong>к</strong>и.<br />

СЕМЕЙСТВО HODOTERMITIDAE DESNEUX, 1904<br />

[nom. transl. Snyder, 1925<br />

(ex Hodotermitini Desneux, 1904)]<br />

Чешуя переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че чешуи з<strong>а</strong>днег<strong>о</strong>;<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ереди­<br />

ны" длины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; R <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; М<br />

пр<strong>о</strong>х<strong>о</strong>дит ближе <strong>к</strong> Си, <strong>с</strong> д<strong>в</strong>умя — пятью <strong>в</strong>ет<strong>в</strong>ями;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шире меди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>.<br />

Гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> четыре. Цер<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие (ри<strong>с</strong>. 263 и 264). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong><br />

пятью р<strong>о</strong>д<strong>а</strong>ми из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы и ми<strong>о</strong>цен<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО RHINOTERMIT1DAE FROGGAT,<br />

1896<br />

[nom. transl. Light, 1921<br />

(ex Rhinotermitinae Froggat, 1896)]<br />

Чешуя переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> длиннее чешуи<br />

з<strong>а</strong>днег<strong>о</strong>; SC нет; R <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями или без них; М без <strong>в</strong>ет<strong>в</strong>ей или<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ями;<br />

меди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле немн<strong>о</strong>г<strong>о</strong> шире р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

Си <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong>; <strong>с</strong>и<strong>с</strong>тем<strong>а</strong><br />

ее <strong>в</strong>ет<strong>в</strong>ей з<strong>а</strong>ним<strong>а</strong>ет п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину ширины <strong>к</strong>рыл<strong>а</strong>.<br />

Ф<strong>о</strong>нт<strong>а</strong>нель е<strong>с</strong>ть. Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> четыре. Цер<strong>к</strong>и<br />

д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 265). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>с</strong> пятью р<strong>о</strong>д<strong>а</strong>ми из третичных <strong>о</strong>тл<strong>о</strong>жений<br />

Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и и плей<strong>с</strong>т<strong>о</strong>цен<strong>а</strong><br />

В.-Афри<strong>к</strong>и.<br />

СЕМЕЙСТВО TERMITIDAE LATRE1LLE, 1802<br />

Чешуя переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; SC нет;<br />

R пр<strong>о</strong>х<strong>о</strong>дит близ<strong>к</strong><strong>о</strong> <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>,<br />

без <strong>в</strong>ет<strong>в</strong>ей; М р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> близ<strong>к</strong><strong>о</strong> <strong>к</strong> Си, мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шире меди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

Си <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, без <strong>в</strong>ет<strong>в</strong>ей, или <strong>в</strong>ет<strong>в</strong>ей <strong>о</strong>дн<strong>а</strong>-д<strong>в</strong>е,<br />

и т<strong>о</strong>гд<strong>а</strong> Си длинн<strong>а</strong>я; <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ей<br />

Си з<strong>а</strong>ним<strong>а</strong>ет б<strong>о</strong>льшую ч<strong>а</strong><strong>с</strong>ть ширины <strong>к</strong>рыл<strong>а</strong>.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> четыре. Стр<strong>о</strong>ят б<strong>о</strong>льшие<br />

гнезд<strong>а</strong> (ри<strong>с</strong>. 266). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Четыре<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong> ше<strong>с</strong>тью<br />

р<strong>о</strong>д<strong>а</strong>ми из э<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и плей<strong>с</strong>т<strong>о</strong>цен<strong>а</strong><br />

В. Афри<strong>к</strong>и.<br />

ОТРЯД DERM APT ERA. КОЖИСТОКРЫЛЫЕ<br />

Крыл<strong>а</strong>тые и бе<strong>с</strong><strong>к</strong>рылые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Передние<br />

<strong>к</strong>рылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, лишенные жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния,<br />

<strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные*, не при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют брюш<strong>к</strong><strong>а</strong>.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong>еер, <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong>д<strong>о</strong>ль и 2 р<strong>а</strong>з<strong>а</strong> п<strong>о</strong>пере<strong>к</strong>, <strong>с</strong><strong>к</strong>рыты п<strong>о</strong>д<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

8 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 113<br />

н<strong>а</strong>д<strong>к</strong>рыльями. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

грызущег<strong>о</strong> тип<strong>а</strong>; <strong>а</strong>нтенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

Н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, бедр<strong>а</strong> р<strong>а</strong><strong>с</strong>ширенные; члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

л<strong>а</strong>п<strong>о</strong><strong>к</strong> три (у мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм четырепять).<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong> члени<strong>с</strong>тыми или пр<strong>о</strong><strong>с</strong>тыми<br />

http://jurassic.ru/

цер<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими <strong>к</strong>лещи; <strong>с</strong>терниты и<br />

тергиты, <strong>с</strong><strong>о</strong>единяя<strong>с</strong>ь, <strong>о</strong>бр<strong>а</strong>зуют <strong>с</strong>б<strong>о</strong><strong>к</strong>у зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зную<br />

линию. Преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> н<strong>о</strong>чные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые,<br />

днем <strong>с</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>с</strong>ырых у<strong>к</strong>р<strong>о</strong>мных<br />

ме<strong>с</strong>т<strong>а</strong>х: п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями, <strong>о</strong>п<strong>а</strong><strong>в</strong>шей ли<strong>с</strong>т<strong>в</strong><strong>о</strong>й, <strong>к</strong><strong>о</strong>р<strong>о</strong>й,<br />

<strong>в</strong> гниющих пнях. В<strong>с</strong>еядны. Юр<strong>а</strong> — ныне.<br />

П<strong>о</strong>д<strong>о</strong>тряды: Archidermaptera, Eudermaptera.<br />

ПОДОТРЯД ARCHIDERMAPTERA<br />

Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> длинные н<strong>а</strong>д<strong>к</strong>рылья, д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong><br />

<strong>с</strong>ередины <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> брюш<strong>к</strong><strong>а</strong>, <strong>с</strong> немн<strong>о</strong>гими<br />

пр<strong>о</strong>д<strong>о</strong>льными ребрыш<strong>к</strong><strong>а</strong>ми, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

жил<strong>к</strong><strong>а</strong>ми. Антенны <strong>с</strong> длинным <strong>в</strong>т<strong>о</strong>рым члени<strong>к</strong><strong>о</strong>м<br />

(р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине третьему). Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

четыре-пять. Цер<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, т<strong>о</strong>н<strong>к</strong>ие,<br />

члени<strong>с</strong>тые. Юр<strong>а</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protodiplatidae.<br />

СЕМЕЙСТВО PROTODIPLATIDAE<br />

MARTYNOV, 1925<br />

Н<strong>а</strong>д<strong>к</strong>рылья нешир<strong>о</strong><strong>к</strong>ие, пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тые; <strong>о</strong>т<br />

четырех д<strong>о</strong> пяти п<strong>о</strong>перечных <strong>к</strong><strong>о</strong>ричне<strong>в</strong>ых п<strong>о</strong>л<strong>о</strong><strong>с</strong>.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 17—18. Члени<strong>к</strong><strong>о</strong><strong>в</strong> цер<strong>о</strong><strong>к</strong><br />

пять-ше<strong>с</strong>ть. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Protodiplatys Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. fortis Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Пер<strong>в</strong>ый члени<strong>к</strong><br />

цер<strong>о</strong><strong>к</strong> р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине <strong>в</strong>т<strong>о</strong>р<strong>о</strong>му и третьему <strong>в</strong>ме<strong>с</strong>те<br />

<strong>в</strong>зятым, чет<strong>в</strong>ертый и пятый зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че, ше<strong>с</strong>т<strong>о</strong>й — рудимент<strong>а</strong>рный. Длин<strong>а</strong> тел<strong>а</strong><br />

20 мм (ри<strong>с</strong>. 267). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

ПОДОТРЯД EUDERMAPTERA<br />

(Forficulina)<br />

Крыл<strong>а</strong>тые и бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы 1<br />

; н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>о</strong>бычн<strong>о</strong> д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> брюш<strong>к</strong><strong>а</strong> (<strong>к</strong>р<strong>о</strong>ме<br />

Semenoviola), <strong>в</strong><strong>с</strong>егд<strong>а</strong> без пр<strong>о</strong>д<strong>о</strong>льных ребрыше<strong>к</strong>;<br />

лишь у немн<strong>о</strong>гих ф<strong>о</strong>рм пр<strong>о</strong>д<strong>о</strong>льный б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>к</strong>иль. Антенны <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им <strong>в</strong>т<strong>о</strong>рым члени<strong>к</strong><strong>о</strong>м<br />

(<strong>к</strong><strong>о</strong>р<strong>о</strong>че третьег<strong>о</strong>). Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

три. Цер<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, б<strong>о</strong>льшие,<br />

пре<strong>в</strong>р<strong>а</strong>щенные <strong>в</strong> <strong>к</strong>лещи. В<strong>к</strong>люч<strong>а</strong>ют п<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющее<br />

б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Юр<strong>а</strong> —<br />

ныне. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>. Protodermaptera, Forficulidea.<br />

НАДСЕМЕЙСТВО PROTODERMAPTERA<br />

Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> е<strong>с</strong>ть гл<strong>а</strong>з<strong>к</strong>и. Генит<strong>а</strong>льный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong> п<strong>а</strong>рным penis; пигидий пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й. Личин<strong>к</strong>и<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>с</strong> члени<strong>с</strong>тыми цер<strong>к</strong><strong>а</strong>ми.<br />

Ми<strong>о</strong>цен — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>,<br />

из них т<strong>о</strong>ль<strong>к</strong><strong>о</strong> Labiduridae (ри<strong>с</strong>. 268)<br />

114<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 267—271. Отряд Dermaptera<br />

267. Protodiplatys fortis Martynov; <strong>о</strong>бщий <strong>в</strong>ид, х 3,3, <strong>в</strong>. г<strong>о</strong>р<strong>а</strong>,<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 268. Labidura riparia (Pallas);<br />

<strong>о</strong>бщий <strong>в</strong>ид, х 2,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Бей-Биен<strong>к</strong><strong>о</strong>, 1936). 269. Semenoviola<br />

obliquotruncata Martynov; <strong>о</strong>бщий <strong>в</strong>ид, х 4,3, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 270. Forficula kaznakovi Semenov; <strong>о</strong>бщий<br />

<strong>в</strong>ид, х 2,6, <strong>с</strong><strong>о</strong><strong>в</strong>рем., К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (Бей-Биен<strong>к</strong><strong>о</strong>, 1936). 271. Forficula<br />

aurtcularia Linnaeus; А — переднее <strong>к</strong>рыл<strong>о</strong>, Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, х 15,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938)

из<strong>в</strong>е<strong>с</strong>тны из третичных <strong>о</strong>тл<strong>о</strong>жений; <strong>о</strong>дин р<strong>о</strong>д из<br />

янт<strong>а</strong>ря Бирмы, друг<strong>о</strong>й — из Аргентины.<br />

НАДСЕМЕЙСТВО FORFICULIDEA<br />

(Eudermaptera)<br />

Гл<strong>а</strong>з<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Генит<strong>а</strong>льный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й; penis неп<strong>а</strong>рный; пилидий ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>в</strong><strong>о</strong><strong>о</strong>ружен шип<strong>а</strong>ми и <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми; цер<strong>к</strong>и не<br />

<strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Labiide,<br />

Forficulidae.<br />

СЕМЕЙСТВО LABIIDAE BURR, 1909<br />

От <strong>о</strong>чень мел<strong>к</strong>их д<strong>о</strong> умеренн<strong>о</strong> <strong>к</strong>рупных<br />

ф<strong>о</strong>рм. Н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> р<strong>а</strong>з<strong>в</strong>иты, ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong> <strong>к</strong>илями. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 10—23. Вт<strong>о</strong>р<strong>о</strong>й<br />

члени<strong>к</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, цилиндриче<strong>с</strong><strong>к</strong>ий. Генит<strong>а</strong>льный<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренными н<strong>а</strong><br />

<strong>в</strong>ершине мет<strong>а</strong>п<strong>а</strong>р<strong>а</strong>мер<strong>а</strong>ми. Юр<strong>а</strong> — ныне. Д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

Labiinae из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

ПОДСЕМЕЙСТВО LABIINAE VERHOEFF, 1902<br />

Мел<strong>к</strong>ие и <strong>о</strong>чень мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Юр<strong>а</strong> —<br />

ныне. Б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

Передние <strong>к</strong>рылья <strong>в</strong> <strong>в</strong>иде пл<strong>о</strong>тных, <strong>в</strong>ыпу<strong>к</strong>лых,<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льных н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>, <strong>с</strong> немн<strong>о</strong>гими прямыми,<br />

ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я жил<strong>к</strong><strong>а</strong>ми. З<strong>а</strong>дние<br />

<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ие, длиннее<br />

передних, <strong>с</strong> <strong>в</strong>ершинным <strong>о</strong>тдел<strong>о</strong>м, <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ющим<strong>с</strong>я<br />

п<strong>о</strong>пере<strong>к</strong> :<strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>к</strong>рылья п<strong>о</strong><strong>к</strong>рыты<br />

н<strong>а</strong>д<strong>к</strong>рыльями. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong><br />

<strong>в</strong> <strong>в</strong>иде мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящих<strong>с</strong>я<br />

жил<strong>о</strong><strong>к</strong>. Тел<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>о</strong>е, пл<strong>о</strong>тн<strong>о</strong>е, <strong>с</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ыми, н<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>а</strong>нтенн<strong>а</strong>ми и без длинных цер<strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 272). Обр<strong>а</strong>з жизни и р<strong>а</strong>з<strong>в</strong>итие неиз<strong>в</strong>е<strong>с</strong>тны.<br />

Пермь. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Protelytridae, Megelytridae,<br />

Blattelytridae, Archelytridae, Elytroneuridae,<br />

из них н<strong>а</strong> террит<strong>о</strong>рии СССР <strong>о</strong>бн<strong>а</strong>ружен<strong>о</strong><br />

пер<strong>в</strong><strong>о</strong>е.<br />

Фил<strong>о</strong>генез изучен нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>: п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

<strong>с</strong><strong>в</strong>яз<strong>а</strong>ны <strong>с</strong> и<strong>с</strong>х<strong>о</strong>дными ф<strong>о</strong>рм<strong>а</strong>ми ух<strong>о</strong><strong>в</strong>ерт<strong>о</strong><strong>к</strong>.<br />

И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие, <strong>в</strong>идим<strong>о</strong>, <strong>о</strong>пределял<strong>о</strong><strong>с</strong>ь<br />

<strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>й <strong>с</strong><strong>о</strong><strong>в</strong>ершенных п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ных н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong><br />

и шир<strong>о</strong><strong>к</strong>их <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong><strong>в</strong>ших<strong>с</strong>я переп<strong>о</strong>нч<strong>а</strong>тых<br />

з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong>. Эт<strong>о</strong>т <strong>о</strong>тряд не<strong>с</strong><strong>о</strong>мненн<strong>о</strong><br />

иллю<strong>с</strong>трирует пример п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

<strong>с</strong> <strong>о</strong>тряд<strong>о</strong>м же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых.<br />

ОТРЯД PROTELYTROPTERA<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

<strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>м Ст<strong>а</strong>р<strong>о</strong>г<strong>о</strong><br />

и Н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> С<strong>в</strong>ет<strong>а</strong>, из них <strong>в</strong> СССР три р<strong>о</strong>д<strong>а</strong>.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин р<strong>о</strong>д.<br />

SemenoviolaMartynov, 1925. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. obliquotruncata Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>. Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>-Т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылья<br />

д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong>,<br />

<strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти зн<strong>а</strong>чительный<br />

з<strong>а</strong><strong>к</strong>ругленный <strong>в</strong>ы<strong>с</strong>туп н<strong>а</strong>ружу, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>уж<strong>а</strong>ют<strong>с</strong>я. Члени<strong>к</strong>и <strong>а</strong>нтенн <strong>о</strong>дн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны,<br />

длин<strong>а</strong> их немн<strong>о</strong>г<strong>о</strong> б<strong>о</strong>льше ширины. Длин<strong>а</strong> тел<strong>а</strong><br />

8,5 мм; н<strong>а</strong>иб<strong>о</strong>льш<strong>а</strong>я ширин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий 1,5 мм<br />

(ри<strong>с</strong>. 269). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО FORFICULIDAE STEPHENS, 1829<br />

Средних р<strong>а</strong>змер<strong>о</strong><strong>в</strong> и <strong>к</strong>рупных, <strong>к</strong>рыл<strong>а</strong>тые<br />

и бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы 1<br />

. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 10—-15,<br />

цилиндриче<strong>с</strong><strong>к</strong>ие. Вт<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширен. Генит<strong>а</strong>льный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>а</strong>мц<strong>а</strong><br />

<strong>с</strong> <strong>о</strong><strong>к</strong>ругленными н<strong>а</strong> <strong>в</strong>ершине мет<strong>а</strong>п<strong>а</strong>р<strong>а</strong>мер<strong>а</strong>ми<br />

(ри<strong>с</strong>. 270 и 271). Мез<strong>о</strong>з<strong>о</strong>й — ныне. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них четыре р<strong>о</strong>д<strong>а</strong> из<br />

третичных <strong>о</strong>тл<strong>о</strong>жений Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

Вне СССР: Mesoforficula Ping, 1928 (мел<br />

Кит<strong>а</strong>я).<br />

Ри<strong>с</strong>. 272—273. Отряд Protelytroptera<br />

272. Protelytron perminum Tillyard (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x 5,8,<br />

н. Пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1933). 273. Uralelytron martynovi<br />

Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, x 7,5, н. пермь, Ур<strong>а</strong>л (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1939)<br />

115 8*<br />

http://jurassic.ru/

СЕМЕЙСТВО PROTELYTRIDAE TILLYARD, 1931<br />

Переднее <strong>к</strong>рыл<strong>о</strong> п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>с</strong> чет<strong>к</strong><strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>йм<strong>о</strong>й; RS х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, без<br />

<strong>в</strong>ет<strong>в</strong>ей. Н. пермь. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Uralelytron Rohdendorf, 1939. Тип р<strong>о</strong>д<strong>а</strong><br />

— U. tnartynovi Rohdendorf, 1939; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). RS<br />

Д<strong>в</strong>е п<strong>а</strong>ры х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итых <strong>к</strong>рылье<strong>в</strong>,<br />

<strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е леж<strong>а</strong>щих пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong> н<strong>а</strong> <strong>с</strong>пине. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; RS рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т R, и ег<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>и идут <strong>к</strong> <strong>в</strong>ершине и з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>;<br />

М <strong>о</strong>бр<strong>а</strong>зует д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и, ред<strong>к</strong><strong>о</strong> пять;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>лы МА и MP <strong>с</strong>л<strong>а</strong>б<strong>о</strong> дифференцир<strong>о</strong><strong>в</strong><strong>а</strong>ны;<br />

CuA CuP<br />

ОТРЯД PROTOBLATTODEA<br />

<strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; М,<br />

Си и А нея<strong>с</strong>ные; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong><strong>о</strong>гнутый.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,5 мм, ширин<strong>а</strong> 2 мм<br />

(ри<strong>с</strong>. 273). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Вне СССР: Protelytron Tillyard, 1931;<br />

Protelytropsis Tillyard, 1931; Permelytropsis<br />

Carpenter 1933.<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

Ри<strong>с</strong>. 274—282. Отряд Protoblattodea<br />

274. Aenigmatella comparabilis Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,9,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 275. Asiopompus<br />

tomicus Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,3, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 276. Sindon uralensis Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 6,5, н. пермь* Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 277. Sindonopsis<br />

subcostalis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>," х 5,8, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). 278. Euryptilon blattoides Martynov;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 4,0,^н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, ,1940). 279.<br />

http://jurassic.ru/<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть передних <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>тделен<strong>а</strong><br />

глуб<strong>о</strong><strong>к</strong><strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й, <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й <strong>в</strong>перед, н<strong>а</strong> дне<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й пр<strong>о</strong>х<strong>о</strong>дит CuP. Между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми имеют<strong>с</strong>я ред<strong>к</strong><strong>а</strong>я <strong>с</strong>еть — <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>а</strong>рхеди<strong>к</strong>тия<br />

или пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итым <strong>а</strong>н<strong>а</strong>льным <strong>в</strong>еер<strong>о</strong>м.<br />

Euryptilodes cascus Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,5, н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 280. Epimastax parvulus<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,5, <strong>в</strong>. Пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1928). 281. Stenoneura fayoli Brongniart; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 1,6, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925). 282. Anrgetus<br />

cubitalis Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 0,9, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1911)

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong>перед и не п<strong>о</strong><strong>к</strong>рыт<strong>а</strong><br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й. Грудь и брюш<strong>к</strong><strong>о</strong> шир<strong>о</strong><strong>к</strong>ие,<br />

упл<strong>о</strong>щенные д<strong>о</strong>р<strong>с</strong><strong>о</strong>-<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>; н<strong>о</strong>ги бег<strong>а</strong>тельные,<br />

т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>ньег<strong>о</strong> тип<strong>а</strong>, <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итым <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>им<br />

<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льным члени<strong>к</strong><strong>о</strong>м, <strong>к</strong>рупным<br />

бедр<strong>о</strong>м и г<strong>о</strong>ленью, <strong>в</strong><strong>о</strong><strong>о</strong>руженными <strong>с</strong>ильными<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными шип<strong>а</strong>ми; длинн<strong>а</strong>я пятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>, пер<strong>в</strong>ый члени<strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й н<strong>а</strong>иб<strong>о</strong>льший,<br />

<strong>с</strong>ледующий з<strong>а</strong> ним п<strong>о</strong> <strong>в</strong>еличине —<br />

пятый, н<strong>а</strong>именьший — чет<strong>в</strong>ертый. С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> яйце<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м,<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим д<strong>а</strong>ле<strong>к</strong><strong>о</strong> з<strong>а</strong> <strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. 22 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Aenigmatodidae,<br />

Asiopompidae, Blattinopsidae, Euryptilonidae,<br />

Epimastacidae, Stenoneuridae, Eoblattidae,<br />

Aetophlebiidae, Asyncritidae, Epideigmatidae,<br />

Cheliphlebiidae, Eucaenidae, Gerapompidae,<br />

Adiphlebiidae, Anthracothremmidae, Taeneopteridae?,<br />

Protophasmidae, Fayoliellidae?, Stenoneurellidae,<br />

Anthracoptilidae, Prostenoneuridae?,<br />

KHveriidae?, из них т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пер<strong>в</strong>ые пять <strong>в</strong> предел<strong>а</strong>х<br />

СССР.<br />

СЕМЕЙСТВО AENIGMATODIDAE<br />

HANDLIRSCH, 1906<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е. R без <strong>в</strong>ет<strong>в</strong>ей<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. RS <strong>о</strong>бр<strong>а</strong>зует <strong>о</strong>дну — три<br />

<strong>в</strong>ет<strong>в</strong>и. Имеют<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и,<br />

т<strong>а</strong><strong>к</strong> и уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и <strong>с</strong> ред<strong>к</strong><strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми. К<strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Aenigmatella Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. comparabilis Sharov, 1961; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Желтый Яр). В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле 15—18 <strong>в</strong>ет<strong>в</strong>ей<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>ты, <strong>о</strong>тх<strong>о</strong>дящих <strong>о</strong>т п<strong>о</strong><strong>с</strong>ледней п<strong>о</strong>д угл<strong>о</strong>м<br />

М т<strong>а</strong><strong>к</strong>же <strong>с</strong>ж<strong>а</strong>т<strong>а</strong>я и <strong>о</strong>бр<strong>а</strong>зует немн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей;<br />

СиА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з или п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>; Ai <strong>к</strong>реп<strong>к</strong><strong>а</strong>я,<br />

Аг дре<strong>в</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я. Пермь. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>.<br />

Euryptilon Martynov, 1940. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Е. blattoides Martynov, 1940; н. пермь. Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> уз<strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем, без<br />

<strong>в</strong>ет<strong>в</strong>ей SC; RS <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями. М из<strong>о</strong>гнут<strong>а</strong><br />

S-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong>лит<strong>а</strong> <strong>с</strong> СиА; СиА п<strong>о</strong><strong>с</strong>ле <strong>о</strong>тх<strong>о</strong>ждения<br />

<strong>о</strong>т М <strong>о</strong>бр<strong>а</strong>зует три пр<strong>о</strong><strong>с</strong>тые п<strong>а</strong>р<strong>а</strong>ллельные<br />

<strong>в</strong>ет<strong>в</strong>и, <strong>а</strong> з<strong>а</strong>тем д<strong>в</strong><strong>а</strong>жды дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; Ai д<strong>в</strong>у- или трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 11 мм (ри<strong>с</strong>. 278; т<strong>а</strong>бл. V,<br />

фиг- 1). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Euryptilodes Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Е. cascus Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льны'м п<strong>о</strong>лем, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ыми <strong>в</strong>ет<strong>в</strong>ями SC <strong>в</strong> нем. RS н<strong>а</strong><br />

<strong>с</strong>ередине <strong>с</strong><strong>в</strong><strong>о</strong>ей длины р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е<br />

<strong>в</strong>ет<strong>в</strong>и, <strong>к</strong><strong>а</strong>жд<strong>а</strong>я из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых, п<strong>о</strong>дх<strong>о</strong>дя <strong>к</strong> <strong>в</strong>ершине<br />

<strong>к</strong>рыл<strong>а</strong>, <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> три <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие жил<strong>к</strong>и. А] не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я и пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> CuP.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 16 мм (ри<strong>с</strong>! 279;<br />

т<strong>а</strong>бл. V, фиг. 2). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>, <strong>о</strong>дин из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых —<br />

Euryptilodes'? horridus Sharov, 1961—<strong>о</strong>пи<strong>с</strong><strong>а</strong>н<br />

п<strong>о</strong> нимфе. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО EPIMASTACIDAE MARTYNOV, 1928<br />

Переднее <strong>к</strong>рыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>о</strong>е;<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong> <strong>с</strong>емью-<strong>в</strong><strong>о</strong><strong>с</strong>емью <strong>в</strong>ет<strong>в</strong>ями <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R <strong>в</strong> <strong>с</strong>редней<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC,<br />

ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; СиА н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м<br />

пр<strong>о</strong>тяжении <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

б<strong>о</strong>льш<strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

(пять-ше<strong>с</strong>ть) <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>. Пермь. Один<br />

р<strong>о</strong>д.<br />

Epimastax Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Е. parvulus Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>. Тихие Г<strong>о</strong>ры). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. Н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> RS <strong>с</strong>ближен<strong>о</strong> <strong>с</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong>м<br />

пер<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и R, идущей <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; между SC и R ше<strong>с</strong>ть п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>;<br />

Ai пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

11 мм (ри<strong>с</strong>. 280). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО STENONEURIDAE HANDLIRSCH, 1906<br />

Передние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, <strong>с</strong> туп<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я<br />

жил<strong>к</strong><strong>а</strong>ми гребня SC; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

R близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>; СиА <strong>о</strong>бширн<strong>а</strong>я;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я,<br />

<strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м (15—16) <strong>в</strong>ет<strong>в</strong>ей А 2, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я; между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми т<strong>о</strong>н<strong>к</strong><strong>а</strong>я ячеи<strong>с</strong>т<strong>а</strong>я <strong>с</strong>еть. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 40—50 мм (ри<strong>с</strong>. 281). К<strong>а</strong>рббн.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО EOBLATTIDAE HANDLIRSCH, 1906<br />

В переднем <strong>к</strong>рыле <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле б<strong>о</strong>лее<br />

уз<strong>к</strong><strong>о</strong>е, чем у предыдущег<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и<br />

SC <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я; между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми<br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong><br />

<strong>с</strong>емь-<strong>в</strong><strong>о</strong><strong>с</strong>емь. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

60 мм (ри<strong>с</strong>. 282). К<strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong>не СССР.<br />

НАДОТРЯД PLECOPTEROIDEA. В Е С Н Я Н К О О Б Р А 3 Н Ы Е<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних <strong>к</strong>рылье<strong>в</strong> з<strong>а</strong>метн<strong>о</strong><br />

мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong>и <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>: передняя <strong>в</strong>ет<strong>в</strong>ь<br />

р<strong>а</strong>ди<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>в</strong> <strong>в</strong>иде<br />

R, з<strong>а</strong>дние <strong>в</strong>ет<strong>в</strong>и — <strong>в</strong> <strong>в</strong>иде RS. Ан<strong>о</strong>юг<strong>а</strong>льный<br />

<strong>в</strong>еер з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> умереных р<strong>а</strong>змер<strong>о</strong><strong>в</strong> и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ями <strong>а</strong>н<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong>. Гетер<strong>о</strong>н<strong>о</strong>мия <strong>к</strong>рылье<strong>в</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>,<br />

<strong>о</strong>бычн<strong>о</strong> лишь <strong>в</strong> р<strong>а</strong><strong>с</strong>ширении <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>еер<strong>а</strong>; пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> м<strong>а</strong>л<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong>т пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и з<strong>а</strong>днег<strong>о</strong>. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мия <strong>к</strong>рылье<strong>в</strong>, бе<strong>с</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ть <strong>о</strong>чень<br />

ред<strong>к</strong><strong>а</strong>. Тел<strong>о</strong> удлиненн<strong>о</strong>е, <strong>в</strong><strong>а</strong>л<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>е, умеренн<strong>о</strong><br />

упл<strong>о</strong>щенн<strong>о</strong>е, н<strong>о</strong>ги бег<strong>а</strong>тельные. Р<strong>а</strong>з<strong>в</strong>итие ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

бы<strong>в</strong><strong>а</strong>ет <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>ми, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых жи<strong>в</strong>ут<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

118<br />

http://jurassic.ru/<br />

неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелые <strong>с</strong>т<strong>а</strong>дии, реже — <strong>с</strong> п<strong>о</strong>ч<strong>в</strong><strong>о</strong>й. Преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong>ымерш<strong>а</strong>я групп<strong>а</strong>: <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне имеют<strong>с</strong>я пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители лишь д<strong>в</strong>ух<br />

<strong>о</strong>тряд<strong>о</strong><strong>в</strong> (эмбий и <strong>в</strong>е<strong>с</strong>нян<strong>о</strong><strong>к</strong>), б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

других — п<strong>о</strong>чти и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие<br />

ф<strong>о</strong>рмы. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>и <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны <strong>с</strong> примити<strong>в</strong>ными<br />

т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными, близ<strong>к</strong>ими <strong>к</strong> <strong>о</strong>тряду<br />

Protoblattodea. И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие <strong>в</strong>е<strong>с</strong>нян<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

<strong>о</strong>пределил<strong>о</strong><strong>с</strong>ь, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, р<strong>а</strong>з<strong>в</strong>итием<br />

<strong>в</strong><strong>о</strong>дн<strong>о</strong>г<strong>о</strong> или п<strong>о</strong>ч<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бр<strong>а</strong>з<strong>а</strong> жизни<br />

неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых <strong>с</strong>т<strong>а</strong>дий. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Пять <strong>о</strong>тряд<strong>о</strong><strong>в</strong>: Paraplecoptera, Embioptera, Plecoptera,<br />

Miomoptera, Caloneurodea.

Д<strong>в</strong>е п<strong>а</strong>ры х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итых <strong>к</strong>рылье<strong>в</strong>. В переднем<br />

<strong>к</strong>рыле SC <strong>с</strong> пр<strong>а</strong><strong>в</strong>ильным ряд<strong>о</strong>м <strong>к</strong><strong>о</strong><strong>с</strong>ых<br />

<strong>в</strong>ет<strong>в</strong>ей, идущих <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong><br />

и ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я. Ст<strong>в</strong><strong>о</strong>л М р<strong>а</strong>н<strong>о</strong> (<strong>о</strong>бычн<strong>о</strong><br />

д<strong>о</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS) <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, н<strong>а</strong> д<strong>в</strong>е,<br />

н<strong>о</strong> ин<strong>о</strong>гд<strong>а</strong> и н<strong>а</strong> три <strong>в</strong>ет<strong>в</strong>и. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й пр<strong>о</strong>х<strong>о</strong>дит CuP,<br />

<strong>о</strong>бычн<strong>о</strong> не <strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я. А 2 <strong>о</strong>бр<strong>а</strong>зует неб<strong>о</strong>льш<strong>о</strong>е<br />

чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей. В з<strong>а</strong>днем <strong>к</strong>рыле М т<strong>а</strong><strong>к</strong>же <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; Ai пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

А 2 гребенч<strong>а</strong>т<strong>а</strong>я.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, тел<strong>о</strong> удлиненн<strong>о</strong>е, упл<strong>о</strong>щенн<strong>о</strong>е,<br />

д<strong>о</strong>р<strong>с</strong><strong>о</strong>-<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>. Крылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е<br />

леж<strong>а</strong>т пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong> н<strong>а</strong> <strong>с</strong>пине. Н<strong>о</strong>ги бег<strong>а</strong>тельные,<br />

<strong>с</strong> ред<strong>к</strong>ими шип<strong>а</strong>ми; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

пять. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит яйце<strong>к</strong>л<strong>а</strong>д.<br />

Цер<strong>к</strong>и длинные, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

К<strong>а</strong>рб<strong>о</strong>н — юр<strong>а</strong>. Ше<strong>с</strong>ть н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Ideliidea,<br />

Cacurgidea, Geraridea, Strephocladidea, Liomopteridea,<br />

Megakhosaridea.<br />

НАДСЕМЕЙСТВО IDELIIDEA<br />

Передние <strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, ред<strong>к</strong><strong>о</strong><br />

переп<strong>о</strong>нч<strong>а</strong>тые, шир<strong>о</strong><strong>к</strong>ие, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным<br />

<strong>а</strong>рхеди<strong>к</strong>тием либ<strong>о</strong> <strong>с</strong> б<strong>о</strong>лее или<br />

менее р<strong>а</strong>зреженн<strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми, <strong>с</strong><strong>о</strong>хр<strong>а</strong>няющей ячеи<strong>с</strong>т<strong>о</strong>е <strong>с</strong>тр<strong>о</strong>ение;<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и (е<strong>с</strong>ли <strong>о</strong>ни имеют<strong>с</strong>я)<br />

<strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я лишь <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>к</strong>рыл<strong>а</strong>, реже — н<strong>а</strong> б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>бычн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

<strong>в</strong>ет<strong>в</strong>ями SC, между <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми или<br />

<strong>с</strong>еть <strong>а</strong>рхеди<strong>к</strong>тия, или п<strong>о</strong>перечные жил<strong>к</strong>и, реже<br />

<strong>о</strong>н<strong>о</strong> уз<strong>к</strong><strong>о</strong>е. R д<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong><br />

и <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бр<strong>а</strong>зует ряд <strong>к</strong><strong>о</strong><strong>с</strong>ых <strong>в</strong>ет<strong>в</strong>ей,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>.<br />

CuA <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> переднему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>, з<strong>а</strong>тем <strong>к</strong>рут<strong>о</strong> п<strong>о</strong><strong>в</strong><strong>о</strong>р<strong>а</strong>чи<strong>в</strong><strong>а</strong>ет <strong>к</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю, <strong>о</strong>бр<strong>а</strong>зуя «г<strong>о</strong>рб», <strong>о</strong>т <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дит не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей,<br />

упир<strong>а</strong>ющих<strong>с</strong>я <strong>в</strong> CuP или идущих п<strong>о</strong>д не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым<br />

угл<strong>о</strong>м <strong>к</strong> CuP; н<strong>а</strong>иб<strong>о</strong>лее пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь CuA не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей;<br />

CuP прям<strong>а</strong>я или <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>в</strong>ытянут<strong>а</strong>я.<br />

Тел<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong> шир<strong>о</strong><strong>к</strong>ими п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми;<br />

н<strong>о</strong>ги бег<strong>а</strong>тельные, <strong>с</strong> <strong>к</strong>рупными, <strong>к</strong><strong>а</strong><strong>к</strong><br />

у т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>, или <strong>с</strong>редней <strong>в</strong>еличины <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льными<br />

члени<strong>к</strong><strong>а</strong>ми. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> длинный яйце<strong>к</strong>л<strong>а</strong>д.<br />

ОТРЯД PARAPLECOPTERA<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

119<br />

http://jurassic.ru/<br />

К<strong>а</strong>рб<strong>о</strong>н — три<strong>а</strong><strong>с</strong>. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Ideliidae,<br />

Archiprobnisidae, Camptoneuritidae, Demopteridae,<br />

Chelopteridae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних — <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО IDELIIDAE М. ZALESSKY, 1929<br />

(Rachimentomidae G. Zalessky, 1939)<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

<strong>в</strong>ет<strong>в</strong>ями SC, мн<strong>о</strong>гие из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я. Между <strong>в</strong>ет<strong>в</strong>ями RS и МА нет <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>о</strong><strong>в</strong>;<br />

МА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

<strong>к</strong>рыл<strong>а</strong>; MP ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; н<strong>а</strong>иб<strong>о</strong>льшее<br />

чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей <strong>о</strong>бр<strong>а</strong>зует CuA; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет <strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти. К<strong>а</strong>рб<strong>о</strong>н — три<strong>а</strong><strong>с</strong>. 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Kortshakolia Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. ideliformis Sharov, 1961; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>в</strong>ерхнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>о</strong>рч<strong>а</strong><strong>к</strong><strong>о</strong>л).<br />

Переднее <strong>к</strong>рыл<strong>о</strong> удлиненн<strong>о</strong>е; RS <strong>о</strong>тх<strong>о</strong>дит<br />

п<strong>о</strong>чти п<strong>о</strong><strong>с</strong>редине длины <strong>к</strong>рыл<strong>а</strong>. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми п<strong>о</strong>перечными<br />

или <strong>о</strong>бр<strong>а</strong>зующими <strong>о</strong>тдельные д<strong>в</strong><strong>о</strong>йные ячей<strong>к</strong>и<br />

жил<strong>к</strong><strong>а</strong>ми; <strong>в</strong>ет<strong>в</strong>ей CuA не б<strong>о</strong>лее ше<strong>с</strong>ти. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 мм (ри<strong>с</strong>. 283). Один<br />

<strong>в</strong>ид. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Sylvldelia Martynov, 1940. Тип р<strong>о</strong>д<strong>а</strong> —•<br />

S. latipennis Martynov, 1940; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Передние<br />

<strong>к</strong>рылья <strong>с</strong> мел<strong>к</strong>им <strong>а</strong>рхеди<strong>к</strong>тием; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е; R п<strong>о</strong>чти прям<strong>а</strong>я;<br />

д<strong>о</strong> <strong>с</strong>еми <strong>в</strong>ет<strong>в</strong>ей RS. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм (ри<strong>с</strong>. 284). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Rachimentomon G. Zalessky, 1939. Тип<br />

р<strong>о</strong>д<strong>а</strong> — R. reticulatum G. Zalessky, 1939;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Передние <strong>к</strong>рылья <strong>с</strong> мел<strong>к</strong>им <strong>а</strong>рхеди<strong>к</strong>тием;<br />

RS з<strong>а</strong>метн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong>перед; <strong>в</strong>ет<strong>в</strong>ей RS<br />

четыре. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 26 мм (ри<strong>с</strong>.<br />

285; т<strong>а</strong>бл. VI, фиг. 1). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Archidelia Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. elongata Sharov, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. Между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми ред<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong>еть п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными <strong>о</strong>дин или д<strong>в</strong><strong>а</strong><br />

р<strong>а</strong>з<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я жил<strong>к</strong><strong>а</strong>ми гребня <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>ты.<br />

Суб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> д<strong>в</strong><strong>о</strong>йным или тр<strong>о</strong>йным<br />

ряд<strong>о</strong>м ячее<strong>к</strong>; CuA <strong>о</strong>бр<strong>а</strong>зует б<strong>о</strong>льше ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> б<strong>о</strong>лее 50 мм (ри<strong>с</strong>.<br />

286; т<strong>а</strong>бл. V, фиг. 3). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.

Ри<strong>с</strong>. 283—294. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Ideliidea<br />

283. Kortshakolta ideliformls Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, x 3, 3,<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 284. Sylvidelia<br />

latipennis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,3, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1940). 285. Rachimentomon reticulatum G. Zalessky; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 1,9, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1939).<br />

286. Archldella ovata Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,4, н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 287. Aenigmidelia incredibilis<br />

Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,4, н. Пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 288. Parldella pusilla Sharov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 2,2, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 289. Meti-<br />

120<br />

http://jurassic.ru/<br />

della kargalensis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,1, пермь, Ю. Приур<strong>а</strong>лье<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 290. Stenaropodites permiakovae (M.<br />

Zalessky); переднее <strong>к</strong>рыл<strong>о</strong>, х 1,7, <strong>в</strong>. пермь Приур<strong>а</strong>лье (М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1929). 291 Permotermopsls rosenl Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 1,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937).<br />

292. Kolvidelia curia G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х2,0, <strong>в</strong>. Пермь,<br />

Приур<strong>а</strong>лье (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1950). 293. Archiprobnls repens Sharov;<br />

А — переднее <strong>к</strong>рыл<strong>о</strong>, х 3,4, Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,4, н. Пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 294. Camptoneurites reticulatus<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,3, <strong>в</strong>. пермь, Приур<strong>а</strong>лье<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). ^

Aenigmidelia Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. incredibilis Sharov, 1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. Между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми<br />

— ред<strong>к</strong><strong>а</strong>я <strong>с</strong>еть п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Имеет<strong>с</strong>я<br />

л<strong>о</strong>жн<strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я из<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong> SC; SC <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS;<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля <strong>с</strong> тр<strong>о</strong>йным<br />

ряд<strong>о</strong>м ячее<strong>к</strong>; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> имеют<strong>с</strong>я<br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50 мм (ри<strong>с</strong>. 287; т<strong>а</strong>бл. V,<br />

фиг. 4). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Paridelia Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> — Р. риsilla<br />

Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. Между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные; д<strong>в</strong>урядные<br />

ячей<strong>к</strong>и имеют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля, <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле, «<strong>а</strong><br />

неб<strong>о</strong>льш<strong>о</strong>м уч<strong>а</strong><strong>с</strong>т<strong>к</strong>е меди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>убмеди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и <strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>лей; MP<br />

<strong>о</strong>бр<strong>а</strong>зует менее пяти <strong>в</strong>ет<strong>в</strong>ей; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> б<strong>о</strong>льш<strong>а</strong>я, удлиненн<strong>а</strong>я; между<br />

Ai и Д<strong>а</strong> три ряд<strong>а</strong> ячее<strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 мм (ри<strong>с</strong>. 288). Один <strong>в</strong>ид. Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Metidelia Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М. kargalensis Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> уз<strong>к</strong><strong>о</strong>е; между<br />

пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми ред<strong>к</strong><strong>а</strong>я <strong>с</strong>ет<strong>к</strong><strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зующ<strong>а</strong>я<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> пр<strong>а</strong><strong>в</strong>ильные<br />

<strong>с</strong><strong>о</strong>т<strong>о</strong><strong>в</strong>идные ячей<strong>к</strong>и; <strong>в</strong>ет<strong>в</strong>и SC пр<strong>о</strong><strong>с</strong>тые; МА<br />

д<strong>в</strong>ух-, трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

24—28 мм (ри<strong>с</strong>. 289). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; пермь Приур<strong>а</strong>лья.<br />

Stenaropodites М <strong>а</strong> г t у п <strong>о</strong> v, 1928 (Idelta<br />

М. Zalessky, 1929). Тип р<strong>о</strong>д<strong>а</strong> — 5. reticulata<br />

Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР<br />

(<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры). Сет<strong>к</strong><strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

мел<strong>к</strong><strong>а</strong>я, приближ<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я <strong>к</strong> <strong>а</strong>рхеди<strong>к</strong>тию; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>ет<strong>в</strong>и SC<br />

<strong>в</strong>ильч<strong>а</strong>тые; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и зн<strong>а</strong>чительн<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ления МА и <strong>о</strong>бр<strong>а</strong>зует<br />

четыре-пять п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; МА<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я гребенч<strong>а</strong>т<strong>о</strong>, <strong>о</strong>бр<strong>а</strong>зуя д<strong>о</strong> четырех <strong>в</strong>ет<strong>в</strong>ей;<br />

MP не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

45 мм (ри<strong>с</strong>. 290; т<strong>а</strong>бл. VI, фиг. 2). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

B. пермь Приур<strong>а</strong>лья.<br />

Permotermopsis Martynov, 1937 Тип р<strong>о</strong>д<strong>а</strong><br />

— P. roseni Martynov, 1937; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Сеть<br />

<strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>зрежен<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле менее шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

чем у предыдущег<strong>о</strong> р<strong>о</strong>д<strong>а</strong>, н<strong>о</strong> шире, чем<br />

121<br />

http://jurassic.ru/<br />

у Metidelia; SC зн<strong>а</strong>чительн<strong>о</strong> не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong><br />

<strong>в</strong>ершины <strong>к</strong>ры'л<strong>а</strong>, <strong>в</strong>ет<strong>в</strong>и ее без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>; R<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>ей RS ше<strong>с</strong>ть-<strong>с</strong>емь; MP <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 47—55 мм<br />

(ри<strong>с</strong>. 291). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Kolvidelia G. Zalessky, 1955. Тип р<strong>о</strong>д<strong>а</strong><br />

— К- curta G. Zalessky, 1955; пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (П<strong>о</strong><strong>к</strong>ч<strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

<strong>в</strong>ет<strong>в</strong>и SC пр<strong>о</strong><strong>с</strong>тые; R <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; MP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 мм (ри<strong>с</strong>. 292).<br />

Один <strong>в</strong>ид. Пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Вне СССР: Austroidelia Riek, 1954 (три<strong>а</strong><strong>с</strong><br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии).<br />

СЕМЕЙСТВО ARCHIPROBNISIDAE SHAROV, 1961<br />

Передние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем. Суб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле т<strong>а</strong><strong>к</strong>же<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> пр<strong>а</strong><strong>в</strong>ильным ряд<strong>о</strong>м пр<strong>о</strong><strong>с</strong>тых или <strong>с</strong><br />

неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ильч<strong>а</strong>тых п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>,<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненных <strong>к</strong> <strong>в</strong>ершине. П<strong>о</strong>ля между<br />

R, RS, МА, MP и CuA <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ие и<br />

з<strong>а</strong>п<strong>о</strong>лнены пр<strong>а</strong><strong>в</strong>ильными ряд<strong>а</strong>ми преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

пр<strong>о</strong><strong>с</strong>тых, близ<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных друг<br />

<strong>к</strong> другу п<strong>а</strong>р<strong>а</strong>ллельных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>.<br />

Чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей МА и MP не<strong>в</strong>ели<strong>к</strong><strong>о</strong>, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляя<br />

<strong>в</strong>ме<strong>с</strong>те не б<strong>о</strong>лее пяти-ше<strong>с</strong>ти. От <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

М <strong>к</strong> CuA <strong>о</strong>тх<strong>о</strong>дит <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong>, <strong>с</strong><strong>о</strong>зд<strong>а</strong>ющ<strong>а</strong>я<br />

<strong>в</strong>печ<strong>а</strong>тление еще <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и М, <strong>с</strong>ли<strong>в</strong>шей<strong>с</strong>я <strong>с</strong><br />

CuA. Вет<strong>в</strong>и CuA <strong>с</strong>мещены <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>, те<strong>с</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены друг <strong>к</strong> другу, причем<br />

<strong>к</strong><strong>а</strong>жд<strong>а</strong>я из них <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я 2 или 3 р<strong>а</strong>з<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зуя<br />

<strong>в</strong> <strong>о</strong>бщей <strong>с</strong>л<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти <strong>о</strong>т <strong>с</strong>еми д<strong>о</strong> трин<strong>а</strong>дц<strong>а</strong>ти<br />

<strong>в</strong>ет<strong>в</strong>ей. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong>чень неб<strong>о</strong>льш<strong>а</strong>я.<br />

Ai пр<strong>о</strong>х<strong>о</strong>дит близ<strong>к</strong><strong>о</strong> <strong>к</strong> CuP; А 2 <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong>д<strong>в</strong>инут<strong>а</strong> <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю и <strong>о</strong>бр<strong>а</strong>зует ше<strong>с</strong>ть —<br />

<strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их, <strong>с</strong>л<strong>а</strong>бых <strong>в</strong>ет<strong>о</strong>че<strong>к</strong>. Один р<strong>о</strong>д.<br />

Archiprobnis Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. repens Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й.<br />

Ячей<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми, уз<strong>к</strong>ие и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>ие.' RS <strong>о</strong>бр<strong>а</strong>зует д<strong>о</strong><br />

пяти <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей; между Ai и А 2 д<strong>в</strong><strong>а</strong> или<br />

три ряд<strong>а</strong> ячее<strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 20—<br />

22 мм (ри<strong>с</strong>. 293). Три <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО CAMPTONEURITIDAE MARTYNOV, 1928<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>в</strong><br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети <strong>к</strong>рыл<strong>а</strong>; имеют<strong>с</strong>я <strong>а</strong>н<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>зы<br />

между <strong>в</strong>ет<strong>в</strong>ями RS, МА и MP близ <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>; CuA <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине,<br />

<strong>с</strong>д<strong>в</strong>инут<strong>а</strong> <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. Пермь. Один<br />

р<strong>о</strong>д.

Camptoneurltes Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong><br />

— С. reticulatus Martynov, 1928; <strong>в</strong>. пермь,<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Передние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>уженные<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>с</strong>ет<strong>к</strong><strong>а</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>зрежен<strong>а</strong>,<br />

<strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и. Вет<strong>в</strong>и RS, МА и MP <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

прямые, п<strong>а</strong>р<strong>а</strong>ллельные пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и<br />

<strong>к</strong>рыл<strong>а</strong>; А] ч<strong>а</strong><strong>с</strong>т<strong>о</strong> д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; четыре или б<strong>о</strong>лее<br />

<strong>в</strong>ет<strong>в</strong>ей А 2. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

16 мм (ри<strong>с</strong>. 294; т<strong>а</strong>бл. VI, фиг. 3). Один <strong>в</strong>ид.<br />

В. пермь Приур<strong>а</strong>лья.<br />

НАДСЕМЕЙСТВО CACURGIDEA<br />

Крылья <strong>о</strong>бычн<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>тые, реже — <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые,<br />

удлиненные; <strong>в</strong>ершин<strong>а</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>трен<strong>а</strong>. В<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и<br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. М<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS, <strong>о</strong>бр<strong>а</strong>зуя<br />

МА и MP; п<strong>о</strong><strong>с</strong>ледняя <strong>в</strong>ет<strong>в</strong>ь <strong>о</strong>чень р<strong>а</strong>н<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> CuA. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong><br />

жил<strong>к</strong>и <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я, п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong> жил<strong>к</strong><strong>а</strong>м Neuroptera.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширены <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

и ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуют неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>еер. (К<strong>а</strong>рб<strong>о</strong>н—три<strong>а</strong><strong>с</strong>?)<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Paoliidae, Stygneidae,<br />

Cacurgidae (ри<strong>с</strong>. 295), Homalophlebiidae<br />

(Anhomalophlebiidae), Narkemidae, Thoronysidae,<br />

Pachytylopsidae, Protokollariidae, Prototettigidae,<br />

Coseliidae, Omaliidae, Stenaroceridae?<br />

Mixotermitidae, Laspeyresiellidae, Adeloneuridae?,<br />

Klebsiellidae?, Homoeodictyidae, Triassomanteidae?,<br />

Xenopteridae?, Mesotitanidae?; <strong>в</strong><strong>с</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>, <strong>к</strong>р<strong>о</strong>ме Narkemidae и Homoeodictyidae,<br />

<strong>в</strong>не СССР.<br />

СЕМЕЙСТВО NARKEMIDAE<br />

HANDLIRSCH, 1911<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> R; д<strong>в</strong>е-три пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные<br />

<strong>в</strong>ет<strong>в</strong>и RS п<strong>а</strong>р<strong>а</strong>ллельны друг другу и не<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я; МА <strong>о</strong>бр<strong>а</strong>зует д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и;<br />

передняя <strong>в</strong>ет<strong>в</strong>ь MP <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> МА или <strong>с</strong> ее<br />

з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>ью; <strong>с</strong>т<strong>в</strong><strong>о</strong>л MP п<strong>о</strong><strong>с</strong>ле <strong>о</strong>тх<strong>о</strong>ждения<br />

<strong>о</strong>т CuA р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>, <strong>а</strong> между МА, MP +<br />

CuA и MP имеет<strong>с</strong>я шир<strong>о</strong><strong>к</strong><strong>о</strong>е з<strong>а</strong>м<strong>к</strong>нут<strong>о</strong>е п<strong>о</strong>ле.<br />

В ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми — пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные. К<strong>а</strong>рб<strong>о</strong>н.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Narkemina Martynov, 1930. Тип р<strong>о</strong>д<strong>а</strong> —<br />

N. angustata Martynov, 1930; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

<strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Верх<strong>о</strong>т<strong>с</strong>м<strong>с</strong><strong>к</strong><strong>о</strong>е). RS берет<br />

н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>в</strong>е-три пр<strong>о</strong><strong>с</strong>тые п<strong>а</strong>р<strong>а</strong>ллельные <strong>в</strong>ет<strong>в</strong>и, идущие<br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, <strong>а</strong> з<strong>а</strong>тем <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и; <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зует 9—11 <strong>в</strong>ет<strong>в</strong>ей;<br />

МА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е п<strong>а</strong>р<strong>а</strong>ллельные жил<strong>к</strong>и приблизительн<strong>о</strong><br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>в</strong>п<strong>а</strong>дения SC <strong>в</strong> R. От<br />

122<br />

http://jurassic.ru/<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> MP <strong>о</strong>тх<strong>о</strong>дят три —<br />

пять <strong>в</strong>ет<strong>в</strong>ей. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

40 мм (ри<strong>с</strong>. 296 и 297; т<strong>а</strong>бл. VI, фиг. 4). Три<br />

<strong>в</strong>ид<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Narkema Handlirsch, 1911.<br />

СЕМЕЙСТВО HOMOEODICTYIDAE<br />

MARTYNOV, 1937<br />

Передние <strong>к</strong>рылья уз<strong>к</strong>ие, длинные, <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женным <strong>а</strong>рхеди<strong>к</strong>тием; SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й; R <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м н<strong>а</strong> <strong>к</strong><strong>о</strong>нце;<br />

CuP т<strong>а</strong><strong>к</strong>же <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; Ai д<strong>в</strong><strong>а</strong>жды дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. Пермь. Один р<strong>о</strong>д.<br />

Homoeodictyon Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Н. elongatum Martynov, 1937; пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Передние <strong>к</strong>рылья<br />

<strong>к</strong>рупные, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>с</strong>емь-<strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>к</strong><strong>о</strong><strong>с</strong>ых <strong>в</strong>ет<strong>в</strong>ей; <strong>в</strong>ет<strong>в</strong>ей<br />

RS пять. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60 мм<br />

(ри<strong>с</strong>. 298). Один <strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

НАДСЕМЕЙСТВО GERARIDEA<br />

Крылья уз<strong>к</strong>ие, удлиненные; RS или CuA<br />

(ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong> RS и CuA) гребенч<strong>а</strong>тые;<br />

М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS,<br />

<strong>о</strong>бр<strong>а</strong>зуя неб<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

передних <strong>к</strong>рылье<strong>в</strong> уз<strong>к</strong><strong>а</strong>я, <strong>с</strong> немн<strong>о</strong>гими<br />

(не б<strong>о</strong>лее четырех) <strong>а</strong>н<strong>а</strong>льными жил<strong>к</strong><strong>а</strong>ми;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong><br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> и не <strong>о</strong>бр<strong>а</strong>зует <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>г<strong>о</strong> <strong>в</strong>еер<strong>а</strong>; <strong>с</strong>ет<strong>к</strong><strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. Тел<strong>о</strong><br />

уз<strong>к</strong><strong>о</strong>е, <strong>в</strong>ытянут<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, <strong>с</strong> <strong>в</strong>ыд<strong>а</strong>ющими<strong>с</strong>я<br />

<strong>в</strong>перед м<strong>а</strong>ндибул<strong>а</strong>ми. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>в</strong> длину <strong>в</strong><strong>с</strong>егд<strong>а</strong> б<strong>о</strong>льше, чем <strong>в</strong> ширину, ин<strong>о</strong>гд<strong>а</strong><br />

чрез<strong>в</strong>ыч<strong>а</strong>йн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> удлиненн<strong>а</strong>я; ее б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

<strong>к</strong>р<strong>а</strong>я (п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>ли) <strong>о</strong>чень уз<strong>к</strong>ие или нез<strong>а</strong>метны;<br />

Ин<strong>о</strong>гд<strong>а</strong> передние н<strong>о</strong>ги <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты и<br />

пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти. Яйце<strong>к</strong>л<strong>а</strong>д<br />

длинный. К<strong>а</strong>рб<strong>о</strong>н — пермь. Де<strong>с</strong>ять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Geraridae, Spanioderidae, Cnemidolestidae,<br />

Hapalopteridae, Cymenophlebiidae, Ischnoneuridae,<br />

Hadentomidae?, Emphylopteridae, KHveriidae,<br />

Tillyardembiidae; <strong>в</strong><strong>с</strong>е, <strong>к</strong>р<strong>о</strong>ме п<strong>о</strong><strong>с</strong>леднег<strong>о</strong>,<br />

<strong>в</strong>не СССР.<br />

СЕМЕЙСТВО GERARIDAE<br />

HANDLIRSCH, 1906<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> С; RS гребенч<strong>а</strong>т<strong>а</strong>я,<br />

<strong>о</strong>бширн<strong>а</strong>я, <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R близ<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>рыл<strong>а</strong>; М х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть CuA <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong>ж<strong>а</strong>т<strong>а</strong>. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 40—65 мм<br />

(ри<strong>с</strong>. 299). Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>в</strong>не СССР.

30i<br />

Ри<strong>с</strong>. 295—304. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Cacurgidea, Geraridea, Strephocladidea<br />

295. Heterologus langfordorum Carpenter; переднее <strong>к</strong>рыл<strong>о</strong>, x 0,8,<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1943). 296. Narkemina angustata<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, x 2,1, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 297. Narkemina angustiformis Sharov;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 1,8, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961). 298. Homoeodictyon elongatumMartynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 0,9, Ю. Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 299. Gerarus danielsi<br />

Handlirsch (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x 0,9, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

(Handlirsch, 1925). 300. Spanindera ambulans Handlirsch (ре<strong>к</strong><strong>о</strong>н­<br />

300<br />

<strong>с</strong>тру<strong>к</strong>ция), x 1,0, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1925).<br />

301. Cnemidolestes woodwardi Brongniart (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x 0,6,<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925). 302. Tillyardembia brevipes<br />

(Martynov), и. пермь, Ур<strong>а</strong>л; A — переднее <strong>к</strong>рыл<strong>о</strong>, х 4,8<br />

(<strong>о</strong>риг. ри<strong>с</strong>.); Б — <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 5,3 (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940).<br />

303. Mesorthopetron locustoides Tillyard; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,4<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1922). 304. Strephoneura robvsia<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,4. н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1940)<br />

123<br />

http://jurassic.ru/

СЕМЕЙСТВО SPANIODER1DAE HANDLIRSCH, 1906<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> R; <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть M <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong>ж<strong>а</strong>т<strong>а</strong>; CuA х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, гребенч<strong>а</strong>т<strong>а</strong>я.<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 26—35 мм (ри<strong>с</strong>.300).<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Б<strong>о</strong>лее де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО CNEMIDOLESTIDAE HANDLIRSCH, 1906<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> С; CuA <strong>о</strong>бширн<strong>а</strong>я.<br />

Передние н<strong>о</strong>ги хищные, зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>льше<br />

<strong>с</strong>редних и з<strong>а</strong>дних, д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> <strong>к</strong>реп<strong>к</strong>их шип<strong>о</strong><strong>в</strong> н<strong>а</strong><br />

<strong>в</strong>нутренней п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти бедр<strong>а</strong> и г<strong>о</strong>лени. Длин<strong>а</strong><br />

передних <strong>к</strong>рылье<strong>в</strong> 50—90 мм (ри<strong>с</strong>. 301).<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО TILLYARDEMBIIDAEG. ZALESSKY, 1938<br />

(Permocapniidae Martynov, 1940)<br />

Не б<strong>о</strong>лее д<strong>в</strong>ух <strong>в</strong>ет<strong>в</strong>ей М; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong><br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> CuA п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует ряд гребенч<strong>а</strong>т<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>к</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> умеренн<strong>о</strong><br />

удлиненн<strong>а</strong>я; бедр<strong>а</strong> передних н<strong>о</strong>г <strong>о</strong>бычн<strong>о</strong> <strong>в</strong>здутые.<br />

Пермь. Один р<strong>о</strong>д,<br />

Tillyardembia G. Zalessky, 1937. (Permocapnia<br />

Martynov, 1938). Тип р<strong>о</strong>д<strong>а</strong> — Т. biarmi<strong>с</strong><strong>а</strong><br />

G. Zalessky, 1937; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong><br />

удлиненн<strong>о</strong>-<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>в</strong>ершин<strong>а</strong> <strong>с</strong>ужен<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей<br />

RS не менее пяти; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й<br />

л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идн<strong>о</strong>й ф<strong>о</strong>рмы. Средне- и з<strong>а</strong>днегрудь<br />

<strong>в</strong> длину б<strong>о</strong>льше, чем <strong>в</strong> ширину, прям<strong>о</strong>уг<strong>о</strong>льные.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 11—12,5 мм<br />

(ри<strong>с</strong>. 302 А, Б). Четыре <strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

НАДСЕМЕЙСТВО STREPHOCLADIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; <strong>в</strong>ет<strong>в</strong>и RS, М и<br />

CuA длинные, п<strong>а</strong>р<strong>а</strong>ллельные и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

не <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я <strong>в</strong>близи <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; CuA гребне<strong>в</strong>идн<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong>перед; передняя <strong>в</strong>ет<strong>в</strong>ь CuA<br />

п<strong>о</strong>чти д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н —<br />

три<strong>а</strong><strong>с</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Strephoneuridae, Strephocladidae<br />

и Mesorthopteridae? (ри<strong>с</strong>. 303); из них<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО STREPHONEURIDAE MARTYNOV, 1940<br />

SC дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти; R <strong>с</strong> гребнем <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их жил<strong>о</strong><strong>к</strong>; тричетыре<br />

гребне<strong>в</strong>идн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong>ет<strong>в</strong>и RS;<br />

<strong>в</strong>ет<strong>в</strong>и гребня CuA пр<strong>о</strong><strong>с</strong>тые. Пермь. Один<br />

р<strong>о</strong>д.<br />

Strephoneura Martynov, 1940. Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. robusta Martynov, 1940; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

124<br />

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Передние <strong>к</strong>рылья уз<strong>к</strong>ие, длинные; SC д<strong>о</strong>х<strong>о</strong>дит<br />

д<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трети <strong>к</strong>рыл<strong>а</strong>; между <strong>в</strong>ет<strong>в</strong>ями<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие п<strong>о</strong>перечные жил<strong>к</strong>и; R <strong>с</strong>ильн<strong>а</strong>я,<br />

прям<strong>а</strong>я; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> CuA и <strong>с</strong><strong>о</strong>единен<strong>а</strong><br />

<strong>с</strong> нею <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й;<br />

<strong>в</strong>ет<strong>в</strong>и М и CuA <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong><br />

не <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 50 мм<br />

(ри<strong>с</strong>. 304). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

НАДСЕМЕЙСВО LIOMOPTER1DEA<br />

Передние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льные, ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й, <strong>о</strong>бычн<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>тые,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> или п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью п<strong>о</strong><strong>к</strong>рытые<br />

ч<strong>а</strong><strong>с</strong>тыми мел<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, реже без<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. М, з<strong>а</strong> ред<strong>к</strong>им и<strong>с</strong><strong>к</strong>лючением, н<strong>а</strong>чин<strong>а</strong>ет<br />

<strong>в</strong>ет<strong>в</strong>ить<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS, причем<br />

MP не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я и не <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong><br />

CuA. Н<strong>а</strong>иб<strong>о</strong>лее пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь CuA,<br />

<strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>ем<strong>а</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> СиА 2, п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong><br />

<strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей CuA и пр<strong>о</strong>х<strong>о</strong>дит<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> CuP. Пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>к</strong>рыл<strong>а</strong>. И<strong>с</strong><strong>к</strong>лючение <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют лишь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Phenopteridae и Sylvaphlebiidae, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых имеют н<strong>а</strong> б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong><br />

д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд ячее<strong>к</strong> между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми. З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленным<br />

<strong>о</strong>бширным <strong>а</strong>н<strong>а</strong>льным <strong>в</strong>еер<strong>о</strong>м.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, переднегрудь <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми (п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>ми),<br />

пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ющими<strong>с</strong>я н<strong>а</strong> передний и з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>я передне<strong>с</strong>пин<strong>к</strong>и и <strong>о</strong>бр<strong>а</strong>зующими <strong>к</strong><strong>о</strong>льц<strong>о</strong><br />

<strong>в</strong><strong>о</strong><strong>к</strong>руг <strong>с</strong><strong>о</strong>б<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> н<strong>о</strong>тум<strong>а</strong> (т<strong>а</strong>бл. VI, фиг. 5—<br />

6); <strong>в</strong> этих л<strong>о</strong>п<strong>а</strong><strong>с</strong>тях ин<strong>о</strong>гд<strong>а</strong> н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я темные<br />

пр<strong>о</strong>жил<strong>к</strong>и, н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющие жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рылье<strong>в</strong>. Яйце<strong>к</strong>л<strong>а</strong>д у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченный, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нный. К<strong>а</strong>рб<strong>о</strong>н — юр<strong>а</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Palaeocixiidae, Roomeriidae, Reculidae,<br />

Phenopteridae, Stereopteridae, Probnisidae, Protembiidae,<br />

Germanopriscidae, Chaulioditidae?,<br />

Liomopteridae, Stegopteridae, Euremiscidae, Sylvaphlebidae,<br />

Lemmatophoridae, Atactophlebiidae,<br />

Tomiidae, Geinitziidae; из них пер<strong>в</strong>ые де<strong>в</strong>ять<br />

<strong>в</strong>не СССР.<br />

СЕМЕЙСТВО LIOMOPTERIDAE SELLARDS, 1909<br />

(Khosaridae Martynov, 1937 (pars);<br />

Climaconeuridae Handlirsch, 1919)<br />

Крылья переп<strong>о</strong>нч<strong>а</strong>тые, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> или п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

п<strong>о</strong><strong>к</strong>рытые мел<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; ин<strong>о</strong>гд<strong>а</strong><br />

имеет<strong>с</strong>я ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> из темных п<strong>о</strong>л<strong>о</strong><strong>с</strong> и пятен.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; MP д<strong>в</strong>ух-, реже<br />

трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Ai <strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, н<strong>о</strong> ин<strong>о</strong>гд<strong>а</strong>

Ри<strong>с</strong>. 305—310. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridae<br />

305. Parapertnula sibirica Sharov; A — переднее <strong>к</strong>рыл<strong>о</strong>, x 2,4;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,4; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961). 306. Abashevia suchovl Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,1, пермь,<br />

Ку ец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 307.^Climaconeurites asiaticus<br />

Sharov; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 3,2; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,2,<br />

н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, _1961). 308. Ornaticosta<br />

magna Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,8, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 309. Liomopterella vulgaris Sharov; А —<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 3,6; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,6; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 310. Kaltanella lata Sharov;<br />

А — переднее <strong>к</strong>рыл<strong>о</strong>, х 3,5; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,5, н. Пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)<br />

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<strong>о</strong>бр<strong>а</strong>зует н<strong>а</strong> <strong>к</strong><strong>о</strong>нце р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; н<strong>а</strong> А 2 не менее<br />

четырех <strong>в</strong>ет<strong>в</strong>ей. В ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>,<br />

з<strong>а</strong> и<strong>с</strong><strong>к</strong>лючением п<strong>о</strong>ля между R и RS, <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. К<strong>а</strong>рб<strong>о</strong>н —<br />

пермь. 19 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Parapermula Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. sibirica Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> яйце<strong>в</strong>идн<strong>о</strong>-<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ы'пу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>с</strong> пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными <strong>в</strong> нем, <strong>о</strong>бычн<strong>о</strong><br />

пр<strong>о</strong><strong>с</strong>тыми <strong>в</strong>ет<strong>в</strong>ями SC; <strong>о</strong>дн<strong>а</strong> или д<strong>в</strong>е жил<strong>к</strong>и<br />

<strong>с</strong>реди них ин<strong>о</strong>гд<strong>а</strong> бы<strong>в</strong><strong>а</strong>ют д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми.<br />

Суб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е и <strong>с</strong><strong>о</strong>держит<br />

10—15 пр<strong>о</strong><strong>с</strong>тых п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> д<strong>в</strong>умя ряд<strong>а</strong>ми<br />

ячее<strong>к</strong>, между <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми имеют<strong>с</strong>я <strong>о</strong>тдельные<br />

из<strong>о</strong>лир<strong>о</strong><strong>в</strong><strong>а</strong>нные ячей<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующие <strong>к</strong><strong>а</strong><strong>к</strong> бы<br />

третий ряд. RS <strong>о</strong>бр<strong>а</strong>зует <strong>о</strong>т д<strong>в</strong>ух д<strong>о</strong> ше<strong>с</strong>ти<br />

<strong>в</strong>ет<strong>в</strong>ей, МА — три — пять, MP — д<strong>в</strong>е-три,<br />

СиА — три — пять <strong>в</strong>ет<strong>в</strong>ей. Крылья ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> ри<strong>с</strong>ун<strong>к</strong><strong>о</strong>м<br />

из <strong>к</strong>рупных пятен и темных п<strong>о</strong>л<strong>о</strong><strong>с</strong> п<strong>о</strong><br />

жил<strong>к</strong><strong>а</strong>м. З<strong>а</strong>дние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им<br />

р<strong>а</strong>ди<strong>а</strong>льным п<strong>о</strong>лем. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

27—31 мм (ри<strong>с</strong>. 305; т<strong>а</strong>бл. VII, фиг. 1). Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Abashevia Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong>—А.<br />

suchovi Sharov, 1961; <strong>в</strong> пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Аб<strong>а</strong>ше<strong>в</strong><strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины, <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>в</strong><br />

б<strong>о</strong>лее пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льных ч<strong>а</strong><strong>с</strong>тях; рез<strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>ругляя<strong>с</strong>ь<br />

у <strong>с</strong><strong>а</strong>м<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния, <strong>о</strong>н<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зует неб<strong>о</strong>льшую<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть. Пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь CuAi<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я ближе <strong>к</strong> н<strong>а</strong>ч<strong>а</strong>лу СиА 2<br />

и пр<strong>о</strong>х<strong>о</strong>дит п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> п<strong>о</strong><strong>с</strong>ледней. Длин<strong>а</strong><br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 27 мм (ри<strong>с</strong>. 306).<br />

Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Climaconeurites Sharov, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — С. asiaticus Sharov, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Ди<strong>с</strong>т<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и SC<br />

и R рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнуты н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>. Вершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> туп<strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>а</strong>я.<br />

RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>рыл<strong>а</strong> и <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>в</strong>ершины.<br />

Передние <strong>в</strong>ет<strong>в</strong>и МА <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ершине<br />

<strong>к</strong>рыл<strong>а</strong>. Е<strong>с</strong>ть д<strong>в</strong><strong>а</strong> гл<strong>а</strong><strong>в</strong>ных <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> МА, передний<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>бр<strong>а</strong>зует н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м уч<strong>а</strong><strong>с</strong>т<strong>к</strong>е<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>с</strong> RS; ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние переднег<strong>о</strong><br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> МА уп<strong>о</strong>д<strong>о</strong>бляет<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>е, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> <strong>с</strong><strong>о</strong>зд<strong>а</strong>ет<strong>с</strong>я <strong>в</strong>печ<strong>а</strong>тление,<br />

чт<strong>о</strong> RS <strong>о</strong>чень р<strong>а</strong>н<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>. Вет<strong>в</strong>и А 2 <strong>в</strong> чи<strong>с</strong>ле четырех-пяти<br />

длинные и <strong>к</strong>реп<strong>к</strong>ие. В з<strong>а</strong>дних <strong>к</strong>рыльях МА н<strong>а</strong><br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 21—22 мм (ри<strong>с</strong>. 307; т<strong>а</strong>бл. VII,<br />

126<br />

http://jurassic.ru/<br />

фиг. 2). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ornaticosta Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> — О.<br />

magna Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>с</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем. Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>с</strong>ильн<strong>о</strong> пигментир<strong>о</strong><strong>в</strong><strong>а</strong>н <strong>в</strong> <strong>в</strong>иде'<br />

<strong>с</strong>пл<strong>о</strong>шн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>ы. RS гребенч<strong>а</strong>тый; MP из<strong>о</strong>гнут<strong>а</strong><br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. Вет<strong>в</strong>и CuAi идут<br />

д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>перед, п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. А 2 <strong>о</strong>бр<strong>а</strong>зует <strong>о</strong>т пяти д<strong>о</strong> <strong>с</strong>еми длинных<br />

п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей. В з<strong>а</strong>днем <strong>к</strong>рыле.<br />

МА не <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> RS. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 30—50 мм (ри<strong>с</strong>. 308). Четыре<strong>в</strong>ид<strong>а</strong>.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Liomopterella Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

L. vulgaris Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Передние<br />

<strong>к</strong>рылья <strong>в</strong>ытянутые, <strong>с</strong> умеренн<strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шире р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>,<br />

между <strong>в</strong>ет<strong>в</strong>ями SC п<strong>о</strong>перечные жил<strong>к</strong>и. В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле и <strong>в</strong> п<strong>о</strong>ле между МА и MP д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й<br />

ряд ячее<strong>к</strong>. RS, МА, и MP <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>. Ai ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зует<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>, А 2 имеет четыре-пять <strong>в</strong>ет<strong>в</strong>ей.<br />

В з<strong>а</strong>дних <strong>к</strong>рыльях МА не <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong><br />

<strong>с</strong> RS. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 20—35 мм-<br />

(ри<strong>с</strong>. 309; т<strong>а</strong>бл. VII, фиг. 3). Три <strong>в</strong>ид<strong>а</strong>. Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kaltanella Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> — К.<br />

lata Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее <strong>к</strong>рыл<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> туп<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й. Имеют<strong>с</strong>я д<strong>в</strong><strong>а</strong><br />

гл<strong>а</strong><strong>в</strong>ных <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> МА. Д<strong>в</strong><strong>о</strong>йные ряды ячее<strong>к</strong> <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>ех<br />

п<strong>о</strong>лях <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; н<strong>а</strong>ряду <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми<br />

п<strong>о</strong>перечными имеют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> У-, Н- и<br />

Л-<strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong>перечные жил<strong>к</strong>и. В з<strong>а</strong>днем <strong>к</strong>рыле<br />

МА не <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> RS. Ан<strong>а</strong>льный<br />

<strong>в</strong>еер <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ий. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 мм (ри<strong>с</strong>. 310; т<strong>а</strong>бл. VII, фиг. 4).<br />

Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Liomopterites Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

L. expletus Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). В переднем<br />

<strong>к</strong>рыле <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уже р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и <strong>с</strong><strong>о</strong>держит 15—25 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей SC, между<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. RS<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>о</strong>дну — четыре <strong>в</strong>ет<strong>в</strong>и, МА д<strong>в</strong>е-три,<br />

MP <strong>о</strong>дну — три, СиА три — пять <strong>в</strong>ет<strong>в</strong>ей; А] не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я, А 2 имеет четыре — <strong>с</strong>емь <strong>в</strong>ет<strong>в</strong>ей. Крыл<strong>о</strong><br />

п<strong>о</strong> <strong>в</strong><strong>с</strong>ей п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> ч<strong>а</strong><strong>с</strong>тыми, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. В з<strong>а</strong>днем <strong>к</strong>рыле МА<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>с</strong> RS. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 8—25 мм (ри<strong>с</strong>. 311). Нимфы <strong>с</strong>тр<strong>о</strong>йные,<br />

<strong>с</strong> неб<strong>о</strong>льшими <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными <strong>к</strong>р<strong>а</strong>ями передне<strong>с</strong>пин<strong>к</strong>и<br />

и неб<strong>о</strong>льшими <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми н<strong>а</strong> брюшных

CuP<br />

Ри<strong>с</strong>. 311—319. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridae<br />

311. Liomopterites comans Sharov; A —переднее <strong>к</strong>рыл<strong>о</strong>, x 5,0; Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1957). 316. Paraliomopterum paulum<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,0, н. Пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,5, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

1961). 312. Liomopterites gracilis Sharov; нимф<strong>а</strong>, х 4,6, н. пермь, (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 317. Khosara permiakovae Martynov; перед­<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 313. Sibirella paucinervis нее <strong>к</strong>рыл<strong>о</strong>, х 3,0, пермь, Ю. Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937).<br />

Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>­ 318. Kazanella rotundipennis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,9,<br />

р<strong>о</strong><strong>в</strong>, 1961). 314. Sarbalopterum ignorablle Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, <strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1930). 319. Liomopterum orna-<br />

х 6,1, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 315. Kaltanytum) Sellards (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x 2,9, и. пермь, С Амери<strong>к</strong><strong>а</strong><br />

mpha thysanuriformis Sharov (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x 1,7, н. пермь.<br />

(Carpenter, 1950)<br />

127<br />

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CuP

<strong>с</strong>егмент<strong>а</strong>х; цер<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>че брюш<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 312).<br />

Ше<strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Sibirella Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> — 5.<br />

paucinervis Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Передний<br />

<strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле п<strong>о</strong>чти т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е,<br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>о</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию. В <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле <strong>к</strong><strong>о</strong><strong>с</strong>ые п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

Вет<strong>в</strong>и RS, М и СиА из<strong>о</strong>гнуты <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю.<br />

Н<strong>а</strong> п<strong>о</strong>перечных жил<strong>к</strong><strong>а</strong>х темные пятн<strong>а</strong>. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 15 мм (ри<strong>с</strong>. 313). Один<br />

<strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Sarbalopterum Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. ignorabile Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>а</strong>рб<strong>а</strong>л<strong>а</strong><br />

II). Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним<br />

<strong>к</strong>р<strong>а</strong>ем. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, рез<strong>к</strong><strong>о</strong> <strong>с</strong>уж<strong>а</strong>ющее<strong>с</strong>я<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию, <strong>с</strong> пр<strong>а</strong><strong>в</strong>ильным ряд<strong>о</strong>м<br />

<strong>с</strong>ильн<strong>о</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненных <strong>в</strong>ет<strong>в</strong>ей SC. П<strong>о</strong>перечные<br />

жил<strong>к</strong>и немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны, и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм<br />

(ри<strong>с</strong>. 314). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kattanympha Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К- thysanuriformis Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Переднегрудь нимфы <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льными<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти длинные, <strong>с</strong> <strong>к</strong>рупным<br />

<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льным члени<strong>к</strong><strong>о</strong>м, р<strong>а</strong><strong>в</strong>ным п<strong>о</strong> длине<br />

бедру. Бедр<strong>а</strong> з<strong>а</strong>дних н<strong>о</strong>г <strong>с</strong> <strong>в</strong>ершинными шип<strong>а</strong>ми.<br />

Цер<strong>к</strong>и <strong>с</strong> длинными члени<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong><br />

нимфы без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> 14 мм (ри<strong>с</strong>. 315; т<strong>а</strong>бл.<br />

VII, фиг. 5). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Paraliomopterum Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. paulum Sharov, 1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> I).<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый.<br />

В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле 15—17 <strong>в</strong>ет<strong>в</strong>ей SC.<br />

В ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля<br />

д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд ячее<strong>к</strong>. RS <strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

<strong>в</strong>ет<strong>в</strong>ей. В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и, <strong>в</strong> меди<strong>а</strong>льн<strong>о</strong>м — д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд<br />

ячее<strong>к</strong>, передние из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong>рупнее и <strong>в</strong>ыше.<br />

Д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд ячее<strong>к</strong> имеет<strong>с</strong>я т<strong>а</strong><strong>к</strong>же между СиА 2<br />

и CuP. Темные пятн<strong>а</strong> преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> п<strong>о</strong><br />

п<strong>о</strong>перечным жил<strong>к</strong><strong>а</strong>м. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 мм (ри<strong>с</strong>. 316). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Khosara Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> — K.h.<br />

permiakovae Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>ым<br />

передним <strong>к</strong>р<strong>а</strong>ем. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong><br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R<br />

близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong> и <strong>о</strong>чень п<strong>о</strong>здн<strong>о</strong> <strong>в</strong>ет­<br />

128<br />

http://jurassic.ru/<br />

<strong>в</strong>ит<strong>с</strong>я, <strong>о</strong>бр<strong>а</strong>зуя д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и; МА н<strong>а</strong>чин<strong>а</strong>ет <strong>в</strong>ет<strong>в</strong>ить<strong>с</strong>я<br />

<strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>;<br />

МА, MP и CuAi <strong>о</strong>бр<strong>а</strong>зуют не б<strong>о</strong>лее чем п<strong>о</strong> д<strong>в</strong>е<br />

<strong>в</strong>ет<strong>в</strong>и. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 25 мм (ри<strong>с</strong>.<br />

317). Один <strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

Kazanella Martynov, 1930. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К- rotundipennis Martynov, 1930; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле зн<strong>а</strong>чительн<strong>о</strong><br />

шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>, рез<strong>к</strong><strong>о</strong> <strong>с</strong>уж<strong>а</strong>ющее<strong>с</strong>я<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию. В <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие п<strong>о</strong>перечные жил<strong>к</strong>и, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

перпенди<strong>к</strong>улярн<strong>о</strong> <strong>к</strong> пр<strong>о</strong>д<strong>о</strong>льным; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> ячее<strong>к</strong>; <strong>в</strong> п<strong>о</strong>ле между МА и MP<br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 17—18 мм (ри<strong>с</strong>. 318). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В.<br />

пермь Приур<strong>а</strong>лья.<br />

Вне СССР: Climaconeura Pruvost, 1912;<br />

Permula Handlirsch, 1919; Liomopterum Sellards,<br />

1909 (ри<strong>с</strong>. 319); Tapopterum Carpenter,<br />

1950; Semopterum Carpenter, 1950; Ideliopsis<br />

Carpenter, 1948.<br />

СЕМЕЙСТВО STEGOPTERIDAE SHAROV, 1961<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>е, <strong>с</strong> буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й; <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> <strong>к</strong>рыльях <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Ст<strong>в</strong><strong>о</strong>л RS и ег<strong>о</strong> <strong>в</strong>ет<strong>в</strong>и из<strong>о</strong>гнуты <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. Общий х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет<br />

т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Liomopteridae.<br />

В<strong>с</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые. Один р<strong>о</strong>д.<br />

Пермь.<br />

Stegopterum Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. hirium Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). В переднем<br />

<strong>к</strong>рыле SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong><br />

де<strong>в</strong>ятью-де<strong>с</strong>ятью <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями SC, <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле пять пр<strong>о</strong><strong>с</strong>тых п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>;<br />

RS гребенч<strong>а</strong>т<strong>а</strong>я. В з<strong>а</strong>днем <strong>к</strong>рыле RS не<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> МА. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 12—13 мм (ри<strong>с</strong>. 320). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. и <strong>в</strong>.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО EUREMISCIDAE G. ZALESSKY, 1951<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е, п<strong>о</strong>чти т<strong>а</strong><strong>к</strong><strong>о</strong>й<br />

же ширины, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

пр<strong>о</strong><strong>с</strong>тыми <strong>в</strong>ет<strong>в</strong>ями SC <strong>в</strong> нем. R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong> з<strong>а</strong>метным изгиб<strong>о</strong>м- RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R<br />

близ <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong>. МА и MP <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближены и пр<strong>о</strong>х<strong>о</strong>дят<br />

<strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>к</strong> R, п<strong>о</strong><strong>в</strong>т<strong>о</strong>ряя ее изгиб. Один р<strong>о</strong>д.<br />

Пермь.<br />

Euremisca G. Zalessky, 1951. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. splendens G. Zalessky, 1951; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>) ;<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е <strong>в</strong> длину, <strong>в</strong> <strong>с</strong>редней

328 Б<br />

Ри<strong>с</strong>. 320—328. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Stegopteridae, Euremiscidae, Sylvaphlebiidae,<br />

Lernrnatophondae, Atactophlebiidae<br />

320. Stegopterum hlrtum Sharov; A — переднее <strong>к</strong>рыл<strong>о</strong>, x 4,5;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>; х 4,5, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961). 321. Euremisca splendens G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, хЗ.О,<br />

н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1951). 322. Sylvaphlebia tuberculoma<br />

Martynov; <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 2,7, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1940). 323. Sylvlodes perloides Martynov; <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>,<br />

х 1,7, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 324. Parasylviodes<br />

tetracladus Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,8, н. пермь, Ур<strong>а</strong>л<br />

9 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 129<br />

http://jurassic.ru/<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 325. Sylvaella paurovenosa Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 4,0, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 326. Paraprlsca<br />

urallca G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,5, н. пермь, Ур<strong>а</strong>л<br />

(Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1952). 327. Lemmatophora typa Sellards (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция),<br />

х 5,5, н. пермь, С.Амери<strong>к</strong><strong>а</strong> (<strong>о</strong>риг. ри<strong>с</strong>.). 328. Atactophle-<br />

Ы<strong>а</strong> termitoldesMartynov; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 2,0; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, х 2,0; <strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928)

ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>чти <strong>с</strong> прямым передним <strong>к</strong>р<strong>а</strong>ем; между<br />

MP и CuA — д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд ячее<strong>к</strong>; <strong>в</strong> других п<strong>о</strong>лях<br />

— пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 14 мм (ри<strong>с</strong>. 321). Один <strong>в</strong>ид. Н.<br />

пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

СЕМЕЙСТВО SYLVAPHLEBHDAE MARTYNOV. 1940<br />

(Sylviodidae Martynov, 1940<br />

Sylvaelidae Martynov, 1940)<br />

Передние <strong>к</strong>рылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong><br />

<strong>к</strong>рыльях <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е.<br />

Вет<strong>в</strong>и CuA те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ж<strong>а</strong>ты; п<strong>о</strong>ле между<br />

СиА 2 и СиР б<strong>о</strong>льше п<strong>о</strong>ля между CuAi и СиА 2.<br />

Четыре р<strong>о</strong>д<strong>а</strong>. Пермь.<br />

Sylvaphlebia Martynov, 1940 (Biarmopteron<br />

G. Zalessky, 1953). Тип р<strong>о</strong>д<strong>а</strong> —S. tuberculata<br />

Martynov, 1940; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> непр<strong>а</strong><strong>в</strong>ильными <strong>к</strong><strong>о</strong><strong>с</strong>ыми<br />

<strong>в</strong>ет<strong>в</strong>ями SC, <strong>с</strong><strong>о</strong>единенными п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми, <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле ч<strong>а</strong><strong>с</strong>тые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и н<strong>а</strong><strong>к</strong>л<strong>о</strong>нены <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong> и<br />

<strong>с</strong><strong>о</strong>единены мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными жил<strong>к</strong><strong>а</strong>ми между<br />

ними. Между <strong>в</strong>ет<strong>в</strong>ями RS, МА и MP — п<strong>о</strong> д<strong>в</strong><strong>а</strong><br />

ряд<strong>а</strong> пр<strong>а</strong><strong>в</strong>ильных <strong>с</strong><strong>о</strong>т<strong>о</strong><strong>в</strong>идных ячее<strong>к</strong>. П<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льные<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти неб<strong>о</strong>льшие. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 15—20 мм (ри<strong>с</strong>. 322). Один <strong>в</strong>ид. Н.<br />

пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Sylviodes Martynov, 1940 (Biarmopterites<br />

G. Zalessky, 1953). Тип р<strong>о</strong>д<strong>а</strong> — S. perloides<br />

Martynov, 1940; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

<strong>в</strong>ет<strong>в</strong>и SC пр<strong>о</strong><strong>с</strong>тые, <strong>с</strong> ред<strong>к</strong><strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й или<br />

пр<strong>о</strong><strong>с</strong>тыми п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми между ними.<br />

В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле — ред<strong>к</strong><strong>а</strong>я <strong>с</strong>еть п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти неб<strong>о</strong>льшие.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 24—26 мм (ри<strong>с</strong>. 323;<br />

т<strong>а</strong>бл. VII, фиг. 6). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Parasylviodes Martynov, 1940. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. tetracladus Martynov, 1940; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

мн<strong>о</strong>гие <strong>в</strong>ет<strong>в</strong>и SC <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong>т R п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>, гребенч<strong>а</strong>т<strong>а</strong>я;<br />

п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти шир<strong>о</strong><strong>к</strong>ие.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 16 мм (ри<strong>с</strong>. 324).<br />

Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Sylvaella Martynov, 1940. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. paurovenosa Martynov, 1940; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong> и <strong>о</strong>бр<strong>а</strong>зует не б<strong>о</strong>лее д<strong>в</strong>ух <strong>в</strong>ет<strong>в</strong>ей;<br />

МА и MP не <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я. Неб<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> пр<strong>о</strong>­<br />

130<br />

http://jurassic.ru/<br />

<strong>с</strong>тых п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

неб<strong>о</strong>льшие. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10 мм<br />

(ри<strong>с</strong>. 325). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

СЕМЕЙСТВО LEMMATOPHORIDAE SELLARDS, 1909<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле передних <strong>к</strong>рылье<strong>в</strong> уз<strong>к</strong><strong>о</strong>е,<br />

не шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; RS <strong>о</strong>бр<strong>а</strong>зует не б<strong>о</strong>лее<br />

трех <strong>в</strong>ет<strong>в</strong>ей, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. Чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей<br />

МА, MP и CuA т<strong>а</strong><strong>к</strong>же не<strong>в</strong>ели<strong>к</strong><strong>о</strong>. А 2 <strong>о</strong>бр<strong>а</strong>зует<br />

не б<strong>о</strong>лее трех <strong>в</strong>ет<strong>в</strong>ей, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не (<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. В переднем<br />

и з<strong>а</strong>днем <strong>к</strong>рыле — ред<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. В з<strong>а</strong>дних<br />

<strong>к</strong>рыльях А 2 <strong>с</strong> м<strong>а</strong>лым чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей (не б<strong>о</strong>лее<br />

ше<strong>с</strong>ти), <strong>о</strong>бр<strong>а</strong>зующих<strong>с</strong>я гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>. Крылья п<strong>о</strong><strong>к</strong>рыты ч<strong>а</strong><strong>с</strong>тыми,<br />

мел<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным<br />

яйце<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м, не <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим з<strong>а</strong> <strong>к</strong><strong>о</strong>нец<br />

брюш<strong>к</strong><strong>а</strong>. Пермь. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Paraprisca Handlirsch, 1919. Тип р<strong>о</strong>д<strong>а</strong><br />

— Prisca fragilis Sellards, 1909; н. .пермь,<br />

С. Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле п<strong>о</strong>чти <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> уже <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

RS берет н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> <strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

МА <strong>о</strong>бычн<strong>о</strong> не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я, реже<br />

д<strong>в</strong>у<strong>в</strong>ег<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, MP д<strong>в</strong>у-трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Цер<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие — <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че брюш<strong>к</strong><strong>а</strong>. Н<strong>о</strong>ги<br />

т<strong>о</strong>н<strong>к</strong>ие, <strong>о</strong>чень длинные. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

10—18 мм (ри<strong>с</strong>. 326). Три <strong>в</strong>ид<strong>а</strong>. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong> и С. Амери<strong>к</strong>и.<br />

Вне СССР: Artinska Sellards, 1909; Lemmatophora<br />

Sellards, 1909 (ри<strong>с</strong>. 327; т<strong>а</strong>бл. VII,<br />

фиг. 7); Lisca Sellards, 1909; Lecorium Sellards,<br />

1909.<br />

СЕМЕЙСТВО ATACTOPHLEBIIDAE MARTYNOV, 1930<br />

Мел<strong>к</strong>ие <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> <strong>к</strong>рыльях <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле передних <strong>к</strong>рылье<strong>в</strong> приблизительн<strong>о</strong><br />

т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е,<br />

<strong>с</strong> ред<strong>к</strong>ими, <strong>с</strong>л<strong>а</strong>быми <strong>в</strong>ет<strong>в</strong>ями CuA, между <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле и ин<strong>о</strong>гд<strong>а</strong> между пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льными<br />

<strong>в</strong>ет<strong>в</strong>ями CuA — д<strong>в</strong><strong>о</strong>йные ряды ячее<strong>к</strong>,<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льных ч<strong>а</strong><strong>с</strong>тях <strong>к</strong>рыл<strong>а</strong> — пр<strong>о</strong><strong>с</strong>тые или<br />

Y-, Н- и К-<strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong>перечные жил<strong>к</strong>и. М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

н<strong>а</strong> три гл<strong>а</strong><strong>в</strong>ных <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, <strong>с</strong>редний из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong><strong>о</strong>гнутый (и<strong>с</strong>тинный MP?). Чи<strong>с</strong>л<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ей RS, М и CuA <strong>с</strong>ильн<strong>о</strong> <strong>в</strong><strong>а</strong>рьирует, причем<br />

у<strong>в</strong>еличение чи<strong>с</strong>л<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ей <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>теме<br />

<strong>с</strong><strong>о</strong>пряжен<strong>о</strong> <strong>с</strong> уменьшением <strong>в</strong> друг<strong>о</strong>й. Ст<strong>в</strong><strong>о</strong>лы<br />

М и CuA, <strong>а</strong> т<strong>а</strong><strong>к</strong>же их <strong>в</strong>ет<strong>в</strong>и ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>зы. Яйце<strong>к</strong>л<strong>а</strong>д неб<strong>о</strong>льш<strong>о</strong>й, н<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющий<br />

з<strong>а</strong> <strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong>. Пермь. Один р<strong>о</strong>д.<br />

Atactophelibia Martynov, 1928- Тип<br />

р<strong>о</strong>д<strong>а</strong> —A. termitoides Martynov, 1928; <strong>в</strong>. пермь,

Ри<strong>с</strong>. 329—336. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomopteridea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tomiidae, Geinitziidae, incertae familiae<br />

329. Kargalella subcostalts Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, x 4,0,<br />

пермь, Ю. Приур<strong>а</strong>лье (<strong>о</strong>риг. ри<strong>с</strong>.). 330. Tomla costalis Martynov;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 3,0, три<strong>а</strong><strong>с</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>о</strong>риг.<br />

ри<strong>с</strong>.). 331. Shurabia ovata Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,5,<br />

и. юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 332. Aetophleblopsls jusca<br />

G. Zalessky; <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 1,6, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1955). 333. Khosarophlebla sylvaensls Martynov; з<strong>а</strong>днее<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие<br />

Г<strong>о</strong>ры). Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле н<strong>а</strong>иб<strong>о</strong>лее<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong>. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. RS берет н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong><br />

<strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й трети <strong>к</strong>рыл<strong>а</strong> и <strong>о</strong>бр<strong>а</strong>зует неб<strong>о</strong>льш<strong>о</strong>е<br />

чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей. Для А. termitoides <strong>о</strong>тмечены<br />

линь<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong>тых <strong>о</strong><strong>с</strong><strong>о</strong>бей (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1957). Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм (ри<strong>с</strong>. 328; т<strong>а</strong>бл. VIII,<br />

фиг. 1). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО TOMIIDAE MARTYNOV, 1936<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>с</strong> пр<strong>а</strong><strong>в</strong>ильным ряд<strong>о</strong>м пр<strong>о</strong><strong>с</strong>тых <strong>в</strong>ет<strong>в</strong>ей SC. SC<br />

пр<strong>о</strong>х<strong>о</strong>дит <strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>к</strong> R, и <strong>с</strong><strong>а</strong>м<strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ее <strong>в</strong>ет<strong>в</strong>ь рез<strong>к</strong><strong>о</strong> п<strong>о</strong><strong>в</strong><strong>о</strong>р<strong>а</strong>чи<strong>в</strong><strong>а</strong>ет <strong>о</strong>т R <strong>к</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю, уп<strong>о</strong>д<strong>о</strong>бляя<strong>с</strong>ь п<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жению<br />

б<strong>о</strong>лее пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льным <strong>в</strong>ет<strong>в</strong>ям. В ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля — пр<strong>а</strong><strong>в</strong>ильный ряд<br />

пр<strong>о</strong><strong>с</strong>тых <strong>в</strong>ет<strong>в</strong>ей R, я<strong>в</strong>ляющий<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> бы пр<strong>о</strong>д<strong>о</strong>л­<br />

333<br />

<strong>к</strong>рыл<strong>о</strong>, х 2,4, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 334. Karealodes<br />

incerta: Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,3, <strong>в</strong>. пермь. Ю. При'<br />

ур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,1937). 335. Haplopterum majus Martynov;<br />

?чfi H<br />

^ e<br />

^;^ PЫЛ<br />

; 0<br />

• / w<br />

, ! ,<br />

eu-<br />

п е<br />

Р<br />

м ь<br />

- Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928).<br />

33b. Мitmovia dubla Sharov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,3, <strong>в</strong>, пермь<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961) '<br />

жением ряд<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ей SC. Пермь — три<strong>а</strong><strong>с</strong>. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>.<br />

Kargalella Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. subcostalis Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

<strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем. В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле — пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и; SC<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> и <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>о</strong> де<strong>с</strong>яти <strong>в</strong>ет<strong>в</strong>ей; RS <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>с</strong> МА; МА, MP, и CuA<br />

<strong>в</strong>ме<strong>с</strong>те <strong>о</strong>бр<strong>а</strong>зуют не б<strong>о</strong>лее <strong>с</strong>еми <strong>в</strong>ет<strong>в</strong>ей- Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 16 мм (ри<strong>с</strong>. 329). Один <strong>в</strong>ид.<br />

Пермь Приур<strong>а</strong>лья.<br />

Tomia Martynov, 1936. Тип р<strong>о</strong>д<strong>а</strong> — Т. costalls<br />

Martynov, 1936; три<strong>а</strong><strong>с</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(м<strong>а</strong>льце<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Б<strong>а</strong>бий К<strong>а</strong>мень). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> удлиненн<strong>о</strong>е, <strong>с</strong><strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле, <strong>в</strong> <strong>с</strong>редней<br />

ег<strong>о</strong> ч<strong>а</strong><strong>с</strong>ти, д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд ячее<strong>к</strong>, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

и н<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> пр<strong>о</strong><strong>с</strong>тые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и; МА <strong>о</strong>бширн<strong>а</strong>я, <strong>о</strong>бр<strong>а</strong>зует д<strong>о</strong><br />

131 9*<br />

http://jurassic.ru/

четырех <strong>в</strong>ет<strong>в</strong>ей; CuAi не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 14 мм (ри<strong>с</strong>. 330). Один<br />

<strong>в</strong>ид. Три<strong>а</strong><strong>с</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО GEINITZIIDAE HANDLIRSCH, 1908<br />

Передние <strong>к</strong>рылья <strong>с</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>й<br />

<strong>с</strong>имметричн<strong>о</strong> п<strong>о</strong> <strong>о</strong>тн<strong>о</strong>шению <strong>к</strong> переднему<br />

и з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ям; R <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю и <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong>ет<strong>в</strong>и RS р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

гребенч<strong>а</strong>т<strong>о</strong> <strong>в</strong>перед и <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> переднем<br />

<strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>. Три<strong>а</strong><strong>с</strong> — юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Shurabia Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Sh. ovata Martynov, 1937; н. юр<strong>а</strong>, Cp- Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Переднее <strong>к</strong>рыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е,<br />

<strong>с</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем;<br />

R <strong>с</strong> д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; RS <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

16—17 мм (ри<strong>с</strong>. 331). Три <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии;<br />

н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: Geinitzia Handlirsch, 1906.<br />

INCERTAE FAMILIAE<br />

Aetophlebiopsis G. Z a 1 e s s k y, 1955. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. fusca G. Zalessky, 1955; н. пермь.<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> уз<strong>к</strong><strong>о</strong>е; SC<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>в</strong><strong>с</strong>ем пр<strong>о</strong>тяжении п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> R; RS<br />

н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 27 мм (ри<strong>с</strong>. 332).. Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Khosarophlebla Martynov, 1940. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Kh. sylvaensis Martynov, 1940; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем;<br />

R <strong>с</strong> пятью <strong>в</strong>ет<strong>в</strong>ями <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; МА не<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> RS. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

23 мм (ри<strong>с</strong>. 333). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Kargalodes Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. incerta Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. SC<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>;<br />

R <strong>с</strong> ше<strong>с</strong>тью-<strong>с</strong>емью длинными <strong>в</strong>ет<strong>в</strong>ями н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце; RS и МА н<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>м уч<strong>а</strong><strong>с</strong>т<strong>к</strong>е <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

15 мм (ри<strong>с</strong>. 334). Один <strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

Haplopterum Martynov, 1928. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Н. majus Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е;<br />

<strong>в</strong>ет<strong>в</strong>и R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

(<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти); МА <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS, не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я.<br />

Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 мм (ри<strong>с</strong>. 335).<br />

В. пермь Приур<strong>а</strong>лья.<br />

Mltinovia Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М. dubia Sharov, 1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Митин<strong>а</strong>). З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>. Вершин<strong>а</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>а</strong>я; <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле 10—11 <strong>в</strong>ет<strong>в</strong>ей SC; <strong>в</strong>ет<strong>в</strong>и длинные, ин<strong>о</strong>гд<strong>а</strong><br />

<strong>в</strong>ильч<strong>а</strong>тые; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м<br />

уч<strong>а</strong><strong>с</strong>т<strong>к</strong>е д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й ряд ячее<strong>к</strong>; МА не <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> RS; МА, MP и CuA имеют п<strong>о</strong><br />

д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 22 мм<br />

(ри<strong>с</strong>. 336), В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Mesonotoperla Riek, 1954; Меsacridites<br />

Riek, 1954; Xenogryllacris Riek, 1955.<br />

132<br />

http://jurassic.ru/<br />

НАДСЕМЕЙСТВО<br />

MEGAKHOSARIDEA<br />

Крылья уз<strong>к</strong>ие, длинные, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими, т<strong>о</strong>л<strong>с</strong>тыми<br />

жил<strong>к</strong><strong>а</strong>ми, рез<strong>к</strong><strong>о</strong> <strong>о</strong>тгр<strong>а</strong>ниченными <strong>о</strong>т<br />

мембр<strong>а</strong>ны. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> неб<strong>о</strong>льшим<br />

чи<strong>с</strong>л<strong>о</strong>м пр<strong>о</strong><strong>с</strong>тых п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

между С и SC. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>,<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, были н<strong>а</strong>иб<strong>о</strong>лее х<strong>о</strong>р<strong>о</strong>шими летун<strong>а</strong>ми<br />

<strong>с</strong>реди п<strong>а</strong>р<strong>а</strong>пле<strong>к</strong><strong>о</strong>птер. Пермь. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО MEGAKHOSARIDAE SHAROV, 1961<br />

[Khosaridae Martynov, 1937 (pars)] '<br />

Передние <strong>к</strong>рылья н<strong>а</strong> <strong>о</strong>тдельных уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>х<br />

п<strong>о</strong><strong>к</strong>рыты мел<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. От М <strong>к</strong> CuA, нед<strong>а</strong>ле<strong>к</strong><strong>о</strong><br />

<strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>тх<strong>о</strong>дит <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>, <strong>с</strong><strong>о</strong>зд<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong>печ<strong>а</strong>тление еще <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющей<strong>с</strong>я <strong>с</strong> CuA. CuP прям<strong>а</strong>я<br />

или <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут<strong>а</strong>я. Между CuA и CuP пр<strong>а</strong><strong>в</strong>ильный<br />

ряд <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных жил<strong>о</strong><strong>к</strong>, рез<strong>к</strong><strong>о</strong><br />

из<strong>о</strong>гнутых нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т ме<strong>с</strong>т<strong>а</strong> <strong>с</strong><strong>о</strong>единения <strong>с</strong><br />

CuP. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть б<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>я;<br />

<strong>в</strong>ет<strong>в</strong>и А 2 мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные, <strong>к</strong>реп<strong>к</strong>ие.<br />

В з<strong>а</strong>днем <strong>к</strong>рыле МА н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит <strong>с</strong> CuA.<br />

Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Megakhosara М <strong>а</strong> г t у п <strong>о</strong> v, 1937 (Syndestnophora<br />

Martynov, 1937). Тип р<strong>о</strong>д<strong>а</strong> — М. fascipennis<br />

Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. RS дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; MP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и имеют пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>е<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение, <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ующее <strong>о</strong> х<strong>о</strong>р<strong>о</strong>ших<br />

летных <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong><strong>а</strong>х <strong>к</strong>рыл<strong>а</strong>- П<strong>о</strong>перечные<br />

жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные, <strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>о</strong>лях <strong>о</strong>бр<strong>а</strong>зуют<br />

д<strong>в</strong><strong>о</strong>йные ряды ячее<strong>к</strong>. Ан<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>зы между<br />

пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т 30 д<strong>о</strong> 60 мм (ри<strong>с</strong>. 337;<br />

т<strong>а</strong>бл. VIII, фиг. 2). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; пермь Приур<strong>а</strong>лья.<br />

Megakhosarella Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М. regressa Sharov, 1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> I). Пе-

338<br />

МА<br />

337<br />

МА<br />

Ри<strong>с</strong>. 337—344. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Megakhosaridea, Paraplecoptera incertae sedis<br />

337. Megakhosara dilucldaSharov; переднее <strong>к</strong>рыл<strong>о</strong>, x 1,8, н. пермь,"<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 338. Megakhosarella regressa<br />

Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,3, <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 339. Termltoldes product us G. Zalessky; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 2,2, н. Пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1955). 340.<br />

Uralotermes permlanus G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,8, н.<br />

Пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1937). 341. Sellardslopsis consplcua<br />

реднее <strong>к</strong>рыл<strong>о</strong>. Ст<strong>в</strong><strong>о</strong>л RS н<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>м <strong>о</strong>трез<strong>к</strong>е<br />

<strong>с</strong>лит <strong>с</strong> МА. MP <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ей длины, нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

МА — MP. Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> и <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнуты. П<strong>о</strong>пе­<br />

Termitoides G. Zalessky, 1955. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. productus G. Zalessky, 1955; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Передние <strong>к</strong>рылья <strong>в</strong>ытянутые, л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 22 мм (ри<strong>с</strong>. 339). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Uralotermes G. Zalessky, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — U. permianus G. Zalessky, 1937;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

У<strong>с</strong>ть-Кишерть). Переднее <strong>к</strong>рыл<strong>о</strong> удлиненн<strong>о</strong>е;<br />

' 340<br />

G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х 5,6, н. пермь, Ур<strong>а</strong>л (Ю.'З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1939). 342. Gurianovella blattoides Martynov "'нимф<strong>а</strong>,<br />

х 3,3, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 343. Dyadozoarium pachypas<br />

Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, х 1,2, пермь, Приур<strong>а</strong>лье<br />

(Handlirsch, 1906). 344. Kaltanopierodes vanus Sharov; з<strong>а</strong>днее<br />

нимф<strong>а</strong>льн<strong>о</strong>е <strong>к</strong>рыл<strong>о</strong>, х 11,4, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий, б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961).<br />

речные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>о</strong>чень ред<strong>к</strong><strong>о</strong>, <strong>о</strong>ни<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые и лишь у <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong> м<strong>о</strong>гут быть Y-<strong>о</strong>бр<strong>а</strong>зными. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 17 мм (ри<strong>с</strong>. 338), Один <strong>в</strong>ид.<br />

В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

PARAPLECOPTERA INCERTAE SEDIS<br />

133<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>с</strong> ячеи<strong>с</strong>т<strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я близ<strong>к</strong><strong>о</strong><br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>, гребенч<strong>а</strong>т<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 20 мм (ри<strong>с</strong>. 340). Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Sellardslopsis G. Zalessky, 1939. Тип<br />

р<strong>о</strong>д<strong>а</strong> S. conspicua G. Zalessky, 1939; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

В переднем <strong>к</strong>рыле R из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> ме<strong>с</strong>те н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS; МА <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; М и CuA <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>литы; CuP длинн<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

8 мм (ри<strong>с</strong>. 341). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

http://jurassic.ru/<br />

343

Gurianovella G. Zalessky, 1939 (Kirkorella<br />

G. Zalessky, 1939; Permella G. Zalessky,<br />

1939; Czekardia Martynov, 1940; Marietta G. Zalessky,<br />

1955). Тип р<strong>о</strong>д<strong>а</strong> — G. silphidoides G. Zalessky,<br />

1939; и. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Нимф<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я,<br />

<strong>а</strong>нтенны длинные, ните<strong>в</strong>идные; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>ырез<strong>к</strong><strong>о</strong>й п<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю;<br />

тел<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е; тергиты <strong>с</strong> б<strong>о</strong>льшими п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льными<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Н<strong>а</strong> <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong> <strong>с</strong>низу<br />

— неб<strong>о</strong>льшие грифель<strong>к</strong>и. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>т 8<br />

д<strong>о</strong> 13 мм (ри<strong>с</strong>. 342; т<strong>а</strong>бл. VIII, фиг. 3). Пять<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Dyadozoarium Handlirsch, 1904. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. pachypus Handlirsch, 1904; пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Крылья уз<strong>к</strong>ие,<br />

Средней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые (длин<strong>а</strong> тел<strong>а</strong><br />

5—35 мм); д<strong>в</strong>е п<strong>а</strong>ры т<strong>о</strong>н<strong>к</strong>их переп<strong>о</strong>нч<strong>а</strong>тых<br />

<strong>к</strong>рылье<strong>в</strong>, <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong> н<strong>а</strong> <strong>с</strong>пине;<br />

передние <strong>к</strong>рылья уз<strong>к</strong>ие и длинные, передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; SC <strong>о</strong>бр<strong>а</strong>зует н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>, передняя <strong>в</strong>ет<strong>в</strong>ь <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> идет <strong>к</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю, <strong>а</strong> з<strong>а</strong>дняя — <strong>к</strong> R. Ее <strong>в</strong>ет<strong>в</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, п<strong>о</strong>х<strong>о</strong>жи н<strong>а</strong> п<strong>о</strong>перечные жил<strong>к</strong>и; RS<br />

<strong>о</strong>бычн<strong>о</strong> н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>о</strong>т <strong>о</strong>дн<strong>о</strong>й д<strong>о</strong> пяти <strong>в</strong>ет<strong>в</strong>ей RS. М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> пр<strong>о</strong>х<strong>о</strong>дит близ<strong>к</strong><strong>о</strong> <strong>к</strong> R, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь;<br />

МА р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и; MP <strong>с</strong>лит<strong>а</strong> <strong>с</strong><br />

СиА, и их <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>о</strong>чень п<strong>о</strong>здн<strong>о</strong>,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зуя не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> гребенч<strong>а</strong>тых <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей. В п<strong>о</strong>лях между МА, MP + СиА и CuP<br />

д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> пр<strong>о</strong><strong>с</strong>тых п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>, реже —<br />

д<strong>в</strong><strong>о</strong>йные ряды ячее<strong>к</strong>; CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и, е<strong>с</strong>ли их мн<strong>о</strong>г<strong>о</strong>, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>пределены.<br />

В з<strong>а</strong>дних <strong>к</strong>рыльях RS <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong><br />

<strong>с</strong> М. Ан<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong>бр<strong>а</strong>зует <strong>в</strong>еер,<br />

реже бы<strong>в</strong><strong>а</strong>ет редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>. Пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льный<br />

уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> RS <strong>о</strong>т R д<strong>о</strong> <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> М и<strong>с</strong>чез,<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> <strong>к</strong><strong>а</strong>жет<strong>с</strong>я, чт<strong>о</strong> R пр<strong>о</strong>х<strong>о</strong>дит <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong><br />

<strong>о</strong>т RS.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызущег<strong>о</strong><br />

тип<strong>а</strong>, нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>ит; <strong>а</strong>нтенны ните<strong>в</strong>идные,<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> 25—100; ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие;<br />

<strong>о</strong>бычн<strong>о</strong> имеют<strong>с</strong>я гл<strong>а</strong>з<strong>к</strong>и. Н<strong>о</strong>ги недлинные,<br />

бег<strong>а</strong>тельные; з<strong>а</strong>дняя п<strong>а</strong>р<strong>а</strong> немн<strong>о</strong>г<strong>о</strong> длиннее<br />

<strong>с</strong>редней и передней п<strong>а</strong>р; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> три.<br />

Брюш<strong>к</strong><strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е, тергиты брюш<strong>к</strong><strong>а</strong> без п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>, п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong> п<strong>о</strong> длине<br />

груди; длинные, <strong>о</strong>бычн<strong>о</strong> члени<strong>с</strong>тые цер<strong>к</strong>и. Не<br />

пит<strong>а</strong>ют<strong>с</strong>я, н<strong>о</strong> <strong>о</strong>х<strong>о</strong>тн<strong>о</strong> пьют. Обит<strong>а</strong>ют п<strong>о</strong> берег<strong>а</strong>м<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>. Яйц<strong>а</strong> <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют <strong>в</strong> <strong>в</strong><strong>о</strong>ду. Нимфы<br />

длинные, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными жил<strong>к</strong><strong>а</strong>ми. Н<strong>о</strong>ги<br />

<strong>к</strong>реп<strong>к</strong>ие, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные. Длин<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong><br />

<strong>с</strong>л<strong>о</strong>женными <strong>к</strong>рыльями 43 мм (ри<strong>с</strong>. 343). Один<br />

<strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

Kaltanopterodes Sharov, 1961- Тип<br />

р<strong>о</strong>д<strong>а</strong> — К. vanus Sharov, 1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Зеленый<br />

Луг). З<strong>а</strong>днее нимф<strong>а</strong>льн<strong>о</strong>е <strong>к</strong>рыл<strong>о</strong>. RS гребенч<strong>а</strong>т<strong>а</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зует пять <strong>в</strong>ет<strong>в</strong>ей; МА не <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> RS и не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; MP д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

СиА <strong>о</strong>бр<strong>а</strong>зует три <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>м <strong>в</strong>еере,<br />

<strong>к</strong>р<strong>о</strong>ме гребенч<strong>а</strong>т<strong>о</strong>й А 2, де<strong>в</strong>ять п<strong>а</strong>р<strong>а</strong>ллельных<br />

жил<strong>о</strong><strong>к</strong> юг<strong>а</strong>льн<strong>о</strong>й (?) <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong><strong>с</strong>е п<strong>о</strong>перечные<br />

жил<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 4,8 мм<br />

(ри<strong>с</strong>. 344). В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

ОТРЯД PLECOPTERA. ВЕСНЯНКИ<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

134<br />

http://jurassic.ru/<br />

<strong>в</strong><strong>о</strong>дные, хищные, жи<strong>в</strong>ут 2—2,5 г<strong>о</strong>д<strong>а</strong>; дыш<strong>а</strong>т<br />

тр<strong>а</strong>хейными ж<strong>а</strong>бр<strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными у<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей р<strong>а</strong>зных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> н<strong>а</strong> р<strong>а</strong>зличных<br />

уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>х тел<strong>а</strong>: н<strong>а</strong> нижней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не' груди, н<strong>а</strong><br />

передних и з<strong>а</strong>днем <strong>с</strong>егмент<strong>а</strong>х брюш<strong>к</strong><strong>а</strong>.<br />

Пермь — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Palaeoperlidae,<br />

Perlopseidae, Taeniopterygidae, Eustheniidae,<br />

Nemouridae, Leuctridae, Perlidae, Perlodidae,<br />

Pteronarcidae, Diamphipnoidae, Penturoperlidae,<br />

Senzillidae, Gripopterygidae, Scopuridae,<br />

Austroperlidae, Peltoperlidae, Capniidae, Chloroperlidae;<br />

пер<strong>в</strong>ые <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, причем Eustheniidae и<br />

Perlodidae — <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО PALAEOPERLIDAE SHAROV, 1961<br />

В <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле четыре-пять <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей SC. R <strong>с</strong> тремя-четырьмя <strong>в</strong>ет<strong>в</strong>ями<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. Имеют<strong>с</strong>я д<strong>в</strong>е п<strong>о</strong>перечные<br />

жил<strong>к</strong>и r-rs и д<strong>в</strong>е r-rm. П<strong>о</strong>перечные<br />

жил<strong>к</strong>и между <strong>в</strong>ет<strong>в</strong>ями RS и МА <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Передняя <strong>в</strong>ет<strong>в</strong>ь MP + СиА не <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> МА 2.<br />

Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть уз<strong>к</strong><strong>а</strong>я, длинн<strong>а</strong>я. А 2 пр<strong>о</strong>х<strong>о</strong>дит<br />

п<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>му з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти и<br />

не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. Между Ai и А 2 <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong><br />

имеет<strong>с</strong>я S-<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>. Пермь.<br />

Один р<strong>о</strong>д.<br />

Palaeoperla Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. exacta Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). RS <strong>о</strong>бр<strong>а</strong>зует<br />

три-четыре <strong>в</strong>ет<strong>в</strong>и; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии не<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> R; <strong>в</strong> меди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>в</strong><strong>о</strong><strong>с</strong>емь п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>, <strong>в</strong> <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м — четыре. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10 мм (ри<strong>с</strong>. 345 и 346). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>, <strong>о</strong>дин из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен нимф<strong>о</strong>й.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.

345. Palaeoperla exacta Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, x 5,5, н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 346. Palaeoperla prisca<br />

Sharov; нимф<strong>а</strong>, х 10,0. и. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961). 347. Perlopsls fillcornts Martynov; А — <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong><br />

г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>, х 3,0 (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940); Б — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 3,3, (<strong>о</strong>риг. ри<strong>с</strong>.); В — л<strong>а</strong>п<strong>к</strong><strong>а</strong>, х 8,3 (<strong>о</strong>риг. ри<strong>с</strong>.); н. пермь,<br />

Ур<strong>а</strong>л. 348. Palaeotaenlopteryx elegans Sharov; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 8,3; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 8,3; В— нимф<strong>а</strong>, х 8,3;<br />

н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 349. Palaeonemoura<br />

tiara Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 6,5, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

Ри<strong>с</strong>. 345—: 54. Отряд Plecoptera<br />

135<br />

(Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)). 350. Mesonemura turanlca Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 5,1, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 351. Mesotaeniopteryx<br />

elongate Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,8, н. юр<strong>а</strong>, Ср,<br />

Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 352. Nemoura ctnerea (Retzlus); А —<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 6,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 6,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(lilies, 1955). 353. Leuctra hlppopus Kempy; A — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

x 7,7; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 7,7; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Mosely,<br />

1932). 354. Perla sp.; <strong>о</strong>бщий <strong>в</strong>ид, х 1,9, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> Азия.<br />

Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1925)<br />

http://jurassic.ru/

СЕМЕЙСТВО PERLOPSEIDAE MARTYNOV, 1940<br />

В <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле три-четыре <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>в</strong>ет<strong>в</strong>и SC; R <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью близ <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

SC; <strong>о</strong>дн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> г-rs, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичн<strong>а</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й r-rs у <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Palaeoperlidae,<br />

и п<strong>о</strong> <strong>к</strong>р<strong>а</strong>йней мере три г-гт, д<strong>в</strong>е из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее r-rs. Между <strong>в</strong>ет<strong>в</strong>ями<br />

МА имеют<strong>с</strong>я п<strong>о</strong>перечные жил<strong>к</strong>и. Антенны<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из длинных члени<strong>к</strong><strong>о</strong><strong>в</strong>, б<strong>а</strong>з<strong>а</strong>льный<br />

члени<strong>к</strong> <strong>в</strong>здутый. Грудн<strong>о</strong>й <strong>о</strong>тдел <strong>о</strong>бширный.<br />

Н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><br />

3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че пер<strong>в</strong><strong>о</strong>г<strong>о</strong> и третьег<strong>о</strong>, р<strong>а</strong><strong>в</strong>ных п<strong>о</strong><br />

длине. Пермь. Один р<strong>о</strong>д.<br />

Perlopsis М <strong>а</strong> г t'y п <strong>о</strong> v, 1940. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Р, filicornis Martynov, 1940; н. пермь, Ур<strong>а</strong>л<br />

(<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине т<strong>а</strong><strong>к</strong><strong>о</strong>й<br />

же ширины, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е, <strong>с</strong> четырьмяпятью<br />

<strong>в</strong>ет<strong>в</strong>ями SC <strong>в</strong> нем; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

r-rs р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC; RS четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

MP + СиА д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Н<strong>а</strong><br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong>е г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рылье<strong>в</strong> пл<strong>о</strong>х<strong>о</strong><br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>о</strong><strong>с</strong>ь. Среди нед<strong>а</strong><strong>в</strong>н<strong>о</strong> н<strong>а</strong>йденных мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> эт<strong>о</strong>г<strong>о</strong> же <strong>в</strong>ид<strong>а</strong> из т<strong>о</strong>г<strong>о</strong><br />

же ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения имеют<strong>с</strong>я э<strong>к</strong>земпляры <strong>с</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличимым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 15—17 мм (ри<strong>с</strong>. 347 А — В;<br />

т<strong>а</strong>бл. VIII, фиг. 4).. Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

СЕМЕЙСТВО TAENIOPTERYGIDAE KLAPALEK, 1905<br />

В <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле не б<strong>о</strong>лее <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и<br />

SC. R <strong>о</strong>бычн<strong>о</strong> без <strong>в</strong>ет<strong>в</strong>ей; е<strong>с</strong>ли <strong>о</strong>дн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь имеет<strong>с</strong>я,<br />

т<strong>о</strong> <strong>о</strong>н<strong>а</strong> уд<strong>а</strong>лен<strong>а</strong> <strong>о</strong>т з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и SC, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

чег<strong>о</strong> Х-<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я фигур<strong>а</strong> <strong>в</strong> эт<strong>о</strong>м ме<strong>с</strong>те <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

RS <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бр<strong>а</strong>зует<br />

ч<strong>а</strong>ще д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и, реже три или четыре. От<br />

MP + СиА <strong>о</strong>тх<strong>о</strong>дит <strong>в</strong>перед <strong>о</strong>т <strong>о</strong>дн<strong>о</strong>й д<strong>о</strong> четырех<br />

<strong>в</strong>ет<strong>в</strong>ей. Пермь — ныне. Четыре <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong> и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Pataeotaeniopteryx Sharov, 1961- Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. elegans Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Отрез<strong>о</strong><strong>к</strong><br />

RS <strong>о</strong>т п<strong>о</strong>перечн<strong>о</strong>й д<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> т<strong>а</strong><strong>к</strong><strong>о</strong>й же<br />

длины 1*ли <strong>с</strong>лег<strong>к</strong><strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет длину <strong>в</strong>ет<strong>в</strong>ей<br />

RS; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> R; между МА и<br />

MP + СиА не б<strong>о</strong>лее четырех п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>;<br />

Ai <strong>с</strong><strong>о</strong>единен<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й<br />

<strong>с</strong> CuP; А 2 р<strong>а</strong>н<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е жил<strong>к</strong>и.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 7 мм<br />

(ри<strong>с</strong>. 348 А, Б). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>, <strong>о</strong>дин из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>н п<strong>о</strong> нимфе (ри<strong>с</strong>. 348В). Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Palaeonemoura Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. clara Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Отрез<strong>о</strong><strong>к</strong> RS<br />

между r-rs и р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>в</strong>ет<strong>в</strong>ей RS; <strong>в</strong>ет­<br />

136<br />

http://jurassic.ru/<br />

<strong>в</strong>и МА <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнуты <strong>к</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю; MP + СиА без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце и из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>йней мере <strong>о</strong>дн<strong>а</strong> из п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>ля между MP + СиА и CuP<br />

упир<strong>а</strong>ет<strong>с</strong>я не <strong>в</strong> CuP, <strong>а</strong> <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>;<br />

CuP из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти имеет <strong>в</strong>ид длинн<strong>о</strong>г<strong>о</strong>,<br />

уз<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong>лин<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

8 мм (ри<strong>с</strong>. 349). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Mesonemura Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — М. maaki,<br />

Brauer, Redtenbacher, Ganglbauer, 1889; н. юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-Б<strong>а</strong>лей).<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>о</strong>чень близ<strong>к</strong><strong>о</strong> п<strong>о</strong>дх<strong>о</strong>дит <strong>к</strong> С, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong><br />

передняя <strong>в</strong>ет<strong>в</strong>ь SC <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> RS между r-rs и р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м <strong>в</strong><br />

2—3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>в</strong>ет<strong>в</strong>ей RS; л<strong>а</strong>п<strong>к</strong><strong>а</strong> длинн<strong>а</strong>я;<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>й и третий члени<strong>к</strong>и р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине, <strong>с</strong><strong>а</strong>мый<br />

длинный — пер<strong>в</strong>ый члени<strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 8—11 мм (ри<strong>с</strong>. 350; т<strong>а</strong>бл. VIII,<br />

фиг. 5). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и<br />

Ср. Азии.<br />

Mesotaeniopteryx М <strong>а</strong> г t у п <strong>о</strong> v, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. elongata Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Передние<br />

<strong>к</strong>рылья уз<strong>к</strong>ие; SC пр<strong>о</strong>х<strong>о</strong>дит п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю, з<strong>а</strong>дняя ее <strong>в</strong>ет<strong>в</strong>ь <strong>о</strong>чень длинн<strong>а</strong>я,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее r-rs; R <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и. MP -f СиА н<strong>а</strong> <strong>к</strong><strong>о</strong>нце<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> три <strong>в</strong>ет<strong>в</strong>и; д<strong>в</strong>е передние р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

близ<strong>к</strong><strong>о</strong> друг <strong>к</strong> другу, длинные, <strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине пр<strong>о</strong>х<strong>о</strong>дят п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю. Между МА, MP + СиА и CuP<br />

б<strong>о</strong>лее де<strong>с</strong>яти п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 15—21 мм (ри<strong>с</strong>. 351). Три <strong>в</strong>ид<strong>а</strong>.<br />

Н. юр<strong>а</strong> Ср. Азии.<br />

СЕМЕЙСТВО NEMOURIDAE KLAPALEK, 1905<br />

В передних и з<strong>а</strong>дних <strong>к</strong>рыльях имеет<strong>с</strong>я <strong>о</strong>дн<strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>ь R, берущ<strong>а</strong>я н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> <strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>о</strong>т ме<strong>с</strong>т<strong>а</strong><br />

<strong>в</strong>п<strong>а</strong>дения з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и SC <strong>в</strong> R, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

чег<strong>о</strong> <strong>в</strong> эт<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е уп<strong>о</strong>мянутые жил<strong>к</strong>и <strong>в</strong>ме<strong>с</strong>те <strong>с</strong><br />

rs-ma и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием RS <strong>о</strong>бр<strong>а</strong>зуют Х-<strong>о</strong>бр<strong>а</strong>зную<br />

фигуру; r-rs р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> <strong>с</strong>имметричн<strong>о</strong> п<strong>о</strong> <strong>о</strong>тн<strong>о</strong>шению<br />

<strong>к</strong> эт<strong>о</strong>й фигуре. MAi и <strong>с</strong>т<strong>в</strong><strong>о</strong>л МА <strong>о</strong>бр<strong>а</strong>зуют<br />

единую прямую жил<strong>к</strong>у, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>о</strong>тх<strong>о</strong>дит<br />

н<strong>а</strong>з<strong>а</strong>д МА 2; ин<strong>о</strong>гд<strong>а</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> МА 2 <strong>в</strong> <strong>в</strong>иде л<strong>о</strong>жн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й.<br />

MP + СиА не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я (ри<strong>с</strong>. 352). Мел — ныне.<br />

Три <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong> из н. мел<strong>а</strong> Кит<strong>а</strong>я (Sinoperla<br />

Ping, 1928; Pertariopsis Ping, 1928; Synonemoura<br />

Ping, 1928) и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).

СЕМЕЙСТВО LEUCTRIDAE KLAPALEK, 1905<br />

В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> не<br />

б<strong>о</strong>лее <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и SC, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>й <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> и имеющей <strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и<br />

между С и SC. SC или <strong>с</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й передней<br />

<strong>в</strong>ет<strong>в</strong>ью, или <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> переднем<br />

<strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>, <strong>а</strong> ее з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и <strong>в</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле- Вет<strong>в</strong>и R <strong>в</strong><br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Вет<strong>в</strong>и RS и<br />

МА прямые, п<strong>а</strong>р<strong>а</strong>ллельные. П<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

<strong>в</strong> п<strong>о</strong>ле между MP + CuA и CuP приблизительн<strong>о</strong><br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше, чем между МА и<br />

MP + CuA. CuP S-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong> и <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине пр<strong>о</strong>х<strong>о</strong>дит близ<strong>к</strong><strong>о</strong> <strong>к</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> и п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> ему д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>перед,<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>я передней трети <strong>к</strong>рыл<strong>а</strong>. В з<strong>а</strong>днем <strong>к</strong>рыле<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льный <strong>в</strong>еер <strong>в</strong> <strong>в</strong>иде неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти <strong>с</strong> пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й Аг (ри<strong>с</strong>. 353). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из<strong>в</strong>е<strong>с</strong>тный т<strong>а</strong><strong>к</strong>же из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий-<br />

Ри<strong>с</strong>. 355—358. Plecoptera incertae sedis<br />

<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и Герм<strong>а</strong>нии. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне.<br />

СЕМЕЙСТВО PERL1DAE STEPHENS, 1835<br />

В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>в</strong>ет<strong>в</strong>и SC <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечных; четырепять<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их* <strong>в</strong>ет<strong>в</strong>ей R <strong>в</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле.<br />

RS <strong>о</strong>бр<strong>а</strong>зует три-четыре <strong>в</strong>ет<strong>в</strong>и, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

гребенч<strong>а</strong>т<strong>о</strong>. Между RS и МА шир<strong>о</strong><strong>к</strong><strong>о</strong>е<br />

п<strong>о</strong>ле, з<strong>а</strong>м<strong>к</strong>нут<strong>о</strong>е <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й rs-ma. MP + CuA из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>,<br />

з<strong>а</strong>мы<strong>к</strong><strong>а</strong>ет п<strong>о</strong>ле между МА и MP + CuA<br />

и <strong>о</strong>бр<strong>а</strong>зует <strong>о</strong>т четырех д<strong>о</strong> ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей,<br />

гребенч<strong>а</strong>т<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дящих н<strong>а</strong>з<strong>а</strong>д. CuP <strong>о</strong>бычн<strong>о</strong><br />

прям<strong>а</strong>я. В з<strong>а</strong>дних <strong>к</strong>рыльях <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льный <strong>в</strong>еер<br />

<strong>о</strong>бширный, <strong>с</strong> гребенч<strong>а</strong>т<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящей<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д А 2<br />

(ри<strong>с</strong>. 354). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и Англии. .<br />

355. Uralonympha varica О. Zalessky; нимф<strong>а</strong>, x 3,3, н. пермь, нимф<strong>а</strong>, х 4,0, н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>.). 358. Platyper-<br />

Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1939). 356. Permoleuctropsis gracilis Marty- la platypodia Brauer, Redtenbacher, Ganglbauer; <strong>в</strong>иыф<strong>а</strong>, x 3,6,<br />

noy; нимф<strong>а</strong>, x 6,0, <strong>в</strong>. Пермь, Ю. Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). и. юр<strong>а</strong>. Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

357. Mesoleuctra gracilis Brauer, Redtenbacher, Ganglbauer;<br />

137<br />

http://jurassic.ru/

PLECOPTERA INCERTAE SEDIS<br />

Uralonympha G. Zalessky, 1939. Тип -<br />

р<strong>о</strong>д<strong>а</strong> — U. varica G. Zalessky, 1939; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Нимф<strong>а</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я. Крыл<strong>о</strong><strong>в</strong>ые<br />

з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и уз<strong>к</strong>ие, длинные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong> <strong>о</strong>тн<strong>о</strong>шению<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>и тел<strong>а</strong> п<strong>о</strong>д угл<strong>о</strong>м <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30°; их<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены, <strong>в</strong>ершины з<strong>а</strong><strong>к</strong>руглены.<br />

Цер<strong>к</strong>и <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Длин<strong>а</strong><br />

тел<strong>а</strong> без цер<strong>о</strong><strong>к</strong> 10 мм (ри<strong>с</strong>. 355). Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Permoleuctropsis Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. gracilis Martynov, 1937; пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Нимф<strong>а</strong>. Крыл<strong>о</strong><strong>в</strong>ые<br />

з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и з<strong>а</strong><strong>о</strong><strong>с</strong>тренные, их <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>с</strong>ближены.<br />

Длин<strong>а</strong> цер<strong>о</strong><strong>к</strong> приблизительн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> длине<br />

брюш<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> без цер<strong>о</strong><strong>к</strong> 5,3 мм<br />

(ри<strong>с</strong>. 356). Один <strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

Mesoleuctra Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — М. gracilis<br />

Brauer, Redtenbacher, Ganglbauer, 1889; н. юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>. У<strong>с</strong>ть-Б<strong>а</strong>лей).<br />

Нимф<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

уз<strong>к</strong><strong>а</strong>я, тр<strong>а</strong>пецие<strong>в</strong>идн<strong>а</strong>я, б<strong>о</strong>лее уз<strong>к</strong><strong>а</strong>я у передне-<br />

PLECOPTEROIDEA<br />

СЕМЕЙСТВО HYPOPERLIDAE MARTYNOV, 1928<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong> переп<strong>о</strong>нч<strong>а</strong>тыми<br />

<strong>к</strong>рыльями, <strong>с</strong>л<strong>о</strong>женными <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>к</strong>р<strong>о</strong><strong>в</strong>ле<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>.<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>реп<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> между С и SC.<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> R, <strong>а</strong> ее един<strong>с</strong>т<strong>в</strong>енн<strong>а</strong>я передняя<br />

<strong>в</strong>ет<strong>в</strong>ь идет <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>.<br />

R <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бр<strong>а</strong>зует д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и.<br />

RS <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я гребенч<strong>а</strong>т<strong>о</strong>. МА н<strong>а</strong>чин<strong>а</strong>ет <strong>в</strong>ет<strong>в</strong>ить<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> RS и <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>о</strong> трех <strong>в</strong>ет<strong>в</strong>ей; MP <strong>с</strong>лит<strong>а</strong> <strong>с</strong> СиА, и их <strong>о</strong>бщий<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л <strong>о</strong>бр<strong>а</strong>зует три-четыре гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>в</strong>ет<strong>в</strong>и, пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, упир<strong>а</strong>я<strong>с</strong>ь <strong>в</strong><br />

CuP. Д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и. Пермь.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>. •<br />

Hypoperla Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Н. elegans Martynov, 1928; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; <strong>в</strong>ет<strong>в</strong>ь <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>ты<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т т<strong>о</strong>ч<strong>к</strong>и <strong>в</strong>п<strong>а</strong>дения SC <strong>в</strong> R; четыре<br />

<strong>в</strong>ет<strong>в</strong>и RS; <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong> MP + СиА четыре-лять.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм (ри<strong>с</strong>. 359).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

г<strong>о</strong> <strong>к</strong><strong>а</strong>я; <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ые з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong>д<br />

<strong>о</strong>чень м<strong>а</strong>лым угл<strong>о</strong>м <strong>к</strong> пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>;<br />

н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> л<strong>а</strong>п<strong>к</strong>и <strong>с</strong><strong>а</strong>мый<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, пер<strong>в</strong>ый немн<strong>о</strong>г<strong>о</strong> длиннее, <strong>с</strong><strong>а</strong>мый<br />

длинный — третий; цер<strong>к</strong>и немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че длины<br />

тел<strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> 18 мм (ри<strong>с</strong>. 357; т<strong>а</strong>бл. IX,<br />

фиг. 1)ч Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; н. мел Кит<strong>а</strong>я.<br />

Platyperla Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — P. platypodia<br />

Brauer, Redtenbacher, Ganglbauer, 1889; н. юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-<br />

Б<strong>а</strong>лей). Нимф<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> прям<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>а</strong>я<br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ые з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong>д неб<strong>о</strong>льшим угл<strong>о</strong>м<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>и тел<strong>а</strong>; н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, бедр<strong>а</strong> и г<strong>о</strong>лени <strong>в</strong><strong>с</strong>ех<br />

трех п<strong>а</strong>р н<strong>о</strong>г упл<strong>о</strong>щенные, шир<strong>о</strong><strong>к</strong>ие; пер<strong>в</strong>ый и<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>в</strong>ме<strong>с</strong>те <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

длинн<strong>о</strong>г<strong>о</strong> третьег<strong>о</strong>; цер<strong>к</strong>и т<strong>о</strong>н<strong>к</strong>ие, длинные.<br />

Длин<strong>а</strong> тел<strong>а</strong> без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> 18 мм (ри<strong>с</strong>. 358;<br />

т<strong>а</strong>бл. IX, фиг. 2). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

INCERTAE SEDIS<br />

Hypoperlopsis G. Zalessky, 1948. Тип<br />

р<strong>о</strong>д<strong>а</strong> — H. splendens G. Zalessky, 1948;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине уже <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>.<br />

Вет<strong>в</strong>ь SC <strong>о</strong>тх<strong>о</strong>дит д<strong>о</strong> <strong>в</strong>п<strong>а</strong>дения SC <strong>в</strong> R; три <strong>в</strong>ет<strong>в</strong>и<br />

RS; <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong> MP + СиА <strong>о</strong>дн<strong>а</strong>-д<strong>в</strong>е. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 14 мм (ри<strong>с</strong>. 360).<br />

Один <strong>в</strong>ид- Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

МА<br />

359—360. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hypoperlidae<br />

359. Hypoperla elegans Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,6, <strong>в</strong><br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). 360. Hypoperlopsis<br />

splendens G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,4, н. пермь, Ур<strong>а</strong>л<br />

/ (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1948)<br />

138<br />

http://jurassic.ru/

ОТРЯД EMBIOPTERA. ЭМБИИ<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Средней <strong>в</strong>еличины и мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

(длин<strong>а</strong> тел<strong>а</strong> 4—20 мм). Бе<strong>с</strong><strong>к</strong>рылые или <strong>с</strong> д<strong>в</strong>умя<br />

п<strong>а</strong>р<strong>а</strong>ми уз<strong>к</strong>их г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong> и <strong>к</strong><strong>о</strong>р<strong>о</strong>че, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти<br />

<strong>к</strong>рыл<strong>а</strong>, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью; М и Си<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>литы, М п<strong>о</strong><strong>с</strong>ле <strong>о</strong>тщепления <strong>о</strong>т Си<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS или р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я, и т<strong>о</strong>гд<strong>а</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> МА н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении <strong>с</strong>лит<strong>а</strong> <strong>с</strong><br />

RS; MP — <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>; СиА делит<strong>с</strong>я н<strong>а</strong><br />

длинную, идущую <strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> CuAi,<br />

и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую СиАг; СиР нет; <strong>о</strong>дн<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> немн<strong>о</strong>г<strong>о</strong>. В з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>в</strong>еер<strong>а</strong> нет.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я; р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

грызущег<strong>о</strong> тип<strong>а</strong>; <strong>а</strong>нтенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, чи<strong>с</strong>л<strong>о</strong><br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> 15—30; ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты,<br />

гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Передне- и з<strong>а</strong>днегрудь <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>с</strong>реднегруди; н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, бедр<strong>а</strong> р<strong>а</strong><strong>с</strong>ширены,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> з<strong>а</strong>дние; пер<strong>в</strong>ый члени<strong>к</strong><br />

передней л<strong>а</strong>п<strong>к</strong>и у<strong>в</strong>еличен и р<strong>а</strong><strong>с</strong>ширен, <strong>в</strong> нем<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>а</strong>утинные железы; члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

л<strong>а</strong>п<strong>о</strong><strong>к</strong> три. Сегмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> де<strong>с</strong>ять, н<strong>а</strong> <strong>к</strong><strong>о</strong>нце<br />

— <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие цер<strong>к</strong>и, <strong>о</strong>бычн<strong>о</strong> прид<strong>а</strong>т<strong>к</strong>и <strong>с</strong><strong>а</strong>мц<strong>а</strong><br />

<strong>а</strong><strong>с</strong>имметричны. Пре<strong>в</strong>р<strong>а</strong>щение неп<strong>о</strong>лн<strong>о</strong>е. Жи<strong>в</strong>ут<br />

<strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями, <strong>к</strong><strong>о</strong>р<strong>о</strong>й. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 70 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

- тр<strong>о</strong>пи<strong>к</strong><strong>о</strong><strong>в</strong> и <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>о</strong><strong>в</strong>. Пермь — ныне. П<strong>о</strong>д<strong>о</strong>тряды:<br />

Sheimioidea, Euembiaria.<br />

ПОДОТРЯД SHEIMIOIDEA<br />

С<strong>а</strong>мцы и <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong> <strong>к</strong>рыльями, р<strong>а</strong><strong>в</strong>ными п<strong>о</strong><br />

длине телу. Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> из<strong>о</strong>гнуты <strong>к</strong>з<strong>а</strong>ди; длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong>; у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий яйце<strong>к</strong>л<strong>а</strong>д, члени<strong>к</strong><strong>о</strong><strong>в</strong> цер<strong>о</strong><strong>к</strong> б<strong>о</strong>лее<br />

д<strong>в</strong>ух. Пермь. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Sheimiidae.<br />

СЕМЕЙСТВО SHEIMIIDAE<br />

О. MARTYNOVA, 1958<br />

Крылья <strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong> тел<strong>о</strong>м. SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> С немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние М длиннее <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

RS и <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS д<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления; RS <strong>о</strong>тх<strong>о</strong>дит<br />

<strong>о</strong>т МА п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>. Пермь<br />

<strong>о</strong>дин р<strong>о</strong>д.<br />

139<br />

Sheimia О. М <strong>а</strong> г t у п <strong>о</strong> v <strong>а</strong>, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Sh. sojanensis О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. R и RS <strong>в</strong> <strong>в</strong>ершине не<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я; МА <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т RS н<strong>а</strong> <strong>с</strong>ередине<br />

длины <strong>к</strong>рыл<strong>а</strong>, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. Яйце<strong>к</strong>л<strong>а</strong>д<br />

Ри<strong>с</strong>. 361—362. Отряд Embioptera<br />

361. Sheimia sojanensis О. Martynova; <strong>о</strong>бщий <strong>в</strong>ид, х 14,5, <strong>в</strong>.<br />

пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 362. OUgotoma<br />

saundersl West wood; переднее <strong>к</strong>рыл<strong>о</strong>, х 8,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Азия<br />

(Handlirsch, 1906)<br />

р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине трем п<strong>о</strong><strong>с</strong>ледним <strong>с</strong>егмент<strong>а</strong>м<br />

брюш<strong>к</strong><strong>а</strong>; члени<strong>к</strong><strong>о</strong><strong>в</strong> цер<strong>о</strong><strong>к</strong> 10—12. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

1,8 мм (ри<strong>с</strong>. 361). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

ПОДОТРЯД EUEMBIARIA<br />

Бе<strong>с</strong><strong>к</strong>рылы; <strong>к</strong>рыл<strong>а</strong>ты т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>а</strong>мцы. Крылья<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че тел<strong>а</strong>. Пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong><br />

прямые; длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину <strong>в</strong><br />

4 р<strong>а</strong>з<strong>а</strong>. Яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong> нет, члени<strong>к</strong><strong>о</strong><strong>в</strong> цер<strong>о</strong><strong>к</strong> не б<strong>о</strong>лее<br />

д<strong>в</strong>ух. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Clothodidae,<br />

Oligotomidae, Notoligotomidae, Embiidae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

http://jurassic.ru/

СЕМЕЙСТВО CLOTHODIDAE TILLYARD, 1937<br />

С<strong>а</strong>мцы <strong>с</strong> <strong>к</strong>рыльями. SC д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>; R <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> 'переднем<br />

<strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я д<strong>о</strong> <strong>с</strong>лияния <strong>с</strong> RS;<br />

МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuAi <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й-д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями.<br />

Цер<strong>к</strong>и д<strong>в</strong>учлени<strong>с</strong>тые, неизмененные; <strong>в</strong>ентр<strong>а</strong>льные<br />

прид<strong>а</strong>т<strong>к</strong>и <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong>имметричные. Не<strong>о</strong>ген—<br />

ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне и <strong>о</strong>дин из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО OLIGOTOM1DAE ENDERLEIN, 1909<br />

С<strong>а</strong>мцы <strong>с</strong> <strong>к</strong>рыльями: SC д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>с</strong>ередины<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; R <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

Д<strong>в</strong>е п<strong>а</strong>ры неб<strong>о</strong>льших г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong><br />

длин<strong>о</strong>й 3,5—10 мм. SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> С <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

ди<strong>с</strong>т<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец ее <strong>с</strong><strong>о</strong>единен п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й<br />

<strong>с</strong> R; R <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м н<strong>а</strong> <strong>к</strong><strong>о</strong>нце: RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>нце <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые или <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м<br />

<strong>в</strong>ет<strong>в</strong>и; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> CuA, <strong>о</strong>тделяет<strong>с</strong>я<br />

<strong>о</strong>т п<strong>о</strong><strong>с</strong>ледней <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; МА<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> теряет <strong>с</strong><strong>в</strong><strong>о</strong>е<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние и, <strong>с</strong><strong>о</strong>едини<strong>в</strong>ши<strong>с</strong>ь <strong>с</strong> п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й<br />

гэз +4 — ma, дел<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>х<strong>о</strong>жей н<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS;<br />

MP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong><br />

<strong>с</strong>ильную CuA <strong>с</strong> шир<strong>о</strong><strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м (у Регmembiidae<br />

нет р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>) и пр<strong>о</strong><strong>с</strong>тую, <strong>с</strong>л<strong>а</strong>бую<br />

CuP; д<strong>в</strong>е <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

немн<strong>о</strong>г<strong>о</strong>. В з<strong>а</strong>дних <strong>к</strong>рыльях <strong>в</strong>еер<strong>а</strong> нет.<br />

Неб<strong>о</strong>льш<strong>а</strong>я г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong>низ: <strong>а</strong>нтенны<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; члени<strong>к</strong><strong>о</strong><strong>в</strong> 12—13. Переднегрудь<br />

уже г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; <strong>с</strong>реднегрудь примерн<strong>о</strong> треуг<strong>о</strong>льн<strong>о</strong>й<br />

ф<strong>о</strong>рмы, п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине переднегруди;<br />

н<strong>о</strong>ги <strong>с</strong>редней длины, бег<strong>а</strong>тельные, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> четыре; из них пер<strong>в</strong>ый<br />

длинный, р<strong>а</strong><strong>в</strong>ный п<strong>о</strong> длине трем ди<strong>с</strong>т<strong>а</strong>льным;<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>.<br />

Тел<strong>о</strong> нимфы упл<strong>о</strong>щен<strong>о</strong> <strong>в</strong> д<strong>о</strong>р<strong>с</strong><strong>о</strong>-<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>м<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лении. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я; гл<strong>а</strong>з<strong>а</strong><br />

<strong>к</strong>рупные; <strong>а</strong>нтенны жгути<strong>к</strong><strong>о</strong><strong>в</strong>идные, мн<strong>о</strong>г<strong>о</strong>члени<strong>с</strong>тые.<br />

Грудн<strong>о</strong>й <strong>о</strong>тдел нерез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т<br />

брюшн<strong>о</strong>г<strong>о</strong>; переднегрудь неб<strong>о</strong>льш<strong>а</strong>я, п<strong>о</strong>чти<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я; <strong>с</strong>редне- и з<strong>а</strong>днегрудь п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ны<br />

п<strong>о</strong> <strong>в</strong>еличине, не<strong>с</strong>ут н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленные н<strong>а</strong>з<strong>а</strong>д <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ые<br />

чехлы; н<strong>о</strong>ги <strong>с</strong> приблизительн<strong>о</strong> р<strong>а</strong><strong>в</strong>ными<br />

п<strong>о</strong> длине бедр<strong>а</strong>ми и г<strong>о</strong>ленями, п<strong>о</strong><strong>с</strong>ледние упл<strong>о</strong>щены,<br />

<strong>с</strong> не<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>им гребнем п<strong>о</strong> <strong>с</strong>редней линии;<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я. Члени<strong>к</strong><strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> де<strong>с</strong>ять,<br />

де<strong>в</strong>ять из них <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>ль-<br />

ОТРЯД MIOMOPTERA<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

140<br />

http://jurassic.ru/<br />

н<strong>а</strong> RS; RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; М не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я д<strong>о</strong><br />

<strong>с</strong>лияния <strong>с</strong> RS; МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; CuA р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

длинн<strong>а</strong>я; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> ее пр<strong>о</strong>х<strong>о</strong>дит<br />

пл<strong>о</strong>тн<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>. Прид<strong>а</strong>т<strong>к</strong>и<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>а</strong><strong>с</strong>имметричные; де<strong>с</strong>ятый тергит р<strong>а</strong>зделен<br />

н<strong>а</strong> д<strong>в</strong>е ч<strong>а</strong><strong>с</strong>ти; <strong>с</strong>л<strong>о</strong>жные прид<strong>а</strong>т<strong>к</strong>и н<strong>а</strong> ле<strong>в</strong><strong>о</strong>м<br />

п<strong>о</strong>лутергите и пер<strong>в</strong><strong>о</strong>м члени<strong>к</strong>е ле<strong>в</strong><strong>о</strong>й цер<strong>к</strong>и;<br />

цер<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 362). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне Азии,<br />

С. Амери<strong>к</strong>и, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии; <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин из плей<strong>с</strong>т<strong>о</strong>цен<strong>а</strong><br />

В. Афри<strong>к</strong>и.<br />

ными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми; н<strong>а</strong> де<strong>с</strong>ят<strong>о</strong>м <strong>с</strong>егменте <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ие цер<strong>к</strong>и, члени<strong>к</strong>и <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong>литы. Видим<strong>о</strong>, <strong>о</strong>бит<strong>а</strong>ли <strong>в</strong> те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>а</strong>х,<br />

<strong>в</strong>ели хищный <strong>о</strong>бр<strong>а</strong>з жизни. Длин<strong>а</strong> тел<strong>а</strong><br />

д<strong>о</strong> 18 мм (ри<strong>с</strong>. 363 А, Б). В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — пермь.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Archaemiopteridae, Palaeomanteidae;<br />

Permembiidae? (ри<strong>с</strong>. 371), п<strong>о</strong><strong>с</strong>леднее <strong>в</strong>не<br />

СССР.<br />

СЕМЕЙСТВО ARCHAEMIOPTERIDAE GL1THORL, 1939<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong>;<br />

<strong>в</strong> передних <strong>к</strong>рыльях <strong>о</strong>бе <strong>в</strong>ет<strong>в</strong>и RS <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м,<br />

М <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т CuA б<strong>а</strong>з<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS, <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — пермь. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>.<br />

Archaemioptera G u t h б г 1, 1939. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. carbonaria Guthorl, 1939; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (низы <strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong>, С<strong>а</strong><strong>а</strong>р).<br />

R прям<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; М <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>,<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

5,3 мм (ри<strong>с</strong>. 364). Один <strong>в</strong>ид. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Tychtodelopterum О. Martynov <strong>а</strong>, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. relictum О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>). R из<strong>о</strong>гнут<strong>а</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

<strong>с</strong>ередине <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA длинный. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3,2 мм (ри<strong>с</strong>. 365). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.

СЕМЕЙСТВО PALAEOMANTEIDAE HANDLIRSCH,<br />

1906<br />

(Delopteridae Sellards, 1909;<br />

Archisialidae Martynov, 1933)<br />

Крылья уз<strong>к</strong>ие, длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину <strong>в</strong><br />

3 р<strong>а</strong>з<strong>а</strong>. З<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; М <strong>о</strong>тх<strong>о</strong>дит<br />

<strong>о</strong>т СиА п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong>м RS.<br />

Пермь. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

363В<br />

Ри<strong>с</strong>. 363—371.<br />

363. Delopterum sp., нимф<strong>а</strong>; А — <strong>о</strong>бщий г<br />

<strong>в</strong>ид, х 3,0; Б — переднее<br />

нимф<strong>а</strong>льн<strong>о</strong>е <strong>к</strong>рыл<strong>о</strong>, х 11,0; В — з<strong>а</strong>днее йимф<strong>а</strong>льн<strong>о</strong>е <strong>к</strong>рыл<strong>о</strong>,<br />

х 11,0; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1956). 364. Агchaemioptera<br />

carbonaria Guthorl; переднее <strong>к</strong>рыл<strong>о</strong>, х 8,7, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Guthorl, 1939). 365. Tychtodelopterum rellctumO. Martynova;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 19,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 366. Miomatoneura frigida Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 8,3, <strong>в</strong>. пермь, 1<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>.).<br />

Miomatoneura Martynov, 1927. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. frigida Martynov, 1927; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

RSi+2 <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; МА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS3+4, <strong>о</strong>бр<strong>а</strong>зуя<br />

третью <strong>в</strong>ет<strong>в</strong>ь RS; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА имеет<br />

<strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и-; Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 5,0—<br />

6,4 мм (ри<strong>с</strong>. 366). Четыре <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Miomatoneurella О. Martynova, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. reducta О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> I). Обе <strong>в</strong>ет<strong>в</strong>и RS пр<strong>о</strong><strong>с</strong>тые; МА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> RS3+4, теряя <strong>с</strong><strong>в</strong><strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 4 мм (ри<strong>с</strong>. 367). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Отряд Miomoptera<br />

141<br />

http://jurassic.ru/<br />

СИА<br />

1<br />

м 371<br />

367. Miomatoneurella reducta О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong><br />

х 11,6, <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 368.<br />

Palaeomantisca lata Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,0, н. пермь;<br />

Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1940). 369. Palaeomantls schmldti Handlirsch<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 10,0, <strong>в</strong>. пермь. Приур<strong>а</strong>лье (<strong>о</strong>риг. ри<strong>с</strong>.). 370.<br />

Delopterum Incertum (Martynov); переднее <strong>к</strong>рыл<strong>о</strong>, х10,0. <strong>в</strong>. пермь,<br />

Приур<strong>а</strong>лье (<strong>о</strong>риг. ри<strong>с</strong>.). 371. Permembia dellcatula Tillyard;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, н. Пермь, С Амери<strong>к</strong><strong>а</strong> (Tillyard, 1937)<br />

Palaeomantisca Martynov, 1940. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. lata Martynov, 1940; н. пермь. Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>, длиннее RS ] + 2, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; RSi+г <strong>с</strong>

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, р<strong>а</strong><strong>в</strong>н<strong>а</strong> ему п<strong>о</strong> длине; М <strong>в</strong> 6 р<strong>а</strong>з <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; МА не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS;<br />

<strong>в</strong>ыем<strong>к</strong>и п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> нет. Длин<strong>а</strong><br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 9,5 мм (ри<strong>с</strong>. 368). Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Palaeomantis Handlirsch, 1904. Тип<br />

р<strong>о</strong>д<strong>а</strong>—P. schmidti Handlirsch,, 1904; <strong>в</strong>. пермь,<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие<br />

Г<strong>о</strong>ры). SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины<br />

<strong>к</strong>рыл<strong>а</strong>; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>че RSi+2,<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>; RSi+2 длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; МА не<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляя<strong>с</strong>ь <strong>с</strong> нею п<strong>о</strong>чти н<strong>а</strong><br />

Д<strong>в</strong>е п<strong>а</strong>ры г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong> <strong>с</strong> п<strong>о</strong>чти<br />

прямыми пр<strong>о</strong>д<strong>о</strong>льными и р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. В передних<br />

<strong>к</strong>рыльях пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й и <strong>к</strong>лин<strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тей нет. SC длинн<strong>а</strong>я; R <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей<br />

RS <strong>о</strong>т <strong>о</strong>дн<strong>о</strong>й д<strong>о</strong> <strong>с</strong>еми; МА <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью или<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; MP, CuA и CuP пр<strong>о</strong><strong>с</strong>тые. В <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е<br />

Enthygrammatidae CuA и CuP <strong>с</strong>ближены,<br />

<strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>х <strong>с</strong>ближены MP и<br />

CuA. MP и CuA <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е<br />

<strong>к</strong>рыл<strong>а</strong>: н<strong>а</strong> <strong>с</strong>ередине ег<strong>о</strong> — у <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> и ди<strong>с</strong>т<strong>а</strong>льнее ее — у перм<strong>с</strong><strong>к</strong>их.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>редней <strong>в</strong>еличины, гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я;<br />

длинные ните<strong>в</strong>идные <strong>а</strong>нтенны <strong>с</strong> б<strong>о</strong>льшим р<strong>а</strong>здутым<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ным члени<strong>к</strong><strong>о</strong>м, переднегрудь<br />

п<strong>о</strong>чти <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я. Н<strong>о</strong>ги длинные, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные;<br />

з<strong>а</strong>дние н<strong>о</strong>ги немн<strong>о</strong>г<strong>о</strong> длиннее, <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширенными бедр<strong>а</strong>ми, н<strong>о</strong> еще не прыг<strong>а</strong>тельные.<br />

Средней <strong>в</strong>еличины брюш<strong>к</strong><strong>о</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

цер<strong>к</strong><strong>а</strong>ми н<strong>а</strong> <strong>к</strong><strong>о</strong>нце. Ср. «<strong>а</strong>рб<strong>о</strong>н— пермь.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Synomaloptilidae, Pleisiogrammatidae,<br />

Paleuthygrammatidae (ри<strong>с</strong>. 372), Euthygrammatidae,<br />

Caloneuridae (ри<strong>с</strong>. 373), Permobiellidae,<br />

Anomalogrammatidae, Genopterygidae,<br />

из них четыре п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО SYNOMALOPTILIDAE<br />

MARTYNOV, 1938<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>, четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; MP и CuA н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>м<br />

пр<strong>о</strong>тяжении <strong>с</strong>литы, <strong>в</strong> <strong>в</strong>ершине р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зуя р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>. Н. пермь. Один р<strong>о</strong>д.<br />

Synomatoptila Martynov, 1938. Тип р<strong>о</strong>д<strong>а</strong><br />

— S. longipennis Martynov, 1938; н. пермь,<br />

ОТРЯД CALONEURODEA<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

142<br />

http://jurassic.ru/<br />

<strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; <strong>в</strong>ыем<strong>к</strong>и п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong><br />

нет. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 6 мм (ри<strong>с</strong>. 369). Один<br />

<strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Delopterum Sellards, 1909 (Leptoneurula<br />

Martynov, 1928; Detopsocus Tillyard, 1928;<br />

Pseudotnantis Martynov, 1930; Archisialis Martynov,<br />

1933). Тип р<strong>о</strong>д<strong>а</strong> — D. minutum Sellards,<br />

1909; н. пермь, С. Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). Отлич<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong>т Palaeomantis б<strong>о</strong>лее уз<strong>к</strong>им <strong>к</strong>рыл<strong>о</strong>м, н<strong>а</strong>личием<br />

неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>ыем<strong>к</strong>и п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong> (между <strong>в</strong>ершин<strong>а</strong>ми CuP и А) и б<strong>о</strong>лее<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й SC. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 3,5—<br />

10 мм (ри<strong>с</strong>. 370). 15 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Пермь Е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти СССР, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>, С. Амери<strong>к</strong>и.<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины «рыл<strong>а</strong>; MP <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ния SC; д<strong>в</strong>е<br />

длинные <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и. Длин<strong>а</strong> «рыл<strong>а</strong> 27,5<br />

мм (ри<strong>с</strong>. 374). Один <strong>в</strong>ид. Н. Пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

СЕМЕЙСТВО PLEISIOGRAMMATIDAE<br />

CARPENTER, 1943<br />

Переднее «рыл<strong>о</strong>. SC длинн<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong>, из<strong>о</strong>гнут<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> и R; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

ед<strong>в</strong><strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>, 'трех-четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; МА<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>с</strong>ближенные MP и<br />

CuA <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льные<br />

жил<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Н. пермь. Один р<strong>о</strong>д.<br />

Pleisiogramma Garpenter, 1943. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. mediate Carpenter, 1943; н. пермь, С. Амери<strong>к</strong><strong>а</strong><br />

(К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

уз<strong>к</strong><strong>о</strong>е, <strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>е; Ai <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; А 2 <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 15—18 мм (ри<strong>с</strong>. 375).<br />

Три <strong>в</strong>ид<strong>а</strong>. Н. пермь С. Амери<strong>к</strong>и и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО PALEUTHYGRAMMATIDAE<br />

CARPENTER, 1943<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC длинн<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong><br />

и немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее ее; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины «рыл<strong>а</strong>, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и; МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м

Ри<strong>с</strong>. 372—381. С<br />

372. Paleuthygramma tenuicorne Martynov (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция);<br />

x 1,5, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938). 373. Caloneura dawsoni<br />

Brongniart; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Carpenter,<br />

1948). 374. SynomalopHla longipennis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 2,0, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938). 375. Pleslogramma mediate<br />

Carpenter; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,6, н. пермь, С. Амери<strong>к</strong><strong>а</strong><br />

(Carpenter, 1943; интерпрет<strong>а</strong>ция жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния О. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й).<br />

376. Vllvia densinervosa G. Zalessky, ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, х 2,0, н. Пермь,<br />

Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1933). 377. Pseudogramma aberrant Marty-<br />

или пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; MP и СиА <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong> и ди<strong>с</strong>т<strong>а</strong>льнее<br />

ее; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е; А 2 длинн<strong>а</strong>я.<br />

Пермь. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Paleuthygramma Martynov, 1930. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. tenuicorne Martynov, 1930; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC прям<strong>а</strong>я, не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong><br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти «рыл<strong>а</strong>; RS <strong>о</strong>бр<strong>а</strong>зует <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; Ai <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 18—22 мм (ри<strong>с</strong>. 372). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>, С. Амери<strong>к</strong>и.<br />

143<br />

эяд Caloneurodea<br />

nov; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,4, <strong>в</strong>. Пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1938). 378. Vtlviopsls extensa Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 2,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938). 379. Ейthygramma<br />

parallellum (Matynov); переднее <strong>к</strong>рыл<strong>о</strong>, х 2,5, <strong>в</strong><br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938). 380. Permoblella<br />

perspicua Tillyard; переднее <strong>к</strong>рыл<strong>о</strong>, х 5,0, н. пермь, С. Амери<strong>к</strong><strong>а</strong><br />

(Carpenter, 1943). 381. Anomalogramma parvum Carpenter; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 6,0, н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1943;<br />

интерпрет<strong>а</strong>ция жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния О. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>о</strong>й)<br />

Vitvia G. Zalessky, 1933. Тип р<strong>о</strong>д<strong>а</strong> —<br />

V. densinervosa G. Zalessky, 1933; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

О<strong>к</strong><strong>о</strong>нч<strong>а</strong>ние SC и р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ление RS н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; RS трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; MP и СиА <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее SC; м<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> гу<strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 30 мм (ри<strong>с</strong>.<br />

376). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Pseudogramma Carpenter, 1943. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Euthygramma aberrans Martynov,<br />

1938; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>о</strong><strong>к</strong><strong>а</strong>н-<br />

http://jurassic.ru/

чи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее, a RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; MP и<br />

CuA <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

SC; А! <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рььл<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 17,5 мм (ри<strong>с</strong>. 377).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Vilviopsis Martynov, 1938. Тип р<strong>о</strong>д<strong>а</strong> —<br />

V. extensa Martynov, 1938; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ее, трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; MP и CuA <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

<strong>с</strong>ередине <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; А- <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 28 мм (ри<strong>с</strong>. 378).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Aspidoneura Carpenter, 1943.<br />

СЕМЕЙСТВО EUTHYGRAMMATIDAE MARTYNOV,<br />

1928<br />

Переднее «рыл<strong>о</strong>. RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине длины «рыл<strong>а</strong>; <strong>с</strong>ближены CuA и<br />

CuP. В. пермь. Один р<strong>о</strong>д.<br />

Enthygramma Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. parallettum Martynov, 1928; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuA и CuP <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; Ai <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее CuA. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 21—<br />

24 мм (ри<strong>с</strong>. 379). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО CALONEURIDAE HANDLIRSCH, 1906<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине длины «рыл<strong>а</strong> или не­<br />

мн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее ее, <strong>о</strong>бр<strong>а</strong>зуя <strong>о</strong>т четырех д<strong>о</strong><br />

<strong>с</strong>еми <strong>в</strong>ет<strong>в</strong>ей; <strong>с</strong>ближенные MP и CuA <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong> или немн<strong>о</strong>г<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее ее. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 28—50 мм<br />

(ри<strong>с</strong>. 373). Ср. — <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы. Четыре<br />

р<strong>о</strong>д<strong>а</strong>.<br />

Вне СССР: Caloneura Brongniart, 1885;<br />

Confusio Handlirsch, 1919; Homaloptila Meunier,<br />

1911; Euthyneura Pruvost, 1919.<br />

СЕМЕЙСТВО PERMOBIELLIDAE TILLYARD, 1937<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; R из<strong>о</strong>гнут<strong>а</strong><br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

ее, трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; MP и<br />

CuA <strong>в</strong> <strong>в</strong>ершине немн<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнуты; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены ред<strong>к</strong><strong>о</strong>, и <strong>с</strong>реди них мн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>ых.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 11 мм (ри<strong>с</strong>. 380). Один р<strong>о</strong>д.<br />

Н. пермь С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО ANOMALOGRAMMATIDAE CARPENTER,<br />

1943<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>;<br />

R п<strong>о</strong>чти прям<strong>а</strong>я, <strong>в</strong> <strong>в</strong>ершине не из<strong>о</strong>гнут<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>в</strong> н<strong>а</strong>ч<strong>а</strong>ле ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и; МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м;<br />

<strong>с</strong>ближенные MP и CuA <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и ред<strong>к</strong>ие, прямые.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 9 мм (ри<strong>с</strong>. 381). Н. пермь<br />

С. Амери<strong>к</strong>и. Один р<strong>о</strong>д Anomalogramma Carpenter,<br />

1943.<br />

НАДОТРЯД ORTHOPTEROIDEA.<br />

П Р Я М О К Р Ы Л О О Б Р А З Н Ы Е<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних <strong>к</strong>рылье<strong>в</strong> мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong>и<br />

<strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>: R <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т RS, п<strong>о</strong><strong>с</strong>ледняя<br />

жил<strong>к</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> гребнем п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>в</strong>ет<strong>в</strong>ей. Гетер<strong>о</strong>н<strong>о</strong>мия <strong>к</strong>рылье<strong>в</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> рез<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>: з<strong>а</strong>дние <strong>к</strong>рылья р<strong>а</strong><strong>с</strong>ширенные,<br />

передние — уз<strong>к</strong>ие и упл<strong>о</strong>тненные. Ан<strong>о</strong>юг<strong>а</strong>льный<br />

<strong>в</strong>еер з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong>рупный<br />

и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен <strong>в</strong>ет<strong>в</strong>ями <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>.<br />

Ф<strong>о</strong>рмы бе<strong>с</strong><strong>к</strong>рылые и «<strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong>. Тел<strong>о</strong> удлиненн<strong>о</strong>е, цилиндриче­<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

144<br />

http://jurassic.ru/<br />

<strong>с</strong><strong>к</strong><strong>о</strong>е, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>д<strong>а</strong><strong>в</strong>ленн<strong>о</strong>е <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>, реже упл<strong>о</strong>щенн<strong>о</strong>е.<br />

Н<strong>о</strong>ги х<strong>о</strong>дильные, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong>дние прыг<strong>а</strong>тельные;<br />

передние и <strong>с</strong>редние, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

з<strong>а</strong>дних. С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итым яйце<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м,<br />

реже яйце<strong>к</strong>л<strong>а</strong>д редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н. Цер<strong>к</strong>и<br />

<strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, нечлени<strong>с</strong>тые.<br />

Дре<strong>в</strong>няя групп<strong>а</strong>, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енн<strong>а</strong>я<br />

примити<strong>в</strong>ным т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>я<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>тщепили<strong>с</strong>ь <strong>в</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>м пери<strong>о</strong>де.<br />

И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие прям<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>з-

ных <strong>о</strong>пределил<strong>о</strong><strong>с</strong>ь, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лением<br />

<strong>с</strong><strong>в</strong>язей <strong>с</strong> р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>тью, <strong>с</strong>реди <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

<strong>с</strong>т<strong>а</strong>ли <strong>о</strong>бит<strong>а</strong>ть эти н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые; <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне прям<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные-—<strong>о</strong>дн<strong>а</strong> из пр<strong>о</strong>ц<strong>в</strong>ет<strong>а</strong>ющих<br />

групп <strong>с</strong>реди н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> неп<strong>о</strong>лным<br />

ОТРЯД PROTORTHOPTERA<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле неб<strong>о</strong>льш<strong>о</strong>е, без <strong>к</strong><strong>а</strong><strong>к</strong>ихлиб<strong>о</strong><br />

<strong>с</strong>лед<strong>о</strong><strong>в</strong> жил<strong>о</strong><strong>к</strong> <strong>в</strong> нем. Чет<strong>к</strong><strong>о</strong> дифференцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я,<br />

з<strong>а</strong>ним<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong><strong>о</strong>гнут<strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение<br />

МАг <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; CuP <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>о</strong>бычн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. Ai и А 2 <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я п<strong>о</strong> <strong>в</strong><strong>с</strong>ей<br />

длине <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, <strong>о</strong>бр<strong>а</strong>зуя б<strong>о</strong>льш<strong>о</strong>е<br />

чи<strong>с</strong>л<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей. Тел<strong>о</strong> упл<strong>о</strong>щен<strong>о</strong> д<strong>о</strong>р<strong>с</strong><strong>о</strong>-<strong>в</strong>ентр<strong>а</strong>ль'н<strong>о</strong>;<br />

<strong>к</strong>рылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е леж<strong>а</strong>т пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>пине. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong> длину.<br />

Бедр<strong>а</strong> з<strong>а</strong>дних н<strong>о</strong>г ин<strong>о</strong>гд<strong>а</strong> ут<strong>о</strong>лщены, н<strong>о</strong>, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

бег<strong>а</strong>тельные или <strong>в</strong> <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong>й <strong>с</strong>тепени<br />

<strong>в</strong>ып<strong>о</strong>лняющие пры-г<strong>а</strong>тельную фун<strong>к</strong>цию.<br />

Г<strong>о</strong>лени з<strong>а</strong>дних н<strong>о</strong>г без пр<strong>а</strong><strong>в</strong>ильных ряд<strong>о</strong><strong>в</strong> шип<strong>о</strong><strong>в</strong><br />

п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю. К<strong>а</strong>рб<strong>о</strong>н — пермь. Одн<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Sthenaropodidae.<br />

СЕМЕЙСТВО STHENAROPODIDAE HANDLIRSCH,<br />

1906<br />

Крылья шир<strong>о</strong><strong>к</strong>ие; передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый. Передняя <strong>в</strong>ет<strong>в</strong>ь<br />

МА <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>с</strong><strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м<br />

пре<strong>в</strong>р<strong>а</strong>щением, н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бильн<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>а</strong>я<br />

<strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>тях <strong>с</strong> теплым и ж<strong>а</strong>р<strong>к</strong>им <strong>к</strong>лим<strong>а</strong>т<strong>о</strong>м.<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. Отряды: Protorthoptera,<br />

Orthoptera (= Saltatoria), Phasmatodea, Glosselytrodea,<br />

из них пер<strong>в</strong>ый и п<strong>о</strong><strong>с</strong>ледний — <strong>в</strong>ымершие.<br />

Aj CuP CuA 2<br />

383<br />

Ри<strong>с</strong>. 382—383. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Sthenaropodidae<br />

RS. Ан<strong>а</strong>СТОМОЗ MP СиА р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен ИЛИ 3 8 2. Adumbratus extentus Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,2, н. пермь,<br />

Н<strong>а</strong> р<strong>а</strong>ВНОМ р<strong>а</strong>ССТОЯ'НИИ ОТ Переднег<strong>о</strong> И З<strong>а</strong>ИНеГО Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961). 383. Kamia angustovenosa<br />

г r<br />

, Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,2, <strong>в</strong>. пермь, Приур<strong>а</strong>лье<br />

Кр<strong>а</strong>е<strong>в</strong> КрЫЛ<strong>а</strong> ИЛИ бЛИЖе К Переднему Кр<strong>а</strong>ю. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1930)<br />

Ст<strong>в</strong><strong>о</strong>л MP + СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong><br />

Ри<strong>с</strong>. 384. Sihenaropoda fischeri Brongniart<br />

ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция; х 1,3, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1960<strong>а</strong>)<br />

10 <strong>о</strong> <strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 145<br />

http://jurassic.ru/

п<strong>о</strong> <strong>о</strong>тн<strong>о</strong>шению <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; <strong>о</strong>бр<strong>а</strong>зует<br />

четыре — ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей. К<strong>а</strong>рб<strong>о</strong>н — пермь. Де<strong>в</strong>ять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Adumbratus Sharov, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. extentus Sharov, 1961; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ными жил<strong>к</strong><strong>а</strong>ми п<strong>о</strong><br />

д<strong>в</strong><strong>а</strong>-три ряд<strong>а</strong> ячее<strong>к</strong>; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>м<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; R <strong>в</strong><br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я; RS<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и; <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з MP —<br />

CuA р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен н<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т переднег<strong>о</strong><br />

и з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong> <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 25—27 мм (ри<strong>с</strong>. 382; т<strong>а</strong>бл. IX,<br />

фиг. 3). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kamia Martynov, 1928 (Spongoneura<br />

Martynov, 1928; Permacridites Martynov, 1930).<br />

Тип р<strong>о</strong>д<strong>а</strong> — К- angustovenosa Martynov, 1928;<br />

<strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Тихие Г<strong>о</strong>ры). Переднее <strong>к</strong>рыл<strong>о</strong>. Между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

SC идет п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; R <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти не б<strong>о</strong>лее чем <strong>с</strong> д<strong>в</strong>умя-тремя<br />

<strong>в</strong>ет<strong>в</strong>ями; RS <strong>о</strong>бр<strong>а</strong>зует д<strong>о</strong> ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей; МА<br />

пр<strong>о</strong>х<strong>о</strong>дит <strong>о</strong>чень близ<strong>к</strong><strong>о</strong> ж R и RS; <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ж<strong>а</strong>т<strong>а</strong>,<br />

<strong>о</strong>бр<strong>а</strong>зуя д<strong>о</strong> четырех гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

<strong>в</strong>ет<strong>в</strong>ей; между пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми<br />

п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> имеют<strong>с</strong>я <strong>в</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

жил<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующие <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> п<strong>о</strong>перечными<br />

ч<strong>а</strong><strong>с</strong>тую <strong>с</strong>еть. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

60—65 мм (ри<strong>с</strong>. 383, т<strong>а</strong>бл. IX, фиг. 4). Один<br />

<strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Вне СССР: Sthenaropoda Brongniart, 1885<br />

(=» Archaeacridites Meunier, 1909) (ри<strong>с</strong>.384),<br />

Macrophlebium Goldenberg, 1869; Berlichia<br />

Schlechtendal, 1913; Acridites Germar, 1842;<br />

Palaeoedischia Meunier, 1914; Nacekomia Richardson,<br />

1956; ?Pseudooedischia Handlirsch,<br />

1919.<br />

ОТРЯД ORTHOPTERA. ПРЯМОКРЫЛЫЕ<br />

(Saltatoria)<br />

Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>бширн<strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

<strong>в</strong>ет<strong>в</strong>ями <strong>к</strong><strong>о</strong><strong>с</strong>ты и пре<strong>к</strong><strong>о</strong><strong>с</strong>ты <strong>в</strong> нем.<br />

Имеет<strong>с</strong>я чет<strong>к</strong><strong>о</strong> дифференцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>а</strong>я<br />

<strong>в</strong><strong>о</strong>гнут<strong>о</strong> МА 2. CuP и Ai не <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я;<br />

А 2 <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>, и ее<br />

<strong>в</strong>ет<strong>в</strong>и п<strong>о</strong>д<strong>о</strong>бны А], имеют <strong>в</strong>ид д<strong>в</strong>ух-трех<br />

пр<strong>о</strong><strong>с</strong>тых п<strong>а</strong>р<strong>а</strong>ллельных жил<strong>о</strong><strong>к</strong>. В з<strong>а</strong>дних<br />

<strong>к</strong>рыльях передняя <strong>в</strong>ет<strong>в</strong>ь А 2 пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, без гребня<br />

жил<strong>о</strong><strong>к</strong>. Тел<strong>о</strong> упл<strong>о</strong>щенн<strong>о</strong>е <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> или <strong>к</strong>ругл<strong>о</strong>е<br />

<strong>в</strong> <strong>с</strong>ечении; <strong>к</strong>рылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е леж<strong>а</strong>т п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

тел<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я. Передне<strong>о</strong>лин<strong>к</strong><strong>а</strong><br />

<strong>с</strong>едл<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, реже <strong>с</strong> длинными <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми.<br />

З<strong>а</strong>дние н<strong>о</strong>ги прыг<strong>а</strong>тельные, их бедр<strong>а</strong><br />

ут<strong>о</strong>лщенные; г<strong>о</strong>лени п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>с</strong><br />

д<strong>в</strong>умя ряд<strong>а</strong>ми шип<strong>о</strong><strong>в</strong>. Яйц<strong>а</strong> при п<strong>о</strong>м<strong>о</strong>щи яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong><br />

<strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>у, реже <strong>в</strong> <strong>с</strong>тебли и<br />

п<strong>а</strong>зухи ли<strong>с</strong>тье<strong>в</strong>. К<strong>а</strong>рб<strong>о</strong>н — ныне. П<strong>о</strong>д<strong>о</strong>тряды:<br />

Ensifera, Caelifera<br />

ПОДОТРЯД ENSIFERA<br />

Антенны ните<strong>в</strong>идные, длиннее тел<strong>а</strong>; чи<strong>с</strong>л<strong>о</strong><br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> б<strong>о</strong>лее 30; члени<strong>к</strong>и <strong>о</strong>тгр<strong>а</strong>ничены <strong>о</strong>дин<br />

<strong>о</strong>т друг<strong>о</strong>г<strong>о</strong> <strong>с</strong>л<strong>а</strong>быми н<strong>а</strong><strong>с</strong>еч<strong>к</strong><strong>а</strong>ми, без рез<strong>к</strong>их<br />

перех<strong>в</strong><strong>а</strong>т<strong>о</strong><strong>в</strong>. З<strong>а</strong>дние бедр<strong>а</strong> гл<strong>а</strong>д<strong>к</strong>ие, без пр<strong>о</strong>д<strong>о</strong>льных<br />

<strong>к</strong>илей. Яйце<strong>к</strong>л<strong>а</strong>д длинный, <strong>с</strong><strong>а</strong>бле<strong>в</strong>идный,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленный <strong>к</strong> пр<strong>о</strong><strong>к</strong><strong>а</strong>лы<strong>в</strong><strong>а</strong>нию или<br />

пр<strong>о</strong>пили<strong>в</strong><strong>а</strong>нию <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>а</strong>, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рый <strong>о</strong>т<strong>к</strong>л<strong>а</strong>­<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

146<br />

http://jurassic.ru/<br />

ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я яйц<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н — ныне. Три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Oedischiidea, Gryllidea, Tettigoniidea.<br />

НАДСЕМЕЙСТВО OEDISCHIIDEA<br />

Передние <strong>к</strong>рылья уз<strong>к</strong>ие, длинные, переп<strong>о</strong>нч<strong>а</strong>тые.<br />

Вет<strong>в</strong>ей МА, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, б<strong>о</strong>лее<br />

трех; <strong>в</strong>ет<strong>в</strong>и MP + CuA, е<strong>с</strong>ли имеют<strong>с</strong>я, т<strong>о</strong><br />

<strong>о</strong>бычн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены гребенч<strong>а</strong>т<strong>о</strong>. З<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т н<strong>а</strong> передних <strong>к</strong>рыльях <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует или<br />

<strong>в</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии. Тимп<strong>а</strong>н<strong>а</strong>льные <strong>о</strong>рг<strong>а</strong>ны<br />

<strong>в</strong> г<strong>о</strong>ленях передних н<strong>о</strong>г <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. К<strong>а</strong>рб<strong>о</strong>н<br />

— юр<strong>а</strong>. Ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Oedischiidae,<br />

Tcholmanvissiidae, Permelcanidae, Elcanidae,<br />

Permorhaphidiidae, Bintoniellidae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних<br />

<strong>в</strong>не СССР.<br />

СЕМЕЙСТВО OEDISCHIIDAE HANDLIRSCH, 1906<br />

(Pruvostitidae M. Zalessky, 1928)<br />

Передняя <strong>в</strong>ет<strong>в</strong>ь MAi <strong>в</strong> переднем <strong>к</strong>рыле <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>с</strong> RS; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льный уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong><br />

RS (д<strong>о</strong> <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong>) длинный, <strong>в</strong> <strong>в</strong>иде я<strong>с</strong>н<strong>о</strong>й<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й жил<strong>к</strong>и — <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RS; между<br />

ним и R, <strong>а</strong> т<strong>а</strong><strong>к</strong>же МА] имеют<strong>с</strong>я п<strong>о</strong>перечные<br />

жил<strong>к</strong>и; MAi и ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS<br />

(п<strong>о</strong><strong>с</strong>ле <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong>) не имеют х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>а</strong> еди-

Ри<strong>с</strong>. 385—389. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Oedischiidae<br />

385. Oedischia williamsoni Brongniart; A — переднее <strong>к</strong>рыл<strong>о</strong>, ЛЫ;<br />

Б — з<strong>а</strong>дние| н<strong>о</strong>ги, х 1,1; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1906). 386. Jasvia reticulata G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, x 1,2,<br />

н. пермь. Ур<strong>а</strong>л (G. Zalessky, 1934). 387. Rlmnosentomon grande<br />

G. Zalessky; фр<strong>а</strong>гмент переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, x 1,4, н. Пермь, Ур<strong>а</strong>л<br />

н<strong>о</strong>й жил<strong>к</strong>и. MP + CuA <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> гребнем <strong>в</strong>ет<strong>в</strong>ей,<br />

идущих <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н—<br />

пермь. Де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Oedischia Brongniart, 1885. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Ое. williamsoni Brongniart, 1885; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Фр<strong>а</strong>нция (<strong>с</strong>теф<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, шир<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>в</strong>ет<strong>в</strong>и<br />

SC пр<strong>о</strong><strong>с</strong>тые; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти R ше<strong>с</strong>ть-<strong>с</strong>емь<br />

пр<strong>о</strong><strong>с</strong>тых <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь MAi<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бр<strong>а</strong>зует три <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>в</strong>ет<strong>в</strong>и; три <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичных <strong>в</strong>ет<strong>в</strong>и <strong>о</strong>бр<strong>а</strong>зует и МА 2;<br />

MP + CuAi <strong>с</strong> пятью <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>к</strong>рут<strong>о</strong> идущими <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; между пр<strong>о</strong>д<strong>о</strong>льными — пр<strong>о</strong><strong>с</strong>тые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

62 мм (ри<strong>с</strong>. 385 А, Б). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Фр<strong>а</strong>нции.<br />

Jasvia G. Z <strong>а</strong> 1 е s s k у,1934. Тип р<strong>о</strong>д<strong>а</strong> — /.<br />

reticulata G. Zalessky, 1934; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Язь<strong>в</strong><strong>а</strong>). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле удлиненн<strong>о</strong>е, уз<strong>к</strong><strong>о</strong>е; <strong>в</strong>ет<strong>в</strong>и SC и R<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; д<strong>о</strong> ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и МАг, МА 2 не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; MP + CuAi <strong>с</strong> пятью п<strong>о</strong>л<strong>о</strong>г<strong>о</strong> идущими<br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>в</strong>ет<strong>в</strong>ями; <strong>в</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют С- и S-<strong>о</strong>бр<strong>а</strong>зные<br />

п<strong>о</strong>перечные жил<strong>к</strong>и, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й — п<strong>о</strong><br />

три-четыре ряд<strong>а</strong> мел<strong>к</strong>их ячее<strong>к</strong> между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

(Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1955). 388. Pruvostites takhtachurensis М. Zalessky;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 1,3, <strong>в</strong>. пермь. Приур<strong>а</strong>лье (М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1928). 389. Metoedischla magnlftca Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 2,2, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928)<br />

61 мм (ри<strong>с</strong>. 386; т<strong>а</strong>бл. X, фиг. 1). Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Rlmnosentomon G. Zalessky, 1955. Тип<br />

р<strong>о</strong>д<strong>а</strong> — R. grande G. Zalessky, 1955; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, М<strong>о</strong>р<strong>о</strong>з<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>).<br />

Вет<strong>в</strong>и R мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны, пр<strong>о</strong>х<strong>о</strong>дят те<strong>с</strong>н<strong>о</strong><br />

друг <strong>к</strong> другу; RS п<strong>о</strong><strong>с</strong>ле <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> MAi<br />

из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь<br />

MAi <strong>с</strong> шир<strong>о</strong><strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>в</strong>ет<strong>в</strong>и MP + CuAi<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближены. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 80 мм (ри<strong>с</strong>. 387).<br />

Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Pruvostites М. Zalessky, 1928. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. takhtachurensis М. Zalessky, 1928; <strong>в</strong>.<br />

пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Т<strong>а</strong>хт<strong>а</strong>шур). Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый. Отдельные <strong>в</strong>ет<strong>в</strong>и SC <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми;<br />

Д<strong>в</strong>е <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные <strong>в</strong>ет<strong>в</strong>и MAf, MP не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; MP + CuAi <strong>о</strong>бр<strong>а</strong>зует ше<strong>с</strong>ть-<strong>с</strong>емь<br />

<strong>в</strong>ет<strong>в</strong>ей. П<strong>о</strong>перечные жил<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые или Y-<strong>о</strong>бр<strong>а</strong>зные,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные те<strong>с</strong>н<strong>о</strong> друг <strong>к</strong> другу.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60 мм (ри<strong>с</strong>.<br />

388). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Metoedischia Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. magnifica Martynov, 1928; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Вет<strong>в</strong>и SC и R без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>; RS гребенч<strong>а</strong>т<strong>а</strong>я;<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь MAj <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; МА 2 не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; MP + CuA пр<strong>о</strong>х<strong>о</strong>дит п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

147 10*<br />

http://jurassic.ru/

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю и <strong>о</strong>бр<strong>а</strong>зует <strong>в</strong><strong>о</strong><strong>с</strong>емь-де<strong>в</strong>ять<br />

<strong>в</strong>ет<strong>в</strong>ей; преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые, ред<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

п<strong>о</strong>перечные жил<strong>к</strong>и; д<strong>в</strong>у- и трехрядные<br />

ячей<strong>к</strong>и между <strong>в</strong>ет<strong>в</strong>ями М близ з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

35 мм (ри<strong>с</strong>. 389; т<strong>а</strong>бл. X, фиг. 2). Три <strong>в</strong>ид<strong>а</strong>.<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н П<strong>о</strong>ртуг<strong>а</strong>лии; <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong>не СССР: ? Genentomum<br />

Scudder, 1885; ? Pro genentomum Handlirsch,<br />

1906; Plesioidischia Schlechtendal, 1913; Permoedischia<br />

Kukalova, 1955.<br />

СЕМЕЙСТВО TCHOLMANV1SS11DAE G. ZALESSKY,<br />

1934<br />

В переднем <strong>к</strong>рыле пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е. В пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>в</strong>ет<strong>в</strong>и SC <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>, п<strong>о</strong>чти перпенди<strong>к</strong>улярных С и SC. SC<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>. R<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong> ше<strong>с</strong>тью — <strong>в</strong><strong>о</strong><strong>с</strong>емью длинными <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>о</strong>тх<strong>о</strong>дящими <strong>о</strong>т R п<strong>о</strong>д неб<strong>о</strong>льшим угл<strong>о</strong>м;<br />

между ними мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные пр<strong>о</strong><strong>с</strong>тые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong>ильн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширен<strong>о</strong> <strong>в</strong> центр<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти, <strong>с</strong> С-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

из<strong>о</strong>гнутыми п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>щенными<br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>о</strong>й <strong>к</strong> <strong>в</strong>ершине, те<strong>с</strong>н<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными друг <strong>к</strong> другу и ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong>единенными<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ыми жил<strong>к</strong><strong>а</strong>ми. RS не <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью МА. MAi и<br />

МА 2 <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я <strong>о</strong>чень п<strong>о</strong>здн<strong>о</strong> и <strong>о</strong>бр<strong>а</strong>зуют п<strong>о</strong> д<strong>в</strong>е<br />

С<br />

<strong>в</strong>ет<strong>в</strong>и, или же MAi <strong>о</strong>бр<strong>а</strong>зует три <strong>в</strong>ет<strong>в</strong>и, <strong>а</strong> МА 2<br />

не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; между MAi и МА 2 <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

S-<strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong>перечные жил<strong>к</strong>и. Пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> MP — CuA имеет<strong>с</strong>я <strong>о</strong>т<br />

д<strong>в</strong>ух д<strong>о</strong> трех <strong>в</strong>ет<strong>в</strong>ей СиА. Три <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и между их <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми и<br />

з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем «рыл<strong>а</strong> С-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнуты <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>о</strong>й <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

между <strong>в</strong>ет<strong>в</strong>ями RS и МА <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>к</strong>рыл<strong>а</strong> и ин<strong>о</strong>гд<strong>а</strong> п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти имеет<strong>с</strong>я д<strong>в</strong><strong>а</strong>-три ряд<strong>а</strong> ячее<strong>к</strong>. В з<strong>а</strong>днем<br />

<strong>к</strong>рыле R и RS т<strong>а</strong><strong>к</strong>же не <strong>о</strong>бр<strong>а</strong>зуют <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong>.<br />

Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненными <strong>к</strong> <strong>в</strong>ершине<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Пермь. Один р<strong>о</strong>д.<br />

Pinegia Martynov, 1928 (Thnetodes<br />

Martynov, 1928; Tcholmanvissia M. Zalessky,<br />

1929; Kamaites M. Zalessky, 1929, Tillyardiella<br />

Martynov, 1930). Тип р<strong>о</strong>д<strong>а</strong> — P. oknowae Martynov,<br />

1928; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е<br />

и длинн<strong>о</strong>е, <strong>в</strong> <strong>с</strong>рединн<strong>о</strong>й трети <strong>с</strong> прямым<br />

передним <strong>к</strong>р<strong>а</strong>ем; длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>в</strong> 6—7 р<strong>а</strong>з. Вет<strong>в</strong>и SC <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong>, и между ними нет пр<strong>а</strong><strong>в</strong>ильных<br />

ряд<strong>о</strong><strong>в</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; лишь изред<strong>к</strong><strong>а</strong><br />

<strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я единичные п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

Вет<strong>в</strong>и MP + CuA р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены те<strong>с</strong>н<strong>о</strong> друг <strong>к</strong><br />

другу и п<strong>о</strong>д <strong>о</strong>чень м<strong>а</strong>лым угл<strong>о</strong>м <strong>к</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 60—<br />

90 мм (ри<strong>с</strong>. 390, 391; т<strong>а</strong>бл. X, фиг. 3—4).<br />

Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>, <strong>в</strong>. пермь Приур<strong>а</strong>лья<br />

и Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Aj CuP CuA 2 MP + CuA, MA 2 MA,<br />

CuA<br />

Ри<strong>с</strong>. 390—391. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tcholmanvissiidae<br />

390. Pinegia longipes (Martynov); переднее <strong>к</strong>рыл<strong>о</strong>, x 1,4; н. пермь, Ур<strong>а</strong>л (<strong>о</strong>риг. ри<strong>с</strong>.).391. Pinegia oknowae<br />

Martynov; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, х 4,4, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928)<br />

148<br />

http://jurassic.ru/<br />

391<br />

390

CuAj<br />

Ри<strong>с</strong>. 392—395. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Permelcanidae и Elcanidae<br />

392. Proelcana uralica Sharov; А — переднее <strong>к</strong>рыл<strong>о</strong>, x 4,6;Б —<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 4,6; н. пермь, Ур<strong>а</strong>л (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1962). 393. Permel<strong>с</strong><strong>а</strong>п<strong>а</strong><br />

sojanense Sharov; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 6,0; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1962). 394. Elcana<br />

СЕМЕЙСТВО PERMELCANIDAE SHAROV, 1962<br />

Мел<strong>к</strong>ие прям<strong>о</strong><strong>к</strong>рылые <strong>с</strong> т<strong>о</strong>н<strong>к</strong>ими переп<strong>о</strong>нч<strong>а</strong>тыми<br />

<strong>к</strong>рыльями. SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не<br />

н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее.<br />

Уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

SC — уз<strong>к</strong>ий, <strong>о</strong>бычн<strong>о</strong> пигментир<strong>о</strong><strong>в</strong><strong>а</strong>н, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными п<strong>о</strong>д угл<strong>о</strong>м 60° и<br />

б<strong>о</strong>лее <strong>в</strong>ет<strong>в</strong>ями R. Передняя <strong>в</strong>ет<strong>в</strong>ь МА <strong>в</strong><br />

переднем и з<strong>а</strong>днем <strong>к</strong>рыльях <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<br />

<strong>с</strong> RS; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льный уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> RS (д<strong>о</strong><br />

<strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong>) <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет <strong>к</strong><strong>о</strong><strong>с</strong>ую<br />

п<strong>о</strong>перечную жил<strong>к</strong>у; между нею и R, <strong>а</strong> т<strong>а</strong><strong>к</strong>же<br />

МА п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле ред<strong>к</strong>ие пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и. MAi и ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS<br />

(п<strong>о</strong><strong>с</strong>ле <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong>) <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют единую жил<strong>к</strong>у.<br />

Ст<strong>в</strong><strong>о</strong>л MP + CuA <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>о</strong>чень п<strong>о</strong>здн<strong>о</strong> и<br />

<strong>о</strong>бр<strong>а</strong>зует не б<strong>о</strong>лее д<strong>в</strong>ух-трех <strong>в</strong>ет<strong>в</strong>ей. Ан<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong>чень длинн<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong>я приблизительн<strong>о</strong><br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими л<strong>о</strong>жными <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>в</strong><strong>о</strong>зни<strong>к</strong>шими из п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. В переднем<br />

и з<strong>а</strong>днем <strong>к</strong>рыльях пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют пр<strong>о</strong><strong>с</strong>тые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и, лишь <strong>в</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong><br />

149<br />

395<br />

reticulata Handlirsch; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,8, н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1925). 395. Elcana sp. (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); х 2,7,н. юр<strong>а</strong>.<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925)<br />

http://jurassic.ru/<br />

имеют<strong>с</strong>я <strong>к</strong>рупные <strong>с</strong><strong>о</strong>т<strong>о</strong><strong>в</strong>идные ячей<strong>к</strong>и. Н. и <strong>в</strong>.<br />

пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Proelcana Sharov, 1962. Тип р<strong>о</strong>д<strong>а</strong> —Р.<br />

uralica Sharov, 1962; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Передний <strong>к</strong>р<strong>а</strong>й<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый. SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS; RS <strong>о</strong>бр<strong>а</strong>зует д<strong>о</strong> четырех <strong>в</strong>ет<strong>в</strong>ей; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь берет н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> близ<strong>к</strong><strong>о</strong> <strong>к</strong> н<strong>а</strong>ч<strong>а</strong>лу<br />

RS, н<strong>а</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии, меньшем ширины р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; д<strong>в</strong>е <strong>с</strong><strong>а</strong>мые ди<strong>с</strong>т<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и идут<br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> изгиб<strong>а</strong>я<strong>с</strong>ь н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; е<strong>с</strong>ть <strong>о</strong>дн<strong>а</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь MAi. В з<strong>а</strong>днем <strong>к</strong>рыле SC<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; <strong>в</strong>ет<strong>в</strong>и<br />

RS имеют <strong>с</strong>х<strong>о</strong>дн<strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение <strong>с</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>в</strong> переднем <strong>к</strong>рыле; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь MAi<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

15 мм (ри<strong>с</strong>. 392). Один <strong>в</strong>ид. Н. пермь<br />

Ур<strong>а</strong>л<strong>а</strong>.<br />

Permelcana Sharov, 1962; Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. sojanense Sharov, 1962; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Передний<br />

<strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й. Пре<strong>к</strong><strong>о</strong>-

<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями<br />

С <strong>в</strong> нем; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут я<strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю и <strong>о</strong>бр<strong>а</strong>зует пять-ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь берет н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> н<strong>а</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>т н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS, пре<strong>в</strong>ыш<strong>а</strong>ющем ширину р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; д<strong>в</strong>е <strong>с</strong><strong>а</strong>мые ди<strong>с</strong>т<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и<br />

р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ляя<strong>с</strong>ь <strong>к</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные <strong>в</strong>ет<strong>в</strong>и MAi <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. В з<strong>а</strong>днем<br />

<strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS; <strong>в</strong>ет<strong>в</strong>и RS и МА имеют т<strong>а</strong><strong>к</strong><strong>о</strong>е же р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и <strong>в</strong> переднем <strong>к</strong>рыле. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 12—13 мм (ри<strong>с</strong>. 393; т<strong>а</strong>бл.<br />

XI, фиг. 1). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

Вне СССР: ? Elcanopsis Tillyard, 1918.<br />

СЕМЕЙСТВО ELCANIDAE HANDLIRSCH, 1906<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>я<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong><strong>о</strong>гнуты, <strong>о</strong>бр<strong>а</strong>зуя перетяж<strong>к</strong>у н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong>. В пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>в</strong>переди<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ты и п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> ей пр<strong>о</strong>х<strong>о</strong>дит еще <strong>о</strong>дн<strong>а</strong><br />

жил<strong>к</strong><strong>а</strong> (пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>?), <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>о</strong>тх<strong>о</strong>дят <strong>в</strong>перед<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>в</strong>ет<strong>в</strong>и. SC <strong>с</strong> гребнем <strong>в</strong>ет<strong>в</strong>ей <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти; <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS. Ст<strong>в</strong><strong>о</strong>лы МА и RS пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют единую<br />

жил<strong>к</strong>у, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>о</strong>тх<strong>о</strong>дят <strong>в</strong> <strong>в</strong>иде гребня<br />

н<strong>а</strong>з<strong>а</strong>д <strong>в</strong>ет<strong>в</strong>и RS и МА. MP + CuAi не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

или <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й-д<strong>в</strong>умя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>о</strong>ч<strong>к</strong><strong>а</strong>ми,<br />

идущими <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> и имеющими<br />

<strong>в</strong>ид п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. СиА <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

CuP близ<strong>к</strong><strong>о</strong> <strong>к</strong> н<strong>а</strong>ч<strong>а</strong>лу СиА 2; п<strong>о</strong><strong>с</strong>ледняя пр<strong>о</strong>х<strong>о</strong>дит<br />

<strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>к</strong> CuP. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>в</strong> длину зн<strong>а</strong>чительн<strong>о</strong> меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины<br />

<strong>к</strong>рыл<strong>а</strong>. В з<strong>а</strong>днем <strong>к</strong>рыле R, RS, МА и их<br />

<strong>в</strong>ет<strong>в</strong>и имеют т<strong>а</strong><strong>к</strong><strong>о</strong>е же п<strong>о</strong>л<strong>о</strong>жение, <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>в</strong> переднем<br />

<strong>к</strong>рыле. СиА <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т СиР <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong>. Ан<strong>а</strong>льный <strong>в</strong>еер т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> <strong>к</strong>рыл<strong>а</strong>. Антенны длинные,<br />

ните<strong>в</strong>идные. З<strong>а</strong>дние н<strong>о</strong>ги <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> ут<strong>о</strong>лщенными<br />

бедр<strong>а</strong>ми; шипы п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю г<strong>о</strong>лени<br />

упл<strong>о</strong>щенные, <strong>в</strong> <strong>в</strong>иде пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>. Длинный<br />

яйце<strong>к</strong>л<strong>а</strong>д, р<strong>а</strong><strong>в</strong>ный п<strong>о</strong> длине брюш<strong>к</strong>у (ри<strong>с</strong>.<br />

394, 395). Юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> (б<strong>о</strong>лее 100 <strong>в</strong>ид<strong>о</strong><strong>в</strong>)<br />

<strong>в</strong>не СССР.<br />

OEDISCHIIDEA INCERTAE SEDIS<br />

Thnetus Handlirsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Т. stuckenbergi Handlirsch, 1904; <strong>в</strong>. пермь, Кир<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Шуня). Фр<strong>а</strong>гмент<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м МА и MP +<br />

+ CuA. У з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми п<strong>о</strong> д<strong>в</strong>е-три интер<strong>к</strong><strong>а</strong>лярные<br />

жил<strong>к</strong>и, <strong>с</strong><strong>о</strong>единенные п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>в</strong> б<strong>о</strong>лее пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти — пр<strong>о</strong><strong>с</strong>тые, Yи<br />

К-<strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong>перечные жил<strong>к</strong>и. Фр<strong>а</strong>гмент<br />

150<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 396. Thnetus stuckenbergi Handlirsch<br />

фр<strong>а</strong>гмент переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, х 2,3, <strong>в</strong>. пермь,<br />

Приур<strong>а</strong>лье (Handlirsch, 1906)<br />

м<strong>о</strong>жет прин<strong>а</strong>длеж<strong>а</strong>ть пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителю <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Oedischiidae, или <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tcholmanvissiidae.<br />

Длин<strong>а</strong> фр<strong>а</strong>гмент<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

13 мм (ри<strong>с</strong>. 396). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Вне СССР: Mesogryllacris Riek, 1955;<br />

?Proparagryllacris Riek, 1956.<br />

НАДСЕМЕЙСТВО GRYLLIDEA<br />

Передние <strong>к</strong>рылья, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, шир<strong>о</strong><strong>к</strong>ие.<br />

Вет<strong>в</strong>ей МА не б<strong>о</strong>лее трех, <strong>о</strong>бычн<strong>о</strong> д<strong>в</strong>е: MAi и<br />

МА 2; <strong>к</strong><strong>а</strong><strong>к</strong> и<strong>с</strong><strong>к</strong>лючение ин<strong>о</strong>гд<strong>а</strong> MAi д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я.<br />

MAi не <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> RS ни <strong>в</strong><br />

передних, ни <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях; ин<strong>о</strong>гд<strong>а</strong> MAi<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>. Вет<strong>в</strong>и MP + CuA, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

не имеют пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>г<strong>о</strong> гребенч<strong>а</strong>т<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жения<br />

п<strong>о</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лению <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>. Три-четыре <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и. Передние<br />

<strong>к</strong>рылья <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>с</strong> з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>ым <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong>м,<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нным CuP и <strong>а</strong>н<strong>а</strong>льными жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнутыми <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ения н<strong>а</strong> пр<strong>а</strong><strong>в</strong><strong>о</strong>м и ле<strong>в</strong><strong>о</strong>м <strong>к</strong>рыльях.<br />

Ин<strong>о</strong>гд<strong>а</strong> з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н.<br />

Г<strong>о</strong>лени передних н<strong>о</strong>г, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong> тимп<strong>а</strong>н<strong>а</strong>льными<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми. Три<strong>а</strong><strong>с</strong>—ныне. Семь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Haglidae, Gryllacrididae, Tchorkuphlebiidae,<br />

Isfaropteridae, Gryllidae, Gryllotalpidae,<br />

Schizodactylidae; п<strong>о</strong><strong>с</strong>леднее <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

СЕМЕЙСТВО HAGLIDAE HANDLIRSCH, 1906<br />

(Pamphagopsidae Martynov, 1925;<br />

Aboilidae Martynov, 1925;<br />

Prophalangopseidae Handlirsch, 1929)<br />

R и RS <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями; <strong>в</strong>ет<strong>в</strong>и RS р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

гребенч<strong>а</strong>т<strong>о</strong> н<strong>а</strong>з<strong>а</strong>д. MAi и МА 2 <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют и д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>.<br />

Вет<strong>в</strong>ей MP + CuA не менее четырех. Вт<strong>о</strong>р<strong>а</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь СиА <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong><br />

<strong>о</strong>т б<strong>о</strong>лее ди<strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей и <strong>о</strong>тх<strong>о</strong>дит<br />

или <strong>о</strong>т ме<strong>с</strong>т<strong>а</strong> <strong>с</strong>лияния MP <strong>с</strong> СиА ИЛИ не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я не д<strong>а</strong>лее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины <strong>к</strong>рыл<strong>а</strong>.<br />

Три<strong>а</strong><strong>с</strong> — ныне. Три п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Haglinae,<br />

Prophalangopsejnae, Cyrtophyllitinae.

397—403. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Haglidae, Gryllacrididae, Tchorkuphlebiidae<br />

97 Hagla gracilis Giebel; передние <strong>к</strong>рылья, x 1,5; A — <strong>с</strong><strong>а</strong>мц<strong>а</strong>; тын<strong>о</strong><strong>в</strong>, 1937). 401. Pseuiohumbtrtiella grandis (Brauer, Redtenba-<br />

Б — <strong>с</strong><strong>а</strong>м<strong>к</strong>и; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Zeuner, 1939). 398. Aboilus colum- cher, Ganglbauer); фр<strong>а</strong>гмент з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, x 2,2, н. юр<strong>а</strong>, Ир<strong>к</strong>утnatus<br />

Martynov; А — переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х 1,5 (<strong>о</strong>риг. ри<strong>с</strong>.); е<strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Brauer, Redtenbacher, Ganglbauer, 1889). 402. Zcune-<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,3 (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925; пере<strong>в</strong>ернут<strong>о</strong> и и<strong>с</strong>- roptera scotica (Zeuner); переднее <strong>к</strong>рыл<strong>о</strong>, х 2,3, третичные <strong>о</strong>тл<strong>о</strong>пр<strong>а</strong><strong>в</strong>лен<strong>о</strong><br />

п<strong>о</strong> <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong>у), <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н. 399. Pseudohagla жения Ш<strong>о</strong>тл<strong>а</strong>ндии (<strong>о</strong>риг. ри<strong>с</strong>. п<strong>о</strong> ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии). 403. Tchorkuph.ltpospelovi<br />

(О. Martynova); переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х 1,9, н. юр<strong>а</strong>, Ы<strong>а</strong> compressa Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,0, н. юр<strong>а</strong>. Ср. Азия<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>о</strong>риг. ри<strong>с</strong>.). 400. Archaboilus shurabicus (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х 1,1, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>р-<br />

ПОДСЕМЕЙСТВО HAGLINAE ZEUNER, 1935<br />

(Prohaglinae Riek, 1954)<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. С х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличим<strong>а</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я;<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и SC р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> С; между С и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>й<br />

ди<strong>с</strong>т<strong>а</strong>льнее пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью SC имеют<strong>с</strong>я<br />

р<strong>а</strong>зличн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> п<strong>о</strong>перечные жил<strong>к</strong>и. Регу­<br />

151<br />

http://jurassic.ru/<br />

лярных интер<strong>к</strong><strong>а</strong>лярных жил<strong>о</strong><strong>к</strong> между <strong>в</strong>ет<strong>в</strong>ями<br />

SC нет. В <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

<strong>к</strong>реп<strong>к</strong><strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> между т<strong>о</strong>ч<strong>к</strong><strong>о</strong>й <strong>с</strong>лияния<br />

MP <strong>с</strong> CuA и <strong>с</strong><strong>а</strong>м<strong>о</strong>й пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью<br />

CuA. Три<strong>а</strong><strong>с</strong> — н. юр<strong>а</strong>. Четыре р<strong>о</strong>д<strong>а</strong>, <strong>в</strong><strong>с</strong>е <strong>в</strong>не<br />

СССР: Prohagla Riek, 1954; Neohagla Riek,<br />

1955; Zalmona Giebel, 1856; Hagla Giebel,<br />

1856 (ри<strong>с</strong>. 397 А, Б).

ПОДСЕМЕЙСТВО PROPHALANGOPSEINAE<br />

CAUDEL, 1911<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong> длинн<strong>а</strong>я, пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ющ<strong>а</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей SC. Регулярные интер<strong>к</strong><strong>а</strong>лярные<br />

жил<strong>к</strong>и между <strong>в</strong>ет<strong>в</strong>ями SC <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

В <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>т ме<strong>с</strong>т<strong>а</strong> <strong>с</strong><strong>о</strong>единения<br />

MP <strong>с</strong> СиА <strong>о</strong>тх<strong>о</strong>дит <strong>к</strong>реп<strong>к</strong><strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я интер<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я н<strong>а</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

<strong>в</strong>ет<strong>в</strong>и CuA. Н. юр<strong>а</strong> — ныне. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Aboilus Martynov, 1925 (Pamphagopsis<br />

Martynov, 1925; Syndesmophyllum Martynov,<br />

1934). Тип р<strong>о</strong>д<strong>а</strong> — A. fasciatus Martynov, 1925;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ьшу<strong>к</strong>лый.<br />

Вт<strong>о</strong>р<strong>а</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь СиА <strong>о</strong>тх<strong>о</strong>дит<br />

<strong>о</strong>т ме<strong>с</strong>т<strong>а</strong> <strong>с</strong><strong>о</strong>единения MP <strong>с</strong> СиА; между<br />

<strong>в</strong>ет<strong>в</strong>ями RS и МА п<strong>о</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> <strong>с</strong><strong>о</strong>т<strong>о</strong><strong>в</strong>идных или<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тных ячее<strong>к</strong>. Ше<strong>с</strong>ть-<strong>с</strong>емь п<strong>о</strong>перечных<br />

темных п<strong>о</strong>л<strong>о</strong><strong>с</strong>, б<strong>о</strong>лее <strong>с</strong>ближенных у <strong>в</strong>ершины.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 45—80 мм (ри<strong>с</strong>. 398<br />

А, Б; т<strong>а</strong>бл. XI, фиг. 2). Четыре <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong><br />

Ср. Азии; <strong>в</strong>. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Pseudohagla Sharov, gen. nov. Тип р<strong>о</strong>д<strong>а</strong><br />

— Hagla pospelovi О. Martynova, 1949; юр<strong>а</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong><strong>о</strong>нгл<strong>о</strong>мер<strong>а</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>о</strong>рч<strong>а</strong><strong>к</strong><strong>о</strong>л). Пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные <strong>в</strong>ет<strong>в</strong>и SC, упир<strong>а</strong>ющие<strong>с</strong>я<br />

<strong>в</strong> С, <strong>с</strong>ильн<strong>о</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>нены <strong>к</strong> SC и из<strong>о</strong>гнуты<br />

<strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>. Вт<strong>о</strong>р<strong>а</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь СиА <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т СиА ди<strong>с</strong>т<strong>а</strong>льнее<br />

ме<strong>с</strong>т<strong>а</strong> <strong>с</strong><strong>о</strong>единения MP <strong>с</strong> СиА. Между <strong>в</strong>ет<strong>в</strong>ями<br />

<strong>в</strong><strong>с</strong>ех жил<strong>о</strong><strong>к</strong>, з<strong>а</strong> и<strong>с</strong><strong>к</strong>лючением <strong>в</strong>ет<strong>в</strong>ей SC,<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. Ри<strong>с</strong>ун<strong>о</strong><strong>к</strong><br />

из мел<strong>к</strong>их темных пятен, бе<strong>с</strong>п<strong>о</strong>ряд<strong>о</strong>чн<strong>о</strong> р<strong>а</strong><strong>с</strong>пределенных<br />

п<strong>о</strong> <strong>к</strong>рылу. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм (ри<strong>с</strong>. 399; т<strong>а</strong>бл. XI, фиг. 3).<br />

Один <strong>в</strong>ид. Юр<strong>а</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Pycnophlebia Deichmuller,<br />

1886; Mesogryllus Handlirsch, 1908; Termitidium<br />

Westwood, 1854; Prophalangopsis Walker,<br />

1871; Cyphoderris Uhler, 1864; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних<br />

р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

ПОДСЕМЕЙСТВО CYRTOPHYLLITINAE ZEUNER, 1935<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong> и ее <strong>в</strong>ет<strong>в</strong>и <strong>в</strong> <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> не <strong>о</strong>тличимы<br />

<strong>о</strong>т <strong>в</strong>ет<strong>в</strong>ей SC, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> <strong>к</strong><strong>а</strong>жет<strong>с</strong>я,<br />

чт<strong>о</strong> С и<strong>с</strong>чезл<strong>а</strong>. Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>зличим<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

— <strong>в</strong> <strong>в</strong>иде уз<strong>к</strong><strong>о</strong>й длинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и, ин<strong>о</strong>гд<strong>а</strong><br />

д<strong>о</strong>х<strong>о</strong>дящей д<strong>о</strong> <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong> или д<strong>а</strong>же не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> регулярными интер<strong>к</strong><strong>а</strong>лярными<br />

жил<strong>к</strong><strong>а</strong>ми между <strong>в</strong>ет<strong>в</strong>ями SC. От МА] <strong>к</strong> RS<br />

ин<strong>о</strong>гд<strong>а</strong> пр<strong>о</strong>х<strong>о</strong>дит длинн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>,<br />

пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ющ<strong>а</strong>я другие п<strong>о</strong>перечные жил<strong>к</strong>и. В<br />

<strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>т ме<strong>с</strong>т<strong>а</strong> <strong>с</strong>лияния MP <strong>с</strong> СиА<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>к</strong>реп<strong>к</strong><strong>а</strong>я интер<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>.<br />

Юр<strong>а</strong>. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

152<br />

http://jurassic.ru/<br />

Archaboilus Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. kisyl-kiensis Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Суб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле без рез<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>ужения <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти. R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я близ<strong>к</strong><strong>о</strong> <strong>о</strong>т н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS и <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>о</strong> пяти <strong>в</strong>ет<strong>в</strong>ей; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R п<strong>о</strong>д<br />

<strong>о</strong><strong>с</strong>трым угл<strong>о</strong>м и <strong>о</strong>бр<strong>а</strong>зует три-четыре <strong>в</strong>ет<strong>в</strong>и<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 45—80 мм (ри<strong>с</strong>. 400;<br />

т<strong>а</strong>бл. XI, фиг. 4). Три <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: Cyrtophyllites Oppenheim, 1888;<br />

? Procyrtophyllites Zeuner, 1935; Liassophyllum<br />

Zeuner, 1935.<br />

HAGLIDAE INCERTAE SEDIS<br />

Pseudohumbertiella Handlirsch, 1906.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Humbertiella grandis Brauer, Redtenbacher,<br />

Ganglbauer, 1889; н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-Б<strong>а</strong>лей).<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> (фр<strong>а</strong>гмент). Вет<strong>в</strong>и R п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельны<br />

переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; RS <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>о</strong> ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей; передняя <strong>в</strong>ет<strong>в</strong>ь МА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м;<br />

между <strong>в</strong>ет<strong>в</strong>ями RS и М <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти д<strong>в</strong><strong>о</strong>йные ряды ячее<strong>к</strong>. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50 мм (ри<strong>с</strong>. 401). Один <strong>в</strong>ид. Н.<br />

юр<strong>а</strong> Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Notopamphagopsis Cabrera,<br />

1928; Zalmonites Handlirsch, 1908; Euspilopteron<br />

Cockerell, 1915; Jurassobatea Zeuner, 1937;<br />

Palaeorehnia Cockerell, 1908.<br />

СЕМЕЙСТВО GRYLLACRIDIDAE STAL, 1874<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. С длинн<strong>а</strong>я, <strong>о</strong>бычн<strong>о</strong> д<strong>о</strong>х<strong>о</strong>дящ<strong>а</strong>я<br />

д<strong>о</strong> <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong> или д<strong>а</strong>же ди<strong>с</strong>т<strong>а</strong>льнее.<br />

Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей SC не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong><br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, упир<strong>а</strong>я<strong>с</strong>ь <strong>в</strong> С, и <strong>о</strong>бычн<strong>о</strong><br />

имеет <strong>о</strong>бли<strong>к</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> между С и<br />

SC. В <strong>к</strong>рыльях <strong>с</strong> н<strong>а</strong>иб<strong>о</strong>лее <strong>а</strong>рх<strong>а</strong>ичным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

(ри<strong>с</strong>. 402) R и RS т<strong>а</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> же тип<strong>а</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е Haglidae, н<strong>о</strong> у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне R и RS<br />

<strong>с</strong> немн<strong>о</strong>гими <strong>в</strong>ет<strong>в</strong>ями или не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. MP +<br />

+ CuA <strong>о</strong>бр<strong>а</strong>зует не б<strong>о</strong>лее трех <strong>в</strong>ет<strong>в</strong>ей. МА,<br />

<strong>в</strong>ет<strong>в</strong>и MP + CuA, CuP и <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и<br />

идут д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>перед, пр<strong>о</strong>х<strong>о</strong>дя п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>чти д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>. Пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют<br />

пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и. З<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> нет. Тимп<strong>а</strong>н<strong>а</strong>льные <strong>о</strong>рг<strong>а</strong>ны н<strong>а</strong><br />

передних г<strong>о</strong>ленях <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, реже<br />

<strong>о</strong>ни рудимент<strong>а</strong>рны или имеют<strong>с</strong>я. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бе<strong>с</strong><strong>к</strong>рылые<br />

ф<strong>о</strong>рмы. С<strong>о</strong><strong>в</strong>ременные <strong>в</strong>иды преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х и <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Palaeorehniinae,<br />

Stenopelmatinae, Rhaphidophorinae, Lezininae,<br />

Gryllacridinae, Henicinae, Deinacridinae, из<br />

них т<strong>о</strong>ль<strong>к</strong><strong>о</strong> три пер<strong>в</strong>ых из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, причем д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых—<strong>в</strong>не СССР.

ПОДСЕМЕЙСТВО PALAEOREHNIINAE ZEUNER, 1937 СЕМЕЙСТВО ISFAROPTERIDAE MARTYNOV, 1937<br />

С <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, не д<strong>о</strong>х<strong>о</strong>дящ<strong>а</strong>я д<strong>о</strong><br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>. R <strong>о</strong>бр<strong>а</strong>зует ше<strong>с</strong>ть и б<strong>о</strong>лее<br />

<strong>в</strong>ет<strong>в</strong>ей. Д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>к</strong>з<strong>а</strong>ди<br />

<strong>о</strong>т <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; между <strong>в</strong>ет<strong>в</strong>ями RS д<strong>в</strong><strong>а</strong><br />

ряд<strong>а</strong> ячее<strong>к</strong>; МА д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; MP + CuAi <strong>о</strong>бр<strong>а</strong>зует<br />

три длинные <strong>в</strong>ет<strong>в</strong>и; CuP <strong>с</strong> рез<strong>к</strong>им изгиб<strong>о</strong>м.<br />

Третичные — ныне. Один р<strong>о</strong>д.<br />

Zeuneroptera Sharov, gen. nov. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Palaeorhenia scotica Zeuner, 1939; третичные<br />

<strong>о</strong>тл<strong>о</strong>жения, Ш<strong>о</strong>тл<strong>а</strong>ндия. Переднее <strong>к</strong>рыл<strong>о</strong>. С<strong>а</strong>м<strong>а</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь R <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я,<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л RS из<strong>о</strong>гнут S-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> и не<strong>с</strong>ет д<strong>о</strong><br />

ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей; MAi и МА 2 <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнуты.<br />

Между MP + CuAi и пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

<strong>в</strong>ет<strong>в</strong>ью CuA шир<strong>о</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

прямыми или S-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнутыми п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; между пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

<strong>в</strong>ет<strong>в</strong>ью CuA и CuP п<strong>о</strong>ле п<strong>о</strong>чти т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины<br />

и <strong>с</strong> <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичными п<strong>о</strong>перечными. Вет<strong>в</strong>и<br />

А 2 те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближенные, п<strong>о</strong>чти прямые. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 32 мм (ри<strong>с</strong>. 402). Один<br />

<strong>в</strong>ид. Третичные <strong>о</strong>тл<strong>о</strong>жения Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

ПОДСЕМЕЙСТВО RHAPHIDIPHORIDAE KIRBY, 1906<br />

Темя между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием <strong>а</strong>нтенн <strong>с</strong> жел<strong>о</strong>б<strong>к</strong><strong>о</strong>м<br />

или <strong>с</strong>ж<strong>а</strong>т<strong>о</strong>е. Тимп<strong>а</strong>н<strong>а</strong>льный <strong>о</strong>рг<strong>а</strong>н <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong>ж<strong>а</strong>ты <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Бе<strong>с</strong><strong>к</strong>рылые<br />

ф<strong>о</strong>рмы. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>о</strong>рме и <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО TCHORKUPHLEBIIDAE MARTYNOV,<br />

1937<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong> не <strong>о</strong>тличим<strong>а</strong> <strong>о</strong>т <strong>в</strong>ет<strong>в</strong>ей SC. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

пр<strong>о</strong><strong>с</strong>тыми <strong>в</strong>ет<strong>в</strong>ями SC. R н<strong>а</strong>чин<strong>а</strong>ет <strong>в</strong>ет<strong>в</strong>ить<strong>с</strong>я<br />

<strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>к</strong> н<strong>а</strong>ч<strong>а</strong>лу RS и <strong>о</strong>бр<strong>а</strong>зует четыре<br />

— ше<strong>с</strong>ть длинных гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

<strong>в</strong>ет<strong>в</strong>ей; RS д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. МА д<strong>а</strong>ет три<br />

п<strong>а</strong>р<strong>а</strong>ллельные <strong>в</strong>ет<strong>в</strong>и. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> MP<br />

(д<strong>о</strong> <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> CuA) или <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и, или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, <strong>с</strong>т<strong>а</strong><strong>в</strong>, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, не <strong>о</strong>тличим<strong>о</strong>й <strong>о</strong>т<br />

<strong>с</strong><strong>о</strong><strong>с</strong>едних. MP + CuAi <strong>о</strong>бр<strong>а</strong>зует четыре—<strong>с</strong>емь<br />

п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных гребенч<strong>а</strong>т<strong>о</strong><br />

<strong>в</strong>перед <strong>о</strong>т <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> и идущих не <strong>в</strong>ет<strong>в</strong>я<strong>с</strong>ь<br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Tchorkuphlebia Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong><br />

— Т. compressa Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (я. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Вет<strong>в</strong>и RS<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong>; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

MP <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; <strong>с</strong>емь <strong>в</strong>ет<strong>в</strong>ей MP + CuA. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 35—40 мм (ри<strong>с</strong>. 403).<br />

Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

153<br />

Крылья у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные; передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый. К<strong>о</strong><strong>с</strong>т<strong>а</strong> и ее<br />

<strong>в</strong>ет<strong>в</strong>и не <strong>о</strong>тличимы <strong>о</strong>т <strong>в</strong>ет<strong>в</strong>ей SC. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; между <strong>в</strong>ет<strong>в</strong>ями SC регулярные<br />

интер<strong>к</strong><strong>а</strong>лярные жил<strong>к</strong>и. R и RS не <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я.<br />

MAi <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, рудимент<strong>а</strong>рн<strong>а</strong>я, не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong><br />

<strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>я<strong>с</strong>ь <strong>с</strong>реди п<strong>о</strong>переч-<br />

406<br />

407<br />

Ри<strong>с</strong>. 404—408. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Isfaropteridae, Gryllidae,<br />

Gryllacrididae<br />

404. Isfaroptera gryllijormts Martynov; переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>,<br />

x 2,9, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 405. Protogrytlus <strong>а</strong><strong>с</strong>иtipennis<br />

Handlirsch; переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х 6,8, н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1925). 406. Protogryllus minor (Bode); переднее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х 6,8, н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Zeuner, 1939). 407. Gryllotalpa<br />

sp., передняя н<strong>о</strong>г<strong>а</strong>, х 4,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Ш<strong>а</strong>рп, 1910).<br />

408. Gryllotalpa prima Cockerell; переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х 6,0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Zeuner, 1939)<br />

http://jurassic.ru/<br />

МА<br />

404

ных жил<strong>о</strong><strong>к</strong> д<strong>о</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS. МА 2 S-<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я. От<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> MP + CuA <strong>о</strong>тх<strong>о</strong>дят <strong>в</strong>перед д<strong>в</strong>е длинные<br />

<strong>в</strong>ет<strong>в</strong>и, <strong>а</strong> н<strong>а</strong>з<strong>а</strong>д — три-четыре <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

из<strong>о</strong>гнутые н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>.<br />

В <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> интер<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> п<strong>о</strong>ле между СиА 2 и CuP <strong>в</strong> <strong>в</strong>иде<br />

ди<strong>а</strong>г<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и, идущей <strong>в</strong> уг<strong>о</strong>л изгиб<strong>а</strong><br />

CuP. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние СиА 2 д<strong>о</strong> интер<strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й<br />

и ди<strong>а</strong>г<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>й жил<strong>о</strong><strong>к</strong> и<strong>с</strong>чезл<strong>о</strong>, пре<strong>в</strong>р<strong>а</strong>ти<strong>в</strong>ши<strong>с</strong>ь,<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>в</strong> п<strong>о</strong>перечную жил<strong>к</strong>у<br />

между СиА и интер<strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й. «Зер<strong>к</strong><strong>а</strong>льце»<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Isfaroptera Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

/. grytliformis Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>о</strong> <strong>к</strong> <strong>в</strong>ершине; R <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>;<br />

MAi <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т МА 2; МА 2 п<strong>о</strong>дх<strong>о</strong>дит<br />

близ<strong>к</strong><strong>о</strong> <strong>к</strong> RS нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> п<strong>о</strong><strong>с</strong>ледней.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong> 26 мм. (ри<strong>с</strong>.<br />

404). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

СЕМЕЙСТВО GRYLLIDAE LATREILLE, 1802.<br />

СВЕРЧКИ<br />

[nom. transl. Stal, 1875<br />

(ex Grylliae Latreille, 1802)]<br />

Передние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные,<br />

з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющие брюш<strong>к</strong><strong>о</strong>, н<strong>о</strong> не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющие<br />

<strong>к</strong><strong>о</strong>нц<strong>о</strong><strong>в</strong> цер<strong>о</strong><strong>к</strong>; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong> и ее <strong>в</strong>ет<strong>в</strong>и не <strong>о</strong>тличимы <strong>о</strong>т <strong>в</strong>ет<strong>в</strong>ей SC.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; <strong>в</strong>ет<strong>в</strong>и SC<br />

длинные, п<strong>о</strong>л<strong>о</strong>гие. R и М <strong>с</strong>ж<strong>а</strong>ты; R не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я;<br />

RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; М <strong>о</strong>бр<strong>а</strong>зует д<strong>в</strong>е-три<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>в</strong>ет<strong>в</strong>и. Кубит<strong>а</strong>льн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>—-з<strong>а</strong>ним<strong>а</strong>ет б<strong>о</strong>лее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>пине. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>-<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м тел<strong>а</strong>, п<strong>о</strong>д прямым угл<strong>о</strong>м<br />

<strong>к</strong> <strong>к</strong>убит<strong>о</strong>-<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти; пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>о</strong>гнут<strong>а</strong> п<strong>о</strong> R и М. В з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>м <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>те<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> ди<strong>а</strong>г<strong>о</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>,<br />

<strong>в</strong>переди <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й имеет<strong>с</strong>я переп<strong>о</strong>нч<strong>а</strong>тый<br />

рез<strong>о</strong>н<strong>а</strong>т<strong>о</strong>р — «зер<strong>к</strong><strong>а</strong>льце»; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние СиА 2 <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует,<br />

и СиА 2 <strong>о</strong>тх<strong>о</strong>дит неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>т<br />

ди<strong>а</strong>г<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и. Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые;<br />

з<strong>а</strong>дние г<strong>о</strong>лени <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими длинными шип<strong>а</strong>ми.<br />

Цер<strong>к</strong>и длинные. Яйце<strong>к</strong>л<strong>а</strong>д <strong>с</strong>тилет<strong>о</strong><strong>в</strong>идный, <strong>с</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными <strong>в</strong>нутренними <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong><strong>а</strong>ми.<br />

Три<strong>а</strong><strong>с</strong>—ныне. 12 п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Protogryllinae,<br />

Gryllinae, Pteroplistinae, Trigonidiinae, Eneopterinae,<br />

Oecanthinae, Mogoplistinae, Myrmecophilinae,<br />

Scleropterinae, Cachoplistinae, Pentacentrinae,<br />

Phalangopsinae, из них ' пер<strong>в</strong>ые<br />

ше<strong>с</strong>ть из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии,<br />

причем Protogryllinae (ри<strong>с</strong>. 405, 406), Oecanthinae<br />

и Trigonidiinae <strong>в</strong>не СССР.<br />

ПОДСЕМЕЙСТВО GRYLLINAE KIRBY, 1906<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я, <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong>ж<strong>а</strong>ты,<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> м<strong>а</strong>лень<strong>к</strong>ий. З<strong>а</strong>дние г<strong>о</strong>ле­<br />

154<br />

http://jurassic.ru/<br />

ни без зубц<strong>о</strong><strong>в</strong>, н<strong>о</strong> <strong>в</strong><strong>о</strong><strong>о</strong>ружены неп<strong>о</strong>д<strong>в</strong>ижными<br />

или п<strong>о</strong>д<strong>в</strong>ижными шип<strong>а</strong>ми п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю.<br />

П<strong>а</strong>ле<strong>о</strong>ген —ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

пять <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле д<strong>в</strong><strong>а</strong> <strong>в</strong><br />

б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре.<br />

ПОДСЕМЕЙСТВО PTEROPLISTINAE CHOPARD, 1936<br />

З<strong>а</strong>дние г<strong>о</strong>лени п<strong>о</strong><strong>к</strong>рыты <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; шипы<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. Яйце<strong>к</strong>л<strong>а</strong>д пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тый.<br />

П<strong>а</strong>ле<strong>о</strong>ген —• ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>, <strong>о</strong>дин из них <strong>в</strong>ымерший,<br />

из<strong>в</strong>е<strong>с</strong>тный из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО ENEOPTERINAE KIRBY, 1906<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим лб<strong>о</strong>м. З<strong>а</strong>дние г<strong>о</strong>лени<br />

з<strong>а</strong>зубрены и <strong>в</strong><strong>о</strong><strong>о</strong>ружены шип<strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь): д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (<strong>о</strong>дин из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря).<br />

СЕМЕЙСТВО GRYLLOTALP1DAE BRUNNER, 1882.<br />

МЕДВЕДКИ<br />

Передние <strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены и з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<br />

лишь <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние брюш<strong>к</strong><strong>а</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong> и ее<br />

<strong>в</strong>ет<strong>в</strong>и не <strong>о</strong>тличимы <strong>о</strong>т <strong>в</strong>ет<strong>в</strong>ей SC. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>. R не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я; RS<br />

при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й не<strong>в</strong>ет<strong>в</strong>ящей<strong>с</strong>я<br />

жил<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующей <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>с</strong> передней<br />

<strong>в</strong>ет<strong>в</strong>ью М. Стр<strong>о</strong>ение з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong><br />

п<strong>о</strong>чти т<strong>а</strong><strong>к</strong><strong>о</strong>е же, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>в</strong>ерч<strong>к</strong><strong>о</strong><strong>в</strong>, н<strong>о</strong> «зер<strong>к</strong><strong>а</strong>льце»<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я; гл<strong>а</strong>з<strong>а</strong><br />

неб<strong>о</strong>льшие; <strong>а</strong>нтенны не длиннее тел<strong>а</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

длинн<strong>а</strong>я, <strong>о</strong>х<strong>в</strong><strong>а</strong>ты<strong>в</strong><strong>а</strong>ющ<strong>а</strong>я переднегрудь<br />

<strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Передние<br />

н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong><strong>о</strong>п<strong>а</strong>тельные (ри<strong>с</strong>. 407).<br />

Цер<strong>к</strong>и длинные; яйце<strong>к</strong>л<strong>а</strong>д редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>; из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> (ри<strong>с</strong>. 408),<br />

друг<strong>о</strong>й из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО TETTIGONIIDEA.<br />

КУЗНЕЧИКОВЫЕ<br />

Передние <strong>к</strong>рылья р<strong>а</strong>зличных <strong>о</strong>черт<strong>а</strong>ний —<br />

<strong>о</strong>т уз<strong>к</strong>их, л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идных д<strong>о</strong> шир<strong>о</strong><strong>к</strong>их, ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идных;<br />

<strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong><strong>в</strong>шим<strong>с</strong>я<br />

<strong>а</strong>рхеди<strong>к</strong>тием между пр<strong>о</strong>д<strong>о</strong>льными<br />

и п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

длинн<strong>о</strong>е, <strong>с</strong> гребенч<strong>а</strong>т<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дящими <strong>о</strong>т С <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>о</strong>бр<strong>а</strong>зующими ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> <strong>в</strong>т<strong>о</strong>ричным<br />

<strong>а</strong>рхеди<strong>к</strong>тием ред<strong>к</strong>ую <strong>с</strong>ет<strong>к</strong>у. R <strong>с</strong> неб<strong>о</strong>льшим<br />

чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. МА и<br />

MP, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, и их ме<strong>с</strong>т<strong>о</strong> з<strong>а</strong>ним<strong>а</strong>ет<br />

гребенч<strong>а</strong>т<strong>а</strong>я MP + CuA. У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> н<strong>а</strong><br />

передних <strong>к</strong>рыльях имеет<strong>с</strong>я з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т,<br />

причем ег<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ение н<strong>а</strong> пр<strong>а</strong><strong>в</strong><strong>о</strong>м и ле<strong>в</strong><strong>о</strong>м<br />

<strong>к</strong>рыльях р<strong>а</strong>зличн<strong>о</strong>: н<strong>а</strong> пр<strong>а</strong><strong>в</strong><strong>о</strong>м <strong>к</strong>рыле, з<strong>а</strong>ним<strong>а</strong>-

ющем нижнее п<strong>о</strong>л<strong>о</strong>жение, имеет<strong>с</strong>я рез<strong>о</strong>нирующ<strong>а</strong>я<br />

мембр<strong>а</strong>н<strong>а</strong> — «зер<strong>к</strong><strong>а</strong>льце», <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ующ<strong>а</strong>я<br />

н<strong>а</strong> <strong>в</strong>ерхнем ле<strong>в</strong><strong>о</strong>м. З<strong>а</strong>дние <strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые,<br />

лишь ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти имеет<strong>с</strong>я<br />

неб<strong>о</strong>льш<strong>о</strong>й уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>г<strong>о</strong> <strong>а</strong>рхеди<strong>к</strong>тия.<br />

Передняя <strong>в</strong>ет<strong>в</strong>ь МА — MAi <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<br />

<strong>с</strong> RS; з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь — ?МА 2 <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я.<br />

Г<strong>о</strong>лени передних н<strong>о</strong>г <strong>с</strong> тимп<strong>а</strong>н<strong>а</strong>льными <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми;<br />

л<strong>а</strong>п<strong>к</strong>и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Яйце<strong>к</strong>л<strong>а</strong>д<br />

<strong>с</strong>ж<strong>а</strong>т <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>; гл<strong>а</strong>д<strong>к</strong>ий, <strong>с</strong><strong>а</strong>бле<strong>в</strong>идный; <strong>в</strong><strong>с</strong>е три<br />

п<strong>а</strong>ры <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Tettigoniidae.<br />

СЕМЕЙСТВО TETTIGONIIDAE KRAUSS,<br />

1902. КУЗНЕЧИКИ<br />

R <strong>о</strong>бр<strong>а</strong>зует н<strong>а</strong> <strong>к</strong><strong>о</strong>нце три-четыре <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>в</strong>ет<strong>в</strong>и или не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я. CuA <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

CuP н<strong>а</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти или<br />

ди<strong>с</strong>т<strong>а</strong>льнее. СиА 2 пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong>д угл<strong>о</strong>м <strong>к</strong> <strong>в</strong>ет<strong>в</strong>ям<br />

MP + CuA, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> пере<strong>с</strong>е<strong>к</strong><strong>а</strong>я их. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. 26 п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Pseudotettigoniinae,<br />

Rammeinae, Conocephalinae, Decticinae,<br />

Phaneropterinae, Tympanophorinae, Ephippigerinae,<br />

Pycnogastrinae, Bradyporinae, Deracanthinae,<br />

Hetrodinae, Acridoxeninae, Pterophyllinae,<br />

Pseudophyllinae, Meconeminae, Mecopodinae,<br />

Phyllophorinae, Tettigoniinae, Saginae, Salomoninae,<br />

Agraepterinae, Prochilinae, Simoderinae,<br />

Aspidonotinae, Moristinae, Phasmodinae; из них<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пер<strong>в</strong>ые ше<strong>с</strong>ть из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, причем пять пер<strong>в</strong>ых <strong>в</strong>не СССР.<br />

ПОДСЕМЕЙСТВО PSEUDOTETTIGONIINAE<br />

SHAROV, SUBFAM. NOV.<br />

Ред<strong>к</strong><strong>а</strong>я <strong>с</strong>еть <strong>а</strong>рхеди<strong>к</strong>тия т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> и у <strong>в</strong>ершины; пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

К<strong>о</strong><strong>с</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> между СиА 2 и CuP (<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

Ри<strong>с</strong>. 409. Pseudotettigonia <strong>а</strong>т<strong>о</strong>еп<strong>а</strong> (Henriksen)<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 1,2, п<strong>а</strong>ле<strong>о</strong>ген, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Henriksen, 1929! интерпрет<strong>а</strong>ция<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния изменен<strong>а</strong>)<br />

CuA) <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, пре<strong>в</strong>р<strong>а</strong>ти<strong>в</strong>ши<strong>с</strong>ь, п<strong>о</strong><strong>в</strong>идим<strong>о</strong>му,<br />

<strong>в</strong> п<strong>о</strong>перечную. В <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

четыре <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и, не <strong>о</strong>бр<strong>а</strong>зующие <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>о</strong><strong>в</strong><br />

друг <strong>с</strong> друг<strong>о</strong>м или <strong>с</strong> CuP, между ними<br />

155<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

П<strong>а</strong>ле<strong>о</strong>ген — не<strong>о</strong>ген. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong>не СССР: Pseudotettigonia<br />

Zeuner, 1937 (ри<strong>с</strong>. 409); Arctolocusta<br />

Zeuner, 1937; Lithymnetes Scudder, 1890.<br />

ПОДСЕМЕЙСТВО TYMPANOPHORINAE<br />

BRUNNER, 1893<br />

[nom. transl. Kirby, 1906<br />

(ex Tympanophoridae Brunner, 1893)1<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и шир<strong>о</strong><strong>к</strong><strong>а</strong>я,<br />

упл<strong>о</strong>щенн<strong>а</strong>я <strong>с</strong><strong>в</strong>ерху между гл<strong>а</strong>з. Гл<strong>а</strong>з<strong>а</strong> ш<strong>а</strong>р<strong>о</strong><strong>в</strong>идные,<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющие; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>едл<strong>о</strong><strong>в</strong>идн<strong>а</strong>я<br />

или п<strong>о</strong>луцилиндриче<strong>с</strong><strong>к</strong><strong>а</strong>я. Передние<br />

г<strong>о</strong>лени <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>с</strong>ильными<br />

шип<strong>а</strong>ми <strong>с</strong>низу. Крылья рудимент<strong>а</strong>рные. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии д<strong>в</strong><strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ымерший.<br />

ПОДОТРЯД CAELIFERA<br />

Антенны, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, не длиннее тел<strong>а</strong>,<br />

чи<strong>с</strong>л<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> меньше 30; члени<strong>к</strong>и чет<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

<strong>о</strong>тделены друг <strong>о</strong>т друг<strong>а</strong> рез<strong>к</strong>ими перех<strong>в</strong><strong>а</strong>т<strong>а</strong>ми;<br />

их длин<strong>а</strong> <strong>в</strong> 2 и б<strong>о</strong>лее р<strong>а</strong>з пре<strong>в</strong>ыш<strong>а</strong>ет<br />

т<strong>о</strong>лщину. З<strong>а</strong>дние бедр<strong>а</strong> <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льными <strong>к</strong>илями.<br />

Чи<strong>с</strong>л<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>к</strong>и <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> н<strong>а</strong><br />

передних, <strong>с</strong>редних и з<strong>а</strong>дних н<strong>о</strong>г<strong>а</strong>х или 3—3—3,<br />

или 2—2—3, или 2—2—1. Яйце<strong>к</strong>л<strong>а</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий,<br />

<strong>к</strong><strong>о</strong>п<strong>а</strong>тельный, <strong>с</strong> зубц<strong>а</strong>ми н<strong>а</strong> <strong>в</strong>ершине<br />

<strong>в</strong>ерхних и нижних <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>о</strong><strong>к</strong>; <strong>в</strong>нутренние <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong>и<br />

рудимент<strong>а</strong>рные; реже яйце<strong>к</strong>л<strong>а</strong>д <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

При <strong>к</strong><strong>о</strong>п<strong>а</strong>нии <strong>в</strong>ерхние и нижние <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong>и<br />

<strong>о</strong>т<strong>в</strong><strong>о</strong>дят<strong>с</strong>я <strong>в</strong> пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong>л<strong>о</strong>жные <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны. Три<strong>а</strong><strong>с</strong>—<br />

ныне. Три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Locustopseidea,<br />

Acrididea, Tridactylidea; п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong> д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерн<strong>о</strong><strong>с</strong>тью не из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

НАДСЕМЕЙСТВО LOCUSTOPSEIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong> н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>м пр<strong>о</strong>тяжении пр<strong>о</strong>х<strong>о</strong>дит<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>о</strong>бр<strong>а</strong>зуя гребень <strong>в</strong>ет<strong>в</strong>ей; ин<strong>о</strong>гд<strong>а</strong> имеет<strong>с</strong>я б<strong>о</strong>лее<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> SC рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>о</strong>. RS гребенч<strong>а</strong>т<strong>а</strong>я;<br />

М <strong>о</strong>бр<strong>а</strong>зует три <strong>в</strong>ет<strong>в</strong>и, з<strong>а</strong>дняя из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими жил<strong>к</strong><strong>а</strong>ми <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трети;<br />

д<strong>в</strong>е или три <strong>в</strong>ет<strong>в</strong>и CuA. Три<strong>а</strong><strong>с</strong> — юр<strong>а</strong>. Одн<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Locustopseidae.<br />

СЕМЕЙСТВО LOCUSTOPSEIDAE<br />

HANDLIRSCH, 1908<br />

Вет<strong>в</strong>и R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, имеющие <strong>о</strong>бли<strong>к</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные упир<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> SC. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми <strong>в</strong> б<strong>о</strong>льшей ег<strong>о</strong> ч<strong>а</strong><strong>с</strong>ти.<br />

http://jurassic.ru/

Ри<strong>с</strong>. 410—413. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Locustopseidae,-Tetrigidae, Acrididae<br />

410. Locustopsis magnified Handlirsch; переднее <strong>к</strong>рыл<strong>о</strong>, x 4,7,<br />

н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925). 411. Parapleurites gracilis<br />

Brauer, Redtenbacher, Ganglbauer; переднее <strong>к</strong>рыл<strong>о</strong>, x 4,2,<br />

н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Brauer, Redtenbacher, Ganglbauer,<br />

Между <strong>в</strong>ет<strong>в</strong>ями RS, М и СиА д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> пр<strong>а</strong><strong>в</strong>ильных<br />

ячее<strong>к</strong> прям<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>й или <strong>с</strong><strong>о</strong>т<strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

ф<strong>о</strong>рмы. Ан<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong> чи<strong>с</strong>ле д<strong>в</strong>ух, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями, <strong>в</strong><strong>о</strong>зни<strong>к</strong>шими из п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Три<strong>а</strong><strong>с</strong> — юр<strong>а</strong>. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Locustopsis Handlirsch, 1908 (Brodi<strong>а</strong>п<strong>а</strong><br />

Zeuner, 1942). Тип р<strong>о</strong>д<strong>а</strong> — L. elegans<br />

Handlirsch, 1908; н. юр<strong>а</strong>, Герм<strong>а</strong>ния (<strong>в</strong>. лей<strong>а</strong><strong>с</strong>,<br />

Ме<strong>к</strong>ленбург). SC п<strong>о</strong>чти д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>; RS имеет д<strong>о</strong> пяти <strong>в</strong>ет<strong>в</strong>ей; между М, перед<br />

н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong>м ее <strong>в</strong>ет<strong>в</strong>ления, и Си — пр<strong>о</strong><strong>с</strong>тые<br />

ИЛИ Y-<strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong>перечные ЖИЛКИ. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т 10 д<strong>о</strong> 30 мм (ри<strong>с</strong>. 410). 19<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и Ср. Азии.<br />

Parapleurites Brauer, Redtenbacher,<br />

Ganglbauer, 1889. Тип р<strong>о</strong>д<strong>а</strong> — P. gracilis<br />

Brauer, Redtenbacher, Ganglbauer; н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-Б<strong>а</strong>лей).<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ления RS. Д<strong>в</strong><strong>о</strong>йные<br />

ряды прям<strong>о</strong>уг<strong>о</strong>льных ячее<strong>к</strong> между <strong>в</strong>ет<strong>в</strong>ями<br />

PS, М и СиА лишь у <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>. Между<br />

М, перед н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong>м ее <strong>в</strong>ет<strong>в</strong>ления, и Си — ДВОЙ­<br />

НОЙ ряд ячее<strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

16 мм (ри<strong>с</strong>. 411). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Вне СССР: Triassolocusta Tillyard, 1922;<br />

Conocephalella Strand, 1928.<br />

156<br />

http://jurassic.ru/<br />

413Б<br />

1889). 412. Tetrix nutans Hagenbach; <strong>о</strong>бщий <strong>в</strong>ид, x 4,9, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Ю. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Бей-Биен<strong>к</strong><strong>о</strong> и Мищен<strong>к</strong><strong>о</strong>, 1951). 413. Schistocerca sp,<br />

А — переднее <strong>к</strong>рыл<strong>о</strong>, х 1,1; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,1; <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Афри<strong>к</strong><strong>а</strong>, ЮЗ Азия, Ю. Амери<strong>к</strong><strong>а</strong> (Handlirsch, 1906)<br />

НАДСЕМЕЙСТВО ACRIDIDEA.<br />

САРАНЧОВЫЕ<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> длинн<strong>а</strong>я,<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я или д<strong>а</strong>же пр<strong>о</strong>х<strong>о</strong>дящ<strong>а</strong>я зн<strong>а</strong>чительную<br />

ч<strong>а</strong><strong>с</strong>ть ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>,<br />

SC и R без <strong>в</strong>ет<strong>в</strong>ей. RS гребенч<strong>а</strong>т<strong>а</strong>я, пр<strong>о</strong>х<strong>о</strong>дит<br />

<strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>к</strong> R, причем ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

RS <strong>с</strong>лит<strong>о</strong> <strong>с</strong> R. Д<strong>в</strong>е <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные <strong>в</strong>ет<strong>в</strong>и М. CuP<br />

прям<strong>а</strong>я. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong>чень длинн<strong>а</strong>я, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я<br />

или ин<strong>о</strong>гд<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ющ<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину<br />

длины <strong>к</strong>рыл<strong>а</strong>. Д<strong>в</strong>е <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и. Ин<strong>о</strong>гд<strong>а</strong><br />

передние <strong>к</strong>рылья рудимент<strong>а</strong>рны. В з<strong>а</strong>днем<br />

<strong>к</strong>рыле МА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; СиА не<br />

<strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я и не <strong>о</strong>бр<strong>а</strong>зует <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong> <strong>с</strong> М или ее<br />

<strong>в</strong>ет<strong>в</strong>ями. Ai пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

(не б<strong>о</strong>лее д<strong>в</strong>ух) <strong>в</strong>ет<strong>в</strong>ей. Ан<strong>о</strong>юг<strong>а</strong>льный <strong>в</strong>еер<br />

шир<strong>о</strong><strong>к</strong>ий. Ин<strong>о</strong>гд<strong>а</strong> бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы. Стре<strong>к</strong><strong>о</strong>т<strong>а</strong>ние<br />

при п<strong>о</strong>м<strong>о</strong>щи трения з<strong>а</strong>дних бедер <strong>о</strong><br />

передние <strong>к</strong>рылья или <strong>с</strong>егменты брюш<strong>к</strong><strong>а</strong>. Тимп<strong>а</strong>н<strong>а</strong>льные<br />

<strong>о</strong>рг<strong>а</strong>ны п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м пер<strong>в</strong><strong>о</strong>г<strong>о</strong> брюшн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong>; ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Л<strong>о</strong>б <strong>в</strong>ерти<strong>к</strong><strong>а</strong>льный;<br />

гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> три. Цер<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Яйц<strong>а</strong> <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>к</strong>уч<strong>к</strong><strong>а</strong>ми <strong>в</strong> <strong>в</strong>иде <strong>к</strong>убыше<strong>к</strong> <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>у.<br />

Третичные — ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Tetrigidae,<br />

Eumastacidae, Acrididae, Pneumoridae, Proscopiidae;<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии т<strong>о</strong>ль<strong>к</strong><strong>о</strong> три<br />

пер<strong>в</strong>ых.

СЕМЕЙСТВО TETRIGIDAE RAMBUR, 1938.<br />

ПРЫГУНЧИКИ<br />

[nom. transl. Walker, 1870<br />

(ex Tetrigides Rambur, 1938)]<br />

Передние <strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены, <strong>в</strong> <strong>в</strong>иде<br />

неб<strong>о</strong>льших чешуе<strong>к</strong>, <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием. З<strong>а</strong>дние '<strong>к</strong>рылья длинные; пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

б<strong>о</strong>льшую ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> з<strong>а</strong>ним<strong>а</strong>ет<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льный <strong>в</strong>еер. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> длинным<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м н<strong>а</strong>з<strong>а</strong>д, при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющим <strong>к</strong>рылья и<br />

брюш<strong>к</strong><strong>о</strong> (ри<strong>с</strong>. 412). Не<strong>о</strong>ген — ныне. С<strong>в</strong>ыше<br />

100 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии д<strong>в</strong><strong>а</strong><br />

из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО EUMASTACIDAE<br />

BURR, 1889<br />

[nom. transl. Burr, 1903<br />

(ex Eumastacides Burr, 1889)]<br />

Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е,<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющее з<strong>а</strong> передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>;<br />

RS <strong>с</strong> немн<strong>о</strong>гими <strong>в</strong>ет<strong>в</strong>ями; <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> (MP)<br />

между М и CuA <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. CuA не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я.<br />

Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы. Антенны <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

<strong>о</strong>бычн<strong>о</strong> «<strong>о</strong>р<strong>о</strong>че передних бедер. Н<strong>а</strong><br />

Д<strong>в</strong>е п<strong>а</strong>ры п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong> <strong>с</strong><br />

прямыми пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми и б<strong>о</strong>льшим<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных ряд<strong>а</strong>ми п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Переднее <strong>к</strong>рыл<strong>о</strong>. Выпу<strong>к</strong>л<strong>а</strong>я пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть и <strong>к</strong>лин<strong>о</strong><strong>в</strong>идн<strong>а</strong>я <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я;<br />

<strong>с</strong>ильные R и CuP пр<strong>о</strong>х<strong>о</strong>дят п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему<br />

и з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ям <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong>ершины их<br />

<strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти, <strong>о</strong><strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>я<br />

<strong>к</strong>рыл<strong>о</strong> (мез<strong>о</strong>з<strong>о</strong>й); MP и CuAi <strong>о</strong>бычн<strong>о</strong><br />

<strong>с</strong>ближены (<strong>к</strong>р<strong>о</strong>ме <strong>к</strong>рупных ф<strong>о</strong>рм), пр<strong>о</strong><strong>с</strong>тые;<br />

СиА 2 <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями; н<strong>а</strong> <strong>в</strong><strong>с</strong>ех жил<strong>к</strong><strong>а</strong>х <strong>с</strong>леды при<strong>к</strong>репления<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>; <strong>в</strong> з<strong>а</strong>днем <strong>к</strong>рыле пре<strong>к</strong><strong>о</strong><strong>с</strong> -<br />

т<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти нет, <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>в</strong> <strong>в</strong>иде<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong> <strong>в</strong>еер<strong>а</strong>; CuP ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> R. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong>низ, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызущег<strong>о</strong> тип<strong>а</strong>; <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> неб<strong>о</strong>льшие<br />

ните<strong>в</strong>идные <strong>а</strong>нтенны. Грудные <strong>с</strong>егменты<br />

р<strong>а</strong>з<strong>в</strong>иты р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>; н<strong>о</strong>ги бег<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>, п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядны.<br />

Жили <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жных у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

и умеренн<strong>о</strong>г<strong>о</strong> <strong>к</strong>лим<strong>а</strong>т<strong>а</strong>. Н. пермь — три<strong>а</strong><strong>с</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Archoglossopteridae, Permoberothidae,<br />

Jurinidae, Glosselytridae, Uskatelytridae, Polycytellidae.<br />

Вне СССР: Permoberothidae.<br />

ОТРЯД GLOSSELYTRODEA<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

пер<strong>в</strong><strong>о</strong>м <strong>с</strong>егменте брюш<strong>к</strong><strong>а</strong> тимп<strong>а</strong>н<strong>а</strong>льный <strong>о</strong>рг<strong>а</strong>н<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Не<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> де<strong>с</strong>ят<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тях, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин<br />

из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong> из<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО ACRIDIDAE<br />

LATREILLE, 1825<br />

[nom. transl. Brunner, 1900<br />

(ex Acriidites Latreille, 1825)]<br />

Крылья х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. В переднем «рыле<br />

RS <strong>с</strong> б<strong>о</strong>льшим' чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей. Между М и<br />

Си <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> —<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние MP; CuA <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зуя д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и. В з<strong>а</strong>днем <strong>к</strong>рыле<br />

М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS (ри<strong>с</strong>. 413). У<strong>с</strong>и<strong>к</strong>и<br />

длиннее передних бедер. Н<strong>а</strong> пер<strong>в</strong><strong>о</strong>м брюшн<strong>о</strong>м<br />

<strong>с</strong>егменте имеет<strong>с</strong>я тимп<strong>а</strong>н<strong>а</strong>льный <strong>о</strong>рг<strong>а</strong>н. С<strong>в</strong>ыше<br />

100 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>в</strong><strong>о</strong><strong>с</strong>емь из третичных и чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и; н<strong>а</strong> террит<strong>о</strong>рии<br />

СССР — <strong>о</strong>дин <strong>в</strong>ид не<strong>о</strong>пределенн<strong>о</strong>й р<strong>о</strong>д<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ти — gen? kudiana (Cockerell,<br />

1927) из нижнеми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых <strong>о</strong>тл<strong>о</strong>жений Прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я (Ам<strong>а</strong>гу). Третичные — ныне.<br />

157<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ARCHOGLOSSOPTERIDAE<br />

О. MARTY NOVA, 1958<br />

Переднее «рыл<strong>о</strong>. Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong> <strong>о</strong>дн<strong>о</strong>й, не <strong>о</strong><strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>ющей ее жил<strong>к</strong><strong>о</strong>й; MP<br />

и CuAi <strong>с</strong>ближены; R у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>, не <strong>о</strong>бр<strong>а</strong>зует<br />

изгиб<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю, ни<br />

<strong>о</strong>дн<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> н<strong>а</strong> R не <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я;<br />

CuP <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong><br />

<strong>с</strong>ередине ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>,<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны, не <strong>о</strong>бр<strong>а</strong>зуют ряд<strong>а</strong> пр<strong>а</strong><strong>в</strong>ильных<br />

ячей. Н. пермь. Один р<strong>о</strong>д.<br />

Archoglossopterum О. Martynov <strong>а</strong>, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — A. shoricum О. Martynova, 1958;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Переднее <strong>к</strong>рыл<strong>о</strong>-. Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть длиннее <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти и <strong>в</strong> 1,3 р<strong>а</strong>з<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че длины н<strong>а</strong>д<strong>к</strong>рылья; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>;<br />

RS и МА <strong>о</strong>дн<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые; <strong>о</strong>дн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь н<strong>а</strong> СиА 2.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 7,5 мм (ри<strong>с</strong>. 414). Один <strong>в</strong>ид.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.

С SC<br />

A<br />

l «18<br />

Ри<strong>с</strong>. 414—418. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archoglossopteridae, Jurinidae<br />

414. Archoglossopterum shortcum О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 8,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958).«415.<br />

Eoglosselutrum kondomense О. Martynova; А — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 13,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 13,0; н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(О. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1952). 416. Surijoka grandlcella О. Martynova;<br />

передняя п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, х 12,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 417. Protojurlna cellalosa О.<br />

Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, х 8,0. <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн;<br />

А — <strong>к</strong>рыл<strong>о</strong>; В — <strong>о</strong>т<strong>о</strong>р<strong>в</strong><strong>а</strong>нн<strong>а</strong>я и пере<strong>в</strong>ернут<strong>а</strong>я пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 418. Jurina marginata Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х6,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1958).<br />

СЕМЕЙСТВО JURINIDAE М. ZALESSKY, 1928<br />

(Eoglosselytridae О. Martynova, 1952)<br />

В пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong>д<strong>о</strong>льных<br />

жил<strong>о</strong><strong>к</strong>, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле и <strong>о</strong><strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>ющ<strong>а</strong>я<br />

ее; MP и CuAi <strong>с</strong>ближены, делят <strong>к</strong>рыл<strong>о</strong> н<strong>а</strong><br />

д<strong>в</strong>е нер<strong>а</strong><strong>в</strong>ные ч<strong>а</strong><strong>с</strong>ти, передняя п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> шире<br />

з<strong>а</strong>дней; R из<strong>о</strong>гнут<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю, н<strong>о</strong> д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong> не д<strong>о</strong>х<strong>о</strong>дит; RS<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> R, CuP <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют б<strong>о</strong>лее или менее<br />

пр<strong>а</strong><strong>в</strong>ильные ряды ячей. Пермь. Четыре<br />

р<strong>о</strong>д<strong>а</strong>.<br />

Eogtosselytrum О. Martynova, 1952.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Е. kondomense О. Martynova, 1952;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Обр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми ячеи р<strong>а</strong>зличны п<strong>о</strong> ф<strong>о</strong>рме; ширин<strong>а</strong> их<br />

не пре<strong>в</strong>ыш<strong>а</strong>ет п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>в</strong>ы<strong>с</strong><strong>о</strong>ты. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий<br />

5—6,5 мм (ри<strong>с</strong>. 415 А, Б). Ше<strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

158<br />

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Surijoka О. Martynova, 1958. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. grandlcella О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Обр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми ячеи <strong>к</strong>рупные, р<strong>а</strong>зличны п<strong>о</strong> ф<strong>о</strong>рме,<br />

ширин<strong>а</strong> их пре<strong>в</strong>ыш<strong>а</strong>ет <strong>в</strong>ы<strong>с</strong><strong>о</strong>ту незн<strong>а</strong>чительн<strong>о</strong><br />

или р<strong>а</strong><strong>в</strong>н<strong>а</strong> ей. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 7 мм<br />

(ри<strong>с</strong>. 416). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Protojurlna О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. cellulosa О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Обр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми ячеи уз<strong>к</strong>ие и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>ие, <strong>в</strong>ы<strong>с</strong><strong>о</strong>т<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

ширину <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий<br />

7—8 мм (ри<strong>с</strong>. 417). Четыре <strong>в</strong>ид<strong>а</strong>. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Jurina М. Zalessky, 1929 (Anorthoneura<br />

Martynov, 1938, <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<strong>о</strong> п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>рылу).<br />

Тип р<strong>о</strong>д<strong>а</strong> — /. scutulata М. Zalessky, 1929; <strong>в</strong>.<br />

пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие<br />

Г<strong>о</strong>ры). П<strong>о</strong>перечные жил<strong>к</strong>и <strong>в</strong><strong>с</strong>е прямые,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>, ширин<strong>а</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

ими ячей <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong><br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> 0,2—0,3 мм при т<strong>а</strong><strong>к</strong><strong>о</strong>й же или немн<strong>о</strong>г<strong>о</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ющей ширину <strong>в</strong>ы<strong>с</strong><strong>о</strong>те. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий<br />

7,2—11 мм (ри<strong>с</strong>. 418). Четыре <strong>в</strong>ид<strong>а</strong>.<br />

В. пермь Приур<strong>а</strong>лья и Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Permoberothella Riek, 1953.<br />

СЕМЕЙСТВО GLOSSELYTRIDAE MARTYNOV, 1938<br />

Переднее «рыл<strong>о</strong>. О<strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>ющей жил<strong>к</strong>и<br />

п<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

нет; пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льные жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>;<br />

MP и CuAi не <strong>с</strong>ближены; <strong>в</strong>ершин<strong>а</strong><br />

R не <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь; CuP <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины «рыл<strong>а</strong>;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют ряды мел<strong>к</strong>их<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тных ячей. В. пермь. Один р<strong>о</strong>д.

CuP 419<br />

CuP<br />

421<br />

Ри<strong>с</strong>. 419—421. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Glosselytridae, Uskatelytridae,<br />

Polycytellidae<br />

419. Qlosselyiron multivenosum Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, x 2,7,<br />

<strong>в</strong>. Пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1938). 420. Uskatelytrum<br />

siblrlcum О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, х 4.0, <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>о</strong>риг. ри<strong>с</strong>.). 421. Mesojurina sogutensls<br />

О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, х 11,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1943)<br />

Glosselytron Martynov, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

G. multivenosum Martynov, 1938; <strong>в</strong> пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

В переднем <strong>к</strong>рыле б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей<br />

CuAi. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 16—19 мм (ри<strong>с</strong>.<br />

419). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Средней <strong>в</strong>еличины и <strong>к</strong>рупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые;<br />

бе<strong>с</strong><strong>к</strong>рылые и <strong>к</strong>рыл<strong>а</strong>тые. У <strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм<br />

<strong>о</strong>дн<strong>а</strong> из п<strong>а</strong>р <strong>к</strong>рылье<strong>в</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> д<strong>о</strong> <strong>в</strong>еличины<br />

чешуе<strong>к</strong>; у мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм <strong>о</strong>бе п<strong>а</strong>ры<br />

<strong>к</strong>рылье<strong>в</strong> р<strong>а</strong>з<strong>в</strong>иты. Переднее <strong>к</strong>рыл<strong>о</strong>. Неб<strong>о</strong>льш<strong>а</strong>я<br />

пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть; н<strong>а</strong> уз<strong>к</strong><strong>о</strong>м и длинн<strong>о</strong>м<br />

<strong>к</strong>рыле прямые, п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельные, м<strong>а</strong>л<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые<br />

жил<strong>к</strong>и; <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>к</strong><strong>о</strong><strong>с</strong>ые <strong>в</strong>ет<strong>в</strong>и<br />

SC и R; <strong>о</strong><strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>ющих <strong>к</strong>рыл<strong>о</strong> и <strong>с</strong>ближенных<br />

жил<strong>о</strong><strong>к</strong> нет. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>. Х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я; р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

СЕМЕЙСТВО USKATELYTRIDAE О. MARTYNOVA,<br />

1952<br />

О<strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>ющ<strong>а</strong>я жил<strong>к</strong><strong>а</strong> п<strong>о</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти е<strong>с</strong>ть; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть зн<strong>а</strong>чительн<strong>о</strong> длиннее пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й;<br />

MP и CuAi не <strong>с</strong>ближены; <strong>в</strong>ершин<strong>а</strong> R не <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь,<br />

передняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

R; <strong>в</strong>ершин<strong>а</strong> CuP не <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные, <strong>о</strong>бр<strong>а</strong>зуют<br />

ряды мел<strong>к</strong>их ячей. В. пермь. Один р<strong>о</strong>д.<br />

Uskatelytrum О. Martynova, 1952. Тип<br />

р<strong>о</strong>д<strong>а</strong> — U. sibiricum О. Martynova, 1952;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. Ан<strong>а</strong>льные<br />

жил<strong>к</strong>и длинные, прямые; <strong>в</strong>ет<strong>в</strong>ей СиАг три;<br />

<strong>в</strong>ет<strong>в</strong>ей RS три. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий 16 мм (ри<strong>с</strong>.<br />

420). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

СЕМЕЙСТВО POLYCYTELLIDAE О. MARTYNOVA,<br />

1952<br />

О<strong>к</strong><strong>о</strong>нтури<strong>в</strong><strong>а</strong>ющ<strong>а</strong>я передний <strong>к</strong>р<strong>а</strong>й пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти жил<strong>к</strong><strong>а</strong> е<strong>с</strong>ть; MP и CuAi <strong>с</strong>лили<strong>с</strong>ь<br />

<strong>в</strong> <strong>о</strong>дну <strong>с</strong>ильную жил<strong>к</strong>у, делящую <strong>к</strong>рыл<strong>о</strong><br />

н<strong>а</strong> д<strong>в</strong>е р<strong>а</strong><strong>в</strong>ные п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины; R и CuP <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong>. .Три<strong>а</strong><strong>с</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Mesojurina О. Martynova, 1943. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. sogjutensis О. Martynova, 1943;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я и <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине; <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми ячеи шир<strong>о</strong><strong>к</strong>ие и низ<strong>к</strong>ие, п<strong>о</strong>д<strong>о</strong>бны т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым<br />

у р<strong>о</strong>д<strong>а</strong> Jurina. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий 4,7 мм<br />

(ри<strong>с</strong>. 421). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Вне СССР: Polycytella Tillyard, 1922.<br />

ОТРЯД PHASMATODEA. ПАЛОЧНИКИ<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

159<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызущег<strong>о</strong> тип<strong>а</strong>; <strong>а</strong>нтенны ните<strong>в</strong>идные;<br />

чи<strong>с</strong>л<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 8—100; ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные<br />

гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие; гл<strong>а</strong>з<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Переднегрудь <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ых<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>; н<strong>о</strong>ги длинные, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленные<br />

<strong>к</strong> л<strong>а</strong>з<strong>а</strong>нью; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>о</strong>бычн<strong>о</strong><br />

пять. Н<strong>а</strong> <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong> нечлени<strong>с</strong>тые цер<strong>к</strong>и;<br />

яйце<strong>к</strong>л<strong>а</strong>д <strong>с</strong><strong>а</strong>м<strong>к</strong>и рудимент<strong>а</strong>рен. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 2000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х. Три<strong>а</strong><strong>с</strong> —<br />

ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Chresmododea, Euphasmatodea.<br />

http://jurassic.ru/

ПОДОТРЯД CHRESMODODEA<br />

Передние <strong>к</strong>рылья х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. Мез<strong>о</strong>з<strong>о</strong>й.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Aerophasmatidae, Necrophasmatidae,<br />

Aeroplanidae, Chresmodidae;<br />

Д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО AEROPHASMATIDAE MARTYNOV, 1928<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>к</strong> <strong>в</strong>ершине не <strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; н<strong>а</strong> R три <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие передние<br />

<strong>в</strong>ет<strong>в</strong>и; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; <strong>о</strong>бе <strong>в</strong>ет<strong>в</strong>и М <strong>с</strong> длинными р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми;<br />

Си <strong>о</strong>дн<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, длинн<strong>а</strong>я; Ai <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> мн<strong>о</strong>г<strong>о</strong>. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Aerophasma Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. prynadai Martynov, 1928; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле передних <strong>к</strong>рылье<strong>в</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>;<br />

R и Aj <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>в</strong>ершины их прямые; А 2 и Аз <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

предыдущих жил<strong>к</strong><strong>а</strong>х, <strong>о</strong>бр<strong>а</strong>зуя петли. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 26 мм (ри<strong>с</strong>. 422). Один <strong>в</strong>ид. В.<br />

юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО NECROPHASMAT1DAE MARTYNOV,<br />

1925<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong> <strong>в</strong>ершине <strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я, з<strong>а</strong><strong>о</strong><strong>с</strong>трен<strong>о</strong>;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>,<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я п<strong>о</strong>чти <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong> и пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и; СиА <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями; CuP<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; Ai <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> немн<strong>о</strong>г<strong>о</strong>.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Necrophasma Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong>—<br />

N. shabarovi Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> уз<strong>к</strong><strong>о</strong>е и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е;<br />

SC д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; R д<strong>о</strong>х<strong>о</strong>дит<br />

д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ние ее<br />

из<strong>о</strong>гнут<strong>о</strong> <strong>к</strong>з<strong>а</strong>ди; н<strong>а</strong> СиА три гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>в</strong>ет<strong>в</strong>и; н<strong>а</strong> А 2 три <strong>в</strong>ет<strong>в</strong>и. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 21,5 мм (ри<strong>с</strong>. 423). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Ри<strong>с</strong>. 422—424. Отряд Phasmatodea<br />

422. Aerophasma prynadai Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 2,3, <strong>в</strong>.<br />

юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). 423. Necrophasma shabarovi<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х 3,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 424. Calvisia atrosignata Stal; <strong>о</strong>бщий <strong>в</strong>ид,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Brunner, 1893)<br />

ПОДОТРЯД EUPHASMATOD ЕА<br />

Передние <strong>к</strong>рылья п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> рудимент<strong>а</strong>рны.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Phyllidae и Phasmatidae; п<strong>о</strong><strong>с</strong>леднее и<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

«<br />

СЕМЕЙСТВО PHASMATIDAE LEACH, 1815<br />

[nom. transl. Serville, 1839<br />

(ex Phasmida Leach, 1815)]<br />

Бе<strong>с</strong><strong>к</strong>рылые или <strong>к</strong>рыл<strong>а</strong>тые ф<strong>о</strong>рмы <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й<br />

д<strong>о</strong> р<strong>а</strong>змер<strong>о</strong><strong>в</strong> чешуи передней п<strong>а</strong>р<strong>о</strong>й<br />

<strong>к</strong>рылье<strong>в</strong>. Грудные <strong>с</strong>егменты длинные и уз<strong>к</strong>ие<br />

(ри<strong>с</strong>. 424). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три р<strong>о</strong>д<strong>а</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и<br />

С. Амери<strong>к</strong>и.<br />

КОГОРТА PARA NEOPTERA. ПАРАНЕОПТЕРЫ<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рылье<strong>в</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>бедненн<strong>о</strong>е;<br />

юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> умеренных<br />

р<strong>а</strong>змер<strong>о</strong><strong>в</strong> и <strong>о</strong>бычн<strong>о</strong> не<strong>с</strong>ет <strong>о</strong>дну <strong>в</strong>ет<strong>в</strong>ящую<strong>с</strong>я<br />

юг<strong>а</strong>льную жил<strong>к</strong>у. З<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> р<strong>а</strong><strong>с</strong>­<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

160<br />

http://jurassic.ru/<br />

ширенные и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> б<strong>о</strong>лее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, чем передние:<br />

гетер<strong>о</strong>н<strong>о</strong>мия <strong>к</strong>рылье<strong>в</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>чти<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я диптеригия. Ин<strong>о</strong>гд<strong>а</strong><br />

передние <strong>к</strong>рылья упл<strong>о</strong>тняют<strong>с</strong>я или при

уменьшении р<strong>а</strong>змер<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых при<strong>о</strong>брет<strong>а</strong>ют<br />

длинные <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и и р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пер<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ть<br />

(птил<strong>о</strong>птеригия). Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong><br />

бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><br />

<strong>с</strong>ильным <strong>с</strong><strong>о</strong><strong>с</strong><strong>а</strong>тельным н<strong>а</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong>м и имеет р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>лющег<strong>о</strong> члени<strong>с</strong>т<strong>о</strong>г<strong>о</strong> х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong><br />

или <strong>в</strong>ид<strong>о</strong>измененных грызущих челю<strong>с</strong>тей.<br />

Пре<strong>в</strong>р<strong>а</strong>щение неп<strong>о</strong>лн<strong>о</strong>е: ин<strong>о</strong>гд<strong>а</strong> у <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

ф<strong>о</strong>рм (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых х<strong>о</strong>б<strong>о</strong>тных и<br />

трип<strong>с</strong><strong>о</strong><strong>в</strong>) <strong>в</strong> инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>м р<strong>а</strong>з<strong>в</strong>итии <strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ет<br />

п<strong>о</strong><strong>к</strong><strong>о</strong>ящ<strong>а</strong>я<strong>с</strong>я <strong>с</strong>т<strong>а</strong>дия. Неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелые<br />

<strong>с</strong>т<strong>а</strong>дии р<strong>а</strong>з<strong>в</strong>ития, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong>х<strong>о</strong>дны <strong>с</strong><strong>о</strong><br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лыми н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мыми и не я<strong>в</strong>ляют<strong>с</strong>я <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нными<br />

личин<strong>к</strong><strong>а</strong>ми. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Три н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>: Rhynchota, Psocopteroidea,<br />

Thysanopteroidea.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения п<strong>а</strong>р<strong>а</strong>не<strong>о</strong>птер<br />

<strong>с</strong> другими <strong>к</strong><strong>о</strong>г<strong>о</strong>рт<strong>а</strong>ми <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых нея<strong>с</strong>ны.<br />

Н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ления и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> п<strong>а</strong>р<strong>а</strong>не<strong>о</strong>птер <strong>о</strong>пределили<strong>с</strong>ь р<strong>а</strong>зличными<br />

ф<strong>о</strong>рм<strong>а</strong>ми <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния п<strong>а</strong>р<strong>а</strong>зитизм<strong>а</strong><br />

и тем <strong>с</strong><strong>а</strong>мым — <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong><strong>в</strong> прием<strong>а</strong> пищи<br />

и <strong>с</strong>тр<strong>о</strong>ения р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. Пер<strong>в</strong>ый н<strong>а</strong>д<strong>о</strong>тряд—<br />

Rhynchota, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующий<strong>с</strong>я <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>лющег<strong>о</strong> члени<strong>с</strong>т<strong>о</strong>г<strong>о</strong> х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong> у р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядных<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, те<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н <strong>с</strong><strong>о</strong><br />

<strong>в</strong>т<strong>о</strong>рым, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, и<strong>с</strong>х<strong>о</strong>дным н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong>м,<br />

именн<strong>о</strong> Psocopteroidea, <strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong>й мере<br />

гетер<strong>о</strong>генным и <strong>в</strong> т<strong>о</strong> же <strong>в</strong>ремя х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризу­<br />

ющим<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> примити<strong>в</strong>ными (грызущие р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

эт<strong>о</strong>г<strong>о</strong> н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>), т<strong>а</strong><strong>к</strong> и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong> <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нными<br />

черт<strong>а</strong>ми (<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>с</strong>тр<strong>о</strong>ения, <strong>о</strong>предели<strong>в</strong>шие<strong>с</strong>я<br />

п<strong>а</strong>р<strong>а</strong>зитизм<strong>о</strong>м у мн<strong>о</strong>гих групп).<br />

Еще б<strong>о</strong>лее <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong> э<strong>в</strong><strong>о</strong>люция п<strong>о</strong><strong>с</strong>леднег<strong>о</strong><br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>, <strong>о</strong>предели<strong>в</strong>ш<strong>а</strong>я<strong>с</strong>я р<strong>а</strong>з<strong>в</strong>итием мел<strong>к</strong>их<br />

р<strong>а</strong>змер<strong>о</strong><strong>в</strong> трип<strong>с</strong><strong>о</strong><strong>в</strong> и <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong><strong>а</strong>ми их пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

пит<strong>а</strong>ния, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong> н<strong>а</strong><br />

<strong>в</strong>ы<strong>с</strong><strong>а</strong><strong>с</strong>ы<strong>в</strong><strong>а</strong>нии пыльце<strong>в</strong>ых зерен р<strong>а</strong><strong>с</strong>тений (ри<strong>с</strong>.<br />

425). Б<strong>о</strong>льш<strong>а</strong>я дре<strong>в</strong>н<strong>о</strong><strong>с</strong>ть и <strong>с</strong><strong>в</strong><strong>о</strong>ебр<strong>а</strong>зие <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляющих<br />

эту <strong>к</strong><strong>о</strong>г<strong>о</strong>рту н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong>, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

п<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют неи<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>еннные <strong>с</strong><strong>в</strong>язи<br />

этих н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых <strong>с</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми групп<strong>а</strong>ми<br />

дре<strong>в</strong>не<strong>к</strong>рылых (именн<strong>о</strong> <strong>с</strong> Palaeodictyoptera,<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ли<br />

члени<strong>с</strong>тым х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м) и т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, <strong>а</strong><br />

<strong>с</strong> друг<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляют <strong>с</strong><strong>о</strong>мне<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong><br />

цел<strong>о</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти и фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>м един<strong>с</strong>т<strong>в</strong>е<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> эт<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

и <strong>о</strong>тряд<strong>о</strong><strong>в</strong>. Оче<strong>в</strong>идн<strong>о</strong>, чт<strong>о</strong> е<strong>с</strong>ли эти <strong>с</strong><strong>о</strong>мнения<br />

и предп<strong>о</strong>л<strong>о</strong>жения <strong>о</strong><strong>к</strong><strong>а</strong>жут<strong>с</strong>я <strong>с</strong>пр<strong>а</strong><strong>в</strong>едли<strong>в</strong>ыми,<br />

т<strong>о</strong> <strong>в</strong><strong>о</strong>зни<strong>к</strong>нет не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>ть пере<strong>с</strong>м<strong>о</strong>треть<br />

<strong>в</strong><strong>с</strong>ю <strong>с</strong>хему <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений<br />

групп н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых <strong>в</strong> цел<strong>о</strong>м; <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее<br />

<strong>в</strong>ремя, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><strong>с</strong>ти фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их<br />

д<strong>а</strong>нных, м<strong>о</strong>жн<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>т<strong>а</strong><strong>в</strong>ить, н<strong>о</strong><br />

еще не реш<strong>а</strong>ть эту пр<strong>о</strong>блему.<br />

НАДОТРЯД RHYNCHOTA. ХОБОТНЫЕ<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> д<strong>в</strong>ур<strong>а</strong>здельным н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м и <strong>с</strong><strong>о</strong><strong>с</strong>ущим<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong>ым <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong>м, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящим из щетин<strong>о</strong><strong>к</strong><br />

пер<strong>в</strong><strong>о</strong>й и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й п<strong>а</strong>ры челю<strong>с</strong>тей, з<strong>а</strong><strong>к</strong>люченных<br />

<strong>в</strong> длинный члени<strong>с</strong>тый х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нный<br />

<strong>с</strong>р<strong>о</strong><strong>с</strong>шими<strong>с</strong>я щуп<strong>а</strong>льц<strong>а</strong>ми нижней<br />

губы. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, четырех- или пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

реже длинные, мн<strong>о</strong>г<strong>о</strong>члени<strong>с</strong>тые.<br />

Обычн<strong>о</strong> д<strong>в</strong>е п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong>, реже з<strong>а</strong>дние <strong>к</strong>рылья<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны: мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые<br />

и бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы. Передние <strong>к</strong>рылья ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> пл<strong>о</strong>тные н<strong>а</strong>д<strong>к</strong>рылья: ин<strong>о</strong>гд<strong>а</strong><br />

при эт<strong>о</strong>м не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е <strong>с</strong>тр<strong>о</strong>ение. Жил<strong>к</strong><strong>а</strong> CuP<br />

прям<strong>а</strong>я и <strong>о</strong>тделяет треуг<strong>о</strong>льн<strong>о</strong>е <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле.<br />

Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные, хищни<strong>к</strong>и, п<strong>а</strong>р<strong>а</strong>зиты<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

11 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 161<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных, немн<strong>о</strong>гие <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н —<br />

ныне. Д<strong>в</strong><strong>а</strong> <strong>о</strong>тряд<strong>а</strong>: Homoptera и Heteroptera.<br />

Пер<strong>в</strong>ый <strong>о</strong>тряд и<strong>с</strong>х<strong>о</strong>дный и из<strong>в</strong>е<strong>с</strong>тен <strong>с</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>,<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>й^, п<strong>о</strong> мнению не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ученых,—<br />

фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь пер<strong>в</strong><strong>о</strong>г<strong>о</strong>, <strong>о</strong>тдели<strong>в</strong>ш<strong>а</strong>я<strong>с</strong>я<br />

<strong>в</strong> перми. И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие х<strong>о</strong>б<strong>о</strong>тных<br />

<strong>о</strong>пределял<strong>о</strong><strong>с</strong>ь р<strong>а</strong><strong>с</strong>ширением и углублением <strong>с</strong><strong>в</strong>язей<br />

<strong>с</strong> р<strong>а</strong><strong>с</strong>тениями, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, и мн<strong>о</strong>г<strong>о</strong>ядн<strong>о</strong><strong>с</strong>тью<br />

и хищниче<strong>с</strong>т<strong>в</strong><strong>о</strong>м, <strong>с</strong> друг<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны.<br />

Р<strong>а</strong>з<strong>в</strong>итие фит<strong>о</strong>ф<strong>а</strong>гии пр<strong>о</strong>х<strong>о</strong>дил<strong>о</strong> <strong>в</strong> <strong>о</strong>тряде р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых<br />

и при<strong>в</strong>ел<strong>о</strong> <strong>к</strong> и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нным<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым — фит<strong>о</strong>ф<strong>а</strong>г<strong>а</strong>м, н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящим<br />

п<strong>а</strong>р<strong>а</strong>зит<strong>а</strong>м р<strong>а</strong><strong>с</strong>тений, <strong>к</strong><strong>о</strong><strong>к</strong>цид<strong>а</strong>м и тлям.<br />

Хищниче<strong>с</strong>т<strong>в</strong><strong>о</strong>, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нн<strong>о</strong>е <strong>с</strong> р<strong>а</strong>з<strong>в</strong>итием п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><strong>с</strong>ти,<br />

<strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ил<strong>о</strong> п<strong>о</strong>я<strong>в</strong>ление <strong>к</strong>л<strong>о</strong>п<strong>о</strong><strong>в</strong>.<br />

http://jurassic.ru/

Ри<strong>с</strong>. 425. Схем<strong>а</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений <strong>о</strong>тряд<strong>о</strong><strong>в</strong> Paraneoptera; н<strong>а</strong> ф<strong>о</strong>не ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>л<strong>о</strong>н<strong>к</strong>и п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>о</strong> <strong>в</strong>ремя<br />

ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нения и фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>с</strong><strong>в</strong>язей.<br />

Об<strong>о</strong>зн<strong>а</strong>чения: Thp—Thysanoptera, Ps — Psocoptera; \Z— Zoraptera, Mai — Malophaga, An — Anoplura, Horn — Homoptera<br />

Het — Heteroptera (Р<strong>о</strong>денд<strong>о</strong>рф, Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,[М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)<br />

ОТРЯД HOMOPTERA. РАВНОКРЫЛЫЕ<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые или <strong>о</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нные,<br />

<strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я <strong>к</strong>рыше<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>,<br />

реже пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>; <strong>о</strong>бычн<strong>о</strong> имеет<strong>с</strong>я чет<strong>к</strong><strong>о</strong> <strong>о</strong>тделенный<br />

жил<strong>к</strong><strong>о</strong>й CuP з<strong>а</strong>дний уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> — <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>;<br />

ин<strong>о</strong>гд<strong>а</strong> переп<strong>о</strong>нч<strong>а</strong>т<strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>тделен<strong>а</strong> т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>о</strong>й делящей<br />

162<br />

http://jurassic.ru/<br />

линией и н<strong>а</strong>дл<strong>о</strong>м<strong>а</strong>ми пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong>. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью;<br />

ин<strong>о</strong>гд<strong>а</strong> п<strong>о</strong><strong>с</strong>ледняя редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>. Неред<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые или бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я или <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, <strong>о</strong>бычн<strong>о</strong><br />

пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong><strong>о</strong>единенн<strong>а</strong>я <strong>с</strong> грудным <strong>о</strong>тдел<strong>о</strong>м,

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я; гулярн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>иты, имеют<strong>с</strong>я гл<strong>а</strong>з<strong>к</strong>и. Ореднегрудь б<strong>о</strong>льше<br />

з<strong>а</strong>днегруди. Н<strong>о</strong>ги бег<strong>а</strong>тельные, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong>дняя<br />

п<strong>а</strong>р<strong>а</strong> прыг<strong>а</strong>тельн<strong>а</strong>я; ин<strong>о</strong>гд<strong>а</strong> н<strong>о</strong>ги <strong>с</strong>ильн<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные; л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>о</strong>дним — тремя члени<strong>к</strong><strong>а</strong>ми.<br />

Брюш<strong>к</strong><strong>о</strong> из <strong>в</strong><strong>о</strong><strong>с</strong>ьми — <strong>о</strong>динн<strong>а</strong>дц<strong>а</strong>ти<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> н<strong>а</strong> брюш<strong>к</strong>е р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>ые и <strong>с</strong>лух<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны. В инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>м<br />

р<strong>а</strong>з<strong>в</strong>итии ин<strong>о</strong>гд<strong>а</strong> пр<strong>о</strong>х<strong>о</strong>дят неп<strong>о</strong>д<strong>в</strong>ижную<br />

<strong>с</strong>т<strong>а</strong>дию. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нные <strong>с</strong> р<strong>а</strong>зличными н<strong>а</strong>земными<br />

би<strong>о</strong>цен<strong>о</strong>з<strong>а</strong>ми. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. Три<br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Auchenorrhyncha, Sternorrhyncha,<br />

Coleorrhyncha.<br />

ПОДОТРЯД AUCHENORRHYNCHA<br />

Крылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> гетер<strong>о</strong>н<strong>о</strong>мны, передние<br />

пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> пл<strong>о</strong>тные н<strong>а</strong>д<strong>к</strong>рылья или переп<strong>о</strong>нч<strong>а</strong>тые;<br />

з<strong>а</strong>дние <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью, реже <strong>о</strong><strong>к</strong>руглые, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й<br />

юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong><strong>о</strong>единен<br />

<strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н—<br />

ныне. Четыре инфр<strong>а</strong><strong>о</strong>тряд<strong>а</strong>: Blattoprosbolomorpha,<br />

Cicadomorpha, Cicadellomorpha,<br />

Fulgoromorpha.<br />

* И НФРАОТРЯД<br />

BLATTOPROSBOLOMORPHA<br />

Жил<strong>к</strong>и н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельны д<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния,<br />

не <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>бщег<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Blattoprosbolidae, Protoprosbolidae.<br />

СЕМЕЙСТВО BLATTOPROSBOLIDAE<br />

BECKER-MIGDISOVA, 1958<br />

В <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья. SC, R, М и CuA<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены; С упл<strong>о</strong>тненн<strong>а</strong>я, <strong>с</strong> <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зным<br />

ут<strong>о</strong>лщением <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; SC длинн<strong>а</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; CuA гребенч<strong>а</strong>т<strong>а</strong>я. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Один<br />

р<strong>о</strong>д.<br />

Ри<strong>с</strong>. 426—431. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Blattoprosbolidae, Protoprosbolidae,<br />

Mesogereonidae и Homoptera insertae sedii<br />

426. Blattoprosbole tomiensls Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

x 3,8, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958).<br />

427. Protoprosbole straeleni Laurentiaux; н<strong>а</strong>д<strong>к</strong>рылье х 4,3,<br />

н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Бельгия (Laurentiaux, 1952). 428. Permodunstania<br />

prosbololdes Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 7,1 ,н.пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 429. Kaltanocicada dunstanioides<br />

Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 14,0, н. пермь,»Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 430. Mesogereon<br />

superbum Tillyard; переднее <strong>к</strong>рыл<strong>о</strong>, х 1,6, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

(Tillyard, 1921). 431. Mesogereon affine Tillyard; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

х 4,6, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1921)<br />

163<br />

http://jurassic.ru/

Blattoprosbole Becker-Migdisova,<br />

1958. Тип р<strong>о</strong>д<strong>а</strong> — В. tomiensis Becker-Migdisova,<br />

1958; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Желтый Яр). SC<br />

<strong>с</strong>ильн<strong>о</strong> упл<strong>о</strong>тнен<strong>а</strong>, прям<strong>а</strong>я и длинн<strong>а</strong>я, д<strong>а</strong>ет<br />

ряд п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей. R и М <strong>с</strong> м<strong>а</strong>лым чи<strong>с</strong>л<strong>о</strong>м<br />

<strong>в</strong>ет<strong>в</strong>ей; СиА гребенч<strong>а</strong>т<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

17,8 мм (ри<strong>с</strong>. 426; т<strong>а</strong>бл. XII, фиг. 1).<br />

Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО PROTOPROSBOLIDAE<br />

LAURENTIAUX, 1952<br />

В <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья SC, R и М <strong>с</strong>ближены,<br />

н<strong>о</strong> не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я. С без <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

ут<strong>о</strong>лщения <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; SC мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

14 мм (ри<strong>с</strong>. 427). Н<strong>а</strong>мюр Бельгии.<br />

Один р<strong>о</strong>д — Protoprosbole Laurentiaux, 1952.<br />

ИНФРАОТРЯД<br />

С I CADOMOR Р НА<br />

Передние <strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>тые, <strong>с</strong><br />

пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми, R и М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й;<br />

tegulae <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, <strong>с</strong>редние т<strong>а</strong>зи<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

причленяют<strong>с</strong>я близ<strong>к</strong><strong>о</strong> друг <strong>к</strong> другу; з<strong>а</strong>дний<br />

<strong>о</strong>тдел н<strong>а</strong>лични<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый; л<strong>о</strong>б неб<strong>о</strong>льш<strong>о</strong>й,<br />

треуг<strong>о</strong>льный; три пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н —<br />

ныне. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Palaeontinidea, Cicadidea.<br />

НАДСЕМЕЙСТВО PALAEONTINIDEA<br />

В передних <strong>к</strong>рыльях SC длинн<strong>а</strong>я; М и СиА<br />

делят<strong>с</strong>я <strong>о</strong>чень р<strong>а</strong>н<strong>о</strong>, <strong>с</strong> длинными <strong>к</strong><strong>о</strong><strong>с</strong>ыми п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е,<br />

без р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Н.<br />

пермь — юр<strong>а</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Dunstaniidae, Mesogereonidae,<br />

Palaeontinidae.<br />

СЕМЕЙСТВО DUNSTANIIDAE TILLYARD, 1916<br />

Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я переп<strong>о</strong>нч<strong>а</strong>т<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>о</strong>тделяет<strong>с</strong>я <strong>о</strong>т <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

ряд<strong>о</strong>м пл<strong>о</strong>тных, длинных, <strong>к</strong><strong>о</strong><strong>с</strong>ых п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; узел<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит; SC длинн<strong>а</strong>я,<br />

д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылий; RS<br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; М д<strong>а</strong>ет <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>зы <strong>с</strong> RS и СиА.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е и <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

<strong>в</strong>ырез<strong>к</strong><strong>о</strong>й переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Н. пермь —<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Вне СССР: Dunstania Tillyard, 1916; Dunstaniopsis<br />

Tillyard, 1918; Paradunstania Tillyard,<br />

1918.<br />

СЕМЕЙСТВО MESOGEREONIDAE<br />

TILLYARD, 1921<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину,<br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е, <strong>с</strong> пл<strong>о</strong>тными, <strong>в</strong>ыпу<strong>к</strong>лыми жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й и м<strong>о</strong>рщин<strong>а</strong>ми н<strong>а</strong><br />

мембр<strong>а</strong>не; <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и делят<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong>, и ди<strong>с</strong>т<strong>а</strong>льные их <strong>в</strong>ет<strong>в</strong>ления <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н; SC,<br />

R2 и RS пр<strong>о</strong>х<strong>о</strong>дят <strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>я близ<strong>к</strong><strong>о</strong> <strong>о</strong>дн<strong>а</strong> <strong>к</strong><br />

друг<strong>о</strong>й, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong>; н<strong>а</strong>дл<strong>о</strong>м жил<strong>о</strong><strong>к</strong> и узел<strong>о</strong><strong>к</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>в</strong>д<strong>в</strong><strong>о</strong>е меньше передних,<br />

<strong>о</strong><strong>к</strong>руглые <strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й. М4 и СиА, <strong>с</strong><strong>о</strong>единяя<strong>с</strong>ь<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>зуют з<strong>а</strong>м<strong>к</strong>нутую<br />

б<strong>а</strong>з<strong>а</strong>льную ячей<strong>к</strong>у. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

44 мм (ри<strong>с</strong>. 430, 431). Три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Один<br />

р<strong>о</strong>д — Mesogereon Tillyard, 1921 (пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>).<br />

СЕМЕЙСТВО PALAEONTINIDAE HANDLIRSCH, 1906<br />

(Cicadomorphidae Evans, 1956)<br />

Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья зн<strong>а</strong>чительн<strong>о</strong><br />

б<strong>о</strong>льше б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е<br />

и уз<strong>к</strong><strong>о</strong>е; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, <strong>с</strong>р<strong>а</strong>зу п<strong>о</strong><strong>с</strong>ле <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления<br />

М делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и: R2 и RS, п<strong>а</strong>р<strong>а</strong>ллельны<br />

переднему <strong>к</strong>р<strong>а</strong>ю; р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и М и<br />

СиА длинные. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> неглуб<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>в</strong>ырез<strong>к</strong><strong>о</strong>й<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я и б<strong>о</strong>льшим г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным<br />

<strong>в</strong><strong>о</strong>з<strong>в</strong>ышением; Mi <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м<br />

пр<strong>о</strong>тяжении <strong>с</strong> RS, <strong>о</strong>бр<strong>а</strong>зуя б<strong>а</strong>з<strong>а</strong>льную<br />

ячей<strong>к</strong>у; М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я. В. три<strong>а</strong><strong>с</strong> —юр<strong>а</strong>. 22 р<strong>о</strong>д<strong>а</strong>.<br />

Cicadomorpha Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong>—<br />

С. punctulata Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылье<br />

уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е; делящ<strong>а</strong>я линия, узел<strong>о</strong><strong>к</strong><br />

и длинн<strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> гщ-<br />

Ри<strong>с</strong>. 432—444. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Palaeontinidea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Palaeontinidae.<br />

432. Cicadomorpha punctulata Martynov; н<strong>а</strong>д<strong>к</strong>рылье, x 1,3, <strong>в</strong>.<br />

юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 433. Palaeontinodes<br />

schabarovi Martynov; н<strong>а</strong>д<strong>к</strong>рылье, x 1,2, н. юр<strong>а</strong>, Ср. Азия<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 434. Phragmatoecicossus shurabensis Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 2,2, н. юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1949). 435. Palaeocossus jurassicus Oppenheim; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 2,6, юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 436. Phragmatoecites<br />

damesi Oppenhein; н<strong>а</strong>д<strong>к</strong>рылье, x 1,6, юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 437. Suljuctocossus prosboloides Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 1,3, юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1949). 438. Pseudocossus zemcuznicovi Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

xl,5,юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я<strong>о</strong>бл.(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 439. Aslocossus costa-<br />

164<br />

http://jurassic.ru/<br />

lis Becker-Migdisova; фр<strong>а</strong>гмент н<strong>а</strong>д<strong>к</strong>рылья, х,4,5, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср-<br />

Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 440. Palaeontinopsis latipennis<br />

Martynov; н<strong>а</strong>д<strong>к</strong>рылье, х 2,2, юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937).<br />

441. Ijacossus suchanovae Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье; х 3,0,<br />

юр<strong>а</strong>. Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1950). 442. Plachutella<br />

picta Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 2,3, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 443. Shurabocossus gigas<br />

Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,6, юр<strong>а</strong>, Средняя Азия<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 444. Suluktaja turkestanensis Becker-<br />

Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 1,5, юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949)

165<br />

http://jurassic.ru/

cua <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; CuA без изгиб<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

М. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong><br />

ф<strong>о</strong>рме тр<strong>а</strong>пеции, <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> шире <strong>с</strong><strong>в</strong><strong>о</strong>ей длины;<br />

<strong>с</strong>редне<strong>с</strong>пин<strong>к</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я; з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щен<strong>а</strong>.<br />

Тел<strong>о</strong> п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

54 мм (ри<strong>с</strong>. 432). Один <strong>в</strong>ид. В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Palaeontinodes Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. schabarovi Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ытянут<strong>о</strong><br />

<strong>в</strong> длину, б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ужен<strong>о</strong>; делящ<strong>а</strong>я линия<br />

и узел<strong>о</strong><strong>к</strong> я<strong>с</strong>ные; между <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>тью<br />

СиА и М не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> длинных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 30—58 мм (ри<strong>с</strong>. 433).<br />

Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Phragmatoecicossus Becker-Migdisova,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong>—Ph. shurabensis Becker-<br />

Migdisova, 1949; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>,<br />

Шур<strong>а</strong>б II). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти не<br />

р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>, <strong>с</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й и прямым<br />

передним <strong>к</strong>р<strong>а</strong>ем; узел<strong>о</strong><strong>к</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; делящ<strong>а</strong>я<br />

линия упл<strong>о</strong>тнен<strong>а</strong>; SC <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>в</strong>ет<strong>в</strong>ей; б<strong>а</strong>з<strong>а</strong>льный изгиб СиА <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong>ершин<strong>о</strong>й <strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием М, <strong>о</strong>бр<strong>а</strong>зует неб<strong>о</strong>льшую<br />

треуг<strong>о</strong>льную б<strong>а</strong>з<strong>а</strong>льную ячей<strong>к</strong>у. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 32 мм (ри<strong>с</strong>. 434). Один <strong>в</strong>ид. Н.<br />

юр<strong>а</strong> Ср. Азии.<br />

Phragmatoecites Oppenheim, 1885. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ph. damesi Oppenheim, 1885; юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-<br />

Б<strong>а</strong>лей). Н<strong>а</strong>д<strong>к</strong>рылье удлиненн<strong>о</strong>е, эллиптиче<strong>с</strong><strong>к</strong><strong>о</strong>е;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>; М б<strong>а</strong>з<strong>а</strong>льнее делящей линии, делит<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; узел<strong>о</strong><strong>к</strong> б<strong>о</strong>лее или менее я<strong>с</strong>ный.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 23 мм (ри<strong>с</strong>. 436). Один<br />

<strong>в</strong>ид. Юр<strong>а</strong> В. Сибири.<br />

Palaeocossus Oppenheim, 1885. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. jurassicus Oppenheim, 1885; н. юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-<br />

Б<strong>а</strong>лей). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>нным<br />

з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужен<strong>а</strong>; передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й;<br />

узел<strong>о</strong><strong>к</strong> я<strong>с</strong>ный; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й SC; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>о</strong>гр<strong>а</strong>ничен<strong>а</strong> пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми, пятиуг<strong>о</strong>льн<strong>а</strong>я.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 38 мм (ри<strong>с</strong>. 435).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Юр<strong>а</strong> В. Сибири и Ср. Азии.<br />

Suljuctocossus Becker-Migdisova,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong> — S. prosboloides Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (лей<strong>а</strong><strong>с</strong>, Сулю'<strong>к</strong>т<strong>а</strong>).<br />

З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong><br />

<strong>с</strong>рез<strong>а</strong>н, <strong>в</strong>ершин<strong>а</strong> <strong>в</strong>ытянут<strong>а</strong>, передний <strong>к</strong>р<strong>а</strong>й<br />

прям<strong>о</strong>й; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е; делящ<strong>а</strong>я<br />

линия упл<strong>о</strong>тнен<strong>а</strong>; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я,<br />

<strong>о</strong>гр<strong>а</strong>ничен<strong>а</strong> <strong>с</strong>л<strong>а</strong>быми жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 45 мм (ри<strong>с</strong>. 437; т<strong>а</strong>бл. XIII,<br />

фиг. 3). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

166<br />

http://jurassic.ru/<br />

Pseudocossus М <strong>а</strong> г t у п <strong>о</strong> v, 1931. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. zemcuznicovi Martynov, 1931; юр<strong>а</strong>,<br />

Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-<br />

Б<strong>а</strong>лей). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е, б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ужен<strong>о</strong>:<br />

узел<strong>о</strong><strong>к</strong> <strong>с</strong> у<strong>с</strong>туп<strong>о</strong>м <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; делящ<strong>а</strong>я<br />

линия я<strong>с</strong>н<strong>а</strong>я; SC <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

41 мм (ри<strong>с</strong>. 438; т<strong>а</strong>бл. XIII, фиг. 5).<br />

Один <strong>в</strong>ид. Юр<strong>а</strong> В. Сибири.<br />

Asiocossus Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> —A. costalis Becker-Migdisova, 1962;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>. И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем<br />

и д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем;<br />

SC <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong><strong>с</strong>редине <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т R;<br />

R + Ми.<strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, р<strong>а</strong><strong>в</strong>ны п<strong>о</strong><br />

длине; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> не з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>, н<strong>о</strong><br />

<strong>о</strong>гр<strong>а</strong>ничен<strong>а</strong> <strong>о</strong>переди ут<strong>о</strong>лщенными жил<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 48 мм (ри<strong>с</strong>. 439).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Palaeontinopsis Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. latipennis Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азии (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Н<strong>а</strong>д<strong>к</strong>рылье шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е; передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый; узел<strong>о</strong><strong>к</strong> нея<strong>с</strong>ный, делящ<strong>а</strong>я<br />

линия упл<strong>о</strong>тнен<strong>а</strong>; <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и <strong>с</strong>ближены.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 38 мм (ри<strong>с</strong>. 440). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong><br />

Н. юр<strong>а</strong> Ср. Азии.<br />

Ijacossus Becker-Migdisova, 1950.<br />

Тип р<strong>о</strong>д<strong>а</strong> — I. suchanovae Becker-Migdisova,<br />

1950; юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, р. Ия). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину;<br />

передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>в</strong>ыпу<strong>к</strong>лый; узел<strong>о</strong><strong>к</strong><br />

имеет<strong>с</strong>я; делящ<strong>а</strong>я линия упл<strong>о</strong>тнен<strong>а</strong>; SC д<strong>а</strong>ет<br />

ряд п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

длинный, RS <strong>о</strong>тх<strong>о</strong>дит <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трети <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; между <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>тью СиА и<br />

М не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>ых жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 55 мм (ри<strong>с</strong>. 441). Один <strong>в</strong>ид. Юр<strong>а</strong> В. Сибири.<br />

Plachutella Becker-Migdisova, 1949.<br />

Тип. р<strong>о</strong>д<strong>а</strong> — P. rotundata Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II).<br />

Передний <strong>к</strong>р<strong>а</strong>й з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>чти прям<strong>о</strong>й, <strong>в</strong>ырез<strong>к</strong><strong>а</strong> <strong>о</strong>чень неб<strong>о</strong>льш<strong>а</strong>я,<br />

г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е <strong>в</strong><strong>о</strong>з<strong>в</strong>ышение <strong>о</strong>чень б<strong>о</strong>льш<strong>о</strong>е;<br />

SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong> углублении<br />

<strong>в</strong>ырез<strong>к</strong>и; Mi+г и Мз+4 <strong>в</strong> <strong>в</strong>иде д<strong>в</strong>ух<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельных <strong>в</strong>ет<strong>в</strong>ей д<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 20—23 мм<br />

(ри<strong>с</strong>. 442, т<strong>а</strong>бл. XIII, фиг. 4, 6). Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Юр<strong>а</strong> Ср. Азии и В. Сибири.<br />

Shurabocossus Becker-Migdisova,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong> — Sh. gigas Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б III).<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>-<strong>в</strong>ытянут<strong>о</strong>е, <strong>в</strong>ырез<strong>к</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я<br />

и глуб<strong>о</strong><strong>к</strong><strong>а</strong>я, г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е <strong>в</strong><strong>о</strong>з<strong>в</strong>ышение<br />

б<strong>о</strong>льш<strong>о</strong>е; SC идет <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь <strong>в</strong>д<strong>о</strong>ль R и R 2 и д<strong>а</strong>-

ет ряд <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей <strong>к</strong> <strong>в</strong>ырез<strong>к</strong>е <strong>к</strong>рыл<strong>а</strong>;<br />

Мз+4 и Мг <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я р<strong>а</strong>нее <strong>с</strong>лияния п<strong>о</strong><strong>с</strong>ледней<br />

<strong>с</strong> Mi и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 37,4 мм (ри<strong>с</strong>. 443). Один <strong>в</strong>ид. Н. юр<strong>а</strong><br />

Ср. Азии.<br />

Suljuktaja Becker-Migdisova, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — S. turkestanensis Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (лей<strong>а</strong><strong>с</strong>, Сулю<strong>к</strong>т<strong>а</strong>).<br />

Вырез<strong>к</strong><strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> глуб<strong>о</strong><strong>к</strong><strong>а</strong>я и<br />

умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е <strong>в</strong><strong>о</strong>з<strong>в</strong>ышение<br />

б<strong>о</strong>льш<strong>о</strong>е; Mi и RS <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>о</strong>чень неб<strong>о</strong>льш<strong>о</strong>м<br />

пр<strong>о</strong>тяжении; Мз+4 <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> Мг<br />

немн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>нее <strong>с</strong>лияния п<strong>о</strong><strong>с</strong>ледней <strong>с</strong> Mi и <strong>в</strong>ме<strong>с</strong>те<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> R. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 29 мм<br />

(ри<strong>с</strong>. 444). Один <strong>в</strong>ид. Юр<strong>а</strong> Ср. Азии.<br />

Ри<strong>с</strong>. 445—453. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Prosbolidae<br />

^45. Prosbole kondomensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 4,2,<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 446.<br />

Prosbole brevialata Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, x 2,7,<strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1941). 447. Orthoprosbole<br />

triangularis Martynov; н<strong>а</strong>д<strong>к</strong>рылье, х 3,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 448. Orthoprosbole congesta Martynov;<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,0, <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1935). 449. Dictyoprosbole membranosa Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 2,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 450. Sojano-<br />

Вне СССР: Palaeontina Butler, 1873; Р<strong>а</strong>chypsyche<br />

Handlirsch, 1906; Cyttonium Westwood,<br />

1854; Limacodites Handlirsch, 1906; Protopsyche<br />

Handlirsch, 1906; Archipsyche Handlirsch,<br />

1906; Polystra Oppenheim, 1888; Eocicada<br />

Oppenheim, 1888; Beloptesis Handlirsch,<br />

1906; Fletcheriana Evans, 1956.<br />

НАДСЕМЕЙСТВО CICADIDEA<br />

В н<strong>а</strong>д<strong>к</strong>рыльях SC д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я или пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и М и CuA <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong> "р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым д<strong>в</strong>ур<strong>а</strong>здельным<br />

н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м. Н. пермь •— ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Prosbolidae, Tettigarctidae, Cicadidae.<br />

167<br />

http://jurassic.ru/<br />

neura stignata Martynov; н<strong>а</strong>д<strong>к</strong>рылье, x 5,4, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 451. Sojanoneura bimaculata Martynov;<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 4,5, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1935). 452. Prosboloneura kondomensis Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 9,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 453. Prosboloneura colorata Becker-Migdisova; фр<strong>а</strong>гмент<br />

н<strong>а</strong>д<strong>к</strong>рылья, х 10,0, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

447

СЕМЕЙСТВО PROSBOLIDAE HANDLIRSCH, 1904<br />

[Scytinopteridae Tillyard, 1926<br />

(pars)]<br />

Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>т<strong>а</strong>я;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC делит<strong>с</strong>я<br />

«<strong>а</strong> SCA, <strong>о</strong>бр<strong>а</strong>зующую <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

изгиб <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и упл<strong>о</strong>тненный <strong>к</strong>р<strong>а</strong>й, и<br />

SCP, идущую <strong>в</strong>д<strong>о</strong>ль R + М и Ri; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

<strong>в</strong>ырез<strong>к</strong><strong>о</strong>й и г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зяым <strong>в</strong><strong>о</strong>з<strong>в</strong>ышением.<br />

К<strong>а</strong>рб<strong>о</strong>н — три<strong>а</strong><strong>с</strong>. 17 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Prosbole Handlirsch, 1904 (Prosbolina<br />

Handlirsch, 1937). Тип р<strong>о</strong>д<strong>а</strong> — P. hirsuta Handlirsch,<br />

1904; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Тихие Г<strong>о</strong>ры). Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>о</strong>бычн<strong>о</strong> б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>рез<strong>а</strong>н; делящ<strong>а</strong>я<br />

линия и узел<strong>о</strong><strong>к</strong> имеют<strong>с</strong>я; R, М и СиА делят<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не. В з<strong>а</strong>днем <strong>к</strong>рыле М <strong>с</strong><br />

четырьмя <strong>в</strong>ет<strong>в</strong>ями и б<strong>о</strong>льше. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

12—35 мм (ри<strong>с</strong>. 445, 446; т<strong>а</strong>бл. XII,<br />

фиг. 4). 22 <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

и <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

Orthoprosbole Martynov, 1935. Тип р<strong>о</strong>д<strong>а</strong>—<br />

О. congesta Martynov, 1935; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Б<strong>а</strong>й<strong>к</strong><strong>а</strong>им<strong>с</strong><strong>к</strong><strong>а</strong>я Ли<strong>с</strong>т<strong>в</strong>яж<strong>к</strong><strong>а</strong>). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>а</strong>, <strong>с</strong> <strong>а</strong><strong>с</strong>имметричн<strong>о</strong><br />

<strong>с</strong>рез<strong>а</strong>нным з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем, делящей линией и<br />

узел<strong>к</strong><strong>о</strong>м; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; R делит<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее,<br />

чем М и СиА. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>с</strong>ильн<strong>о</strong><br />

<strong>в</strong>ытянут<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью и мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми,<br />

р<strong>а</strong>н<strong>о</strong> делящими<strong>с</strong>я М и СиА. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 23 мм (ри<strong>с</strong>. 447, 448). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Dictyoprosbole Martynov, 1935. Тип р<strong>о</strong>д<strong>а</strong>—<br />

D. membranosa Martynov, 1935; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>).<br />

Н<strong>а</strong>д<strong>к</strong>рылье переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е, <strong>с</strong> мел<strong>к</strong><strong>о</strong>й<br />

<strong>с</strong>ет<strong>к</strong><strong>о</strong>й; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть шир<strong>о</strong><strong>к</strong><strong>а</strong>я; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; R, М и СиА делят<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не; RS, М и СиА мн<strong>о</strong>н<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>ты; R 2 и RS<br />

из<strong>о</strong>гнуты. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 32 мм (ри<strong>с</strong>. 449).<br />

Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ри<strong>с</strong>. 454—466. О<br />

454. Kaltanetta nigra a Becker-Migdisova; Л—н<strong>а</strong>д<strong>к</strong>рылье, х 7,5;<br />

Б —з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 10,0; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 455. Permocicada nigrita Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 10,6, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 456. Permocicada kuznetskiensis Becker-Migdisova;<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 12,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 457. Permocicada integra Becker-Migdisova<br />

(ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция), х 4,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1940). 458. Kaltanopsis ornata Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 9,7, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 459. Orthoscytlnaskokt Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 9,2, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961).<br />

460. Orthoscytlna longijurcata Becker-Migdisova; з<strong>а</strong>днее, <strong>к</strong>рыл<strong>о</strong>,<br />

168<br />

Sojanoneura Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. edemskyi Martynov, 1928; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>. С<strong>о</strong>ян<strong>а</strong>).<br />

Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong>ругл<strong>о</strong> <strong>с</strong>рез<strong>а</strong>н<strong>а</strong>;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е; R, М и СиА делят<strong>с</strong>я н<strong>а</strong><br />

<strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>в</strong> 0,57—0,65 длины н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>о</strong>т ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния. М имеет три-четыре <strong>в</strong>ет<strong>в</strong>и.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong> п<strong>о</strong>л<strong>о</strong>гим г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным<br />

<strong>в</strong><strong>о</strong>з<strong>в</strong>ышением; М имеет д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 6—11 мм (ри<strong>с</strong>. 450, 451; т<strong>а</strong>бл.<br />

XII, фиг. 8). 12 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

Prosboloneura Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — P. colorata Becker-Migdisova,<br />

1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>,' К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>к</strong>ругл<strong>о</strong> <strong>с</strong>рез<strong>а</strong>н<strong>о</strong><br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е; R и<br />

СиА делят<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м-ур<strong>о</strong><strong>в</strong>не, <strong>в</strong> 0,57 длины<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>,<br />

делит<strong>с</strong>я <strong>в</strong> 0,66—0,77 длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6—7 мм (ри<strong>с</strong>.<br />

452, 453; т<strong>а</strong>бл. XII, фиг. 9). Три <strong>в</strong>ид<strong>а</strong>. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kaltanetta Becker-Migdisova, 196L<br />

Тип р<strong>о</strong>д<strong>а</strong> — К. nigra Becker-Migdisova, 1961;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ытянут<strong>о</strong>, <strong>о</strong>тн<strong>о</strong>шение<br />

длины <strong>к</strong> ширине 3,0 : 1; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ытянут<strong>а</strong><br />

и <strong>с</strong>имметричн<strong>о</strong> <strong>о</strong><strong>к</strong>руглен<strong>а</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

бедн<strong>о</strong>е; R идет <strong>в</strong>пл<strong>о</strong>тную <strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>я и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> нег<strong>о</strong> р<strong>а</strong>нее <strong>в</strong>ершины н<strong>а</strong>д<strong>к</strong>рылья; птер<strong>о</strong><strong>с</strong>тигмы<br />

нет; ячей<strong>к</strong><strong>а</strong> r 2-rs <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>а</strong>я; R *<br />

делит<strong>с</strong>я <strong>в</strong> 0,47 длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния,<br />

<strong>а</strong> М — <strong>в</strong> 0,66; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>в</strong>ытянут<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью, глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

и шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й и п<strong>о</strong>л<strong>о</strong>гим г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным<br />

<strong>в</strong><strong>о</strong>з<strong>в</strong>ышением; ячей<strong>к</strong><strong>а</strong> r 2-rs шир<strong>о</strong><strong>к</strong><strong>а</strong>я;<br />

М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

8,1 мм (ри<strong>с</strong>. 454 А, Б; т<strong>а</strong>бл. XII, фиг. 5). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permocicada Martynov, 1928(Permocicadopsis<br />

Becker-Migdisova, 1940). Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. nigronervosa Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> 0,44—0,50,<br />

<strong>а</strong> М — <strong>в</strong> 0,50—0,77 длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния;<br />

М трех- или четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. З<strong>а</strong>дние<br />

:ей<strong>с</strong>т<strong>в</strong><strong>о</strong> Prosbolidae<br />

х 9,0, <strong>в</strong>. першь,' Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961>._<br />

461. Archeglyphis crassinervis Martynov; н<strong>а</strong>д<strong>к</strong>рылье, х 2,0, <strong>с</strong>р.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931). 462. Kondomoprosbole<br />

picta Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 11,2, н. Пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 463, Pervestigiafumosa<br />

Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 9,2, н. Пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 464. Pervestigia veteris<br />

Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 8,2, н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 465. Mesocicada verrucosa-<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 21,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961).466. Evanscicadaspeciosa Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

xJ3,2, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961)<br />

http://jurassic.ru/

4Б9<br />

http://jurassic.ru/

<strong>к</strong>рылья <strong>с</strong> п<strong>о</strong>л<strong>о</strong>гим <strong>в</strong>ырез<strong>о</strong>м; R, М и СиА <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я<br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины <strong>в</strong>ырез<strong>к</strong>и. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 4—16 мм (ри<strong>с</strong>. 455—457; т<strong>а</strong>бл. XII,<br />

фиг. 7). 23 <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

и <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

Kaltanopsis Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — К. ornata Becker-Migdisova, 1961;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья эллиптиче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ытянут<strong>а</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е;<br />

SCP длинн<strong>а</strong>я; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я и <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я жил<strong>к</strong>и <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ие;<br />

R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> 0,4 длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

и зн<strong>а</strong>чительн<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее, чем М и<br />

•CuA. М <strong>с</strong> тремя-четырьмя <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 6,9 мм (ри<strong>с</strong>. 458). Один <strong>в</strong>ид. Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Orthoscytina Tillyard, 1926 (Anomalos-<br />

•cytina Evans, 1943, non. Davis, 1942) Тип р<strong>о</strong>д<strong>а</strong>—<br />

О. mitchelli Tillyard, 1926, <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ытянуты, <strong>о</strong>тн<strong>о</strong>шение длины<br />

<strong>к</strong> ширине 2,6—2,9; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>,<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong><br />

Ю,3—0,4 длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния;<br />

М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> 0,6—0,7 длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину;<br />

М и СиА <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я р<strong>а</strong>н<strong>о</strong>, н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>ты.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 7,5—8,5 мм (ри<strong>с</strong>.<br />

459, 460; т<strong>а</strong>бл. XII, фиг. 1). 15 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> (д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>).<br />

Archeglyphis Martynov, 1931. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. crassinervis Martynov, 1931; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, <strong>а</strong>лы<strong>к</strong><strong>а</strong>е<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я п<strong>о</strong>д<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Верх<strong>о</strong>т<strong>о</strong>м<strong>с</strong><strong>к</strong><strong>о</strong>е).<br />

Крыл<strong>о</strong> <strong>с</strong> прямым передним <strong>к</strong>р<strong>а</strong>ем и уз<strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; R и R 2<br />

<strong>в</strong> <strong>в</strong>иде прям<strong>о</strong>й г<strong>о</strong>риз<strong>о</strong>нт<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и; RS <strong>с</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; М и СиА мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 29 мм (ри<strong>с</strong>. 461). Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kondomoprosbole Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — К. pictata Becker-Migdisova,<br />

1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Передний <strong>к</strong>р<strong>а</strong>й з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый, <strong>с</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>в</strong>ырез<strong>к</strong><strong>о</strong>й и б<strong>о</strong>льшим<br />

т<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным <strong>в</strong><strong>о</strong>з<strong>в</strong>ышением; R 2— прям<strong>о</strong>е<br />

пр<strong>о</strong>д<strong>о</strong>лжение R; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R зн<strong>а</strong>чительн<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong>рыл<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>в</strong>ырез<strong>к</strong>и; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>и ее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 6 мм (ри<strong>с</strong>. 462). Один <strong>в</strong>ид. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Pervestigia Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. veteris Becker-Migdisova, 1961;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Крыл<strong>о</strong><strong>в</strong><strong>а</strong>я <strong>в</strong>ырез<strong>к</strong><strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е <strong>в</strong><strong>о</strong>з<strong>в</strong>ышение<br />

неб<strong>о</strong>льш<strong>о</strong>е, п<strong>о</strong>л<strong>о</strong>г<strong>о</strong>е; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е;<br />

170<br />

http://jurassic.ru/<br />

ячей<strong>к</strong><strong>а</strong> r 2-rs д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я: М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я.<br />

Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 6,7 мм (ри<strong>с</strong>. 463,<br />

464; т<strong>а</strong>бл. XII, фиг. 6). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong>'<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Mesocicada Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. verrucosa Becker-Migdisova,<br />

1962; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль).<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, жил<strong>к</strong>и пл<strong>о</strong>тные,<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние н<strong>а</strong>д<strong>к</strong>рылья п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми;<br />

R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее, чем<br />

М и СиА, и р<strong>а</strong>нее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья; узел<strong>о</strong><strong>к</strong><br />

и делящ<strong>а</strong>я линия <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; М четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА неб<strong>о</strong>льш<strong>о</strong>й. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 5 мм (ри<strong>с</strong>. 465). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Evanscicada Becker-Migdisova nom.<br />

nov. (Evansia Becker-Migdisova, 1961,<br />

nom. nud.). Тип р<strong>о</strong>д<strong>а</strong> — E. speciosa Becker-Migdisova,<br />

1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е,<br />

<strong>о</strong>тн<strong>о</strong>шение длины <strong>к</strong> ширине <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 2,5 : 1: б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е <strong>к</strong>рыл<strong>о</strong><br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е; RS <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья и зн<strong>а</strong>чительн<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее Ri; М делит<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не <strong>с</strong> R, СиА н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> Ri; <strong>с</strong>леды<br />

<strong>а</strong>рхеди<strong>к</strong>тия <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тые, <strong>к</strong>рупные ячей<strong>к</strong>и или межжил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я<br />

г<strong>о</strong>фрир<strong>о</strong><strong>в</strong><strong>к</strong><strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 22—<br />

28 мм (ри<strong>с</strong>. 466; т<strong>а</strong>бл. XII, фиг. 2—3). Четыре<br />

<strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Mitchelloneura Tillyard, 1929;<br />

Permodiphtera Tillyard, 1926; Austroprosbole<br />

Evans, 1943.<br />

СЕМЕЙСТВО TETTIGARCTIDAE<br />

BECKER-MIGDISOVA, 1949<br />

Н<strong>а</strong>д<strong>к</strong>рылье пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>е, <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нными<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным и треуг<strong>о</strong>льным <strong>а</strong>н<strong>а</strong>льным<br />

п<strong>о</strong>лями, <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й; SC<br />

делит<strong>с</strong>я н<strong>а</strong> SCA, <strong>о</strong>бр<strong>а</strong>зующую <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

изгиб <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и пл<strong>о</strong>тный <strong>к</strong>р<strong>а</strong>й, и SCP,<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ющую<strong>с</strong>я <strong>с</strong> R + М и R; <strong>в</strong>ершины ди<strong>с</strong>т<strong>а</strong>льных<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> жил<strong>о</strong><strong>к</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й<br />

прям<strong>о</strong>й. В з<strong>а</strong>днем <strong>к</strong>рыле <strong>в</strong>ыем<strong>к</strong><strong>а</strong> и г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е<br />

<strong>в</strong><strong>о</strong>з<strong>в</strong>ышение <strong>о</strong>чень м<strong>а</strong>лы. З<strong>а</strong>ч<strong>а</strong>т<strong>о</strong><strong>к</strong> з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong> <strong>в</strong> <strong>в</strong>иде <strong>к</strong>ругл<strong>о</strong>й г<strong>о</strong>фрир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й<br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м брюш<strong>к</strong><strong>а</strong>. Х<strong>о</strong>рд<strong>о</strong>т<strong>о</strong>н<strong>а</strong>льный<br />

<strong>о</strong>рг<strong>а</strong>н <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Три<strong>а</strong><strong>с</strong> — ныне. Де<strong>с</strong>ять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>о</strong>дин <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Cicadoprosbole Becker-Migdisova,<br />

1947. Тип р<strong>о</strong>д<strong>а</strong> — С. sogutensis Becker-Migdisova,<br />

1947; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong><strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>а</strong><strong>с</strong>имметричн<strong>о</strong> <strong>с</strong>рез<strong>а</strong>нным<br />

з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен 0,75<br />

длины R + М; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> 0,37, <strong>а</strong> М — <strong>в</strong> 0,5

Ри<strong>с</strong>. 467—470. СеЙей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tettigarctidae<br />

467. • Cicadoprosbole sogutensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, plachutai Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 19,0, юр<strong>а</strong>, Cp Азия<br />

x 4,7, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Cp Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1947). 468. Тиги- (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 470. Kisylia psylloides Martynov;<br />

tanovia karaiavica Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 8,7, <strong>в</strong>. юр<strong>а</strong>, н<strong>а</strong>д<strong>к</strong>рылье, x 3,7, юр<strong>а</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 469. Shuraboprosbole '<br />

длины н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>т ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния; R 2 гребенч<strong>а</strong>т<strong>а</strong>я;<br />

узел<strong>о</strong><strong>к</strong> я<strong>с</strong>ный; делящ<strong>а</strong>я линия упл<strong>о</strong>тнен<strong>а</strong>;<br />

<strong>в</strong>ершины ди<strong>с</strong>т<strong>а</strong>льных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> жил<strong>о</strong><strong>к</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены приблизительн<strong>о</strong> н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не;<br />

з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь CuA п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong> передней.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 17,8 мм (ри<strong>с</strong>. 467; т<strong>а</strong>бл. XIII,<br />

фиг. 2). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Turutanovia Becker-Migdisova, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. karaiavica Becker-Migdisova,<br />

1949; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм. К<strong>а</strong>р<strong>а</strong>г<strong>а</strong>у).<br />

Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>имметричн<strong>а</strong>,<br />

эллиптиче<strong>с</strong><strong>к</strong>и <strong>в</strong>ытянут<strong>а</strong>; R 2 <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями; делящ<strong>а</strong>я линия упл<strong>о</strong>тнен<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и жил<strong>о</strong><strong>к</strong> длинные, их <strong>в</strong>ершины р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

не н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, <strong>а</strong> п<strong>о</strong> п<strong>о</strong>перечн<strong>о</strong>й<br />

н<strong>а</strong><strong>к</strong>л<strong>о</strong>нн<strong>о</strong>й прям<strong>о</strong>й; з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

CuA <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передней. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 14 мм (ри<strong>с</strong>. 468). Один <strong>в</strong>ид. В.<br />

юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Shuraboprosbole Becker-Migdisova,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong> — Sh. plachutai Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б<br />

II). В н<strong>а</strong>д<strong>к</strong>рыльях <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины R + М; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее,<br />

<strong>а</strong> М — н<strong>а</strong> <strong>с</strong>ередине длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

CuA <strong>в</strong> ме<strong>с</strong>те изгиб<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию М<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>с</strong>ледней н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>межут<strong>к</strong>е;<br />

б<strong>а</strong>з<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец CuA <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>ден, из<strong>о</strong>гнут<br />

<strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну CuP. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 35—<br />

40 мм (ри<strong>с</strong>. 469). Один <strong>в</strong>ид. Юр<strong>а</strong> Ср. Азии.<br />

171<br />

Kisylia Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К- psylloides Martynov, 1937; н'. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Делящ<strong>а</strong>я линия и узел<strong>о</strong><strong>к</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен трем<br />

чет<strong>в</strong>ертям длины R + М; RS <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее,<br />

<strong>а</strong> М — н<strong>а</strong> <strong>с</strong>ередине длины н<strong>а</strong>д<strong>к</strong>рылья;<br />

Ri <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; RS не <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; R 2 и RS п<strong>а</strong>р<strong>а</strong>ллельны;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA неб<strong>о</strong>льш<strong>о</strong>й, <strong>в</strong>ет<strong>в</strong>и<br />

ег<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й длины. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 17—20 мм<br />

(ри<strong>с</strong>. 470). Один <strong>в</strong>ид. Юр<strong>а</strong> Ср. Азии.<br />

Вер<strong>о</strong>ятн<strong>о</strong>, <strong>к</strong> эт<strong>о</strong>му же <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я<br />

р<strong>о</strong>д Diphtheropsis Martynov, 1937 из н. юры<br />

Ср. Азии.<br />

Вне СССР: Hylaeoneura Lameere et Severin,<br />

1897; Eotettigarcta Zeuner, 1944; Mesodiphthera<br />

Tillyard, 1919; Meshemipteron Cockerell,<br />

1916, <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>с</strong><strong>о</strong><strong>в</strong>ременный<br />

рели<strong>к</strong>т<strong>о</strong><strong>в</strong>ый р<strong>о</strong>д Tettigarcta Wife.<br />

СЕМЕЙСТВО CICADIDAE<br />

LATREILLE, 1802<br />

[nom. transl. Westwood, 1940<br />

(ex Cicadae Latreille, 1802, 1804)]<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> — R 2 <strong>с</strong>д<strong>в</strong>инут<strong>а</strong><br />

<strong>в</strong>пл<strong>о</strong>тную <strong>к</strong> <strong>к</strong>р<strong>а</strong>ю; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

или пре<strong>в</strong>р<strong>а</strong>щен<strong>о</strong> <strong>в</strong> хитин<strong>о</strong><strong>в</strong>ую п<strong>о</strong>л<strong>о</strong><strong>с</strong>у;<br />

<strong>в</strong>ершины ди<strong>с</strong>т<strong>а</strong>льных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> жил<strong>о</strong><strong>к</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>й прям<strong>о</strong>й; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, зн<strong>а</strong>чительн<strong>о</strong> меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины<br />

н<strong>а</strong>д<strong>к</strong>рылья; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> без <strong>в</strong>ырез<strong>к</strong>и и г<strong>о</strong>рб<strong>о</strong>-<br />

http://jurassic.ru/

Ри<strong>с</strong>. 471—472. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cicadidae<br />

471. Tymoclcada gorbunovi Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 2,3, ми<strong>о</strong>цен, З.Сибирь (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1954). 472".<br />

Pomponia merula Distant; <strong>о</strong>бщий <strong>в</strong>ид, 10,75, М<strong>а</strong>л<strong>а</strong>й<strong>с</strong><strong>к</strong>ий <strong>а</strong>рхипел<strong>а</strong>г (Distant, 1912)<br />

<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>. З<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й и х<strong>о</strong>рд<strong>о</strong>т<strong>о</strong>н<strong>а</strong>льный<br />

<strong>о</strong>рг<strong>а</strong>ны имеют<strong>с</strong>я. Третичные — ныне.<br />

Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Cicadinae, Platypleurinae,<br />

Tettigadinae, Tibicininae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йдены.<br />

ПОДСЕМЕЙСТВО PLATYPLEURINAE HANDLIRSCH,<br />

1925<br />

Тимп<strong>а</strong>н<strong>а</strong>льные п<strong>о</strong><strong>к</strong>рыш<strong>к</strong>и р<strong>а</strong>з<strong>в</strong>иты, з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<br />

<strong>в</strong>е<strong>с</strong>ь з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 20—60 мм (ри<strong>с</strong>. 471). В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии три р<strong>о</strong>д<strong>а</strong>: <strong>о</strong>дин<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и, <strong>о</strong>дин из ми<strong>о</strong>цен<strong>а</strong><br />

3. Сибири и <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ПОДСЕМЕЙСТВО CICADINAE LATREILLE<br />

Тимп<strong>а</strong>н<strong>а</strong>льные п<strong>о</strong><strong>к</strong>рыш<strong>к</strong>и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты,<br />

з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют не <strong>в</strong>е<strong>с</strong>ь з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10—70 мм (ри<strong>с</strong>. 472). Не<strong>о</strong>ген<br />

— ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии три р<strong>о</strong>д<strong>а</strong>:<br />

<strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> Cicadoidea incertae sedis <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я<br />

Liassocicada Bode, 1953 из <strong>в</strong>. лей<strong>а</strong><strong>с</strong><strong>а</strong><br />

Герм<strong>а</strong>нии.<br />

CICADOMORPHA INCERTAE SEDIS<br />

Permodunstania Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — P. prosboloides Becker-Mig­<br />

172<br />

http://jurassic.ru/<br />

disova, 1961; H. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> бедным<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; SC длинн<strong>а</strong>я, не<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

RS <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; R2, RS и М <strong>с</strong> изгиб<strong>о</strong>м н<strong>а</strong> гр<strong>а</strong>нице<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти и <strong>с</strong> длинными, пл<strong>о</strong>тными,<br />

<strong>к</strong><strong>о</strong><strong>с</strong>ыми п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми — упл<strong>о</strong>тненн<strong>о</strong>й<br />

делящей линией; М 4 <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, не<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong> Мз. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 16—<br />

20 мм (ри<strong>с</strong>. 428). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kattanocicada Becker-Migdisova,.<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — К- dunstanioides Becker-Migdisova,<br />

1961; н пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью и шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

<strong>в</strong>ырез<strong>к</strong><strong>о</strong>й переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; R2 — прям<strong>о</strong>е<br />

пр<strong>о</strong>д<strong>о</strong>лжение R; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> (гг-rs)<br />

<strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>а</strong>я; RS <strong>с</strong>ильн<strong>о</strong> изгиб<strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю; <strong>к</strong><strong>о</strong>нцы R2 и RS, <strong>а</strong><br />

т<strong>а</strong><strong>к</strong>же <strong>в</strong>ет<strong>в</strong>ей М ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

СиА ДЛИННЫЙ. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 7,0 мм (ри<strong>с</strong>. 429). Один <strong>в</strong>ид. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

И НФРАОТРЯД<br />

CICADELLOMORPHA<br />

(Jassidomorpha)<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>быми<br />

жил<strong>к</strong><strong>а</strong>ми; R и М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

<strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; tegulae <strong>о</strong>т<strong>с</strong>ут-

<strong>с</strong>т<strong>в</strong>ует, <strong>с</strong>редние т<strong>а</strong>зи<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, причленяют<strong>с</strong>я<br />

близ<strong>к</strong><strong>о</strong> друг <strong>к</strong> другу; з<strong>а</strong>дний <strong>о</strong>тдел н<strong>а</strong>лични<strong>к</strong><strong>а</strong><br />

•б<strong>о</strong>льш<strong>о</strong>й, не <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не <strong>о</strong>тделен<br />

<strong>о</strong>т<strong>о</strong> лб<strong>а</strong>; лишь д<strong>в</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. Н.<br />

пермь — ныне. Д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Cicadellidea<br />

и Cercopidea.<br />

НАДСЕМЕЙСТВО CICADELLIDEA<br />

(Jassoidea)<br />

Н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нные, реже переп<strong>о</strong>нч<strong>а</strong>тые<br />

<strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; <strong>в</strong>ет<strong>в</strong>и М и CuA<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> з<strong>а</strong>м<strong>к</strong>нутых <strong>в</strong>нутренних<br />

ячее<strong>к</strong> между R, М и CuA. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

<strong>о</strong>чень неб<strong>о</strong>льшим <strong>в</strong><strong>о</strong>з<strong>в</strong>ышением <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я или без нег<strong>о</strong>; RS <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я, CuA пр<strong>о</strong>­<br />

<strong>с</strong>т<strong>а</strong>я; Ai и Аг <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong>им н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющим<strong>с</strong>я<br />

<strong>с</strong><strong>о</strong> лб<strong>о</strong>м; передние ру<strong>к</strong>и тент<strong>о</strong>риум<strong>а</strong> не<br />

<strong>с</strong><strong>о</strong>единяют<strong>с</strong>я <strong>с</strong> з<strong>а</strong>дними. З<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и п<strong>о</strong>перечные,<br />

<strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> р<strong>а</strong><strong>с</strong>ширены; з<strong>а</strong>дние г<strong>о</strong>лени<br />

без шп<strong>о</strong>р, <strong>в</strong><strong>о</strong><strong>о</strong>ружены шип<strong>а</strong>ми. Н. пермь —<br />

ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Scytinopteridae, Biturritidae,<br />

Cicadellidae, Eurymelidae, Membracidae, Aetalionidae,<br />

Hylicidae.<br />

СЕМЕЙСТВО SCYTINOPTERIDAE HANDLIRSCH, 1904<br />

Н<strong>а</strong>д<strong>к</strong>рылье п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, SC нея<strong>с</strong>н<strong>а</strong>. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

без г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong> и без <strong>в</strong>ырез<strong>к</strong>и.<br />

Пермь — три<strong>а</strong><strong>с</strong>. Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Scytinopterinae, Ivaiinae, Permojassinae, Ingruinae.<br />

Ри<strong>с</strong>. 473—478. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Scytinopteridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Scytinopterinae<br />

473. Scytinoptera kaltanica Becker-Migdisova; А — н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 11,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 11,0; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 474. Scytinoptera sp.; нимф<strong>а</strong>, х 15,0;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 475.<br />

Scytinoptera picturata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 9,7, н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 476.<br />

173<br />

Sarbaloptera sarbalensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 8,8, н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 477. Регmolamproptera<br />

grandis Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 6,3,<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 478.<br />

Tychtoscytina kuznetskiensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 10,5,<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1952)<br />

http://jurassic.ru/

ПОДСЕМЕЙСТВО SCYTINOPTERINAE<br />

BECKER-MIGDISOVA, 1960<br />

В н<strong>а</strong>д<strong>к</strong>рылье SCP <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, идет лишь<br />

<strong>в</strong>д<strong>о</strong>ль R + М. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью;<br />

SC идет <strong>в</strong>пл<strong>о</strong>тную <strong>в</strong>д<strong>о</strong>ль переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я, <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>о</strong><strong>с</strong>ую жил<strong>к</strong>у, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленную<br />

<strong>о</strong>т Ri+2 н<strong>а</strong>з<strong>а</strong>д (ри<strong>с</strong>. 473 А, Б).<br />

Н. пермь — три<strong>а</strong><strong>с</strong>. Де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Scytinoptera Handlirsch, 1904 (Anomoscyta<br />

Martynov, 1928; Permocixius Martynov,<br />

1928; Scytinopterula Handlirsch, 1937). Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. kokeni Handlirsch, 1904; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

В н<strong>а</strong>д<strong>к</strong>рылье SCP <strong>с</strong> неб<strong>о</strong>льшим, н<strong>о</strong><br />

рез<strong>к</strong>им изгиб<strong>о</strong>м н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти<br />

R + М; Ми СиА, деля<strong>с</strong>ь ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>, <strong>о</strong>бр<strong>а</strong>зуют<br />

ряд неб<strong>о</strong>льших ячее<strong>к</strong>; <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми; щит<strong>о</strong><strong>к</strong> <strong>о</strong>чень длинный.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,2—8,4 мм (ри<strong>с</strong>. 474,<br />

475; т<strong>а</strong>бл. XIV, фиг. 1, 2). В<strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

из <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong> Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

и Приур<strong>а</strong>лья и из н. и <strong>в</strong>. перми Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл., Приур<strong>а</strong>лья<br />

и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Sarbaloptera Becker - Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — S. sarbalensis Becker-<br />

Migdisova, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>а</strong>рб<strong>а</strong>л<strong>а</strong> II). Н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>к</strong>рупные, шир<strong>о</strong><strong>к</strong>ие, <strong>о</strong>тн<strong>о</strong>шение длины <strong>к</strong> ширине<br />

1,8—1,9; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 8,8 мм (ри<strong>с</strong>. 476). Один <strong>в</strong>ид. Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permolamproptera Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — P. grandis Becker-Migdisova,<br />

1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>. Верхний К<strong>а</strong>лт<strong>а</strong>н). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>е<br />

— <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм длины. Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong><br />

неб<strong>о</strong>льшим <strong>в</strong><strong>о</strong>з<strong>в</strong>ышением <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>к</strong>ругленн<strong>а</strong>я; Ri <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я;<br />

R2 и RS дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнуты (ри<strong>с</strong>. 477;<br />

т<strong>а</strong>бл. XIV, фиг. 3). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tychtoscytina Becker-Migdisova,<br />

1952. Тип р<strong>о</strong>д<strong>а</strong> — Т. kuznetskiensis Becker-<br />

Migdisova, 1952; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья, М —<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее, пяти<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 5,8 мм (ри<strong>с</strong>. 478). Один <strong>в</strong>ид. В.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Homaloscytina Tillyard, 1926;<br />

Elliptoscarta Tillyard, 1926; Anomaloscytina Darn's,<br />

1942; (non Anomaloscytina Evans, 1943);<br />

174<br />

http://jurassic.ru/<br />

Triassoscytina Evans, 1956. Си<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

п<strong>о</strong>л<strong>о</strong>жение р<strong>о</strong>д<strong>а</strong> Stenoscytina Tillyard, 1926 нея<strong>с</strong>н<strong>о</strong>.<br />

ПОДСЕМЕЙСТВО IVAIINAE BECKER-MIGDISOVA,<br />

1960<br />

Н<strong>а</strong>д<strong>к</strong>рылье. SCP <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; M и CuP <strong>о</strong>бр<strong>а</strong>зуют<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии л<strong>о</strong>жный пере<strong>к</strong>ре<strong>с</strong>т;<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть М и СиА нея<strong>с</strong>н<strong>а</strong>; СиР <strong>о</strong>чень<br />

<strong>с</strong>л<strong>а</strong>б<strong>а</strong>я. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong>' <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тью и р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью.<br />

В. пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Ivaia Becker-Migdisova, 1960. Тип<br />

р<strong>о</strong>д<strong>а</strong> — /. indistincta Becker-Migdisova, 1960;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). М и СиА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бр<strong>а</strong>зуют<br />

л<strong>о</strong>жный пере<strong>к</strong>ре<strong>с</strong>т; ч<strong>а</strong><strong>с</strong>ть М нея<strong>с</strong>н<strong>а</strong>; CuP<br />

<strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,3 мм<br />

(ри<strong>с</strong>. 479). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Anaprosbole Becker-Migdisova, 1946.<br />

Тип р<strong>о</strong>д<strong>а</strong> —• A. ivensis Becker-Migdisova, 1946;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>а</strong><strong>с</strong>имметричн<strong>о</strong> <strong>с</strong><strong>к</strong><strong>о</strong>шенным<br />

передним <strong>к</strong>р<strong>а</strong>ем; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

R 2 <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и М и СиА <strong>с</strong>л<strong>а</strong>бые; М<br />

четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong><br />

д<strong>о</strong> неб<strong>о</strong>льш<strong>о</strong>й чешуй<strong>к</strong>и. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

10,8 мм (ри<strong>с</strong>. 480). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Вне СССР: Triassoscelis Evans, 1956; Triassoscytinopsis<br />

Evans, 1956; Mesonirvana<br />

Evans, 1956; Mesothymbris Evans, 1956.<br />

ПОДСЕМЕЙСТВО PERMOJASSINAE<br />

BECKER-MIGDISOVA, 1960<br />

Н<strong>а</strong>д<strong>к</strong>рылье. SCP длинн<strong>а</strong>я, идет <strong>в</strong>д<strong>о</strong>ль <strong>в</strong><strong>с</strong>ей<br />

длины R + М, R и Ri — д<strong>о</strong> <strong>в</strong>п<strong>а</strong>дения <strong>в</strong> передний<br />

<strong>к</strong>р<strong>а</strong>й. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> прямым передним<br />

<strong>к</strong>р<strong>а</strong>ем, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью<br />

и длинн<strong>о</strong>й SC, идущей близ<strong>к</strong><strong>о</strong> <strong>к</strong> R + М<br />

и R; SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong>; R <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й у <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. В. пермь. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>.<br />

Surijokovia Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — S. lata Becker-Migdisova, 1961; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем; SCP пр<strong>о</strong>х<strong>о</strong>дит н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>т R + М и RS и з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> ме<strong>с</strong>те<br />

<strong>в</strong>ет<strong>в</strong>ления R п<strong>о</strong>д прямым угл<strong>о</strong>м <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; R2 и RS длинные. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4,9 мм (ри<strong>с</strong>. 481). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.

Ри<strong>с</strong>. 479—484. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Scytinopteridae, п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Ivaiinae, Permojassinae, Ingruinae<br />

479. Ivaia Indistincta Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 14,4.<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 480.<br />

Anaprosbole ivensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, и<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 10,8, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1960). 481. Surijokovia lata Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 12,5. <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Миг­<br />

Вне СССР: Perniojassus Tillyard, 1926<br />

(Permojassula Handlirsch, 1937).<br />

ПОДСЕМЕЙСТВО INGRUINAE BECKER-MIGDISOVA,<br />

1946<br />

I [nom. transl. Becker-Migdisova, 1952<br />

(ex Ingruidae Becker-Migdisova, 1946)1<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong>чень неб<strong>о</strong>льш<strong>о</strong>е и уз<strong>к</strong><strong>о</strong>е, <strong>с</strong> уз<strong>к</strong>им<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем; SCP <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, н<strong>о</strong> я<strong>с</strong>н<strong>а</strong>я<br />

<strong>в</strong>д<strong>о</strong>ль R + М, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R; М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее, чем R. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

<strong>с</strong> <strong>с</strong>уженн<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью и <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й; передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми.<br />

ди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 482. Ingruo smolenskyl Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>ры.<br />

лье, х 15,7, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1960). 483. Kaltanospes kuznetskiensis Becker-Migdisova; н<strong>а</strong>д-'<br />

<strong>к</strong>рылья и тел<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>, х 12,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн,<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961) .484. Permododa membracoides Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 8,7, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн.<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Щит<strong>о</strong><strong>к</strong> шир<strong>о</strong><strong>к</strong>ий — <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> шире длины.<br />

Пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Ingruo Becker-Migdisova, 1960. Тип<br />

р<strong>о</strong>д<strong>а</strong> — /. lanceolatus Becker-Migdisova, 1960;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). В н<strong>а</strong>д<strong>к</strong>рылье RS и CuA делят<strong>с</strong>я п<strong>о</strong>чти<br />

н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA <strong>о</strong>чень б<strong>о</strong>льш<strong>о</strong>й,<br />

М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее, чем CuA. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 3—3,5 мм (ри<strong>с</strong>. 482). Четыре <strong>в</strong>ид<strong>а</strong>.<br />

В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Kaltanospes Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> —• К- kuznetskiensis Beckter-Migdisova,.<br />

1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). В н<strong>а</strong>д<strong>к</strong>рылье М и CuA делят~<br />

175<br />

http://jurassic.ru/

<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; R зн<strong>а</strong>чительн<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА шир<strong>о</strong><strong>к</strong>ий, н<strong>о</strong> умеренн<strong>о</strong>й<br />

длины. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4,3—5,1 мм (ри<strong>с</strong>.483;<br />

т<strong>а</strong>бл. XIV, фиг. 4). Три <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permododa Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. membracoides Becker-Migdisova,<br />

1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е,<br />

<strong>с</strong> уз<strong>к</strong><strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й и б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

ячее<strong>к</strong> между RS и М, <strong>а</strong> т<strong>а</strong><strong>к</strong>же между М и СиА.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,25 мм. (ри<strong>с</strong>. 484; т<strong>а</strong>бл.<br />

XIV, фиг. 5). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО BITURRITIDAE<br />

EVANS, 1953<br />

(Lampropteridae, Evans, 1948, nom. nud.)<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> без <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>ймы; М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>дин ряд неб<strong>о</strong>льших<br />

ди<strong>с</strong>т<strong>а</strong>льных ячее<strong>к</strong>; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong><br />

р<strong>а</strong>зделен<strong>а</strong> н<strong>а</strong> д<strong>в</strong>е п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> неб<strong>о</strong>льшим <strong>в</strong><strong>о</strong>з<strong>в</strong>ышением<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА <strong>о</strong>чень<br />

б<strong>о</strong>льш<strong>о</strong>й; М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; R <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> передним<br />

<strong>к</strong>р<strong>а</strong>ем. Темя д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>с</strong>ередины лице<strong>в</strong><strong>о</strong>й п<strong>о</strong>­<br />

<strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми, щит<strong>о</strong><strong>к</strong> б<strong>о</strong>льш<strong>о</strong>й — <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong><br />

длиннее ширины. Три<strong>а</strong><strong>с</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин р<strong>о</strong>д.<br />

Absoluta Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong>—A. distincta Becker-Migdisova, 1962;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>о</strong>чень т<strong>о</strong>л<strong>с</strong>тыми жил<strong>к</strong><strong>а</strong>ми;<br />

R2 <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> <strong>к</strong>р<strong>а</strong>ем <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти;<br />

R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее, чем М и СиА, п<strong>о</strong><strong>с</strong>ледняя<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R + М или идет<br />

<strong>в</strong>пл<strong>о</strong>тную <strong>в</strong>д<strong>о</strong>ль нее. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

4 мм (ри<strong>с</strong>. 485). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО CICADELLIDAE<br />

LATREILLE, 1825<br />

(Jassidae Spinola, 1850)<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, без <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>ймы или лишь <strong>с</strong> р<strong>а</strong>нт<strong>о</strong>м <strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. Мн<strong>о</strong>г<strong>о</strong> <strong>к</strong>рупных ячее<strong>к</strong>; М<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА шир<strong>о</strong><strong>к</strong>ий. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ужен<strong>о</strong>, <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью и <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й; передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>с</strong> неб<strong>о</strong>льшим <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong>м (ри<strong>с</strong>. 486;<br />

Ри<strong>с</strong>. 485—489. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Biturritidae, Cicadellidae, Eurymelidae<br />

485. Absoluta distincta Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, x 15,0 тын<strong>о</strong><strong>в</strong>, 1926). 488. Karabasla paucinervis Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 486. Durgades х 13,6, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 489. Mesojassus<br />

miocenica Becker-Migdisova; тел<strong>о</strong> и <strong>к</strong>рылья, х 11,2. ми<strong>о</strong>цен, ipsvlclensts Tillyard; н<strong>а</strong>д<strong>к</strong>рылье, х 11,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1951). 487. Karajassus crasslner- (Evans, 1956)<br />

nisMartyno v; тел<strong>о</strong> и <strong>к</strong>рылья, х 20,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Map-<br />

176<br />

http://jurassic.ru/<br />

485<br />

88

т<strong>а</strong>бл. XIV, фиг. 6). Юр<strong>а</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 17 п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> и б<strong>о</strong>лее 5000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>;<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Karajassus Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. crassinervis Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>в</strong>ытянут<strong>о</strong>; <strong>в</strong>ершин<strong>а</strong> <strong>о</strong><strong>к</strong>руглен<strong>а</strong> эллип<strong>с</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>;<br />

R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины 1<br />

н<strong>а</strong>д<strong>к</strong>рылья;<br />

М ди<strong>с</strong>т<strong>а</strong>льнее ее; R <strong>о</strong>бр<strong>а</strong>зует ряд <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> трехл<strong>о</strong>п<strong>а</strong><strong>с</strong>тн<strong>о</strong>е; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен н<strong>а</strong> гр<strong>а</strong>нице <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,7 мм (ри<strong>с</strong>. 487). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Karabasia Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. paucinervis Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, R пр<strong>о</strong>х<strong>о</strong>дит близ<strong>к</strong><strong>о</strong> <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю. Ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> ячей<strong>к</strong>и <strong>в</strong>ытянуты <strong>в</strong> длину;<br />

между RS и М б<strong>о</strong>льш<strong>а</strong>я, <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я центр<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong>; между М и CuA и CuA и CuP группы<br />

мел<strong>к</strong>их ячее<strong>к</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>в</strong><br />

2 р<strong>а</strong>з<strong>а</strong> шире длины. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

2,75 мм (ри<strong>с</strong>. 488). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Вне СССР: Eurymelidium Tillyard, 1919;<br />

Triassqjassus Tillyard, 1919; Mesoledra Evans,<br />

1956 (= Mesojassus Handlirsch, 1939; non. Mesojassus<br />

Tillyard, 1916); Jassites Handlirsch,<br />

1906; Mesojassoides Oman, 1937; Acocephalites<br />

Meunier, 1904; Cicadettium Westwood, 1854;<br />

Homopterulum Handlirsch, 1907.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и С. Амери<strong>к</strong>и, ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и<br />

и три р<strong>о</strong>д<strong>а</strong> из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong><br />

и В. Сибири.<br />

СЕМЕЙСТВО EURYMELIDAE AMYOT<br />

ET SERVILLE, 1843<br />

[nom. transl. Evans, 1934<br />

(ex Eurymelides Amyot et Serviille, 1843)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья пр<strong>о</strong>зр<strong>а</strong>чные или <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нные,<br />

<strong>с</strong> пр<strong>о</strong><strong>с</strong>ты'м или <strong>с</strong>етч<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я или х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я;<br />

R 2 <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей; М делит<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>: Mi+ 2 и Мз+4, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющих<br />

<strong>в</strong> <strong>в</strong>ершину CuA <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

<strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й neala и <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью;<br />

R 2 и RS имеют<strong>с</strong>я; М делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е<br />

<strong>в</strong>ет<strong>в</strong>и (ри<strong>с</strong>. 489). Три<strong>а</strong><strong>с</strong>—ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 25<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин — Mesojassus Tillyard, 1916 из три<strong>а</strong><strong>с</strong><strong>а</strong><br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО MEMBRACIDAE GERMAR, 1821<br />

[nom. transl. Dohrn, 1859<br />

(ex Membracides Germar, 1821)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>тые, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми,<br />

чет<strong>к</strong>ими пр<strong>о</strong>д<strong>о</strong>льными и п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми и <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>йм<strong>о</strong>й; ячей<strong>к</strong>и я<strong>с</strong>н<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленные и ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

непр<strong>а</strong><strong>в</strong>ильные; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е;<br />

М или <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, или <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

R + М <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

CuA. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> и щит<strong>о</strong><strong>к</strong> <strong>с</strong> п<strong>а</strong>рными и неп<strong>а</strong>рными<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Три<strong>а</strong><strong>с</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 2500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> теплых и<br />

умеренных <strong>о</strong>бл<strong>а</strong><strong>с</strong>тях, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

Амери<strong>к</strong>е. 13 п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Oxyrhachinae,<br />

Smiliinae, Darninae, Hyphinoinae, Terentiinae,<br />

Stegaspinae, Tolaniinae, Heteronotinae,<br />

Polyglyptinae, Darniopseinae, Ceresopseinae,<br />

Membracinae, Centrotinae; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

п<strong>о</strong><strong>с</strong>ледние четыре.<br />

ПОДСЕМЕЙСТВО DARNIOPSEINAE<br />

BECKER-MIGDISOVA, 1958<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й,<br />

ячей<strong>к</strong>и <strong>с</strong>д<strong>в</strong>инуты <strong>к</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>в</strong>д<strong>о</strong>ль <strong>в</strong><strong>с</strong>ей ег<strong>о</strong> ширины; <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных<br />

ячее<strong>к</strong> четыре; <strong>а</strong> 4 р<strong>а</strong>зделен<strong>а</strong> перетяж<strong>к</strong><strong>о</strong>й н<strong>а</strong> д<strong>в</strong>е<br />

ячей<strong>к</strong>и; RS длинн<strong>а</strong>я; М и CuA <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong><br />

<strong>в</strong> <strong>о</strong>бщий <strong>с</strong>тебель; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле б<strong>о</strong>льш<strong>о</strong>е,<br />

треуг<strong>о</strong>льн<strong>о</strong>е, <strong>с</strong> Ai и А 2; ег<strong>о</strong> з<strong>а</strong>дний уг<strong>о</strong>л<br />

<strong>с</strong>мещен ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>. В. три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Darniopsis Becker-Migdisova, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — D. tragopea Becker-Migdisova, 1958;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину; передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый, <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

<strong>с</strong> уз<strong>к</strong><strong>о</strong>й, упл<strong>о</strong>тненн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong>й; <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я<br />

<strong>к</strong><strong>а</strong>йм<strong>а</strong> з<strong>а</strong>ним<strong>а</strong>ет ди<strong>с</strong>т<strong>а</strong>льную треть н<strong>а</strong>д<strong>к</strong>рылья;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; R2 <strong>о</strong>бры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии;<br />

RS длинн<strong>а</strong>я; <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные ячей<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>­<br />

л<strong>о</strong>жены 1<br />

12 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 177<br />

http://jurassic.ru/<br />

<strong>в</strong>ерти<strong>к</strong><strong>а</strong>льн<strong>о</strong> <strong>в</strong> ряд; ai б<strong>о</strong>льш<strong>а</strong>я, m<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, пятиуг<strong>о</strong>льн<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4,7 мм (ри<strong>с</strong>. 490). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

ПОДСЕМЕЙСТВО CERESOPSEINAE<br />

BECKER-MIGDISOVA, 1958<br />

Н<strong>а</strong>д<strong>к</strong>рылья р<strong>а</strong><strong>с</strong>ширены; <strong>к</strong>рыл<strong>о</strong><strong>в</strong><strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong><br />

<strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>а</strong>я; <strong>в</strong><strong>с</strong>е ячей<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong><br />

центре <strong>к</strong>рыл<strong>а</strong>; передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>о</strong><br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong>й; Ri <strong>о</strong>тх<strong>о</strong>дит <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>, М и R <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

<strong>о</strong>дин <strong>с</strong>т<strong>в</strong><strong>о</strong>л; три <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные ячей<strong>к</strong>и. В. три<strong>а</strong><strong>с</strong>.<br />

Один р<strong>о</strong>д.

490. Darniopsis tragopea Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 12,7,<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 491. Ceresopsis<br />

costalls Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 10,7, <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958).492. Sphongophoriella reticulata Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 50,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигдн<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1958). 493. Mtnuta heteropterata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 19,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958).<br />

Ceresopsis Becker-Migdisova, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — С. costalis Becker-Migdisova, 1958;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>о</strong><strong>к</strong>руглен<strong>о</strong>; <strong>к</strong>рыл<strong>о</strong><strong>в</strong><strong>а</strong>я<br />

<strong>к</strong><strong>а</strong>йм<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> чет<strong>в</strong>ерти длины н<strong>а</strong>д<strong>к</strong>рылья.<br />

Ri <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R н<strong>а</strong> гр<strong>а</strong>нице б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети<br />

н<strong>а</strong>д<strong>к</strong>рылья и з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>; ai п<strong>о</strong>чти треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> (rm) п<strong>о</strong> длине р<strong>а</strong><strong>в</strong>н<strong>а</strong> трети длины<br />

н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,6 мм<br />

(ри<strong>с</strong>. 491). Три <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы'<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

ПОДСЕМЕЙСТВО MEMBRACINAE STAL, 1866<br />

В н<strong>а</strong>д<strong>к</strong>рылье ячей<strong>к</strong><strong>а</strong> mi п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> р<strong>а</strong>зделен<strong>а</strong><br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; ячей<strong>к</strong>и г и mi из<strong>о</strong>гнуты<br />

п<strong>о</strong>д тупым угл<strong>о</strong>м и рез<strong>к</strong><strong>о</strong> <strong>с</strong>уж<strong>а</strong>ют<strong>с</strong>я <strong>к</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию; жил<strong>к</strong>и R и М идут п<strong>о</strong>чти <strong>в</strong>пл<strong>о</strong>тную<br />

<strong>о</strong>дн<strong>а</strong> <strong>к</strong> друг<strong>о</strong>й; <strong>в</strong>ет<strong>в</strong>и R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, Ri, R2<br />

и RS п<strong>а</strong>р<strong>а</strong>ллельны, или <strong>к</strong><strong>о</strong>нцы <strong>в</strong>ет<strong>в</strong>ей <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зуя непр<strong>а</strong><strong>в</strong>ильные ячей<strong>к</strong>и; М и СиА<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дны ИЛИ лишь <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии,<br />

н<strong>о</strong> не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л. Щит<strong>о</strong><strong>к</strong><br />

<strong>о</strong>бычн<strong>о</strong> при<strong>к</strong>рыт <strong>в</strong>ытянут<strong>о</strong>й н<strong>а</strong>з<strong>а</strong>д передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й;<br />

передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong> р<strong>а</strong><strong>с</strong>ширении <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong><br />

прип<strong>о</strong>днят<strong>а</strong>, ее б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>ль-<br />

Ри<strong>с</strong>. 490—498. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Membracidae, Aetalionidae<br />

178<br />

http://jurassic.ru/<br />

492<br />

494. Maguvlopsls kotschnevi Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 18,0, н. юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1953). 495. Aetallon<br />

reticulatum (Linnaeus); г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, х 10,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Ю. Амери<strong>к</strong><strong>а</strong><br />

(Evans, 1946). 496. A. reticulatum (Linnaeus); з<strong>а</strong>дняя н<strong>о</strong>г<strong>а</strong>,<br />

х 15,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Ю. Амери<strong>к</strong><strong>а</strong> (Evans, 1946). 497. A. viticolle<br />

Stal; н<strong>а</strong>д<strong>к</strong>рылье, х 5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Ю. Амери<strong>к</strong><strong>а</strong> (Evans, 1946).<br />

498. Darthula hardwickit Gray; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

ЮВ Азия (Evans, 1946)<br />

ными <strong>к</strong>илями н<strong>а</strong> нижнем <strong>к</strong>р<strong>а</strong>е, плече<strong>в</strong>ые <strong>к</strong>или<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. В. три<strong>а</strong><strong>с</strong> — ныне. Д<strong>в</strong>е трибы:<br />

Hypsoprorini и Membracini; п<strong>о</strong><strong>с</strong>ледняя т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

ТРИБА HYPSOPRORINI HAUPT, 1929<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; ячей<strong>к</strong>и <strong>в</strong>ытянут<strong>о</strong>-<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й<br />

ф<strong>о</strong>рмы; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля уз<strong>к</strong>ие.<br />

В. три<strong>а</strong><strong>с</strong> — ныне. С<strong>о</strong><strong>в</strong>ременные р<strong>о</strong>ды <strong>в</strong><br />

не<strong>о</strong>тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Один и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емый<br />

р<strong>о</strong>д.<br />

Sphongophoriella Becker-Migdisova,<br />

1958. Тип р<strong>о</strong>д<strong>а</strong> — 5. reticulata Becker-Migdisova,<br />

1958; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Н<strong>а</strong>д<strong>к</strong>рылье л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>;<br />

три <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные (<strong>а</strong>) и пре<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong>и (р<strong>с</strong>) <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong> <strong>в</strong>ытянуты; гит <strong>о</strong>чень<br />

длинные, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии не з<strong>а</strong>м<strong>к</strong>нуты; SCP идет<br />

<strong>в</strong>пл<strong>о</strong>тную <strong>к</strong> R. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 1,2 мм<br />

(ри<strong>с</strong>. 492). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

ПОДСЕМЕЙСТВО CENTROTINAE STAL, 1866<br />

Из <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ых<strong>о</strong>дят д<strong>в</strong><strong>а</strong><br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> лишь <strong>о</strong>дин; SC, R и М б<strong>о</strong>лее

или менее п<strong>а</strong>р<strong>а</strong>ллельны; М] <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м,<br />

'<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине <strong>с</strong> CuA; <strong>а</strong> 3 р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong><br />

у <strong>в</strong>ершины; ячей<strong>к</strong><strong>а</strong> mi <strong>о</strong>бычн<strong>о</strong> не р<strong>а</strong>зделен<strong>а</strong><br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. Щит<strong>о</strong><strong>к</strong> б<strong>о</strong>лее или<br />

менее я<strong>с</strong>н<strong>о</strong> <strong>в</strong>иден, <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> не при<strong>к</strong>рыт <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми<br />

передне<strong>с</strong>пин<strong>к</strong>и; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й передне<strong>с</strong>пин<strong>к</strong>и<br />

<strong>с</strong> длинным <strong>с</strong>пинным шип<strong>о</strong>м, ред<strong>к</strong><strong>о</strong> <strong>с</strong> р<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными<br />

р<strong>а</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ниями; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> ин<strong>о</strong>гд<strong>а</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>-<strong>к</strong>уп<strong>о</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я или <strong>с</strong><strong>о</strong> <strong>с</strong>рединным<br />

<strong>с</strong>теблем, пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ющим<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>о</strong>рне<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е<br />

или <strong>с</strong><strong>о</strong> <strong>в</strong>здутиями <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния. Бедр<strong>а</strong> <strong>с</strong>переди<br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>ие, их б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>к</strong>р<strong>а</strong>я <strong>о</strong><strong>к</strong>руглены. Б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е<br />

<strong>в</strong>ид<strong>а</strong>ми п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, <strong>о</strong>бит<strong>а</strong>ющее <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тях. В. три<strong>а</strong><strong>с</strong> — ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Minuta Becker-Migdisova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. heteropterata Becker-Migdisova, 1958;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> <strong>в</strong>д<strong>о</strong>ль переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я д<strong>о</strong><br />

Mi+2 <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти и д<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния СиА 2<br />

<strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>;<br />

<strong>в</strong><strong>с</strong>е жил<strong>к</strong>и (R, М, CuAi и СиА 2) <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> CuP; R <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м;<br />

М делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и;<br />

<strong>в</strong>ет<strong>в</strong>и CuA длинные, п<strong>а</strong>р<strong>а</strong>ллельные; <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные<br />

ячей<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

2 мм (ри<strong>с</strong>. 493). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Maguviopsis Becker-Migdisova, 1953.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. kotchnevi Becker-Migdisova,<br />

1953; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, М<strong>а</strong>дыген).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong><strong>к</strong>ругл<strong>о</strong>-из<strong>о</strong>гнут<strong>о</strong>е,<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>нн<strong>о</strong>е п<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю;<br />

<strong>в</strong>е<strong>с</strong>ь передний <strong>к</strong>р<strong>а</strong>й д<strong>о</strong> М <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н; R,<br />

М и CuP <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии пр<strong>о</strong>х<strong>о</strong>дят близ<strong>к</strong><strong>о</strong> друг<br />

<strong>к</strong> другу, не <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь; М прям<strong>а</strong>я пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я жил<strong>к</strong><strong>а</strong>,<br />

пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong><strong>с</strong>редине н<strong>а</strong>д<strong>к</strong>рылья; R делит<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>нее <strong>с</strong>ередины, <strong>в</strong>ет<strong>в</strong>и ее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и<br />

п<strong>а</strong>р<strong>а</strong>ллельны М; CuA <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> CuP немн<strong>о</strong>г<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 3,3 мм (ри<strong>с</strong>. 494). Н. юр<strong>а</strong> Ср. Азии.<br />

СЕМЕЙСТВО AETALIONIDAE SP1NOLA, 1850<br />

[nom. transl. Dohrn, 1859<br />

(ex Aethalionoideae Spinola, 1850)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> чет<strong>к</strong>ими жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими<br />

б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> ячее<strong>к</strong> или <strong>с</strong>ет<strong>к</strong>у; R пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

не делит<strong>с</strong>я н<strong>а</strong> я<strong>с</strong>ные <strong>с</strong>т<strong>в</strong><strong>о</strong>лы 1<br />

, д<strong>а</strong>ет ряд<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>о</strong>че<strong>к</strong> <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю, <strong>к</strong> <strong>в</strong>ершине<br />

<strong>к</strong>рыл<strong>а</strong>; М трех-четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>ильн<strong>о</strong> изменчи<strong>в</strong><strong>о</strong>. В з<strong>а</strong>днем<br />

<strong>к</strong>рыле R не<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, не<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет з<strong>а</strong> <strong>к</strong>р<strong>а</strong>й передне<strong>с</strong>пин<strong>к</strong>и; передний<br />

<strong>о</strong>тдел н<strong>а</strong>лични<strong>к</strong><strong>а</strong> не <strong>о</strong>тделен <strong>о</strong>т неб<strong>о</strong>льш<strong>о</strong>г<strong>о</strong><br />

з<strong>а</strong>днег<strong>о</strong> <strong>о</strong>тдел<strong>а</strong>; л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти уздеч<strong>к</strong>и б<strong>о</strong>льшие;<br />

з<strong>а</strong>днел<strong>о</strong>бный и л<strong>о</strong>бный ш<strong>в</strong>ы 1<br />

<strong>о</strong>бр<strong>а</strong>зуют <strong>с</strong>пл<strong>о</strong>шную<br />

линию п<strong>о</strong>з<strong>а</strong>ди лб<strong>а</strong>; <strong>а</strong>нтенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

179<br />

пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>к</strong>и леж<strong>а</strong>т <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

ш<strong>в</strong><strong>а</strong>. Г<strong>о</strong>лени прямые, цилиндриче<strong>с</strong><strong>к</strong>ие, з<strong>а</strong>дние<br />

— длинные, упл<strong>о</strong>щенные, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> ряд<strong>о</strong>м<br />

щетин<strong>о</strong><strong>к</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й ее из<strong>о</strong>гнут, з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>у<br />

и <strong>в</strong>ытянут д<strong>о</strong> уз<strong>к</strong><strong>о</strong>г<strong>о</strong>, длинн<strong>о</strong>г<strong>о</strong> щит<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 5—28 мм (ри<strong>с</strong>. 495—498).<br />

Взр<strong>о</strong><strong>с</strong>лые и нимфы <strong>в</strong>едут <strong>о</strong>бще<strong>с</strong>т<strong>в</strong>енный <strong>о</strong>бр<strong>а</strong>з<br />

жизни и п<strong>о</strong><strong>с</strong>ещ<strong>а</strong>ют<strong>с</strong>я мур<strong>а</strong><strong>в</strong>ьями. С<strong>а</strong>м<strong>к</strong>и <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<br />

яйц<strong>а</strong> н<strong>а</strong> р<strong>а</strong><strong>с</strong>тения <strong>в</strong> <strong>с</strong>пеци<strong>а</strong>льные<br />

<strong>в</strong>ыделения и умир<strong>а</strong>ют; тел<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>лужит<br />

з<strong>а</strong>щит<strong>о</strong>й для ямц. П<strong>о</strong>лиф<strong>а</strong>ги. П<strong>а</strong>ле<strong>о</strong>ген — ны'не.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне из ЮВ<br />

Азии и тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й Амери<strong>к</strong>и, <strong>а</strong> т<strong>а</strong><strong>к</strong>же из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО CERCOPIDEA<br />

(Aphrophoroideae Spinola, 1850;<br />

Handlirsch, 1925)<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны; SC, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, длинн<strong>а</strong>я; ячей<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>о</strong>бычн<strong>о</strong><br />

<strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>я, или <strong>о</strong>ни мел<strong>к</strong>ие и з<strong>а</strong>ним<strong>а</strong>ют<br />

<strong>в</strong><strong>с</strong>ю ди<strong>с</strong>т<strong>а</strong>льную ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья; М и CuA <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я или <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я<br />

длинн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; <strong>в</strong><strong>о</strong>з<strong>в</strong>ышение переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>; М <strong>в</strong><strong>с</strong>егд<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, CuA <strong>с</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; Ai и А 2 <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

лишь <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти. Передние и з<strong>а</strong>дние ру<strong>к</strong>и<br />

тент<strong>о</strong>риум<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я. З<strong>а</strong>дние н<strong>о</strong>ги <strong>с</strong><strong>о</strong> шп<strong>о</strong>р<strong>а</strong>ми<br />

и без шип<strong>о</strong><strong>в</strong>; т<strong>а</strong>зи<strong>к</strong>и их <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ие.<br />

Три<strong>а</strong><strong>с</strong> — ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Eoscarterellidae,<br />

Procercopidae, Cercopidae, Clastopteridae,<br />

Machaerotidae; п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йден<strong>о</strong>.<br />

СЕМЕЙСТВО EOSCARTERELLIDAE EVANS, 1956<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong>руглые; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

R 2 мн<strong>о</strong>г<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong> делит<strong>с</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>с</strong>д<strong>в</strong>инут<strong>о</strong> н<strong>а</strong>з<strong>а</strong>д, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>-<br />

Ри<strong>с</strong>. 499—500. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eoscarterellidae<br />

449. Eoscarterella media Evans; н<strong>а</strong>д<strong>к</strong>рылье, x 10,5, <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Evans, 1956). 500. Prosbolopsites tillyardi Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 9,6, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960)<br />

http://jurassic.ru/<br />

12*

502Б<br />

602Л<br />

501<br />

Ри<strong>с</strong>. 501—508. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Procercopidae, Cercopidae, Clastopteridae, Archijassidae, Hylicellidae<br />

501. Procercopis longipenna Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 5,7,<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 502. Procercopina<br />

aslatica Martynov; A — н<strong>а</strong>д<strong>к</strong>рылье, x 6,4; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

X 5,6; н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 503. Megacercopis<br />

optima Cockerell; н<strong>а</strong>д<strong>к</strong>рылье, х 3,2, ми<strong>о</strong>цен, Прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(Cockerell, 1921). 504. Philagra kudiana Cockerell; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

ля и ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong><br />

<strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не (ри<strong>с</strong>. 499, 500). В. три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Три р<strong>о</strong>д<strong>а</strong>: Eoscarterella Evans, 1956; Eoscartoides<br />

Evans, 1956; Prosbolopsites Becker-<br />

Migdisova, 1961.<br />

СЕМЕЙСТВО PROCERCOPIDAE HANDLIRSCH, 1906<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

ширину; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R н<strong>а</strong> гр<strong>а</strong>нице б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

трети; М и СиА делят<strong>с</strong>я н<strong>а</strong> гр<strong>а</strong>нице<br />

180<br />

http://jurassic.ru/<br />

х 3,2, ми<strong>о</strong>цен, Прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Cockerell, 1921). 505. LocrU<br />

areata Walker; <strong>о</strong>бщий <strong>в</strong>ид, х 4,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Lallemand. 1912).<br />

506. Clastoptera proteus Fritch; <strong>о</strong>бщий <strong>в</strong>ид, х 9,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Lallemand,<br />

1912). 507. Archtjassus heerl Handlirsch; н<strong>а</strong>д<strong>к</strong>рылье,<br />

x 9,0, н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1906). 508. Hylicella colorata<br />

Evans; н<strong>а</strong>д<strong>к</strong>рылье, x 6,8, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Evans, 1955)<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трети н<strong>а</strong>д<strong>к</strong>рылья, <strong>в</strong>ет<strong>в</strong>и их <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й, <strong>в</strong>ытянут<strong>о</strong><br />

<strong>в</strong> длину, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью; СиА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и М и СиА уз<strong>к</strong>ие и длинные. В.<br />

пермь — н. юр<strong>а</strong>. Три р<strong>о</strong>д<strong>а</strong>.<br />

Procercopis Н a n d 1 i г s |С h, 1906. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. alutacea Handlirsch, 1906; н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья п<strong>о</strong>чти <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong><br />

б<strong>о</strong>льше ширины; R 2 и М <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

507

н<strong>а</strong>д<strong>к</strong>рылья д<strong>а</strong>ют не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей<br />

и <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми.<br />

В з<strong>а</strong>днем <strong>к</strong>рыле передний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти<br />

прям<strong>о</strong>й. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 14 мм (ри<strong>с</strong>. 501).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>: <strong>о</strong>дин из н. юры 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин<br />

из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Procercopina М <strong>а</strong> г t у п <strong>о</strong> v, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. asiatica Martynov, 1937, н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

п<strong>о</strong>чти <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

бедн<strong>о</strong>е, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>лений м<strong>а</strong>л<strong>о</strong>, <strong>к</strong><strong>а</strong>ждые д<strong>в</strong>е жил<strong>к</strong>и<br />

<strong>с</strong><strong>о</strong>единены <strong>о</strong>дн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. В з<strong>а</strong>днем<br />

<strong>к</strong>рыле ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я и б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ти <strong>в</strong>ыпу<strong>к</strong>лые.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 12 мм (ри<strong>с</strong>. 502).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> — н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: Stenoglyphis Evans, 1947.<br />

СЕМЕЙСТВО CERCOP1DAE LEACH, 1815<br />

[nom. transl. Leach, 1818<br />

(ex Cercopida Leach, 1815)]<br />

(Tomaspididae Melander-Brues, 1932;<br />

Metcalf, 1934; Poisson-Pesson, 1951)<br />

Н<strong>а</strong>д<strong>к</strong>рылья без я<strong>с</strong>н<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>ймы; RS <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинн<strong>а</strong>я;<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> у <strong>в</strong>ершины з<strong>а</strong><strong>о</strong><strong>с</strong>трен. В з<strong>а</strong>дних <strong>к</strong>рыльях<br />

RS р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. Щит<strong>о</strong><strong>к</strong> <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>че, чем<br />

передне<strong>с</strong>пин<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5—25 мм<br />

(ри<strong>с</strong>. 503—505). Б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е <strong>в</strong>ид<strong>а</strong>ми <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>:<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> р<strong>а</strong>зны'х <strong>о</strong>бл<strong>а</strong><strong>с</strong>тях. В. три<strong>а</strong><strong>с</strong><br />

— ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии 25 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Из -<strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Trifidella<br />

Evans, 1956 и Aldotrifidus Evans, 1956. Кр<strong>о</strong>ме<br />

т<strong>о</strong>г<strong>о</strong>, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>в</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и,<br />

Гренл<strong>а</strong>ндии и 3. Е<strong>в</strong>р<strong>о</strong>пы, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> пяти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С.<br />

К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>, В. Сибири и С. Амери<strong>к</strong>и, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>в</strong>яти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> из 3. Е<strong>в</strong>р<strong>о</strong>пы, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong> и В. Сибири.<br />

СЕМЕЙСТВО CLASTOPTERIDAE DOHRN, 1859<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, <strong>с</strong><strong>о</strong>единяющей<br />

ди<strong>с</strong>т<strong>а</strong>льные <strong>к</strong><strong>о</strong>нцы 1<br />

<strong>в</strong><strong>с</strong>ех жил<strong>о</strong><strong>к</strong>, и <strong>с</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленным<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong>ым уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> у <strong>в</strong>ершины<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong><strong>к</strong>руглен. В з<strong>а</strong>днем <strong>к</strong>рыле RS<br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> R, <strong>о</strong>бр<strong>а</strong>зуя <strong>о</strong><strong>к</strong>руглую ячей<strong>к</strong>у. Щит<strong>о</strong><strong>к</strong> <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>и<br />

длиннее, чем у передне<strong>с</strong>пин<strong>к</strong>и<br />

(ри<strong>с</strong>. 506). Не<strong>о</strong>ген — ныне. Бедн<strong>о</strong>е <strong>в</strong>ид<strong>а</strong>ми<br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

CERCOPIDEA INCERTAE SEDIS<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> Cercopidea <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я р<strong>о</strong>д<br />

Mesojassuta Evans, 1956 из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии<br />

(из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>).<br />

181<br />

CICADELLIDOMORPHA<br />

INCERTAE SEDIS<br />

СЕМЕЙСТВО ARCHIJASSIDAE<br />

BECKER-MIGDISOVA, 1962<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>им, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> треуг<strong>о</strong>льным<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем и пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ющей<br />

ег<strong>о</strong> длинн<strong>о</strong>й SC; RS <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинн<strong>а</strong>я; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья ряд п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> и <strong>о</strong>бычн<strong>о</strong> групп<strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных ячее<strong>к</strong>;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, треуг<strong>о</strong>льн<strong>о</strong>е. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 5—9,4 мм (ри<strong>с</strong>. 507). В. три<strong>а</strong><strong>с</strong> —<br />

н. юр<strong>а</strong>. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и 3. Е<strong>в</strong>р<strong>о</strong>пе:<br />

Archijassus Handlirsch, 1906 (Cercopidium Geinitz,<br />

1880; Atitizon Handlirsch. 1939; Eojassus<br />

Handlirsch, 1939; Liojassus Handlirsch, 1939;<br />

Mesoscytina Tillyard, 1919).<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> эт<strong>о</strong>му же <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у, <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я Mesocixiodes orthoclada Tillyard,<br />

1922 из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО HYLICELLIDAE EVANS, 1956<br />

В ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья CuAi ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М; п<strong>о</strong><strong>с</strong>ледняя трех-четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

д<strong>в</strong>е-три з<strong>а</strong>м<strong>к</strong>нутые ячей<strong>к</strong>и между<br />

<strong>в</strong>ет<strong>в</strong>ями R и М. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,5—<br />

3,8 мм (ри<strong>с</strong>. 508). В. три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>: Hylicelta Evans, 1956 и Hylicellites Becker-Migdisova,<br />

1962.<br />

ИНФРАОТРЯД<br />

FULGOROMORPHА<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, реже <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые,<br />

<strong>с</strong> длинн<strong>о</strong>й SC, ч<strong>а</strong>ще <strong>с</strong> б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

<strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong> и <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; R 2 у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

передний <strong>к</strong>р<strong>а</strong>й р<strong>а</strong>нее <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; tegulae<br />

имеют<strong>с</strong>я; <strong>с</strong>редние т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong>ытянуты, причленяют<strong>с</strong>я<br />

ближе <strong>к</strong> н<strong>а</strong>ружн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю, чем т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

передних н<strong>о</strong>г, и <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бны <strong>к</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым д<strong>в</strong>ижениям;<br />

з<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и неп<strong>о</strong>д<strong>в</strong>ижны; н<strong>а</strong>лични<strong>к</strong><br />

не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет ур<strong>о</strong><strong>в</strong>ня <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з, пл<strong>о</strong><strong>с</strong><strong>к</strong>ий;<br />

л<strong>о</strong>бн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong> неп<strong>а</strong>рным<br />

пр<strong>о</strong><strong>с</strong>тым гл<strong>а</strong>з<strong>к</strong><strong>о</strong>м, или же <strong>о</strong>н<strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

п<strong>а</strong>рные- пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>к</strong>и'р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям<br />

<strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з. Пермь — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 6500 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. 18 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Mundidae,<br />

Pereboriidae, Dictyopharidae, Fulgoridae,<br />

Fulgoridiidae, Lophopidae, Cixiidae, Achilidae,<br />

Ricaniidae, Flatidae, Delphacidae, Issidae, Tettigomotridae,<br />

Derbidae, Achilixiidae, Acanaloniidae,<br />

Tropiduchidae, Eurybrachiidae; п<strong>о</strong><strong>с</strong>ледние<br />

ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

неиз<strong>в</strong>е<strong>с</strong>тны.<br />

http://jurassic.ru/

' 610<br />

612<br />

509. Mundus nodosus Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, xll.O, <strong>в</strong>.<br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 5 1 0. Реreboria<br />

bella M. Zalessky; н<strong>а</strong>д<strong>к</strong>рылье, х 1,5, <strong>в</strong>. пермь, Приур<strong>а</strong>лье<br />

(<strong>о</strong>ригин<strong>а</strong>льный ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> п<strong>о</strong> ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий)<br />

511. Permopibrocha ratnosa Martynov, н<strong>а</strong>д<strong>к</strong>рылье, х 4,5, <strong>в</strong>.<br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong>ь<strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 512. Scytophara<br />

СЕМЕЙСТВО MUNDIDAE<br />

BECKER-MIGDISOVA, I960<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>-мембр<strong>а</strong>н<strong>о</strong>зн<strong>о</strong>е, без<br />

<strong>с</strong><strong>к</strong>ульптуры. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е; жил<strong>к</strong>и <strong>в</strong>ыпу<strong>к</strong>лые,<br />

<strong>с</strong> б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ыпу<strong>к</strong>лый;<br />

СиА делит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е.<br />

В. пермь. Один р<strong>о</strong>д.<br />

Mundus В е <strong>с</strong> k е r-M i g d i s <strong>о</strong> v a, 1960. Тип<br />

р<strong>о</strong>д<strong>а</strong>—M. nodosus Becker-Migdisova, 1960;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й, <strong>а</strong><strong>с</strong>имметричн<strong>о</strong><br />

<strong>с</strong>рез<strong>а</strong>нн<strong>о</strong>й <strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тью; R делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья;<br />

R2 и RS п<strong>а</strong>р<strong>а</strong>ллельны; М и СиА делят<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее,<br />

чем R; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и т<strong>о</strong>л<strong>с</strong>тые; <strong>в</strong><strong>с</strong>е<br />

жил<strong>к</strong>и <strong>с</strong> б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,1 мм<br />

Ри<strong>с</strong>. 509—514. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Mundidae, Pereboriidae<br />

182<br />

http://jurassic.ru/<br />

extensa Martynov; н<strong>а</strong>д<strong>к</strong>рылье х 9,0, <strong>в</strong>. пермь, Ю. Приур<strong>а</strong>лье<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 513. Neuropibrocha ramisubcostalis Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 3,3, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 514. Kalianopibr<strong>о</strong><strong>с</strong>п<strong>а</strong> boreoscytoides<br />

Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 10,3, н. пермь. Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

(ри<strong>с</strong>. 509). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

СЕМЕЙСТВО PEREBORIIDAE<br />

М. ZALESSKY, 1930<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; R 2, RS, М и СиА д<strong>а</strong>ют мн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей,<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> нея<strong>с</strong>ных; R 2 <strong>с</strong> ряд<strong>о</strong>м <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей; М и R делят<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее, чем СиА.<br />

Н. пермь — <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Pereboria М. Z <strong>а</strong> 1 е s s <strong>к</strong> у, 1930. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. bella М. Zalessky, 1930; <strong>в</strong>. пермь, К<strong>о</strong>ми АССР<br />

(<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, р. Переб<strong>о</strong>р<strong>а</strong>). RS гребенч<strong>а</strong>т<strong>а</strong>я<br />

<strong>с</strong> б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей;<br />

М мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, гребенч<strong>а</strong>т<strong>а</strong>я; мн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>в</strong>ыше<br />

40 мм (ри<strong>с</strong>. 510). Один <strong>в</strong>ид. В. пермь С. Ур<strong>а</strong>л<strong>а</strong>.

Permopibrocha Martynov, 1935. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. ramosa Martynov, 1935; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины;<br />

ег<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый; М и R <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине н<strong>а</strong>д<strong>к</strong>рылья, CuA — ди<strong>с</strong>т<strong>а</strong>льнее;<br />

R2 <strong>о</strong>бр<strong>а</strong>зует ряд п<strong>а</strong>р<strong>а</strong>ллельных <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 13 мм (ри<strong>с</strong>. 511). Один <strong>в</strong>ид.<br />

В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Scytophara Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

5. extensa Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> 2,5<br />

р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины, передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й;<br />

М и R <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья,<br />

CuA — ди<strong>с</strong>т<strong>а</strong>льнее; R2 гребенч<strong>а</strong>т<strong>а</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зует длинные п<strong>а</strong>р<strong>а</strong>ллельные <strong>в</strong>ет<strong>в</strong>и, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленные<br />

<strong>к</strong> <strong>в</strong>ершине <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

7,5 мм (ри<strong>с</strong>. 512). Один <strong>в</strong>ид..Пермь<br />

Приур<strong>а</strong>лья.<br />

Neuropibrocha Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — N. ramisubcostalis Becker-<br />

Migdisova, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>о</strong><strong>к</strong>руглен<strong>о</strong>, передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый;<br />

SC п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong>лил<strong>а</strong><strong>с</strong>ь <strong>с</strong> R, <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<br />

н<strong>а</strong> <strong>в</strong><strong>с</strong>ем <strong>с</strong><strong>в</strong><strong>о</strong>ем пр<strong>о</strong>тяжении <strong>с</strong>леды гребенч<strong>а</strong>т<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ения; н<strong>а</strong> гр<strong>а</strong>нице ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти между<br />

R, М и CuA длинные п<strong>о</strong>перечные жил<strong>к</strong>и; R2<br />

<strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных<br />

<strong>к</strong> <strong>в</strong>ершине; RS я<strong>с</strong>н<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, делит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

меди<strong>а</strong>льн<strong>о</strong>е зн<strong>а</strong>чительн<strong>о</strong> уже, <strong>о</strong>б<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми<br />

<strong>к</strong>рупн<strong>о</strong>ячеи<strong>с</strong>т<strong>о</strong>г<strong>о</strong> <strong>а</strong>рхеди<strong>к</strong>тия, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> перех<strong>о</strong>дящег<strong>о</strong><br />

<strong>в</strong> ячей<strong>к</strong>и; М <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>в</strong>ет<strong>в</strong>ей; CuA делит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее, чем М и R.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 26—38 мм (ри<strong>с</strong>. 513;<br />

т<strong>а</strong>бл. XIV, фиг. 7, 8). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kaltanopibrocha Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — К. boreoscytinoid.es Becker-<br />

Migdisova, 1961; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Передний <strong>к</strong>р<strong>а</strong>й<br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> п<strong>о</strong>чти прям<strong>о</strong>й; R 2 из<strong>о</strong>гнут<strong>а</strong>я,<br />

гребенч<strong>а</strong>т<strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>и ее п<strong>а</strong>р<strong>а</strong>ллельны Ri, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены<br />

<strong>к</strong> <strong>в</strong>ершине; RS <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; М гребенч<strong>а</strong>т<strong>а</strong>я;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть не р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>, <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я.<br />

Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм<br />

(ри<strong>с</strong>. 514). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Permobrachus Evans, 1943 из<br />

<strong>в</strong>. перми А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>,<br />

<strong>с</strong>юд<strong>а</strong> же <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я р<strong>о</strong>ды Crossbella Evans,<br />

1956; Mesocicadella Evans, 1956 и Permoglyphis<br />

Tillyard, 1926.<br />

183<br />

СЕМЕЙСТВО FULGORIDAE LETREILLE, 1807<br />

[nom. transl. Stephens, 1929<br />

(ex Fulgorellae Latreille, 1807)]<br />

(Laternariidae Haupt, 1929;<br />

Lystridae Pasco], 1882;<br />

Rhinortharia Distant, 1911)<br />

Н<strong>а</strong>д<strong>к</strong>рылья и з<strong>а</strong>дние <strong>к</strong>рылья ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширеньг, <strong>с</strong> <strong>о</strong>чень б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием,<br />

п<strong>о</strong><strong>к</strong>рыты гу<strong>с</strong>т<strong>о</strong>й <strong>с</strong>етью пр<strong>о</strong>д<strong>о</strong>льных и п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. В н<strong>а</strong>д<strong>к</strong>рыльях SC <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>,<br />

гребенч<strong>а</strong>т<strong>а</strong>я, <strong>с</strong> ряд<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных жил<strong>о</strong><strong>к</strong> п<strong>о</strong><br />

Ри<strong>с</strong>. 515—520. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Fulgoridae, Fulgoridiidae<br />

615. Nyctophylax uhleri Scudder; <strong>о</strong>бщий <strong>в</strong>ид, x 2,5, ми<strong>о</strong>цен, С<br />

Амери<strong>к</strong><strong>а</strong> (Scudder, 1890). 516. Fulgoridium breviradiatum Handlirsch;<br />

н<strong>а</strong>д<strong>к</strong>рылье, x 6,9, н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1939).<br />

517. Fulgorldiella rhaetica Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 8,7,<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 518. Fulgoropsis<br />

dubiosa Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, x 4,0, н, юр<strong>а</strong>. Ср. Азия<br />

М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 519. Eofulgoridlum klzylklensls Martynov; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 4,3, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 520. Eofulgoridium<br />

proximum Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,6, н. юр<strong>а</strong>, Ср.<br />

Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

http://jurassic.ru/

<strong>в</strong><strong>с</strong>ей длине, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю,<br />

и <strong>с</strong> пуч<strong>к</strong><strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных RS <strong>в</strong>ет<strong>в</strong>ей у <strong>в</strong>ершины;<br />

R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти н<strong>а</strong>д<strong>к</strong>рылья;<br />

СиА делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>.<br />

В з<strong>а</strong>днем <strong>к</strong>рыле <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> <strong>с</strong>ет<strong>к</strong><strong>о</strong>й.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными р<strong>о</strong>г<strong>а</strong>ми, шип<strong>а</strong>ми<br />

или <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Крупные ф<strong>о</strong>рмы: длин<strong>а</strong><br />

тел<strong>а</strong> д<strong>о</strong> 75 мм (ри<strong>с</strong>. 515). Тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>,<br />

бедн<strong>о</strong>е р<strong>а</strong>д<strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>: <strong>о</strong>дин из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), три<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. Амери<strong>к</strong>и<br />

и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии, д<strong>в</strong><strong>а</strong> из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО FULGORIDIIDAE HANDLIRSCH, 1939<br />

[nom. transl. Becker-Migdisova, 1961<br />

(ex Fulgoridiinae Handlirsch, 1939)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью; SC пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, не<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

пр<strong>о</strong>х<strong>о</strong>дит <strong>в</strong>д<strong>о</strong>ль переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong><br />

и <strong>в</strong>д<strong>о</strong>ль Ri; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья<br />

или немн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>нее; СиА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

трети или р<strong>а</strong>нее; <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong><br />

делят<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>чень<br />

ред<strong>к</strong>и; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> без<br />

теменн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>. Н. юр<strong>а</strong> — ныне. В<strong>о</strong><strong>с</strong>емь<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Fulgoridium Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Phryganidium balticum Geinitz, 1880; н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния. Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> ри<strong>с</strong>ун<strong>к</strong><strong>о</strong>м или пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>е,<br />

<strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; SC<br />

длинн<strong>а</strong>я, идет близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>к</strong>р<strong>а</strong>ю, не<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

R д<strong>а</strong>ет ряд <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей; СиА делит<strong>с</strong>я <strong>в</strong><br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти н<strong>а</strong>д<strong>к</strong>рылья, <strong>с</strong> тремя длинными<br />

<strong>в</strong>ет<strong>в</strong>ями; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>.<br />

8 з<strong>а</strong>дних <strong>к</strong>рыльях RS <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями;<br />

СиА трех-четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>; М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее,<br />

чем СиА. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6—<br />

9 мм (ри<strong>с</strong>. 516). 108 <strong>в</strong>ид<strong>о</strong><strong>в</strong> из 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

<strong>о</strong>дин <strong>в</strong>ид из И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. В. три<strong>а</strong><strong>с</strong> —<br />

н. юр<strong>а</strong>.<br />

Fulgoridiella Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> — F. rhaetica Becker-Migdisova, 1962;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> ри<strong>с</strong>ун<strong>к</strong><strong>о</strong>м, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; Ri д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> длинн<strong>а</strong>я, из<strong>о</strong>гнут<strong>а</strong>я,<br />

не п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им <strong>в</strong>ет<strong>в</strong>ям R 2.<br />

RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; М и СиА <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими ди<strong>с</strong>т<strong>а</strong>льными<br />

<strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 10 мм<br />

(ри<strong>с</strong>. 517). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Fulgoropsis Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

F. dubiosa Martynov, 1937; н. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). В з<strong>а</strong>днем <strong>к</strong>рыле R и<br />

RS <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближены; М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не <strong>с</strong> R, б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины, <strong>о</strong>бр<strong>а</strong>зует<br />

184<br />

http://jurassic.ru/<br />

<strong>с</strong>рединную ячей<strong>к</strong>у; <strong>в</strong>ет<strong>в</strong>и СиА длинные; жил<strong>к</strong>и<br />

из<strong>о</strong>гнутые. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 11 мм<br />

(ри<strong>с</strong>. 518). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: Fulgoridulum Handlirsch, 1939;<br />

Parafulgoridium Handlirsch, 1939; Metafulgoridium<br />

Handlirsch, 1939; Margaroptilon Handlirsch,<br />

1906, Cixiites Handlirsch, 1906.<br />

СЕМЕЙСТВО LOPHOPIDAE STAL, 1866<br />

[nom. transl. Melichar, 1903<br />

(ex Lophopida Stal, 1866)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; <strong>в</strong>ет<strong>в</strong>и<br />

длинные, <strong>с</strong>ближенные; SC <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, длинн<strong>а</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; R, М и СиА <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я р<strong>а</strong>нее <strong>с</strong>ередины<br />

н<strong>а</strong>д<strong>к</strong>рылья, R — не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее,<br />

чем М и СиА; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> уже, чем передне<strong>с</strong>пин<strong>к</strong><strong>а</strong>, <strong>с</strong><br />

длинным теменным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м; н<strong>а</strong>лични<strong>к</strong> <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>к</strong>илями. Юр<strong>а</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 28 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>о</strong>дин и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емый.<br />

Eofulgoridium Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong><br />

— Е. kisylkiense Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Н<strong>а</strong>д<strong>к</strong>рылье<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й ширины; SC пр<strong>о</strong>х<strong>о</strong>дит п<strong>о</strong><strong>с</strong>редине<br />

между С и R; R делит<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети<br />

н<strong>а</strong>д<strong>к</strong>рылья, М —н<strong>а</strong> <strong>с</strong>ередине, СиА — н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не<br />

R; н<strong>а</strong> СиА и RS мн<strong>о</strong>г<strong>о</strong> гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

<strong>в</strong>ет<strong>в</strong>ей; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. З<strong>а</strong>днее <strong>к</strong>ры 1<br />

-<br />

л<strong>о</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью;<br />

R <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; Ri е<strong>с</strong>ть; М и СиА<br />

трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые, делят<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 12,5 мм (ри<strong>с</strong>. 519—520). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ср. Азии.<br />

СЕМЕЙСТВО CIXIIDAE SPINOLA, 1850<br />

[nom. transl. Fieber, 1872<br />

(ex Cixioideae Spinola, 1850)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong>зр<strong>а</strong>чные, <strong>с</strong> умеренн<strong>о</strong><br />

б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием и пл<strong>о</strong>тными пр<strong>о</strong>д<strong>о</strong>льными<br />

и п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми<br />

н<strong>а</strong> них, <strong>с</strong>мы<strong>к</strong><strong>а</strong>ют<strong>с</strong>я п<strong>о</strong> <strong>в</strong><strong>с</strong>ему з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю; SC не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R; R<br />

делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья или б<strong>а</strong>з<strong>а</strong>льнее,<br />

М — ди<strong>с</strong>т<strong>а</strong>льнее; СиА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии прям<strong>а</strong>я.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью, пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми и <strong>в</strong>ырез<strong>к</strong><strong>о</strong>й<br />

<strong>в</strong>д<strong>о</strong>ль переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не <strong>в</strong>ытянут<strong>а</strong><br />

<strong>в</strong>перед. Б<strong>о</strong>лее 780 <strong>в</strong>ид<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 100 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Пермь — ныне.<br />

Mesocixietla Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. asiatica Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). Передний <strong>к</strong>р<strong>а</strong>й<br />

н<strong>а</strong>д<strong>к</strong>рылья прям<strong>о</strong>й; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>и<br />

R 2 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен или б<strong>о</strong>льше<br />

длины Ri +2; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R р<strong>а</strong>нее <strong>с</strong>ередины<br />

н<strong>а</strong>д<strong>к</strong>рылья; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий.

521. Mesocixiella asiatica Martynov; н<strong>а</strong>д<strong>к</strong>рылье, x 8,0, н.<br />

юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 522. Cycloscytina extensa<br />

Martynov; A — н<strong>а</strong>д<strong>к</strong>рылье, x 9,0, Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 6,5; н.<br />

юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 523. Surijokocixius tomiensis<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 14,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 524. Permocixiella<br />

venosa Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, х 6,5, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 525. Scytoclxius mendax Martynov;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 12,0, <strong>в</strong>. пермь, Ю. Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,5 мм (ри<strong>с</strong>. 521; т<strong>а</strong>бл. XV,<br />

фиг. 1). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл., н. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Cycloscytina Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong>—<br />

С. delutinervis Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину; пе­<br />

Ри<strong>с</strong>. 521—530. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Cixiidae, Achilidae<br />

185<br />

http://jurassic.ru/<br />

1937). 526. Boreocixius sibiricus Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 13,0, н. три<strong>а</strong><strong>с</strong>, С Сибирь (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1955). 527. Cixiella<br />

reducta Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 14,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 528. Vitreacixius elliptlcus Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 8,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1962). 529. Asiocixius fulgoroides Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 6,8, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 530. Helicoptera<br />

sp.; <strong>о</strong>бщий <strong>в</strong>ид, х 8,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Haupt, 1935)<br />

редний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>с</strong> ряд<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей;<br />

RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R р<strong>а</strong>нее <strong>с</strong>ередины;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong><br />

п<strong>о</strong>л<strong>о</strong>гим <strong>в</strong>ырез<strong>о</strong>м переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; М <strong>с</strong> д<strong>в</strong>умя<br />

длинными <strong>в</strong>ет<strong>в</strong>ями: передней гребенч<strong>а</strong>т<strong>о</strong>й и<br />

з<strong>а</strong>дней пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA длинный. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 9 мм (ри<strong>с</strong>. 522). 17 <strong>в</strong>ид<strong>о</strong><strong>в</strong>.

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и <strong>в</strong>. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Surijokocixius Becker - Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — 5. tomiensis Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Н<strong>а</strong>д<strong>к</strong>рылье ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым упл<strong>о</strong>тненным<br />

передним <strong>к</strong>р<strong>а</strong>ем; <strong>в</strong>ет<strong>в</strong>и R 2 и RS п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA длинный и из<strong>о</strong>гнутый.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 4,3 мм<br />

(ри<strong>с</strong>. 523). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permocixiella Becker - Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — P. venosa Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е,<br />

<strong>с</strong> пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми, <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину;<br />

передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; R, Ri+г и Ri —<br />

прям<strong>о</strong>е пр<strong>о</strong>д<strong>о</strong>лжение <strong>о</strong>дн<strong>а</strong> друг<strong>о</strong>й; М делит<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> CuA; е<strong>с</strong>ть делящ<strong>а</strong>я линия;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA <strong>с</strong> длинными прямыми <strong>в</strong>ет<strong>в</strong>ями.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 10,5 мм (ри<strong>с</strong>. 524). Один<br />

<strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Scytocixius Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

5. mendax Martynov, 1937; пермь, Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ужен<strong>о</strong>,<br />

передний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й; R и RS<br />

дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнутые, пр<strong>о</strong><strong>с</strong>тые; R и CuA <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, М — ди<strong>с</strong>т<strong>а</strong>льнее; <strong>в</strong>ет<strong>в</strong>и<br />

CuA длинные, прямые. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 5,5 мм (ри<strong>с</strong>. 525). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь<br />

Приур<strong>а</strong>лья и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Boreocixius Becker-Migdisova, 1955.<br />

Тип р<strong>о</strong>д<strong>а</strong> — В. sibiricus Becker-Migdisoya, 1955;<br />

н. три<strong>а</strong><strong>с</strong>, С. Сибирь. Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>к</strong>ругленн<strong>о</strong>е,<br />

<strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым и упл<strong>о</strong>тненным передним <strong>к</strong>р<strong>а</strong>ем;<br />

<strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; RS и Ri + 2 <strong>с</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA длинный,<br />

из<strong>о</strong>гнутый. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5,5 мм<br />

(ри<strong>с</strong>. 526). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. три<strong>а</strong><strong>с</strong> С. Сибири.<br />

Cixiella Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> — С. reducba Becker-Migdisova, 1962;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть пр<strong>о</strong>зр<strong>а</strong>чн<strong>а</strong>я, <strong>о</strong><strong>к</strong>ругл<strong>о</strong> <strong>с</strong>рез<strong>а</strong>н<strong>а</strong>;<br />

передний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>к</strong><strong>о</strong>р<strong>о</strong>че,<br />

чем Ri + R 2, RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины н<strong>а</strong>д<strong>к</strong>рылья. М <strong>с</strong>ильн<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, <strong>с</strong>рединн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льный<br />

изгиб CuA длинный, п<strong>о</strong>л<strong>о</strong>гий. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5,1 мм (ри<strong>с</strong>. 527). Один <strong>в</strong>ид.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Vitreacixius Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> — V. elliptica Becker-Migdisova, 1962;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-.<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, эл-.<br />

лип<strong>с</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы 1<br />

, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним<br />

186<br />

http://jurassic.ru/<br />

<strong>к</strong>р<strong>а</strong>ем; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

.немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее, чем у Cixiella;<br />

М мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>, делит<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины н<strong>а</strong>д<strong>к</strong>рылья; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA<br />

неб<strong>о</strong>льш<strong>о</strong>й. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 9 мм (ри<strong>с</strong>. 528).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Asiocixius Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> — A. fulgoroides Becker-Migdisova,<br />

1962; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Н<strong>а</strong>д<strong>к</strong>рылье эллиптиче<strong>с</strong><strong>к</strong>и <strong>о</strong><strong>к</strong>ругленн<strong>о</strong>е,<br />

пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>е, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е; жил<strong>к</strong>и <strong>с</strong>ближены; <strong>с</strong>т<strong>в</strong><strong>о</strong>л<br />

R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, меньше длины Ri +2; Mn, R 2 гребенч<strong>а</strong>ты.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 10,9 мм (ри<strong>с</strong>. 529).<br />

Один р<strong>о</strong>д. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и <strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong><br />

В. Сибири.<br />

Вне СССР: Triassocixius Tillyard, 1919; Меsocixioides<br />

Tillyard, 1919; Mesocixius Tillyard,<br />

1919 из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и,<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и и четыре р<strong>о</strong>д<strong>а</strong> из не<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>к</strong> эт<strong>о</strong>му же <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я<br />

р<strong>о</strong>д Cixioides Handlirsch, 1907 из <strong>в</strong>. юры<br />

Англии и, <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong> Triassocotis Evans, 1956<br />

из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО ACHILIDAE STAL, 1866<br />

[nom. transl. Melichar, 1903<br />

(ex Tympanophoridae Brunner, 1893)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> мембр<strong>а</strong>н<strong>о</strong>зные, <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я г<strong>о</strong>риз<strong>о</strong>нт<strong>а</strong>льн<strong>о</strong> или пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>-<strong>к</strong>рыше<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>;<br />

ди<strong>с</strong>т<strong>а</strong>льные ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылий з<strong>а</strong><br />

<strong>в</strong>ершин<strong>о</strong>й <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширены и н<strong>а</strong>х<strong>о</strong>дят при<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>нии <strong>о</strong>дн<strong>а</strong> н<strong>а</strong> другую; SC <strong>о</strong>бычн<strong>о</strong> без<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> у <strong>в</strong>ершины з<strong>а</strong><strong>о</strong><strong>с</strong>трен. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> уз<strong>к</strong><strong>а</strong>я;<br />

н<strong>а</strong>лични<strong>к</strong> <strong>с</strong> я<strong>с</strong>ными б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>к</strong>илями; д<strong>в</strong><strong>а</strong><br />

пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>а</strong>; п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong><br />

удлинен. Б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>к</strong>или передне<strong>с</strong>пин<strong>к</strong>и из<strong>о</strong>гнуты.<br />

Длин<strong>а</strong> тел<strong>а</strong> 6—12 мм (ри<strong>с</strong>. 530)^ П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные (преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие) р<strong>о</strong>ды. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

три р<strong>о</strong>д<strong>а</strong>: <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и д<strong>в</strong><strong>а</strong> из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО R1CANIIDAE AMYOT ET SERVILLE,<br />

1843<br />

[nom. transl. Dohrn, 1859<br />

(ex Ricanides Amyot et Serville, 1843)]<br />

. Н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong>рупные, <strong>о</strong>бычн<strong>о</strong> непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й<br />

или треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы <strong>с</strong> <strong>в</strong>ытянут<strong>о</strong>й <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й; мн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>д<strong>о</strong>льных и п<strong>о</strong>переч-

Ри<strong>с</strong>. 531—533. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Ricaniidae, Flatidae<br />

1 Tritophania patruelis Jakobi; <strong>о</strong>бщий <strong>в</strong>ид, x 13,0, п<strong>а</strong>ле<strong>о</strong>ген, (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 533. Ormenis sp., <strong>о</strong>бщий <strong>в</strong>ид, x 3,9<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Jakobi, 1937). 532. Ludibrium lu- <strong>с</strong><strong>о</strong><strong>в</strong>рем., Ю. Амери<strong>к</strong><strong>а</strong> (Obenberger, 1957)<br />

dus Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 6,3, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия<br />

ных жил<strong>о</strong><strong>к</strong>; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти ряд п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не, <strong>о</strong>бр<strong>а</strong>зует п<strong>о</strong>перечную линию; SC<br />

длинн<strong>а</strong>я, <strong>с</strong> ряд<strong>о</strong>м мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных, п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>в</strong>ет<strong>в</strong>ей; R и М <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я <strong>с</strong>р<strong>а</strong>зу п<strong>о</strong><strong>с</strong>ле<br />

<strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления М <strong>о</strong>т R; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> неб<strong>о</strong>льш<strong>о</strong>е, без р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

передне<strong>с</strong>пин<strong>к</strong>е (ри<strong>с</strong>. 531). В. три<strong>а</strong><strong>с</strong> — ныне.<br />

Мн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне тр<strong>о</strong>пи<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

де<strong>с</strong>ять и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Ludibrium Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong> — L. ludus Becker-Migdisova, 1962; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, без г<strong>о</strong>рб<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; R пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я (<strong>в</strong><br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь лишь <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я<br />

из<strong>о</strong>гнут<strong>а</strong>я RS; R2 редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>); М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины;<br />

СиА делит<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и:<br />

пр<strong>о</strong><strong>с</strong>тую и гребенч<strong>а</strong>тую. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

7,3 мм (ри<strong>с</strong>. 532). Один <strong>в</strong>ид. В, три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, ше<strong>с</strong>ть <strong>в</strong>ымерших р<strong>о</strong>д<strong>о</strong><strong>в</strong>: <strong>о</strong>дин<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), д<strong>в</strong><strong>а</strong><br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и, <strong>о</strong>дин из не<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и и Ricaniites Handlirsch, 1907 из<br />

юры 3. Е<strong>в</strong>р<strong>о</strong>пы, <strong>а</strong> т<strong>а</strong><strong>к</strong>же три р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны: <strong>о</strong>дин из третичных <strong>о</strong>тл<strong>о</strong>жений А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии,<br />

<strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО FLATIDAE SPINOLA, 1839<br />

[nom. transl. Dohrn, 1859<br />

(ex Flatoides Spinola, 1839)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й ф<strong>о</strong>рмы, шир<strong>о</strong><strong>к</strong>ие,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>переди <strong>с</strong>рез<strong>а</strong>ны; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле я<strong>с</strong>н<strong>о</strong><br />

187<br />

http://jurassic.ru/<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>е; SC длинн<strong>а</strong>я, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями <strong>к</strong> <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>; R делит<strong>с</strong>я близ <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния н<strong>а</strong> R 2 и <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тую<br />

RS; М <strong>о</strong>тх<strong>о</strong>дит немн<strong>о</strong>г<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее <strong>в</strong>ет<strong>в</strong>ления<br />

R, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием RS;<br />

СиА прям<strong>а</strong>я, д<strong>в</strong>ух<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, идет <strong>в</strong>пл<strong>о</strong>тную<br />

<strong>в</strong>д<strong>о</strong>ль CuP; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен ряд п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong><br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти или непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й ф<strong>о</strong>рмы<br />

(ри<strong>с</strong>. 533). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 11 и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

<strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>с</strong>емь <strong>в</strong>ымерших р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и, д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и третичных<br />

<strong>о</strong>тл<strong>о</strong>жений Ю. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО DELPHACIDAE LEACH, 1815<br />

[nom. transl. Dohrn, 1859<br />

(ex Delphacida Leach, 1815)1<br />

Araeopidae Metcalf, 1938)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья уз<strong>к</strong>ие, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть эллиптиче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ытянут<strong>а</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, жил<strong>к</strong>и<br />

т<strong>о</strong>л<strong>с</strong>тые;, ряд чет<strong>к</strong>их п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>о</strong>тделяет<br />

менее пл<strong>о</strong>тную ди<strong>с</strong>т<strong>а</strong>льную ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> п<strong>о</strong>чти прямым передним<br />

<strong>к</strong>р<strong>а</strong>ем; из<strong>о</strong>гнут<strong>а</strong>я RS и б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ближенные<br />

М и СиА. З<strong>а</strong>дние г<strong>о</strong>лени <strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й ли<strong>с</strong>т<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й<br />

шп<strong>о</strong>р<strong>о</strong>й (ри<strong>с</strong>. 534). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне умеренн<strong>о</strong>й и х<strong>о</strong>л<strong>о</strong>дн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тей мн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> —<strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 137 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

и б<strong>о</strong>лее 1000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии

Ри<strong>с</strong>. 534—538. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Delphacidae, Issidae, Dictyopharidae<br />

M»L? a<br />

r ta fcff o^nc Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 21,0 <strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 537. Thanatodidya vtshneviensis Becker-<br />

К<br />

/53?? ; Л/,Г;^ "S 8<br />

Б е <strong>к</strong> <strong>к</strong> е<br />

. < Р-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>) 535. Etasmosce- Migdisova; тел<strong>о</strong> и'<strong>к</strong>рылья, x 4,4, ми<strong>о</strong>цен. С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (Бе<strong>к</strong><strong>к</strong>ер-Миг-<br />

Д И С 0 В <strong>а</strong> )<br />

Sta^ Т В ? ' • Т - 5 3 8<br />

- ОШу<strong>о</strong>рШ<strong>а</strong> sp.; з<strong>а</strong>днее <strong>к</strong>рьТл<strong>о</strong>, х 10,5, ми<strong>о</strong>цен,<br />

<strong>с</strong>т<strong>а</strong>н (м<strong>а</strong>ртын<strong>о</strong><strong>в</strong>. 1926). 536. MesotubUusirium asiatlcum Becker- С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Migdisova; н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е <strong>с</strong>б<strong>о</strong><strong>к</strong>у, <strong>к</strong> 12,5, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Бе<strong>к</strong>- л<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з ^е<strong>к</strong><strong>к</strong>ер тигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong><br />

прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти и <strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный<br />

р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong> и п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря и янт<strong>а</strong>ря Бирмы; <strong>о</strong>дин <strong>в</strong>ымерший<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и; <strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и и<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ISSIDAE SPINOLA, 1839<br />

[nom. transl. Schaum, 1850<br />

(ex Issiites Spinola, 1839)1<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нные и <strong>в</strong>ыпу<strong>к</strong>лые,<br />

<strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й ф<strong>о</strong>рмы; SC не <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; RS<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или гребенч<strong>а</strong>т<strong>а</strong>я, <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R <strong>в</strong> ег<strong>о</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> ин<strong>о</strong>гд<strong>а</strong> делит<strong>с</strong>я н<strong>а</strong><br />

три р<strong>а</strong><strong>в</strong>ные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти: ди<strong>с</strong>т<strong>а</strong>льную, <strong>а</strong>н<strong>а</strong>льную и<br />

юг<strong>а</strong>льную; п<strong>о</strong><strong>с</strong>ледняя <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>с</strong>ужен<strong>а</strong>. Г<strong>о</strong>-,<br />

л<strong>о</strong><strong>в</strong><strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> ширине груди или шире. Р<strong>а</strong>змеры<br />

3—7 мм. Мн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

<strong>в</strong>. юр<strong>а</strong> — ныне; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии четыре<br />

р<strong>о</strong>д<strong>а</strong>.<br />

Elasmoscelidium Martynov, 1926. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Е. rotundatum Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть р<strong>а</strong><strong>с</strong>ширен<strong>а</strong><br />

и <strong>о</strong><strong>к</strong>руглен<strong>а</strong>; SC п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> переднему <strong>к</strong>р<strong>а</strong>ю,<br />

<strong>о</strong>бр<strong>а</strong>зует упл<strong>о</strong>тненную п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>у; R 2 длинн<strong>а</strong>я,<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; RS и М <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; CuA <strong>с</strong><br />

длинными <strong>в</strong>ет<strong>в</strong>ями; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> з<strong>а</strong>гнут<strong>а</strong>;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>в</strong>ершины<br />

н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 7 мм<br />

188<br />

http://jurassic.ru/<br />

(ри<strong>с</strong>. 535). Четыре <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong><br />

и н. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Mesotubilustrium Becker-Migdisova,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong>—М. asiaticum Becker-Migdisova,<br />

1949; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм,<br />

К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Н<strong>а</strong>д<strong>к</strong>рылье д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, лишь<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> длиннее брюш<strong>к</strong><strong>а</strong>, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>;<br />

R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья или лишь<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льнее; R и RS <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые; CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

М <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я. Брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 4,7 мм (ри<strong>с</strong>. 536). Один <strong>в</strong>ид. В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы,<br />

<strong>о</strong>дин из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря, <strong>о</strong>дин из<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО DICTYOPHARIDAE SPINOLA, 1839<br />

[nom. transl. Kirschberg, 1868<br />

(ex Dictyopharoides Spinola. 1839)<br />

[Fulgorinae Haupt, 1929; Pseudophanides<br />

Amyot et Serville, 1843 (pars);<br />

Dohrn, 1859 (pars)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, жил<strong>к</strong>и пл<strong>о</strong>тные<br />

и ред<strong>к</strong>ие; SC не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; R делит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины; М и CuA н<strong>а</strong> <strong>с</strong>ередине; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти,<br />

<strong>о</strong>риентир<strong>о</strong><strong>в</strong><strong>а</strong>ны <strong>в</strong>ерти<strong>к</strong><strong>а</strong>льн<strong>о</strong>. З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong><br />

прямым передним <strong>к</strong>р<strong>а</strong>ем и без <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong>и.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong>ытянут<strong>а</strong>, <strong>с</strong> длинным теменным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> д<strong>в</strong>умя-тремя пр<strong>о</strong>д<strong>о</strong>льными<br />

<strong>к</strong>илями; д<strong>в</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. Н<strong>о</strong>ги длинные.

Длин<strong>а</strong> тел<strong>а</strong> 6—15 мм (ри<strong>с</strong>. 537—538). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Немн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> теплых и ж<strong>а</strong>р<strong>к</strong>их<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тях. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии три р<strong>о</strong>д<strong>а</strong>:<br />

<strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

и д<strong>в</strong><strong>а</strong> из не<strong>о</strong>ген<strong>а</strong> С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>.<br />

FULGOROMORPHA INCERTAE<br />

SEDIS<br />

Reticulocicada Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — R. brachyptera Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>о</strong>, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми; передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>н <strong>к</strong> <strong>в</strong>ершине; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; неб<strong>о</strong>льш<strong>о</strong>й узел<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

птер<strong>о</strong><strong>с</strong>тигмы; R 2 д<strong>а</strong>ет <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>в</strong>ет<strong>в</strong>и<br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти между<br />

RS, М и СиА гу<strong>с</strong>т<strong>а</strong>я <strong>с</strong>еть ячее<strong>к</strong>. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 6—7 мм (ри<strong>с</strong>. 539; т<strong>а</strong>бл. XV,<br />

фиг. 2). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ри<strong>с</strong>. 539—541. Fulgoridea incertae sedis и Auchenorrhyncha<br />

incertae sedis (<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dismorphoptilidae)<br />

539. Reticulocicada brachyptera Becker-Migdisova; фр<strong>а</strong>гмент н<strong>а</strong>д<strong>к</strong>рылья,<br />

x 10,4, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 540. Dysmorphoptiloldes elongata Evans; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 4,4, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Evans, 1956). 541. Mesoatracis reducta<br />

Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 10,0, н. юр<strong>а</strong>. Ср. Азия<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949)<br />

К Fulgoromorpha incertae sedis <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я<br />

т<strong>а</strong><strong>к</strong>же р<strong>о</strong>ды Pertnofulgor Tillyard, 1917; Н<strong>о</strong>mopterites<br />

Handlirsch, 1906; Pendodelphax Handlirsch,<br />

1907; Petropteron Cockerell, 1912; четы­<br />

541<br />

189<br />

http://jurassic.ru/<br />

ре р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и, <strong>а</strong> т<strong>а</strong><strong>к</strong>же<br />

<strong>о</strong>дин <strong>в</strong>ид нея<strong>с</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жения<br />

(Cicadellium psocus Westwood, 1854).<br />

AUCHENORRHYNCHA INCERTAE<br />

SEDIS<br />

СЕМЕЙСТВО DYSMORPHOPTIL1DAE HANDLIRSCH,<br />

1906<br />

Н<strong>а</strong>д<strong>к</strong>рылье непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й ф<strong>о</strong>рмы <strong>с</strong> рез<strong>к</strong>им<br />

р<strong>а</strong><strong>с</strong>ширением <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

н<strong>о</strong> <strong>с</strong> я<strong>с</strong>ным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; SC <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> R, н<strong>о</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я ее п<strong>а</strong>р<strong>а</strong>ллельные<br />

<strong>в</strong>ет<strong>в</strong>и, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленные <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; RS <strong>о</strong>чень длинн<strong>а</strong>я, <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R р<strong>а</strong>нее<br />

<strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья; М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> <strong>в</strong>ершину; СиА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, без<br />

рез<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> изгиб<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию М;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными Ai и А 2. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 10—16 мм (ри<strong>с</strong>. 540). Си<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

п<strong>о</strong>л<strong>о</strong>жение не <strong>в</strong>ыя<strong>с</strong>нен<strong>о</strong>: Handlirsch<br />

(1939) <strong>о</strong>тн<strong>о</strong><strong>с</strong>ил Dysmorphoptilidae <strong>к</strong> Jassidea,<br />

Э<strong>в</strong><strong>а</strong>н<strong>с</strong> (1956)—<strong>к</strong> Cercopidea; Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong><br />

(1949) <strong>о</strong>тн<strong>о</strong><strong>с</strong>ил<strong>а</strong> фр<strong>а</strong>гмент <strong>к</strong>рыл<strong>а</strong> Mesoatracus<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии ег<strong>о</strong> непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й ф<strong>о</strong>рмы<br />

<strong>к</strong> Flatidae. Три<strong>а</strong><strong>с</strong> — юр<strong>а</strong>. Три р<strong>о</strong>д<strong>а</strong>.<br />

Mesoatracis Becker-Migdisova, 1949. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. reducta, 1949; н. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>чень<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>к</strong>рыл<strong>о</strong> <strong>с</strong>р<strong>а</strong>зу рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширяет<strong>с</strong>я <strong>к</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию; <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь <strong>с</strong>леды <strong>в</strong>ет<strong>в</strong>ей SC; RS<br />

<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й <strong>с</strong> R 2; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>и п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й длины. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм (ри<strong>с</strong>. 541). Один <strong>в</strong>ид. Н.<br />

юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: Dysmorphoptila Handlirsch,<br />

1906; Dysmorphoptitoides Evans, 1956.<br />

СЕМЕЙСТВО IPSVIC1IDAE TILLYARD, 1919<br />

(Stenoviciidae Evans, 1956)<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью, <strong>с</strong><br />

рез<strong>к</strong><strong>о</strong>й з<strong>а</strong><strong>к</strong>р<strong>а</strong>ин<strong>о</strong>й (р<strong>а</strong>нт<strong>о</strong>м) <strong>в</strong>д<strong>о</strong>ль <strong>в</strong><strong>с</strong>ег<strong>о</strong> переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я д<strong>о</strong> <strong>в</strong>ершины н<strong>а</strong>д<strong>к</strong>рылья; SCP ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

пре<strong>в</strong>р<strong>а</strong>щен<strong>а</strong> <strong>в</strong> н<strong>о</strong>д<strong>а</strong>льную линию; RS или R 2<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, е<strong>с</strong>ли же имеет<strong>с</strong>я, т<strong>о</strong> М <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> СиА; е<strong>с</strong>ли СиА <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>,<br />

т<strong>о</strong> R 2 ИЛИ RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>. В з<strong>а</strong>днем <strong>к</strong>рыле<br />

R <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й — тремя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями; М пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й <strong>с</strong><br />

R и прям<strong>о</strong>й — <strong>с</strong> CuA; Ai и А 2 <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии,<br />

и А] нея<strong>с</strong>н<strong>а</strong>я; юг<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>тделяет<br />

неб<strong>о</strong>льшую м<strong>о</strong>рщини<strong>с</strong>тую юг<strong>а</strong>льную <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть.<br />

Эт<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Тильярд<strong>а</strong> Э<strong>в</strong><strong>а</strong>н<strong>с</strong> (Evans,<br />

1950, 1956) р<strong>а</strong>зделил н<strong>а</strong> три: Paraknightiidae<br />

(Paraknightia), Ipsviciidae (Ipsvicia, Ipsiviciopsis)<br />

и Stenoviciidae <strong>с</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льными р<strong>о</strong>д<strong>а</strong>ми.<br />

Пер<strong>в</strong>ые д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>о</strong>н <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит <strong>к</strong> Heteropte-

г<strong>а</strong>, п<strong>о</strong><strong>с</strong>леднее — <strong>к</strong> Cicadellidomorpha. С<strong>о</strong>гл<strong>а</strong>ш<strong>а</strong>я<strong>с</strong>ь<br />

<strong>с</strong> <strong>в</strong>ыделением Paraknightiidae <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>,<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляем п<strong>о</strong><strong>к</strong><strong>а</strong> (<strong>в</strong><strong>в</strong>иду <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ия<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> тел<strong>а</strong>) <strong>о</strong><strong>с</strong>т<strong>а</strong>льных Ipsviciidae <strong>в</strong> прежнем<br />

п<strong>о</strong>ним<strong>а</strong>нии (Evans, 1943). Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

6—14,2 мм (ри<strong>с</strong>. 542). В. пермь —<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>. 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Tomioscarta Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. surijokovensis Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Н<strong>а</strong>д<strong>к</strong>рылье эллиптиче<strong>с</strong><strong>к</strong>и-<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е.<br />

Ri, R2 и RS р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й<br />

т<strong>о</strong>ч<strong>к</strong>е; между М, CuA и RS <strong>о</strong>дн<strong>а</strong> или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

з<strong>а</strong>м<strong>к</strong>нутых ячее<strong>к</strong>; <strong>в</strong>ет<strong>в</strong>и CuA <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 6,8 мм (ри<strong>с</strong>. 543). Один <strong>в</strong>ид.<br />

В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tychlicola Becker-Migdisova, 1952.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. longipenna Becker-Migdisova,<br />

1952; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е,<br />

<strong>с</strong> <strong>в</strong>ытянут<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й; RS длинн<strong>а</strong>я, п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

R 2; п<strong>о</strong><strong>с</strong>ледняя и<strong>с</strong>чез<strong>а</strong>ет, не д<strong>о</strong>х<strong>о</strong>дя<br />

д<strong>о</strong> <strong>в</strong>ершины; <strong>в</strong>ет<strong>в</strong>и М длинные; CuA <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

б<strong>а</strong>з<strong>а</strong>льнее, чем М. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 7 мм<br />

(ри<strong>с</strong>. 544). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tychtoscarta Becker-Migdisova, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. sokolovensis Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е,<br />

длинн<strong>о</strong>е; R 2 <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; R, RS, Ri прямые, <strong>о</strong>бр<strong>а</strong>зуют<br />

шир<strong>о</strong><strong>к</strong>ий р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я М и CuA<br />

<strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,2 мм (ри<strong>с</strong>. 545;<br />

т<strong>а</strong>бл. XV, фиг. 3). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ipsviciella Becker-Migdisova, 1962.<br />

Тип р<strong>о</strong>д<strong>а</strong>—Л asiaticz Becker-Migdisova, 1962;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е; R прям<strong>а</strong>я, <strong>о</strong>т<strong>с</strong>ыл<strong>а</strong>ет<br />

п<strong>о</strong> <strong>в</strong><strong>с</strong>ей длине ряд п<strong>а</strong>р<strong>а</strong>ллельных <strong>в</strong>ет<strong>о</strong>че<strong>к</strong><br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; Ri, R 2 и RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

М ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> д<strong>а</strong>ет не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>о</strong>че<strong>к</strong>;<br />

CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> М.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм (ри<strong>с</strong>. 546).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Apheloscyta Tillyard, 1922;<br />

Permoscarta Tillyard, 1917; Stanleyana Evans,<br />

1943; Permovicia Evans, 1943; Palaeovicia<br />

Evans, 1943; Stenovicia Evans, 1943; Permagra<br />

Evans, 1943; Permocentrus Evans, 1956;<br />

Ipsvicia Tillyard, 1918 (Pseudipsvicia Handlirsch,<br />

1939), Ipsviciopsis Tillyard, 1922.<br />

INCERTAE FAMILIE<br />

Borisrohdendorfia Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong> — В. picturata Becker-Migdiso­<br />

190<br />

http://jurassic.ru/<br />

va, 1959; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>к</strong>ружен<strong>о</strong><br />

пл<strong>о</strong>тн<strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>-<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, <strong>с</strong> темными пятн<strong>а</strong>ми, не<br />

<strong>о</strong>тделен<strong>а</strong> длинными п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми;<br />

<strong>с</strong>и<strong>с</strong>темы жил<strong>о</strong><strong>к</strong> R, М и CuA ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> трудн<strong>о</strong><br />

р<strong>а</strong>зличимы; SC + R <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> ряд<strong>о</strong>м<br />

п<strong>а</strong>р<strong>а</strong>ллельных жил<strong>о</strong><strong>к</strong>, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>к</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и RS, М и CuA дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны<br />

и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>к</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю;<br />

R <strong>с</strong> четырьмя — пятью <strong>в</strong>ет<strong>в</strong>ями; М <strong>с</strong> д<strong>в</strong>умя<br />

— четырьмя <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 мм (ри<strong>с</strong>. 547; т<strong>а</strong>бл. XIV,<br />

фиг. 9). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> Auchenorrhyncha incertae<br />

sedis <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я Permoglyphis belmontensis Tillyard,<br />

1926; (Homopteron?) hahni Westwood,<br />

1854; Permodiphtheroides dubitans Martynov,<br />

1928; Parajassus hattorfensis Bode, 1953 и Triassoscarta<br />

Tillyard, 1919.<br />

ПОДОТРЯД STERNORRHYNCHA<br />

Крылья гетер<strong>о</strong>н<strong>о</strong>мны или г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мны. Передние<br />

<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, реже <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

или <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые. З<strong>а</strong>дние <strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые;<br />

юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>, или <strong>в</strong><strong>с</strong>е з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> лишь немн<strong>о</strong>г<strong>о</strong><br />

шире переднег<strong>о</strong>. Антенны длинные, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

И<strong>с</strong>х<strong>о</strong>дн<strong>а</strong>я групп<strong>а</strong> — дре<strong>в</strong>ние Агchescytinidea<br />

<strong>с</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мными <strong>к</strong>рыльями тип<strong>а</strong><br />

не<strong>в</strong>р<strong>о</strong>птеригии, утеря<strong>в</strong>шими юг<strong>а</strong>льный з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong><strong>к</strong>.<br />

Отдельные группы шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены <strong>в</strong><br />

перми и мез<strong>о</strong>з<strong>о</strong>е. В <strong>к</strong><strong>а</strong>йн<strong>о</strong>з<strong>о</strong>е (<strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> р<strong>а</strong>з<strong>в</strong>итием<br />

ц<strong>в</strong>ет<strong>к</strong><strong>о</strong><strong>в</strong>ых) эт<strong>о</strong>т п<strong>о</strong>д<strong>о</strong>тряд д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нения и д<strong>а</strong>ет ряд<br />

<strong>к</strong>рупных инфр<strong>а</strong><strong>о</strong>тряд<strong>о</strong><strong>в</strong> и н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, д<strong>о</strong><strong>с</strong>тигших<br />

<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>тепени <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ции. Н.<br />

пермь — ныне. Четыре инфр<strong>а</strong><strong>о</strong>тряд<strong>а</strong>: Archescytinomorpha,<br />

Aphidomorpha, Coccidomorpha,<br />

Psyllomorpha.<br />

И НФРАОТРЯД<br />

ARCHESCYTINOMORPHA<br />

В передних <strong>к</strong>рыльях CuA <strong>с</strong> рез<strong>к</strong>им изгиб<strong>о</strong>м<br />

<strong>к</strong> R + М и длинным ее б<strong>а</strong>з<strong>а</strong>льным <strong>о</strong>трез<strong>к</strong><strong>о</strong>м.<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, переп<strong>о</strong>нч<strong>а</strong>тые, <strong>в</strong>ытянутые<br />

<strong>в</strong> длину. З<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т передних не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>о</strong>льшей ширин<strong>о</strong>й и<br />

н<strong>а</strong>личием <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> CuA без изгиб<strong>а</strong> <strong>к</strong><br />

R + М, или же з<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е,<br />

<strong>а</strong> передние ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширены. К<strong>о</strong>нец<br />

брюш<strong>к</strong><strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> длинными прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Пермь. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archescytinidae и В<strong>о</strong>reoscytidae.

Ри<strong>с</strong>. 542—547. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Ipsviciidae и Auchenorrhyncha incertae sedis<br />

542. Ipsvicia jonesi Tillyard; A — н<strong>а</strong>д<strong>к</strong>рылье, x 5,0, три<strong>а</strong><strong>с</strong>,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Evans, 1956). 543. Tomioscarta surijokovensis Becker-<br />

Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, x 8,8, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 544. Tychticola longipenna Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 9,7, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

СЕМЕЙСТВО ARCHESCYT1NIDAE TILLYARD, 1926<br />

(Maueriidae G. Zalessky, 1937;<br />

Uraloscytinidae G. Zalessky, 1939;<br />

Permoscytinopsidae G. Zalessky, 1939;<br />

Maripsocidae G. Zalessky, 1939)<br />

Передняя и з<strong>а</strong>дняя п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мны;<br />

передние <strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, <strong>о</strong>бычн<strong>о</strong><br />

<strong>в</strong>ытянуты <strong>в</strong> длину, <strong>с</strong> эллиптиче<strong>с</strong><strong>к</strong>и-<strong>о</strong><strong>к</strong>ругленн<strong>о</strong>й,<br />

<strong>в</strong>ытянут<strong>о</strong>й ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле длинн<strong>о</strong>е; R 2 и RS п<strong>а</strong>р<strong>а</strong>ллельны, пр<strong>о</strong>х<strong>о</strong>дят<br />

д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; RS <strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

реже <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

неб<strong>о</strong>льш<strong>о</strong>е. З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong>лег<strong>к</strong><strong>а</strong> б<strong>о</strong>лее р<strong>а</strong><strong>с</strong>­<br />

191<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1952). 545. Tychtoscarta sokolovensis Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 11,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 546. Ipsviciella asiatica Becker-Migdisova<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 4,8, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962),<br />

547. Borisrohdendorfia picturata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

х 2,9, н. Пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959.<br />

ширены; б<strong>а</strong>з<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец СиА без изгиб<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

М, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>. Н<strong>а</strong> <strong>к</strong><strong>о</strong>нце<br />

брюш<strong>к</strong><strong>а</strong> у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

длинные прид<strong>а</strong>т<strong>к</strong>и. Пермь. 16 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Uraloscytina G. Zalessky, 1939. Тип<br />

р<strong>о</strong>д<strong>а</strong> — U. prosbolioides G. Zalessky, 1939;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> длинн<strong>о</strong>е, длин<strong>а</strong><br />

ег<strong>о</strong> <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; R + М дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я; <strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong><br />

между R 2 и RS п<strong>о</strong>чти <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> меньше длины<br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong> между Ri и R 2; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R д<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления<br />

р<strong>а</strong><strong>в</strong>ен длине Ri+г и меньше длины <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

М; М мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>, имеет<strong>с</strong>я <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь Ms; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>в</strong> 4—6 р<strong>а</strong>з <strong>к</strong><strong>о</strong>р<strong>о</strong>че длины<br />

http://jurassic.ru/

<strong>в</strong><strong>с</strong>ей <strong>с</strong>и<strong>с</strong>темы М; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong> изгиб<strong>а</strong><br />

<strong>к</strong> М б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины R + М. Ф<strong>о</strong>рмы<br />

<strong>к</strong>рупные, длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 12—20 мм<br />

(ри<strong>с</strong>. 548). Три <strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong> и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Maueria G. Zalessky, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М. sylvensis G. Zalessky, 1937; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Переднее <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, длин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> 3,5 р<strong>а</strong>з<strong>а</strong><br />

б<strong>о</strong>льше ширины; R + М <strong>с</strong>л<strong>а</strong>б<strong>о</strong>дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, длин<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> R меньше<br />

длины <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М; пять <strong>в</strong>ет<strong>в</strong>ей М; М4 <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я,<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й М5 нет; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М меньше трети длины<br />

<strong>в</strong><strong>с</strong>ей <strong>с</strong>и<strong>с</strong>темы М. Ф<strong>о</strong>рмы мел<strong>к</strong>ие; 4—6 мм<br />

(ри<strong>с</strong>. 549). Четыре <strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Sarbaloscytina Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong> — S. angustipennis Becker-<br />

Migdisova, 1959; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>а</strong>рб<strong>а</strong>л<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong><br />

<strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е: <strong>о</strong>тн<strong>о</strong>шение длины <strong>к</strong> ширине б<strong>о</strong>лее<br />

трех; R + М п<strong>о</strong>чти прям<strong>а</strong>я; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я;<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между R2 и RS <strong>в</strong>д<strong>в</strong><strong>о</strong>е<br />

меньше <strong>о</strong>трез<strong>к</strong><strong>а</strong> между Ri и R2; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен<br />

или <strong>к</strong><strong>о</strong>р<strong>о</strong>че, чем Ri+г, и <strong>в</strong> 1,5—2 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М; три длинные <strong>в</strong>ет<strong>в</strong>и М; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М р<strong>а</strong><strong>в</strong>ен<br />

трети длины <strong>в</strong><strong>с</strong>ей <strong>с</strong>и<strong>с</strong>темы М; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong> изгиб<strong>а</strong> <strong>к</strong> М б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины R + М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 12 мм<br />

(ри<strong>с</strong>. 550). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Archescytina Tillyard, 1926; Permoscytinopsis<br />

G. Zalessky, 1939). Тип р<strong>о</strong>д<strong>а</strong>—A. permiana<br />

Tillyard, 1926; н. пермь, С. Амери<strong>к</strong><strong>а</strong>.<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Длин<strong>а</strong> <strong>в</strong> 2,5—3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше<br />

ширины; R + М дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>, птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

длинн<strong>а</strong>я и шир<strong>о</strong><strong>к</strong><strong>а</strong>я; <strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между<br />

R 2 и RS <strong>в</strong> 1,7—2,5 меньше <strong>о</strong>трез<strong>к</strong><strong>а</strong> между Ri и<br />

R 2; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>о</strong>дн<strong>о</strong>й длины или немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че, чем<br />

Ri+2, и р<strong>а</strong><strong>в</strong>ен п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М; п<strong>о</strong><strong>с</strong>ледний<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ен длине ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

8—20 мм (ри<strong>с</strong>. 551, 552, 553). 11 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.; н. пермь С. Амери<strong>к</strong>и.<br />

548. Uraloscytina multinervosa Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х 5,3, н. пермь, Ур<strong>а</strong>л (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 549. Maueria<br />

sylvensis G. Zalessky; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х 4,8, н. пермь,<br />

Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1936). 550. Sarbaloscytina angustipennis<br />

Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>, х 6,0, н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 551. Archescytina gundersenl<br />

Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,3, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 552. A. gundersenl Becker-<br />

Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 4,3, <strong>в</strong>. Пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 553. A. lusca Becker-Migdisova;<br />

д _ переднее <strong>к</strong>рыл<strong>о</strong>, х 5,2; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 5,2; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 554. Kaltanoscy-<br />

Una nigra Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>, х 9,5, н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 555. Permopsylla<br />

Ри<strong>с</strong>. 548—561. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archescytinidae<br />

192<br />

http://jurassic.ru/<br />

Kaltanoscyttna Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong> — К- nigra Becker-Migdisova,<br />

1959; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее <strong>к</strong>рыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е: <strong>о</strong>тн<strong>о</strong>шение<br />

длины <strong>к</strong> ширине 2,2—2,6; R + М дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я; <strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong><br />

между R 2 и RS <strong>в</strong> 1,5—2 р<strong>а</strong>з<strong>а</strong> меньше<br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong> между Ri и R 2; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен длине Ri<br />

и <strong>в</strong> 2—2,5 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>в</strong> М и Ri +2; три<br />

<strong>в</strong>ет<strong>в</strong>и М, <strong>с</strong>т<strong>в</strong><strong>о</strong>л М немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong> изгиб<strong>а</strong> <strong>к</strong> М<br />

б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины R + М. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 6—10 мм (ри<strong>с</strong>. 554). Три <strong>в</strong>ид<strong>а</strong>.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permopsylla Tillyard, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. americana Tillyard, 1926; н. пермь, С. Амери<strong>к</strong><strong>а</strong>.<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> перетяж<strong>к</strong><strong>о</strong>й <strong>в</strong> <strong>с</strong>ередине;<br />

R + М дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> умеренн<strong>о</strong>й<br />

длины и ширины; <strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между<br />

R 2 и RS <strong>в</strong> 2—2,5 р<strong>а</strong>з<strong>а</strong> меньше <strong>о</strong>трез<strong>к</strong><strong>а</strong> между<br />

Ri и R2; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й ширины;<br />

R 2 и RS п<strong>а</strong>р<strong>а</strong>ллельны п<strong>о</strong> <strong>в</strong><strong>с</strong>ей длине;<br />

М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>, длин<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

длине ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong> изгиб<strong>а</strong> <strong>к</strong> М меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины R + М. Ф<strong>о</strong>рмы мел<strong>к</strong>и: 2—6 мм<br />

(ри<strong>с</strong>. 555, т<strong>а</strong>бл. XV, фиг. 4). В<strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Н. пермь Приур<strong>а</strong>лья, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>,<br />

С. Амери<strong>к</strong>и и <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Sojanos-cytina Martynov, 1933 (Ivascytina<br />

Martynov, 1933). Тип р<strong>о</strong>д<strong>а</strong> — S. grandis<br />

Martynov, 1933; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>в</strong> 3—3,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; R + М<br />

дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я и длинн<strong>а</strong>я;<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между R2 и RS <strong>в</strong><strong>а</strong>рьирует,<br />

<strong>в</strong> 2,2—5 р<strong>а</strong>з меньше <strong>о</strong>трез<strong>к</strong><strong>а</strong> между Ri и R 2;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л R зн<strong>а</strong>чительн<strong>о</strong> меньше длины Ri +2;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М п<strong>о</strong>чти <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че ее ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

четырех- или пяти<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти и <strong>в</strong> 2—3 р<strong>а</strong>з<strong>а</strong><br />

длиннее <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> R; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong><br />

<strong>в</strong>ершины изгиб<strong>а</strong> <strong>к</strong> М б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины<br />

R + М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 12—20 мм<br />

(ри<strong>с</strong>. 556, 557). Ше<strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

kuzbassensls Becker-Migdisova; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х 27,0, и.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 556. Sojanoscytina<br />

grandis Martynov: переднее <strong>к</strong>рыл<strong>о</strong>, х 5,0, <strong>в</strong>. пермь-<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933). 557. S. ramosa Becker,<br />

Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 4,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 558. Permopsyllopsis rossica G.<br />

Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х 12,0, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1939) 559. Paleoscytina sibirica Becker-Migdisova; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 13,5, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 560. P. brevistigmata Carpenter; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, х 13,0, н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1933). 561.<br />

Maripsocus ambiguus G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, x 8,0, и.<br />

пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1939).

13 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

http://jurassic.ru/

Maripsocus G. Zalessky. 1939. Тип р<strong>о</strong>д<strong>а</strong>—<br />

M. ambiguus G. Zalessky, 1939; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> немн<strong>о</strong>г<strong>о</strong> б<strong>о</strong>лее чем<br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; R + М п<strong>о</strong>чти прям<strong>а</strong>я;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong> <strong>к</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ен Ri+г и длиннее<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> меньше длины ее<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти, р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и<br />

М шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены; длин<strong>а</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

СиА д<strong>о</strong> <strong>в</strong>ершины изгиб<strong>а</strong> <strong>к</strong> М б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины R + М; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 7 мм (ри<strong>с</strong>. 561). Один<br />

<strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Paleoscytina Carpenter, 1933. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. brevistigmata Carpenter, 1933; н- пермь,<br />

С. Амери<strong>к</strong><strong>а</strong>. Переднее <strong>к</strong>рыл<strong>о</strong> без пережим<strong>а</strong> <strong>в</strong><br />

<strong>с</strong>ередине; R + М дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>о</strong>чень неб<strong>о</strong>льш<strong>а</strong>я; <strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между<br />

R2 и RS лишь немн<strong>о</strong>г<strong>о</strong> б<strong>о</strong>льше или меньше<br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong> между R! и R2; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>;<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong> <strong>в</strong>ершины<br />

изгиб<strong>а</strong> <strong>к</strong> М меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины R + М;<br />

СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, ИЛИ <strong>в</strong>ет<strong>в</strong>и ее <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближены<br />

и передняя <strong>в</strong>ет<strong>в</strong>ь нея<strong>с</strong>н<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 3—4 мм (ри<strong>с</strong>. 559, 560). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

Н. пермь С. Амери<strong>к</strong>и и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permopsyllopsis G. Zalessky, 1939.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. rossica G. Zalessky, 1939;<br />

н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> без пережим<strong>а</strong> <strong>в</strong><br />

<strong>с</strong>ередине; R + М дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я, без р<strong>а</strong><strong>с</strong>ширения; <strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я<br />

<strong>к</strong>рыл<strong>а</strong> между R2 и RS <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> меньше <strong>о</strong>трез<strong>к</strong><strong>а</strong><br />

между Ri и R 2; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й<br />

ширины; <strong>с</strong>т<strong>в</strong><strong>о</strong>л R р<strong>а</strong><strong>в</strong>ен длине Ri+г и меньше<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>;<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> между Мз+4 и<br />

передней <strong>в</strong>ет<strong>в</strong>ью СиА <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ий — <strong>в</strong><br />

3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше <strong>о</strong>трез<strong>к</strong><strong>а</strong> между <strong>в</strong>ет<strong>в</strong>ями СиА;<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть СиА д<strong>о</strong> <strong>в</strong>ершины изгиб<strong>а</strong> <strong>к</strong> М<br />

б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины R + М. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 5 мм (ри<strong>с</strong>. 558). Один <strong>в</strong>ид.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Вне СССР: Austroscytina Evans, 1943; Bekkerscytina<br />

Evans, 1958; Eoscytina Evans, 1958;<br />

Lithoscytina Carpenter, 1933; Permoscytina<br />

Tillyard, 1926; Triassopsylla Tillyard, 1917.<br />

СЕМЕЙСТВО BOREOSCYTIDAE<br />

BECKER-MIGDISOVA, 1949<br />

Передние <strong>к</strong>рылья <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>ильн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширены, <strong>с</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>й мембр<strong>а</strong>н<strong>о</strong>й и <strong>с</strong>л<strong>а</strong>быми<br />

жил<strong>к</strong><strong>а</strong>ми, б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ужены <strong>с</strong> пл<strong>о</strong>тными <strong>в</strong>ыпу<strong>к</strong>лыми<br />

жил<strong>к</strong><strong>а</strong>ми; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е;<br />

R2 и RS ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> неп<strong>а</strong>р<strong>а</strong>ллельны; R 2 <strong>в</strong>п<strong>а</strong>д<strong>а</strong>­<br />

194<br />

http://jurassic.ru/<br />

ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong><strong>с</strong>егд<strong>а</strong> р<strong>а</strong>нее <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я; RS <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

ин<strong>о</strong>гд<strong>а</strong> гребенч<strong>а</strong>т<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>о</strong>чень неб<strong>о</strong>льш<strong>о</strong>е. З<strong>а</strong>дние <strong>к</strong>рылья, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют или пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> прид<strong>а</strong>т<strong>к</strong>и<br />

тип<strong>а</strong> жужж<strong>а</strong>лец. Р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны Archescytinidae.<br />

В. пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Boreoscyta Becker-Migdisova, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — В. mirabilis Martynov, 1937;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>.<br />

С<strong>о</strong>ян<strong>а</strong>). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е;<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я прям<strong>а</strong>я;<br />

R2 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>; RS гребенч<strong>а</strong>т<strong>а</strong>я, <strong>с</strong> б<strong>о</strong>льшим<br />

чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей; СиА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я р<strong>а</strong>н<strong>о</strong>, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

<strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ий, делит<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 8—9 мм (ри<strong>с</strong>. 562). Три<br />

<strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Ри<strong>с</strong>. 562—563. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Boreoscytidae<br />

562. Boreoscyta mirabilis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, Хб,8, <strong>в</strong>.<br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 563.<br />

Archescytinopsis smolenskii Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х7,5, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949)<br />

Archescytinopsis Becker-Migdisova,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong> — Sojanoscytina latipennis Martynov,<br />

1933; н. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е; R 2 <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>;<br />

RS и М <strong>с</strong> м<strong>а</strong>лым чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей; СиА делит<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> шир<strong>о</strong><strong>к</strong>ий. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 6—7 мм (ри<strong>с</strong>. 563). Три <strong>в</strong>ид<strong>а</strong>.<br />

В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

ИНФРАОТРЯД APHIDOMORPHA<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, Е. Е. Айзенберг)<br />

Передние <strong>к</strong>рылья <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>ны, <strong>с</strong> упл<strong>о</strong>тненным<br />

передним <strong>к</strong>р<strong>а</strong>ем и <strong>о</strong>блегченным з<strong>а</strong>дним;<br />

з<strong>а</strong>дние <strong>к</strong>рылья зн<strong>а</strong>чительн<strong>о</strong> меньше пе-

редних, ин<strong>о</strong>гд<strong>а</strong> пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> жужж<strong>а</strong>льц<strong>а</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>, л<strong>о</strong>бный <strong>с</strong><strong>к</strong>лерит не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен,<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти переднег<strong>о</strong> <strong>о</strong>тдел<strong>а</strong> н<strong>а</strong>лични<strong>к</strong><strong>а</strong><br />

не <strong>о</strong>тчленены. У <strong>к</strong>рыл<strong>а</strong>тых ф<strong>о</strong>рм <strong>с</strong>реднегрудь<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. Н<strong>о</strong>ги длинные, бег<strong>а</strong>тельные.<br />

К<strong>о</strong>нец брюш<strong>к</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми труб<strong>о</strong>ч<strong>к</strong><strong>а</strong>ми,<br />

де<strong>в</strong>ятый тергит ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>о</strong>м. Тел<strong>о</strong><br />

неред<strong>к</strong><strong>о</strong> <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми желез<strong>а</strong>ми. Антенны из<br />

трех — ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong> рин<strong>а</strong>риями. В.<br />

пермь — ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Permaphidopseidae,<br />

Pincombeidae, Genaphididae, Phylloxeridae,<br />

Aphididae.<br />

СЕМЕЙСТВО PERMAPHIDOPSEIDAE<br />

BECKER-MIGDISOVA, 1960<br />

Переднее <strong>к</strong>рыл<strong>о</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>о</strong>льше з<strong>а</strong>днег<strong>о</strong>;<br />

SCP <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R + М и R и <strong>о</strong>бр<strong>а</strong>зует<br />

упл<strong>о</strong>тненную жил<strong>к</strong>у; М,СиА и CuP <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют<br />

<strong>в</strong> R + SCP примерн<strong>о</strong> н<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дн<strong>а</strong> <strong>о</strong>т друг<strong>о</strong>й п<strong>о</strong>д угл<strong>о</strong>м 50—70°; М д<strong>в</strong>ухили<br />

трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>; CuA <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м или пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние ее <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R + М. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

цилиндриче<strong>с</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong>реднегрудь х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я. Де<strong>в</strong>ятый члени<strong>к</strong><br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong> х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>о</strong>м; <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые труб<strong>о</strong>ч<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Н<strong>о</strong>ги. длинные, бег<strong>а</strong>тельные, <strong>с</strong><br />

длинными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми. Пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Pertnaphidopsis Becker-Migdisova,<br />

1960. Тип р<strong>о</strong>д<strong>а</strong> — P. sojanensis Becker-Migdi­<br />

564. Pertnaphidopsis sojanensis Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>;<br />

x 10,2. <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1960). 565. Tshekardaella tshekardaensis Becker-Migdisova; <strong>к</strong>рылья<br />

и тел<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>, х9,7, н. пермь, Ур<strong>а</strong>л (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1960). 566. Kaltanaphis permiensis Becker-Migdisova; переднее<br />

sova, 1960; <strong>в</strong> пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong>ужен<strong>о</strong>;<br />

RS и R 2 дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>с</strong>редней<br />

<strong>в</strong>еличины; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>в</strong>д<strong>в</strong><strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че ее <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; <strong>в</strong>ет<strong>в</strong>и<br />

М и CuA <strong>с</strong>л<strong>а</strong>бые; CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, ее передняя<br />

<strong>в</strong>ет<strong>в</strong>ь <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М 3 +4. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

5,9 мм (ри<strong>с</strong>. 564). Один <strong>в</strong>ид. В. пермь<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Тshekardaella Becker-Migdisova, 1960.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Tsh. tshekardaensis Becker-Midgisova,<br />

1960; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужен<strong>а</strong>, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>о</strong><strong>к</strong>ругл<strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я;<br />

R + М <strong>с</strong>лег<strong>к</strong><strong>а</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льные<br />

<strong>к</strong><strong>о</strong>нцы <strong>в</strong><strong>с</strong>ех жил<strong>о</strong><strong>к</strong> чет<strong>к</strong>ие; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

умеренн<strong>о</strong>й <strong>в</strong>еличины; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> уз<strong>к</strong><strong>а</strong>я;<br />

М <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями; CuA <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5,5 мм<br />

(ри<strong>с</strong>. 565). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Kaltanaphis Becker-Migdisova, 1959.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Kaltanaphis permiensis Becker-Migdisova,<br />

1959; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Переднее <strong>к</strong>рыл<strong>о</strong><br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е; R + М п<strong>о</strong>чти прям<strong>а</strong>я;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я и длинн<strong>а</strong>я; R2 и RS<br />

длинные, п<strong>а</strong>р<strong>а</strong>ллельные н<strong>а</strong> <strong>в</strong><strong>с</strong>ем пр<strong>о</strong>тяжении;<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> длинн<strong>а</strong>я; М, RS и CuA<br />

Ри<strong>с</strong>. 564—568. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Permaphidopseidae, Pincombeidae<br />

<strong>к</strong>рыл<strong>о</strong>, X 1,7, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1959). 567. Pincombea mirabilis Tillyard; переднее <strong>к</strong>рыл<strong>о</strong>, X 24,0,<br />

<strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1922). 568. Eopincombea postica<br />

Davis; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 14,0, <strong>в</strong>. Пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Davis,<br />

1942)<br />

195 13*<br />

http://jurassic.ru/

Ри<strong>с</strong>. 569—575. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Genaphididae, Phylloxeridae, Aphididae<br />

669. Genaphis valdensis (Brodie); переднее <strong>к</strong>рыл<strong>о</strong>, X 12,7,'<strong>в</strong>. юр<strong>а</strong>,<br />

Англия (Handlirsch, 1906). 570. Adelges abietis (Linnaeus); A —<br />

переднее <strong>к</strong>рыл<strong>о</strong>, X 45,0; Б —з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,X45,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (М<strong>о</strong>рд<strong>в</strong>ил<strong>к</strong><strong>о</strong>,<br />

1934). 571. Aslphumtremulae Degeer; А — переднее <strong>к</strong>рыл<strong>о</strong><br />

X 30,0; Б—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 30,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (М<strong>о</strong>рд<strong>в</strong>ил<strong>к</strong><strong>о</strong>,<br />

1934). 572. Paratrichosiphum tattakanum Takahashi; <strong>о</strong>бщий <strong>в</strong>ид<br />

<strong>с</strong><strong>в</strong>ерху, X 30,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (М<strong>о</strong>рд<strong>в</strong>ил<strong>к</strong><strong>о</strong>, 1934). 573. Cinara pinicola<br />

Kaltenbach; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 30,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

<strong>о</strong>тх<strong>о</strong>дят <strong>о</strong>т R + М H;R н<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии, н<strong>о</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния их <strong>с</strong>ближены; М <strong>с</strong> д<strong>в</strong>умя длинными<br />

<strong>в</strong>ет<strong>в</strong>ями, делит<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти <strong>в</strong><strong>с</strong>ей ее<br />

длины; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние М нея<strong>с</strong>н<strong>о</strong>; СиА <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м и <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5,2 мм (ри<strong>с</strong>. 566; т<strong>а</strong>бл. XV,<br />

фиг. .5). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО PINCOMBEIDAE<br />

TILLYARD, 1922<br />

Переднее <strong>к</strong>рыл<strong>о</strong> зн<strong>а</strong>чительн<strong>о</strong> длиннее з<strong>а</strong>днег<strong>о</strong>,<br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию; М и СиА<br />

<strong>о</strong>тх<strong>о</strong>дят <strong>о</strong>т R + М <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>ль­<br />

,196<br />

http://jurassic.ru/<br />

X 30,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (М<strong>о</strong>рд<strong>в</strong>ил<strong>к</strong><strong>о</strong>, 1934). 574. Mindarus magnus Backer;<br />

А — переднее и з<strong>а</strong>днее <strong>к</strong>рылья, х 18,0; Б — <strong>а</strong>нтенн<strong>а</strong>,<br />

X 100,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Baker, 1922);<br />

В —- Mindarus transparens Germar; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, X 21,0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Germar, 1856). 575. Calaphis<br />

scudderi Backer; A — переднее <strong>к</strong>рыл<strong>о</strong>, X 18,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

X 100,0; В — <strong>а</strong>нтенн<strong>а</strong>, X 100,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Backer, 1922)<br />

н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; М и СиА <strong>о</strong>тх<strong>о</strong>дят <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й<br />

т<strong>о</strong>ч<strong>к</strong>е. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 4—5 мм<br />

(ри<strong>с</strong>. 567, 568). В. пермь А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Три р<strong>о</strong>д<strong>а</strong>:<br />

Pincombea Tillyard, 1922; Protopincombea<br />

Evans, 1943; Eupincombea Davis, 1942.<br />

СЕМЕЙСТВО GENAPHIDIDAE<br />

HANDLIRSCH, 1907<br />

В переднем <strong>к</strong>рыле птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти птер<strong>о</strong><strong>с</strong>тигмы;<br />

М и СиА не<strong>в</strong>ерти<strong>к</strong><strong>а</strong>льные, п<strong>о</strong>л<strong>о</strong>гие, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> Д; М <strong>с</strong> тремя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2 мм<br />

(ри<strong>с</strong>. 569). В. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д— Genaphis<br />

Handlirsch, 1907.

СЕМЕЙСТВО PHYLLOXERIDAE<br />

HERRICH-SCHAEFFER IN KOCH, 1857<br />

[nom. transl. Dreyfus, 1889<br />

(ex Phylloxeridae Herrich-Schaeffer in Koch, 1857)]<br />

(Chermesidae Herrich-Schaeffer in Koch, 1857;<br />

Adelgidae C. Borner, 1930)<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>,<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; <strong>о</strong>т R <strong>о</strong>тх<strong>о</strong>дят перпенди<strong>к</strong>улярн<strong>о</strong><br />

три пр<strong>о</strong><strong>с</strong>тые жил<strong>к</strong>и: RS, М, CuA<br />

(или М, CuA и CuP). В з<strong>а</strong>днем <strong>к</strong>рыле <strong>о</strong>дн<strong>а</strong><br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> или <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й.<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, трех-четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

реже члени<strong>к</strong><strong>о</strong><strong>в</strong> пять. Х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong> и <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые труб<strong>о</strong>ч<strong>к</strong>и<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>а</strong> брюш<strong>к</strong>е пять-ше<strong>с</strong>ть<br />

п<strong>а</strong>р <strong>с</strong>тигм. Де<strong>в</strong><strong>с</strong>т<strong>в</strong>енницы <strong>к</strong>рыл<strong>а</strong>тые и бе<strong>с</strong><strong>к</strong>рылые,<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> яйце<strong>к</strong>л<strong>а</strong>дущие. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>о</strong><strong>с</strong><strong>о</strong>би <strong>с</strong><br />

х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м или без нег<strong>о</strong>. Н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>н<strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нн<strong>о</strong>е и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> <strong>с</strong> дере<strong>в</strong>ьями.<br />

Чет<strong>в</strong>ертичные — ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Adelginae и Phylloxerinae; п<strong>о</strong><strong>с</strong>леднее <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

ПОДСЕМЕЙСТВО ADELGINAE С. BORNER,<br />

1930. ХЕРМЕСЫ<br />

Крылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>к</strong>р<strong>о</strong><strong>в</strong>ле<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>;<br />

н<strong>а</strong> з<strong>а</strong>дних <strong>к</strong>рыльях <strong>о</strong>дн<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong>.<br />

У <strong>к</strong>рыл<strong>а</strong>тых ф<strong>о</strong>рм пять члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн <strong>с</strong> тремя<br />

рин<strong>а</strong>риями. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые. ф<strong>о</strong>рмы <strong>к</strong>рыл<strong>а</strong>тые, <strong>с</strong><br />

х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> н<strong>а</strong> х<strong>в</strong><strong>о</strong>йных. Очень<br />

мел<strong>к</strong>ие ф<strong>о</strong>рмы. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 0,50—<br />

1,67 мм (ри<strong>с</strong>. 570). Чет<strong>в</strong>ертичные — ныне. Один<br />

р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны из чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений<br />

(<strong>к</strong><strong>о</strong>п<strong>а</strong>л Афри<strong>к</strong>и).<br />

СЕМЕЙСТВО APHIDIDAE LATREILLE, 1802<br />

[nom. transl. Herrich-Schaeffer in Koch, 1857<br />

(ex Aphidii Latreille, 1802)]<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> четырьмя <strong>к</strong><strong>о</strong><strong>с</strong>ыми жил<strong>к</strong><strong>а</strong>ми:<br />

RS, М (пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или ч<strong>а</strong>ще д<strong>в</strong>ух-трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я),<br />

CuA и CuP. У <strong>к</strong>рыл<strong>а</strong>тых ф<strong>о</strong>рм <strong>о</strong>бычн<strong>о</strong><br />

пять-ше<strong>с</strong>ть члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн <strong>с</strong> д<strong>в</strong>умя рин<strong>а</strong>риями;<br />

у бе<strong>с</strong><strong>к</strong>рылых — четыре — ше<strong>с</strong>ть, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> у бе<strong>с</strong><strong>к</strong>рылых мн<strong>о</strong>гОф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чны,<br />

<strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых групп<strong>а</strong>х трехф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные<br />

буг<strong>о</strong>р<strong>к</strong>и. У мн<strong>о</strong>гих «<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые» п<strong>о</strong>ры, буг<strong>о</strong>р<strong>к</strong>и<br />

или <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые труб<strong>о</strong>ч<strong>к</strong>и н<strong>а</strong> пят<strong>о</strong>м тергите, <strong>а</strong> н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong> х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong>. Н<strong>а</strong> брюш<strong>к</strong>е <strong>с</strong>емь п<strong>а</strong>р<br />

<strong>с</strong>тигм. Де<strong>в</strong><strong>с</strong>т<strong>в</strong>енницы жи<strong>в</strong><strong>о</strong>р<strong>о</strong>дящи, п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые<br />

ф<strong>о</strong>рмы яйце<strong>к</strong>л<strong>а</strong>дущие. Н<strong>а</strong> дере<strong>в</strong>яни<strong>с</strong>тых и тр<strong>а</strong><strong>в</strong>яни<strong>с</strong>тых<br />

р<strong>а</strong><strong>с</strong>тениях. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Четырн<strong>а</strong>дц<strong>а</strong>ть<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Schizoneurinae, Greenideinae,<br />

Lachninae, Mindarinae, Drepanosiphinae,<br />

Aphidinae, Phloeomyzinae, Anoeciinae,<br />

Thelaxinae, Chaitophorinae, Atheroidinae, Pterocommatinae,<br />

Cervaphidinae, Schoutedeniinae;<br />

п<strong>о</strong><strong>с</strong>ледние <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

неиз<strong>в</strong>е<strong>с</strong>тны.<br />

ПОДСЕМЕЙСТВО SCHIZONEURINAE<br />

HERRICH-SCHAEFFER IN KOCH, 1857<br />

[nom. transl. Obenberger, 1857<br />

(ex Schizoneuriden Koch, 1857)]<br />

(Pemphigiden Koch, 1857; Schizoneurinae<br />

Buckton, 1883; Eriosomatinae Baker, 1920;<br />

Borner, 1930, 1952; Shaposhnikov, 1953;<br />

Aizenberg, 1956)<br />

В переднем <strong>к</strong>рыле M пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти не <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я, реже <strong>с</strong> д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями.<br />

Обычн<strong>о</strong> ше<strong>с</strong>ть члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн у <strong>к</strong>рыл<strong>а</strong>тых,<br />

четыре — ше<strong>с</strong>ть у бе<strong>с</strong><strong>к</strong>рылых; <strong>а</strong>нтенны не<br />

длинее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы <strong>с</strong> грудью. Бе<strong>с</strong><strong>к</strong>рылые де<strong>в</strong><strong>с</strong>т<strong>в</strong>енницы<br />

и м<strong>о</strong>л<strong>о</strong>дые <strong>о</strong><strong>с</strong><strong>о</strong>би <strong>о</strong>бычн<strong>о</strong> <strong>в</strong>ыделяют н<strong>а</strong><br />

<strong>с</strong>пин<strong>к</strong>е белый <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й пуш<strong>о</strong><strong>к</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>о</strong><strong>с</strong><strong>о</strong>би<br />

личин<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зны, без х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong>. У <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

ф<strong>о</strong>рм х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong> нет. С<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые труб<strong>о</strong>ч<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

или рудимент<strong>а</strong>рны. Н<strong>а</strong> дере<strong>в</strong>ьях и <strong>к</strong>у<strong>с</strong>т<strong>а</strong>рни<strong>к</strong><strong>а</strong>х,<br />

пере<strong>с</strong>еленцы н<strong>а</strong> тр<strong>а</strong><strong>в</strong>яни<strong>с</strong>тых р<strong>а</strong><strong>с</strong>тениях.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,4—4,5 мм<br />

(ри<strong>с</strong>. 571). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь); <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и три из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО GREENIDEINAE WILSON, 1910<br />

[nom. transl. Takahashi, 1931<br />

(ex Greenideini Wilson, 1910)]<br />

В передних <strong>к</strong>рыльях M <strong>с</strong> тремя <strong>в</strong>ет<strong>в</strong>ями.<br />

Обычн<strong>о</strong> и <strong>с</strong><strong>а</strong>мцы и <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong>тые. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

т<strong>о</strong>н<strong>к</strong>ий, п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> ег<strong>о</strong> <strong>в</strong>тянут. Члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

брюш<strong>к</strong><strong>а</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емь-де<strong>в</strong>ять, п<strong>о</strong>з<strong>а</strong>ди <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

труб<strong>о</strong>че<strong>к</strong> <strong>о</strong>ни <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены, т<strong>а</strong><strong>к</strong> чт<strong>о</strong> <strong>с</strong>иф<strong>о</strong>ны<br />

длинные, п<strong>о</strong><strong>к</strong>рытые длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>тх<strong>о</strong>дят <strong>к</strong><strong>а</strong><strong>к</strong> бы <strong>о</strong>т <strong>к</strong><strong>о</strong>нц<strong>а</strong> брюш<strong>к</strong><strong>а</strong>. Х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong><br />

нет. Жи<strong>в</strong>ут н<strong>а</strong> ли<strong>с</strong>т<strong>в</strong>енных и <strong>в</strong>ечн<strong>о</strong>зеленых дере<strong>в</strong>ьях.<br />

Групп<strong>а</strong> <strong>о</strong>чень дре<strong>в</strong>няя. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3—4 мм (ри<strong>с</strong>. 572). Не<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие<br />

р<strong>о</strong>ды Ст<strong>а</strong>р<strong>о</strong>г<strong>о</strong> С<strong>в</strong>ет<strong>а</strong>, из них <strong>о</strong>дин р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из не<strong>о</strong>ген<strong>а</strong> Ю. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ПОДСЕМЕЙСТВО LACHNINAE<br />

HERRICH-SCHAEFFER IN KOCH, 1857<br />

[nom. transl. Passerini, 1862<br />

(ex Lachniden Herrich-Schaeffer in Koch, 1857)]<br />

(Cinarinae Mordvilko, 1934)<br />

В переднем <strong>к</strong>рыле птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> линейн<strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>,<br />

RS <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я и прям<strong>а</strong>я,<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> птер<strong>о</strong><strong>с</strong>тиг-<br />

т<br />

http://jurassic.ru/

мы, реже <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я; три <strong>в</strong>ет<strong>в</strong>и М,<br />

реже д<strong>в</strong>е или <strong>о</strong>дн<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. Гл<strong>а</strong>з<strong>а</strong> бе<strong>с</strong><strong>к</strong>рылых<br />

ф<strong>о</strong>рм <strong>о</strong>бычн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чны. Щ<br />

С<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые буг<strong>о</strong>р<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> \<br />

р<strong>а</strong>з<strong>в</strong>иты, у<strong>с</strong>еченн<strong>о</strong> <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идные или <strong>к</strong>уп<strong>о</strong>л<strong>о</strong><strong>в</strong>идные,<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>с</strong>тен<strong>к</strong>и п<strong>о</strong><strong>к</strong>рыты <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

Ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong>ыделяют лишь <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ую<br />

пыль, н<strong>о</strong> не пуш<strong>о</strong><strong>к</strong>. У <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм<br />

х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong> нет. С<strong>в</strong>яз<strong>а</strong>ны <strong>с</strong> дре<strong>в</strong>е<strong>с</strong>н<strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>тью:<br />

<strong>с</strong><strong>о</strong><strong>с</strong>ут н<strong>а</strong> <strong>к</strong><strong>о</strong>ре и <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>х<br />

х<strong>в</strong><strong>о</strong>йных, ред<strong>к</strong><strong>о</strong> н<strong>а</strong> других дере<strong>в</strong>ьях и н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>рнях тр<strong>а</strong><strong>в</strong>яни<strong>с</strong>тых д<strong>в</strong>уд<strong>о</strong>льных. Не мигрируют.<br />

С<strong>а</strong>мые <strong>к</strong>рупные тли д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют<br />

б—8 мм (ри<strong>с</strong>. 573). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО MINDARINAE<br />

TULLGREN, 1909<br />

[nom. transl. Backer, 1920<br />

(ex Mindarina Tullgren, 1909)1<br />

i[Rhyllaphidinae Mordvilko, 1948 (pars);<br />

Telaxidae Вбгпег, 1952 (pars);<br />

Thelaxinae Obenberger, 1957 (pars)]<br />

В переднем <strong>к</strong>рыле птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> длинн<strong>а</strong>я,<br />

<strong>с</strong>ерп<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>; RS<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы, <strong>о</strong>чень б<strong>о</strong>льш<strong>а</strong>я;<br />

М д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, <strong>к</strong>рылья <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>к</strong>р<strong>о</strong><strong>в</strong>ле<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. У<strong>с</strong>и<strong>к</strong>и, ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

шпицем. У 'бе<strong>с</strong><strong>к</strong>рылых де<strong>в</strong><strong>с</strong>т<strong>в</strong>енниц<br />

гл<strong>а</strong>з<strong>а</strong> трехф<strong>а</strong><strong>с</strong>ет<strong>о</strong>ч<strong>в</strong>ые. Бе<strong>с</strong><strong>к</strong>рылые и м<strong>о</strong>л<strong>о</strong>дые<br />

ф<strong>о</strong>рмы <strong>с</strong> ше<strong>с</strong>тью ряд<strong>а</strong>ми <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых желез, <strong>в</strong>ыделяющих<br />

<strong>с</strong>из<strong>о</strong>-белый пуш<strong>о</strong><strong>к</strong>. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> х<strong>в</strong><strong>о</strong>йных<br />

(Abietinae) не мигрируют. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3 мм (ри<strong>с</strong>. 574 <strong>а</strong>, б). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В<strong>с</strong>ег<strong>о</strong> <strong>о</strong>дин р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

и не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО DREPANOSIPHINAE<br />

HERRICH-SCHAEFFER IN KOCH, 1857<br />

[nom. transl. Aizenberg, 1956<br />

(ex Drepanosiphiden Herrich-Schaeffer in Koch, 1857)]<br />

(Callipteriden Herrich-Schaeffer in Koch, 1857;<br />

Callipterinae Mordvilko, 1895;<br />

Phyllaphidinae Aizenberg, 1932;<br />

Mordvilko, 1948; Callaphinidae Вбгпег, 1952)<br />

В переднем <strong>к</strong>рыле RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т з<strong>а</strong>днен<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong><br />

угл<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

Mi+2 и Mi рез<strong>к</strong><strong>о</strong> з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю,<br />

ин<strong>о</strong>гд<strong>а</strong> з<strong>а</strong>мещ<strong>а</strong>я RS, <strong>а</strong> М 3 +4 — <strong>к</strong> з<strong>а</strong>днему.<br />

Крылья <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>к</strong>рыше<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. Антен-<br />

198<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 576—577. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Aphidinae<br />

576. Yezabura malifolii Fitch; A — <strong>к</strong>рыл<strong>а</strong>тый <strong>с</strong><strong>а</strong>мец; Б —бе<strong>с</strong><strong>к</strong>рыл<strong>а</strong>я<br />

де<strong>в</strong><strong>с</strong>т<strong>в</strong>енниц<strong>а</strong>; В — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> бе<strong>с</strong><strong>к</strong>рыл<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>о</strong><strong>с</strong><strong>о</strong>би; <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Beier, 1938). 577. Aphis gurnetensis Cockerell; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Х12.8, <strong>о</strong>лиг<strong>о</strong>цен, С. Амери<strong>к</strong><strong>а</strong> (Cockerell, 1921)<br />

ны няти-ше<strong>с</strong>ти чле<strong>в</strong>и<strong>к</strong><strong>о</strong><strong>в</strong>ые, шпиц <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий<br />

или <strong>в</strong>ытянутый. С<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые труб<strong>о</strong>ч<strong>к</strong>и, буг<strong>о</strong>р<strong>к</strong>и или<br />

п<strong>о</strong>ры имеют<strong>с</strong>я. Х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong> <strong>к</strong><strong>о</strong>лб<strong>о</strong><strong>в</strong>идный, п<strong>о</strong>луш<strong>а</strong>р<strong>о</strong><strong>в</strong>идный,<br />

реже <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. У бе<strong>с</strong><strong>к</strong>рылых де<strong>в</strong><strong>с</strong>т<strong>в</strong>енниц<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т переднегруди,<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> мн<strong>о</strong>г<strong>о</strong>ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чны. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong><br />

ли<strong>с</strong>т<strong>в</strong>енных, <strong>о</strong>бычн<strong>о</strong> п<strong>о</strong>ряд<strong>к</strong><strong>а</strong> Fagales, реже<br />

н<strong>а</strong> тр<strong>а</strong><strong>в</strong><strong>а</strong>х. Не мигрируют. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2—4 мм (ри<strong>с</strong>. 575). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В<strong>с</strong>ег<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 100 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

ПОДСЕМЕЙСТВО APHIDINAE KOCH, 1857<br />

[nom. transl. Mordvilko, 1896<br />

(ex Aphiden Koch, 1857)]<br />

(Siphonophorini Ashmead, 1881)<br />

В переднем <strong>к</strong>рыле M <strong>с</strong> д<strong>в</strong>умя-тремя <strong>в</strong>ет<strong>в</strong>ями.<br />

Пять-шееть члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн. Гл<strong>а</strong>з<strong>а</strong><br />

у бе<strong>с</strong><strong>к</strong>рылых де<strong>в</strong><strong>с</strong>т<strong>в</strong>енниц мн<strong>о</strong>г<strong>о</strong>ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные.<br />

Ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong>ыделяют <strong>с</strong>ер<strong>о</strong>-бурую <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ую пыль.<br />

Х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>- или <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>треуг<strong>о</strong>льный,<br />

п<strong>а</strong>льце<strong>в</strong>идный, <strong>с</strong><strong>а</strong>бле<strong>в</strong>идный, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жен. С<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые труб<strong>о</strong>ч<strong>к</strong>и б<strong>о</strong>льшей<br />

ч<strong>а</strong><strong>с</strong>тью х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, реже <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их

буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>. Ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>й тергит брюш<strong>к</strong><strong>а</strong> не<strong>с</strong>ет<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т, н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющий х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong>. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2—5 мм (ри<strong>с</strong>. 576 <strong>а</strong>, б; 577).<br />

Мел — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>:<br />

<strong>о</strong>дин р<strong>о</strong>д Canadaphis Essig, 1937 из мел<strong>а</strong><br />

С. Амери<strong>к</strong>и (<strong>к</strong><strong>а</strong>н<strong>а</strong>д<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

APH1DOMORPHA INCERTAE SEDIS<br />

К н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я т<strong>а</strong><strong>к</strong>же Triassoaphis<br />

Evans, 1956 из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и<br />

Aphidulum Handlirsch, 1939 из юры 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ИНФРАОТРЯД<br />

COCCIDOMORPHA<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы. Д<strong>в</strong>е п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong>, з<strong>а</strong>дние<br />

зн<strong>а</strong>чительн<strong>о</strong> меньше или лишь передние<br />

<strong>к</strong>рылья, <strong>а</strong> з<strong>а</strong>дние пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> жужж<strong>а</strong>льцы;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, четыре-пять пр<strong>о</strong>д<strong>о</strong>льных<br />

жил<strong>о</strong><strong>к</strong> ин<strong>о</strong>гд<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м (R,<br />

RS,M, CuA и CuP). У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong><strong>а</strong>м<strong>к</strong>и и ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong><strong>а</strong>мцы бе<strong>с</strong><strong>к</strong>рылые. Антенны <strong>к</strong>рыл<strong>а</strong>тых ф<strong>о</strong>рм<br />

мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> у и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

ф<strong>о</strong>рм и у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>; бе<strong>с</strong><strong>к</strong>рылые<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и ин<strong>о</strong>гд<strong>а</strong> лишь <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми гл<strong>а</strong>з, у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

групп <strong>с</strong><strong>а</strong>мцы лишь <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми гл<strong>а</strong>з<strong>а</strong>ми.<br />

З<strong>а</strong>дние н<strong>о</strong>ги бег<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. Личин<strong>к</strong><strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>,<br />

реже <strong>с</strong>идяч<strong>а</strong>я. В. пермь — ныне. Д<strong>в</strong><strong>а</strong><br />

н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Protopsyllidiidea и Coccidea.<br />

НАДСЕМЕЙСТВО PROTOPSYLLIDIIDEA<br />

Д<strong>в</strong>е п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong>, з<strong>а</strong>дние зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че. В переднем <strong>к</strong>рыле R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я, М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> CuA, <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле неб<strong>о</strong>льш<strong>о</strong>е,<br />

упл<strong>о</strong>тненн<strong>о</strong>е, <strong>с</strong> я<strong>с</strong>ными <strong>а</strong>н<strong>а</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>тделен<strong>о</strong> прям<strong>о</strong>й CuP. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чными<br />

гл<strong>а</strong>з<strong>а</strong>ми и треуг<strong>о</strong>льным л<strong>о</strong>бным<br />

<strong>с</strong><strong>к</strong>лерит<strong>о</strong>м. Пермь — мез<strong>о</strong>з<strong>о</strong>й. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

— Protopsyllidiidae.<br />

СЕМЕЙСТВО PROTOPSYLLIDIIDAE<br />

CARPENTER, 1931<br />

(Permopsyllidae Tillyard, 1926)<br />

В н<strong>а</strong>д<strong>к</strong>рылье М и CuA <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong><strong>о</strong> <strong>с</strong>рединным ш<strong>в</strong><strong>о</strong>м. У нимфы г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>,<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, треуг<strong>о</strong>льный<br />

л<strong>о</strong>бный <strong>с</strong><strong>к</strong>лерит и <strong>с</strong>рединный ш<strong>о</strong><strong>в</strong>, пр<strong>о</strong>­<br />

д<strong>о</strong>лж<strong>а</strong>ющий<strong>с</strong>я <strong>в</strong> <strong>с</strong>рединный ш<strong>о</strong><strong>в</strong> <strong>с</strong>пин<strong>к</strong>и. П<strong>о</strong><br />

<strong>с</strong>рединн<strong>о</strong>му ш<strong>в</strong>у тре<strong>с</strong><strong>к</strong><strong>а</strong>л<strong>а</strong><strong>с</strong>ь ш<strong>к</strong>ур<strong>к</strong><strong>а</strong> при линь<strong>к</strong>е.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> без п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льных <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>,<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong>е; з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> меньше;<br />

брюш<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>с</strong>т<strong>о</strong>е (ри<strong>с</strong>. 578). В.<br />

пермь — юр<strong>а</strong>. 18 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Propatrix Becker-Migdisova, 1960.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. psylloides Becker-Migdisova,<br />

1960; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> прямым передним<br />

<strong>к</strong>р<strong>а</strong>ем, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> длинн<strong>а</strong>я; R делит<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья; R и RS п<strong>а</strong>р<strong>а</strong>ллельны<br />

переднему <strong>к</strong>р<strong>а</strong>ю; три <strong>в</strong>ет<strong>в</strong>и М; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

CuA шир<strong>о</strong><strong>к</strong>ий; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 6 мм (ри<strong>с</strong>. 579).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Cicadellopsis Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

С. incertaMartynov, 1937; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем, <strong>к</strong> <strong>в</strong>ершине <strong>с</strong>ужен<strong>о</strong>;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>о</strong>чень<br />

м<strong>а</strong>л<strong>а</strong>; R делит<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья; R<br />

и RS ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я; М делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

н<strong>а</strong>д<strong>к</strong>рылья н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA<br />

неб<strong>о</strong>льш<strong>о</strong>й. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5,8 мм<br />

(ри<strong>с</strong>. 580). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> — н. юр<strong>а</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

199<br />

http://jurassic.ru/<br />

Cicadopsyllidium Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong> — С. elongatum Becker-Migdisova,<br />

1959; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е и длинн<strong>о</strong>е,<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти не р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; жил<strong>к</strong>и<br />

пл<strong>о</strong>тные; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

R делит<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети н<strong>а</strong>д<strong>к</strong>рылья;<br />

R и RS п<strong>о</strong>чти прямые; М и CuA <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети; д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и М; CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> гщ-<strong>с</strong>и<strong>а</strong> длинн<strong>а</strong>я. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2,7 мм (ри<strong>с</strong>. 581)'. Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Asiopsyllidiutn Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong>—A. unicum Becker-Migdisova,<br />

1959, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Н<strong>а</strong>д<strong>к</strong>рылье переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е,<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; R <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

<strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

не<strong>в</strong>ели<strong>к</strong><strong>а</strong> или не<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; М делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

н<strong>а</strong>д<strong>к</strong>рылья; CuA <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не <strong>в</strong>ет<strong>в</strong>ления R; CuA делит<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее,<br />

чем М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

5 мм (ри<strong>с</strong>. 582). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Tomiopsyllidium Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong> — Т. iljinskiense Becker-Migdi-

Ри<strong>с</strong>. 578—584. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protopsyllidiidae<br />

578. Protopsyllidiidae sp.; нимф<strong>а</strong>, X 21,0, <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

(Evans, 1943). 579. Propatrix psylloides Becker-Migdisova; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у,X 9,1, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1960). 580. Cicadellopsls incerta Martynov; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 10,5, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 581. Cicadopsyllldlum<br />

elongatum Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,X 16,6,<br />

три<strong>а</strong><strong>с</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 582. Asiopsylli-<br />

sova, 1959; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Н<strong>а</strong>д<strong>к</strong>рылье пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>е,<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е; передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; R<br />

делит<strong>с</strong>я у гр<strong>а</strong>ницы <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й трети н<strong>а</strong>д<strong>к</strong>рылья;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> умеренн<strong>о</strong>й длины; R и RS из<strong>о</strong>гнутые,<br />

их ди<strong>с</strong>т<strong>а</strong>льные <strong>к</strong><strong>о</strong>нцы р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я; три <strong>в</strong>ет<strong>в</strong>и<br />

М; CuA <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее, чем М, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> ней н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении; CuA<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> CuP. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2 мм (ри<strong>с</strong>. 583). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>,<br />

Mesaleuropsis Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong>—М. venosa Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong> <strong>в</strong>иде уз<strong>к</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>ы; R идет <strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>я;<br />

RS <strong>о</strong>тх<strong>о</strong>дит <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>;<br />

М делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья; д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и;<br />

CuA пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я, <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я<br />

<strong>с</strong> М 3 + 4 п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> М. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong>д<strong>в</strong><strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че переднег<strong>о</strong>;<br />

д<strong>в</strong>е пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и М и CuA. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2,3 мм (ри<strong>с</strong>. 584). Один <strong>в</strong>ид.<br />

Н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из н. перми А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии—<br />

Protopsyllidium Tillyard, 1926; Permothea<br />

Tillyard, 1926; Permopsyllidium Tillyard,<br />

200<br />

http://jurassic.ru/<br />

dium unicum Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>, X10.4, <strong>в</strong>. три<strong>а</strong><strong>с</strong>.<br />

Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,» 1959). 583. Tomiopsyllidium iljinsklense<br />

Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>, X 28,5, <strong>в</strong>. Пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 584. Mesaleuropsis<br />

venosa Martynov; переднее и з<strong>а</strong>днее <strong>к</strong>рылья, X 28,0; н. юр<strong>а</strong><br />

Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, J937)<br />

1926; Clavopsyllidium Davis, 1942; Pertnotheella<br />

Davis, 1942; Permopsyllidops Davis, 1942;<br />

Psocoscytina Davis, 1942; Psocopsyllidium Davis,<br />

1942 (Psyllidiana Evans, 1943, Protopsyllops<br />

Evans, 1943); Belpsylla Evans, 1943; Eopsyllidium<br />

Davis, 1942; Psyllidella Evans, 1943;<br />

Permopsylloides Evans, 1943; Triassothea Evans,<br />

1956 из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

НАДСЕМЕЙСТВО COCCIDEA<br />

Ф<strong>о</strong>рмы <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным п<strong>о</strong>л<strong>о</strong><strong>в</strong>ым дим<strong>о</strong>рфизм<strong>о</strong>м.<br />

С<strong>а</strong>мцы <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й п<strong>а</strong>р<strong>о</strong>й <strong>к</strong>рылье<strong>в</strong>,<br />

реже бе<strong>с</strong><strong>к</strong>рылые; з<strong>а</strong>дние <strong>к</strong>рылья пре<strong>в</strong>р<strong>а</strong>щены<br />

<strong>в</strong> жужж<strong>а</strong>льц<strong>а</strong>; <strong>в</strong> переднем <strong>к</strong>рыле пять —три<br />

пр<strong>о</strong><strong>с</strong>тых жил<strong>о</strong><strong>к</strong>, R, М и CuA не <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть пр<strong>о</strong>зр<strong>а</strong>чн<strong>а</strong>я, <strong>о</strong>блегченн<strong>а</strong>я, не <strong>о</strong>тделен<strong>а</strong><br />

прям<strong>о</strong>й CuP; ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> имеют<strong>с</strong>я,<br />

<strong>а</strong>нтенны мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>ит; л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я, реже д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я,<br />

<strong>с</strong> <strong>о</strong>дним <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м. С<strong>а</strong>м<strong>к</strong>и бе<strong>с</strong><strong>к</strong>рылые;<br />

ин<strong>о</strong>гд<strong>а</strong> пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>идящие, <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями; тел<strong>о</strong> не имеет я<strong>с</strong>н<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>членения<br />

н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>у, грудь и брюш<strong>к</strong><strong>о</strong>; п<strong>о</strong><strong>к</strong>рыты<br />

<strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыделениями или <strong>о</strong>г<strong>о</strong>лены и д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>а</strong>я<br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью или ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> переднегрудью;<br />

<strong>а</strong>нтенны, гл<strong>а</strong>з<strong>а</strong> и р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти тел<strong>а</strong>. В <strong>с</strong><strong>о</strong><strong>в</strong>ре-

Ри<strong>с</strong>. 585—590. Ь<br />

585. Drostcha turkestanica Archangelskaja; <strong>с</strong><strong>а</strong>мец, X 18,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Б<strong>о</strong>рх<strong>с</strong>ениу<strong>с</strong>, 1950). 586. Orthezia insignis Douglas; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong> яйце<strong>в</strong>ым<br />

меш<strong>к</strong><strong>о</strong>м, X 7,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Б<strong>о</strong>рх<strong>с</strong>ениу<strong>с</strong>, 1950). 587. Coccus<br />

hesperldum Linnaeus; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>, х 10,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Б<strong>о</strong>рх<strong>с</strong>ениу<strong>с</strong>, 1950).<br />

588. Aspidiotus hederae (Vallot); щит<strong>к</strong>и <strong>с</strong><strong>а</strong>мц<strong>а</strong>, <strong>с</strong><strong>а</strong>м<strong>к</strong>и и личин<strong>о</strong><strong>к</strong><br />

менн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3000 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>с</strong><strong>а</strong>м<strong>а</strong>я <strong>с</strong>е<strong>в</strong>ерн<strong>а</strong>я и южн<strong>а</strong>я<br />

гр<strong>а</strong>ницы те же, чт<strong>о</strong> и у р<strong>а</strong><strong>с</strong>тений. Пермь —<br />

ныне. 17 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

Margarodidae, Ortheziidae, Coccidae, Diaspididae.<br />

201<br />

ц<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Coccidea<br />

1-г<strong>о</strong> и 2-г<strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>, X 15,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Б<strong>о</strong>рх<strong>с</strong>ениу<strong>с</strong>,<br />

1950). 589. Aspidiotus sp.; <strong>к</strong>рыл<strong>а</strong>тый <strong>с</strong><strong>а</strong>мец <strong>с</strong> длинным <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ным<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong>м, X 20,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1896). 590. Mesococcus<br />

asiaticus Becker-Migdisova; тел<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>, X 69,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>.<br />

Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959)<br />

СЕМЕЙСТВО MARGARODIDAE NEUSTEAD, 1901<br />

ГИГАНТСКИЕ ЧЕРВЕЦЫ<br />

[nom. transl. Enderlein, 1920<br />

(ex Margarodinae Neustead, 1901)]<br />

С<strong>а</strong>мцы <strong>о</strong>бычн<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>тые, передние <strong>к</strong>рылья<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong><strong>с</strong>ширены, <strong>с</strong> четырьмя-пятью<br />

http://jurassic.ru/

пр<strong>о</strong>д<strong>о</strong>льными, не<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тыми жил<strong>к</strong><strong>а</strong>ми (R, М,<br />

CuA, CuP); з<strong>а</strong>дние <strong>к</strong>рылья пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> жужж<strong>а</strong>льц<strong>а</strong>,<br />

ин<strong>о</strong>гд<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> и передние, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>л<strong>о</strong>жные и пр<strong>о</strong><strong>с</strong>тые, ред<strong>к</strong><strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>с</strong>тые;<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>тдельные члени<strong>к</strong>и н<strong>о</strong>г <strong>с</strong>литы;<br />

предп<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><br />

групп<strong>а</strong>ми или ряд<strong>а</strong>ми цилиндриче<strong>с</strong><strong>к</strong>их желез и<br />

<strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми нитями; <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. С<strong>а</strong>м<strong>к</strong>и п<strong>о</strong>д<strong>в</strong>ижны или пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>идячие;<br />

д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тел<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я,<br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я или <strong>с</strong> хитин<strong>о</strong><strong>в</strong>ыми пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми;<br />

<strong>с</strong>егмент<strong>а</strong>ция тел<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> я<strong>с</strong>н<strong>а</strong>я, ин<strong>о</strong>гд<strong>а</strong><br />

з<strong>а</strong>метны перетяж<strong>к</strong>и между тергит<strong>а</strong>ми груди;<br />

брюш<strong>к</strong><strong>о</strong> из <strong>в</strong><strong>о</strong><strong>с</strong>ьми я<strong>с</strong>ных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

д<strong>в</strong>ух-трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый или нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>ит, ин<strong>о</strong>гд<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; н<strong>о</strong>ги х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, ин<strong>о</strong>гд<strong>а</strong><br />

ч<strong>а</strong><strong>с</strong>ть члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я, реже н<strong>о</strong>ги редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; з<strong>а</strong>дний <strong>к</strong><strong>о</strong>нец тел<strong>а</strong> <strong>с</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й труб<strong>к</strong><strong>о</strong>й или т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> хитин<strong>о</strong><strong>в</strong>ым<br />

<strong>к</strong><strong>о</strong>льц<strong>о</strong>м без щитин<strong>о</strong><strong>к</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>, н<strong>а</strong>иб<strong>о</strong>лее б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>г<strong>о</strong><br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ими ф<strong>о</strong>рм<strong>а</strong>ми, <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех<br />

ч<strong>а</strong><strong>с</strong>тях <strong>с</strong><strong>в</strong>ет<strong>а</strong>, <strong>о</strong>бит<strong>а</strong>ют н<strong>а</strong> р<strong>а</strong>зличных р<strong>а</strong><strong>с</strong>тениях<br />

и их <strong>к</strong><strong>о</strong>рнях. Длин<strong>а</strong> тел<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и 1,5—3,5 мм<br />

(ри<strong>с</strong>. 585). В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 240 <strong>в</strong>ид<strong>о</strong><strong>в</strong>;<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ORTHEZIIDAE AMYOT ЕТ<br />

SERVILLE, 1843<br />

[nom. transl. Enderlein, 1920<br />

(ex Orthezides Amyot et Serville, 1843)]<br />

С<strong>а</strong>мец <strong>с</strong> длинными, уз<strong>к</strong>ими <strong>к</strong>рыльями; <strong>а</strong>нтенны<br />

де<strong>в</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; н<strong>о</strong>ги длинные; предп<strong>о</strong><strong>с</strong>ледний<br />

члени<strong>к</strong> брюш<strong>к</strong><strong>а</strong> <strong>с</strong> групп<strong>а</strong>ми цилиндриче<strong>с</strong><strong>к</strong>их<br />

желез и <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми нитями; <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т из<strong>о</strong>гнут <strong>к</strong>низу. Тел<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми, не<br />

<strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong> и п<strong>о</strong>чти пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е<br />

<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>; н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н;<br />

<strong>с</strong>егмент<strong>а</strong>ция тел<strong>а</strong> я<strong>с</strong>н<strong>а</strong>я; брюш<strong>к</strong><strong>о</strong> из<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьми члени<strong>к</strong><strong>о</strong><strong>в</strong>; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>о</strong>дн<strong>о</strong>- или д<strong>в</strong>ухчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый;<br />

н<strong>о</strong>ги длиннее тел<strong>а</strong>, <strong>в</strong>ертлуг и бедр<strong>о</strong><br />

<strong>с</strong>литы, ин<strong>о</strong>гд<strong>а</strong> т<strong>а</strong><strong>к</strong>же г<strong>о</strong>лень и л<strong>а</strong>п<strong>к</strong><strong>а</strong>; л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>о</strong><strong>к</strong>ружен<strong>о</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>им хитин<strong>о</strong><strong>в</strong>ым <strong>к</strong><strong>о</strong>льц<strong>о</strong>м <strong>с</strong> п<strong>о</strong>р<strong>а</strong>ми и<br />

ше<strong>с</strong>тью щетин<strong>к</strong><strong>а</strong>ми; яйц<strong>а</strong> <strong>в</strong> пл<strong>о</strong>тн<strong>о</strong>м <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>м<br />

меш<strong>к</strong>е. Жи<strong>в</strong>ут н<strong>а</strong> н<strong>а</strong>дземных ч<strong>а</strong><strong>с</strong>тях тр<strong>а</strong><strong>в</strong>яни<strong>с</strong>тых<br />

и <strong>к</strong>у<strong>с</strong>т<strong>а</strong>рни<strong>к</strong><strong>о</strong><strong>в</strong>ых, реже дре<strong>в</strong>е<strong>с</strong>ных р<strong>а</strong><strong>с</strong>тений.<br />

Длин<strong>а</strong> тел<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и 1,5—3,5 мм (ри<strong>с</strong>. 586).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

45 <strong>в</strong>ид<strong>о</strong><strong>в</strong> и пять шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

из них <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

202<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО COCCIDAE FALLEN, 1814.<br />

л<strong>о</strong>жн<strong>о</strong>щит<strong>о</strong><strong>в</strong><strong>к</strong>и<br />

[nom. transl. Stephens, 1829<br />

(ex decades Fallen, 1814)1<br />

(Lecaniidae Balachnovsky, 1915;<br />

Lecanococcidae Lindinger, 1889)<br />

С<strong>а</strong>мец <strong>к</strong>рыл<strong>а</strong>тый, жужж<strong>а</strong>льц<strong>а</strong> р<strong>а</strong>зличн<strong>о</strong>й<br />

ф<strong>о</strong>рмы; <strong>а</strong>нтенны де<strong>с</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; гл<strong>а</strong>з<strong>а</strong> пр<strong>о</strong><strong>с</strong>тые,<br />

<strong>в</strong> чи<strong>с</strong>ле д<strong>в</strong>ух — ше<strong>с</strong>ти п<strong>а</strong>р; н<strong>о</strong>ги длинные;<br />

<strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т р<strong>а</strong>зличн<strong>о</strong>й длины,<br />

<strong>к</strong> <strong>в</strong>ершине з<strong>а</strong><strong>о</strong><strong>с</strong>трен. Тел<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е<br />

или яйце<strong>в</strong>идн<strong>о</strong>е, д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е<br />

и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; <strong>с</strong>егмент<strong>а</strong>ция<br />

тел<strong>а</strong> п<strong>о</strong>чти п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

грудь и брюш<strong>к</strong><strong>о</strong> <strong>с</strong>литы; <strong>а</strong>нтенны ше<strong>с</strong>ти-<strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

ин<strong>о</strong>гд<strong>а</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны, гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ый;<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я; з<strong>а</strong>дний <strong>к</strong><strong>о</strong>нец<br />

р<strong>а</strong>зд<strong>в</strong><strong>о</strong>ен <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й щелью, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии ее <strong>с</strong><br />

д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>е<br />

н<strong>а</strong> <strong>в</strong>ершине <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й труб<strong>к</strong>и и <strong>о</strong><strong>к</strong>руженн<strong>о</strong>е<br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>им <strong>а</strong>н<strong>а</strong>льным <strong>к</strong><strong>о</strong>льц<strong>о</strong>м <strong>с</strong> п<strong>о</strong>р<strong>а</strong>ми<br />

и ше<strong>с</strong>тью — <strong>в</strong><strong>о</strong><strong>с</strong>емью щетин<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бычны<br />

т<strong>а</strong><strong>к</strong>же <strong>а</strong>н<strong>а</strong>льные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и. В<strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

р<strong>а</strong>зличных р<strong>а</strong><strong>с</strong>тениях и их <strong>к</strong><strong>о</strong>рнях, <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех ч<strong>а</strong><strong>с</strong>тях<br />

<strong>с</strong><strong>в</strong>ет<strong>а</strong> (ри<strong>с</strong>. 587). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне н<strong>а</strong><strong>с</strong>читы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я б<strong>о</strong>лее 800 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные<br />

р<strong>о</strong>ды, из них <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО DIASPIDIDAE SIGNORET, 1868.<br />

ЩИТОВКИ<br />

[nom. transl. Maskell, 1878<br />

(ex Diaspides Signoret, 1868)]<br />

С<strong>а</strong>мцы <strong>о</strong>бычн<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>тые; <strong>а</strong>нтенны ните<strong>в</strong>идные,<br />

де<strong>с</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; <strong>о</strong>т четырех д<strong>о</strong> ше<strong>с</strong>ти<br />

пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з; <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т уз<strong>к</strong>ий,<br />

р<strong>а</strong>зличн<strong>о</strong>й длины; брюш<strong>к</strong><strong>о</strong> без <strong>в</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых нитей.<br />

Тел<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>й р<strong>а</strong>зличн<strong>о</strong>й ф<strong>о</strong>рмы, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> щит<strong>к</strong><strong>о</strong>м<br />

из <strong>с</strong>бр<strong>о</strong>шенных ш<strong>к</strong>ур<strong>о</strong><strong>к</strong>, <strong>с</strong><strong>к</strong>репленных <strong>с</strong>е<strong>к</strong>рет<strong>о</strong>м;<br />

<strong>а</strong>нтенны редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны д<strong>о</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong> <strong>с</strong><br />

щетин<strong>к</strong><strong>а</strong>ми; гл<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ый; н<strong>о</strong>ги <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

<strong>с</strong>егмент<strong>а</strong>ция тел<strong>а</strong> я<strong>с</strong>н<strong>а</strong>я; брюш<strong>к</strong><strong>о</strong><br />

<strong>о</strong>бычн<strong>о</strong> делит<strong>с</strong>я н<strong>а</strong> переднюю <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нную<br />

ч<strong>а</strong><strong>с</strong>ть и з<strong>а</strong>днюю, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую из пигидия;<br />

IX, X и XI <strong>с</strong>егменты <strong>о</strong>бр<strong>а</strong>зуют н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пигидия уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong>, <strong>о</strong><strong>к</strong>руж<strong>а</strong>ющий <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие. Н<strong>а</strong> р<strong>а</strong>зличных дере<strong>в</strong>ьях и <strong>к</strong>у<strong>с</strong>т<strong>а</strong>рни<strong>к</strong><strong>а</strong>х,<br />

реже н<strong>а</strong> тр<strong>а</strong><strong>в</strong>яни<strong>с</strong>тых р<strong>а</strong><strong>с</strong>тениях.<br />

Длин<strong>а</strong> тел<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и 0,6—2 мм (ри<strong>с</strong>. 588, 589).<br />

Не<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong><strong>в</strong>ыше<br />

1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> и б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

<strong>о</strong>дин <strong>в</strong> не<strong>о</strong>гене 3. Е<strong>в</strong>р<strong>о</strong>пы.

COCCIDEA INCERTAE SEDIS<br />

Mesococcus Becker-Migdisova, 1959.<br />

Тип р<strong>о</strong>д<strong>а</strong> — M. asiaticus Becker-Migdisova,<br />

1959; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Бе<strong>с</strong><strong>к</strong>рыл<strong>а</strong>я ф<strong>о</strong>рм<strong>а</strong>, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>. Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>гнут<strong>а</strong> п<strong>о</strong>д<br />

грудь, нея<strong>с</strong>н<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует,<br />

гл<strong>а</strong>з<strong>а</strong> не р<strong>а</strong>з<strong>в</strong>иты. Переднегрудь б<strong>о</strong>льше,<br />

чем <strong>с</strong>редне- и з<strong>а</strong>днегрудь. Пле<strong>в</strong>р<strong>а</strong>льные <strong>с</strong><strong>к</strong>лериты<br />

груди б<strong>о</strong>льшие, нея<strong>с</strong>н<strong>о</strong> <strong>о</strong>тчлененные. Н<strong>о</strong>ги<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны д<strong>о</strong> незн<strong>а</strong>чительных прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>ближены. Брюш<strong>к</strong><strong>о</strong> из де<strong>в</strong>яти<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, де<strong>с</strong>ятый редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н <strong>в</strong> <strong>в</strong>иде неб<strong>о</strong>льш<strong>о</strong>й<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> 1—2 мм (ри<strong>с</strong>.590).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, четыре р<strong>о</strong>д<strong>а</strong> нея<strong>с</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>л<strong>о</strong>жения из мел<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы: С<strong>о</strong><strong>с</strong>codes<br />

Fritsch, 1901; Puccinites Velenowski, 1889;<br />

Variolina Fritsch, 1901; Pteridophyllites Renger,<br />

1866.<br />

ИНФРАОТРЯД PSYLLOMORPHA<br />

Передние и з<strong>а</strong>дние <strong>к</strong>рылья п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

длины <strong>с</strong><strong>о</strong> <strong>с</strong>х<strong>о</strong>дным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием. Передние<br />

<strong>к</strong>рылья <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>ны <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>рг<strong>а</strong>н п<strong>о</strong>лет<strong>а</strong>,<br />

<strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>-переп<strong>о</strong>нч<strong>а</strong>тые, ин<strong>о</strong>гд<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены<br />

и <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны или <strong>к</strong><strong>о</strong>жи<strong>с</strong>ты и п<strong>о</strong>чти<br />

г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мны. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я или <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я.<br />

Брюш<strong>к</strong><strong>о</strong> <strong>о</strong>тделен<strong>о</strong> <strong>о</strong>т груди перетяж<strong>к</strong><strong>о</strong>й.<br />

Среднегрудь х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. З<strong>а</strong>дние н<strong>о</strong>ги<br />

прыг<strong>а</strong>тельные; <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> яйце<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м. Личин<strong>к</strong>и рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм; т<strong>а</strong>зи<strong>к</strong>и у личин<strong>о</strong><strong>к</strong><br />

<strong>с</strong>ближены. Пермь •— ныне. Три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Aleurodidea, Cicadopsyllidea, Psyllidea.<br />

НАДСЕМЕЙСТВО ALEURODIDEA<br />

Крылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные; передние<br />

<strong>к</strong>рылья б<strong>о</strong>лее шир<strong>о</strong><strong>к</strong>ие, з<strong>а</strong>дние б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong><br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужены, не имеют <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong> для<br />

<strong>с</strong>цепления <strong>с</strong> передним <strong>к</strong>рыл<strong>о</strong>м; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>о</strong>чень бедн<strong>о</strong>е, <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>дн<strong>а</strong>-д<strong>в</strong>е пр<strong>о</strong>д<strong>о</strong>льных<br />

жил<strong>к</strong>и; R и М р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; СиА <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я<br />

ИЛИ <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; <strong>в</strong><strong>с</strong>е ЖИЛКИ <strong>с</strong>л<strong>а</strong>бые, не д<strong>о</strong>х<strong>о</strong>дят<br />

д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>рт<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я;<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong> переж<strong>а</strong>ты; д<strong>в</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>тых<br />

гл<strong>а</strong>з<strong>к</strong><strong>а</strong>; л<strong>о</strong>бный <strong>с</strong><strong>к</strong>лерит не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен;<br />

теменные <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ти и щечные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Пер<strong>в</strong>ый члени<strong>к</strong> брюш<strong>к</strong><strong>а</strong> <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>о</strong><strong>к</strong>руглый <strong>с</strong>тебеле<strong>к</strong>; ч<strong>а</strong><strong>с</strong>ть <strong>с</strong><strong>к</strong>лерит<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong><br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м. Т<strong>а</strong>зи<strong>к</strong>и з<strong>а</strong>дних<br />

н<strong>о</strong>г <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>и <strong>в</strong>ытянуты. Личин<strong>к</strong>и пр<strong>о</strong>х<strong>о</strong>дят<br />

<strong>с</strong>т<strong>а</strong>дию неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> пуп<strong>а</strong>рия. Длин<strong>а</strong> тел<strong>а</strong><br />

Ри<strong>с</strong>. 591—593. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Aleurodidea<br />

591. Permaleurodes rofundatus Becker-Migdisova; нимф?, X 16,1,<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 592. Aleurolobus<br />

olivlnus Silvestri; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, X 30,0 <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Silvestri, 1914). 593. Aleurodes sp.; А— переднее <strong>к</strong>рыл<strong>о</strong>, X 30,0;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 30,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch, 1925)<br />

203<br />

http://jurassic.ru/<br />

1—5 мм. В. пермь — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 200 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Permaleurodidae и Aleurodidae.<br />

СЕМЕЙСТВО PERMALEURODIDAE<br />

BECKER-MIGDISOVA, 1959<br />

Пуп<strong>а</strong>рий <strong>о</strong><strong>к</strong>руглый, длиннее ширины. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong>.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> п<strong>о</strong>луди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы;<br />

<strong>с</strong>редне-, з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> и <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ые з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и их<br />

<strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>в</strong>еличины; <strong>с</strong>рединные <strong>с</strong><strong>к</strong>лериты

передне-, <strong>с</strong>редне- и з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong>и <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены и<br />

р<strong>а</strong>зделены п<strong>о</strong>п<strong>о</strong>л<strong>а</strong>м <strong>с</strong>рединным ш<strong>в</strong><strong>о</strong>м. Брюш<strong>к</strong><strong>о</strong><br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>с</strong>т<strong>о</strong>е, <strong>с</strong> я<strong>с</strong>н<strong>о</strong>й <strong>с</strong>егмент<strong>а</strong>цией.<br />

В. пермь. Один р<strong>о</strong>д.<br />

Permateurodes Becker-Migdisova,<br />

1959. Тип р<strong>о</strong>д<strong>а</strong>—P. rotundatus Becker-Migdisova,<br />

1959; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Длин<strong>а</strong> передне<strong>с</strong>пин<strong>к</strong>и<br />

<strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> меньше ширины; <strong>с</strong>рединный <strong>с</strong><strong>к</strong>лерит<br />

не д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; у <strong>к</strong>р<strong>а</strong>я передне<strong>с</strong>пин<strong>к</strong>и<br />

д<strong>в</strong>е ям<strong>к</strong>и —<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние щетин<strong>о</strong><strong>к</strong>. Длин<strong>а</strong><br />

тел<strong>а</strong> 3 мм (ри<strong>с</strong>. 591; т<strong>а</strong>бл. XV, фиг. 9). Один<br />

<strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО ALEURODIDAE WESTWOOD, 1840<br />

Пуп<strong>а</strong>рий <strong>о</strong><strong>к</strong>руглый или <strong>о</strong><strong>к</strong>ругл<strong>о</strong> <strong>в</strong>ытянутый<br />

д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>. Средне- и з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>литы, или<br />

з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> уз<strong>к</strong><strong>а</strong>я; <strong>с</strong>рединные <strong>с</strong><strong>к</strong>лериты <strong>с</strong>пин<strong>к</strong>и<br />

не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены, <strong>с</strong>рединный ш<strong>о</strong><strong>в</strong> я<strong>с</strong>ный, <strong>с</strong>егмент<strong>а</strong>ция<br />

брюш<strong>к</strong><strong>а</strong> нея<strong>с</strong>н<strong>а</strong>я, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь лишь<br />

меди<strong>а</strong>льн<strong>о</strong>. Длин<strong>а</strong> тел<strong>а</strong> 1—3 мм (ри<strong>с</strong>. 592,<br />

593). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО CICADOPSYLLIDEA<br />

Гетер<strong>о</strong>н<strong>о</strong>мия <strong>к</strong>рылье<strong>в</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>.<br />

Передние <strong>к</strong>рылья <strong>о</strong><strong>в</strong><strong>а</strong>льные, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>ны<br />

<strong>к</strong> п<strong>о</strong>лету, н<strong>о</strong> <strong>с</strong> пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>-переп<strong>о</strong>нч<strong>а</strong>тые или реже <strong>с</strong>ильн<strong>о</strong><br />

у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены и <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими р<strong>а</strong><strong>с</strong>плы<strong>в</strong>ч<strong>а</strong>тыми<br />

жил<strong>к</strong><strong>а</strong>ми; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужены б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние их <strong>с</strong>х<strong>о</strong>дн<strong>о</strong> <strong>с</strong> передними, лишь CuA<br />

без б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> изгиб<strong>а</strong> <strong>к</strong> М; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

уже и передний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong> или <strong>о</strong>рт<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> не<br />

р<strong>а</strong>зделены; теменные <strong>в</strong><strong>о</strong>з<strong>в</strong>ышения неб<strong>о</strong>льшие;<br />

л<strong>о</strong>бные (щечные) <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

з<strong>а</strong>днег<strong>о</strong> <strong>о</strong>тдел<strong>а</strong> н<strong>а</strong>лични<strong>к</strong><strong>а</strong> не <strong>о</strong>тчленены,<br />

л<strong>о</strong>бный <strong>с</strong><strong>к</strong>лерит не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен. Т<strong>а</strong>зи<strong>к</strong>и з<strong>а</strong>дних<br />

н<strong>о</strong>г б<strong>о</strong>льшие, <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>и <strong>в</strong>ытянуты или меньших<br />

р<strong>а</strong>змер<strong>о</strong><strong>в</strong>, <strong>с</strong>лег<strong>к</strong><strong>а</strong> п<strong>о</strong>перечные. Н. пермь —<br />

ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Coleoscytidae и Cicadopsyllidae.<br />

СЕМЕЙСТВО COLEOSCYTIDAE<br />

MARTYNOV, 1935<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> и б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong><br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>нн<strong>о</strong>е; жил<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> лишь <strong>в</strong><br />

<strong>в</strong>иде теней или углублений; R b R 2 и RS п<strong>о</strong>чти<br />

р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине, р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong> р<strong>а</strong>зные <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны;<br />

<strong>в</strong>ет<strong>в</strong>и М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA шир<strong>о</strong><strong>к</strong>ий; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть CuA <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и п<strong>о</strong>л<strong>о</strong>г<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле длинн<strong>о</strong>е; Ai и А 2 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> длинный <strong>с</strong>тебель. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>ль­<br />

204<br />

http://jurassic.ru/<br />

н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е и р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е; <strong>в</strong>ет<strong>в</strong>и R и<br />

М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA шир<strong>о</strong><strong>к</strong>ий. З<strong>а</strong>дние<br />

н<strong>о</strong>ги прыг<strong>а</strong>тельные, т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>лег<strong>к</strong><strong>а</strong> п<strong>о</strong>перечные.<br />

Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Coleoscyta Martynov, 1935. Тип р<strong>о</strong>д<strong>а</strong> —<br />

С. rotundata Martynov, 1935; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, без <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

пер<strong>в</strong>ичн<strong>о</strong>й <strong>с</strong>ет<strong>к</strong>и (<strong>а</strong>рхеди<strong>к</strong>тия) и без <strong>с</strong><strong>к</strong>ульптуры;<br />

жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>бые, р<strong>а</strong><strong>с</strong>плы<strong>в</strong>ч<strong>а</strong>тые <strong>в</strong> <strong>в</strong>иде теней;<br />

<strong>в</strong>ет<strong>в</strong>и М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. В з<strong>а</strong>днем <strong>к</strong>рыле р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

CuA <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ий, ег<strong>о</strong> з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь <strong>в</strong> 2—2,5<br />

р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передней. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 7—<br />

7,5 мм (ри<strong>с</strong>. 594, 595). Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь<br />

Приур<strong>а</strong>лья и Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Kattanoscyta Becker-Migdisova, 1959.<br />

Тип р<strong>о</strong>д<strong>а</strong> — К. reticulata Becker-Migdisova,<br />

1959; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>леритиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>,<br />

п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> мел<strong>к</strong><strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>зующей<br />

<strong>с</strong><strong>к</strong>ульптуру, <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е; <strong>в</strong>ершин<strong>а</strong> <strong>с</strong>д<strong>в</strong>инут<strong>а</strong><br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; жил<strong>к</strong>и т<strong>о</strong>н<strong>к</strong>ие, я<strong>с</strong>ные;<br />

<strong>в</strong>ет<strong>в</strong>и М длинные. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5—6 мм<br />

(ри<strong>с</strong>. 596; т<strong>а</strong>бл. XV. фиг. 6). Один <strong>в</strong>ид.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Sojanopsylla Becker-Migdisova, 1960.<br />

Тип р<strong>о</strong>д<strong>а</strong> — 5. brevipennis Becker-Migdisova,<br />

I960; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е; передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й, б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> рез<strong>к</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>н; М и<br />

CuA делят<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, ди<strong>с</strong>т<strong>а</strong>льнее, чем<br />

R; <strong>в</strong>ет<strong>в</strong>ей М четыре, <strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я ч<strong>а</strong><strong>с</strong>ть длиннее<br />

ее <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, Mi делит<strong>с</strong>я; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA умеренн<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong>ий. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 11,5 мм (ри<strong>с</strong>. 597;<br />

т<strong>а</strong>бл. XV, фиг. 7). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО CICADOPSYLLIDAE<br />

MARTYNOV, 1931<br />

[nom. transl. Martynov, 1935<br />

(ex Cicadopsyllinae Martynov, 1931)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е или <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>-переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>е;<br />

R и М <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; CuA<br />

д<strong>а</strong>ет изгиб <strong>к</strong> М и <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> CuP.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong> п<strong>о</strong> длине переднему,<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ытянут<strong>о</strong>; R и RS п<strong>а</strong>р<strong>а</strong>ллельны, длинные<br />

и п<strong>о</strong>чти прямые; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA неб<strong>о</strong>льш<strong>о</strong>й,<br />

б<strong>а</strong>з<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>ден. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>рт<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я,<br />

н<strong>а</strong>лични<strong>к</strong> <strong>с</strong> не<strong>о</strong>тчлененными л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями;<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>с</strong> <strong>к</strong>рупными ред<strong>к</strong>ими ф<strong>а</strong><strong>с</strong>ет<strong>к</strong><strong>а</strong>ми;<br />

неб<strong>о</strong>льшие теменные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты, л<strong>о</strong>бные<br />

(щечные) <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. З<strong>а</strong>дние н<strong>о</strong>ги<br />

прыг<strong>а</strong>тельные, т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличены, <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ие,<br />

<strong>в</strong>ертлуг б<strong>о</strong>льш<strong>о</strong>й, бедр<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширены,<br />

л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Пермь. Четыре<br />

р<strong>о</strong>д<strong>а</strong>.

Ри<strong>с</strong>. 594—603. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Coli oscytidae[|H Cicadopsyllidae<br />

594. Coteoscyta occa/afo-Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 8,3,<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.» (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 595.<br />

С. elltrata Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 9,2, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, I960). 596. Kaltanoscyta reticulata<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 8,7, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959). 597. Sojanopsylla brevtpetinis<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 6,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1960). 598. Clcadopsylla permlana<br />

Martynov; н<strong>а</strong>д<strong>к</strong>рылье, X 6,0, <strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

http://jurassic.ru/<br />

1931). 599 Scytoneura eltlptica Martynov (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция),X 11,8<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й),<br />

600. S. vltrtpenna Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 13,0, н.пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 601. Scytoneura<br />

sp.; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 7,0. <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1960). 602. Scytoneurella major Q. Zalessky;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 8,0, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1939). 603.<br />

Ctcadopsls rugosipenna Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 9, 3,н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959)

Cicadopsylla Martynov, 1931. . Тип<br />

р<strong>о</strong>д<strong>а</strong> — С. permiana Martynov, 1931; <strong>в</strong>. пермь;<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие<br />

Г<strong>о</strong>ры). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е <strong>с</strong> <strong>с</strong>йльн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым<br />

передним <strong>к</strong>р<strong>а</strong>ем, мембр<strong>а</strong>н<strong>о</strong>зн<strong>о</strong>-<strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>'е; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е; R, Ri+г и R 2 <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>с</strong>пл<strong>о</strong>шную дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зную жил<strong>к</strong>у; М<br />

трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, делит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее, чем R, ее<br />

<strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я ч<strong>а</strong><strong>с</strong>ть длиннее <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; CuA делит<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> М, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> уз<strong>к</strong>ий.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 13,5 мм (ри<strong>с</strong>. 598).<br />

Один <strong>в</strong>ид. В;, пермь Приур<strong>а</strong>лья.<br />

Scytoneura Martynov, 1935. Тип р<strong>о</strong>д<strong>а</strong> —<br />

5. elliptica Martynov, 1935; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е, <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>-мембр<strong>а</strong>н<strong>о</strong>зн<strong>о</strong>е;<br />

передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; <strong>в</strong>ет<strong>в</strong>ей М три, <strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть р<strong>а</strong><strong>в</strong>н<strong>а</strong> или <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

М, делит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее CuA; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA уз<strong>к</strong>ий,<br />

из<strong>о</strong>гнутый. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> уже переднег<strong>о</strong>,<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; передний <strong>к</strong>р<strong>а</strong>й<br />

прям<strong>о</strong>й, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA шир<strong>о</strong><strong>к</strong>ий. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 4,6—8,5 мм (ри<strong>с</strong>. 599—601). Три<br />

<strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и <strong>в</strong>. пермь<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Scytoneurella G. Zalessky, 1939. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. major G. Zalessky, 1939; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Н<strong>а</strong>д<strong>к</strong>рылье мембр<strong>а</strong>н<strong>о</strong>зн<strong>о</strong>е, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> эллиптиче<strong>с</strong><strong>к</strong>и-<strong>о</strong><strong>к</strong>ругл<strong>о</strong>е;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>о</strong>,<br />

передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong> <strong>с</strong>ередине <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

М <strong>с</strong> тремя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями, делит<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее, чем CuA; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA шир<strong>о</strong><strong>к</strong>ий.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 4—6 мм (ри<strong>с</strong>. 602).<br />

Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Cicadopsis Becker-Migdisova, 1959.<br />

Тип р<strong>о</strong>д<strong>а</strong> — С. rugosipenna Becker-Migdisova,<br />

1959; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье мембр<strong>а</strong>н<strong>о</strong>зн<strong>о</strong>е,<br />

<strong>с</strong> м<strong>о</strong>рщин<strong>к</strong><strong>а</strong>ми; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ытянут<strong>а</strong>,<br />

<strong>в</strong>ершин<strong>а</strong> <strong>с</strong>д<strong>в</strong>инут<strong>а</strong> <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; Ri+г и<br />

R 2 — прям<strong>о</strong>е пр<strong>о</strong>д<strong>о</strong>лжение R; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М р<strong>а</strong><strong>в</strong>ен<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й ее ч<strong>а</strong><strong>с</strong>ти, делит<strong>с</strong>я н<strong>а</strong><br />

<strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> CuA; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA неб<strong>о</strong>льш<strong>о</strong>й;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>е, <strong>с</strong>ильн<strong>о</strong> упл<strong>о</strong>тненн<strong>о</strong>е,<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют ребри<strong>с</strong>тые <strong>в</strong><strong>о</strong>з<strong>в</strong>ышения.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 7,3 мм (ри<strong>с</strong>. 603;<br />

т<strong>а</strong>бл. XV, фиг. 8). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

НАДСЕМЕЙСТВО PSYLLIDEA<br />

Гетер<strong>о</strong>н<strong>о</strong>мия <strong>к</strong>рылье<strong>в</strong> я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>. Передние<br />

<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые или <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>-переп<strong>о</strong>нч<strong>а</strong>тые,<br />

шир<strong>о</strong><strong>к</strong>ие, <strong>о</strong><strong>в</strong><strong>а</strong>льные, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми,<br />

пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ытянуты, <strong>с</strong> прямым пе­<br />

206<br />

http://jurassic.ru/<br />

редним <strong>к</strong>р<strong>а</strong>ем и <strong>с</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong>м для <strong>с</strong>цепления<br />

<strong>с</strong> передним <strong>к</strong>рыл<strong>о</strong>м. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>; л<strong>о</strong>бный<br />

<strong>с</strong><strong>к</strong>лерит <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен; теменные <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ти<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, л<strong>о</strong>бные (щечные) <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>иты; три пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>а</strong>; <strong>а</strong>нтенны<br />

длинные, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Пер<strong>в</strong>ый члени<strong>к</strong><br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> з<strong>а</strong>днегрудью, <strong>о</strong>бр<strong>а</strong>зуя<br />

пр<strong>о</strong>межут<strong>о</strong>чный <strong>с</strong>егмент; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> <strong>о</strong>бр<strong>а</strong>зует<br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>ий <strong>с</strong>тебеле<strong>к</strong>. Т<strong>а</strong>зи<strong>к</strong>и з<strong>а</strong>дних н<strong>о</strong>г п<strong>о</strong>перечные.<br />

В<strong>с</strong>е личин<strong>о</strong>чные <strong>с</strong>т<strong>а</strong>дии п<strong>о</strong>д<strong>в</strong>ижны.<br />

Юр<strong>а</strong> — ныне. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Psyllidae.<br />

СЕМЕЙСТВО PSYLLIDAE LATREILLE, 1807<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, реже <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые;<br />

жил<strong>к</strong>и <strong>о</strong>чень пл<strong>о</strong>тные, <strong>в</strong>ыпу<strong>к</strong>лые; имеет<strong>с</strong>я <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>; R, М и CuA ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong> д<strong>в</strong>умя<br />

<strong>в</strong>ет<strong>в</strong>ями <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>в</strong> <strong>о</strong>дин <strong>с</strong>т<strong>в</strong><strong>о</strong>л.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че и <strong>о</strong>бычн<strong>о</strong> уже переднег<strong>о</strong>,<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>; R и М пр<strong>о</strong><strong>с</strong>тые;<br />

CuA <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, <strong>в</strong><strong>с</strong>е три жил<strong>к</strong>и <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. Антенны де<strong>в</strong>яти-де<strong>с</strong>ятичле<strong>в</strong>и<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Л<strong>а</strong>п<strong>к</strong><strong>а</strong> д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>й. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тях, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> теплых и умеренных.<br />

Обычн<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны <strong>с</strong> <strong>о</strong>пределенными<br />

р<strong>а</strong><strong>с</strong>тениями-х<strong>о</strong>зяе<strong>в</strong><strong>а</strong>ми. Юр<strong>а</strong> — ныне. В<strong>о</strong><strong>с</strong>емь<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Aphalarinae, Psyllinae, Triozinae,<br />

Liviinae, Pauropsyllinae, Ciriacreminae, Spondyliaspinae,<br />

Carsidarinae; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

три п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>.<br />

ПОДСЕМЕЙСТВО APHALARINAE LOW, 1879<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые или <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые,<br />

р<strong>а</strong>зличн<strong>о</strong>й ф<strong>о</strong>рмы; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> м<strong>о</strong>жет <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ть.<br />

Темя пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>нн<strong>о</strong>,<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет <strong>в</strong>переди <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>а</strong>нтенны; л<strong>о</strong>бный<br />

<strong>с</strong><strong>к</strong>лерит уз<strong>к</strong>ий, длинный, не з<strong>а</strong><strong>к</strong>рыт <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми;<br />

<strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ие л<strong>о</strong>бные (щечные) <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

неп<strong>а</strong>рный гл<strong>а</strong>з<strong>о</strong><strong>к</strong> <strong>в</strong>иден <strong>с</strong><strong>в</strong>ерху; три<br />

пер<strong>в</strong>ых члени<strong>к</strong><strong>а</strong> <strong>а</strong>нтенн удлинены. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

н<strong>а</strong><strong>к</strong>л<strong>о</strong>нн<strong>а</strong>я, не длинее темени; <strong>а</strong><strong>к</strong><strong>с</strong>иллярные<br />

<strong>с</strong><strong>к</strong>лериты имеют<strong>с</strong>я; tegulae <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Длин<strong>а</strong> тел<strong>а</strong> 1—4 мм. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы,<br />

<strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле <strong>о</strong>дин из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.<br />

ПОДСЕМЕЙСТВО LIADOPSYLLINAE<br />

MARTYNOV, 1926<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

неб<strong>о</strong>льш<strong>о</strong>е; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я и длинн<strong>а</strong>я;<br />

Ri, R 2 и RS <strong>о</strong>тх<strong>о</strong>дят <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е. Юр<strong>а</strong>. Один<br />

р<strong>о</strong>д.<br />

Liadopsylla Handlirsch, 1925. Тип<br />

р<strong>о</strong>д<strong>а</strong> — L. geinitzi Handlirsch, 1925; юр<strong>а</strong>, Герм<strong>а</strong>ния.<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>че переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>. Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее ширины;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>в</strong> 7 р<strong>а</strong>з длиннее ширины; р<strong>а</strong>ди<strong>а</strong>ль-

н<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> прям<strong>а</strong>я; R 2 и RS длинные, прямые,<br />

п<strong>а</strong>р<strong>а</strong>ллельные; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л R <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М длинный; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА шир<strong>о</strong><strong>к</strong>ий,<br />

передняя ег<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ь лишь <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,5—2 мм (ри<strong>с</strong>. 605).<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Юр<strong>а</strong> Ср. Азии и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ПОДСЕМЕЙСТВО PSYLLINAE LOW, 1879<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> д<strong>в</strong>умя м<strong>а</strong>ргин<strong>а</strong>льными ячей<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>зуемыми р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми М и CuA; RS<br />

не <strong>с</strong><strong>о</strong>единен<strong>а</strong> п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й <strong>с</strong> М, п<strong>о</strong><strong>с</strong>ледняя<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong> СиА <strong>в</strong> <strong>о</strong>бщий <strong>с</strong>тебеле<strong>к</strong>,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рый <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>тебля R; R 2 длинн<strong>а</strong>я; н<strong>а</strong>д-<br />

v<br />

Ри<strong>с</strong>. 604—607. Семей<strong>с</strong>тЕ<strong>о</strong> Psyllidae<br />

604. Agonoscena marmorea Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 21,0,<br />

ми<strong>о</strong>цен, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962). 605. Liadopsylla<br />

tenuicornts Martynov; н<strong>а</strong>д<strong>к</strong>рылье, X 17,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 606. Psylla nigrlta Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 28,0, ми<strong>о</strong>цен, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962).<br />

607. Trioza karaganlca Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье и г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

X 15,0, ми<strong>о</strong>цен, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962)<br />

<strong>к</strong>рылья <strong>с</strong> <strong>а</strong><strong>к</strong><strong>с</strong>иллярными <strong>с</strong><strong>к</strong>лерит<strong>а</strong>ми <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

и <strong>с</strong> tegulae. Л<strong>о</strong>бный <strong>с</strong><strong>к</strong>лерит з<strong>а</strong><strong>к</strong>рыт <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ими<br />

л<strong>о</strong>бными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми, неп<strong>а</strong>рный гл<strong>а</strong>з<strong>о</strong><strong>к</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен н<strong>а</strong> <strong>с</strong><strong>о</strong>единении лб<strong>а</strong> и ще<strong>к</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 2—6 мм (ри<strong>с</strong>. 604—606). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные<br />

р<strong>о</strong>ды, из них Psylla Goeffroy,<br />

207<br />

1762 <strong>с</strong> б<strong>о</strong>льшим <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>в</strong>ид<strong>о</strong><strong>в</strong> и Agonoscena<br />

Enderleiw, из ми<strong>о</strong>цен<strong>а</strong> С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>, <strong>о</strong>дин<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и,<br />

<strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО TRIOZINAE LOW, 1872<br />

Н<strong>а</strong>д<strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, у <strong>в</strong>ершины угл<strong>о</strong><strong>в</strong><strong>а</strong>тые;<br />

R, М и СиА р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е;<br />

<strong>о</strong>бщий <strong>с</strong>тебель М и СиА <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; Ri+ 2<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Л<strong>о</strong>б <strong>с</strong><strong>к</strong>рыт <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ими л<strong>о</strong>бными<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми; неп<strong>а</strong>рный гл<strong>а</strong>з<strong>о</strong><strong>к</strong> н<strong>а</strong> гр<strong>а</strong>нице<br />

<strong>с</strong><strong>о</strong>единения лб<strong>а</strong> и ще<strong>к</strong>. Длин<strong>а</strong> тел<strong>а</strong> 1,5—4 мм<br />

(ри<strong>с</strong>. 607). Не<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех <strong>о</strong>бл<strong>а</strong><strong>с</strong>тях, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле р<strong>о</strong>д<br />

Trioza Forster, 1848 из ми<strong>о</strong>цен<strong>а</strong> С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>.<br />

STENORRHYNCHA INCERTAE SEDIS<br />

Вне СССР: р<strong>о</strong>д Tripsyllidium Evans, 1956<br />

из <strong>в</strong>. три<strong>а</strong><strong>с</strong><strong>а</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

ПОДОТРЯД COLEORRHYNCHA<br />

Крылья гетер<strong>о</strong>н<strong>о</strong>мны. Н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>ты'е,<br />

без т<strong>о</strong>чечн<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптуры, <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

<strong>с</strong>пин<strong>к</strong>е пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>, <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я п<strong>о</strong> прям<strong>о</strong>й линии или<br />

н<strong>а</strong>лег<strong>а</strong>ют <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>дн<strong>о</strong> н<strong>а</strong> друг<strong>о</strong>е,<br />

<strong>в</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>луч<strong>а</strong>е ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>бычн<strong>о</strong><br />

рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>, <strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>с</strong>д<strong>в</strong>инут. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я,<br />

gula <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, з<strong>а</strong>дний <strong>о</strong>тдел н<strong>а</strong>лични<strong>к</strong><strong>а</strong> <strong>с</strong><br />

не<strong>о</strong>тчлененными л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 608. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Peloridiidae<br />

Hemiodoecus veitchi Hacker; тел<strong>о</strong> и <strong>к</strong>рылья н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>,<br />

X 22, 0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Т<strong>а</strong><strong>с</strong>м<strong>а</strong>ния (Hacker, 1932)

Ан<strong>а</strong>льн<strong>а</strong>я труб<strong>к</strong><strong>а</strong> у <strong>с</strong><strong>а</strong>мц<strong>а</strong> и <strong>с</strong><strong>а</strong>м<strong>к</strong>и цилиндриче<strong>с</strong><strong>к</strong><strong>а</strong>я;<br />

XI <strong>с</strong>егмент <strong>в</strong>ытянут, <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong>и яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong><br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

тремя рели<strong>к</strong>т<strong>о</strong><strong>в</strong>ыми р<strong>о</strong>д<strong>а</strong>ми и <strong>в</strong><strong>о</strong><strong>с</strong>емью<br />

<strong>в</strong>ид<strong>а</strong>ми н<strong>а</strong> <strong>к</strong>р<strong>а</strong>йнем юге или н<strong>а</strong> зн<strong>а</strong>чительных<br />

<strong>в</strong>ы<strong>с</strong><strong>о</strong>т<strong>а</strong>х (2000—3000 м) умеренн<strong>о</strong>й з<strong>о</strong>ны<br />

Ю. Амери<strong>к</strong>и, Ю. А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии, Н<strong>о</strong><strong>в</strong><strong>о</strong>й Зел<strong>а</strong>ндии,<br />

Т<strong>а</strong><strong>с</strong>м<strong>а</strong>нии и прилег<strong>а</strong>ющих <strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong>. От Coleorrhyncha,<br />

<strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>, пр<strong>о</strong>из<strong>о</strong>шли пер<strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

<strong>о</strong>тряд<strong>а</strong> Heteroptera. Одн<strong>о</strong> рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

Actinoscytinidae (Cicadocoridae Becker-<br />

Migdisova) Э<strong>в</strong><strong>а</strong>н<strong>с</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит <strong>к</strong> Heteroptera.<br />

СЕМЕЙСТВО PELORIDIIDAE BREDDIN, 1897<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>с</strong>пин<strong>к</strong>е пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>,<br />

<strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я п<strong>о</strong> прям<strong>о</strong>й линии или н<strong>а</strong>х<strong>о</strong>дят<br />

СЕМЕЙСТВО PROSBOLOPSEIDAE<br />

BECKER-MIGDISOVA, 1946<br />

HOMOPTERA INCERTAE SEDIS<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны или <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; передний<br />

<strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ыпу<strong>к</strong>лый; R 2 п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong><br />

RS, д<strong>а</strong>ет ряд <strong>в</strong>ет<strong>в</strong>ей; Ai и А 2 <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>с</strong>тебеле<strong>к</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong> или п<strong>о</strong>луп<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>;<br />

•х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> длинный, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния з<strong>а</strong>дних<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong>; з<strong>а</strong>дний <strong>о</strong>тдел н<strong>а</strong>лични<strong>к</strong><strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>й, <strong>с</strong><br />

не<strong>о</strong>тчленными л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и шир<strong>о</strong><strong>к</strong><strong>а</strong>я; щит<strong>о</strong><strong>к</strong> р<strong>а</strong><strong>в</strong>ен ей п<strong>о</strong> ширине,<br />

треуг<strong>о</strong>льный. Брюш<strong>к</strong><strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

труб<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Пермь — три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

<strong>о</strong>дн<strong>о</strong> н<strong>а</strong> друг<strong>о</strong>е, <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, <strong>с</strong> т<strong>о</strong>л<strong>с</strong>тыми жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>зующими непр<strong>а</strong><strong>в</strong>ильные, з<strong>а</strong>м<strong>к</strong>нутые<br />

ячей<strong>к</strong>и п<strong>о</strong> <strong>в</strong><strong>с</strong>ей п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>с</strong> ряд<strong>о</strong>м б<strong>о</strong>лее мел<strong>к</strong>их ячее<strong>к</strong>; передний <strong>к</strong>р<strong>а</strong>й<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ыпу<strong>к</strong>лый, з<strong>а</strong>дний — прям<strong>о</strong>й. Х<strong>о</strong>д<br />

жил<strong>о</strong><strong>к</strong> М и CuA нея<strong>с</strong>ен; М ме<strong>с</strong>т<strong>а</strong>ми <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> R и CuA, a CuA — <strong>с</strong> М и CuP. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

б<strong>о</strong>льш<strong>а</strong>я, треуг<strong>о</strong>льн<strong>а</strong>я, п<strong>о</strong>луп<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я.<br />

У <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых л<strong>о</strong>бн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м,<br />

у нимф <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я л<strong>о</strong>бн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong><br />

и не<strong>о</strong>тчлененные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти н<strong>а</strong>лични<strong>к</strong><strong>а</strong>. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми,<br />

щит<strong>о</strong><strong>к</strong> неб<strong>о</strong>льш<strong>о</strong>й. Ан<strong>а</strong>льн<strong>а</strong>я труб<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я.<br />

Длин<strong>а</strong> тел<strong>а</strong> 4—5 мм (ри<strong>с</strong>. 608). В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне четыре р<strong>о</strong>д<strong>а</strong>, р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные н<strong>а</strong> юге<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии, Ю. Амери<strong>к</strong>и, <strong>в</strong> Н<strong>о</strong><strong>в</strong><strong>о</strong>й Зел<strong>а</strong>ндии и<br />

Т<strong>а</strong><strong>с</strong>м<strong>а</strong>нии.<br />

Prosbotopsis Martynov 1935. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. ovalis Martynov, 1935; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем, <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым<br />

<strong>о</strong><strong>к</strong>руглым передним <strong>к</strong>р<strong>а</strong>ем, <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong><br />

прямым з<strong>а</strong>дним и <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>а</strong><strong>с</strong>имметричн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й;<br />

R + М длинн<strong>а</strong>я; R, М и CuA делят<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья;<br />

М трех-пяти<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

8,5—9 мм (ри<strong>с</strong>. 609—611). Один <strong>в</strong>ид. В. пермь<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong> <strong>к</strong> Homoptera incertae sedis <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я<br />

Eochiliocycla Davis, 1942 (<strong>в</strong>. пермь А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии)<br />

.<br />

ОТРЯД HETEROPTERA. РАЗНОКРЫЛЫЕ, ИЛИ КЛОПЫ<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>, пре<strong>в</strong>р<strong>а</strong>щены<br />

<strong>в</strong> гемиэлитры и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, р<strong>а</strong>зделены<br />

ш<strong>в</strong><strong>а</strong>ми и н<strong>а</strong>дл<strong>о</strong>м<strong>а</strong>ми н<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> я<strong>с</strong>н<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных<br />

ч<strong>а</strong><strong>с</strong>тей: переп<strong>о</strong>нч<strong>а</strong>тую — мембр<strong>а</strong>ну и<br />

<strong>к</strong><strong>о</strong>жи<strong>с</strong>тые — <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> и <strong>к</strong><strong>о</strong>риум; ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> <strong>к</strong><strong>о</strong>риуме<br />

<strong>в</strong>ыделяют<strong>с</strong>я <strong>к</strong>унеу<strong>с</strong>, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующий птер<strong>о</strong><strong>с</strong>тигме<br />

других групп, и эмб<strong>о</strong>лиум, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующий<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му п<strong>о</strong>лю; у немн<strong>о</strong>гих<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья цельные,<br />

неп<strong>о</strong>др<strong>а</strong>зделенные; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние н<strong>а</strong> упл<strong>о</strong>тненных<br />

ч<strong>а</strong><strong>с</strong>тях <strong>к</strong>рыл<strong>а</strong> нея<strong>с</strong>н<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>,<br />

п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>', <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>.<br />

Средне<strong>с</strong>пин<strong>к</strong><strong>а</strong> б<strong>о</strong>льше з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong>и и пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong><br />

нею <strong>с</strong>р<strong>а</strong>щен<strong>а</strong>; щит<strong>о</strong><strong>к</strong> <strong>о</strong>бычн<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит, у<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм н<strong>а</strong> груди и у личин<strong>о</strong><strong>к</strong> н<strong>а</strong> тер­<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

208<br />

http://jurassic.ru/<br />

гит<strong>а</strong>х брюш<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>а</strong>хучих желез. Мн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>й и прибрежн<strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>тью<br />

и д<strong>а</strong>же <strong>в</strong>едут пел<strong>а</strong>гиче<strong>с</strong><strong>к</strong>ий<br />

<strong>о</strong>бр<strong>а</strong>з жизни. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные, хищни<strong>к</strong>и, н<strong>а</strong>ружные<br />

п<strong>а</strong>р<strong>а</strong>зиты п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных, <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги и<br />

ф<strong>о</strong>рмы <strong>с</strong> <strong>к</strong><strong>о</strong>мбинир<strong>о</strong><strong>в</strong><strong>а</strong>нным пит<strong>а</strong>нием. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>и<br />

<strong>с</strong><strong>в</strong>яз<strong>а</strong>ны <strong>с</strong> примити<strong>в</strong>ными р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылыми,<br />

<strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>тделили<strong>с</strong>ь, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>в</strong><br />

перми. Об<strong>о</strong><strong>с</strong><strong>о</strong>бление и ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ние р<strong>а</strong>зн<strong>о</strong><strong>к</strong>рылых<br />

<strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>или<strong>с</strong>ь н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е перех<strong>о</strong>д<strong>а</strong> <strong>о</strong>т<br />

р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядн<strong>о</strong><strong>с</strong>ти <strong>к</strong> <strong>к</strong><strong>о</strong>мбинир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>му пит<strong>а</strong>нию<br />

и хищниче<strong>с</strong>т<strong>в</strong>у и <strong>с</strong><strong>в</strong>яз<strong>а</strong>нн<strong>о</strong>й <strong>с</strong> этим б<strong>о</strong>льш<strong>о</strong>й<br />

п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><strong>с</strong>ти ф<strong>о</strong>рм. Эт<strong>о</strong> <strong>с</strong><strong>о</strong>дей<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>л<strong>о</strong><br />

б<strong>о</strong>льшему р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зию и шир<strong>о</strong>те з<strong>а</strong>ним<strong>а</strong>емых<br />

би<strong>о</strong>т<strong>о</strong>п<strong>о</strong><strong>в</strong> и <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> тем б<strong>о</strong>льшему р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нению<br />

п<strong>о</strong> Земле. Пермь — ныне.

Ри<strong>с</strong>. 609—611. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Prosbolopseidae<br />

609. Prosbolopsis ovalis Martynov; <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong> тел<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>- X 6,3, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 611.<br />

р<strong>о</strong>ны, х 4,7, <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, Prosbolopsis sp.; <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong> тел<strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны.Х 6,3, <strong>в</strong>.<br />

1960). 610. Prosbolopsis sp.; <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong> тел<strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935)<br />

С<strong>о</strong><strong>в</strong>ременные р<strong>а</strong>зн<strong>о</strong><strong>к</strong>рылые р<strong>а</strong><strong>с</strong>пределяют<strong>с</strong>я<br />

<strong>с</strong>реди д<strong>в</strong>ух п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> — н<strong>а</strong>земных <strong>к</strong>л<strong>о</strong>п<strong>о</strong><strong>в</strong>,<br />

Geocorisae, и <strong>в</strong><strong>о</strong>дных <strong>к</strong>л<strong>о</strong>п<strong>о</strong><strong>в</strong>, Hydrocorisae;<br />

<strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>а</strong>я х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong><strong>а</strong> этих<br />

групп <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> т<strong>а</strong><strong>к</strong>их <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тях<br />

<strong>с</strong>тр<strong>о</strong>ения тел<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong>бычн<strong>о</strong> не м<strong>о</strong>гут быть<br />

лег<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трены н<strong>а</strong> неп<strong>о</strong>лн<strong>о</strong>м и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

м<strong>а</strong>тери<strong>а</strong>ле. П<strong>о</strong>эт<strong>о</strong>му <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние р<strong>а</strong>зн<strong>о</strong><strong>к</strong>рылых н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

прям<strong>о</strong> <strong>с</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряды и н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

не р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ют<strong>с</strong>я. Отряд <strong>в</strong><strong>к</strong>люч<strong>а</strong>ет<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

и 16 <strong>в</strong>ымерших мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены 40 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

СЕМЕЙСТВО APOPNIDAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> <strong>о</strong>чень б<strong>о</strong>льшим <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м;<br />

<strong>к</strong><strong>о</strong>риум б<strong>о</strong>лее пл<strong>о</strong>тный, <strong>о</strong>тделен <strong>о</strong>т мембр<strong>а</strong>ны<br />

дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й линией, <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й н<strong>а</strong>ружу, <strong>с</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми четырех гл<strong>а</strong><strong>в</strong>ных жил<strong>о</strong><strong>к</strong>; <strong>в</strong> мембр<strong>а</strong>не<br />

жил<strong>к</strong>и нея<strong>с</strong>ны. Щит<strong>о</strong><strong>к</strong> б<strong>о</strong>льш<strong>о</strong>й. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4 мм (ри<strong>с</strong>. 612). В<strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>, р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны<br />

Notonectidae. Н. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Один<br />

р<strong>о</strong>д — Apopnus Handlirsch, 1939.<br />

СЕМЕЙСТВО APHLEBOCOR1DAE<br />

HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>им, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м, угл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим, <strong>с</strong> я<strong>с</strong>н<strong>о</strong>й<br />

Аь <strong>к</strong><strong>о</strong>риум и мембр<strong>а</strong>н<strong>а</strong> нерез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены,<br />

<strong>к</strong><strong>о</strong>риум б<strong>о</strong>льш<strong>о</strong>й, пл<strong>о</strong>тный, пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>нный,<br />

<strong>с</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями R и М; мембр<strong>а</strong>н<strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>в</strong><br />

<strong>в</strong>иде шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>ймы без жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

5 мм (ри<strong>с</strong>. 613). П<strong>о</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>теру н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong><br />

н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют Naucoridae, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

пред<strong>к</strong>и п<strong>о</strong><strong>с</strong>ледних. Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Один<br />

р<strong>о</strong>д — Aphlebocoris Handlirsch, 1906.<br />

14 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 209<br />

609<br />

^Ри<strong>с</strong>. 612—614. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Apopnidae, Aphlebocoridae,<br />

Probascanionidae<br />

612. Apopnus magnlclavus Handlirsch; н<strong>а</strong>д<strong>к</strong>рылье,X 13,7, н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния (Handlirsch, 1939). 613. Aphlebocoris punctata Handlirsch;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 11,0, н. юр<strong>а</strong>. Герм<strong>а</strong>ния (Handlirsch, 1939).<br />

614. Probascanion megacephalum Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид, X 9,6,<br />

н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1939)<br />

http://jurassic.ru/<br />

613

Ри<strong>с</strong>. 615—619. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archegocimicidae, Progonocimicidae,<br />

Eonabidae, Eocimicidae<br />

615. Archegocimex geinlizl Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид, xl0,2, н.<br />

юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1906). 616. Progonocoris pictus<br />

Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид, xl4,5, н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch,<br />

1939). 617.. Progonocimex jurasslcus Handlirsch; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X7,6, н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1906). 618. Eonabis prlmttiva<br />

Handlirsch; н<strong>а</strong>д<strong>к</strong>рылье, x 13,8, н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch,<br />

1939). 619. Eocimex liasinus Handlirsch; н<strong>а</strong>д<strong>к</strong>рылье, X 10,0,<br />

н. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1906)<br />

210<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PROBASCANIONIDAE<br />

HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>о</strong><strong>к</strong>руглые, <strong>с</strong><strong>о</strong> <strong>с</strong>д<strong>в</strong>инутым<br />

б<strong>о</strong>льшим и угл<strong>о</strong><strong>в</strong><strong>а</strong>тым <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

б<strong>о</strong>льш<strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>а</strong>я; гл<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льшие, <strong>в</strong>ыпу<strong>к</strong>лые.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, цилиндриче<strong>с</strong><strong>к</strong><strong>а</strong>я, <strong>в</strong><br />

2 р<strong>а</strong>з<strong>а</strong> шире длины. Щит<strong>о</strong><strong>к</strong> неб<strong>о</strong>льш<strong>о</strong>й, р<strong>а</strong><strong>в</strong>н<strong>о</strong>бедренный,<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренный, з<strong>а</strong>ним<strong>а</strong>ет ед<strong>в</strong><strong>а</strong><br />

б<strong>о</strong>лее трети длины тел<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong><br />

6 мм (ри<strong>с</strong>. 614). П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны<br />

Naucoridae. Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Рг<strong>о</strong>bascanion<br />

Handlirsch, 1939.<br />

СЕМЕЙСТВО ARCHEGOCIMICIDAE<br />

HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>й мембр<strong>а</strong>н<strong>о</strong>й, не <strong>о</strong>тделенн<strong>о</strong>й<br />

я<strong>с</strong>н<strong>о</strong>й делящей линией или р<strong>а</strong>зры<strong>в</strong><strong>о</strong>м<br />

<strong>о</strong>т <strong>к</strong><strong>о</strong>риум<strong>а</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й, или жил<strong>к</strong>и<br />

не д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; SC длинн<strong>а</strong>я; R ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

гребенч<strong>а</strong>т<strong>а</strong>я, трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

нея<strong>с</strong>н<strong>а</strong> или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong> тремя-четырьмя<br />

<strong>в</strong>ет<strong>в</strong>ями, делит<strong>с</strong>я н<strong>а</strong> гр<strong>а</strong>нице<br />

<strong>к</strong><strong>о</strong>риум<strong>а</strong>. Передняя <strong>в</strong>ет<strong>в</strong>ь CuA длинн<strong>а</strong>я, з<strong>а</strong>дняя<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong>.<br />

З<strong>а</strong>дний уг<strong>о</strong>л <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>а</strong> <strong>с</strong>д<strong>в</strong>инут <strong>в</strong> диет<strong>а</strong>льную ег<strong>о</strong><br />

треть. Тел<strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е, передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong><br />

шире длины; щит<strong>о</strong><strong>к</strong> б<strong>о</strong>льш<strong>о</strong>й, длин<strong>а</strong> ег<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

ширине. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong>идящ<strong>а</strong>я.<br />

Близ<strong>к</strong>и п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению <strong>к</strong> Actinoscytinidae ( = Cicadocoridae).<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4—6 мм<br />

(ри<strong>с</strong>. 615, 616). Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>:<br />

Archegocimex Handlirsch, 1906; Archegocoris<br />

Handlirsch, 1939; Anosmus Handlirsch, 1939;<br />

Deraiocoris Bode, 1953; Progonocoris Handlirsch,<br />

1939; Megacoris Bode, 1953; Ophthalmocoris<br />

Bode, 1953.<br />

СЕМЕЙСТВО PROGONOCIMICIDAE<br />

HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> нея<strong>с</strong>ными гр<strong>а</strong>ниц<strong>а</strong>ми мембр<strong>а</strong>ны<br />

и <strong>к</strong><strong>о</strong>риум<strong>а</strong>, без делящей линии; жил<strong>к</strong>и <strong>к</strong><strong>о</strong>риум<strong>а</strong><br />

нея<strong>с</strong>ны. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>переди <strong>в</strong>ырез<strong>а</strong>н<strong>а</strong>,<br />

<strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти<br />

прям<strong>о</strong>й. Щит<strong>о</strong><strong>к</strong> <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> шире длины, ф<strong>о</strong>рмы<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>бедренн<strong>о</strong>г<strong>о</strong> треуг<strong>о</strong>льни<strong>к</strong><strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> умеренн<strong>о</strong>й<br />

<strong>в</strong>еличины. Длин<strong>а</strong> тел<strong>а</strong> 7 мм (ри<strong>с</strong>.617).<br />

Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Progonocimex<br />

Handlirsch, 1906.<br />

СЕМЕЙСТВО EONABIDAE HANDLIRSCH, 1939<br />

В н<strong>а</strong>д<strong>к</strong>рыльях <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и мембр<strong>а</strong>н<strong>а</strong><br />

нерез<strong>к</strong><strong>о</strong> р<strong>а</strong>зделены, не р<strong>а</strong>злич<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong> <strong>с</strong><strong>к</strong>ульптуре;<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й; <strong>о</strong><strong>к</strong>руглый; SC х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, <strong>о</strong>тгр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет п<strong>о</strong>д<strong>о</strong>бие линейн<strong>о</strong>г<strong>о</strong><br />

эмб<strong>о</strong>лиум<strong>а</strong>; R и М <strong>с</strong>литы б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> д<strong>о</strong> <strong>с</strong>ереди-

ны н<strong>а</strong>д<strong>к</strong>рылья, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дны'; R <strong>в</strong> мембр<strong>а</strong>не<br />

д<strong>а</strong>ет ряд <strong>в</strong>ет<strong>в</strong>ей; СиА д<strong>а</strong>ет д<strong>в</strong>е длинные<br />

п<strong>а</strong>р<strong>а</strong>ллельные <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>;<br />

CuP прям<strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> з<strong>а</strong>днюю <strong>в</strong>ет<strong>в</strong>ь<br />

СиА, <strong>с</strong>луж<strong>а</strong>щую ег<strong>о</strong> пр<strong>о</strong>д<strong>о</strong>лжением, п<strong>о</strong><strong>с</strong>ледняя<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ую жил<strong>к</strong>у. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4,5 мм (ри<strong>с</strong>. 618). Н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Nabidae. Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>:<br />

Eonabis Handlirsch, 1939; Copidopus Handlirsch,<br />

1906; Pronabis Bode, 1953; Engynabis Bode,<br />

1953; Eogerridium Bode, 1953.<br />

СЕМЕЙСТВО EOCIMICIDAE HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong> р<strong>а</strong>зделены 1<br />

ребр<strong>а</strong>ми н<strong>а</strong><br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и мембр<strong>а</strong>ну; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й,<br />

п<strong>о</strong>лу<strong>к</strong>руглый; <strong>к</strong><strong>о</strong>риум <strong>о</strong>бр<strong>а</strong>зует р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>о</strong>нний<br />

треуг<strong>о</strong>льни<strong>к</strong>, <strong>с</strong> единичны'ми жил<strong>к</strong><strong>а</strong>ми R,<br />

М <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> СиА; СиА делит<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong> д<strong>в</strong>е длинные <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong> мембр<strong>а</strong>не<br />

<strong>в</strong><strong>с</strong>е жил<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде ребер и <strong>о</strong>бры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я у<br />

делящег<strong>о</strong> ребр<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,5 мм<br />

(ри<strong>с</strong>. 619). Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д—Eociтех<br />

Handlirsch, 1906.<br />

СЕМЕЙСТВО HADROCORIDAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья р<strong>а</strong>зделены н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум<br />

и мембр<strong>а</strong>ну; <strong>к</strong><strong>о</strong>риум <strong>с</strong> я<strong>с</strong>ными гл<strong>а</strong><strong>в</strong>ными жил­<br />

<strong>к</strong><strong>а</strong>ми SC, R, М и СиА; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> уз<strong>к</strong>ий, длинный,,<br />

угл<strong>о</strong><strong>в</strong><strong>а</strong>тый; мембр<strong>а</strong>н<strong>а</strong> <strong>о</strong>тделен<strong>а</strong> пл<strong>о</strong>тными п<strong>о</strong>перечными.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>, <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong>перед,<br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я. Гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

б<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong>лег<strong>к</strong><strong>а</strong> тр<strong>а</strong>пецие<strong>в</strong>идн<strong>а</strong>я, <strong>с</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>им пр<strong>о</strong>д<strong>о</strong>льным <strong>в</strong>д<strong>а</strong><strong>в</strong>лением. Щит<strong>о</strong><strong>к</strong> б<strong>о</strong>льш<strong>о</strong>й,<br />

треуг<strong>о</strong>льный, <strong>в</strong>ытянут <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong> длину,<br />

<strong>с</strong> груше<strong>о</strong>бр<strong>а</strong>зным <strong>в</strong>д<strong>а</strong><strong>в</strong>лением п<strong>о</strong><strong>с</strong>редине. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 9 мм (ри<strong>с</strong>. 620). Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Hadrocoris Handlirsch, 1939;<br />

Liasocoris Wendt, 1940.<br />

СЕМЕЙСТВО PACHYMERID1IDAE<br />

HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ужены, <strong>с</strong> уз<strong>к</strong>им <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м,<br />

не <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим угл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>; <strong>к</strong><strong>о</strong>риум<br />

ед<strong>в</strong><strong>а</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>с</strong>ередины длины н<strong>а</strong>д<strong>к</strong>рылья и<br />

нерез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т мембр<strong>а</strong>ны, пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>н,<br />

<strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими я<strong>с</strong>ными жил<strong>к</strong><strong>а</strong>ми; мембр<strong>а</strong>н<strong>а</strong> <strong>с</strong><br />

де<strong>с</strong>ятью прямьгми п<strong>а</strong>р<strong>а</strong>ллельными нея<strong>с</strong>ными<br />

жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6 мм (ри<strong>с</strong>. 621).<br />

Юр<strong>а</strong>. 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Pachymeridium<br />

Geinitz, 1880.<br />

СЕМЕЙСТВО PROTOCORIDAE HANDLIRSCH, 1906<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленными <strong>к</strong><strong>о</strong>риум<strong>о</strong>м,<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м и мембр<strong>а</strong>н<strong>о</strong>й, п<strong>о</strong><strong>с</strong>ледняя уме-<br />

Ри<strong>с</strong>. 620-627. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Hadrocoridae, Pachymeridiidae, Protocoridae, Psychocoridae, Cuneocoridae f<br />

Sisyrocoridae, Diatilidae '<br />

211<br />

http://jurassic.ru/<br />

627<br />

Герм<strong>а</strong>ния<br />

н<strong>а</strong>д<strong>к</strong>рылье,<br />

iatllus debt-<br />

(Handlirsch,<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 9,3,<br />

14*

ренн<strong>о</strong>й <strong>в</strong>еличины; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и шир<strong>о</strong><strong>к</strong>ий;<br />

SC я<strong>с</strong>н<strong>а</strong>я; имеют<strong>с</strong>я М и CuA; мембр<strong>а</strong>н<strong>а</strong> без<br />

жил<strong>о</strong><strong>к</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> умеренн<strong>о</strong>й <strong>в</strong>еличины, <strong>с</strong>переди<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими гл<strong>а</strong>з<strong>а</strong>ми. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong>лег<strong>к</strong><strong>а</strong> тр<strong>а</strong>пецие<strong>в</strong>идн<strong>о</strong>й ф<strong>о</strong>рмы<br />

и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ырез<strong>а</strong>нным передним <strong>к</strong>р<strong>а</strong>ем. Щит<strong>о</strong><strong>к</strong><br />

неб<strong>о</strong>льш<strong>о</strong>й, ф<strong>о</strong>рмы р<strong>а</strong><strong>в</strong>н<strong>о</strong>бедренн<strong>о</strong>г<strong>о</strong> треуг<strong>о</strong>льни<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 10 мм (ри<strong>с</strong>. 622). Юр<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Protocoris Неег, 1865.<br />

СЕМЕЙСТВО PSYCHOCORIDAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>ры'лья <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленными <strong>к</strong><strong>о</strong>риум<strong>о</strong>м,<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м и мембр<strong>а</strong>н<strong>о</strong>й, имеют<strong>с</strong>я т<strong>а</strong><strong>к</strong>же<br />

нечет<strong>к</strong>ие эмб<strong>о</strong>лиум и <strong>к</strong>унеу<strong>с</strong>; четыре пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и <strong>к</strong><strong>о</strong>риум<strong>а</strong> <strong>о</strong>бры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я, не перех<strong>о</strong>дя<br />

<strong>в</strong> мембр<strong>а</strong>ну; п<strong>о</strong><strong>с</strong>ледняя <strong>с</strong> непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й<br />

из мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных жил<strong>о</strong><strong>к</strong>; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> угл<strong>о</strong><strong>в</strong><strong>а</strong>тый,<br />

з<strong>а</strong>мы<strong>к</strong><strong>а</strong>ет умеренн<strong>о</strong>й <strong>в</strong>еличины щит<strong>о</strong><strong>к</strong>. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 5 мм (ри<strong>с</strong>. 623). Н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременные Miridae и Anthocoridae.<br />

Юр<strong>а</strong>. 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Psychocoris<br />

Handlirsch, 1939.<br />

СЕМЕЙСТВО CUNEOCORIDAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> б<strong>о</strong>льшим <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong><strong>о</strong>м и нерез<strong>к</strong><strong>о</strong><br />

<strong>о</strong>тгр<strong>а</strong>ниченными <strong>к</strong><strong>о</strong>риум<strong>о</strong>м и мембр<strong>а</strong>н<strong>о</strong>й; имеет<strong>с</strong>я<br />

нея<strong>с</strong>ный <strong>к</strong>унеу<strong>с</strong>; мембр<strong>а</strong>н<strong>а</strong> <strong>с</strong> пятью-ше<strong>с</strong>тью<br />

нерез<strong>к</strong>ими жил<strong>к</strong><strong>а</strong>ми. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> умеренн<strong>о</strong>й<br />

<strong>в</strong>еличины, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми гл<strong>а</strong>з<strong>а</strong>ми. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>в</strong> ф<strong>о</strong>рме тр<strong>а</strong>пеции, <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> шире<br />

длины. Щит<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, треуг<strong>о</strong>льный, н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong><br />

уже передне<strong>с</strong>пин<strong>к</strong>и. Мел<strong>к</strong>ие ф<strong>о</strong>рмы 1<br />

. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 3 мм (ри<strong>с</strong>. 624). Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы/.<br />

Один р<strong>о</strong>д — Cuneocoris Handlirsch, 1939.<br />

СЕМЕЙСТВО SISYROCORIDAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й мембр<strong>а</strong>н<strong>о</strong>й, без<br />

рез<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тгр<strong>а</strong>ничения <strong>о</strong>т <strong>к</strong><strong>о</strong>риум<strong>а</strong>; п<strong>о</strong><strong>с</strong>ледний<br />

<strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong>ч<strong>а</strong>т<strong>о</strong>й <strong>с</strong><strong>к</strong>ульп­<br />

THassoZls avails' Tillyard; тел<strong>о</strong> и н<strong>а</strong>д<strong>к</strong>рылья, X 7,6, <strong>в</strong>. три1С,<br />

тур<strong>о</strong>й, ме<strong>с</strong>т<strong>а</strong>ми м<strong>о</strong>рщини<strong>с</strong>тый; SC <strong>о</strong>тделяет<br />

п<strong>о</strong>д<strong>о</strong>бие линейн<strong>о</strong>г<strong>о</strong> эмб<strong>о</strong>лиум<strong>а</strong>; R <strong>с</strong>ильн<strong>а</strong>я, <strong>с</strong><br />

неб<strong>о</strong>льшими р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми (п<strong>о</strong>д<strong>о</strong>бие <strong>к</strong>унеу<strong>с</strong><strong>а</strong>)<br />

н<strong>а</strong> гр<strong>а</strong>нице мембр<strong>а</strong>ны; М б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> нея<strong>с</strong>н<strong>а</strong>,<br />

<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине <strong>с</strong> R п<strong>о</strong>перечн<strong>о</strong>й, <strong>а</strong><br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> и <strong>с</strong> CuA; п<strong>о</strong><strong>с</strong>ледняя <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

длины 1<br />

Ри<strong>с</strong> 628—630. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Triassocoridae, Hypocimicidae<br />

н<strong>а</strong>д<strong>к</strong>рылья; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> умеренн<strong>о</strong>й <strong>в</strong>еличи­<br />

ны, угл<strong>о</strong><strong>в</strong><strong>а</strong>т. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5 мм (ри<strong>с</strong>. 625).<br />

Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Р<strong>о</strong>д Sisyrocoris Handlirsch,<br />

1939; <strong>к</strong> ним же Б<strong>о</strong>де <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит 15 <strong>с</strong>ледующих<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>: Acromocoris Bode, 1953; Adelocoris Bode,<br />

1953; Corynecoris Bode, 1953; Dichaspis Bode,<br />

1953; Emgerrophorus Bode, 1953; Ensphingocoris<br />

Bode, 1953; Entomecoris Bode, 1953;<br />

Euraspidium Bode, 1953; Eurynotis Bode, 1953;<br />

Ischnocoris Bode, 1953; Macropterocoris Bode,<br />

1953; Mesomphalocoris Bode, 1953; Somatocoris<br />

Bode, 1953; Stiphroschema Bode, 1953; Strobilocoris<br />

Bode, 1953, Trachycoris Bode, 1953.<br />

СЕМЕЙСТВО DIATILIDAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong><strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

мембр<strong>а</strong>н<strong>о</strong>й, <strong>о</strong>тделенн<strong>о</strong>й <strong>о</strong>т <strong>к</strong><strong>о</strong>риум<strong>а</strong> делящей<br />

линией; <strong>к</strong><strong>о</strong>риум не <strong>о</strong>чень пл<strong>о</strong>тный, без <strong>с</strong><strong>к</strong>ульптуры;<br />

гл<strong>а</strong><strong>в</strong>ные жил<strong>к</strong>и перех<strong>о</strong>дят из <strong>к</strong><strong>о</strong>риум<strong>а</strong><br />

<strong>в</strong> мембр<strong>а</strong>ну; SC я<strong>с</strong>н<strong>а</strong>я; R <strong>с</strong>ильн<strong>а</strong>я, д<strong>а</strong>ет ряд<br />

<strong>в</strong>ет<strong>в</strong>ей, <strong>с</strong><strong>о</strong>единен<strong>а</strong> <strong>с</strong> CuA дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й;<br />

М <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь лишь <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти;<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>тгр<strong>а</strong>ничен, р<strong>а</strong><strong>в</strong>ен 0,5 длины<br />

н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5—7 мм<br />

(ри<strong>с</strong>. 626, 627). Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы. Три р<strong>о</strong>д<strong>а</strong>:<br />

Diatilus Handlirsch, 1939; Apsicoria Handlirsch,<br />

1939; Cathalus Handlirsch, 1939.<br />

СЕМЕЙСТВО TRIASSOCORIDAE TILLYARD, 1922<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну, имеет<strong>с</strong>я лишь з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong><strong>к</strong> эмб<strong>о</strong>лиум<strong>а</strong><br />

(<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, <strong>о</strong>тделенн<strong>а</strong>я жил<strong>к</strong><strong>о</strong>й); <strong>к</strong><strong>о</strong>риум пр<strong>о</strong>тяги<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

д<strong>а</strong>лее <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>ры'лья, <strong>о</strong>тделя-<br />

212 -<br />

http://jurassic.ru/

ет<strong>с</strong>я <strong>о</strong>т уз<strong>к</strong><strong>о</strong>й мембр<strong>а</strong>ны дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й,<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>й <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>; гл<strong>а</strong><strong>в</strong>ные <strong>с</strong>т<strong>в</strong><strong>о</strong>лы М<br />

и СиА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии я<strong>с</strong>ные, <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти мембр<strong>а</strong>ны<br />

R и М д<strong>а</strong>ют <strong>с</strong>ерию р<strong>а</strong>ди<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей,<br />

пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ющих дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зную <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ую жил<strong>к</strong>у;<br />

<strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, <strong>с</strong>д<strong>в</strong>инут б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>, <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющий,<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет лишь <strong>с</strong>ередины 1<br />

CuP; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылий н<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют <strong>о</strong>дн<strong>а</strong> другую.<br />

Щит<strong>о</strong><strong>к</strong> треуг<strong>о</strong>льный, <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире длины. Бедр<strong>о</strong><br />

и г<strong>о</strong>лень передних н<strong>о</strong>г <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и шир<strong>о</strong><strong>к</strong>ие.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий 4—5 мм (ри<strong>с</strong>. 628,<br />

629). Пред<strong>к</strong>и б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>ти <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Hydrocorisae. Три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Один р<strong>о</strong>д —<br />

Triassocoris Tillyard, 1922.<br />

СЕМЕЙСТВО HYPOCIM1C1DAE HANDLIRSCH, 1939<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong><strong>о</strong>риум, <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> и<br />

мембр<strong>а</strong>ну; эмб<strong>о</strong>лиум нея<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен, и имеет<strong>с</strong>я<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е п<strong>о</strong>д<strong>о</strong>бие <strong>к</strong>унеу<strong>с</strong><strong>а</strong>; мембр<strong>а</strong>н<strong>а</strong><br />

<strong>о</strong>тделен<strong>а</strong> <strong>к</strong><strong>о</strong>нцентриче<strong>с</strong><strong>к</strong>ими, п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны'ми линиями; <strong>в</strong> <strong>к</strong><strong>о</strong>риуме<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь R, <strong>о</strong>тх<strong>о</strong>дящ<strong>а</strong>я <strong>о</strong>т нег<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

М и СиА; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, шир<strong>о</strong><strong>к</strong>ий,<br />

угл<strong>о</strong><strong>в</strong><strong>а</strong>тый; <strong>в</strong> мембр<strong>а</strong>не единичные жил<strong>к</strong>и.<br />

631. Corixa praeusfa Fieber; <strong>о</strong>бщий <strong>в</strong>ид, X 1,75, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Jensen-Haarup, 1925). 632. С. punctata llliger; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 2,9; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 2,9; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch,<br />

1925). 633. С. rhenana Statz; <strong>о</strong>бщий <strong>в</strong>ид, X 6,5, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Герм<strong>а</strong>ния (Statz, 1950). 634. ranatra linearis Linnaeus; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 2,6; Б—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 2,6; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch.<br />

Щит<strong>о</strong><strong>к</strong> неб<strong>о</strong>льш<strong>о</strong>й. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 6,5 мм<br />

(ри<strong>с</strong>. 630). Н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют Triassocoridae, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>ны и ближе<br />

<strong>к</strong> Naucoridae, <strong>о</strong>т п<strong>о</strong><strong>с</strong>ледних <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong>личием<br />

з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong><strong>а</strong> <strong>к</strong>унеу<strong>с</strong><strong>а</strong>, чт<strong>о</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong><br />

для не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Geocorisae. Юр<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д — Hypocimex Handlirsch,<br />

1939.<br />

СЕМЕЙСТВО CORIXIDAE LEACH, 1815. ГРЕБЛЯКИ<br />

[nom. transl. Kirby, 1837<br />

(ex Corixida Leach, 18151J<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум, линейный<br />

эмб<strong>о</strong>лиум и переп<strong>о</strong>н<strong>о</strong>ч<strong>к</strong>у без жил<strong>о</strong><strong>к</strong>.<br />

Л<strong>о</strong>б <strong>с</strong>ильн<strong>о</strong> з<strong>а</strong>гнут н<strong>а</strong>з<strong>а</strong>д. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> нер<strong>а</strong><strong>с</strong>члененный<br />

или д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ый, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong><br />

<strong>с</strong><strong>к</strong>рытый п<strong>о</strong>д г<strong>о</strong>л<strong>о</strong><strong>в</strong>ны'м щит<strong>к</strong><strong>о</strong>м. Антенны<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, при<strong>к</strong>реплены п<strong>о</strong>д гл<strong>а</strong>з<strong>а</strong>ми. Средне- и<br />

з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> <strong>с</strong>л<strong>о</strong>жные, п<strong>о</strong><strong>с</strong>ледняя <strong>с</strong> п<strong>а</strong>р<strong>а</strong>пле<strong>в</strong>р<strong>а</strong>ми.<br />

Передние н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> л<strong>о</strong>п<strong>а</strong>т<strong>о</strong>ч<strong>к</strong>у (pala), у<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> у<strong>с</strong><strong>а</strong>жены зубчи<strong>к</strong><strong>а</strong>ми, <strong>с</strong>луж<strong>а</strong>щими <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми<br />

з<strong>в</strong>у<strong>к</strong><strong>а</strong>; <strong>с</strong>редние н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, пр<strong>о</strong><strong>с</strong>тые,<br />

Ри<strong>с</strong>. 631 — 637. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Corixidae, Nepidae, Belostomatidae<br />

213<br />

http://jurassic.ru/<br />

1925). 635. Nepa cinerea Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 1,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Stichel, 1925). 636. Mesobelostomum deperditum (Germar);<br />

ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция, x 0,66; <strong>в</strong>. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch. 1925). 637.<br />

Limnogeton fteberl Mayr; А — переднее <strong>к</strong>рыл<strong>о</strong>.Х 1,5; Б— з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>,XI,5; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch, 1925)

з<strong>а</strong>дние — пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные, их <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Брюш<strong>к</strong><strong>о</strong> у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>с</strong>низу ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>а</strong><strong>с</strong>имметричн<strong>о</strong>.<br />

Жи<strong>в</strong>ут <strong>в</strong> <strong>с</strong>т<strong>о</strong>чных или медленн<strong>о</strong> те<strong>к</strong>ущих,<br />

ред<strong>к</strong><strong>о</strong> <strong>в</strong> бы<strong>с</strong>тр<strong>о</strong> те<strong>к</strong>ущих пре<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>дных<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х. Пит<strong>а</strong>ют<strong>с</strong>я <strong>в</strong><strong>о</strong>д<strong>о</strong>р<strong>о</strong><strong>с</strong>лями. Длин<strong>а</strong> тел<strong>а</strong><br />

2—16 мм (ри<strong>с</strong>. 631—633). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Неб<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е р<strong>о</strong>д<strong>а</strong>ми <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, из них <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы, не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и и чет<strong>в</strong>ертичных<br />

<strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО NEPIDAE LATRE1LLE, 1802.<br />

ВОДЯНЫЕ СКОРПИОНЫ<br />

[nom. transl. Stephens, 1829<br />

(ex Nepariae Latreille, 1802)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну; эмб<strong>о</strong>лиум нея<strong>в</strong><strong>с</strong>т<strong>в</strong>енный; мембр<strong>а</strong>н<strong>а</strong><br />

<strong>с</strong> жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими <strong>с</strong>еть. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong>ыд<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong>перед. Гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый,<br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Антенны <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми, <strong>с</strong><strong>к</strong>рытые и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

п<strong>о</strong>д гл<strong>а</strong>з<strong>а</strong>ми. Передние т<strong>а</strong>зи<strong>к</strong>и при<strong>к</strong>реплены<br />

у переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я переднегруд<strong>к</strong>и или близ нег<strong>о</strong>.<br />

Передние н<strong>о</strong>ги х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные, <strong>с</strong>редние и з<strong>а</strong>дние<br />

— х<strong>о</strong>дильные; т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ющие<strong>с</strong>я. Тел<strong>о</strong><br />

<strong>с</strong>ильн<strong>о</strong> упл<strong>о</strong>щенн<strong>о</strong>е <strong>с</strong> длинными <strong>в</strong><strong>о</strong>здушными<br />

труб<strong>к</strong><strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>с</strong> <strong>о</strong>дним<br />

члени<strong>к</strong><strong>о</strong>м. Хищни<strong>к</strong>и. Длин<strong>а</strong> тел<strong>а</strong> 15—<br />

65 мм (ри<strong>с</strong>. 634, 635). Юр<strong>а</strong> — ныне. Неб<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е<br />

р<strong>о</strong>д<strong>а</strong>ми <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны 1<br />

Mesonepa Handlirsch,<br />

1906 из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы 1<br />

, <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин из чет<strong>в</strong>ертичных<br />

<strong>о</strong>тл<strong>о</strong>жений С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО BELOSTOMAT1DAE LEACH, 1815<br />

[nom. transl. Uhler, 1886<br />

(ex Belostomida Leach, 1815)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье делит<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну, п<strong>о</strong><strong>с</strong>ледняя <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> я<strong>с</strong>ными жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>зующими <strong>с</strong>еть, и <strong>с</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющ<strong>а</strong>я. Гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый, <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>о</strong>б<strong>а</strong> <strong>с</strong>редних члени<strong>к</strong><strong>а</strong><br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> пильч<strong>а</strong>тые. Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong><br />

<strong>с</strong>л<strong>о</strong>жные. Передние т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющие.<br />

Передние н<strong>о</strong>ги х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные, з<strong>а</strong>дние —<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong>редних и з<strong>а</strong>дних н<strong>о</strong>г <strong>с</strong><br />

д<strong>в</strong>умя члени<strong>к</strong><strong>а</strong>ми и д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. В<strong>о</strong>здушные<br />

труб<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные, д<strong>в</strong>ухчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющие з<strong>а</strong> <strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong><br />

или <strong>с</strong><strong>к</strong>рытые. Хищни<strong>к</strong>и. Обит<strong>а</strong>ют <strong>в</strong> нижнем<br />

течении ре<strong>к</strong> <strong>в</strong>пл<strong>о</strong>ть д<strong>о</strong> приб<strong>о</strong>йн<strong>о</strong>й з<strong>о</strong>ны м<strong>о</strong>ря.<br />

Длин<strong>а</strong> тел<strong>а</strong> 35—90 мм (ри<strong>с</strong>. 636, 637). Юр<strong>а</strong> —<br />

214<br />

ныне. Неб<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е р<strong>о</strong>д<strong>а</strong>ми <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, из них <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии Mesobelostomum Н<strong>а</strong><strong>а</strong>se,<br />

1820 из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы 1<br />

, <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин р<strong>о</strong>д из<br />

не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО NAUCORIDAE FALLEN, 1814. ПЛАВТЫ<br />

[nom. transl. Samuelle, 1819<br />

(ex Naucoridei Fallen, 1814)]<br />

Н<strong>а</strong>д<strong>к</strong>ры'лья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум, эмб<strong>о</strong>лиум<br />

и мембр<strong>а</strong>ну, п<strong>о</strong><strong>с</strong>ледняя без жил<strong>о</strong><strong>к</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я. Гл<strong>а</strong>з<strong>к</strong>и рудимент<strong>а</strong>рные.<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>­<br />

л<strong>о</strong><strong>в</strong>ы 1<br />

http://jurassic.ru/<br />

, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, пр<strong>о</strong><strong>с</strong>тые, <strong>с</strong><strong>к</strong>рытые.<br />

Передние т<strong>а</strong>зи<strong>к</strong>и при<strong>к</strong>реплены у переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я переднегруд<strong>к</strong>и. Передние н<strong>о</strong>ги х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные,<br />

з<strong>а</strong>дние — пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные; п<strong>о</strong> <strong>к</strong>р<strong>а</strong>йней<br />

мере з<strong>а</strong>дние л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> д<strong>в</strong>умя<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. Тел<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е, без <strong>в</strong><strong>о</strong>здушные<br />

труб<strong>о</strong><strong>к</strong>. Жи<strong>в</strong>ут <strong>в</strong> <strong>в</strong><strong>о</strong>де. Хищни<strong>к</strong>и. Длин<strong>а</strong><br />

тел<strong>а</strong> 10—30 мм (ри<strong>с</strong>. 638, 639). Юр<strong>а</strong> —ныне.<br />

Из<strong>в</strong>е<strong>с</strong>тны р<strong>о</strong>ды Palaeoheteroptera Meunier,<br />

1900 и Nepidium Westwood, 1854 из <strong>в</strong>. юры<br />

3. Е<strong>в</strong>р<strong>о</strong>пы, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и<br />

не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы' и д<strong>в</strong><strong>а</strong> из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и. Юр<strong>а</strong> — ныне.<br />

СЕМЕЙСТВО NOTONECTIDAE LEACH, 1815.<br />

ГЛАДЫШИ <<br />

[nom. transl. Samuelle, 1819<br />

(ex Notonectida Leach, 1815)]<br />

(TO. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум,<br />

эмб<strong>о</strong>лиум и переп<strong>о</strong>н<strong>о</strong>ч<strong>к</strong>у без жил<strong>о</strong><strong>к</strong>, <strong>к</strong>рыше<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>с</strong><strong>о</strong>гнутую, <strong>с</strong> <strong>в</strong>ырез<strong>к</strong><strong>о</strong>й п<strong>о</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю. Четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>а</strong>нтенны <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

при<strong>к</strong>репленные п<strong>о</strong>д гл<strong>а</strong>з<strong>а</strong>ми, <strong>с</strong><strong>к</strong>рытые.<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет лишь передних<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong>, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет.<br />

Передние н<strong>о</strong>ги нех<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные, з<strong>а</strong>дние — пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные;<br />

передние т<strong>а</strong>зи<strong>к</strong>и при<strong>к</strong>реплены<br />

у з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я переднегруд<strong>к</strong>и, з<strong>а</strong>дние п<strong>о</strong>д<strong>в</strong>ижны<br />

<strong>в</strong> <strong>о</strong>дн<strong>о</strong>й пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>с</strong>ти, л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

ин<strong>о</strong>гд<strong>а</strong> у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> передние <strong>с</strong><br />

<strong>о</strong>дним члени<strong>к</strong><strong>о</strong>м. Брюш<strong>к</strong><strong>о</strong> без <strong>в</strong><strong>о</strong>здушных труб<strong>о</strong><strong>к</strong>.<br />

Жи<strong>в</strong>ут <strong>в</strong> <strong>в</strong><strong>о</strong>де, <strong>в</strong> <strong>с</strong>т<strong>о</strong>ячих или медленн<strong>о</strong><br />

те<strong>к</strong>ущих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х. Хищни<strong>к</strong>и. Длин<strong>а</strong> тел<strong>а</strong> 11—<br />

18 мм (ри<strong>с</strong>. 640 А, Б). В. юр<strong>а</strong> — ныне.<br />

Asionecta J. Popov, gen. nov. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. curtipes J. Popoy, sp. nov.; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Тел<strong>о</strong> удлиненн<strong>о</strong>-<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е.<br />

Бедр<strong>а</strong> <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ие, <strong>с</strong>ильн<strong>о</strong><br />

ут<strong>о</strong>лщенные, немн<strong>о</strong>г<strong>о</strong> длиннее г<strong>о</strong>леней; <strong>в</strong>ертлуги<br />

б<strong>о</strong>льшие, <strong>о</strong><strong>к</strong>руглые, <strong>с</strong> <strong>в</strong>ытянут<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й;<br />

л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; пер<strong>в</strong>ый члени<strong>к</strong><br />

л<strong>а</strong>п<strong>к</strong>и передних и <strong>с</strong>редних н<strong>о</strong>г п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ен<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>му, пер<strong>в</strong>ый члени<strong>к</strong> з<strong>а</strong>дней л<strong>а</strong>п<strong>к</strong>и р<strong>а</strong><strong>в</strong>ен

638. Iliocoris cimicoides (Linnaeus); A — переднее <strong>к</strong>рыл<strong>о</strong>, X 7,0<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 7,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925).<br />

639. Naucoris rottensis Statz, A — <strong>о</strong>бщий <strong>в</strong>ид<strong>с</strong><strong>в</strong>ерху.Х 20,8; Б —<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, X 20,8; п<strong>а</strong>ле<strong>о</strong>ген, Герм<strong>а</strong>ния (Statz. 1950).<br />

п<strong>о</strong>чти п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины з<strong>а</strong>дних г<strong>о</strong>леней. Стернит<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из<br />

д<strong>в</strong>ух гл<strong>а</strong>д<strong>к</strong>их пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> тел<strong>а</strong> 9,0—<br />

9,5 мм (ри<strong>с</strong>. 641). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, Notonectites Handlirsch, 1906<br />

из <strong>в</strong>. юры Герм<strong>а</strong>нии, <strong>о</strong>дин/ р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong>, не<strong>о</strong>ген<strong>а</strong><br />

и чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО TINGIDAE LAPORTE, 1832.<br />

КРУЖЕВНИЦЫ<br />

[nom. transl. Fieber, 1861 (ex Tigidites Laporte, 1832)]<br />

(E. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> <strong>с</strong>етью ячее<strong>к</strong> п<strong>о</strong> <strong>в</strong><strong>с</strong>ей п<strong>о</strong><strong>в</strong>ерх-<br />

Ри<strong>с</strong>. 638—641. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Naucoridae, Notonectidae<br />

640. Notonecta glauca Linnaeus; А — <strong>о</strong>бщий <strong>в</strong>ид, X 1,6; Б — пе-<br />

? |еднее <strong>к</strong>рыл<strong>о</strong>, X 2,4; В — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 2,4; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handirsch,<br />

1925). 641. Asionecta curtipes J. Popov; <strong>о</strong>бщ<strong>а</strong>й <strong>в</strong>ид,<br />

X 7,4, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

я <strong>о</strong><strong>с</strong>ти; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует или <strong>с</strong>лит <strong>с</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тью; деления н<strong>а</strong>д<strong>к</strong>рылья н<strong>а</strong> <strong>к</strong><strong>о</strong>риум<br />

и мембр<strong>а</strong>ну нет. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong><strong>к</strong>ул<strong>о</strong><strong>в</strong>ыми<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми, не пр<strong>о</strong>д<strong>в</strong>инутыми <strong>в</strong>перед, х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый; у<strong>с</strong>и<strong>к</strong>и длиннее г<strong>о</strong>­<br />

л<strong>о</strong><strong>в</strong>ы 1<br />

, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, ме<strong>с</strong>т<strong>о</strong> их при<strong>к</strong>реп­<br />

ления <strong>в</strong>идн<strong>о</strong> <strong>с</strong><strong>в</strong>ерху. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> пятиуг<strong>о</strong>льн<strong>о</strong>й<br />

ф<strong>о</strong>рмы, <strong>с</strong> длинным, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющим<br />

щит<strong>о</strong><strong>к</strong> <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м. Средне- и з<strong>а</strong>днегрудь<br />

цельные, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят <strong>к</strong><strong>а</strong>жд<strong>а</strong>я из <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> нер<strong>а</strong>зделенн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лерит<strong>а</strong>. Л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и<br />

<strong>с</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> тел<strong>а</strong> 1,5—5 мм<br />

(ри<strong>с</strong>. 642). Юр<strong>а</strong> — ныне. С<strong>в</strong>ыше 160 <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Tingiopsis Becker-Migdisova, 1953.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. reticulata Becker-Migdisova,<br />

1953; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (м<strong>а</strong>дыген<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

М<strong>а</strong>дыген). Н<strong>а</strong>д<strong>к</strong>рылье п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> чет<strong>к</strong><strong>о</strong>й <strong>в</strong>ыпу<strong>к</strong>-<br />

215<br />

http://jurassic.ru/

Ри<strong>с</strong>. 642—648. Tingidae, Saldidae, Dipsocoridae, Anthocoridae<br />

642. Phantoma baltica Drake; <strong>о</strong>бщий <strong>в</strong>ид, X 16,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Drake, 1950). 643. Tingiopsls reticulata<br />

Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 12,5, н. юр<strong>а</strong>, Ср. Азия<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1953). 644. Oligosaldtna aquatllls Statz;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 8,5, п<strong>а</strong>ле<strong>о</strong>ген, Герм<strong>а</strong>ния (Statz, 1950). 645. Сгурtostemma<br />

alienum Herrich-Schaeffer; <strong>о</strong>бщий <strong>в</strong>ид, X 24,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

л<strong>о</strong>й <strong>с</strong>ет<strong>к</strong><strong>о</strong>й; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong> <strong>с</strong>рез<strong>а</strong>нн<strong>а</strong>я; SC <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; R и RS<br />

<strong>о</strong>чень длинные; е<strong>с</strong>ть п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs;<br />

М <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти; е<strong>с</strong>ть rs-m.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,4 мм (ри<strong>с</strong>. 643). Один<br />

<strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы 1<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С.<br />

Амери<strong>к</strong>и, д<strong>в</strong><strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и и <strong>о</strong>дин из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и.<br />

(Киричен<strong>к</strong><strong>о</strong>,1951).646. Ceratocombus hurdi Wygodzinsky; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 60,0, не<strong>о</strong>ген (янт<strong>а</strong>рь). Ме<strong>к</strong><strong>с</strong>и<strong>к</strong><strong>а</strong> (Wygodzinsky, 1959).<br />

647. Orlus minutus (Linnaeus); <strong>о</strong>бщий <strong>в</strong>ид, X 22,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch,<br />

1925). 648. Anthocoris silvestris Linnaeus; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 14,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,]Х 14,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch,<br />

1925)<br />

216<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО SALDIDAE AMYOT ET SERVILLE, 1843.<br />

САЛЬДЫ, ИЛИ ПРИБРЕЖНЫЕ ПРЫГУНЫ<br />

[nom. trans. Costa, 1852<br />

(ex Saldides Amyot et Serville, 1843)]<br />

(Acanthiidae Stephens, 1829)<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну; п<strong>о</strong><strong>с</strong>ледняя <strong>с</strong> пятью-ше<strong>с</strong>тью пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими ячей<strong>к</strong>и,<br />

д<strong>а</strong>ле<strong>к</strong><strong>о</strong> не д<strong>о</strong>х<strong>о</strong>дящие д<strong>о</strong> ее <strong>в</strong>ершины. Гл<strong>а</strong>з<strong>а</strong><br />

<strong>о</strong>чень б<strong>о</strong>льшие, <strong>в</strong>ыпу<strong>к</strong>лые, <strong>с</strong>з<strong>а</strong>ди <strong>с</strong> <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й;

д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>к</strong><strong>а</strong> н<strong>а</strong> буг<strong>о</strong>р<strong>к</strong>е. Антенны 1<br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

н<strong>а</strong>*мн<strong>о</strong>г<strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый,<br />

д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>в</strong>ершины <strong>с</strong>реднегруд<strong>к</strong>и.<br />

Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ьш, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Жи<strong>в</strong>ут п<strong>о</strong> берег<strong>а</strong>м ручье<strong>в</strong>, ре<strong>к</strong>, <strong>о</strong>зер,<br />

м<strong>о</strong>рей. Хищни<strong>к</strong>и. Длин<strong>а</strong> тел<strong>а</strong> 3—7 мм<br />

(ри<strong>с</strong>. 644). П<strong>а</strong>ле<strong>о</strong>ген —ныне. Один р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и чет<strong>в</strong>ертичных<br />

<strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО DIPSOCORIDAE DOHRN, 1859<br />

(Ceratocombidae Puton, 1869;<br />

Cryptostemmatidae Hedicke, 1935)<br />

(70. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылье делит<strong>с</strong>я н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>й <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>,<br />

<strong>к</strong><strong>о</strong>риум и мембр<strong>а</strong>ну, <strong>к</strong>унеу<strong>с</strong> <strong>о</strong>бычн<strong>о</strong> имеет<strong>с</strong>я.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong>ытянут<strong>а</strong>я или <strong>с</strong>лег<strong>к</strong><strong>а</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненн<strong>а</strong>я:<br />

гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> м<strong>а</strong>лень<strong>к</strong>ие, пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>к</strong>и р<strong>а</strong>з<strong>в</strong>иты;<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый; у<strong>с</strong>и<strong>к</strong>и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

пер<strong>в</strong>ый и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong>и их<br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, п<strong>о</strong><strong>с</strong>ледние члени<strong>к</strong>и длинные,<br />

т<strong>о</strong>н<strong>к</strong>ие, п<strong>о</strong><strong>к</strong>рытые <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, третий члени<strong>к</strong><br />

у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния ут<strong>о</strong>лщенный. Грудь пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия<br />

п<strong>а</strong>хучих желез <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong><br />

тел<strong>а</strong> 0,8—3,0 мм (ри<strong>с</strong>. 645, 646). Жи<strong>в</strong>ут <strong>в</strong><strong>о</strong><br />

мху, п<strong>о</strong>д <strong>о</strong>п<strong>а</strong><strong>в</strong>шими ли<strong>с</strong>тьями, <strong>в</strong> мур<strong>а</strong><strong>в</strong>ейни<strong>к</strong><strong>а</strong>х<br />

и п<strong>о</strong> берег<strong>а</strong>м ре<strong>к</strong>. Не<strong>о</strong>ген —ныне. Неб<strong>о</strong>льш<strong>о</strong>е<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д из<br />

не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО ANTHOCORIDAE AMYOT ЕТ<br />

SERVILLE, 1843<br />

[nom. transl. Dallas, 1852<br />

(ex Anthocorides Amyot et Serville, 1843)]<br />

(E. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong> р<strong>а</strong>згр<strong>а</strong>ничены 1<br />

н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>,<br />

<strong>к</strong><strong>о</strong>риум, <strong>к</strong>унеу<strong>с</strong>, эмб<strong>о</strong>лиум и мембр<strong>а</strong>ну; п<strong>о</strong><strong>с</strong>ледняя<br />

у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>с</strong> <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>й треуг<strong>о</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong><strong>о</strong>й, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>й <strong>в</strong>д<strong>о</strong>ль ш<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>риум<strong>а</strong><br />

и не д<strong>о</strong>х<strong>о</strong>дящей д<strong>о</strong> <strong>в</strong>ершины. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

г<strong>о</strong>риз<strong>о</strong>нт<strong>а</strong>льн<strong>а</strong>я, н<strong>а</strong> <strong>в</strong>ершине <strong>о</strong>брубленн<strong>а</strong>я.<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Антенны четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Средне- и з<strong>а</strong>днегрудь <strong>с</strong>л<strong>о</strong>жные.<br />

Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Длин<strong>а</strong> тел<strong>а</strong> 1,5—<br />

4,0 мм (ри<strong>с</strong>. 647, 648). Хищни<strong>к</strong>и, мн<strong>о</strong>гие пит<strong>а</strong>ют<strong>с</strong>я<br />

тлями, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые — <strong>к</strong><strong>о</strong>р<strong>о</strong>ед<strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген,—<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 70 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>о</strong>дин р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО MIRIDAE HAHN, 1831. СЛЕПНЯКИ<br />

f<br />

' (Capsidae Burmeister, 1835)<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум,<br />

<strong>к</strong>унеу<strong>с</strong> и мембр<strong>а</strong>ну; п<strong>о</strong><strong>с</strong>ледняя б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью<br />

<strong>с</strong> <strong>о</strong>дн<strong>о</strong>й — д<strong>в</strong>умя ячей<strong>к</strong><strong>а</strong>ми. Гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет,<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Антенны н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong><br />

длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Среднеи<br />

з<strong>а</strong>днегрудь <strong>с</strong>л<strong>о</strong>жные. З<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и п<strong>о</strong>д<strong>в</strong>ижны<br />

<strong>в</strong> <strong>о</strong>дн<strong>о</strong>й пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>с</strong>ти. Л<strong>а</strong>п<strong>к</strong>и тречлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и <strong>с</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми. Пер<strong>в</strong>ые <strong>с</strong>егменты<br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong>низу н<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>с</strong>ледующих<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Н<strong>а</strong>ружные п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> <strong>о</strong>бы<strong>к</strong>н<strong>о</strong><strong>в</strong>енн<strong>о</strong><br />

нежные и мяг<strong>к</strong>ие, пл<strong>о</strong>х<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няющие<strong>с</strong>я.<br />

Б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> и <strong>в</strong>ид<strong>о</strong><strong>в</strong>,<br />

(ри<strong>с</strong>. 649). Юр<strong>а</strong> — ныне. С<strong>в</strong>ыше 600 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Miridoides Becker-Migdisova, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. mesozoicus Becker-Migdisova, sp.<br />

nov.; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>ильн<strong>о</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненн<strong>а</strong>я, п<strong>о</strong>перечн<strong>а</strong>я,<br />

ее ширин<strong>а</strong> между гл<strong>а</strong>з<strong>а</strong>ми менее ширины з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я передне<strong>с</strong>пин<strong>к</strong>и. У<strong>с</strong>и<strong>к</strong>и т<strong>о</strong>н<strong>к</strong>ие, <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

длины тел<strong>а</strong>; пер<strong>в</strong>ый члени<strong>к</strong> <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ий, з<strong>а</strong>х<strong>о</strong>дит<br />

з<strong>а</strong> <strong>в</strong>ершину г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й <strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее<br />

пер<strong>в</strong><strong>о</strong>г<strong>о</strong>, п<strong>о</strong><strong>с</strong>ледние члени<strong>к</strong>и <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятые<br />

лишь немн<strong>о</strong>г<strong>о</strong> длиннее <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong>. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> т<strong>о</strong>н<strong>к</strong>ий,<br />

д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>редних н<strong>о</strong>г. Гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие<br />

<strong>с</strong>идячие, <strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

тр<strong>а</strong>пецие<strong>в</strong>идн<strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong>и. Н<strong>а</strong>д<strong>к</strong>рылья д<strong>о</strong>х<strong>о</strong>дят<br />

д<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> брюш<strong>к</strong><strong>а</strong>, непр<strong>о</strong>зр<strong>а</strong>чные, передний<br />

и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>я их дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые<br />

<strong>в</strong>перед. CuP <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>д<strong>а</strong><strong>в</strong>лен<strong>а</strong>, н<strong>а</strong> <strong>к</strong><strong>о</strong>риуме т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

д<strong>в</strong>е жил<strong>к</strong>и, переп<strong>о</strong>н<strong>к</strong><strong>а</strong> непр<strong>о</strong>зр<strong>а</strong>чн<strong>а</strong>я; <strong>к</strong><strong>о</strong>неу<strong>с</strong><br />

немн<strong>о</strong>г<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния.<br />

Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, г<strong>о</strong>лени прямые; з<strong>а</strong>дние бедр<strong>а</strong> не<br />

з<strong>а</strong>х<strong>о</strong>дят з<strong>а</strong> <strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong>, немн<strong>о</strong>г<strong>о</strong> длиннее и<br />

т<strong>о</strong>лще <strong>с</strong>редних бедер. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 5 мм<br />

(ри<strong>с</strong>. 650). 'Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 18 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы 1<br />

, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле 11 из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>­<br />

ря, 7 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО NABIDAE COSTA, 1852<br />

[nom. transl. Dohrn, 1859<br />

(ex Nabini Costa, 1852)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong><strong>о</strong>риум, <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> и<br />

мембр<strong>а</strong>ну, ин<strong>о</strong>гд<strong>а</strong> имеет<strong>с</strong>я эмб<strong>о</strong>лиум; <strong>в</strong> мембр<strong>а</strong>не<br />

три-четыре б<strong>о</strong>льшие ячей<strong>к</strong>и, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>тх<strong>о</strong>дят п<strong>а</strong>р<strong>а</strong>ллельные жил<strong>к</strong>и, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ю­<br />

щие <strong>в</strong>ершины 1<br />

217<br />

http://jurassic.ru/<br />

н<strong>а</strong>д<strong>к</strong>рылий. Теменн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет <strong>в</strong>переди гл<strong>а</strong>з, <strong>с</strong>переди<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> из<strong>о</strong>гнутый, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый,<br />

пер<strong>в</strong>ый члени<strong>к</strong> ег<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Антенны<br />

<strong>о</strong>бычн<strong>о</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Переднегруд<strong>к</strong><strong>а</strong><br />

без з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong>. Средне- и<br />

з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> цельные. Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Д<strong>в</strong><strong>а</strong> генит<strong>а</strong>льных <strong>с</strong>егмент<strong>а</strong><br />

у <strong>с</strong><strong>а</strong>м<strong>к</strong>и п<strong>о</strong><strong>с</strong>редине р<strong>а</strong><strong>с</strong><strong>с</strong>еченные и з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ют<br />

<strong>в</strong> <strong>с</strong>ебе яйце<strong>к</strong>л<strong>а</strong>д. Немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленн<strong>о</strong>е

49. Calocoris antcnnatus Statz; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, x 5.8, п<strong>а</strong>ле<strong>о</strong>ен,<br />

Герм<strong>а</strong>ния (Statz, 1950). 650. Miridoides mesozoicus Beccker-Migdisova;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х10.8, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong> 1963) 651. Karanabis kiritshenkoiBeckeT-<br />

Migdisova, <strong>о</strong>бщий <strong>в</strong>ид, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Бе<strong>к</strong><strong>к</strong>ер-Мигди­<br />

Ри<strong>с</strong>. 649—653. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Miridae, Nabidae, Reduviidae<br />

218<br />

http://jurassic.ru/<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1963). 652. Platymeris, insignis (Germar); <strong>о</strong>бщий <strong>в</strong>иц,<br />

X 5,6, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Handlirsch,<br />

1952). 653. Rhinocoris michalki Statz; <strong>о</strong>бщий <strong>в</strong>ид,x 5,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Герм<strong>а</strong>ния (Statz, 1950)

р<strong>о</strong>д<strong>а</strong>ми <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>. Длин<strong>а</strong> тел<strong>а</strong> 5—10 мм.<br />

Юр<strong>а</strong> — ныне.<br />

Karanabis Becker-Migdisova, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К- kiritshenkoi Becker-Migdisova, sp.<br />

nov. <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Тел<strong>о</strong> удлиненн<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> умеренн<strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>я,<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong><strong>а</strong>я. Антенны четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

длинные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены у <strong>с</strong>ередины<br />

предгл<strong>а</strong>зничн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> т<strong>о</strong>ньше пер<strong>в</strong><strong>о</strong>г<strong>о</strong>, третий и чет<strong>в</strong>ертый<br />

т<strong>о</strong>н<strong>к</strong>ие; пер<strong>в</strong>ый — третий члени<strong>к</strong>и п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

длины, третий члени<strong>к</strong> длинный. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, т<strong>о</strong>н<strong>к</strong>ий, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> из<strong>о</strong>гнут.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong><strong>а</strong>я, <strong>в</strong>переди<br />

<strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ужен<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зуя я<strong>с</strong>ную шею, з<strong>а</strong>дняя<br />

д<strong>о</strong>ля ее дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>, <strong>с</strong>ередин<strong>а</strong> ди<strong>с</strong><strong>к</strong><strong>а</strong><br />

упл<strong>о</strong>щен<strong>а</strong>. Щит<strong>о</strong><strong>к</strong> неб<strong>о</strong>льш<strong>о</strong>й. Н<strong>о</strong>ги длинные,<br />

т<strong>а</strong>зи<strong>к</strong>и передних н<strong>о</strong>г <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ытянуты; передние<br />

бедр<strong>а</strong> р<strong>а</strong>зн<strong>о</strong>мерн<strong>о</strong>й ширины и без зубц<strong>о</strong><strong>в</strong>,<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>редних, передние г<strong>о</strong>лени<br />

р<strong>а</strong><strong>в</strong>ны длине бедер. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong> я<strong>с</strong>ным, <strong>о</strong>б<strong>о</strong>д<strong>к</strong><strong>о</strong>м,<br />

<strong>о</strong>тделенным <strong>с</strong>низу <strong>о</strong>т <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти<br />

брюш<strong>к</strong><strong>а</strong> я<strong>с</strong>ным <strong>в</strong>д<strong>а</strong><strong>в</strong>лением. Ф<strong>о</strong>рмы <strong>с</strong>редней<br />

<strong>в</strong>еличины — длин<strong>а</strong> тел<strong>а</strong> 5 мм (ри<strong>с</strong>. 651). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО REDUVIIDAE LATREILLE, 1807.<br />

ХИЩНИКИ<br />

(nom. transl. Stephens, 1829<br />

(ex Reduvini Latreille, 1807)1<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну; <strong>в</strong> мембр<strong>а</strong>не д<strong>в</strong>е-три <strong>к</strong>лет<strong>к</strong>и у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

и д<strong>в</strong>е-три <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные жил<strong>к</strong>и. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

из<strong>о</strong>гнутый, <strong>с</strong>ильный. Антенны четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чными<br />

члени<strong>к</strong><strong>а</strong>ми, <strong>к</strong> <strong>в</strong>ершине ут<strong>о</strong>нч<strong>а</strong>ющие<strong>с</strong>я. Гл<strong>а</strong>з<strong>к</strong>и<br />

н<strong>а</strong> <strong>в</strong><strong>о</strong>з<strong>в</strong>ышении. Переднегруд<strong>к</strong><strong>а</strong> <strong>с</strong> жел<strong>о</strong>б<strong>к</strong><strong>о</strong>м,<br />

и<strong>с</strong>черченным ч<strong>а</strong><strong>с</strong>тыми п<strong>о</strong>перечными рубчи<strong>к</strong><strong>а</strong>ми,<br />

<strong>с</strong>луж<strong>а</strong>щими з<strong>в</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>ым <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong>м. Щит<strong>о</strong><strong>к</strong><br />

м<strong>а</strong>лень<strong>к</strong>ий или умеренн<strong>о</strong>й <strong>в</strong>еличины. Среднеи<br />

з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> цельные. Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

ред<strong>к</strong><strong>о</strong> д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, ин<strong>о</strong>гд<strong>а</strong> передние<br />

<strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

<strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н<strong>а</strong>х, <strong>в</strong> п<strong>а</strong>ле<strong>а</strong>р<strong>к</strong>ти<strong>к</strong>е<br />

немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> хищни<strong>к</strong>и.<br />

Длин<strong>а</strong> тел<strong>а</strong> 4 мм (ри<strong>с</strong>. 652, 653). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. С<strong>в</strong>ыше 300 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> пяти р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> ше<strong>с</strong>ти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и и<br />

<strong>о</strong>дин из чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО VELIIDAE DUGLAS ET SCOTT, 1865.<br />

ВЕЛИИ<br />

В н<strong>а</strong>д<strong>к</strong>рыльях <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и мембр<strong>а</strong>н<strong>а</strong><br />

<strong>с</strong>литы <strong>в</strong> <strong>о</strong>дн<strong>о</strong> цел<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>к</strong>переди<br />

<strong>с</strong>уж<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я и н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненн<strong>а</strong>я. Гл<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я передне<strong>с</strong>пин<strong>к</strong>и; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет<br />

или нея<strong>в</strong><strong>с</strong>т<strong>в</strong>енные. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый.<br />

Антенны зн<strong>а</strong>чительн<strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>з<strong>а</strong>ди <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong><br />

треуг<strong>о</strong>льный <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>к</strong>, з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющий щит<strong>о</strong><strong>к</strong>.<br />

Передне- и <strong>с</strong>реднегруд<strong>к</strong><strong>а</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й длины,<br />

цельные. Средние т<strong>а</strong>зи<strong>к</strong>и при<strong>к</strong>реплены п<strong>о</strong>чти<br />

н<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т передне- и з<strong>а</strong>днегруд<strong>к</strong>и.<br />

З<strong>а</strong>дние н<strong>о</strong>ги длиннее передних, бедр<strong>а</strong><br />

их ут<strong>о</strong>лщенные; у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>ух-трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и при<strong>к</strong>реплены перед их <strong>в</strong>ершин<strong>о</strong>й.<br />

Жи<strong>в</strong>ут н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>в</strong><strong>о</strong>ды бы<strong>с</strong>тр<strong>о</strong>те<strong>к</strong>ущих<br />

и <strong>с</strong>т<strong>о</strong>ячих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>. Длин<strong>а</strong> тел<strong>а</strong> 2—8 мм<br />

(ри<strong>с</strong>. 654, 655). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пе и <strong>о</strong>дин из не<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО HEN1COCEPHALIDAE STAL, 1865<br />

Гп<strong>о</strong>т. transl. Berg, 1896<br />

(ex Henicocephalida Stal, 1865)]<br />

Н<strong>а</strong>д<strong>к</strong>рылье без я<strong>с</strong>н<strong>о</strong>г<strong>о</strong> деления н<strong>а</strong> <strong>к</strong><strong>о</strong>риум<br />

и мембр<strong>а</strong>ну, жил<strong>к</strong>и пл<strong>о</strong>тные, не р<strong>а</strong>з<strong>о</strong>р<strong>в</strong><strong>а</strong>ны<br />

н<strong>а</strong> <strong>с</strong>ередине и <strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong> центре <strong>к</strong>рыл<strong>а</strong> <strong>к</strong>рупные<br />

ячей<strong>к</strong>и; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> я<strong>с</strong>н<strong>о</strong> <strong>о</strong>тделен, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> длинн<strong>а</strong>я,<br />

я<strong>с</strong>н<strong>о</strong> <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong>, пр<strong>о</strong><strong>с</strong>тые<br />

гл<strong>а</strong>з<strong>к</strong>и я<strong>с</strong>ные. Антенны четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Пле<strong>в</strong>ры <strong>с</strong>редне- и<br />

з<strong>а</strong>днегруд<strong>к</strong>и нея<strong>с</strong>н<strong>о</strong> р<strong>а</strong>зделены; переднегруд<strong>к</strong><strong>а</strong><br />

<strong>с</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й для <strong>с</strong>тре<strong>к</strong><strong>о</strong>т<strong>а</strong>ния. Л<strong>а</strong>п<strong>к</strong>и передних<br />

н<strong>о</strong>г <strong>с</strong> <strong>о</strong>дним, <strong>а</strong> з<strong>а</strong>дних — <strong>с</strong> тремя члени<strong>к</strong><strong>а</strong>ми,<br />

без <strong>а</strong>р<strong>о</strong>лий; передние г<strong>о</strong>лени <strong>к</strong> <strong>к</strong><strong>о</strong>нцу р<strong>а</strong><strong>с</strong>ширены.<br />

П<strong>а</strong>хучие железы <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong><br />

тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 мм (ри<strong>с</strong>. 656, 657 А—В). Хищни<strong>к</strong>и.<br />

П<strong>а</strong>ле<strong>о</strong>ген •— ныне. Из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Ю. Азии (янт<strong>а</strong>рь Бирмы).<br />

СЕМЕЙСТВО GERRIDAE LEACH, 1815.<br />

ВОДОМЕРКИ<br />

[nom. transl. Dohrn, 1859<br />

(ex Gerrida Leach, 1815)]<br />

В н<strong>а</strong>д<strong>к</strong>рылье <strong>к</strong><strong>о</strong>риум, <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> и мембр<strong>а</strong>н<strong>а</strong><br />

<strong>с</strong>литы <strong>в</strong> <strong>о</strong>дн<strong>о</strong> цел<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> ед<strong>в</strong><strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

ширины. Гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>идят у ее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния и п<strong>о</strong>чти<br />

<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я передне<strong>с</strong>пин<strong>к</strong>и.<br />

Гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> д<strong>в</strong><strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> нея<strong>с</strong>ные. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый.<br />

Щит<strong>о</strong><strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> <strong>с</strong><strong>к</strong>рыт п<strong>о</strong>д<br />

<strong>в</strong>ытянутым н<strong>а</strong>з<strong>а</strong>д треуг<strong>о</strong>льным <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м передне<strong>с</strong>пин<strong>к</strong>и.<br />

Средне- и з<strong>а</strong>днегрудь цельные,<br />

<strong>с</strong>реднегрудь <strong>о</strong>чень б<strong>о</strong>льш<strong>а</strong>я, з<strong>а</strong>днегрудь <strong>к</strong><strong>о</strong>-<br />

219<br />

http://jurassic.ru/

657А<br />

Ри<strong>с</strong>. 654—661. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Veliidae, Henicocephalidae, Gerridae, Hydrometridae<br />

654. Velia brachialis Stal; <strong>о</strong>бщий <strong>в</strong>ид.Х 6,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем., С. и Ю. Амери<strong>к</strong><strong>а</strong><br />

(Blatchley, 1926). 655. V. rivulorum Fabricius; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 3,6; Б —з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 3,6; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch,<br />

1925). 656. Henicocephalus formicinus Uhler; <strong>о</strong>бщий <strong>в</strong>ид, X 2,0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем., С. Амери<strong>к</strong><strong>а</strong> (Beier, 1938). 657. Н. fossilis Cockerell;<br />

А — ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья, X 2,5; Б — передняя н<strong>о</strong>г<strong>а</strong>;<br />

р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. З<strong>а</strong>дние и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>с</strong>редние н<strong>о</strong>ги зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее передних; т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>редних и<br />

з<strong>а</strong>дних н<strong>о</strong>г <strong>с</strong>ближенные. З<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и длин-ные,<br />

цилиндриче<strong>с</strong><strong>к</strong>ие, <strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ющие<strong>с</strong>я, при<strong>к</strong>репленные<br />

п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м тел<strong>а</strong> и д<strong>а</strong>ле<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные.<br />

Л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

при<strong>к</strong>репленные у <strong>в</strong>ершины. Жи<strong>в</strong>ут н<strong>а</strong><br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>в</strong><strong>о</strong>ды, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые ф<strong>о</strong>рмы <strong>о</strong><strong>к</strong>е<strong>а</strong>ниче<strong>с</strong><strong>к</strong>ие.<br />

Длин<strong>а</strong> тел<strong>а</strong> 6—18 мм (ри<strong>с</strong>. 658, 659).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Из<strong>в</strong>е<strong>с</strong>тен <strong>о</strong>дин р<strong>о</strong>д . из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и<br />

<strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ге<strong>в</strong><strong>а</strong>, не<strong>о</strong>ген<strong>а</strong> и чет<strong>в</strong>ертичных<br />

<strong>о</strong>тл<strong>о</strong>жений 3. Е<strong>в</strong>р<strong>о</strong>пы, З<strong>а</strong><strong>к</strong><strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>зья и<br />

С. Амери<strong>к</strong>и.<br />

220<br />

http://jurassic.ru/<br />

657 Б<br />

В — у<strong>с</strong>и<strong>к</strong>и; янт<strong>а</strong>рь, Бирм<strong>а</strong> (Cockerell, 1916). 658. Gem's argentata<br />

Schiimmel; <strong>о</strong>бщий <strong>в</strong>ид, X 5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Beier, 1938)<br />

659. G. paludum Fabricius; A — переднее <strong>к</strong>рыл<strong>о</strong>, X 7,9; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, х 7,9; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch, 1925). 660. Hydrometra<br />

martini Kirkaldy; <strong>о</strong>бщий <strong>в</strong>ид, X 5,3, <strong>с</strong><strong>о</strong><strong>в</strong>рем., С. Амери<strong>к</strong><strong>а</strong> (Blatchley,<br />

1926). 661. Н. stagnorum Linnaeus', А — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Х6,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 6,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., (Handlirsch, 1925)<br />

658<br />

СЕМЕЙСТВО HYDROMETRIDAE BILLBERG, 1820.<br />

ПАЛОЧКОВИДНЫЕ ВОДОМЕРКИ<br />

[nom. transl. Stephens, 1829<br />

(ex Hydrometraedes Billberg, 1820)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>дн<strong>о</strong>р<strong>о</strong>дные, без деления н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>риум и мембр<strong>а</strong>ну; пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют бе<strong>с</strong><strong>к</strong>рылые<br />

ф<strong>о</strong>рмы или <strong>с</strong> рудимент<strong>а</strong>ми <strong>к</strong>рылье<strong>в</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> удлиненн<strong>а</strong>я, н<strong>а</strong> <strong>в</strong>ершине р<strong>а</strong><strong>с</strong>ширенн<strong>а</strong>я.<br />

Гл<strong>а</strong>з<strong>а</strong> д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т<strong>о</strong>д<strong>в</strong>инуты <strong>о</strong>т переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

передне<strong>с</strong>пин<strong>к</strong>и, гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> трехчле-

Ри<strong>с</strong>. 662—668. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Aradidae, Lygaeidae, Coreidae, Pyrrhocoridae, Berytidae<br />

€62. Aradus superstes Germar; <strong>о</strong>бщий <strong>в</strong>ид, X 10,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Germar, 1856). 663. Lygeosoma wagneri<br />

Statz; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу,X 9,5, п<strong>а</strong>ле<strong>о</strong>ген, Герм<strong>а</strong>ния (Statz,<br />

1950) 664. Acanthocephala femorata Fabricius; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху,<br />

ХЗ.О, <strong>с</strong><strong>о</strong><strong>в</strong>рем., С. Амери<strong>к</strong><strong>а</strong> (Blatchley, 1926). 665. Karatavocons<br />

asiatica Becker-Migdisova, <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, X 8,2, <strong>в</strong>. юр<strong>а</strong>,<br />

ни<strong>к</strong><strong>о</strong><strong>в</strong>ый, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Щит<strong>к</strong><strong>а</strong> нет или <strong>о</strong>чень м<strong>а</strong>л.<br />

Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> цельные. Н<strong>о</strong>ги <strong>о</strong>чень<br />

т<strong>о</strong>н<strong>к</strong>ие, ните<strong>в</strong>идные; л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и н<strong>а</strong> <strong>в</strong>ершине их, без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Жи<strong>в</strong>ут<br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>в</strong><strong>о</strong>ды и п<strong>о</strong> берег<strong>а</strong>м <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 9—25 мм (ри<strong>с</strong>. 660, 661). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Из<strong>в</strong>е<strong>с</strong>тен <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

и не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы'<br />

и С. Амери<strong>к</strong>и.<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1963). 666. Odontopus nlgncornis<br />

Stal; <strong>о</strong>бщий <strong>в</strong>ид, X 3,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Индия (Beier, 1938). 667.<br />

Largus rufipennis Laporte; A — переднее <strong>к</strong>рыл<strong>о</strong>, X 5,8; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, X 5,8; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch, 1925). 668. Neides ttpularius<br />

(Linnaeus); A — переднее <strong>к</strong>рыл<strong>о</strong>,X 9,2; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong><br />

Аш Х9,2; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Handlirsch, 1925)<br />

221<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ARADIDAE BR., 1835<br />

ПОДКОРНИКИ<br />

(nom. trausl. Costa, 1852<br />

(ex Aradiens Br., 1835)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну; <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong> <strong>к</strong> <strong>в</strong>ершине п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> <strong>с</strong>уж<strong>а</strong>ющий<strong>с</strong>я<br />

и ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не з<strong>а</strong>х<strong>о</strong>дящий з<strong>а</strong> <strong>в</strong>ершину<br />

щит<strong>к</strong><strong>а</strong>, мембр<strong>а</strong>н<strong>а</strong> <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими непр<strong>а</strong><strong>в</strong>ильными<br />

и <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>а</strong>м<strong>о</strong>зирующими жил<strong>к</strong><strong>а</strong>ми,

ред<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> без жил<strong>о</strong><strong>к</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> г<strong>о</strong>риз<strong>о</strong>нт<strong>а</strong>льн<strong>а</strong>я,<br />

между у<strong>с</strong>и<strong>к</strong><strong>а</strong>ми <strong>в</strong>ытянут<strong>а</strong>я <strong>в</strong> длинный<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>к</strong>. Буг<strong>о</strong>р<strong>к</strong>и, п<strong>о</strong>ддержи<strong>в</strong><strong>а</strong>ющие <strong>а</strong>нтенны,<br />

<strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итые, з<strong>а</strong><strong>о</strong><strong>с</strong>тренные. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый,<br />

пер<strong>в</strong>ый члени<strong>к</strong> <strong>о</strong>чень м<strong>а</strong>л.<br />

Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> цельные. Л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и без прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Тел<strong>о</strong> <strong>с</strong><strong>в</strong>ерху<br />

и <strong>с</strong>низу пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е. Жи<strong>в</strong>ут п<strong>о</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>й дере<strong>в</strong>ье<strong>в</strong><br />

и н<strong>а</strong> трут<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong>х, пит<strong>а</strong>я<strong>с</strong>ь грибным мицелием<br />

и <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>м р<strong>а</strong><strong>с</strong>тений. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 2—<br />

5 мм (ри<strong>с</strong>. 662). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

40 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и<br />

не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО LYGAEIDAE SCHILLER, 1829<br />

[nom. transl. Herrich-Schaeffer, 1835<br />

(ex Lygaeides Schiller, 1829)1<br />

(Myodochidae Kirkaldy, 1899)<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну <strong>с</strong> пятью пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми.<br />

Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> без перетяж<strong>к</strong>и<br />

перед гл<strong>а</strong>з<strong>а</strong>ми. Буг<strong>о</strong>р<strong>к</strong>и, п<strong>о</strong>ддержи<strong>в</strong><strong>а</strong>ющие<br />

<strong>а</strong>нтенны, з<strong>а</strong>ним<strong>а</strong>ют б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение. Пр<strong>о</strong><strong>с</strong>тые<br />

гл<strong>а</strong>з<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong> е<strong>с</strong>ть. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый.<br />

У<strong>с</strong>и<strong>к</strong>и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> цельные.<br />

Д<strong>в</strong><strong>а</strong> генит<strong>а</strong>льных <strong>с</strong>егмент<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и р<strong>а</strong><strong>с</strong>­<br />

<strong>с</strong>ечены 1<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й щелью, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>с</strong><strong>к</strong>рыт<br />

яйце<strong>к</strong>л<strong>а</strong>д. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные и <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги.<br />

Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 2,5—20 мм (ри<strong>с</strong>. 663).<br />

Б<strong>о</strong>лее 100 р<strong>о</strong>д<strong>о</strong><strong>в</strong>. Мел — ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны: Mesolygaeus<br />

Ping, 1928 из мел<strong>а</strong> Кит<strong>а</strong>я, д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы' (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

12 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и,<br />

четыре р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из<br />

не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО COREIDAE LEACH, 1815.<br />

КРАЕВИКИ<br />

[nom. transl. Stephens, 1829<br />

(ex Coreides Leach, 1815)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну. Кл<strong>а</strong><strong>в</strong>у<strong>с</strong> п<strong>о</strong>з<strong>а</strong>ди щит<strong>к</strong><strong>а</strong> <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>о</strong>бщий ш<strong>о</strong><strong>в</strong>; мембр<strong>а</strong>н<strong>а</strong> <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong>единяющими<strong>с</strong>я жил<strong>к</strong><strong>а</strong>ми. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не<br />

<strong>о</strong>бр<strong>а</strong>зует щит<strong>к</strong><strong>а</strong> и без п<strong>о</strong>перечн<strong>о</strong>г<strong>о</strong> <strong>в</strong>д<strong>а</strong><strong>в</strong>ления<br />

перед гл<strong>а</strong>з<strong>а</strong>ми. Гл<strong>а</strong>з<strong>к</strong>и е<strong>с</strong>ть, х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый.<br />

Антенны н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы,<br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, ме<strong>с</strong>т<strong>о</strong> их при<strong>к</strong>репления<br />

<strong>в</strong>идн<strong>о</strong> <strong>с</strong><strong>в</strong>ерху. Щит<strong>о</strong><strong>к</strong> умеренн<strong>о</strong>й <strong>в</strong>еличины,<br />

222<br />

http://jurassic.ru/<br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че брюш<strong>к</strong><strong>а</strong>. Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong><br />

цельные. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и<br />

<strong>с</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные. Длин<strong>а</strong><br />

тел<strong>а</strong> 5 мм (ри<strong>с</strong>. 664). В. юр<strong>а</strong> — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 200 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Karatavocoris Becker - Migdisova,<br />

gen. nov. Тип р<strong>о</strong>д<strong>а</strong> — К. asiatica Becker-Migdisova,<br />

sp. nov.; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм<br />

К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тел<strong>а</strong>, у<strong>с</strong>и<strong>к</strong>и и н<strong>о</strong>ги г<strong>о</strong>лые.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> уже передне<strong>с</strong>пин<strong>к</strong>и, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

ее ширин<strong>а</strong> <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> гл<strong>а</strong>з<strong>а</strong>ми немн<strong>о</strong>г<strong>о</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ет ее длину; гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>идячие; <strong>с</strong><strong>к</strong>ул<strong>о</strong><strong>в</strong>ые<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и не пр<strong>о</strong>д<strong>в</strong>инуты <strong>в</strong>перед. Пер<strong>в</strong>ый члени<strong>к</strong><br />

у<strong>с</strong>и<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, не з<strong>а</strong>х<strong>о</strong>дит з<strong>а</strong> <strong>в</strong>ершину<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й и третий неут<strong>о</strong>лщенные, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й<br />

немн<strong>о</strong>г<strong>о</strong> длиннее третьег<strong>о</strong>; чет<strong>в</strong>ертый члени<strong>к</strong><br />

удлиненн<strong>о</strong>-яйце<strong>в</strong>идный, <strong>в</strong> 1,6 <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> и<br />

<strong>в</strong> 1,4 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че третьег<strong>о</strong> члени<strong>к</strong><strong>а</strong>. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

п<strong>о</strong>чти д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>редних н<strong>о</strong>г. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong> цельными б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>к</strong>р<strong>а</strong>ями; щит<strong>о</strong><strong>к</strong><br />

<strong>о</strong>т<strong>к</strong>рытый, у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>в</strong>д<strong>а</strong><strong>в</strong>ленный. Т<strong>а</strong>зи<strong>к</strong>и<br />

з<strong>а</strong>дних н<strong>о</strong>г р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные, бедр<strong>а</strong> не <strong>в</strong><strong>о</strong><strong>о</strong>ружены,<br />

г<strong>о</strong>лени з<strong>а</strong>дних н<strong>о</strong>г п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ны длине<br />

з<strong>а</strong>дних бедер, п<strong>о</strong>чти прямые, неи<strong>с</strong><strong>к</strong>ри<strong>в</strong>ленные<br />

и не <strong>в</strong><strong>о</strong><strong>о</strong>ружены шип<strong>а</strong>ми. Брюш<strong>к</strong><strong>о</strong> эллиптиче<strong>с</strong><strong>к</strong>и<br />

<strong>о</strong><strong>к</strong>ругленн<strong>о</strong>е, нешир<strong>о</strong><strong>к</strong><strong>о</strong>е; ше<strong>с</strong>т<strong>о</strong>й <strong>с</strong>тернит<br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и н<strong>а</strong>п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину р<strong>а</strong><strong>с</strong><strong>с</strong>ечен пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й<br />

щелью. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 8 мм (ри<strong>с</strong>. 665).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Вне СССР: <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьми р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и, <strong>о</strong>дин р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. Амери<strong>к</strong>и и Ю.<br />

Амери<strong>к</strong>и и <strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и и<br />

чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений Афри<strong>к</strong>и.<br />

СЕМЕЙСТВО PYRRHOCORIDAE<br />

AMYOT ET SERVILLE, 1843.<br />

КРАСНОКЛОПЫ, СОЛДАТИКИ<br />

[nom. transl. Dohrn, 1859<br />

(ex Pyrrhocorides Amyot et Serville, 1843)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну; п<strong>о</strong><strong>с</strong>ледняя б<strong>о</strong>лее чем <strong>с</strong> пятью жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми<br />

или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми,<br />

п<strong>о</strong>ддержи<strong>в</strong><strong>а</strong>ющими у<strong>с</strong>и<strong>к</strong>и и з<strong>а</strong>ним<strong>а</strong>ющими б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е<br />

п<strong>о</strong>л<strong>о</strong>жение. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый.<br />

Антенны четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>тые.<br />

Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и <strong>с</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> тел<strong>а</strong> 7—8 мм (ри<strong>с</strong>. 666,667).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин из<br />

не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.

Ри<strong>с</strong>. 669—671. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pentatomidae, Cydnidae, Heteroptera incertae sedis<br />

669. Aspongopus nubtlis Westwood; <strong>о</strong>бщий <strong>в</strong>ид, X 1,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Афри<strong>к</strong><strong>а</strong> (Schouteden, 1913). 670. Cydnopsis vishneviensis<br />

J.Popov; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, X 15,7; Б —<strong>о</strong>бщий <strong>в</strong>ид<br />

СЕМЕЙСТВО BERYTIDAE F1EBER, 1851.<br />

ПАЛОЧКОВИДЫ КОЛЕНЧАТОУСЫЕ<br />

[nom. transl. Fieber, 1861 (ex Beritidea Fieber, 1851)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум<br />

и мембр<strong>а</strong>ну; мембр<strong>а</strong>н<strong>а</strong> <strong>с</strong> четырьмя-пятью<br />

жил<strong>к</strong><strong>а</strong>ми. Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>тые.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> перетяж<strong>к</strong><strong>о</strong>й, гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>д<strong>в</strong>инуты <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы 1<br />

, пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>к</strong>и е<strong>с</strong>ть. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Антенны четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые, н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы,<br />

пер<strong>в</strong>ый члени<strong>к</strong> длинный. Щит<strong>о</strong><strong>к</strong> <strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong>ий.<br />

Н<strong>о</strong>ги длинные, бедр<strong>а</strong> у <strong>в</strong>ершины бул<strong>а</strong><strong>в</strong><strong>о</strong>­<br />

223<br />

http://jurassic.ru/<br />

<strong>с</strong>низу, X 15,7, не<strong>о</strong>ген, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (<strong>о</strong>риг. ри<strong>с</strong>. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

671. Lygenocoris prynadai J. Popov, <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, X 17,0, <strong>в</strong>.<br />

юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Ю. П<strong>о</strong>п<strong>о</strong><strong>в</strong>, 1961)<br />

<strong>в</strong>идные, л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и <strong>с</strong><br />

прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> тел<strong>а</strong> 6—12 мм (ри<strong>с</strong>.668).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д — Berytinus sp.<br />

Menge, 1856 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО PENTATOMIDAE LEACH, 1815.<br />

ЩИТНИКИ<br />

[nom. transl. Samuelle, 1819<br />

(ex Pentatomidea Leach, 1815)]<br />

Н<strong>а</strong>д<strong>к</strong>рылья делят<strong>с</strong>я н<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>в</strong>у<strong>с</strong>, <strong>к</strong><strong>о</strong>риум и<br />

мембр<strong>а</strong>ну. Кл<strong>а</strong><strong>в</strong>у<strong>с</strong> <strong>к</strong> <strong>в</strong>ершине <strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я и не<br />

<strong>о</strong>бр<strong>а</strong>зует п<strong>о</strong>з<strong>а</strong>ди щит<strong>к</strong><strong>а</strong> <strong>о</strong>бщег<strong>о</strong> ш<strong>в</strong><strong>а</strong>, или эт<strong>о</strong>т

ш<strong>о</strong><strong>в</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; мембр<strong>а</strong>н<strong>а</strong> <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

жил<strong>к</strong><strong>а</strong>ми. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong> <strong>в</strong>иде щит<strong>к</strong><strong>а</strong>, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong><br />

н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м и <strong>с</strong><strong>к</strong>ул<strong>о</strong><strong>в</strong>ыми пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми.<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ый; <strong>а</strong>нтенны<br />

пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, мн<strong>о</strong>г<strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы,<br />

при<strong>к</strong>реплены <strong>с</strong>низу т<strong>а</strong><strong>к</strong>, чт<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние их<br />

<strong>с</strong><strong>в</strong>ерху не <strong>в</strong>идн<strong>о</strong>. Средне- и з<strong>а</strong>днегруд<strong>к</strong><strong>а</strong> дельные.<br />

К<strong>о</strong>г<strong>о</strong>т<strong>к</strong>и <strong>с</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные,<br />

реже хищни<strong>к</strong>и. Длин<strong>а</strong> тел<strong>а</strong> 20 мм (ри<strong>с</strong>.<br />

669). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды<br />

и <strong>в</strong>иды. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и С. Амери<strong>к</strong>и, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из<br />

не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, Прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и С. Амери<strong>к</strong>и,<br />

и четыре р<strong>о</strong>д<strong>а</strong> из чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и З<strong>а</strong><strong>к</strong><strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>зья.<br />

HETEROPTERA<br />

Lygaenocoris J. Popov, 1961. Тип р<strong>о</strong>д<strong>а</strong> — L.<br />

prynadai J. Popov, 1961; <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Тел<strong>о</strong> удлиненн<strong>о</strong>-<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

п<strong>о</strong>лу<strong>к</strong>ругл<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong><br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>редних н<strong>о</strong>г. Грудь пл<strong>о</strong>тн<strong>а</strong>я,' шир<strong>о</strong><strong>к</strong><strong>а</strong>я,<br />

б<strong>о</strong>ч<strong>к</strong>и з<strong>а</strong>днегруд<strong>к</strong>и п<strong>о</strong>з<strong>а</strong>ди прямые.<br />

Брюш<strong>к</strong><strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>о</strong>е.<br />

Сегменты брюш<strong>к</strong><strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужены <strong>в</strong> <strong>с</strong>редней<br />

ч<strong>а</strong><strong>с</strong>ти, <strong>к</strong>р<strong>о</strong>ме <strong>с</strong>едьм<strong>о</strong>г<strong>о</strong>. Стерниты <strong>с</strong>едьм<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью р<strong>а</strong>зделены <strong>с</strong>терн<strong>а</strong>льными<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>. Стернит<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит и при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<br />

де<strong>в</strong>ятый <strong>с</strong>егмент. Длин<strong>а</strong> тел<strong>а</strong> 5,6 мм<br />

(ри<strong>с</strong>. 671). В. юр<strong>а</strong> Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО PARAKNIGHTIIDAE EVANS, 1950<br />

[Ipsviciidae (pars) Evans, 1953;<br />

non Ipsviciidae Tillyard, 1919]<br />

(E. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им, н<strong>о</strong> шир<strong>о</strong><strong>к</strong>им эмб<strong>о</strong>лиум,<br />

пр<strong>о</strong>тяги<strong>в</strong><strong>а</strong>ющим<strong>с</strong>я д<strong>о</strong> <strong>с</strong>лияния R + М и<br />

без н<strong>о</strong>д<strong>а</strong>льн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong>и. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итыми п<strong>а</strong>р<strong>а</strong>н<strong>о</strong>т<strong>а</strong>льными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми.<br />

У <strong>с</strong><strong>а</strong>м<strong>к</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итый яйце<strong>к</strong>л<strong>а</strong>д<br />

(ри<strong>с</strong>. 672). Три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Один р<strong>о</strong>д — Р<strong>а</strong>-<br />

^raknightia Evans, 1943.<br />

СЕМЕЙСТВО ACTINOSCYTINIDAE EVANS, 1956<br />

(Cicadocoridae Becker-Migdisova, 1958)<br />

(E. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылья <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, без т<strong>о</strong>чечн<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>ульптуры, <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>с</strong>пине пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>,<br />

<strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я п<strong>о</strong> прям<strong>о</strong>й линии, или ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

СЕМЕЙСТВО CYDNIDAE BILLBERG, 1820<br />

[nom. transl. Fieber, 1861 (ex Cydnidea Billberg, 1820)]<br />

(Thyreocoridae Van Duzee, 1907)<br />

(70. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>)<br />

Н<strong>а</strong>д<strong>к</strong>рылье п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет брюш<strong>к</strong><strong>о</strong>,<br />

<strong>к</strong><strong>о</strong>риум п<strong>о</strong>чти треуг<strong>о</strong>льный, мембр<strong>а</strong>н<strong>а</strong> без жил<strong>о</strong><strong>к</strong>.<br />

Вт<strong>о</strong>р<strong>о</strong>й <strong>с</strong>тернит брюш<strong>к</strong><strong>а</strong> <strong>в</strong>е<strong>с</strong>ь или п<strong>о</strong>чти<br />

<strong>в</strong>е<strong>с</strong>ь з<strong>а</strong><strong>к</strong>рыт з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем з<strong>а</strong>днегруд<strong>к</strong>и, т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ше<strong>с</strong>ть тергит<strong>о</strong><strong>в</strong> <strong>с</strong> <strong>в</strong>идимым брюшным<br />

<strong>о</strong>б<strong>о</strong>д<strong>к</strong><strong>о</strong>м. Г<strong>о</strong>лени н<strong>о</strong>г гу<strong>с</strong>т<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты шип<strong>а</strong>ми.<br />

Л<strong>а</strong>п<strong>к</strong><strong>а</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я. Длин<strong>а</strong> тел<strong>а</strong> 3—20 мм<br />

(ри<strong>с</strong>. 670А, Б). Жи<strong>в</strong>ут <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е у <strong>к</strong><strong>о</strong>рней р<strong>а</strong><strong>с</strong>тений.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. Амери<strong>к</strong>и,<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> пяти р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

из не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и и <strong>о</strong>дин<br />

р<strong>о</strong>д из чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений З<strong>а</strong><strong>к</strong><strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>зья.<br />

INCERTAE SEDIS<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, Ю. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>)<br />

224<br />

http://jurassic.ru/<br />

ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong>лег<strong>а</strong>ет н<strong>а</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ую друг<strong>о</strong>г<strong>о</strong>,<br />

<strong>в</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>луч<strong>а</strong>е з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>с</strong>д<strong>в</strong>иг<strong>а</strong>ет<strong>с</strong>я <strong>в</strong><strong>в</strong>ерх; жил<strong>к</strong>и шир<strong>о</strong><strong>к</strong>ие;<br />

Ри<strong>с</strong>. 672—674. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Paraknightiidae, Actinoscytinidae<br />

672. Parakn ghtla magnified Evans; <strong>о</strong>бщий <strong>в</strong>ид.х 4,2, <strong>в</strong>. Пермь,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Evans, 1943), 673. Cicadocorls kuliki Becker-<br />

MijrdisovaJ(peKOHCTpyKiiHH);i X 9,6, <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 674. Olgamartynavia turanica Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 18,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958)

Ри<strong>с</strong>. 675—677. Heteroptera incertae sedis<br />

675. Shurabella lepyroniopsis Becker-Migdisova; A — н<strong>а</strong>д<strong>к</strong>рылье, <strong>с</strong>т<strong>а</strong>н (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 677. Coleopteropsis dollchoptera<br />

X17.0; Б — н<strong>а</strong>д<strong>к</strong>рылья и щит<strong>о</strong><strong>к</strong>, X 17,0; н. юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong>- Becker-Migdisova; А — н<strong>а</strong>д<strong>к</strong>рылье, X 14,0; Б — н<strong>а</strong>д<strong>к</strong>рылья и<br />

<strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 676. Karataviella brachyptera Becker-Migdi- щит<strong>о</strong><strong>к</strong>. X 14,0; н. юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949).<br />

ova; н<strong>а</strong>д<strong>к</strong>рылья и ч<strong>а</strong><strong>с</strong>ти тел<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, X 6,5, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х-<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>плы<strong>в</strong>ч<strong>а</strong>тые или теряющие<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии;<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья б<strong>о</strong>лее<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, жил<strong>к</strong>и ее б<strong>о</strong>лее пл<strong>о</strong>тные, передний<br />

<strong>к</strong>р<strong>а</strong>й упл<strong>о</strong>тнен шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й<br />

С; Ri, R 2 и RS <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, р<strong>а</strong><strong>в</strong>ные п<strong>о</strong> длине, п<strong>а</strong>р<strong>а</strong>ллельные.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, уже переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я передне<strong>с</strong>пин<strong>к</strong>и; п<strong>о</strong><strong>с</strong>ледняя <strong>к</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>е<br />

<strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я; щит<strong>о</strong><strong>к</strong> <strong>о</strong>чень б<strong>о</strong>льш<strong>о</strong>й, треуг<strong>о</strong>льный;<br />

передний <strong>к</strong>р<strong>а</strong>й ег<strong>о</strong> р<strong>а</strong><strong>в</strong>ен ширине<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния передне<strong>с</strong>пин<strong>к</strong>и; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я труб<strong>к</strong><strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. В. три<strong>а</strong><strong>с</strong> — <strong>в</strong>. юр<strong>а</strong>. Де<strong>с</strong>ять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Cicadocoris Becker-Migdisova, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — С. kuliki Becker-Migdisova, 1958;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, передний <strong>к</strong>р<strong>а</strong>й<br />

<strong>в</strong>ыпу<strong>к</strong>лый; М трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, Мз+4 пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

CuA пре<strong>в</strong>р<strong>а</strong>щен<strong>о</strong> <strong>в</strong> п<strong>о</strong>перечную, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

<strong>с</strong> п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми между Мз+4, CuA<br />

и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями Мз+4, М1+2 и Mi <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>с</strong>пл<strong>о</strong>шную дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зную линию <strong>о</strong>т <strong>в</strong>ершины<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля д<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я н<strong>а</strong>д<strong>к</strong>рылья.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4—6 мм (ри<strong>с</strong>. 673). Три <strong>в</strong>ид<strong>а</strong>.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Olgamartynovia Becker-Migdisova,<br />

1958. Тип р<strong>о</strong>д<strong>а</strong> — О. turanica Becker-Migdisova,<br />

1958; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>о</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый; М д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я или трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

М3+4 пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> cuai —<br />

<strong>с</strong>и<strong>а</strong>г <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, СиА <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> СиР <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong>.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3—5 мм (ри<strong>с</strong>. 674).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Archicercopis Handlirsch, 1939;<br />

Eocercopis. Handlirsch, 1939; Cercoprisca<br />

16 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

Handlirsch, 1939; Cercopinus Handlirsch, 1939;<br />

Actynoscytina Tillyard, 1926; Heteroscytina<br />

Evans, 1956; PlatyscytinellaEvans, 1956; Triscytina<br />

Evans, 1956.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> Heteroptera incertae sedis <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я<br />

<strong>с</strong>ледующие р<strong>о</strong>ды:<br />

Shurabella Becker-Migdisova, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Sh. lepyroniopsis Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б<br />

II). Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны,<br />

передний <strong>к</strong>р<strong>а</strong>й дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

и<strong>с</strong>чезл<strong>о</strong>, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong><strong>с</strong>ь лишь R <strong>в</strong> <strong>в</strong>иде углубления<br />

<strong>в</strong>д<strong>о</strong>ль <strong>в</strong><strong>с</strong>ей длины н<strong>а</strong>д<strong>к</strong>рылья и CuP,<br />

<strong>о</strong>тделяющ<strong>а</strong>я т<strong>а</strong><strong>к</strong>же <strong>в</strong> <strong>в</strong>иде углубления <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле. Scutellum неб<strong>о</strong>льш<strong>о</strong>й, треуг<strong>о</strong>льный, <strong>с</strong><br />

<strong>в</strong>ытянут<strong>о</strong>й <strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>к</strong><strong>о</strong>нечн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 3,5—4,9 мм (ри<strong>с</strong>. 675 А, Б). Один<br />

<strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Karataviella Becker-Migdisova, 1949 '.<br />

Тип р<strong>о</strong>д<strong>а</strong> — К. brachyptera Becker-Migdisova,<br />

1949; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идн<strong>о</strong>-<strong>в</strong>ытянут<strong>о</strong>е,<br />

не з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

брюш<strong>к</strong><strong>а</strong>, брюш<strong>к</strong><strong>о</strong> де<strong>в</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е. Scutellum<br />

б<strong>о</strong>льш<strong>о</strong>й, <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> шире длины, треуг<strong>о</strong>льный,<br />

<strong>в</strong>ершин<strong>а</strong> ег<strong>о</strong> туп<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я. Передне- и <strong>с</strong>редне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

т<strong>а</strong><strong>к</strong>же шир<strong>о</strong><strong>к</strong>ие, ширин<strong>а</strong> пер<strong>в</strong><strong>о</strong>й <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ет длину. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4,5—6,2 мм, длин<strong>а</strong> тел<strong>а</strong> 8,0—9,0 мм (ри<strong>с</strong>. 676).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

1<br />

П<strong>о</strong> н<strong>о</strong><strong>в</strong>ым д<strong>а</strong>нным р<strong>о</strong>д Karataviella В.-М. <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я<br />

<strong>к</strong> <strong>с</strong>ем. <strong>в</strong><strong>о</strong>дных п<strong>о</strong>луже<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых Corixidac<br />

(Ю. А. П<strong>о</strong>п<strong>о</strong><strong>в</strong>).<br />

225<br />

http://jurassic.ru/

Coleopteropsis Becker-Migdisova,<br />

1949; Тип р<strong>о</strong>д<strong>а</strong> — С. dolichoptera Becker-Migdisova,<br />

1949; н. юр<strong>а</strong>, Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б<br />

II).' Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны,<br />

передний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й, жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние и<strong>с</strong>чезл<strong>о</strong>,<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь лишь R и CuP <strong>в</strong> <strong>в</strong>иде пр<strong>о</strong>д<strong>о</strong>льных<br />

б<strong>о</strong>р<strong>о</strong>зд. Scutellum длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины,<br />

непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й мн<strong>о</strong>г<strong>о</strong>гр<strong>а</strong>нн<strong>о</strong>й ф<strong>о</strong>рмы, <strong>в</strong>ершин<strong>а</strong><br />

<strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>к</strong><strong>о</strong>нечн<strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>, н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е<br />

треуг<strong>о</strong>льн<strong>а</strong>я <strong>в</strong>ырез<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,8—<br />

5,0 мм (ри<strong>с</strong>. 677). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР: из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы Scylacocoris<br />

Handlirsch, 1906; Hemipteron Westwood,<br />

1854; Dimeropterum Handlirsch, 1906; Cimicidiutn<br />

Westwood, 1854; Galerucites Oppenheim,<br />

НАДОТРЯД PSOCOPTEROIDEA.<br />

СЕНОЕД00БРАЗНЫЕ<br />

Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны грызущие, реже <strong>к</strong><strong>о</strong>лющие,<br />

<strong>в</strong>ид<strong>о</strong>измененные; <strong>о</strong>бычн<strong>о</strong> имеет<strong>с</strong>я р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й н<strong>а</strong><strong>с</strong><strong>о</strong><strong>с</strong>,<br />

упр<strong>а</strong><strong>в</strong>ляемый м<strong>о</strong>щными мышц<strong>а</strong>ми, ме<strong>с</strong>т<strong>о</strong><br />

при<strong>к</strong>репления <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>тр<strong>а</strong>жен<strong>о</strong> <strong>с</strong>ильным<br />

<strong>в</strong>здутием н<strong>а</strong>лични<strong>к</strong><strong>а</strong>. Гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие. Крылья,<br />

е<strong>с</strong>ли имеют<strong>с</strong>я, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong> гетер<strong>о</strong>н<strong>о</strong>мные,<br />

и <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют я<strong>с</strong>н<strong>о</strong>й диптеригией,<br />

реже п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

или <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> пер<strong>о</strong><strong>к</strong>рылые (<strong>с</strong>н<strong>а</strong>бжены длинными<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю — птил<strong>о</strong>птеригия).<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>бедненн<strong>о</strong>е, п<strong>о</strong>перечные жил<strong>к</strong>и<br />

немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные. Пермь — ныне. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>в</strong>язи н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong> н<strong>а</strong>меч<strong>а</strong>ют<strong>с</strong>я <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й<br />

групп<strong>о</strong>й нижнеперм<strong>с</strong><strong>к</strong>их н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых<br />

Archaescytinidae, близ<strong>к</strong><strong>о</strong>й <strong>к</strong> ф<strong>о</strong>рме <strong>с</strong>ен<strong>о</strong>ед<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

и р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых. Отряды: Psocoptera,<br />

Д<strong>в</strong>е п<strong>а</strong>ры гетер<strong>о</strong>н<strong>о</strong>мных, реже г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных<br />

переп<strong>о</strong>нч<strong>а</strong>тых <strong>к</strong>рылье<strong>в</strong> <strong>с</strong> пл<strong>о</strong>тными жил<strong>к</strong><strong>а</strong>ми;<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. SC<br />

р<strong>а</strong>з<strong>о</strong>р<strong>в</strong><strong>а</strong>н<strong>а</strong>, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть ее з<strong>а</strong>мы<strong>к</strong><strong>а</strong>ет птер<strong>о</strong><strong>с</strong>тигму,<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я — <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я или <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> R. М и СиА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии или<br />

<strong>с</strong><strong>о</strong>единяют<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. CuP<br />

прям<strong>а</strong>я, <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я,<br />

реже пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong><br />

р<strong>а</strong>з<strong>в</strong>иты, у <strong>к</strong>рыл<strong>а</strong>тых три пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>а</strong>;<br />

з<strong>а</strong>дний <strong>о</strong>тдел н<strong>а</strong>лични<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый, <strong>в</strong>ерхняя губ<strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я; <strong>в</strong>ерхние челю<strong>с</strong>ти грызущег<strong>о</strong> тип<strong>а</strong>,<br />

нижние — <strong>с</strong> длинн<strong>о</strong>й, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й, <strong>с</strong><strong>к</strong><strong>о</strong>бля­<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, В. Н. Вишня<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

ОТРЯД PSOCOPTERA. СЕНОЕДЫ<br />

1888; Ап<strong>а</strong><strong>с</strong><strong>о</strong>ЬрШ<strong>а</strong> Handlirsch, 1906; Ischyopteron<br />

Oppenheim 1888. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, р<strong>о</strong>ды и <strong>о</strong>тдельные<br />

<strong>в</strong>иды из <strong>в</strong>. юры 3. Е<strong>в</strong>р<strong>о</strong>пы, <strong>о</strong>пи<strong>с</strong><strong>а</strong>нные<br />

<strong>к</strong><strong>а</strong><strong>к</strong> Cimicidae Westwood, 1854; Phillips, 1871,<br />

Brodie 1845; Pentatomidae Brodie, 1845; Nepidae<br />

Westwood, 1854; ? Citnex Brodie, 1854; ?<br />

Corita Oppenheim, 1888; Hydrometra Brodie,<br />

1845; Velia Brodie, 1845; Kleidocerys (=Pachimerus)<br />

Brodie, 1845; ? Cimicidae priscus Giebel,<br />

1856; Pachimerus zucholdi Giebel, 1856; Velia ?<br />

cormita Weyenberg, 1879, Protocorus ovalis<br />

Heer, 1865; <strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> из<br />

не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

Perielytrodea, Anoplura, Mallophaga, Rhynchophthiraptera<br />

и Zoraptera; из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны три пер<strong>в</strong>ых, причем<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель Anoplura (<strong>о</strong>дин <strong>в</strong>ид) <strong>о</strong>бн<strong>а</strong>ружен<br />

<strong>в</strong> плей<strong>с</strong>т<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых <strong>о</strong>тл<strong>о</strong>жениях Сибири.<br />

И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong>ен<strong>о</strong>ед<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

еще м<strong>а</strong>л<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>. П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

пит<strong>а</strong>ния (мицет<strong>о</strong>ф<strong>а</strong>гия, <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>гия и<br />

п<strong>а</strong>р<strong>а</strong>зитизм) н<strong>а</strong>ряду <strong>с</strong> уменьшением р<strong>а</strong>змер<strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> и ух<strong>о</strong>д<strong>о</strong>м <strong>в</strong> те<strong>с</strong>ные, <strong>с</strong><strong>к</strong>рытые <strong>с</strong>т<strong>а</strong>ции (п<strong>о</strong>д<br />

<strong>к</strong><strong>о</strong>ру, <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>у и т. д.), был<strong>о</strong> <strong>в</strong><strong>а</strong>жнейшим н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лением<br />

э<strong>в</strong><strong>о</strong>люции эт<strong>о</strong>й группы. В не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>с</strong>луч<strong>а</strong>ях р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь и «<strong>о</strong>бще<strong>с</strong>т<strong>в</strong>енный» <strong>о</strong>бр<strong>а</strong>з<br />

жизни (т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы з<strong>о</strong>р<strong>а</strong>итеры)

t<br />

Ри<strong>с</strong>. 678—681. П<strong>о</strong>д<br />

878. Dlchentomum sofanense Becker-Migdisova <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у<br />

X 11,5 <strong>в</strong> пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961).<br />

679. Dlchentomum ttnctum Tillyard (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); X 10,0,<br />

Н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1933). 680. Martynopsocus ar-<br />

ПОДОТРЯД PERMOPSOCIDA<br />

Крылья г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мны. В переднем "<strong>к</strong>рыле SC<br />

длинн<strong>а</strong>я, птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, RS<br />

д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, М <strong>с</strong> четырьмя <strong>в</strong>ет<strong>в</strong>ями. Ан<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> упл<strong>о</strong>щенным н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м.<br />

Л<strong>а</strong>п<strong>к</strong><strong>а</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я. Цер<strong>к</strong>и е<strong>с</strong>ть. Пермь.<br />

Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Psocidiidae, Martynopsocidae,<br />

Permopsocidae.<br />

СЕМЕЙСТВО PSOCIDIIDAE TILLYARD, 1926<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong> <strong>в</strong> длину;<br />

SC длинн<strong>а</strong>я, дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R <strong>в</strong> <strong>о</strong><strong>с</strong>­<br />

1яд Permopsocida.<br />

678<br />

cuatus (Martynov); переднее <strong>к</strong>рыл<strong>о</strong>, X Б.ВЛ<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). 681. Permopsocus congener Tillyard,<br />

переднее <strong>к</strong>рыл<strong>о</strong>, Х16.0, н. Пермь, С. Амери<strong>к</strong><strong>а</strong> (Tillyard, 1926).<br />

н<strong>о</strong><strong>в</strong><strong>а</strong>нии и ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

<strong>в</strong>ет<strong>в</strong>ей RS д<strong>в</strong>е, реже три, <strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

<strong>в</strong>ет<strong>в</strong>ей М четыре или б<strong>о</strong>лее; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA<br />

б<strong>о</strong>льш<strong>о</strong>й, ширин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> 2—3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше <strong>в</strong>ы<strong>с</strong><strong>о</strong>ты;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и уз<strong>к</strong><strong>о</strong>е; д<strong>в</strong>е п<strong>о</strong>перечные<br />

жил<strong>к</strong>и r-rs и rs-m. Пермь. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Dlchentomum Tillyard, 1926 (Psocidium<br />

Tillyard, 1926; Chaetopsocidium Tillyard, 1926;<br />

Metapsocidium Tillyard, 1926; Pentapsocidium<br />

Tillyard, 1926; Permentomum Tillyard, 1926; Parapsocidium<br />

G. Zalessky, 1937). Тип р<strong>о</strong>д<strong>а</strong> —<br />

D. tinctum Tillyard, 1926; н. пермь, С. Амери<strong>к</strong><strong>а</strong>.<br />

(К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

227 15*<br />

http://jurassic.ru/

прям<strong>о</strong>й; SC <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R немн<strong>о</strong>г<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее<br />

птер<strong>о</strong><strong>с</strong>тигмы; R, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, R 2, RS и М <strong>о</strong>чень<br />

длинные, р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти<br />

<strong>к</strong>рыл<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями;<br />

<strong>о</strong>дн<strong>а</strong> межр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

r-rs; <strong>в</strong>ет<strong>в</strong>ей М четыре. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

4,0—5,0 мм (ри<strong>с</strong>. 678, 679). Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н.<br />

пермь Ур<strong>а</strong>л<strong>а</strong> и С. Амери<strong>к</strong>и, <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Вне СССР: Austropsocidium Tillyard, 1935 и<br />

Stenopsocidium Tillyard, 1935. В<strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие пл<strong>о</strong>х<strong>о</strong>й<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ти <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение<br />

р<strong>о</strong>д<strong>а</strong> Megapsocidium Tillyard, 1935 нея<strong>с</strong>н<strong>о</strong>.<br />

СЕМЕЙСТВО MARTYNOPSOCIDAE KARNY, 1930<br />

(Dinopsocidae Martynov, 1928)<br />

Переднее <strong>к</strong>рыл<strong>о</strong> умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>в</strong>ытянут<strong>о</strong>е;<br />

SC длинн<strong>а</strong>я дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> <strong>с</strong> R; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей RS три, длинные;<br />

<strong>в</strong>ет<strong>в</strong>ей М три; <strong>с</strong>т<strong>в</strong><strong>о</strong>л СиА ДЛИННЫЙ; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

СиА неб<strong>о</strong>льш<strong>о</strong>й; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е<br />

и длинн<strong>о</strong>е; не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

<strong>о</strong>бр<strong>а</strong>зует <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> з<strong>а</strong>м<strong>к</strong>нутые<br />

ячей<strong>к</strong>и. В. пермь. Один р<strong>о</strong>д.<br />

Martynopsocus Karny, 1930 (Dinopsocus<br />

Martynov, 1928; Idelopsocus M. Zalessky, 1929).<br />

Тип р<strong>о</strong>д<strong>а</strong> — Dinopsocus arcuatus Martynov,<br />

1928; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). В переднем <strong>к</strong>рыле SC <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> R н<strong>а</strong> <strong>с</strong>ередине «<strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует ряд <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; R прям<strong>а</strong>я, <strong>с</strong> лег<strong>к</strong>им<br />

изгиб<strong>о</strong>м <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти птер<strong>о</strong><strong>с</strong>тигмы, RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

R <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети <strong>к</strong>рыл<strong>а</strong>, ее передняя <strong>в</strong>ет<strong>в</strong>ь<br />

и Мз+4 <strong>в</strong>ет<strong>в</strong>ят<strong>с</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 11 мм<br />

(ри<strong>с</strong>. 680). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

СЕМЕЙСТВО PERMOPSOCIDAE TILLYARD, 1926<br />

Переднее <strong>к</strong>рыл<strong>о</strong> умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC<br />

прям<strong>а</strong>я, длинн<strong>а</strong>я, <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> R; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей М четыре; ширин<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

СиА меньше ег<strong>о</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong>ты; передняя <strong>в</strong>ет<strong>в</strong>ь<br />

СиА рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> между <strong>в</strong>ет<strong>в</strong>ями R и М; <strong>в</strong><strong>с</strong>егд<strong>а</strong> е<strong>с</strong>ть п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> Шз +4-<strong>с</strong>и<strong>а</strong> (ри<strong>с</strong>. 681). Пермь. Четыре<br />

р<strong>о</strong>д<strong>а</strong>.<br />

Вне СССР: Progonopsocus Tillyard, 1926,<br />

Permopsocus Tillyard, 1926 (Ancylopsocus Tillyard,<br />

1926), Lithopsocidium Carpenter, 1932, Orthopsocus<br />

Carpenter, 1932.<br />

ПОДОТРЯД PARAPSOCIDA<br />

Крыл<strong>а</strong>тые и бе<strong>с</strong><strong>к</strong>рылые. Крылья гетер<strong>о</strong>мные.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних <strong>к</strong>рылье<strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>о</strong><br />

примити<strong>в</strong>ные черты: передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>к</strong><strong>о</strong>-<br />

228<br />

http://jurassic.ru/<br />

<strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>н, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, жил<strong>к</strong>и<br />

б<strong>о</strong>лее прямые, <strong>с</strong> длинными <strong>в</strong>ет<strong>в</strong>ями. З<strong>а</strong>дние<br />

зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних, <strong>с</strong> <strong>о</strong>бедненным<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е, <strong>о</strong>ни <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные<br />

или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я у <strong>в</strong>ершины <strong>в</strong> <strong>о</strong>бщий <strong>с</strong>тебель,<br />

или <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>а</strong> и <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

у <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым н<strong>а</strong>лични<strong>к</strong><strong>о</strong>м.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Пермь — ныне. Одинн<strong>а</strong>дц<strong>а</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Surijocopsocidae,<br />

Lophioneuridae, Zygopsocidae, Pachytroctidae,<br />

Sphaeropsocidae, Liposcelidae, Amphientomidae,<br />

Lepidopsocidae, Empheriidae,<br />

Psyllipsocidae, Archipsyllidae, Archipsocidae,<br />

Psoquillidae; п<strong>о</strong><strong>с</strong>леднее т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО SURIJOKOPSOCIDAE<br />

BECKER-MIGDISOVA, 1961<br />

Передние <strong>к</strong>рылья <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти рез<strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширены, пр<strong>о</strong>зр<strong>а</strong>чные, жил<strong>к</strong>и пл<strong>о</strong>тные; передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й, <strong>с</strong>лег<strong>к</strong><strong>а</strong> упл<strong>о</strong>тнен; SC прям<strong>а</strong>я,<br />

<strong>с</strong>ближен<strong>а</strong> <strong>с</strong> R + М, длинн<strong>а</strong>я, н<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рылье<strong>в</strong> я<strong>с</strong>н<strong>а</strong>я б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong><br />

(В), <strong>о</strong>гр<strong>а</strong>ниченн<strong>а</strong>я ут<strong>о</strong>лщенными <strong>о</strong>трез<strong>к</strong><strong>а</strong>ми<br />

жил<strong>о</strong><strong>к</strong> R+M, СиА (д<strong>о</strong> изгиб<strong>а</strong> <strong>к</strong> М), <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

М (д<strong>о</strong> ее рез<strong>к</strong><strong>о</strong>г<strong>о</strong> изгиб<strong>а</strong> п<strong>о</strong>д прямым угл<strong>о</strong>м)<br />

и п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й m-cua; пять <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ет<strong>в</strong>ей М; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ужен<strong>о</strong>.<br />

В. пермь. Один р<strong>о</strong>д.<br />

Surijokopsocus В е <strong>с</strong> k е r-M i g d i s <strong>о</strong> v a,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — 5. radtshenkoi Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). В переднем <strong>к</strong>рыле<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>, четырехгр<strong>а</strong>нн<strong>а</strong>я,<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние ее рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>о</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

р<strong>а</strong><strong>с</strong>ширен<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; <strong>с</strong>лег<strong>к</strong><strong>а</strong> упл<strong>о</strong>тнен<strong>о</strong>; RS<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> R; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья; <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>а</strong>я<br />

М5;1б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть СиА <strong>о</strong>бр<strong>а</strong>зует неб<strong>о</strong>льш<strong>о</strong>й<br />

изгиб <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию М, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дит<br />

п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> m-cua; <strong>с</strong>тебель СиА <strong>о</strong>чень<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; ширин<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> СиА <strong>в</strong> 2—3 р<strong>а</strong>з<strong>а</strong><br />

б<strong>о</strong>льше ег<strong>о</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong>ты; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нных,<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> лет<strong>а</strong>ющих ф<strong>о</strong>рм, <strong>о</strong>чень<br />

уз<strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 9 мм<br />

(ри<strong>с</strong>. 682; т<strong>а</strong>бл. XVI, фиг. 1). Один <strong>в</strong>ид.<br />

В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО LOPHIONEURIDAE TILLYARD, 1921<br />

(Cyphoneuridae Carpenter, 1932;<br />

Zoropsocidae Tillyard, 1935)<br />

Передние <strong>к</strong>рылья пр<strong>о</strong>зр<strong>а</strong>чные, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенные или л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные;<br />

SC <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> С<br />

б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>, реже <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

R пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; RS <strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>, <strong>с</strong> длинным<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>в</strong>ет<strong>в</strong>ей М <strong>в</strong><strong>с</strong>егд<strong>а</strong> д<strong>в</strong>е; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

б<strong>о</strong>лее ИЛИ менее из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й

Ри<strong>с</strong>. 682—691. П<strong>о</strong><br />

682. Surijokopsocus radtshenkoi Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 10,7, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 683. Lophioneurodes sarbalensis Becker-Migdisova; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 15,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1953). 684. ZoropsocusfomfensisBecker-Migdisova (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция);<br />

Х20.0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961).<br />

685. Surtjokocypha surijokovensis Becker-Migdisova; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 33,7, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961). 686. Vitriala nlgriapex Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Х15.2, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961).<br />

1тряд Parapsocida<br />

687. Psococlcadellopsis primitiva Becker-Msovigdia; передней<br />

<strong>к</strong>рыл<strong>о</strong>,X 32,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 688.<br />

Zygopsocus permianus Tillyard; переднее <strong>к</strong>рыл<strong>о</strong>, XI 1,5, <strong>в</strong>. пермь,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1935). 689. Palaeotroctes succinicus Hagen;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 7,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Enderlein,<br />

1911). 690. Sphaeropsocus kanowi Hagen; А — <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 62,0; Б — л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>с</strong>редней н<strong>о</strong>ги, X 300,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Enderlein, 1911). 691. Llposelis atavus<br />

Enderlein; <strong>о</strong>бщий <strong>в</strong>ид, X 50,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Enderlein, 1911)<br />

http://jurassic.ru/

ч<strong>а</strong><strong>с</strong>ти или <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>й з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>ью, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> CuP <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии или <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> нею <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й (изгиб CuA <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии), либ<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong><br />

<strong>о</strong>т <strong>с</strong><strong>о</strong>единения, пр<strong>о</strong>х<strong>о</strong>дит п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

М и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R+M; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong><br />

у мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>. З<strong>а</strong>дние <strong>к</strong>рылья н<strong>а</strong><br />

<strong>о</strong>дну треть <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних; SC нет; Ri+г п<strong>о</strong>чти<br />

прям<strong>а</strong>я. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, цилиндриче<strong>с</strong><strong>к</strong><strong>а</strong>я;<br />

<strong>с</strong>редне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми<br />

<strong>с</strong><strong>к</strong>лерит<strong>а</strong>ми; з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее.<br />

Пермь — три<strong>а</strong><strong>с</strong>. Де<strong>с</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Lophioneurodes Becker-Migdisova,<br />

1953. Тип р<strong>о</strong>д<strong>а</strong> — L. sarbalensis Becker-Migdisova,<br />

1953; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>а</strong>рб<strong>а</strong>л<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е,<br />

длинн<strong>о</strong>е; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS длиннее р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

М; CuA п<strong>о</strong>чти прям<strong>а</strong>я, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>а</strong> <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong>е<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние, <strong>с</strong><strong>о</strong>единяющее<strong>с</strong>я <strong>с</strong> CuP, п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> г + m-cua я<strong>с</strong>н<strong>а</strong>я, длинн<strong>а</strong>я, п<strong>о</strong>чему CuA<br />

<strong>к</strong><strong>а</strong>жет<strong>с</strong>я п<strong>о</strong>теря<strong>в</strong>шей <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние и <strong>о</strong>тх<strong>о</strong>дящей <strong>о</strong>т<br />

R+M. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 3,0 мм (ри<strong>с</strong>.<br />

683). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Zoropsocus Tillyard, 1935. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Z. delicatulus Tillyard, 1935; <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Переднее <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е, <strong>в</strong>ершин<strong>а</strong><br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, птер<strong>о</strong><strong>с</strong>типм<strong>а</strong> <strong>о</strong><strong>к</strong>р<strong>а</strong>шен<strong>а</strong>. Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

RS длиннее р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> М; CuA не <strong>с</strong>лит<strong>а</strong> <strong>с</strong> М, <strong>с</strong><strong>о</strong>единен<strong>а</strong><br />

<strong>с</strong> нею б<strong>о</strong>лее или менее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й или <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> ее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием,<br />

<strong>о</strong>тделяя б<strong>а</strong>з<strong>а</strong>льную ячей<strong>к</strong>у; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е, треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,8—3,0 мм (ри<strong>с</strong>. 684; т<strong>а</strong>бл.<br />

XVI, фиг. 2). Четыре <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Surijokocypha Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — S. surijokovensis Becker-Migdisova,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий 'б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Переднее <strong>к</strong>рыл<strong>о</strong> <strong>в</strong><br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти не<strong>с</strong>имметричн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>о</strong>,<br />

мембр<strong>а</strong>н<strong>а</strong> т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>" <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong><br />

з<strong>а</strong>темнением; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е;<br />

R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии из<strong>о</strong>гнут<strong>а</strong>, ди<strong>с</strong>т<strong>а</strong>льнее п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS <strong>к</strong><strong>о</strong>р<strong>о</strong>че р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,6—1,8 мм<br />

(ри<strong>с</strong>. 685). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Vitriala Becker-Migdisova, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — V. nigriapex Becker-Migdisova, 1961;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Переднее <strong>к</strong>рыл<strong>о</strong> пр<strong>о</strong>зр<strong>а</strong>чн<strong>о</strong>е, <strong>с</strong> м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong>й<br />

мембр<strong>а</strong>н<strong>о</strong>й; жил<strong>к</strong>и т<strong>о</strong>н<strong>к</strong>ие; R из<strong>о</strong>гнут<strong>а</strong><br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>;<br />

RS <strong>с</strong> длинным р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; М п<strong>о</strong>чти прям<strong>а</strong>я,<br />

без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; CuA <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 3,5 мм<br />

(ри<strong>с</strong>. 686). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Psococicadellopsis Becker-Migdisova,<br />

1961. Тип р<strong>о</strong>д<strong>а</strong> — P. primitiva Becker-Migdisova,<br />

1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(рэт. И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). В переднем <strong>к</strong>рыле SC <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

R + М и R пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют <strong>о</strong>дну прямую,<br />

<strong>с</strong>ильн<strong>о</strong> упл<strong>о</strong>тненную жил<strong>к</strong>у; RS без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>;<br />

М <strong>с</strong> длинным р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuA <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>, не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я и не <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й <strong>с</strong> М, из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти, <strong>с</strong> нея<strong>с</strong>н<strong>о</strong>й <strong>в</strong><strong>а</strong>д<strong>в</strong>ей <strong>в</strong>ет<strong>в</strong>ью р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>;<br />

CuP <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я; А я<strong>с</strong>н<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле длинн<strong>о</strong>е.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,8—2,0 мм (ри<strong>с</strong>. 687).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

230<br />

http://jurassic.ru/<br />

Вне СССР: Cyphoneura Carpenter, 1932;<br />

Cyphoneurodes Becker-Migdisova, 1953; Lophioneura<br />

Tillyard, 1922; Lophiocypha Tillyard, 1935;<br />

Austrocypha Tillyard, 1935.<br />

СЕМЕЙСТВО ZYGOPSOCIDAE TILLYARD, 1935<br />

Передние <strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые, <strong>с</strong> т<strong>о</strong>л<strong>с</strong>тыми<br />

жил<strong>к</strong><strong>а</strong>ми; SC длинн<strong>а</strong>я, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>;<br />

RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии CuA <strong>с</strong>лит<strong>а</strong> <strong>с</strong><br />

М; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA уз<strong>к</strong>ий и длинный; е<strong>с</strong>ть п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> r-m; CuP из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти; д<strong>в</strong>е <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 4,7 мм (ри<strong>с</strong>. 688). В. пермь А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Один р<strong>о</strong>д.<br />

СЕМЕЙСТВО PACHYTROCTIDAE ENDERLEIN, 1903<br />

[nom. transl. Pearman, 1936<br />

(ex Pachytroctinae Enderlein, 1903)]<br />

Бе<strong>с</strong><strong>к</strong>рылые или <strong>к</strong>рыл<strong>а</strong>тые, <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием, <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я лишь<br />

д<strong>в</strong>е-три пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и. У<strong>с</strong>и<strong>к</strong>и 15-члени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, бедр<strong>а</strong> т<strong>о</strong>н<strong>к</strong>ие.<br />

Тел<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Pachytroctinae, Tapinellinae;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

ПОДСЕМЕЙСТВО PACHYTROCTINAE<br />

ENDERLEIN, 1905<br />

Бе<strong>с</strong><strong>к</strong>рылые. Средне- и з<strong>а</strong>днегрудь р<strong>а</strong>зделены<br />

ш<strong>в</strong><strong>о</strong>м. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы <strong>с</strong><strong>к</strong>ульптурир<strong>о</strong><strong>в</strong><strong>а</strong>нные. Длин<strong>а</strong><br />

тел<strong>а</strong> 0,6—1,4 мм (ри<strong>с</strong>. 689). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Три р<strong>о</strong>д<strong>а</strong>, из них <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО SPHAEROPSOCIDAE MENON, 1942<br />

Передние <strong>к</strong>рылья у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные, элитр<strong>о</strong><strong>в</strong>ид-<br />

«ые, п<strong>о</strong>лу<strong>с</strong>фериче<strong>с</strong><strong>к</strong>ие, <strong>с</strong><strong>к</strong>ульптурир<strong>о</strong><strong>в</strong><strong>а</strong>нные.<br />

З<strong>а</strong>дние <strong>к</strong>рылья редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. У<strong>с</strong>и<strong>к</strong>и 15-члени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Длин<strong>а</strong> те-

л<strong>а</strong> 0,8 мм (ри<strong>с</strong>. 690). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО LIPOSCELIDAE ENDERLEIN, 1911<br />

(Troctidae Kolbe, 1882)<br />

Бе<strong>с</strong><strong>к</strong>рылые. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong>редне-<br />

и з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>литы. У<strong>с</strong>и<strong>к</strong>и 15-члени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Бедр<strong>а</strong> з<strong>а</strong>дних н<strong>о</strong>г ут<strong>о</strong>лщенные. Тел<strong>о</strong> эллиптиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

или яйце<strong>в</strong>идн<strong>о</strong>е, д<strong>о</strong>р<strong>с</strong><strong>о</strong>-<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong> упл<strong>о</strong>щен<strong>о</strong>.<br />

Л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Длин<strong>а</strong> тел<strong>а</strong> 1,0—<br />

2,0 мм (ри<strong>с</strong>. 691). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

р<strong>о</strong>д<strong>а</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>дин из<strong>в</strong>е<strong>с</strong>тен <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и из Афри<strong>к</strong>и<br />

(<strong>к</strong><strong>о</strong>п<strong>а</strong>л).<br />

СЕМЕЙСТВО AMPHIENTOMIDAE ENDERLEIN, 1903<br />

Передние <strong>к</strong>рылья и тел<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты чешуй<strong>к</strong><strong>а</strong>ми<br />

или передние <strong>к</strong>рылья <strong>с</strong> ми<strong>к</strong>р<strong>о</strong>трихиями; SC<br />

б<strong>о</strong>лее или менее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> не<br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей М три; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> CuA<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; длин<strong>а</strong> з<strong>а</strong>дней '<strong>в</strong>ет<strong>в</strong>и CuA <strong>к</strong><strong>о</strong>р<strong>о</strong>че ее<br />

<strong>с</strong>тебля; передняя <strong>в</strong>ет<strong>в</strong>ь CuA не <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong><br />

М п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е;<br />

Ai <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> CuP у <strong>в</strong>ершины; Аг <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>;<br />

реже Ai и Аг <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 13. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong><strong>в</strong>ерху не <strong>в</strong>идн<strong>а</strong> или пл<strong>о</strong>х<strong>о</strong> <strong>в</strong>идн<strong>а</strong>. Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

три. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Amphientominae, Electrentominae, Tineomorphiп<strong>а</strong>е.<br />

П<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йден<strong>о</strong>.<br />

ПОДСЕМЕЙСТВО AMPHIENTOMINAE<br />

ENDERLEIN, 1903<br />

Передние <strong>к</strong>рылья и тел<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты чешуй<strong>к</strong><strong>а</strong>ми,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong>рез<strong>а</strong>ны или <strong>с</strong> д<strong>в</strong>умя<br />

зубц<strong>а</strong>ми; SC прям<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>о</strong>тделены<br />

<strong>о</strong>дн<strong>а</strong> <strong>о</strong>т друг<strong>о</strong>й, реже <strong>с</strong>ближены, н<strong>о</strong> не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>дин <strong>с</strong>т<strong>в</strong><strong>о</strong>л. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong><strong>в</strong>ерху не<br />

<strong>в</strong>идн<strong>а</strong>. К<strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>с</strong> д<strong>в</strong>умя зубц<strong>а</strong>ми. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2—4 мм (ри<strong>с</strong>. 692). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них лишь <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО ELECTRENTOMINAE<br />

ENDERLEIN, 1911<br />

Крылья п<strong>о</strong><strong>к</strong>рыты ми<strong>к</strong>р<strong>о</strong>трих<strong>а</strong>ми ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

или п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью, чешуе<strong>к</strong> нет. SC <strong>в</strong> переднем <strong>к</strong>рыле<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую дугу, ди<strong>с</strong>т<strong>а</strong>льный ее <strong>к</strong><strong>о</strong>нец<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R+M; <strong>к</strong><strong>о</strong>нцы Ai и Аг <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>-<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я,<br />

<strong>с</strong><strong>в</strong>ерху ед<strong>в</strong><strong>а</strong> лишь <strong>в</strong>идн<strong>а</strong>.<br />

К<strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>с</strong> <strong>о</strong>дним зубц<strong>о</strong>м. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2,2 мм (ри<strong>с</strong>. 693 А, Б). П<strong>а</strong>ле<strong>о</strong>ген.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО LEPIDOPSOCIDAE ENDERLEIN, 1903<br />

(Lepidillidae Ribaga, 1905)<br />

Крылья и тел<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми или чешуй<strong>к</strong><strong>а</strong>ми.<br />

В переднем <strong>к</strong>рыле SC длинн<strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>ей<br />

М три; <strong>в</strong>ет<strong>в</strong>и М и CuA длинные; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

CuA <strong>о</strong>чень длинный и уз<strong>к</strong>ий; СиА 2 <strong>в</strong> 2—3 р<strong>а</strong>з<strong>а</strong><br />

длиннее <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> CuA; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>а</strong>, <strong>к</strong><br />

<strong>в</strong>ершине не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> CuP. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн<br />

21—47. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong>идн<strong>а</strong> <strong>с</strong><strong>в</strong>ерху <strong>в</strong> <strong>в</strong>иде<br />

уз<strong>к</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> три. Юр<strong>а</strong> —<br />

ныне. Пять п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Perientominae, Thylacinae,<br />

Asientorainae, Echinoprocinae, Lepidopsocinae;<br />

п<strong>о</strong><strong>с</strong>ледние д<strong>в</strong><strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне.<br />

ПОДСЕМЕЙСТВО PERIENTOMINAE KOLBE, 1884<br />

[nom. transl. Enderlein, 1903<br />

(ex Perientomini Kolbe, 1884)]<br />

Крылья и тел<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты чешуй<strong>к</strong><strong>а</strong>ми; <strong>в</strong>ершины<br />

передних и з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> <strong>в</strong>ытянуты. Щетин<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong>д<strong>о</strong>ль переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong>, или их нет. Между R и М д<strong>в</strong>е<br />

п<strong>о</strong>перечные жил<strong>к</strong>и. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> уз<strong>к</strong><strong>о</strong>й з<strong>а</strong>м<strong>к</strong>нут<strong>о</strong>й<br />

ячей<strong>к</strong><strong>о</strong>й между R + М и CuA; Ri <strong>о</strong>тх<strong>о</strong>дит<br />

между <strong>в</strong>ет<strong>в</strong>ями М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

2—3 мм (ри<strong>с</strong>. 694). Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

де<strong>с</strong>яти тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong> Индии и<br />

Афри<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО THYLACINAE ENDERLEIN, 1903<br />

Чешуе<strong>к</strong> н<strong>а</strong> теле и <strong>к</strong>рыльях нет; переднее<br />

<strong>к</strong>рыл<strong>о</strong> <strong>с</strong> щетин<strong>к</strong><strong>а</strong>ми <strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>я или без них;<br />

<strong>в</strong>ершины переднег<strong>о</strong> и з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рылье<strong>в</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>трены,<br />

<strong>в</strong> переднем <strong>к</strong>рыле д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и<br />

г-т. З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, <strong>с</strong> уз<strong>к</strong><strong>о</strong>й з<strong>а</strong>м<strong>к</strong>нут<strong>о</strong>й ячей<strong>к</strong><strong>о</strong>й<br />

между М и CuA. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

1,6 мм (ри<strong>с</strong>. 695А, Б). Чет<strong>в</strong>ертичные ? — ныне.<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из <strong>к</strong><strong>о</strong>п<strong>а</strong>л<strong>а</strong> Афри<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО ASIENTOMINAE MARTYNOV, 1926<br />

[nom. transl. Beeker-Mjgdisova, 1961, hie<br />

(ex Asientomidae Martynov, 1926)]<br />

(Lithentomidae Martynov, 1926;<br />

Lithopsocidae Karny, 1930)<br />

Вершины передних и з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong><br />

<strong>с</strong>рез<strong>а</strong>нные. В переднем <strong>к</strong>рыле ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть SC ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R; б<strong>а</strong>з<strong>а</strong>ль*<br />

231<br />

http://jurassic.ru/

н<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть R прям<strong>а</strong>я, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>,<br />

<strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нную птер<strong>о</strong><strong>с</strong>тигму;<br />

RS <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>в</strong>ет<strong>в</strong>ей<br />

М д<strong>в</strong>е; <strong>с</strong>тебель М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; СиА прям<strong>а</strong>я, не<br />

<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть уз<strong>к</strong><strong>а</strong>я и длинн<strong>а</strong>я.<br />

В з<strong>а</strong>днем <strong>к</strong>рыле RS и М <strong>о</strong>бр<strong>а</strong>зуют неб<strong>о</strong>льшие<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Asientomum Martynov, 1926 (Lithentomum<br />

Martynov, 1926, пе<strong>с</strong>. Scudder, 1867; Lithopsocus<br />

Karny, 1930). Тип р<strong>о</strong>д<strong>а</strong> — Lithentomum praecox<br />

Martynov, 1926; IB. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). В переднем <strong>к</strong>рыле <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; R <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я у <strong>в</strong>ершины<br />

<strong>к</strong>рыл<strong>а</strong>; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>, из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> ме<strong>с</strong>те р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жения<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> r-rs и rs-mi+2;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я б<strong>а</strong>з<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>.<br />

В з<strong>а</strong>днем <strong>к</strong>рыле ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть R нея<strong>с</strong>н<strong>а</strong>;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS <strong>в</strong>ерти<strong>к</strong><strong>а</strong>льн<strong>о</strong>; М н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м<br />

пр<strong>о</strong>тяжении <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; СиА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, прям<strong>а</strong>я.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2 мм (ри<strong>с</strong>. 696). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО EMPHERIIDAE<br />

ENDERLEIN, 1911<br />

Крылья <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые, <strong>в</strong>ершины передних и<br />

з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> б<strong>о</strong>лее или менее <strong>о</strong><strong>к</strong>руглены.<br />

В переднем <strong>к</strong>рыле SC <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> R; СиА д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, CuP и А не <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я<br />

у <strong>в</strong>ершины <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е, <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные.<br />

У<strong>с</strong>и<strong>к</strong>и мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> неп<strong>о</strong><strong>с</strong>т<strong>о</strong>янным чи<strong>с</strong>л<strong>о</strong>м<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Empheriinae и Tapinellinae; п<strong>о</strong><strong>с</strong>леднее<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йден<strong>о</strong>.<br />

ПОДСЕМЕЙСТВО EMPHERIINAE KOLBE, 1884<br />

[nom. transl. Enderlein, 1908<br />

(ex Empheriidae Kolbe, 1884)]<br />

В переднем <strong>к</strong>рыле <strong>о</strong>дн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

r-rs; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА ДЛИННЫЙ И уз<strong>к</strong>ий, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

<strong>с</strong>теблем. В з<strong>а</strong>днем <strong>к</strong>рыле RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием М<br />

и CUA; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния R, М и СиА <strong>с</strong>литы. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2—4 мм (ри<strong>с</strong>. 697). П<strong>а</strong>ле<strong>о</strong>­<br />

692. Amphientomum paradoxum Pictet; A — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 13,5; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 13,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Enderlein, 1911).693. Electrentomum klebsianum Enderlein;<br />

A — переднее <strong>к</strong>рыл<strong>о</strong>, X 29,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 29,0;<br />

g _ з<strong>а</strong>дняя н<strong>о</strong>г<strong>а</strong>, х 150,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Enderlein, 1911). 694. Perientomum ceylonicum Enderlein;<br />

A — переднее <strong>к</strong>рыл<strong>о</strong>, X 20,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 20,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Enderlein, 1906). 695. Thylacella eversiana Enderlein; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 41,4; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 41,4; В — у<strong>с</strong>и<strong>к</strong>,<br />

Х41.4; Г — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong>, X 41,4; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Enderlein, 1911). 696. Asientomum praecox (Martynov);<br />

Ри<strong>с</strong>. 692—700. П<strong>о</strong>д<strong>о</strong>тряд Parapsocida<br />

232<br />

http://jurassic.ru/<br />

ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны;<br />

Empheria Hagen, 1856, Bebiosis Enderlein,<br />

1911, и <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО PSYLLIPSOCIDAE<br />

KOLBE, 1884<br />

[nom. transl. Enderlein, 1903<br />

(ex Psyllipsocini Kolbe, 1884)]<br />

Крылья и тел<strong>о</strong> не п<strong>о</strong><strong>к</strong>рыты чешуй<strong>к</strong><strong>а</strong>ми;<br />

<strong>в</strong>ет<strong>в</strong>ей М д<strong>в</strong>е — три; SC длинн<strong>а</strong>я, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong>з<strong>о</strong>р<strong>в</strong><strong>а</strong>нн<strong>а</strong>я, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R; CuP и<br />

А у <strong>в</strong>ершины <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я или <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближены.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 22—25. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong><strong>в</strong>ерху я<strong>с</strong>н<strong>о</strong> <strong>в</strong>идн<strong>а</strong>; <strong>с</strong>редне- и з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>а</strong> я<strong>с</strong>н<strong>о</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> три. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2,5—5,5 мм (ри<strong>с</strong>. 698). Не<strong>о</strong>ген<br />

— ныне. Четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из них <strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д из<strong>в</strong>е<strong>с</strong>тен из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Ю. Азии (янт<strong>а</strong>рь Бирмы).<br />

СЕМЕЙСТВО ARCHIPSYLLIDAE<br />

HANDLIRSCH, 1925<br />

Передние <strong>к</strong>рылья б<strong>о</strong>лее или менее <strong>в</strong>ытянутые;<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й SC нет: б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> С, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> — <strong>с</strong> R; R ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>,<br />

<strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет я<strong>с</strong>ную, шир<strong>о</strong><strong>к</strong>ую птер<strong>о</strong><strong>с</strong>тигму;<br />

RS длинн<strong>а</strong>я, <strong>с</strong> неб<strong>о</strong>льшим р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м,<br />

н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей М четыре;<br />

СиА п<strong>о</strong>л<strong>о</strong>г<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Archipsytta Handlirsch, 1907. Тип р<strong>о</strong>д<strong>а</strong>—Л.<br />

primitiva Handlirsch, 1907; н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния. Переднее <strong>к</strong>рыл<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е,<br />

длин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; RS<br />

н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; R<br />

<strong>о</strong>чень длинн<strong>а</strong>я, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>в</strong>перед <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> угл<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины'<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>с</strong>т<strong>в</strong><strong>о</strong>л СиА ДЛИННЫЙ, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> шир<strong>о</strong><strong>к</strong>ий,<br />

СиА 2 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

4,0—5,0 мм (ри<strong>с</strong>. 699). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из н. юры 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы и <strong>в</strong>. юры К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

А — переднее <strong>к</strong>рыл<strong>о</strong>, X 26,2; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 26,2, <strong>в</strong>. юр<strong>а</strong>,<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 697. Trichempheria yillosa<br />

Hagen; А — переднее <strong>к</strong>рыл<strong>о</strong>, X 36,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 36,0;<br />

В— з<strong>а</strong>дняя н<strong>о</strong>г<strong>а</strong>, х 100,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Enderlain, 1911). 698. Psyllipsocus banksi Cockerell; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 30,5; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 30,5; В— у<strong>с</strong>и<strong>к</strong>, X 30,5;<br />

Г — л<strong>а</strong>п<strong>к</strong><strong>а</strong>, х 100,0; Бирм<strong>а</strong>, янт<strong>а</strong>рь (Cockerell, 1916). 699. Агchipsylla<br />

turanica Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 11,8, <strong>в</strong>. юр<strong>а</strong>,<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 700. Archipsocus puber Hagen;<br />

А— переднее <strong>к</strong>рыл<strong>о</strong>, X 28,7; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 28,7; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Enderlein, 1911).

695А<br />

http://jurassic.ru/

СЕМЕЙСТВО ARCHIPSOCIDAE ENDERLEIN, 1903<br />

[nom. transl. Pearman, 1936<br />

(ex Archipsocinae Enderlein, 1903)]<br />

С<strong>а</strong>мцы' бе<strong>с</strong><strong>к</strong>рылые; мембр<strong>а</strong>н<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>бедненн<strong>о</strong>е:<br />

<strong>в</strong> переднем <strong>к</strong>рыле М н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дный<br />

и пл<strong>о</strong><strong>с</strong><strong>к</strong>ий; <strong>в</strong> з<strong>а</strong>днем <strong>к</strong>рыле СиА не <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 13. Переднегрудь <strong>с</strong><strong>в</strong>ерху<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>а</strong>. Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> у личин<strong>о</strong><strong>к</strong>,<br />

т<strong>а</strong><strong>к</strong> и у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых д<strong>в</strong><strong>а</strong>. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 1,5—1,6 мм (ри<strong>с</strong>. 700). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Три <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>а</strong>, из них <strong>о</strong>дин из<strong>в</strong>е<strong>с</strong>тен<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДОТРЯД EUPSOCIDA<br />

Крылья гетер<strong>о</strong>н<strong>о</strong>мные, з<strong>а</strong>дние зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че передних, <strong>с</strong> <strong>о</strong>бедненным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием.<br />

Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, <strong>в</strong>ет<strong>в</strong>и СиА нер<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

длины: СиА] зн<strong>а</strong>чительн<strong>о</strong> длиннее СиАг.<br />

Ан<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>а</strong>. У<strong>с</strong>и<strong>к</strong>и 13-члени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong><strong>а</strong>-три. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

Philotarsidae, Elipsqcidae, Epipsocidae,<br />

Caeciliidae, Peripsocidae, Psocidae.<br />

СЕМЕЙСТВО PHILOTARSIDAE PEARMAN, 1936<br />

Обе п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые. З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й<br />

передних <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>пушен д<strong>в</strong><strong>о</strong>йным ряд<strong>о</strong>м <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>.<br />

R и М передних и з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong><br />

<strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении. Л<strong>а</strong>п<strong>к</strong>и<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лых трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми<br />

г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физ. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

1,5—3,5 мм (ри<strong>с</strong>. 701). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>дин из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ELIPSOC1DAE PEARMAN, 1936<br />

Кр<strong>а</strong>я передних <strong>к</strong>рылье<strong>в</strong> и жил<strong>к</strong>и <strong>о</strong>пушены.<br />

З<strong>а</strong>дние <strong>к</strong>рылья не <strong>о</strong>пущены или <strong>о</strong>пущен<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> между <strong>в</strong>ет<strong>в</strong>ями RS. R и<br />

М передних <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м<br />

пр<strong>о</strong>тяжении, R и М з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й r-m. Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

д<strong>в</strong>ух- и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми<br />

г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физ. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,8—<br />

3,7 мм (ри<strong>с</strong>. 702 А, Б). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> пяти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>дин из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО EPIPSOCIDAE PEARMAN, 1936<br />

Крыл<strong>а</strong>тые и бе<strong>с</strong><strong>к</strong>рылые. Обе п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong>,<br />

е<strong>с</strong>ли имеют<strong>с</strong>я,— <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые. R и М передних<br />

<strong>к</strong>рылье<strong>в</strong> <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й r-m.<br />

R и М з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м<br />

пр<strong>о</strong>тяжении. С<strong>а</strong>м<strong>к</strong>и <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й — д<strong>в</strong>умя<br />

п<strong>а</strong>р<strong>а</strong>ми г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физ. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Epipsocinae, Neurostigminae, Gojinae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

неиз<strong>в</strong>е<strong>с</strong>тны 1<br />

.<br />

234<br />

http://jurassic.ru/<br />

ПОДСЕМЕЙСТВО EPIPSOCINAE ROESLER, 1940<br />

Ан<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> без <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>.<br />

С<strong>а</strong>м<strong>к</strong><strong>а</strong> бе<strong>с</strong><strong>к</strong>рыл<strong>а</strong>я. Lacinia <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью —<br />

де<strong>с</strong>ятью <strong>в</strong>ершинными зубц<strong>а</strong>ми. Н<strong>а</strong>ружные г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физЫ<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong> 5 мм. (ри<strong>с</strong>. 703). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CAECILIIDAE KOLBE, 1884<br />

Крылья <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые, ин<strong>о</strong>гд<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены<br />

у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>. R и М передних и з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong><br />

<strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении. Л<strong>а</strong>п<strong>к</strong>и<br />

д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Н<strong>а</strong>ружные г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физы! <strong>с</strong><strong>а</strong>м<strong>к</strong>и<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2—4 мм<br />

(ри<strong>с</strong>. 704 и 705). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

четыре р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), д<strong>в</strong><strong>а</strong> из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ымершие.<br />

СЕМЕЙСТВО PERIPSOCIDAE KOLBE, 1882<br />

[nom. transl. Pearman, 1936 (ex Peripsocinae Kolbe, 1882)]<br />

Крылья ин<strong>о</strong>гд<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, без<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. R и М передних <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е или н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении.<br />

СиА без <strong>в</strong>ет<strong>в</strong>ей. Л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. С<strong>а</strong>м<strong>к</strong>и<br />

<strong>с</strong> <strong>о</strong>дн<strong>о</strong>й — тремя п<strong>а</strong>р<strong>а</strong>ми г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физ. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2—4 мм (ри<strong>с</strong>. 706).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>дин из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Ю. Азии (<strong>к</strong><strong>о</strong>п<strong>а</strong>л<br />

Бирмы).<br />

СЕМЕЙСТВО PSOCIDAE LEACH, 1815<br />

[nom. transl. Stephans, 1836 (ex Psocida Leach, 1815)]<br />

Крылья ин<strong>о</strong>гд<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, без<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. R и М передних <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е или н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении<br />

или <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й r-m.<br />

CuAi <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> б<strong>о</strong>лее или менее б<strong>о</strong>льш<strong>о</strong>м<br />

пр<strong>о</strong>тяжении <strong>с</strong> М или <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> нею<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й m-cuai. Л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые-.<br />

Три п<strong>а</strong>ры г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физ у <strong>с</strong><strong>а</strong>м<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2,0—7,0 мм (ри<strong>с</strong>. 707). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны 1<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).

W 4 4 V 4 4<br />

CUP Cu'A ' ' 4<br />

701. Philotarsus antiauus Kplbe; A — переднее <strong>к</strong>рыл<strong>о</strong>, 25,0; Б —<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 25,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Enderlein, 1911). 702. Elipsocus abnormis Hagen; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 38,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 38,0; В— у<strong>с</strong>и<strong>к</strong>, X 38,0;<br />

Г— з<strong>а</strong>дняя н<strong>о</strong>г<strong>а</strong>, X 90,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> '(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Enderlein, 1911). 703. Epipsocus ciliaius Pictet et Hagen; A —<br />

переднее <strong>к</strong>рыл<strong>о</strong>, X 16,4; Б—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х?16,4; В—у<strong>с</strong>и<strong>к</strong>,<br />

X 6,4; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Enderlein, 1911).<br />

704. Caeeilius debeles Pictet et Hagen; А — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

CuP<br />

Ри<strong>с</strong>. 701—707. П<strong>о</strong>д<strong>о</strong>тряд Eupsocida<br />

CuA<br />

707 Б<br />

701А<br />

707А<br />

X 16,4; Б— з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 16,4; В—у<strong>с</strong>и<strong>к</strong>, X 16,4; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Enderlein, 1911). 705. Palaeopsocus<br />

tener Hagen; A —переднее <strong>к</strong>рыл<strong>о</strong>, X 41,8; Б—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

Х41,8; В — у<strong>с</strong>и<strong>к</strong>, Х41.8; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Enderlein, 1911). 706. Pcripsocus phaeopterus Stephens; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 18,6; Б— з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х18,6; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Enderlein,<br />

1927); 707. Psocus picteti Enderlein; A —переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 24,2; Б— з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 24,2; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Enderlein, 1911)<br />

http://jurassic.ru/

PSOCOPTERA INCERTAE SEDIS<br />

К эт<strong>о</strong>му <strong>о</strong>тряду <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые и<strong>с</strong><strong>к</strong><strong>о</strong>- ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и из <strong>о</strong>лиг<strong>о</strong>п<strong>а</strong>емы'е,<br />

т<strong>о</strong>чн<strong>о</strong>е <strong>о</strong>пределение <strong>к</strong><strong>о</strong>т<strong>о</strong>рых з<strong>а</strong>труд- цен<strong>а</strong> С. Амери<strong>к</strong>и и Lithentomum Scudder, 1867<br />

нительн<strong>о</strong>: Parapsocus Scudder, 1890 из п<strong>а</strong>ле<strong>о</strong>- из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.'<br />

Крылья <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

тел<strong>а</strong>. Передние <strong>к</strong>рылья <strong>о</strong><strong>в</strong><strong>а</strong>льные, тип<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылий:<br />

пл<strong>о</strong>тные, <strong>с</strong><strong>о</strong> <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й <strong>в</strong> <strong>в</strong>иде мел<strong>к</strong><strong>о</strong>й<br />

<strong>с</strong>ет<strong>к</strong>и; пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде <strong>в</strong>ыпу<strong>к</strong>лых<br />

ребер, я<strong>в</strong>ляющих<strong>с</strong>я <strong>к</strong><strong>а</strong><strong>к</strong> бы <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тью <strong>с</strong>ет<strong>к</strong>и; п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, треуг<strong>о</strong>льн<strong>о</strong>е; <strong>с</strong>имметричн<strong>о</strong><br />

ему р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть,<br />

<strong>о</strong>тделенн<strong>а</strong>я <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й. З<strong>а</strong>дние <strong>к</strong>рылья приблизительн<strong>о</strong><br />

т<strong>а</strong><strong>к</strong>их же р<strong>а</strong>змер<strong>о</strong><strong>в</strong> и <strong>о</strong>черт<strong>а</strong>ний, <strong>к</strong><strong>а</strong><strong>к</strong> и передние,<br />

н<strong>о</strong> т<strong>о</strong>н<strong>к</strong>ие, переп<strong>о</strong>нч<strong>а</strong>тые, <strong>с</strong> немн<strong>о</strong>гими<br />

жил<strong>к</strong><strong>а</strong>ми.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я; р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые прид<strong>а</strong>т<strong>к</strong>и<br />

жующег<strong>о</strong> тип<strong>а</strong>. Антенны <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из 12—14<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong>, ди<strong>а</strong>метр <strong>к</strong><strong>о</strong>т<strong>о</strong>рых рез<strong>к</strong><strong>о</strong> уменьш<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лении; члени<strong>к</strong>и <strong>в</strong><strong>о</strong><strong>о</strong>ружены<br />

<strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими щетин<strong>к</strong><strong>а</strong>ми.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong>едл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я.<br />

Н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; г<strong>о</strong>лени передних и <strong>с</strong>редних н<strong>о</strong>г<br />

<strong>с</strong> м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й; л<strong>а</strong>п<strong>к</strong>и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>е.<br />

Тергиты брюш<strong>к</strong><strong>а</strong> без <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м.<br />

П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, имеют<strong>с</strong>я <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие цер<strong>к</strong>и и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий<br />

яйце<strong>к</strong>л<strong>а</strong>д. Пермь. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> —<br />

Perielytridae.<br />

СЕМЕЙСТВО PERIELYTRIDAE G. ZALESSKY, 1948<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

рез<strong>к</strong><strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>я<strong>с</strong>ь п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. R т<strong>а</strong><strong>к</strong>же из<strong>о</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д,<br />

ОТРЯД PERIELYTRODEA<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

<strong>с</strong> неб<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей. RS <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й трети <strong>к</strong>рыл<strong>а</strong>. В <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле <strong>о</strong>дн<strong>а</strong> не<strong>в</strong>ет<strong>в</strong>ящ<strong>а</strong>я<strong>с</strong>я <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>.<br />

Пермь. Один р<strong>о</strong>д.<br />

Ри<strong>с</strong>. 708. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Perielytridae.<br />

708. Perielytron mirabile G. Zalessky (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); н. пермь,<br />

Ур<strong>а</strong>л (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

Perielytron Q. Zalessky, 1948. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. mirabile G. Zalessky, 1948; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. RS <strong>о</strong>бр<strong>а</strong>зует д<strong>в</strong>е-три <strong>в</strong>ет<strong>в</strong>и,<br />

теряющие <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<br />

жил<strong>о</strong><strong>к</strong> и <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я <strong>с</strong> <strong>с</strong>етью <strong>с</strong><strong>к</strong>ульптуры<br />

<strong>к</strong>рыл<strong>а</strong>; М <strong>о</strong>бр<strong>а</strong>зует три-четыре <strong>в</strong>ет<strong>в</strong>и, т<strong>а</strong><strong>к</strong>же<br />

теряющие<strong>с</strong>я <strong>с</strong>реди <strong>с</strong>ет<strong>к</strong>и. СиА гребенч<strong>а</strong>т<strong>а</strong>я,<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

7—8 мм (ри<strong>с</strong>. 708; т<strong>а</strong>бл. XVI, фиг. З) 1<br />

. Один<br />

<strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

1<br />

В 1959 г. <strong>в</strong> Че<strong>к</strong><strong>а</strong>рде мн<strong>о</strong>ю были н<strong>а</strong>йдены д<strong>в</strong><strong>а</strong> н<strong>о</strong><strong>в</strong>ых<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong> Perielytron, п<strong>о</strong>з<strong>в</strong><strong>о</strong>ли<strong>в</strong>шие д<strong>а</strong>ть б<strong>о</strong>лее дет<strong>а</strong>льную<br />

ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>цию эт<strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>, чем эт<strong>о</strong> <strong>с</strong>дел<strong>а</strong>н<strong>о</strong><br />

<strong>в</strong> р<strong>а</strong>б<strong>о</strong>те Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> (1948).<br />

НАДОТРЯД THYSANOPTEROIDEA.<br />

ТРИПСООБРАЗНЫЕ<br />

Тел<strong>о</strong> рез<strong>к</strong><strong>о</strong> удлиненн<strong>о</strong>е. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны<br />

<strong>к</strong><strong>о</strong>люще-<strong>с</strong><strong>о</strong><strong>с</strong>ущег<strong>о</strong> тип<strong>а</strong>; <strong>с</strong>тр<strong>о</strong>ение их <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я,<br />

длинн<strong>а</strong>я, гл<strong>а</strong>з<strong>а</strong> м<strong>а</strong>лы 1<br />

. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, из<br />

немн<strong>о</strong>гих члени<strong>к</strong><strong>о</strong><strong>в</strong>, чет<strong>к</strong><strong>о</strong><strong>в</strong>идные. Крылья уз<strong>к</strong>ие<br />

и длинные, п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные и п<strong>о</strong>-'<br />

<strong>к</strong>рытые п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми: пер<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ть<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>бед­<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

236<br />

http://jurassic.ru/<br />

ненн<strong>о</strong>е — не б<strong>о</strong>лее д<strong>в</strong>ух пр<strong>о</strong>д<strong>о</strong>льных жил<strong>о</strong><strong>к</strong>.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения нея<strong>с</strong>ны. И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong>е<br />

<strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зие <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции трип<strong>с</strong><strong>о</strong><strong>в</strong><br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет <strong>с</strong><strong>о</strong>мне<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong> их прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ти<br />

<strong>к</strong> п<strong>а</strong>р<strong>а</strong>не<strong>о</strong>птер<strong>а</strong>м, <strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым <strong>о</strong>ни <strong>в</strong><strong>с</strong>ет<strong>а</strong><strong>к</strong>и<br />

ближе, чем <strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong>им-либ<strong>о</strong> другим н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мым.<br />

Пермь — ныне. Отряд Thysanoptera.

ОТРЯД THYSANOPTERA. ПУЗЫРЕНОГИЕ, ИЛИ ТРИПСЫ<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые (0,5—5 мм длин<strong>о</strong>й).<br />

Крылья <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong> н<strong>а</strong> <strong>с</strong>пине; мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы. П<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> не б<strong>о</strong>лее пяти; длинные <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуют б<strong>а</strong>хр<strong>о</strong>му; ин<strong>о</strong>гд<strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть н<strong>а</strong> з<strong>а</strong>дних <strong>к</strong>рыльях. Р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>к</strong><strong>о</strong>люще-<strong>с</strong><strong>о</strong><strong>с</strong>ущег<strong>о</strong> тип<strong>а</strong>, <strong>а</strong><strong>с</strong>имметричный;<br />

ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> и <strong>о</strong>бычн<strong>о</strong> три гл<strong>а</strong>з<strong>к</strong><strong>а</strong>,<br />

<strong>а</strong>нтенны 1<br />

длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, 13 члени<strong>к</strong><strong>о</strong><strong>в</strong>. Н<strong>о</strong>ги<br />

бег<strong>а</strong>тельные; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>о</strong>дин — д<strong>в</strong><strong>а</strong>; меж<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> изменен<strong>а</strong> <strong>в</strong> пузыре<strong>к</strong><br />

(при<strong>с</strong><strong>а</strong><strong>с</strong>ы<strong>в</strong><strong>а</strong>тельн<strong>о</strong>е при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бление). Сегмент<strong>о</strong><strong>в</strong><br />

брюш<strong>к</strong><strong>а</strong> де<strong>с</strong>ять.<br />

Жи<strong>в</strong>ут н<strong>а</strong> р<strong>а</strong>зличных р<strong>а</strong><strong>с</strong>тениях, <strong>с</strong><strong>о</strong>ц<strong>в</strong>етиях,<br />

ц<strong>в</strong>ет<strong>а</strong>х, ли<strong>с</strong>тьях, гниющих р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>х.<br />

Пит<strong>а</strong>ют<strong>с</strong>я <strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>ми р<strong>а</strong><strong>с</strong>тений (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>в</strong>иды <strong>в</strong>редят), реже хищни<strong>к</strong>и. Пермь — ныне.<br />

П<strong>о</strong>д<strong>о</strong>тряды: Terebrantia, Tubulifera.<br />

ПОДОТРЯД TEREBRANTIA.<br />

^ЯЙЦЕКЛАДНЫЕ ТРИПСЫ<br />

Крыл<strong>а</strong>т<strong>о</strong>е ф<strong>о</strong>рмы; передние <strong>к</strong>рылья длиннее<br />

з<strong>а</strong>дних; <strong>в</strong> передних <strong>к</strong>рыльях <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>дн<strong>а</strong>д<strong>в</strong>е<br />

пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и; мембр<strong>а</strong>н<strong>а</strong> п<strong>о</strong><strong>к</strong>рыт<strong>а</strong><br />

ми<strong>к</strong>р<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>пиче<strong>с</strong><strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. П<strong>о</strong><strong>с</strong>ледний<br />

<strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ий, <strong>с</strong> яйце­<br />

709. Permothrips longipennis Martynov; <strong>о</strong>бщий <strong>в</strong>ид, X 8,0, н.<br />

пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 710. Liassothrtps crassips (Martynov);<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 18,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1927). 711. Archankothrips pugionifer Priesner; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 50,0; Б — у<strong>с</strong>и<strong>к</strong>и,X 50,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Priesner, 1924). 712. Protothrips speratus Priesner;<br />

<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м, у <strong>с</strong><strong>а</strong>мц<strong>а</strong> шир<strong>о</strong><strong>к</strong>ий, з<strong>а</strong><strong>к</strong>ругленный.<br />

Пермь — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Permothripidae,<br />

Liassothripidae, Aeolothripidae, Heterothripidae,<br />

Thripidae, Merothripidae.<br />

СЕМЕЙСТВО PERMOTHRIPIDAE<br />

MARTYNOV, 1935<br />

Сл<strong>о</strong>женные <strong>к</strong>рылья длиннее брюш<strong>к</strong><strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>; передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>уж<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong>переди, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, длин<strong>а</strong> ее меньше п<strong>о</strong>­<br />

л<strong>о</strong><strong>в</strong>ины 1<br />

длины птер<strong>о</strong>т<strong>о</strong>р<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong>; н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

Пермь. Один р<strong>о</strong>д.<br />

Permothrips Martynov, 1935. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. longipennis Martynov, 1935; н. пермь, Перм<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я, четырехуг<strong>о</strong>льн<strong>а</strong>я; <strong>а</strong>нтенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

и т<strong>о</strong>л<strong>с</strong>тые. Бедр<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>здуты 1<br />

, г<strong>о</strong>лени<br />

т<strong>о</strong>н<strong>к</strong>ие и длиннее их. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3 мм<br />

(ри<strong>с</strong>. 709). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Ри<strong>с</strong>. 709—715. Отряд Thysanoptera<br />

СЕМЕЙСТВО LIASSOTHRIPIDAE<br />

PRIESNER, 1949<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> уз<strong>к</strong><strong>а</strong>я; <strong>а</strong>нтенны т<strong>о</strong>н<strong>к</strong>ие, из <strong>с</strong>еми —<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьми члени<strong>к</strong><strong>о</strong><strong>в</strong>. Передние бедр<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширены,<br />

г<strong>о</strong>лени т<strong>о</strong>н<strong>к</strong>ие. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> и переднегрудь, X 40,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

237<br />

http://jurassic.ru/

Liassothrips Priesner, 1949. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Mesothrips crassicornis Martynov, 1927; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Крылья<br />

не <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

и <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. Длин<strong>а</strong> тел<strong>а</strong> 2—2,5 мм<br />

(ри<strong>с</strong>. 710). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО AEOLOTHRIPIDAE HALIDAY, 1836<br />

Вершин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен<strong>а</strong>,<br />

д<strong>в</strong>е пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и, <strong>о</strong>т трех д<strong>о</strong> пяти<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн де<strong>в</strong>ять.<br />

Яйце<strong>к</strong>л<strong>а</strong>д з<strong>а</strong>гнут <strong>к</strong><strong>в</strong>ерху (ри<strong>с</strong>. 711). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и,<br />

из них четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong>ымершие.<br />

СЕМЕЙСТВО HETEROTHRIPIDAE BAGNAL, 1912<br />

Крылья з<strong>а</strong><strong>о</strong><strong>с</strong>трены; <strong>в</strong> передних <strong>к</strong>рыльях д<strong>в</strong>е<br />

пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и, п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> нет.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн де<strong>в</strong>ять, ин<strong>о</strong>гд<strong>а</strong> де<strong>с</strong>ять; третий<br />

и чет<strong>в</strong>ертый члени<strong>к</strong>и длиннее <strong>о</strong><strong>с</strong>т<strong>а</strong>льных,<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зны; н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем члени<strong>к</strong>е з<strong>а</strong><strong>о</strong><strong>с</strong>тренный<br />

грифеле<strong>к</strong>. Яйце<strong>к</strong>л<strong>а</strong>д н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен <strong>в</strong>низ.<br />

К<strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> н<strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>м члени<strong>к</strong>е<br />

передней л<strong>а</strong>п<strong>к</strong>и (ри<strong>с</strong>. 712). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне и <strong>о</strong>дин<br />

<strong>в</strong>ымерший р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО THR1PIDAE STEPHENS, 1829<br />

Крылья уз<strong>к</strong>ие, з<strong>а</strong><strong>о</strong><strong>с</strong>тренные; п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> нет. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн <strong>о</strong>т ше<strong>с</strong>ти д<strong>о</strong><br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьми; п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> <strong>с</strong> шип<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны'м <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>м.<br />

Яйце<strong>к</strong>л<strong>а</strong>д з<strong>а</strong>гнут <strong>в</strong>низ (ри<strong>с</strong>. 713).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды<br />

СЕМЕЙСТВО UNINERVIDAE<br />

BECKER-MIGDISOVA, 1951<br />

Крылья п<strong>о</strong>чти <strong>с</strong>тр<strong>о</strong>г<strong>о</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, уз<strong>к</strong>ие,<br />

длинные, <strong>в</strong> <strong>в</strong>ершине <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужены и <strong>о</strong><strong>к</strong>руглены:<br />

переп<strong>о</strong>н<strong>к</strong><strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong><strong>а</strong>я. Н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong>дн<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

<strong>о</strong>тх<strong>о</strong>дят четыре з<strong>а</strong>дние <strong>в</strong>ет<strong>в</strong>и, идущие<br />

<strong>к</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. Р<strong>а</strong>змеры 1<br />

PARANEOPTERA INCERTAE SEDIS<br />

(Е. Э. Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

мел<strong>к</strong>ие — <strong>к</strong>рылья <strong>о</strong>т<br />

2 д<strong>о</strong> 3 мм. Стр<strong>о</strong>ение тел<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

В. пермь — <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

238<br />

http://jurassic.ru/<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), из них<br />

четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong>ымершие.<br />

СЕМЕЙСТВО MEROTHRIPIDAE HOOD, 1912<br />

Крылья з<strong>а</strong><strong>о</strong><strong>с</strong>трены; д<strong>в</strong>е пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и,<br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> нет. Антенны чет<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, без з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>г<strong>о</strong> грифель<strong>к</strong><strong>а</strong><br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце. Передние и з<strong>а</strong>дние бедр<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><br />

ут<strong>о</strong>лщены. Яйце<strong>к</strong>л<strong>а</strong>д редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н (ри<strong>с</strong>. 714).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы* (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь), из них <strong>о</strong>дин <strong>в</strong>ымерший.<br />

ПОДОТРЯД TUBULIFERA.<br />

ТРУБКОХВОСТЫЕ ТРИПСЫ<br />

Крыл<strong>а</strong>тые и бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы. Передние<br />

и з<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>дн<strong>о</strong>й длины; <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й жил<strong>к</strong>и<br />

нет; пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>; н<strong>а</strong><br />

мембр<strong>а</strong>не ми<strong>к</strong>р<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong> нет. П<strong>о</strong><strong>с</strong>ледний<br />

<strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> труб<strong>к</strong><strong>о</strong><strong>в</strong>идный; яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong><br />

нет. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> РЫ<strong>о</strong>eothripidae.<br />

СЕМЕЙСТВО PHLOEOTHRIP1DAE HALIDAY, 1836<br />

Крыл<strong>а</strong>тые ф<strong>о</strong>рмы. Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн <strong>с</strong>емь —<br />

<strong>в</strong><strong>о</strong><strong>с</strong>емь; нижнечелю<strong>с</strong>тные щупи<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Средние т<strong>а</strong>зи<strong>к</strong>и н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

<strong>с</strong>ильнее, чем т<strong>а</strong>зи<strong>к</strong>и д<strong>в</strong>ух других п<strong>а</strong>р н<strong>о</strong>г.<br />

П<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong> — <strong>с</strong>уженн<strong>а</strong>я<br />

<strong>к</strong> <strong>к</strong><strong>о</strong>нцу труб<strong>к</strong><strong>а</strong>: ширин<strong>а</strong> де<strong>в</strong>ят<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

немн<strong>о</strong>г<strong>о</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ег<strong>о</strong> длину (ри<strong>с</strong>. 715).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и, из них пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong>ымершие.<br />

Uninervus Becker-Migdisova, 1951.<br />

Тип р<strong>о</strong>д<strong>а</strong> — U. zorapteroides Becker-Migdisova,<br />

1951; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). Передние <strong>к</strong>рылья<br />

л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идн<strong>о</strong>й ф<strong>о</strong>рмы, передний их <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти<br />

прям<strong>о</strong>й. П<strong>о</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> пр<strong>о</strong>х<strong>о</strong>дит пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>с</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>нн<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дящими <strong>о</strong>т нее RS,<br />

М, СиА и СиР; ЖИЛКИ R и RS <strong>о</strong>бр<strong>а</strong>зуют р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющий <strong>в</strong> <strong>в</strong>ершину <strong>к</strong>рыл<strong>а</strong>. З<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong> немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че переднег<strong>о</strong>: р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>

R — RS зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3 мм (ри<strong>с</strong>. 716). Один <strong>в</strong>ид.<br />

В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Mononeura Becker-Migdisova, 1951.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. angustipiennis Becker-Migdisova,<br />

1951; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong> <strong>с</strong>ередине <strong>в</strong>ыпу<strong>к</strong>лый, <strong>с</strong> ут<strong>о</strong>лщенн<strong>о</strong>й С; <strong>с</strong>рединн<strong>а</strong>я<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>с</strong>д<strong>в</strong>инут<strong>а</strong> <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; М <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т нее п<strong>о</strong>чти п<strong>о</strong>д прямым<br />

угл<strong>о</strong>м; CuA н<strong>а</strong><strong>к</strong>л<strong>о</strong>нн<strong>а</strong>я. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2—3 мм (ри<strong>с</strong>. 717). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong><br />

Ср. Азии.<br />

Ри<strong>с</strong>. 716—717. Paraneoptera incertae sedis<br />

716. Uninervus zorapieroldes Becker-Migdisova; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 22,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 22,0; <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1951). 717. Mononeura angustipennis<br />

Becker-Migdisova; <strong>к</strong>рылья, X 24,0, <strong>в</strong>. трн<strong>а</strong><strong>с</strong>, Ср. Азия<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1951).<br />

716Б<br />

КОГОРТА HOLOMETABOLA (OLIGONEOPTERA)<br />

НАСЕКОМЫЕ С ПОЛНЫМ ПРЕВРАЩЕНИЕМ,<br />

ИЛИ МАЛОЖИЛКОВЫЕ НОВОКРЫЛЫЕ<br />

Юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>а</strong><br />

и не<strong>с</strong>ет <strong>о</strong>дну пр<strong>о</strong><strong>с</strong>тую, не<strong>в</strong>ет<strong>в</strong>ящую<strong>с</strong>я жил<strong>к</strong>у.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние и ф<strong>о</strong>рм<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>чень р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны:<br />

из<strong>в</strong>е<strong>с</strong>тны преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> р<strong>а</strong>зличные<br />

ф<strong>о</strong>рмы гетер<strong>о</strong>н<strong>о</strong>мии <strong>к</strong>рылье<strong>в</strong>, н<strong>о</strong> мн<strong>о</strong>г<strong>о</strong> и п<strong>о</strong>чти<br />

г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мн<strong>о</strong><strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е грызущий,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>в</strong>ид<strong>о</strong>измененный <strong>в</strong> р<strong>а</strong>зных групп<strong>а</strong>х<br />

<strong>в</strong> р<strong>а</strong>зличные ф<strong>о</strong>рмы <strong>с</strong><strong>о</strong><strong>с</strong>ущих р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> п<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м<br />

н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong><strong>к</strong><strong>о</strong>ящ<strong>а</strong>я<strong>с</strong>я <strong>с</strong>т<strong>а</strong>дия, <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong>,<br />

не приним<strong>а</strong>ющ<strong>а</strong>я пищи и <strong>о</strong>бычн<strong>о</strong> м<strong>а</strong>л<strong>о</strong>п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я.<br />

Р<strong>а</strong>нние <strong>с</strong>т<strong>а</strong>дии р<strong>а</strong>з<strong>в</strong>ития <strong>в</strong> <strong>в</strong>иде<br />

личин<strong>о</strong><strong>к</strong>, <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ющих рез<strong>к</strong><strong>о</strong> <strong>о</strong>тличн<strong>о</strong>й <strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мы'х <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>цией: <strong>в</strong> их <strong>с</strong>тр<strong>о</strong>ении<br />

имеют<strong>с</strong>я р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные личин<strong>о</strong>чные при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления,<br />

<strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>ленные <strong>о</strong>бит<strong>а</strong>нием эт<strong>о</strong>й <strong>с</strong>т<strong>а</strong>дии<br />

<strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бых у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях, п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>тличных<br />

<strong>о</strong>т т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н —<br />

ныне. Четыре н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>: Coleopteroidea,<br />

Neuropteroidea, Mecopteroidea, Hymenopteroidea.<br />

Пер<strong>в</strong>ый пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель <strong>к</strong><strong>о</strong>г<strong>о</strong>рты 1<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

(<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н-<br />

ниц<strong>а</strong> Metropator pusillus Handlirsch), <strong>о</strong>бн<strong>а</strong>руженный<br />

<strong>в</strong> <strong>о</strong>дн<strong>о</strong>м из <strong>с</strong><strong>а</strong>мых дре<strong>в</strong>них ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> нижнег<strong>о</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Аме­<br />

239<br />

ри<strong>к</strong>и, д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р еще <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong> изучен и из<strong>в</strong>е<strong>с</strong>тен<br />

лишь п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>му пл<strong>о</strong>х<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шему<strong>с</strong>я<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong>у <strong>к</strong>рыл<strong>а</strong>. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> н. перми мы <strong>в</strong><strong>с</strong>треч<strong>а</strong>ем<strong>с</strong>я<br />

<strong>в</strong>пер<strong>в</strong>ые <strong>с</strong> р<strong>а</strong>зличными групп<strong>а</strong>ми н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нницы,<br />

же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые и <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные; н<strong>а</strong>йдены<br />

т<strong>а</strong><strong>к</strong>же и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые другие нижнеперм<strong>с</strong><strong>к</strong>ие<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители групп, д<strong>о</strong><strong>с</strong>тигшие б<strong>о</strong>льшег<strong>о</strong><br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зия лишь <strong>с</strong> <strong>в</strong>ерхней перми. С н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

мез<strong>о</strong>з<strong>о</strong>я п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong> три<strong>а</strong><strong>с</strong>е — д<strong>в</strong>у<strong>к</strong>рылые и<br />

<strong>с</strong> нижней юры — переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые. Уже<br />

<strong>с</strong> юры н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong><strong>а</strong>мыми мн<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными и з<strong>а</strong>ним<strong>а</strong>ют<br />

гл<strong>а</strong><strong>в</strong>ен<strong>с</strong>т<strong>в</strong>ующее ме<strong>с</strong>т<strong>о</strong> <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х <strong>к</strong><strong>а</strong><strong>к</strong> п<strong>о</strong> чи<strong>с</strong>лу <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

единиц, т<strong>а</strong><strong>к</strong> и п<strong>о</strong> <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>у инди<strong>в</strong>ид<strong>о</strong><strong>в</strong>. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие<br />

<strong>о</strong>тн<strong>о</strong>шения эт<strong>о</strong>й <strong>к</strong><strong>о</strong>г<strong>о</strong>рты <strong>с</strong> другими<br />

д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью не я<strong>с</strong>ны. Отн<strong>о</strong>шения <strong>о</strong>тдельных<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя н<strong>а</strong>меч<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>бщих черт<strong>а</strong>х. Дре<strong>в</strong>ние ме<strong>к</strong><strong>о</strong>птер<strong>о</strong>иды,<br />

именн<strong>о</strong> нижнеперм<strong>с</strong><strong>к</strong>ие Protomecoptera,<br />

я<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>в</strong>язы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>я<strong>с</strong>ь, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, их дери<strong>в</strong><strong>а</strong>т<strong>а</strong>ми. Нея<strong>с</strong>ны<br />

и<strong>с</strong>х<strong>о</strong>дные ф<strong>о</strong>рмы для <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

и же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных: <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие д<strong>а</strong>нных п<strong>о</strong><br />

http://jurassic.ru/

Ри<strong>с</strong>. 718. Схем<strong>а</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений <strong>о</strong>тряд<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением (Holometabola)<br />

Н<strong>а</strong> ф<strong>о</strong>не ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>л<strong>о</strong>н<strong>к</strong>и п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>о</strong> <strong>в</strong>ремя п<strong>о</strong>я<strong>в</strong>ления, р<strong>а</strong>з- tera, Neu — Neuroptera.'-R — Raphidioptera, St — Strepsiptera,<br />

<strong>в</strong>ития и р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нения <strong>о</strong>тдельных <strong>о</strong>тряд<strong>о</strong><strong>в</strong>. С<strong>о</strong><strong>к</strong>р<strong>а</strong>щения: Aph— Tr — Trichoptera, CI —JColeopteroidea, М<strong>с</strong> — Mecopteroidena<br />

Aphaniptera, Col — Coleoptera, Dip — Diptera, Hym — Hyme- (Р<strong>о</strong>денд<strong>о</strong>рф и др., 1961)<br />

noptera, Lep — Lepidoptera, Mec — Mecoptera, Mgl — Megalop-<br />

дре<strong>в</strong>нейшим ф<strong>а</strong>ун<strong>а</strong>м и глуб<strong>о</strong><strong>к</strong>ие р<strong>а</strong>зличия этих<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют ме<strong>с</strong>т<strong>о</strong> д<strong>а</strong>же для <strong>с</strong><strong>о</strong>мнений<br />

<strong>в</strong> их р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>е. Б<strong>о</strong>лее я<strong>с</strong>ны <strong>с</strong><strong>в</strong>язи переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

<strong>с</strong> другими н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мыми<br />

<strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением, именн<strong>о</strong> <strong>с</strong> <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными;<br />

<strong>в</strong>ы<strong>с</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь д<strong>а</strong>же предп<strong>о</strong>л<strong>о</strong>жения<br />

<strong>о</strong> неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>м р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>е переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых<br />

<strong>с</strong> <strong>о</strong>тряд<strong>о</strong>м <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц, чт<strong>о</strong>,<br />

<strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong>, не<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong> <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем ин<strong>о</strong>г<strong>о</strong><br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>а</strong> <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ции лет<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong><br />

(р<strong>а</strong>нней диптеригии) и <strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых. В цел<strong>о</strong>м<br />

и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> п<strong>о</strong>лным<br />

пре<strong>в</strong>р<strong>а</strong>щением <strong>о</strong>пределил<strong>о</strong><strong>с</strong>ь <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

<strong>о</strong>нт<strong>о</strong>генез<strong>а</strong>. Выр<strong>а</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong> п<strong>о</strong>лн<strong>о</strong>г<strong>о</strong> пре<strong>в</strong>р<strong>а</strong>щения,<br />

при <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м личин<strong>к</strong>и <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бны <strong>о</strong>бит<strong>а</strong>ть<br />

<strong>в</strong> <strong>с</strong><strong>а</strong>мых р<strong>а</strong>зных у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях, д<strong>а</strong>ле<strong>к</strong>их <strong>о</strong>т у<strong>с</strong>л<strong>о</strong><strong>в</strong>ий<br />

<strong>о</strong>бит<strong>а</strong>ния <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм,— <strong>в</strong><strong>с</strong>е эт<strong>о</strong> <strong>с</strong>дел<strong>а</strong>л<strong>о</strong><br />

г<strong>о</strong>л<strong>о</strong>мет<strong>а</strong>б<strong>о</strong>лию <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенным при<strong>о</strong>бретением<br />

<strong>в</strong> э<strong>в</strong><strong>о</strong>люции этих н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и д<strong>а</strong>л<strong>о</strong> им<br />

и<strong>с</strong><strong>к</strong>лючительные премуще<strong>с</strong>т<strong>в</strong><strong>а</strong> п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нению<br />

<strong>с</strong> другими, тем <strong>с</strong><strong>а</strong>мым <strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>и<strong>в</strong> их шир<strong>о</strong>ч<strong>а</strong>йший<br />

р<strong>а</strong><strong>с</strong>ц<strong>в</strong>ет.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения <strong>в</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>рте<br />

Holometabola п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ны н<strong>а</strong> ри<strong>с</strong>. 718.<br />

НАДОТРЯД COLEOPTEROIDEA.<br />

ЖЕСТКОКРЫЛООБРАЗНЫЕ<br />

Очень рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> гетер<strong>о</strong>н<strong>о</strong>мия <strong>о</strong>б<strong>о</strong>их<br />

п<strong>а</strong>р <strong>к</strong>рылье<strong>в</strong>. Передние <strong>к</strong>рылья пре<strong>в</strong>р<strong>а</strong>щены<br />

<strong>в</strong> пл<strong>о</strong>тные п<strong>о</strong><strong>к</strong>рыш<strong>к</strong>и (элитры или н<strong>а</strong>д<strong>к</strong>рылья),<br />

п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем не <strong>с</strong><strong>о</strong><strong>в</strong>ерш<strong>а</strong>ющие <strong>в</strong>зм<strong>а</strong>х<strong>а</strong> <strong>в</strong> пр<strong>о</strong>­<br />

240<br />

http://jurassic.ru/<br />

це<strong>с</strong><strong>с</strong>е п<strong>о</strong>лет<strong>а</strong>; ин<strong>о</strong>гд<strong>а</strong> передние <strong>к</strong>рылья п<strong>о</strong>чти<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних<br />

<strong>к</strong>рылье<strong>в</strong> <strong>в</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е <strong>с</strong>луч<strong>а</strong>е<strong>в</strong> нер<strong>а</strong>зличим<strong>о</strong>е,<br />

реже <strong>в</strong> <strong>в</strong>иде <strong>к</strong>реп<strong>к</strong>их, м<strong>а</strong>л<strong>о</strong><strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я, п<strong>а</strong>р<strong>а</strong>л-

лельных жил<strong>о</strong><strong>к</strong>. З<strong>а</strong>дние <strong>к</strong>рылья переп<strong>о</strong>нч<strong>а</strong>тые,<br />

<strong>с</strong> чет<strong>к</strong>им жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием, <strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

и я<strong>в</strong>ляющие<strong>с</strong>я <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ным <strong>о</strong>рг<strong>а</strong>н<strong>о</strong>м<br />

<strong>в</strong>зм<strong>а</strong>х<strong>а</strong> <strong>в</strong> п<strong>о</strong>лете. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны грызущие,<br />

примити<strong>в</strong>н<strong>о</strong> п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енные, ред<strong>к</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong>измененные<br />

и <strong>в</strong> т<strong>о</strong>й или друг<strong>о</strong>й <strong>с</strong>тепени редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные.<br />

Личин<strong>к</strong>и <strong>к</strong><strong>а</strong>мп<strong>о</strong>дее<strong>в</strong>идные, реже эру<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

<strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й, <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми н<strong>о</strong>г и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

дых<strong>а</strong>льц<strong>а</strong>ми. Обит<strong>а</strong>ют <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х,<br />

<strong>в</strong> т<strong>к</strong><strong>а</strong>нях р<strong>а</strong><strong>с</strong>тений и гниющих <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong>х;<br />

реже жи<strong>в</strong>ут <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong> н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях или<br />

п<strong>а</strong>р<strong>а</strong>зитируют у р<strong>а</strong>зличных жи<strong>в</strong><strong>о</strong>тных.<br />

Н. пермь — ныне. Д<strong>в</strong><strong>а</strong> <strong>о</strong>тряд<strong>а</strong>: Coleoptera,<br />

Strepsiptera.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>в</strong>язи <strong>с</strong> другими н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>ми<br />

д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р не <strong>в</strong>ыя<strong>с</strong>нены. Же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные<br />

я<strong>в</strong>ляют<strong>с</strong>я н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

групп<strong>о</strong>й <strong>с</strong>реди <strong>в</strong><strong>с</strong>ех н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> п<strong>о</strong>лным<br />

пре<strong>в</strong>р<strong>а</strong>щением. Они х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>бым<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лением и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития — <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ением<br />

<strong>к</strong>р<strong>а</strong>йних ф<strong>о</strong>рм п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ных при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блений<br />

<strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении <strong>к</strong>рылье<strong>в</strong> и <strong>в</strong><strong>с</strong>ег<strong>о</strong> тел<strong>а</strong>.<br />

П<strong>о</strong>д<strong>о</strong>бные при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены<br />

и <strong>в</strong> и<strong>с</strong>т<strong>о</strong>рии <strong>о</strong>б<strong>о</strong>их других <strong>к</strong><strong>о</strong>г<strong>о</strong>рт н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> у Polyneoptera (Blattodea, Dermaptera<br />

и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых других), т<strong>а</strong><strong>к</strong> и у Paraneoptera<br />

(<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у Heteroptera). Н<strong>о</strong> у<br />

Holometabola п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ть р<strong>а</strong>з<strong>в</strong>ил<strong>а</strong><strong>с</strong>ь<br />

ц<strong>о</strong>чти и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> у же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых. В<strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ение<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>с</strong>ти у н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением <strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>л<strong>о</strong><strong>с</strong>ь <strong>к</strong>р<strong>а</strong>йне<br />

п<strong>о</strong>лезным при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блением <strong>в</strong> э<strong>в</strong><strong>о</strong>люции этих<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>о</strong>бе<strong>с</strong>печи<strong>в</strong>шим же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылым шир<strong>о</strong>ч<strong>а</strong>йшее<br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зие:<br />

ныне <strong>о</strong>тряд Coleoptera н<strong>а</strong>иб<strong>о</strong>лее р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

и р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненный <strong>с</strong>реди <strong>в</strong><strong>с</strong>ех н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

ОТРЯД COLEOPTERA. ЖЕСТКОКРЫЛЫЕ, ИЛИ ЖУКИ<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф, А. Г. П<strong>о</strong>н<strong>о</strong>м<strong>а</strong>рен<strong>к</strong><strong>о</strong>)<br />

Упл<strong>о</strong>тненные передние <strong>к</strong>рылья, р<strong>о</strong><strong>в</strong>н<strong>о</strong> <strong>с</strong><strong>о</strong>- прит<strong>о</strong>м <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> н<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>я<strong>с</strong>ь<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ими з<strong>а</strong>дними <strong>к</strong>р<strong>а</strong>ями, з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>- нительн<strong>о</strong>-м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниях и не<br />

ют з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong>у, <strong>в</strong>ерхнюю п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть брюш<strong>к</strong><strong>а</strong> п<strong>о</strong>д<strong>к</strong>реплял<strong>с</strong>я д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ими<br />

и переп<strong>о</strong>нч<strong>а</strong>тые з<strong>а</strong>дние <strong>к</strong>рылья. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong><strong>а</strong>ми. Же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые <strong>в</strong><strong>о</strong>зни<strong>к</strong>ли,<br />

грызущие. Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> неб<strong>о</strong>льшие. Грудн<strong>о</strong>й <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>м пери<strong>о</strong>де <strong>о</strong>т<br />

<strong>о</strong>тдел <strong>в</strong>ели<strong>к</strong>: переднегрудь <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен- <strong>к</strong><strong>а</strong><strong>к</strong>их-т<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зных <strong>о</strong>рт<strong>о</strong>птер<strong>о</strong>ид<strong>о</strong><strong>в</strong>, пред<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>о</strong>й <strong>с</strong>пин<strong>к</strong><strong>о</strong>й, <strong>с</strong>реднегрудь неб<strong>о</strong>льш<strong>а</strong>я и <strong>с</strong>лит<strong>а</strong>я <strong>в</strong><strong>с</strong>ех н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением. Эти<br />

<strong>с</strong> з<strong>а</strong>днегрудью, <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ющей б<strong>о</strong>льшим <strong>с</strong>тер- и<strong>с</strong>х<strong>о</strong>дные ф<strong>о</strong>рмы н<strong>а</strong>ряду <strong>с</strong> <strong>в</strong>ыр<strong>а</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>й п<strong>о</strong>лн<strong>о</strong>н<strong>а</strong>льным<br />

<strong>о</strong>тдел<strong>о</strong>м. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из р<strong>а</strong>злич- г<strong>о</strong> пре<strong>в</strong>р<strong>а</strong>щения при<strong>о</strong>брели <strong>с</strong><strong>о</strong><strong>в</strong>ершенные п<strong>о</strong>н<strong>о</strong>г<strong>о</strong><br />

чи<strong>с</strong>л<strong>а</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и лишен<strong>о</strong> длинных при- <strong>к</strong>р<strong>о</strong><strong>в</strong>ные при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления, н<strong>а</strong>д<strong>к</strong>рылья. Д<strong>о</strong> <strong>с</strong>их<br />

д<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong> п<strong>о</strong>лным пре<strong>в</strong>р<strong>а</strong>щением: п<strong>о</strong>р эти пред<strong>к</strong><strong>о</strong><strong>в</strong>ые, перех<strong>о</strong>дные ф<strong>о</strong>рмы между<br />

личин<strong>к</strong>и <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й и, <strong>к</strong><strong>а</strong><strong>к</strong> прям<strong>о</strong><strong>к</strong>рылыми и жу<strong>к</strong><strong>а</strong>ми не <strong>о</strong>бн<strong>а</strong>ружены,<br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми грудных н<strong>о</strong>г: имеют<strong>с</strong>я Впер<strong>в</strong>ые же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong> нижн<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющие<br />

цер<strong>к</strong>и, н<strong>о</strong> лишь <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичные ней перми Ю. Сибири и Ур<strong>а</strong>л<strong>а</strong>, прит<strong>о</strong>м <strong>с</strong>р<strong>а</strong>зу<br />

прид<strong>а</strong>т<strong>к</strong>и н<strong>а</strong> <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong>. Суб<strong>с</strong>тр<strong>а</strong>ты <strong>о</strong>бит<strong>а</strong>- <strong>в</strong> б<strong>о</strong>льш<strong>о</strong>м р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зии. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы пред<strong>с</strong>т<strong>а</strong><strong>в</strong>иния<br />

и <strong>о</strong>бр<strong>а</strong>з жизни личин<strong>о</strong><strong>к</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны, тели не менее чем ше<strong>с</strong>ти <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> — Tshekardo-<br />

Пит<strong>а</strong>ние же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых <strong>о</strong>чень р<strong>а</strong>зличн<strong>о</strong>: coleidae, Asiocoleidae, Kaltanocoleidae, Permarхищниче<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

и п<strong>а</strong>р<strong>а</strong>зитизм, пит<strong>а</strong>ние р<strong>а</strong>зл<strong>а</strong>г<strong>а</strong>- rhaphidae, Cupidae, Taldycupidae, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

ющими<strong>с</strong>я жи<strong>в</strong><strong>о</strong>тными и р<strong>а</strong><strong>с</strong>тительными <strong>в</strong>еще<strong>с</strong>т- три п<strong>о</strong><strong>с</strong>ледних перех<strong>о</strong>дят и <strong>в</strong> <strong>в</strong>ерхнеперм<strong>с</strong><strong>к</strong>ую<br />

<strong>в</strong><strong>а</strong>ми, р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядн<strong>о</strong><strong>с</strong>ть. Си<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong><strong>а</strong> и<strong>с</strong>- ф<strong>а</strong>уну, еще б<strong>о</strong>лее б<strong>о</strong>г<strong>а</strong>тую. С три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> пеп<strong>о</strong>льзует<br />

для р<strong>а</strong>зличения <strong>в</strong>ы<strong>с</strong>ших <strong>к</strong><strong>а</strong>тег<strong>о</strong>рий ри<strong>о</strong>д<strong>а</strong> же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з-<br />

(н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>) <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти жил<strong>к</strong><strong>о</strong>- н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зия, <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>я<strong>с</strong>ь <strong>о</strong>дним из н<strong>а</strong>иб<strong>о</strong>лее б<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong>, <strong>с</strong>егмент<strong>а</strong>цию брюш<strong>к</strong><strong>а</strong>, г<strong>а</strong>тых <strong>в</strong>ид<strong>а</strong>ми <strong>о</strong>тряд<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Следует<br />

<strong>с</strong>тр<strong>о</strong>ение н<strong>о</strong>г, грудн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тдел<strong>а</strong> и <strong>а</strong>нтенн, м<strong>о</strong>р- <strong>о</strong>тметить, чт<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong><strong>а</strong> мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их, <strong>о</strong>чень<br />

ф<strong>о</strong>л<strong>о</strong>гию ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития. Р<strong>о</strong>ды и <strong>в</strong>иды х<strong>а</strong>р<strong>а</strong><strong>к</strong>те- р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных жу<strong>к</strong><strong>о</strong><strong>в</strong> (из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 250<br />

ризуют<strong>с</strong>я дет<strong>а</strong>льным <strong>с</strong>тр<strong>о</strong>ением <strong>с</strong><strong>а</strong>мых р<strong>а</strong>злич- р<strong>о</strong>д<strong>о</strong><strong>в</strong>) <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я еще <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong>р<strong>а</strong>зр<strong>а</strong>б<strong>о</strong>т<strong>а</strong>нных<br />

<strong>о</strong>тдел<strong>о</strong><strong>в</strong> тел<strong>а</strong> и <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. Следует <strong>о</strong>тметить, н<strong>о</strong>й. Же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые третичн<strong>о</strong>г<strong>о</strong> пери<strong>о</strong>д<strong>а</strong> предчт<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ение н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> и<strong>с</strong>п<strong>о</strong>льзует<strong>с</strong>я <strong>в</strong> <strong>с</strong>и- <strong>с</strong>т<strong>а</strong><strong>в</strong>лены <strong>в</strong>ид<strong>а</strong>ми р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненны'х <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ре<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>е<br />

м<strong>а</strong>л<strong>о</strong>: эт<strong>о</strong> <strong>с</strong><strong>о</strong>зд<strong>а</strong>ет б<strong>о</strong>льшие труд- менн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60) и <strong>в</strong><strong>о</strong>льн<strong>о</strong><strong>с</strong>ти<br />

при изучении и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>ры- шей ч<strong>а</strong><strong>с</strong>тью и тех же р<strong>о</strong>д<strong>о</strong><strong>в</strong> (<strong>с</strong><strong>в</strong>ыше 650).<br />

лых, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленных преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т- Нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>е зн<strong>а</strong>ние <strong>с</strong>тр<strong>о</strong>ения дре<strong>в</strong>них же<strong>к</strong><strong>а</strong>ми<br />

н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>. Фил<strong>о</strong>генез м<strong>а</strong>л<strong>о</strong>из<strong>в</strong>е<strong>с</strong>тен, <strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых не д<strong>а</strong>ет <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти изучить<br />

16 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 241<br />

http://jurassic.ru/

Ри<strong>с</strong>. 719—723. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tshekardocoleidae, Sojanocoleidae, Permarrhaphidae, Asiocoleidae, Kaltanocoleidae<br />

719. Tschekardocoleus magnus Rohdendorf; A — <strong>о</strong>бщий <strong>в</strong>ид, x 4,2;<br />

Б— <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, х 60,0; н. пермь, Ур<strong>а</strong>л (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1944)., 720. Sojanocoleus reticulatus Martynov; пр<strong>а</strong><strong>в</strong><strong>о</strong>е н<strong>а</strong>д<strong>к</strong>рылье,<br />

XS.0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1932). 721.<br />

Permarrhaphus venosus Martynov; н<strong>а</strong>д<strong>к</strong>рылье, X 4,5, <strong>в</strong>. пермь.<br />

<strong>с</strong>и<strong>с</strong>тему <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> и б<strong>о</strong>лее <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>их <strong>к</strong><strong>а</strong>тег<strong>о</strong>рий<br />

перм<strong>с</strong><strong>к</strong>их и б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>ти мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их жу<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

П<strong>о</strong>эт<strong>о</strong>му х<strong>а</strong>р<strong>а</strong><strong>к</strong>тери<strong>с</strong>ти<strong>к</strong>и н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, "<strong>с</strong>ерий<br />

и п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> для <strong>о</strong>пи<strong>с</strong>ы<strong>в</strong><strong>а</strong>емых и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емы'х<br />

же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых не м<strong>о</strong>гут быть д<strong>а</strong>ны.<br />

СЕМЕЙСТВО TSHEKARDOCOLEIDAE<br />

ROHDENDORF, 1944<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты'м жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящим из <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

жил<strong>о</strong><strong>к</strong>: трех р<strong>а</strong>ди<strong>а</strong>льных (R, RSi, RS 2), <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й<br />

SC, д<strong>в</strong>ух М, <strong>о</strong>дн<strong>о</strong>й Си и трех А. Между<br />

пр<strong>о</strong>д<strong>о</strong>льными б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> б<strong>о</strong>лее <strong>с</strong>л<strong>а</strong>бых<br />

п<strong>о</strong>перечных, <strong>о</strong>бр<strong>а</strong>зующих д<strong>в</strong><strong>о</strong>йные ряды<br />

ячее<strong>к</strong>. Н. пермь. Один р<strong>о</strong>д.<br />

Tshekardocoleus Rohdendorf, 1944. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. magnus Pohdendorf, 1944; и. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

242<br />

http://jurassic.ru/<br />

Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1930). 722. Astocoleus ncvojtlovi Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 11,0, н. пермь, Кузнец<strong>к</strong>ий! б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 723. Kaltanecoleus pospelovt Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 16,5, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961)<br />

Между R и RSi ячей<strong>к</strong>и ме<strong>с</strong>т<strong>а</strong>ми <strong>о</strong>дн<strong>о</strong>рядные,<br />

между RSi я RS 2 трехрядные. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

15 мм, ширин<strong>а</strong> 2,5 мм (ри<strong>с</strong>. 719 А,Б).<br />

Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

СЕМЕЙСТВО SOJANOCOLEIDAE<br />

MARTYNOV, 1932<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

из <strong>к</strong>реп<strong>к</strong>их пр<strong>о</strong>д<strong>о</strong>льных и п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>:<br />

SC, д<strong>в</strong>ух р<strong>а</strong>ди<strong>а</strong>льных, <strong>к</strong>репн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> М <strong>с</strong> непр<strong>а</strong><strong>в</strong>ильными,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями, <strong>о</strong>дн<strong>о</strong>й Си и<br />

<strong>о</strong>дн<strong>о</strong>й длинн<strong>о</strong>й А, идущих п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> д<strong>о</strong><br />

<strong>в</strong>ершины. В. пермь. Один р<strong>о</strong>д.<br />

Sojanocoleus Martynov, 1932. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. reticulatus Martynov, 1932; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>}.<br />

Между <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>ми R и М, <strong>а</strong> т<strong>а</strong><strong>к</strong>же между М и С<br />

непр<strong>а</strong><strong>в</strong>ильные жил<strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующие мн<strong>о</strong>г<strong>о</strong>ряд-

ные ячей<strong>к</strong>и. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 12 мм,<br />

ширин<strong>а</strong> 5 мм (ри<strong>с</strong>. 720). Один <strong>в</strong>ид. В. пермь<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО PERMARRHAPHIDAE<br />

MARTYNOV, 1930<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

из непр<strong>а</strong><strong>в</strong>ильные пр<strong>о</strong>д<strong>о</strong>льных и п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>: д<strong>в</strong>ух SC, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й и длинн<strong>о</strong>й, д<strong>в</strong>ух<br />

р<strong>а</strong>ди<strong>а</strong>льных, <strong>о</strong>дн<strong>о</strong>й М, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й Си и д<strong>в</strong>ух А,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я длинн<strong>а</strong>я д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong>. В. пермь. Один р<strong>о</strong>д.<br />

Permarrhaphus Martynov, 1930. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. venosus Martynov, 1930; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

П<strong>о</strong>перечные жил<strong>к</strong>и <strong>к</strong>реп<strong>к</strong>ие, н<strong>о</strong> непр<strong>а</strong><strong>в</strong>ильные;<br />

<strong>в</strong><strong>с</strong>е жил<strong>к</strong>и темные, <strong>к</strong>рыл<strong>о</strong> без <strong>с</strong><strong>к</strong>ульптуры, <strong>к</strong><strong>о</strong>жи<strong>с</strong>т<strong>о</strong>е.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм, ширин<strong>а</strong><br />

3 мм (ри<strong>с</strong>. 721). Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong> эт<strong>о</strong>му <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я р<strong>о</strong>д Uralocoleus G. Zalessky,<br />

1947, нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> п<strong>о</strong>лн<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>нный из н. ПЕРми<br />

Ур<strong>а</strong>л<strong>а</strong> (ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждение Крут<strong>а</strong>я К<strong>а</strong>туш<strong>к</strong><strong>а</strong><br />

н<strong>а</strong> р. Б<strong>а</strong>рде).<br />

СЕМЕЙСТВО ASIOCOLE1DAE<br />

ROHDENDORF, 1961<br />

М и Си н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зуя рез<strong>к</strong><strong>о</strong> <strong>о</strong>гр<strong>а</strong>ниченный <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з <strong>в</strong> <strong>в</strong>иде<br />

бу<strong>к</strong><strong>в</strong>ы X. Н<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье б<strong>о</strong>лее де<strong>с</strong>яти пр<strong>о</strong>д<strong>о</strong>льных<br />

ряд<strong>о</strong><strong>в</strong> <strong>к</strong>руглых ячее<strong>к</strong>. R, М, Си <strong>к</strong>реп<strong>к</strong>ие<br />

<strong>в</strong>ыпу<strong>к</strong>лые. О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>трез<strong>о</strong><strong>к</strong> М и б<strong>а</strong>з<strong>а</strong>льные<br />

<strong>в</strong>ет<strong>в</strong>и RS <strong>к</strong><strong>о</strong><strong>с</strong>ые. А <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, н<strong>о</strong> я<strong>с</strong>н<strong>а</strong>я. Н.<br />

пермь. Один р<strong>о</strong>д.<br />

Asiocoleus Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. novojitovi Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> 12 ряд<strong>а</strong>ми <strong>о</strong><strong>к</strong>руглых ячее<strong>к</strong>,<br />

интер<strong>в</strong><strong>а</strong>лы между ряд<strong>а</strong>ми р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые,<br />

<strong>с</strong> <strong>с</strong>ильн<strong>о</strong> буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тью; RS<br />

<strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, имеет <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

четыре гребне<strong>в</strong>идн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong>ет<strong>в</strong>и; Си<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong> <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т М<br />

и п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю, п<strong>о</strong><strong>с</strong>ле <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з<strong>а</strong><br />

<strong>с</strong> М Си рез<strong>к</strong><strong>о</strong> изгиб<strong>а</strong>ет<strong>с</strong>я <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> н<strong>а</strong>з<strong>а</strong>д;<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong> переднег<strong>о</strong> и з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 7,4 мм, ширин<strong>а</strong><br />

1,6 мм (ри<strong>с</strong>. 722). Один <strong>в</strong>ид.Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО KALTANOCOLEIDAE<br />

ROHDENDORF, 1961<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими ше<strong>с</strong>ть пр<strong>о</strong>д<strong>о</strong>льных<br />

ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>. В<strong>с</strong>е жил<strong>к</strong>и примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

т<strong>о</strong>лщины. Р<strong>а</strong>ди<strong>а</strong>льный <strong>с</strong>т<strong>в</strong><strong>о</strong>л <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й R и <strong>в</strong>ил<strong>к</strong>и д<strong>в</strong>ух жил<strong>о</strong><strong>к</strong>, RSi и RS 2.<br />

М и Си пр<strong>о</strong><strong>с</strong>тые, прямые, <strong>с</strong><strong>о</strong>единенные <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

чет<strong>в</strong>ерти н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong> неп<strong>а</strong>рный <strong>с</strong>т<strong>в</strong><strong>о</strong>л; А<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я <strong>к</strong><strong>о</strong>нц<strong>а</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й трети<br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я н<strong>а</strong>д<strong>к</strong>рылья. Между <strong>в</strong><strong>с</strong>еми жил<strong>к</strong><strong>а</strong>ми<br />

пр<strong>о</strong><strong>с</strong>тые ряды <strong>к</strong>рупных ячее<strong>к</strong>. Брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> и имеет пять <strong>в</strong>идимых <strong>с</strong>тернит<strong>о</strong><strong>в</strong>.<br />

Щит<strong>о</strong><strong>к</strong> <strong>к</strong>рупный. Н. пермь. Один р<strong>о</strong>д.<br />

Kaltanocoteus Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

К. pospelovi Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>о</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> уз<strong>к</strong><strong>о</strong>й, упл<strong>о</strong>щенн<strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>йм<strong>о</strong>й; <strong>в</strong>ершины н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренные<br />

и р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я. В<strong>с</strong>ег<strong>о</strong> <strong>с</strong>емь жил<strong>о</strong><strong>к</strong>.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4,5—4,6 мм, ширин<strong>а</strong> 0,78—<br />

0,85 мм (ри<strong>с</strong>. 723). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО CUPIDAE LATRE1LLE, 1825<br />

Имеют<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льные ряды 1<br />

<strong>к</strong>рупных и чет<strong>к</strong>их<br />

ячее<strong>к</strong>, р<strong>а</strong>згр<strong>а</strong>ниченных пр<strong>о</strong>д<strong>о</strong>льными ребр<strong>а</strong>ми-жил<strong>к</strong><strong>а</strong>ми<br />

нер<strong>а</strong><strong>в</strong>н<strong>о</strong>й т<strong>о</strong>лщины, <strong>к</strong>реп<strong>к</strong>ими<br />

и т<strong>о</strong>н<strong>к</strong>ими: между <strong>к</strong>реп<strong>к</strong>ими ребр<strong>а</strong>ми-жил<strong>к</strong><strong>а</strong>ми<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие эт<strong>о</strong>г<strong>о</strong> д<strong>в</strong><strong>о</strong>йные или<br />

тр<strong>о</strong>йные ряды ячее<strong>к</strong>. Пермь — ныне. Семь<br />

перм<strong>с</strong><strong>к</strong>их, <strong>о</strong>дин юр<strong>с</strong><strong>к</strong>ий и д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>а</strong>йн<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их<br />

р<strong>о</strong>д<strong>а</strong>; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

Archicupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. jacobsoni Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Жил<strong>к</strong>и <strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> и <strong>в</strong>ых<strong>о</strong>дят н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й:<br />

Си — <strong>в</strong> 3<br />

/б длины <strong>к</strong>рыл<strong>а</strong>, М — <strong>в</strong> <strong>с</strong><strong>а</strong>мую нершину<br />

<strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,9—5,4 мм, ширин<strong>а</strong><br />

0,75—1 мм (ри<strong>с</strong>. 724). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Eocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Е. lukjanovitshi Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>о</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong>ыпу<strong>к</strong>лый <strong>о</strong>бщий<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий <strong>с</strong>т<strong>в</strong><strong>о</strong>л R+ М и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние Си;<br />

<strong>в</strong> центре <strong>в</strong><strong>о</strong><strong>с</strong>емь ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong> п<strong>о</strong>чти пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й<br />

<strong>к</strong>ругл<strong>о</strong>й ф<strong>о</strong>рмы. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4,7 мм, ширин<strong>а</strong><br />

1,0 (ри<strong>с</strong>. 725). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kaltanicupes R <strong>о</strong> h d е n d or f, 1961. Тип р<strong>о</strong>д<strong>а</strong><br />

— К. richteri Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

К<strong>о</strong>нец н<strong>а</strong>д<strong>к</strong>рылья з<strong>а</strong><strong>о</strong><strong>с</strong>трен и <strong>о</strong>ттянут <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну;<br />

жил<strong>к</strong>и не<strong>к</strong><strong>о</strong><strong>с</strong>Ые и неп<strong>а</strong>р<strong>а</strong>ллельные; RS<br />

и М, <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти и Си <strong>к</strong>репче других; де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

ячее<strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,6 мм, ширин<strong>а</strong><br />

1,1 мм (ри<strong>с</strong>. 726). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Palaeocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. kaltanicus Rohdendorf, 1961; н. пермь,<br />

243 16*<br />

http://jurassic.ru/

729А<br />

724. Archicupes jacobsoni Rohdendorf; A — н<strong>а</strong>д<strong>к</strong>рылье, X 15,0;<br />

Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, Х60.0; н. Пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1958). 725. Eocupes lukjanovitshi Rohdendorf;<br />

А — н<strong>а</strong>д<strong>к</strong>рылье, X 16,5; Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, X 50,0,<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1958). 726. Kaltanicupes<br />

richteri Rohdendorf; A— н<strong>а</strong>д<strong>к</strong>рылье, X 120,0, Б —<strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья, х 70,0; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>ден­<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong><strong>о</strong><strong>с</strong>емь ряд<strong>о</strong><strong>в</strong><br />

<strong>к</strong>рупных ячее<strong>к</strong>. Жил<strong>к</strong>и зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные; R и<br />

М <strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бщий<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л, д<strong>а</strong>лее р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящий<strong>с</strong>я <strong>в</strong> <strong>в</strong>иде <strong>в</strong>ил<strong>к</strong>и, RS<br />

не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; М д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>в</strong>ершины н<strong>а</strong>д<strong>к</strong>рылья;<br />

Си <strong>к</strong>реп<strong>к</strong><strong>а</strong>я д<strong>о</strong> <strong>с</strong>ередины н<strong>а</strong>д<strong>к</strong>рылья,<br />

д<strong>а</strong>льше <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я. А <strong>к</strong>реп<strong>к</strong><strong>а</strong>я; пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные; н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е,<br />

730. Protocupes martynovi Rohdendorf; Л—н<strong>а</strong>д<strong>к</strong>рылье, X 14,7;<br />

Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, X 60,0; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 731. Tomiocupes carinatus Rohdendorf;<br />

А — н<strong>а</strong>д<strong>к</strong>рылье, X 22,5; Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, х 80,0;<br />

<strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 732. Tricupes<br />

acer Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье, X 24,2; Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья, х 95,0; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

728<br />

) O 0 Q G G < |<br />

Ри<strong>с</strong>. 724—729. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cupidae<br />

Ри<strong>с</strong>. 730—735. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cupidae<br />

244<br />

http://jurassic.ru/<br />

724Б<br />

729Б<br />

726Б<br />

и и <strong>о</strong> и 1<br />

O O O t M<br />

Ь О : С О С<br />

725Б<br />

д<strong>о</strong>рф, 1958). 727. Pataeocupes kaltanicus Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 14,5 н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 728'. Суtocupes<br />

angustus Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X 17,0, н. пермь. Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 729. Protocupoides plavilstshikovl<br />

Rohdendorf; A—н<strong>а</strong>д<strong>к</strong>рылье, X 15,8; Б —<strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья,<br />

X 80,0; н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961)<br />

<strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим н<strong>а</strong> <strong>с</strong>ередине передним<br />

<strong>к</strong>р<strong>а</strong>ем и <strong>с</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>й <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

н<strong>а</strong>д<strong>к</strong>рылья буг<strong>о</strong>рч<strong>а</strong>т<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

5,2 мм, ширин<strong>а</strong> 0,9 мм (ри<strong>с</strong>. 727).<br />

Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Cytocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

С. angustus Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья де<strong>с</strong>ять ряд<strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong>руг-<br />

1961). 733. Permocupoides skoki Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье<br />

Х15.0; Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, X 60,0; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 734. Permocupes sojanensls Ponomarenko;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 10,0 <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг.<br />

ри<strong>с</strong>.). 735. Mesocupes primitivus Martynov; <strong>о</strong>бщий <strong>в</strong>ид, X 7,5.<br />

<strong>в</strong>, юр<strong>а</strong> ? Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926)

730А<br />

731А<br />

732A<br />

733A<br />

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руте<br />

730 Б<br />

ooo'q<br />

^ QQ.Q. 731Б<br />

OOCC<br />

732Б

л<strong>о</strong>-прям<strong>о</strong>уг<strong>о</strong>льны'х ячее<strong>к</strong>. Жил<strong>к</strong>и прямые;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные <strong>с</strong>т<strong>в</strong><strong>о</strong>лы их нея<strong>с</strong>ные; <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

н<strong>а</strong>д<strong>к</strong>рылья рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыделяют<strong>с</strong>я жил<strong>к</strong>и М и Си;<br />

н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья четыре <strong>в</strong>ыпу<strong>к</strong>лые жил<strong>к</strong>и,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong>ыделяют<strong>с</strong>я RS и М.<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е: <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>о</strong>чень н<strong>а</strong><strong>к</strong>л<strong>о</strong>нен<strong>о</strong> <strong>в</strong>перед. Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющий, <strong>с</strong>н<strong>а</strong>бжен шипи<strong>к</strong><strong>а</strong>ми. Вершин<strong>а</strong><br />

<strong>о</strong><strong>с</strong>тр<strong>а</strong>я. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я, бле<strong>с</strong>тящ<strong>а</strong>я.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,1 мм, ширин<strong>а</strong><br />

0,9 мм (ри<strong>с</strong>. 728). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Protocupoides Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. plavilstshikovi Rohdendorf, 1961; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> четырьмя прямыми<br />

<strong>к</strong>реп<strong>к</strong>ими жил<strong>к</strong><strong>а</strong>ми (R, RS, М, Си). А <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

нея<strong>с</strong>н<strong>а</strong>я. Н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья 12 пр<strong>о</strong>д<strong>о</strong>льных<br />

ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>. В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м и <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>лях тр<strong>о</strong>йные ряды ячее<strong>к</strong>. Передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый, перед <strong>в</strong>ершинн<strong>о</strong>й третью <strong>с</strong><strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>бым изгиб<strong>о</strong>м; <strong>в</strong>ершин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

туп<strong>а</strong>я. Передний и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>я н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>с</strong> грубыми буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong> 4,8 мм,<br />

ширин<strong>а</strong> 1,1 мм (ри<strong>с</strong>. 729А, Б). Один <strong>в</strong>ид. Н.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Protocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. martynovi Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong> уз<strong>к</strong><strong>о</strong>й, н<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метн<strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>йм<strong>о</strong>й из р<strong>о</strong>мбиче<strong>с</strong><strong>к</strong>их ячее<strong>к</strong>. Н<strong>а</strong> <strong>с</strong>ередине<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>в</strong><strong>о</strong><strong>с</strong>емь-де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong> ячее<strong>к</strong>: жил<strong>к</strong>и<br />

R, RS, М, Си прямые, <strong>с</strong>тр<strong>о</strong>г<strong>о</strong> п<strong>а</strong>р<strong>а</strong>ллельные,<br />

<strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют д<strong>в</strong><strong>о</strong>йные ряды ячее<strong>к</strong>; пр<strong>о</strong>межут<strong>о</strong>чные<br />

пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и т<strong>а</strong><strong>к</strong>же п<strong>а</strong>р<strong>а</strong>ллельны.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,6—4,6 мм, ширин<strong>а</strong><br />

0,65—0,90 мм (ри<strong>с</strong>. 730). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tomiocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. carinatus Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Передняя <strong>к</strong><strong>а</strong>йм<strong>а</strong> пл<strong>о</strong>х<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>.<br />

Н<strong>а</strong> <strong>с</strong>ередине де<strong>с</strong>ять ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>: жил<strong>к</strong>и R,<br />

RS, М и Си из<strong>о</strong>гнутые дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, п<strong>а</strong>р<strong>а</strong>ллельные,<br />

<strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ющие д<strong>в</strong><strong>о</strong>йные ряды' ячее<strong>к</strong>;<br />

пр<strong>о</strong>межут<strong>о</strong>чные пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>бее гл<strong>а</strong><strong>в</strong>ных, <strong>в</strong> <strong>в</strong>иде зигз<strong>а</strong>г<strong>о</strong><strong>в</strong>. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 3 мм, ширин<strong>а</strong> 0,8 мм (ри<strong>с</strong>. 731А,<br />

Б). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tricupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Т. acer Rohdendorf, 1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II).<br />

Н<strong>а</strong>д<strong>к</strong>рылье уз<strong>к</strong><strong>о</strong>е, <strong>в</strong> <strong>с</strong>ередине <strong>с</strong> тремя <strong>к</strong>реп<strong>к</strong>ими<br />

пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми; <strong>в</strong><strong>с</strong>ег<strong>о</strong> де<strong>в</strong>ять ряд<strong>о</strong><strong>в</strong><br />

угл<strong>о</strong><strong>в</strong><strong>а</strong>тых ячее<strong>к</strong>, р<strong>а</strong>згр<strong>а</strong>ниченных <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ими,<br />

прямыми пр<strong>о</strong>межут<strong>о</strong>чными пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,10 мм, ширин<strong>а</strong><br />

246<br />

http://jurassic.ru/<br />

0,55 мм (ри<strong>с</strong>. 732А, Б). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permocupoides Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. distinctus Martynov, 1933; <strong>в</strong>. пермь.<br />

Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие<br />

Г<strong>о</strong>ры). R, RS, М, Си п<strong>а</strong>р<strong>а</strong>ллельны, <strong>с</strong>редние три<br />

жил<strong>к</strong>и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>к</strong>реп<strong>к</strong>ие; пр<strong>о</strong>межут<strong>о</strong>чные жил<strong>к</strong>и<br />

зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, з<strong>а</strong>метн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее гл<strong>а</strong><strong>в</strong>ных.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5 мм, ширин<strong>а</strong> б<strong>о</strong>льше 1 мм<br />

(ри<strong>с</strong>. 733 А, Б). Три <strong>в</strong>ид<strong>а</strong>. В. пермь Приур<strong>а</strong>лья<br />

и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permocupes Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. semenovi Martynov. 1933; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

В<strong>с</strong>е жил<strong>к</strong>и <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья п<strong>а</strong>р<strong>а</strong>ллельны,<br />

н<strong>а</strong>иб<strong>о</strong>лее <strong>к</strong>реп<strong>к</strong>ие А 2, Си и М; пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и прямые, зн<strong>а</strong>чительн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее<br />

гл<strong>а</strong><strong>в</strong>ных. Передняя <strong>к</strong><strong>а</strong>йм<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>я. В<strong>о</strong><strong>с</strong>емь ряд<strong>о</strong><strong>в</strong><br />

ячее<strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,8 мм, ширин<strong>а</strong><br />

0,9 мм (ри<strong>с</strong>. 734). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Приур<strong>а</strong>лья<br />

и Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Mesocupes Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М. primitivus Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н.<br />

Не менее <strong>в</strong><strong>о</strong><strong>с</strong>ьми ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>. Пр<strong>о</strong>межут<strong>о</strong>чные<br />

жил<strong>к</strong>и м<strong>а</strong>л<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т гл<strong>а</strong><strong>в</strong>ных,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> те, т<strong>а</strong><strong>к</strong> и другие прямые. Н<strong>а</strong>д<strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие,<br />

<strong>с</strong>н<strong>а</strong>бженные т<strong>о</strong>чечн<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й.<br />

Длин<strong>а</strong> тел<strong>а</strong> 7,5 мм, н<strong>а</strong>д<strong>к</strong>рылья 5,6 мм, ширин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1,6 мм (ри<strong>с</strong>. 735). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО TALDYCUPIDAE ROHDENDORF, 1961<br />

Н<strong>а</strong>д<strong>к</strong>рылье без <strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных<br />

жил<strong>о</strong><strong>к</strong>, <strong>с</strong> <strong>с</strong>емью — де<strong>с</strong>ятью п<strong>а</strong>р<strong>а</strong>ллельными<br />

ряд<strong>а</strong>ми ячее<strong>к</strong>, р<strong>а</strong>зделенных пр<strong>о</strong>д<strong>о</strong>льными<br />

и б<strong>о</strong>лее <strong>с</strong>л<strong>а</strong>быми п<strong>о</strong>перечными интер<strong>в</strong><strong>а</strong>л<strong>а</strong>ми.<br />

В<strong>с</strong>е пр<strong>о</strong>д<strong>о</strong>льные ребр<strong>а</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>чень пл<strong>о</strong>тный, и т<strong>о</strong>л<strong>с</strong>тый,<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> ячей<strong>к</strong>и <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong>ильн<strong>о</strong><br />

уменьшенными и б<strong>о</strong>лее я<strong>с</strong>н<strong>о</strong> з<strong>а</strong>метны лишь <strong>с</strong><br />

<strong>с</strong> нижней п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья. Брюш<strong>к</strong><strong>о</strong><br />

имеет ше<strong>с</strong>ть х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женных <strong>с</strong>тернит<strong>о</strong><strong>в</strong>.<br />

Н. пермь — три<strong>а</strong><strong>с</strong>. 13 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Schizotaldycupes Rohdendorf, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong>. — 5. ananjevi Rohdendorf, 1961; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> я<strong>с</strong>ным, н<strong>о</strong> не<br />

<strong>о</strong><strong>с</strong>трым, <strong>а</strong> <strong>с</strong>гл<strong>а</strong>женным пр<strong>о</strong>д<strong>о</strong>льным ребр<strong>о</strong>м <strong>в</strong><br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти Ri. В р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>.<br />

Н<strong>а</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья де<strong>с</strong>ять <strong>о</strong><strong>к</strong>ругл<strong>о</strong>-угл<strong>о</strong><strong>в</strong><strong>а</strong>тых<br />

ячее<strong>к</strong>. Вершин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>с</strong> г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> б<strong>о</strong>лее мел<strong>к</strong>ими и ред<strong>к</strong>ими<br />

ячей<strong>к</strong><strong>а</strong>ми, чем <strong>в</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья.<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>й<br />

эпипле<strong>в</strong>р<strong>о</strong>й, гл<strong>а</strong>д<strong>к</strong><strong>о</strong>й п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю и гу<strong>с</strong>т<strong>о</strong>т<strong>о</strong>чечн<strong>о</strong>й<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>с</strong><strong>в</strong><strong>о</strong>ей ч<strong>а</strong><strong>с</strong>ти. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья

736. Schtzotaldycupes) ananjevl Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X12.6, н.<br />

пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>ф, 1961). 737. Ichthyocupes<br />

tyzhnovi Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X 26,5, н. пермь, Кузнец<strong>к</strong>ий<br />

7,6 мм, ширин<strong>а</strong> 2,3 мм (ри<strong>с</strong>. 736). Один <strong>в</strong>ид.<br />

Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ichthyocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

/. tyzhnovi Rohdendorf, 1961; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> грубыми буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми<br />

и непр<strong>а</strong><strong>в</strong>ильными ячей<strong>к</strong><strong>а</strong>ми, <strong>о</strong>чень рез<strong>к</strong><strong>о</strong> бле<strong>с</strong>тящ<strong>а</strong>я.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>метн<strong>о</strong>. Передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>в</strong> д<strong>в</strong>ух <strong>с</strong><strong>в</strong><strong>о</strong>их б<strong>а</strong>з<strong>а</strong>льных третях п<strong>о</strong>чти<br />

прям<strong>о</strong>й, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый.<br />

Ри<strong>с</strong>. 736—738, 740. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Taldycupidae<br />

247<br />

http://jurassic.ru/<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 738. Carinicupes beckermlgdlsovae<br />

Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X 24,0, н. Пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). HO.Tecticupes heckeri Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 250, <strong>в</strong>.пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,» 1961)<br />

Вершин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> ут<strong>о</strong>нч<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я,<br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>о</strong><strong>с</strong>тр<strong>а</strong>я н<strong>а</strong> <strong>к</strong><strong>о</strong>нце. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

3,6 мм, ширин<strong>а</strong> 0,9 мм (ри<strong>с</strong>. 737).<br />

Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Carinicupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong> —<br />

С. beckermigdisovae Rohdendorf, 1961; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Н<strong>а</strong>д<strong>к</strong>рылье не<strong>с</strong>ет пр<strong>о</strong>д<strong>о</strong>льную <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>у-ребр<strong>о</strong>,<br />

<strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>о</strong><strong>с</strong>ую и <strong>о</strong><strong>с</strong>трую.<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья груб<strong>о</strong>буг<strong>о</strong>рч<strong>а</strong>т<strong>а</strong>я, <strong>с</strong> 12

Ри<strong>с</strong>. 739, 741. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Taldycupidae<br />

739. Stegocupes fedotovi Rohdendorf; А —<strong>о</strong>бщий <strong>в</strong>ид, X 18,0; Б— <strong>к</strong>рылье, X 26,0; Б— <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>л<strong>к</strong>пылья X 100 <strong>о</strong>* <strong>в</strong> пепмь<br />

<strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, х 100,0; <strong>в</strong> пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн. Кузнец<strong>к</strong>ий Й<strong>а</strong><strong>с</strong><strong>с</strong>ей<strong>в</strong>i (Р<strong>о</strong>ленлппА' I9fin Р<br />

Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 741. Taldycupes khalfini Rohdendorf; А— н<strong>а</strong>д- лузнец<strong>к</strong>ии б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961).<br />

пр<strong>о</strong>д<strong>о</strong>льными ряд<strong>а</strong>ми <strong>к</strong>рупных <strong>о</strong><strong>к</strong>руглых, н<strong>о</strong><br />

неглуб<strong>о</strong><strong>к</strong>их ячее<strong>к</strong>. Передний <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>'и <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й<br />

п<strong>о</strong>чти прям<strong>о</strong>й. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4,75 мм, ширин<strong>а</strong><br />

1,20 мм (ри<strong>с</strong>. 738). Один <strong>в</strong>ид. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Stegocupes R <strong>о</strong> h d е n d <strong>о</strong> г f, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. fedotovi Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>).<br />

В <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>емь ряд<strong>о</strong><strong>в</strong><br />

ячее<strong>к</strong>. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья м<strong>о</strong>рщини<strong>с</strong>т<strong>а</strong>я,<br />

ячей<strong>к</strong>и непр<strong>а</strong><strong>в</strong>ильные. Ше<strong>с</strong>ть <strong>с</strong>тернит<strong>о</strong><strong>в</strong><br />

брюш<strong>к</strong><strong>а</strong>. З<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и п<strong>о</strong>перечные, <strong>с</strong>редние —<br />

ш<strong>а</strong>р<strong>о</strong><strong>в</strong>идные. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 2,85—3,50 мм,<br />

ширин<strong>а</strong> 0,65—0,70 мм, длин<strong>а</strong> <strong>в</strong><strong>с</strong>ег<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong><br />

4 мм (ри<strong>с</strong>. 739). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tecticupes R <strong>о</strong> h d е n d <strong>о</strong> г f, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. heckeri Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>).<br />

Де<strong>с</strong>ять пр<strong>о</strong>д<strong>о</strong>льных ряд<strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong>руглых<br />

ячее<strong>к</strong>. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> груб<strong>о</strong>й зерни<strong>с</strong>т<strong>о</strong><strong>с</strong>тью.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,0—3,15 мм, ширин<strong>а</strong><br />

0,65—0,85 мм (ри<strong>с</strong>. 740). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В.<br />

пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Taldycupes R <strong>о</strong> h d е n d <strong>о</strong> г f, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. khalfini Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Де<strong>с</strong>ять ряд<strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong>руглых ячее<strong>к</strong>.<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть ребер гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я, без зерни<strong>с</strong>т<strong>о</strong><strong>с</strong>ти<br />

или м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong><strong>с</strong>ти. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 2,6—<br />

248<br />

739А<br />

3,6 мм, ширин<strong>а</strong> 1,0—1,6 мм (ри<strong>с</strong>. 741А, Б).<br />

Четыре <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tatdycupidium Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. bergi Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Де<strong>с</strong>ять ряд<strong>о</strong><strong>в</strong> <strong>о</strong>чень мел<strong>к</strong>их ячее<strong>к</strong>,<br />

нерез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных <strong>о</strong>т пл<strong>о</strong><strong>с</strong><strong>к</strong>их интер<strong>в</strong><strong>а</strong>л<strong>о</strong><strong>в</strong>.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 1,95—2,10 мм, ширин<strong>а</strong> 0,72—<br />

0,74 мм (ри<strong>с</strong>. 742). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Cryptocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

С. rjabinini Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Н<strong>а</strong>д<strong>к</strong>рылье <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е, гл<strong>а</strong>д<strong>к</strong><strong>о</strong>е, <strong>с</strong><br />

<strong>в</strong><strong>о</strong><strong>с</strong>емью ряд<strong>а</strong>ми нечет<strong>к</strong>их <strong>к</strong>рупных <strong>о</strong><strong>к</strong>руглых<br />

ячее<strong>к</strong>, н<strong>а</strong> дне <strong>к</strong><strong>о</strong>т<strong>о</strong>рых имеет<strong>с</strong>я <strong>с</strong>рединный чет<strong>к</strong>ий<br />

буг<strong>о</strong>р<strong>о</strong><strong>к</strong>. Передний и з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>я н<strong>а</strong>д<strong>к</strong>рылья<br />

умеренн<strong>о</strong> и п<strong>о</strong>чти <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые.<br />

Вершин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья угл<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>о</strong><strong>с</strong>тр<strong>а</strong>я,<br />

не<strong>в</strong>ытянут<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 1,9 мм,<br />

ширин<strong>а</strong> 0,5 мм (ри<strong>с</strong>. 743). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tychticupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. radtschenkoi Rohdendorf, 1961; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). Пр<strong>о</strong>д<strong>о</strong>льные ребр<strong>а</strong>-жил<strong>к</strong>и не<br />

<strong>в</strong>ыд<strong>а</strong>ют<strong>с</strong>я. В центре н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>емь — <strong>в</strong><strong>о</strong><strong>с</strong>емь<br />

пр<strong>о</strong>д<strong>о</strong>льных ряд<strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong>руглых или угл<strong>о</strong><strong>в</strong><strong>а</strong>тых<br />

ячее<strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 2,5—3,4 мм, ширин<strong>а</strong><br />

0,6—0,8 мм (ри<strong>с</strong>. 744). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

http://jurassic.ru/

Tychtocupoides Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. grjazevi Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Пр<strong>о</strong>д<strong>о</strong>льные ребр<strong>а</strong>-жил<strong>к</strong>и умерен-<br />

7446<br />

н<strong>о</strong> <strong>в</strong>ыд<strong>а</strong>ют<strong>с</strong>я; <strong>в</strong> <strong>с</strong>ередине н<strong>а</strong>д<strong>к</strong>рылья де<strong>с</strong>ять<br />

пр<strong>о</strong>д<strong>о</strong>льных ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>, имеющих <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тные<br />

или прям<strong>о</strong>уг<strong>о</strong>льные <strong>о</strong>черт<strong>а</strong>ния и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> друг пр<strong>о</strong>ти<strong>в</strong> друг<strong>а</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4,5 м, ширин<strong>а</strong> 1,0 (ри<strong>с</strong>. 745А, Б).<br />

Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Uskaticupes Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — U. javorskyi Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Де<strong>в</strong>ять ряд<strong>о</strong><strong>в</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тых прям<strong>о</strong>уг<strong>о</strong>льных<br />

ячее<strong>к</strong>. Пр<strong>о</strong>д<strong>о</strong>льные ребр<strong>а</strong>-жил<strong>к</strong>и<br />

я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ют н<strong>а</strong> <strong>в</strong><strong>с</strong>ем пр<strong>о</strong>тяжении. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 1,8 мм, ширин<strong>а</strong> 0,5 мм (ри<strong>с</strong>. 746А,<br />

Б). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Mesothoris Tillyard, 1916 (pars);<br />

Simmondsia Dunstan, 1924.<br />

СЕМЕЙСТВО RHOMBOCOLEIDAE<br />

ROHDENDORF, 1961<br />

В б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине н<strong>а</strong>д<strong>к</strong>рылья ег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных<br />

2<br />

/s длины имеют<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льные ряды<br />

мел<strong>к</strong>их ячее<strong>к</strong> и 11—14 пр<strong>о</strong>д<strong>о</strong>льных ребер, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>гл<strong>а</strong>жены. В. пермь.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Phombocoleus Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Rh. andreae Rohdendorf, 1961; <strong>в</strong>. пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>.<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). 12—13 ряд<strong>о</strong><strong>в</strong> <strong>о</strong>чень уз<strong>к</strong>их ячее<strong>к</strong>,<br />

<strong>о</strong>бр<strong>а</strong>зующих пр<strong>о</strong>д<strong>о</strong>льные б<strong>о</strong>р<strong>о</strong>зды. Плече<strong>в</strong><strong>о</strong>й<br />

буг<strong>о</strong>р<strong>о</strong><strong>к</strong> рез<strong>к</strong>ий. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,0 мм, ширин<strong>а</strong><br />

1,2 мм (ри<strong>с</strong>. 747А, Б). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Erunakicupes Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Е. kryshtofovichi Rohdendorf, 1961; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). Де<strong>с</strong>ять ряд<strong>о</strong><strong>в</strong> ячее<strong>к</strong>, имеющих<br />

<strong>в</strong>ид глуб<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> пун<strong>к</strong>тир<strong>а</strong>, р<strong>а</strong>згр<strong>а</strong>ниченных<br />

11 пр<strong>о</strong>д<strong>о</strong>льными ребр<strong>а</strong>ми. Плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong><br />

б<strong>о</strong>лее <strong>с</strong>л<strong>а</strong>бый. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 1,75—<br />

3,75 мм, ширин<strong>а</strong> 0,75—1,10 мм (ри<strong>с</strong>. 748, 749).<br />

Четыре <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО SCHIZOCOLE1DAE<br />

ROHDENDORF, 1961<br />

В <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти жил<strong>к</strong>и RS н<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье имеет<strong>с</strong>я<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>,<br />

имеющ<strong>а</strong>я <strong>в</strong>ид и<strong>с</strong><strong>к</strong>у<strong>с</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> <strong>в</strong>д<strong>а</strong><strong>в</strong>ле-<br />

Ри<strong>с</strong>. 742—746. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Taldycupidae<br />

742. Taldycupidium bergi Rohdendorf; A — н<strong>а</strong>д<strong>к</strong>рылье, X 38,0;<br />

Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, х 200,0; <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

( Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 743. Cryptocupes rjabinini Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 38,0; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 744. Tychticupes radtshenkoi Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье,<br />

Х27.0; Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, X 70,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 745. Tychtocupoides grjazevi<br />

Rohdendorf; А— н<strong>а</strong>д<strong>к</strong>рылье, X 15,0; Б — <strong>с</strong><strong>к</strong>ульпур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья,<br />

Х38.0; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 746.<br />

Uskaticupes javorskyi Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье, X 41,5;<br />

Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, X 150,0; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961)<br />

249<br />

http://jurassic.ru/

7476 749<br />

Ри<strong>с</strong>. 747—749. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhombocoleidae<br />

747. Rhombocoleus andreae Rohdendorf; A — н<strong>а</strong>д<strong>к</strong>рылье, X 25,0;<br />

Б— <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, х 108,0; <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 748. Erunakicupes kryshtofovlchi Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, х 32,3, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

ния. Эт<strong>а</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> глуб<strong>о</strong><strong>к</strong><strong>а</strong>я и рез<strong>к</strong><strong>а</strong>я,<br />

реже <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я. Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й: ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> нежн<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong><strong>с</strong>тью или <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льными<br />

ряд<strong>а</strong>ми буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>, т<strong>о</strong>чечных углублений или<br />

т<strong>о</strong>н<strong>к</strong>их ребер. Брюш<strong>к</strong><strong>о</strong> имеет пять чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных<br />

<strong>с</strong>тернит<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>о</strong><strong>с</strong>ледний<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong>ели<strong>к</strong>. Пермь. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Rossocoleus Rohdendorf, 1961 (Curcutiopsis<br />

Martynov, 1937, пе<strong>с</strong> Curculiopsis Handlirsch,<br />

1906). Тип р<strong>о</strong>д<strong>а</strong> — R. grandis Rohdendorf,<br />

1961; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> I). Выпу<strong>к</strong>л<strong>о</strong>е<br />

н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> ряд<strong>а</strong>ми х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метных буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>в</strong> 3,2—4,2 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины: длин<strong>а</strong><br />

2,3—4,9 мм, ширин<strong>а</strong> 0,70—1,35 мм (ри<strong>с</strong>. 750).<br />

Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. и <strong>в</strong>. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

и Приур<strong>а</strong>лья.<br />

Schizocoleus Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. kuznetskiensis Rohdendorf, 1961;<br />

250<br />

http://jurassic.ru/<br />

748<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 749. Е. schmalhauseni Rohdendorf; <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья, х 132,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961)<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> II). Н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лые,<br />

<strong>в</strong>пл<strong>о</strong>ть д<strong>о</strong> <strong>в</strong>ершинн<strong>о</strong>й трети п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><strong>к</strong>р<strong>а</strong>йние,<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>тренные лишь н<strong>а</strong> <strong>к</strong><strong>о</strong>нце.<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть т<strong>о</strong>н<strong>к</strong><strong>о</strong>- и непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>м<strong>о</strong>рщини<strong>с</strong>т<strong>а</strong>я.<br />

Б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> чет<strong>к</strong><strong>а</strong>я. Плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> нея<strong>с</strong>ный.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 7,2 мм, ширин<strong>а</strong> 2,0 мм<br />

(ри<strong>с</strong>. 751 А, Б). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Schizocupes Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. obrutshevi Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Зеленый<br />

Луг). Сильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> рез<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женным плече<strong>в</strong>ым буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м, <strong>в</strong> 2,9 р<strong>а</strong>з<strong>а</strong><br />

длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины. Б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии д<strong>о</strong> де<strong>с</strong>яти<br />

нея<strong>с</strong>ных пр<strong>о</strong>д<strong>о</strong>льных ряд<strong>о</strong><strong>в</strong> мел<strong>к</strong>их ям<strong>о</strong><strong>к</strong>. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> груб<strong>о</strong>й м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong><strong>с</strong>тью.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 3,2 мм, ширин<strong>а</strong> 1,1 мм<br />

(ри<strong>с</strong>. 752 А, Б). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.

750. Rossocoleus avus Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X116.0, н. пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961)! 751.Schizocoleus kuznetskiensis<br />

Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье, X 10,0; 5 — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья, х 40,0; В—<strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, X 160,0; <strong>в</strong>.<br />

пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 752. Schlzocupes<br />

obrutshevl Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье, X 24,0; Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья, X 150,0; <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 753. Karakanocoleus lebedevl Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье,<br />

765<br />

Ри<strong>с</strong>. 750—756. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Schizocoleidae<br />

751Б<br />

761B<br />

762Б<br />

764Б<br />

X 25,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 754.<br />

Karakanocoleodes latlssimus Rohdendorf; А — н<strong>а</strong>д<strong>к</strong>рылье, X 16,5;<br />

Б — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, х 50,0; <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 755. Uskatocoleus zalesskyl Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 23,0, <strong>в</strong>. пермь Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 756. Aenigmocoleus borlsstaki Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 16,0, <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961).<br />

http://jurassic.ru/

Ри<strong>с</strong>. 757—763. Се*<br />

757. Palademosyne martynovae Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X 25,0,<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 758. Stegosyne<br />

rugosa Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье,X 38,0, <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 759. Tychtocoleus neuburgae Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 42,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 760. Ademosyne siblrica Rohdendorf; н<strong>а</strong>д<strong>к</strong>рылье, X 50,0,<br />

Karakanocoleus Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К. lebedevi Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Очень <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong> <strong>о</strong><strong>к</strong>руглым<br />

плече<strong>в</strong>ым буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м, удлиненн<strong>о</strong>й ф<strong>о</strong>рмы,<br />

<strong>в</strong> 3,3—3,7 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины. Б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong><br />

изменчи<strong>в</strong><strong>о</strong>й ф<strong>о</strong>рмы 1<br />

. Имеют<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льные ряды<br />

<strong>о</strong>чень нежных т<strong>о</strong>че<strong>к</strong> или т<strong>о</strong>н<strong>к</strong>их ребер, р<strong>а</strong>зличн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 2,6—4,6<br />

мм, ширин<strong>а</strong> 0,80—1,35 мм (ри<strong>с</strong>. 753). Ше<strong>с</strong>ть<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

liarakanocoleodes Rohdendorf, 1961.<br />

Тип р<strong>о</strong>д<strong>а</strong> — К. latissimus Rohdendorf, 1961; <strong>в</strong>.<br />

252<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong> Permosynidae<br />

http://jurassic.ru/<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 761. Ademosynoides<br />

asiaticus Martynov; н<strong>а</strong>д<strong>к</strong>рылье,X 11,0, три<strong>а</strong><strong>с</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1936). 762. Polysltum kuznetskien.se Rohdendorf;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 37,0; <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 763. Permocrossos elongatus Martynov; н<strong>а</strong>д<strong>к</strong>рылье, X 10,0,<br />

<strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1932)<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). Сл<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>с</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыд<strong>а</strong>ющим<strong>с</strong>я плече<strong>в</strong>ым буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м, шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

лишь <strong>в</strong> 2,75 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины.<br />

Б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> <strong>в</strong> б<strong>о</strong>льш<strong>о</strong>м углублении.<br />

Имеют<strong>с</strong>я ряды <strong>о</strong>чень мел<strong>к</strong>их п<strong>о</strong>перечных ям<strong>о</strong><strong>к</strong><br />

и <strong>в</strong>ыпу<strong>к</strong>лые пр<strong>о</strong>д<strong>о</strong>льные ребр<strong>а</strong> <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти<br />

н<strong>а</strong>д<strong>к</strong>рылья. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 4,4 мм, ширин<strong>а</strong><br />

1,6 мм (ри<strong>с</strong>. 754). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Uskatocoleus Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — U. zalesskyi Rohdendorf, 1961; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). Н<strong>а</strong>д<strong>к</strong>рылья умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>-

лые, рез<strong>к</strong><strong>о</strong> <strong>с</strong>уж<strong>а</strong>ющие<strong>с</strong>я <strong>к</strong> <strong>в</strong>ершине. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

п<strong>о</strong>чти гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я, <strong>с</strong>л<strong>а</strong>б<strong>о</strong>м<strong>о</strong>рщини<strong>с</strong>т<strong>а</strong>я.<br />

Плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленный. Длин<strong>а</strong><br />

н<strong>а</strong>д<strong>к</strong>рылья 2,25—4,60 мм, ширин<strong>а</strong> 0,75—1,30 мм<br />

(ри<strong>с</strong>. 755). 13 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Aenigmocoteus Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. borissiaki Rohdendorf, 1961; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong>ужен<strong>о</strong>, <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим плече<strong>в</strong>ым<br />

буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м. Передний <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й, з<strong>а</strong>дний<br />

— <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый. Вершин<strong>а</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

<strong>о</strong><strong>с</strong>тр<strong>а</strong>я. Щеле<strong>в</strong>идн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и чет<strong>к</strong><strong>а</strong>я.<br />

Н<strong>а</strong>д<strong>к</strong>рылье незн<strong>а</strong>чительн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е,<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, т<strong>о</strong>н<strong>к</strong><strong>о</strong>е, п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть ег<strong>о</strong> <strong>с</strong> гу<strong>с</strong>т<strong>о</strong>й<br />

мел<strong>к</strong><strong>о</strong>й буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong><strong>с</strong>тью. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

4,8 мм, ширин<strong>а</strong> 1,2 мм (ри<strong>с</strong>. 756). Один <strong>в</strong>ид.<br />

В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО PERMOSYN1DAE TILLYARD, 1924<br />

Н<strong>а</strong>д<strong>к</strong>рылья без з<strong>а</strong>метн<strong>о</strong>г<strong>о</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния, <strong>с</strong><br />

гл<strong>а</strong>д<strong>к</strong><strong>о</strong>й или <strong>с</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тью,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>ми,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>н<strong>а</strong>бженными <strong>с</strong>л<strong>а</strong>быми ям<strong>к</strong><strong>а</strong>ми. Р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong> («щель») н<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Брюш<strong>к</strong><strong>о</strong> имеет четыре или пять<br />

<strong>с</strong>тернит<strong>о</strong><strong>в</strong>. В. пермь — три<strong>а</strong><strong>с</strong>. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Palademosyne Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. martynovae Rohdendorf, 1961; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Н<strong>а</strong>д<strong>к</strong>рылье умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е,<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние туп<strong>о</strong>е, зн<strong>а</strong>чительн<strong>о</strong> уже м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й<br />

ширины. Передний <strong>к</strong>р<strong>а</strong>й рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый<br />

и непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> из<strong>о</strong>гнутый перед <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

переднее п<strong>о</strong>ле <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong> и р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>т<strong>о</strong>е<br />

<strong>в</strong>перед. З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> и <strong>с</strong>ильн<strong>о</strong><br />

дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им, ут<strong>о</strong>лщенным<br />

р<strong>а</strong>нт<strong>о</strong>м. Вершин<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, угл<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я и<br />

<strong>о</strong><strong>с</strong>тр<strong>а</strong>я. О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я,<br />

б<strong>о</strong>лее пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я. С<strong>к</strong>ульптур<strong>а</strong> <strong>в</strong> <strong>в</strong>иде т<strong>о</strong>н<strong>к</strong><strong>о</strong>й<br />

буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong><strong>с</strong>ти, пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 2,20 мм, ширин<strong>а</strong><br />

0,88 мм (ри<strong>с</strong>. 757). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Stegosyne R <strong>о</strong> h d е n d <strong>о</strong> г f, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

S. rugosa Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Чи<strong>с</strong>т<strong>а</strong>я<br />

Гри<strong>в</strong><strong>а</strong>). Н<strong>а</strong>д<strong>к</strong>рылье умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е, шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

притуплённ<strong>о</strong>е, незн<strong>а</strong>чительн<strong>о</strong> уже <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й ширины. Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый,<br />

<strong>с</strong> чет<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тью перед <strong>с</strong>ередин<strong>о</strong>й н<strong>а</strong>д<strong>к</strong>рылья;<br />

переднее п<strong>о</strong>ле <strong>о</strong>пущен<strong>о</strong> <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong>. З<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым, не<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> уз<strong>к</strong>им р<strong>а</strong>нт<strong>о</strong>м. Вершин<strong>а</strong> умеренн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, угл<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я и <strong>о</strong><strong>с</strong>тр<strong>а</strong>я. Н<strong>а</strong>д<strong>к</strong>рылье п<strong>о</strong><strong>к</strong>рыт<strong>о</strong><br />

груб<strong>о</strong>й и непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong><strong>с</strong>тью,<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>. Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong><br />

<strong>с</strong>ильным бле<strong>с</strong><strong>к</strong><strong>о</strong>м. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 1,80 мм,<br />

ширин<strong>а</strong> 0,82 мм (ри<strong>с</strong>. 758). Один <strong>в</strong>ид. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tychtocoleus Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. neuburgae Rohdendorf, 1961; <strong>в</strong>. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

II). Н<strong>а</strong>д<strong>к</strong>рылье <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>е.<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый,<br />

переднее п<strong>о</strong>ле <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> <strong>о</strong>пущенн<strong>о</strong>е, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

н<strong>а</strong>д<strong>к</strong>рылья п<strong>о</strong>чти <strong>в</strong>ерти<strong>к</strong><strong>а</strong>льн<strong>о</strong>е. З<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong> и р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым<br />

р<strong>а</strong>нт<strong>о</strong>м. Н<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылье де<strong>в</strong>ять п<strong>а</strong>р<strong>а</strong>ллельных<br />

шир<strong>о</strong><strong>к</strong>их и <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лых ребер, р<strong>а</strong>зделенных<br />

де<strong>с</strong>ятью т<strong>о</strong>н<strong>к</strong>ими глуб<strong>о</strong><strong>к</strong>ими б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми,<br />

не<strong>с</strong>ущими мел<strong>к</strong>ие ям<strong>к</strong>и- П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>а</strong>д<strong>к</strong>рылья<br />

шер<strong>о</strong>х<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й штрих<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong><strong>с</strong>тью,<br />

з<strong>а</strong>метн<strong>о</strong>й при <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>м <strong>о</strong><strong>с</strong><strong>в</strong>ещении. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

1,80—1,82 мм, ширин<strong>а</strong> 0,52—0,55 мм<br />

(ри<strong>с</strong>. 759). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Ademosyne Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. major Handlirsch, 1961; три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен.<br />

Семь — де<strong>с</strong>ять чет<strong>к</strong>их б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong><strong>к</strong>, не<strong>с</strong>ущих з<strong>а</strong>метные,<br />

н<strong>о</strong> неб<strong>о</strong>льшие и мел<strong>к</strong>ие ям<strong>к</strong>и. Отн<strong>о</strong>шение<br />

ширины <strong>к</strong> длине 1 : 1,9—3,5. Ребр<strong>о</strong> перед<br />

ш<strong>о</strong><strong>в</strong>ным <strong>к</strong>р<strong>а</strong>ем <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

1,5—6,2 мм, ширин<strong>а</strong> 0,55—3,0 мм (ри<strong>с</strong>. 760).<br />

16 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и<br />

три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Ademosynoides Dunstan, 1924. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Ademosyne minor Handlirsch, 1906; три<strong>а</strong><strong>с</strong>,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия. Плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит.<br />

В<strong>о</strong><strong>с</strong>емь — де<strong>с</strong>ять б<strong>о</strong>р<strong>о</strong>зд, лишенных т<strong>о</strong>че<strong>к</strong> или<br />

ям<strong>о</strong><strong>к</strong>. Отн<strong>о</strong>шение длины н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> <strong>к</strong> их ширине<br />

1:2, 4—3,0. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й без <strong>к</strong><strong>а</strong>ймы,<br />

ребр<strong>а</strong> перед ш<strong>о</strong><strong>в</strong>ным <strong>к</strong>р<strong>а</strong>ем нет. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

2,0—8,5 мм, ширин<strong>а</strong> 0,8—2,9 мм (ри<strong>с</strong>.<br />

761). В<strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Три<strong>а</strong><strong>с</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong><br />

(Б<strong>а</strong>бий К<strong>а</strong>мень) и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Polysitum Dunstan, 1924. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. punctatum Dunstan, 1924; три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит. Б<strong>о</strong>р<strong>о</strong>зды<br />

п<strong>о</strong>чти не р<strong>а</strong>з<strong>в</strong>иты, имеют<strong>с</strong>я лишь их з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong>и,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> н<strong>а</strong> передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

м<strong>о</strong>рщини<strong>с</strong>т<strong>а</strong>я. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й <strong>с</strong> уз<strong>к</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 1,7—6,9 мм, ширин<strong>а</strong><br />

0,75—2,70 мм (ри<strong>с</strong>. 762). Три <strong>в</strong>ид<strong>а</strong>. В. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Permocrossos Martynov, 1932. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. elongatus Martynov, 1932; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Имеет<strong>с</strong>я <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>к</strong><strong>а</strong>йм<strong>а</strong>,<br />

253<br />

http://jurassic.ru/

Ри<strong>с</strong>. 7«-„,. <strong>с</strong>е.* С^.е, O g ^ ^ ^ H - ^ Pa u s s l t o,<br />

764. Carabopteron punctatollneatum Martynov; н<strong>а</strong>д<strong>к</strong>рылье у ч 4<br />

пл^ Р<br />

К <strong>а</strong> з <strong>а</strong> х <strong>с</strong> т <strong>а</strong><br />

- Я <strong>а</strong> / <strong>а</strong> е <strong>о</strong><br />

5'| J?" н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926) 765 Copt^cTatalonk linn «*'''""<br />

poda Ping; личин<strong>к</strong><strong>а</strong> (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция)- XI 5 <strong>в</strong> n г « ^<br />

5<br />

sr"ga/us Schlechtendal; <strong>о</strong>бщий <strong>в</strong>ил<br />

Х'0.0, п<strong>а</strong>ле<strong>о</strong>ген, Герм<strong>а</strong>ния (Hatch, 1928). 769 Paussui Л, •'<br />

<strong>о</strong>бщий <strong>в</strong>ид.х 7 О <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 170 PhUonthL tn<br />

<strong>о</strong>бщий <strong>в</strong>ид, х 16,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 19 5) 77 . Bythnus s£' :<br />

<strong>о</strong>бщий <strong>в</strong>ид, х 16,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915) Р<br />

"*<br />

<strong>о</strong>бр<strong>а</strong>зующ<strong>а</strong>я плече<strong>в</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>ширение; н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящий<br />

плече<strong>в</strong><strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен. Б<strong>о</strong>р<strong>о</strong>зды без<br />

ям<strong>о</strong><strong>к</strong> или т<strong>о</strong>че<strong>к</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 5,4 мм, ширин<strong>а</strong><br />

2 мм (ри<strong>с</strong>. 763). Один <strong>в</strong>ид. В. пермь<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Вне СССР: Permosyne Tillyard, 1924; Platycrossos<br />

Dunstan, 1924; Tryoniopsis Dunstan,<br />

1924; Ulomites Tillyard, 1916; Reeveana Dunstan,<br />

1924; Grammositum Dunstan, 1924; Shepherdia<br />

Dunstan, 1924.<br />

СЕМЕЙСТВО CARABIDAE LATREILLE, 1825.<br />

ЖУЖЕЛИЦЫ<br />

Крупные или <strong>с</strong>редней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

Н<strong>а</strong>д<strong>к</strong>рылье <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льными<br />

ряд<strong>а</strong>ми буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> или ям<strong>о</strong><strong>к</strong>; чи<strong>с</strong>л<strong>о</strong> т<strong>о</strong>чечных<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong><strong>к</strong> изменчи<strong>в</strong><strong>о</strong>: <strong>о</strong>т 9 д<strong>о</strong> 32. Ди<strong>а</strong>гн<strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong><br />

п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> з<strong>а</strong>труднительн<strong>а</strong>.<br />

В. юр<strong>а</strong> — ныне. Одн<strong>о</strong> из <strong>к</strong>рупнейших <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны, б<strong>о</strong>г<strong>а</strong>т<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>о</strong>е <strong>в</strong><br />

третичных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х (<strong>с</strong><strong>в</strong>ыше<br />

50 р<strong>о</strong>д<strong>о</strong><strong>в</strong>).<br />

254<br />

http://jurassic.ru/<br />

Carabopteron М <strong>а</strong> г t у п <strong>о</strong> v, 1926. Тип р<strong>о</strong>д<strong>а</strong>—<br />

С. punctatotineatum Martynov, 1926;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Имеет<strong>с</strong>я четыре гл<strong>а</strong><strong>в</strong>ных пр<strong>о</strong>д<strong>о</strong>льных ребр<strong>а</strong><br />

<strong>с</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми и п<strong>о</strong> четыре <strong>в</strong>т<strong>о</strong>ричных ребрыш<strong>к</strong><strong>а</strong><br />

между гл<strong>а</strong><strong>в</strong>ными ребр<strong>а</strong>ми; ин<strong>о</strong>гд<strong>а</strong> ребр<strong>а</strong> <strong>с</strong>гл<strong>а</strong>женные.<br />

Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 17,5—21,0 мм, ширин<strong>а</strong><br />

6,5—6,8 мм (ри<strong>с</strong>. 764). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

К эт<strong>о</strong>му <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я, <strong>к</strong>р<strong>о</strong>ме мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

третичных р<strong>о</strong>д<strong>о</strong><strong>в</strong>, еще не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong>'<br />

юр<strong>с</strong><strong>к</strong>их, н<strong>а</strong>пример, Procalosoma Meunier, 1895,<br />

Nebrioides Handlirsch, 1906 и, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, мн<strong>о</strong>гие<br />

другие.<br />

СЕМЕЙСТВО CICINDELIDAE LATREILLE, 1802.<br />

СКАКУНЫ<br />

Средней <strong>в</strong>еличины или <strong>к</strong>рупные жу<strong>к</strong>и<br />

<strong>с</strong> длинными бег<strong>а</strong>тельными н<strong>о</strong>г<strong>а</strong>ми, шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> б<strong>о</strong>льшими челю<strong>с</strong>тями и <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими<br />

гл<strong>а</strong>з<strong>а</strong>ми. Н<strong>а</strong>д<strong>к</strong>рылья гл<strong>а</strong>д<strong>к</strong>ие. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Три р<strong>о</strong>д<strong>а</strong>.

СЕМЕЙСТВО COPTOCLAVIDAE<br />

PONOMARENKO, 1961<br />

Крупные <strong>в</strong><strong>о</strong>дные жу<strong>к</strong>и. Тел<strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е; <strong>с</strong>редние<br />

и з<strong>а</strong>дние н<strong>о</strong>ги пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные, упл<strong>о</strong>щенные,<br />

<strong>с</strong> р<strong>а</strong><strong>с</strong>ширенными члени<strong>к</strong><strong>а</strong>ми и без пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Личин<strong>к</strong>и <strong>с</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong>тельными н<strong>о</strong>г<strong>а</strong>ми<br />

и грызущими <strong>в</strong>ерхними челю<strong>с</strong>тями.<br />

В. юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Coptoclava Ping, 1928. Тип р<strong>о</strong>д<strong>а</strong> — С. longipoda<br />

Ping, 1928; <strong>в</strong>. юр<strong>а</strong>, Кит<strong>а</strong>й (Л<strong>а</strong>йян).<br />

Жу<strong>к</strong>: г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>в</strong>ерхние челю<strong>с</strong>ти<br />

грызущие, передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> длинн<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

длины и ширины, брюш<strong>к</strong><strong>о</strong> <strong>с</strong> ше<strong>с</strong>тью <strong>с</strong>тернит<strong>а</strong>ми,<br />

<strong>с</strong>редние и з<strong>а</strong>дние г<strong>о</strong>лени и л<strong>а</strong>п<strong>к</strong>и <strong>о</strong>чень<br />

шир<strong>о</strong><strong>к</strong>ие; длин<strong>а</strong> 34—41 мм. Личин<strong>к</strong><strong>а</strong>: <strong>в</strong>ерхние<br />

челю<strong>с</strong>ти трехзубч<strong>а</strong>тые, передние н<strong>о</strong>ги х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные,<br />

<strong>с</strong>редние и з<strong>а</strong>дние — шир<strong>о</strong><strong>к</strong>ие, пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные;<br />

длин<strong>а</strong> 10—35 мм (ри<strong>с</strong>. 765). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> В. Азии (З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье, М<strong>о</strong>нг<strong>о</strong>лия,<br />

Кит<strong>а</strong>й).<br />

СЕМЕЙСТВО DYTISC1DAE LATRE1LLE, 1825.<br />

ПЛАВУНЦЫ<br />

Крупные или <strong>с</strong>редней <strong>в</strong>еличины. Н<strong>а</strong>д<strong>к</strong>рылья<br />

гл<strong>а</strong>д<strong>к</strong>ие или <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>ми.<br />

Тел<strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е, з<strong>а</strong>дние н<strong>о</strong>ги пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные.<br />

Юр<strong>а</strong> — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 2300 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> 90 р<strong>о</strong>д<strong>а</strong>м, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 766). Мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong><strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

изучен<strong>а</strong>.<br />

СЕМЕЙСТВО HALIPL1DAE AUBE, 1836<br />

Мел<strong>к</strong>ие, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им тел<strong>о</strong>м<br />

и <strong>о</strong>чень б<strong>о</strong>льшими з<strong>а</strong>дними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми и пл<strong>а</strong><strong>в</strong><strong>а</strong>тельными<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х. В<strong>о</strong>дные жу<strong>к</strong>и<br />

(ри<strong>с</strong>. 767). Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 130 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> трем р<strong>о</strong>д<strong>а</strong>м.<br />

Один <strong>в</strong>ид р<strong>о</strong>д<strong>а</strong> Haliplus Latreille,<br />

1825—<strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> <strong>в</strong>. ми<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PALAEOGYRINIDAE<br />

SCHLECHTENDAL, 1894<br />

В<strong>о</strong>дные жу<strong>к</strong>и. Среднегруд<strong>к</strong><strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я,<br />

<strong>с</strong>редние и з<strong>а</strong>дние н<strong>о</strong>ги упл<strong>о</strong>щенные, пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные,<br />

члени<strong>к</strong>и их л<strong>а</strong>п<strong>о</strong><strong>к</strong> неп<strong>о</strong>перечные- В. <strong>о</strong>лиг<strong>о</strong>цен<br />

Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д.<br />

Palaeogyrinus Schlechtendal, 1894.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. strigatus Schlechtendal, 1894;<br />

<strong>в</strong>. <strong>о</strong>лиг<strong>о</strong>цен 3. Е<strong>в</strong>р<strong>о</strong>пы (Р<strong>о</strong>тт). Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> п<strong>о</strong>перечн<strong>а</strong>я, брюш<strong>к</strong><strong>о</strong> <strong>с</strong> ше<strong>с</strong>тью <strong>с</strong>тернит<strong>а</strong>ми,<br />

н<strong>а</strong>д<strong>к</strong>рылья <strong>с</strong> де<strong>в</strong>ятью б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>ми, г<strong>о</strong>лени<br />

и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные члени<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ных<br />

длины и ширины (ри<strong>с</strong>. 768). Один <strong>в</strong>ид. В. <strong>о</strong>лиг<strong>о</strong>цен<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

255<br />

СЕМЕЙСТВО GYRINIDAE LATREILLE, 1810.<br />

ВЕРТЯЧКИ<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong>им<br />

тел<strong>о</strong>м, пл<strong>а</strong><strong>в</strong><strong>а</strong>тельными <strong>с</strong>редними и з<strong>а</strong>дними<br />

н<strong>о</strong>г<strong>а</strong>ми, <strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличенн<strong>о</strong>й <strong>с</strong>реднегруд<strong>к</strong><strong>о</strong>й,<br />

р<strong>а</strong>зделенными гл<strong>а</strong>з<strong>а</strong>ми, у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными <strong>а</strong>нтенн<strong>а</strong>ми.<br />

Юр<strong>а</strong> — ныне. С<strong>в</strong>ыше 500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> де<strong>в</strong>яти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; 4 р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х. Из<strong>в</strong>е<strong>с</strong>тны еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> изученные<br />

<strong>в</strong>иды из юры.<br />

СЕМЕЙСТВО PAUSSIDAE LATREILLE, 1807<br />

Мел<strong>к</strong>ие жу<strong>к</strong>и <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличенными и<br />

ут<strong>о</strong>лщенными <strong>а</strong>нтенн<strong>а</strong>ми, жи<strong>в</strong>ущие <strong>в</strong> мур<strong>а</strong><strong>в</strong>ейни<strong>к</strong><strong>а</strong>х<br />

(ри<strong>с</strong>. 769). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

350 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я<br />

примерн<strong>о</strong> <strong>к</strong> 30 р<strong>о</strong>д<strong>а</strong>м; <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>о</strong>бн<strong>а</strong>ружены<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО STAPHILINIDAE LATREILLE, 1802.<br />

ХИЩНИКИ<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редних р<strong>а</strong>змер<strong>о</strong><strong>в</strong> жу<strong>к</strong>и<br />

<strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными н<strong>а</strong>д<strong>к</strong>рыльями, зн<strong>а</strong>чительн<strong>о</strong><br />

б<strong>о</strong>лее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими, чем длинн<strong>о</strong>е, гиб<strong>к</strong><strong>о</strong>е<br />

брюш<strong>к</strong><strong>о</strong> (ри<strong>с</strong>. 770). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Одн<strong>о</strong> из<br />

<strong>о</strong>бширнейших <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны,<br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющее <strong>с</strong><strong>в</strong>ыше 20 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> примерн<strong>о</strong><br />

900 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; 42 р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО PSELAPHIDAE LATREILLE, 1802<br />

Мел<strong>к</strong>ие, <strong>с</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными н<strong>а</strong>д<strong>к</strong>рыльями и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им, <strong>в</strong>здутым тел<strong>о</strong>м, <strong>о</strong>бычн<strong>о</strong> удлиненн<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й и у<strong>в</strong>еличенными м<strong>а</strong><strong>к</strong><strong>с</strong>иллярными щуп<strong>а</strong>льц<strong>а</strong>ми<br />

(ри<strong>с</strong>. 771). П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше<br />

4000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 500 р<strong>о</strong>д<strong>о</strong><strong>в</strong> г<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

23 р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО SCYDMAENIDAE LEACH, 1815<br />

Мел<strong>к</strong>ие, жи<strong>в</strong>ущие <strong>в</strong> мур<strong>а</strong><strong>в</strong>ейни<strong>к</strong><strong>а</strong>х, <strong>с</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными<br />

удлиненными з<strong>а</strong>дними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми,<br />

длинны'ми н<strong>а</strong>д<strong>к</strong>рыльями и щуп<strong>а</strong>льц<strong>а</strong>ми (ри<strong>с</strong>.<br />

772). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1300 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

60 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО SILPHIDAE LATREILLE, 1807.<br />

МЕРТВОЕДЫ<br />

Изменчи<strong>в</strong>ых р<strong>а</strong>змер<strong>о</strong><strong>в</strong>, <strong>с</strong> бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идными<br />

<strong>а</strong>нтенн<strong>а</strong>ми, уз<strong>к</strong>ими, нер<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными з<strong>а</strong>дними<br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми и длинными, <strong>в</strong>ыд<strong>а</strong>ющими<strong>с</strong>я пе-<br />

http://jurassic.ru/

редними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 773). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. С<strong>в</strong>ыше 2О0 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, 200 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО PTILIIDAE HEER, 1844<br />

И<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> мел<strong>к</strong>ие (0,25—1,40 мм),<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им тел<strong>о</strong>м, <strong>к</strong>рупн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й и <strong>о</strong>бычн<strong>о</strong><br />

пери<strong>с</strong>тыми <strong>к</strong>рыльями (ри<strong>с</strong>. 774). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 350 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. Обн<strong>а</strong>ружены <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО SCAPHIDIIDAE LATREILLE, 1806<br />

Мел<strong>к</strong>ие, <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>здутые жуч<strong>к</strong>и <strong>с</strong> длинными<br />

н<strong>а</strong>д<strong>к</strong>рыльями, <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими передними<br />

и р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными <strong>с</strong>редними и з<strong>а</strong>дними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 775). П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше<br />

550 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; три<br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичн<strong>о</strong>й ф<strong>а</strong>уне-<br />

СЕМЕЙСТВО HYDROPHILIDAE LATREILLE, 1802.<br />

ВОДОЛЮБЫ<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины, реже <strong>к</strong>рупные<br />

жу<strong>к</strong>и <strong>с</strong> бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong> <strong>в</strong>здутыми <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>а</strong>нтенн<strong>а</strong>ми, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>перед г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й переднегрудью и длинными, <strong>о</strong>бычн<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем не у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными н<strong>а</strong>д<strong>к</strong>рыльями. Н<strong>о</strong>ги<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong>тельными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми или <strong>с</strong><br />

р<strong>а</strong><strong>с</strong>ширенными л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 776). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. С<strong>в</strong>ыше 1700 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 130 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

256<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО HISTERIDAE GYLLENHAL, 1808.<br />

КАРАПУЗИКИ<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong><br />

рез<strong>к</strong><strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными и притуплённым <strong>с</strong>з<strong>а</strong>ди<br />

и <strong>с</strong>переди тел<strong>о</strong>м, ред<strong>к</strong><strong>о</strong> удлиненные, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми и н<strong>а</strong>д<strong>к</strong>рыльями, не<br />

п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими <strong>к</strong><strong>о</strong>нц<strong>а</strong> брюш<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 777).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>лее 3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 130 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; р<strong>о</strong>д Hister Linnaeus <strong>в</strong><br />

третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

Ри<strong>с</strong>. 772—779. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Scydmaenidae, у Silphidae,<br />

Ptiliidae, Scaphidiidae, Hydrophilidae, Histeridae,<br />

Scarabaeidae<br />

772. Scydmaenus sp.; <strong>о</strong>бщий <strong>в</strong>ид.х 12,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915),<br />

773. Blitophaga vetusta Lomnicki; <strong>о</strong>бщий <strong>в</strong>ид, X 5,0, чет<strong>в</strong>ерт.,<br />

3. У<strong>к</strong>р<strong>а</strong>ин<strong>а</strong> (Lomnicki, 1894). 774. Ptenldium ьр.; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х28.1, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 775. Scaphidium sp.; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х4,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 776. Helophorus polonicus Lomnicki;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 15,5, чет<strong>в</strong>ерт., 3. У<strong>к</strong>р<strong>а</strong>ин<strong>а</strong> (Lomnicki, 1894).<br />

777. Hister sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 778.<br />

Aphodius rhinocerontis Lomnicki; <strong>о</strong>бщий <strong>в</strong>ид, х 6,5, чет<strong>в</strong>ерт.<br />

3. У<strong>к</strong>р<strong>а</strong>ин<strong>а</strong> (Lomnicki, 1894). 779. Cetonia aurata Linnaeus; <strong>о</strong>бщий<br />

<strong>в</strong>ид, X 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н. 1915)

СЕМЕЙСТВО SCARABAEIDAE LATREILLE, 1802.<br />

ПЛАСТИНЧАТОУСЫЕ<br />

Средней <strong>в</strong>еличины, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупные,<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енные жу<strong>к</strong>и <strong>с</strong> <strong>а</strong>нтенн<strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>зующими бул<strong>а</strong><strong>в</strong>у из упл<strong>о</strong>щенных, ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идных<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong>. Тел<strong>о</strong> <strong>о</strong>бычн<strong>о</strong> притуплённ<strong>о</strong>е,<br />

<strong>с</strong> цеп<strong>к</strong>ими или <strong>к</strong><strong>о</strong>п<strong>а</strong>тельными н<strong>о</strong>г<strong>а</strong>ми, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итыми, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong>рыльями, <strong>к</strong>рупн<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й и грудным <strong>о</strong>тдел<strong>о</strong>м (ри<strong>с</strong>. 778, 779).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 20 ООО <strong>в</strong>ид<strong>о</strong><strong>в</strong> 1600 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; 31 р<strong>о</strong>д <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х; <strong>в</strong> юре Proteroscarabeus Grabau, 1921.<br />

СЕМЕЙСТВО LUCANIDAE LATREILLE, 1802.<br />

ЖУКИ-ОЛЕНИ<br />

Крупные, реже <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong><br />

бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идными пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми и<br />

<strong>к</strong>рупными челю<strong>с</strong>тями, ин<strong>о</strong>гд<strong>а</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющими<br />

у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> б<strong>о</strong>льших р<strong>а</strong>змер<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 780). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 900, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> 80 р<strong>о</strong>д<strong>а</strong>м;<br />

четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО DASC1LLIDAE GUERIN, 1823<br />

Средней <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щими <strong>в</strong>перед<br />

.челю<strong>с</strong>тями, ните<strong>в</strong>идными или <strong>с</strong>л<strong>а</strong>б<strong>о</strong>пиль*<br />

ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, шир<strong>о</strong><strong>к</strong><strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й,<br />

<strong>с</strong>ближенными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми, р<strong>а</strong><strong>с</strong>ширенными члени<strong>к</strong><strong>а</strong>ми<br />

л<strong>а</strong>п<strong>о</strong><strong>к</strong>. Юр<strong>а</strong> — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 400 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>, <strong>в</strong> юре <strong>о</strong>дин р<strong>о</strong>д.<br />

Mesodascilla Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М. jacobsoni Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я.<br />

Антенны не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы и передне<strong>с</strong>пин<strong>к</strong>и,<br />

т<strong>о</strong>л<strong>с</strong>тые и пильч<strong>а</strong>тые. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

12 мм (ри<strong>с</strong>. 781). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО HELODIDAE LECONTE, 1862<br />

Мел<strong>к</strong>ие, <strong>в</strong>ыпу<strong>к</strong>лые жуч<strong>к</strong>и <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им <strong>о</strong><strong>в</strong><strong>а</strong>льным<br />

тел<strong>о</strong>м, <strong>о</strong>т<strong>в</strong>е<strong>с</strong>н<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, ните<strong>в</strong>идными <strong>а</strong>нтенн<strong>а</strong>ми,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 782). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

650 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х-<br />

Ри<strong>с</strong>. 780—787. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Lucanidae, DascilHdae, Helodidae,<br />

Byrrhidae, Nosodendridae, Dermestidae, Cantharididae,<br />

Lampyridae<br />

780. Platyceruscaraboides Linneaus; <strong>о</strong>бщий <strong>в</strong>ид, X 2,7,<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915). 781. Mesodascilla jacobsoni Martynov; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X,5,4, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 782. Scirtes hemisphaericus<br />

Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 8,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915). 783. Byrrhus sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 5,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915). 784. Nosodendron fasciculare Olivier; <strong>о</strong>бщий <strong>в</strong>ид, X 8,5,<br />

«<strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 785. Attagenus pellio (Linnaeus); <strong>о</strong>бщий<br />

<strong>в</strong>ид,х 4,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 786. Cantharis sp.; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х3,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 787. Lampyrus noctiluca Linnaeus;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915)<br />

1 7 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие j-267<br />

http://jurassic.ru/

СЕМЕЙСТВО PSEPHENIDAE LACORDAIRE, 1854<br />

Мел<strong>к</strong>ие, упл<strong>о</strong>щенные, пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льные<br />

жу<strong>к</strong>и <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, длинными,<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>пильч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, <strong>к</strong>реп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми,<br />

<strong>с</strong><strong>о</strong> <strong>в</strong>здутыми бедр<strong>а</strong>ми и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми. Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я<br />

<strong>к</strong> немн<strong>о</strong>гим р<strong>о</strong>д<strong>а</strong>м; рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне групп<strong>а</strong>, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нн<strong>а</strong>я <strong>с</strong> г<strong>о</strong>рными<br />

п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>а</strong>ми, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong> ф<strong>а</strong>уне ми<strong>о</strong>цен<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО BYRRHIDAE LATREILLE, 1806<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льные,<br />

рез<strong>к</strong><strong>о</strong> <strong>в</strong>здутые же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й,<br />

глуб<strong>о</strong><strong>к</strong><strong>о</strong> <strong>в</strong>тянут<strong>о</strong>й <strong>в</strong> переднегрудь г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

<strong>к</strong>реп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> пяти- или четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 783). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 450 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО NOSODENDRIDAE ERICHSON, 1846<br />

Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> мел<strong>к</strong>ие, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льные, <strong>с</strong>ильн<strong>о</strong><br />

<strong>в</strong>здутые жу<strong>к</strong>и <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, рез<strong>к</strong><strong>о</strong> п<strong>о</strong>перечными<br />

передними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми, <strong>о</strong>чень б<strong>о</strong>льш<strong>о</strong>й з<strong>а</strong>дне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 784). П<strong>а</strong>ле<strong>о</strong>ген — ныне. 40 <strong>в</strong>ид<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> д<strong>в</strong>ум р<strong>о</strong>д<strong>а</strong>м <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО DERMESTIDAE LATREILLE, 1802.<br />

КОЖЕЕДЫ<br />

Не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины,<br />

умеренн<strong>о</strong> удлиненные или <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льные<br />

жу<strong>к</strong>и, без бле<strong>с</strong><strong>к</strong><strong>а</strong>, <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей <strong>в</strong>низ г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми; н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong><strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными передними и <strong>с</strong>редними<br />

и <strong>с</strong>ближенными з<strong>а</strong>дними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 785). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 700 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО CANTHARIDIDAE HEYDEN,<br />

REITTER, WE1SE, 1883.<br />

МЯГКОТЕЛО<br />

Средней <strong>в</strong>еличины, реже мел<strong>к</strong>ие или <strong>к</strong>рупные,<br />

м<strong>а</strong>л<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лые, удлиненные жу<strong>к</strong>и <strong>с</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

переднегрудью, <strong>с</strong> <strong>с</strong>емью — <strong>в</strong><strong>о</strong><strong>с</strong>емью<br />

<strong>с</strong>тернит<strong>а</strong>ми брюш<strong>к</strong><strong>а</strong> и <strong>с</strong>л<strong>а</strong>быми н<strong>о</strong>г<strong>а</strong>ми<br />

(ри<strong>с</strong>. 786)- П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1300 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

258<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО LAMPYRIDAE LEACH, 1817.<br />

СВЕТЛЯКИ<br />

Средней <strong>в</strong>еличины, удлиненные, не<strong>в</strong>ыпу<strong>к</strong>лые<br />

жу<strong>к</strong>и, <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, <strong>с</strong><strong>к</strong>рыт<strong>о</strong>й п<strong>о</strong>д н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющей<br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, не<strong>с</strong>ущей пильч<strong>а</strong>тые <strong>а</strong>нтенны,<br />

<strong>с</strong> длинными у\<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> н<strong>а</strong>д<strong>к</strong>рыльями, не<strong>с</strong>ущими<br />

эпиле<strong>в</strong>ры; н<strong>о</strong>ги <strong>с</strong><strong>о</strong> <strong>в</strong>здутыми бедр<strong>а</strong>ми и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми. С<strong>а</strong>м<strong>к</strong>и чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зны<br />

(ри<strong>с</strong>. 787). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 18 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

60 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

<strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО LYCIDAE LACORDAIRE, 1857<br />

Средней или <strong>к</strong>рупн<strong>о</strong>й <strong>в</strong>еличины, умеренн<strong>о</strong><br />

удлиненные, пл<strong>о</strong><strong>с</strong><strong>к</strong>ие жу<strong>к</strong>и <strong>с</strong> м<strong>а</strong>лень<strong>к</strong><strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

<strong>с</strong><strong>к</strong>рыт<strong>о</strong>й п<strong>о</strong>д передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> пильч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> б<strong>о</strong>льшими пл<strong>о</strong><strong>с</strong><strong>к</strong>ими н<strong>а</strong>д<strong>к</strong>рыльями,<br />

не<strong>с</strong>ущими <strong>с</strong>етч<strong>а</strong>т<strong>о</strong>е жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние, и<br />

н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенными бедр<strong>а</strong>ми и <strong>к</strong>рупными<br />

<strong>в</strong>ертлуг<strong>а</strong>ми (ри<strong>с</strong>. 788). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>лее<br />

2800, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> 53 р<strong>о</strong>д<strong>а</strong>м; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО MELYRIDAE NEWMANN, 1834.<br />

МАЛАШКИ<br />

Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> мел<strong>к</strong>ие, умеренн<strong>о</strong> удлиненные<br />

жу<strong>к</strong>и <strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>перед,<br />

<strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й,<br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>ими н<strong>а</strong>д<strong>к</strong>рыльями, ин<strong>о</strong>гд<strong>а</strong><br />

у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными, ше<strong>с</strong>тью-<strong>с</strong>емью <strong>с</strong>тернит<strong>а</strong>ми<br />

брюш<strong>к</strong><strong>а</strong>, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (передние<br />

л<strong>а</strong>п<strong>к</strong>и ин<strong>о</strong>гд<strong>а</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 789).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 1800 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО CLERIDAE LATREILLE, 1802.<br />

ПЕСТРЯКИ<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины, удлиненные,<br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые жу<strong>к</strong>и <strong>с</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>перед<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, пильч<strong>а</strong>тыми или г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й переднегрудью, <strong>к</strong>реп<strong>к</strong>ими<br />

и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> длинными н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми и <strong>с</strong> пятью-ше<strong>с</strong>тью <strong>с</strong>тернит<strong>а</strong>ми<br />

брюш<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 790). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 2500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 200 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ELATER1DAE LEACH, 1815.<br />

ЩЕЛКУНЫ<br />

Средней или <strong>к</strong>рупн<strong>о</strong>й <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тым<br />

тел<strong>о</strong>м, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>тянут<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й упл<strong>о</strong>щенн<strong>о</strong>й переднегрудью, п<strong>о</strong>д-

<strong>в</strong>ижн<strong>о</strong>й <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>с</strong>реднегруди, ните<strong>в</strong>идными,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> пильч<strong>а</strong>тыми или гребенч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными <strong>в</strong>переди гл<strong>а</strong>з, ниже<br />

<strong>к</strong>р<strong>а</strong>я лб<strong>а</strong>. Юр<strong>а</strong> — ныне. Обширн<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>:<br />

<strong>с</strong><strong>в</strong>ыше 7000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> 350 р<strong>о</strong>д<strong>а</strong>м<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 80 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я<br />

примерн<strong>о</strong> <strong>к</strong> 20 р<strong>о</strong>д<strong>а</strong>м <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х; не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пл<strong>о</strong>х<strong>о</strong> изученных р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

юре 3. Е<strong>в</strong>р<strong>о</strong>пы; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> юре К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Tersus Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Т. crassicornis Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ей длины, п<strong>о</strong>перечн<strong>а</strong>я. Н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong><strong>к</strong>. Г<strong>о</strong>лени т<strong>о</strong>н<strong>к</strong>ие и пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>ребри<strong>с</strong>тые.<br />

Длин<strong>а</strong> тел<strong>а</strong> 12,25 мм, длин<strong>а</strong> передне<strong>с</strong>пин<strong>к</strong>и<br />

2,15 мм (ри<strong>с</strong>. 791). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО MELASIDAE LEACH, 1817<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и, п<strong>о</strong>х<strong>о</strong>жие<br />

н<strong>а</strong> щел<strong>к</strong>ун<strong>о</strong><strong>в</strong>. Антенны р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

между гл<strong>а</strong>з<strong>а</strong>ми, ните<strong>в</strong>идные или пери<strong>с</strong>тые,<br />

<strong>с</strong>ближенные <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями (ри<strong>с</strong>. 792). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1100 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я<br />

<strong>к</strong> 150 р<strong>о</strong>д<strong>а</strong>м <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> третичн<strong>о</strong>й<br />

ф<strong>а</strong>уне из<strong>в</strong>е<strong>с</strong>тны три р<strong>о</strong>д<strong>а</strong>.<br />

СЕМЕЙСТВО THROSCIDAE LACORDAIRE, 1857<br />

Мел<strong>к</strong>ие, пл<strong>о</strong>тные жуч<strong>к</strong>и <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>тянут<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong>им тел<strong>о</strong>м н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> (ри<strong>с</strong>. 793). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

200 <strong>в</strong>ид<strong>о</strong><strong>в</strong> немн<strong>о</strong>гих р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО BUPRESTIDAE LEACH, 1815. ЗЛАТКИ<br />

Средней <strong>в</strong>еличины, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>к</strong>рупные, ред<strong>к</strong><strong>о</strong><br />

мел<strong>к</strong>ие жу<strong>к</strong>и, <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тым тел<strong>о</strong>м,<br />

<strong>в</strong>тянут<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> ните<strong>в</strong>идными или пильч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong><strong>о</strong>единенными, неп<strong>о</strong>д<strong>в</strong>ижными<br />

р<strong>а</strong>здел<strong>а</strong>ми груди, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими, <strong>к</strong>реп<strong>к</strong>ими<br />

н<strong>о</strong>г<strong>а</strong>ми и <strong>в</strong>ыпу<strong>к</strong>лыми н<strong>а</strong>д<strong>к</strong>рыльями<br />

(ри<strong>с</strong>. 794). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Обширн<strong>о</strong>е, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>; <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 12 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 400 р<strong>о</strong>д<strong>о</strong><strong>в</strong>;<br />

17 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

Ри<strong>с</strong>. 788—795. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Lycidae, Melyridae, Cleridae,<br />

Elateridae, Melasidae, Throscidae, Buprestidae, Lymexylidae<br />

788. Lycostomus sp.; <strong>о</strong>бщий <strong>в</strong>ид. X 2,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915),<br />

789. Malachius bipustulatus Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 4,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 790. Eucnemis capucina Ahrens, <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х5,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 791. Tersus crassicornis Martynov;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х4,6, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926).<br />

792. Trichodes sp.; <strong>о</strong>бщий <strong>в</strong>ид, Х3,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915).<br />

793. Throscus dermestoides Linaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 9,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 794. Buprestis sp.; <strong>о</strong>бщий <strong>в</strong>ид, Х2.7, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н 1915). 795. Lymexylon navale\Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х3,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915)<br />

259<br />

http://jurassic.ru/

Ри<strong>с</strong>. 796—802. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Bostrychidae, Lyctiriae, Anobiidae,<br />

Ptinidae, Cisidae, Ostomatidae<br />

796. Apate sp.; <strong>о</strong>бщий <strong>в</strong>ид, X2,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 797. Indus<br />

sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 798. Ап<strong>о</strong>Ыит<br />

rujipes Fabricius; <strong>о</strong>бщий <strong>в</strong>ид,; Х8,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. /(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915).<br />

799. Ptinus sp.; <strong>о</strong>бщий <strong>в</strong>ид, Х9,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 800.<br />

Cis boleti Scopoli; <strong>о</strong>бщий <strong>в</strong>ид, XI 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915).<br />

801. Ostoma jerrugineum Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, Х3,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

неу<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными н<strong>а</strong>д<strong>к</strong>рыльями, <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими<br />

эпипле<strong>в</strong>р<strong>а</strong>ми и <strong>с</strong>л<strong>а</strong>быми н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 800).' П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 350 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 25 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО OSTOMATIDAE REITTER, 1882<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong><br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>перед, <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong><strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, п<strong>о</strong>перечными<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными передними т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бычн<strong>о</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>им и шир<strong>о</strong><strong>к</strong>им тел<strong>о</strong>м <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенными<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong>ыми <strong>к</strong><strong>а</strong>ем<strong>к</strong><strong>а</strong>ми длинных н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> и<br />

передне<strong>с</strong>пин<strong>к</strong>и-: Н<strong>о</strong>ги <strong>с</strong>л<strong>а</strong>бые <strong>с</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

пер<strong>в</strong>ым члени<strong>к</strong><strong>о</strong>м л<strong>а</strong>п<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 801). Юр<strong>а</strong> —<br />

ныне. Б<strong>о</strong>лее 600 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х; <strong>о</strong><strong>с</strong><strong>о</strong>бый<br />

р<strong>о</strong>д <strong>в</strong> юре.<br />

Lithostoma Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

L. expansion Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Антенны <strong>с</strong> <strong>о</strong>чень<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным <strong>к</strong><strong>о</strong>нце<strong>в</strong>ым <strong>в</strong>здутием; передний<br />

<strong>к</strong>р<strong>а</strong>й передне<strong>с</strong>пин<strong>к</strong>и р<strong>о</strong><strong>в</strong>ный, без <strong>в</strong>ыем<strong>к</strong>и;<br />

н<strong>а</strong>д<strong>к</strong>рылья без <strong>в</strong>ыпу<strong>к</strong>лых пр<strong>о</strong>д<strong>о</strong>льных ребер.<br />

Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 6 мм (ри<strong>с</strong>. 802). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО NITIDULIDAE LATREILLE, 1802<br />

Мел<strong>к</strong>ие, умеренн<strong>о</strong> удлиненные или <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

тел<strong>о</strong>м, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые жу<strong>к</strong>и <strong>с</strong><br />

<strong>о</strong>бычн<strong>о</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>перед г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>к</strong>реп<strong>к</strong>ими, н<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми (<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>) р<strong>а</strong><strong>с</strong>ширенными<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми, <strong>о</strong>чень ч<strong>а</strong><strong>с</strong>т<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными<br />

н<strong>а</strong>д<strong>к</strong>рыльями (ри<strong>с</strong>. 803). Юр<strong>а</strong> — ныне.<br />

С<strong>в</strong>ыше 2400 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 160 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х; <strong>о</strong>дин р<strong>о</strong>д<br />

<strong>в</strong> юре-<br />

Nitidulina Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

N. eclavata Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным <strong>в</strong>здутием;<br />

передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong> п<strong>о</strong>лт<strong>о</strong>р<strong>а</strong> р<strong>а</strong>з<strong>а</strong> шире длины.<br />

Длин<strong>а</strong> тел<strong>а</strong> 4,9 мм (ри<strong>с</strong>. 804). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Ри<strong>с</strong>. 803—811. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Nitidulidae, Rhizophagidae, Cucujidae,<br />

Phalacridae, Erotylidae, Lathridiidae, Mycetophagidae,<br />

Colydiidae<br />

803. Nitidula blpunctata Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X8.2, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 804. Nitidulina eclavata Martynov; A — <strong>о</strong>бщий<br />

<strong>в</strong>ид, X 1,5; Б — <strong>а</strong>нтенн<strong>а</strong>, x 2,8; <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1926). 805 Rhlzophagus bipustulatus Fabricius <strong>о</strong>бщий <strong>в</strong>ид,<br />

XI 1,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 806. Silvanus sp.; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 12,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 807. Phalacrus sp.; <strong>о</strong>бщий <strong>в</strong>ид,<br />

х8,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 808. Tritoma sp.; <strong>о</strong>бщий <strong>в</strong>ид, Х7,5,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 809. Lathridius sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 16,5,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 810. Trtphyllus bicolor Fabricius; <strong>о</strong>бщий<br />

<strong>в</strong>ид, X 10,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 811. Bothrideres sp. <strong>о</strong>бщий<br />

<strong>в</strong>ид, Хб,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915)<br />

http://jurassic.ru/

СЕМЕЙСТВО RHIZOPHAGIDAE<br />

REDTENBACHER, 1845<br />

Мел<strong>к</strong>ие, удлиненные жу<strong>к</strong>и <strong>с</strong> <strong>к</strong>рупн<strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й<br />

<strong>в</strong>перед г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> умеренн<strong>о</strong> <strong>в</strong>здутыми<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими<br />

н<strong>о</strong>г<strong>а</strong>ми, <strong>в</strong>ытянут<strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й и<br />

длинными н<strong>а</strong>д<strong>к</strong>рыльями, не п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими<br />

<strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 805). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong><strong>в</strong>ыше 40 <strong>в</strong>ид<strong>о</strong><strong>в</strong> трех р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>дин из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>бн<strong>а</strong>ружен <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре).<br />

СЕМЕЙСТВО CUCUJIDAE LATREILLE, 1802<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины, удлиненные<br />

и упл<strong>о</strong>щенные жу<strong>к</strong>и <strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й, т<strong>о</strong>рч<strong>а</strong>щей<br />

<strong>в</strong>перед г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> удлиненными г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

или ните<strong>в</strong>идными <strong>а</strong>нтенн<strong>а</strong>ми, не з<strong>а</strong>м<strong>к</strong>нутыми<br />

<strong>с</strong><strong>в</strong>ерху ям<strong>к</strong><strong>а</strong>ми <strong>с</strong>редних т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong>, длинными<br />

н<strong>а</strong>д<strong>к</strong>рыльями, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не<strong>с</strong>ущими, <strong>к</strong><strong>а</strong><strong>к</strong> и передне<strong>с</strong>пин<strong>к</strong><strong>а</strong>,<br />

пр<strong>о</strong>д<strong>о</strong>льные ребр<strong>а</strong> или буг<strong>о</strong>р<strong>к</strong>и<br />

(ри<strong>с</strong>. 806). П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 1300 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

100 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; пять р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО PHALACRIDAE LEACH, 1815<br />

Мел<strong>к</strong>ие, <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лые<br />

жу<strong>к</strong>и <strong>с</strong><strong>о</strong> <strong>в</strong>тянут<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми,<br />

<strong>с</strong> гл<strong>а</strong>д<strong>к</strong>ими н<strong>а</strong>д<strong>к</strong>рыльями, ин<strong>о</strong>гд<strong>а</strong><br />

не<strong>с</strong>ущими т<strong>о</strong>н<strong>к</strong>ие б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong>и и п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими<br />

<strong>в</strong><strong>с</strong>е брюш<strong>к</strong><strong>о</strong>, и <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми (ри<strong>с</strong>. 807).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕСТВО EROTYLIDAE LATREILLE, 1802<br />

Мел<strong>к</strong>ие, <strong>с</strong>редние, реже <strong>к</strong>рупные жу<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

<strong>в</strong>тянут<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными ш<strong>а</strong>р<strong>о</strong><strong>в</strong>идными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми, <strong>с</strong><br />

з<strong>а</strong>м<strong>к</strong>нутыми т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong>ыми ям<strong>к</strong><strong>а</strong>ми, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими и<br />

<strong>к</strong>реп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми и <strong>с</strong> длинными н<strong>а</strong>д<strong>к</strong>рыльями<br />

(ри<strong>с</strong>. 808). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 3800 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

230 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО LATHRIDIIDAE REDTENBACHER, 1845<br />

Мел<strong>к</strong>ие или <strong>о</strong>чень мел<strong>к</strong>ие пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тые<br />

жуч<strong>к</strong>и <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> уз<strong>к</strong><strong>о</strong>й переднегрудью, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не<strong>с</strong>ущей<br />

рез<strong>к</strong>ую <strong>с</strong><strong>к</strong>ульптуру, <strong>с</strong> длинными н<strong>а</strong>д<strong>к</strong>рыльями<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льными ребр<strong>а</strong>ми<br />

(ри<strong>с</strong>. 809). П<strong>а</strong>ле<strong>о</strong>ген—• ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 550 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и <strong>в</strong><br />

<strong>о</strong>лиг<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

262<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО MYCETOPHAGIDAE LEACH, 1815<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины <strong>о</strong><strong>в</strong><strong>а</strong>льные<br />

жу<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, <strong>в</strong>тянут<strong>о</strong>й <strong>в</strong> переднегрудь,<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, длинными н<strong>а</strong>д<strong>к</strong>рыльями<br />

и д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> т<strong>о</strong>н<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми!)<br />

(ри<strong>с</strong>. 810). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 200 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

<strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> н. <strong>о</strong>лиг<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО COLYD11DAE ERICHSON, 1845<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянутые<br />

или упл<strong>о</strong>щенные жуч<strong>к</strong>и <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й <strong>в</strong> <strong>в</strong>иде буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> и ребер,<br />

<strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й т<strong>о</strong>рч<strong>а</strong>щей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, длинными н<strong>а</strong>д<strong>к</strong>рыльями <strong>с</strong> ребр<strong>а</strong>ми<br />

и ям<strong>к</strong><strong>а</strong>ми, <strong>с</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong>реп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 811). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1400 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 100 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ENDOMYCHIDAE LEACH, 1810<br />

Мел<strong>к</strong>ие <strong>о</strong><strong>в</strong><strong>а</strong>льные, умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые жуч<strong>к</strong>и<br />

<strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>тянут<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> пр<strong>о</strong><strong>с</strong>тыми<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ых л<strong>а</strong>п<strong>к</strong><strong>а</strong>х<br />

(или трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ых) л<strong>а</strong>п<strong>к</strong><strong>а</strong>х (ри<strong>с</strong>. 812). Жи<strong>в</strong>ут<br />

<strong>в</strong> гриб<strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1000<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> 90 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО COCCINELLIDAE LATREILLE, 1807<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льные,<br />

ин<strong>о</strong>гд<strong>а</strong> п<strong>о</strong>чти <strong>о</strong><strong>к</strong>руглые <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лые<br />

жу<strong>к</strong>и. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенными<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии или <strong>с</strong>н<strong>а</strong>бженными зубц<strong>а</strong>ми<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ых, ред<strong>к</strong><strong>о</strong> четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>х (ри<strong>с</strong>. 813). Жи<strong>в</strong>ут н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 3400 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

240 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО OEDEMERIDAE LATREILLE, 1810<br />

Средней <strong>в</strong>еличины или мел<strong>к</strong>ие, <strong>с</strong>ильн<strong>о</strong> удлиненные<br />

жу<strong>к</strong>и <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щей <strong>в</strong>перед <strong>в</strong>ытянут<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> <strong>к</strong>рупными гл<strong>а</strong>з<strong>а</strong>ми, длинными, т<strong>о</strong>н<strong>к</strong>ими<br />

<strong>а</strong>нтенн<strong>а</strong>ми, уз<strong>к</strong><strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, <strong>с</strong> длинными,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я <strong>в</strong>ершин<strong>а</strong>ми н<strong>а</strong>д<strong>к</strong>рыльями<br />

и <strong>к</strong>рупными т<strong>о</strong>рч<strong>а</strong>щими т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми.<br />

Средние и передние л<strong>а</strong>п<strong>к</strong>и пятичлени<strong>к</strong><strong>о</strong>-

<strong>в</strong>ые, з<strong>а</strong>дние — четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 814).<br />

Юр<strong>а</strong> — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 800 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 70 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> юре, <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Necromera Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

JV. baeckmani Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

р<strong>а</strong><strong>с</strong>ширен<strong>а</strong> <strong>в</strong> передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине и зн<strong>а</strong>чительн<strong>о</strong><br />

уже <strong>о</strong>б<strong>о</strong>их н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые не<strong>с</strong>ут<br />

нея<strong>с</strong>ные п<strong>о</strong>л<strong>о</strong><strong>с</strong>ы. Антенны <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> длиннее<br />

передне<strong>с</strong>пин<strong>к</strong>и. Длин<strong>а</strong> тел<strong>а</strong> 7,75 мм (ри<strong>с</strong>. 815).<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>. Один <strong>в</strong>ид.<br />

СЕМЕЙСТВО PYTHIDAE SOLSKY, 1834<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины, умеренн<strong>о</strong> удлиненные<br />

жу<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й, т<strong>о</strong>рч<strong>а</strong>щей и <strong>в</strong>ытянут<strong>о</strong>й<br />

<strong>в</strong>перед г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> т<strong>о</strong>н<strong>к</strong>ими, реже пильч<strong>а</strong>тыми<br />

или <strong>с</strong>л<strong>а</strong>б<strong>о</strong>г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми.<br />

Л<strong>а</strong>п<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> у предыдущег<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

(ри<strong>с</strong>. 816). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 300 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

46 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong><br />

<strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО PYROCHROIDAE LATREILLE, 1807<br />

Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины<br />

жу<strong>к</strong>и <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щей, рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>о</strong>й <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> пильч<strong>а</strong>тыми или пери<strong>с</strong>тыми, ред<strong>к</strong><strong>о</strong><br />

ните<strong>в</strong>идными <strong>а</strong>нтенн<strong>а</strong>ми. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> м<strong>а</strong>л<strong>а</strong>,<br />

шире г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, н<strong>о</strong> уже н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

р<strong>а</strong><strong>с</strong>ширяют<strong>с</strong>я <strong>к</strong> з<strong>а</strong>днему <strong>к</strong><strong>о</strong>нцу и п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<br />

<strong>в</strong><strong>с</strong>е брюш<strong>к</strong><strong>о</strong>. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные <strong>с</strong> б<strong>о</strong>льшими<br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми передними<br />

и <strong>с</strong>редними и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми з<strong>а</strong>дними<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 817). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

150 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин<br />

р<strong>о</strong>д <strong>в</strong>'п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ADERIDAE AUCT.<br />

Мел<strong>к</strong>ие или <strong>о</strong>чень мел<strong>к</strong>ие, умеренн<strong>о</strong> <strong>в</strong>ытянутые<br />

<strong>в</strong>ыпу<strong>к</strong>лые жуч<strong>к</strong>и <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й<br />

<strong>в</strong>низ и <strong>в</strong>тянут<strong>о</strong>й <strong>в</strong> переднегрудь, не<strong>с</strong>ущей<br />

<strong>к</strong>рупные гл<strong>а</strong>з<strong>а</strong> и т<strong>о</strong>н<strong>к</strong>ие чет<strong>к</strong><strong>о</strong><strong>в</strong>идные и <strong>с</strong>л<strong>а</strong>б<strong>о</strong>г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тые<br />

<strong>а</strong>нтенны. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> уже<br />

длинных и гл<strong>а</strong>д<strong>к</strong>их н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>. Передние и<br />

<strong>с</strong>редние т<strong>а</strong>зи<strong>к</strong>и т<strong>о</strong>рч<strong>а</strong>щие, л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченным<br />

предп<strong>о</strong><strong>с</strong>ледним члени<strong>к</strong><strong>о</strong>м, пяти- и че-<br />

Ри<strong>с</strong>. 812—819. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Endomychidae, Coccinellidae,<br />

Oedemeridae, Pythidae, Pyrochroidae, Aderidae, Anthicidae<br />

812. Lycoperdina succincta Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 6,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 813. Coccinella sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 6,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 814. Oedemera lurida Marsham; <strong>о</strong>бщий <strong>в</strong>ид,<br />

SC,6,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 815. Necromera baeckmani Martynov;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х5,2, <strong>в</strong>.. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925).<br />

816. Mycterus curculionoides Illiger; <strong>о</strong>бщий <strong>в</strong>ид, Х,3,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

827<br />

264<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ANTHICIDAE LATREILLE, 1825<br />

Мел<strong>к</strong>ие, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>с</strong>тр<strong>о</strong>йные жуч<strong>к</strong>и <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щей<br />

<strong>к</strong>рупн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

<strong>о</strong>т переднегруди и не<strong>с</strong>ущей чет<strong>к</strong><strong>о</strong><strong>в</strong>идные <strong>а</strong>нтенны.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>з<strong>а</strong>ди рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>а</strong> и<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>. Т<strong>а</strong>зи<strong>к</strong>и т<strong>о</strong>рч<strong>а</strong>щие,<br />

л<strong>а</strong>п<strong>к</strong>и пяти- и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 819).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 2200 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и <strong>в</strong> <strong>о</strong>лиг<strong>о</strong>цене<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО MORDELLIDAE LATREILLE, 1802<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и, з<strong>а</strong>метн<strong>о</strong><br />

<strong>с</strong>ж<strong>а</strong>тые <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>, <strong>с</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренным <strong>с</strong>з<strong>а</strong>ди тел<strong>о</strong>м,<br />

неб<strong>о</strong>льш<strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>низ г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, ните<strong>в</strong>идными<br />

или пильч<strong>а</strong>тыми <strong>а</strong>нтенн<strong>а</strong>ми, шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, р<strong>а</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong> ширине н<strong>а</strong>д<strong>к</strong>рыльям,<br />

<strong>с</strong> <strong>с</strong>уж<strong>а</strong>ющими<strong>с</strong>я и не п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими <strong>в</strong><strong>с</strong>е<br />

брюш<strong>к</strong><strong>о</strong> н<strong>а</strong>д<strong>к</strong>рыльями. Пять-ше<strong>с</strong>ть <strong>с</strong>тернит<strong>о</strong><strong>в</strong><br />

брюш<strong>к</strong><strong>а</strong>. З<strong>а</strong>дние л<strong>а</strong>п<strong>к</strong>и* длиннее г<strong>о</strong>леней<br />

(ри<strong>с</strong>. 820). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1000 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

24 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и<br />

ми<strong>о</strong>цене Сицилии.<br />

СЕМЕЙСТВО PRAEMORDELLIDAE<br />

SCEGOLEVA-BAROVSKAJA, 1929<br />

|[nom. transl. Rohdendorf, hie<br />

(ex Praemordellinae Scegoleva-Barovskaja, 1929)]<br />

Средней <strong>в</strong>еличины <strong>с</strong>ж<strong>а</strong>тые <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> жу<strong>к</strong>и<br />

<strong>с</strong> рез<strong>к</strong><strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренным брюш<strong>к</strong><strong>о</strong>м, не<strong>с</strong>ущим <strong>с</strong>емь<br />

<strong>с</strong>тернит<strong>о</strong><strong>в</strong> и не п<strong>о</strong><strong>к</strong>рытым н<strong>а</strong>д<strong>к</strong>рыльями д<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>.<br />

З<strong>а</strong>дние л<strong>а</strong>п<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>леней. Близ<strong>к</strong>и <strong>к</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у<br />

Mordellidae, <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных <strong>с</strong>тернит<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

з<strong>а</strong>дними л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Praemordelta Scegoleva-Barovskaja,<br />

1929. Тип р<strong>о</strong>д<strong>а</strong> — P. martynovi Scegolevy-Barovskaja,<br />

1929; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Г<strong>о</strong>лени <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й шп<strong>о</strong>р<strong>о</strong>й.<br />

Члени<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>в</strong>еличины.<br />

Длин<strong>а</strong> тел<strong>а</strong> 7,3 мм (ри<strong>с</strong>. 821). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Ри<strong>с</strong>. 820—827. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Mordellidae, Praemordellidae,<br />

Scraptiidae, Serropalpidae, Meloidae, Lagriidae, Alleculidae,<br />

Tenebrionidae<br />

820. Anaspis sp.; <strong>о</strong>бщий <strong>в</strong>ид, X7,0,* <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915).<br />

821. Praemordella martynovi Scegoleva-Barovskaya; <strong>о</strong>бщий <strong>в</strong>ид.<br />

X 6,6, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Щег<strong>о</strong>ле<strong>в</strong><strong>а</strong>-Б<strong>а</strong>р<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я, 1929). 822.<br />

Scraptia sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 14,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н. 1915. 823-.<br />

Orchesla micans Р<strong>а</strong>пгег;<strong>о</strong>бЩий <strong>в</strong>ид, Хб,6, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915).<br />

824. Lytta sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 1,9, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 825.<br />

Lagria hirta Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 3,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н.<br />

1915). 826. Pseudocistella cerambycoides Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 4,8, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 827. Helops sp.; <strong>о</strong>бщий <strong>в</strong>ид.<br />

X 3,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915)

СЕМЕЙСТВО SCRAPTIIDAE MULSANT, 1856<br />

Мел<strong>к</strong>ие жуч<strong>к</strong>и <strong>с</strong> умеренн<strong>о</strong> удлиненным тел<strong>о</strong>м,<br />

<strong>к</strong>рупн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>низ и<br />

<strong>в</strong>перед, ните<strong>в</strong>идными <strong>а</strong>нтенн<strong>а</strong>ми, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й<br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, лишь ед<strong>в</strong><strong>а</strong> б<strong>о</strong>лее уз<strong>к</strong><strong>о</strong>й, чем<br />

ширин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья, п<strong>о</strong><strong>с</strong>ледние длинные и п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<br />

<strong>в</strong><strong>с</strong>е брюш<strong>к</strong><strong>о</strong> (ри<strong>с</strong>. 822). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 125 <strong>в</strong>ид<strong>о</strong><strong>в</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО SERROPALPIDAE<br />

LATREILLE, 1829<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины <strong>о</strong><strong>в</strong><strong>а</strong>льные<br />

или удлиненные жу<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>низ и не<strong>с</strong>ущей ните<strong>в</strong>идные или<br />

умеренн<strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тые <strong>а</strong>нтенны, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й, т<strong>о</strong>н<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми <strong>с</strong> д<strong>в</strong>умя шп<strong>о</strong>р<strong>а</strong>ми<br />

и пяти- и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 823). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 430 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

90 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>-ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО MELOIDAE GYLLENHAL, 1810.<br />

НАРЫВНИКИ<br />

Средней <strong>в</strong>еличины или д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong>рупные<br />

жу<strong>к</strong>и <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й <strong>о</strong>т переднегруди<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> умеренн<strong>о</strong> удлиненными, р<strong>а</strong>зличн<strong>о</strong><br />

п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енными <strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й переднегрудью,<br />

б<strong>о</strong>лее уз<strong>к</strong><strong>о</strong>й, чем н<strong>а</strong>д<strong>к</strong>рылья, и <strong>к</strong>реп<strong>к</strong>ими,<br />

цеп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми (ри<strong>с</strong>. 824). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 2300 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 90 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х-<br />

СЕМЕЙСТВО RHIPIPHORIDAE COSTA, 1850<br />

Средней <strong>в</strong>еличины пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тые жу<strong>к</strong>и <strong>с</strong><br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тью тел<strong>а</strong>, <strong>с</strong><br />

неб<strong>о</strong>льш<strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>о</strong>й <strong>в</strong>низ и н<strong>а</strong>з<strong>а</strong>д г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>й передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й и <strong>о</strong>бычн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными<br />

н<strong>а</strong>д<strong>к</strong>рыльями. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 350 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО LAGRIIDAE LATREILLE, 1825<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и <strong>с</strong> умеренн<strong>о</strong><br />

удлиненным тел<strong>о</strong>м, <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong> т<strong>о</strong>рч<strong>а</strong>щей<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong> ните<strong>в</strong>идными или <strong>с</strong>л<strong>а</strong>б<strong>о</strong>пильч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й, п<strong>о</strong>чти цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

переднегрудью, зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>лее уз<strong>к</strong><strong>о</strong>й, чем<br />

н<strong>а</strong>д<strong>к</strong>рылья, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют <strong>в</strong><strong>с</strong>е брюш<strong>к</strong><strong>о</strong> и<br />

р<strong>а</strong><strong>с</strong>ширяют<strong>с</strong>я <strong>с</strong>з<strong>а</strong>ди. Н<strong>о</strong>ги длинные, л<strong>а</strong>п<strong>к</strong>и<br />

пяти- и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 825). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 650 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 60 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

265<br />

СЕМЕЙСТВО ALLECUL1DAE MULSANT, 1856.<br />

ПЫЛЬЦЕЕДЫ<br />

Средней <strong>в</strong>еличины или мел<strong>к</strong>ие жу<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

т<strong>о</strong>рч<strong>а</strong>щей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>лее<br />

уз<strong>к</strong><strong>о</strong>й, чем передне<strong>с</strong>пин<strong>к</strong><strong>а</strong>, и не<strong>с</strong>ущей ните<strong>в</strong>идные,<br />

б<strong>о</strong>лее или менее пильч<strong>а</strong>тые <strong>а</strong>нтенны. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong><br />

<strong>с</strong>з<strong>а</strong>ди шир<strong>о</strong><strong>к</strong><strong>а</strong>я. Н<strong>а</strong>д<strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong><br />

п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют <strong>в</strong><strong>с</strong>е брюш<strong>к</strong><strong>о</strong>. Н<strong>о</strong>ги длинные, л<strong>а</strong>п<strong>к</strong>и<br />

<strong>с</strong> пятью и четырьмя члени<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 826).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1400 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 120 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>рменн<strong>о</strong>й ф<strong>а</strong>уне; пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО TENEBRIONIDAE LATREILLE, 1802.<br />

ЧЕРНОТЕЛКИ<br />

Средней <strong>в</strong>еличины или <strong>к</strong>рупные, ред<strong>к</strong><strong>о</strong> мел<strong>к</strong>ие<br />

жу<strong>к</strong>и. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не<strong>с</strong>ет шир<strong>о</strong><strong>к</strong>ую л<strong>о</strong>бную пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у<br />

<strong>в</strong> <strong>в</strong>иде щит<strong>к</strong><strong>а</strong>, п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющег<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>а</strong>нтенн; <strong>а</strong>нтенны ните<strong>в</strong>идные, чет<strong>к</strong><strong>о</strong><strong>в</strong>идные или<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тые. Средние и з<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и б<strong>о</strong>лее<br />

или менее р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные. Н<strong>а</strong>д<strong>к</strong>рылья п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<br />

<strong>в</strong><strong>с</strong>е брюш<strong>к</strong><strong>о</strong>. Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие и длинные,<br />

л<strong>а</strong>п<strong>к</strong>и пяти- и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 827).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 14 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО CERAMBYCIDAE LATREILLE, 1802.<br />

ДРОВОСЕКИ, ИЛИ УСАЧИ<br />

Средней <strong>в</strong>еличины или <strong>к</strong>рупные жу<strong>к</strong>и <strong>с</strong> удлиненным<br />

тел<strong>о</strong>м, умеренн<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong><br />

<strong>с</strong>ильн<strong>о</strong> удлиненными ните<strong>в</strong>идными или чет<strong>к</strong><strong>о</strong><strong>в</strong>идными<br />

<strong>а</strong>нтенн<strong>а</strong>ми, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, длинными<br />

н<strong>а</strong>д<strong>к</strong>рыльями, ин<strong>о</strong>гд<strong>а</strong> не<strong>с</strong>ущими з<strong>а</strong>метные <strong>с</strong>леды<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния, <strong>к</strong>реп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми и л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми<br />

<strong>с</strong> <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенным чет<strong>в</strong>ертым члени<strong>к</strong><strong>о</strong>м и<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие эт<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>а</strong>жущими<strong>с</strong>я четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

(ри<strong>с</strong>. 828). Юр<strong>а</strong> — ныне. Очень <strong>о</strong>бширн<strong>о</strong>е<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>: <strong>с</strong><strong>в</strong>ыше<br />

17 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> примерн<strong>о</strong> 2400 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

31 р<strong>о</strong>д <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> юре.<br />

Parandrexis Martynov, 1926. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. parvula Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я,<br />

п<strong>о</strong>перечн<strong>а</strong>я, <strong>с</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong>перед <strong>в</strong>ерхними<br />

челю<strong>с</strong>тями, р<strong>а</strong><strong>в</strong>ными или пре<strong>в</strong><strong>о</strong><strong>с</strong>х<strong>о</strong>дящими г<strong>о</strong>л<strong>о</strong><strong>в</strong>у,<br />

<strong>с</strong> длинными чет<strong>к</strong><strong>о</strong><strong>в</strong>идными <strong>а</strong>нтенн<strong>а</strong>ми и<br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми. Длин<strong>а</strong> тел<strong>а</strong> 4,5—<br />

5,5 мм, <strong>а</strong>нтенн 3,5—5,2 мм (ри<strong>с</strong>. 829). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО CHRYSOMELIDAE LATRELLE, 1802.<br />

ЛИСТОГРЫЗЫ<br />

Мел<strong>к</strong>ие или <strong>с</strong>редней <strong>в</strong>еличины жу<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>льных или д<strong>а</strong>же <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льных, реже<br />

пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тых <strong>о</strong>черт<strong>а</strong>ний," <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лые,<br />

<strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>иль-<br />

http://jurassic.ru/

266<br />

http://jurassic.ru/<br />

н<strong>о</strong> <strong>в</strong>тянут<strong>о</strong>й <strong>в</strong> переднегрудь, не<strong>с</strong>ущей чет<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

ред<strong>к</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>пильч<strong>а</strong>тые или г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тые<br />

<strong>а</strong>нтенны. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> уже н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>,<br />

п<strong>о</strong><strong>с</strong>ледние, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют <strong>в</strong><strong>с</strong>е<br />

брюш<strong>к</strong><strong>о</strong>. Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие, цеп<strong>к</strong>ие, л<strong>а</strong>п<strong>к</strong>и <strong>к</strong><strong>а</strong>жут<strong>с</strong>я<br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми (ри<strong>с</strong>. 830, 831). Юр<strong>а</strong> —<br />

ныне. Гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне:<br />

<strong>с</strong><strong>в</strong>ыше 24 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 1200 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х; из юры из<strong>в</strong>е<strong>с</strong>тны<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Meseumolpites Ponomarenko, nom. nov.<br />

(Eumolpites Martynov, 1926, nec Eumolpites<br />

Heer, 1865, syn. nov). Тип р<strong>о</strong>д<strong>а</strong> — Eumolpites<br />

jurassicus Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>в</strong>ерху не <strong>в</strong>идн<strong>а</strong>.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>переди <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>с</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренными<br />

з<strong>а</strong>дними б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми угл<strong>а</strong>ми и <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong>й, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong>й т<strong>о</strong>чечными углублениями.<br />

Н<strong>а</strong>д<strong>к</strong>рылья шире передне<strong>с</strong>пин<strong>к</strong>и.<br />

Длин<strong>а</strong> тел<strong>а</strong> 10,6 мм, н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong> 7,75 мм<br />

(ри<strong>с</strong>. 832). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Mesosagrites Martynov, 1935. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. multipunctatus Martynov, 1935;<br />

н. юр<strong>а</strong>, Челябин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (н. лей<strong>а</strong><strong>с</strong>, Сух<strong>о</strong>ме<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>).<br />

Имеет<strong>с</strong>я я<strong>с</strong>н<strong>а</strong>я <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ть <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

трети н<strong>а</strong>д<strong>к</strong>рылья, не<strong>с</strong>ущег<strong>о</strong> 14 ряд<strong>о</strong><strong>в</strong> т<strong>о</strong>чечных<br />

углублений; между ряд<strong>а</strong>ми т<strong>о</strong>че<strong>к</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я м<strong>о</strong>рщини<strong>с</strong>т<strong>о</strong><strong>с</strong>ть. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья<br />

11,5 мм, ширин<strong>а</strong> 4 мм (ри<strong>с</strong>. 833). Один<br />

<strong>в</strong>ид. Ср. юр<strong>а</strong> 3. Сибири.<br />

СЕМЕЙСТВО BRUCHIDAE LATREILLE, 1802.<br />

ЗЕРНОВКИ<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины жуч<strong>к</strong>и <strong>с</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им, <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лым тел<strong>о</strong>м, з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>й<br />

<strong>с</strong>переди и <strong>о</strong>пущенн<strong>о</strong>й <strong>в</strong>низ г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, не<strong>с</strong>ущей<br />

ните<strong>в</strong>идные, пильч<strong>а</strong>тые или <strong>с</strong>л<strong>а</strong>б<strong>о</strong>г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тые<br />

<strong>а</strong>нтенны, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>з<strong>а</strong>ди передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й,<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong> ширине н<strong>а</strong>д<strong>к</strong>рыльям, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены и не п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ют <strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong>. Н<strong>о</strong><br />

ги длинные и <strong>к</strong>реп<strong>к</strong>ие <strong>с</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми, <strong>к</strong><strong>а</strong>жущими<strong>с</strong>я<br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ыми (ри<strong>с</strong>. 834). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 900 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 13 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х четыре р<strong>о</strong>д<strong>а</strong>-<br />

СЕМЕЙСТВО ANTHRIBIDAE BILLBERGH, 1820<br />

Очень мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины, умеренн<strong>о</strong><br />

удлиненные жуч<strong>к</strong>и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>-<br />

Ри<strong>с</strong>. 828—834. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Cerambycidae, Chrysomelidae,<br />

Bruchidae<br />

828. Cerambyx cerdo Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 0,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915). 829. Parandrexis parvula Martynov; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 5,5, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 830. Donacia sp.;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х3,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 831. Chrysomela<br />

sp.; <strong>о</strong>бщий <strong>в</strong>ид, Х3,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 832. Meseumolpites<br />

jurassicus (Martynov); <strong>о</strong>бщий <strong>в</strong>ид, х 13,3, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 833. Mesosagrites multipunctatus Martynov;<br />

н<strong>а</strong>д<strong>к</strong>рылье, Х5,2, н. юр<strong>а</strong>. 3. Сибирь (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935).<br />

834. Bruchus rufipes Herbst; <strong>о</strong>бщий <strong>в</strong>ид, Х7,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915)

ленн<strong>о</strong>й <strong>в</strong>низ <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, не<strong>с</strong>ущей рез<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющую <strong>в</strong>ерхнюю' губу, длинные, не<strong>с</strong>тебельч<strong>а</strong>тые,<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тые <strong>а</strong>нтенны, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>з<strong>а</strong>ди<br />

передне<strong>с</strong>пин<strong>к</strong><strong>о</strong>й и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными<br />

н<strong>а</strong>д<strong>к</strong>рыльями (ри<strong>с</strong>. 835). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 2200 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 280 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

•ф<strong>а</strong>уне; 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО CURCULIONIDAE LATREILLE, 1802.<br />

ДОЛГОНОСИКИ<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины или <strong>к</strong>рупные<br />

жу<strong>к</strong>и <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>зующей б<strong>о</strong>лее или<br />

менее <strong>в</strong>ытянутую г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>труб<strong>к</strong>у, <strong>с</strong> <strong>к</strong><strong>о</strong>лнеч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми, имеющими рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленный <strong>с</strong>тебеле<strong>к</strong><br />

(ри<strong>с</strong>. 836, 837). Юр<strong>а</strong> — ныне. Н<strong>а</strong>иб<strong>о</strong>льшее<br />

п<strong>о</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зию <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых: <strong>с</strong><strong>в</strong>ыше 40 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>их<br />

ты<strong>с</strong>яч р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; б<strong>о</strong>лее<br />

130 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х; <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> юре<br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Archaeorrhynchus Martynov, 1926. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. tenuicorne Martynov, 1926; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>труб<strong>к</strong><strong>а</strong><br />

прям<strong>а</strong>я, не менее чем <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, <strong>с</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>ми <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> <strong>а</strong>нтенн.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, н<strong>а</strong>д<strong>к</strong>рылья<br />

длинные, бедр<strong>а</strong> <strong>в</strong>здутые. Длин<strong>а</strong> тел<strong>а</strong> 6,2 мм<br />

^ри<strong>с</strong>. 838). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО IPIDAE HEYDEN,<br />

REITTER, WEISE, 1833. КОРОЕДЫ<br />

Мел<strong>к</strong>ие или <strong>о</strong>чень мел<strong>к</strong>ие жуч<strong>к</strong>и <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>цилиндриче<strong>с</strong><strong>к</strong>им<br />

тел<strong>о</strong>м, притуплённым <strong>с</strong>з<strong>а</strong>ди.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я, <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>тянут<strong>а</strong>я <strong>в</strong> передне<strong>с</strong>пин<strong>к</strong>у,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленн<strong>а</strong>я <strong>в</strong>низ, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими г<strong>о</strong>л<strong>о</strong><strong>в</strong>ч<strong>а</strong>тыми<br />

<strong>а</strong>нтенн<strong>а</strong>ми. Н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, н<strong>о</strong> <strong>к</strong>реп<strong>к</strong>ие,<br />

н<strong>а</strong>д<strong>к</strong>рылья длинные, <strong>с</strong>з<strong>а</strong>ди ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зующие<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>в</strong>ыем<strong>к</strong>и и зубцы, <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ющие<br />

б<strong>о</strong>льш<strong>о</strong>е углубление н<strong>а</strong> <strong>к</strong><strong>о</strong>нце тел<strong>а</strong> («т<strong>а</strong>ч<strong>к</strong><strong>а</strong>»<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>ед<strong>о</strong><strong>в</strong>) (ри<strong>с</strong>. 839). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

2000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 130 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х.<br />

В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящую <strong>с</strong><strong>в</strong><strong>о</strong>д<strong>к</strong>у не были <strong>в</strong><strong>к</strong>лючены<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие р<strong>о</strong>ды же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых,<br />

прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ть <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong> <strong>о</strong>пределенным<br />

•<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>м нельзя д<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> т<strong>о</strong>чн<strong>о</strong> <strong>о</strong>пределить<br />

из-з<strong>а</strong> пл<strong>о</strong>хих <strong>о</strong>пи<strong>с</strong><strong>а</strong>ний и из<strong>о</strong>бр<strong>а</strong>жений.<br />

836<br />

Ри<strong>с</strong>. 835—839.Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Anthribidae.Circulionidae, Ipidae.<br />

835. Anthribus albinus Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915). 836. Ceutorhynchus sp.; <strong>о</strong>бщий <strong>в</strong>ид, х9,3, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915). 837. Apion sp.; <strong>о</strong>бщий <strong>в</strong>ид, Х9,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н,<br />

1915). 838. Archaeorrhynchus tenuicornis Martynov; <strong>о</strong>бщий<br />

<strong>в</strong>ид, Х8,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1926). 839. Hylesinus<br />

fraxini Panzer; <strong>о</strong>бщий^<strong>в</strong>ид, Xl2,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Я<strong>к</strong><strong>о</strong>б<strong>с</strong><strong>о</strong>н, 1915)<br />

С<strong>в</strong>едения п<strong>о</strong> этим ф<strong>о</strong>рм<strong>а</strong>м при<strong>в</strong>едены Г<strong>а</strong>ндлиршем<br />

(1906, 1938). В<strong>с</strong>ег<strong>о</strong> из три<strong>а</strong><strong>с</strong><strong>а</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<strong>о</strong><br />

36 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из юры 200 р<strong>о</strong>д<strong>о</strong><strong>в</strong> и из мел<strong>а</strong> 18 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, мн<strong>о</strong>гим <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых<br />

<strong>в</strong><strong>о</strong><strong>о</strong>бще не был<strong>о</strong> д<strong>а</strong>н<strong>о</strong> н<strong>а</strong>учн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>з<strong>в</strong><strong>а</strong>ния.<br />

267<br />

http://jurassic.ru/<br />

837

ОТРЯД STREPSIPTERA. ВЕЕРОКРЫЛЫЕ<br />

Неб<strong>о</strong>льшие (1,5—4 мм) н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong> <strong>о</strong>чень<br />

рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным п<strong>о</strong>л<strong>о</strong><strong>в</strong>ым дим<strong>о</strong>рфизм<strong>о</strong>м.<br />

С<strong>а</strong>мцы <strong>к</strong>рыл<strong>а</strong>тые; передние <strong>к</strong>рылья рудимен-<br />

Ри<strong>с</strong>. 840. Отряд Strepsiptera. Mengea tertiaria (Menge);<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 9,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Menge, 1866)<br />

т<strong>а</strong>рны, пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, уз<strong>к</strong>ие, из<strong>о</strong>гнутые<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и; з<strong>а</strong>дние <strong>к</strong>рылья х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты,<br />

шир<strong>о</strong><strong>к</strong>ие, <strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>;<br />

жил<strong>к</strong>и пр<strong>о</strong>д<strong>о</strong>льные, немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные.<br />

Крупные ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong>; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет; р<strong>о</strong>т<strong>о</strong>­<br />

(О. И. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

<strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н; <strong>а</strong>нтенны четырех<strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

третий члени<strong>к</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м. Передне- и <strong>с</strong>реднегрудь <strong>о</strong>чень<br />

м<strong>а</strong>лы; з<strong>а</strong>днегрудь <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, ее тергит<br />

при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние брюш<strong>к</strong><strong>а</strong>. Л<strong>а</strong>п<strong>к</strong><strong>а</strong> из<br />

д<strong>в</strong>ух — пяти члени<strong>к</strong><strong>о</strong><strong>в</strong>, брюш<strong>к</strong><strong>о</strong> из де<strong>с</strong>яти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>-<br />

С<strong>а</strong>м<strong>к</strong><strong>а</strong> п<strong>о</strong>х<strong>о</strong>ж<strong>а</strong> н<strong>а</strong> личин<strong>к</strong>у, без <strong>к</strong>рылье<strong>в</strong>,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> д<strong>а</strong>же без н<strong>о</strong>г и других <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей.<br />

У гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>г<strong>о</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и неп<strong>о</strong>д<strong>в</strong>ижны<br />

и н<strong>а</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong>нутри тел<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong>х<strong>о</strong>зяин<strong>а</strong>,<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м жил<strong>а</strong> личин<strong>к</strong><strong>а</strong> — энд<strong>о</strong>п<strong>а</strong>р<strong>а</strong>зит<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых р<strong>а</strong>зных <strong>о</strong>тряд<strong>о</strong><strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Семь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых лишь <strong>о</strong>дн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

— Mengeidae.<br />

СЕМЕЙСТВО MENGEIDAE PIERCE, 1908<br />

У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>а</strong>нтенны <strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; л<strong>а</strong>п<strong>к</strong>и<br />

пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 840).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один <strong>в</strong>ид <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

С О L Е О Р Т Е R О I D Е А I N С Е R Т А Е S Е D I S<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

СЕМЕЙСТВО PROTOCOLEIDAE TILLYARD, 1924<br />

Передние <strong>к</strong>рылья <strong>в</strong> <strong>в</strong>иде шир<strong>о</strong><strong>к</strong>их, н<strong>о</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

пл<strong>о</strong><strong>с</strong><strong>к</strong>их н<strong>а</strong>д<strong>к</strong>рылье<strong>в</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние их<br />

или д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>е, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящее из мн<strong>о</strong>гих,<br />

<strong>в</strong> цел<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельных жил<strong>о</strong><strong>к</strong>, или же нея<strong>с</strong>н<strong>о</strong>е<br />

и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из един<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й (? <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й)<br />

жил<strong>к</strong>и. Стр<strong>о</strong>ение з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> и <strong>в</strong><strong>с</strong>ег<strong>о</strong><br />

тел<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>. Длин<strong>а</strong> н<strong>а</strong>д<strong>к</strong>рылья 8—15 мм.<br />

В. пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Protocoleus Tillyard, 1924. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. mitchelli Tillyard, 1924; <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

(Бельм<strong>о</strong>нт). Передний (?) <strong>к</strong>р<strong>а</strong>й н<strong>а</strong>д<strong>к</strong>рылья<br />

<strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый и т<strong>о</strong>н<strong>к</strong>ий. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящих<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льных<br />

жил<strong>о</strong><strong>к</strong>, п<strong>о</strong>чти без п<strong>о</strong>перечных; р<strong>а</strong>зличимы<br />

п<strong>о</strong>чти <strong>в</strong><strong>с</strong>е <strong>с</strong>и<strong>с</strong>темы жил<strong>о</strong><strong>к</strong> (SC, R, М, Си<br />

и А); з<strong>а</strong>дний (?) <strong>к</strong>р<strong>а</strong>й п<strong>о</strong>чти прям<strong>о</strong>й, <strong>с</strong>ильн<strong>о</strong><br />

ут<strong>о</strong>лщенный. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 15 мм<br />

(ри<strong>с</strong>- 841). Один <strong>в</strong>ид. В. пермь А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Arctocoleus Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. ivensis Martynov, 1933; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><br />

<strong>к</strong> <strong>а</strong> я<br />

„ ~ , * , , , °бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>-<br />

Ри<strong>с</strong>. 841-842. Coleoptero.dea incertae sedis ш е н е я <strong>с</strong> н <strong>о</strong> е ; з <strong>а</strong> д н и й ( ? ) <strong>к</strong> р <strong>а</strong> й п р я м <strong>о</strong> й и у т <strong>о</strong> л.<br />

841. Protocoleus mitchelli Tillyard; н<strong>а</strong>д<strong>к</strong>рылье, х5,0, <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>че- щ е ННЫЙ. Длин<strong>а</strong> НЭДКрЫЛЬЯ 9—10 ММ, ШИрИНЭння<br />

<strong>о</strong>бычные, <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1924). 842. Arctocoleus<br />

ivensis Martynov; н<strong>а</strong>д<strong>к</strong>рылье, Х2,2, <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

0 К 0 Л 0<br />

.<br />

4 М М<br />

, <strong>о</strong>лп\ г\ г> _<br />

(Р<br />

л„<br />

ИС 8 4 2<br />

' ) • °Д ВИ<br />

Д- В<br />

- П е М Ь<br />

Р<br />

А<br />

Р"<br />

<strong>о</strong>бл. (м<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933) х<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>и <strong>о</strong>бл.<br />

268<br />

http://jurassic.ru/

НАДОТРЯД NEUROPTEROIDEA.<br />

СЕТЧАТ0КРЫЛ00БРАЗНЫЕ<br />

Крылья удлиненные, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные,<br />

<strong>о</strong>чень ред<strong>к</strong><strong>о</strong> з<strong>а</strong>дние <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>чень б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я<br />

жил<strong>к</strong><strong>а</strong>ми п<strong>о</strong>чти <strong>в</strong><strong>с</strong>ех <strong>с</strong>и<strong>с</strong>тем. Ред<strong>к</strong><strong>о</strong><br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, причем н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я рез<strong>к</strong><strong>а</strong>я<br />

гетер<strong>о</strong>н<strong>о</strong>мия (диптеригия Coniopterygidae).<br />

Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны грызущег<strong>о</strong> тип<strong>а</strong>: ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>в</strong>язи<br />

<strong>с</strong> <strong>а</strong>ф<strong>а</strong>гией <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны <strong>с</strong>л<strong>а</strong>бы и нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. Личин<strong>к</strong>и<br />

<strong>к</strong><strong>а</strong>мп<strong>о</strong>дее<strong>в</strong>идные, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими и <strong>к</strong>рупными<br />

челю<strong>с</strong>тями, хищни<strong>к</strong>и, н<strong>а</strong>п<strong>а</strong>д<strong>а</strong>ющие н<strong>а</strong> других<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых или <strong>к</strong>лещей, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>бит<strong>а</strong>ю­<br />

щие н<strong>а</strong> <strong>с</strong>уше, реже <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рылье<strong>в</strong> дре<strong>в</strong>нейших <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

чрез<strong>в</strong>ыч<strong>а</strong>йн<strong>о</strong> н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных. П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>о</strong>ни пр<strong>о</strong>из<strong>о</strong>шли<br />

<strong>о</strong>т <strong>к</strong><strong>а</strong><strong>к</strong><strong>о</strong>й-т<strong>о</strong> группы т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных;<br />

их и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие <strong>о</strong>пределил<strong>о</strong><strong>с</strong>ь р<strong>а</strong><strong>с</strong>ширением<br />

и <strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием хищниче<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

личин<strong>о</strong><strong>к</strong> и р<strong>а</strong>з<strong>в</strong>итием у них <strong>в</strong>не<strong>к</strong>ишечн<strong>о</strong>г<strong>о</strong><br />

пище<strong>в</strong><strong>а</strong>рения. С<strong>о</strong><strong>в</strong>ременн<strong>а</strong>я ф<strong>а</strong>ун<strong>а</strong> з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ет<br />

б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> рели<strong>к</strong>т<strong>о</strong><strong>в</strong>ых ф<strong>о</strong>рм. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н —<br />

ныне. Отряды: Megaloptera, Raphidioptera,<br />

Neuroptera.<br />

ОТРЯД MEGALOPTERA. ВИСЛОКРЫЛЫЕ<br />

Средней <strong>в</strong>еличины и <strong>к</strong>рупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>с</strong><br />

д<strong>в</strong>умя п<strong>а</strong>р<strong>а</strong>ми шир<strong>о</strong><strong>к</strong>их г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong>,<br />

п<strong>о</strong><strong>к</strong>рытых мел<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, <strong>с</strong> мех<strong>а</strong>ниче<strong>с</strong><strong>к</strong>и<br />

не<strong>с</strong><strong>о</strong><strong>в</strong>ершенным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием. Переднее <strong>к</strong>рыл<strong>о</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; передний <strong>к</strong>р<strong>а</strong>й<br />

<strong>в</strong>ыпу<strong>к</strong>лый; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>те или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> R; <strong>в</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>луч<strong>а</strong>е мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

<strong>в</strong>ет<strong>в</strong>и SC перех<strong>о</strong>дят н<strong>а</strong> R; <strong>в</strong>ет<strong>в</strong>ей RS три —<br />

де<strong>в</strong>ять; д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и М или <strong>с</strong> д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чными<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; СиА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

три <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и; ч<strong>а</strong>ще н<strong>а</strong> А 2 д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чный<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; птер<strong>о</strong><strong>с</strong>тигмы нет. Пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и не <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong><br />

п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>. В п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>к</strong>рылья <strong>с</strong>л<strong>о</strong>жены <strong>к</strong>рыше<strong>о</strong>бр<strong>а</strong>зи<strong>о</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong> ните<strong>в</strong>идными<br />

мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ыми у<strong>с</strong>и<strong>к</strong><strong>а</strong>ми и ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чными,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленными гл<strong>а</strong>з<strong>а</strong>ми.<br />

Р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызущег<strong>о</strong> тип<strong>а</strong>. Грудные<br />

<strong>с</strong>егменты п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные; члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

пять; чет<strong>в</strong>ертый члени<strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, д<strong>в</strong>ул<strong>о</strong>п<strong>а</strong><strong>с</strong>тн<strong>о</strong>й.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> де<strong>с</strong>ять, н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

цер<strong>к</strong>и; у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> яйце<strong>к</strong>л<strong>а</strong>д <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Взр<strong>о</strong><strong>с</strong>лые не пит<strong>а</strong>ют<strong>с</strong>я, <strong>в</strong>едут н<strong>о</strong>чн<strong>о</strong>й <strong>о</strong>бр<strong>а</strong>з<br />

жизни <strong>в</strong>близи <strong>с</strong>т<strong>о</strong>ячих или медленн<strong>о</strong> те<strong>к</strong>ущих<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong> <strong>с</strong> или<strong>с</strong>тым дн<strong>о</strong>м; жи<strong>в</strong>ут не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

дней. Яйц<strong>а</strong> <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют <strong>в</strong> м<strong>а</strong><strong>с</strong><strong>с</strong>е н<strong>а</strong> н<strong>а</strong>д<strong>в</strong><strong>о</strong>дные<br />

ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>тений.<br />

Личин<strong>к</strong><strong>а</strong> <strong>в</strong><strong>о</strong>дн<strong>а</strong>я, хищн<strong>а</strong>я, жи<strong>в</strong>ет д<strong>в</strong><strong>а</strong> г<strong>о</strong>д<strong>а</strong>,<br />

<strong>в</strong>н<strong>а</strong>ч<strong>а</strong>ле <strong>в</strong> <strong>в</strong><strong>о</strong>де, з<strong>а</strong>тем з<strong>а</strong>ры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> ил. Н<strong>а</strong><br />

брюш<strong>к</strong>е <strong>с</strong>емь — де<strong>в</strong>ять п<strong>а</strong>р тр<strong>а</strong>хейных ж<strong>а</strong>бер;<br />

<strong>в</strong>н<strong>а</strong>ч<strong>а</strong>ле тр<strong>а</strong>хейные ж<strong>а</strong>бры нечлени<strong>с</strong>тые, з<strong>а</strong>тем<br />

п<strong>о</strong>я<strong>в</strong>ляет<strong>с</strong>я члени<strong>с</strong>т<strong>о</strong><strong>с</strong>ть. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ред<strong>к</strong>и, н<strong>о</strong><strong>с</strong>ят рели<strong>к</strong>т<strong>о</strong><strong>в</strong>ый х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер.<br />

Пермь —ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Permosialidae, Sialidae,<br />

Corydalidae; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>е <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

не н<strong>а</strong>йден<strong>о</strong>.<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

СЕМЕЙСТВО PERMOSIALIDAE<br />

MARTYNOV, 1928<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>те, н<strong>а</strong> <strong>с</strong>ередине <strong>в</strong>ершинн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong> или немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее,<br />

<strong>в</strong>ет<strong>в</strong>ей SC ше<strong>с</strong>ть — де<strong>с</strong>ять; н<strong>а</strong> RS<br />

д<strong>в</strong>е — четыре <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong>ет<strong>в</strong>и М пр<strong>о</strong><strong>с</strong>тые; СиА и<br />

М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>м пр<strong>о</strong>тяжении<br />

269<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 843—844. Отряд Megaloptera<br />

843. Permosialis bifasciata Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, Х6,0,<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>.). 844. Permosialis<br />

sp.; личин<strong>к</strong><strong>а</strong>, X 2,5, пермь, Ю. Приур<strong>а</strong>лье (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1953)<br />

<strong>с</strong>литы или <strong>с</strong><strong>о</strong>единены <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й; СиА <strong>с</strong> пр<strong>о</strong><strong>с</strong>тым р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>й

из <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>, ч<strong>а</strong>ще н<strong>а</strong> А 2, р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и ред<strong>к</strong>и, п<strong>о</strong>чти <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

ин<strong>о</strong>гд<strong>а</strong> н<strong>а</strong> <strong>к</strong>рыле ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечных<br />

п<strong>о</strong>л<strong>о</strong><strong>с</strong>. Пермь. Один р<strong>о</strong>д.<br />

Permosialis Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. paucinervis Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры). SC<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> С; <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong> R нет; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>нце <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; длин<strong>а</strong><br />

<strong>к</strong>рылье<strong>в</strong> 10—18 мм. Личин<strong>к</strong><strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>а</strong>я,<br />

<strong>с</strong> <strong>к</strong>рупными ж<strong>в</strong><strong>а</strong>л<strong>а</strong>ми; л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми; де<strong>в</strong>ять п<strong>а</strong>р<br />

члени<strong>с</strong>тых тр<strong>а</strong>хейных ж<strong>а</strong>бер; длин<strong>а</strong> личин<strong>к</strong>и<br />

25 мм (ри<strong>с</strong>. 843, 844). 16 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл., Приур<strong>а</strong>лья, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО CORYDALIDAE LEACH, 1815<br />

[nom. transl. Burmeister, 1838 (ex Corydalida Leach, 1815)1<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC длинн<strong>а</strong>я, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

R; <strong>в</strong>ет<strong>в</strong>ей SC 16—17; д<strong>в</strong>е — три <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong> R;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; М не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong> CuA; <strong>в</strong>ет<strong>в</strong>и М пр<strong>о</strong><strong>с</strong>тые и <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; <strong>в</strong><strong>с</strong>е<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; мн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; н<strong>а</strong> <strong>к</strong>рыльях мн<strong>о</strong>г<strong>о</strong> ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>.<br />

Крупные-—длин<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> 30—62 мм.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны, р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м<br />

<strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х и <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х <strong>о</strong>б<strong>о</strong>их п<strong>о</strong>луш<strong>а</strong>рий,<br />

из них <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

Вне СССР; <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из К<strong>о</strong>л<strong>о</strong>р<strong>а</strong>д<strong>о</strong>.<br />

ОТРЯД RAPHIDIOPTERA. ВЕРБЛЮДКИ<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Д<strong>в</strong>е н<strong>а</strong>ры уз<strong>к</strong>их, п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong>.<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong> (<strong>к</strong>р<strong>о</strong>ме <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Sojanoraphidiidae);<br />

<strong>в</strong>ет<strong>в</strong>ей SC <strong>о</strong>т ше<strong>с</strong>ти д<strong>о</strong> трин<strong>а</strong>дц<strong>а</strong>ти; RS <strong>с</strong> м<strong>а</strong>лым<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей; RS и МА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении <strong>с</strong>литы; <strong>в</strong> з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА длинн<strong>о</strong>е или<br />

п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong> п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>е; Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; CuA мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я;<br />

CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е — три;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют ряд х<strong>а</strong>р<strong>а</strong><strong>к</strong>терных<br />

ячей; <strong>в</strong><strong>с</strong>егд<strong>а</strong> е<strong>с</strong>ть <strong>о</strong><strong>к</strong>р<strong>а</strong>шенн<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>.<br />

Грудные <strong>с</strong>егменты р<strong>а</strong>з<strong>в</strong>иты нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>, переднегрудь<br />

у перм<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителя <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>с</strong>редне- и з<strong>а</strong>днегруди, у мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их и <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

<strong>о</strong>н<strong>а</strong> длиннее <strong>с</strong>редне- и з<strong>а</strong>днегруди. Г<strong>о</strong>­<br />

МА<br />

л<strong>о</strong><strong>в</strong><strong>а</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я, пр<strong>о</strong>гн<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>а</strong>я, ч<strong>а</strong><strong>с</strong>ть, леж<strong>а</strong>щ<strong>а</strong>я<br />

п<strong>о</strong>з<strong>а</strong>ди гл<strong>а</strong>з <strong>в</strong>ытянут<strong>а</strong>; р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<br />

грызущег<strong>о</strong> тип<strong>а</strong>; гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие, ин<strong>о</strong>гд<strong>а</strong> при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

гл<strong>а</strong>з<strong>к</strong>и. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> длинный яйце<strong>к</strong>л<strong>а</strong>д.<br />

Взр<strong>о</strong><strong>с</strong>лые и личин<strong>к</strong>и хищные. Личин<strong>к</strong><strong>а</strong> <strong>к</strong><strong>а</strong>мп<strong>о</strong>дее<strong>в</strong>идн<strong>а</strong>я,<br />

жи<strong>в</strong>ет <strong>в</strong> трещин<strong>а</strong>х <strong>к</strong><strong>о</strong>ры х<strong>в</strong><strong>о</strong>йных,<br />

<strong>в</strong> ле<strong>с</strong>н<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е, мхе. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Fatjanopteridae, Letopalopteridae,<br />

Sojanoraphidiidae, Mesoraphidiidae, Raphidiidae,<br />

Inocelliidae.<br />

СЕМЕЙСТВО FATJANOPTERIDAE О. MARTY NOVA, 1961<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый; SC из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong>перед, пр<strong>о</strong>х<strong>о</strong>дит<br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уже д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong><br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS д<strong>о</strong><br />

12 ячей; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления MP, <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>е и длин-<br />

270<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 845—847. Отряд Raphidioptera; <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Fatjanopteridae,<br />

Letopalopteridae, Sojanoraphidiidae<br />

845. Fatjanoptera mnemonica O. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X3,0;<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Тунгу<strong>с</strong><strong>с</strong><strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 846. Letopaloptera<br />

albardiana О. Martynova; переднее и з<strong>а</strong>днее <strong>к</strong>рылья,<br />

X 7,5, <strong>в</strong>. Пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 847.<br />

Sojanoraphidia rossica О. Martynova; <strong>о</strong>бщий <strong>в</strong>ид, Х7,5, <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1952).<br />

SC

н<strong>о</strong>е; CuA не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М, птер<strong>о</strong><strong>с</strong>тигмы нет.<br />

Крыл<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>е, <strong>о</strong>тн<strong>о</strong>шение длины <strong>к</strong>рыл<strong>а</strong> <strong>к</strong> ег<strong>о</strong><br />

ширине р<strong>а</strong><strong>в</strong>н<strong>о</strong> 2,8. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Один р<strong>о</strong>д.<br />

Fatjanoptera О. Martynova, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — F. mnemonica О. Martynova, 1961; <strong>в</strong>.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Тунгу<strong>с</strong><strong>с</strong><strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (бургу<strong>к</strong>лин<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Ф<strong>а</strong>тьяних<strong>а</strong>). Н<strong>а</strong> SC 15—16 <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong><br />

R д<strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьми <strong>в</strong>ет<strong>в</strong>ей; ширин<strong>а</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля н<strong>а</strong>иб<strong>о</strong>льш<strong>а</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; три<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и, <strong>в</strong><strong>с</strong>е <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 23—25 мм (ри<strong>с</strong>. 845). Один <strong>в</strong>ид.<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Тунгу<strong>с</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО LETOPALOPTERIDAE<br />

О. MARTYNOVA, 1961<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, прям<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле немн<strong>о</strong>г<strong>о</strong> шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

уже р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> (ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS н<strong>а</strong> <strong>в</strong><strong>с</strong>ем пр<strong>о</strong>тяжении <strong>о</strong>дн<strong>о</strong>й ширины); RS<br />

н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я незн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>е и длинн<strong>о</strong>е, <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т М перед <strong>с</strong><strong>а</strong>мым<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>лением MP, пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS; СиА п<strong>о</strong>чти не <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М, длинн<strong>а</strong>я;<br />

птер<strong>о</strong><strong>с</strong>тигмы нет; <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е и длинн<strong>о</strong>е, <strong>о</strong>тн<strong>о</strong>шение<br />

длины <strong>к</strong>рыл<strong>а</strong> <strong>к</strong> ег<strong>о</strong> ширине р<strong>а</strong><strong>в</strong>н<strong>о</strong> четырем.<br />

Переднегрудь длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Очень<br />

длинные н<strong>о</strong>ги. В. пермь. Один р<strong>о</strong>д.<br />

Letopaloptera О. Martynova, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong>—L. albardiana О. Martynova, 1961; <strong>в</strong>,<br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). Вет<strong>в</strong>ей SC пять — ше<strong>с</strong>ть; н<strong>а</strong> R <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьми <strong>в</strong>ет<strong>в</strong>ей; RSi +2, МА, MP] и МР 2 <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем<br />

прямые, их <strong>в</strong>ет<strong>в</strong>и зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны, п<strong>о</strong>х<strong>о</strong>жи н<strong>а</strong><br />

интер<strong>к</strong><strong>а</strong>лярные жил<strong>к</strong>и; Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>,<br />

СиА пр<strong>о</strong>х<strong>о</strong>дит п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

10 мм (ри<strong>с</strong>. 846). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти.<br />

СЕМЕЙСТВО SOJANORAPH1DIIDAE<br />

О. MARTYNOVA, 1952<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й;<br />

SC прям<strong>а</strong>я, длинн<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля <strong>о</strong>дн<strong>о</strong>й ширины; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле д<strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьми<br />

ячей; МА <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т М ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; СиА не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М; птер<strong>о</strong><strong>с</strong>тигмы<br />

нет; <strong>к</strong>рыл<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и уз<strong>к</strong><strong>о</strong>е, <strong>о</strong>тн<strong>о</strong>шение длины<br />

<strong>к</strong> ег<strong>о</strong> ширине р<strong>а</strong><strong>в</strong>н<strong>о</strong> трем. Переднегрудь<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. В. пермь. Один р<strong>о</strong>д.<br />

Sojanoraphidia О. Martynova, 1952. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. rossica О. Martynova, 1952; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

'CuA I CuP А 850А<br />

РИС. 848—851. Отряд Raphidioptera; <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Mesoraphidiidae, Raphidiidae, Inocelliidae<br />

848. Mesoraphidia pterostigmalls O. Martynova; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X3,5, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1947). 849. Proraphidla<br />

turkestanica О. Martynova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х6,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1947). 850. Raphidia ophiopsis Linnaeus;<br />

А\СиР<br />

8 5 0 6<br />

А — переднее <strong>к</strong>рыл<strong>о</strong>, Х6,0,; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х6,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (<strong>о</strong>риг. ри<strong>с</strong>.). 851, Fibla exusta (Cockerell et Custer); з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, X5,0, не<strong>о</strong>ген, К<strong>о</strong>л<strong>о</strong>р<strong>а</strong>д<strong>о</strong> (Carpenter, 1936)<br />

271<br />

http://jurassic.ru/

Вет<strong>в</strong>ей SC ше<strong>с</strong>ть; н<strong>а</strong> R д<strong>в</strong>е —три <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong><strong>с</strong>е<br />

<strong>в</strong>ет<strong>в</strong>и незигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны; Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

п<strong>о</strong>чти <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>; д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>а</strong>н<strong>а</strong>льные<br />

жил<strong>к</strong>и. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 5 мм (ри<strong>с</strong>. 847).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти.<br />

СЕМЕЙСТВО MESORAPHIDIIDAE<br />

MARTYNOV, 1925<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му;<br />

SC прям<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

длины <strong>к</strong>рыл<strong>а</strong> и ди<strong>с</strong>т<strong>а</strong>льнее; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

зн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле три ячеи; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т М немн<strong>о</strong>г<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления MP или <strong>в</strong> ме<strong>с</strong>те<br />

ее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления; СиА <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й т<strong>о</strong>ч<strong>к</strong>е <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> М; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> е<strong>с</strong>ть, длинн<strong>а</strong>я, не пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. В. з<strong>а</strong>днем<br />

<strong>к</strong>рыле <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА длинн<strong>о</strong>е, <strong>к</strong><strong>а</strong><strong>к</strong><br />

<strong>в</strong> <strong>к</strong>рыле <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> Raphidia. Переднегрудь<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, н<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong><strong>о</strong> <strong>с</strong>реднегрудью.<br />

Юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Mesoraphidia Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong>—<br />

M. grandis Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я: длин<strong>а</strong> ее р<strong>а</strong><strong>в</strong>н<strong>а</strong> трети длины <strong>к</strong>рыл<strong>а</strong>;<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я гр<strong>а</strong>ниц<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы и п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> r-rs р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 5,5—16 мм<br />

(ри<strong>с</strong>. 848). Семь <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> и<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Proraphidia О. Martynova, 1947. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. turkestanica О. Martynova, 1947; <strong>в</strong>.<br />

юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). М<strong>а</strong>лень<strong>к</strong><strong>а</strong>я<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>, б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я гр<strong>а</strong>ниц<strong>а</strong> ее<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льнее п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и<br />

r-rs и <strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 11—12 мм (ри<strong>с</strong>. 849). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong><br />

1<br />

Из <strong>в</strong>. юры З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лья из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Baisopteridae<br />

О. Mart., 1961 <strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м и д<strong>в</strong>умя <strong>в</strong>ид<strong>а</strong>ми;<br />

<strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т Mesoraphidiidae б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м ячей<br />

<strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле (<strong>о</strong>пи<strong>с</strong><strong>а</strong>ние <strong>в</strong>ышл<strong>о</strong> п<strong>о</strong><strong>с</strong>ле <strong>с</strong>д<strong>а</strong>чи ру<strong>к</strong><strong>о</strong>пи<strong>с</strong>и<br />

<strong>в</strong> н<strong>а</strong>б<strong>о</strong>р).<br />

СЕМЕЙСТВО RAPHIDIIDAE LATREILLE, 1810<br />

[nom. transl. Stephens, 1836 (ex Raphidiinae Latreille, 1810)]<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый; SC прям<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; SC н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле три ячеи; МА <strong>о</strong>тх<strong>о</strong>дит не <strong>о</strong>т М, <strong>а</strong> <strong>о</strong>т передней<br />

<strong>в</strong>ет<strong>в</strong>и MP, т. е. ее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>ме<strong>с</strong>тил<strong>о</strong><strong>с</strong>ь<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>; СиА н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> М; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> е<strong>с</strong>ть, <strong>о</strong>н<strong>а</strong> <strong>о</strong>гр<strong>а</strong>ничен<strong>а</strong><br />

и пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>тн<strong>о</strong>шение длины <strong>к</strong>рыл<strong>а</strong> <strong>к</strong> ег<strong>о</strong> ширине р<strong>а</strong><strong>в</strong>н<strong>о</strong><br />

2,8—3,2, н<strong>о</strong> ч<strong>а</strong>ще 2,8; д<strong>в</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие <strong>а</strong>н<strong>а</strong>льные<br />

жил<strong>к</strong>и. Переднегрудь <strong>о</strong>дн<strong>о</strong>й длины или немн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы и <strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее <strong>с</strong>реднегруди.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, <strong>с</strong> тремя гл<strong>а</strong>з<strong>к</strong><strong>а</strong>ми. Не<strong>о</strong>ген —<br />

ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

<strong>о</strong>дин <strong>в</strong> третичн<strong>о</strong>й ф<strong>а</strong>уне.<br />

Raphidia Linnaeus, 1735. Тип р<strong>о</strong>д<strong>а</strong> — R.<br />

ophiopsis Linnaeus, 1735 (Subilla Navas, 1916;<br />

Alena Navas, 1915; Lesna Navas, 1915; Erma<br />

Navas, 1918); <strong>с</strong><strong>о</strong><strong>в</strong>ременные, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. В з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА длинн<strong>о</strong>е;<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я гр<strong>а</strong>ниц<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы лежит н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>льн<strong>о</strong>й ячеи. Длин<strong>а</strong><br />

<strong>к</strong>рылье<strong>в</strong> 7,5—12 мм (ри<strong>с</strong>. 850). О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин <strong>в</strong><br />

С. Амери<strong>к</strong>е; <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong> из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

Ми<strong>о</strong>цен — ныне.<br />

СЕМЕЙСТВО INOCELLIIDAE NAVAS, 1916<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т предыдущег<strong>о</strong> птер<strong>о</strong><strong>с</strong>тигм<strong>о</strong>й,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я не <strong>о</strong>гр<strong>а</strong>ничен<strong>а</strong> и не пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми. Переднегрудь длиннее <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ых<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> груди и <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й длины<br />

или немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не <strong>с</strong>ужен<strong>а</strong><br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них <strong>о</strong>дин <strong>в</strong> третичн<strong>о</strong>й<br />

ф<strong>а</strong>уне.<br />

"Fibla Navas, 1915 (Burcha Navas, 1915). Тип<br />

р<strong>о</strong>д<strong>а</strong> — F. hesperica Navas, 1915 {Inocellia maclachlani<br />

Navas, nec Albarda, 1908); <strong>с</strong><strong>о</strong><strong>в</strong>ременные,<br />

П<strong>о</strong>ртуг<strong>а</strong>лия. В з<strong>а</strong>днем <strong>к</strong>рыле <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА длинн<strong>о</strong>е. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 9—<br />

14 мм (ри<strong>с</strong>. 851). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один <strong>в</strong>ид<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и С. Амери<strong>к</strong>и.<br />

ОТРЯД NEUROPTERA. СЕТЧАТОКРЫЛЫЕ<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

Д<strong>в</strong>е п<strong>а</strong>ры п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных (<strong>к</strong>р<strong>о</strong>ме Nemopteridae)<br />

<strong>к</strong>рылье<strong>в</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние б<strong>о</strong>г<strong>а</strong>т<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong><br />

(редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е Coniopterygidae);<br />

н<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>ч­<br />

ные ди<strong>с</strong>т<strong>а</strong>льные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и; б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>ей SC ч<strong>а</strong><strong>с</strong>т<strong>о</strong> перех<strong>о</strong>дящих н<strong>а</strong> R; <strong>в</strong>ет<strong>в</strong>и<br />

RS р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены гребенч<strong>а</strong>т<strong>о</strong>; передняя <strong>в</strong>ет<strong>в</strong>ь<br />

М у мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их и <strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм <strong>о</strong>бычн<strong>о</strong><br />

272<br />

http://jurassic.ru/

SC.<br />

852. Solenoptilon martynovi O. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X3.2,<br />

<strong>в</strong>. юр<strong>а</strong>. Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1949). 853. Sog/uta speclosa<br />

О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 7,7, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия<br />

<strong>с</strong>лит<strong>а</strong> н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении (<strong>к</strong>р<strong>о</strong>ме Chrysopidae)<br />

<strong>с</strong> R или RS. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, <strong>с</strong><br />

грызущим р<strong>о</strong>т<strong>о</strong><strong>в</strong>ым <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong>м; гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>л<strong>о</strong>жные;<br />

<strong>о</strong>бычн<strong>о</strong> н<strong>а</strong> лбу три гл<strong>а</strong>з<strong>к</strong><strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>ни <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

<strong>а</strong>нтенны ните<strong>в</strong>идные, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> ут<strong>о</strong>лщением<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце, умеренн<strong>о</strong>й длины. Грудь примити<strong>в</strong>н<strong>а</strong>я;<br />

н<strong>о</strong>ги бег<strong>а</strong>тельные, <strong>с</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

л<strong>а</strong>п<strong>к</strong><strong>о</strong>й. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong> нет<br />

(<strong>к</strong>р<strong>о</strong>ме Dilar). Личин<strong>к</strong>и <strong>к</strong><strong>а</strong>мп<strong>о</strong>дее<strong>в</strong>идные, хищные,<br />

<strong>с</strong> <strong>в</strong>ытянутыми челю<strong>с</strong>тями. Сетч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые<br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>в</strong>п<strong>о</strong>лне <strong>с</strong>ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong><strong>в</strong>ший<strong>с</strong>я <strong>о</strong>тряд из<strong>в</strong>е<strong>с</strong>тны<br />

<strong>с</strong> нижнеперм<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени, <strong>о</strong>че<strong>в</strong>идн<strong>о</strong>,<br />

<strong>в</strong><strong>о</strong>зни<strong>к</strong>ли <strong>в</strong> б<strong>о</strong>лее дре<strong>в</strong>нее ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>в</strong>ремя.<br />

Принципи<strong>а</strong>льн<strong>о</strong>е <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рылье<strong>в</strong> <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых и н<strong>а</strong>иб<strong>о</strong>лее дре<strong>в</strong>них и<br />

<strong>а</strong>рх<strong>а</strong>ичных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц д<strong>а</strong>ет <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>чит<strong>а</strong>ть<br />

эт<strong>о</strong>т <strong>о</strong>тряд р<strong>о</strong>д<strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льным для <strong>в</strong><strong>с</strong>ех <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

ме<strong>к</strong><strong>о</strong>птер<strong>о</strong>идн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>. Пермь —<br />

ныне. Четыре н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Myrmeleontidea,<br />

Polystoechotidea, Hemerobiidea, Coniopterygidea.<br />

18 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

MA<br />

Ри<strong>с</strong>. 852—854. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Myrmeleontidea<br />

273<br />

SC+R<br />

SC+R<br />

853<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 854. Chr ysoleonites ocellatus Martynov;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, X 5,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1949)<br />

НАДСЕМЕЙСТВО MYRMELEONTIDEA<br />

Крылья длинные (д<strong>о</strong> 80 мм), <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

уз<strong>к</strong>ие; ди<strong>с</strong>т<strong>а</strong>льные ч<strong>а</strong><strong>с</strong>ти SC и R <strong>с</strong>литы; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь М <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R; MP и CuA п<strong>о</strong>чти<br />

п<strong>а</strong>р<strong>а</strong>ллельны; MP <strong>о</strong>дн<strong>о</strong>-д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь CuA т<strong>о</strong>лще <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей;<br />

CuP <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Юр<strong>а</strong> — ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Solenoptilidae, Nymphitidae, Myrmeleonidae,<br />

Ascalaphidae, Nemopteridae.<br />

СЕМЕЙСТВО SOLENOPTILIDAE<br />

HANDLIRSCH, 1908<br />

В переднем <strong>к</strong>рыле R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, без изгиб<strong>а</strong><br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS <strong>с</strong> изгиб<strong>о</strong>м п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует ряд пр<strong>о</strong><strong>с</strong>тых <strong>к</strong><strong>о</strong><strong>с</strong>ых <strong>в</strong>ет<strong>в</strong>ей.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Solenoptilon Handlirsch, 1908. Тип р<strong>о</strong>д<strong>а</strong>—<br />

АЫ<strong>а</strong> kochi Geinitz, 1888; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

http://jurassic.ru/

(<strong>в</strong>. лей<strong>а</strong><strong>с</strong>, Герм<strong>а</strong>ния). Передний <strong>к</strong>р<strong>а</strong>й <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут <strong>к</strong>з<strong>а</strong>ди, з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

33—40 мм (ри<strong>с</strong>. 852). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong><br />

и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО NYMPHITIDAE HANDLIRSCH, 1908<br />

SC и R д<strong>о</strong> <strong>с</strong>лияния <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong>;<br />

SC 4- R пл<strong>а</strong><strong>в</strong>н<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

R б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>, длинн<strong>а</strong>я, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong><br />

<strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; н<strong>а</strong> MP д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и;<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь СиА нерез<strong>к</strong><strong>о</strong> <strong>о</strong>тличн<strong>а</strong> <strong>о</strong>т<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ет<strong>в</strong>ей; MP и СиА не <strong>с</strong>ближены.<br />

Юр<strong>а</strong> — ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и Н<strong>о</strong><strong>в</strong><strong>о</strong>й Г<strong>в</strong>инеи; ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых.<br />

Sogjuta О. Martynova, 1958. Тип р<strong>о</strong>д<strong>а</strong>—<br />

5. speciosa О. Martynova, 1958; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). В переднем<br />

<strong>к</strong>рыле р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и уже <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти и н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong>дн<strong>о</strong>й ширины; не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>ет<strong>в</strong>и SC <strong>с</strong> д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чными<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; <strong>в</strong>ет<strong>в</strong>ей RS <strong>в</strong><strong>о</strong><strong>с</strong>емь, прямые;<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь RS <strong>о</strong>тх<strong>о</strong>дит пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; д<strong>в</strong><strong>а</strong> <strong>с</strong>тупенч<strong>а</strong>тых<br />

ряд<strong>а</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 12 мм<br />

(ри<strong>с</strong>. 853). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Chrysoleonites Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong><br />

— Ch. ocellatus Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). В передних<br />

<strong>к</strong>рыльях р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й ширины <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти шире,<br />

чем н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и<br />

SC пр<strong>о</strong><strong>с</strong>тые; де<strong>с</strong>ять <strong>в</strong>ет<strong>в</strong>ей RS, из<strong>о</strong>гнутых <strong>в</strong> ме<strong>с</strong>те<br />

пр<strong>о</strong>х<strong>о</strong>ждения пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> ряд<strong>а</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь RS <strong>о</strong>тх<strong>о</strong>дит<br />

н<strong>а</strong> <strong>с</strong>редине длины <strong>к</strong>рыл<strong>а</strong>; д<strong>в</strong><strong>а</strong> <strong>с</strong>тупенч<strong>а</strong>тых ряд<strong>а</strong><br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 15—17 мм<br />

(ри<strong>с</strong>. 854). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Вне СССР: Epigambria Handlirsch, 1939;<br />

Mesonymphes Carpenter, 1929; Nymphites Haase,<br />

1890; Sialium Westwood, 1854.<br />

СЕМЕЙСТВО MYRMELEONIDAE BURMEISTER, 1839.<br />

МУРАВЬИНЫЕ ЛЬВЫ<br />

SC и R д<strong>о</strong> <strong>с</strong>лияния <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong>;<br />

SC + R пл<strong>а</strong><strong>в</strong>н<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и SC пр<strong>о</strong><strong>с</strong>тые,<br />

SC + R — <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; MP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>с</strong>ближен<strong>а</strong> <strong>с</strong> СиА; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь СиА рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чен<strong>а</strong>; <strong>а</strong>нтенны <strong>в</strong> не­<br />

274<br />

http://jurassic.ru/<br />

<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>а</strong>з <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>к</strong>рылье<strong>в</strong>, р<strong>а</strong><strong>с</strong>ширены н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет. Личин<strong>к</strong><strong>а</strong> <strong>с</strong> шир<strong>о</strong><strong>к</strong>им тел<strong>о</strong>м,<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й и длинными челю<strong>с</strong>тями.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Обширн<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>: <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1200 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я<br />

<strong>к</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленным р<strong>о</strong>д<strong>а</strong>м; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ASCALAPHIDAE LEFROY, 1842<br />

[nom. transl. Schenider, 1845<br />

(ex Ascalaphides Lefroy, 1842)]<br />

SC и R п<strong>а</strong>р<strong>а</strong>ллельны, перед <strong>с</strong><strong>а</strong>мым <strong>с</strong>лиянием<br />

<strong>к</strong>рут<strong>о</strong> из<strong>о</strong>гнуты; SC + R прям<strong>а</strong>я, <strong>с</strong> R <strong>о</strong>бр<strong>а</strong>зует<br />

изл<strong>о</strong>м; <strong>в</strong>ет<strong>в</strong>и SC и <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти птер<strong>о</strong><strong>с</strong>тигмы<br />

пр<strong>о</strong><strong>с</strong>тые, SC + R — <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми и п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; MP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь СиА рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чен<strong>а</strong>. Антенны р<strong>а</strong><strong>с</strong>ширены<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце, длиннее <strong>к</strong>рылье<strong>в</strong> или немн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че их у <strong>к</strong>рупных ф<strong>о</strong>рм. Личин<strong>к</strong>и н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют<br />

личин<strong>к</strong>и мур<strong>а</strong><strong>в</strong>ьиных ль<strong>в</strong><strong>о</strong><strong>в</strong>. Не<strong>о</strong>ген —<br />

ныне. Обширн<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>: <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

р<strong>о</strong>ды, н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бильны <strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х, из них<br />

<strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong> Фр<strong>а</strong>нции.<br />

СЕМЕЙСТВО NEMOPTERIDAE HAGEN, 1866<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>в</strong>ытянутые, лент<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные;<br />

<strong>в</strong> передних <strong>к</strong>рыльях SC и R <strong>с</strong>ближ<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong>;<br />

SC + R пл<strong>а</strong><strong>в</strong>н<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и SC<br />

пр<strong>о</strong><strong>с</strong>тые, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены ч<strong>а</strong>ще;<br />

RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>; MP и СиА <strong>с</strong>ближены. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong>ытянут<strong>а</strong><br />

<strong>в</strong> <strong>к</strong>лю<strong>в</strong>, <strong>а</strong>нтенны неб<strong>о</strong>льшие, <strong>к</strong><strong>о</strong>р<strong>о</strong>че или<br />

р<strong>а</strong><strong>в</strong>ны <strong>к</strong>рыльям, без р<strong>а</strong><strong>с</strong>ширения н<strong>а</strong> <strong>к</strong><strong>о</strong>нце. Не<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 25<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 100 тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>), из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> ми<strong>о</strong>цене С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО POLYSTOECHOTIDEA<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

п<strong>о</strong>чти <strong>в</strong>тр<strong>о</strong>е; <strong>в</strong> передних <strong>к</strong>рыльях SC <strong>с</strong>лит<strong>а</strong><br />

<strong>с</strong> R; МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS<br />

у <strong>с</strong><strong>о</strong><strong>в</strong>ременных и мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм и <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>а</strong><br />

у п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их; MP делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е<br />

гл<strong>а</strong><strong>в</strong>ные <strong>в</strong>ет<strong>в</strong>и; СиА и CuP длинные, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими,<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>к</strong>з<strong>а</strong>ди <strong>в</strong>ет<strong>в</strong>ями; три<br />

<strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> мн<strong>о</strong>г<strong>о</strong>,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>ни <strong>о</strong>бр<strong>а</strong>зуют <strong>с</strong>тупенч<strong>а</strong>тые ряды.<br />

Пермь — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Permithonidae, Агcheosmylidae,<br />

Osmylitidae, Mesopolystoechotidae,<br />

Osmylidae, из них п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йден<strong>о</strong>.

855. Permegalomus maculipennis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

x4,5, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1952). 856.<br />

Eopsychops sojanensis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 4,6, <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1952). 857. Permithonopsis<br />

obscura О.Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X,5,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1952), 858. Petrushevskla borisi О. Martynova;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, X 4,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

СЕМЕЙСТВО PERMITHONIDAE TILLYARD, 1922<br />

(Permegalomidae О. Martynova, 1952;<br />

Permopsychopsidae Riek, 1953)<br />

В передних <strong>к</strong>рыльях р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шире<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong>ет<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; SC <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> R п<strong>о</strong>чти н<strong>а</strong><br />

<strong>с</strong>ередине ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; МА мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я;<br />

CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong>ч<strong>к</strong><strong>о</strong>м; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и пр<strong>а</strong><strong>в</strong>ильных ряд<strong>о</strong><strong>в</strong> не <strong>о</strong>бр<strong>а</strong>зуют.<br />

Пермь. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

R SC<br />

Ри<strong>с</strong>. 855—861. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Polystoechctidae<br />

1958). 859. Kirgisellodes ornatus (Martynov); переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Х4,6, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.) 860. Mesopolystoechus<br />

apicalis Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х4,0, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1949). 861. Lithosmylus columblanus (Cockerell); А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, Х2,5; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х2,5; не<strong>о</strong>ген, С. Амери<strong>к</strong><strong>а</strong><br />

(Carpenter, 1935)<br />

Permegalomus Martynov, 1930. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. regularis Martynov, 1930; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

Апи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>к</strong>руглый,<br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; <strong>в</strong>ет<strong>в</strong>и SC <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS и ди<strong>с</strong>т<strong>а</strong>льнее<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления Си. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 10 мм (ри<strong>с</strong>.<br />

855). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Приур<strong>а</strong>лья и Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Eopsychops Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. sojanensis Martynov, 1933; <strong>в</strong>. пермь, Арх<strong>а</strong>н-<br />

275 18*<br />

http://jurassic.ru/

гель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Апи<strong>к</strong><strong>а</strong>льный<br />

<strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>к</strong>руглый; ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong> передний <strong>к</strong>р<strong>а</strong>й<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>в</strong>ыем<strong>к</strong>у; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й из<strong>о</strong>гнут угл<strong>о</strong>м<br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>в</strong>ыем<strong>к</strong>и 'переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>и SC<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине пр<strong>о</strong><strong>с</strong>тые; длин<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

MP б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины МА; п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong>единяет МА <strong>с</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью<br />

RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 13 мм (ри<strong>с</strong>. 856). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Permithonopsis Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. ivensis Martynov, 1933; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

Апи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong> <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ченн<strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й не <strong>о</strong>бр<strong>а</strong>зует<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>г<strong>о</strong> угл<strong>а</strong>; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и SC <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS; длин<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> MP <strong>в</strong>д<strong>в</strong><strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> МА<br />

или еще <strong>к</strong><strong>о</strong>р<strong>о</strong>че. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10—<br />

16 мм (ри<strong>с</strong>. 857). 12 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: Permithone Tillyard, 1922; Permorapisma<br />

Tillyard, 1926; Pertnosmylus Tillyard,<br />

1926; Permopsychops Tillyard, 1926.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archeosmylidae Riek, 1953, близ<strong>к</strong><strong>о</strong>е<br />

<strong>к</strong> Permithonidae, из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>му р<strong>о</strong>ду<br />

Archeosmylus Riek, 1953 из <strong>в</strong>. перми и три<strong>а</strong><strong>с</strong><strong>а</strong><br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО OSMYLITIDAE О. MARTYNOVA, 1949<br />

В передних <strong>к</strong>рыльях <strong>с</strong>у'б<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ля уз<strong>к</strong>ие, п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й ширины; SC<br />

и R <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

зн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, <strong>с</strong>лит<strong>а</strong><br />

<strong>с</strong> RS; MP <strong>о</strong>бр<strong>а</strong>зует д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и; СиА<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; н<strong>а</strong> CuP и Ai (ин<strong>о</strong>гд<strong>а</strong><br />

А 2) ряд <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>в</strong>ет<strong>в</strong>ей. В. три<strong>а</strong><strong>с</strong> — <strong>в</strong>. юр<strong>а</strong>. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Petrushevskia О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. borisi О. Martynova, 1958; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Апи<strong>к</strong><strong>а</strong>льный<br />

<strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> з<strong>а</strong><strong>к</strong>руглен,<br />

<strong>в</strong>ершин<strong>а</strong> не <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чен<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей RS 9—11; ди<strong>с</strong>т<strong>а</strong>льный<br />

<strong>к</strong><strong>о</strong>нец CuP р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен п<strong>о</strong>л<strong>о</strong>г<strong>о</strong>; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля прямые;<br />

ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечных п<strong>о</strong>л<strong>о</strong><strong>с</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 16 мм (ри<strong>с</strong>. 858). Один <strong>в</strong>ид.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Kirgisellodes Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Kirgisella ornata Martynov, 1925; юр<strong>а</strong>.<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Апи<strong>к</strong><strong>а</strong>льный<br />

<strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ченн<strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й; SC и R <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я без п<strong>о</strong>­<br />

276<br />

http://jurassic.ru/<br />

<strong>с</strong>тепенн<strong>о</strong>г<strong>о</strong> <strong>с</strong>ближения; <strong>в</strong>ет<strong>в</strong>и SC <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> реже,<br />

чем <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ег<strong>о</strong> ч<strong>а</strong><strong>с</strong>ти; RS <strong>с</strong> 19 <strong>в</strong>ет<strong>в</strong>ями;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и RS длинные; п<strong>о</strong>перечные жил<strong>к</strong>и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>к</strong><strong>о</strong><strong>с</strong>ые; н<strong>а</strong> <strong>к</strong>рыле <strong>к</strong><strong>о</strong>ричне<strong>в</strong>ый<br />

ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> пятн<strong>а</strong>ми. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

20 мм (ри<strong>с</strong>. 859). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Вне СССР: Osmylites Haase, 1890; Tetanoption<br />

Bode, 1953; Mesosmylina Bode, 1953.<br />

СЕМЕЙСТВО MESOPOLYSTOECHOTIDAE<br />

O. MARTYNOVA, 1949<br />

Вершин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>с</strong>тр<strong>а</strong>я; SC и R,<br />

п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> <strong>с</strong>ближ<strong>а</strong>я<strong>с</strong>ь, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и<br />

RS без длинных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>; МА пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я,<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS; д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и MP; ряд<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их гребенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных <strong>в</strong>ет<strong>в</strong>ей<br />

н<strong>а</strong> СиА; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; д<strong>в</strong><strong>а</strong> <strong>с</strong>тупенч<strong>а</strong>тых ряд<strong>а</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>; ди<strong>с</strong>т<strong>а</strong>льный ряд <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ях<br />

СиА. Юр<strong>а</strong>. ОДИН р<strong>о</strong>д.<br />

Mesopolystoechus Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. apicalis Martynov, 1937; н. юр<strong>а</strong>, Ср.<br />

Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б I). В переднем <strong>к</strong>рыле<br />

неб<strong>о</strong>льшие р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ях SC + R; <strong>к</strong><strong>о</strong>нечные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> SC, <strong>в</strong>ет<strong>в</strong>ях RS, М и СиА<br />

<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льный ряд п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

п<strong>а</strong>р<strong>а</strong>ллелен з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зует<br />

б<strong>о</strong>рт. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

25 мм (ри<strong>с</strong>. 860). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

СЕМЕЙСТВО OSMYLIDAE LEACH, 1815<br />

[nom. transl. Brauer, 1851 (ex Osmylida Leach, 1815)]<br />

SC и R <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>; н<strong>а</strong> <strong>в</strong><strong>с</strong>ех<br />

<strong>в</strong>ет<strong>в</strong>ях гл<strong>а</strong><strong>в</strong>ных жил<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и; д<strong>в</strong><strong>а</strong><br />

ряд<strong>а</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>; ди<strong>с</strong>т<strong>а</strong>льный ряд<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ях MP; б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле и <strong>в</strong><br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>. Гл<strong>а</strong>з<strong>к</strong>и е<strong>с</strong>ть. Три<strong>а</strong><strong>с</strong>—<br />

ныне. 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уне ше<strong>с</strong>ти п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>,<br />

из них три — Osmylinae, Protosmylinae,<br />

Kempyninae — из<strong>в</strong>е<strong>с</strong>тны из третичных <strong>о</strong>тл<strong>о</strong>жений,<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

ПОДСЕМЕЙСТВО OSMYLINAE KROGER, 1913<br />

В переднем <strong>к</strong>рыле ди<strong>с</strong>т<strong>а</strong>льный ряд п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из 19 жил<strong>о</strong><strong>к</strong> и пр<strong>о</strong>тянут<br />

<strong>о</strong>т SC + R д<strong>о</strong> MP (ри<strong>с</strong>. 861). П<strong>а</strong>ле<strong>о</strong>ген —ныне.<br />

Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне и <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).

НАДСЕМЕЙСТВО HEMEROBIIDEA<br />

(Psychopsidoidea О. Martynova, 1952)<br />

Крылья <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> шир<strong>о</strong><strong>к</strong>ие, длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

ширину <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> и меньше; SC не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> R; SC, R и RS<br />

<strong>с</strong>ближены (<strong>к</strong>р<strong>о</strong>ме Chrysopidae и Mesochrysopidae),<br />

п<strong>а</strong>р<strong>а</strong>ллельны; <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ля <strong>в</strong>ме<strong>с</strong>те уже <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> (<strong>к</strong>р<strong>о</strong>ме Chrysopidae).<br />

У мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их и <strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм<br />

МА <strong>с</strong>лит<strong>а</strong> н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м пр<strong>о</strong>тяжении <strong>с</strong> R или<br />

<strong>с</strong> RS, у п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм МА <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я.<br />

Пермь — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Psychopidae, Kalligrammatidae,<br />

Brongniartiellidae, Palaemerobiidae,<br />

Sialidopseidae, Hemerobiidae, Sisyridae,<br />

Berothidae, Mesochrysopidae, Chrysopidae, Prohemerobiidae,<br />

Proberothidae, Osmylopsychopidae;<br />

три п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО PSYCHOPIDAE HANDLIRSCH, 1906<br />

[nom. transl. Handlirsch, 1908 (ex Psychopsinae Handlirsch,<br />

1906)]<br />

Б<strong>о</strong>льшие треуг<strong>о</strong>льные <strong>к</strong>рылья; <strong>в</strong> передних<br />

<strong>к</strong>рыльях <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>лей <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых;<br />

RS<br />

862—864. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Psychopidae<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я плече<strong>в</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

<strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льные ч<strong>а</strong><strong>с</strong>ти SC, R и RS <strong>с</strong><strong>о</strong>единены<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми т<strong>а</strong><strong>к</strong><strong>о</strong>й же т<strong>о</strong>лщины,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong><strong>о</strong>единяемые ими пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и;<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong><strong>о</strong>единения п<strong>о</strong><strong>с</strong>ледние дел<strong>а</strong>ют<strong>с</strong>я<br />

т<strong>о</strong>н<strong>к</strong>ими; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть п<strong>о</strong><br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зует б<strong>о</strong>рт, <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичный<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>му п<strong>о</strong>лю; МА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и или р<strong>а</strong>збр<strong>о</strong><strong>с</strong><strong>а</strong>ны п<strong>о</strong> <strong>к</strong>рылу<br />

бе<strong>с</strong>п<strong>о</strong>ряд<strong>о</strong>чн<strong>о</strong>, или <strong>о</strong>бр<strong>а</strong>зуют пр<strong>а</strong><strong>в</strong>ильные ряды;<br />

з<strong>а</strong>дние <strong>к</strong>рылья уже передних. Юр<strong>а</strong> — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии, Ю. Афри<strong>к</strong>и<br />

и Ю. Азии <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>; и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых де<strong>в</strong>ять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Angaropsychops О. Martynova, 1949.Тип<br />

р<strong>о</strong>д<strong>а</strong>—A. turgensis О. Martynova, 1949; <strong>в</strong>. юр<strong>а</strong>,<br />

Читин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье, тургин<strong>о</strong>-<strong>в</strong>итим<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Быр<strong>к</strong><strong>а</strong>). SC, R и RS <strong>в</strong>ыгнуты <strong>к</strong>з<strong>а</strong>ди;<br />

MP делит<strong>с</strong>я п<strong>о</strong>чти <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, <strong>о</strong>бр<strong>а</strong>зуя<br />

ряд длинных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>в</strong>д<strong>в</strong><strong>о</strong>е уже <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

мн<strong>о</strong>г<strong>о</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены бе<strong>с</strong>п<strong>о</strong>ряд<strong>о</strong>чн<strong>о</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 40 мм (ри<strong>с</strong>. 862). Один <strong>в</strong>ид. В.<br />

юр<strong>а</strong> З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лья.<br />

862 Angaropsychops turgensis О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, <strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.; <strong>с</strong>дел<strong>а</strong>н п<strong>о</strong> ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>). 864. Gramme-<br />

Х2 0 <strong>в</strong> юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1949), 863. Embaneura psychops lebedevi О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, Х2,0, <strong>в</strong>. мел,<br />

vachrameevi G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, xl,5, <strong>в</strong>. мел, 3. К<strong>а</strong>з<strong>а</strong>х- Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>яр<strong>с</strong><strong>к</strong>ий <strong>к</strong>р<strong>а</strong>й (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1954)<br />

277<br />

http://jurassic.ru/

Embaneura G. Zalessky, 1953. Тип р<strong>о</strong>д<strong>а</strong>—<br />

E. vachrameevi G. Zalessky, 1953; <strong>в</strong>. мел,<br />

А<strong>к</strong>тюбин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>с</strong>ен<strong>о</strong>м<strong>а</strong>н, Эмб<strong>а</strong>). SC, R и<br />

RS <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыгнуты <strong>к</strong>з<strong>а</strong>ди; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ля п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й ширины; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и п<strong>о</strong>чти <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; <strong>к</strong>рыл<strong>о</strong> темн<strong>о</strong>е<br />

<strong>с</strong> тремя <strong>с</strong><strong>в</strong>етлыми <strong>о</strong><strong>к</strong>руглыми пятн<strong>а</strong>ми. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50 мм (ри<strong>с</strong>. 863). Один<br />

<strong>в</strong>ид. В. мел 3. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Grammopsychops О. Martynova, 1954.<br />

Тип р<strong>о</strong>д<strong>а</strong> — G. lebedevi О. Martynova, 1954; <strong>в</strong>.<br />

мел, Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>яр<strong>с</strong><strong>к</strong>ий <strong>к</strong>р<strong>а</strong>й (<strong>с</strong>уч<strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

р. Кемь). SC, R и RS не <strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong>ыгиб<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди;<br />

пять р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных гребенч<strong>а</strong>т<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей М<br />

д<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления ее н<strong>а</strong> МА и MP; МА <strong>с</strong>лит<strong>а</strong><br />

<strong>в</strong> RS; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле немн<strong>о</strong>г<strong>о</strong> уже <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>к</strong><strong>о</strong>нцентричн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных ряд<strong>о</strong><strong>в</strong>, пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ющих<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 44 мм (ри<strong>с</strong>. 864). Один <strong>в</strong>ид. В.<br />

мел В. Сибири.<br />

Вне СССР: Apeirophlebia Handlirsch, 1908;<br />

Liassopsychops Bode, 1953; Archepsychops Tillyard,<br />

1919; Triassopsychops Tillyard, 1922; Protopsychopsis<br />

Tillyard, 1917 и <strong>о</strong>дин из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря и ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО KALLIGRAMMATIDAE<br />

HANDLIRSCH, 1906<br />

Крупные <strong>к</strong>рылья (80—120 мм <strong>в</strong> длину);<br />

з<strong>а</strong>дние <strong>к</strong>рылья уже передних; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

<strong>в</strong>д<strong>в</strong><strong>о</strong>е шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых или р<strong>а</strong><strong>в</strong>н<strong>о</strong> им; <strong>с</strong>ильн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> sc-r; п<strong>о</strong>д<strong>о</strong>бие <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля п<strong>о</strong><br />

з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> е<strong>с</strong>ть, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е; мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я<br />

MP; з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь MP <strong>о</strong>бр<strong>а</strong>зует ряд<br />

н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>в</strong>перед <strong>в</strong>ет<strong>в</strong>ей; б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет <strong>в</strong><strong>с</strong>е <strong>к</strong>рыл<strong>о</strong>. В. юр<strong>а</strong>. Три р<strong>о</strong>д<strong>а</strong>.<br />

Kalligramma Walter, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К. haeckeli Walter, 1904; <strong>в</strong>. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

Ри<strong>с</strong>. 865. 867. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Kalligrammatidae, Brongniartiellidae<br />

865. Kalligramma haeckeli Walter; A - переднее <strong>к</strong>рыл<strong>о</strong>, X0.6; <strong>в</strong>. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1919). 867. Epactinophlebia karaffi(T<br />

3<br />

Ж„ е<br />

,? рыл0<br />

," X<br />

¥ ;<br />

и' ю<br />

- р<strong>а</strong><br />

' г<br />

м <strong>а</strong><br />

£Р , н и я<br />

(Handlirsch, 1908). basica Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 3,0, <strong>в</strong>. юр<strong>а</strong> Ю К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

л 5<br />

866. Kalhgrammula senkenbergiana Handlirsch; <strong>к</strong>рылья, Х0.75, (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1927)<br />

278<br />

http://jurassic.ru/

(м<strong>а</strong>льм, З<strong>о</strong>льнг<strong>о</strong>фен). В передних <strong>к</strong>рыльях<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> п<strong>о</strong>лт<strong>о</strong>р<strong>а</strong> р<strong>а</strong>з<strong>а</strong> шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>лей <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых;<br />

<strong>в</strong>ет<strong>в</strong>ей RS 8—12; <strong>к</strong>ругл<strong>о</strong>е д<strong>в</strong>уц<strong>в</strong>етн<strong>о</strong>е пятн<strong>о</strong><br />

<strong>в</strong> н<strong>а</strong>ч<strong>а</strong>ле ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 90—120 мм (ри<strong>с</strong>. 865). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Kalligrammula Handlirsch, 1919. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К. senkenbergiana Handlirsch, 1919; <strong>в</strong>.<br />

юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (м<strong>а</strong>льм, З<strong>о</strong>льнг<strong>о</strong>фен). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й ширины <strong>с</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

и р<strong>а</strong>ди<strong>а</strong>льным п<strong>о</strong>лями <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятыми;<br />

<strong>в</strong>ет<strong>в</strong>ей RS пять — <strong>в</strong><strong>о</strong><strong>с</strong>емь; пятен н<strong>а</strong> <strong>к</strong>рыле нет.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 70—80 мм (ри<strong>с</strong>. 866).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Вне СССР: Meioneurites Handlirsch, 1908.<br />

СЕМЕЙСТВО BRONGNIARTIELLIDAE<br />

О. MARTYNOVA, 1949<br />

Крылья треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и sc-r и r-rs <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; <strong>в</strong>ет<strong>в</strong>ей RS 16—26; MP р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>; CuA и CuP <strong>с</strong> б<strong>о</strong>льшим<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong><br />

нет. Юр<strong>а</strong>. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Epactinophlebia Martynov, 1927. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. karabasica Martynov, 1927; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). В б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> п<strong>о</strong>лт<strong>о</strong>р<strong>а</strong><br />

р<strong>а</strong>з<strong>а</strong> шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>лей <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых; <strong>в</strong>ет<strong>в</strong>ей RS 27; MP <strong>о</strong>бр<strong>а</strong>зует<br />

д<strong>в</strong>е гл<strong>а</strong><strong>в</strong>ные <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong>ет<strong>в</strong>и CuA и CuP р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

гребенч<strong>а</strong>т<strong>о</strong>; <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong>д<strong>в</strong><strong>о</strong>е<br />

шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

21,5 мм (ри<strong>с</strong>. 867). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у.<br />

Вне СССР: Actinophlebia Handlirsch, 1908;<br />

Brongniartiella Meunier, 1897; Mesopsychopsis<br />

Handlirsch, 1908; Pterinoblattlna Scudder, 1885;<br />

Par actinophlebia Handlirsch, 1908.<br />

СЕМЕЙСТВО PALAEMEROBIIDAE MARTYNOV, 1928<br />

Крылья неб<strong>о</strong>льшие (8—12 мм <strong>в</strong> длину),<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льные; <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

з<strong>а</strong><strong>к</strong>руглен; передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и <strong>в</strong> <strong>к</strong><strong>о</strong>нце; н<strong>а</strong> RS<br />

не менее четырех <strong>в</strong>ет<strong>в</strong>ей; <strong>о</strong>бе <strong>в</strong>ет<strong>в</strong>и М <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дны;<br />

МА не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS; МА и MP р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong>е гл<strong>а</strong><strong>в</strong>ные <strong>в</strong>ет<strong>в</strong>и, при эт<strong>о</strong>м р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ление<br />

МА р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления<br />

MP. CuA р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я д<strong>в</strong><strong>а</strong>жды.<br />

CuP <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; три <strong>а</strong>н<strong>а</strong>льные<br />

жил<strong>к</strong>и, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; пр<strong>о</strong>д<strong>о</strong>льные<br />

жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> р<strong>а</strong>зных р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>яниях<br />

<strong>о</strong>т <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; ше<strong>с</strong>ть — <strong>с</strong>емь п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

между <strong>в</strong><strong>с</strong>еми пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

бе<strong>с</strong>п<strong>о</strong>ряд<strong>о</strong>чн<strong>о</strong>. Пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Palaemerobius Martynov, 1928. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. proavitus Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

В переднем <strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong><br />

<strong>с</strong>ередине ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; пять —<br />

ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей SC перех<strong>о</strong>дят н<strong>а</strong> R; ширин<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля р<strong>а</strong><strong>в</strong>н<strong>а</strong> Vs ширины <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>в</strong>ет<strong>в</strong>ей RS пять — ше<strong>с</strong>ть; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; МА <strong>в</strong>ет<strong>в</strong>ит<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

8,0—8,5 мм (ри<strong>с</strong>. 868). Три <strong>в</strong>ид<strong>а</strong>. Пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл., Приур<strong>а</strong>лья, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Tychtobius О. Martynova, 1958. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. brevicostatus О. Martynova, 1958; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий 'б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). В переднем <strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>; ширин<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля р<strong>а</strong><strong>в</strong>н<strong>а</strong> 1,8 ширины<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>; де<strong>с</strong>ять <strong>в</strong>ет<strong>в</strong>ей SC (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми) перех<strong>о</strong>дят н<strong>а</strong> R; <strong>в</strong>ет<strong>в</strong>ей RS<br />

ше<strong>с</strong>ть; МА р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины<br />

<strong>к</strong>рыл<strong>а</strong>; MP — немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее. Ан<strong>а</strong>льные<br />

жил<strong>к</strong>и <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 10,75 мм (ри<strong>с</strong>. 869). Один <strong>в</strong>ид. Пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Bianchia О. Martynova, 1952; Тип р<strong>о</strong>д<strong>а</strong><br />

— В. spectabilis О. Martynova, 1952; <strong>в</strong>.<br />

пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

С<strong>о</strong>ян<strong>а</strong>). В переднем <strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>о</strong>дн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь SC перех<strong>о</strong>дит н<strong>а</strong> R; <strong>в</strong>ет<strong>в</strong>ей<br />

RS ше<strong>с</strong>ть; МА р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

длины <strong>к</strong>рыл<strong>а</strong>; <strong>с</strong>т<strong>в</strong><strong>о</strong>л MP п<strong>о</strong>чти <strong>в</strong>д<strong>в</strong><strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> МА. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 11 мм<br />

(ри<strong>с</strong>. 870). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

СЕМЕЙСТВО SIAL1DOPSEIDAE<br />

М. ZALESSKY, 1926<br />

(Permosisyridae Martynov, 1933)<br />

Крылья <strong>о</strong><strong>в</strong><strong>а</strong>льные (длин<strong>а</strong> 7,6—13,0 мм); <strong>в</strong><br />

переднем <strong>к</strong>рыле <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й з<strong>а</strong><strong>к</strong>руглен;<br />

передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния и <strong>в</strong><br />

<strong>к</strong><strong>о</strong>нце <strong>к</strong>рыл<strong>а</strong>; S.C <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и<br />

SC пр<strong>о</strong><strong>с</strong>тые, перех<strong>о</strong>дят н<strong>а</strong> R; <strong>в</strong>ет<strong>в</strong>ей RS четыре;<br />

пр<strong>о</strong>д<strong>о</strong>льные <strong>в</strong>ет<strong>в</strong>и <strong>о</strong>бр<strong>а</strong>зуют р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong><br />

р<strong>а</strong>зн<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>ые п<strong>о</strong>перечные<br />

жил<strong>к</strong>и имеют<strong>с</strong>я <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле,<br />

и немн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> р<strong>а</strong>збр<strong>о</strong><strong>с</strong><strong>а</strong>н<strong>о</strong> п<strong>о</strong><br />

<strong>к</strong>рылу. Пермь. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

http://jurassic.ru/

868. Palaemerobius proavitus Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, Хб,5,<br />

<strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928). 869. Tychtobius brevicostatus<br />

О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, Хб,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 870. Bianchia spectabilis O.Martynova;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х5,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1952). 871. Slaltdopsis sojanensis О. Martynova; переднее<br />

Sialidopsis М. Zalessky, 1926. Тип р<strong>о</strong>д<strong>а</strong><br />

— S. kargalensis М. Zalessky, 1926; <strong>в</strong>. пермь,<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). В переднем<br />

<strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>о</strong>нце третьей чет<strong>в</strong>ерти<br />

длины <strong>к</strong>рыл<strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не<br />

<strong>с</strong>ередины <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RS. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 10,5—12,5 мм (ри<strong>с</strong>. 871). Три <strong>в</strong>ид<strong>а</strong>. В.<br />

пермь Приур<strong>а</strong>лья и Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Permosisyra Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. latipennis Martynov, 1933; <strong>в</strong>. пермь,<br />

Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>).<br />

В переднем <strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

третьей чет<strong>в</strong>ерти длины <strong>к</strong>рыл<strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

RS; CuA <strong>с</strong> пр<strong>о</strong><strong>с</strong>тым р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 7,5—10,5 мм (ри<strong>с</strong>. 872). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Parasisyra G. Zalessky, 1933. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. kargalica G. Zalessky, 1933; <strong>в</strong>. пермь.<br />

Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (К<strong>а</strong>рг<strong>а</strong>л<strong>а</strong>). В переднем<br />

<strong>к</strong>рыле SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; де<strong>с</strong>ять <strong>в</strong>ет<strong>в</strong>ей SC перех<strong>о</strong>дит<br />

н<strong>а</strong> R; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле д<strong>в</strong>е п<strong>о</strong>перечные<br />

жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм<br />

(ри<strong>с</strong>. 873). Один <strong>в</strong>ид. Пермь Приур<strong>а</strong>лья.<br />

Ри<strong>с</strong>. 868—873. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Palaemerobiidae, Sialidopseidae<br />

280<br />

http://jurassic.ru/<br />

<strong>к</strong>рыл<strong>о</strong>, х6,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1952).<br />

872. Permosisyra latipennis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 7,2,<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933). 873. Parasisyra<br />

kargalica G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, х7,0, <strong>в</strong>. пермь, Ю. Приур<strong>а</strong>лье<br />

(Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий)<br />

СЕМЕЙСТВО HEMEROBIIDAE<br />

LATREILLE, 1802<br />

[nom. transl. Westwood, 1840<br />

(ex Hemerobini Latreille, 1802)]<br />

Длин<strong>а</strong> <strong>к</strong>рылье<strong>в</strong> 3—15 мм, пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют р<strong>а</strong>змеры<br />

8—10 мм; <strong>в</strong> передних <strong>к</strong>рыльях плече<strong>в</strong><strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> длинн<strong>а</strong>я, из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>рыл<strong>а</strong>; SC длинн<strong>а</strong>я, н<strong>а</strong> R перех<strong>о</strong>дят д<strong>в</strong>е — четыре<br />

<strong>в</strong>ет<strong>в</strong>и SC; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рыл<strong>а</strong>; МА и д<strong>в</strong>е — четыре пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные<br />

<strong>в</strong>ет<strong>в</strong>и RS (ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и RS, <strong>к</strong><strong>а</strong><strong>к</strong> у р<strong>о</strong>д<strong>а</strong><br />

Drepanopteryx) <strong>с</strong>литы <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> R и <strong>о</strong>тх<strong>о</strong>дят<br />

<strong>о</strong>т нее; MP <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е<br />

длинные <strong>в</strong>ет<strong>в</strong>и; CuA <strong>с</strong> гребенч<strong>а</strong>тым ряд<strong>о</strong>м <strong>в</strong>ет<strong>в</strong>ей<br />

<strong>к</strong>з<strong>а</strong>ди; CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я (<strong>к</strong>р<strong>о</strong>ме <strong>к</strong>рупных<br />

ф<strong>о</strong>рм); три <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и; п<strong>о</strong>перечные жил<strong>к</strong>и<br />

<strong>о</strong>бр<strong>а</strong>зуют д<strong>в</strong><strong>а</strong> <strong>с</strong>тупенч<strong>а</strong>тых ряд<strong>а</strong> (ри<strong>с</strong>. 874).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

25 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных р<strong>о</strong>д<strong>о</strong><strong>в</strong>; четыре<br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО S1SYRIDAE HANDLIRSCH, 1906<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 5—6 мм; <strong>в</strong> передних<br />

<strong>к</strong>рыльях <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля уз<strong>к</strong><strong>о</strong>е,<br />

плече<strong>в</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й

874. Hemerobius humuli Linnaeus; A — переднее <strong>к</strong>рыл<strong>о</strong>, X7,0;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х7,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Comstock, 1918).<br />

875. Sisyra flavicornis Linnaeus; A — переднее <strong>к</strong>рыл<strong>о</strong>, X10.0;<br />

Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х 10,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (<strong>о</strong>риг. ри<strong>с</strong>.). 876.<br />

Spermophorella sp.; А — переднее <strong>к</strong>рыл<strong>о</strong>, х 3,0; Б — з<strong>а</strong>днее <strong>к</strong>ры-<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> sc-r;<br />

МА <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> R и <strong>о</strong>тх<strong>о</strong>дит<br />

<strong>о</strong>т RS; MP делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не<br />

н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й МА, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> ma-mpi; CuA <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями;<br />

CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong>,<br />

ряд<strong>о</strong><strong>в</strong> их нет (ри<strong>с</strong>. 875). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных <strong>в</strong>ид<strong>о</strong><strong>в</strong>), из них <strong>о</strong>дин<br />

р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО BEROTHIDAE HANDLIRSCH, 1906<br />

{nom. transl. Handlirsch, 1908<br />

(ex Berothinae Handlirsch, 1906)]<br />

Ри<strong>с</strong>. 874—878. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hemerobiidea<br />

В передних <strong>к</strong>рыльях плече<strong>в</strong><strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я;<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; н<strong>а</strong> R перех<strong>о</strong>дят<br />

пять — ше<strong>с</strong>ть <strong>в</strong>ет<strong>в</strong>ей SC; М, МА, MP <strong>с</strong>литы <strong>в</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> R; МА <strong>о</strong>тх<strong>о</strong>дит уже <strong>о</strong>т RS, MP —<br />

<strong>о</strong>т R; <strong>в</strong>ет<strong>в</strong>и RS не <strong>с</strong>литы <strong>с</strong> R; СиА <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями;<br />

CuP <strong>с</strong> длинным р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; н<strong>а</strong> <strong>к</strong>рыльях ряды<br />

281<br />

http://jurassic.ru/<br />

л<strong>о</strong>, Х3,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем., К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (<strong>о</strong>риг. ри<strong>с</strong>.). 877. Mesypochrysa<br />

latipennis Martynov; <strong>о</strong>бщий <strong>в</strong>ид, Х3,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 878. Palaeochrysa stricta Scudder; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 3,5; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 3,5; не<strong>о</strong>ген, С. Амери<strong>к</strong><strong>а</strong><br />

(Carpenter, 1935)<br />

чешуе<strong>к</strong> (ри<strong>с</strong>. 876). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне де<strong>с</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong> (тр<strong>о</strong>пи<strong>к</strong>и, <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong>и);<br />

<strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО MESOCHRYSOPIDAE<br />

HANDLIRSCH, 1908<br />

К<strong>а</strong>жд<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>с</strong>тым<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>в</strong> переднем <strong>к</strong>рыле<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; R д<strong>о</strong>х<strong>о</strong>дит д<strong>о</strong> <strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong>,<br />

из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди; <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле п<strong>о</strong>перечные<br />

жил<strong>к</strong>и, чи<strong>с</strong>л<strong>о</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует чи<strong>с</strong>лу<br />

<strong>в</strong>ет<strong>в</strong>ей RS; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS, <strong>о</strong>бе <strong>в</strong>ет<strong>в</strong>и <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> з<strong>а</strong>днем<br />

<strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>; Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее М;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>о</strong>бр<strong>а</strong>зуют д<strong>в</strong><strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных<br />

ряд<strong>а</strong>. В. юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Mesypochrysa М <strong>а</strong> г t у п <strong>о</strong> v, 1927. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. latipennis Martynov, 1927; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). В переднем<br />

<strong>к</strong>рыле SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти птер<strong>о</strong>-

<strong>с</strong>тигмы, пере<strong>с</strong>еченн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

<strong>в</strong>ет<strong>в</strong>ями; <strong>в</strong>ет<strong>в</strong>ей RS <strong>с</strong>емь. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 12 мм (ри<strong>с</strong>. 877). Один <strong>в</strong>ид. В. юр<strong>а</strong>.<br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Вне СССР: Mesochrysopa Handlirsch,<br />

1906.<br />

СЕМЕЙСТВО CHRYSOPIDAE HAGEN, 1866<br />

В переднем <strong>к</strong>рыле SC и R из<strong>о</strong>гнуты <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

изгибу <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>, д<strong>о</strong><br />

<strong>в</strong>ершины <strong>к</strong>рыл<strong>а</strong> не д<strong>о</strong>х<strong>о</strong>дят; р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых;<br />

чи<strong>с</strong>л<strong>о</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует чи<strong>с</strong>лу <strong>в</strong>ет<strong>в</strong>ей RS; М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>лит<strong>а</strong> <strong>с</strong> R; MP, <strong>о</strong>бр<strong>а</strong>зуя петлю, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> МА. MP + МА <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями<br />

RS и не д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; Ai и А 2 <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми; А 3 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; д<strong>в</strong><strong>а</strong><br />

пр<strong>о</strong>д<strong>о</strong>льных ряд<strong>а</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> (ри<strong>с</strong>.<br />

878). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

40 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных р<strong>о</strong>д<strong>о</strong><strong>в</strong>; четыре<br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

и С. Амери<strong>к</strong>и.<br />

Ри<strong>с</strong>. 879—881. Hemerobiidea incertae sedis<br />

879. Archedilaropsis furcata Martynov; фр<strong>а</strong>гмент <strong>к</strong>рыл<strong>о</strong>, X 3,2,<br />

не<strong>о</strong>ген, 3. Сибирь (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1935). 880. Dilarites incertus Martynov;<br />

фр<strong>а</strong>гмент, хЗ,5, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925).<br />

£81. Eopsychopsis permiana Martynov; фр<strong>а</strong>гмент переднег<strong>о</strong><br />

_ <strong>к</strong>рыл<strong>а</strong>, х.9,0. <strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928)<br />

282<br />

http://jurassic.ru/<br />

HEMEROBIIDEA INCERTAE SEDIS<br />

Archedilaropsis Martynov, 1935 (ри<strong>с</strong>.<br />

879); <strong>в</strong>. три<strong>а</strong><strong>с</strong> — н. юр<strong>а</strong>, <strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>чный <strong>с</strong><strong>к</strong>л<strong>о</strong>н<br />

Ур<strong>а</strong>л<strong>а</strong>; Dilarites Martynov, 1925 (ри<strong>с</strong>. 880); <strong>в</strong>.<br />

юр<strong>а</strong>, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у; Eopsychopsis Martynov, 1928<br />

(ри<strong>с</strong>. 881); н. пермь, Тихие Г<strong>о</strong>ры.<br />

Вне СССР: юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы — Archegetes<br />

Handlirsch, 1906; Creagroptera Handlirsch, 1906;<br />

Dicranoptila Handlirsch, 1906; Gigantotermes<br />

Haase, 1890; Ineptiae Handlirsch, 1906; Loxophleps<br />

Handlirsch, 1939; Megapolystoechus Tillyard,<br />

1933; Melamnous Handlirsch, 1939; Melaneimon<br />

Handlirsch, 1939; Mesoleon Handlirsch,<br />

1906; Osmylopsis Handlirsch, 1906; Palparites<br />

Handlirsch, 1906; Pseudomyrmeleon<br />

Handlirsch, 1906; мел Кит<strong>а</strong>я — Mesohetnerobius<br />

Ping, 1928.<br />

НАДСЕМЕЙСТВО CONIOPTERYGIDEA<br />

Очень мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые: 3—10 мм <strong>в</strong> р<strong>а</strong>зм<strong>а</strong>хе.<br />

Крылья п<strong>о</strong><strong>к</strong>рыты бел<strong>о</strong>й пыльц<strong>о</strong>й, жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>о</strong>бедненн<strong>о</strong>е, без д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

з<strong>а</strong>дние <strong>к</strong>рылья уже и <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних. Юр<strong>а</strong> —<br />

ныне. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Coniopterygidae.<br />

СЕМЕЙСТВО CONIOPTERYGIDAE<br />

BURMEISTER, 1839<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>,<br />

не <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь <strong>с</strong> R; RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> <strong>о</strong>дним<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; М и Си р<strong>а</strong>зделяют<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е<br />

Ри<strong>с</strong>. 882. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Coniopterygidea. Semidalis alcurodiformis<br />

Stephens; <strong>к</strong>рылья, X 15,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Handlirsch, 1925)<br />

пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и (ри<strong>с</strong>. 882). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>; четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и 3. Е<strong>в</strong>р<strong>о</strong>пы.

НАДОТРЯД MECOPTEROIDEA. МЕКОПТЕРОИДЫ<br />

Крылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> переп<strong>о</strong>нч<strong>а</strong>тые, <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>н<strong>с</strong>и<strong>с</strong>тенции, ин<strong>о</strong>гд<strong>а</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, ч<strong>а</strong>ще же передние<br />

б<strong>о</strong>льше з<strong>а</strong>дних; п<strong>о</strong><strong>с</strong>ледние ин<strong>о</strong>гд<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че передних или же <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют:<br />

диптеригия р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong><strong>о</strong> мн<strong>о</strong>гих<br />

групп<strong>а</strong>х. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние бедн<strong>о</strong>е, п<strong>о</strong>перечные жил<strong>к</strong>и<br />

<strong>о</strong>бычн<strong>о</strong> немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны. Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е грызущег<strong>о</strong> тип<strong>а</strong>: ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>ни пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>ущий или <strong>к</strong><strong>о</strong>лющий х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения. Личин<strong>к</strong>и эру<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й; грудные<br />

н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, реже длинные, ин<strong>о</strong>гд<strong>а</strong> имеют<strong>с</strong>я<br />

л<strong>о</strong>жн<strong>о</strong>н<strong>о</strong>ж<strong>к</strong>и н<strong>а</strong> брюш<strong>к</strong>е, или н<strong>о</strong>ги <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е <strong>о</strong>т<strong>с</strong>ут-<br />

Д<strong>в</strong>е п<strong>а</strong>ры п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных <strong>к</strong>рылье<strong>в</strong><br />

(<strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряде Paratrichoptera н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

уменьшение з<strong>а</strong>дней п<strong>а</strong>ры <strong>к</strong>рылье<strong>в</strong>);<br />

RS и М дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и делят<strong>с</strong>я н<strong>а</strong> четыре<br />

гл<strong>а</strong><strong>в</strong>ные <strong>в</strong>ет<strong>в</strong>и; <strong>в</strong>ет<strong>в</strong>ей RS четыре —<br />

д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>ть; <strong>в</strong>ет<strong>в</strong>ей М четыре — <strong>в</strong><strong>о</strong><strong>с</strong>емь;<br />

Си р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong><br />

д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и, CuA бы<strong>в</strong><strong>а</strong>ет <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, CuP<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; <strong>о</strong>бычн<strong>о</strong> три <strong>а</strong>н<strong>а</strong>льные<br />

жил<strong>к</strong>и, д<strong>о</strong>х<strong>о</strong>дящие д<strong>о</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>;<br />

три — четыре ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чных <strong>о</strong>рг<strong>а</strong>н<strong>а</strong> <strong>с</strong><br />

п<strong>о</strong><strong>с</strong>т<strong>о</strong>янным ме<strong>с</strong>т<strong>о</strong>п<strong>о</strong>л<strong>о</strong>жением.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, <strong>с</strong> жующим р<strong>о</strong>т<strong>о</strong><strong>в</strong>ым<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>о</strong>м, <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong> <strong>в</strong>иде <strong>к</strong>лю<strong>в</strong><strong>а</strong><br />

(<strong>к</strong>р<strong>о</strong>ме п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Paratrichoptera), у<strong>с</strong>и<strong>к</strong>и<br />

ните<strong>в</strong>идные, не пре<strong>в</strong>ыш<strong>а</strong>ющие длину<br />

<strong>к</strong>рыл<strong>а</strong>; б<strong>о</strong>льшие <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>, три гл<strong>а</strong>з<strong>к</strong><strong>а</strong>.<br />

Переднегрудь немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>реднеи<br />

з<strong>а</strong>днегруди. Н<strong>о</strong>ги г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, бег<strong>а</strong>тельные;<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я, <strong>с</strong> д<strong>в</strong>умя<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем члени<strong>к</strong>е (у Bittacidae<br />

<strong>о</strong>дин <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>о</strong><strong>к</strong>). Брюш<strong>к</strong><strong>о</strong> немн<strong>о</strong>г<strong>о</strong><br />

длиннее <strong>к</strong>рылье<strong>в</strong>, <strong>с</strong> р<strong>а</strong>здутым де<strong>в</strong>ятым генит<strong>а</strong>льным<br />

<strong>с</strong>егмент<strong>о</strong>м, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми<br />

г<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>а</strong>ми; <strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряде Eumecoptera <strong>с</strong><br />

мез<strong>о</strong>з<strong>о</strong>я н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я удлинение ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong>,<br />

<strong>с</strong>едьм<strong>о</strong>г<strong>о</strong> и <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые цер<strong>к</strong>и. Личин<strong>к</strong>и<br />

эру<strong>к</strong><strong>о</strong><strong>в</strong>идные, <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> гнилых пнях<br />

и <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й ле<strong>с</strong>н<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е. Взр<strong>о</strong><strong>с</strong>лые<br />

и личин<strong>о</strong>чные <strong>с</strong>т<strong>а</strong>дии пит<strong>а</strong>ют<strong>с</strong>я гниющими<br />

<strong>о</strong>рг<strong>а</strong>ниче<strong>с</strong><strong>к</strong>ими <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>ми, ц<strong>в</strong>ет<strong>о</strong>чн<strong>о</strong>й<br />

пыльц<strong>о</strong>й, яйц<strong>а</strong>ми Lepidoptera, ред<strong>к</strong><strong>о</strong><br />

<strong>с</strong><strong>в</strong>ежими мерт<strong>в</strong>ыми н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мыми, <strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е<br />

Bittacidae — хищни<strong>к</strong>и. П<strong>о</strong>д<strong>о</strong>тряд<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

<strong>с</strong>т<strong>в</strong>уют. Обит<strong>а</strong>ют <strong>в</strong> <strong>с</strong><strong>а</strong>мых р<strong>а</strong>зличных у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях<br />

— <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, <strong>в</strong> <strong>в</strong><strong>о</strong>де, <strong>в</strong> гниющих <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong>х,<br />

т<strong>к</strong><strong>а</strong>нях р<strong>а</strong><strong>с</strong>тений, п<strong>а</strong>р<strong>а</strong>зитируют <strong>в</strong> теле жи<strong>в</strong><strong>о</strong>тных<br />

или жи<strong>в</strong>ут <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong> н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях. С<strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

пре<strong>в</strong>р<strong>а</strong>щения (<strong>а</strong>д<strong>а</strong>пти<strong>в</strong>ные изменения<br />

ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития), пит<strong>а</strong>ния (изменение<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>) и л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>ции (р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong><br />

лет<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong>) —<strong>в</strong><strong>с</strong>е<br />

эти при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления <strong>о</strong>пределили р<strong>а</strong>з<strong>в</strong>итие и<br />

ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ние ме<strong>к</strong><strong>о</strong>птер<strong>о</strong>ид<strong>о</strong><strong>в</strong>—ныне н<strong>а</strong>иб<strong>о</strong>лее<br />

мн<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong> <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. Отряды: Mecoptera,<br />

Trichoptera, Lepidoptera, Diptera, Aphaniptera.<br />

ОТРЯД MECOPTERA. СКОРПИОННИЦЫ<br />

(О. M. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

CuA CuP<br />

884<br />

Ри<strong>с</strong>. 883—884. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Platychoristidae<br />

883. Platychorista venosa Tillyard (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция), X11.7,<br />

н. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1930). 884. Marimerobius sp/endens<br />

G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, x7,0, н. пермь, Ур<strong>а</strong>л (<strong>о</strong>р<strong>к</strong>г.<br />

ри<strong>с</strong>. <strong>с</strong> ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

283<br />

http://jurassic.ru/

Protomecoptera д<strong>а</strong>л н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>в</strong>ет<strong>в</strong>и Trichoptera — Lepidoptera; п<strong>о</strong>д<strong>о</strong>тряд Р<strong>а</strong>ratrichoptera<br />

— Hymenoptera, Diptera, Aphaniptera<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1959). H. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне.<br />

П<strong>о</strong>д<strong>о</strong>тряды: Protomecoptera, Eumecoptera, Р<strong>а</strong>ratrichoptera.<br />

ПОДОТРЯД PROTOMECOPTERA<br />

В передних и з<strong>а</strong>дних <strong>к</strong>рыльях <strong>в</strong>ет<strong>в</strong>ей SC<br />

ше<strong>с</strong>ть и б<strong>о</strong>льше; СиА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м. Пермь —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне Амери<strong>к</strong>и и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии<br />

три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>; <strong>в</strong>ымершие <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Р1<strong>а</strong>tychoristidae;<br />

Kaltanidae, Cycloristidae, Cyclopteridae,<br />

Tomiochoristidae, Permomeropidae, Archipanorpidae,<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних — <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО PLATYCHORISTIDAE<br />

CARPENTER, 1930<br />

(Protomeropidae Tillyard, 1926)<br />

В переднем <strong>к</strong>рыле передний <strong>к</strong>р<strong>а</strong>й <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>в</strong>ершин<strong>а</strong> SC <strong>с</strong><strong>о</strong>единен<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й <strong>с</strong>ильн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й <strong>с</strong> R, пр<strong>о</strong>из<strong>в</strong><strong>о</strong>дит <strong>в</strong>печ<strong>а</strong>тление <strong>с</strong>ли<strong>в</strong>шей<strong>с</strong>я<br />

<strong>с</strong> R; птер<strong>о</strong><strong>с</strong>тигмы нет; <strong>в</strong>ет<strong>в</strong>ей SC 12 и б<strong>о</strong>льше;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей RS де<strong>в</strong>ять —<br />

д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>ть; RSi+г <strong>в</strong>д<strong>в</strong><strong>о</strong>е и б<strong>о</strong>лее чем <strong>в</strong>д<strong>в</strong><strong>о</strong>е<br />

длиннее RS3-H; <strong>в</strong>ет<strong>в</strong>ей М <strong>в</strong><strong>о</strong><strong>с</strong>емь—де<strong>в</strong>ять; ди<strong>с</strong>т<strong>а</strong>льные<br />

ч<strong>а</strong><strong>с</strong>ти <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны <strong>к</strong><strong>о</strong><strong>с</strong>ыми<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми, и <strong>в</strong>ершины<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>, ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong><strong>в</strong>язы<strong>в</strong><strong>а</strong>ющих их<br />

п<strong>о</strong>перечных, т<strong>о</strong>ньше, чт<strong>о</strong> д<strong>а</strong>ет <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong><br />

<strong>а</strong>н<strong>а</strong>льными жил<strong>к</strong><strong>а</strong>ми передних <strong>к</strong>рылье<strong>в</strong> ручейни<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>тлич<strong>а</strong>я<strong>с</strong>ь <strong>о</strong>т п<strong>о</strong><strong>с</strong>ледних <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием непере<strong>с</strong>еченн<strong>о</strong>г<strong>о</strong><br />

жил<strong>к</strong><strong>а</strong>ми шир<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5,6—<br />

10 мм (ри<strong>с</strong>. 883). Н. пермь Ур<strong>а</strong>л<strong>а</strong> и С. Амери<strong>к</strong>и.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Marimerobius G. Zalessky, 1946. Тип р<strong>о</strong>д<strong>а</strong><br />

— М. splendens G. Zalessky, 1946; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я незн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; RS 1 + 2<br />

б<strong>о</strong>лее чем <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее RS 3 + 4; Мп +2 <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

М 3 + 4, п<strong>о</strong><strong>с</strong>ледняя <strong>с</strong> пр<strong>о</strong><strong>с</strong>тым р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; <strong>в</strong>ершины<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> не п<strong>о</strong>х<strong>о</strong>жи н<strong>а</strong> п<strong>о</strong>перечные<br />

жил<strong>к</strong>и. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10 мм<br />

(ри<strong>с</strong>. 884). Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>.<br />

Вне СССР: Platychorista Tillyard, 1926.<br />

СЕМЕЙСТВО KALTANIDAE О. MARTYNOVA, 1958<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>в</strong>ершин<strong>а</strong> SC <strong>с</strong><strong>о</strong>единен<strong>а</strong> <strong>с</strong> R прям<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й, не п<strong>о</strong>х<strong>о</strong>жей н<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющую<strong>с</strong>я<br />

<strong>с</strong> R; птер<strong>о</strong><strong>с</strong>тигмы нет; <strong>в</strong>ет<strong>в</strong>ей SC ше<strong>с</strong>ть —<br />

284<br />

http://jurassic.ru/<br />

Си А А,<br />

sc R<br />

Ри<strong>с</strong>. 885—889. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Kaltanidae<br />

885. АН<strong>а</strong>/<strong>о</strong>р<strong>а</strong>п<strong>о</strong>гр<strong>а</strong> pilosa (О. Martynova;) переднее <strong>к</strong>рыл<strong>о</strong>, Х10,0,<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 886. Pinnachorista<br />

sarbalensis О. Martynova; {переднее <strong>к</strong>рыл<strong>о</strong>, Х10,0, н. пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,1958). 887. Megachorista khalfini.<br />

О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, хб.О, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 888. Cyclorista convexicosta О.Martynova;.<br />

переднее <strong>к</strong>рыл<strong>о</strong>, Х8,6, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1958). 889. Cycloptera autumnalis О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Х4,0, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958).<br />

д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>ть; RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong> или пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

ее; <strong>в</strong>ет<strong>в</strong>ей <strong>с</strong>емь — де<strong>с</strong>ять; RSi +2 немн<strong>о</strong>г<strong>о</strong> длиннее<br />

RS3+4, ч<strong>а</strong>ще <strong>о</strong>дн<strong>о</strong>й длины; <strong>в</strong>ет<strong>в</strong>ей М<br />

ше<strong>с</strong>ть — де<strong>в</strong>ять; ди<strong>с</strong>т<strong>а</strong>льные ч<strong>а</strong><strong>с</strong>ти <strong>а</strong>н<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong> не <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми; бу-

г<strong>о</strong>р<strong>к</strong>и н<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х е<strong>с</strong>ть или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Пермь. Три р<strong>о</strong>д<strong>а</strong>.<br />

Altajopanorpa О. Martynova, 1948 (Kaltana<br />

О. Martynova, 1958). Тип р<strong>о</strong>д<strong>а</strong>—A. kaltanica<br />

О. Martynova, 1948; н. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н). Вет<strong>в</strong>ей<br />

SC не б<strong>о</strong>лее де<strong>в</strong>яти, ч<strong>а</strong>ще ше<strong>с</strong>ть — <strong>с</strong>емь; <strong>в</strong>ет<strong>в</strong>ей<br />

М ше<strong>с</strong>ть, р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> Mi и Мг; Mi+г р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS, длиннее<br />

М3+4; пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и п<strong>о</strong><strong>к</strong>рыты буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 5,7—8 мм (ри<strong>с</strong>.<br />

885). Де<strong>с</strong>ять <strong>в</strong>ид<strong>о</strong><strong>в</strong> из н. перми и <strong>о</strong>дин из <strong>в</strong>.<br />

перми Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Pinnachorista О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. sarbalensis О. Martynova, 1958; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>а</strong>рб<strong>а</strong>л<strong>а</strong>). Вет<strong>в</strong>ей SC не менее де<strong>в</strong>яти; <strong>в</strong>ет<strong>в</strong>ей<br />

М б<strong>о</strong>льше ше<strong>с</strong>ти; буг<strong>о</strong>р<strong>к</strong>и н<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х<br />

е<strong>с</strong>ть у ф<strong>о</strong>рм из н. перми. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 5,25—19 мм (ри<strong>с</strong>. 886). В<strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

из н. перми и <strong>о</strong>дин из <strong>в</strong>. перми Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Megachorista О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. khalfini О. Martynova, 1958; <strong>в</strong>.<br />

лермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Кемер<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.). Вет<strong>в</strong>ей SC де<strong>в</strong>ять; <strong>в</strong>ет<strong>в</strong>ей<br />

М ше<strong>с</strong>ть, р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> Mi и М4; Mi+г р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS и <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

Мз+4; н<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong><br />

нет. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 13,0—13,5 мм<br />

(ри<strong>с</strong>. 887). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО CYCLORISTIDAE<br />

О. MARTYNOVA, 1958<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong> п<strong>о</strong>лт<strong>о</strong>р<strong>а</strong> р<strong>а</strong>з<strong>а</strong> шире<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> е<strong>с</strong>ть; <strong>в</strong>ет<strong>в</strong>ей SC<br />

<strong>с</strong>емь; R <strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей RS <strong>в</strong><strong>о</strong><strong>с</strong>емь;<br />

<strong>в</strong>ет<strong>в</strong>ей М ше<strong>с</strong>ть; н<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х буг<strong>о</strong>р<strong>к</strong>и.<br />

Пермь. Один р<strong>о</strong>д.<br />

Cyclorista О. Martynova, 1958. Тип р<strong>о</strong>д<strong>а</strong><br />

— С. convexicosta О. Martynova, 1958; н.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и Mi и Мз; CuA<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления М.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 7 мм (ри<strong>с</strong>. 888). Один<br />

<strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО CYCLOPTERIDAE<br />

О. MARTYNOVA, 1958<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле б<strong>о</strong>лее чем <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> е<strong>с</strong>ть; <strong>в</strong>ет<strong>в</strong>ей<br />

SC 12—13; R из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong>,<br />

ди<strong>с</strong>т<strong>а</strong>льнее же дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong>пеперед,<br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю<br />

285<br />

http://jurassic.ru/<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей RS де<strong>в</strong>ять; <strong>в</strong>ет<strong>в</strong>ей М <strong>в</strong><strong>о</strong><strong>с</strong>емь; н<strong>а</strong><br />

пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> нет. Пермь.<br />

Один р<strong>о</strong>д.<br />

Cycloptera О. Martynova, 1958. Тип р<strong>о</strong>д<strong>а</strong><br />

— С. autumnalis О. Martynova, 1958; <strong>в</strong>.<br />

пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong>р<strong>в</strong><strong>о</strong>). SC и R <strong>в</strong>ыгнуты <strong>в</strong>перед; М 3 пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

CuA и М4 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не;<br />

д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> <strong>с</strong>тупенч<strong>а</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 14,5—<br />

15 мм (ри<strong>с</strong>. 889). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО TOMIOCHORIST1DAE<br />

О. MARTYNOVA, 1958<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый;<br />

птер<strong>о</strong><strong>с</strong>тигмы нет; <strong>в</strong>ет<strong>в</strong>ей SC <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>в</strong>яти;<br />

<strong>в</strong>ет<strong>в</strong>ей RS четыре; <strong>в</strong>ет<strong>в</strong>ей М четыре —<br />

<strong>с</strong>емь; буг<strong>о</strong>р<strong>к</strong>и н<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льных жил<strong>к</strong><strong>а</strong>х е<strong>с</strong>ть. Н.<br />

пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Tomiochorista О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. nubila О. Martynova, 1958; н. пермь,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, К<strong>а</strong>лт<strong>а</strong>н).<br />

В передних <strong>к</strong>рыльях <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>ль-<br />

Ри<strong>с</strong>. 890—892. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tomiochoristidae, Permomeropidae<br />

890 Tomiochorista nubila О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 12,5,<br />

н пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 891. Kaltanochorista<br />

griasevi О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 7,5, н. пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 892. Permomerope australis<br />

Tillyard; переднее <strong>к</strong>рыл<strong>о</strong>, Хб,8, <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

(Tillyard, 1926)<br />

RS

н<strong>о</strong>е п<strong>о</strong>ля <strong>о</strong>дн<strong>о</strong>й ширины; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS немн<strong>о</strong>г<strong>о</strong><br />

длиннее RSi+г и RS3+4, или <strong>о</strong>ни р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине;<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния М5 нет; <strong>в</strong>ет<strong>в</strong>ей М<br />

б<strong>о</strong>льше четырех. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

4,47—5,10 мм (ри<strong>с</strong>. 890). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Kaltanochorista О. Martynova, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — К- grjasevi О. Martynova, 1958;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). В передних <strong>к</strong>рыльях <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; RS <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

RSi +2 и RS 3-h; RSi+г длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>;<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние М5 е<strong>с</strong>ть; <strong>в</strong>ет<strong>в</strong>ей М четыре.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 8 мм (ри<strong>с</strong>. 891). Один<br />

<strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО PERMOMEROPIDAE RIEK, 1953<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> п<strong>о</strong>чти<br />

прям<strong>о</strong>й; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, немн<strong>о</strong>г<strong>о</strong> длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины <strong>к</strong>рыл<strong>а</strong>, <strong>с</strong> <strong>с</strong>емью — <strong>в</strong><strong>о</strong><strong>с</strong>емью <strong>в</strong>ет<strong>в</strong>ями;<br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й птер<strong>о</strong><strong>с</strong>тигмы <strong>о</strong>гр<strong>а</strong>ничен R;<br />

<strong>в</strong>ет<strong>в</strong>ей RS де<strong>с</strong>ять; <strong>в</strong>ет<strong>в</strong>ей М <strong>в</strong><strong>о</strong><strong>с</strong>емь; Mi + 2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и <strong>в</strong>ет<strong>в</strong>ей RS и М <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>о</strong>й длины (ри<strong>с</strong>. 892). В. пермь<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Pertnomerope Tillyard,<br />

1926; Aphryganoneura Tillyard, 1926.<br />

ПОДОТРЯД EUMECOPTERA<br />

(Paramecoptera)<br />

В передних <strong>к</strong>рыльях <strong>в</strong>ет<strong>в</strong>ей SC не б<strong>о</strong>лее<br />

трех; R из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти (<strong>к</strong>р<strong>о</strong>ме Metropatridae); <strong>в</strong>ет<strong>в</strong>ей RS четыре—<br />

де<strong>с</strong>ять; <strong>в</strong>ет<strong>в</strong>ей М четыре — <strong>в</strong><strong>о</strong><strong>с</strong>емь; СиА<br />

без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>. Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Metropatridae, Permopanorpidae,<br />

Permochoristidae, Anormochoristidae,<br />

Tychtopsychidae, Permocentropidae, Orthophlebiidae,<br />

Neorthophlebiidae, Panorpidae, Bittacidae,<br />

Nannochoristidae, Choristidae, из них четыре<br />

п<strong>о</strong><strong>с</strong>ледних <strong>с</strong><strong>о</strong><strong>в</strong>ременные.<br />

Вне СССР: Metropatridae, Anormochoristidae,<br />

Nannochoristidae, Choristidae.<br />

СЕМЕЙСТВО METROPATRIDAE HANDLIRSCH, 1906<br />

Переднее <strong>к</strong>рыл<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>в</strong><strong>с</strong>ех <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> п<strong>о</strong>чти прямыми SC и<br />

R, шир<strong>о</strong><strong>к</strong>им р<strong>а</strong>ди<strong>а</strong>льным п<strong>о</strong>лем (шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых). Р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и<br />

н<strong>а</strong> RSi, RS2 и М 4; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong>; мн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных бе<strong>с</strong><strong>с</strong>и<strong>с</strong>темн<strong>о</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 8,9 мм (ри<strong>с</strong>. 893). Верхи н.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>. Один р<strong>о</strong>д Metropator Handlirsch, 1906<br />

из С. Амери<strong>к</strong>и.<br />

286<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PERMOPANORPIDAE TILLYARD, 1926<br />

Переднее <strong>к</strong>рыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, длинн<strong>о</strong>е, длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

ширину <strong>в</strong> 3,5 р<strong>а</strong>з<strong>а</strong> и б<strong>о</strong>лее; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

не длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины <strong>к</strong>рыл<strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

е<strong>с</strong>ть; н<strong>а</strong> R три — <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>в</strong>ет<strong>в</strong>ей; RS<br />

мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и бы<strong>в</strong><strong>а</strong>ют<br />

н<strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>в</strong>ет<strong>в</strong>ях; <strong>в</strong>ет<strong>в</strong>ей М ше<strong>с</strong>ть — <strong>с</strong>емь;<br />

М5 <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, длинн<strong>а</strong>я; п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> немн<strong>о</strong>г<strong>о</strong>.<br />

Пермь — три<strong>а</strong><strong>с</strong>. Три р<strong>о</strong>д<strong>а</strong>.<br />

Permopanorpa Tillyard, 1926. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. formosa Tillyard, 1926; н. пермь, С.<br />

Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). RS 1+2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong><br />

<strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>, длиннее RS3+4; р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и<br />

н<strong>а</strong> Mi и Мз ин<strong>о</strong>гд<strong>а</strong> д<strong>в</strong><strong>о</strong>йные; п<strong>о</strong>перечные<br />

жил<strong>к</strong>и <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле и между <strong>в</strong>ет<strong>в</strong>ями<br />

М. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 4—11 мм (ри<strong>с</strong>.<br />

894). Ше<strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и н. пермь С. Амери<strong>к</strong>и.<br />

Вне СССР: Neopermopanorpa Riek, 1955;<br />

Xenopanorpa Riek, 1955.<br />

СЕМЕЙСТВО PERMOCHORISTIDAE<br />

TILLYARD, 1917<br />

(Mesopanorpidae Tillyard, 1918;<br />

Belmontiidae Tillyard, 1919;<br />

Mesochoristidae Tillyard, 1925;<br />

Agetopanorpidae Carpenter, 1930;<br />

Idelopanorpidae M. Zalessky, 1929;<br />

Xenochoristidae Riek, 1953)<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

ег<strong>о</strong> ширину менее чем <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong>. SC длинн<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>;<br />

R без <strong>в</strong>ет<strong>в</strong>ей или <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью; <strong>в</strong>ет<strong>в</strong>ей<br />

RS четыре — де<strong>с</strong>ять; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные <strong>в</strong>ет<strong>в</strong>и бы<strong>в</strong><strong>а</strong>ют<br />

н<strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>в</strong>ет<strong>в</strong>ях RS; <strong>в</strong>ет<strong>в</strong>ей М <strong>о</strong>бычн<strong>о</strong> не б<strong>о</strong>лее<br />

ше<strong>с</strong>ти (и<strong>с</strong><strong>к</strong>лючение Sylvopanorpa); М5 <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я;<br />

п<strong>о</strong>перечные жил<strong>к</strong>и <strong>в</strong> умеренн<strong>о</strong>м <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е<br />

(ри<strong>с</strong>. 895). Пермь — юр<strong>а</strong>. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Protochoristinae и Permochoristinae.<br />

ПОДСЕМЕЙСТВО PROTOCHORISTINAE<br />

О. MARTYNOVA, SUBFAM. NOV.<br />

Вет<strong>в</strong>ей RS б<strong>о</strong>лее четырех; <strong>в</strong>ет<strong>в</strong>ей М ше<strong>с</strong>ть.<br />

Пермь. 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Sylvopanorpa Martynov, 1940. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. carpenteri Martynov, 1940; н. пермь,<br />

Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл- (<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong>).<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; три п<strong>о</strong>л<strong>о</strong>гие длинные <strong>в</strong>ет<strong>в</strong>и<br />

SC, н<strong>а</strong>чин<strong>а</strong>ющие<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>'ч<strong>а</strong>л<strong>а</strong> RS;<br />

длин<strong>а</strong> SC 2 и SC3 п<strong>о</strong>чти <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; <strong>в</strong>ет<strong>в</strong>ей RS де<strong>с</strong>ять; RSi+2<br />

и RS3+4 <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; <strong>к</strong>\ <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10,5 мм (ри<strong>с</strong>. 896)<br />

Один <strong>в</strong>ид. Н. пермь Ур<strong>а</strong>л<strong>а</strong>

Ри<strong>с</strong>. 893—897. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Metropatridae, Permochoristidae<br />

893. Metropator puslllus Handlirsch; переднее <strong>к</strong>рыл<strong>о</strong>, x7,0. нее <strong>к</strong>рыл<strong>о</strong>, x2,7, <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Tillyard, 1919). 896.<br />

н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Tillyard, 1926). 894. Permopanorpa angus- Sylvopanorpa carpenterl Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 6,0, н.<br />

tata Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X6,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940). 897. Agetochorista tillyardi Marty<strong>о</strong>бл.<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1948). 895. Belmontia mitsheli Tillyard; перед- nov (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); x5,3, н. пермь., Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940).<br />

Agetochorista. Martynov, 1933 (Neoageta<br />

Riek, 1953; Agetochoristella Riek, 1953). Тип<br />

р<strong>о</strong>д<strong>а</strong> — Agetochorista ornata Martynov, 1933;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong> 1,5—2 р<strong>а</strong>з<strong>а</strong> шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>, SC2<br />

п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>я, <strong>в</strong> 2—2,5 р<strong>а</strong>з<strong>а</strong> длиннее ширины <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; <strong>в</strong>ет<strong>в</strong>ей RS пять; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RS4, М 2 и М 4; RSi +2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>, длиннее<br />

RS3+4. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 4,2—<br />

14 мм (ри<strong>с</strong>. 897). 15 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>,<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл., Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Kamopanorpa Martynov, 1928. Тип<br />

р<strong>о</strong>д<strong>а</strong> — К. lata Martynov, 1928; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры).<br />

RSi+2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>,<br />

RS 3+4 — пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RS b RS 2, Mi и М 3. Длин<strong>а</strong><br />

передних <strong>к</strong>рылье<strong>в</strong> 4,8—5,5 мм (ри<strong>с</strong>. 898). Три<br />

<strong>в</strong>ид<strong>а</strong>. Н. пермь Ур<strong>а</strong>л<strong>а</strong>; <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл. и Приур<strong>а</strong>лья.<br />

Oochorista Martynov, 1933. Тип р<strong>о</strong>д<strong>а</strong> —<br />

О. gunderseni Martynov, 1933; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Пе­<br />

287<br />

http://jurassic.ru/<br />

реднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ля <strong>о</strong>дн<strong>о</strong>й ширины; SC 2 <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> длиннее<br />

ширины <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; RSi + 2 длиннее<br />

RS3+4, <strong>о</strong>бе р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong><br />

RS2, RS 4, Мг и М 4; темные п<strong>о</strong>перечные п<strong>о</strong>л<strong>о</strong><strong>с</strong>ы.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 12,2 мм (ри<strong>с</strong>. 899).<br />

Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Parachorista Tillyard, 1926 (Phipoides<br />

Riek, 1953). Тип р<strong>о</strong>д<strong>а</strong> — Parachorista pincombeae<br />

Tillyard, 1926; <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия. Переднее<br />

<strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уже <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

<strong>о</strong>дн<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь SC; R <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RSi,<br />

Mi, М 4; п<strong>о</strong>перечные п<strong>о</strong>л<strong>о</strong><strong>с</strong>ы. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 5—14 мм (ри<strong>с</strong>. 900). Ше<strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong>; <strong>в</strong>- пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Protopanorpoides О. Martynova, gen.<br />

nov. Тип р<strong>о</strong>д<strong>а</strong> — Protopanorpa elongata Martynov,<br />

1933; <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. SC<br />

из<strong>о</strong>гнут<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле д<strong>о</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> SC 3 уже<br />

<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>, <strong>в</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти SC2 — <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире;<br />

SC2 <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее н<strong>а</strong>именьшей ширины <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RS 2,

898. Kamopanorpa rotundipennis Martynov; <strong>о</strong>бщий <strong>в</strong>ид, X 6,3,<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933). 899. Oochorista<br />

gunderseni Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, Хб,0, <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933). 900. Parachorista uralensls Martynov;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х 11,0, н. пермь, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940)<br />

М 2 и М 3; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

RS. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 9 мм<br />

(ри<strong>с</strong>. 901). Один <strong>в</strong>ид. В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

Stigmorista О. Martynova, 1958. Тип.<br />

р<strong>о</strong>д<strong>а</strong> — 5. captiosa О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Суриё<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>). Переднее <strong>к</strong>рыл<strong>о</strong>. Передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля<br />

<strong>о</strong>дн<strong>о</strong>й ширины, уз<strong>к</strong>ие; SC2 и SC3 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

RSi+г зн<strong>а</strong>чительн<strong>о</strong> длиннее RS 3+4; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RS 4, М 2 и М 4- Выделяет<strong>с</strong>я <strong>с</strong>реди<br />

других р<strong>о</strong>д<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> н<strong>а</strong>личием <strong>к</strong>рупн<strong>о</strong>й<br />

ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идн<strong>о</strong>й птер<strong>о</strong><strong>с</strong>тигмы, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

пр<strong>о</strong>х<strong>о</strong>дит <strong>в</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле ближе <strong>к</strong><br />

RS]. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 7 мм (ри<strong>с</strong>. 902).<br />

Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Вне СССР: ProtochoristaTiWyard, 1926; Agetopanorpa<br />

Carpenter, 1930; Protopanorpa Tillyard,<br />

1926; Betmontia Tillyard, 1919; Parabelmontia<br />

Tillyard, 1922.<br />

ПОДСЕМЕЙСТВО PERMOCHOPJSTINAE<br />

O. MARTYNOVA, SUBFAM. NOV.<br />

Вет<strong>в</strong>ей RS четыре; <strong>в</strong>ет<strong>в</strong>ей M четыре —<br />

ше<strong>с</strong>ть. Пермь — юр<strong>а</strong>. Де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Petromantis Handlirsch, 1904 (Idelopanorpa<br />

M. Zalessky, 1929; Martynoviella Handlirsch,<br />

1937). Тип р<strong>о</strong>д<strong>а</strong> — P. rossica Handlirsch,<br />

1904); <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Тихие Г<strong>о</strong>ры). Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>ль-<br />

Ри<strong>с</strong>. 898—902. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Permochoristidae<br />

288<br />

http://jurassic.ru/<br />

901<br />

901. Protopanorpoides elongata (Martynov); переднее <strong>к</strong>рыл<strong>о</strong><br />

Х,6,8, <strong>в</strong>. пермь. Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933). 902. Stigmorista<br />

captiosa О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 10,0. <strong>в</strong>. Пермь.<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958)<br />

н<strong>о</strong>е п<strong>о</strong>ле ди<strong>с</strong>т<strong>а</strong>льнее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния SC 3 шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

SC2 и SC3 длинные; SC 2 п<strong>о</strong>л<strong>о</strong>г<strong>а</strong>я, <strong>в</strong> 3,5—4 р<strong>а</strong>з<strong>а</strong><br />

длиннее н<strong>а</strong>именьшей ширины <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>ля; RSi+г и RS34.4 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, р<strong>а</strong><strong>в</strong>ны или п<strong>о</strong>чти<br />

р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине и <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>их<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 4,5—<br />

12,5 мм (ри<strong>с</strong>. 903). 19 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>; <strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.,<br />

Приур<strong>а</strong>лья и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Asiachorista О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong>—A. neuburgae О. Martynova, 1958;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Переднее <strong>к</strong>рыл<strong>о</strong>. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния SC2 шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC 2 длинн<strong>а</strong>я,<br />

<strong>в</strong> 3—5 р<strong>а</strong>з длиннее н<strong>а</strong>именьшей ширины<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; SC 3 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, п<strong>о</strong>д<strong>о</strong>бн<strong>а</strong><br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>е; RSi +2<br />

и RS 3 + 4 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, р<strong>а</strong><strong>в</strong>ны или п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ны п<strong>о</strong><br />

длине и <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>их р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 5—10 мм (ри<strong>с</strong>. 904).<br />

Де<strong>в</strong>ять <strong>в</strong>ид<strong>о</strong><strong>в</strong> из н. перми и четыре <strong>в</strong>ид<strong>а</strong> из<br />

<strong>в</strong>. перми Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Callietheira О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — С. khalfini О. Martynova, 1958;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

К<strong>а</strong>лт<strong>а</strong>н). Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т Asiachorista б<strong>о</strong>лее<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ченн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й <strong>к</strong>рыл<strong>а</strong> и з<strong>а</strong>гнутыми<br />

<strong>в</strong>перед ди<strong>с</strong>т<strong>а</strong>льными <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ниями <strong>в</strong>ет<strong>в</strong>ей<br />

RS. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 7,5—9,5 мм<br />

(ри<strong>с</strong>. 905). Три <strong>в</strong>ид<strong>а</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Petrochorista Martynov, 1930 (Neopet-

903. Petromantis kaltanica О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 8,5; н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1961).<br />

904. Asiachorista neuburgae О. Martynova; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 6,7, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1958). 905. Callietheira khalfini О. Martynova; переднее<br />

<strong>к</strong>рыл<strong>о</strong>. х8,0, н. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958).<br />

906. Petrochorista elegantula Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х8,0,<br />

romantis Riek, 1953). Тип р<strong>о</strong>д<strong>а</strong> — P. elegantula<br />

Martynov, 1930; <strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР<br />

(<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Тихие Г<strong>о</strong>ры). Переднее <strong>к</strong>рыл<strong>о</strong>.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, не шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>;<br />

SC длинн<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ет<strong>в</strong>ями;<br />

н<strong>а</strong> R не б<strong>о</strong>лее <strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и; RSi +2 и RS 3+4<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине, примерн<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й длины<br />

<strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ими р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>ми и длиннее RS, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 3,6—10 мм (ри<strong>с</strong>. 906).<br />

18 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>;<br />

<strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл., Приур<strong>а</strong>лья, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Mesochorista Т i 11 у <strong>а</strong> г d, 1916 (Permochorista<br />

Tillyard, 1918; Caenoptllon G. Zalessky,<br />

1933; Prochorlstella Riek, 1953). Тип р<strong>о</strong>д<strong>а</strong> —<br />

Mesochorista proavita Tillyard, 1916; три<strong>а</strong><strong>с</strong>,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (К<strong>в</strong>ин<strong>с</strong>ленд). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е,<br />

не шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; SC длинн<strong>а</strong>я, <strong>в</strong>ет<strong>в</strong>и<br />

ее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; RSi +2 зн<strong>а</strong>чительн<strong>о</strong> длиннее<br />

RS3+4; RS3-H <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS и н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 3,3—14 мм (ри<strong>с</strong>. 907). 48 <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

19 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

РИС. 903—909. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Permochoristidae<br />

CuA<br />

-Си<br />

<strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1930). 907. Mesochorista oblonga<br />

О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 12,3, <strong>в</strong>. пермь, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 908. Lithopanorpa pusilla (Tillyard);<br />

переднее <strong>к</strong>рыл<strong>о</strong>, X 18,0, и. пермь, С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1930).<br />

909. Liassochorista asiatica О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 10,0<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1948).<br />

Н. пермь Ур<strong>а</strong>л<strong>а</strong> и Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>;<br />

<strong>в</strong>. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл., Приур<strong>а</strong>лья, Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии; три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Lithopanorpa Carpenter, 1930. Тип<br />

р<strong>о</strong>д<strong>а</strong> Protopanorpa pusilla Tillyard, 1926;<br />

н. пермь, С. Амери<strong>к</strong><strong>а</strong> (К<strong>а</strong>нз<strong>а</strong><strong>с</strong>). М<strong>а</strong>лень<strong>к</strong>ие,<br />

уз<strong>к</strong>ие <strong>к</strong>рылья; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>;<br />

н<strong>а</strong> R д<strong>в</strong>е — три <strong>в</strong>ет<strong>в</strong>и; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>; RS '<strong>к</strong><strong>о</strong>р<strong>о</strong>че RS3+4; RSi+ 2<br />

немн<strong>о</strong>г<strong>о</strong> длиннее RS 3+ 4 и зн<strong>а</strong>чительн<strong>о</strong> длиннее<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 4—5 мм (ри<strong>с</strong>. 908). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. пермь<br />

С. Амери<strong>к</strong>и; <strong>в</strong>. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Liassochorista Tillyard, 1933. Тип<br />

р<strong>о</strong>д<strong>а</strong> — L. anglicana Tillyard, 1933; н- лей<strong>а</strong><strong>с</strong>,<br />

Англия. От Mesochorista <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием<br />

д<strong>в</strong>ух — трех <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong> R при н<strong>а</strong>личии<br />

длинн<strong>о</strong>й SC. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 5—<br />

7,5 мм (ри<strong>с</strong>. 909). В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.; н. юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Вне СССР: Mesopanorpodes Tillyard, 1918.<br />

289<br />

http://jurassic.ru/

Ри<strong>с</strong>. 910—911. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tychtopsychidae, Permocentropidae<br />

910. Tychtopsyche beljanini O. Martynova: переднее <strong>к</strong>рыл<strong>о</strong>,<br />

XlO.O, <strong>в</strong>. пермь. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 911.<br />

Permocentropus philopotamoides Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 10,0, <strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1933)<br />

СЕМЕЙСТВО TYCHTOPSYCHIDAE<br />

О. MARTYNOVA, 1958<br />

Переднее <strong>к</strong>рыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

ширину <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong>; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние уз<strong>к</strong><strong>о</strong>е, п<strong>о</strong>чти <strong>в</strong><br />

3 р<strong>а</strong>з<strong>а</strong> уже <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; передний<br />

<strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый; <strong>в</strong>ершин<strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>а</strong>я; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; R <strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>;<br />

н<strong>а</strong> RS и М п<strong>о</strong> четыре <strong>в</strong>ет<strong>в</strong>и. В. пермь. Один<br />

р<strong>о</strong>д.<br />

Tychtopsyche О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. beljanini О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> II). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле д<strong>о</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

SC 2 уже <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>, ди<strong>с</strong>т<strong>а</strong>льнее SC2<br />

<strong>о</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>ширяет<strong>с</strong>я; SC 2 длиннее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й SC 3;<br />

RS <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong>я п<strong>о</strong> длине RS 3+4, RS ] + 2<br />

незн<strong>а</strong>чительн<strong>о</strong> длиннее RS 3 + 4; мн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 6,2 мм<br />

(ри<strong>с</strong>. 910). Один <strong>в</strong>ид. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

СЕМЕЙСТВО PERMOCENTROPIDAE<br />

MARTYNOV, 1933<br />

Передний <strong>к</strong>р<strong>а</strong>й переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый<br />

и из<strong>о</strong>гнутый, длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>в</strong> 2,8 р<strong>а</strong>з<strong>а</strong>; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong>, из<strong>о</strong>гнут<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю,<br />

<strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью; <strong>о</strong>дн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь н<strong>а</strong> R; RS<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>, длиннее<br />

RSi +2 и RS 3 + 4; <strong>в</strong>ет<strong>в</strong>ей RS четыре; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> Mi и М 2. В. пермь. Один |<br />

р<strong>о</strong>д. £*<br />

290<br />

http://jurassic.ru/<br />

Permocentropus Martynov, 1933. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. philopotamoides Martynov, 1933;<br />

<strong>в</strong>. пермь, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, С<strong>о</strong>ян<strong>а</strong>). RSi+г <strong>к</strong><strong>о</strong>р<strong>о</strong>че RS3+4, и <strong>о</strong>бе <strong>о</strong>ни<br />

п<strong>о</strong>чти <strong>в</strong>тр<strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>их р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 6,3 мм (ри<strong>с</strong>. 911). Один <strong>в</strong>ид.<br />

В. пермь Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО NANNOCHORISTIDAE<br />

TILLYARD, 1917<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Вет<strong>в</strong>ей RS три, RSi +2 не<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я; М и СиА н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении<br />

<strong>с</strong>литы. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>с</strong> пятью <strong>в</strong>ид<strong>а</strong>ми <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии- В. пермь А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Nannochoristella Riek, 1953 и<br />

Neochoristella Riek, 1953.<br />

СЕМЕЙСТВО ORTHOPHLEBIIDAE<br />

HANDLIRSCH, 1906<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей SC не б<strong>о</strong>лее <strong>о</strong>дн<strong>о</strong>й; R пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;<br />

н<strong>а</strong> RS <strong>о</strong>т пяти д<strong>о</strong> де<strong>в</strong>яти <strong>в</strong>ет<strong>в</strong>ей; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

<strong>в</strong>ет<strong>в</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены гребенч<strong>а</strong>т<strong>о</strong> н<strong>а</strong> RSi;<br />

RSi+г р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ния SC д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> н<strong>а</strong><br />

М 4; М5 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, п<strong>о</strong>чти прям<strong>а</strong>я. Э<strong>в</strong><strong>о</strong>люция<br />

Orthophlebiidae шл<strong>а</strong> <strong>в</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лении <strong>с</strong>ужения<br />

<strong>к</strong>рылье<strong>в</strong> и упр<strong>о</strong>щения их жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния. Вер<strong>о</strong>ятн<strong>о</strong>,<br />

<strong>о</strong>т эт<strong>о</strong>г<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> пр<strong>о</strong>из<strong>о</strong>шли <strong>с</strong><strong>о</strong><strong>в</strong>ременные<br />

Panorpidae. Три<strong>а</strong><strong>с</strong> — не<strong>о</strong>ген. Три р<strong>о</strong>д<strong>а</strong>.<br />

r sc<br />

Щ Ри<strong>с</strong>. 912—1914. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Orthophlebiidae<br />

912. Orthophlebia liassica (Mantell); переднее <strong>к</strong>рыл<strong>о</strong>, Х4,0, н. юр<strong>а</strong>,<br />

Англия (Tillyard, 1938). 913. Protorthophlebia latipennis Tillyard;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, х7,3, н. юр<strong>а</strong>, Англия (Tillyard, 1933). 914.<br />

Mesopanorpa hartungi Brauer, Redtenbacher, Ganglbauer; перед-<br />

, нее <strong>к</strong>рыл<strong>о</strong>, X 5,5, н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1927)

Orthophlebia Westwood, 1843. (Orthophlebioides<br />

Handlirsch, 1906; Orthophlebiites<br />

Handlirsch, 1939; Synorthophlebia Handlirsch,<br />

1939, pars; Dinopanorpa Cockerell, 1924; Neoparachorista<br />

Riek, 1955, pars). Тип р<strong>о</strong>д<strong>а</strong> —<br />

Panorpa liassica Mantell, 1844; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(н. лей<strong>а</strong><strong>с</strong>, Англия). RSi +2 и RS 3 +4 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; RSi мн<strong>о</strong>г<strong>о</strong><strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

<strong>в</strong>ет<strong>в</strong>ей <strong>в</strong><strong>с</strong>егд<strong>а</strong> б<strong>о</strong>лее трех. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 7—35 мм (ри<strong>с</strong>. 912). 29 <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии;<br />

юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>, Ср. Азии и 3- Е<strong>в</strong>р<strong>о</strong>пы; ми<strong>о</strong>цен<br />

В. Сибири (Кудя).<br />

Protorthophlebia Tillyard, 1933 (Choristopanorpa<br />

Riek, 1950; Neoparachorista Riek,<br />

1955, pars). Тип р<strong>о</strong>д<strong>а</strong> — P. latipennis Tillyard,<br />

1933; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (н. лей<strong>а</strong><strong>с</strong>, Англия).<br />

RSi+2 и RS3+4 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не; RSi <strong>с</strong> д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 7,5—12 мм (ри<strong>с</strong>. 913). Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии;<br />

н. юр<strong>а</strong> Англии.<br />

Mesopanorpa Handlirsch, 1906 (Synorthophlebia<br />

Handlirsch, 1939, pars; Neoparachorista<br />

Riek, 1955, pars). Тип р<strong>о</strong>д<strong>а</strong> — Panorpa<br />

hartungi Brauer, Redtenbacher, Ganglbauer,<br />

1889); н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (черемх<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, У<strong>с</strong>ть-Б<strong>а</strong>лей). RSi +2 длиннее RS3-1-4 <strong>в</strong><br />

2 р<strong>а</strong>з<strong>а</strong> и б<strong>о</strong>лее; <strong>в</strong>ет<strong>в</strong>ей RS три и б<strong>о</strong>льше. Длин<strong>а</strong><br />

передних <strong>к</strong>рылье<strong>в</strong> 7—31 мм (ри<strong>с</strong>. 914).<br />

23 <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии;<br />

юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>, Ср. Азии<br />

и В. Сибири.<br />

СЕМЕЙСТВО PANORPIDAE М. LEAY, 1821<br />

[nom. transl. Stephens, 1836 (ex Panorpina M. Leay, 1821]<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong>; <strong>о</strong>дн<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь SC; R без<br />

<strong>в</strong>ет<strong>в</strong>ей; <strong>о</strong>т пяти д<strong>о</strong> <strong>с</strong>еми <strong>в</strong>ет<strong>в</strong>ей RS; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

<strong>в</strong>ет<strong>в</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены гребенч<strong>а</strong>т<strong>о</strong> н<strong>а</strong> RSi;<br />

RS1+2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> ди<strong>с</strong>т<strong>а</strong>льным<br />

<strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>нием SC; М четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я,<br />

д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> нет; М5 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

прям<strong>а</strong>я (ри<strong>с</strong>. 915). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Пять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 150 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии; из них д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных 1<br />

.<br />

1<br />

П<strong>о</strong> мнению пр<strong>о</strong>ф. К<strong>а</strong>рпентер<strong>а</strong> р<strong>о</strong>ды Holcorpa<br />

Scudd., 1878 и Panorpodes McL., 1875 <strong>с</strong>ледует <strong>в</strong>ыделить<br />

<strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>.<br />

Ри<strong>с</strong>. 915—919. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Panorpidae, Neorthophlebiidae, Bittacidae<br />

915. Panorpa sibirica Esb.-P.; <strong>к</strong>рылья, <strong>с</strong><strong>о</strong><strong>в</strong>рем. 916. Neorthophtebia<br />

robusta Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 3,0, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1937). 917. Protobittacus liassicus TUIyard; А— переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Хб.О; Б — з<strong>а</strong>дняя н<strong>о</strong>г<strong>а</strong>, X 14,0; н. юр<strong>а</strong>, Англия (Tillyard,<br />

1933). 918. Probittacus avitus Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, х4 0,<br />

<strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1927). 919. Biitacus succlnus<br />

Carpenter; переднее <strong>к</strong>рыл<strong>о</strong>, Х4,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Carpenter, 1954)<br />

http://jurassic.ru/<br />

291 19*

СЕМЕЙСТВО NEORTHOPHLEBIIDAE<br />

HANDLIRSCH, 1921<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong>; SC у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>, без <strong>в</strong>ет<strong>в</strong>ей, <strong>о</strong>дн<strong>а</strong><br />

<strong>в</strong>ет<strong>в</strong>ь н<strong>а</strong> R; ди<strong>с</strong>т<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец R не <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>к</strong><strong>о</strong><strong>в</strong>ше<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыгиб<strong>а</strong>; четыре <strong>в</strong>ет<strong>в</strong>и RS;<br />

RSi + 2 <strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее RS 3 + 4; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

н<strong>а</strong> М 4; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е; <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и незигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны.<br />

Neorthophlebiidae я<strong>в</strong>ляют<strong>с</strong>я пред<strong>к</strong><strong>а</strong>ми<br />

Bittacidae, у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong>рылья б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>ны.<br />

Три<strong>а</strong><strong>с</strong> — юр<strong>а</strong>. Три р<strong>о</strong>д<strong>а</strong>-<br />

Neorthophlebia Handlirsch, 1908 (Меsobittacus<br />

Handlirsch, 1939). Тип р<strong>о</strong>д<strong>а</strong> —<br />

Orthophlebia megapolitana Geinitz, 1884;<br />

н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (<strong>в</strong>. лей<strong>а</strong><strong>с</strong>, Герм<strong>а</strong>ния). Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> хитинизир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>, не у<strong>в</strong>еличен<strong>а</strong><br />

<strong>в</strong> р<strong>а</strong>змере; М и СиА п<strong>о</strong><strong>с</strong>ле СЛИЯНИЯ р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS; SC не <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong><br />

6—32 мм (ри<strong>с</strong>. 916). 14 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.; н. юр<strong>а</strong> Ср. Азии и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Bittacopanorpa G. Zalessky, 1935. Тип<br />

р<strong>о</strong>д<strong>а</strong> — В. javorskii G. Zalessky, 1935; три<strong>а</strong><strong>с</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (м<strong>а</strong>льце<strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Б<strong>а</strong>бий<br />

К<strong>а</strong>мень)- Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т Neorthophlebia<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й, не д<strong>о</strong>х<strong>о</strong>дящей д<strong>о</strong> <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong> SC и р<strong>а</strong><strong>с</strong>х<strong>о</strong>ждением М и СиА ди<strong>с</strong>т<strong>а</strong>льнее<br />

н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

11 мм. Один <strong>в</strong>ид. Три<strong>а</strong><strong>с</strong> Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Protobittacus Tillyard, 1933 (Archebittacus<br />

Riek, 1955). Тип р<strong>о</strong>д<strong>а</strong> — P. liassicus Tillyard,<br />

1933; н. юр<strong>а</strong>, Англия. Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т<br />

Neorthophlebia упл<strong>о</strong>тненн<strong>о</strong>й, шир<strong>о</strong><strong>к</strong><strong>о</strong>й (д<strong>о</strong>х<strong>о</strong>дит<br />

д<strong>о</strong> RSi +2) птер<strong>о</strong><strong>с</strong>тигм<strong>о</strong>й, н<strong>а</strong>личием <strong>с</strong>ильн<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и rs-m и р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> М 4 без <strong>с</strong>тебель<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 15,7 мм<br />

(ри<strong>с</strong>. 917). Четыре <strong>в</strong>ид<strong>а</strong>. Три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии;<br />

н. юр<strong>а</strong> Англии.<br />

Вне СССР: Auxobittacus Bode, 1953<br />

(Protobittacus?); Haplobittacus Bode, 1953;<br />

Metaxibittacus Bode, 1953 (Neorthophlebia?);<br />

Pleobittacus Bode, 1953 (Neorthophlebia?); Polydicrobiltacus<br />

Bode, 1953.<br />

СЕМЕЙСТВО BITTACIDAE ENDERLEIN, 1910<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т Neorthophlebiidae н<strong>а</strong>личием<br />

<strong>в</strong> переднем <strong>к</strong>рыле <strong>к</strong><strong>о</strong><strong>в</strong>ше<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> изгиб<strong>а</strong><br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> R, <strong>с</strong><strong>о</strong>единенн<strong>о</strong>г<strong>о</strong> <strong>с</strong> RS <strong>о</strong>дн<strong>о</strong>й-д<strong>в</strong>умя<br />

п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми, зигз<strong>а</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зными<br />

пр<strong>о</strong>д<strong>о</strong>льными жил<strong>к</strong><strong>а</strong>ми <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong> и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием <strong>о</strong>бычн<strong>о</strong> д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чных<br />

<strong>в</strong>ет<strong>в</strong>ей М. Юр<strong>а</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я <strong>к</strong> ше<strong>с</strong>ти<br />

р<strong>о</strong>д<strong>а</strong>м; б<strong>о</strong>льш<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нен<strong>а</strong><br />

<strong>в</strong> Южн<strong>о</strong>м п<strong>о</strong>луш<strong>а</strong>рии; четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

292<br />

http://jurassic.ru/<br />

Probittacus Martynov, 1927. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. avitus Martynov, 1927; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у)- RS 3 + 4 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины RSi +2; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> н<strong>а</strong> М4; <strong>о</strong>дн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rsi+2 р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее птер<strong>о</strong><strong>с</strong>тигмы. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 16,5 мм (ри<strong>с</strong>. 918). Один <strong>в</strong>ид, В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> и <strong>в</strong> Е<strong>в</strong>р<strong>о</strong>пе (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Вне СССР: Protobittacus Tillyard, 1933;<br />

Paleobittacus Carpenter, 1928; Bittacus Latreille,<br />

1805 (ри<strong>с</strong>. 919).<br />

. ПОДОТРЯД PARATRICHOPTERA<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т других п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong><br />

прям<strong>о</strong>й <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии R, <strong>с</strong>ближенными<br />

СиА и СиР и н<strong>а</strong>личием т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дн<strong>о</strong>й-д<strong>в</strong>ух<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>к</strong>ругл<strong>а</strong>я, не <strong>в</strong>ытянут<strong>а</strong><br />

«<strong>к</strong>лю<strong>в</strong><strong>о</strong>м»; <strong>а</strong>нтенны не пре<strong>в</strong>ыш<strong>а</strong>ют длину<br />

<strong>к</strong>рылье<strong>в</strong>. Н<strong>о</strong>ги п<strong>о</strong>чти г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные; л<strong>а</strong>п<strong>к</strong>и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Вер<strong>о</strong>ятн<strong>о</strong>, из эт<strong>о</strong>г<strong>о</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> путем<br />

д<strong>а</strong>льнейшей <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

и реду<strong>к</strong>ции з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> р<strong>а</strong>з<strong>в</strong>или<strong>с</strong>ь д<strong>в</strong><strong>а</strong><br />

пр<strong>о</strong>гре<strong>с</strong><strong>с</strong>и<strong>в</strong>ных <strong>о</strong>тряд<strong>а</strong> — Diptera и Hymenoptera<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1959). В. пермь—юр<strong>а</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Permotipulidae, Pseudopolycentropidae,<br />

Pseudodipteridae, Choristopsychidae, Mesopsychidae,<br />

Permotanyderidae, Dobbertiniidae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО PERMOTIPULIDAE TILLYARD, 1937<br />

(Robinjohniidae О. Martynova, 1948)<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы. Г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные <strong>к</strong>рылья <strong>с</strong> уз»<br />

<strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем и уз<strong>к</strong>им <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ег<strong>о</strong> ширины; SC<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>ей<br />

RS три — четыре; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS; СиА и<br />

CuP <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближены; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong><br />

<strong>о</strong>дн<strong>а</strong> — д<strong>в</strong>е* В. пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Permotipula Tillyard, 1929. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. patricia Tillyard, 1929; <strong>в</strong>. пермь, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>. RSi +2 без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; <strong>а</strong>н<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 4,8—5 мм (ри<strong>с</strong>. 920).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> и<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

Вне СССР-.Robinjohnia О. Martynova, 1948.<br />

СЕМЕЙСТВО PSEUDOPOLYCENTROPODIDAE<br />

HANDLIRSCH, 1921<br />

З<strong>а</strong>дние <strong>к</strong>рылья зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних<br />

( 3<br />

Д их длины); ф<strong>о</strong>рм<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong><br />

треуг<strong>о</strong>льн<strong>а</strong>я; <strong>в</strong>ершин<strong>а</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>а</strong>я, н<strong>а</strong>иб<strong>о</strong>лее<br />

р<strong>а</strong><strong>с</strong>ширен<strong>о</strong> <strong>к</strong>рыл<strong>о</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ния М4 (tornus); жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рылье<strong>в</strong> <strong>о</strong>чень

920. Permotipula borealis О. Martynova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х10,0,<br />

<strong>в</strong>. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1961). 921 .Pseudopolycentropus<br />

latipennis Martynov, <strong>к</strong>рылья, X7,5, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1927). 922. Pseudodiptera gallica Laurentiaux<br />

et Grauvogel; <strong>к</strong>рылья, x5,7, три<strong>а</strong><strong>с</strong>, Фр<strong>а</strong>нция (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1959)<br />

923. Choristopsyche tenuinervis Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, Хб,3,<br />

н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>тын<strong>о</strong><strong>в</strong>, 1937). 924. Sogdapsyche elongata<br />

близ<strong>к</strong><strong>о</strong>; н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; SC <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти<br />

<strong>к</strong>рыл<strong>а</strong>; RSi+2 <strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее RS3+4", р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

н<strong>а</strong> М 2; RS и М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong>;<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> д<strong>в</strong>е. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Pseudopolycentropus Handlirsch, 1906.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Phryganidium perlaeformis Geinitz,<br />

1884; н. юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>- В з<strong>а</strong>дних <strong>к</strong>рыльях<br />

три <strong>в</strong>ет<strong>в</strong>и М. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 6—<br />

8 мм (ри<strong>с</strong>. 921). Четыре <strong>в</strong>ид<strong>а</strong>. Юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong><br />

и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PSEUDODIPTERIDAE<br />

О. MARTYNOVA, 1961<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны (<strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины передних) <strong>с</strong> измененным<br />

реду<strong>к</strong>цией жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; передние <strong>к</strong>рылья<br />

треуг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы; н<strong>а</strong>иб<strong>о</strong>льшее р<strong>а</strong><strong>с</strong>ширение<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>н-<br />

Ри<strong>с</strong>. 920—927. П<strong>о</strong>д<strong>о</strong>тряд Paratrichoptera<br />

293<br />

Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 3,6, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1937). 925. Ferganopsyche rotundata Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

X 6,5, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937). 926. Ptychopteropsis<br />

mirabills Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 4,0, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>рг<br />

тын<strong>о</strong><strong>в</strong>, 1937). 927. Тuranopsyche venosa Martynov; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

Х4,5, н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

ч<strong>а</strong>ния CuA; <strong>в</strong>ершин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> уз<strong>к</strong><strong>а</strong>я;<br />

SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>в</strong> третьей чет<strong>в</strong>ерти; RSi+г немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

RS 3+ 4; р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и RS и М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие (ри<strong>с</strong>. 922).<br />

Три<strong>а</strong><strong>с</strong> Фр<strong>а</strong>нции. Один р<strong>о</strong>д — Pseudodiptera<br />

Laurentiaux et Grauvogel, 1953<br />

1<br />

Еще <strong>о</strong>дин р<strong>о</strong>д эт<strong>о</strong>г<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Ijapsyche Kolosn. et<br />

О. Mart., 1961 из<strong>в</strong>е<strong>с</strong>тен из <strong>в</strong>. юры З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лья.<br />

СЕМЕЙСТВО CHORISTOPSYCH1DAE<br />

MARTYNOV, 1937<br />

З<strong>а</strong>дние <strong>к</strong>рылья неиз<strong>в</strong>е<strong>с</strong>тны; переднее <strong>к</strong>рыл<strong>о</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы, шир<strong>о</strong><strong>к</strong><strong>о</strong>е, длин<strong>а</strong> <strong>в</strong>д<strong>в</strong><strong>о</strong>е<br />

б<strong>о</strong>льше ширины; передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый; SC<br />

длинн<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти<br />

<strong>к</strong>рыл<strong>а</strong>, <strong>с</strong> д<strong>в</strong>умя длинными <strong>в</strong>ет<strong>в</strong>ями; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS шире <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

и <strong>о</strong>дн<strong>о</strong>й ширины <strong>с</strong> р<strong>а</strong>ди<strong>а</strong>льным;<br />

RSi +2 немн<strong>о</strong>г<strong>о</strong> длиннее RS 3+ 4; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее, чем RS; <strong>в</strong>ет<strong>в</strong>ей М<br />

http://jurassic.ru/

пять; р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и М зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>их<br />

<strong>с</strong>теблей. Н. юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Choristopsyche Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Ch. tenuinervis Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). RSi + 2 длиннее<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; RS3+4 немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> н<strong>а</strong> М2. Длин<strong>а</strong><br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 9,5 мм (ри<strong>с</strong>. 923). Один <strong>в</strong>ид.<br />

Н. юр<strong>а</strong> Ср. Азии.<br />

СЕМЕЙСТВО MESOPSYCHIDAE TILLYARD, 1917<br />

Крылья длинные и уз<strong>к</strong>ие; передний <strong>к</strong>р<strong>а</strong>й<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й; SC у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong>, н<strong>о</strong><br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>;<br />

RS1+2 зн<strong>а</strong>чительн<strong>о</strong> длиннее RS3+4 и <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; RS и М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong>;<br />

<strong>в</strong>ет<strong>в</strong>ей М четыре; Mi + 2 и М 3+ 4 <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>их<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>. В. три<strong>а</strong><strong>с</strong> — н. юр<strong>а</strong>. Четыре<br />

р<strong>о</strong>д<strong>а</strong>.<br />

Sogdopsyche Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — S. elongata Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н- лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины; RSi + 2<br />

<strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> и <strong>в</strong>тр<strong>о</strong>е длиннее<br />

RS 3+4; RSi+2 п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> R; М и<br />

RS 3 + 4 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не. Длин<strong>а</strong><br />

передних <strong>к</strong>рылье<strong>в</strong> 13—18 мм (ри<strong>с</strong>. 924). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Ср. Азии.<br />

Вне СССР.: Mesopsyche Tillyard, 1917 (Aristopsyche<br />

Tillyard, 1919); Neuropsyche Tillyard,<br />

1919; Triassopsyche Tillyard, 1917.<br />

Д<strong>в</strong>е п<strong>а</strong>ры п<strong>о</strong><strong>к</strong>рытых <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми <strong>к</strong>рылье<strong>в</strong>.<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью (у дре<strong>в</strong>них ф<strong>о</strong>рм б<strong>о</strong>льше<br />

<strong>в</strong>ет<strong>в</strong>ей); RS и М делят<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зуя<br />

п<strong>о</strong> д<strong>в</strong><strong>а</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> (у перм<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм ин<strong>о</strong>гд<strong>а</strong><br />

е<strong>с</strong>ть д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и); СиА <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м,<br />

CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; ди<strong>с</strong>т<strong>а</strong>льные <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ния CuP<br />

и Ai <strong>с</strong>ближены или <strong>с</strong>литы; А 2 <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

Аь А 3 — н<strong>а</strong> Аг, <strong>о</strong>бр<strong>а</strong>зуя з<strong>а</strong>м<strong>к</strong>нутые ячеи и<br />

п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле. З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong> <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льным<br />

<strong>в</strong>еер<strong>о</strong>м или реже без нег<strong>о</strong>; <strong>а</strong>н<strong>а</strong>льные<br />

жил<strong>к</strong>и <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>; п<strong>о</strong>перечных<br />

жил<strong>о</strong><strong>к</strong> немн<strong>о</strong>г<strong>о</strong>. В<strong>с</strong>егд<strong>а</strong> н<strong>а</strong> <strong>к</strong>рыльях е<strong>с</strong>ть д<strong>в</strong><strong>а</strong><br />

ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чных <strong>о</strong>рг<strong>а</strong>н<strong>а</strong> — <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

RS 3 +4 и <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти тириди<strong>а</strong>льн<strong>о</strong>й ячеи<br />

(tc, меди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле).<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я; р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т жующег<strong>о</strong><br />

тип<strong>а</strong>, не фун<strong>к</strong>ци<strong>о</strong>нирует; х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>и­<br />

ОТРЯД TRICHOPTERA. РУЧЕЙНИКИ<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

294<br />

http://jurassic.ru/<br />

PARATRICHOPTERA INCERTAE<br />

SEDIS<br />

Ferganopsyche Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — F. rotundata Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>.<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>в</strong>ыгнут; SC<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; R <strong>с</strong><br />

длинным р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м, <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти из<strong>о</strong>гнут<strong>а</strong>;<br />

RSi+г из<strong>о</strong>гнут<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> R, длиннее<br />

RS 3+4 и <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

RS; <strong>в</strong>ет<strong>в</strong>ей М четыре. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 7 мм (ри<strong>с</strong>. 925). Один <strong>в</strong>ид. Н. юр<strong>а</strong><br />

Ср. Азии.<br />

Ptychopteropsis Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. mirabilis Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>.<br />

Ф<strong>о</strong>рм<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, передний <strong>к</strong>р<strong>а</strong>й р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>лый; R немн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыгнут<strong>а</strong> <strong>в</strong>перед;<br />

RSi+г без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; RS 3 + 4 <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления<br />

RS н<strong>а</strong> четыре <strong>в</strong>ет<strong>в</strong>и. Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> И мм (ри<strong>с</strong>. 926). Н. юр<strong>а</strong> Ср. Азии.<br />

Turanopsyche Martynov, 1937. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. venosa Martynov, 1937; н. юр<strong>а</strong>,<br />

Ср. Азия (н. лей<strong>а</strong><strong>с</strong>, Шур<strong>а</strong>б II). З<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>.<br />

Передний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS; RSi+г п<strong>о</strong>чти<br />

<strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> и п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й<br />

длины <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м RS 3+ 4; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS, <strong>с</strong> четырьмя<br />

<strong>в</strong>ет<strong>в</strong>ями; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> между RS4 и<br />

Мь Длин<strong>а</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 12 мм (ри<strong>с</strong>. 927).<br />

Один <strong>в</strong>ид. Н. юр<strong>а</strong> Ср. Азии.<br />

ты челю<strong>с</strong>тные щупи<strong>к</strong>и и гип<strong>о</strong>ф<strong>а</strong>рин<strong>к</strong><strong>с</strong>, при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленный<br />

<strong>к</strong> <strong>с</strong>лизы<strong>в</strong><strong>а</strong>нию жид<strong>к</strong><strong>о</strong><strong>с</strong>ти; мн<strong>о</strong>г<strong>о</strong>члени<strong>с</strong>тые<br />

ните<strong>в</strong>идные <strong>а</strong>нтенны; б<strong>о</strong>льшие<br />

ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong>, у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> три<br />

гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. Переднегрудь уз<strong>к</strong><strong>а</strong>я, гу<strong>с</strong>т<strong>о</strong> п<strong>о</strong><strong>к</strong>рыт<strong>а</strong>я<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; <strong>с</strong>редне- и з<strong>а</strong>днегрудь б<strong>о</strong>льшие;<br />

н<strong>о</strong>ги бег<strong>а</strong>тельные, л<strong>а</strong>п<strong>к</strong>и пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, н<strong>а</strong><br />

г<strong>о</strong>ленях шп<strong>о</strong>ры.<br />

Личин<strong>к</strong>и <strong>к</strong><strong>а</strong>мп<strong>о</strong>дее<strong>в</strong>идные или эру<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

ше<strong>с</strong>тин<strong>о</strong>гие, без л<strong>о</strong>жн<strong>о</strong>н<strong>о</strong>же<strong>к</strong> н<strong>а</strong> <strong>с</strong>редних <strong>с</strong>егмент<strong>а</strong>х<br />

брюш<strong>к</strong><strong>а</strong>; п<strong>а</strong>р<strong>а</strong> <strong>с</strong>н<strong>а</strong>бженных <strong>к</strong>рюч<strong>к</strong><strong>а</strong>ми<br />

<strong>а</strong>н<strong>а</strong>льных н<strong>о</strong>же<strong>к</strong> н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>егменте брюш<strong>к</strong><strong>а</strong>;<br />

<strong>о</strong>бычн<strong>о</strong> жи<strong>в</strong>ут <strong>в</strong> перен<strong>о</strong><strong>с</strong>ных д<strong>о</strong>ми<strong>к</strong><strong>а</strong>х, п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енных<br />

из нитей шел<strong>к</strong><strong>о</strong><strong>о</strong>тделительн<strong>о</strong>й железы,<br />

<strong>с</strong> <strong>о</strong>б<strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong>ми из р<strong>а</strong><strong>с</strong>тительных или минер<strong>а</strong>льных<br />

ч<strong>а</strong><strong>с</strong>тиц; н<strong>а</strong><strong>с</strong>еляют те<strong>к</strong>учие и <strong>с</strong>т<strong>о</strong>ячие <strong>в</strong><strong>о</strong>д<strong>о</strong>емы<br />

р<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. Пит<strong>а</strong>ние р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е:<br />

р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>ть, детрит, реже хищниче<strong>с</strong>т<strong>в</strong><strong>о</strong>:

<strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> <strong>в</strong><strong>о</strong>дн<strong>а</strong>я. От группы ручейни<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

пит<strong>а</strong><strong>в</strong>ших<strong>с</strong>я, <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>, прибрежн<strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>тью,<br />

пр<strong>о</strong>из<strong>о</strong>шел <strong>о</strong>тряд Lepidoptera.<br />

Пермь — ныне. П<strong>о</strong>д<strong>о</strong>тряды: Permotrichoptera,<br />

Annulipalpia, Integripalpia.<br />

ПОДОТРЯД PERMOTRICHOPTERA.<br />

Д Р ЕВН ЕЩУ ПИКОВ Ы Е<br />

Переднее <strong>к</strong>рыл<strong>о</strong>- Н<strong>а</strong> RS и М п<strong>о</strong> ше<strong>с</strong>ти <strong>в</strong>ет<strong>в</strong>ей;<br />

ди<strong>с</strong>т<strong>а</strong>льные ч<strong>а</strong><strong>с</strong>ти CuP и Ai <strong>с</strong>ближены,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>, пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины ег<strong>о</strong> длины. Пермь. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Microptysmatidae.<br />

СЕМЕЙСТВО MICROPTYSMATIDAE<br />

О. MARTYNOVA, 1958<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. SC <strong>в</strong>ыгнут<strong>а</strong> <strong>в</strong>перед, п<strong>о</strong>чти<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> переднему <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong>ершин<strong>а</strong><br />

ее <strong>с</strong><strong>о</strong>единен<strong>а</strong> п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й <strong>с</strong> R; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RS b RS2, Mi и М 3; RS <strong>о</strong>тх<strong>о</strong>дит<br />

<strong>о</strong>т R п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>в</strong>д<strong>в</strong><strong>о</strong>е шире<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. Пермь. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Ри<strong>с</strong>. 928—929. П<strong>о</strong>д <strong>о</strong>тряд Permotrichoptera<br />

928. BAficroptysma sibiricum О. Martynova; переднее <strong>к</strong>рыл<strong>о</strong><br />

XI 1,0, н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958). 929.<br />

Microptysmodes uralicus О Martynova; переднее <strong>к</strong>рыл<strong>о</strong>, X 12,0<br />

<strong>в</strong>. пермь, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>, 1958)<br />

Microptysma О. Martynova, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. sibiricum О. Martynova, 1958;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

С<strong>а</strong>рб<strong>а</strong>л<strong>а</strong> II). Вершин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

туп<strong>а</strong>я; н<strong>а</strong>иб<strong>о</strong>льш<strong>а</strong>я ширин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> прих<strong>о</strong>дит<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередину ег<strong>о</strong> длины; М 4 <strong>с</strong>лит<strong>а</strong> <strong>с</strong> пе­<br />

295<br />

http://jurassic.ru/<br />

редней <strong>в</strong>ет<strong>в</strong>ью CuA, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние ее п<strong>о</strong>х<strong>о</strong>же н<strong>а</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong>ую п<strong>о</strong>перечную жил<strong>к</strong>у, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

ручейни<strong>к</strong><strong>о</strong><strong>в</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я и меди<strong>а</strong>льн<strong>а</strong>я ячеи<br />

з<strong>а</strong>м<strong>к</strong>нуты п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

4 мм (ри<strong>с</strong>. 928). Один <strong>в</strong>ид. Н. пермь Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Microptysmodes О. Martynova, 1958.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. uralicus О. Martynova, 1958;<br />

<strong>в</strong>. пермь, Т<strong>а</strong>т<strong>а</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я АССР (<strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>,<br />

Тихие Г<strong>о</strong>ры). Переднее <strong>к</strong>рыл<strong>о</strong> <strong>с</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>о</strong><strong>с</strong>тр<strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й; н<strong>а</strong>иб<strong>о</strong>льш<strong>а</strong>я ширин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> прих<strong>о</strong>дит<strong>с</strong>я<br />

зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины ег<strong>о</strong><br />

длины; М 4 не <strong>с</strong>лит<strong>а</strong> <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

CuA; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я и меди<strong>а</strong>льн<strong>а</strong>я ячеи <strong>о</strong>т<strong>к</strong>рытые.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5 мм (ри<strong>с</strong>. 929).<br />

Один <strong>в</strong>ид. В. пермь Приур<strong>а</strong>лья.<br />

ПОДОТРЯД ANNULIPALPIA.<br />

КОЛЬЧАТОЩУПИКОВЫЕ<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Обычн<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>уют <strong>в</strong><strong>с</strong>е<br />

пять <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>, т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у мел<strong>к</strong>их<br />

ф<strong>о</strong>рм <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong><br />

меньше (Agapetus, Hydroptilidae); пер<strong>в</strong>ый<br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong>тебельч<strong>а</strong>тый<br />

(т. е. RSi + 2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и, з<strong>а</strong>мы<strong>к</strong><strong>а</strong>ющей ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льную<br />

ячею), или ег<strong>о</strong> нет; <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з не <strong>в</strong>ыр<strong>а</strong>жен.<br />

З<strong>а</strong>дние <strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних, <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льный<br />

<strong>в</strong>еер <strong>о</strong>бычн<strong>о</strong> не<strong>в</strong>ели<strong>к</strong> (<strong>к</strong>р<strong>о</strong>ме Macronematinae).<br />

П<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> нижнечелю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>тый или з<strong>а</strong><strong>о</strong><strong>с</strong>тренный. Личин<strong>к</strong>и <strong>к</strong><strong>а</strong>мп<strong>о</strong>дее<strong>в</strong>идные,<br />

<strong>о</strong>бычн<strong>о</strong> не <strong>с</strong>тр<strong>о</strong>ят д<strong>о</strong>ми<strong>к</strong><strong>а</strong>, <strong>о</strong>бит<strong>а</strong>ют<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>а</strong>х. В не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

черт<strong>а</strong>х близ<strong>к</strong>и <strong>к</strong> <strong>о</strong>тряду Mecoptera.<br />

В. пермь — ныне. 13 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Cladochoristidae,<br />

Prorhyacophilidae, Prosepididontidae, Necrotaulidae,<br />

Rhyacophilidae, Glossosomatidae, Hydroptilidae,<br />

Philopotamidae, Stenopsychidae, Psychomyidae,<br />

Ecnomidae, Polycentropodidae,<br />

Hydropsychidae; пер<strong>в</strong>ые четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, <strong>в</strong>не<br />

СССР.<br />

СЕМЕЙСТВО CLADOCHORISTIDAE RIEK, 1953<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый.<br />

Вет<strong>в</strong>ей SC ше<strong>с</strong>ть — <strong>с</strong>емь; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чные<br />

<strong>в</strong>ет<strong>в</strong>и н<strong>а</strong> R; RS четырех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; RSi + 2 и RS3+4<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й длины; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>; М и CuA делят<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS; Mi + 2 <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; меди<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; <strong>о</strong>бщ<strong>а</strong>я<br />

<strong>в</strong>ершин<strong>а</strong> <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 12 мм. В- пермь и три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Д<strong>в</strong><strong>а</strong>

Ри<strong>с</strong>. 930—939. П<strong>с</strong><br />

930. Necrotaulius furcatus (Giebel); переднее <strong>к</strong>рыл<strong>о</strong>, X 7,5,<br />

н. три<strong>а</strong><strong>с</strong>, Англия (Tillyard, 1917). 931. Rhyacophila projusa UIглег;<br />

А — переднее <strong>к</strong>рыл<strong>о</strong>, Х10,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 10,0;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912), 932. Electragapetus<br />

scitulus Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>, X 10,0; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, X 10,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer,<br />

1912). 933. EUctroirichia subtilis Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X22.0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х22,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Ulmer, 1912). 934. Electracanthinus klebsi Ulmer,<br />

A —переднее <strong>к</strong>рыл<strong>о</strong>, X8,6; Б—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, х8,6; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912). 935. Stenopsyche<br />

м<strong>о</strong>н<strong>о</strong>типных р<strong>о</strong>д<strong>а</strong> {Ctadochorista Tillyard, 1926<br />

и Cladochoristella Riek, 1955).<br />

СЕМЕЙСТВО PRORHYACOPHILIDAE RIEK, 1955<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й.<br />

Вет<strong>в</strong>ей SC три; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чных <strong>в</strong>ет<strong>в</strong>ей н<strong>а</strong> R<br />

нет; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>ры­<br />

296<br />

1тряд Annulipalpia<br />

http://jurassic.ru/<br />

imitata Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>, ХЗ.О; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>;<br />

Х3,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912).<br />

936. Lype sericea (Pictet); A — переднее <strong>к</strong>рыл<strong>о</strong>, x 17,0; Б — з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, X 17,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) •(Ulmer,<br />

1912). 937. Ecnomus contlnentalis Ulmer; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

Х7,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.; А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Mosely, 1953). 938. Neureclipsis geniculata<br />

Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>, x 8,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

х8,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912).<br />

939. Potamyia nitida Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>, X 5,3; Б —<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х5,3; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Ulmer, 1912)<br />

л<strong>а</strong>; RSi +2 зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че длинн<strong>о</strong>й RS 3+4;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; Mi+г и М 3 +4<br />

зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>их р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>; меди<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; <strong>о</strong>бщ<strong>а</strong>я <strong>в</strong>ершин<strong>а</strong><br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5 мм.<br />

В. три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии. Один р<strong>о</strong>д.

СЕМЕЙСТВО PROSEPIDIDONTIDAE<br />

HANDLIRSCH, 1939<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Выпу<strong>к</strong>лый передний <strong>к</strong>р<strong>а</strong>й,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле; SC <strong>с</strong> д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; R без д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чных <strong>в</strong>ет<strong>в</strong>ей; RS<br />

трех<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; М и CuA р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS и <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>; М делит пл<strong>а</strong><strong>с</strong>тину <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> уз<strong>к</strong>ую переднюю<br />

и шир<strong>о</strong><strong>к</strong>ую з<strong>а</strong>днюю ч<strong>а</strong><strong>с</strong>ти; <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> Mi+2, меди<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я;<br />

<strong>о</strong>бщ<strong>а</strong>я <strong>в</strong>ершин<strong>а</strong> <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong><br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 10 мм.<br />

Один р<strong>о</strong>д. Н. юр<strong>а</strong> Герм<strong>а</strong>нии.<br />

СЕМЕЙСТВО NECROTAULIDAE<br />

HANDLIRSCH, 1906<br />

(Liassophilidae Tillyard, 1933)<br />

Обычн<strong>о</strong> пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют мел<strong>к</strong>ие ф<strong>о</strong>рмы <strong>с</strong> длин<strong>о</strong>й<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 3,5—6 мм, близ<strong>к</strong>и <strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>му<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у Rhyacophilidae, <strong>о</strong>тлич<strong>а</strong>я<strong>с</strong>ь<br />

<strong>о</strong>т нег<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>лением RS, М, CuA и <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>нием <strong>о</strong>бщей<br />

<strong>в</strong>ершины <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 3,5—9 мм, т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у Liassophila —<br />

21 мм (ри<strong>с</strong>. 930)- Три<strong>а</strong><strong>с</strong> и юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

11 р<strong>о</strong>д<strong>о</strong><strong>в</strong>: Epididontus Handlirsch, 1939; Liadotaulius<br />

Handlirsch, 1939; Mesotrichopteridium<br />

Handlirsch, 1906; Metarchitaulius Handlirsch,<br />

1939; Metatrichopteridium Handlirsch, 1939;<br />

Nercotaulius Handlirsch, 1906; Necrotauliodes<br />

Handlirsch, 1939; Parataulius Handlirsch, 1939;<br />

Palaeotaulius Handlirsch, 1906; Liassophila Tillyard,<br />

1933; Pseudoorthophlebla Handlirsch,<br />

1906.<br />

СЕМЕЙСТВО RHYACOPHILIDAE STEPHENS, 1836<br />

Средней и неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>еличины ф<strong>о</strong>рмы,<br />

реже <strong>к</strong>рупные. Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть передних<br />

<strong>к</strong>рылье<strong>в</strong> треуг<strong>о</strong>льн<strong>о</strong>-<strong>в</strong>ытянут<strong>а</strong>я; <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>т<strong>о</strong><strong>с</strong>ть<br />

<strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, б<strong>а</strong>хр<strong>о</strong>мы нет; dc <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я;<br />

М и RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не, п<strong>о</strong>чти<br />

н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong> или немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее,<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> Мз+4 (чет<strong>в</strong>ертый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный)<br />

<strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong><strong>о</strong> <strong>с</strong>теблем или д<strong>а</strong>же длиннее.<br />

З<strong>а</strong>дние <strong>к</strong>рылья немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних,<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>в</strong>еер<strong>а</strong> нет. Антенны длиннее<br />

<strong>к</strong>рылье<strong>в</strong>; д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых члени<strong>к</strong><strong>а</strong> челю<strong>с</strong>тных<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и т<strong>о</strong>л<strong>с</strong>тые, третий длиннее<br />

и т<strong>о</strong>лще их, пятый не<strong>к</strong><strong>о</strong>льч<strong>а</strong>тый, з<strong>а</strong><strong>о</strong><strong>с</strong>трен н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце. Шп<strong>о</strong>р 3.4.4. Личин<strong>к</strong>и <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные, жи<strong>в</strong>ут<br />

<strong>в</strong> те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>и .у<strong>к</strong>рыты <strong>в</strong> при<strong>к</strong>репленных<br />

<strong>к</strong> <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>ту д<strong>о</strong>ми<strong>к</strong><strong>а</strong>х-пещер<strong>к</strong><strong>а</strong>х,<br />

п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енных из пе<strong>с</strong>чин<strong>о</strong><strong>к</strong> и <strong>к</strong><strong>а</strong>меш<strong>к</strong><strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 931). Третичные — ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны — Rhyacophila Pictet, 1834 <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО GLOSSOSOMATIDAE<br />

WALLENGREN, 1891<br />

Ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я, меди<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бе ее <strong>в</strong>ет<strong>в</strong>и — я <strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не, р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и их <strong>с</strong>идячие, М делит<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее, чем RS; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М 3 + 4 (чет<strong>в</strong>ертый<br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льный) зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> <strong>с</strong>тебля.<br />

Личин<strong>к</strong>и жи<strong>в</strong>ут <strong>в</strong> перен<strong>о</strong><strong>с</strong>ных чехли<strong>к</strong><strong>а</strong>х.<br />

Шп<strong>о</strong>р 2.4.4^ Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 3—<br />

3,75 мм (ри<strong>с</strong>. 932). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО HYDROPTILIDAE<br />

STEPHENS, 1836<br />

Очень мел<strong>к</strong>ие ф<strong>о</strong>рмы: длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 2,6—3,7 мм; <strong>к</strong>рылья уз<strong>к</strong>ие, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные,<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>трены н<strong>а</strong> <strong>к</strong><strong>о</strong>нце, гу<strong>с</strong>т<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты длинными<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими б<strong>а</strong>хр<strong>о</strong>му, длин<strong>а</strong><br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях зн<strong>а</strong>чительн<strong>о</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ет их ширину; жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>идны,<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> длинный, у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

RS 2, теряя <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> RS 3 + 4; н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящей<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ячеи нет; меди<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления RS, ин<strong>о</strong>гд<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>; чет<strong>в</strong>ертый<br />

и пятый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Ф<strong>о</strong>рмул<strong>а</strong> шп<strong>о</strong>р р<strong>а</strong>зн<strong>а</strong>я. Личин<strong>к</strong>и <strong>в</strong> р<strong>а</strong>нних<br />

<strong>с</strong>т<strong>а</strong>диях без д<strong>о</strong>ми<strong>к</strong><strong>а</strong>; <strong>в</strong> п<strong>о</strong><strong>с</strong>ледней (пят<strong>о</strong>й) <strong>с</strong>тр<strong>о</strong>ят<br />

перен<strong>о</strong><strong>с</strong>н<strong>о</strong>й д<strong>о</strong>ми<strong>к</strong> из <strong>в</strong>ыделений шел<strong>к</strong><strong>о</strong><strong>о</strong>тделительн<strong>о</strong>й<br />

железы, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> <strong>о</strong>б<strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>о</strong>й из<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>р<strong>о</strong><strong>с</strong>лей или пе<strong>с</strong>чин<strong>о</strong><strong>к</strong>; б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>о</strong>бит<strong>а</strong>ет<br />

<strong>в</strong> те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>а</strong>х (ри<strong>с</strong>. 933). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне мн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

150 <strong>в</strong>ид<strong>о</strong><strong>в</strong>; три р<strong>о</strong>д<strong>а</strong> из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.<br />

СЕМЕЙСТВО PHILOPOTAMIDAE<br />

STEPHENS, 1836<br />

Передние <strong>к</strong>рылья <strong>в</strong>ытянутые, яйце<strong>в</strong>идные;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я и меди<strong>а</strong>льн<strong>а</strong>я ячеи з<strong>а</strong>м<strong>к</strong>нуты;<br />

тириди<strong>а</strong>льн<strong>а</strong>я ячея <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>че р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

CuA, р<strong>а</strong><strong>в</strong>н<strong>а</strong> или <strong>к</strong><strong>о</strong>р<strong>о</strong>че меди<strong>а</strong>льн<strong>о</strong>й ячеи. RS<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я примерн<strong>о</strong> н<strong>а</strong> <strong>с</strong>ередине длины<br />

<strong>к</strong>рыл<strong>а</strong>, М — пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

<strong>с</strong>идячий или <strong>с</strong>тебельч<strong>а</strong>тый; CuP и <strong>о</strong>бщ<strong>а</strong>я <strong>в</strong>ершин<strong>а</strong><br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong>дн<strong>о</strong>й<br />

т<strong>о</strong>ч<strong>к</strong>е, п<strong>о</strong>эт<strong>о</strong>му р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>яние между <strong>в</strong>ершин<strong>а</strong>ми<br />

з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и CuA и CuP б<strong>о</strong>льшее, чем между<br />

другими жил<strong>к</strong><strong>а</strong>ми; п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шире<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. Антенны <strong>с</strong>ильные, н<strong>о</strong> не длинные,<br />

297<br />

http://jurassic.ru/

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими члени<strong>к</strong><strong>а</strong>ми; гл<strong>а</strong>з<strong>к</strong>и е<strong>с</strong>ть; пер<strong>в</strong>ый<br />

члени<strong>к</strong> нижнечелю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий,<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>й длиннее, <strong>с</strong>н<strong>а</strong>бжен щетин<strong>к</strong><strong>а</strong>ми; чет<strong>в</strong>ертый<br />

р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине <strong>в</strong>т<strong>о</strong>р<strong>о</strong>му, третий длиннее<br />

их; пятый <strong>к</strong><strong>о</strong>льч<strong>а</strong>тый, р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине третьему<br />

и чет<strong>в</strong>ерт<strong>о</strong>му <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятым. Шп<strong>о</strong>р 2.4.4. Личин<strong>к</strong>и<br />

<strong>с</strong>тр<strong>о</strong>ят длинные труб<strong>к</strong>и, <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> г<strong>о</strong>рных<br />

п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>а</strong>х или ручьях <strong>с</strong> х<strong>о</strong>л<strong>о</strong>дн<strong>о</strong>й <strong>в</strong><strong>о</strong>д<strong>о</strong>й<br />

(ри<strong>с</strong>- 934). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В<strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; три р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и С. Амери<strong>к</strong>и, из<br />

них <strong>о</strong>дин <strong>в</strong>ымерший.<br />

СЕМЕЙСТВО STENOPSYCHIDAE MARTYNOV, 1924<br />

Крупные ф<strong>о</strong>рмы: длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong><br />

20 мм и б<strong>о</strong>льше; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>я,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS;<br />

пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> (RSi+г) <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

третьег<strong>о</strong> (RS 3 +4); меди<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>,<br />

длиннее ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й, <strong>к</strong><strong>о</strong>р<strong>о</strong>че тириди<strong>а</strong>льн<strong>о</strong>й<br />

ячеи; п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е и длинн<strong>о</strong>е;<br />

ширин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>а</strong>з пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>н<strong>а</strong>льные ячеи<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. З<strong>а</strong>дние <strong>к</strong>рылья <strong>с</strong> <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льным <strong>в</strong>еер<strong>о</strong>м.<br />

Антенны зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>к</strong>рылье<strong>в</strong>,<br />

пильч<strong>а</strong>тые; пятый члени<strong>к</strong> челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>тый, длиннее <strong>в</strong><strong>с</strong>ех <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ме<strong>с</strong>те<br />

<strong>в</strong>зятых. Шп<strong>о</strong>р 3.4.5. Личин<strong>к</strong>и <strong>с</strong> <strong>о</strong>чень длинн<strong>о</strong>й<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, жи<strong>в</strong>ут <strong>в</strong> г<strong>о</strong>рных п<strong>о</strong>т<strong>о</strong><strong>к</strong><strong>а</strong>х <strong>с</strong> г<strong>а</strong>лечным<br />

дн<strong>о</strong>м, <strong>с</strong>тр<strong>о</strong>ят пр<strong>о</strong><strong>с</strong>тые д<strong>о</strong>ми<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде длинных<br />

труб<strong>о</strong><strong>к</strong>; ми<strong>к</strong>р<strong>о</strong>ф<strong>а</strong>ги (ри<strong>с</strong>. 935). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

<strong>о</strong>дин <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре.<br />

СЕМЕЙСТВО PSYCHOMYIDAE KOLENATI, 1859<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы, длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong><br />

3—8 мм. Переднее <strong>к</strong>рыл<strong>о</strong>. Вет<strong>в</strong>ей н<strong>а</strong> R нет; переднег<strong>о</strong><br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> нет (RSi +2 пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я);<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея п<strong>о</strong>чти <strong>в</strong><br />

4 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>. RS з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>, <strong>в</strong>д<strong>в</strong><strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

меди<strong>а</strong>льн<strong>о</strong>й ячеи; тириди<strong>а</strong>льн<strong>а</strong>я ячея р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

меди<strong>а</strong>льн<strong>о</strong>й, ч<strong>а</strong>ще <strong>к</strong><strong>о</strong>р<strong>о</strong>че ее, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> или <strong>в</strong> ме<strong>с</strong>те ее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong>;<br />

Ms немн<strong>о</strong>г<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

СиА; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и шир<strong>о</strong><strong>к</strong>ий; п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, уже <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че и уже передних. Антенны<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>к</strong>рылье<strong>в</strong>; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong><br />

длиннее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>г<strong>о</strong> пер<strong>в</strong><strong>о</strong>г<strong>о</strong>, пятый —<br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>тый, р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине третьему и чет<strong>в</strong>ерт<strong>о</strong>му<br />

<strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятым. Шп<strong>о</strong>р 2-4.4. Личин<strong>к</strong>и ми<strong>к</strong>р<strong>о</strong>ф<strong>а</strong>ги,<br />

жи<strong>в</strong>ут <strong>в</strong> р<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х<br />

(ри<strong>с</strong>. 936). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>о</strong>дин из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.<br />

298<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ECNOMIDAE ULMER, 1903<br />

[nom. transl. Lepneva, 1956 (ex Ecnominae Ulmer, 1903)]<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы. Передние <strong>к</strong>рылья. SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

н<strong>а</strong> R <strong>в</strong>ет<strong>в</strong>ь; пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

е<strong>с</strong>ть (RSi+г р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я), <strong>с</strong>тебельч<strong>а</strong>тый;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины<br />

<strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS, з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>, <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

меди<strong>а</strong>льн<strong>о</strong>й ячеи; тириди<strong>а</strong>льн<strong>а</strong>я ячея п<strong>о</strong>чти<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине меди<strong>а</strong>льн<strong>о</strong>й, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> или <strong>в</strong> ме<strong>с</strong>те ее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong>;<br />

Мб длинн<strong>а</strong>я, зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния СиА; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий;<br />

п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е. Пер<strong>в</strong>ый и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й<br />

члени<strong>к</strong>и челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> р<strong>а</strong><strong>в</strong>ны п<strong>о</strong><br />

длине и немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че третьег<strong>о</strong>, пятый —<br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>тый, р<strong>а</strong><strong>в</strong>ен п<strong>о</strong> длине трем предыдущим<br />

<strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятым. Шп<strong>о</strong>р 3.4.4. Личин<strong>к</strong>и хищные,<br />

жи<strong>в</strong>ут <strong>в</strong> труб<strong>к</strong><strong>а</strong>х примити<strong>в</strong>н<strong>о</strong>г<strong>о</strong> п<strong>с</strong>их<strong>о</strong>миидн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong> (ри<strong>с</strong>. 937). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы( б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО POLYCENTROPODIDAE ULMER, 1903<br />

[nom. transl. Ulmer, 1906 (ex Polycentropinae, 1903)]<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы: длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong><br />

3—8 мм. Переднее <strong>к</strong>рыл<strong>о</strong>. R без <strong>в</strong>ет<strong>в</strong>ей; RSi +2<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>тебельч<strong>а</strong>тым р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м или пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея р<strong>а</strong><strong>в</strong>н<strong>а</strong>я п<strong>о</strong> длине <strong>с</strong>т<strong>в</strong><strong>о</strong>лу RS<br />

и меди<strong>а</strong>льн<strong>о</strong>й ячеи; тириди<strong>а</strong>льн<strong>а</strong>я ячея длиннее<br />

меди<strong>а</strong>льн<strong>о</strong>й и ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ячей, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> ме<strong>с</strong>те н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> меди<strong>а</strong>льн<strong>о</strong>й ячеи; Мб<br />

зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

СиА; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> СиА ДЛИННЫЙ И шир<strong>о</strong><strong>к</strong>ий; п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong><strong>с</strong>ширены <strong>в</strong><br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>че или немн<strong>о</strong>г<strong>о</strong><br />

длиннее <strong>к</strong>рылье<strong>в</strong>; д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых члени<strong>к</strong><strong>а</strong><br />

челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, пятый — 1<br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>тый, длиннее <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых.<br />

Шп<strong>о</strong>р 3.4.4. Личин<strong>к</strong>и жи<strong>в</strong>ут <strong>в</strong> те<strong>к</strong>учих, реже <strong>в</strong><br />

<strong>с</strong>т<strong>о</strong>ячих <strong>в</strong><strong>о</strong>д<strong>а</strong>х; хищни<strong>к</strong>и, <strong>с</strong>тр<strong>о</strong>ят л<strong>о</strong><strong>в</strong>чие <strong>с</strong>ети<br />

(ри<strong>с</strong>. 938). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

<strong>с</strong>емь (из них д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong>) из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО HYDROPSYCHIDAE CURTIS, 1835<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 6—10 мм; передние<br />

<strong>к</strong>рылья <strong>в</strong>ытянуты, <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>ны п<strong>о</strong> з<strong>а</strong>днему<br />

<strong>к</strong>р<strong>а</strong>ю <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти; SC длинн<strong>а</strong>я; R без<br />

<strong>в</strong>ет<strong>в</strong>ей; имеют<strong>с</strong>я <strong>в</strong><strong>с</strong>е пять <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

из них пер<strong>в</strong>ый, третий, пятый п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>с</strong>тебельч<strong>а</strong>тые; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>в</strong> 2—3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS, з<strong>а</strong>м<strong>к</strong>нут<strong>а</strong>я; меди<strong>а</strong>льн<strong>а</strong>я<br />

ячея немн<strong>о</strong>г<strong>о</strong> длиннее ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>ль-

н<strong>о</strong>й, з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>; тириди<strong>а</strong>льн<strong>а</strong>я ячея длинн<strong>а</strong>я, п<strong>о</strong>чти<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине <strong>с</strong>т<strong>в</strong><strong>о</strong>лу RS, ин<strong>о</strong>гд<strong>а</strong> длиннее,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> меди<strong>а</strong>льн<strong>о</strong>й<br />

ячеи; п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле немн<strong>о</strong>г<strong>о</strong> шире <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>. З<strong>а</strong>дние <strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че и шире передних,<br />

<strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть немн<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>; чет<strong>в</strong>ертый<br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Ан- '<br />

тенны т<strong>о</strong>н<strong>к</strong>ие, ин<strong>о</strong>гд<strong>а</strong> длиннее <strong>к</strong>рылье<strong>в</strong>; челю<strong>с</strong>тные<br />

щупи<strong>к</strong>и: пер<strong>в</strong>ый члени<strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й<br />

длинный, третий и чет<strong>в</strong>ертый <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> и<br />

р<strong>а</strong><strong>с</strong>ширены <strong>с</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, пятый р<strong>а</strong><strong>в</strong>ен<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льным <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятым, <strong>к</strong><strong>о</strong>льч<strong>а</strong>тый. Шп<strong>о</strong>р<br />

2.4.4. Личин<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, <strong>в</strong><br />

т<strong>о</strong>м чи<strong>с</strong>ле <strong>в</strong> б<strong>о</strong>льших м<strong>о</strong>щных речных <strong>а</strong>ртериях,<br />

<strong>с</strong>тр<strong>о</strong>ят неб<strong>о</strong>льшие д<strong>о</strong>ми<strong>к</strong>и-<strong>к</strong><strong>а</strong>меры <strong>с</strong> пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й<br />

ячеи<strong>с</strong>т<strong>о</strong><strong>с</strong>тью, у<strong>к</strong>репленные н<strong>а</strong> т<strong>в</strong>ерд<strong>о</strong>м<br />

<strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>те (ри<strong>с</strong>- 939). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>, <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong>дн<strong>о</strong> из<br />

них — Hydropsychinae —• из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), где пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong><br />

четырьмя р<strong>о</strong>д<strong>а</strong>ми (<strong>о</strong>дин <strong>в</strong>ымерший).<br />

ПОДОТРЯД INTEGRIPALPIA<br />

ЦЕЛЬНОЩУПИКОВЫЕ<br />

Переднее <strong>к</strong>рыл<strong>о</strong>. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует чет<strong>в</strong>ертый<br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>, т. е. М 3 +4 не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я;<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют и другие р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и;<br />

пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong>идячий,<br />

и<strong>с</strong><strong>к</strong>лючение <strong>в</strong><strong>с</strong>треч<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е<br />

Leptoceridae; <strong>а</strong>н<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>з х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>в</strong><strong>с</strong>егд<strong>а</strong> з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>. З<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних; <strong>а</strong>н<strong>о</strong>юг<strong>а</strong>льный <strong>в</strong>еер<br />

е<strong>с</strong>ть. П<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong><br />

целый, не з<strong>а</strong><strong>о</strong><strong>с</strong>трен. Личин<strong>к</strong>и эру<strong>к</strong><strong>о</strong><strong>в</strong>идные или<br />

<strong>с</strong>убэру<strong>к</strong><strong>о</strong><strong>в</strong>идные, <strong>о</strong>бит<strong>а</strong>ют преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong><br />

<strong>с</strong>т<strong>о</strong>ячих и медленн<strong>о</strong> те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х. Близ<strong>к</strong>и<br />

<strong>к</strong> <strong>о</strong>тряду Lepidoptera. Мел — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Kalophryganeidae, Phryganeidae, Limnophilidae,<br />

Leptoceridae, Calamoceridae, Odontoceratidae,<br />

Molannidae, Sericostomatidae: пер<strong>в</strong><strong>о</strong>е,<br />

<strong>в</strong>ымершее, из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО PHRYGANEIDAE BURMEISTER, 1839<br />

Крупные: длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> д<strong>о</strong><br />

30 мм; передние <strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> шир<strong>о</strong><strong>к</strong>ие, и<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льные; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, шире<br />

п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong>; ди<strong>с</strong>т<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец R из<strong>о</strong>гнут;<br />

у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует чет<strong>в</strong>ертый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея длинн<strong>а</strong>я, <strong>о</strong>дн<strong>о</strong>й<br />

длины <strong>с</strong><strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м RS или длиннее; пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> <strong>о</strong>тх<strong>о</strong>дит п<strong>о</strong>чти <strong>о</strong>т <strong>с</strong>ередины<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ячеи, т. е. RSi + 2 р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

зн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления<br />

RS34.4; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й и третий <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и<br />

е<strong>с</strong>ть; <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют п<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>в</strong><strong>о</strong>лни<strong>с</strong>т<strong>о</strong>й линией. Антенны т<strong>о</strong>л<strong>с</strong>тые,<br />

<strong>о</strong>дн<strong>о</strong>й длины или <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>к</strong>рылье<strong>в</strong>; гл<strong>а</strong>з<strong>к</strong>и<br />

е<strong>с</strong>ть; члени<strong>к</strong><strong>о</strong><strong>в</strong> нижнечелю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> у<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong> четыре, у <strong>с</strong><strong>а</strong>м<strong>к</strong>и пять. Шп<strong>о</strong>р 2.4.4. Мел —<br />

ныне- В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м<br />

<strong>в</strong> Се<strong>в</strong>ерн<strong>о</strong>м п<strong>о</strong>луш<strong>а</strong>рии; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>й<br />

ф<strong>а</strong>уне.<br />

Phryganea Linnaeus, 1740. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Ph. grandis Linnaeus, 1740; <strong>с</strong><strong>о</strong><strong>в</strong>ременные, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

Передние <strong>к</strong>рылья п<strong>а</strong>р<strong>а</strong>б<strong>о</strong>личе<strong>с</strong><strong>к</strong>и <strong>с</strong>рез<strong>а</strong>ны<br />

<strong>с</strong>з<strong>а</strong>ди; R из<strong>о</strong>гнут<strong>а</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>ль-<br />

Ри<strong>с</strong>. 940—944. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Phryganeidae, Limnophilidae<br />

Phryganea tavrushini Cockerell; переднее <strong>к</strong>рыл<strong>о</strong>, <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>, х 3,5, dontetla, не<strong>о</strong>ген, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (<strong>о</strong>риг. ри<strong>с</strong>.). 943. Limnophilidae?<br />

не<strong>о</strong>ген, Прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>.). 941. Limnophilus kaspievi д<strong>о</strong>ми<strong>к</strong> личин<strong>к</strong>и из р<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ин Ostracooa; <strong>в</strong>. юр<strong>а</strong>. З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (Reis,<br />

(О. Martynova); переднее <strong>к</strong>рыл<strong>о</strong>, Х5,3, не<strong>о</strong>ген, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong> (<strong>о</strong>риг. 1909). 944. Indusia sequoia Cockerell; д<strong>о</strong>ми<strong>к</strong> личин<strong>к</strong>и нз х<strong>в</strong><strong>о</strong>и<br />

ри<strong>с</strong>.). 952. Limnophilus sp.; д<strong>о</strong>ми<strong>к</strong> личин<strong>к</strong>и из р<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ин Spanlo- Sequoia lansdorffi, X 3,0, не<strong>о</strong>ген. Прим<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

299<br />

http://jurassic.ru/<br />

942<br />

944

н<strong>о</strong>й ячеи; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>. М — п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея длинн<strong>а</strong>я и уз<strong>к</strong><strong>а</strong>я, <strong>в</strong> з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Личин<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong><br />

<strong>с</strong>т<strong>о</strong>ячих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, <strong>с</strong>тр<strong>о</strong>ят цилиндриче<strong>с</strong><strong>к</strong>ие д<strong>о</strong>ми<strong>к</strong>и<br />

из <strong>с</strong>пир<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных мел<strong>к</strong>их р<strong>а</strong><strong>с</strong>тительных<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong><br />

9—22 мм (ри<strong>с</strong>. 940). Мел — ныне. 11 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и<br />

не<strong>о</strong>гене К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> и В. Сибири. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные д<strong>о</strong>ми<strong>к</strong>и личин<strong>о</strong><strong>к</strong> из мел<strong>а</strong> Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии<br />

и третичных <strong>о</strong>тл<strong>о</strong>жений Е<strong>в</strong>р<strong>о</strong>пы и<br />

Азии.<br />

СЕМЕЙСТВО LIMNOPHILIDAE KOLENATI, 1848<br />

[nom. transl. Kolenati, 1859<br />

(ex Limnophiloidea Kolenati, 1848)]<br />

Средних и <strong>к</strong>рупных р<strong>а</strong>змер<strong>о</strong><strong>в</strong>. Передние<br />

<strong>к</strong>рылья <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>рез<strong>а</strong>ны п<strong>о</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й<br />

<strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я б<strong>о</strong>лее чем<br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льный<br />

<strong>к</strong><strong>о</strong>нец R из<strong>о</strong>гнут; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины, М — мн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

п<strong>о</strong> длине <strong>с</strong>т<strong>в</strong><strong>о</strong>лу RS или незн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

или длиннее ег<strong>о</strong> и мн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че тириди<strong>а</strong>льн<strong>о</strong>й<br />

ячеи; пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

<strong>с</strong>идячий, чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong> нет; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

гпз-cua <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я. Антенны п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ны длине<br />

передних <strong>к</strong>рылье<strong>в</strong>; гл<strong>а</strong>з<strong>к</strong>и е<strong>с</strong>ть; члени<strong>к</strong><strong>о</strong><strong>в</strong> нижнечелю<strong>с</strong>тных<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong> у <strong>с</strong><strong>а</strong>мц<strong>а</strong> три, у <strong>с</strong><strong>а</strong>м<strong>к</strong>и —<br />

четыре. Шп<strong>о</strong>р 0.3.4, 1.3.4. Личин<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong><br />

р<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong> <strong>с</strong>т<strong>о</strong>ячих<br />

<strong>в</strong><strong>о</strong>д<strong>а</strong>х, <strong>с</strong>тр<strong>о</strong>ят р<strong>а</strong>зличн<strong>о</strong>й ф<strong>о</strong>рмы д<strong>о</strong>ми<strong>к</strong>и из<br />

р<strong>а</strong><strong>с</strong>тительных ч<strong>а</strong><strong>с</strong>тиц или из детрит<strong>а</strong>, т<strong>а</strong><strong>к</strong>же из<br />

пе<strong>с</strong>чин<strong>о</strong><strong>к</strong>, мел<strong>к</strong>их р<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ин<strong>о</strong><strong>к</strong>. Не<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне ше<strong>с</strong>ть п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>с</strong><strong>в</strong>ыше<br />

680 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong>е<strong>в</strong>ерн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>луш<strong>а</strong>рия, б<strong>о</strong>лее 30<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> Ю. Амери<strong>к</strong>е и <strong>о</strong>дин <strong>в</strong>ид <strong>в</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии;<br />

лишь Limnophilinae из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

ПОДСЕМЕЙСТВО LIMNOPHILINAE ULMER, 1903<br />

SC <strong>в</strong> передних <strong>к</strong>рыльях <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет прям<strong>о</strong> <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льный<br />

<strong>к</strong>р<strong>а</strong>й, <strong>а</strong> не <strong>в</strong> п<strong>о</strong>перечную жил<strong>к</strong>у <strong>с</strong>-г,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> у <strong>в</strong>ид<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Apataniinae. Шп<strong>о</strong>р<br />

1.3.4, 0.3.4. (ри<strong>с</strong>. 941—944). Не<strong>о</strong>ген —ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; пять р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии, из них три <strong>в</strong>ымершие.<br />

Крылья Limnophilinae, <strong>о</strong>тн<strong>о</strong><strong>с</strong>имые <strong>к</strong> трем<br />

р<strong>о</strong>д<strong>а</strong>м, из<strong>в</strong>е<strong>с</strong>тны из ми<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>,<br />

В. Сибири и С. Амери<strong>к</strong>и. Д<strong>о</strong>ми<strong>к</strong>и личин<strong>о</strong><strong>к</strong>,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>имых <strong>к</strong> трем р<strong>о</strong>д<strong>а</strong>м, из<strong>в</strong>е<strong>с</strong>тны<br />

из ми<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы, С. К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong> (Pectinariopsis<br />

Andrusov, 1916), В. Сибири и С. Амери<strong>к</strong>и.<br />

300<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО LEPTOCERIDAE LEACH, 1815<br />

Передние <strong>к</strong>рылья длинные и уз<strong>к</strong>ие, длин<strong>а</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ет ширину <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong>; ди<strong>с</strong>т<strong>а</strong>льный <strong>к</strong><strong>о</strong>нец<br />

R прям<strong>о</strong>й; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее М;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong> RS;<br />

меди<strong>а</strong>льн<strong>о</strong>й ячеи нет; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я и тириди<strong>а</strong>льн<strong>а</strong>я<br />

ячеи <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м<br />

ур<strong>о</strong><strong>в</strong>не; <strong>в</strong><strong>с</strong>егд<strong>а</strong> при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют пер<strong>в</strong>ый <strong>с</strong>тебельч<strong>а</strong>тый<br />

и пятый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

и третьег<strong>о</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> нет; п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> cua-cup прям<strong>а</strong>я. Антенны <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие,<br />

<strong>в</strong> 2—3 р<strong>а</strong>з<strong>а</strong> длиннее <strong>к</strong>рылье<strong>в</strong>; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> нижнечелю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> пять, <strong>о</strong>ни<br />

л<strong>о</strong>хм<strong>а</strong>тые, п<strong>о</strong><strong>к</strong>рыты длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. Личин<strong>к</strong>и<br />

<strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> <strong>с</strong>т<strong>о</strong>ячих и те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х,<br />

<strong>с</strong>тр<strong>о</strong>ят д<strong>о</strong>ми<strong>к</strong>и из пе<strong>с</strong>чин<strong>о</strong><strong>к</strong>, реже из детрит<strong>а</strong> и<br />

р<strong>а</strong><strong>с</strong>тительных ч<strong>а</strong><strong>с</strong>тиц. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Triplectidinae,<br />

Leptocerinae.<br />

ПОДСЕМЕЙСТВО TRIPLECTIDINAE ULMER, 1906<br />

В з<strong>а</strong>дних <strong>к</strong>рыльях ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>;<br />

при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и:<br />

пер<strong>в</strong>ый, третий и пятый- Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 8—12 мм (ри<strong>с</strong>. 945). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО LEPROCERINAE ULMER, 1903<br />

В з<strong>а</strong>дних <strong>к</strong>рыльях ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>;<br />

при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют <strong>а</strong>пи<strong>к</strong><strong>а</strong>льные р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и:<br />

пер<strong>в</strong>ый и пятый. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 4—<br />

7 мм (ри<strong>с</strong>. 946). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CALAMOCERATIDAE ULMER, 1905<br />

[nom. transl. Ulmer, 1906 (ex Calamoceratinae Ulmer, 1905)}<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

10—13 мм. В передних <strong>к</strong>рыльях ди<strong>с</strong>т<strong>а</strong>льный<br />

<strong>к</strong><strong>о</strong>нец R из<strong>о</strong>гнут, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> RSi, не<br />

д<strong>о</strong>х<strong>о</strong>дя д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления М; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея<br />

п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й длины <strong>с</strong> RS; тириди<strong>а</strong>льн<strong>а</strong>я ячея<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее меди<strong>а</strong>льн<strong>о</strong>й; <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> пять, чет<strong>в</strong>ертый ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong>тебельч<strong>а</strong>тый. Антенны п<strong>о</strong>чти <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее<br />

<strong>к</strong>рылье<strong>в</strong>; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет; члени<strong>к</strong><strong>о</strong><strong>в</strong> нижнечелю<strong>с</strong>тны'х<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong> пять, <strong>с</strong>ильн<strong>о</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые.<br />

Шп<strong>о</strong>р 2.4.4. Личин<strong>к</strong>и жи<strong>в</strong>ут <strong>в</strong> <strong>с</strong>т<strong>о</strong>ячих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х;<br />

д<strong>о</strong>ми<strong>к</strong>и <strong>с</strong>тр<strong>о</strong>ят из <strong>к</strong>рупных р<strong>а</strong><strong>с</strong>тительных<br />

ч<strong>а</strong><strong>с</strong>тиц (ри<strong>с</strong>. 947). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне Азии и Амери<strong>к</strong>и, из<br />

них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).

945. Triplectides rudis Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>, X 5,0; Б —<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 5,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Ulmer, 1912). 946. Eroiesis aequalis Ulmer; А— переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 9,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 9,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Ulmer, 1912). 947. Ganonema regulare Ulmer; A — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X4; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х4; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Ulmer, 1912). 948. Electrocerum pedestre Ulmer;<br />

А—переднее <strong>к</strong>рыл<strong>о</strong>, X5,0; Б—з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х5.0; п<strong>а</strong>ле<strong>о</strong>ген,'Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912). 949. Molanodes dubia Ulmer;<br />

A — переднее <strong>к</strong>рыл<strong>о</strong>, X4,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 4.0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer,1912). 950. Molanna angustata<br />

Ulmer; A — переднее <strong>к</strong>рыл<strong>о</strong>, X7,7; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>.<br />

СЕМЕЙСТВО ODONTOCERIDAE<br />

WALLENGREN, 1891<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

7—12 мм. Переднее <strong>к</strong>рыл<strong>о</strong>. R прям<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> RSi, не д<strong>о</strong>х<strong>о</strong>дя д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong><br />

Ри<strong>с</strong>. 945—955. П<strong>о</strong>д<strong>о</strong>тряд Integripalpia<br />

301<br />

http://jurassic.ru/<br />

X 7,7; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912).<br />

951. Molanna angustata Ulmer; д<strong>о</strong>ми<strong>к</strong> личин<strong>к</strong>и, X 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (П<strong>а</strong><strong>в</strong>л<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий, Лепне<strong>в</strong><strong>а</strong>, 1948). 952. Goera gracilicornis<br />

Ulmer; переднее <strong>к</strong>рыл<strong>о</strong>, Х8,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Ulmer, 1912). 953. Brachycentrus lablalls Hagen;.4 —<br />

переднее <strong>к</strong>рыл<strong>о</strong>.х 10,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 10,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912).<br />

954. Archaeocrunoecla tenuicornis Ulmer; A —переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X12.0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, Х12.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Ulmer, 1912). 955. Hellcopsyche typica Ulmer; А — переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 12,0; Б — з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 12,0; п<strong>а</strong>ле<strong>о</strong>ген?<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Ulmer, 1912)<br />

или <strong>в</strong>ершины; SC, R RSi <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны п<strong>о</strong>перечными<br />

жил<strong>к</strong><strong>а</strong>ми; RS и М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляют<strong>с</strong>я п<strong>о</strong>чти н<strong>а</strong><br />

<strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея длиннее<br />

RS; тириди<strong>а</strong>льн<strong>а</strong>я ячея <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> и б<strong>о</strong>лее длин-

нее ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ячеи; меди<strong>а</strong>льн<strong>о</strong>й ячеи нет;<br />

пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> <strong>с</strong>идячий, чет<strong>в</strong>ертый<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>. Антенны 1<br />

длиннее<br />

<strong>к</strong>рылье<strong>в</strong> п<strong>о</strong>чти <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong>, пильч<strong>а</strong>тые; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong><br />

нет; нижнечелю<strong>с</strong>тные щупи<strong>к</strong>и длинные, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>с</strong>ильн<strong>о</strong><strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые. Шп<strong>о</strong>р 2.4.4<br />

(ри<strong>с</strong>. 948). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 25 <strong>в</strong>ид<strong>о</strong><strong>в</strong>); три р<strong>о</strong>д<strong>а</strong><br />

из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО MOLANNIDAE<br />

WALLENGREN, 1891<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

7—14 мм; уз<strong>к</strong>ие, з<strong>а</strong><strong>к</strong>ругленные <strong>в</strong> <strong>в</strong>ершине,<br />

<strong>в</strong> п<strong>о</strong><strong>к</strong><strong>о</strong>е <strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> труб<strong>о</strong>ч<strong>к</strong>у; SC и R<br />

прямые, п<strong>а</strong>р<strong>а</strong>ллельные; <strong>в</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

из<strong>в</strong>р<strong>а</strong>щен<strong>о</strong> у р<strong>а</strong>зных р<strong>о</strong>д<strong>о</strong><strong>в</strong> и <strong>в</strong>ид<strong>о</strong><strong>в</strong> п<strong>о</strong>р<strong>а</strong>зн<strong>о</strong>му;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й и меди<strong>а</strong>льн<strong>о</strong>й ячей<br />

нет; з<strong>а</strong>дние <strong>к</strong>рылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че и шире передних.<br />

Антенны <strong>к</strong>реп<strong>к</strong>ие, <strong>о</strong>дн<strong>о</strong>й длины или <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>к</strong>рылье<strong>в</strong>; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет; члени<strong>к</strong><strong>о</strong><strong>в</strong> нижнечелю<strong>с</strong>тных<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong> пять, <strong>о</strong>ни <strong>с</strong>ильн<strong>о</strong><strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тые.<br />

Шп<strong>о</strong>р 2.4.4. Личин<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> <strong>о</strong>зер<strong>а</strong>х или<br />

э<strong>с</strong>ту<strong>а</strong>риях ре<strong>к</strong>; д<strong>о</strong>ми<strong>к</strong>и из пе<strong>с</strong>чин<strong>о</strong><strong>к</strong>, имеют<br />

ф<strong>о</strong>рму щит<strong>к</strong><strong>а</strong>, <strong>с</strong> труб<strong>к</strong><strong>о</strong>й н<strong>а</strong> нижней п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

(ри<strong>с</strong>. 949—951). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>, С. Амери<strong>к</strong><strong>а</strong>,<br />

Яп<strong>о</strong>ния и Индия); <strong>о</strong>б<strong>а</strong> р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь; им<strong>а</strong>г<strong>о</strong>)<br />

и ми<strong>о</strong>цен<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong><strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти СССР (д<strong>о</strong>ми<strong>к</strong><br />

личин<strong>к</strong>и)..<br />

СЕМЕЙСТВО SERICOSTOMATIDAE<br />

STEPHENS, 1836<br />

Крылья <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> шир<strong>о</strong><strong>к</strong>ие и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие:<br />

длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину <strong>в</strong> 2,5—3 р<strong>а</strong>з<strong>а</strong>; п<strong>о</strong><strong>к</strong>рыты<br />

гу<strong>с</strong>тыми <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong>ми<br />

и чешуй<strong>к</strong><strong>а</strong>ми. Переднее <strong>к</strong>рыл<strong>о</strong>. R п<strong>о</strong>чти<br />

прям<strong>а</strong>я; <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> С; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

немн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее, <strong>а</strong> М — немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине RS; пер<strong>в</strong>ый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> <strong>с</strong>идячий, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т <strong>с</strong>ередины<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ячеи; чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> нет; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>с</strong>и<strong>а</strong> 2<strong>с</strong>ир<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я. Антенны п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong>й<br />

длины" <strong>с</strong> <strong>к</strong>рыльями, у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния гу<strong>с</strong>т<strong>о</strong> п<strong>о</strong><strong>к</strong>рыты<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми; гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> нет; челю<strong>с</strong>тные щупи<strong>к</strong>и<br />

у<strong>с</strong>тр<strong>о</strong>ены р<strong>а</strong>зличн<strong>о</strong>. Личин<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>о</strong>бычн<strong>о</strong><br />

<strong>в</strong> бы<strong>с</strong>тр<strong>о</strong>те<strong>к</strong>учих <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, реже <strong>с</strong>т<strong>о</strong>ячих. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Goerinae,<br />

Brachycentrinae, Lepidostomatinae, Sericostomatinae;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

не из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

302<br />

• ПОДСЕМЕЙСТВО GOERINAE ULMER, 1903<br />

В передних и з<strong>а</strong>дних <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> е<strong>с</strong>ть<br />

четыре <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>: 1,2,3,5; <strong>в</strong> з<strong>а</strong>дних<br />

<strong>к</strong>рыльях ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>. Антенны<br />

не длиннее переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, б<strong>а</strong>з<strong>а</strong>льный<br />

члени<strong>к</strong> <strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы, <strong>с</strong>ильн<strong>о</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>т;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> у <strong>с</strong><strong>а</strong>мц<strong>а</strong><br />

три, у <strong>с</strong><strong>а</strong>м<strong>к</strong>и — пять, п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> длинный,<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тый, з<strong>а</strong>гнут <strong>к</strong><strong>в</strong>ерху.-Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 5—9 мм (ри<strong>с</strong>. 952). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО BRACHYCENTRINAE<br />

ULMER, 1903<br />

В передних <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>а</strong> четыре <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>: 1,2,3,5, <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях д<strong>в</strong><strong>а</strong>;<br />

<strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> щупи<strong>к</strong><strong>о</strong><strong>в</strong> у <strong>с</strong><strong>а</strong>мц<strong>а</strong> три, <strong>о</strong>ни<br />

з<strong>а</strong>гнуты <strong>в</strong><strong>в</strong>ерх. Шп<strong>о</strong>р 2.3.3. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 6—9 мм (ри<strong>с</strong>. 953). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пьГ (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО LEPIDOSTOMATINAE<br />

ULMER, 1903<br />

В передних <strong>к</strong>рыльях четыре <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>: 1,2,3,5, <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях три: 1,2,5;:<br />

<strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я.<br />

Антенны <strong>о</strong>дн<strong>о</strong>й длины 1<br />

<strong>с</strong> передними <strong>к</strong>рыльями;<br />

пер<strong>в</strong>ый члени<strong>к</strong> длинный, ин<strong>о</strong>гд<strong>а</strong> длиннеег<strong>о</strong>л<strong>о</strong><strong>в</strong>ы,<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>тый, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> чешуй<strong>к</strong><strong>а</strong>ми; члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong> у <strong>с</strong><strong>а</strong>мц<strong>а</strong> три. Шп<strong>о</strong>р<br />

2.4.4. Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> 4,5—8 мм<br />

(ри<strong>с</strong>. 954). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть <strong>в</strong>ымерших:<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО HELICOPSYCHIDAE ULMER, 1912<br />

'[nom. transl. Kimmins, 1952<br />

(ex Helicopsychinae Ulmer, 1912)]<br />

В передних и з<strong>а</strong>дних <strong>к</strong>рыльях п<strong>о</strong> три <strong>а</strong>пи<strong>к</strong><strong>а</strong>льных<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>: 1, 4, 5; ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> з<strong>а</strong>дних<br />

<strong>к</strong>рыльях <strong>в</strong>ып<strong>а</strong>д<strong>а</strong>ет чет<strong>в</strong>ертый <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я. Антенны<br />

не длиннее <strong>к</strong>рылье<strong>в</strong>; б<strong>а</strong>з<strong>а</strong>льный члени<strong>к</strong> р<strong>а</strong><strong>в</strong>ен<br />

длине г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы; члени<strong>к</strong><strong>о</strong><strong>в</strong> челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong><br />

три—• четыре. Шп<strong>о</strong>р 2.2.4. Длин<strong>а</strong> передних<br />

<strong>к</strong>рылье<strong>в</strong> 4—6 мм (ри<strong>с</strong>. 955). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Три р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь), <strong>о</strong>дин из них <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> еще <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> ручейни<strong>к</strong><strong>о</strong><strong>в</strong><br />

нея<strong>с</strong>н<strong>о</strong>г<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жения.<br />

http://jurassic.ru/

ОТРЯД LEPIDOPTERA. БАБОЧКИ, ИЛИ ЧЕШУЕКРЫЛЫЕ<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных р<strong>а</strong>змер<strong>о</strong><strong>в</strong>, <strong>с</strong><br />

д<strong>в</strong>умя п<strong>а</strong>р<strong>а</strong>ми х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итых <strong>к</strong>рылье<strong>в</strong>, п<strong>о</strong><strong>к</strong>рытых<br />

<strong>о</strong><strong>к</strong>р<strong>а</strong>шенны'ми чешуй<strong>к</strong><strong>а</strong>ми (<strong>в</strong>ид<strong>о</strong>измененными<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми); SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><br />

<strong>о</strong>дн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью; <strong>в</strong>ет<strong>в</strong>ей RS три — четыре; у <strong>а</strong>рх<strong>а</strong>ичных<br />

ф<strong>о</strong>рм RS делит<strong>с</strong>я дих<strong>о</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>и, у <strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

ф<strong>о</strong>рм <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>тх<strong>о</strong>дят <strong>о</strong>т<br />

<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; три — четыре <strong>в</strong>ет<strong>в</strong>и М (у три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> пять); ч<strong>а</strong><strong>с</strong>т<strong>о</strong> М и Си <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>литы 1<br />

, <strong>о</strong>бр<strong>а</strong>зуя <strong>к</strong>рупную ячею; CuA<br />

<strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; CuP пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong><br />

<strong>о</strong>дн<strong>а</strong> — три; А2 <strong>о</strong>бр<strong>а</strong>зует <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую петлю, <strong>в</strong><br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь <strong>с</strong> Аь А3 редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong><br />

д<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong>й жил<strong>к</strong>и или<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>о</strong><strong>с</strong>ущег<strong>о</strong><br />

тип<strong>а</strong> или <strong>а</strong>тр<strong>о</strong>фир<strong>о</strong><strong>в</strong><strong>а</strong>н у непит<strong>а</strong>ющих<strong>с</strong>я<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм; б<strong>о</strong>льшие <strong>в</strong>ыпу<strong>к</strong>лые гл<strong>а</strong>з<strong>а</strong>,<br />

неред<strong>к</strong><strong>о</strong> д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>к</strong><strong>а</strong>; <strong>а</strong>нтенны умеренн<strong>о</strong>й длины,<br />

р<strong>а</strong>зн<strong>о</strong>й ф<strong>о</strong>рмы: ните<strong>в</strong>идные, гребенч<strong>а</strong>тые,<br />

бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идные. Переднегрудь <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>с</strong>реднеи<br />

з<strong>а</strong>днегруди; н<strong>о</strong>ги длинные, х<strong>о</strong>дильные, ин<strong>о</strong>гд<strong>а</strong><br />

передние н<strong>о</strong>ги редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>к</strong>рылье<strong>в</strong>, <strong>с</strong>егмент<strong>о</strong><strong>в</strong> де<strong>с</strong>ять. Личин<strong>к</strong>и эру<strong>к</strong><strong>о</strong><strong>в</strong>идные<br />

— гу<strong>с</strong>еницы. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я, р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызущег<strong>о</strong> тип<strong>а</strong>; три п<strong>а</strong>ры грудных<br />

и пять п<strong>а</strong>р брюшных н<strong>о</strong>же<strong>к</strong>. Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у<br />

<strong>а</strong>рх<strong>а</strong>ичных Micropterygidae. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

П<strong>о</strong>д<strong>о</strong>тряды.- Jugata, Frenata.<br />

ПОДОТРЯД JUGATA. НИЗШИЕ<br />

РАВНОКРЫЛЫЕ БАБОЧКИ<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних и з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong><br />

<strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>; SC и R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>литы;<br />

<strong>в</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong>рыльях <strong>в</strong>ет<strong>в</strong>ей RS три — четыре, <strong>в</strong>ет<strong>в</strong>ей<br />

М — три; Ai <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>. Ж<strong>в</strong><strong>а</strong>лы<br />

б<strong>о</strong>лее или менее р<strong>а</strong>з<strong>в</strong>иты; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> — Micropterygidae и<br />

Eriocraniidae — из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong> п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong>.<br />

СЕМЕЙСТВО MICROPTERYGIDAE COTES, 1889.<br />

ЗУБЧАТЫЕ ПЕРВИЧНЫЕ МОЛИ<br />

[nom. transl. Comstok, 1893<br />

(ex Micropteryginae Cotes, 1889)1<br />

Очень мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые — не б<strong>о</strong>лее 12 мм<br />

<strong>в</strong> р<strong>а</strong>зм<strong>а</strong>хе; <strong>к</strong>рылья л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные, <strong>в</strong>ершин<strong>а</strong><br />

з<strong>а</strong><strong>о</strong><strong>с</strong>трен<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; н<strong>а</strong> SC<br />

<strong>в</strong>ет<strong>в</strong>ь; <strong>в</strong>ет<strong>в</strong>ей RS четыре; р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> RS <strong>с</strong><strong>в</strong><strong>о</strong>­<br />

(А. С. Д<strong>а</strong>ниле<strong>в</strong><strong>с</strong><strong>к</strong>ий и О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

б<strong>о</strong>дные; з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея. Ж<strong>в</strong><strong>а</strong>лы<br />

е<strong>с</strong>ть. Гу<strong>с</strong>еницы <strong>с</strong> длинными <strong>а</strong>нтенн<strong>а</strong>ми:<br />

н<strong>о</strong>ги н<strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>с</strong>егмент<strong>а</strong>х брюш<strong>к</strong><strong>а</strong> (тип<strong>а</strong> Ме<strong>с</strong><strong>о</strong>рtera);<br />

м<strong>а</strong><strong>к</strong><strong>с</strong>иллы <strong>с</strong> л<strong>а</strong>циниями; <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я<br />

(ри<strong>с</strong>. 956). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семь шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

R sc<br />

Ри<strong>с</strong>. 956—957. П<strong>о</strong>д<strong>о</strong>тряд Jugata<br />

956. Micropteryx sp.; ле<strong>в</strong>ые <strong>к</strong>рылья, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Handlirsch, 1925),<br />

957. Electrocrania immensipalpa Kusnezov; <strong>о</strong>бщий <strong>в</strong>ид, Xl2,0.<br />

п<strong>а</strong>ле<strong>о</strong>ген. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Кузнец<strong>о</strong><strong>в</strong>, 1941)<br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из них <strong>о</strong>дин из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и ми<strong>о</strong>цен<strong>а</strong> Ю. Азии (янт<strong>а</strong>рь<br />

Бирмы).<br />

СЕМЕЙСТВО ERIOCRANIIDAE REBEL, 1901.<br />

ПЕРВИЧНЫЕ МОЛИ<br />

Отлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т Micropterygidae б<strong>о</strong>лее ди<strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>л<strong>о</strong>жением н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>е<strong>к</strong>т<strong>о</strong>р<strong>а</strong> (н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>). Ж<strong>в</strong><strong>а</strong>л<br />

нет. Гу<strong>с</strong>еницы <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>а</strong>нтенн<strong>а</strong>ми; н<strong>о</strong>г<br />

нет; м<strong>а</strong><strong>к</strong><strong>с</strong>иллы <strong>с</strong> рудимент<strong>а</strong>ми л<strong>а</strong>циний; <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong><br />

<strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я (ри<strong>с</strong>. 957). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>дин <strong>в</strong>ымерший<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

303<br />

http://jurassic.ru/

ПОДОТРЯД FRENATA<br />

РАЗНОКРЫЛЫЕ БАБОЧКИ<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних и з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong><br />

р<strong>а</strong>зн<strong>о</strong>е; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях <strong>о</strong>дн<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я R; RS<br />

е<strong>с</strong>ть; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>о</strong>бычн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н;<br />

<strong>в</strong>ет<strong>в</strong>ей М три; Ai <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

длины <strong>к</strong>рыл<strong>а</strong> или немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее. Ж<strong>в</strong><strong>а</strong>лы<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны, х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> р<strong>а</strong>з<strong>в</strong>ит или <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н. П<strong>а</strong>ле<strong>о</strong>ген — ныне. П<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющее<br />

чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> чешуе<strong>к</strong>рылых <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я <strong>к</strong> эт<strong>о</strong>му п<strong>о</strong>д<strong>о</strong>тряду. Инфр<strong>а</strong><strong>о</strong>тряды:<br />

Microfrenata, Macrofrenata.<br />

И НФРАОТРЯД<br />

MICROFRENATA<br />

Б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью мел<strong>к</strong>ие или <strong>о</strong>чень мел<strong>к</strong>ие<br />

б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong>и. З<strong>а</strong>дние <strong>к</strong>рылья ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> б<strong>а</strong>хр<strong>о</strong>м<strong>к</strong><strong>о</strong>й,<br />

длин<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>о</strong>бычн<strong>о</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ширину<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М ч<strong>а</strong><strong>с</strong>т<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит, неред<strong>к</strong><strong>о</strong> е<strong>с</strong>ть<br />

меди<strong>а</strong>льн<strong>а</strong>я ячея; <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>о</strong>дн<strong>а</strong> —<br />

д<strong>в</strong>е. Гу<strong>с</strong>еницы, з<strong>а</strong> ред<strong>к</strong>им и<strong>с</strong><strong>к</strong>лючением, <strong>в</strong>едут<br />

<strong>с</strong><strong>к</strong>рытый <strong>о</strong>бр<strong>а</strong>з жизни. Брюшные н<strong>о</strong>ги <strong>в</strong>енечн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>; щетин<strong>к</strong>и т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пер<strong>в</strong>ичные. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>,<br />

из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong> третичн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени:<br />

Tineidea, Tortricidea, Gelechiidea, Pyralidea.<br />

НАДСЕМЕЙСТВО TINEIDEA.<br />

МОЛЕОБРАЗНЫЕ<br />

Общий <strong>с</strong>т<strong>в</strong><strong>о</strong>л М при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, или имеют<strong>с</strong>я<br />

<strong>с</strong>леды ег<strong>о</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong> т<strong>о</strong>рч<strong>а</strong>щих <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong>идных<br />

чешуй<strong>к</strong><strong>а</strong>х; челю<strong>с</strong>тные щупи<strong>к</strong>и р<strong>а</strong>з<strong>в</strong>иты (<strong>к</strong>р<strong>о</strong>ме<br />

Psychidae); ди<strong>с</strong>т<strong>а</strong>льны'й члени<strong>к</strong> нижнегубных<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong> туп<strong>о</strong>й. Гу<strong>с</strong>еницы <strong>о</strong>бычн<strong>о</strong> <strong>в</strong> чехли<strong>к</strong><strong>а</strong>х<br />

или п<strong>а</strong>утинных х<strong>о</strong>д<strong>а</strong>х. Ку<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong> <strong>с</strong> шип<strong>а</strong>ми<br />

н<strong>а</strong> тергит<strong>а</strong>х, <strong>в</strong>ыд<strong>в</strong>иг<strong>а</strong>ет<strong>с</strong>я из <strong>к</strong><strong>о</strong><strong>к</strong><strong>о</strong>н<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них четыре из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong> третичн<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>ремени: Incurvariidae, Adelidae, Tineidae,<br />

Psychidae.<br />

СЕМЕЙСТВО INCURVARIIDAE SPULER, 1910.<br />

МИННОЧЕХЛИКОВЫЕ МОЛИ<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием<br />

н<strong>а</strong> мембр<strong>а</strong>не <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong>р<strong>о</strong>ме чешуи, шипи<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

В<strong>с</strong>е жил<strong>к</strong>и е<strong>с</strong>ть; х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жены р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

и меди<strong>а</strong>льн<strong>а</strong>я ячеи. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит,<br />

<strong>а</strong>ф<strong>а</strong>ги; челю<strong>с</strong>тные щупи<strong>к</strong>и длиннее губных.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

(<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 70 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong>е<strong>в</strong>ерн<strong>о</strong>м умеренн<strong>о</strong>м п<strong>о</strong>я<strong>с</strong>е),<br />

из них <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

304<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ADELIDAE SPULER, 1910<br />

[nom. transl. Meyrick, 1912 (ex Adelinae Spuler, 1910)]<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние не <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

Incurvariidae, н<strong>о</strong> н<strong>а</strong> мембр<strong>а</strong>не <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

шипи<strong>к</strong>и; <strong>к</strong>рылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> мет<strong>а</strong>лличе<strong>с</strong><strong>к</strong>им<br />

бле<strong>с</strong><strong>к</strong><strong>о</strong>м. В <strong>о</strong>тличие <strong>о</strong>т других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

имеют т<strong>о</strong>н<strong>к</strong>ие длинные <strong>а</strong>нтенны (<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong><br />

длиннее <strong>к</strong>рылье<strong>в</strong>), н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м члени<strong>к</strong>е <strong>а</strong>нтенн<br />

гребен<strong>к</strong>и <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong> нет; темя <strong>с</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми;<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит; челю<strong>с</strong>тные щупи<strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че губных. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 200 шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных ле<strong>с</strong>ных <strong>в</strong>ид<strong>о</strong><strong>в</strong>), из них<br />

<strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО TINEIDAE LINNAEUS, 1758.<br />

НАСТОЯЩИЕ МОЛИ<br />

[nom. transl. Leach, 1819 (ex Tineae Linnaeus, 1758)]<br />

Н<strong>а</strong> мембр<strong>а</strong>не <strong>к</strong>рыл<strong>а</strong> шипи<strong>к</strong><strong>о</strong><strong>в</strong> нет; б<strong>а</strong>хр<strong>о</strong>м<strong>к</strong><strong>а</strong><br />

длинн<strong>а</strong>я, длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины ширины <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>о</strong>бычн<strong>о</strong> <strong>к</strong>рылья л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их,<br />

<strong>с</strong>т<strong>о</strong>ящих <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>х; <strong>а</strong>нтенны п<strong>о</strong> длине<br />

р<strong>а</strong><strong>в</strong>ны <strong>к</strong>рыльям, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й их члени<strong>к</strong> р<strong>а</strong><strong>с</strong>ширен,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> гребен<strong>к</strong><strong>о</strong>й; нижнечелю<strong>с</strong>тные щупи<strong>к</strong>и<br />

т<strong>о</strong>н<strong>к</strong>ие, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; нижнегубные<br />

щупи<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие (ри<strong>с</strong>. 958). Детрит<strong>о</strong>ф<strong>а</strong>ги,<br />

н<strong>а</strong> лиш<strong>а</strong>йни<strong>к</strong><strong>а</strong>х, гриб<strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1800 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, н<strong>а</strong>иб<strong>о</strong>лее<br />

<strong>о</strong>бильных <strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>: п<strong>а</strong>ле<strong>о</strong>ген Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

и 3. Е<strong>в</strong>р<strong>о</strong>пы; не<strong>о</strong>ген С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО PSYCHIDAE BOISDUVAL, 1929.<br />

МЕШЕЧНИЦЫ<br />

Н<strong>а</strong> мембр<strong>а</strong>не <strong>к</strong>рыл<strong>а</strong> шипи<strong>к</strong><strong>о</strong><strong>в</strong> нет; <strong>к</strong>рылья<br />

шир<strong>о</strong><strong>к</strong>ие, б<strong>а</strong>хр<strong>о</strong>м<strong>к</strong><strong>а</strong> з<strong>а</strong>дних <strong>к</strong>рылье<strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я.<br />

В передних <strong>к</strong>рыльях А] и А 2 <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>м пр<strong>о</strong>тяжении, з<strong>а</strong>тем р<strong>а</strong>зделяют<strong>с</strong>я, <strong>о</strong>бр<strong>а</strong>зуя<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>. Х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong> нет; челю<strong>с</strong>тные и<br />

губные щупи<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. С<strong>а</strong>м<strong>к</strong>и<br />

б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью без <strong>к</strong>рылье<strong>в</strong>, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> без н<strong>о</strong>г.<br />

Личин<strong>к</strong>и <strong>в</strong> чехли<strong>к</strong><strong>а</strong>х. Не<strong>о</strong>ген —• ныне. Неб<strong>о</strong>льш<strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м <strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х<br />

(<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 400 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне). Чехли<strong>к</strong>и<br />

личин<strong>о</strong><strong>к</strong> из ми<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО TORTRICIDEA.<br />

ЛИСТОВЕРТКООБРАЗНЫЕ<br />

З<strong>а</strong>дние <strong>к</strong>рылья т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины' или шире<br />

передних, <strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й SC; М <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я<br />

<strong>в</strong> <strong>в</strong>иде <strong>с</strong>лед<strong>о</strong><strong>в</strong> или х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> (Cossidae);<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong> прилег<strong>а</strong>ющих чешуй<strong>к</strong><strong>а</strong>х; нижнечелю<strong>с</strong>тных<br />

щупи<strong>к</strong><strong>о</strong><strong>в</strong> нет, или е<strong>с</strong>ть т<strong>о</strong>ль<strong>к</strong><strong>о</strong> их <strong>с</strong>ле-

RS, R sc<br />

Ри<strong>с</strong>. 958—964. П<strong>о</strong>д<strong>о</strong>тряд Frenata — Microfrenata<br />

58. Pa/aeoscardiites mordvilkoi Kusnezov; <strong>о</strong>бщий <strong>в</strong>ид, X8,0,<br />

<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Кузнец<strong>о</strong><strong>в</strong>, 1941). 962. Symmocites rohdendorfi Kus­<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Кузнец<strong>о</strong><strong>в</strong>, 1941). 959.<br />

nezov; <strong>о</strong>бщий <strong>в</strong>ид, х 8,7, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

Xyleutites miocenicus Kozhantshikov; переднее <strong>к</strong>рыл<strong>о</strong>, х 4,0,<br />

(Кузнец<strong>о</strong><strong>в</strong>, 1941). 963. Electresia zalesskii Kusnezov; <strong>о</strong>бщий <strong>в</strong>ид)<br />

п<strong>а</strong>ле<strong>о</strong>ген, С К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (<strong>о</strong>риг. ри<strong>с</strong>. <strong>с</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>), 960. Prolyonetia<br />

Х13.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Кузнец<strong>о</strong><strong>в</strong>, 1941),<br />

cockerelli Kusnezov; переднее <strong>к</strong>рыл<strong>о</strong>, х 19,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

964. Glendotricha olgae Kusnezov; <strong>о</strong>бщий <strong>в</strong>ид, х 8,5, п<strong>а</strong>ле<strong>о</strong>ген'<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Кузнец<strong>о</strong><strong>в</strong>, 1941). 961. Glesseumeyrickia hen-<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Кузнец<strong>о</strong><strong>в</strong>, 1941)<br />

rikseni Kusnezov; <strong>о</strong>бщий <strong>в</strong>ид, х 9,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий-<br />

ды; ди<strong>с</strong>т<strong>а</strong>льный члени<strong>к</strong> челю<strong>с</strong>тных щупи<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, цилиндриче<strong>с</strong><strong>к</strong>ий или туп<strong>о</strong>й. Гу<strong>с</strong>еницы<br />

<strong>с</strong><strong>к</strong>рыт<strong>о</strong>жи<strong>в</strong>ущие, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong> <strong>с</strong><strong>в</strong>ернутых ли<strong>с</strong>тьях<br />

или <strong>в</strong>нутри р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>й т<strong>к</strong><strong>а</strong>ни. Ку<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong> <strong>с</strong><br />

д<strong>в</strong>умя ряд<strong>а</strong>ми шипи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м тергите.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии:<br />

Cossidae, Tortricidae.<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>ги» 305<br />

СЕМЕЙСТВО COSSIDAE LEACH, 1815.<br />

ДРЕВОТОЧЦЫ<br />

[nom. transl. Walker, '1855 (ex Cossida Leach, 1815)]<br />

Крылья д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong>ие, <strong>с</strong> нез<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й; <strong>с</strong>т<strong>в</strong><strong>о</strong>л М е<strong>с</strong>ть; <strong>в</strong>ет<strong>в</strong>ей М три;<br />

меди<strong>а</strong>льн<strong>а</strong>я ячея х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>; р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея е<strong>с</strong>ть, <strong>в</strong>ершин<strong>а</strong> ее р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

меди<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и; д<strong>о</strong>-<br />

http://jurassic.ru/

б<strong>а</strong><strong>в</strong><strong>о</strong>чный р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>й из Вет<strong>в</strong>ей RS;<br />

CuP х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> й У-<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я фигур<strong>а</strong>.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 15—80 мм (ри<strong>с</strong>.959).<br />

Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. Один <strong>в</strong>ымерший р<strong>о</strong>д из ми<strong>о</strong>цен<strong>а</strong><br />

К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>.<br />

СЕМЕЙСТВО TORTRICIDAE LINNAEUS, 1758.<br />

ЛИСТОВЕРТКИ<br />

[nom. transl. Stephens, 1829 (ex Tortrices Linnaeus, 1758)]<br />

Вершины р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й и меди<strong>а</strong>льн<strong>о</strong>й ячей<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не; д<strong>о</strong>б<strong>а</strong><strong>в</strong><strong>о</strong>чных<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ях RS <strong>о</strong>бычн<strong>о</strong> нет; CuP <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует,<br />

или <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я ее <strong>с</strong>леды <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>; У-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й фигуры не<br />

<strong>в</strong>идн<strong>о</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong><br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong>: Eucosminae, Phaloniinae.<br />

ПОДСЕМЕЙСТВО EUCOSMINAE DURRENT, 1918<br />

В передних <strong>к</strong>рыльях CuA р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не третьей чет<strong>в</strong>ерти длины меди<strong>а</strong>льн<strong>о</strong>-<strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й<br />

ячеи; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях н<strong>а</strong> CuA гребен<strong>к</strong><strong>а</strong><br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>; <strong>в</strong>ершин<strong>а</strong> CuP <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я<br />

(ри<strong>с</strong>. 960). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненн<strong>о</strong>е п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>,<br />

б<strong>о</strong>лее де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО PHALONIINAE REBEL, 1901<br />

[nom transl. Rebel, 1935 (ex Phaloniidae Rebel, 1901)]<br />

В передних <strong>к</strong>рыльях CuA р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>в</strong>близи ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> угл<strong>а</strong> меди<strong>а</strong>льн<strong>о</strong>-<strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й<br />

ячеи, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й чет<strong>в</strong>ерти ее<br />

длины; CuP <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong>е<strong>в</strong>ерн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>луш<strong>а</strong>рия (<strong>с</strong> неб<strong>о</strong>льшим <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>); <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО GELECHIIDEA<br />

Близ<strong>к</strong><strong>о</strong> п<strong>о</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong> Tortricidea. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong> прилег<strong>а</strong>ющими чешуй<strong>к</strong><strong>а</strong>ми, гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я; нижнегубные<br />

щупи<strong>к</strong>и <strong>с</strong><strong>а</strong>бле<strong>в</strong>идные, <strong>с</strong> длинным,<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>тренным ди<strong>с</strong>т<strong>а</strong>льным члени<strong>к</strong><strong>о</strong>м. Гу<strong>с</strong>еницы<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> <strong>с</strong><strong>к</strong>рытн<strong>о</strong> жи<strong>в</strong>ущие, ин<strong>о</strong>гд<strong>а</strong> минеры.<br />

Ку<strong>к</strong><strong>о</strong>л<strong>к</strong>и без шипи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> тергит<strong>а</strong>х, не <strong>в</strong>ыд<strong>в</strong>иг<strong>а</strong>ют<strong>с</strong>я<br />

из <strong>к</strong><strong>о</strong><strong>к</strong><strong>о</strong>н<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Четыре<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны: Hyponomeutidae,<br />

Oecophoridae, Gelechiidae, Lyonetiidae.<br />

СЕМЕЙСТВО HYPONOMEUTIDAE STAINTON, 1854.<br />

ГОРНОСТАЕВЫЕ МОЛИ<br />

Крылья уз<strong>к</strong>ие, л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

з<strong>а</strong>дние; н<strong>а</strong> передних <strong>к</strong>рыльях ч<strong>а</strong><strong>с</strong>т<strong>о</strong> е<strong>с</strong>ть птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>;<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>чень пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>,<br />

306<br />

http://jurassic.ru/<br />

<strong>о</strong>т<strong>к</strong>л<strong>о</strong>няет<strong>с</strong>я <strong>о</strong>т R, <strong>о</strong>бр<strong>а</strong>зуя уг<strong>о</strong>л; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и RS<br />

н<strong>а</strong>чин<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т <strong>в</strong>ершины р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й ячеи; RS 4<br />

упир<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>а</strong>пи<strong>к</strong><strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>; CuP<br />

е<strong>с</strong>ть. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или <strong>с</strong> нер<strong>о</strong><strong>в</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

чешуй<strong>к</strong><strong>а</strong>ми; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й члени<strong>к</strong> <strong>а</strong>нтенн<br />

р<strong>а</strong><strong>с</strong>ширен, ин<strong>о</strong>гд<strong>а</strong> ут<strong>о</strong>лщен; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> б<strong>о</strong>льшей<br />

ч<strong>а</strong><strong>с</strong>тью р<strong>а</strong>з<strong>в</strong>ит; нижнечелю<strong>с</strong>тные щупи<strong>к</strong>и редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны'.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 800<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>); три <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО OECOPHORIDAE COTES, 1889<br />

[nom. transl. Meyrick, 1895<br />

(ex Oecophorinae Cotes, 1889)]<br />

В передних <strong>к</strong>рыльях RS3 и RS4 н<strong>а</strong> длинн<strong>о</strong>м<br />

<strong>с</strong>тебель<strong>к</strong>е или <strong>с</strong>литы; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует:<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>-<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я ячея шир<strong>о</strong><strong>к</strong><strong>а</strong>я;<br />

з<strong>а</strong>дние <strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, <strong>о</strong><strong>в</strong><strong>а</strong>льные или менее<br />

л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные; <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т Gelechiidae <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием<br />

<strong>в</strong>ыем<strong>к</strong>и п<strong>о</strong> <strong>в</strong>нешнему <strong>к</strong>р<strong>а</strong>ю; SC <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я;<br />

R и М п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельны. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong><strong>к</strong>рыт<strong>а</strong><br />

приж<strong>а</strong>тыми или <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными чешуй<strong>к</strong><strong>а</strong>ми;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й члени<strong>к</strong> <strong>а</strong>нтенн <strong>с</strong> гребен<strong>к</strong><strong>о</strong>й <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>;<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бы<strong>в</strong><strong>а</strong>ет; нижнечелю<strong>с</strong>тные<br />

щупи<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; губные<br />

щупи<strong>к</strong>и длинные, из<strong>о</strong>гнутые (ри<strong>с</strong>. 961). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>), из них п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong><br />

<strong>а</strong><strong>в</strong><strong>с</strong>тр<strong>а</strong>лий<strong>с</strong><strong>к</strong>ие; четыре р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО LYONETIIDAE STAINTON, 1854.<br />

УЗКОКРЫЛЫЕ МИНЕРЫ<br />

Очень мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Кры'лья уз<strong>к</strong>ие,<br />

л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идные, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> з<strong>а</strong>дние; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; М п<strong>о</strong>чти <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

CuP х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>а</strong>. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий;<br />

нижнечелю<strong>с</strong>тные щупи<strong>к</strong>и <strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны (ри<strong>с</strong>. 962). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

900 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных <strong>в</strong>ид<strong>о</strong><strong>в</strong>); <strong>о</strong>дин<br />

<strong>в</strong>ымерший р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО GELECHIIDAE STAINTON. 1854.<br />

ВЫЕМЧАТОКРЫЛЫЕ МОЛИ<br />

Крылья б<strong>о</strong>лее или менее шир<strong>о</strong><strong>к</strong>ие, р<strong>а</strong>зн<strong>о</strong>й<br />

ф<strong>о</strong>рмы; <strong>в</strong> передних <strong>к</strong>рыльях р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й ячеи<br />

нет; RS3 и RS4 н<strong>а</strong> <strong>с</strong>тебель<strong>к</strong>е или <strong>с</strong>литы, RS 4<br />

<strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong>переди <strong>в</strong>ершины; <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л<br />

М <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong>; CuP ч<strong>а</strong><strong>с</strong>т<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>;<br />

SC <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>а</strong> п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й <strong>с</strong> R;<br />

з<strong>а</strong>дние <strong>к</strong>рылья ч<strong>а</strong><strong>с</strong>т<strong>о</strong> тр<strong>а</strong>пец<strong>о</strong>идн<strong>о</strong>й ф<strong>о</strong>рмы, <strong>с</strong><br />

<strong>в</strong>ыем<strong>к</strong><strong>о</strong>й п<strong>о</strong>д <strong>в</strong>ершин<strong>о</strong>й п<strong>о</strong> <strong>в</strong>нешнему <strong>к</strong>р<strong>а</strong>ю.

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й члени<strong>к</strong> <strong>а</strong>нтенн <strong>с</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми;<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит; нижние челю<strong>с</strong>тные<br />

щупи<strong>к</strong>и прямые, или их нет; губные<br />

щупи<strong>к</strong>и длинные, из<strong>о</strong>гнутые, п<strong>о</strong>дняты <strong>к</strong><strong>в</strong>ерху<br />

(ри<strong>с</strong>. 963). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>); д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО PYRALIDIDEA.<br />

ОГНЕВКООБРАЗНЫЕ<br />

Передние <strong>к</strong>рылья <strong>с</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящей р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>меди<strong>а</strong>льн<strong>о</strong>й<br />

ячеей, т. е. <strong>с</strong>т<strong>в</strong><strong>о</strong>л М п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н;<br />

з<strong>а</strong>дние <strong>к</strong>рылья <strong>о</strong>бычн<strong>о</strong> шире передних;<br />

SC и R <strong>с</strong>литы <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я<br />

з<strong>а</strong> предел<strong>а</strong>ми ячеи, ред<strong>к</strong><strong>о</strong> <strong>с</strong>литы цели<strong>к</strong><strong>о</strong>м <strong>в</strong><br />

<strong>о</strong>дну жил<strong>к</strong>у. Челю<strong>с</strong>тные щупи<strong>к</strong>и х<strong>о</strong>р'<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты';<br />

нижнегубные щупи<strong>к</strong>и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них <strong>с</strong> третичн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

<strong>о</strong>дн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pyralididae.<br />

СЕМЕЙСТВО PYRALIDIDAE LINNAEUS, 1758.<br />

ОГНЕВКИ<br />

rnom. transl. Leach, 1819 (ex Pyralides Linnaeus, 1758)]<br />

Крылья треуг<strong>о</strong>льные; <strong>в</strong> передних <strong>к</strong>рыльях<br />

RS 2, RS3 и RS 4 н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м <strong>с</strong>тебле; CuP <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует,<br />

или <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я лишь <strong>с</strong>леды ее; Ai<br />

длинн<strong>а</strong>я, Аг нет (ри<strong>с</strong>. 964). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Б<strong>о</strong>лее 300 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

10 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м инд<strong>о</strong><strong>а</strong><strong>в</strong><strong>с</strong>тр<strong>а</strong>лий<strong>с</strong><strong>к</strong>их<br />

и не<strong>о</strong>тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их); д<strong>в</strong><strong>а</strong> <strong>в</strong>ы'мерших<br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

и из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ИНФРАОТРЯД MACROFRENATA<br />

Б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью <strong>к</strong>рупные или <strong>с</strong>редней <strong>в</strong>еличины<br />

б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong>и; з<strong>а</strong>дние <strong>к</strong>рылья б<strong>о</strong>лее или<br />

менее шир<strong>о</strong><strong>к</strong>ие, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й б<strong>а</strong>хр<strong>о</strong>м<strong>о</strong>й; <strong>о</strong>бщий<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н; меди<strong>а</strong>льн<strong>о</strong>й ячеи нет;<br />

<strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> <strong>о</strong>дн<strong>а</strong> — д<strong>в</strong>е. Гу<strong>с</strong>еницы, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>в</strong>едут <strong>о</strong>т<strong>к</strong>рытый <strong>о</strong>бр<strong>а</strong>з жизни. Брюшные<br />

н<strong>о</strong>ги <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льным ряд<strong>о</strong>м <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>. П<strong>а</strong>­<br />

Одн<strong>а</strong> передняя п<strong>а</strong>р<strong>а</strong> <strong>к</strong>рылье<strong>в</strong>. Передний<br />

<strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й. SC без <strong>в</strong>ет<strong>в</strong>и или <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й<br />

передней <strong>в</strong>ет<strong>в</strong>ью <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й. СиА<br />

и СиР пр<strong>о</strong><strong>с</strong>тые, <strong>в</strong><strong>с</strong>егд<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>с</strong>ближенные и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong> <strong>в</strong>иде те<strong>с</strong>н<strong>о</strong>й <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и. З<strong>а</strong>дние<br />

<strong>к</strong>рылья пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идные <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния—<br />

жужж<strong>а</strong>льц<strong>а</strong>. С<strong>о</strong><strong>с</strong>ущий или <strong>к</strong><strong>о</strong>лю­<br />

ле<strong>о</strong>ген — ныне. Семь н<strong>а</strong>д<strong>с</strong>ёмей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них три — Rhopalocera, Sphingodea,<br />

Noctuidea — из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong> третичн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени;<br />

пер<strong>в</strong>ые д<strong>в</strong><strong>а</strong> <strong>в</strong>не СССР.<br />

НАДСЕМЕЙСТВО NOCTUIDEA.<br />

СОВКООБРАЗНЫЕ<br />

В передних <strong>к</strong>рыльях <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния М 2 и Мз<br />

<strong>с</strong>ближены'; CuP редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях<br />

RS не <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>а</strong> <strong>с</strong> R. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них <strong>с</strong><br />

третичн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ремени из<strong>в</strong>е<strong>с</strong>тны Amatidae, Noctuidae,<br />

Arctiidae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО AMATIDAE JANSE, 1917<br />

(Syntomidae Н. Schafer, 1847)<br />

Средней <strong>в</strong>еличины пе<strong>с</strong>трые б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong>и; з<strong>а</strong>дние<br />

<strong>к</strong>рылья <strong>в</strong><strong>с</strong>егд<strong>а</strong> меньше передних; <strong>в</strong>ет<strong>в</strong>и RS<br />

з<strong>а</strong> предел<strong>а</strong>ми ячей, <strong>в</strong> передних <strong>к</strong>рыльях н<strong>а</strong>.<br />

Ри<strong>с</strong>. 965. П<strong>о</strong>д<strong>о</strong>тряд Frenata — Macrofrenata.<br />

965. Oligamatites martynovi Kusnezov; переднее <strong>к</strong>рыл<strong>о</strong>, Xl,3,<br />

п<strong>а</strong>ле<strong>о</strong>ген, В. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Кузнец<strong>о</strong><strong>в</strong>, 1928)<br />

<strong>о</strong>дн<strong>о</strong>м <strong>с</strong>тебле; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях SC + R <strong>о</strong>т<strong>с</strong>ут-';<br />

<strong>с</strong>т<strong>в</strong>ует, <strong>с</strong>лит<strong>а</strong> <strong>с</strong> RS; <strong>в</strong>ет<strong>в</strong>и М <strong>к</strong><strong>а</strong><strong>к</strong> бы <strong>о</strong>тх<strong>о</strong>дят<br />

<strong>о</strong>т передней <strong>в</strong>ет<strong>к</strong>и СиА, <strong>о</strong>бр<strong>а</strong>зуя «гребен<strong>к</strong>у»,<br />

<strong>в</strong>ет<strong>в</strong>ей М три. Гл<strong>а</strong>з<strong>к</strong>и редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

и губные щупи<strong>к</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты (ри<strong>с</strong>. 965).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

ОТРЯД D1PTERA. ДВУКРЫЛЫЕ<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

щий х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>. Среднегрудь <strong>о</strong>чень <strong>в</strong>ели<strong>к</strong><strong>а</strong>, передне-<br />

и з<strong>а</strong>днегрудь м<strong>а</strong>лы'; л<strong>а</strong>п<strong>к</strong>и п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Личин<strong>к</strong>и безн<strong>о</strong>гие, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><br />

пл<strong>о</strong>х<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong> у<strong>к</strong>рытых<br />

<strong>с</strong>ред<strong>а</strong>х: п<strong>о</strong>ч<strong>в</strong>е, пре<strong>с</strong>ных <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, т<strong>к</strong><strong>а</strong>нях<br />

р<strong>а</strong><strong>с</strong>тений и жи<strong>в</strong><strong>о</strong>тных. Крыл<strong>а</strong>т<strong>а</strong>я ф<strong>а</strong>з<strong>а</strong><br />

те<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>а</strong> <strong>с</strong> <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>а</strong>ми <strong>о</strong>бит<strong>а</strong>ния личи-<br />

307 20*<br />

http://jurassic.ru/

Ри<strong>с</strong>. 966 Схем<strong>а</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>о</strong>тн<strong>о</strong>шений инфр<strong>а</strong><strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>а</strong> Diptera<br />

Н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>л<strong>о</strong>н<strong>к</strong>и <strong>о</strong>т<strong>о</strong>бр<strong>а</strong>жены <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>л<strong>о</strong><strong>с</strong><br />

р<strong>а</strong>зличн<strong>о</strong>й ширины инфр<strong>а</strong><strong>о</strong>тряды д<strong>в</strong>у<strong>к</strong>рылых, <strong>в</strong>ремя их <strong>в</strong><strong>о</strong>зни<strong>к</strong>н<strong>о</strong><strong>в</strong>ения<br />

и <strong>в</strong>ымир<strong>а</strong>ния, и<strong>с</strong>х<strong>о</strong>дные группы и <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong>е р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зие<br />

(п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>нн<strong>о</strong>е ширин<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>). Преры<strong>в</strong>и<strong>с</strong>т<strong>о</strong>й линией п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>о</strong><br />

<strong>в</strong>ремя <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong><strong>в</strong> пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong>. Ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы<strong>а</strong><br />

— <strong>в</strong>ерхнетри<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ый из Ср. Азии; 6 — нижнеюр<strong>с</strong><strong>к</strong>ий из 3 Е<strong>в</strong>р<strong>о</strong>пы;<br />

<strong>в</strong> — <strong>в</strong>ерхнеюр<strong>с</strong><strong>к</strong>ий из К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> (К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у); г — <strong>в</strong>ерхнеюр<strong>с</strong><strong>к</strong>ий<br />

из З.Е<strong>в</strong>р<strong>о</strong>пы; д — п<strong>а</strong>ле<strong>о</strong>ген<strong>о</strong><strong>в</strong>ый из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря-<br />

н<strong>о</strong><strong>к</strong>, ц<strong>в</strong>ет<strong>к</strong><strong>о</strong><strong>в</strong>ыми р<strong>а</strong><strong>с</strong>тениями и п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чными<br />

жи<strong>в</strong><strong>о</strong>тными. Пит<strong>а</strong>ние <strong>о</strong>чень р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. Фил<strong>о</strong>генез<br />

еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тен (ри<strong>с</strong>. 966).<br />

Три<strong>а</strong><strong>с</strong> — ныне. В <strong>к</strong><strong>о</strong>нце три<strong>а</strong><strong>с</strong><strong>а</strong> были р<strong>а</strong>з<strong>в</strong>иты<br />

<strong>о</strong>б<strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: дре<strong>в</strong>ний, и<strong>с</strong>х<strong>о</strong>дный — Archidiptera,<br />

уже т<strong>о</strong>гд<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленный немн<strong>о</strong>гими<br />

групп<strong>а</strong>ми, и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й — Eudiptera, <strong>в</strong> д<strong>а</strong>льнейшем<br />

д<strong>о</strong><strong>с</strong>тигший шир<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития.<br />

ПОДОТРЯД ARCHIDIPTERA<br />

Очень неб<strong>о</strong>льшие <strong>к</strong>рылья, не пре<strong>в</strong>ыш<strong>а</strong>ющие<br />

4 мм, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>перечными п<strong>о</strong>чти<br />

<strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех п<strong>о</strong>лях. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е:<br />

пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и нер<strong>а</strong><strong>в</strong>н<strong>о</strong>й т<strong>о</strong>лщины, <strong>к</strong>реп<strong>к</strong>ие<br />

С, R, CuA и т<strong>о</strong>н<strong>к</strong>ие RS, М, CuP и А. Мез<strong>о</strong>­<br />

308<br />

http://jurassic.ru/<br />

инфр<strong>а</strong><strong>о</strong>тряды д<strong>в</strong>у<strong>к</strong>рылых.- 1 - Diplopolyneuromorpha; 2 - Dic-<br />

£nrnh^ r<br />

°T rph<br />

З Si и ~ Nymphomylomorpha; 4 - Deuterophleb omprpha,<br />

5 — Blephariceromorpha; 6 - Tipulomorpha; 7-<br />

Bibionornorpha; 8 - Musidoromorpha; 9 - Phoromorpha; 10 -<br />

Termitoxenfomorpha, // - Asilomorpha, 12 - Myiomorpha;13-<br />

Braulomorpha; 14 - Nycteribiomorpha; 15 - Strebfomorpha<br />

(Р<strong>о</strong>денд<strong>о</strong>рф и др. 1961). Н<strong>а</strong> <strong>с</strong>хеме <strong>с</strong>т<strong>в</strong><strong>о</strong>л ф<strong>о</strong>р<strong>о</strong>м<strong>о</strong>рф <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<br />

<strong>о</strong>тх<strong>о</strong>дящим <strong>о</strong>т <strong>а</strong>зил<strong>о</strong>м<strong>о</strong>рф <strong>в</strong> юре; <strong>о</strong>н <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ил<strong>с</strong>я <strong>о</strong>т и<strong>с</strong>х<strong>о</strong>дных<br />

мий<strong>о</strong>м<strong>о</strong>рф <strong>в</strong> мелу.<br />

з<strong>о</strong>й. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Dictyodipteridea, Hyperpolyneuridea,<br />

Dyspolyneuridea.<br />

НАДСЕМЕЙСТВО DICTYODIPTERIDEA<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

из<strong>о</strong>гнутый, без фр<strong>а</strong>гмы и изл<strong>о</strong>м<strong>а</strong>.<br />

CuA <strong>о</strong>чень <strong>к</strong>реп<strong>к</strong><strong>а</strong>я. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итыми п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми. Три<strong>а</strong><strong>с</strong>.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО DICTYODIPTERIDAE<br />

ROHDENDORF, 1961<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

прям<strong>о</strong>й; С, SC и R ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> п<strong>а</strong>р<strong>а</strong>ллельны,

<strong>с</strong>ближены; RS и М <strong>с</strong> немн<strong>о</strong>гими <strong>в</strong>ет<strong>в</strong>ями, т<strong>о</strong>н<strong>к</strong>ие;<br />

Си р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>с</strong>редней линии<br />

<strong>к</strong>рыл<strong>а</strong>. Три<strong>а</strong><strong>с</strong>. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Dictyodiptera Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. multinervis Rohdendorf, 1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль).<br />

Между СиА и СиР имеют<strong>с</strong>я п<strong>о</strong>перечные; <strong>о</strong>бе<br />

А п<strong>о</strong>чти П<strong>а</strong>р<strong>а</strong>ллельны. Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, б<strong>о</strong>лее чем<br />

<strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

2,8 мм (ри<strong>с</strong>. 967). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Paradictyodiptera Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. trianalis Rohdendorf, 1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Между<br />

СиА и СиР п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong> нет; А неп<strong>а</strong>р<strong>а</strong>ллельные,<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е трех т<strong>о</strong>н<strong>к</strong>их жил<strong>о</strong><strong>к</strong>;<br />

между С и SC е<strong>с</strong>ть п<strong>о</strong>перечные жил<strong>к</strong>и.<br />

Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е — <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3,5 мм (ри<strong>с</strong>. 968). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Dipterodictya Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

D. tipuloides Rohdendorf, 1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Между<br />

СиА и СиР п<strong>о</strong>перечных нет; MP <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong><br />

<strong>в</strong> <strong>в</strong>иде п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>й жил<strong>к</strong>и перед СиА<br />

не <strong>с</strong><strong>о</strong>единенн<strong>о</strong>й <strong>с</strong> п<strong>о</strong><strong>с</strong>ледней п<strong>о</strong>перечными.<br />

Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, б<strong>о</strong>лее чем <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

ширины. Длин<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3,5 мм (ри<strong>с</strong>. 970).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

НАДСЕМЕЙСТВО HYPERPOLYNEURIDEA<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> <strong>с</strong> рез<strong>к</strong>им<br />

изл<strong>о</strong>м<strong>о</strong>м, <strong>о</strong>бр<strong>а</strong>зующим з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чную фр<strong>а</strong>гму.<br />

Крыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> <strong>к</strong>реп<strong>к</strong><strong>о</strong>й передней <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong><br />

без п<strong>о</strong>перечных жил<strong>о</strong><strong>к</strong>. Три<strong>а</strong><strong>с</strong>. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО HYPERPOLYNEURIDAE<br />

ROHDENDORF, 1961<br />

Передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

прям<strong>о</strong>й. RS и R <strong>с</strong>ближенные и п<strong>а</strong>р<strong>а</strong>ллельные.<br />

Не менее трех <strong>в</strong>ет<strong>в</strong>ей RS и не менее ше<strong>с</strong>ти<br />

<strong>в</strong>ет<strong>в</strong>ей М, <strong>в</strong>ых<strong>о</strong>дящих з<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й. Р<strong>а</strong>змеры<br />

мел<strong>к</strong>ие: <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3 мм. Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Hyperpolyneura Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Н. phryganeoides Rohdendorf, 1961; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Между MP, <strong>к</strong>реп<strong>к</strong><strong>о</strong>й СиА имеют<strong>с</strong>я п<strong>о</strong>перечные;<br />

MP <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>с</strong>ледней <strong>в</strong>ет<strong>в</strong>ью<br />

МА; SC н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> R. А ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я, <strong>о</strong>бр<strong>а</strong>зуя з<strong>а</strong>м<strong>к</strong>нутую ячей<strong>к</strong>у.<br />

Крыл<strong>о</strong> примерн<strong>о</strong> <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

ширины. Длин<strong>а</strong> ег<strong>о</strong> 2,8 мм (ри<strong>с</strong>. 971). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

309<br />

Ри<strong>с</strong>. 967—972. П<strong>о</strong>д<strong>о</strong>тряд Archidiptera<br />

967. Dictyodiptera multinervis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x 21,0, <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, Ср. Азия (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 968. Paradictyodiptera trianalis<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х25.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1961). 969. Nymphomyia alba Tokunaga; <strong>к</strong>рыл<strong>о</strong>, х 20,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Tokunaga, 1.935). 970. Dipterodictya tipuloides Rohdendorf;<br />

<strong>к</strong>рыл<strong>о</strong>, x 17,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 971. Hyperpolyneura<br />

phryganeoides Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х24,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>;<br />

Ср. Азия (Р<strong>о</strong>денд<strong>о</strong>рф, 1961). 972. Dyspolyneura longipennis Rohdendorf;<br />

<strong>к</strong>рыл<strong>о</strong>, x 20,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (Р<strong>о</strong>денд<strong>о</strong>рф, 1961)<br />

http://jurassic.ru/

НАДСЕМЕЙСТВО DYSPOLYNEURIDEA<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и <strong>с</strong> изл<strong>о</strong>м<strong>о</strong>м<br />

и фр<strong>а</strong>гм<strong>о</strong>й. Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> ут<strong>о</strong>лщенн<strong>о</strong>й<br />

<strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й. Ан<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

без п<strong>о</strong>перечных. Три<strong>а</strong><strong>с</strong>. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО DYSPOLYNEURIDAE<br />

ROHDENDORF, 1961<br />

MP рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я <strong>о</strong>т МА и <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляющ<strong>а</strong>я<strong>с</strong>я<br />

<strong>о</strong>т CuA, <strong>с</strong><strong>о</strong>единенн<strong>а</strong>я <strong>с</strong> МА <strong>к</strong>реп<strong>к</strong><strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й. RS <strong>в</strong> <strong>в</strong>иде <strong>к</strong>реп<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>, <strong>о</strong>т<strong>с</strong>ыл<strong>а</strong>ющег<strong>о</strong><br />

<strong>в</strong>ет<strong>в</strong>и <strong>в</strong>перед. Си <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong>реп<strong>к</strong>их р<strong>а</strong>ди<strong>о</strong>меди<strong>а</strong>льных<br />

п<strong>о</strong>перечных. Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Dyspolyneura Rohdendorf, 1961. Тип р<strong>о</strong>д<strong>а</strong>—<br />

D. longipennis Rohdendorf, 1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). SC<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; между С и R имеют<strong>с</strong>я<br />

п<strong>о</strong>перечные; не менее пяти <strong>в</strong>ет<strong>в</strong>ей RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющих<br />

<strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> MP<br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>ближен<strong>а</strong> и п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> CuA; п<strong>о</strong><strong>с</strong>ледняя<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> п<strong>о</strong>чти<br />

<strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ширины, р<strong>а</strong><strong>в</strong>н<strong>а</strong> 3 мм<br />

(ри<strong>с</strong>. 972). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

ПОДОТРЯД EUDIPTERA<br />

Крылья <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>уженные. В<strong>с</strong>егд<strong>а</strong><br />

имеют<strong>с</strong>я х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я р<strong>а</strong>ди<strong>о</strong>меди<strong>а</strong>льн<strong>а</strong>я<br />

п<strong>о</strong>перечн<strong>а</strong>я и немн<strong>о</strong>гие интермеди<strong>а</strong>льные:<br />

другие п<strong>о</strong>перечные, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

или р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>. Р<strong>а</strong>змеры<br />

р<strong>а</strong>зличны: <strong>о</strong>т 0,5 д<strong>о</strong> 50 мм. Три<strong>а</strong><strong>с</strong> — ныне. Из<strong>в</strong>е<strong>с</strong>тны<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители инфр<strong>а</strong><strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

Diplopolyneuromorpha, Tipulomorpha, Bibionomorpha,<br />

Asilomorpha, Phoromorpha, Musidoromorpha,<br />

Myiomorpha; <strong>о</strong><strong>с</strong>т<strong>а</strong>льные <strong>с</strong>емь из<strong>в</strong>е<strong>с</strong>тных<br />

инфр<strong>а</strong><strong>о</strong>тряд<strong>о</strong><strong>в</strong> д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р <strong>о</strong>бн<strong>а</strong>ружен<strong>а</strong><br />

лишь <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

ИНФРАОТРЯД<br />

DIPLOPOLYNEUROMORPHA<br />

Крыл<strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из д<strong>в</strong>ух <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных р<strong>а</strong>здел<strong>о</strong><strong>в</strong>:<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными ячей<strong>к</strong><strong>а</strong>ми<br />

и п<strong>а</strong>р<strong>а</strong>ллельными жил<strong>к</strong><strong>а</strong>ми и пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong> п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong> <strong>с</strong>ближенными пр<strong>о</strong>д<strong>о</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми, лишенными п<strong>о</strong>перечных. Три<strong>а</strong><strong>с</strong>.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Diplopolyneuridae.<br />

СЕМЕЙСТВО DIPLOPOLYNEURIDAE<br />

ROHDENDORF, 1961<br />

R и SG <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> трети <strong>к</strong>рыл<strong>а</strong>, <strong>с</strong>ближенные<br />

т<strong>а</strong><strong>к</strong> же, <strong>к</strong><strong>а</strong><strong>к</strong> МА и MP, CuA и CuP, А] и А 2.<br />

Ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> не менее чем <strong>с</strong> че-<br />

310<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 973. Инфр<strong>а</strong><strong>о</strong>тряд Diplopolyneuromorpha. Diplopolyneura<br />

mirabilis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X 17,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср.<br />

Азия (Р<strong>о</strong>денд<strong>о</strong>рф, 1961)<br />

тырьмя жил<strong>к</strong><strong>а</strong>ми. А 2 <strong>с</strong> д<strong>в</strong>умя <strong>в</strong>ет<strong>в</strong>ями. Три<strong>а</strong><strong>с</strong>.<br />

Один р<strong>о</strong>д.<br />

Diplopolyneura Rohdendorf, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. mirabilis Rohdendorf, 1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). R,<br />

МА, CuA и С <strong>к</strong>реп<strong>к</strong>ие, рез<strong>к</strong><strong>о</strong> <strong>о</strong>тличные <strong>о</strong>т других<br />

жил<strong>о</strong><strong>к</strong>; <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л М и CuA т<strong>о</strong>н<strong>к</strong>ий. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4,5 мм (ри<strong>с</strong>. 973). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong><br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

ИНФРАОТРЯД<br />

TIPULOMORPHA<br />

RS <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющими <strong>в</strong> <strong>в</strong>ершинный <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>.<br />

Крыл<strong>о</strong> удлиненн<strong>о</strong>е <strong>с</strong> неб<strong>о</strong>льшим <strong>в</strong>ершинным<br />

<strong>о</strong>тдел<strong>о</strong>м. Антенны <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> г<strong>о</strong>м<strong>о</strong>н<strong>а</strong>льными члени<strong>к</strong><strong>а</strong>ми.<br />

Р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дит преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> <strong>в</strong><strong>о</strong>дн<strong>о</strong>й <strong>с</strong>реде. 13 н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Tipulidodictyidea,<br />

Eopolyneuridea, Tipulidea, Eoptychopteridea,<br />

Tanyderophryneidea, Dixidea, Culicidea,<br />

Psychodidea, Chironomidea, Rhaetomyiidea, Mesophantasmatidea,<br />

Thaumaleidea и Pachyneuridea;<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители д<strong>в</strong>ух п<strong>о</strong><strong>с</strong>ледних д<strong>о</strong> <strong>с</strong>их<br />

п<strong>о</strong>р из<strong>в</strong>е<strong>с</strong>тны лишь <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>е <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны.<br />

НАДСЕМЕЙСТВО TIPULODICTYIDEA<br />

З<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь М <strong>с</strong>ближен<strong>а</strong> <strong>с</strong> CuA <strong>о</strong>бр<strong>а</strong>зуя<br />

<strong>с</strong> нею и <strong>с</strong> CuP <strong>в</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>л<strong>о</strong>жную<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>у. RS <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ллельными <strong>в</strong>ет<strong>в</strong>ями.<br />

Имеют<strong>с</strong>я <strong>с</strong>л<strong>а</strong>бые п<strong>о</strong>перечные <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле<br />

и между р<strong>а</strong>ди<strong>а</strong>льными <strong>в</strong>ет<strong>в</strong>ями. Три<strong>а</strong><strong>с</strong>. Одн<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО TIPULODICTYIDAE<br />

ROHDENDORF, Fam. nov.<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние R <strong>в</strong><strong>о</strong>лн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнут<strong>о</strong>е, без<br />

фр<strong>а</strong>гмы. SC не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> д<strong>а</strong>льше <strong>с</strong>ередины 1<br />

<strong>к</strong>рыл<strong>а</strong><br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С. М. <strong>с</strong>л<strong>а</strong>бые. Три<strong>а</strong><strong>с</strong>. Один<br />

р<strong>о</strong>д.

Ри<strong>с</strong> 974—977. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tipulodictyidea, Eopolyneuridea<br />

Tipulodictya Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. minima Rohdendorf sp. nov. <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Р<strong>а</strong>ди<strong>о</strong>меди<strong>а</strong>льн<strong>а</strong>я<br />

п<strong>о</strong>перечн<strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льнее ур<strong>о</strong><strong>в</strong>ня<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> SC; <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле не менее де<strong>с</strong>ят<strong>к</strong><strong>а</strong><br />

<strong>с</strong>л<strong>а</strong>бых п<strong>о</strong>перечных. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

2,5 мм (ри<strong>с</strong>. 974). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

НАДСЕМЕЙСТВО EOPOLYNEURIDEA<br />

Жил<strong>к</strong>и М и SC <strong>с</strong>л<strong>а</strong>бые, зн<strong>а</strong>чительн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее<br />

р<strong>а</strong>ди<strong>а</strong>льных. Вет<strong>в</strong>ления RS <strong>с</strong>ближенные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>. Имеют<strong>с</strong>я <strong>с</strong>л<strong>а</strong>бые<br />

п<strong>о</strong>перечные между <strong>в</strong>ет<strong>в</strong>ями RS и М. Три<strong>а</strong><strong>с</strong>.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Eopolyneuridae и Musidoromimidae.<br />

СЕМЕЙСТВО EOPOLYNEURIDAE<br />

ROHDENDORF, Fam. nov.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>-<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>о</strong>е<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рьгл<strong>а</strong>, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>с</strong>уж<strong>а</strong>ющее<strong>с</strong>я<br />

<strong>к</strong> <strong>в</strong>ершине. SC не <strong>с</strong><strong>о</strong>единяющ<strong>а</strong>я<strong>с</strong>я <strong>с</strong> С. О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

<strong>о</strong>тдел RS зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че ди<strong>с</strong>т<strong>а</strong>льных<br />

ег<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей. MP не <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>. Три<strong>а</strong><strong>с</strong>. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>.<br />

Eopolyneura Rahdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Е. tenuinervis Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

311<br />

http://jurassic.ru/<br />

Куль). Между SC и R п<strong>о</strong>перечных нет; <strong>к</strong><strong>о</strong>нец<br />

передней <strong>в</strong>ет<strong>в</strong>и RS р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен ближе <strong>к</strong> <strong>с</strong>редней<br />

<strong>в</strong>ет<strong>в</strong>и RS, чем <strong>к</strong> R. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

3 мм (ри<strong>с</strong>. 975). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Pareopolyneura R <strong>о</strong> h d е n d <strong>о</strong> г f, 1961. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. costalis Rohdendorf, 1961; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Между<br />

SC и R имеют<strong>с</strong>я п<strong>о</strong>перечные. К<strong>о</strong>нец передней<br />

<strong>в</strong>ет<strong>в</strong>и RS <strong>с</strong>ближен <strong>с</strong> <strong>к</strong><strong>о</strong>нц<strong>о</strong>м R. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>о</strong>льше 3 мм (ри<strong>с</strong>. 976). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО MUSIDOROMIMIDAE<br />

ROHDENDORF, Fam. nov.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>-<strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й<br />

ширины, не <strong>с</strong>уж<strong>а</strong>ющее<strong>с</strong>я. SC т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, я<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>о</strong>единяющ<strong>а</strong>я<strong>с</strong>я<br />

<strong>с</strong> С. О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS длинный,<br />

п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ный передней <strong>в</strong>ет<strong>в</strong>и. MP з<strong>а</strong>метн<strong>а</strong><br />

<strong>в</strong> <strong>в</strong>иде т<strong>о</strong>н<strong>к</strong><strong>о</strong>й жил<strong>к</strong>и, п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>й <strong>с</strong><br />

CuA. Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Musidoromima Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Н, crassinervis Rohdendorf, sp,<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). CuA <strong>с</strong> я<strong>с</strong>ным <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м перед<br />

<strong>в</strong>ершин<strong>о</strong>й. Имеет<strong>с</strong>я <strong>к</strong>реп<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я <strong>в</strong> <strong>в</strong>иде<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и R. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3 мм (ри<strong>с</strong>. 977). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong><br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.

НАДСЕМЕЙСТВО TIPULIDEA<br />

Жил<strong>к</strong>и <strong>в</strong><strong>с</strong>ех <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных <strong>с</strong>и<strong>с</strong>тем р<strong>а</strong><strong>в</strong>н<strong>о</strong>й т<strong>о</strong>лщины.<br />

Вет<strong>в</strong>ления р<strong>а</strong>ди<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> не<strong>с</strong>ближенные.<br />

Интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> м<strong>а</strong>л<strong>а</strong> или<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; SC <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинн<strong>а</strong>я, б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong><strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. Личин<strong>к</strong>и<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> <strong>в</strong><strong>о</strong>де. Три<strong>а</strong><strong>с</strong> — ныне. В<strong>о</strong><strong>с</strong>емь<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

978. Dictyotipula densa Rohdendorf; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, X 30,0, <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 979. Dlplarchitiputa multimedialis<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X20.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг.<br />

ри<strong>с</strong>.). 980. Architipula radiata Rohdendorf; <strong>к</strong>рыл<strong>о</strong> X 11,0,<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 981. Eollmnobia geinitzi<br />

Handlirsch; <strong>к</strong>рыл<strong>о</strong>, X 5,7, и. юр<strong>а</strong>, Герм<strong>а</strong>ния (Handlirsch, 1906).<br />

982. Trichocera sp.; <strong>к</strong>рыл<strong>о</strong>, Х5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran,<br />

1934). 983. Tricyphona sp.; <strong>к</strong>рыл<strong>о</strong>, х 3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Cur­<br />

Ри<strong>с</strong>. 978—988 Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tipulidea, Eoptychopteridea<br />

312<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ARCHITIPULIDAE HANDLIRSCH, 1906<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> м<strong>а</strong>л<strong>о</strong><strong>с</strong>уженн<strong>о</strong>е. Вет<strong>в</strong>ления<br />

RS и М, п<strong>о</strong>перечн<strong>а</strong>я rm <strong>с</strong>ближены и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; SC<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. Мел<strong>к</strong>ие, реже <strong>с</strong>редней<br />

<strong>в</strong>еличины ф<strong>о</strong>рмы", не <strong>с</strong><strong>в</strong>ыше 9 мм. Три<strong>а</strong><strong>с</strong>. Семь<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Dictyotipula Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — D. densa Rohdendorf, sp. nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

ran, 1934). 984. Tipula sp.; <strong>к</strong>рыл<strong>о</strong>, X 1,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran,<br />

1934). 985. Phalacocera sp.; <strong>к</strong>рыл<strong>о</strong>, X 2,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Curran, 1934). 986. Ptychoptera sp.; <strong>к</strong>рыл<strong>о</strong> X 4,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Curran, 1934). 987. Macrochile spectrum Loew; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х7,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Crampton, 1926).<br />

988. Proptychoptera maculata Handlirsch; <strong>к</strong>рыл<strong>о</strong>, X 12,0, н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния (Hanslirsch, 1939)

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). Ai и<br />

А 2 <strong>с</strong>ближены <strong>в</strong> <strong>в</strong>иде те<strong>с</strong>н<strong>о</strong>й <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

и <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ля <strong>с</strong> <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>быми, н<strong>о</strong><br />

з<strong>а</strong>метными п<strong>о</strong>перечными. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

3 мм (ри<strong>с</strong>. 978). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Diplarchitipula Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — D. muttimedialis Rohdendorf, sp.<br />

nov. <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль)<br />

; MP <strong>в</strong> <strong>в</strong>иде т<strong>о</strong>н<strong>к</strong><strong>о</strong>й <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и <strong>с</strong>з<strong>а</strong>ди<br />

МА; А неп<strong>а</strong>р<strong>а</strong>ллельные; не менее пяти <strong>в</strong>ет<strong>в</strong>ей<br />

М. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4 мм (ри<strong>с</strong>. 979). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Architipula Н a n d 1 i г s <strong>с</strong> h, 1906. Тип р<strong>о</strong>д<strong>а</strong>—Л.<br />

seebachiana Handlirsch, 1906; н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния. Четыре <strong>в</strong>ет<strong>в</strong>и М, передние из них<br />

<strong>с</strong><strong>о</strong>единены <strong>о</strong>бщим <strong>с</strong>тебель<strong>к</strong><strong>о</strong>м. Об<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я<br />

MP <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Одн<strong>а</strong> р<strong>а</strong>ди<strong>о</strong>меди<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т 3 д<strong>о</strong> 9 мм (ри<strong>с</strong>. 980).<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 30 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл., юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong> и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Вне СССР: Eotipula Handlirsch, 1906; Protipula<br />

Handlirsch, 1906; Mesotipula Handlirsch,<br />

1939; Liassotipula Tillyard, 1925.<br />

СЕМЕЙСТВО EOLIMNOBIIDAE ROHDENDORF,<br />

FAM. NOV.<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> м<strong>а</strong>л<strong>о</strong><strong>с</strong>уженн<strong>о</strong>е. П<strong>о</strong>перечн<strong>а</strong>я<br />

rm н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>; SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С;<br />

интермеди<strong>а</strong>льных п<strong>о</strong>перечных нет; три чет<strong>к</strong>ие<br />

А. Средней <strong>в</strong>еличины. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Eolimnobia Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. geinitzi Handlirsch, 1906; н. юр<strong>а</strong>, Герм<strong>а</strong>ния.<br />

Крыл<strong>о</strong> <strong>в</strong> 3,2 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины,<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>е 10,5 мм. С примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> 2<br />

/з длины<br />

<strong>к</strong>рыл<strong>а</strong> (ри<strong>с</strong>. 981). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Герм<strong>а</strong>нии.<br />

СЕМЕЙСТВО TRICHOCERIDAE EDWARDS, 1928<br />

. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е. Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

М и п<strong>о</strong>перечн<strong>а</strong>я rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены зн<strong>а</strong>чительн<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины 1<br />

<strong>к</strong>рыл<strong>а</strong>; имеют<strong>с</strong>я<br />

интермеди<strong>а</strong>льные п<strong>о</strong>перечные, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бляющие<br />

ячей<strong>к</strong>у; А2 у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>а</strong>я (ри<strong>с</strong>. 982). Личин<strong>к</strong>и<br />

<strong>с</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>к</strong><strong>а</strong>п<strong>с</strong>ул<strong>о</strong>й, <strong>в</strong> <strong>в</strong><strong>о</strong>де.<br />

Средней <strong>в</strong>еличины <strong>к</strong>рылья. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Четыре р<strong>о</strong>д<strong>а</strong>, <strong>о</strong>дин из них из<strong>в</strong>е<strong>с</strong>тен <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО L1MONIIDAE HENDEL, 1936<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>уженн<strong>о</strong>е. SC <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я<br />

<strong>с</strong> С; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М, п<strong>о</strong>перечные rm и mcu<br />

<strong>с</strong>ближены и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины<br />

<strong>к</strong>рыл<strong>а</strong>; А 2 длинн<strong>а</strong>я (ри<strong>с</strong>. 983). Г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>а</strong>я<br />

<strong>к</strong><strong>а</strong>п<strong>с</strong>ул<strong>а</strong> личин<strong>о</strong><strong>к</strong> не<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я. Преимуще­<br />

<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины. Личин<strong>к</strong>и<br />

<strong>в</strong> <strong>в</strong><strong>о</strong>де и других <strong>в</strong>л<strong>а</strong>жных <strong>с</strong>ред<strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Б<strong>о</strong>г<strong>а</strong>т<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне: <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из третичных<br />

<strong>о</strong>тл<strong>о</strong>жений, из них 24 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО TIPULIDAE LEACH, 1815<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е. SC н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М, rm и mcu р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

д<strong>а</strong>ле<strong>к</strong><strong>о</strong> з<strong>а</strong> <strong>с</strong>ередин<strong>о</strong>й <strong>к</strong>рыл<strong>а</strong>; А 2<br />

длинн<strong>а</strong>я (ри<strong>с</strong>. 984). Крупные, реже <strong>с</strong>редней<br />

<strong>в</strong>еличины. Личин<strong>к</strong>и <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х и <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Б<strong>о</strong>г<strong>а</strong>т<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне — примерн<strong>о</strong> де<strong>с</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из<br />

них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CYLINDROTOMIDAE OSTEN-SACKEN,<br />

1869<br />

[nom. transl. Czizek, 1931<br />

(ex Cylindrotomina Osten-Sacken, 1869)]<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>уженн<strong>о</strong>е. SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong> <strong>в</strong> переп<strong>о</strong>н<strong>к</strong>е <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М и п<strong>о</strong>перечные rm и mcu р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>; R <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> переднюю <strong>в</strong>ет<strong>в</strong>ь RS; CuA н<strong>а</strong> <strong>к</strong><strong>о</strong>нце рез<strong>к</strong><strong>о</strong> з<strong>а</strong>гнут<strong>а</strong><br />

н<strong>а</strong>з<strong>а</strong>д (ри<strong>с</strong>. 985). Г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>а</strong>я <strong>к</strong><strong>а</strong>п<strong>с</strong>ул<strong>а</strong> личин<strong>о</strong><strong>к</strong><br />

не<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я. Средней <strong>в</strong>еличины 1<br />

.<br />

Личин<strong>к</strong>и н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х и <strong>в</strong>л<strong>а</strong>жных<br />

луг<strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Бедн<strong>о</strong>е <strong>в</strong>ид<strong>а</strong>ми <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>о</strong>дин р<strong>о</strong>д из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

и не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО LIRIOPEIDAE HENDEL, 1936<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е. SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> С; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М и п<strong>о</strong>перечные rm и mcu<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>; интермеди<strong>а</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong>и нет (ри<strong>с</strong>. 986). Г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>а</strong>я <strong>к</strong><strong>а</strong>п<strong>с</strong>ул<strong>а</strong><br />

личин<strong>к</strong>и рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я. Личин<strong>к</strong>и <strong>в</strong><br />

<strong>в</strong><strong>о</strong>де. П<strong>а</strong>ле<strong>о</strong>ген —• ныне. Бедн<strong>о</strong>е <strong>в</strong>ид<strong>а</strong>ми, рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>; <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

из<strong>в</strong>е<strong>с</strong>тен <strong>о</strong>дин р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

СЕМЕЙСТВО TANYDERIDAE OSTEN-SACKEN, 1879<br />

[nom. transl. Alexander, 1920<br />

(ex Tanyderina Osten-Sacken, 1879)]<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рыл<strong>а</strong> умеренн<strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е. SC<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С и <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я п<strong>о</strong>перечн<strong>о</strong>й <strong>с</strong> R;<br />

имеют<strong>с</strong>я четыре <strong>в</strong>ет<strong>в</strong>и RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющие <strong>в</strong> <strong>к</strong>р<strong>а</strong>й<br />

<strong>к</strong>рыл<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М и п<strong>о</strong>перечные rm и mcu<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> или пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее;<br />

А 2 нея<strong>с</strong>н<strong>а</strong> (ри<strong>с</strong>. 987). Личин<strong>к</strong>и <strong>в</strong><strong>о</strong><br />

313<br />

http://jurassic.ru/

<strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й п<strong>о</strong>ч<strong>в</strong>е. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Бедн<strong>о</strong>е <strong>в</strong>и«<br />

д<strong>а</strong>ми, рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>;<br />

<strong>о</strong>дин р<strong>о</strong>д из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО EOPTYCHOPTERIDEA<br />

Жил<strong>к</strong>и <strong>в</strong><strong>с</strong>ех <strong>с</strong>и<strong>с</strong>тем р<strong>а</strong><strong>в</strong>н<strong>о</strong>й т<strong>о</strong>лщины'; <strong>в</strong>ет<strong>в</strong>ления<br />

R не<strong>с</strong>ближе<strong>в</strong>ные; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупн<strong>а</strong>я, п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я;<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

<strong>к</strong>рыл<strong>а</strong>. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>. Юр<strong>а</strong>.<br />

СЕМЕЙСТВО EOPTYCHOPTERIDAE HANDLIRSCH,<br />

1906<br />

Крыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, примерн<strong>о</strong> <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> длиннее<br />

ширины, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием (ри<strong>с</strong>. 988),<br />

4—б мм. Юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>. Вне СССР: Eoptychoptera<br />

Handlirsch, 1906 и Propty chapter <strong>а</strong><br />

Handlirsch, 1939.<br />

НАДСЕМЕЙСТВО TANYDEROPHRYNEIDEA<br />

M, RS и Си <strong>с</strong>л<strong>а</strong>бее С, SC и R; <strong>в</strong>ет<strong>в</strong>ления R<br />

не<strong>с</strong>ближенные; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>к</strong>реп<strong>к</strong>ий, <strong>о</strong>тх<strong>о</strong>дящий<br />

<strong>о</strong>т R; MP <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т МА; интермеди<strong>а</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong>и нет; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Н<strong>о</strong>ги бег<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>. Юр<strong>а</strong>.<br />

СЕМЕЙСТВО TANYDEROPHRYNEIDAE<br />

ROHDENDORF, FAM. NOV.<br />

Крыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, б<strong>о</strong>лее чем <strong>в</strong> 2,75 р<strong>а</strong>з<strong>а</strong><br />

длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии уз<strong>к</strong><strong>о</strong>е;<br />

SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С, <strong>с</strong><strong>о</strong>единяя<strong>с</strong>ь н<strong>а</strong> <strong>к</strong><strong>о</strong>нце и <strong>с</strong> R<br />

п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и; пять <strong>в</strong>ет<strong>в</strong>ей<br />

RS и не менее четырех <strong>в</strong>ет<strong>в</strong>ей М; СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я.<br />

Юр<strong>а</strong>. ОДИН р<strong>о</strong>д.<br />

Tanyderophryne Rohdendorf, gen. nov.<br />

ТИП р<strong>о</strong>д<strong>а</strong> — Т. multinervis Rohdendorf, sp. nov.;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

З<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>о</strong>бъединен<strong>а</strong> <strong>с</strong> <strong>с</strong>и<strong>с</strong>тем<strong>о</strong>й М;<br />

четыре передние <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>бр<strong>а</strong>зуют д<strong>в</strong>е п<strong>а</strong>ры<br />

п<strong>а</strong>р<strong>а</strong>ллельных жил<strong>о</strong><strong>к</strong>; MP <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т<br />

СиА и <strong>о</strong>бр<strong>а</strong>зует не менее д<strong>в</strong>ух <strong>в</strong>ет<strong>в</strong>ей: эт<strong>а</strong><br />

жил<strong>к</strong><strong>а</strong> не <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> МА. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3,5 мм, тел<strong>а</strong> — <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4,2 мм (ри<strong>с</strong>. 989). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

НАДСЕМЕЙСТВО DIXIDEA<br />

Крыл<strong>о</strong> <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины,<br />

умеренн<strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. Р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ление<br />

М и п<strong>о</strong>перечн<strong>а</strong>я жилй<strong>а</strong> rm <strong>с</strong>ближенные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>с</strong>р<strong>а</strong>зу з<strong>а</strong> <strong>с</strong>ередин<strong>о</strong>й <strong>к</strong>рыл<strong>а</strong>. Д<strong>в</strong>е<br />

<strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>бр<strong>а</strong>зуют п<strong>а</strong>ру п<strong>а</strong>р<strong>а</strong>ллельных, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

з<strong>а</strong>гнутых н<strong>а</strong>з<strong>а</strong>д жил<strong>о</strong><strong>к</strong>. Юр<strong>а</strong> — ныне.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Dixamimidae, Dixidae.<br />

314<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО DIXAMIMIDAE ROHDENDORF, 1957<br />

SC длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>. Имеет<strong>с</strong>я х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong>.<br />

С и R з<strong>а</strong>метн<strong>о</strong> т<strong>о</strong>лще других жил<strong>о</strong><strong>к</strong>. Н<strong>о</strong>ги<br />

<strong>к</strong>реп<strong>к</strong>ие, бег<strong>а</strong>тельные. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Dixamima Rohdendorf, 1951. Тип р<strong>о</strong>д<strong>а</strong>—<br />

D. villosa- Rohdendorf, 1951; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R прям<strong>а</strong>я;<br />

rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> у <strong>с</strong><strong>а</strong>м<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния з<strong>а</strong>дней<br />

<strong>в</strong>ет<strong>в</strong>и RS; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и М имеет<br />

<strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 5 мм<br />

(ри<strong>с</strong>. 990<strong>а</strong>, б). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО DIXIDAE VAN DER VULP, 1877<br />

[nom. transl. Brauer, 1880<br />

(ex Dixinae van der Vulp, 1877)]<br />

SC меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины' <strong>к</strong>рыл<strong>а</strong>; интермеди<strong>а</strong>льные<br />

п<strong>о</strong>перечные <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, и ячей<strong>к</strong><strong>а</strong> не<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>й т<strong>о</strong>лщины;<br />

н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong><strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. Личин<strong>к</strong>и <strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х<br />

(ри<strong>с</strong>. 991). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, <strong>о</strong>дин из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО CULICIDEA<br />

Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, не менее чем <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> длиннее<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ей ширины, умеренн<strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии;<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ление М и п<strong>о</strong>перечн<strong>а</strong>я rm не <strong>с</strong>ближены;<br />

д<strong>в</strong>е передние <strong>в</strong>ет<strong>в</strong>и RS не <strong>о</strong>бр<strong>а</strong>зуют п<strong>а</strong>ры<br />

из<strong>о</strong>гнутых п<strong>а</strong>р<strong>а</strong>ллельных жил<strong>о</strong><strong>к</strong>. Личин<strong>к</strong>и <strong>в</strong><br />

<strong>в</strong><strong>о</strong>де. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Chaoboridae,<br />

Culicidae.<br />

СЕМЕЙСТВО CHAOBORIDAE HENDEL, 1936<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и мяг<strong>к</strong>ий. Крылья без<br />

шир<strong>о</strong><strong>к</strong>их чешуе<strong>к</strong>, лишь <strong>с</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>.992).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Немн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), <strong>о</strong>дин (Chaoborus Lichtwardt)<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Ю. Фр<strong>а</strong>нции и <strong>о</strong>дин <strong>в</strong> не<strong>о</strong>гене<br />

Ю. Азии (янт<strong>а</strong>рь Бирмы 1<br />

).<br />

СЕМЕЙСТВО CULICIDAE BILLBERGH, 1820<br />

[nom. transl. Stephens, 1829 (ex Culicides Billbergh, 1820)]<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong>ильн<strong>о</strong> удлиненный; <strong>к</strong>рылья п<strong>о</strong><br />

жил<strong>к</strong><strong>а</strong>м п<strong>о</strong><strong>к</strong>рыты шир<strong>о</strong><strong>к</strong>ими чешуй<strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 993). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них три <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

3. Е<strong>в</strong>р<strong>о</strong>пы 1<br />

и С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО PSYCHODIDEA<br />

Крылья <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем, <strong>с</strong>уженные<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, п<strong>о</strong><strong>к</strong>рытые гу<strong>с</strong>тыми <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми-щетин<strong>к</strong><strong>а</strong>ми;<br />

SC <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

1<br />

<strong>к</strong>рыл<strong>а</strong>; п<strong>о</strong>перечн<strong>а</strong>я rm пл<strong>о</strong>х<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен-

Ри<strong>с</strong>. 989—993. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tanyderophryneidea, Dixidea, Culicidea<br />

989. Tanyderophryne multinervis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X 15,0, <strong>в</strong>. <strong>о</strong>бщий <strong>в</strong>ид, х5,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, 1934). 992. Mochlonyx<br />

юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.). 990. Dlxamima villosa Roh- sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 7,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, 1934). 993. Culex<br />

dendorf; А _ <strong>о</strong>бщий <strong>в</strong>Ид и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>г<strong>о</strong> X 7,0; Б — ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, pipiens<br />

7<br />

Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, X 0. <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Lindner,<br />

X 20,0; <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1951). 991. Dixa sp.! 1932)<br />

315<br />

http://jurassic.ru/

Ри<strong>с</strong>. 994—998. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Psychodidea<br />

994. Psychoda sp.; <strong>к</strong>рыл<strong>о</strong>, X29.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hendel, 1928) 1934). 997. Phlebotomus sp.; <strong>к</strong>рыл<strong>о</strong>, 15,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Brues, Melan-<br />

995. Trichomyia sp.; <strong>к</strong>рыл<strong>о</strong>, X23.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, der, Carpenter, 1954). 998. Bruchomyla sp.; <strong>к</strong>рыл<strong>о</strong>, Xl3,0,<br />

1934). 996. Sycorax sp.; <strong>к</strong>рыл<strong>о</strong>, X 16,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, <strong>с</strong><strong>о</strong><strong>в</strong>рем., А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Brues, Melander, Carpenter, 1954)<br />

п<strong>а</strong>я, пр<strong>о</strong>д<strong>о</strong>льные жил<strong>к</strong>и не<strong>с</strong>мещенные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> н<strong>а</strong> <strong>в</strong><strong>с</strong>ей пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Psychodidae, Trichomyiidae,<br />

Phlebotomidae и Nemopalpidae.<br />

СЕМЕЙСТВО PSYCHODIDAE NEWMAN, 1834<br />

[nom. transl. Bigot, 1854 (ex Psychodites Newman, 1834)]<br />

Крылья шир<strong>о</strong><strong>к</strong>ие, н<strong>а</strong> <strong>в</strong>ершине з<strong>а</strong><strong>о</strong><strong>с</strong>тренные<br />

или угл<strong>о</strong><strong>в</strong><strong>а</strong>тые; СиА длинн<strong>а</strong>я, п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я<br />

М 4 . SC п<strong>о</strong>чти п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я<br />

(ри<strong>с</strong>. 994). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Личин<strong>к</strong>и <strong>в</strong><strong>о</strong><br />

<strong>в</strong>л<strong>а</strong>жных <strong>с</strong>ред<strong>а</strong>х. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО TRICHOMYIIDAE TONNIOR, 1922<br />

[nom. transl. Rohdendorf, hie<br />

(ex Trichomyinae Tonnior, 1922)]<br />

Крылья шир<strong>о</strong><strong>к</strong>ие, н<strong>а</strong> <strong>в</strong>ершине притуплённые.<br />

RS <strong>с</strong> д<strong>в</strong>умя или тремя <strong>в</strong>ет<strong>в</strong>ями; СиА <strong>о</strong>чень<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я ИЛИ удлиненн<strong>а</strong>я; SC х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метн<strong>а</strong>я,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>с</strong> С и <strong>с</strong><strong>о</strong>единенн<strong>а</strong>я <strong>с</strong> R п<strong>о</strong>перечн<strong>о</strong>й<br />

(ри<strong>с</strong>. 995, 996). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янг<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО PHLEBOTOMIDAE RONDANI, 1840<br />

[nom. transl. Rohdendorf, hie<br />

(ex Phlebotominae Rondani, 1840)]<br />

Крылья уз<strong>к</strong>ие, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренные н<strong>а</strong> <strong>в</strong>ершине;<br />

RS <strong>с</strong> четырьмя <strong>в</strong>ет<strong>в</strong>ями; СиА <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

316<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рыл<strong>а</strong>;<br />

SC х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метн<strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С и<br />

<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> R п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й (ри<strong>с</strong>.997).<br />

Очень мел<strong>к</strong>ие, <strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong>ущие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. Личин<strong>к</strong>и<br />

<strong>в</strong> гниющих <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong>х и п<strong>о</strong>ч<strong>в</strong>е. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО NEMOPALPIDAE EDWARDS, 1921<br />

[nom. transl. Rohdendorf, hie<br />

(ex Nemopalpinae Edwards, 1921)]<br />

Крылья не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> уз<strong>к</strong>ие, <strong>с</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>й<br />

<strong>в</strong>ершин<strong>о</strong>й, длинн<strong>о</strong>й SC, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> меньшей<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й СиА, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющей<br />

п<strong>о</strong>д прямым угл<strong>о</strong>м <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> (ри<strong>с</strong>. 998).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Средней <strong>в</strong>еличины, <strong>о</strong>чень<br />

ред<strong>к</strong>ие рели<strong>к</strong>т<strong>о</strong><strong>в</strong>ые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Южн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>луш<strong>а</strong>рия.<br />

Р<strong>а</strong>з<strong>в</strong>итие неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>. Один р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы 1<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО CHIRONOMIDEA<br />

Крылья <strong>с</strong> прямым передним <strong>к</strong>р<strong>а</strong>ем и рез<strong>к</strong><strong>о</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; имеет<strong>с</strong>я х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я фр<strong>а</strong>гм<strong>а</strong>; передние жил<strong>к</strong>и (С<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> R) з<strong>а</strong>метн<strong>о</strong> б<strong>о</strong>лее <strong>к</strong>реп<strong>к</strong>ие, чем<br />

з<strong>а</strong>дние (М, Си и А); жил<strong>к</strong><strong>а</strong> С не з<strong>а</strong>х<strong>о</strong>дит н<strong>а</strong><br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>. Личин<strong>к</strong>и <strong>в</strong> <strong>в</strong><strong>о</strong>де, реже<br />

<strong>с</strong>реди <strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>ти. Три<strong>а</strong><strong>с</strong> — ныне.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Architendipedidae, Protendipedidae,<br />

Chironomidae, Ceratopogonidae, Simuliidae.<br />

http://jurassic.ru/

СЕМЕЙСТВО ARCH ITENDIPED-I DAE<br />

ROHDENDORF, FAM. NOV.<br />

Крылья удлиненные, <strong>с</strong> чет<strong>к</strong>им, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличимым<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием. Имеют<strong>с</strong>я п<strong>о</strong>перечные<br />

жил<strong>к</strong>и между R и М, М и СиА <strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

п<strong>о</strong>ле и между <strong>в</strong>ет<strong>в</strong>ями RS. Три<strong>а</strong><strong>с</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Architendipes Rohdendorf, gen. nov.<br />

Тип. р<strong>о</strong>д<strong>а</strong> — A. tshernovskiji Rohdendorf, sp.<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль)<br />

. Середин<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля <strong>с</strong> п<strong>о</strong>перечными;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные ч<strong>а</strong><strong>с</strong>ти R и М <strong>к</strong>реп<strong>к</strong>ие, <strong>с</strong>литые;<br />

RS <strong>о</strong>бр<strong>а</strong>зует длинную переднюю <strong>в</strong>ет<strong>в</strong>ь,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую из д<strong>в</strong>ух длинных п<strong>а</strong>р<strong>а</strong>ллельных<br />

<strong>в</strong>ет<strong>в</strong>ей. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 4,5 мм (ри<strong>с</strong>. 999). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Palaeotendipes Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. alexii Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Середин<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля без п<strong>о</strong>перечных;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные ч<strong>а</strong><strong>с</strong>ти М и RS <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие,<br />

нея<strong>с</strong>ные; передняя <strong>в</strong>ет<strong>в</strong>ь RS из<strong>о</strong>гнут<strong>а</strong>я, <strong>с</strong>н<strong>а</strong>бженн<strong>а</strong>я<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце неб<strong>о</strong>льшим |р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 5 мм (ри<strong>с</strong>. 1000). Один <strong>в</strong>ид.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

•999. Architendipes tshernovskiji Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х15.0, <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 1000. Palaeotendipes alexii<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х12.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>. Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.).<br />

1001. Protendipes dasypterus Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид, х 14,0,<br />

<strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.). 1002. Anatopynia sp.;<br />

Ри<strong>с</strong>. 999—1005. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Chironomidea<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PROTENDIPEDIDAE ROHDENDORF,<br />

FAM. NOV.<br />

Крылья удлиненные, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>бым, нер<strong>а</strong>зличимым<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием и гу<strong>с</strong>тым п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong>м из темны'х<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, <strong>о</strong>чень длинные,<br />

передний мет<strong>а</strong>т<strong>а</strong>рз <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Protendipes Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. dasypterus Rohdendorf, sp. nov.; <strong>в</strong>.<br />

юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Брюш<strong>к</strong><strong>о</strong> удлиненн<strong>о</strong>е, <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>е. З<strong>а</strong>дние бедр<strong>а</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> длиннее з<strong>а</strong>дних г<strong>о</strong>леней. Длин<strong>а</strong><br />

тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3,25 мм, <strong>к</strong>рылье<strong>в</strong> — 3,5 мм, з<strong>а</strong>дних<br />

бедер— 1,35 мм (ри<strong>с</strong>. 1001). Один <strong>в</strong>ид. В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО CHIRONOMIDAE MACQUART, 1838<br />

[nom. transl. Haliday, 1851<br />

(ex Chironomides Macquart, 1838)]<br />

(Tendipedidae Grflnberg, 1910; Hendel, 1928)<br />

Крылья удлиненные, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>бым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием,<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метным <strong>в</strong>близи переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я, г<strong>о</strong>лые или <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими и т<strong>о</strong>н<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

Н<strong>о</strong>ги, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> передние, <strong>о</strong>чень длин-<br />

100Б<br />

<strong>к</strong>рыл<strong>о</strong>, х9,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Brues, Melander, Carpenter,<br />

1954). 1003. Chironomus sp.; <strong>к</strong>рыл<strong>о</strong>, X6,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Brues,<br />

Melander, Carpenter, 1954). 1004. Helea sp.; <strong>к</strong>рыл<strong>о</strong>, X 17,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, 1934). 1005. Slmulium sp.; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 14,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Рубц<strong>о</strong><strong>в</strong>, 1959)

ные и т<strong>о</strong>н<strong>к</strong>ие. В<strong>с</strong>ег<strong>о</strong> д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и М (Mi и М4),<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передняя пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я и <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я<br />

<strong>о</strong>т з<strong>а</strong>дней. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>г<strong>а</strong>т<strong>о</strong>е <strong>в</strong>ид<strong>а</strong>ми<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненн<strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>; из <strong>с</strong>еми п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> три.<br />

ПОДСЕМЕЙСТВО TANYPODINAE SKUSE, 1889<br />

[nom. transl. Enderlein, 1920 (ex Tanypina Skuse, 1889)]<br />

(Pelopiina Hendel, 1928)<br />

П<strong>о</strong>перечн<strong>а</strong>я mcu имеет<strong>с</strong>я; RS х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong><br />

и <strong>с</strong><strong>о</strong>единен<strong>а</strong> <strong>с</strong> R (ри<strong>с</strong>. 1002). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ПОДСЕМЕЙСТВО CHIRONOMINAE MACQUART, 1838<br />

[nom. transl. Kieffer, 1906<br />

(ex Chironomides Macquart, 1838)]<br />

П<strong>о</strong>перечн<strong>а</strong>я mcu <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, M4 <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong><br />

<strong>о</strong>т Mi и имеет <strong>в</strong>ид <strong>в</strong>ет<strong>в</strong>и CuA; передний мет<strong>а</strong>т<strong>а</strong>рз<br />

длиннее г<strong>о</strong>лени; п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> генит<strong>а</strong>льных<br />

прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен н<strong>а</strong>з<strong>а</strong>д<br />

(ри<strong>с</strong>. 1003). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Мн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), 3. Е<strong>в</strong>р<strong>о</strong>пы,<br />

С. Амери<strong>к</strong>и, и не<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ПОДСЕМЕЙСТВО ORTHOCLADIINAE KIEFFER, 1911<br />

П<strong>о</strong>перечн<strong>а</strong>я mcu <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; передний мет<strong>а</strong>т<strong>а</strong>рз<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>лени; п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> генит<strong>а</strong>льных<br />

прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong> з<strong>а</strong>гнут <strong>в</strong>перед. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CERATOPOGONIDAE SKUSE, 1889<br />

[nom. transl. Malloch, 1917<br />

(ex Ceratopogonina Skuse, 1889)]<br />

(Heleidae Speiser, 1910)<br />

Умеренн<strong>о</strong> удлиненные, реже длинные<br />

<strong>к</strong>рылья <strong>с</strong> <strong>о</strong>чень рез<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием:<br />

р<strong>а</strong>ди<strong>а</strong>льные жил<strong>к</strong>и зн<strong>а</strong>чительн<strong>о</strong> т<strong>о</strong>лще<br />

других; три <strong>в</strong>ет<strong>в</strong>и М, д<strong>в</strong>е передние <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>в</strong>ил<strong>к</strong>у; н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и цеп<strong>к</strong>ие, <strong>с</strong> длинными<br />

<strong>к</strong><strong>о</strong>гтями и <strong>в</strong>здутыми бедр<strong>а</strong>ми. Личин<strong>к</strong>и<br />

<strong>в</strong> <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х и <strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й п<strong>о</strong>ч<strong>в</strong>е (ри<strong>с</strong>. 1004). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненн<strong>о</strong>е <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>; <strong>о</strong>дин р<strong>о</strong>д из<strong>в</strong>е<strong>с</strong>тен<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Герм<strong>а</strong>нии.<br />

СЕМЕЙСТВО SIMULIIDAE NEWMAN, 1834<br />

[nom. transl. Williston, 1908<br />

(ex Simuliites Newman, 1834)]<br />

Б<strong>о</strong>льшие, <strong>о</strong>чень шир<strong>о</strong><strong>к</strong>ие <strong>в</strong>еер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные<br />

<strong>к</strong>рылья <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием. Три<br />

<strong>в</strong>ет<strong>в</strong>и М, <strong>с</strong>л<strong>а</strong>бые; р<strong>а</strong>ди<strong>а</strong>льные жил<strong>к</strong>и <strong>к</strong>реп<strong>к</strong>ие,<br />

318<br />

<strong>с</strong>ближенные. Тел<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и<br />

цеп<strong>к</strong>ие (ри<strong>с</strong>. 1005). Личин<strong>к</strong>и <strong>в</strong> бы<strong>с</strong>тр<strong>о</strong>те<strong>к</strong>ущих<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х, <strong>к</strong>рыл<strong>а</strong>тые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые — <strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong>ы<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО RHAETOMYIIDEA<br />

Крылья <strong>с</strong> прямым передним <strong>к</strong>р<strong>а</strong>ем и р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пределенным п<strong>о</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е <strong>к</strong>реп<strong>к</strong>имжил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием.<br />

Х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> фр<strong>а</strong>гм<strong>а</strong>, <strong>с</strong><strong>о</strong>единяющ<strong>а</strong>я<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>лы R и CuA; жил<strong>к</strong><strong>а</strong> С з<strong>а</strong>х<strong>о</strong>дит<br />

н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й; п<strong>о</strong>перечные rm и mcu <strong>о</strong>чень,<br />

<strong>к</strong>реп<strong>к</strong>ие; А <strong>с</strong>л<strong>а</strong>бые, п<strong>о</strong>чти редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные.<br />

Три<strong>а</strong><strong>с</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhaetomyiidae.<br />

СЕМЕЙСТВО RHAETOMYIIDAE ROHDENDORF,<br />

FAM. NOV.<br />

SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С; RS<br />

<strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC и д<strong>а</strong>ет три<br />

<strong>в</strong>ет<strong>в</strong>и, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передние <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>бщий<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>; М д<strong>а</strong>ет д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

SC; CuA <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>с</strong> изл<strong>о</strong>м<strong>о</strong>м. Три<strong>а</strong><strong>с</strong>. Один<br />

р<strong>о</strong>д,<br />

Rhaetomyia Rohdendorf, gen. nov. Типр<strong>о</strong>д<strong>а</strong>—<br />

Rh. necopinata Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>с</strong>л<strong>а</strong>бее R и CuA; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь М <strong>о</strong>бр<strong>а</strong>зует р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>, з<strong>а</strong>дняя<br />

з<strong>с</strong><br />

CuA<br />

Ри<strong>с</strong>. 1006. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhaetomyiidea. Rhaetomyia necopinata<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X 17,5, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия-<br />

(<strong>о</strong>риг. ри<strong>с</strong>.)<br />

пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; CuP т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, идущ<strong>а</strong>я <strong>в</strong>пл<strong>о</strong>ть д<strong>о</strong> <strong>с</strong>ередины<br />

п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> CuA. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3,5 мм, ширин<strong>а</strong> 1,6 мм (ри<strong>с</strong>. 1006). Один <strong>в</strong>ид.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

НАДСЕМЕЙСТВО MESOPHANTASMATIDEA<br />

Крылья <strong>о</strong>чень уз<strong>к</strong>ие, <strong>с</strong> прямым передним<br />

<strong>к</strong>р<strong>а</strong>ем и <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>с</strong>уженн<strong>о</strong>й б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью,,<br />

имеющей <strong>в</strong>ид <strong>с</strong>тебель<strong>к</strong><strong>а</strong>; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>реп<strong>к</strong><strong>о</strong>е,.<br />

http://jurassic.ru/

не<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; <strong>в</strong>ет<strong>в</strong>ления жил<strong>о</strong><strong>к</strong> не<strong>с</strong>ближенные;<br />

R и СиА т<strong>о</strong>лще других жил<strong>о</strong><strong>к</strong>". Юр<strong>а</strong>.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesophantasmatidae.<br />

СЕМЕЙСТВО MESOPHANTASMATIDAE<br />

ROHDENDORF, FAM. NOV.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>бх<strong>о</strong>дит <strong>в</strong><strong>с</strong>ё <strong>к</strong>рыл<strong>о</strong>;<br />

<strong>к</strong>рылыш<strong>к</strong><strong>о</strong> не <strong>в</strong>ыр<strong>а</strong>жен<strong>о</strong>, <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть м<strong>а</strong>л<strong>а</strong>,<br />

<strong>в</strong>ершин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> туп<strong>а</strong>я; А <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Mesophantasma Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. tipuliforme Rohdendorf, sp.<br />

RS<br />

R sc<br />

Ри<strong>с</strong>. 1007. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesophantasmatidea. Mesophantasma<br />

tipuliforme Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x 9,2, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(<strong>о</strong>риг. ри<strong>с</strong>.)<br />

nov.; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>^<br />

т<strong>а</strong>у); RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> и<br />

имеет д<strong>в</strong><strong>о</strong>йн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние: <strong>о</strong>бр<strong>а</strong>зует три <strong>в</strong>ет<strong>в</strong>и;<br />

М делит<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> и ди<strong>с</strong>т<strong>а</strong>льнее,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из трех <strong>в</strong>ет<strong>в</strong>ей; СиР т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, н<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

з<strong>а</strong>метн<strong>а</strong>я. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 8 мм (ри<strong>с</strong>. 1007).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

И НФРАОТРЯД<br />

В I В I О N О М О R Р Н А<br />

RS <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями, передние из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> или <strong>в</strong> R. Крыл<strong>о</strong><br />

м<strong>а</strong>л<strong>о</strong>удлиненн<strong>о</strong>е, <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным,<br />

<strong>к</strong>рупным <strong>в</strong>ершинным <strong>о</strong>тдел<strong>о</strong>м. Антенны <strong>с</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мными<br />

члени<strong>к</strong><strong>а</strong>ми. Р<strong>а</strong>з<strong>в</strong>итие не <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong><br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>ми. Три<strong>а</strong><strong>с</strong> — ныне. Де<strong>в</strong>ять н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Pleciodictyidea, Protoligoneuridea, Fungivoridea,<br />

Bibionidea, Bolitophilidea, Scatopsidea,<br />

Rhyphidea, Phragmoligoneuridea и Cecidomyiidea.<br />

НАДСЕМЕЙСТВО PLECIODICTYIDEA<br />

Между M и CuP имеют<strong>с</strong>я пр<strong>а</strong><strong>в</strong>ильные п<strong>о</strong>перечные<br />

жил<strong>к</strong>и; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>л<strong>а</strong>б<strong>о</strong>е, лишь R и<br />

СиР чет<strong>к</strong>ие и я<strong>с</strong>ные. Три<strong>а</strong><strong>с</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pleciodictyidae.<br />

СЕМЕЙСТВО PLECIODICTY1DAE ROHDENDORF,<br />

FAM. NOV.<br />

RS не<strong>с</strong>ет ряд <strong>в</strong>ет<strong>в</strong>ей, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленных <strong>в</strong>перед<br />

и имеющих <strong>в</strong>ид непр<strong>а</strong><strong>в</strong>ильных п<strong>о</strong>перечных;<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong> М б<strong>о</strong>г<strong>а</strong>т<strong>а</strong>я, н<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я и пл<strong>о</strong>х<strong>о</strong> р<strong>а</strong>зличим<strong>а</strong>я.<br />

Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Pleciodictya Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. tnodesta Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Не менее трех <strong>в</strong>ет<strong>в</strong>ей RS <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> R;<br />

п<strong>о</strong>перечные rm <strong>в</strong> чи<strong>с</strong>ле трех <strong>к</strong><strong>о</strong><strong>с</strong>ых жил<strong>о</strong><strong>к</strong>; SC<br />

<strong>с</strong>л<strong>а</strong>б<strong>а</strong>я. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,2 мм (ри<strong>с</strong>. 1008). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

НАДСЕМЕЙСТВО PROTOLIGONEURIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле <strong>с</strong> рез<strong>к</strong>им <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong>м; имеет<strong>с</strong>я<br />

з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чн<strong>а</strong>я фр<strong>а</strong>гм<strong>а</strong>; <strong>в</strong>ет<strong>в</strong>и RS <strong>с</strong>л<strong>о</strong>жные и<br />

нея<strong>с</strong>ные; д<strong>в</strong>е чет<strong>к</strong>ие rm, <strong>о</strong>тд<strong>а</strong>ленные друг <strong>о</strong>т<br />

друг<strong>а</strong>. Три<strong>а</strong><strong>с</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Protoligoneuridae.<br />

СЕМЕЙСТВО PROTOLIGONEURIDAE ROHDENDORF,.<br />

FAM. NOV.<br />

M <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из трех чет<strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей, <strong>в</strong>ых<strong>о</strong>дящих<br />

из <strong>о</strong>бщег<strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>; А д<strong>в</strong>е, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты.<br />

Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Protoligoneura Rohdendorf, gen, nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. fusicosta Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). SC т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л Mi+2 <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

2,5 мм (ри<strong>с</strong>. 1009). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong><br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл..<br />

НАДСЕМЕЙСТВО FUNGIVORIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>. Фр<strong>а</strong>гм<strong>а</strong> нер<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

б<strong>а</strong>з<strong>а</strong>льный уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> СиР не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен.<br />

Р<strong>а</strong>змеры мел<strong>к</strong>ие. Н<strong>о</strong>ги бег<strong>а</strong>тельные или т<strong>о</strong>н<strong>к</strong>ие.<br />

Антенны т<strong>о</strong>н<strong>к</strong>ие и <strong>с</strong> г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мными члени<strong>к</strong><strong>а</strong>ми.<br />

Интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

М из трех <strong>в</strong>ет<strong>в</strong>ей (Mi, М 2 и М 4). Три<strong>а</strong><strong>с</strong> — ныне.<br />

16 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них лишь <strong>о</strong>дн<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>в</strong>.<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PALAEOPLECIIDAE ROHDENDORF,<br />

FAM. NOV.<br />

R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии прям<strong>а</strong>я; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т<br />

R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й чет<strong>в</strong>ерти <strong>к</strong>рыл<strong>а</strong> и не<strong>с</strong>ет длиннуюпереднюю<br />

<strong>в</strong>ет<strong>в</strong>ь, п<strong>а</strong>р<strong>а</strong>ллельную <strong>к</strong><strong>о</strong>нцу R; между<br />

С, R и передней <strong>в</strong>ет<strong>в</strong>ью М имеют<strong>с</strong>я чет<strong>к</strong>ие<br />

п<strong>о</strong>перечные; з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь М прям<strong>а</strong>я и я<strong>с</strong>н<strong>а</strong>я.<br />

Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Palaeoplecia Rohdendorf, gen. nov. Типр<strong>о</strong>д<strong>а</strong>—<br />

P. rhaetica Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>. И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). SC р<strong>а</strong><strong>в</strong>н<strong>а</strong> примерн<strong>о</strong> трети длины <strong>к</strong>ры-<br />

319<br />

http://jurassic.ru/

Ри<strong>с</strong>. 1008—1011. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pleciodictyidea, Protoligoneuridea, Fungivoridea<br />

1008. Pleciodirtya modesta Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X24.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, rhaetica Rohdendorf; <strong>к</strong>рыл<strong>о</strong>х19,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong> Ср. Азия (<strong>о</strong>риг.<br />

Ср. Азия) <strong>о</strong>риг. ри<strong>с</strong>.). 1009.Protolig oneura fusicosta Rohdendor; ри<strong>с</strong>.). 1011. Rhaetofungivora reticulata Rohdendorf; <strong>к</strong>рыл<strong>о</strong>,<br />

<strong>к</strong>рыл<strong>о</strong>, Х24.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>рип. ри<strong>с</strong>.). 1010. Palaeoplecla X 35,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.).<br />

л<strong>а</strong>,т<strong>о</strong>н<strong>к</strong><strong>а</strong>я;з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь <strong>о</strong>бр<strong>а</strong>зует уз<strong>к</strong>ий р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong>;<br />

передние меди<strong>а</strong>льные жил<strong>к</strong>и непр<strong>а</strong><strong>в</strong>ильные<br />

и т<strong>о</strong>н<strong>к</strong>ие. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4 мм<br />

(ри<strong>с</strong>. 1010). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл.<br />

СЕМЕЙСТВО PLECIOFUNGIVOR1DAE ROHDENDORF,<br />

1946<br />

R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong> изл<strong>о</strong>м<strong>о</strong>м или рез<strong>к</strong>им изгиб<strong>о</strong>м;<br />

RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й трети<br />

<strong>к</strong>рыл<strong>а</strong> и <strong>в</strong><strong>с</strong>егд<strong>а</strong> не<strong>с</strong>ет р<strong>а</strong>зличн<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> передних<br />

<strong>в</strong>ет<strong>в</strong>ей; чет<strong>к</strong>ие п<strong>о</strong>перечные между С, <strong>к</strong><strong>о</strong>нц<strong>о</strong>м<br />

R и <strong>в</strong>ет<strong>в</strong>ями RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; з<strong>а</strong>дняя<br />

<strong>в</strong>ет<strong>в</strong>ь М не т<strong>о</strong>лще других <strong>в</strong>ет<strong>в</strong>ей М. Три<strong>а</strong><strong>с</strong> —<br />

юр<strong>а</strong>. 16 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Rhaetofungivora Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Rh. reticulata Rohdendorf, sp. nov.;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Имеют<strong>с</strong>я я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

<strong>а</strong>рхеди<strong>к</strong>тия <strong>в</strong> <strong>в</strong>иде <strong>с</strong>л<strong>а</strong>бых п<strong>о</strong>перечных: rm<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong> чи<strong>с</strong>ле д<strong>в</strong>ух или б<strong>о</strong>лее жил<strong>о</strong><strong>к</strong>, р<strong>а</strong><strong>в</strong>н<strong>о</strong><br />

<strong>к</strong><strong>а</strong><strong>к</strong> и чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ей RS; <strong>в</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле<br />

<strong>к</strong>реп<strong>к</strong>их п<strong>о</strong>перечных нет. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т 1,5<br />

д<strong>о</strong> 3,25 мм (ри<strong>с</strong>. 1011). Б<strong>о</strong>лее 15 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. три<strong>а</strong><strong>с</strong><br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Rhaetofungivorella Rohdendorf, gen.<br />

nov. Тип р<strong>о</strong>д<strong>а</strong> — Rh. subcosta Rohdendorf, sp.<br />

320<br />

http://jurassic.ru/<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, .И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль).<br />

Имеют<strong>с</strong>я мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные <strong>с</strong>л<strong>а</strong>бые<br />

п<strong>о</strong>перечные жил<strong>к</strong>и или <strong>в</strong>ет<strong>в</strong>и SC, R, RS и между<br />

<strong>в</strong>ет<strong>в</strong>ями М; rm <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong> чи<strong>с</strong>ле не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>их<br />

жил<strong>о</strong><strong>к</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,8—2,0 мм (ри<strong>с</strong>. 1012).<br />

Четыре <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Transversiptecia Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Т. transversinervis Rohdendorf, 1946; <strong>в</strong>.<br />

юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). О<strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>а</strong>рхеди<strong>к</strong>тия нет; имеют<strong>с</strong>я <strong>к</strong>реп<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>с</strong>уб<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле и д<strong>в</strong>е п<strong>о</strong>перечные<br />

между М и СиА; <strong>о</strong>дн<strong>а</strong> передняя <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3,5 мм (ри<strong>с</strong>. 1013). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Polyneurisca Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. atavina Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я'<strong>о</strong>бл.<br />

(м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Имеют<strong>с</strong>я передние<br />

<strong>в</strong>ет<strong>в</strong>и RS <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечных, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющих<br />

<strong>в</strong> R и С; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь RS рез<strong>к</strong><strong>о</strong> з<strong>а</strong>гнут<strong>а</strong><br />

н<strong>а</strong>з<strong>а</strong>д; между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием Mi+2, М4 и СиА<br />

имеют<strong>с</strong>я п<strong>о</strong>перечные. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,75 мм<br />

(ри<strong>с</strong>. 1014). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Rhaetofungivorodes Rohdendorf, gen.<br />

nov. Тип р<strong>о</strong>д<strong>а</strong> — Rh. defectivus Rohdendorf, sp.

Ри<strong>с</strong>. 1012—1016. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Fungivoridea<br />

1012. Rhaetofungivorella analis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, |X36,0, <strong>в</strong>. X 26,0, <strong>в</strong>.'юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1015. Rhaeto<br />

три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 1013. Transversiplecia trans- fungivorodes defectivus Rohdendorf; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, X 17,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>,<br />

versinervis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, х17,5, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>- Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 1016. Protallactoneura turanica Rohденд<strong>о</strong>рф,<br />

1946). 1014. Polyneurisca atavina Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, dendorf; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>,X26,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср.Азия (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль).<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е; SC<br />

длинн<strong>а</strong>я, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я ур<strong>о</strong><strong>в</strong>ня <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния т<strong>о</strong>н<strong>к</strong><strong>о</strong>й<br />

и <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й передней <strong>в</strong>ет<strong>в</strong>и RS; имеет<strong>с</strong>я <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями R и М + CuA;<br />

М <strong>с</strong>л<strong>а</strong>бые. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 3 мм (ри<strong>с</strong>. 1015).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Protallactoneura Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. turanica Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле умеренн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е;<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>чень нежные и нея<strong>с</strong>ные,<br />

<strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>ети. Б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong><br />

RS <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

2,3 мм (ри<strong>с</strong>. 1016). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Archihesperinus Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — А. phryneoid.es Rohdendorf, sp.<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль).<br />

SC н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ями; <strong>о</strong>дн<strong>а</strong> из<strong>о</strong>гнут<strong>а</strong>я,<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я передняя <strong>в</strong>ет<strong>в</strong>ь RS; имеют<strong>с</strong>я<br />

т<strong>о</strong>н<strong>к</strong>ие п<strong>о</strong>перечные жил<strong>к</strong>и между М 4 и CuA;<br />

21 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 321<br />

http://jurassic.ru/<br />

CuA<br />

б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> RS б<strong>о</strong>лее чем <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong><br />

длиннее rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,6 мм (ри<strong>с</strong>. 1017).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Archipleciofungivora Rohdendorf, gen.<br />

nov. Тип р<strong>о</strong>д<strong>а</strong> — A. blnerva Rohdendorf, sp.<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль)<br />

. Имеет<strong>с</strong>я чет<strong>к</strong><strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

<strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле; <strong>о</strong>дн<strong>а</strong> <strong>к</strong>реп<strong>к</strong><strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь RS,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; д<strong>в</strong>е rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

2 мм (ри<strong>с</strong>. 1018). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Archipleciomima Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — A. obtusipennis Rohdendorf, sp.<br />

nov.; <strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>б чл. (рэт,<br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль). SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong> <strong>в</strong> переп<strong>о</strong>н<strong>к</strong>е;<br />

<strong>о</strong>дн<strong>а</strong> передняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>в</strong> <strong>в</strong>иде прям<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й между RS и R; б<strong>а</strong>з<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> RS б<strong>о</strong>лее чем <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> длиннее rm;<br />

передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> прям<strong>о</strong>й, <strong>в</strong>ершин<strong>а</strong> ег<strong>о</strong> туп<strong>а</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,9 мм (ри<strong>с</strong>. 1019). Один<br />

<strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.

R SC SC R<br />

http://jurassic.ru/

Eopachyneura Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

E. trisectoralis Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Д<strong>в</strong>е <strong>к</strong><strong>о</strong><strong>с</strong>ые<br />

передние <strong>в</strong>ет<strong>в</strong>и RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющие <strong>в</strong> С; SC<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С ди<strong>с</strong>т<strong>а</strong>льнее ур<strong>о</strong><strong>в</strong>ня rm; С пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>в</strong>п<strong>а</strong>дения <strong>в</strong> нее SC <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> RS <strong>к</strong><strong>о</strong>р<strong>о</strong>че rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3 мм<br />

(ри<strong>с</strong>. 1020). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Pleciofungivora Rohdendorf, 1938. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. latipennis Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Одн<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, м<strong>а</strong>л<strong>о</strong>из<strong>о</strong>гнут<strong>а</strong>я передняя <strong>в</strong>ет<strong>в</strong>ь<br />

RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> RS <strong>в</strong><br />

1,5—2 р<strong>а</strong>з<strong>а</strong> длиннее rm. SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2—3 мм (ри<strong>с</strong>. 1021).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Pteciofungivorella Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. binerva Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чи<strong>к</strong>мент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Д<strong>в</strong>е <strong>с</strong>л<strong>а</strong>бые,<br />

п<strong>о</strong>чти прямые и м<strong>а</strong>л<strong>о</strong>н<strong>а</strong><strong>к</strong>л<strong>о</strong>ненные передние<br />

<strong>в</strong>ет<strong>в</strong>и RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющие <strong>в</strong> R и С; б<strong>а</strong>з<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> RS <strong>в</strong> 2—2,5 р<strong>а</strong>з<strong>а</strong> длиннее rm; SC <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> С н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не или ди<strong>с</strong>т<strong>а</strong>льнее <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления<br />

RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,5—1,9 мм (ри<strong>с</strong>. 1022). Три<br />

<strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Palaeohesperinus Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> -— P. longipennis Rohdendorf, sp. nov.;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-<br />

Куль). Одн<strong>а</strong> длинн<strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, т<strong>о</strong>н<strong>к</strong><strong>а</strong>я передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong><br />

RS <strong>в</strong>ели<strong>к</strong>, <strong>в</strong> 3,5—5 р<strong>а</strong>з длиннее <strong>к</strong>реп<strong>к</strong><strong>о</strong>й rm. SC<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2—3 мм (ри<strong>с</strong>.<br />

1023). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Prohesperinus Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. abdominalis Rohdendorf, 1946; <strong>в</strong>.<br />

юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Одн<strong>а</strong> <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я <strong>к</strong><strong>о</strong>нцу R<br />

передняя <strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; б<strong>а</strong>з<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> RS <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> длиннее rm; SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3,3 мм (ри<strong>с</strong>. 1024). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Allactoneurites Rohdendorf, 1938. Тип<br />

р<strong>о</strong>д<strong>а</strong> —Л. jurassicus Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Одн<strong>а</strong><br />

<strong>к</strong>реп<strong>к</strong><strong>а</strong>я из<strong>о</strong>гнут<strong>а</strong>я передняя <strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ю­<br />

Ри<strong>с</strong>. 1017—1029. Н<strong>а</strong><br />

1017. Krchihesperinus phryneoides Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х23.0,<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 1018. Archipleciofungivora<br />

binerva Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X20.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг.<br />

ри<strong>с</strong>.). 1019. Archipleciomima obtusipennis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>,<br />

X28.0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 1020. Eopachyneura<br />

trisectoralis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>,X 17,0, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1946). 1021. Pleciofungivora major Rohdendorf; <strong>к</strong>рыл<strong>о</strong><br />

Xl6,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1022. Pleciofungivorella<br />

binerva Rohdendorf; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, Х35.0, <strong>в</strong>. юр<strong>а</strong>,<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1023. Palaeohesperinus longioennis<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>,X 17,0, <strong>в</strong>. три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг.<br />

щ<strong>а</strong>я <strong>в</strong> С и <strong>о</strong><strong>к</strong>руженн<strong>а</strong>я нтер<strong>о</strong><strong>с</strong>тигм<strong>о</strong>й; б<strong>а</strong>з<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> RS р<strong>а</strong><strong>в</strong>ен rm; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М <strong>с</strong>ильн<strong>о</strong><br />

уд<strong>а</strong>лен <strong>о</strong>т rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,5 мм (ри<strong>с</strong>. 1025).<br />

Один. <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Eohesperinus Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. martynovi Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Одн<strong>а</strong><br />

<strong>к</strong>реп<strong>к</strong><strong>а</strong>я <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С и<br />

<strong>о</strong><strong>к</strong>руженн<strong>а</strong>я 'птер<strong>о</strong><strong>с</strong>тигм<strong>о</strong>й; б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong><br />

RS <strong>в</strong> 1,5 р<strong>а</strong>з<strong>а</strong> длиннее rm; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М м<strong>а</strong>л<strong>о</strong><br />

уд<strong>а</strong>лен <strong>о</strong>т rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3 мм (ри<strong>с</strong>. 1026).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО FUNGIVORITIDAE ROHDENDORF, 1957<br />

R <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>чень длинн<strong>а</strong>я, не менее трех чет<strong>в</strong>ертей<br />

<strong>к</strong>рыл<strong>а</strong>, прям<strong>а</strong>я или з<strong>а</strong>гнут<strong>а</strong>я н<strong>а</strong>з<strong>а</strong>д; RS<br />

лишен<strong>а</strong> передних <strong>в</strong>ет<strong>в</strong>ей, или же <strong>о</strong>ни имеют<strong>с</strong>я<br />

<strong>в</strong> <strong>в</strong>иде един<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и между<br />

RS и R; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т R п<strong>о</strong>д <strong>о</strong><strong>с</strong>трым утл<strong>о</strong>м,<br />

и ее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние не имеет <strong>в</strong>ид<strong>а</strong> п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и; rm <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, ин<strong>о</strong>гд<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и М р<strong>а</strong><strong>в</strong>н<strong>о</strong>й т<strong>о</strong>лщины; д<strong>о</strong>п<strong>о</strong>лнительные<br />

п<strong>о</strong>перечные <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Юр<strong>а</strong>. Пять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Mimalycoria Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. allactoneuroides Rohdendorf, 1946;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

rm п<strong>о</strong>перечн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>а</strong>я, уд<strong>а</strong>ленн<strong>а</strong>я <strong>о</strong>т<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> R + RS; SC н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я<br />

<strong>в</strong> С н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong><strong>а</strong>м<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RS; <strong>к</strong><strong>о</strong>нцы<br />

R и RS <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнуты н<strong>а</strong>з<strong>а</strong>д. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

2,75 мм (ри<strong>с</strong>. 1027). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Eoboletina Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. gracilis Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). rm <strong>с</strong> <strong>о</strong>чень<br />

<strong>с</strong>л<strong>а</strong>бым н<strong>а</strong><strong>к</strong>л<strong>о</strong>н<strong>о</strong>м; б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> RS примерн<strong>о</strong><br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее rm; R длинн<strong>а</strong>я, б<strong>о</strong>лее<br />

0,9 длины <strong>к</strong>рыл<strong>а</strong>; С <strong>с</strong>лег<strong>к</strong><strong>а</strong> з<strong>а</strong>х<strong>о</strong>дит з<strong>а</strong> ее <strong>к</strong><strong>о</strong>нец;<br />

SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

ур<strong>о</strong><strong>в</strong>ня rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 4,5 мм (ри<strong>с</strong>. 1028).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Fungivorites Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

F. latimedius Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). rm <strong>с</strong><br />

<strong>о</strong>чень <strong>с</strong>л<strong>а</strong>бым н<strong>а</strong><strong>к</strong>л<strong>о</strong>н<strong>о</strong>м, р<strong>а</strong><strong>в</strong>н<strong>а</strong> или <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong><br />

:емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Fungivoridea<br />

ри<strong>с</strong>.). 1024. Prohesperinus abdominalis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>.<br />

Х13.0, <strong>в</strong>. юр<strong>а</strong>, Ю.К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1025. Allactoneurites<br />

jurassicus Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид.хП.О, <strong>в</strong>. юр<strong>а</strong>, Ю.К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1026. Eohesperinus martynovi Rohdendorf;<br />

<strong>к</strong>рыл<strong>о</strong>, Х20.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946).<br />

1027. Mimalycoria allactoneuroides Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, х19,0,<br />

<strong>в</strong>. юр<strong>а</strong>. Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1028. Eoboletina gracilis<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X 12,0, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1946). 1029. Fungivorites indistinctus Rohdendorf; ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>,<br />

ХЮ,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946)<br />

323 21*<br />

http://jurassic.ru/

<strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> RS; R длинн<strong>а</strong>я, п<strong>о</strong>чти<br />

прям<strong>а</strong>я; С з<strong>а</strong>х<strong>о</strong>дит з<strong>а</strong> <strong>к</strong><strong>о</strong>нец RS; SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я,<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3—5 мм (ри<strong>с</strong>. 1029). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Mesosciophila Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. venosa Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). П<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> rm <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong>я <strong>о</strong>трез<strong>к</strong>у RS; имеет<strong>с</strong>я<br />

передняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>в</strong> <strong>в</strong>иде непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й.<br />

Крыл<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е, лишь <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее<br />

ширины; длин<strong>а</strong> ег<strong>о</strong> 4,5 мм (ри<strong>с</strong>. 1030). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Mesosciophilodes Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. angustipennis Rohdendorf, 1946;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

П<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> rm <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я, незн<strong>а</strong>чительн<strong>о</strong><br />

длиннее или <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>перечн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> RS; имеет<strong>с</strong>я передняя <strong>в</strong>ет<strong>в</strong>ь<br />

RS <strong>в</strong> <strong>в</strong>иде <strong>к</strong>реп<strong>к</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й. Крыл<strong>о</strong> уз<strong>к</strong><strong>о</strong>е,<br />

не менее чем <strong>в</strong> 2,5 р<strong>а</strong>з<strong>а</strong> длиннее ширины; ег<strong>о</strong><br />

длин<strong>а</strong> 4—4,5 мм (ри<strong>с</strong>. 1031). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО SCIARIDAE BILLBERGH, 1820<br />

[nom. transl. Bigot, 1852 (ex Sciaraedes Billbergh, 1820)]<br />

(Lycoriidae Hendel, 1928)<br />

R у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>а</strong>я, не б<strong>о</strong>льше д<strong>в</strong>ух третей <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че, ин<strong>о</strong>гд<strong>а</strong> менее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; RS лишен<strong>а</strong> передних <strong>в</strong>ет<strong>в</strong>ей, <strong>с</strong> рез<strong>к</strong><strong>о</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленным б<strong>а</strong>з<strong>а</strong>льным <strong>о</strong>трез<strong>к</strong><strong>о</strong>м, имеющим<br />

<strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й; н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящ<strong>а</strong>яп<strong>о</strong>перечн<strong>а</strong>я rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong><br />

<strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RS; М и Си <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

зн<strong>а</strong>чительн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее р<strong>а</strong>ди<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> (ри<strong>с</strong>.<br />

1032). Т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г умеренн<strong>о</strong> у<strong>в</strong>еличенные.<br />

Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>литы <strong>с</strong>з<strong>а</strong>ди <strong>а</strong>нтенн. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

из них де<strong>с</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong> из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО MANOTIDAE HENDEL, 1936<br />

Сильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>й,<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й н<strong>а</strong> <strong>к</strong><strong>о</strong>нце SC, <strong>с</strong> пре<strong>в</strong>р<strong>а</strong>щенным<br />

<strong>в</strong> п<strong>о</strong>перечную б<strong>а</strong>з<strong>а</strong>льным <strong>о</strong>трез<strong>к</strong><strong>о</strong>м RS,<br />

1030. Mesosciophila venosa Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X 13,0,<strong>в</strong>. юр<strong>а</strong>,<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1031. Mesosciophilodes similis<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х12.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

1032. Lycoria bicolor Meigen; <strong>к</strong>рыл<strong>о</strong>, xll,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong><br />

п<strong>а</strong> (Hennig, 1954). 1033. Manota sp.; <strong>к</strong>рыл<strong>о</strong>, xll,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hendel, 1936). 1034. Zygomyia notata Stannius; <strong>к</strong>рыл<strong>о</strong><br />

X14.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954). 1035. Zeltnira fasciata<br />

Meigen; <strong>к</strong>рыл<strong>о</strong>, X9,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954), 1036. Mac<br />

rocera tutea Meigen; <strong>к</strong>рыл<strong>о</strong>,X9,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954)<br />

1037. Lygistorrhina sp.; <strong>к</strong>рыл<strong>о</strong>, X7,6, <strong>с</strong><strong>о</strong><strong>в</strong>рем., А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (Hen<br />

del, 1936). 1038. Diadocidia jerruginea Meigen; <strong>к</strong>рыл<strong>о</strong>, х9Д<br />

Ри<strong>с</strong>. 1030—1044. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Fungivoridea<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я имеет <strong>в</strong>ид пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й жил<strong>к</strong>и, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющей<br />

<strong>в</strong> передний <strong>к</strong>р<strong>а</strong>й. Антенны р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><br />

н<strong>а</strong> темени; х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий (ри<strong>с</strong>. 1033).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

324<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО FUNGIVORIDAE LATREILLE, 1809<br />

[nom. transl. Hendel, 1928 (ex Fungivorae Latreille, 1809)]<br />

R изменчи<strong>в</strong><strong>о</strong>й длины; RS лишен<strong>а</strong> передних<br />

<strong>в</strong>ет<strong>в</strong>ей или <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й <strong>в</strong>ет<strong>к</strong><strong>о</strong>й <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й;<br />

'б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> RS рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен п<strong>о</strong>перечн<strong>о</strong>; жил<strong>к</strong>и М ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>л<strong>а</strong>бые<br />

и ме<strong>с</strong>т<strong>а</strong>ми редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные. Т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличенные.<br />

Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленные (ри<strong>с</strong>.<br />

1034). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Обширн<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>,<br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющее <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1500<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> б<strong>о</strong>лее 60 р<strong>о</strong>д<strong>о</strong><strong>в</strong> д<strong>в</strong>ух п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> — Sciophilinae<br />

и Fungivorinae. В третичных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х <strong>о</strong>бн<strong>а</strong>ружены пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

45 р<strong>о</strong>д<strong>о</strong><strong>в</strong> (<strong>с</strong><strong>в</strong>ыше 170 <strong>в</strong>ид<strong>о</strong><strong>в</strong>), еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

изученных.<br />

СЕМЕЙСТВО CEROPLATIDAE RONDANI, 1856<br />

[nom. transl. Hendel, 1928<br />

(ex Ceroplatina Rondani, 1856)]<br />

R у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>а</strong>я, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> длиннее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; rm, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

<strong>с</strong>лияния <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти М <strong>с</strong> RS н<strong>а</strong> <strong>с</strong>ередине<br />

<strong>к</strong>рыл<strong>а</strong>, реже имеет<strong>с</strong>я; М4 и СиА р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я<br />

<strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния. Антенны рез<strong>к</strong><strong>о</strong> <strong>с</strong>плющенные<br />

<strong>в</strong> <strong>в</strong>иде х<strong>а</strong>р<strong>а</strong><strong>к</strong>терных мн<strong>о</strong>г<strong>о</strong>члени<strong>с</strong>тых пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>,<br />

реже цилиндриче<strong>с</strong><strong>к</strong>ие (ри<strong>с</strong>. 1035). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. 17 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

из<strong>в</strong>е<strong>с</strong>тны три р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО MACROCERATIDAE RONDANI, 1856<br />

[nom. transl. Malloch, 1917<br />

(ex Macroceratina Rondani, 1856)]<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние т<strong>а</strong><strong>к</strong><strong>о</strong>е же, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>в</strong> предыдущем<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е; М4 <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong> СиА.<br />

Антенны <strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличены, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> длиннее тел<strong>а</strong>,<br />

т<strong>о</strong>н<strong>к</strong>ие (ри<strong>с</strong>. 1036). П<strong>а</strong>ле<strong>о</strong>ген. Один <strong>о</strong>бширный<br />

р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<strong>в</strong>е<strong>с</strong>тный из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954). 1039. Mycetobia patlipes Meigen;<br />

<strong>к</strong>рыл<strong>о</strong>, Хб,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954). 1040. Archizelmira<br />

kazachstanica Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X20, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(<strong>о</strong>риг. ри<strong>с</strong>.). 1041. Megalycoriomima magnipennis Rohdendorf;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х18,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг<br />

ри<strong>с</strong>.). 1042. Lycoriotnima ventralis Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, х32,0,<br />

<strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1043. Paralycoriomima<br />

sororcula Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х30.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1946). 1044. Lycorioplecia elongataRohdendorf, <strong>к</strong>рыл<strong>о</strong>,<br />

Х25,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946).

http://jurassic.ru/<br />

CuA

СЕМЕЙСТВО LYGISTORRHINIDAE HENDEL, 1936 СЕМЕЙСТВО PLECIOMIMIDAE ROHDENDORF, 1946<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; RS п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т R; SC <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я;<br />

жил<strong>к</strong>и <strong>с</strong>и<strong>с</strong>темы М лишь <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>нечных<br />

<strong>о</strong>тдел<strong>о</strong><strong>в</strong>. Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> рез<strong>к</strong><strong>о</strong> удлиненный (ри<strong>с</strong>.<br />

1037). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уне и <strong>о</strong>дин из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО DIADOCIDIIDAE WINNERTZ, 1863<br />

[nom. transl. Hendel, 1928<br />

(ex Diadocidinae Winnertz, 1863)]<br />

SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; RS <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong>т R п<strong>о</strong>д <strong>о</strong><strong>с</strong>трым угл<strong>о</strong>м, без <strong>в</strong>ет<strong>в</strong>ей; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н; rm и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние М4<br />

имеют<strong>с</strong>я, примерн<strong>о</strong> н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не (ри<strong>с</strong>.<br />

1038). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО MYCETOB11DAE WINNERTZ, 1863<br />

[nom. transl. Hendel, 1928<br />

(ex Mycetobinae Winnertz, 1863)]<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>х<strong>о</strong>дн<strong>о</strong> <strong>с</strong> предыдущим <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>м:<br />

имеет<strong>с</strong>я <strong>к</strong>реп<strong>к</strong><strong>а</strong>я и длинн<strong>а</strong>я передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS; жил<strong>к</strong>и СиА и А р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я<br />

(ри<strong>с</strong>. 1039). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один<br />

р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ARCHIZELMIRIDAE ROHDENDORF,<br />

FAM. NOV.<br />

R длинн<strong>а</strong>я, б<strong>о</strong>лее трех чет<strong>в</strong>ертей <strong>к</strong>рыл<strong>а</strong>; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

RS <strong>с</strong>л<strong>а</strong>б<strong>о</strong>е: имеют<strong>с</strong>я д<strong>в</strong>е передние <strong>в</strong>ет<strong>в</strong>и<br />

RS, <strong>о</strong>дн<strong>а</strong> из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й, друг<strong>а</strong>я<br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет .<strong>в</strong> С; Mi + 2 <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я; rm имеет<strong>с</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Archizelmira Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong>—A. kazachsianica Rohdendorf, sp. nov.;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

SC т<strong>о</strong>н<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

М. Р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3 мм (ри<strong>с</strong>. 1040). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, не б<strong>о</strong>лее трех чет<strong>в</strong>ертей <strong>к</strong>рыл<strong>а</strong>,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong>реп<strong>к</strong><strong>о</strong>е;<br />

RS <strong>в</strong><strong>с</strong>егд<strong>а</strong> без <strong>в</strong>ет<strong>в</strong>ей; п<strong>о</strong>перечные между жил<strong>к</strong><strong>а</strong>ми<br />

М 4 и СиА <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Крылья шир<strong>о</strong><strong>к</strong>ие,<br />

<strong>о</strong>бычн<strong>о</strong> н<strong>а</strong> <strong>в</strong>ершине з<strong>а</strong><strong>к</strong>ругленные. Юр<strong>а</strong>. 13 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Megalycoriomima Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — М. magnipennis Rohdendorf, sp. nov.;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

R длинн<strong>а</strong>я, прям<strong>а</strong>я, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> трех чет<strong>в</strong>ертей длины<br />

<strong>к</strong>рыл<strong>а</strong>: rm нея<strong>с</strong>н<strong>а</strong>я; RS <strong>с</strong>ильн<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д<br />

<strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей <strong>в</strong>ершинн<strong>о</strong>й трети; Mi+2 <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я,<br />

нея<strong>с</strong>н<strong>а</strong>я. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,6 мм (ри<strong>с</strong>. 1041).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Lycoriotnima Rohdendorf, 1946.Тип р<strong>о</strong>д<strong>а</strong>—<br />

L. ventralis Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R н<strong>а</strong> <strong>в</strong>ершине<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед, длин<strong>а</strong> ее не б<strong>о</strong>лее д<strong>в</strong>ух<br />

третей длины <strong>к</strong>рыл<strong>а</strong>; rm <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong> RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>в</strong><strong>с</strong>я п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> и<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д; М и SC <strong>с</strong>л<strong>а</strong>бые; СиР<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метн<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,75 мм (ри<strong>с</strong>.<br />

1042). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Paralycoriomima Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. sororcula Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я О'бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R п<strong>о</strong>чти<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем прям<strong>а</strong>я, длин<strong>а</strong> ее р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong>чти д<strong>в</strong>ум третям<br />

длины <strong>к</strong>рыл<strong>а</strong>; rm <strong>в</strong> 1,5 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong> RS; п<strong>о</strong><strong>с</strong>ледняя п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong>з<strong>а</strong>д; М и SC не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бые, СиР<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,75 мм (ри<strong>с</strong>.<br />

1043). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Lycorioplecia Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

L. elongata Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R з<strong>а</strong>гнут<strong>а</strong>я<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>в</strong>перед, длин<strong>а</strong> ее з<strong>а</strong>ним<strong>а</strong>ет <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> д<strong>в</strong>ух<br />

третей длины <strong>к</strong>рыл<strong>а</strong>; rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> близ<strong>к</strong><strong>о</strong><br />

<strong>о</strong>т ме<strong>с</strong>т<strong>а</strong> <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong>ильн<strong>о</strong> и<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> з<strong>а</strong>гиб<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д; М и SC не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>бые; СиР нея<strong>с</strong>н<strong>а</strong>; имеет<strong>с</strong>я <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,5 мм (ри<strong>с</strong>. 1044).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Lycoriomimod.es Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — L. deformatus Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R лишь<br />

н<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м <strong>к</strong><strong>о</strong>нце <strong>с</strong>лег<strong>к</strong><strong>а</strong> з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед,<br />

Ри<strong>с</strong>. 1045—1055. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Fungivoridea<br />

1045. Lycoriomimodes deformatus Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х30.0,<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1051. Antiquamedia tenuipes Rohdendorf;<br />

<strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1046. Lycoriomimelta<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х15.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938).<br />

minor Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, х48,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н.'(Р<strong>о</strong>ден­<br />

1052. Archilycoria magna Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х15.0, <strong>в</strong>. юр<strong>а</strong>,<br />

д<strong>о</strong>рф, 1946). 1047. Pleciomima secunda Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, Х25,0,<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1053. Paritonida brachyptera<br />

<strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1048. Pleciomimella karata-<br />

Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид, Х16.0, <strong>в</strong>. юр<strong>а</strong>.Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>нЦР<strong>о</strong>денд<strong>о</strong>рф,<br />

vica Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид, Х14.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>­ 1946). 1054. Tipuloplecia breviventris Rohdendorf; ч<strong>а</strong><strong>с</strong>ти <strong>к</strong>рылье<strong>в</strong>,<br />

денд<strong>о</strong>рф, 1946). 1049. Mimallactoneura vetusta Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X 15,0; А — ле<strong>в</strong><strong>о</strong>е <strong>к</strong>рыл<strong>о</strong>; Б — пр<strong>а</strong><strong>в</strong><strong>о</strong>е <strong>к</strong>рыл<strong>о</strong>, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х­<br />

Х18,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1050. Antefun<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.). 1055. Sinemedia angustipennis Rohdendorf;<br />

givora prima Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x32,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н ч<strong>а</strong><strong>с</strong>ть <strong>к</strong>рыл<strong>а</strong>, Х24,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.).<br />

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ft

р<strong>а</strong><strong>в</strong>н<strong>а</strong> д<strong>в</strong>ум третям <strong>к</strong>рыл<strong>а</strong>; rm лишь <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong>ильн<strong>о</strong><br />

з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д; М и SC не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бые;<br />

птер<strong>о</strong><strong>с</strong>тигмы нет; СиР нея<strong>с</strong>н<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

1,6 мм (ри<strong>с</strong>. 1045). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Lycoriomimetta Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — L. minor Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><strong>в</strong><strong>о</strong>лни<strong>с</strong>т<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> д<strong>в</strong>ух третей <strong>к</strong>рыл<strong>а</strong>; rm <strong>в</strong><br />

1,5 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че RS, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть п<strong>о</strong><strong>с</strong>ледней<br />

п<strong>о</strong>чти прям<strong>а</strong>я; SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, з<strong>а</strong>гнут<strong>а</strong>я н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,25 мм (ри<strong>с</strong>. 1046). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Pleciomima Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. secunda Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R п<strong>о</strong>чти прям<strong>а</strong>я,<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong>я д<strong>в</strong>ум третям <strong>к</strong>рыл<strong>а</strong>; rm <strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong>ильн<strong>о</strong><br />

з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине;<br />

М и SC <strong>к</strong>реп<strong>к</strong>ие. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,25—2,50 мм<br />

(ри<strong>с</strong>. 1047). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Pleciomimella Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. karatavica Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R п<strong>о</strong>чти<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем прям<strong>а</strong>я, ед<strong>в</strong><strong>а</strong> з<strong>а</strong>гнут<strong>а</strong>я н<strong>а</strong> <strong>в</strong>ершине; rm<br />

лишь <strong>в</strong> 1,75 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong><br />

RS, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong>;<br />

SC н<strong>а</strong> <strong>в</strong>ершине <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я; М не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

т<strong>о</strong>н<strong>к</strong>ие. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,1 мм (ри<strong>с</strong>. 1048). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Mimallactoneura Rohdendorf, 1946. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. vetusta Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R п<strong>о</strong>чти<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем прям<strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> RS <strong>в</strong> 5 р<strong>а</strong>з<br />

длиннее rm; ди<strong>с</strong>т<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> RS <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

з<strong>а</strong>гнут н<strong>а</strong>з<strong>а</strong>д; SC <strong>о</strong>чень <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, нея<strong>с</strong>н<strong>а</strong>я; М<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метные, т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и СиР. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 3 мм (ри<strong>с</strong>. 1049). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Antefungivora Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. prima Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R прям<strong>а</strong>я; rm<br />

б<strong>о</strong>лее чем <strong>в</strong> 6 р<strong>а</strong>з <strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong><br />

RS; ди<strong>с</strong>т<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong> эт<strong>о</strong>й жил<strong>к</strong>и <strong>в</strong> <strong>в</strong>ершинн<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> з<strong>а</strong>гнут н<strong>а</strong>з<strong>а</strong>д <strong>в</strong> <strong>в</strong>иде<br />

дуги; SC <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С;<br />

М4 длинн<strong>а</strong>я, <strong>с</strong><strong>о</strong>единенн<strong>а</strong>я <strong>с</strong> Mi+2. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

1,5 мм (ри<strong>с</strong>. 1050). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Antiquamedia Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. tenuipes Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong><br />

<strong>в</strong>перед; rm нея<strong>с</strong>н<strong>а</strong>я; RS <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> R; М 4 длинн<strong>а</strong>я, <strong>с</strong><strong>о</strong>е­<br />

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динен<strong>а</strong> <strong>с</strong> М. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2 мм (ри<strong>с</strong>. 1051).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Archilycoria Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. magna Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R прям<strong>а</strong>я,<br />

длинн<strong>а</strong>я; rm <strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong><br />

RS; RS р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д;<br />

SC 'нея<strong>с</strong>н<strong>а</strong>; М 4 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>в</strong> <strong>в</strong>иде <strong>в</strong>ет<strong>в</strong>и СиА.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,6 мм (ри<strong>с</strong>. 1052). Один <strong>в</strong>ид,<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Paritonida Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P.'brachyptera Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

з<strong>а</strong>гнут<strong>а</strong> н<strong>а</strong> <strong>в</strong>ершине; rm б<strong>о</strong>лее чем <strong>в</strong> 6 р<strong>а</strong>з <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> RS, ди<strong>с</strong>т<strong>а</strong>льный <strong>о</strong>трез<strong>о</strong><strong>к</strong><br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> з<strong>а</strong>гнут н<strong>а</strong>з<strong>а</strong>д; SC <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>; M 4 <strong>в</strong> <strong>в</strong>иде длинн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и СиА,<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т Mi +2. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 1,75 мм (ри<strong>с</strong>.<br />

1053). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО TIPULOPLECIIDAE ROHDENDORF,<br />

FAM. NOV.<br />

Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие. Крылья зн<strong>а</strong>чительн<strong>о</strong> длиннее<br />

брюш<strong>к</strong><strong>а</strong>, шир<strong>о</strong><strong>к</strong>ие. Имеет<strong>с</strong>я не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> передних<br />

<strong>в</strong>ет<strong>в</strong>ей RS; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние нежн<strong>о</strong>е, <strong>к</strong>р<strong>о</strong>ме<br />

<strong>к</strong>реп<strong>к</strong><strong>о</strong>й С. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Tipuloptecia Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Т. breviventris Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). R т<strong>о</strong>н<strong>к</strong><strong>а</strong>я,<br />

прям<strong>а</strong>я; д<strong>о</strong> ше<strong>с</strong>ти передних <strong>в</strong>ет<strong>в</strong>ей RS, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

б<strong>о</strong>льш<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть имеет <strong>в</strong>ид п<strong>о</strong>перечных между<br />

RS и R; лишь немн<strong>о</strong>гие <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> С; RS<br />

<strong>с</strong>ильн<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong>я. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,25 мм (ри<strong>с</strong>.<br />

1054). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО SINEMEDIIDAE ROHDENDORF,<br />

FAM. NOV.<br />

Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие, бег<strong>а</strong>тельные. Крылья уз<strong>к</strong>ие,<br />

не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> длинные. Передние <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>реп<strong>к</strong><strong>о</strong>е; е<strong>с</strong>ть <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я<br />

интермеди<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я между передними<br />

<strong>в</strong>ет<strong>в</strong>ями М. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Sinemedia Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — 5. angustipennis Rohdendorf, sp. nov.;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

R <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнут<strong>а</strong>я <strong>в</strong>перед; rm <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я,<br />

<strong>в</strong> 3 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> RS; ди<strong>с</strong>т<strong>а</strong>льный<br />

<strong>о</strong>трез<strong>о</strong><strong>к</strong> эт<strong>о</strong>й жил<strong>к</strong>и <strong>с</strong>л<strong>а</strong>б<strong>о</strong> з<strong>а</strong>гнут<br />

н<strong>а</strong>з<strong>а</strong>д; жил<strong>к</strong>и М не <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бые. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2,25 мм (ри<strong>с</strong>. 1055). Один <strong>в</strong>ид. В. юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

НАДСЕМЕЙСТВО BIBIONIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>; фр<strong>а</strong>гм<strong>а</strong> не<br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. Н<strong>о</strong>ги цеп<strong>к</strong>ие, реже бег<strong>а</strong>тельные, <strong>а</strong>нтенны<br />

т<strong>о</strong>н<strong>к</strong>ие или у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; RS п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong>

CuA<br />

1056. Eoplecia primitiva Handlirsch; <strong>к</strong>рыл<strong>о</strong>, X12.0, н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния (Handlirsch, 1939). 1057. Paraxymyia quadrirtdialls<br />

Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x27,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф,<br />

1946). 1058. Protoplecia Hasina (Geinitz); <strong>к</strong>рыл<strong>о</strong>, X10.0, н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния (Handlirsch, 1906). 1059. Mesopleciella minor Rohdendorf;<br />

<strong>к</strong>рыл<strong>о</strong>, Xl6,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946).<br />

1060. Mesoplecia stigma Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид, х9,0, <strong>в</strong>. юр<strong>а</strong>,<br />

передними <strong>в</strong>ет<strong>в</strong>ями; е<strong>с</strong>ли передних <strong>в</strong>ет<strong>в</strong>ей RS<br />

нет, т<strong>о</strong> М и CuA зн<strong>а</strong>чительн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее и <strong>с</strong><strong>в</strong>етлее<br />

R и р<strong>а</strong>змеры <strong>к</strong>рупные; SC <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинн<strong>а</strong>я, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>. Р<strong>а</strong>змеры<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong>редние или <strong>к</strong>рупные, ред<strong>к</strong><strong>о</strong> мел<strong>к</strong>ие.<br />

Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Eopleciidae, Р<strong>а</strong>г<strong>а</strong>хуmyiidae,<br />

Protopleciidae, Penthetriidae, Hesperinidae,<br />

Bibionidae, Axymyiidae; п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не н<strong>а</strong>йден<strong>о</strong>.<br />

СЕМЕЙСТВО EOPLECIIDAE ROHDENDORF, 1946<br />

Д<strong>в</strong>е <strong>к</strong>реп<strong>к</strong>ие и длинные <strong>в</strong>ет<strong>в</strong>и RS, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

п<strong>о</strong> <strong>о</strong>бе <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>о</strong>т rm; mcu р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

между CuA и М 4; д<strong>в</strong>е А. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

3,5 мм (ри<strong>с</strong>. 1056). Н. юр<strong>а</strong> Герм<strong>а</strong>нии. Один р<strong>о</strong>д.<br />

Ри<strong>с</strong>. 1056—1064. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>" Bibionidea<br />

CuA<br />

1058<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг. ри<strong>с</strong>.). 1061. Penthetria funebris Meigen;<br />

<strong>к</strong>рыл<strong>о</strong>, X3,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954). 1062. Hesperinus<br />

imbecillus Loew; <strong>к</strong>рыл<strong>о</strong>, X5,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig,<br />

1954). 1063. Protobibio jurassicus Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X30.0,<br />

<strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1946). 1964. Bibio marci<br />

Linnaeus; <strong>к</strong>рыл<strong>о</strong>, Х3,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954)<br />

СЕМЕЙСТВО PARAXYMYIIDAE ROHDENDORF, 1946<br />

Д<strong>в</strong>е <strong>к</strong>реп<strong>к</strong>ие <strong>в</strong>ет<strong>в</strong>и и длинные <strong>в</strong>ет<strong>в</strong>и RS, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

п<strong>о</strong> <strong>о</strong>бе <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны rm; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>трез<strong>о</strong><strong>к</strong><br />

М редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н; SC меньше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Paraxymyia Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. quadriradialis Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>.<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Имеет<strong>с</strong>я<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; SC т<strong>о</strong>н<strong>к</strong><strong>а</strong>я; передняя <strong>в</strong>ет<strong>в</strong>ь RS п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong><br />

R. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,25 мм (ри<strong>с</strong>.<br />

1057). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО PROTOPLECIIDAE ROHDENDORF, 1946<br />

Одн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льнее<br />

rm; mcu р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й между<br />

CuA и М; SC изменчи<strong>в</strong><strong>о</strong>г<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жения. Юр<strong>а</strong>.<br />

Три р<strong>о</strong>д<strong>а</strong>.<br />

329<br />

http://jurassic.ru/

Protoplecia Handlirsch, 1906. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. Hasina (Geinitz, 1884); н. юр<strong>а</strong> Герм<strong>а</strong>нии.<br />

SC не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> Mi+2 р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен ди<strong>с</strong>т<strong>а</strong>льнее ур<strong>о</strong><strong>в</strong>ня<br />

<strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления <strong>в</strong>ет<strong>в</strong>и RS. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,5 мм<br />

(ри<strong>с</strong>. 1058). Один <strong>в</strong>ид. Н. юр<strong>а</strong> Герм<strong>а</strong>нии.<br />

Mesopleciella Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

М. minor Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). SC зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> Mi+2 р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее ур<strong>о</strong><strong>в</strong>ня р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

RS; mcu <strong>с</strong><strong>о</strong>единяет СиА и <strong>о</strong>бщий <strong>с</strong>т<strong>в</strong><strong>о</strong>л<br />

М. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,75 мм (ри<strong>с</strong>. 1059). Один<br />

<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Mesoplecia Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong><br />

— М. jurassica Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). SC примерн<strong>о</strong><br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> Mi+2<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен зн<strong>а</strong>чительн<strong>о</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong><br />

RS; mcu <strong>с</strong><strong>о</strong>единяет СиА и М 4. Крупнее,<br />

длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 7—8,2 мм (ри<strong>с</strong>. 1060). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО PENTHETRIIDAE ROHDENDORF, 1946<br />

Одн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS. Р<strong>а</strong>змеры <strong>с</strong>редние или <strong>к</strong>рупные.<br />

Жил<strong>к</strong>и <strong>в</strong><strong>с</strong>ех <strong>с</strong>и<strong>с</strong>тем примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

т<strong>о</strong>лщины. SC <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинн<strong>а</strong>я; mcu р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> и имеет <strong>в</strong>ид <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> М 4, <strong>в</strong> т<strong>о</strong><br />

<strong>в</strong>ремя <strong>к</strong><strong>а</strong><strong>к</strong> и<strong>с</strong>тинн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние М 4 <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й.<br />

Н<strong>о</strong>ги без шип<strong>о</strong><strong>в</strong>. Антенны у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные,<br />

меньше длины груди (ри<strong>с</strong>. 1061). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, н<strong>а</strong>йденные<br />

т<strong>а</strong><strong>к</strong>же <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО HESPERINIDAE HENDEL, 1928<br />

Одн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS; жил<strong>к</strong>и примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

т<strong>о</strong>лщины; SC <strong>о</strong>чень длинн<strong>а</strong>я, б<strong>о</strong>льше п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>трез<strong>о</strong><strong>к</strong> М 4 имеет <strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и. Антенны длинные и т<strong>о</strong>н<strong>к</strong>ие,<br />

р<strong>а</strong><strong>в</strong>ные длине труди. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

5—7 мм (ри<strong>с</strong>. 1062). Не<strong>о</strong>ген — ныне. Один<br />

р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне и <strong>о</strong>дин р<strong>о</strong>д из не<strong>о</strong>ген<strong>а</strong><br />

Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО PROTOBIBIONIDAE ROHDENDORF,<br />

1946<br />

Одн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими р<strong>а</strong>ди<strong>а</strong>льными<br />

жил<strong>к</strong><strong>а</strong>ми и <strong>с</strong>л<strong>а</strong>быми, нея<strong>с</strong>ными М<br />

и СиА. Р<strong>а</strong>змеры <strong>о</strong>чень м<strong>а</strong>лы — длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

1,5 мм. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Protobibio Rohdendorf, 1946. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. jurassicus Rohdendorf, 1946; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). SC <strong>к</strong><strong>о</strong>р<strong>о</strong>­<br />

330<br />

http://jurassic.ru/<br />

че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; rm длинн<strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я. Крыл<strong>о</strong><br />

уз<strong>к</strong><strong>о</strong>е (ри<strong>с</strong>. 1063). Один.<strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО BIBIONIDAE NEWMAN, 1834<br />

[nom. transl. Kirby, 1837<br />

(ex Bibionides Newman, 1834)]<br />

RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я, без <strong>в</strong>ет<strong>в</strong>ей. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>ильн<strong>о</strong><br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; р<strong>а</strong>ди<strong>а</strong>льные жил<strong>к</strong>и, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

М и rm <strong>к</strong>реп<strong>к</strong>ие, темные, рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ющие<strong>с</strong>я<br />

<strong>о</strong>т бледных з<strong>а</strong>дних жил<strong>о</strong><strong>к</strong> М и CuA; rm<br />

имеет <strong>в</strong>ид <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>о</strong>трез<strong>к</strong><strong>а</strong> RS, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>с</strong>лияния М <strong>с</strong> RS; М 4<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т Mi+2 и я<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong>ет<strong>в</strong>ью<br />

СиА: ее <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние п<strong>о</strong>х<strong>о</strong>дит н<strong>а</strong> п<strong>о</strong>перечную<br />

жил<strong>к</strong>у, <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>мещенную ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong><br />

(ри<strong>с</strong>. 1064). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong><br />

третичных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х.<br />

НАДСЕМЕЙСТВО BOLITOPHILIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>; фр<strong>а</strong>гм<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

<strong>в</strong>ме<strong>с</strong>т<strong>о</strong> нее н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong>рыл<strong>а</strong> имеет<strong>с</strong>я<br />

<strong>с</strong>ильн<strong>о</strong> ут<strong>о</strong>лщенный б<strong>а</strong>з<strong>а</strong>льный уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong><br />

жил<strong>к</strong>и СиА. Р<strong>а</strong>змеры мел<strong>к</strong>ие. Антенны т<strong>о</strong>н<strong>к</strong>ие<br />

и длинные. Интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>.<br />

Три <strong>в</strong>ет<strong>в</strong>и М. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Bolitophilidae, Arachnocampidae, п<strong>о</strong><strong>с</strong>леднее<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО BOL1THOPH1L1DAE WINNERTZ, 1863<br />

[nom. transl. Malloch, 1917<br />

(ex Bolithophilinae Winnertz, 1863)]<br />

JRS <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью; mcu р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ления RS; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н; SC н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong> (ри<strong>с</strong>. 1065). П<strong>а</strong>ле<strong>о</strong>ген. Один<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО SCATOPSIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й,<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й н<strong>а</strong> <strong>к</strong><strong>о</strong>нце SC; фр<strong>а</strong>гм<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

или имеет<strong>с</strong>я. Тел<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е и <strong>к</strong>реп<strong>к</strong><strong>о</strong>е;<br />

н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, цеп<strong>к</strong>ие; <strong>а</strong>нтенны т<strong>о</strong>н<strong>к</strong>ие, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные. Интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>.<br />

Три <strong>в</strong>ет<strong>в</strong>и М. Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Protoscatopsidae, Scatopsidae, Canthyloscelididae,<br />

Synneuridae: д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних из<strong>в</strong>е<strong>с</strong>тны<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PROTOSCATOPSIDAE ROHDENDORF,<br />

1946<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, ед<strong>в</strong><strong>а</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние рез<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е: б<strong>о</strong>льш<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и <strong>к</strong>рыл<strong>а</strong> лишен<strong>а</strong> жил<strong>о</strong><strong>к</strong>. SC <strong>о</strong>т-

CuA<br />

Ри<strong>с</strong>- 1065—1073. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> В<strong>о</strong> litophilidea, Scatopsidea, Rhyphidea<br />

1065. Bolitophila hybrida Meigen; <strong>к</strong>рыл<strong>о</strong>, Хб,4, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

СЕМЕЙСТВО OLIGOPHRYNEIDAE ROHDENDORF,<br />

FAM. NOV.<br />

Д<strong>в</strong>е передние <strong>в</strong>ет<strong>в</strong>и RS: длинн<strong>а</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я<br />

и <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я, имеющ<strong>а</strong>я<br />

<strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Имеют<strong>с</strong>я<br />

три интермеди<strong>а</strong>льных п<strong>о</strong>перечных; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>а</strong>. Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Oligophryne Rohdendorf, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — О. fungivoroides Rohdendorf, sp. nov.<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, С<strong>о</strong>гюты).<br />

SC н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я; R <strong>к</strong>реп<strong>к</strong><strong>а</strong>я. Длин<strong>а</strong><br />

<strong>к</strong>рыл<strong>а</strong> 2 мм (ри<strong>с</strong>. 1068). Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong><br />

И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО PROTOLBIOGASTRIDAE ROHDENDORF,<br />

FAM. NOV.<br />

Одн<strong>а</strong> длинн<strong>а</strong>я, из<strong>о</strong>гнут<strong>а</strong>я передняя <strong>в</strong>ет<strong>в</strong>ь<br />

RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е; <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>с</strong>л<strong>о</strong>жн<strong>а</strong>я фр<strong>а</strong>гм<strong>а</strong>; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>о</strong>чень <strong>о</strong><strong>с</strong>л<strong>а</strong>блен; имеют<strong>с</strong>я д<strong>о</strong>п<strong>о</strong>лнительные<br />

п<strong>о</strong>перечные между RS и М, между<br />

М 3 и М 4; SC длинн<strong>а</strong>я, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я ур<strong>о</strong><strong>в</strong>ня<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> интермеди<strong>а</strong>льн<strong>о</strong>й ячей<strong>к</strong>и. Три<strong>а</strong><strong>с</strong>.<br />

Один р<strong>о</strong>д.<br />

Protolbiogaster Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. rhaetica Rohdendorf, sp. nov.;<br />

<strong>в</strong>. три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, С<strong>о</strong>гюты).<br />

Ди<strong>с</strong>т<strong>а</strong>льнее передней <strong>в</strong>ет<strong>в</strong>и имеют<strong>с</strong>я <strong>с</strong>леды<br />

других <strong>в</strong>ет<strong>в</strong>ей; жил<strong>к</strong>и <strong>с</strong>и<strong>с</strong>темы М я<strong>в</strong>н<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее<br />

R и СиА. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 4,7 мм (ри<strong>с</strong>. 1069).<br />

Один <strong>в</strong>ид. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

СЕМЕЙСТВО OLBIOGASTR1DAE HENNIG, 1948<br />

Одн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С или <strong>в</strong> R;<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я <strong>с</strong><strong>а</strong>м<strong>о</strong>е б<strong>о</strong>льшее<br />

пер<strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления М; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л М не<br />

<strong>о</strong><strong>с</strong>л<strong>а</strong>блен; д<strong>о</strong>п<strong>о</strong>лнительных п<strong>о</strong>перечных нет<br />

(ри<strong>с</strong>. 1070). Юр<strong>а</strong> — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны и д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из юры 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО RHYPH1DAE NEWMAN, 1834<br />

[nom. transl. Macquart, 1838<br />

(ex Rhyphites Newman, 1834)]<br />

(Phryneidae Hendel, 1928)<br />

Одн<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; SC длинн<strong>а</strong>я,<br />

<strong>к</strong><strong>о</strong>нц<strong>о</strong>м з<strong>а</strong>х<strong>о</strong>дящ<strong>а</strong>я з<strong>а</strong> ур<strong>о</strong><strong>в</strong>ень rm; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л М <strong>к</strong>реп<strong>к</strong>ий (ри<strong>с</strong>. 1071). Юр<strong>а</strong> — ныне.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

СЕМЕЙСТВО PROTORHYPHIDAE HANDLIRSCH, 1906<br />

Д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и RS, <strong>о</strong>тх<strong>о</strong>дящие п<strong>о</strong> <strong>о</strong>бе <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

rm и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющие <strong>в</strong> С; SC <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, ед<strong>в</strong><strong>а</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я<br />

ур<strong>о</strong><strong>в</strong>ня <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> интермеди­<br />

332<br />

http://jurassic.ru/<br />

<strong>а</strong>льн<strong>о</strong>й ячей<strong>к</strong>и; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>с</strong>т<strong>в</strong><strong>о</strong>л М ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong><strong>с</strong>л<strong>а</strong>блен.<br />

Три<strong>а</strong><strong>с</strong> — юр<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Archirhyphus Handlirsch, 1939. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. geinitzi Handlirsch, 1939; н. юр<strong>а</strong>,<br />

Герм<strong>а</strong>ния. Mi и Мг <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>бщий <strong>с</strong>тебеле<strong>к</strong>,<br />

<strong>в</strong>ых<strong>о</strong>дящий из ячей<strong>к</strong>и. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3 мм<br />

(ри<strong>с</strong>. 1072). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. Н. юр<strong>а</strong> Герм<strong>а</strong>нии и юр<strong>а</strong><br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Protorhyphus Handlirsch, 1906. Три р<strong>о</strong>д<strong>а</strong>—<br />

Phryganidium simplex Geinitz, 1887; н.<br />

юр<strong>а</strong>, Герм<strong>а</strong>ния. Mi и М 2 <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong> <strong>в</strong>ых<strong>о</strong>дят<br />

из интермеди<strong>а</strong>льн<strong>о</strong>й ячей<strong>к</strong>и, не <strong>о</strong>бр<strong>а</strong>зуя<br />

<strong>с</strong>тебель<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,5—3 мм (ри<strong>с</strong>. 1073).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл. и<br />

н. юр<strong>а</strong> Герм<strong>а</strong>нии.<br />

НАДСЕМЕЙСТВО<br />

PHRAGMOLIGONEURIDEA<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле без <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>; имеет<strong>с</strong>я х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я фр<strong>а</strong>гм<strong>а</strong>; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>; М из трех <strong>в</strong>ет<strong>в</strong>ей. Три<strong>а</strong><strong>с</strong>.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phragmoligoneuridae.<br />

СЕМЕЙСТВО PHRAGMOLIGONEURIDAE<br />

ROHDENDORF, FAM. NOV.<br />

Б<strong>а</strong>з<strong>а</strong>льные <strong>о</strong>трез<strong>к</strong>и RS и М редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные,<br />

RS <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й передней <strong>в</strong>ет<strong>в</strong>ью <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й.<br />

Три<strong>а</strong><strong>с</strong>. Один р<strong>о</strong>д.<br />

Phragmoligoneura Rohdendorf, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Ph. incerta Rohdendorf, sp. nov.; <strong>в</strong>.<br />

три<strong>а</strong><strong>с</strong>, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (рэт, С<strong>о</strong>гюты)<br />

SC <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С; Mi + 2 нед<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>о</strong>т<br />

rm. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 2,4 мм (ри<strong>с</strong>. 1074). Один <strong>в</strong>ид.<br />

В. три<strong>а</strong><strong>с</strong> И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

НАДСЕМЕЙСТВО CECIDOMY1IDEA<br />

(Itonidoidea)<br />

Р<strong>а</strong>змеры мел<strong>к</strong>ие. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, небег<strong>а</strong>тельные.<br />

Фр<strong>а</strong>гм<strong>а</strong> не р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>. Антенны <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> п<strong>о</strong>др<strong>а</strong>зделенными члени<strong>к</strong><strong>а</strong>ми или<br />

<strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, рез<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенн<strong>о</strong>е; <strong>к</strong>рылья <strong>с</strong>л<strong>а</strong>бые,<br />

без <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Lestremiidae, Cecidomyiidae, Heteropezidae.<br />

СЕМЕЙСТВО LESTREMIIDAE RONDANI, 1841<br />

[nom. transl. Rohdendorf, hie<br />

(ex Lestriminae Rondani, 1841)]<br />

Mi + 2 <strong>в</strong><strong>с</strong>егд<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>, <strong>о</strong>бычн<strong>о</strong> <strong>в</strong> <strong>в</strong>иде <strong>в</strong>ил<strong>к</strong>и;<br />

А з<strong>а</strong>метн<strong>а</strong>; пер<strong>в</strong>ый члени<strong>к</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> длиннее<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong>. Пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>к</strong>и имеют<strong>с</strong>я (ри<strong>с</strong>.

Ри<strong>с</strong>. 1074—1083. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Phragmol igoneuridea, Cecidomyiidea, Stratiomyiidea<br />

1074. Phragmoligoneura incerta Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, x22,0, <strong>в</strong>. з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1080. Palaeostratiomyia pygmaea<br />

три<strong>а</strong><strong>с</strong>, Ср. Азия (<strong>о</strong>риг. ри<strong>с</strong>.). 1075. Catocha sp.; <strong>к</strong>рыл<strong>о</strong>, Rohdendorf; A — <strong>к</strong>рыл<strong>о</strong>, X24.0; Б — <strong>о</strong>бщий <strong>в</strong>ид, xll.O. <strong>в</strong>. юр<strong>а</strong>,<br />

Х10.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., С Амери<strong>к</strong><strong>а</strong> (Hennig, 1954). 1076. Asphondylia Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1081. Eomyia veterrima Roh­<br />

sarothamni Loew; <strong>к</strong>рыл<strong>о</strong>, Xl5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, dendorf; <strong>о</strong>бщий <strong>в</strong>ид, Х27.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (<strong>о</strong>риг.<br />

1954). 1077. Miastor hastatus Kieffer; <strong>к</strong>рыл<strong>о</strong>, Xl4,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., ри<strong>с</strong>.). 1082. Protocyrtus jurassicus Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954). 1078. Protobrachyceron liasinum Hand­ Х7,5, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1083. Hermione<br />

lirsch; <strong>к</strong>рыл<strong>о</strong>, X9,5, и. юр<strong>а</strong>, Герм<strong>а</strong>ния (Hennig, 1954). 1079. Ar- locuples Loew; <strong>к</strong>рыл<strong>о</strong>, хб,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hennig, 1954)<br />

chisargus pulcher Rohdendorf; <strong>о</strong>бщий <strong>в</strong>ид, x2,3, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>-<br />

1075). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CECIDOMYIIDAE NEWMAN, 1834<br />

[nom. transl. Macquart, 1938<br />

(ex Cecidomiites Newman, 1834)]<br />

(Itonididae Griinberg, 1910)<br />

Mi + 2 <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует или имеет<strong>с</strong>я <strong>в</strong> <strong>в</strong>иде пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й<br />

жил<strong>к</strong>и. SC з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чн<strong>а</strong>я и н<strong>а</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е<br />

<strong>к</strong>рыл<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Пер<strong>в</strong>ый члени<strong>к</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

333<br />

М Ю78<br />

<strong>о</strong>бычн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, реже имеет<strong>с</strong>я (ри<strong>с</strong>. 1076).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Обширн<strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>: <strong>с</strong><strong>в</strong>ыше<br />

3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> и б<strong>о</strong>лее 100 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО HETEROPEZIDAE KERTESZ, 1902<br />

[nom. transl. Hendel, 1928<br />

(ex Heteropezinae Kertesz, 1908)]<br />

Пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Пер<strong>в</strong>ый члени<strong>к</strong><br />

л<strong>а</strong>п<strong>о</strong><strong>к</strong> длиннее <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong>, или л<strong>а</strong>п<strong>к</strong>и из<br />

д<strong>в</strong>ух — четырех члени<strong>к</strong><strong>о</strong><strong>в</strong>; М <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует или<br />

http://jurassic.ru/

<strong>в</strong> <strong>в</strong>иде пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и (ри<strong>с</strong>. 1077). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; из них три р<strong>о</strong>д<strong>а</strong>—Miastor Meinert, Неteropeza<br />

Winnertz, Frirenia Kieffer — <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ИНФРАОТРЯД ASILOMORPHA<br />

В<strong>с</strong>егд<strong>а</strong> имеет<strong>с</strong>я п<strong>о</strong> <strong>к</strong>р<strong>а</strong>йней мере <strong>о</strong>дн<strong>а</strong> длинн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь RS, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ющ<strong>а</strong>я <strong>в</strong> С и <strong>о</strong>тх<strong>о</strong>дящ<strong>а</strong>я<br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее rm; <strong>в</strong>ершинный <strong>о</strong>тдел <strong>к</strong>рыл<strong>а</strong><br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен. Антенны п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>с</strong>л<strong>о</strong>жн<strong>о</strong> п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енный <strong>в</strong>ершинный <strong>о</strong>тдел,<br />

т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емый «третий» члени<strong>к</strong>. Юр<strong>а</strong> —<br />

ныне. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Stratiomyiidea, Tabanidea,<br />

Asilidea, Bombilidea, Empididea.<br />

НАДСЕМЕЙСТВО STRATIOMYIIDEA<br />

Пр<strong>о</strong>пле<strong>в</strong>риты <strong>с</strong>литы <strong>с</strong> пр<strong>о</strong><strong>с</strong>тернит<strong>о</strong>м, з<strong>а</strong>мы<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong>переди ям<strong>к</strong>у передних т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong>. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; С не <strong>о</strong>бх<strong>о</strong>дит<br />

<strong>в</strong>е<strong>с</strong>ь <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>; интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>а</strong>я. Эмп<strong>о</strong>дий шир<strong>о</strong><strong>к</strong>ий, <strong>в</strong><br />

<strong>в</strong>иде пуль<strong>в</strong>иллы. Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Рг<strong>о</strong>tobrachyceridae,<br />

Archisargidae, Palaeostratiomyiidae,<br />

Eomyiidae, Protocyrtidae, Stratiomyiidae,<br />

Xylophagidae, Rachiceridae, Acroceridae, Solvidae<br />

и Chiromyzidae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>о</strong>бн<strong>а</strong>ружены<br />

лишь <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PROTOBRACHYCERIDAE<br />

ROHDENDORF, FAM. NOV.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние не<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и<br />

<strong>к</strong>реп<strong>к</strong>ие; М 4 и Мз <strong>в</strong>ых<strong>о</strong>дят из интермеди<strong>а</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong>и и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ют <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>, з<strong>а</strong>метн<strong>о</strong><br />

<strong>с</strong>х<strong>о</strong>дя<strong>с</strong>ь; интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong>ытянут<strong>а</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 4,5 мм (ри<strong>с</strong>. 1078). Юр<strong>а</strong>. Один<br />

р<strong>о</strong>д <strong>в</strong> н. юре Герм<strong>а</strong>нии. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным Solvidae.<br />

СЕМЕЙСТВО ARCHISARGIDAE ROHDENDORF,<br />

FAM. NOV.<br />

Крыл<strong>о</strong> длинн<strong>о</strong>е и уз<strong>к</strong><strong>о</strong>е, н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>уженн<strong>о</strong>е.<br />

SC и R <strong>к</strong>репче других жил<strong>о</strong><strong>к</strong>, п<strong>о</strong><strong>с</strong>ледняя<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й <strong>в</strong>ет<strong>в</strong>ью; RS <strong>с</strong>л<strong>а</strong>бые,<br />

пл<strong>о</strong>х<strong>о</strong> р<strong>а</strong>зличимые, <strong>в</strong> чи<strong>с</strong>ле трех <strong>в</strong>ет<strong>в</strong>ей. Тел<strong>о</strong><br />

удлиненн<strong>о</strong>е, р<strong>а</strong>змеры <strong>к</strong>рупные. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Archisargus Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong><br />

— A. pulcher Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). SC б<strong>о</strong>льше<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>; R п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем прям<strong>а</strong>я. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

т<strong>а</strong><strong>к</strong><strong>о</strong>й же ширины, <strong>к</strong><strong>а</strong><strong>к</strong> грудь; гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>чень<br />

<strong>в</strong>ели<strong>к</strong>и. Грудь пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я, щит<strong>о</strong><strong>к</strong> притуплённый,<br />

угл<strong>о</strong><strong>в</strong><strong>а</strong>тый. Сегменты брюш<strong>к</strong><strong>а</strong> <strong>с</strong> темным<br />

ри<strong>с</strong>ун<strong>к</strong><strong>о</strong>м. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

20 мм, <strong>к</strong>рыл<strong>а</strong>— 16 мм (ри<strong>с</strong>. 1079). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

334<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PALAEOSTRATIOMYIIDAE<br />

ROHDENDORF, 1951<br />

Крыл<strong>о</strong> притуплённ<strong>о</strong>е, шир<strong>о</strong><strong>к</strong><strong>о</strong>е, н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong>уженн<strong>о</strong>е. В<strong>с</strong>е жил<strong>к</strong>и примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>й<br />

т<strong>о</strong>лщины, лишь SC и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти М <strong>с</strong>л<strong>а</strong>бее<br />

других; <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и М <strong>в</strong>ых<strong>о</strong>дят из интермеди<strong>а</strong>льн<strong>о</strong>й<br />

ячей<strong>к</strong>и. Тел<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченн<strong>о</strong>е; брюш<strong>к</strong><strong>о</strong> притуплённ<strong>о</strong>е,<br />

яйце<strong>в</strong>идн<strong>о</strong>е. Р<strong>а</strong>змеры мел<strong>к</strong>ие. Юр<strong>а</strong>.<br />

Один р<strong>о</strong>д.<br />

Palaeostratiomyia Rohdendorf, 1938.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. pygmaea Rohdendorf, 1938;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины <strong>к</strong>рыл<strong>а</strong>; R <strong>с</strong>л<strong>а</strong>б<strong>о</strong><br />

з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед; пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь RS <strong>с</strong>ильн<strong>о</strong><br />

из<strong>о</strong>гнут<strong>а</strong>я, ди<strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>ь п<strong>о</strong>чти прям<strong>а</strong>я,<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>а</strong>я <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>му <strong>с</strong>т<strong>в</strong><strong>о</strong>лу RS; rm н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> уже груди;<br />

гл<strong>а</strong>з<strong>а</strong> умеренн<strong>о</strong>й <strong>в</strong>еличины. Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие.<br />

Брюш<strong>к</strong><strong>о</strong> из х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метных четырех <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

<strong>с</strong><strong>в</strong>етлее груди. Длин<strong>а</strong> тел<strong>а</strong> 2,75, <strong>к</strong>рыл<strong>а</strong> —<br />

2,5 мм (ри<strong>с</strong>. 1080 А, Б). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО EOMYIIDAE ROHDENDORF, FAM. NOV.<br />

Крыл<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>с</strong>лег<strong>к</strong><strong>а</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>е.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние нежн<strong>о</strong>е, пл<strong>о</strong>х<strong>о</strong> р<strong>а</strong>зличим<strong>о</strong>е;<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метн<strong>а</strong> лишь С п<strong>о</strong> переднему <strong>к</strong>р<strong>а</strong>ю.<br />

Тел<strong>о</strong> <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й <strong>с</strong>пин<strong>к</strong><strong>о</strong>й, неб<strong>о</strong>льш<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>а</strong>нтенн<strong>а</strong>ми и неб<strong>о</strong>льшими <strong>с</strong>л<strong>о</strong>жными<br />

гл<strong>а</strong>з<strong>а</strong>ми и брюш<strong>к</strong><strong>о</strong>м, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящим из <strong>с</strong>еми<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие, цеп<strong>к</strong>ие. Р<strong>а</strong>змеры<br />

мел<strong>к</strong>ие. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Eomyia Rohdendorf, gen nov. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Е. veterrima Rohdendorf, sp. nov.; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Ще<strong>к</strong>и<br />

лишь ед<strong>в</strong><strong>а</strong> уже <strong>в</strong>ы<strong>с</strong><strong>о</strong>ты гл<strong>а</strong>з<strong>а</strong>. Антенны <strong>с</strong> <strong>о</strong><strong>к</strong>руглым<br />

третьим члени<strong>к</strong><strong>о</strong>м, не<strong>с</strong>ущим <strong>к</strong><strong>о</strong>нце<strong>в</strong>ую<br />

<strong>а</strong>ри<strong>с</strong>ту: <strong>к</strong><strong>о</strong>нец брюш<strong>к</strong><strong>а</strong> притуплённый.<br />

Длин<strong>а</strong> тел<strong>а</strong> 2,25 мм, <strong>к</strong>рыл<strong>а</strong>—1,5 мм (ри<strong>с</strong>.<br />

1081). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО PROTOCYRTIDAE ROHDENDORF, 195&<br />

Тел<strong>о</strong> и <strong>к</strong>рыл<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>реп<strong>к</strong><strong>о</strong>е,<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличим<strong>о</strong>е; R прям<strong>а</strong>я; RS без<br />

<strong>в</strong>ет<strong>в</strong>ей, дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я; М д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>т<strong>а</strong>я. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я; грудь <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я; брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е;<br />

н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие, цеп<strong>к</strong>ие. Р<strong>а</strong>змеры неб<strong>о</strong>льшие.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Protocyrtus Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. jurassicus Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). П<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>. Ще<strong>к</strong><strong>а</strong> <strong>о</strong>чень шир<strong>о</strong><strong>к</strong><strong>а</strong>я,<br />

<strong>в</strong>здут<strong>а</strong>я <strong>с</strong>з<strong>а</strong>ди. Сегмент<strong>а</strong>ция брюш<strong>к</strong><strong>а</strong><br />

нея<strong>с</strong>н<strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> 5,0 мм, <strong>к</strong>рыл<strong>а</strong> — 3,5 мм<br />

(ри<strong>с</strong>. 1082). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.

Ри<strong>с</strong>. 1084—1092. Ин<br />

1084. Erinna atra Fabricius; <strong>о</strong>бщий <strong>в</strong>ид, х4,2, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Verrall, 1909), 1085. Rachicerus sp.; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>в</strong> пр<strong>о</strong>филь, х18,0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем., С. Амери<strong>к</strong><strong>а</strong> (Curran, 1934). 1086. Acrocera sanguined<br />

Meigen; <strong>о</strong>бщий <strong>в</strong>ид, Х6,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Sack, 1936). 1087.<br />

Eostratiomyia avia Rohdendorf; <strong>к</strong>рыл<strong>о</strong>, X7,6, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1951). 1088. Rhagionempis tabanicornis Rohdendorf;<br />

А — <strong>к</strong>рыл<strong>о</strong>, Х15.0; Б — <strong>о</strong>бщий <strong>в</strong>ид, Х14,0, <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>-<br />

335<br />

<strong>а</strong><strong>о</strong>тряд Asilomorpha<br />

з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1089. Archirhagio obscurus Rohdendorf;<br />

А — <strong>к</strong>рыл<strong>о</strong>, х4,0; Б — <strong>о</strong>бщий <strong>в</strong>ид, Х2,4, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1090. Protorhagio capitatus Rohdendorf;<br />

<strong>к</strong>рыл<strong>о</strong>, х9,3, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938).<br />

1091. Rhagiophryne bianalis RohdendorfjKpbwio, X 12,0, <strong>в</strong>. юр<strong>а</strong>,.<br />

Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1951). 1092. Silvius viiuli Fabricius;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Ол<strong>с</strong>уфье<strong>в</strong>, 1937)<br />

http://jurassic.ru/

СЕМЕЙСТВО STRATIOMYIIDAE NEWMAN, 1834<br />

(nom. transl. Rohdendorf, hie<br />

(ex Stratiomites Newman, 1834)]<br />

С не з<strong>а</strong>х<strong>о</strong>дит н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; передние жил<strong>к</strong>и <strong>к</strong>репче<br />

з<strong>а</strong>дних и <strong>с</strong>д<strong>в</strong>инуты <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Тел<strong>о</strong> упл<strong>о</strong>щенн<strong>о</strong>е.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я, х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий.<br />

Р<strong>а</strong>змеры <strong>с</strong>редние или <strong>к</strong>рупные, ред<strong>к</strong><strong>о</strong> мел<strong>к</strong>ие<br />

(ри<strong>с</strong>. 1083). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>г<strong>а</strong>т<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (<strong>с</strong><strong>в</strong>ыше 1600 <strong>в</strong>ид<strong>о</strong><strong>в</strong>)<br />

; <strong>в</strong> третичных ф<strong>а</strong>уни<strong>с</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>х<br />

<strong>о</strong>бн<strong>а</strong>ружен<strong>о</strong> ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

СЕМЕЙСТВО XYLOPHAGIDAE FALLEN, 1810<br />

[nom. transl. Stephens, 1829<br />

(ex Xylophagei Fallen, 1810)]<br />

(Erinnidae, Kertesz, 1908)<br />

С не з<strong>а</strong>х<strong>о</strong>дит н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

не<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, не <strong>с</strong>д<strong>в</strong>инут<strong>о</strong>е <strong>к</strong> переднему<br />

<strong>к</strong>р<strong>а</strong>ю; интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я; М 4<br />

<strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т mcu и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>.<br />

Передняя г<strong>о</strong>лень <strong>с</strong> <strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>о</strong>й шп<strong>о</strong>р<strong>о</strong>й. Тел<strong>о</strong> удлиненн<strong>о</strong>е,<br />

р<strong>а</strong>змеры <strong>с</strong>редние или <strong>к</strong>рупные<br />

(ри<strong>с</strong>. 1084). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны, из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО RACHICERIDAE HANDLIRSCH, 1907<br />

С не з<strong>а</strong>х<strong>о</strong>дит н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

не<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е; интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я; М 4 <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т mcu и <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

М 3. Передние г<strong>о</strong>лени без шп<strong>о</strong>р. Антенны мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из 20—36 члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1085). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны из<strong>в</strong>е<strong>с</strong>тен из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ACROCERIDAE LEACH, 1815<br />

[nom. transl. Leach, 1819 (ex Acrocerides Leach, 1815)]<br />

(Cyrtidae Hendel, 1928)<br />

С <strong>о</strong>бх<strong>о</strong>дит <strong>в</strong><strong>с</strong>е <strong>к</strong>рыл<strong>о</strong> или <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

ег<strong>о</strong> <strong>в</strong>ершине. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние изменчи<strong>в</strong><strong>о</strong>е, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е<br />

или р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>пределенн<strong>о</strong>е<br />

п<strong>о</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>а</strong>; брюш<strong>к</strong><strong>о</strong><br />

<strong>с</strong>ильн<strong>о</strong> у<strong>в</strong>еличенн<strong>о</strong>е. Антенны <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные,<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Очень <strong>к</strong>рупные грудные<br />

чешуй<strong>к</strong>и (ри<strong>с</strong>. 1086). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

<strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО TABANIDEA<br />

Пр<strong>о</strong>пле<strong>в</strong>риты <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены <strong>о</strong>т пр<strong>о</strong><strong>с</strong>тернит<strong>а</strong>,<br />

не з<strong>а</strong>мы<strong>к</strong><strong>а</strong>я переднюю т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong>ую ям<strong>к</strong>у. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

п<strong>о</strong>лн<strong>о</strong>е, не<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е. С п<strong>о</strong>чти<br />

336<br />

http://jurassic.ru/<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>бх<strong>о</strong>дит <strong>в</strong>е<strong>с</strong>ь <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я. Эмп<strong>о</strong>дий <strong>в</strong> <strong>в</strong>иде пуль<strong>в</strong>иллы,<br />

шир<strong>о</strong><strong>к</strong>ий. Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Е<strong>о</strong>stratiomyiidae,<br />

Rhagionempididae, Rhagionidae,<br />

Coenomyiidae, Tabanidae, Nemestrinidae, Acanthomeridae;<br />

п<strong>о</strong><strong>с</strong>леднее пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> лишь <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО EOSTRATIOMYIIDAE ROHDENDORF,<br />

1951<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины <strong>к</strong>рыл<strong>а</strong>, <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я;<br />

имеют<strong>с</strong>я п<strong>о</strong>перечные жил<strong>к</strong>и между R, RSi +2 и<br />

RS 3+ 4- Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>перечных rm. Крыл<strong>о</strong> <strong>с</strong><br />

прямым передним <strong>к</strong>р<strong>а</strong>ем и чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленным<br />

<strong>а</strong>пе<strong>к</strong><strong>с</strong><strong>о</strong>м. М 4 <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>т mcu; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

RS3+4 длинный и уз<strong>к</strong>ий. Р<strong>а</strong>змеры <strong>к</strong>рупные.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Eostratiomyia Rohdendorf, 1951. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Е. avia Rohdendorf, 1951; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Четыре п<strong>о</strong>перечные<br />

жил<strong>к</strong>и rm; п<strong>о</strong>перечные между R и<br />

RSp+2 и между RSi+2 и RS 3+ 4 р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не интер<strong>в</strong><strong>а</strong>л<strong>а</strong> между п<strong>о</strong>перечными жил<strong>к</strong><strong>а</strong>ми<br />

rmi и rm 2 неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ния<br />

SC; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS 3+ 4 н<strong>а</strong> <strong>к</strong><strong>о</strong>нце рез<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширенный;<br />

<strong>в</strong><strong>с</strong>е ячей<strong>к</strong>и М <strong>о</strong>т<strong>к</strong>рытые, т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong><br />

и ячей<strong>к</strong><strong>а</strong> С. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 7 мм, тел<strong>а</strong> — <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

10 мм (ри<strong>с</strong>. 1087). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО RHAGIONEMPIDIDAE ROHDENDORF,<br />

FAM. NOV.<br />

SC <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong>реп<strong>к</strong><strong>а</strong>я; д<strong>о</strong>п<strong>о</strong>лнительных<br />

п<strong>о</strong>перечных нет; передний <strong>к</strong>р<strong>а</strong>й<br />

<strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лый, <strong>а</strong>пе<strong>к</strong><strong>с</strong> нерез<strong>о</strong><strong>к</strong>; М 4 <strong>о</strong>т<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong>т ячей<strong>к</strong>и; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS 3+ 4 <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и шир<strong>о</strong><strong>к</strong>ий.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> длинн<strong>а</strong>я, <strong>с</strong> шир<strong>о</strong><strong>к</strong>им лб<strong>о</strong>м и<br />

у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными <strong>а</strong>нтенн<strong>а</strong>ми <strong>с</strong> шир<strong>о</strong><strong>к</strong>им третьим<br />

члени<strong>к</strong><strong>о</strong>м и <strong>а</strong>пи<strong>к</strong><strong>а</strong>льн<strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. Р<strong>а</strong>змеры мел<strong>к</strong>ие.<br />

Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Rhagionempis Rohdendorf, 1938. Тип<br />

р<strong>о</strong>д<strong>а</strong>—Rh. tabanlcornis Rohdendorf, 1938; <strong>в</strong>.<br />

юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм,'К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). К<strong>о</strong>нец<br />

SC д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет ур<strong>о</strong><strong>в</strong>ня rm; <strong>в</strong><strong>с</strong>е ячей<strong>к</strong>и М<br />

<strong>о</strong>т<strong>к</strong>рытые. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 3,5 мм, тел<strong>а</strong> — 3,75 мм<br />

(ри<strong>с</strong>. 1088А, Б). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО RHAGIONIDAE LATREILLE, 1802<br />

[nom. transl. Bezzi, 1913 (ex Rhagionides Latreille, 1802)]<br />

SC <strong>к</strong>реп<strong>к</strong><strong>а</strong>я, <strong>в</strong><strong>с</strong>егд<strong>а</strong> не <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины<br />

<strong>к</strong>рыл<strong>а</strong>; <strong>а</strong>пе<strong>к</strong><strong>с</strong> <strong>к</strong>рыл<strong>а</strong> нея<strong>с</strong>н<strong>о</strong> <strong>о</strong>тгр<strong>а</strong>ниченный;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS 3 + 4 длинный, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящий<strong>с</strong>я;<br />

третий члени<strong>к</strong> <strong>а</strong>нтенн <strong>в</strong>здутый, нечлени<strong>с</strong>тый.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, л<strong>о</strong>б <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

уз<strong>к</strong>ий. Р<strong>а</strong>змеры <strong>с</strong>редние, ред<strong>к</strong><strong>о</strong> мел<strong>к</strong>ие. Юр<strong>а</strong> —

ныне. Три юр<strong>с</strong><strong>к</strong>их и <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них пять <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Archirhagio Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

A. obscurus Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Крыл<strong>о</strong><br />

уз<strong>к</strong><strong>о</strong>е и длинн<strong>о</strong>е, без <strong>к</strong>рылыш<strong>к</strong><strong>а</strong>; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

RS 3 + 4 <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и шир<strong>о</strong><strong>к</strong>ий. О<strong>с</strong>н<strong>о</strong><strong>в</strong>ные<br />

р<strong>а</strong>зделы R, М и CuA длинные; р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong><br />

М р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 21,5 мм, <strong>к</strong>рыл<strong>а</strong>—13 мм (ри<strong>с</strong>.<br />

1089<strong>а</strong>). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Protorhagio Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. capitatus Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Крыл<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е. О<strong>с</strong>н<strong>о</strong><strong>в</strong>ные р<strong>а</strong>зделы R, RS и М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>; R и RSi п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> из<strong>о</strong>гнутые,<br />

<strong>с</strong>ильн<strong>о</strong> <strong>с</strong>ближенные; rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> SC; Ai и CuA не <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я.<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> 5 мм (ри<strong>с</strong>. 1090). Один <strong>в</strong>ид.<br />

В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Rhagiophryne Rohdendorf, 1951. Тип<br />

р<strong>о</strong>д<strong>а</strong>— Rh. bianalis Rohdendorf, 1951; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Крыл<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные р<strong>а</strong>зделы R, RS и М <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине <strong>к</strong>рыл<strong>а</strong>;<br />

RSi и R п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я, их <strong>к</strong><strong>о</strong>нцы<br />

уд<strong>а</strong>лены друг <strong>о</strong>т друг<strong>а</strong>; rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>к</strong><strong>о</strong>нц<strong>а</strong> SC; Ai и CuA <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я<br />

перед <strong>к</strong>р<strong>а</strong>ем <strong>к</strong>рыл<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong><br />

4,5 мм (ри<strong>с</strong>. 1091). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО COENOMYIIDAE WESTWOOD, 1840<br />

Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS3+4 <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящий<strong>с</strong>я, <strong>о</strong>х<strong>в</strong><strong>а</strong>ты<strong>в</strong><strong>а</strong>ющий<br />

<strong>в</strong>ершину <strong>к</strong>рыл<strong>а</strong>. Средняя г<strong>о</strong>лень<br />

<strong>с</strong><strong>о</strong> шп<strong>о</strong>р<strong>о</strong>й. Грудн<strong>а</strong>я чешуй<strong>к</strong><strong>а</strong> м<strong>а</strong>л<strong>а</strong>. В<strong>с</strong>е ячей<strong>к</strong>и<br />

М <strong>о</strong>т<strong>к</strong>рытые. Антенны <strong>с</strong> т<strong>о</strong>н<strong>к</strong>им, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ым<br />

третьим члени<strong>к</strong><strong>о</strong>м. Р<strong>а</strong>змеры <strong>к</strong>рупные.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО TABANIDAE LEACH, 1819. СЛЕПНИ.<br />

Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS34.4 д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, рез<strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящий<strong>с</strong>я и <strong>о</strong>х<strong>в</strong><strong>а</strong>ты<strong>в</strong><strong>а</strong>ющий <strong>в</strong>ершину <strong>к</strong>рыл<strong>а</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>о</strong>лющим у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м<br />

и <strong>а</strong>нтенн<strong>а</strong>ми <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенным или<br />

<strong>с</strong>н<strong>а</strong>бженным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми третьим члени<strong>к</strong><strong>о</strong>м.<br />

Х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты грудные чешуй<strong>к</strong>и. Р<strong>а</strong>змеры<br />

<strong>к</strong>рупные, реже <strong>с</strong>редние (ри<strong>с</strong>. 1092). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из них три <strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>гене 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

22 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 337<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО NEMESTRINIDAE MACQUART, 1834<br />

Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> RS3+4 <strong>о</strong>чень уз<strong>к</strong>ий, <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>в</strong>ет<strong>в</strong>ями, <strong>в</strong>ы<strong>в</strong><strong>о</strong>дящими н<strong>а</strong> передний <strong>к</strong>р<strong>а</strong>й,<br />

т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>о</strong>бе передние <strong>в</strong>ет<strong>в</strong>и М. Имеет<strong>с</strong>я<br />

<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>а</strong>я «ди<strong>а</strong>г<strong>о</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>», <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>а</strong>я<br />

<strong>о</strong>трез<strong>к</strong><strong>а</strong>ми RS и М; н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ых<strong>о</strong>дят<br />

лишь CuA и н<strong>а</strong>з<strong>в</strong><strong>а</strong>нн<strong>а</strong>я «ди<strong>а</strong>г<strong>о</strong>н<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>», ин<strong>о</strong>гд<strong>а</strong> не д<strong>о</strong>х<strong>о</strong>дящ<strong>а</strong>я д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Р<strong>а</strong>змеры<br />

<strong>к</strong>рупные (ри<strong>с</strong>. 1093). Юр<strong>а</strong> — ныне. С<strong>в</strong>ыше<br />

200 <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>в</strong>ид<strong>о</strong><strong>в</strong> 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

три пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены немн<strong>о</strong>гими <strong>в</strong>ид<strong>а</strong>ми <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы и не<strong>о</strong>гене С. Амери<strong>к</strong>и. Один р<strong>о</strong>д<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>н из <strong>в</strong>. юры Герм<strong>а</strong>нии.<br />

НАДСЕМЕЙСТВО ASILIDEA<br />

Пр<strong>о</strong>пле<strong>в</strong>риты, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong>литы <strong>с</strong> пр<strong>о</strong><strong>с</strong>тернит<strong>о</strong>м.<br />

Эмп<strong>о</strong>дий т<strong>о</strong>н<strong>к</strong>ий, щетин<strong>к</strong><strong>о</strong><strong>в</strong>идный.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние п<strong>о</strong>лн<strong>о</strong>е, не<strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенн<strong>о</strong>е. Интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я. Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Protomphralidae, Therevidae, Mydaidae,<br />

Asilidae, Omphralidae- и Apioceridae; д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong><strong>с</strong>ледних т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PROTOMPHRALIDAE ROHDENDORF,<br />

1938<br />

Жил<strong>к</strong>и RS <strong>с</strong>д<strong>в</strong>инуты 1<br />

<strong>к</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong> и<br />

<strong>о</strong>бр<strong>а</strong>зуют четыре <strong>в</strong>ет<strong>в</strong>и, <strong>в</strong>ых<strong>о</strong>дящие н<strong>а</strong> передний<br />

<strong>к</strong>р<strong>а</strong>й. Антенны трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, причем<br />

п<strong>о</strong><strong>с</strong>ледний члени<strong>к</strong> <strong>к</strong>рупный и лишенный <strong>а</strong>ри<strong>с</strong>ты.<br />

М и Си <strong>с</strong>л<strong>а</strong>бее передних жил<strong>о</strong><strong>к</strong>. Юр<strong>а</strong>.<br />

Один р<strong>о</strong>д.<br />

Protomphrale Rohdendorf, 1938. Тип р<strong>о</strong>д<strong>а</strong><br />

— P. martynovi Rohdendorf, 1938; <strong>в</strong>. юр<strong>а</strong>,<br />

Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Брюш<strong>к</strong><strong>о</strong><br />

<strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е, ут<strong>о</strong>нч<strong>а</strong>ющее<strong>с</strong>я <strong>к</strong> <strong>к</strong><strong>о</strong>нцу. Крылья<br />

уз<strong>к</strong>ие, <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лым передним <strong>к</strong>р<strong>а</strong>ем. Н<strong>о</strong>ги<br />

цеп<strong>к</strong>ие (ри<strong>с</strong>. 1094). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО THEREVIDAE NEWMAN, 1834<br />

[nom. transl. Westwood, 1840<br />

(ex Therevites Newman, 1834)]<br />

Жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> <strong>к</strong>рыле р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>,<br />

не<strong>с</strong>д<strong>в</strong>инутые; четыре <strong>в</strong>ет<strong>в</strong>и М, <strong>в</strong>ых<strong>о</strong>дящие<br />

н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й. Темя <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м, <strong>а</strong>нтенн<strong>а</strong>ми <strong>с</strong> <strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й.<br />

Тел<strong>о</strong> без щетин<strong>о</strong><strong>к</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> т<strong>о</strong>н<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, бег<strong>а</strong>тельные. Р<strong>а</strong>змеры<br />

<strong>с</strong>редние (ри<strong>с</strong>. 1095). П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

25 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>о</strong>дин из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО MYDAIDAE LATREILLE, 1806<br />

[nom. transl. Coquillet, 1901 (ex Mydasii Latreille, 1806)]<br />

Н<strong>а</strong> з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong> <strong>в</strong>ых<strong>о</strong>дит <strong>о</strong>дн<strong>а</strong>, ред<strong>к</strong><strong>о</strong><br />

д<strong>в</strong>е жил<strong>к</strong>и: <strong>в</strong><strong>с</strong>е <strong>в</strong>ет<strong>в</strong>и р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>темы<br />

и п<strong>о</strong>чти <strong>в</strong><strong>с</strong>е М з<strong>а</strong>гнуты' <strong>в</strong>перед и <strong>в</strong>ых<strong>о</strong>дят н<strong>а</strong>

1003. Nemestrinus jasciatus (Olivier); <strong>о</strong>бщий <strong>в</strong>ид, x2,7,<strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Sack, 1933). 1094. Protomphrale martynovi Rohdendorf;<br />

°бщий <strong>в</strong>ид, X15.0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Р<strong>о</strong>денд<strong>о</strong>рф, 1938)<br />

1095. Thereva nobilitata (Fabricius); <strong>о</strong>бщий <strong>в</strong>ид, X 5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Verrall, 1909). 1096. Philonicus albiceps Meigen; <strong>о</strong>бщий<br />

<strong>в</strong>ид, Х3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Verrall, 1909). 1097. Brachystoma<br />

CuA+A<br />

Ри<strong>с</strong>. 1093—1101. Инфр<strong>а</strong><strong>о</strong>тряд Asilomorpha<br />

338<br />

http://jurassic.ru/<br />

sp.; <strong>к</strong>рыл<strong>о</strong>, х5,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., С. Амери<strong>к</strong><strong>а</strong> (Curran, 1934). 1098.<br />

Empis sp.; <strong>к</strong>рыл<strong>о</strong>, Х3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. 1099. Tachypeza sp;<br />

<strong>к</strong>рыл<strong>о</strong>, Х14.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, 1934). 1100. Hilarimorpha<br />

sp.; <strong>к</strong>рыл<strong>о</strong>, x7,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, 1934). 1101.<br />

Dolichopus popularis Wiedemann; <strong>о</strong>бщий <strong>в</strong>ид, x8,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hen el, 1936)

передний <strong>к</strong>р<strong>а</strong>й. Антенны длинные, <strong>в</strong>здутые н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я, шире груди. Тел<strong>о</strong> без<br />

щетин<strong>о</strong><strong>к</strong>. Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие. Р<strong>а</strong>змеры <strong>к</strong>рупные<br />

(15—50 мм). О<strong>к</strong><strong>о</strong>л<strong>о</strong> 250 <strong>в</strong>ид<strong>о</strong><strong>в</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и из<strong>в</strong>е<strong>с</strong>тны<br />

еще не изученные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители эт<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>.<br />

СЕМЕЙСТВО ASIL1DAE LATREILLE, 1802<br />

[nom. transl. Leach, 1819 (ex Asilici Latreille, 1802)]<br />

Жил<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены 1<br />

н<strong>а</strong> <strong>к</strong>рыле р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>,<br />

не<strong>с</strong>д<strong>в</strong>инутые, ред<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>нцы М <strong>о</strong><strong>с</strong>л<strong>а</strong>бленные.<br />

Темя <strong>в</strong><strong>о</strong>гнут<strong>о</strong>е, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>е ниже <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющих<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong> гл<strong>а</strong>з. Тел<strong>о</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

щетин<strong>к</strong><strong>а</strong>ми. Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие, <strong>с</strong> <strong>к</strong>рупными<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми, цеп<strong>к</strong>ие. Антенны <strong>с</strong> <strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. Р<strong>а</strong>змеры <strong>с</strong>редние или <strong>к</strong>рупные<br />

(ри<strong>с</strong>. 1096). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Б<strong>о</strong>лее 4500 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и не<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО BOMBYLIIDEA<br />

Пр<strong>о</strong>пле<strong>в</strong>риты <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенные, не <strong>с</strong>литые <strong>с</strong><strong>о</strong><br />

<strong>с</strong>тернит<strong>о</strong>м. Эмп<strong>о</strong>дий <strong>в</strong> <strong>в</strong>иде т<strong>о</strong>н<strong>к</strong><strong>о</strong>й щетин<strong>к</strong>и;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенн<strong>о</strong>е; М <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong><br />

чи<strong>с</strong>ле лишь трех или д<strong>а</strong>же д<strong>в</strong>ух жил<strong>о</strong><strong>к</strong>; и<strong>к</strong>термеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие реду<strong>к</strong>ции <strong>в</strong>ет<strong>в</strong>ей М. Н<strong>о</strong>ги<br />

<strong>с</strong>л<strong>а</strong>бые, т<strong>о</strong>н<strong>к</strong>ие. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Bombyliidae.<br />

СЕМЕЙСТВО BOMBYLIIDAE LATREILLE, 1802<br />

[nom. transl. van der Wulp, 1877<br />

(ex Bombylarii Latreille, 1802)]<br />

Антенны <strong>с</strong> удлиненным или ут<strong>о</strong>лщенным<br />

третьим члени<strong>к</strong><strong>о</strong>м, <strong>с</strong>н<strong>а</strong>бженным <strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й<br />

или пуч<strong>к</strong><strong>о</strong>м щетин<strong>о</strong><strong>к</strong>. Крылья <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итым <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием, п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> имеющим<br />

фр<strong>а</strong>гму. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> имеет<strong>с</strong>я удлиненный х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> ш<strong>а</strong>р<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong>здут<strong>а</strong>я<br />

<strong>с</strong>з<strong>а</strong>ди: гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>з<strong>а</strong>ди ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й и <strong>с</strong> делящей<br />

линией. П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 3000<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х немн<strong>о</strong>гие,<br />

еще м<strong>а</strong>л<strong>о</strong> изученные <strong>в</strong>иды пяти р<strong>о</strong>д<strong>о</strong>з.<br />

НАДСЕМЕЙСТВО EMPIDIDEA<br />

Пр<strong>о</strong>пле<strong>в</strong>риты <strong>с</strong>литы <strong>с</strong><strong>о</strong> <strong>с</strong>тернит<strong>о</strong>м, реже <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенные.<br />

Эмп<strong>о</strong>дий <strong>в</strong> <strong>в</strong>иде щетин<strong>к</strong>и. З<strong>а</strong>дняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я или <strong>с</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м;<br />

<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

ред<strong>к</strong><strong>о</strong> длинн<strong>а</strong>я; д<strong>в</strong>е или три <strong>в</strong>ет<strong>в</strong>и М;<br />

интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong>рупн<strong>а</strong>я,<br />

реже <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Empididae, Hilarimorphidae, Dolichopodidae.<br />

СЕМЕЙСТВО EMPIDIDAE LATREILLE, 1804<br />

[nom. transl. Leach, 1819 (ex Empides Latreille, 1804)]<br />

Х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> удлиненный. Кубит<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, ред<strong>к</strong><strong>о</strong> удлиненн<strong>а</strong>я. Ин<strong>о</strong>гд<strong>а</strong><br />

имеют<strong>с</strong>я три <strong>в</strong>ет<strong>в</strong>и М, н<strong>о</strong> <strong>в</strong> эт<strong>о</strong>м <strong>с</strong>луч<strong>а</strong>е <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че з<strong>а</strong>дней <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й. Спин<strong>к</strong><strong>а</strong><br />

груди ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>здут<strong>а</strong>я (ри<strong>с</strong>. 1097—1099). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены и <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

СЕМЕЙСТВО HILARIMORPHIDAE HENDEL, 1936<br />

Пр<strong>о</strong>пле<strong>в</strong>риты <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенные; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я длинн<strong>а</strong>я,<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> (ри<strong>с</strong>. 1100). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Немн<strong>о</strong>гие <strong>в</strong>иды р<strong>о</strong>д<strong>а</strong> Hilarimorpha<br />

Krober <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>дин <strong>в</strong>ид <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО DOL1CHOPOD1DAE LATREILLE, 1809<br />

[nom. transl. Gerstacker, 1863<br />

(ex Dolichopodes Latreille, 1809)]<br />

Пр<strong>о</strong>пле<strong>в</strong>риты <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щенные; интермеди<strong>а</strong>льн<strong>а</strong>я<br />

ячей<strong>к</strong><strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я, <strong>с</strong>лит<strong>а</strong>я <strong>с</strong> з<strong>а</strong>дней <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й;<br />

<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, п<strong>о</strong>чти <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ующ<strong>а</strong>я;<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> лишь д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и RS и, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, д<strong>в</strong>е <strong>в</strong>ет<strong>в</strong>и М. Грудн<strong>о</strong>й <strong>о</strong>тдел <strong>о</strong>чень<br />

<strong>к</strong>рупный, <strong>в</strong>здутый. Н<strong>о</strong>ги бег<strong>а</strong>тельные. О<strong>к</strong>р<strong>а</strong><strong>с</strong><strong>к</strong><strong>а</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, мет<strong>а</strong>лличе<strong>с</strong><strong>к</strong>и зелен<strong>а</strong>я. Тел<strong>о</strong> <strong>с</strong><br />

<strong>к</strong>реп<strong>к</strong>ими щетин<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1101). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. С<strong>в</strong>ыше 3600 <strong>в</strong>ид<strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

ИНФРАОТРЯД PHOROMORPHA<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>в</strong><strong>с</strong>егд<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

причем <strong>в</strong><strong>с</strong>е жил<strong>к</strong>и RS <strong>в</strong>ых<strong>о</strong>дят н<strong>а</strong> передний<br />

<strong>к</strong>р<strong>а</strong>й. Антенны трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>в</strong>близи <strong>к</strong>р<strong>а</strong>я рт<strong>а</strong>. Л<strong>о</strong>б <strong>в</strong><strong>с</strong>егд<strong>а</strong> шир<strong>о</strong><strong>к</strong>ий,<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тизир<strong>о</strong><strong>в</strong><strong>а</strong>нный, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими<br />

щетин<strong>к</strong><strong>а</strong>ми. Юр<strong>а</strong> — ныне. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> —<br />

Phoridea.<br />

НАДСЕМЕЙСТВО PHORIDEA<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние рез<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>с</strong>мещенн<strong>о</strong>е<br />

<strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю. Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Передние жил<strong>к</strong>и R и RS зн<strong>а</strong>чительн<strong>о</strong><br />

т<strong>о</strong>лще других. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Phoridae, Aenigmatiidae, Thaumatoxeniidae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

не н<strong>а</strong>йдены.<br />

339 22*<br />

http://jurassic.ru/

СЕМЕЙСТВО PHORIDAE NEWMAN, 1834<br />

[nom. transl. Haliday, 1851 (ex Phorites Newman, 1834)]<br />

Интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> и р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> н<strong>а</strong>М<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; п<strong>о</strong>перечн<strong>а</strong>я rm' нея<strong>с</strong>н<strong>а</strong>, т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong><br />

<strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>а</strong> и <strong>с</strong>мещен<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию<br />

<strong>к</strong>рыл<strong>а</strong>. Грудн<strong>о</strong>й <strong>о</strong>тдел не н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ет н<strong>а</strong>д г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й.<br />

Н<strong>о</strong>ги <strong>к</strong>реп<strong>к</strong>ие (ри<strong>с</strong>. 1103). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> мн<strong>о</strong>гих р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

ИНФРАОТРЯД<br />

М U S I D О R О М О R Р Н А<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>е<br />

н<strong>а</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е <strong>к</strong>рыл<strong>а</strong>, без <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>ции;<br />

С <strong>о</strong>бх<strong>о</strong>дит <strong>в</strong><strong>с</strong>ё <strong>к</strong>рыл<strong>о</strong> п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю. Лиц<strong>о</strong> пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>е,<br />

л<strong>о</strong>б шир<strong>о</strong><strong>к</strong>ий. Антенны трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong><br />

т<strong>о</strong>н<strong>к</strong><strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

— Lonchopteridae.<br />

Ри<strong>с</strong>. 1102—1104. Инфр<strong>а</strong><strong>о</strong>тряды Phoromorpha, Musidoromorpha<br />

1102. Palaeophora ancestris (Rohdendorf); <strong>о</strong>бщий <strong>в</strong>ид, x25,0, rail, 1909). 1104.-Musidora lutea Panzer; <strong>о</strong>бщий <strong>в</strong>ид, X7,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем.,<br />

<strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н|(Р<strong>о</strong>денд<strong>о</strong>рф, 1938). 1103. Chaetoneuropho- Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Verrall, 1909)<br />

г<strong>а</strong> callginosa Meigen; <strong>о</strong>бщий <strong>в</strong>ид, Х14,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Ver-<br />

PHOROMORPHA INCERTAE<br />

SEDIS<br />

СЕМЕЙСТВО PALAEOPHORIDAE ROHDENDORF,<br />

1951<br />

Интермеди<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> е<strong>с</strong>ть; имеют<strong>с</strong>я<br />

р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong>и н<strong>а</strong> М 3 и RSi+г; д<strong>в</strong>е п<strong>о</strong>перечные rm.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я; грудь <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>здут<strong>а</strong>я, н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющ<strong>а</strong>я<br />

н<strong>а</strong>д г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й; н<strong>о</strong>ги <strong>с</strong>л<strong>а</strong>бые; брюш<strong>к</strong><strong>о</strong><br />

мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е, <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong><strong>о</strong>е. Юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Palaeophora Rohdendorf, 1951 (Archiphora<br />

Rohdendorf, 1938, пе<strong>с</strong> Schmitz, 1929). Тип<br />

р<strong>о</strong>д<strong>а</strong> — Archiphora ancestrix Rohdendorf, 1938;<br />

<strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у).<br />

Р<strong>а</strong>з<strong>в</strong>ил<strong>о</strong><strong>к</strong> М 3 р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен ди<strong>с</strong>т<strong>а</strong>льнее ур<strong>о</strong><strong>в</strong>ня<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и между RS и Mi+г; CuA дуг<strong>о</strong><strong>в</strong>идн<strong>о</strong><br />

из<strong>о</strong>гнут<strong>а</strong>я, не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 2,0 мм, <strong>к</strong>рыл<strong>а</strong>—1,9 мм (ри<strong>с</strong>.<br />

1102). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

340<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО LONCHOPTERIDAE MACQUART, 1835<br />

{nom. transl. Walker, 1851<br />

(ex Lonchopterinae Macquart, 1835)]<br />

(Musidoridae Kertesz, 1909)<br />

Крылья н<strong>а</strong> <strong>в</strong>ершине я<strong>с</strong>н<strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренные. Тел<strong>о</strong><br />

<strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими щетин<strong>к</strong><strong>а</strong>ми. Н<strong>о</strong>ги бег<strong>а</strong>тельные,<br />

<strong>к</strong>реп<strong>к</strong>ие (ри<strong>с</strong>. 1104). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Немн<strong>о</strong>гие<br />

<strong>в</strong>иды р<strong>о</strong>д<strong>а</strong> Musidora Meigen <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне и <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре.<br />

ИНФРАОТРЯД MYIOMORPHA<br />

В<strong>с</strong>е три жил<strong>к</strong>и <strong>с</strong>и<strong>с</strong>темы R <strong>в</strong>ых<strong>о</strong>дят н<strong>а</strong> передний<br />

<strong>к</strong>р<strong>а</strong>й. С ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ет<strong>с</strong>я д<strong>а</strong>лее<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> Mi, т. е. не з<strong>а</strong>х<strong>о</strong>дит з<strong>а</strong> <strong>в</strong>ершину<br />

<strong>к</strong>рыл<strong>а</strong>. Антенны <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>тделены <strong>о</strong>т <strong>к</strong>р<strong>а</strong>я рт<strong>а</strong><br />

б<strong>о</strong>льшим лиц<strong>о</strong>м, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. Личин<strong>к</strong>и безг<strong>о</strong>л<strong>о</strong><strong>в</strong>ые, мет<strong>а</strong>-

и <strong>а</strong>мфипне<strong>в</strong><strong>с</strong>тиче<strong>с</strong><strong>к</strong>ие. Третичные — ныне. Обширн<strong>а</strong>я<br />

и р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я групп<strong>а</strong> д<strong>в</strong>у<strong>к</strong>рылых,<br />

<strong>в</strong>пер<strong>в</strong>ые п<strong>о</strong>я<strong>в</strong>и<strong>в</strong>ш<strong>а</strong>я<strong>с</strong>я г<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене и <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее<br />

<strong>в</strong>ремя з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющ<strong>а</strong>я <strong>с</strong><strong>в</strong>ыше 30 ООО <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

ше<strong>с</strong>ти н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Platypezidea, Syrphidea,<br />

Conopidea, Muscidea, Hippoboscidea, Gastrophilidea;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

1108<br />

1105. Clythia picia Meigen; <strong>о</strong>бщий <strong>в</strong>ид, X8,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Verrall, 1909). 1106. Chrysotoxum jestivum Linnaeus; <strong>о</strong>бщий<br />

НАДСЕМЕЙСТВО PLATYPEZIDEA<br />

Антенны <strong>с</strong> <strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. З<strong>а</strong>дние г<strong>о</strong>лени<br />

и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния з<strong>а</strong>дних л<strong>а</strong>п<strong>о</strong><strong>к</strong> б<strong>о</strong>лее или менее<br />

р<strong>а</strong><strong>с</strong>ширены; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> rm <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> <strong>с</strong>д<strong>в</strong>инут<strong>а</strong> <strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нию <strong>к</strong>рыл<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Platypezidae, Sciadoceridae;<br />

п<strong>о</strong><strong>с</strong>леднее т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО PLATYPEZIDAE FALLEN, 1817<br />

[nom. transl. Walker, 1851<br />

* (ex Platypezinae Fallen, 1817)]<br />

(Clythiidae Kertesz, 1910)<br />

Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличим<strong>а</strong>,<br />

ин<strong>о</strong>гд<strong>а</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> б<strong>о</strong>льш<strong>а</strong>я; SC <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С, длинн<strong>а</strong>я; интермеди<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, имеет<strong>с</strong>я (ри<strong>с</strong>. 1105). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. С<strong>в</strong>ыше 120 <strong>в</strong>ид<strong>о</strong><strong>в</strong> примерн<strong>о</strong><br />

де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них три<br />

р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> и не<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле и из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО PLATYPEZIDEA<br />

Антенны, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й.<br />

З<strong>а</strong>дние г<strong>о</strong>лени и л<strong>а</strong>п<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, не р<strong>а</strong><strong>с</strong>ширены;<br />

rm р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> н<strong>а</strong> <strong>с</strong>ередине <strong>к</strong>рыл<strong>а</strong>.<br />

Ри<strong>с</strong>. 1105—1107. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Platypezidea, Syrphidea<br />

341<br />

Щетин<strong>к</strong>и н<strong>а</strong> теле немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> без лун<strong>к</strong>и н<strong>а</strong>д <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием <strong>а</strong>нтенн.<br />

Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я, п<strong>о</strong>чти<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющ<strong>а</strong>я <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>. Н<strong>а</strong>лет <strong>о</strong>бычн<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит <strong>с</strong>л<strong>а</strong>б<strong>о</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Syrphidae, Pipunculidae.<br />

1107<br />

<strong>в</strong>ид, x2,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Verrall, 1909). 1107. Dorylas campestris<br />

Latreille; <strong>о</strong>бщий <strong>в</strong>ид,5х7,0, [<strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Verrall, 1909)<br />

СЕМЕЙСТВО SYRPHIDAE LATREILLE, 1802<br />

[nom. transl. Leach, 1819 (ex Syrphine Latreille, 1802)]<br />

Передняя <strong>в</strong>ет<strong>в</strong>ь M <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> RS3 +4, <strong>о</strong>бр<strong>а</strong>зуя<br />

з<strong>а</strong>м<strong>к</strong>нутую ячей<strong>к</strong>у; между RS и М, пере<strong>с</strong>е<strong>к</strong><strong>а</strong>я<br />

п<strong>о</strong>перечную rm, идет <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>а</strong>я <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong> — «л<strong>о</strong>жн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong>». Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> умеренн<strong>о</strong>й <strong>в</strong>еличины, <strong>с</strong><br />

шир<strong>о</strong><strong>к</strong>им лиц<strong>о</strong>м, <strong>в</strong> пр<strong>о</strong>филь я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим<br />

перед гл<strong>а</strong>з<strong>а</strong>ми. Р<strong>а</strong>змеры р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>к</strong>рупные (ри<strong>с</strong>. 1106). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 400 <strong>в</strong>ид<strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>; 22 р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PIPUNCULIDAE ZETTERSTEDT, 1844<br />

[nom. transl. Curtis, Walker, 1851<br />

(ex Pipunculini Zetterstedt, 1844)]<br />

(Dorylaidae Kertesz, 1910)<br />

Передняя <strong>в</strong>ет<strong>в</strong>ь M <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> С или <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует — з<strong>а</strong>м<strong>к</strong>нут<strong>о</strong>й ячей<strong>к</strong>и не <strong>о</strong>бр<strong>а</strong>зует<strong>с</strong>я.<br />

Д<strong>о</strong>п<strong>о</strong>лнительн<strong>о</strong>й пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и —<br />

«л<strong>о</strong>жн<strong>о</strong>й жил<strong>к</strong>и» — нет. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong><br />

<strong>о</strong>чень <strong>в</strong>ели<strong>к</strong><strong>а</strong>; л<strong>о</strong>б и лиц<strong>о</strong> <strong>о</strong>чень уз<strong>к</strong>ие, <strong>в</strong> пр<strong>о</strong>филь<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем не <strong>в</strong>ыд<strong>а</strong>ющие<strong>с</strong>я. Р<strong>а</strong>змеры мел<strong>к</strong>ие<br />

(ри<strong>с</strong>. 1107). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 350 <strong>в</strong>ид<strong>о</strong><strong>в</strong> немн<strong>о</strong>гих р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

http://jurassic.ru/

НАДСЕМЕЙСТВО CONOPIDEA<br />

(Acalyptrata; Holometopa)<br />

Антенны п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й,<br />

реже <strong>о</strong>н<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> н<strong>а</strong> <strong>к</strong><strong>о</strong>нце третьег<strong>о</strong> члени<strong>к</strong><strong>а</strong><br />

или <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> п<strong>о</strong>перечн<strong>о</strong>й<br />

щелью и лун<strong>к</strong><strong>о</strong>й н<strong>а</strong>д <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием <strong>а</strong>нтенн.<br />

Грудн<strong>а</strong>я и <strong>к</strong>рыл<strong>о</strong><strong>в</strong><strong>а</strong>я чешуй<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

или <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. Mi, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет<br />

<strong>в</strong> <strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong><strong>с</strong>егд<strong>а</strong> без угл<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>г<strong>о</strong> изгиб<strong>а</strong>.<br />

Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, ред<strong>к</strong><strong>о</strong><br />

длинн<strong>а</strong>я. Преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> бле<strong>с</strong>тящие, лишенные<br />

н<strong>а</strong>лет<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Очень р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я<br />

групп<strong>а</strong>, <strong>в</strong><strong>к</strong>люч<strong>а</strong>ющ<strong>а</strong>я не менее 13 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Conopidae, Ortalididae, Neriidae, Sepseidae,<br />

Sciomyzidae, Lauxaniidae, Drosophilidae,<br />

Chloropidae; <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pyrgotidae, Tachiniscidae,<br />

Agromyzidae, Celyphidae и Milichiidae <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых ф<strong>а</strong>ун<strong>а</strong>х д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р не <strong>о</strong>бн<strong>а</strong>ружены.<br />

Си<strong>с</strong>тем<strong>а</strong> эт<strong>о</strong>г<strong>о</strong> н<strong>а</strong>деемей<strong>с</strong>т<strong>в</strong><strong>а</strong> изучен<strong>а</strong> еще<br />

м<strong>а</strong>л<strong>о</strong>.<br />

СЕМЕЙСТВО CONOPIDAE LATREILLE, 1802<br />

[nom. transl. Stephens, 1829<br />

(ex Conopariae Latreille, 1802)]<br />

Mi <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> RS3-H или <strong>с</strong>ильн<strong>о</strong><br />

<strong>с</strong> нею <strong>с</strong>ближен<strong>а</strong>. Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, длинн<strong>а</strong>я. С<strong>к</strong>улы и ще<strong>к</strong>и шир<strong>о</strong><strong>к</strong>ие, х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> длинный. Ари<strong>с</strong>т<strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>а</strong>я. П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 600 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

примерн<strong>о</strong> 25 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>дин<br />

р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ORTALIDIDAE FALLEN, 1810<br />

[nom. transl. Macquart, 1835 (ex Ortalides Fallen, 1810)]<br />

(Trypetidae schiner, 1863; Trypaneidae<br />

Bezzi, 1911; Otitidae Hennig, 1950)<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong> <strong>в</strong> <strong>в</strong>иде б<strong>о</strong>лее или<br />

менее удлиненн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong>. Ячей<strong>к</strong>и<br />

<strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я и з<strong>а</strong>дняя <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены. Н<strong>о</strong>ги нет<strong>о</strong>н<strong>к</strong>ие, бег<strong>а</strong>тельные<br />

или цеп<strong>к</strong>ие, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными щетин<strong>к</strong><strong>а</strong>ми.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

б<strong>о</strong>лее 4200 <strong>в</strong>ид<strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>пределяющих<strong>с</strong>я <strong>с</strong>реди<br />

мн<strong>о</strong>гих п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, триб и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>их <strong>с</strong><strong>о</strong>тен<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>; пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО NERIIDAE HALIDAY, 1839<br />

[nom. transl. Hendel, 1922 (ex Neriides Haliday, 1839)]<br />

(Tylidae Hendel, 1928)<br />

Яйце<strong>к</strong>л<strong>а</strong>д мяг<strong>к</strong>ий, <strong>в</strong>тяги<strong>в</strong><strong>а</strong>ющий<strong>с</strong>я <strong>в</strong> брюш<strong>к</strong><strong>о</strong>,<br />

без б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

нередуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е. Н<strong>о</strong>ги рез<strong>к</strong><strong>о</strong> удлиненные,<br />

х<strong>о</strong>дуле<strong>о</strong>бр<strong>а</strong>зные. С без р<strong>а</strong>зры<strong>в</strong><strong>о</strong><strong>в</strong>. П<strong>а</strong>­<br />

342<br />

http://jurassic.ru/<br />

ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 550 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их; <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong>бн<strong>а</strong>ружены<br />

еще не изученные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители трибы<br />

Trepidariini.<br />

СЕМЕЙСТВО SEPSIDAE MACQUART, 1835<br />

Яйце<strong>к</strong>л<strong>а</strong>д без у<strong>в</strong>еличенн<strong>о</strong>г<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>.<br />

С <strong>с</strong> р<strong>а</strong>зры<strong>в</strong><strong>о</strong>м или цельн<strong>а</strong>я. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

п<strong>о</strong>лн<strong>о</strong>е. Преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> мел<strong>к</strong>ие мухи<br />

черн<strong>о</strong>й, бле<strong>с</strong>тящей <strong>о</strong><strong>к</strong>р<strong>а</strong><strong>с</strong><strong>к</strong>и, имеющие <strong>о</strong>бли<strong>к</strong><br />

мур<strong>а</strong><strong>в</strong>ье<strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. С<strong>в</strong>ыше 500 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны ше<strong>с</strong>ти п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>в</strong>з<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шения<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> изучены';<br />

четыре р<strong>о</strong>д<strong>а</strong> <strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО SCIOMYZIDAE FALLEN, 1820<br />

[nom. transl. Macquart, 1835<br />

(ex Sciomyzides Fallen, 1820)]<br />

Яйце<strong>к</strong>л<strong>а</strong>д <strong>в</strong>тянутый. С, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, без<br />

р<strong>а</strong>зры<strong>в</strong><strong>о</strong><strong>в</strong>. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние п<strong>о</strong>лн<strong>о</strong>е, без реду<strong>к</strong>ций;<br />

з<strong>а</strong>тыл<strong>о</strong>чные щетин<strong>к</strong>и р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я. Средней<br />

<strong>в</strong>еличины, ин<strong>о</strong>гд<strong>а</strong> <strong>к</strong>рупные мухи, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>с</strong><strong>в</strong>етл<strong>о</strong>й <strong>о</strong><strong>к</strong>р<strong>а</strong><strong>с</strong><strong>к</strong>и и <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> б<strong>о</strong>лее или менее<br />

р<strong>а</strong>з<strong>в</strong>итым м<strong>а</strong>т<strong>о</strong><strong>в</strong>ым н<strong>а</strong>лет<strong>о</strong>м н<strong>а</strong> теле, <strong>к</strong>реп<strong>к</strong>ими<br />

н<strong>о</strong>г<strong>а</strong>ми и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им брюш<strong>к</strong><strong>о</strong>м. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

три м<strong>а</strong>л<strong>о</strong> изученных р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО LAUXANIIDAE MACQUART, 1835<br />

Яйце<strong>к</strong>л<strong>а</strong>д <strong>в</strong>тянутый. С цельн<strong>а</strong>я или <strong>с</strong> р<strong>а</strong>зры<strong>в</strong><strong>о</strong>м;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние п<strong>о</strong>лн<strong>о</strong>е. З<strong>а</strong>тыл<strong>о</strong>чные щетин<strong>к</strong>и<br />

<strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я, пере<strong>к</strong>рещи<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я. Мел<strong>к</strong>ие<br />

мухи р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й <strong>о</strong><strong>к</strong>р<strong>а</strong><strong>с</strong><strong>к</strong>и, <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>о</strong><br />

<strong>с</strong>л<strong>а</strong>быми н<strong>о</strong>г<strong>а</strong>ми, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им тел<strong>о</strong>м. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1900<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> ше<strong>с</strong>ти п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> и б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> чи<strong>с</strong>л<strong>а</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, еще <strong>о</strong>чень м<strong>а</strong>л<strong>о</strong> изученных, <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х.<br />

СЕМЕЙСТВО DROSOPHILIDAE RONDANI, 1856<br />

[nom. transl. Osten-Sacken, 1878<br />

(ex Drosophilina Rondani, 1856)]<br />

Яйце<strong>к</strong>л<strong>а</strong>д <strong>в</strong>тянутый. С <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> <strong>о</strong>дним или<br />

д<strong>в</strong>умя р<strong>а</strong>зры<strong>в</strong><strong>а</strong>ми; SC <strong>в</strong> <strong>в</strong>иде <strong>с</strong>л<strong>а</strong>б<strong>о</strong>й <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и.<br />

З<strong>а</strong>дняя <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я и <strong>к</strong>убит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

<strong>с</strong>литы <strong>с</strong> <strong>с</strong><strong>о</strong><strong>с</strong>едними или редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. З<strong>а</strong>тыл<strong>о</strong>чные<br />

щетин<strong>к</strong>и <strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я или р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я.<br />

Мел<strong>к</strong>ие, реже <strong>с</strong>редней <strong>в</strong>еличины р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>о</strong><strong>к</strong>р<strong>а</strong>шенные мухи <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми<br />

и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им тел<strong>о</strong>м; щетин<strong>к</strong>и н<strong>а</strong> теле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>иты. П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных р<strong>о</strong>д<strong>о</strong><strong>в</strong> четырех п<strong>о</strong>д<strong>с</strong>е-

мей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене и не<strong>о</strong>гене 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CHLOROPIDAE SCHINER, 1864<br />

{nom. transl. Verall, 1901 (exChloropinae Schiner, 1864)]<br />

Яйце<strong>к</strong>л<strong>а</strong>д <strong>в</strong>тянутый. С <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong> р<strong>а</strong>зры<strong>в</strong><strong>о</strong>м,<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я. Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует,<br />

з<strong>а</strong>дняя б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я <strong>с</strong>лит<strong>а</strong> <strong>с</strong> интермеди<strong>а</strong>льн<strong>о</strong>й.<br />

З<strong>а</strong>тыл<strong>о</strong>чные щетин<strong>к</strong>и <strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я.<br />

Мел<strong>к</strong>ие, ред<strong>к</strong><strong>о</strong> <strong>с</strong>редней <strong>в</strong>еличины р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>о</strong><strong>к</strong>р<strong>а</strong>шенные муш<strong>к</strong>и, п<strong>о</strong>чти <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем лишенные<br />

щетин<strong>о</strong><strong>к</strong> н<strong>а</strong> теле. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 1200 <strong>в</strong>ид<strong>о</strong><strong>в</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы 1<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Ри<strong>с</strong>. 1108—1111. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Muscidea, Hippobpscidea<br />

НАДСЕМЕЙСТВО MUSCIDEA<br />

Антенны <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong> чет<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й щелью и<br />

лун<strong>к</strong><strong>о</strong>й н<strong>а</strong>д <strong>а</strong>нтенн<strong>а</strong>ми. Грудные и <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ые<br />

чешуй<strong>к</strong>и п<strong>о</strong>чти <strong>в</strong><strong>с</strong>егд<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, <strong>к</strong>рупные.<br />

Mi ч<strong>а</strong><strong>с</strong>т<strong>о</strong> рез<strong>к</strong><strong>о</strong> з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед, з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>я<strong>с</strong>ь<br />

н<strong>а</strong> <strong>к</strong>р<strong>а</strong>ю <strong>к</strong>рыл<strong>а</strong> ряд<strong>о</strong>м <strong>с</strong> <strong>к</strong><strong>о</strong>нц<strong>о</strong>м RS3+4,<br />

ин<strong>о</strong>гд<strong>а</strong> д<strong>а</strong>же <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь <strong>с</strong> эт<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>зуя<br />

з<strong>а</strong>м<strong>к</strong>нутую ячей<strong>к</strong>у. Кубит<strong>а</strong>льн<strong>а</strong>я ячей<strong>к</strong><strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Тел<strong>о</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> м<strong>а</strong>т<strong>о</strong><strong>в</strong>ым<br />

н<strong>а</strong>лет<strong>о</strong>м и мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными щетин<strong>к</strong><strong>а</strong>ми,<br />

ред<strong>к</strong><strong>о</strong> бле<strong>с</strong>тящее или г<strong>о</strong>л<strong>о</strong>е. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Обширн<strong>а</strong>я групп<strong>а</strong>, <strong>о</strong>бъединяющ<strong>а</strong>я не<br />

менее 6000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. 16 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong><br />

третичных ф<strong>а</strong>ун<strong>а</strong>х <strong>о</strong>бн<strong>а</strong>ружены лишь ше<strong>с</strong>ть:<br />

Cordyluridae, Anthomyiidae, Muscidae, Hypodermatidae,<br />

Sarcophagidae и Tachinidae.<br />

1108. Scopeuma sp.; <strong>о</strong>бщий <strong>в</strong>ид, X3,7, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Curran, x7,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Зимин, 1951). 1111. Glossina ollgocoena<br />

1934). 1109. Phorbia antiqua (Meigen); X6,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем., Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Scudder); <strong>о</strong>бщий <strong>в</strong>ид (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция), Х5,2, ми<strong>о</strong>цен, С. Аме-<br />

(Шт<strong>а</strong><strong>к</strong>ельберг, 1933). 1110. Musca domemtica Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, ' ри<strong>к</strong><strong>а</strong> (Piveteau, 1954)<br />

343<br />

http://jurassic.ru/

СЕМЕЙСТВО CORDYLUR1DAE MACQUART, 1835<br />

Грудны'е и <strong>к</strong>рыл<strong>о</strong><strong>в</strong>ые чешуй<strong>к</strong>и <strong>о</strong>чень м<strong>а</strong>лы.<br />

Тел<strong>о</strong> удлиненн<strong>о</strong>е. Гип<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льные щетин<strong>к</strong>и<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Mi прям<strong>а</strong>я, не <strong>с</strong>ближенн<strong>а</strong>я <strong>с</strong><br />

RS 3+4. Брюш<strong>к</strong><strong>о</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>е, тел<strong>о</strong> и н<strong>о</strong>ги <strong>с</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными т<strong>о</strong>рч<strong>а</strong>щими щетин<strong>к</strong><strong>а</strong>ми. Средней<br />

<strong>в</strong>еличины (ри<strong>с</strong>. 1108). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<br />

них три <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

<strong>о</strong>лиг<strong>о</strong>цене и ми<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ANTHOMY1IDAE RONDANI, 1856<br />

(п<strong>о</strong>т. transl. Low, 1862<br />

(ex Anthomyna Rondani, 1856]<br />

Чешуй<strong>к</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. Тел<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>в</strong>здутым<br />

грудным <strong>о</strong>тдел<strong>о</strong>м. Mi прям<strong>а</strong>я, без изгиб<strong>а</strong>. Гип<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льные<br />

щетин<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Средней<br />

<strong>в</strong>еличины (ри<strong>с</strong>. 1109). П<strong>а</strong>ле<strong>о</strong>ген •— ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong><br />

1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х из<strong>в</strong>е<strong>с</strong>тны м<strong>а</strong>л<strong>о</strong> изученные <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и.<br />

СЕМЕЙСТВО MUSCIDAE FALLEN, 1810<br />

{nom. transl. Robineau Desvoidy, 1830<br />

(ex Muscides Fallen, 1810)]<br />

Чешуй<strong>к</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. Тел<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е<br />

<strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им брюш<strong>к</strong><strong>о</strong>м. Mi<br />

з<strong>а</strong>гнут<strong>а</strong>я <strong>в</strong>перед <strong>в</strong> <strong>в</strong>иде дуги или уг<strong>о</strong>л<strong>к</strong><strong>а</strong>. Гип<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льные<br />

щетин<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Р<strong>а</strong>змеры<br />

<strong>о</strong>чень р<strong>а</strong>зличны (ри<strong>с</strong>. 1110). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 400 <strong>в</strong>ид<strong>о</strong><strong>в</strong> примерн<strong>о</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне. У<strong>к</strong><strong>а</strong>з<strong>а</strong>ния н<strong>а</strong> н<strong>а</strong>х<strong>о</strong>ждение<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей «р<strong>о</strong>д<strong>а</strong>» Musca <strong>в</strong> третичных<br />

ф<strong>а</strong>ун<strong>а</strong>х д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р не п<strong>о</strong>дт<strong>в</strong>ерждены.<br />

СЕМЕЙСТВО HYPODERMATIDAE RONDANI, 1856<br />

[nom. transl. Townsend, 1916<br />

(ex Hypodermina Rondani, 1856)]<br />

Чешуй<strong>к</strong>и умеренн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итые. Тел<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>е,<br />

удлиненн<strong>о</strong>е, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong>е гу<strong>с</strong>тыми <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми,<br />

реже г<strong>о</strong>л<strong>о</strong>е. Mi з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед. Гип<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льные<br />

щетин<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде т<strong>о</strong>н<strong>к</strong>их <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. Пуп<strong>а</strong>рии<br />

<strong>о</strong>чень <strong>к</strong>рупные. П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 30 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х<br />

С. Амери<strong>к</strong>и <strong>о</strong>бн<strong>а</strong>ружены <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и пуп<strong>а</strong>рие<strong>в</strong>,<br />

д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р не изученные.<br />

СЕМЕЙСТВО SARCOPHAGI DAE MACQUART, 1835<br />

[nom. transl. Brauer et Bergenstamm, 1889<br />

(ex Sarcophagii Macquart, 1835)]<br />

Чешуй<strong>к</strong>и <strong>к</strong>рупные. Тел<strong>о</strong> щетини<strong>с</strong>т<strong>о</strong>е, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>т<strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й. Mj <strong>в</strong><strong>с</strong>егд<strong>а</strong> угл<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong><br />

з<strong>а</strong>гнут<strong>а</strong>я <strong>в</strong>перед. Гип<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льные щетин<strong>к</strong>и<br />

<strong>к</strong>реп<strong>к</strong>ие. Пуп<strong>а</strong>рии <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

344<br />

http://jurassic.ru/<br />

ям<strong>к</strong><strong>о</strong>й <strong>с</strong>з<strong>а</strong>ди. Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне. В не<strong>о</strong>ген<strong>о</strong><strong>в</strong>ых и чет<strong>в</strong>ертичных<br />

<strong>о</strong>тл<strong>о</strong>жениях <strong>о</strong>бн<strong>а</strong>ружены <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

пуп<strong>а</strong>рие<strong>в</strong>, еще нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> изученные.<br />

СЕМЕЙСТВО TACHIN1DAE ROBINEAU DESVOIDY,<br />

1830<br />

[nom. transl. Loew, 11862' (ex Tachinariae Robineau,<br />

Desvoidy, 1830)]<br />

Чешуй<strong>к</strong>и <strong>о</strong>чень <strong>к</strong>рупные. Тел<strong>о</strong> <strong>с</strong> длинными<br />

и <strong>к</strong>реп<strong>к</strong>ими щетин<strong>к</strong><strong>а</strong>ми, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> г<strong>о</strong>л<strong>о</strong>й <strong>а</strong>ри<strong>с</strong>т<strong>о</strong>й.<br />

Mi, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, рез<strong>к</strong><strong>о</strong> з<strong>а</strong>гнут<strong>а</strong>я <strong>в</strong> <strong>в</strong>иде<br />

угл<strong>а</strong> или дуги <strong>в</strong>перед. Гип<strong>о</strong>пле<strong>в</strong>р<strong>а</strong>льные щетин<strong>к</strong>и<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> имеют<strong>с</strong>я. Пуп<strong>а</strong>рии без ям<strong>к</strong>и <strong>с</strong>з<strong>а</strong>ди.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне ; В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

б<strong>о</strong>лее 5000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Имеющие<strong>с</strong>я у<strong>к</strong><strong>а</strong>з<strong>а</strong>ния н<strong>а</strong> н<strong>а</strong>х<strong>о</strong>ждения<br />

<strong>в</strong> третичных ф<strong>а</strong>ун<strong>а</strong>х пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> не пр<strong>о</strong><strong>в</strong>ерены.<br />

НАДСЕМЕЙСТВО HIPPOBOSCIDEA<br />

Ари<strong>с</strong>т<strong>а</strong> п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> <strong>в</strong>ет<strong>в</strong>ящими<strong>с</strong>я <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>. Стерниты<br />

брюш<strong>к</strong><strong>а</strong> (<strong>к</strong>р<strong>о</strong>ме пер<strong>в</strong><strong>о</strong>г<strong>о</strong>) <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные;<br />

мембр<strong>а</strong>н<strong>а</strong> брюш<strong>к</strong><strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я,<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>т<strong>а</strong>я. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

<strong>с</strong>д<strong>в</strong>инут<strong>о</strong>е <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю; ин<strong>о</strong>гд<strong>а</strong><br />

<strong>к</strong>рылья редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нные. Жи<strong>в</strong><strong>о</strong>р<strong>о</strong>дящие (<strong>к</strong>у<strong>к</strong>л<strong>о</strong>р<strong>о</strong>дные).<br />

Облиг<strong>а</strong>тные <strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong>ы н<strong>а</strong> мле<strong>к</strong><strong>о</strong>пит<strong>а</strong>ющих<br />

и птиц<strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Glossinidae, Hippoboscidae.<br />

СЕМЕЙСТВО GLOSSINIDAE TOWNSEND, 1935<br />

Крылья б<strong>о</strong>льшие, умеренн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нные:<br />

имеет<strong>с</strong>я х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я <strong>в</strong>ершинн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть жил<strong>к</strong>и Mi+2, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бляющ<strong>а</strong>я ячей<strong>к</strong>у R5.<br />

Тел<strong>о</strong> не <strong>с</strong>плющенн<strong>о</strong>е <strong>с</strong><strong>в</strong>ерху <strong>в</strong>низ, <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щими<br />

<strong>а</strong>нтенн<strong>а</strong>ми и т<strong>о</strong>н<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми. Р<strong>а</strong>змеры'<br />

<strong>к</strong>рупные (ри<strong>с</strong>. 1111). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 <strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>о</strong>д<strong>а</strong> Glossiп<strong>а</strong><br />

Wiedemann <strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й Афри<strong>к</strong>е; <strong>о</strong>дин<br />

<strong>в</strong>ид эт<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> ми<strong>о</strong>цене С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО HIPPOBOSCIDAE LEACH, 1815<br />

Крылья <strong>с</strong> <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>а</strong>цией<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния, ин<strong>о</strong>гд<strong>а</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные<br />

или д<strong>а</strong>же <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ующие. Вершинн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть<br />

М) + 2 не<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я и не з<strong>а</strong>гнут<strong>а</strong>я <strong>в</strong>перед.<br />

Тел<strong>о</strong> упл<strong>о</strong>щенн<strong>о</strong>е, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими,<br />

цеп<strong>к</strong>ими н<strong>о</strong>г<strong>а</strong>ми. Р<strong>а</strong>змеры <strong>с</strong>редние. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong><strong>в</strong>ыше 100<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>о</strong>дин <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Герм<strong>а</strong>нии.<br />

.

Вт<strong>о</strong>ричн<strong>о</strong>бе<strong>с</strong><strong>к</strong>рылые мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленные <strong>к</strong> п<strong>а</strong>р<strong>а</strong>зитизму н<strong>а</strong> тепл<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ных<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных. Тел<strong>о</strong> <strong>с</strong>плю<strong>с</strong>нут<strong>о</strong> <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> и<br />

п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными н<strong>а</strong>з<strong>а</strong>д щетин<strong>к</strong><strong>а</strong>ми,<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми или гребнями шип<strong>о</strong><strong>в</strong>. Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые прид<strong>а</strong>т<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>лющих<br />

<strong>с</strong>тилет<strong>о</strong><strong>в</strong>. Н<strong>о</strong>ги прыг<strong>а</strong>тельные: т<strong>а</strong>зи<strong>к</strong>и б<strong>о</strong>льшие,<br />

л<strong>а</strong>п<strong>к</strong>и длинные. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>о</strong>дн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО HYSTRICHOPSYLLIDAE TIRABOSCHI,<br />

1904<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong> гребнями шип<strong>о</strong><strong>в</strong> н<strong>а</strong> ще<strong>к</strong><strong>а</strong>х. Тергиты<br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong> един<strong>с</strong>т<strong>в</strong>енным ряд<strong>о</strong>м щетин<strong>о</strong><strong>к</strong>.<br />

Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> имеют<strong>с</strong>я (ри<strong>с</strong>. 1112). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Из мн<strong>о</strong>гих р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

ОТРЯД APHANIPTERA.^OXH<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

Ри<strong>с</strong>. 1112. Aphaniptera. Pataeopsylla clebsiana Dampf;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 20,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь<br />

(Handlirsch, 1925)<br />

НАДОТРЯД HYMENOPTEROIDEA.<br />

ПЕРЕПОНЧАТОКРЫЛООБРАЗНЫЕ<br />

Крылья переп<strong>о</strong>нч<strong>а</strong>тые, рез<strong>к</strong><strong>о</strong> гетер<strong>о</strong>н<strong>о</strong>мные,<br />

п<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ющие х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женную диптеригию:<br />

передние <strong>к</strong>рылья длиннее з<strong>а</strong>дних, <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нные. Ф<strong>о</strong>рм<strong>а</strong> <strong>о</strong>б<strong>о</strong>их<br />

<strong>к</strong>рылье<strong>в</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>тличн<strong>а</strong>я; переднее <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><br />

чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й <strong>в</strong>ершинн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женным термин<strong>а</strong>льным <strong>к</strong>р<strong>а</strong>ем. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

бедн<strong>о</strong>е, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими жил<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими<br />

<strong>к</strong>рупные ячей<strong>к</strong>и, и <strong>о</strong>бычн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>е,<br />

лишенн<strong>о</strong>е жил<strong>о</strong><strong>к</strong>, п<strong>о</strong>ле у термин<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong>р<strong>а</strong>я.<br />

Р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны грызущие, при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленные<br />

и для прием<strong>а</strong> жид<strong>к</strong><strong>о</strong>й пищи. Личин<strong>к</strong>и эру<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

или чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зные, <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими н<strong>о</strong>­<br />

(Б. Б. Р<strong>о</strong>денд<strong>о</strong>рф)<br />

г<strong>а</strong>ми или безн<strong>о</strong>гие. Жи<strong>в</strong>ут <strong>в</strong> т<strong>к</strong><strong>а</strong>нях р<strong>а</strong><strong>с</strong>тений<br />

или н<strong>а</strong> их п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти, мн<strong>о</strong>гие п<strong>а</strong>р<strong>а</strong>зиты других<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит ух<strong>о</strong>д з<strong>а</strong> п<strong>о</strong>т<strong>о</strong>м<strong>с</strong>т<strong>в</strong><strong>о</strong>м.<br />

Н<strong>а</strong>иб<strong>о</strong>лее близ<strong>к</strong>и <strong>к</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным.<br />

Три<strong>а</strong><strong>с</strong> — ныне. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>в</strong>язи <strong>с</strong> другими групп<strong>а</strong>ми н<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>рылых<br />

нея<strong>с</strong>ны; предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ем<strong>о</strong>е р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong> ме<strong>к</strong><strong>о</strong>птер<strong>о</strong>ид<strong>а</strong>ми<br />

м<strong>а</strong>л<strong>о</strong><strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>. И<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие<br />

эт<strong>о</strong>й группы, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>, <strong>о</strong>пределил<strong>о</strong><strong>с</strong>ь<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием р<strong>а</strong>змн<strong>о</strong>жения — прежде<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> ух<strong>о</strong>д<strong>о</strong>м з<strong>а</strong> п<strong>о</strong>т<strong>о</strong>м<strong>с</strong>т<strong>в</strong><strong>о</strong>м. Спр<strong>а</strong><strong>в</strong>едли<strong>в</strong><strong>о</strong> эту<br />

группу <strong>к</strong>рыл<strong>а</strong>тых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong>чит<strong>а</strong>ть н<strong>а</strong>иб<strong>о</strong>лее<br />

пр<strong>о</strong>гре<strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>о</strong>й.<br />

ОТРЯД HYMENOPTERA. ПЕРЕПОНЧАТОКРЫЛЫЕ<br />

Д<strong>в</strong>е п<strong>а</strong>ры переп<strong>о</strong>нч<strong>а</strong>тых, рез<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

<strong>к</strong>рылье<strong>в</strong>; бе<strong>с</strong><strong>к</strong>рылые ф<strong>о</strong>рмы 1<br />

— мур<strong>а</strong><strong>в</strong>ьи,<br />

нем<strong>к</strong>и и др.— ред<strong>к</strong>и; з<strong>а</strong>дние <strong>к</strong>рылья<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че передних, при п<strong>о</strong>лете <strong>с</strong>цепляют<strong>с</strong>я<br />

<strong>с</strong> передними. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>ильн<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> и <strong>о</strong>бычн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>с</strong>д<strong>в</strong>инут<strong>о</strong> <strong>к</strong><br />

переднему <strong>к</strong>р<strong>а</strong>ю; т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряде <strong>с</strong>идячебрюхих<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я <strong>в</strong> т<strong>о</strong>м или ин<strong>о</strong>м <strong>в</strong>иде SC,<br />

у п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их и ж<strong>а</strong>лящих переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых<br />

<strong>о</strong>н<strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> р<strong>а</strong>диу<strong>с</strong><strong>о</strong>м; RS делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong>е<br />

(О. М. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong>)<br />

345<br />

http://jurassic.ru/<br />

<strong>в</strong>ет<strong>в</strong>и, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передняя ин<strong>о</strong>гд<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я,<br />

<strong>а</strong> з<strong>а</strong>дняя н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>м пр<strong>о</strong>тяжении <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью меди<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong><strong>о</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины<br />

<strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> д<strong>в</strong>е пр<strong>о</strong><strong>с</strong>тые <strong>в</strong>ет<strong>в</strong>и, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передняя,<br />

МА, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>ью RS,<br />

причем <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние МА <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е;<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> е<strong>с</strong>ть п<strong>о</strong>перечные жил<strong>к</strong>и r-rs и rs-m<br />

(б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>); у п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>.

длинн<strong>а</strong>я и, т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS <strong>о</strong>бычн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>,<br />

<strong>с</strong><strong>о</strong>единяет М <strong>с</strong> R; R и М <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>с</strong>литы; CuA <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я и пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я; у Symphyta<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я CuP <strong>в</strong> <strong>в</strong>иде нея<strong>с</strong>н<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и; <strong>о</strong>дн<strong>а</strong> — д<strong>в</strong>е <strong>а</strong>н<strong>а</strong>льные жил<strong>к</strong>и; между<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием р<strong>а</strong>диу<strong>с</strong><strong>а</strong> и М н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея, г<strong>с</strong>; ее <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л RS и б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>;<br />

между р<strong>а</strong>диу<strong>с</strong><strong>о</strong>м и RS н<strong>а</strong>х<strong>о</strong>дят<strong>с</strong>я р<strong>а</strong>ди<strong>а</strong>льные<br />

ячеи Гь гг. г 3; ячея, <strong>о</strong>гр<strong>а</strong>ниченн<strong>а</strong>я б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й и ч<strong>а</strong><strong>с</strong>тями RS и МА,—<br />

<strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея (sr).<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>бычн<strong>о</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я; р<strong>о</strong>г<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т грызуще-лижущег<strong>о</strong> тип<strong>а</strong>; б<strong>о</strong>льшие<br />

ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные гл<strong>а</strong>з<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы, реже<br />

<strong>о</strong><strong>к</strong>руглые; н<strong>а</strong> темени <strong>о</strong>бычн<strong>о</strong> три гл<strong>а</strong>з<strong>к</strong><strong>а</strong>; <strong>а</strong>нтенны<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й ф<strong>о</strong>рмы <strong>с</strong> тремя и б<strong>о</strong>льше<br />

члени<strong>к</strong><strong>а</strong>ми; пер<strong>в</strong>ые д<strong>в</strong><strong>а</strong> члени<strong>к</strong><strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льных (жгути<strong>к</strong>) ф<strong>о</strong>рм<strong>о</strong>й и <strong>в</strong>еличин<strong>о</strong>й.<br />

Сегменты груди <strong>о</strong>бычн<strong>о</strong> пл<strong>о</strong>тн<strong>о</strong> <strong>с</strong>литы между<br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>ю; н<strong>а</strong>иб<strong>о</strong>лее р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> <strong>с</strong>реднегрудь. Передний<br />

члени<strong>к</strong> брюш<strong>к</strong><strong>а</strong> <strong>с</strong>лит <strong>с</strong> з<strong>а</strong>днегрудью,<br />

у <strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>брюхих <strong>в</strong>х<strong>о</strong>дит <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> пр<strong>о</strong>межут<strong>о</strong>чн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> груди; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong><br />

брюш<strong>к</strong><strong>а</strong> <strong>о</strong>бр<strong>а</strong>зует р<strong>а</strong>зн<strong>о</strong>й длины <strong>с</strong>тебеле<strong>к</strong>.<br />

Обычн<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> пять, реже три — четыре;<br />

н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем члени<strong>к</strong>е д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong> <strong>с</strong> п<strong>о</strong>душеч<strong>к</strong><strong>о</strong>й<br />

между ними.<br />

У б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> личин<strong>к</strong>и чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зные, лишены<br />

гл<strong>а</strong>з и н<strong>о</strong>г; у примити<strong>в</strong>ных Symphyta личин<strong>к</strong>и<br />

<strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми грудных н<strong>о</strong>г. Ку<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. Кр<strong>о</strong>ме <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong>и, <strong>в</strong><strong>с</strong>е<br />

ф<strong>а</strong>зы м<strong>о</strong>гут зим<strong>о</strong><strong>в</strong><strong>а</strong>ть. Взр<strong>о</strong><strong>с</strong>лые пит<strong>а</strong>ют<strong>с</strong>я не<strong>к</strong>т<strong>а</strong>р<strong>о</strong>м,<br />

<strong>с</strong><strong>а</strong>х<strong>а</strong>ри<strong>с</strong>тыми <strong>в</strong>ыделениями тлей, чер<strong>в</strong>ец<strong>о</strong><strong>в</strong><br />

и др., ред<strong>к</strong><strong>о</strong> жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>й пищей. Личин<strong>к</strong>и пит<strong>а</strong>ют<strong>с</strong>я<br />

р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>й или жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>й пищей.<br />

Три<strong>а</strong><strong>с</strong> — ныне. П<strong>о</strong>д<strong>о</strong>тряды: Symphyta, Apocrita.<br />

ПОДОТРЯД SYMPHYTA.<br />

СИДЯЧЕБРЮХИЕ<br />

ПЕРЕПОНЧАТОКРЫЛЫЕ<br />

Крылья <strong>о</strong>бычн<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

передние. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лн<strong>о</strong>е <strong>в</strong> <strong>о</strong>тряде;<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> е<strong>с</strong>ть SC или ее рудименты <strong>в</strong> <strong>в</strong>иде<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я и ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я; <strong>а</strong>н<strong>а</strong>льных<br />

жил<strong>о</strong><strong>к</strong> <strong>о</strong>дн<strong>а</strong> — д<strong>в</strong>е; <strong>в</strong> з<strong>а</strong>дних <strong>к</strong>рыльях неб<strong>о</strong>льш<strong>а</strong>я<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть. Вертлуг <strong>в</strong><strong>с</strong>егд<strong>а</strong> д<strong>в</strong>учлени<strong>с</strong>тый;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong> пять. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>идячее,<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong> яйце<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м р<strong>а</strong>зличн<strong>о</strong>й ф<strong>о</strong>рмы;<br />

<strong>о</strong>бычн<strong>о</strong> <strong>о</strong>н <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из д<strong>в</strong>ух <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

з<strong>а</strong>зубренных пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>. Личин<strong>к</strong>и р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные,<br />

<strong>с</strong> грудными и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льными<br />

н<strong>о</strong>г<strong>а</strong>ми. Три<strong>а</strong><strong>с</strong> — ныне. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Megalodontidea, Siricidea, Orussidea, Сеphidea,<br />

Tenthredinidea.<br />

346<br />

http://jurassic.ru/<br />

НАДСЕМЕЙСТВО MEGALODONTIDEA<br />

В передних <strong>к</strong>рыльях SC ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я;<br />

между R и RS три п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ные п<strong>о</strong> р<strong>а</strong>змеру<br />

р<strong>а</strong>ди<strong>а</strong>льные ячеи; р<strong>а</strong>зделяющие их п<strong>о</strong>перечные<br />

жил<strong>к</strong>и прямые и п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ные п<strong>о</strong> длине;<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> е<strong>с</strong>ть. Три<strong>а</strong><strong>с</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Anaxyelidae, Xyelidae, Pamphilidae, Меgalodontidae;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

не из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

СЕМЕЙСТВО ANAXYELIDAE MARTYNOV, 1925<br />

В передних <strong>к</strong>рыльях <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й SC нет; RSi<br />

без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>; ячея г 3 длинн<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> длине<br />

д<strong>в</strong>ум пер<strong>в</strong>ым ячеям <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятьгм. Личин<strong>к</strong><strong>а</strong><br />

неиз<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>. В. юр<strong>а</strong>. Один р<strong>о</strong>д.<br />

Anaxyela Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong> —<br />

A. gracilis Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). В <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м п<strong>о</strong>ле переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> четыре п<strong>о</strong>перечные жил<strong>к</strong>и;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>а</strong>я; п<strong>о</strong>перечные жил<strong>к</strong>и r-rs<br />

п<strong>а</strong>р<strong>а</strong>ллельны; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и<br />

RS <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 7 мм,<br />

яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong> — 4,6 мм (ри<strong>с</strong>. 1113). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

B. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО XYELIDAE HALIDAY, 1840<br />

В предел<strong>а</strong>х <strong>к</strong>рыльях SC <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>о</strong>м; ячеи ri и г 3 п<strong>о</strong>чти<br />

<strong>о</strong>дн<strong>о</strong>й длины, пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тые, длиннее Гг; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея и <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я ячея между <strong>в</strong>ет<strong>в</strong>ями<br />

RS р<strong>а</strong><strong>в</strong>ны' между <strong>с</strong><strong>о</strong>б<strong>о</strong>й и имеют удлиненную<br />

ф<strong>о</strong>рму. Антенны 9—12-тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; <strong>о</strong>т<br />

других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я <strong>с</strong>ильн<strong>о</strong> удлиненным<br />

и ут<strong>о</strong>лщенным третьим члени<strong>к</strong><strong>о</strong>м. Яйце<strong>к</strong>л<strong>а</strong>д<br />

умеренн<strong>о</strong>й длины или <strong>о</strong>чень длинный.<br />

У личин<strong>о</strong><strong>к</strong> н<strong>о</strong>ги н<strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льных<br />

<strong>с</strong>егмент<strong>а</strong>х; жи<strong>в</strong>ут <strong>в</strong> ц<strong>в</strong>ет<strong>а</strong>х и н<strong>а</strong> ли<strong>с</strong>тьях. Пять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне г<strong>о</strong>л<strong>а</strong>р<strong>к</strong>ти<strong>к</strong>и. Три<strong>а</strong><strong>с</strong><br />

— ныне. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

Liadoxyela Martynov, 1937. Тип р<strong>о</strong>д<strong>а</strong> —<br />

L. praecox Martynov, 1937; н. юр<strong>а</strong>, Ср. Азия<br />

(н. лей<strong>а</strong><strong>с</strong>, Кизил-Кия). В передних <strong>к</strong>рыльях<br />

RS н<strong>а</strong>чин<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>о</strong>дн<strong>о</strong>м ур<strong>о</strong><strong>в</strong>не <strong>с</strong> <strong>в</strong>ершин<strong>о</strong>й<br />

SC; <strong>в</strong>ет<strong>в</strong>и р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong> передней <strong>в</strong>ет<strong>в</strong>и RS не з<strong>а</strong>гнуты<br />

<strong>в</strong>перед; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

(rs-m) <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; ячеи dc и п<strong>о</strong><strong>с</strong>ледующ<strong>а</strong>я между<br />

<strong>в</strong>ет<strong>в</strong>ями RS удлиненные. Длин<strong>а</strong> переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong> 12 мм (ри<strong>с</strong>. 1114). Один <strong>в</strong>ид. Н. юр<strong>а</strong><br />

Азии.<br />

Вне СССР: Archexyela Riek, 1955 из три<strong>а</strong><strong>с</strong><strong>а</strong><br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы,<br />

<strong>о</strong>дин — из ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.

Ри<strong>с</strong>. 1113—1119.<br />

1113. Anaxyela gracilis Martynov; <strong>о</strong>бщий <strong>в</strong>ид, x5,0, <strong>в</strong>. юр<strong>а</strong>,<br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 1114. Liadoxyela praecox Martynov;<br />

переднее <strong>к</strong>рыл<strong>о</strong>, Х5,0, н. юр<strong>а</strong>,- Ср. Азия. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937).<br />

1115. Lyda nemoralis Thomson; <strong>о</strong>бщий <strong>в</strong>ид, Х3,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем."(Гу<strong>с</strong><strong>с</strong><strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий,<br />

1935). 1116. Sirex gigas Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид, xl,0,<br />

СЕМЕЙСТВО PAMPHILIDAE DALLA-TORRE, 1894.<br />

П ИЛ ИЛ ЫЦИКИ-ТКАЧ И<br />

[nom. transl. Viereck, 1916<br />

(ex Pamphilinae Dalla-Torre, 1894)]<br />

В передних <strong>к</strong>рыльях SC <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS без р<strong>а</strong>з<strong>в</strong>ил<strong>к</strong><strong>а</strong>, з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед;<br />

ячея г 3 немн<strong>о</strong>г<strong>о</strong> длиннее ячеи п; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея немн<strong>о</strong>г<strong>о</strong> длиннее <strong>с</strong>ледующей ячеи<br />

rs2. Личин<strong>к</strong>и <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми грудными<br />

н<strong>о</strong>г<strong>а</strong>ми; <strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льных н<strong>о</strong>же<strong>к</strong> нет. Жи<strong>в</strong>ут<br />

347<br />

зд<strong>о</strong>тряд Symphyta<br />

http://jurassic.ru/<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Гу<strong>с</strong><strong>с</strong><strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий, 1935). 1117. Paroryssus extensus Martynov;<br />

<strong>о</strong>бщий <strong>в</strong>ид, Х4,0, <strong>в</strong>. юр<strong>а</strong>, Ю.К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925),<br />

1118. Cephas pygmaeus Linnaeus; <strong>о</strong>бщий <strong>в</strong>ид,X5,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Гу<strong>с</strong><strong>с</strong><strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий,<br />

1935). 1119. Pteronidea ribesit Scopoli; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х5,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем.(Гу<strong>с</strong><strong>с</strong><strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий,из «Определителя н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых», 1948)<br />

<strong>о</strong>бще<strong>с</strong>т<strong>в</strong><strong>а</strong>ми н<strong>а</strong> х<strong>в</strong><strong>о</strong>йных и ли<strong>с</strong>т<strong>в</strong>енных дере<strong>в</strong>ьях,<br />

<strong>в</strong> п<strong>а</strong>утине или <strong>с</strong><strong>в</strong>ернутых ли<strong>с</strong>тьях (ри<strong>с</strong>.<br />

1115). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В<strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне г<strong>о</strong>л<strong>а</strong>р<strong>к</strong>ти<strong>к</strong>и; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) и ми<strong>о</strong>цене С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО SIRICIDEA<br />

Три р<strong>а</strong>ди<strong>а</strong>льные ячеи р<strong>а</strong>зных р<strong>а</strong>змер<strong>о</strong><strong>в</strong>;<br />

П меньше г 2; п<strong>о</strong>перечные жил<strong>к</strong>и r-rs <strong>к</strong><strong>о</strong><strong>с</strong>ые,<br />

неп<strong>а</strong>р<strong>а</strong>ллельные, пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я <strong>к</strong><strong>о</strong>р<strong>о</strong>че ди-

<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й; <strong>о</strong>дн<strong>а</strong> <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>. Яйце<strong>к</strong>л<strong>а</strong>д <strong>в</strong><br />

<strong>в</strong>иде иглы или р<strong>о</strong>г<strong>а</strong>. Юр<strong>а</strong> — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Syntexidae, Xiphydriidae, Siricidae, Pseudosiricidae;<br />

д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не<br />

из<strong>в</strong>е<strong>с</strong>тны; п<strong>о</strong><strong>с</strong>леднее <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> из юры 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО SIRICIDAE KIRBY, 1937. РОГОХВОСТЫ<br />

В передних <strong>к</strong>рыльях <strong>с</strong>леды SC; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

RS и б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>о</strong>дну прямую линию; <strong>в</strong>ершин<strong>а</strong> передней <strong>в</strong>ет<strong>в</strong>и<br />

R5 з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние з<strong>а</strong>дней<br />

<strong>в</strong>ет<strong>в</strong>и RS длинн<strong>о</strong>е; ячея dc п<strong>о</strong>чти <strong>в</strong>д<strong>в</strong><strong>о</strong>е<br />

длиннее ячеи гэг. Тел<strong>о</strong> черн<strong>о</strong>е <strong>с</strong> желтым ри<strong>с</strong>ун<strong>к</strong><strong>о</strong>м.<br />

Личин<strong>к</strong>и жи<strong>в</strong>ут <strong>в</strong> дре<strong>в</strong>е<strong>с</strong>ине, лишены<br />

гл<strong>а</strong>з; грудные н<strong>о</strong>ги <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны, <strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льных<br />

н<strong>о</strong>г нет. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ле<strong>с</strong>н<strong>о</strong>й з<strong>о</strong>ны (ри<strong>с</strong>. 1116).<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и ми<strong>о</strong>цене<br />

Ю. Е<strong>в</strong>р<strong>о</strong>пы.<br />

НАДСЕМЕЙСТВО ORUSSIDEA<br />

В передних <strong>к</strong>рыльях <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й SC нет; д<strong>в</strong>е<br />

р<strong>а</strong>ди<strong>а</strong>льные ячеи; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние передней <strong>в</strong>ет<strong>в</strong>и<br />

RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я,<br />

<strong>в</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong> переднюю <strong>в</strong>ет<strong>в</strong>ь RS, <strong>о</strong>бр<strong>а</strong>зуя<br />

единую <strong>в</strong>ы'гнутую <strong>к</strong>з<strong>а</strong>ди жил<strong>к</strong>у; RS 2 + Mi т<strong>о</strong>же<br />

<strong>в</strong>ыгнут<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди; Си и А <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong>литы.<br />

Личин<strong>к</strong>и <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>пеци<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>а</strong>р<strong>а</strong>зиты<br />

жу<strong>к</strong><strong>о</strong><strong>в</strong> Buprestidae и Cerambycidae, жи<strong>в</strong>ут <strong>в</strong><br />

дре<strong>в</strong>е<strong>с</strong>ине. Шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненн<strong>а</strong>я рели<strong>к</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я<br />

групп<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне. Юр<strong>а</strong> —<br />

ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Paroryssidae, Orussidae.<br />

СЕМЕЙСТВО PARORYSSIDAE MARTYNOV, 1925<br />

В переднем <strong>к</strong>рыле между <strong>в</strong>ет<strong>в</strong>ями RS д<strong>в</strong>е<br />

<strong>с</strong>ближенные п<strong>о</strong>перечные жил<strong>к</strong>и; <strong>о</strong>т <strong>в</strong>ет<strong>в</strong>ей М<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь лишь <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

п<strong>о</strong>д прямым угл<strong>о</strong>м <strong>к</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>лу М. В. юр<strong>а</strong>. Один<br />

р<strong>о</strong>д.<br />

Paroryssus Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. extensus Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). К<strong>о</strong><strong>с</strong>т<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й<br />

переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong>реплен; <strong>с</strong>ли<strong>в</strong>шие<strong>с</strong>я<br />

SC и R <strong>о</strong>бр<strong>а</strong>зуют т<strong>о</strong>л<strong>с</strong>тую жил<strong>к</strong>у; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я. Длин<strong>а</strong> тел<strong>а</strong> <strong>с</strong> яйце<strong>к</strong>л<strong>а</strong>д<strong>о</strong>м 11 мм;<br />

длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 3,4 мм (ри<strong>с</strong>. 1117).<br />

Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

НАДСЕМЕЙСТВО CEPHIDEA<br />

В передних <strong>к</strong>рыльях <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й SC нет; три<br />

р<strong>а</strong>ди<strong>а</strong>льные ячеи (п-з), из них <strong>с</strong>редняя <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

д<strong>в</strong>ух других; п<strong>о</strong>перечные жил<strong>к</strong>и r-rs <strong>к</strong><strong>о</strong><strong>с</strong>ые,<br />

п<strong>а</strong>р<strong>а</strong>ллельные, р<strong>а</strong><strong>в</strong>ны п<strong>о</strong> длине, <strong>с</strong>ближенные;<br />

348<br />

http://jurassic.ru/<br />

<strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>а</strong>. Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong>рупные ф<strong>о</strong>рмы,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> ут<strong>о</strong>лщенными н<strong>а</strong> <strong>в</strong>ершине <strong>а</strong>нтенн<strong>а</strong>ми.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cephidac<br />

СЕМЕЙСТВО CEPHIDAE HALIDAY, 1840.<br />

ХЛЕБНЫЕ ПИЛИЛЬЩИКИ<br />

В передних <strong>к</strong>рыльях <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и rs-m; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS не з<strong>а</strong>гнут<strong>а</strong> <strong>в</strong>перед, пр<strong>о</strong><strong>с</strong>т<strong>а</strong>я;,<br />

ячея ri <strong>к</strong><strong>о</strong>р<strong>о</strong>че ячеи г 3; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея<br />

(dc) удлиненн<strong>а</strong>я, длиннее ячеи rs 2. Личин<strong>к</strong>и<br />

<strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нных грудных н<strong>о</strong>г;<br />

<strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льных н<strong>о</strong>г нет; члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн четыре—<br />

пять, е<strong>с</strong>ть гл<strong>а</strong>з<strong>к</strong>и. Жи<strong>в</strong>ут <strong>в</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>х<br />

р<strong>а</strong><strong>с</strong>тений (ри<strong>с</strong>. 1118). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне г<strong>о</strong>л<strong>а</strong>р<strong>к</strong>ти<strong>к</strong>и,<br />

<strong>о</strong>дин р<strong>о</strong>д н<strong>а</strong> М<strong>а</strong>д<strong>а</strong>г<strong>а</strong><strong>с</strong><strong>к</strong><strong>а</strong>ре; четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и ми<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО TENTHREDINIDEA<br />

В переднем <strong>к</strong>рыле SC <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и; р<strong>а</strong>ди<strong>а</strong>льных ячей д<strong>в</strong>е — три; ячея rj<br />

<strong>о</strong>чень неб<strong>о</strong>льш<strong>а</strong>я; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs <strong>о</strong>чень<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; RS <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т R ди<strong>с</strong>т<strong>а</strong>льнее б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и, т. е. б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> r-m, <strong>а</strong> не rs-m; Си <strong>о</strong>бр<strong>а</strong>зуют<br />

петлю <strong>в</strong> <strong>о</strong>беих п<strong>а</strong>р<strong>а</strong>х <strong>к</strong>рылье<strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген—ныне.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tenthredinidae, Cimbicldae, Diprionidae<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>в</strong>не СССР.<br />

СЕМЕЙСТВО TENTHREDINIDAE LEACH, 1817.<br />

НАСТОЯЩИЕ ПИЛИЛЬЩИКИ<br />

[nom. transl. Leach, 1819 (ex Tenthredinidea Leach, 1817)]<br />

В передних <strong>к</strong>рыльях <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й упл<strong>о</strong>тнен;<br />

<strong>к</strong>рупн<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>в</strong>д<strong>в</strong><strong>о</strong>е длиннее <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния Rb;<br />

передняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>в</strong>ыгнут<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди; ячея п зн<strong>а</strong>чительн<strong>о</strong><br />

меньше ячеи г 2; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея<br />

<strong>в</strong>д<strong>в</strong><strong>о</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>че ячеи г 2; Личин<strong>к</strong>и <strong>о</strong>чень п<strong>о</strong>х<strong>о</strong>жи<br />

н<strong>а</strong> гу<strong>с</strong>ениц, <strong>с</strong> тремя п<strong>а</strong>р<strong>а</strong>ми грудных и <strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льными<br />

н<strong>о</strong>г<strong>а</strong>ми н<strong>а</strong> 2—8 и 10 <strong>с</strong>егмент<strong>а</strong>х<br />

брюш<strong>к</strong><strong>а</strong>. Жи<strong>в</strong>ут н<strong>а</strong> ли<strong>с</strong>тьях, <strong>в</strong> пл<strong>о</strong>д<strong>а</strong>х, г<strong>а</strong>лл<strong>а</strong>х,<br />

нем<strong>а</strong>л<strong>о</strong> минер<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1119). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 4000 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong>еми п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), ми<strong>о</strong>цене<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО CIMBICIDAE LEACH, 1817.<br />

[nom. transl. Kirby, 1837 (ex Cimbicinae Leach, 1817)<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

реду<strong>к</strong>цией <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния RSi. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), ми<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы и<br />

С. Амери<strong>к</strong>и.

СЕМЕЙСТВО D1PR10NIDAE ROHWER, 1911<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

н<strong>а</strong>личием зубц<strong>о</strong><strong>в</strong> или <strong>о</strong>перенн<strong>о</strong><strong>с</strong>ти б<strong>о</strong>лее<br />

чем н<strong>а</strong> де<strong>в</strong>яти члени<strong>к</strong><strong>а</strong>х <strong>а</strong>нтенн. Личин<strong>к</strong>и жи<strong>в</strong>ут<br />

н<strong>а</strong> х<strong>в</strong><strong>о</strong>йных. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

ми<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

ПОДОТРЯД APOCRITA.<br />

СТЕБЕЛЬЧАТОБРЮХИЕ<br />

ПЕРЕПОНЧАТОКРЫЛЫЕ<br />

Крылья х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты или рудимент<strong>а</strong>рны;<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние ч<strong>а</strong><strong>с</strong>т<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>й<br />

SC ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не бы<strong>в</strong><strong>а</strong>ет; р<strong>а</strong>ди<strong>а</strong>льных ячей не<br />

б<strong>о</strong>лее д<strong>в</strong>ух; е<strong>с</strong>ли <strong>в</strong> .переднем <strong>к</strong>рыле при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

<strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я и ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячеи, т<strong>о</strong><br />

<strong>о</strong>ни не <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я (Parasitica) или <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>м пр<strong>о</strong>тяжении (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

Aculeata); <strong>а</strong>н<strong>а</strong>льных жил<strong>о</strong><strong>к</strong> нет; з<strong>а</strong>дние<br />

<strong>к</strong>рылья без <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти.<br />

Брюш<strong>к</strong><strong>о</strong> не <strong>с</strong>идячее; <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й члени<strong>к</strong> брюш<strong>к</strong><strong>а</strong><br />

уже других, ин<strong>о</strong>гд<strong>а</strong> удлинен; тип брюш<strong>к</strong><strong>а</strong><br />

<strong>с</strong>тебельч<strong>а</strong>тый или <strong>в</strong>и<strong>с</strong>ячий; е<strong>с</strong>ли у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

ф<strong>о</strong>рм брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>а</strong>жет<strong>с</strong>я <strong>с</strong>идячим, т<strong>о</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рылье<strong>в</strong> <strong>о</strong>чень упр<strong>о</strong>щен<strong>о</strong>. Яйце<strong>к</strong>л<strong>а</strong>д игл<strong>о</strong><strong>в</strong>идный,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>тянут <strong>в</strong> брюш<strong>к</strong><strong>о</strong> и пре<strong>в</strong>р<strong>а</strong>щен<br />

<strong>в</strong> ж<strong>а</strong>л<strong>о</strong>. Личин<strong>к</strong>и без н<strong>о</strong>г, чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зны, лишены<br />

гл<strong>а</strong>з, <strong>о</strong>бычн<strong>о</strong> м<strong>а</strong>л<strong>о</strong>п<strong>о</strong>д<strong>в</strong>ижны. В. юр<strong>а</strong> —<br />

ныне. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Ichneumonidea, Cynipidea,<br />

Chalcididea, Proctotrupidea, Bethylidea,<br />

'Cretavidea, Scoliidea, Formicidea, Pompilidea,<br />

Apidea, Vespidea; п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

<strong>в</strong>не СССР.<br />

НАДСЕМЕЙСТВО ICHNEUMONIDEA<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние передних <strong>к</strong>рылье<strong>в</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

п<strong>о</strong>лн<strong>о</strong>е; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления<br />

М; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong><br />

и <strong>с</strong>лит<strong>о</strong> <strong>в</strong> <strong>о</strong>дну линию <strong>с</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й; <strong>в</strong><strong>с</strong>егд<strong>а</strong> е<strong>с</strong>ть птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>. Р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ление<br />

М и ее <strong>о</strong>бе <strong>в</strong>ет<strong>в</strong>и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идны'; СиА<br />

е<strong>с</strong>ть. Антенны <strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые, <strong>с</strong> 10 и б<strong>о</strong>льшим<br />

чи<strong>с</strong>л<strong>о</strong>м члени<strong>к</strong><strong>о</strong><strong>в</strong>. Вертлуг д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ый. Личин<strong>к</strong>и<br />

<strong>о</strong>бычн<strong>о</strong> п<strong>а</strong>р<strong>а</strong>зиты, гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м н<strong>а</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Одн<strong>о</strong> из н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бширных н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>—<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

16 000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. юр<strong>а</strong> — ныне.<br />

В<strong>о</strong><strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Pelecinopteridae,<br />

Ephialtitidae, Braconidae, Megalyridae, Evaniidae,<br />

Aulacidae, Stephanidae, Trigonalidae,<br />

Ichneumonidae; из них т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пер<strong>в</strong>ые д<strong>в</strong><strong>а</strong> <strong>в</strong>ымершие.<br />

СЕМЕЙСТВО PELECINOPTERIDAE BRUES, 1933<br />

Длин<strong>а</strong> передних <strong>к</strong>рылье<strong>в</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 4 мм, <strong>в</strong> 2<br />

р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>о</strong>чень длинн<strong>о</strong>г<strong>о</strong> брюш<strong>к</strong><strong>а</strong>; <strong>в</strong> пе­<br />

349<br />

http://jurassic.ru/<br />

редних <strong>к</strong>рыльях птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> нея<strong>с</strong>н<strong>а</strong>я, п<strong>о</strong>х<strong>о</strong>ж<strong>а</strong>я<br />

н<strong>а</strong> ут<strong>о</strong>лщение жил<strong>к</strong>и; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н;<br />

RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я ди<strong>с</strong>т<strong>а</strong>льнее птер<strong>о</strong><strong>с</strong>тигмы;<br />

<strong>в</strong>ершин<strong>а</strong> передней <strong>в</strong>ет<strong>в</strong>и из<strong>о</strong>гнут<strong>а</strong> <strong>к</strong>з<strong>а</strong>ди; ячея<br />

dc <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и не<br />

<strong>в</strong>идн<strong>о</strong>; меди<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>. Крупн<strong>а</strong>я г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong> б<strong>о</strong>льшими гл<strong>а</strong>з<strong>а</strong>ми и тремя гл<strong>а</strong>з<strong>к</strong><strong>а</strong>ми;<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 13; грудь длинн<strong>а</strong>я. Брюш<strong>к</strong><strong>о</strong><br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и из ше<strong>с</strong>ти п<strong>о</strong>чти р<strong>а</strong><strong>в</strong>ных п<strong>о</strong> длине <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

яйце<strong>к</strong>л<strong>а</strong>д не <strong>в</strong>ытянут (ри<strong>с</strong>. 1120). П<strong>а</strong>ле<strong>о</strong>ген.<br />

Один р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО BRACONIDAE BURMEISTER, 1829.<br />

НАЕЗДНИКИ-БРАКОНИДЫ<br />

'[nom. transl. Kirby, 1837<br />

(ex Braconoides Burmeister, 1829)]<br />

Длин<strong>а</strong> <strong>к</strong>рыл<strong>а</strong> немн<strong>о</strong>г<strong>о</strong> пре<strong>в</strong>ыш<strong>а</strong>ет длину<br />

брюш<strong>к</strong><strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>в</strong> передних <strong>к</strong>рыльях я<strong>с</strong>н<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS <strong>о</strong>бычн<strong>о</strong> не редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н,<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не птер<strong>о</strong><strong>с</strong>тигмы;<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и RS <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

<strong>в</strong>ершин<strong>а</strong> передней <strong>в</strong>ет<strong>в</strong>и RS <strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнут<strong>а</strong><br />

<strong>в</strong>перед или прям<strong>а</strong>я; ячея dc з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я,<br />

неб<strong>о</strong>льш<strong>а</strong>я, меньше ячей sr, ri и г 2; ячеи mi<br />

нет. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я или <strong>к</strong>убиче<strong>с</strong><strong>к</strong><strong>а</strong>я, три<br />

гл<strong>а</strong>з<strong>к</strong><strong>а</strong>, <strong>а</strong>нтенны длинные, ните<strong>в</strong>идные. Брюш<strong>к</strong><strong>о</strong><br />

причленяет<strong>с</strong>я <strong>к</strong> нижней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong>межут<strong>о</strong>чн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong>, <strong>с</strong><strong>о</strong>членение между <strong>в</strong>т<strong>о</strong>рым и третьим<br />

<strong>с</strong>егмент<strong>а</strong>ми неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>е (ри<strong>с</strong>. 1121). Личин<strong>к</strong>и<br />

п<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong> <strong>в</strong><strong>с</strong>ех н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, ч<strong>а</strong>ще н<strong>а</strong><br />

же<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых. П<strong>а</strong>ле<strong>о</strong>ген — ныне. 16 п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Braconinae, Spathiinae,<br />

Doryctinae, Rhogadinae, Cheloninae, Triaspinae,<br />

Neoneurinae, Microgasterinae, Blacinae,<br />

Euphorinae, Meteorinae, Aphidiinae, Alysiinae<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

3. Е<strong>в</strong>р<strong>о</strong>пы, С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО MEGALYRIDAE<br />

SCHLETTERER, 1889<br />

Передние <strong>к</strong>рылья немн<strong>о</strong>г<strong>о</strong> длиннее брюш<strong>к</strong><strong>а</strong>;<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, нея<strong>с</strong>н<strong>а</strong>я;<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет неб<strong>о</strong>льшую, п<strong>о</strong>чти <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тную<br />

<strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льную ячею (sr), <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я немн<strong>о</strong>г<strong>о</strong><br />

меньше нижележ<strong>а</strong>щей <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й <strong>к</strong>убит<strong>а</strong>льн<strong>о</strong>й<br />

ячеи; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние передней <strong>в</strong>ет<strong>в</strong>и RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>,<br />

<strong>о</strong>н<strong>а</strong> п<strong>о</strong>х<strong>о</strong>ж<strong>а</strong> н<strong>а</strong> и<strong>с</strong><strong>к</strong>ри<strong>в</strong>ленную длинную<br />

r-rs; з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, не д<strong>о</strong>х<strong>о</strong>дит<br />

д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; dc и mi нет. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>е;<br />

яйце<strong>к</strong>л<strong>а</strong>д ч<strong>а</strong><strong>с</strong>т<strong>о</strong> длиннее тел<strong>а</strong> (ри<strong>с</strong>. 1122).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне (19 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии и <strong>о</strong>дин <strong>в</strong>ид <strong>в</strong><br />

Ю. Амери<strong>к</strong>е); <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).

СЕМЕЙСТВО EVANIIDAE LEACH, 1812<br />

[nom. transl. Westwood, 1840 (ex Evaniades Leach, 1812)]<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>в</strong> передних <strong>к</strong>рыльях я<strong>с</strong>н<strong>а</strong>я,<br />

пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>с</strong>я;<br />

р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы;<br />

передняя <strong>в</strong>ет<strong>в</strong>ь RS <strong>в</strong> <strong>в</strong>ершине <strong>к</strong>рут<strong>о</strong><br />

из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong>перед, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> переднем<br />

<strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>; ячея dc з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>, <strong>к</strong>рупн<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

п<strong>о</strong> длине ячее sr, н<strong>о</strong> шире ее; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> длинн<strong>а</strong>я; ячеи mi нет. Члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>а</strong>нтенн 13—14. Брюш<strong>к</strong><strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, <strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>е,<br />

причленяет<strong>с</strong>я <strong>к</strong> <strong>в</strong>ерхней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пр<strong>о</strong>межут<strong>о</strong>чн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong>, мн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыше <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

з<strong>а</strong>дних н<strong>о</strong>г; яйце<strong>к</strong>л<strong>а</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, п<strong>о</strong>чти не <strong>в</strong>ыд<strong>а</strong>ет<strong>с</strong>я<br />

з<strong>а</strong> <strong>в</strong>ершину брюш<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 1123). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong>т <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>в</strong> р<strong>а</strong>зных ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>их шир<strong>о</strong>т<strong>а</strong>х,<br />

п<strong>а</strong>р<strong>а</strong>зиты' <strong>о</strong><strong>о</strong>те<strong>к</strong> т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>; д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО AULACIDAE SCHUCKARD, 1841<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>а</strong>, пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л RS длинный, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и RS нет, <strong>в</strong>ершин<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong><br />

RS <strong>с</strong>лит<strong>а</strong> <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью М; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS прям<strong>а</strong>я, или <strong>в</strong>ершин<strong>а</strong> ее немн<strong>о</strong>г<strong>о</strong><br />

из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong>перед; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs длинн<strong>а</strong>я,<br />

з<strong>а</strong>мы<strong>к</strong><strong>а</strong>ет <strong>к</strong>рупную ячею п; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong>бр<strong>а</strong>зует длинную и<br />

уз<strong>к</strong>ую ячею sr; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я (dc) и меди­<br />

<strong>а</strong>льн<strong>а</strong>я ячеи з<strong>а</strong><strong>к</strong>рытые. Антенны 1<br />

13—14-члени-<br />

<strong>к</strong><strong>о</strong><strong>в</strong>ые. Брюш<strong>к</strong><strong>о</strong> причленяет<strong>с</strong>я <strong>к</strong> <strong>в</strong>ерхней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

пр<strong>о</strong>межут<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>, мн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыше<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния з<strong>а</strong>дних н<strong>о</strong>г, длиннее груди; яйце<strong>к</strong>л<strong>а</strong>д<br />

я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыд<strong>а</strong>ет<strong>с</strong>я з<strong>а</strong> <strong>в</strong>ершину брюш<strong>к</strong><strong>а</strong>, неред<strong>к</strong><strong>о</strong><br />

длиннее <strong>в</strong><strong>с</strong>ег<strong>о</strong> тел<strong>а</strong>. П<strong>а</strong>р<strong>а</strong>зиты личин<strong>о</strong><strong>к</strong> жу<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>бит<strong>а</strong>ющих <strong>в</strong> дре<strong>в</strong>е<strong>с</strong>ине; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> п<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> личин<strong>к</strong><strong>а</strong>х Xiphidriidae. В. юр<strong>а</strong> — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 100 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>; де<strong>в</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

Mesaulacinus Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong><br />

— М. oviformis Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Передние<br />

<strong>к</strong>рылья шир<strong>о</strong><strong>к</strong>ие, з<strong>а</strong><strong>к</strong>ругленные; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея р<strong>о</strong>мб<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, <strong>к</strong>рупнее треуг<strong>о</strong>льн<strong>о</strong>й<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й ячеи (dc); <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь руди­<br />

менты <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и между <strong>в</strong>ет<strong>в</strong>ями<br />

RS. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 2,1 мм<br />

(ри<strong>с</strong>. 1124). Один <strong>в</strong>ид. В. юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Paraulacus Ping, 1928 из мел<strong>а</strong> Кит<strong>а</strong>я;<br />

<strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> третичных <strong>о</strong>тл<strong>о</strong>жениях Е<strong>в</strong>р<strong>о</strong>пы<br />

(ри<strong>с</strong> 1125), Азии, Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО STEPHANIDAE LEACH, 1815<br />

[nom. transl. Haliday, 1840 (ex Stephanidia Leach, 1815)}<br />

Р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны п<strong>о</strong> <strong>в</strong>еличине: <strong>о</strong>т 4 д<strong>о</strong> 36 мм<br />

длин<strong>о</strong>й. Крылья <strong>к</strong><strong>о</strong>р<strong>о</strong>че тел<strong>а</strong>, птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>, пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й ширины; RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

при <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ении <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью<br />

М, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины длины птер<strong>о</strong><strong>с</strong>тигмы;<br />

<strong>в</strong>ет<strong>в</strong>и RS не из<strong>о</strong>гнуты; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> птер<strong>о</strong><strong>с</strong>тигмы;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я, б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет р<strong>о</strong>мб<strong>о</strong><strong>в</strong>идную<br />

<strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льную ячею; меди<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я. Длинные 30-члени<strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>а</strong>нтенны.<br />

Яйце<strong>к</strong>л<strong>а</strong>д длиннее тел<strong>а</strong>. П<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> <strong>с</strong><strong>в</strong>ерлящих дре<strong>в</strong>е<strong>с</strong>ину н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (ри<strong>с</strong>.<br />

1126). Б<strong>о</strong>лее 100 <strong>в</strong>ид<strong>о</strong><strong>в</strong> де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> ж<strong>а</strong>р<strong>к</strong><strong>о</strong>м<br />

<strong>к</strong>лим<strong>а</strong>те, д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и ми<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО TRIGONALIDAE CRESSON, 1867<br />

Средней <strong>в</strong>еличины н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, <strong>к</strong>рылья<br />

длиннее брюш<strong>к</strong><strong>а</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, не шире<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS из<strong>о</strong>гнут п<strong>о</strong>д прямым<br />

угл<strong>о</strong>м; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я;<br />

RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины<br />

птер<strong>о</strong><strong>с</strong>тигмы, <strong>в</strong>ет<strong>в</strong>и ег<strong>о</strong> не из<strong>о</strong>гнуты; п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> r-rs длинн<strong>а</strong>я, <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет <strong>к</strong>рупную<br />

р<strong>а</strong>ди<strong>а</strong>льную ячею (ri), между <strong>в</strong>ет<strong>в</strong>ями<br />

RS д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея длинн<strong>а</strong>я и уз<strong>к</strong><strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет тр<strong>а</strong>пецие<strong>в</strong>идную <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льную<br />

ячею (sr); меди<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я.<br />

Брюш<strong>к</strong><strong>о</strong> причленяет<strong>с</strong>я <strong>к</strong> нижней ч<strong>а</strong><strong>с</strong>ти<br />

пр<strong>о</strong>межут<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>, н<strong>а</strong>д <strong>с</strong><strong>а</strong>мым <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

з<strong>а</strong>дних н<strong>о</strong>г. Яйц<strong>а</strong> <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют н<strong>а</strong> ли<strong>с</strong>тьях,<br />

личин<strong>к</strong>и п<strong>о</strong>ед<strong>а</strong>ют л<strong>о</strong>жн<strong>о</strong>гу<strong>с</strong>ениц пилильщи<strong>к</strong><strong>о</strong><strong>в</strong><br />

или п<strong>а</strong>р<strong>а</strong>зит<strong>о</strong><strong>в</strong> гу<strong>с</strong>ениц б<strong>а</strong>б<strong>о</strong>че<strong>к</strong> (ри<strong>с</strong>.<br />

1127). Не<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> ше<strong>с</strong>ти р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> не<strong>о</strong>гене<br />

Ю. Азии (янт<strong>а</strong>рь Бирмы) и Герм<strong>а</strong>нии.<br />

Ри<strong>с</strong>. 1120—1129. П<strong>о</strong>д<strong>о</strong>тряд Apocrita, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Ichneumonidea, Cynipidea<br />

1120. Pelecmopteron tubuliforme Brues; <strong>о</strong>бщий <strong>в</strong>ид, x4,5, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Ь<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Brues, 1933). 1121. Microti/-<br />

B r ?£ т " e s<br />

<strong>о</strong> б щ и й ВИ<br />

х 1 8<br />

4 Д- -°- п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)( Brues, 1933). 1122. Prodinapsis minor Brues-<br />

°;'шч?Л ИД<br />

http://jurassic.ru/<br />

U 24 Б

СЕМЕЙСТВО ICHNEUMON I DAE LATREILLE, 1802.<br />

НАСТОЯЩИЕ НАЕЗДНИКИ<br />

[nom. transl. Leach, 1817<br />

(ex lchneumonides Latreille 1802)]<br />

Птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>в</strong> передних <strong>к</strong>рыльях я<strong>с</strong>н<strong>а</strong>я,<br />

треуг<strong>о</strong>льн<strong>а</strong>я; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля нет; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> н<strong>а</strong>з<strong>а</strong>д, <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й, д<strong>а</strong>лее редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>;<br />

<strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея <strong>с</strong>лит<strong>а</strong> <strong>с</strong> пер<strong>в</strong><strong>о</strong>й<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й, з<strong>а</strong><strong>к</strong>рыт<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong><strong>о</strong>й; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs длинн<strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я,<br />

перех<strong>о</strong>дит <strong>в</strong> переднюю <strong>в</strong>ет<strong>в</strong>ь RS; меди<strong>а</strong>льн<strong>а</strong>я<br />

ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я, <strong>к</strong>рупн<strong>а</strong>я. П<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> <strong>в</strong><strong>с</strong>ех н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, н<strong>о</strong> преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

н<strong>а</strong> гу<strong>с</strong>ениц<strong>а</strong>х и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, жи<strong>в</strong>ущих <strong>в</strong> дре<strong>в</strong>е<strong>с</strong>ине<br />

(ри<strong>с</strong>. 1128). П<strong>а</strong>ле<strong>о</strong>ген — ныне. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Ichneumoninae, Cryptinae, Pimplinae,<br />

Ophioninae, Tryphoninae, из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО CYN1PIDEA<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, 1—4 мм длин<strong>о</strong>й; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле е<strong>с</strong>ть; птер<strong>о</strong><strong>с</strong>тигмы нет; <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong>т Ichneuraonidea <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти меди<strong>а</strong>льн<strong>о</strong>й жил<strong>к</strong>и <strong>с</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>лением и<br />

п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й Си. П<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых, д<strong>в</strong>у<strong>к</strong>рылых, тлях,<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые из них г<strong>а</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>тели (ри<strong>с</strong>.<br />

1129). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Ibaliidae,<br />

Figitidae, Eucoilidae, Charipidae, Cynipidae;<br />

п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО CYNIPIDAE LINNAEUS, 1758.<br />

ОРЕХОТВОРКИ<br />

[nom. transl. Haliday, 1840 (ex Cynipes Linnaeus, 1758)]<br />

Пер<strong>в</strong><strong>а</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея (ri) з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>,<br />

длинн<strong>а</strong>я, длин<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет ее ширину <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong>.<br />

Антенны пр<strong>о</strong><strong>с</strong>тые, не б<strong>о</strong>лее 16 члени<strong>к</strong><strong>о</strong><strong>в</strong>. Р<strong>а</strong>нние<br />

<strong>с</strong>т<strong>а</strong>дии жи<strong>в</strong>ут <strong>в</strong> р<strong>а</strong><strong>с</strong>тительных т<strong>к</strong><strong>а</strong>нях и <strong>о</strong>бр<strong>а</strong>зуют<br />

г<strong>а</strong>ллы. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Де<strong>с</strong>ять п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>дн<strong>о</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО CHALCIDIDEA.<br />

ХАЛЬЦИДЫ<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые р<strong>а</strong>змер<strong>о</strong>м <strong>о</strong>т менее 1 мм п<strong>о</strong>чти<br />

д<strong>о</strong> 20 мм; <strong>с</strong>редние р<strong>а</strong>змеры 2—3 мм, ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong> б<strong>о</strong>лее или менее у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченными <strong>к</strong>рыльями.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> д<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й жил<strong>к</strong>и<br />

SC -f- R. Антенны <strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые, <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>льце<strong>в</strong>идными<br />

члени<strong>к</strong><strong>а</strong>ми между <strong>в</strong>т<strong>о</strong>рым члени­<br />

352<br />

http://jurassic.ru/<br />

<strong>к</strong><strong>о</strong>м и жгути<strong>к</strong><strong>о</strong>м. Л<strong>а</strong>п<strong>к</strong>и трех-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

П<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, изред<strong>к</strong><strong>а</strong> н<strong>а</strong> других<br />

члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих (<strong>к</strong>лещи, п<strong>а</strong>у<strong>к</strong>и). Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> и <strong>о</strong>тдельные р<strong>о</strong>ды фит<strong>о</strong>ф<strong>а</strong>ги. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 27 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Chalcididae, Torymidae, Mymaridae, Eurytomidae,<br />

Agaonidae, Perilampidae, Pteromalidae,<br />

Eulophidae, Encyrtidae; п<strong>о</strong><strong>с</strong>ледние ше<strong>с</strong>ть <strong>в</strong>не<br />

СССР.<br />

Ри<strong>с</strong>. 1130—1132. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Chalcididea<br />

ИЗО. Brachymeria'intermedia Nees; <strong>о</strong>бщий <strong>в</strong>ид, х12,0, <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем<br />

<strong>в</strong>иГ^пТ?' 1952)<br />

'AI 31<br />

" Me<br />

easiigmus iuniperi Nikolska <strong>а</strong>;<strong>о</strong>бщий<br />

In^^uL?°. Pe<br />

«-(Ни<strong>к</strong><strong>о</strong>ль<strong>с</strong><strong>к</strong><strong>а</strong>я, 1952). 1132. Gonatocerus cicadellae<br />

Nikolskaja; <strong>о</strong>бщий <strong>в</strong>ид, x20,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Ни<strong>к</strong><strong>о</strong>ль<strong>с</strong><strong>к</strong><strong>а</strong>я, 1952)

СЕМЕЙСТВО CHALCIDIDAE WALKER, 1835<br />

Д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong>рупные ф<strong>о</strong>рмы, ред<strong>к</strong><strong>о</strong> менее<br />

5 мм длин<strong>о</strong>й. З<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и длинные, цилиндриче<strong>с</strong><strong>к</strong>ие;<br />

з<strong>а</strong>дние бедр<strong>а</strong> ут<strong>о</strong>лщенные, <strong>с</strong> зубц<strong>а</strong>ми<br />

п<strong>о</strong> н<strong>а</strong>ружн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю; з<strong>а</strong>дние г<strong>о</strong>лени из<strong>о</strong>гнуты.<br />

Яйце<strong>к</strong>л<strong>а</strong>д е<strong>с</strong>ли <strong>в</strong>ыд<strong>а</strong>ет<strong>с</strong>я, т<strong>о</strong> при<strong>к</strong>рыт<br />

п<strong>о</strong><strong>с</strong>ледним <strong>в</strong>ытянутым тергит<strong>о</strong>м (ри<strong>с</strong>. ИЗО).<br />

Не<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне ше<strong>с</strong>ть<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> и б<strong>о</strong>лее 100 р<strong>о</strong>д<strong>о</strong><strong>в</strong>: из них четыре<br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> ми<strong>о</strong>цене 3. Е<strong>в</strong>р<strong>о</strong>пы 1<br />

и чет<strong>в</strong>ертичных<br />

<strong>о</strong>тл<strong>о</strong>жениях В. Афри<strong>к</strong>и.<br />

СЕМЕЙСТВО TORYMIDAE WALKER, 1833<br />

(Callimomidae Viereck, 1916)<br />

Р<strong>а</strong>змеры тел<strong>а</strong> <strong>о</strong>т 1,5 д<strong>о</strong> 7 мм. От предыдущег<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я меньшими р<strong>а</strong>змер<strong>а</strong>ми.<br />

З<strong>а</strong>дние т<strong>а</strong>зи<strong>к</strong>и трехгр<strong>а</strong>нные, <strong>о</strong>бычн<strong>о</strong> з<strong>а</strong>дние<br />

бедр<strong>а</strong> неут<strong>о</strong>лщенные. Яйце<strong>к</strong>л<strong>а</strong>д б<strong>о</strong>лее<br />

или менее длинный (ри<strong>с</strong>. 1131). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 60 р<strong>о</strong>д<strong>о</strong><strong>в</strong> и<br />

б<strong>о</strong>лее 500 <strong>в</strong>ид<strong>о</strong><strong>в</strong>; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

1<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>гене С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО MYMARIDAE HALIDAY, 1833<br />

[norn. transl. Haliday, 1840 (ex Mymares Haliday, 1833)]<br />

Мел<strong>к</strong>ие ф<strong>о</strong>рмы, <strong>о</strong>бычн<strong>о</strong> менее 1 мм <strong>в</strong> длину,<br />

б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью яйцееды. Крылья уз<strong>к</strong>ие, <strong>с</strong><br />

длинн<strong>о</strong>й б<strong>а</strong>хр<strong>о</strong>м<strong>о</strong>й п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю; жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>р<strong>а</strong>йне<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

R; з<strong>а</strong>дние <strong>к</strong>рылья ин<strong>о</strong>гд<strong>а</strong> щетин<strong>к</strong><strong>о</strong><strong>в</strong>идные.<br />

Члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн 8—13, у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong><br />

бул<strong>а</strong><strong>в</strong><strong>о</strong>й. Л<strong>а</strong>п<strong>к</strong>и четырех-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

(ри<strong>с</strong>. 1132). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong> и б<strong>о</strong>лее 200 <strong>в</strong>ид<strong>о</strong><strong>в</strong>; де<strong>в</strong>ять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых (из них д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших)<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

С. Амери<strong>к</strong>и, В. Афри<strong>к</strong>е (<strong>к</strong><strong>о</strong>п<strong>а</strong>л).<br />

НАДСЕМЕЙСТВО PROCTOTRUPIDEA<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, неред<strong>к</strong><strong>о</strong> без <strong>к</strong>рылье<strong>в</strong>.<br />

Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние ин<strong>о</strong>гд<strong>а</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> д<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й<br />

жил<strong>к</strong>и, т<strong>о</strong>гд<strong>а</strong> SC + R <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и не <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я<br />

<strong>с</strong> С; е<strong>с</strong>ли жил<strong>о</strong><strong>к</strong> б<strong>о</strong>льше, т<strong>о</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

(dc) ячея <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я, <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея (sr) <strong>в</strong><strong>с</strong>егд<strong>а</strong> м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я, <strong>о</strong>бычн<strong>о</strong> треуг<strong>о</strong>льн<strong>о</strong>й<br />

ф<strong>о</strong>рмы; ячеи не <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я <strong>о</strong>дн<strong>а</strong> <strong>с</strong> друг<strong>о</strong>й;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупн<strong>а</strong>я. Антенны<br />

пр<strong>о</strong><strong>с</strong>тые, е<strong>с</strong>ли <strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые, т<strong>о</strong> без <strong>к</strong><strong>о</strong>льце<strong>в</strong>идных<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong>. Яйце<strong>к</strong>л<strong>а</strong>д <strong>о</strong>тх<strong>о</strong>дит <strong>о</strong>т<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong>й <strong>в</strong>ершины брюш<strong>к</strong><strong>а</strong>. П<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong><br />

члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих или н<strong>а</strong> их яйц<strong>а</strong>х. В. юр<strong>а</strong> — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 12 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Megaspilidae,<br />

Heloridae, Proctotrupidae, Scelionidae, Serphitidae.<br />

23 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

СЕМЕЙСТВО MEGASPILIDAE ASHMEAD, 1893<br />

Inom. transl. Ashmead, 1903<br />

(ex Megaspilini Ashmead, 1893)]<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, длин<strong>а</strong> 0,6—3,3 мм, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

бе<strong>с</strong><strong>к</strong>рылые. В переднем <strong>к</strong>рыле С, SC, R<br />

<strong>с</strong>литы <strong>в</strong> <strong>о</strong>дну жил<strong>к</strong>у, <strong>о</strong><strong>с</strong>т<strong>а</strong>льные жил<strong>к</strong>и <strong>в</strong> <strong>в</strong>иде<br />

рудимент<strong>о</strong><strong>в</strong>; неб<strong>о</strong>льш<strong>о</strong>е из<strong>о</strong>гнут<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

RS и <strong>о</strong>т нег<strong>о</strong> б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong>, пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ющ<strong>а</strong>я<br />

<strong>в</strong><strong>с</strong>ё <strong>к</strong>рыл<strong>о</strong> д<strong>о</strong> з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> r-rs перех<strong>о</strong>дит <strong>в</strong> переднюю<br />

353<br />

http://jurassic.ru/<br />

Ри<strong>с</strong>. 1133—1137. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Proctotrupidea<br />

1133. Conostigmus succinalis Brues; <strong>о</strong>бщий <strong>в</strong>ид, х24,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Brues, 1940). 1134. Mesohelorus<br />

muchini Martynov; <strong>о</strong>бщий <strong>в</strong>ид, х9,0, <strong>в</strong>. юр<strong>а</strong>, Ю. К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1925). 1135.. Cryptoserphus pinorus Brues; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х22.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ийянт<strong>а</strong>рь) (Brues, 1940). 1136.<br />

Ceratoteleia proleptica Brues; <strong>о</strong>бщий <strong>в</strong>ид, Х15.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Brues, 1940). 1137. Serphites paradoxus<br />

Brues; переднее <strong>к</strong>рыл<strong>о</strong>, Х39.0, С Амери<strong>к</strong><strong>а</strong>, <strong>в</strong>. мел (<strong>к</strong><strong>а</strong>н<strong>а</strong>д<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Brues, 1937)

<strong>в</strong>ет<strong>в</strong>ь RS, <strong>о</strong>бр<strong>а</strong>зуя <strong>о</strong>дну из<strong>о</strong>гнутую жил<strong>к</strong>у;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> е<strong>с</strong>ть. П<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong> тлях, щит<strong>о</strong><strong>в</strong><strong>к</strong><strong>а</strong>х,<br />

д<strong>в</strong>у<strong>к</strong>рылых (ри<strong>с</strong>. 1133). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне мн<strong>о</strong>г<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных р<strong>о</strong>д<strong>о</strong><strong>в</strong>; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий и <strong>к</strong><strong>а</strong>н<strong>а</strong>д<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО HELORIDAE FORSTER, 1856<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые длин<strong>о</strong>й 3—6,5 мм. Передние<br />

<strong>к</strong>рылья <strong>с</strong><strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> б<strong>о</strong>г<strong>а</strong>тым жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS между R и б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong><strong>о</strong>й ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея треуг<strong>о</strong>льн<strong>а</strong>я; пер<strong>в</strong><strong>а</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея шир<strong>о</strong><strong>к</strong><strong>а</strong>я; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> е<strong>с</strong>ть. П<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых. В. юр<strong>а</strong> — ныне.<br />

Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне Се<strong>в</strong>ерн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>луш<strong>а</strong>рия и <strong>о</strong>дин <strong>в</strong>ымерший р<strong>о</strong>д.<br />

Mesohelorus Martynov, 1925. Тип р<strong>о</strong>д<strong>а</strong> —<br />

М.' muchini Martynov, 1925; <strong>в</strong>. юр<strong>а</strong>, Чим<strong>к</strong>ент<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (м<strong>а</strong>льм, К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у). Переднее <strong>к</strong>рыл<strong>о</strong><br />

<strong>с</strong> удлиненн<strong>о</strong>й птер<strong>о</strong><strong>с</strong>тигм<strong>о</strong>й и уз<strong>к</strong>им <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным<br />

п<strong>о</strong>лем; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>с</strong>я п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью;<br />

<strong>в</strong>ет<strong>в</strong>и RS прямые (не из<strong>о</strong>гнуты); рудименты<br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и, з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющей ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льную<br />

ячею. Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 5 мм<br />

(ри<strong>с</strong>. 1134). Один <strong>в</strong>ид. В- юр<strong>а</strong> К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

СЕМЕЙСТВО PROCTOTRUPIDAE CRESSON, 1887<br />

(Serphidae Kieffer, 1900')<br />

Длин<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых 2—7 мм. Передние<br />

<strong>к</strong>рылья <strong>с</strong> д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным<br />

жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я SC + R<br />

и <strong>к</strong>рупн<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; RS <strong>о</strong>бычн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>,<br />

е<strong>с</strong>ли <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я (у третичных<br />

ф<strong>о</strong>рм), т<strong>о</strong> передняя ег<strong>о</strong> <strong>в</strong>ет<strong>в</strong>ь <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong>л<strong>о</strong>няет<strong>с</strong>я<br />

<strong>в</strong>перед; RS приближен<strong>а</strong> <strong>к</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

птер<strong>о</strong><strong>с</strong>тигмы и п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs<br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я; <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

М д<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея<br />

м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я, пер<strong>в</strong><strong>а</strong>я реди<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>чень <strong>к</strong>рупн<strong>а</strong>я.<br />

П<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong> д<strong>в</strong>у<strong>к</strong>рылых и жу<strong>к</strong><strong>а</strong>х (ри<strong>с</strong>.<br />

1135). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные р<strong>о</strong>ды <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне. Три р<strong>о</strong>д<strong>а</strong> из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и не<strong>о</strong>гене С. Амери<strong>к</strong>и;<br />

<strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уны <strong>в</strong> чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жениях В. Афри<strong>к</strong>и.<br />

СЕМЕЙСТВО SCELIONIDAE HALIDAY, 1840<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые длин<strong>о</strong>й 0,45—5 мм. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

бе<strong>с</strong><strong>к</strong>рылые; н<strong>а</strong> передних <strong>к</strong>рыльях птер<strong>о</strong><strong>с</strong>тигмы<br />

нет; SC + R е<strong>с</strong>ть, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее<br />

<strong>о</strong>тх<strong>о</strong>дит <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

r-rs; п<strong>о</strong><strong>с</strong>ледняя ин<strong>о</strong>гд<strong>а</strong> перех<strong>о</strong>дит <strong>в</strong> рудимент<strong>а</strong>рную,<br />

не д<strong>о</strong>х<strong>о</strong>дящую д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> переднюю<br />

<strong>в</strong>ет<strong>в</strong>ь RS; <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние реду­<br />

354<br />

http://jurassic.ru/<br />

цир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> д<strong>о</strong> <strong>с</strong>лед<strong>о</strong><strong>в</strong>. П<strong>а</strong>р<strong>а</strong>зитируют гл<strong>а</strong><strong>в</strong>ным<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м н<strong>а</strong> яйц<strong>а</strong>х чешуе<strong>к</strong>рылых, переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых<br />

и прям<strong>о</strong><strong>к</strong>рылых (ри<strong>с</strong>. 1136)- П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные р<strong>о</strong>ды; 26 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

не<strong>о</strong>гене 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и, и <strong>в</strong> В. Афри<strong>к</strong>е<br />

(<strong>к</strong><strong>о</strong>п<strong>а</strong>л).<br />

СЕМЕЙСТВО SERPHITIDAE BRUES, 1937<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т Proctotrupidae н<strong>а</strong>личием <strong>в</strong><br />

передних <strong>к</strong>рыльях не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>й птер<strong>о</strong><strong>с</strong>тигмы,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>й н<strong>а</strong> <strong>с</strong>ередине длины<br />

<strong>к</strong>рыл<strong>а</strong>, з<strong>а</strong>ним<strong>а</strong>ющей п<strong>о</strong>чти п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину ширины<br />

<strong>к</strong>рыл<strong>а</strong>; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием RS, <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>.<br />

Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 1,2 мм<br />

(ри<strong>с</strong>. 1137). П<strong>а</strong>ле<strong>о</strong>ген. Один р<strong>о</strong>д Serphites,<br />

Brues, 1937 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и (<strong>к</strong><strong>а</strong>н<strong>а</strong>д<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО BETHYLIDEA<br />

В передних <strong>к</strong>рыльях б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> неред<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong>единяет меди<strong>а</strong>льную жил<strong>к</strong>у<br />

<strong>с</strong> р<strong>а</strong>диу<strong>с</strong><strong>о</strong>м, <strong>а</strong> не <strong>с</strong><strong>о</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong>м RS; <strong>с</strong>т<strong>в</strong><strong>о</strong>л RS<br />

длинный, е<strong>с</strong>ли р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ление <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я, т<strong>о</strong><br />

<strong>о</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления М;<br />

ч<strong>а</strong>ще <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RSi утр<strong>а</strong>чен<strong>о</strong> и передняя <strong>в</strong>ет<strong>в</strong>ь<br />

RS <strong>о</strong>тх<strong>о</strong>дит <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>о</strong>д<strong>о</strong>лжение п<strong>о</strong>перечн<strong>о</strong>й<br />

жил<strong>к</strong>и r-rs и из<strong>о</strong>гнут<strong>а</strong> <strong>в</strong>перед; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

уз<strong>к</strong><strong>а</strong>я, п<strong>о</strong>х<strong>о</strong>ж<strong>а</strong> н<strong>а</strong> ут<strong>о</strong>лщенную жил<strong>к</strong>у;<br />

меди<strong>а</strong>льн<strong>а</strong>я ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; Си е<strong>с</strong>ть. Антенны<br />

из 10—28 члени<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>с</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым чи<strong>с</strong>л<strong>о</strong>м у<br />

<strong>о</strong>б<strong>о</strong>их п<strong>о</strong>л<strong>о</strong><strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген—ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии Embolemidae, Bethylidae, Chrysididae,<br />

Dryinidae.<br />

СЕМЕЙСТВО EMBOLEMIDAE FORSTER, 1856<br />

[nom. transl. Handlirsch, 1925<br />

(ex Embolemoidae Forster, 1856)]<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, <strong>с</strong><strong>а</strong>м<strong>к</strong>и бе<strong>с</strong><strong>к</strong>рылые. В передних<br />

<strong>к</strong>рыльях птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>о</strong>чень м<strong>а</strong>л<strong>а</strong>, п<strong>о</strong>чти<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние передней <strong>в</strong>ет<strong>в</strong>и RS редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> и<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея <strong>с</strong>лит<strong>а</strong> <strong>с</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>к</strong>ругл<strong>а</strong>я, гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я, <strong>с</strong>ужен<strong>а</strong> <strong>к</strong>низу, <strong>с</strong><br />

мет<strong>а</strong>лличе<strong>с</strong><strong>к</strong>им бле<strong>с</strong><strong>к</strong><strong>о</strong>м; члени<strong>к</strong><strong>о</strong><strong>в</strong> <strong>а</strong>нтенн де<strong>с</strong>ять,<br />

т<strong>о</strong>н<strong>к</strong>ие, при<strong>к</strong>реплены <strong>в</strong>ыше н<strong>а</strong>лични<strong>к</strong><strong>а</strong>.<br />

Передние л<strong>а</strong>п<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые, без <strong>к</strong>леще<strong>о</strong>бр<strong>а</strong>зных<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>. Брюш<strong>к</strong><strong>о</strong> <strong>в</strong>и<strong>с</strong>ячее, пер<strong>в</strong>ый <strong>с</strong>егмент<br />

<strong>с</strong>низу не <strong>о</strong>тделен <strong>о</strong>т <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>о</strong>й<br />

или перетяж<strong>к</strong><strong>о</strong>й. Длин<strong>а</strong> 3—5 мм (ри<strong>с</strong> 1138).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне г<strong>о</strong>л<strong>а</strong>р<strong>к</strong>ти<strong>к</strong>и (<strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong>), из них д<strong>в</strong><strong>а</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).

1138. Embolemus breviscapus Brues; <strong>о</strong>бщий <strong>в</strong>ид, X8,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Brues, 1933). 1139. Holepyrls<br />

praecursor Brues; <strong>о</strong>бщий <strong>в</strong>ид, Х23,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Brues, 1933). 1140. Pentachrysis Imperiosa<br />

СЕМЕЙСТВО BETHYLIDAE HAL1DAY, 1840<br />

Мел<strong>к</strong>ие и <strong>с</strong>редней <strong>в</strong>еличины, <strong>с</strong><strong>а</strong>м<strong>к</strong>и бе<strong>с</strong><strong>к</strong>рылые.<br />

В передних <strong>к</strong>рыльях <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле <strong>о</strong>чень уз<strong>к</strong><strong>о</strong>е, ед<strong>в</strong><strong>а</strong> р<strong>а</strong>зличим<strong>о</strong>е; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

<strong>в</strong> <strong>в</strong>иде ут<strong>о</strong>лщения жил<strong>к</strong>и, <strong>о</strong>т <strong>к</strong><strong>о</strong>нц<strong>а</strong> ее, н<strong>а</strong><br />

<strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>тх<strong>о</strong>дит п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> r-rs, перех<strong>о</strong>дящ<strong>а</strong>я <strong>в</strong> переднюю <strong>в</strong>ет<strong>в</strong>ь<br />

RS, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женную п<strong>о</strong>лу<strong>к</strong>руг<strong>о</strong>м, <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тью <strong>к</strong>з<strong>а</strong>ди; <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS нет, ег<strong>о</strong> з<strong>а</strong>дней <strong>в</strong>ет<strong>в</strong>и<br />

т<strong>о</strong>же; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> (r-m) прям<strong>а</strong>я;<br />

М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины<br />

длины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бе ее <strong>в</strong>ет<strong>в</strong>и не д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; Си перех<strong>о</strong>дит <strong>в</strong> з<strong>а</strong>днюю <strong>в</strong>ет<strong>в</strong>ь<br />

М. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> пр<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я; <strong>а</strong>нтенны при<strong>к</strong>реплены<br />

низ<strong>к</strong><strong>о</strong>, <strong>с</strong> 12—13 члени<strong>к</strong><strong>а</strong>ми. П<strong>а</strong>р<strong>а</strong>зиты личин<strong>о</strong><strong>к</strong><br />

жу<strong>к</strong><strong>о</strong><strong>в</strong>, гу<strong>с</strong>ениц чешуе<strong>к</strong>рылых (ри<strong>с</strong>. 1139). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 100 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне (700 <strong>в</strong>ид<strong>о</strong><strong>в</strong>), 23 р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Ри<strong>с</strong>. 1138—1141. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bethylidea<br />

355<br />

Sm.; <strong>о</strong>бщий <strong>в</strong>ид, Хб,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Tillyard, 1926). 1141. Ustodryinus<br />

mortuorum Brues; <strong>о</strong>бщий <strong>в</strong>ид, X10,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>о</strong><strong>с</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Brues, 1933)<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), п<strong>а</strong>ле<strong>о</strong>гене<br />

3. Е<strong>в</strong>р<strong>о</strong>пы, не<strong>о</strong>гене Ю, Азии (янт<strong>а</strong>рь Бирмы)<br />

и д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>а</strong> из чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жений<br />

В. Афри<strong>к</strong>и.<br />

СЕМЕЙСТВО CHRYSIDIDAE LATREILLE, 1802.<br />

ОСЫ БЛЕСТЯНКИ<br />

[nom. transl. В<strong>о</strong>тег, 1919 (ex Chrysidides Latreille, 1802)]<br />

Мел<strong>к</strong>ие и <strong>с</strong>редних р<strong>а</strong>змер<strong>о</strong><strong>в</strong>, длин<strong>а</strong> 2—•<br />

18 мм, <strong>о</strong><strong>к</strong>р<strong>а</strong>шенные, мет<strong>а</strong>лличе<strong>с</strong><strong>к</strong>и бле<strong>с</strong>тящие.<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, н<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>зличи*<br />

м<strong>о</strong>; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> я<strong>с</strong>н<strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> (r-m) из<strong>о</strong>гнут<strong>а</strong> п<strong>о</strong>лу<strong>к</strong>руг<strong>о</strong>м, <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong><br />

р<strong>а</strong>диу<strong>с</strong> <strong>в</strong> ме<strong>с</strong>те н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> RS; п<strong>о</strong><strong>с</strong>ледний<br />

длинный, пл<strong>а</strong><strong>в</strong>н<strong>о</strong> из<strong>о</strong>гнутый <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>тью <strong>к</strong>з<strong>а</strong>ди,<br />

<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> передней <strong>в</strong>ет<strong>в</strong>ью М; передняя<br />

<strong>в</strong>ет<strong>в</strong>ь RS <strong>с</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нным <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием,<br />

<strong>о</strong>тх<strong>о</strong>дит неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>т п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и,<br />

http://jurassic.ru/<br />

23*

-rs немн<strong>о</strong>г<strong>о</strong> ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины <strong>к</strong>рыл<strong>а</strong>, <strong>в</strong>ершин<strong>а</strong><br />

ее прям<strong>а</strong>я; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>ередине<br />

длины <strong>к</strong>рыл<strong>а</strong>, <strong>о</strong>бе ее <strong>в</strong>ет<strong>в</strong>и не д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

<strong>к</strong>рыл<strong>а</strong>; Си <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong><br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления М. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я,<br />

<strong>о</strong>дн<strong>о</strong>й ширины <strong>с</strong> грудью; <strong>а</strong>нтенны при<strong>к</strong>реплены<br />

низ<strong>к</strong><strong>о</strong>, из 13 члени<strong>к</strong><strong>о</strong><strong>в</strong>. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

из трех, реже из четырех <strong>с</strong>егмент<strong>о</strong><strong>в</strong> (у<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> три — пять), <strong>о</strong><strong>с</strong>т<strong>а</strong>льные <strong>с</strong>егменты пре<strong>в</strong>р<strong>а</strong>щены<br />

<strong>в</strong> трубч<strong>а</strong>тый яйце<strong>к</strong>л<strong>а</strong>д, <strong>в</strong>тяги<strong>в</strong><strong>а</strong>ющий<strong>с</strong>я<br />

<strong>в</strong> брюш<strong>к</strong><strong>о</strong>. П<strong>а</strong>р<strong>а</strong>зитируют <strong>в</strong> гнезд<strong>а</strong>х <strong>в</strong>е<strong>с</strong>п<strong>о</strong>идей<br />

и <strong>с</strong>фе<strong>к</strong><strong>о</strong>идей (ри<strong>с</strong>. 1140). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

(1500 <strong>в</strong>ид<strong>о</strong><strong>в</strong>), р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х и <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х; три р<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и ми<strong>о</strong>цене<br />

С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО DRYINIDAE HALIDAY, 1833<br />

Мел<strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, п<strong>о</strong>х<strong>о</strong>жие н<strong>а</strong> мур<strong>а</strong><strong>в</strong>ье<strong>в</strong><br />

(2,4—7 мм); <strong>с</strong><strong>а</strong>м<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бе<strong>с</strong><strong>к</strong>рылые. В передних<br />

<strong>к</strong>рыльях <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле уз<strong>к</strong><strong>о</strong>е, н<strong>о</strong> я<strong>с</strong>н<strong>о</strong>е;<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я; RS <strong>о</strong>тх<strong>о</strong>дит<br />

<strong>о</strong>т R пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>;<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л RS <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>й, р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong><br />

п<strong>о</strong>перечн<strong>о</strong>й жил<strong>к</strong>и r-rs, п<strong>о</strong><strong>с</strong>ледняя <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> бы перех<strong>о</strong>дящ<strong>а</strong>я <strong>в</strong> из<strong>о</strong>гнутую переднюю<br />

<strong>в</strong>ет<strong>в</strong>ь RS; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние передней <strong>в</strong>ет<strong>в</strong>и<br />

RS п<strong>о</strong>х<strong>о</strong>же н<strong>а</strong> п<strong>о</strong>перечную жил<strong>к</strong>у; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

и пер<strong>в</strong><strong>а</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячеи удлиненные,<br />

<strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong><br />

ди<strong>с</strong>т<strong>а</strong>льнее н<strong>а</strong>ч<strong>а</strong>л<strong>а</strong> RS, <strong>о</strong>бе ее <strong>в</strong>ет<strong>в</strong>и д<strong>о</strong>х<strong>о</strong>дят<br />

д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong>; Си перех<strong>о</strong>дит <strong>в</strong> з<strong>а</strong>днюю<br />

<strong>в</strong>ет<strong>в</strong>ь М. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я, п<strong>о</strong>перечн<strong>а</strong>я, гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я;<br />

<strong>а</strong>нтенны при<strong>к</strong>реплены у д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>ти н<strong>а</strong>лични<strong>к</strong><strong>а</strong>, де<strong>с</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Переднегрудь<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>ытянут<strong>а</strong>я; бедр<strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии р<strong>а</strong>здуты;<br />

передние л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> <strong>к</strong>леще<strong>о</strong>бр<strong>а</strong>зными члени<strong>к</strong><strong>о</strong>м<br />

и <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м. Брюш<strong>к</strong><strong>о</strong> <strong>в</strong>и<strong>с</strong>ячее. П<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> нимф<strong>а</strong>х и <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых Fulgoridae и<br />

Cicadellidae (ри<strong>с</strong>. 1141). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> 16 шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них три <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО CRETAVIDEA<br />

Отлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т <strong>в</strong><strong>с</strong>ех из<strong>в</strong>е<strong>с</strong>тных переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шими<strong>с</strong>я р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>лениями <strong>в</strong>ет<strong>в</strong>ей<br />

RS и М, чт<strong>о</strong> дел<strong>а</strong>ет их п<strong>о</strong>х<strong>о</strong>жими н<strong>а</strong> пред<strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц. Пер<strong>в</strong><strong>а</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> н<strong>а</strong>д <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>с</strong>д<strong>в</strong>инут<strong>а</strong> <strong>о</strong>дн<strong>а</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> друг<strong>о</strong>й; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>, н<strong>а</strong> <strong>о</strong>чень неб<strong>о</strong>льш<strong>о</strong>м<br />

пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong>е <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й<br />

ячеей. Мел. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cretavidae.<br />

356<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО CRETAVIDAE SHAROV, 1957<br />

К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>о</strong>, шире <strong>о</strong>бычн<strong>о</strong>г<strong>о</strong>,<br />

неб<strong>о</strong>льш<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; <strong>к</strong><strong>о</strong><strong>с</strong><strong>а</strong>я б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> перех<strong>о</strong>дит,<br />

<strong>о</strong>бр<strong>а</strong>зуя <strong>о</strong>дну линию, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS;<br />

между <strong>в</strong>ет<strong>в</strong>ями RS д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и; из<br />

них пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>а</strong>я перех<strong>о</strong>дит неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> п<strong>о</strong>перечную жил<strong>к</strong>у r-rs; М р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я<br />

п<strong>о</strong>чти н<strong>а</strong> <strong>с</strong>ередине длины <strong>к</strong>рыл<strong>а</strong>; д<strong>в</strong>е п<strong>о</strong>перечные<br />

жил<strong>к</strong>и между М и Си; Си <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления<br />

RS. Мел. Один р<strong>о</strong>д.<br />

Cretavus Sharov, 1957. Тип р<strong>о</strong>д<strong>а</strong> — С. sibiricus<br />

Sharov, 1957; н. мел, Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>яр<strong>с</strong><strong>к</strong>ий<br />

<strong>к</strong>р<strong>а</strong>й (<strong>с</strong>ен<strong>о</strong>н, Б. К<strong>а</strong><strong>с</strong>). RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я немн<strong>о</strong>г<strong>о</strong><br />

пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льнее <strong>с</strong>лияния з<strong>а</strong>дней ег<strong>о</strong> <strong>в</strong>ет<strong>в</strong>и <strong>с</strong><br />

передней <strong>в</strong>ет<strong>в</strong>ью М; д<strong>в</strong>е ячеи между <strong>в</strong>ет<strong>в</strong>ями<br />

RS; ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея треуг<strong>о</strong>льн<strong>а</strong>я, уз<strong>к</strong><strong>а</strong>я,<br />

меньше <strong>с</strong>ледующей ячеи; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея<br />

<strong>к</strong>рупнее ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й и немн<strong>о</strong>г<strong>о</strong> меньше пер<strong>в</strong><strong>о</strong>й<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й ячеи; п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> r-rs<br />

и з<strong>а</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющ<strong>а</strong>я ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льную ячею <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>о</strong>дну линию- Длин<strong>а</strong> переднег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong> 10 мм<br />

(ри<strong>с</strong>. 1142). Н. мел В. Сибири.<br />

НАДСЕМЕЙСТВО SCOLIIDEA<br />

Обычн<strong>о</strong> <strong>к</strong>рупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые, <strong>с</strong><strong>а</strong>м<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

бе<strong>с</strong><strong>к</strong>рылые. К<strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля нет, неб<strong>о</strong>льш<strong>а</strong>я<br />

птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; д<strong>в</strong>е п<strong>о</strong>перечные жил<strong>к</strong>и между<br />

<strong>в</strong>ет<strong>в</strong>ями RS, н<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние передней <strong>в</strong>ет<strong>в</strong>и RS<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я и ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячеи не <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я; меди<strong>а</strong>льн<strong>а</strong>я ячея<br />

з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>в</strong>ытянут<strong>а</strong>я <strong>в</strong> ширину. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Scoliidae (<strong>в</strong>не СССР) и Mutillidae.<br />

СЕМЕЙСТВО MUTILLIDAE LATREILLE, 1802.<br />

НЕМКИ<br />

[nom. transl. Stephens, 1829<br />

(ex Mutillariae Latreille, 1802)]<br />

Р<strong>а</strong>зличн<strong>о</strong>й <strong>в</strong>еличины: <strong>о</strong>т 5 д<strong>о</strong> 30 мм длин<strong>о</strong>й;<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и бе<strong>с</strong><strong>к</strong>рылые. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> гип<strong>о</strong>гн<strong>а</strong>тн<strong>а</strong>я; члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>а</strong>нтенн 12—13, пр<strong>о</strong><strong>с</strong>тые, при<strong>к</strong>реплены низ<strong>к</strong><strong>о</strong>.<br />

Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>в</strong>ыемч<strong>а</strong>тым з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем.<br />

Средние т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>ближены, <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я;<br />

передние бедр<strong>а</strong> не ут<strong>о</strong>лщены; пер<strong>в</strong>ый члени<strong>к</strong><br />

з<strong>а</strong>дних л<strong>а</strong>п<strong>о</strong><strong>к</strong> цилиндриче<strong>с</strong><strong>к</strong>ий. Брюш<strong>к</strong><strong>о</strong><br />

<strong>в</strong>и<strong>с</strong>ячее, <strong>с</strong>егмент<strong>о</strong><strong>в</strong> ше<strong>с</strong>ть — <strong>с</strong>емь; пер<strong>в</strong>ый <strong>с</strong>егмент<br />

<strong>с</strong>низу <strong>о</strong>тделен <strong>о</strong>т <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>о</strong>й<br />

или перетяж<strong>к</strong><strong>о</strong>й. П<strong>а</strong>р<strong>а</strong>зиты личин<strong>о</strong><strong>к</strong> и<br />

<strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>о</strong><strong>к</strong> пчел, <strong>о</strong><strong>с</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> и<strong>с</strong><strong>к</strong>лючение — жу<strong>к</strong><strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1143). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong> и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ты<strong>с</strong>яч <strong>в</strong>ид<strong>о</strong><strong>в</strong>, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х и <strong>с</strong>убтр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х; <strong>о</strong>дин<br />

<strong>в</strong>ымерший р<strong>о</strong>д из<strong>в</strong>е<strong>с</strong>тен из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.

Ри<strong>с</strong>. 1142—1149. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

1142. Cretavus sibiricus Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, Х4,7, <strong>в</strong>. мел.<br />

Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>яр<strong>с</strong><strong>к</strong>ий <strong>к</strong>р<strong>а</strong>й (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1956). 1143. Promutilla megaiophthalma<br />

Brues; <strong>о</strong>бщий <strong>в</strong>ид, хЮ,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Brues, 1933). 1 144. Р<strong>о</strong>пег<strong>а</strong> umbrata Popov; <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х10,0, ми<strong>о</strong>цен, К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (П<strong>о</strong>п<strong>о</strong><strong>в</strong>, 1932). 1 145. Dolichoderus antiquus<br />

Carpenter; <strong>о</strong>бщий <strong>в</strong>ид, х6,0, ми<strong>о</strong>цен, С. Амери<strong>к</strong><strong>а</strong> (Carpenter,<br />

НАДСЕМЕЙСТВО FORMICIDEA.<br />

МУРАВЬИ<br />

Р<strong>а</strong>зн<strong>о</strong>й <strong>в</strong>еличины: <strong>о</strong>т 2 д<strong>о</strong> 25 мм длин<strong>о</strong>й.<br />

Р<strong>а</strong>зделяют<strong>с</strong>я н<strong>а</strong> <strong>к</strong>рыл<strong>а</strong>тых <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> и <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> и<br />

нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>итых бе<strong>с</strong><strong>к</strong>рылых <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>-р<strong>а</strong>б<strong>о</strong>чих.<br />

Крылья <strong>с</strong> <strong>о</strong>бедненным жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нием; <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

п<strong>о</strong>ле е<strong>с</strong>ть; птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я<br />

жил<strong>к</strong><strong>а</strong> <strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> RS, <strong>о</strong>бр<strong>а</strong>зует приблизительн<strong>о</strong><br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тную <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льную ячею,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> бы «п<strong>о</strong>д<strong>в</strong>ешенную» з<strong>а</strong> уг<strong>о</strong>л, <strong>с</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>о</strong>й<br />

ячеей не <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ет<strong>с</strong>я; между <strong>в</strong>ет<strong>в</strong>ями<br />

RS <strong>о</strong>дн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> или ее нет; меди<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; Си <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е <strong>к</strong>рыл<strong>а</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>тебельч<strong>а</strong>т<strong>о</strong>е;<br />

<strong>с</strong>тебеле<strong>к</strong> из пер<strong>в</strong>ых д<strong>в</strong>ух члени<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

1146<br />

Crefavidea, Formicidea, Scoliidea<br />

1930). Н46. Monotnorium minimum Buckley; <strong>о</strong>бщий <strong>в</strong>ид, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Essig, 1942). 1147. Aeromyrma sophiae Emery; <strong>о</strong>бщий <strong>в</strong>ид'<br />

Xl2,0, <strong>с</strong>р. ми<strong>о</strong>цен, янт<strong>а</strong>рь Сицилии (Emery, 1890). 1 148. Lasiu s<br />

martynovi Popov; <strong>о</strong>бщий <strong>в</strong>ид, x 3,7, п<strong>а</strong>ле<strong>о</strong>ген, К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з (П<strong>о</strong>п<strong>о</strong><strong>в</strong>,<br />

1932). 1 149. Gaesomyrmex corniger Emery; <strong>о</strong>бщий <strong>в</strong>ид, х7,5,<br />

<strong>с</strong>р. ми<strong>о</strong>цен, янт<strong>а</strong>рь Сицилии (Emery, 1890)<br />

е<strong>с</strong>ли <strong>в</strong> <strong>с</strong>тебель<strong>к</strong>е <strong>о</strong>дин члени<strong>к</strong>, т<strong>о</strong> н<strong>а</strong> нем т<strong>о</strong>рч<strong>а</strong>щий<br />

<strong>в</strong><strong>в</strong>ерх <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Одн<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Formicidae (б<strong>о</strong>лее 3500 <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>в</strong><br />

тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х), р<strong>а</strong>зделенн<strong>о</strong>е н<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Ponerinae, Dolichoderinae, Myrmiciп<strong>а</strong>е,<br />

Formicinae, Pseudomyrmicinae; п<strong>о</strong><strong>с</strong>леднее<br />

<strong>в</strong>не СССР.<br />

ПОДСЕМЕЙСТВО PONERINAE SMITH, 1851<br />

![nom. transl. Dalla-Torre, 1893<br />

(ex Myrmicidae Smith, 1851)]<br />

Стебеле<strong>к</strong> брюш<strong>к</strong><strong>а</strong> <strong>к</strong><strong>а</strong>жет<strong>с</strong>я <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящим из<br />

<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> пети<strong>о</strong>лю<strong>с</strong><strong>а</strong>, <strong>с</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>й чешуй<strong>к</strong><strong>о</strong>й;<br />

п<strong>о</strong><strong>с</strong>тпети<strong>о</strong>лю<strong>с</strong> <strong>о</strong>чень <strong>в</strong>ели<strong>к</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> и третий<br />

357<br />

http://jurassic.ru/

<strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong>, <strong>о</strong>тделен <strong>о</strong>т п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> перетяж<strong>к</strong><strong>о</strong>й.<br />

В передних <strong>к</strong>рыльях б<strong>о</strong>льш<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>;<br />

<strong>в</strong>ет<strong>в</strong>и RS и М прямые, п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>лдельные;<br />

ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>, длинн<strong>а</strong>я<br />

(ри<strong>с</strong>. 1144). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х,<br />

немн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> Се<strong>в</strong>ерн<strong>о</strong>м п<strong>о</strong>луш<strong>а</strong>рии; 13 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (<strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

янт<strong>а</strong>ре), К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>, С. и Ю. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО DOLICHODERINAE FOREL, 1878<br />

[nom. transl. Dalla-Torre, 1893<br />

(ex Dolichoderidae Forel, 1878)]<br />

Одни из мел<strong>к</strong>их мур<strong>а</strong><strong>в</strong>ье<strong>в</strong>. Стебеле<strong>к</strong> брюш<strong>к</strong><strong>а</strong><br />

из <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>, ж<strong>а</strong>л<strong>о</strong> рудимент<strong>а</strong>рн<strong>о</strong>е.<br />

В передних <strong>к</strong>рыльях неб<strong>о</strong>льш<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>;<br />

RS <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> М д<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления, и<br />

п<strong>о</strong>эт<strong>о</strong>му з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячея <strong>к</strong><strong>а</strong>жет<strong>с</strong>я<br />

<strong>с</strong>идящей н<strong>а</strong> <strong>с</strong>тебель<strong>к</strong>е; <strong>в</strong>ет<strong>в</strong>и RS и М<br />

из<strong>о</strong>гнутые и не д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong><br />

(ри<strong>с</strong>. 1145). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> тр<strong>о</strong>пи<strong>к</strong><strong>а</strong>х; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии 17 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), п<strong>а</strong>ле<strong>о</strong>гене 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО MYRMICINAE SMITH, 1851<br />

[nom. transl. Dalla-Torre, 1893 (ex Myrmicidae Smith,<br />

1851)]<br />

Н<strong>а</strong>иб<strong>о</strong>лее <strong>к</strong>рупные и шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные<br />

мур<strong>а</strong><strong>в</strong>ьи. Стебеле<strong>к</strong> из д<strong>в</strong>ух <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

ж<strong>а</strong>л<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>. В передних <strong>к</strong>рыльях д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong><br />

<strong>к</strong>рупн<strong>а</strong>я птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong>; RS <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

М д<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ления, н<strong>о</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я<br />

ячея <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я; б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong><br />

<strong>в</strong>п<strong>а</strong>д<strong>а</strong>ет <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние RS; <strong>о</strong>бе <strong>в</strong>ет<strong>в</strong>и RS<br />

д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong>рыл<strong>а</strong> (ри<strong>с</strong>. 1146, 1147). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии 36 р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и и <strong>в</strong> чет<strong>в</strong>ертичных <strong>о</strong>тл<strong>о</strong>жениях<br />

В. Афри<strong>к</strong>и.<br />

ПОДСЕМЕЙСТВО FORMICINAE MAYR, 1855<br />

[nom. transl. Ashmead, 1901 (ex Formicidae Mayr, 1855)]<br />

Длин<strong>а</strong> мур<strong>а</strong><strong>в</strong>ье<strong>в</strong> <strong>о</strong>т 2 д<strong>о</strong> 20 мм. Стебеле<strong>к</strong><br />

из <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>; <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>к</strong>л<strong>о</strong><strong>а</strong><strong>к</strong>и, <strong>в</strong>ытянут<strong>о</strong>е<br />

<strong>в</strong> труб<strong>о</strong>ч<strong>к</strong>у, <strong>о</strong><strong>к</strong>ружен<strong>о</strong> <strong>в</strong>енчи<strong>к</strong><strong>о</strong>м <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>;<br />

ж<strong>а</strong>л<strong>а</strong> нет. Обычн<strong>о</strong> жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>;<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют или передняя <strong>в</strong>ет<strong>в</strong>ь RS<br />

(Lasius), или з<strong>а</strong>дняя <strong>в</strong>ет<strong>в</strong>ь М (Camponotus);<br />

<strong>в</strong> пер<strong>в</strong><strong>о</strong>м <strong>с</strong>луч<strong>а</strong>е <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея (ри<strong>с</strong>- 1148, 1149). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненных <strong>в</strong>ид<strong>о</strong><strong>в</strong> (пре<strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют<br />

терм<strong>о</strong>филы); <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

15 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь),<br />

К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>пы, Яп<strong>о</strong>нии, С. Амери<strong>к</strong>и.<br />

358<br />

http://jurassic.ru/<br />

НАДСЕМЕЙСТВО POMPILIDEA.<br />

ДОРОЖНЫЕ ОСЫ<br />

Средней <strong>в</strong>еличины и <strong>к</strong>рупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые,<br />

длин<strong>а</strong> 5—27 мм. Р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й р<strong>а</strong><strong>с</strong>ц<strong>в</strong>ет<strong>к</strong>и и<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> мет<strong>а</strong>лличе<strong>с</strong><strong>к</strong>им бле<strong>с</strong><strong>к</strong><strong>о</strong>м. В переднем<br />

<strong>к</strong>рыле х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идны <strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е п<strong>о</strong>ле, птер<strong>о</strong><strong>с</strong>тигм<strong>а</strong><br />

и б<strong>а</strong>з<strong>а</strong>льн<strong>а</strong>я п<strong>о</strong>перечн<strong>а</strong>я жил<strong>к</strong><strong>а</strong> rs-m;<br />

RS р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляет<strong>с</strong>я д<strong>о</strong> <strong>с</strong>лияния <strong>с</strong> <strong>в</strong>ет<strong>в</strong>ью М, п<strong>о</strong>эт<strong>о</strong>му<br />

<strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я и ди<strong>с</strong><strong>к</strong><strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я ячеи<br />

<strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я; <strong>с</strong>убр<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я ячея длинн<strong>а</strong>я,<br />

длиннее ячей dc и rs 2 <strong>в</strong>ме<strong>с</strong>те <strong>в</strong>зятых и длиннее<br />

пер<strong>в</strong><strong>о</strong>й р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й ячеи; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>а</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>а</strong>я<br />

ячея, между R и М, <strong>к</strong><strong>о</strong>р<strong>о</strong>че п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины длины<br />

<strong>к</strong>рыл<strong>а</strong>; меди<strong>а</strong>льн<strong>а</strong>я ячея з<strong>а</strong><strong>к</strong>рыт<strong>а</strong>я; Си <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льнее <strong>с</strong>ередины длины <strong>к</strong>рыл<strong>а</strong>,<br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong> ширине передне<strong>с</strong>пин<strong>к</strong>е<br />

или шире ее; <strong>а</strong>нтенны длинные;<br />

гл<strong>а</strong>з<strong>а</strong> без рез<strong>к</strong><strong>о</strong>й <strong>в</strong>ырез<strong>к</strong>и п<strong>о</strong> <strong>в</strong>нутреннему<br />

<strong>к</strong>р<strong>а</strong>ю. Н<strong>о</strong>ги длинные, з<strong>а</strong>дние г<strong>о</strong>лени з<strong>а</strong>х<strong>о</strong>дят<br />

з<strong>а</strong> <strong>в</strong>ершину брюш<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 1150). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Pompilidae, Rhopalosomatidae;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии не<br />

н<strong>а</strong>йден<strong>о</strong>.<br />

•RS,+ м,<br />

Ри<strong>с</strong>. 1150—1151. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pompilidea, Apidea<br />

1150. Calllcurgus hyalinatus Fabricius; <strong>о</strong>бщий <strong>в</strong>ид, X4,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Ш<strong>а</strong>рп, 1910). 1151. Andrena sp.; <strong>о</strong>бщий <strong>в</strong>ид,х5,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Essig, 1942)

СЕМЕЙСТВО POMPILIDAE LEACH, 1815<br />

[nom. transl. Leach, 1819 (ex Pompilides Leach, 1815)]<br />

Брюш<strong>к</strong><strong>о</strong> н<strong>а</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>м <strong>с</strong>тебель<strong>к</strong>е. П<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> п<strong>а</strong>у<strong>к</strong><strong>а</strong>х и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong> (3000 <strong>в</strong>ид<strong>о</strong><strong>в</strong>), н<strong>а</strong>иб<strong>о</strong>лее <strong>о</strong>бильны<br />

<strong>в</strong> ж<strong>а</strong>р<strong>к</strong><strong>о</strong>м п<strong>о</strong>я<strong>с</strong>е; <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь), 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и<br />

НАДСЕМЕЙСТВО APIDEA.<br />

ПЧЕЛИНЫЕ<br />

Средней <strong>в</strong>еличины и <strong>к</strong>рупные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

Обычн<strong>о</strong> тел<strong>о</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong>и<strong>с</strong>т<strong>о</strong>е, реже г<strong>о</strong>л<strong>о</strong>е. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>к</strong>рылье<strong>в</strong> <strong>о</strong>чень близ<strong>к</strong><strong>о</strong> <strong>к</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>му у<br />

Pompilidae, <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я менее х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным<br />

<strong>к</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>льным п<strong>о</strong>лем и б<strong>о</strong>лее длинн<strong>о</strong>й<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>й ячеей (г), р<strong>а</strong><strong>в</strong>н<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

длины <strong>к</strong>рыл<strong>а</strong> и б<strong>о</strong>льше. Антенны <strong>к</strong><strong>о</strong>ленч<strong>а</strong>тые,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Передне<strong>с</strong>пин<strong>к</strong><strong>а</strong> <strong>с</strong> я<strong>с</strong>н<strong>о</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленными, <strong>в</strong>ыд<strong>а</strong>ющими<strong>с</strong>я плече<strong>в</strong>ыми бугр<strong>а</strong>ми;<br />

пер<strong>в</strong>ый члени<strong>к</strong> з<strong>а</strong>дних л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>о</strong>бычн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширен, н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не гу<strong>с</strong>т<strong>о</strong> п<strong>о</strong><strong>к</strong>рыт<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми <strong>в</strong> <strong>в</strong>иде щет<strong>о</strong>ч<strong>к</strong>и (ри<strong>с</strong>. 1151). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне; <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Apidae.<br />

СЕМЕЙСТВО APIDAE LATREILLE, 1802. ПЧЕЛЫ<br />

[nom. transl. Leach, 1817 (ex Apiariae Latreille, 1802)]<br />

Антенн<strong>а</strong>льн<strong>а</strong>я ям<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong>единен<strong>а</strong> пр<strong>о</strong><strong>с</strong>тым<br />

ш<strong>в</strong><strong>о</strong>м <strong>с</strong><strong>о</strong> ш<strong>в</strong><strong>о</strong>м fronto-clypeus. Губные щупи<strong>к</strong>и<br />

<strong>с</strong> д<strong>в</strong>умя удлиненными члени<strong>к</strong><strong>а</strong>ми; ширин<strong>а</strong><br />

labrum б<strong>о</strong>льше ее длины. П<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ие и<br />

<strong>о</strong>бще<strong>с</strong>т<strong>в</strong>енные н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), В. Сибири, Ю. Азии<br />

(янт<strong>а</strong>рь Бирмы) и С- Амери<strong>к</strong>и.<br />

http://jurassic.ru/

А яд ру<strong>с</strong><strong>о</strong> <strong>в</strong> Н. il916. Труб<strong>к</strong>и чер<strong>в</strong>ей из <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Amphictenidae <strong>в</strong> .ру<strong>с</strong><strong>а</strong><strong>к</strong><strong>о</strong>м ми<strong>о</strong>цене. Из<strong>в</strong>. ими. А<strong>к</strong><strong>а</strong>д.<br />

н<strong>а</strong>у<strong>к</strong>, СПб., <strong>с</strong>тр. 227—232.<br />

Бей-Биен<strong>к</strong><strong>о</strong> Г. Я. 1936. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>к</strong><strong>о</strong>жиет<strong>о</strong><strong>к</strong>рылые.<br />

Ф<strong>а</strong>ун<strong>а</strong> СССР. Изд. З<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР, н<strong>о</strong><strong>в</strong>. <strong>с</strong>ер.,<br />

№ 5, <strong>с</strong>тр. 1—239.— 4950. Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые т<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>ые. Ф<strong>а</strong>ун<strong>а</strong><br />

СССР. Изд. З<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР, н<strong>о</strong><strong>в</strong>. <strong>с</strong>ер., № 40.<br />

Бе<strong>к</strong> <strong>к</strong> ер Э. Г. 1950—1954. К <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>у <strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении<br />

<strong>к</strong>рыл<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, ч. 1. 'Ве<strong>с</strong>ти. М<strong>о</strong><strong>с</strong><strong>к</strong>. ун-т<strong>а</strong>, 1952, № 9,<br />

<strong>с</strong>тр. '59—68; ч. 2, 1954, № 5, <strong>с</strong>тр. 1>19—130. 'Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong> Е. Э. '1940. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые перм<strong>с</strong><strong>к</strong>ие ци<strong>к</strong><strong>а</strong>ды<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Prosbolidae <strong>с</strong> р. С<strong>о</strong>яны. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

ин-т<strong>а</strong> АН СССР, т. 9, <strong>в</strong>ып. 2, <strong>с</strong>тр. 5—79.—1946—1948.<br />

Очер<strong>к</strong>и п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>й м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гии <strong>с</strong><strong>о</strong><strong>в</strong>ременных и<br />

перм<strong>с</strong><strong>к</strong>их Homoptera, ч. 1. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л.,<br />

№ 6 (1946), <strong>с</strong>тр. 741—766; ч. 2, <strong>с</strong>ер. би<strong>о</strong>л., № 1 (1948),<br />

<strong>с</strong>тр. 123'—1412.— 1947. Cicadoprosbole gen. et sp. nov.—<br />

перех<strong>о</strong>дн<strong>а</strong>я ф<strong>о</strong>рм<strong>а</strong> между перм<strong>с</strong><strong>к</strong>ими Prosbolidae и<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременными Cicadidae. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 5, № 5,<br />

<strong>с</strong>тр. 445—448.— Г948. Типы изменчи<strong>в</strong><strong>о</strong><strong>с</strong>ти жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong>рылье<strong>в</strong> Homoptera. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л., № 2,<br />

<strong>с</strong>тр. 87—92.— 1948. Пермшие ци<strong>к</strong><strong>а</strong>ды <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Scyti*<br />

ncpteridae. <strong>с</strong> р. С<strong>о</strong>яны. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР,<br />

т. 15, <strong>в</strong>ыи. 2, <strong>с</strong>тр. il—42.— 1949<strong>а</strong>. Мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие Homoptera<br />

Средней Азии. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 22,<br />

<strong>с</strong>тр. '1—68.— 19496. Н<strong>о</strong><strong>в</strong><strong>о</strong>е перм<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Boreoscytidae<br />

.и <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong> <strong>о</strong> фил<strong>о</strong>гении пред<strong>к</strong><strong>о</strong><strong>в</strong> Homoptera. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

ин-т<strong>а</strong>. АН СССР, т. 20, <strong>с</strong>тр. 171—182.— 19(50.<br />

Юр<strong>с</strong><strong>к</strong>ие Palaeontinidae н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения н<strong>а</strong> р.<br />

Ии. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 71, № 6, <strong>с</strong>тр. 1(105—1108.—1951.<br />

С<strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зные н<strong>о</strong><strong>в</strong>ые и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из труппы<br />

п<strong>а</strong>р-<strong>а</strong>не<strong>о</strong>лтер. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 78, № 6. <strong>с</strong>тр. 1207—<br />

1210.—1952. Н<strong>о</strong><strong>в</strong>ые .р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые из перми Кузб<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> и<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые з<strong>а</strong>меч<strong>а</strong>ния <strong>о</strong>б ип<strong>с</strong><strong>в</strong>ичиид<strong>а</strong>х. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

ин-т<strong>а</strong> АН СССР, т. 40, <strong>с</strong>тр. '177—1,86—19533. Обз<strong>о</strong>р<br />

ф<strong>а</strong>уны р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых и <strong>с</strong>ен<strong>о</strong>ед<strong>о</strong><strong>в</strong> ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й и <strong>к</strong>узнец<strong>к</strong><strong>о</strong>й<br />

<strong>с</strong><strong>в</strong>ит Кузб<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 90, № 1, <strong>с</strong>тр. 97—<br />

100.—19536. Пер<strong>в</strong><strong>а</strong>я н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong> <strong>с</strong>ен<strong>о</strong>ед<strong>а</strong> (Psocoptera) из<br />

Куэнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. '90, № 2,<br />

<strong>с</strong>тр. 279—282.—1953<strong>в</strong>. Д<strong>в</strong><strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителя п<strong>о</strong>луже<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из ур<strong>о</strong>чищ<strong>а</strong> М<strong>а</strong>дыген. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 90, № 3, <strong>с</strong>тр. 461—464.—^1964. Н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong><br />

южн<strong>о</strong>й ци<strong>к</strong><strong>а</strong>ды <strong>в</strong> ми<strong>о</strong>цене З<strong>а</strong>п<strong>а</strong>дн<strong>о</strong>й Сибири. В<strong>о</strong>пр<strong>о</strong><strong>с</strong>ы<br />

ге<strong>о</strong>л<strong>о</strong>гии Азии, т. I. Изд. АН СССР, <strong>с</strong>тр. 799—^05.—<br />

1955. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из три<strong>а</strong><strong>с</strong><strong>а</strong> Сибири. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 105, № 5, <strong>с</strong>тр. 1100—<strong>Л1</strong>03.—19157. К <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>у<br />

<strong>о</strong>б э<strong>в</strong><strong>о</strong>люции р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылых (Homoptera). Тези<strong>с</strong>ы д<strong>о</strong><strong>к</strong>л<strong>а</strong>д<strong>о</strong><strong>в</strong><br />

III <strong>с</strong><strong>о</strong><strong>в</strong>ет. В<strong>с</strong>е<strong>с</strong>. энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б-<strong>в</strong><strong>а</strong>. 1967 г., ч. V—<br />

П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гия и фил<strong>о</strong>гения. Изд. АН СССР, <strong>с</strong>тр. 103—<br />

105.—il958. Н<strong>о</strong><strong>в</strong>ые и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые, ч. I. 1. Н<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

дре<strong>в</strong>нее <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>е р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рыл<strong>о</strong>е из 'Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. 2. Н<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Cicadocoridae и 'дре<strong>в</strong>ние<br />

Coleorrhyncha. 3. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Membracidae<br />

из <strong>в</strong>ерхнег<strong>о</strong> три<strong>а</strong><strong>с</strong><strong>а</strong> Средней Азии. М<strong>а</strong>тери<strong>а</strong>лы <strong>к</strong><br />

«О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>м п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», <strong>в</strong>ып. 2, <strong>с</strong>тр. 57—67.—1959.<br />

Н<strong>о</strong><strong>в</strong>ый р<strong>о</strong>д Borisrohdendorfia (Homoptera) из Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. Журн., № 3', <strong>с</strong>тр. 138'—140.—•<br />

1960<strong>а</strong>. Н<strong>о</strong><strong>в</strong>ые перм<strong>с</strong><strong>к</strong>ие р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые Е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong><strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

СССР. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 76, <strong>с</strong>тр. 1 —<br />

Л ИТЕРАТУРА<br />

112.— 19606. Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые н<strong>о</strong><strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители группы<br />

<strong>с</strong>терн<strong>о</strong>ринх из перми и мез<strong>о</strong>з<strong>о</strong>я СССР. М<strong>а</strong>тери<strong>а</strong>лы <strong>к</strong> «О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>м<br />

п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», <strong>в</strong>ып. 3, <strong>с</strong>тр. 109'—116.—' 1960<strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>к</strong>рылые СССР и <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>ы фил<strong>о</strong>гении <strong>о</strong>тряд<strong>а</strong>.<br />

П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. Журн., № 3, <strong>с</strong>тр. 28—42.— 1960. Die<br />

Archescytinidae als vermutliche Vorfahren der Blattlause.<br />

XI. Intern. Kongress f. Entomol., Verhandlungen, Bd. 1,<br />

S. 298—301.— 1962. Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые н<strong>о</strong><strong>в</strong>ые п<strong>о</strong>луже<strong>с</strong>т<strong>к</strong><strong>о</strong><strong>к</strong>рылые<br />

и <strong>с</strong>ен<strong>о</strong>еды. П<strong>а</strong>ле<strong>о</strong>нт. журн. № 1, <strong>с</strong>тр. 89—104. Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong> Э. Е. и М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong> О. М. 1951. Ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждение<br />

ми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> Тер<strong>с</strong><strong>к</strong><strong>о</strong>м Ал<strong>а</strong>т<strong>а</strong>у<br />

и ци<strong>к</strong><strong>а</strong>д<strong>к</strong><strong>а</strong> из нег<strong>о</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 78, № 4, <strong>с</strong>тр. 761 —<br />

763,— 1961. См. Р<strong>о</strong>денд<strong>о</strong>рф Б. Б., Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong><br />

Е. Э., М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong> О. М., Ш<strong>а</strong>р<strong>о</strong><strong>в</strong> А. Г.<br />

Берг Л. С. 1928. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>а</strong>я б<strong>а</strong>б<strong>о</strong>ч<strong>к</strong><strong>а</strong> из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong><br />

Иртыш<strong>а</strong>. «Прир<strong>о</strong>д<strong>а</strong>», № 10, <strong>с</strong>тр. 929. Б<strong>о</strong>г<strong>а</strong>ч е<strong>в</strong> А. В.<br />

194<strong>в</strong>. Ф<strong>а</strong>ун<strong>а</strong> бин<strong>а</strong>г<strong>а</strong>дине<strong>к</strong>их Кир<strong>о</strong><strong>в</strong>ых пл<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>. Жу<strong>к</strong>и С<strong>о</strong>leoptera.<br />

Тр. Е<strong>с</strong>т.-и<strong>с</strong>т<strong>о</strong>р. музея АН Азерб.ССР, <strong>в</strong>ып. I и<br />

II, <strong>с</strong>тр. 137—160. Б<strong>о</strong>рх<strong>с</strong>ениу<strong>с</strong> Н. С. 1950. Чер<strong>в</strong>ецы и<br />

гд<strong>а</strong>т<strong>о</strong><strong>в</strong><strong>к</strong>и СССР. Изд. АН СССР, <strong>с</strong>тр. 1—1250. Б р <strong>а</strong> у е р,<br />

Редтенб<strong>а</strong>хер и Г<strong>а</strong>нгльб<strong>а</strong>уэр (Brauer F., Redtenbacher<br />

J., Ganglbauer L.). 1889. Fossile Insekten aus<br />

der Juraformation Ost-Sibiriens. З<strong>а</strong>п. имп. <strong>а</strong><strong>к</strong><strong>а</strong>д. н<strong>а</strong>у<strong>к</strong><br />

СПб., <strong>с</strong>ер. 7, т. 36, № 15, <strong>с</strong>тр. 1—22.<br />

З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий М. Д. (Zalessky М.). 1926. Observation<br />

sur un nouvel insecte fossile du permien de Kargala.<br />

Bull. Soc. Geol. France, ser. 4, t. 26, p. 75—84,— 1928a.<br />

О н<strong>о</strong><strong>в</strong><strong>о</strong>м и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>м из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений<br />

К<strong>а</strong>рг<strong>а</strong>лы. Из<strong>в</strong>. Ге<strong>о</strong>л. <strong>к</strong><strong>о</strong>м-т<strong>а</strong>, т. 46, № 7, <strong>с</strong>тр. 693—699.—<br />

19286. Sur un nouvel insecte neuropteroide du permien du<br />

bassin de Kama. Bull. Soc. Entomol. France, 4 ser., t.<br />

28, p. 381—385.—1929a. О н<strong>о</strong><strong>в</strong>ых и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых из перм<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>тл<strong>о</strong>жений б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>о</strong><strong>в</strong> ре<strong>к</strong> К<strong>а</strong>мы, Вят<strong>к</strong>и и Бел<strong>о</strong>й.<br />

Тр. Об-<strong>в</strong><strong>а</strong> е<strong>с</strong>те<strong>с</strong>т<strong>в</strong><strong>о</strong>и<strong>с</strong>п. при К<strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>. ун-те, т. 52, зып. 1,<br />

<strong>с</strong>тр. 3—50.—19296. Sur un nouveau Prothorthoptera du<br />

permien de Kama. Ann. Soc. Geol. Nord., t. 54, p. 20—<br />

26.—1930. Sur deux representants nouveaux des Paleohemipteres<br />

du permien de la Kama et du Perebore dans<br />

le bassin de la Petchora. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л.,<br />

<strong>с</strong>тр. 1017—11027.—1931. О н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, н<strong>а</strong>йденных <strong>в</strong> углен<strong>о</strong><strong>с</strong>ных<br />

<strong>о</strong>тл<strong>о</strong>жениях Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> <strong>к</strong> <strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>те<br />

этих <strong>о</strong>тл<strong>о</strong>жений н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии энт<strong>о</strong>м<strong>о</strong>ф<strong>а</strong>уны. Бюлл.<br />

М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды, <strong>о</strong>тд. би<strong>о</strong>л., № 3—4, <strong>с</strong>тр.<br />

399—406.— 1932<strong>а</strong>. Observation sur les nouveaux insectes<br />

permiens de l'Europe Orientale. Bull. Soc. Geol. France,<br />

ser. 5, t. 2, p. 183—210.—19326. Observation sur les insectes<br />

trouves dans les depots a charbon du bassin de<br />

Kcusnetzk et sur l'age de ces derniers d'apres la faune<br />

entomologique. Bull. Soc. Geol. France, б ser., t. 2, p.<br />

209—1218. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий Ю. M. (Zalessky G.). 1931a.<br />

Observation sur un nouvel insecte libelluloide du permien<br />

du bassin du fleuve Kama. Ann. Soc. Geol. Nord, v.<br />

56, p. 36—41.—. 19316. О н<strong>о</strong><strong>в</strong><strong>о</strong>м <strong>с</strong>тре<strong>к</strong><strong>о</strong>з<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>м н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>м-<br />

из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> р. К<strong>а</strong>мы. Из<strong>в</strong>.<br />

АН СССР, <strong>с</strong>ер. би<strong>о</strong>л., <strong>с</strong>тр. 855—861.—1932. О жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong>рылье<strong>в</strong> <strong>с</strong>тре<strong>к</strong><strong>о</strong>з и п<strong>о</strong>ден<strong>о</strong><strong>к</strong> и их фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

р<strong>а</strong>з<strong>в</strong>итии. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л., <strong>с</strong>тр. 713—733.—<br />

1933. Sur deux nouveaux insectes permiens. Ann. Soc.<br />

Geol. Nord, t. 57, p. 136—144.— 1934. Sur deux represen-<br />

360<br />

http://jurassic.ru/

tants permiens nouveaux de l'ordre des Protorthopteres.<br />

Ann. Soc. Entomol. France, t. 108', p. 149—1617.— 1936. Sur<br />

deux restes d'insectes fossiles provenant du bassin de<br />

Kouznetzk et sur l'Sge geologique des depots qui les<br />

renferment. Bull. Soc. Geol. France, ser. 5, t. 5, p. 687—<br />

696.—11937a. Ancestors of some groups of the Present-dayinsects.<br />

«Nature», v. 140, p. '847—849'.— 19376. Nouveaux<br />

representants des Protohymenopteres et des Archodonates<br />

piovenant du terrain permien du bassin de la riviere Kama<br />

et leurs liens de parente. Ann. Soc. entomol. France,<br />

t. 106, p. 101—114.— 1937B. Перм<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые p. Сыл<strong>в</strong>ы<br />

и <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>ы э<strong>в</strong><strong>о</strong>люции <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, ч. I.<br />

«Пр<strong>о</strong>бл. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», т. 2—3, <strong>с</strong>тр. 601—607.—• 1937 г.<br />

Ч. II. «Пр<strong>о</strong>бл. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», т. 2—3, <strong>с</strong>ир. 609 1<br />

—G13.—<br />

1937д. Sur un representant d'un nouvel ordre d'insectes<br />

permiens. Ann. Soc. Geol. Nord (1936), t. 60, p. 50—71.—<br />

1938. Nouveaux insectes permiens de l'ordre des Embiodea.<br />

Ann. Soc. Geol. Nord, t. 63, p. 62—®1.—11939. Н<strong>о</strong><strong>в</strong>ые<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители Protohymenoptera, Homoptera, Hemipsocoptera,<br />

Psocoptera, Protoperlaria, Isoptera и Protoblattoidea.<br />

«Пр<strong>о</strong>бл. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», т. 5, <strong>с</strong>тр. 33—91.—1940.<br />

О яеи<strong>о</strong>т<strong>о</strong>рых н<strong>о</strong><strong>в</strong>ых н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong>х и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из<br />

б<strong>а</strong>рдин<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong> перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Ур<strong>а</strong>л<strong>а</strong>. Тр. XVII<br />

<strong>с</strong>е<strong>с</strong><strong>с</strong>ии междун<strong>а</strong>р. ге<strong>о</strong>л.. <strong>к</strong><strong>о</strong>нгре<strong>с</strong><strong>с</strong><strong>а</strong>. М., т. 6, <strong>с</strong>тр. 193—•<br />

196.—1943<strong>а</strong>. Кр<strong>а</strong>т<strong>к</strong><strong>о</strong>е изл<strong>о</strong>жение <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний <strong>к</strong> изменению<br />

термин<strong>о</strong>л<strong>о</strong>гии жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong>рылье<strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. З<strong>о</strong><strong>о</strong>л.<br />

журн., т. 22, <strong>в</strong>ып. 3, <strong>с</strong>тр. 154—169.—19436. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель<br />

н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, не<strong>с</strong>ущих элигры. «Прир<strong>о</strong>д<strong>а</strong>»,<br />

№ 3, <strong>с</strong>тр. 70—'7il.—11944. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель н<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

группы н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Ур<strong>а</strong>л<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 44, № 8, <strong>с</strong>тр. 370—372.—'1946<strong>а</strong>. О н<strong>о</strong><strong>в</strong><strong>о</strong>м<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителе Palaeodictyoptera из <strong>в</strong>ерхнег<strong>о</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>я<br />

Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пьгт. прир<strong>о</strong>ды,<br />

<strong>о</strong>тд. би<strong>о</strong>л., т. 51, № 445, <strong>с</strong>тр. 58—82'.—19466. Об<br />

<strong>о</strong>дн<strong>о</strong>м пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителе н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тр<strong>а</strong>д<strong>а</strong> п<strong>о</strong>лу<strong>с</strong>тре<strong>к</strong><strong>о</strong><strong>в</strong> i(Hemiodonata)<br />

из перми Ур<strong>а</strong>л<strong>а</strong>. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пьгт.<br />

прир<strong>о</strong>ды, <strong>о</strong>тд. би<strong>о</strong>л., <strong>с</strong>тр. 60—70.— 194б<strong>в</strong>. О н<strong>о</strong><strong>в</strong><strong>о</strong>й п<strong>о</strong>ден<strong>к</strong>е<br />

из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Ур<strong>а</strong>л<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН OOGP,<br />

т. 54, № 4, <strong>с</strong>тр. 363—365.— 19416 г. Н<strong>о</strong><strong>в</strong>ый пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель<br />

перм<strong>с</strong><strong>к</strong>их <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых. Д<strong>о</strong><strong>к</strong>л. АН ССОР, т. 51, № 7,<br />

<strong>с</strong>тр. 543—644.—1947. О д<strong>в</strong>ух н<strong>о</strong><strong>в</strong>ых перм<strong>с</strong><strong>к</strong>их жу<strong>к</strong><strong>а</strong>х.<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 6, № 8, <strong>с</strong>тр. 857—1860.—1948<strong>а</strong>. О н<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>в</strong>е<strong>с</strong>нян<strong>к</strong>е из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Ур'<strong>а</strong>л<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 60, № 6, <strong>с</strong>тр. 104.1-4043.—19486. О пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителе<br />

н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ющих элитр<strong>а</strong>ми.<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 59, № 2, <strong>с</strong>тр. 317—320.—4948<strong>в</strong>.<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений С<strong>о</strong>ли<strong>к</strong><strong>а</strong>м<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> и<br />

Чердыне<strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>й<strong>о</strong>н<strong>о</strong><strong>в</strong>. В<strong>о</strong>г<strong>а</strong>р. те<strong>о</strong>р. и при<strong>к</strong>л. ге<strong>о</strong>л., № 5,<br />

<strong>с</strong>тр. 44—62.—1949. Н<strong>о</strong><strong>в</strong>ый третичный мур<strong>а</strong><strong>в</strong>ей. «С<strong>о</strong><strong>в</strong>ет<strong>с</strong><strong>к</strong>.<br />

ге<strong>о</strong>л.», №40, <strong>с</strong>тр. 50—54.—1960 !<br />

197—1966<strong>а</strong>. Н<strong>о</strong><strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>о</strong>тряд<strong>о</strong><strong>в</strong> Protoblattoidea<br />

и Protorthoptera из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Ур<strong>а</strong>л<strong>а</strong>.<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 101, № 2, <strong>с</strong>тр. 347—350.—19556. Н<strong>о</strong><strong>в</strong>ые<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители п<strong>а</strong>ле<strong>о</strong>энт<strong>о</strong>м<strong>о</strong>ф<strong>а</strong>ун Ур<strong>а</strong>л<strong>а</strong> и их ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий<br />

<strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т. Ежег<strong>а</strong>дн. В<strong>с</strong>е<strong>с</strong>. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. <strong>о</strong>б-<strong>в</strong><strong>а</strong>, т. 15,<br />

<strong>с</strong>тр. 274—304.—1955<strong>в</strong>. О д<strong>в</strong>ух н<strong>о</strong><strong>в</strong>ых перм<strong>с</strong><strong>к</strong>их <strong>с</strong>гре<strong>к</strong><strong>о</strong>з<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бяых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>тр<strong>а</strong>д<strong>а</strong> Permodonata. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 101, № 4, <strong>с</strong>тр. 630-*33.<br />

<strong>а</strong>. Перм<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong> р. Сыл<strong>в</strong>ы и <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>ы э<strong>в</strong><strong>о</strong>люции <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

IV. Н<strong>о</strong><strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители группы Epiembiodea и<br />

э<strong>в</strong><strong>о</strong>люция эмбий. «В<strong>о</strong>пр<strong>о</strong><strong>с</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», т. I, <strong>с</strong>тр.<br />

41—60.— 19506. К <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>у <strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>те <strong>с</strong><strong>в</strong>иты С<strong>о</strong>ли<strong>к</strong><strong>а</strong>м<strong>с</strong><strong>к</strong>их<br />

плитня<strong>к</strong><strong>о</strong><strong>в</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 70, № 4, <strong>с</strong>тр. 683—<br />

685.— 1950<strong>в</strong>. Н<strong>о</strong><strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>о</strong>тряд<strong>а</strong> Protodonata. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды,<br />

<strong>о</strong>тд. ге<strong>о</strong>л., т. 25, № 4, <strong>с</strong>тр. 98-—108.— 1951 <strong>а</strong>. Н<strong>о</strong><strong>в</strong>ый пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель<br />

<strong>а</strong>рх<strong>о</strong>д<strong>о</strong>н<strong>а</strong>т. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 81, № 2, <strong>с</strong>тр. 269—<br />

27k— 19516. Н<strong>о</strong><strong>в</strong>ые перм<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые <strong>о</strong>тряд<strong>а</strong> предпрям<strong>о</strong><strong>к</strong>рылых<br />

Protorthoptera. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 78, № 5,<br />

<strong>с</strong>тр. 1005—1007.—1961<strong>в</strong>. Н<strong>о</strong><strong>в</strong>ый пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель перм<strong>с</strong><strong>к</strong>их<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>тряд<strong>а</strong> пред<strong>в</strong>е<strong>с</strong>нян<strong>о</strong><strong>к</strong> — Protoperlaria. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 81, № 1, <strong>с</strong>тр. 81—84'.—1950. О пред<strong>в</strong>е<strong>с</strong>нян<strong>к</strong><strong>а</strong>х<br />

из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Ур<strong>а</strong>л<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР,<br />

т. 82, № 6, <strong>с</strong>тр. 985—989.—1953<strong>а</strong>. Н<strong>о</strong><strong>в</strong>ые ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

мел<strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> П<strong>о</strong><strong>в</strong><strong>о</strong>лжье, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>не и> З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье.<br />

Д<strong>о</strong><strong>к</strong>л. АН ССОР, т. 89, № 1, <strong>с</strong>тр. 1163—166 —<br />

19636. Н<strong>о</strong><strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители перм<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>о</strong>гр<strong>а</strong>д<strong>а</strong><br />

Protoperlaria. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды, <strong>о</strong>тд.<br />

би<strong>о</strong>л., т. 58, <strong>в</strong>ып. 2, <strong>с</strong>тр. 42—47.—1954. Стр<strong>о</strong>ение г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы<br />

перм<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> Perielytron mirabile G. Zalessky.<br />

Ге<strong>о</strong>л. <strong>с</strong>б. Ль<strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>. те<strong>о</strong>л. <strong>о</strong>б-<strong>в</strong><strong>а</strong>, № 1, <strong>с</strong>тр. 194—<br />

К <strong>а</strong><strong>в</strong><strong>а</strong>ль И. (Kawall, J. Н.). 1876. Organische Einschlusse<br />

im Bergkristall. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды,<br />

т. 51, № 3, <strong>с</strong>тр. 170—173. К<strong>о</strong>ле<strong>с</strong>ницы н<strong>а</strong> Г. Р.,<br />

М <strong>а</strong> р т ы н <strong>о</strong> <strong>в</strong> <strong>а</strong> О. М. 1961. Н<strong>о</strong><strong>в</strong>ый юр<strong>с</strong><strong>к</strong>ий р<strong>о</strong>д Ijapsyche<br />

(Mecoptera, Paratrichoptera) из В<strong>о</strong><strong>с</strong>т<strong>о</strong>чн<strong>о</strong>й Сибири. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

журн., № 4, <strong>с</strong>тр. 162—164. Кузнец<strong>о</strong><strong>в</strong> Н. Н.<br />

1926. О пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении 'Пу<strong>с</strong>тынн<strong>о</strong>й ф<strong>а</strong>уны Тур<strong>к</strong>е<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

Ру<strong>с</strong><strong>с</strong><strong>к</strong>. з<strong>о</strong><strong>о</strong>л. журн., т. 6, <strong>в</strong>ып. 1, <strong>с</strong>тр. 61—82. Кузнец<strong>о</strong><strong>в</strong><br />

Н. Я. 1928. Oligamites martynovi, gen. et. sp. nn. a fossil<br />

Amatid Lepidopteron from the Oligocene beds of Central<br />

Asia. Д<strong>о</strong><strong>к</strong>л. AH OOGP, № 20—21, <strong>с</strong>тр. 431—436.— 1941.<br />

Чешуе<strong>к</strong>рылые янт<strong>а</strong>ря. Изд. АН СССР, <strong>с</strong>тр. 1—104.<br />

Лебеде<strong>в</strong> И. 1<br />

В. I960. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из<br />

юр<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений центр<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>й<strong>о</strong>н<strong>а</strong> Кузб<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>. Из<strong>в</strong>.<br />

Т<strong>о</strong>м<strong>с</strong><strong>к</strong>, п<strong>о</strong>литехи, ин-т<strong>а</strong>, т. 66, <strong>в</strong>ып. 2, <strong>с</strong>тр. 141—143.<br />

Л е п н е <strong>в</strong> <strong>а</strong> С. Г. 1956. М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>о</strong><strong>о</strong>тн<strong>о</strong>шения<br />

п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Psychomyinae, Ecnominae и Polycentropinae<br />

(Trichoptera, Annulipalpia) <strong>в</strong> иреим<strong>а</strong>гин<strong>а</strong>лыных ф<strong>а</strong>з<strong>а</strong>х.<br />

Энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>бюзр., т. 35, № 1, <strong>с</strong>тр. 8—07.<br />

М<strong>а</strong>лы ш е <strong>в</strong> С. И. 1947. Пути и у<strong>с</strong>л<strong>о</strong><strong>в</strong>ия э<strong>в</strong><strong>о</strong>люции<br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылых <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong><strong>о</strong>м <strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении<br />

н<strong>а</strong>ездни<strong>к</strong><strong>о</strong><strong>в</strong> (Hymenoptera). Д<strong>о</strong><strong>к</strong>л. АН СССР, т.<br />

58, № 6, <strong>с</strong>тр. 1207—1209. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong> А. В. (Martynov<br />

А. В.). 1923. О д<strong>в</strong>ух <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных тип<strong>а</strong>х <strong>к</strong>рылье<strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

и их зн<strong>а</strong>чении для <strong>о</strong>бщей <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Тр. I В'Сер<strong>о</strong><strong>с</strong><strong>с</strong>. <strong>с</strong>ъезд<strong>а</strong> з<strong>о</strong><strong>о</strong>л., <strong>а</strong>н<strong>а</strong>т. и ги<strong>с</strong>т<strong>о</strong>л. <strong>в</strong><br />

Петр<strong>о</strong>гр<strong>а</strong>де 15—21. XII 1922 г., <strong>с</strong>тр. «8—89.— 1924<strong>а</strong>.<br />

К п<strong>о</strong>ним<strong>а</strong>нию жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ния и тр<strong>а</strong>хе<strong>а</strong>ции <strong>к</strong>рылье<strong>в</strong> <strong>с</strong>тре<strong>к</strong><strong>о</strong>з<br />

и п<strong>о</strong>ден<strong>о</strong><strong>к</strong>. Ру<strong>с</strong><strong>о</strong><strong>к</strong>. энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б<strong>о</strong>зр., т. 18, <strong>с</strong>тр. 146—174.—•<br />

19246. Пр<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong><strong>а</strong>я энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гия, <strong>в</strong>ып. 5—Ручейни<strong>к</strong>и,<br />

<strong>с</strong>тр. 1—388.—1926<strong>а</strong>. О не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых результ<strong>а</strong>т<strong>а</strong>х изучения<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>с</strong>л<strong>а</strong>нце<strong>в</strong> Тур<strong>к</strong>е<strong>с</strong>т<strong>а</strong>н<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, <strong>с</strong>тр.<br />

105—108.— 19256. Т<strong>о</strong> the knowledge of fossil insects from<br />

Jurassic beds in Turkestan. I. Raphidioptera. Из<strong>в</strong>. Pooc.<br />

AH, № 2, <strong>с</strong>тр. 233—246.—1925B. TO the knowledge of<br />

fossil insects from Jurassic beds in Turkestan. 2. Raphidioptera<br />

(cent.), Orthoptera (s. 1.), Odonata, Neuroptera.<br />

Из<strong>в</strong>. Р<strong>о</strong><strong>с</strong><strong>с</strong>. А<strong>к</strong><strong>а</strong>д. н<strong>а</strong>у<strong>к</strong>, <strong>с</strong>тр. 569 1<br />

—598.— 19Е5 г. Т<strong>о</strong> the knowleage<br />

of fossil insects from Jurassic beds in Turkestan.<br />

Hymenoptera, Mecoptera. Из<strong>в</strong>. АН СССР, <strong>с</strong>тр. 753 1<br />

—<br />

762.—1925д. Ober zwei Grundtypen der Flugel bei den<br />

Insekten und ihre Evolution. Ztschr. Morph. Okol. d. Tiere,<br />

Bid. 4, H. 3, S. 465—601.— 1925e. Об <strong>о</strong>дн<strong>о</strong>й н<strong>о</strong><strong>в</strong><strong>о</strong>й ф<strong>о</strong>рме<br />

Orthopteroidea из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Южн<strong>о</strong>-У<strong>с</strong><strong>с</strong>урий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я. Д<strong>о</strong><strong>к</strong>л. Р<strong>о</strong><strong>с</strong><strong>с</strong>. АН, <strong>с</strong>тр. 41—44.—1925ж.<br />

Об <strong>о</strong>дн<strong>о</strong>м интере<strong>с</strong>н<strong>о</strong>м и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м жу<strong>к</strong>е из юр<strong>с</strong><strong>к</strong>их <strong>с</strong>л<strong>а</strong>нце<strong>в</strong><br />

<strong>с</strong>е<strong>в</strong>ерн<strong>о</strong>г<strong>о</strong>' Тур<strong>к</strong>е<strong>с</strong>т<strong>а</strong>н<strong>а</strong>. Ру<strong>с</strong><strong>с</strong><strong>к</strong>. энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б<strong>о</strong>зр.,<br />

т. 19, <strong>с</strong>тр. 73—78.— 1926<strong>а</strong>. К п<strong>о</strong>зн<strong>а</strong>нию и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

юр<strong>с</strong><strong>к</strong>их <strong>с</strong>л<strong>а</strong>нце<strong>в</strong> Тур<strong>к</strong>е<strong>с</strong>т<strong>а</strong>н<strong>а</strong>. 5. О не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ф<strong>о</strong>рм<strong>а</strong>х<br />

жу<strong>к</strong><strong>о</strong><strong>в</strong> (Coleoptera). Ежег<strong>о</strong>дн. Ру<strong>с</strong><strong>с</strong><strong>к</strong>. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

<strong>о</strong>б-<strong>в</strong><strong>а</strong> (1925), т. 5, ч. 1, <strong>с</strong>тр. 1—38.— 19266. Jurassic fossil<br />

insects from Turkestan. 6. Homoptera and Psocoptera.<br />

Из<strong>в</strong>. АН СССР, <strong>с</strong>тр. 1349—«1366.—1927a. Jurassic fossil<br />

insects from Turkestan. 7. Some Odonata, Neuroptera,<br />

Thysanoptera. Из<strong>в</strong>. АН СССР, <strong>с</strong>тр. 757—768— 19276.<br />

О д<strong>в</strong>ух и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых третичных <strong>с</strong>тре<strong>к</strong><strong>о</strong>з<strong>а</strong>х <strong>с</strong> .К<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>.<br />

Ру<strong>с</strong><strong>с</strong><strong>к</strong>. энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б<strong>о</strong>зр., т. 21, № 1—2, <strong>с</strong>тр. 1—5.—<br />

1927<strong>в</strong>. Jurassic fossil Mecoptera and Paratrichoptera<br />

from Turkestan and Ust j<br />

Balei (Siberia). Из<strong>в</strong>. AH OOGP,<br />

<strong>с</strong>тр. 651—666.— 1927. Ober eine neue Ordnung der fossilen<br />

Insekten, Miomoptera nov. Zool. Anz. Bd. 22, H.<br />

3/4, S. 99—109 — 1928a. A new fossil form of Phasmatodea<br />

from Galkino (Turkestan) and on Mesozoic Phasmids<br />

in general. Ann. and Mag. Nat. History, ser. 10, v. 1, p.<br />

319—328.—19286. Permian fossil insects of North-East<br />

361<br />

http://jurassic.ru/

Europe. Тр. Ге<strong>о</strong>л. Муз. АН СССР, т. 4, <strong>с</strong>тр. 1—418 —<br />

1928<strong>в</strong>. О не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых х<strong>а</strong>р<strong>а</strong><strong>к</strong>терных черт<strong>а</strong>х групп <strong>в</strong>ымир<strong>а</strong>ющих<br />

и их з<strong>а</strong>мещ<strong>а</strong>ющих <strong>в</strong> мире н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Тр. III<br />

В<strong>с</strong>ер<strong>о</strong><strong>с</strong><strong>с</strong>. <strong>с</strong>ъезд<strong>а</strong> з<strong>о</strong><strong>о</strong>л., <strong>а</strong>н<strong>а</strong>т. и ги<strong>с</strong>т<strong>о</strong>л. <strong>в</strong> Ленингр<strong>а</strong>де<br />

14—20/XII 1927 г., <strong>с</strong>тр. 143—'144.—1929<strong>а</strong>. Об и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых третичных <strong>о</strong>тл<strong>о</strong>жений Ашут<strong>а</strong>е<strong>а</strong>, З<strong>а</strong>й<strong>с</strong><strong>а</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

уезд<strong>а</strong>. Тр. Ге<strong>о</strong>л. муз. АН СССР, т. 5, <strong>с</strong>тр.<br />

173—216—19296. Permian Entomofauna of North Rossia<br />

and its relation to that of Kansas. 4 Intern. Congr. of<br />

Ent. Ithaca, 1928, v. 2, p. 595—598.— 1930a. О н<strong>о</strong><strong>в</strong>ых и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых Тихих Г<strong>о</strong>р. Отдел Neoptera (без<br />

Miomoptera). Тр. Ге<strong>о</strong>л. Муз. АН СССР, т. 8, <strong>с</strong>тр. 149—<br />

212.—19306. О д<strong>в</strong>ух н<strong>о</strong><strong>в</strong>ых прям<strong>о</strong><strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из<br />

перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Кунгур<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> уезд<strong>а</strong>. Перм<strong>с</strong><strong>к</strong><strong>о</strong>й губ.<br />

Ежег<strong>о</strong>дн. Ру<strong>с</strong><strong>с</strong><strong>к</strong>. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. <strong>о</strong>б-<strong>в</strong><strong>а</strong>, т. 8, <strong>с</strong>тр. 36—47.—<br />

1930<strong>в</strong>. Permian fossil insects from Tikhije Gory. Order<br />

Miomoptera. Pt. I. The new fam. Atactophlebiidae and its<br />

relations. Из<strong>в</strong>. АН СССР, № 9, <strong>с</strong>тр. 951—975.— 1930г.<br />

Permian fossil insects from Tikhije Gory. Pt. II. The fam.<br />

Lemmatophorklae and Palaeomantidae. Из<strong>в</strong>. АН СССР,<br />

№ 9, <strong>с</strong>тр. 1*116—1134.—1930Д. New Permian insects from<br />

Tikhie Gory Kazan province. I. Palaeoptera. Тр. Ге<strong>о</strong>л.<br />

Муз. А<strong>к</strong><strong>а</strong>д. н<strong>а</strong>у<strong>к</strong> СССР, т. 6, <strong>с</strong>тр. 69—86.—1931<strong>а</strong>. On<br />

some new remarkable Odonata from the Permian of<br />

Archangelsk district. Из<strong>в</strong>. АН СССР, № 1, <strong>с</strong>тр. 141—<br />

147.—19316. О п<strong>о</strong>д<strong>о</strong>тряде Permanisoptera nom. nov.<br />

(Odonata) и ег<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жении. Д<strong>о</strong><strong>к</strong>л. АН СССР, <strong>с</strong>тр.246'—<br />

247.—1931 <strong>в</strong>. К п<strong>о</strong>зн<strong>а</strong>нию юр<strong>с</strong><strong>к</strong>их Palaeontinidae Handl.,<br />

м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия, <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение и <strong>о</strong>пи<strong>с</strong><strong>а</strong>ние н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

р<strong>о</strong>д<strong>а</strong> из У<strong>с</strong>ть-Б<strong>а</strong>лея. Ежег<strong>о</strong>дн. Ру<strong>с</strong><strong>с</strong><strong>к</strong>. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

<strong>о</strong>б-<strong>в</strong><strong>а</strong>, т. 9, (1930), <strong>с</strong>тр. 93—122.— 1931 г. О п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> 'б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Из<strong>в</strong>. ге<strong>о</strong>л.-р<strong>а</strong>з<strong>в</strong>. упр.,<br />

т. 49, № 10, <strong>с</strong>тр. 73—100.—1932<strong>а</strong>. On the wing venation<br />

in the fam. Meganeuridae (Meganisoptera). Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, <strong>с</strong>тр. 42—44.—19326. New Permian Palaeoptera<br />

with the discussion of some problems of theier evolution.<br />

Тр. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР (1991), т. I, <strong>с</strong>тр. 1—<br />

44—1933<strong>а</strong>. К <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong>у <strong>о</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>те п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>н<strong>о</strong><strong>с</strong>ных<br />

<strong>о</strong>тл<strong>о</strong>жений Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН<br />

СССР. № 3, <strong>с</strong>тр. 34—37.—19336. On the Permian family<br />

Archescytinidae (Homoptera) and its relationships. Из<strong>в</strong>.<br />

АН СССР, <strong>с</strong>тр. 883—894.—19336. Permian fossil insects<br />

from the Arkhangelsk district. Pt. 1. The order Mecoptera.<br />

Тр. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР (1932), т. 2, <strong>с</strong>тр. 28—<br />

62.— 1938 г. Перм<strong>с</strong><strong>к</strong>ие и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я. Ч. II. Сетч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые, <strong>в</strong>и<strong>с</strong>л<strong>о</strong><strong>к</strong>рылые и жу<strong>к</strong>и,<br />

<strong>с</strong> прил<strong>о</strong>жением <strong>о</strong>пи<strong>с</strong><strong>а</strong>ния д<strong>в</strong>ух н<strong>о</strong><strong>в</strong>ых жу<strong>к</strong><strong>о</strong><strong>в</strong> из Тихих<br />

Гер. Тр. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР (1932), т. 2, <strong>с</strong>тр.<br />

63—96.—1934. Кл<strong>а</strong><strong>с</strong><strong>с</strong> Insecta (Hexapoda). Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые.<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. К. Циттель. Ч. I. Бе<strong>с</strong>п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чные,<br />

М.—Л., <strong>с</strong>тр. 986—11026.—1935<strong>а</strong>. З<strong>а</strong>мет<strong>к</strong><strong>а</strong> <strong>о</strong>б и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Челябин<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

бур<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Тр. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong><br />

АН СССР, т. 4, <strong>с</strong>тр. 37—48.—19356. О н<strong>а</strong>х<strong>о</strong>ждении<br />

глип<strong>с</strong><strong>о</strong><strong>в</strong> (Thysanoptera) <strong>в</strong> перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жениях. Д<strong>о</strong><strong>к</strong>л.<br />

АН СССР, т. 3, № 7, <strong>с</strong>тр. 333—336.—1935<strong>в</strong>. О не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>их<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из <strong>к</strong><strong>о</strong>льчугин<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong><strong>в</strong>иты Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>.<br />

Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л., № 3, <strong>с</strong>тр. 441—448.—<br />

1935 г. Permian fossil insects from Arkhangelsk district. Pt.<br />

5. Homoptera. Тр. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 4, <strong>с</strong>тр.<br />

1—35.— 1936. О не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>о</strong><strong>в</strong>ых м<strong>а</strong>тери<strong>а</strong>л<strong>а</strong>х члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

жи<strong>в</strong><strong>о</strong>тных из Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Из<strong>в</strong>. АН<br />

СССР, <strong>с</strong>ер. би<strong>о</strong>л., № 6, <strong>с</strong>тр. 1251—1264.—1937<strong>а</strong>. Ли<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ые<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Шур<strong>а</strong>б<strong>а</strong> и Кизил-Кии. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

ин-т<strong>а</strong> АН СССР, т. 7, <strong>в</strong>ып. 1, <strong>с</strong>тр. 5—232.—19876. Перм<strong>с</strong><strong>к</strong>ие<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые К<strong>а</strong>рг<strong>а</strong>лы и их <strong>о</strong>тн<strong>о</strong>шения.<br />

Т(.. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong>, т. 7, <strong>в</strong>ып. 2, <strong>с</strong>тр. 8—91.—1937<strong>в</strong>.<br />

О <strong>к</strong>рыльях термит<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> <strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong><strong>о</strong>м фил<strong>о</strong>гении<br />

эт<strong>о</strong>й и <strong>с</strong><strong>о</strong><strong>с</strong>едних групп н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Сб. <strong>в</strong> че<strong>с</strong>ть Н. В. Н<strong>а</strong><strong>с</strong><strong>о</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>.<br />

Изд. АН СССР, <strong>с</strong>тр. 83—1150.—1988<strong>а</strong>. Ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждения<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых <strong>в</strong> предел<strong>а</strong>х СССР. Тр.<br />

П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong>, т. 7, <strong>в</strong>ып. 3, <strong>с</strong>тр. 6—28.—19386. О н<strong>о</strong><strong>в</strong><strong>о</strong>м<br />

перм<strong>с</strong><strong>к</strong><strong>о</strong>м <strong>о</strong>тряде прям<strong>о</strong><strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых Glosse­<br />

362<br />

http://jurassic.ru/<br />

lytrodea. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л. № 1, <strong>с</strong>тр. 187—<br />

206.— 1938<strong>в</strong>. Перм<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Ч. V. Сем. Euthygrammidae и ег<strong>о</strong> <strong>о</strong>тн<strong>о</strong>шения (<strong>с</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>нием<br />

<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> и <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> из Че<strong>к</strong><strong>а</strong>рды). Тр.<br />

П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 7, <strong>в</strong>ып. 3, <strong>с</strong>тр. 69—80.—<br />

1938г. Очер<strong>к</strong>и ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й и<strong>с</strong>т<strong>о</strong>рии и фил<strong>о</strong>гении <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (Pterygota). Ч. I. Palaeoptera и Neoptera<br />

— Polyneoptera. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong>, т. 7, <strong>в</strong>ып. 4,<br />

<strong>с</strong>тр. 5—149.—1940<strong>а</strong>. Перм<strong>с</strong><strong>к</strong>ие и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

Че<strong>к</strong><strong>а</strong>рды. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong>, т. 11, <strong>в</strong>ып. 1, <strong>с</strong>тр. 5—<br />

62.—19406. Ср<strong>а</strong><strong>в</strong>нительный <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых п<strong>о</strong><br />

р. С<strong>о</strong>яне, у Тихих Г<strong>о</strong>р и близ Че<strong>к</strong><strong>а</strong>рды т<strong>о</strong> р. Сыл<strong>в</strong>е. М<strong>а</strong>тери<strong>а</strong>лы<br />

п<strong>о</strong> ге<strong>о</strong>л<strong>о</strong>гии перм<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>и<strong>с</strong>темы Е<strong>в</strong>р<strong>о</strong>пей<strong>с</strong><strong>к</strong><strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти<br />

СССР. Г<strong>о</strong><strong>с</strong>т<strong>о</strong>итехизи<strong>а</strong>т, <strong>с</strong>тр. 86—89. М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong><br />

О. М.—1939. Miopsyche kaspievi, sp. п., н<strong>о</strong><strong>в</strong>ый <strong>в</strong>ид .ручейни<strong>к</strong><strong>а</strong><br />

из ми<strong>о</strong>цен<strong>о</strong><strong>в</strong>ых <strong>о</strong>тл<strong>о</strong>жений Ордж<strong>о</strong>ни<strong>к</strong>идзе<strong>в</strong>'<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я. Тр. В<strong>о</strong>р<strong>о</strong>шил. Г<strong>о</strong><strong>с</strong>. пед. ин-т<strong>а</strong>, т. I, <strong>с</strong>тр. 91—93.—<br />

1941. Перм<strong>с</strong><strong>к</strong>ие Mecoptera <strong>с</strong> р. С<strong>о</strong>яны. Рефер<strong>а</strong>ты р<strong>а</strong>б<strong>о</strong>т<br />

У'чр. <strong>о</strong>тд. би<strong>о</strong>л. н<strong>а</strong>у<strong>к</strong> АН СССР з<strong>а</strong> 1940 г., <strong>с</strong>тр. 266.—<br />

1942. Перм<strong>с</strong><strong>к</strong>ие Mecoptera из Че<strong>к</strong><strong>а</strong>рды и К<strong>а</strong>рг<strong>а</strong>лы. Из<strong>в</strong>.<br />

АН СССР, <strong>с</strong>ер. би<strong>о</strong>л., № 1—2, <strong>с</strong>тр. 133—149.—1943.<br />

Glosselytrodea из юр<strong>с</strong><strong>к</strong>их <strong>с</strong>л<strong>а</strong>нце<strong>в</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong><br />

ме<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ждения С<strong>о</strong>гюты. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 39, № 7,<br />

<strong>с</strong>тр. 312—313.—1945. Юр<strong>с</strong><strong>к</strong>ие Mecoptera из углен<strong>о</strong><strong>с</strong>ных<br />

<strong>с</strong>л<strong>а</strong>нце<strong>в</strong> С<strong>о</strong>гюты. Реф. р<strong>а</strong>б<strong>о</strong>т учр. <strong>о</strong>тд. би<strong>о</strong>л. н<strong>а</strong>у<strong>к</strong><br />

АН СССР з<strong>а</strong> 1941—1943 гг., <strong>с</strong>тр. 225.—.1947<strong>а</strong>. Д<strong>в</strong>е н<strong>о</strong><strong>в</strong>ые<br />

<strong>в</strong>ерблюд<strong>к</strong>и из юр<strong>с</strong><strong>к</strong>и<strong>к</strong> <strong>с</strong>л<strong>а</strong>нце<strong>в</strong> К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у. Д<strong>о</strong><strong>к</strong>л.<br />

ЛН СССР, н<strong>о</strong><strong>в</strong>. <strong>с</strong>ер., т. 56, № 6, <strong>с</strong>тр. 636-^637.-19476.<br />

О прир<strong>о</strong>де труб<strong>о</strong><strong>к</strong> Pectinariopsis Andr. (Trichoptera, п<strong>о</strong>п<br />

Polychaeta). Энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б<strong>о</strong>зр., т. 29, № 3, <strong>с</strong>тр. 152—<br />

153—1947<strong>в</strong>. Kalligrarnmidae (<strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые) из юр<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>л<strong>а</strong>нце<strong>в</strong> К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 58, № 9,<br />

<strong>с</strong>тр. 2055—2058.—1948<strong>а</strong>. Д<strong>в</strong><strong>а</strong> н<strong>о</strong><strong>в</strong>ых <strong>в</strong>ид<strong>а</strong> перм<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

из Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 60,<br />

Ki 1, <strong>с</strong>тр. 113—114.—19486. Инди<strong>в</strong>иду<strong>а</strong>льн<strong>а</strong>я изменчи<strong>в</strong><strong>о</strong><strong>с</strong>ть<br />

<strong>к</strong>рылье<strong>в</strong> Mecoptera. Из<strong>в</strong>. АН ССОР, № 2,<br />

<strong>с</strong>тр. 193—198.—1948<strong>в</strong>. М<strong>а</strong>тери<strong>а</strong>лы п<strong>о</strong> э<strong>в</strong><strong>о</strong>люции Mecoptera.<br />

Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong>, т. 14, <strong>в</strong>ып. 1, <strong>с</strong>тр. 1—76.—<br />

1949<strong>а</strong>. Мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые СССР, Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

ин-т<strong>а</strong>, т. 20, <strong>с</strong>тр. 150—170.—19496. Пер<strong>в</strong><strong>а</strong>я н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong><br />

юр<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> .<strong>в</strong> Кузнец<strong>к</strong><strong>о</strong>м б<strong>а</strong><strong>с</strong><strong>с</strong>ейне. Д<strong>о</strong><strong>к</strong>л. АН<br />

СССР, т. 66, № 5, <strong>с</strong>тр. 928—924.^1949». Ф<strong>а</strong><strong>с</strong>ет<strong>о</strong>чные <strong>о</strong>рг<strong>а</strong>ны<br />

н<strong>а</strong> <strong>к</strong>рыльях <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц (Mecoptera). Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>.<br />

<strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды, <strong>о</strong>тд. ге<strong>о</strong>л., т. 24, <strong>в</strong>ып. 4, <strong>с</strong>тр. 93—<br />

95.—1951<strong>а</strong>. Д<strong>в</strong><strong>а</strong> н<strong>о</strong><strong>в</strong>ых нижнелей<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из<br />

Кизил-Кии. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 78, № 5, <strong>с</strong>тр. 1009—<br />

1011.—19516. Ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждение перм<strong>с</strong><strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых у<br />

дере<strong>в</strong>ни С<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>в</strong> Кузнец<strong>к</strong><strong>о</strong>м б<strong>а</strong><strong>с</strong><strong>с</strong>ейне. Д<strong>о</strong><strong>к</strong>л. АН<br />

СССР, т. 79, № 1, <strong>с</strong>тр. 149—151.—1952<strong>а</strong>. Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые перм<strong>с</strong><strong>к</strong>ие<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из <strong>к</strong>узб<strong>а</strong><strong>с</strong><strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>.<br />

<strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды, <strong>о</strong>тд. ге<strong>о</strong>л., т. 27, <strong>в</strong>ып. 3, <strong>с</strong>тр. 92.—<br />

19626. Отряд Glosselytrodea из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Кемер<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР,<br />

т. 40, <strong>с</strong>тр. 187—196.—1952<strong>в</strong>. Перм<strong>с</strong><strong>к</strong>ие <strong>с</strong>етч<strong>а</strong>т<strong>о</strong><strong>к</strong>рылые<br />

СССР. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 40, <strong>с</strong>тр. 197—<br />

237.—1963<strong>а</strong>. Н<strong>а</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ление для <strong>с</strong>б<strong>о</strong>р<strong>о</strong><strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Изд. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, <strong>с</strong>тр. 1—14—<br />

195Э6. С<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> ф<strong>а</strong>уны <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нниц из ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждений<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых безуг<strong>о</strong>льн<strong>о</strong>й или <strong>к</strong>узнец<strong>к</strong><strong>о</strong>й <strong>с</strong><strong>в</strong>иты углен<strong>о</strong><strong>с</strong>н<strong>о</strong>й<br />

т<strong>о</strong>лщи Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 89,<br />

№ 4, <strong>с</strong>тр. 727—729.— 1954. Сетч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е из<br />

мел<strong>о</strong><strong>в</strong>ых <strong>о</strong>тл<strong>о</strong>жений Сибири. Д<strong>о</strong><strong>к</strong>л. АН СССР. т. 94,<br />

№ 6, <strong>с</strong>тр. 1167—1169.—1958. Н<strong>о</strong><strong>в</strong>ые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые из перм<strong>с</strong><strong>к</strong>их<br />

и мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений СССР. М<strong>а</strong>тери<strong>а</strong>лы <strong>к</strong><br />

«О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>м п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», <strong>в</strong>ып. 2, <strong>с</strong>тр. 69—94.— 1959.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>в</strong>з<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шения н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых ме<strong>к</strong><strong>о</strong>птер<strong>о</strong>идн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong>. Тр. Ин-т<strong>а</strong> м<strong>о</strong>рф<strong>о</strong>л. жи<strong>в</strong><strong>о</strong>тных АН<br />

ССОР, <strong>в</strong>ып. 27, <strong>с</strong>тр. 221—230.—1961. С<strong>о</strong><strong>в</strong>ременные и <strong>в</strong>ымершие<br />

<strong>в</strong>ерблюд<strong>к</strong>и. (Insecta, Raphidioptera). П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

журн. №3, <strong>с</strong>тр. 73—83.—1961<strong>а</strong>. См. Р<strong>о</strong>денд<strong>о</strong>рф<br />

Б. Б., Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong> Е. Э., М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong><br />

О. М., Ш<strong>а</strong>р<strong>о</strong><strong>в</strong> А. Г., М <strong>а</strong> х<strong>о</strong>ти н А. А. 1952. Вз<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шения<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных групп прыг<strong>а</strong>ющих прям<strong>о</strong><strong>к</strong>рылых и

м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия их яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong>. Энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б<strong>о</strong>зр., т. 32,<br />

<strong>с</strong>тр. 126—436.—1963. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>в</strong>з<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шения<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных групп прыг<strong>а</strong>ющих прям<strong>о</strong><strong>к</strong>рылых и м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия<br />

их яйце<strong>к</strong>л<strong>а</strong>д<strong>а</strong>. Тр. Ин-т<strong>а</strong> м<strong>о</strong>рф<strong>о</strong>л. жи<strong>в</strong><strong>о</strong>тных.<br />

АН СССР, <strong>в</strong>ып. 8, <strong>с</strong>тр. 5—61. Меш<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong> Н. П. 1961.<br />

О личин<strong>к</strong><strong>а</strong>х Ephemeropsis trisetalis Eichwald (Insecta).<br />

П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. журн., № 4, <strong>с</strong>тр. 164—168.<br />

П<strong>а</strong>р<strong>а</strong>м<strong>о</strong>н<strong>о</strong><strong>в</strong> С. Я. 1936. Д<strong>о</strong>елщи н<strong>а</strong>д ент<strong>о</strong>м<strong>о</strong>ф<strong>а</strong>ун<strong>о</strong>ю<br />

янт<strong>а</strong>р1<strong>в</strong>.— Гн<strong>с</strong>тит. з<strong>о</strong><strong>о</strong>л. т<strong>а</strong> би<strong>о</strong>л. АН УССР, № 18,<br />

<strong>с</strong>тр. 53—64.— 1939. Kritische Obersicht der gegenwartigen<br />

und fossilen Bombyliiden-Gattungen der ganzen<br />

Welt. IHCTHT. з<strong>о</strong><strong>о</strong>л. т<strong>а</strong> би<strong>о</strong>л. АН УССР, № 23, <strong>с</strong>тр. 23— •<br />

50. П<strong>о</strong>н<strong>о</strong>м<strong>а</strong>рен<strong>к</strong><strong>о</strong> А. Г. 1961. О <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

п<strong>о</strong>л<strong>о</strong>жении Coptoclava longipoda Ping. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л.<br />

журн., № 3, <strong>с</strong>тр. 67—72. П<strong>о</strong>п<strong>о</strong><strong>в</strong>. В. В. 1933. Two new<br />

fossil ants from Caucasus (Hymenoptera, Formicidae).<br />

Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. и.н-т<strong>а</strong> АН СССР, т. 2, <strong>с</strong>тр. 17—21.<br />

Рей<strong>с</strong> О. М. 1910. Ф<strong>а</strong>ун<strong>а</strong> рыбных <strong>с</strong>л<strong>а</strong>нце<strong>в</strong> З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти. Ге<strong>о</strong>л.-и<strong>с</strong><strong>с</strong>лед. и р<strong>а</strong>з<strong>в</strong>ед. р<strong>а</strong>б<strong>о</strong>ты п<strong>о</strong> линии<br />

Сиб. ж. д. СПб., <strong>в</strong>ып. 29, <strong>с</strong>тр. 1—68. Рихтер<br />

А. А. 4935. О жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>а</strong>д<strong>к</strong>рылий жу<strong>к</strong><strong>о</strong><strong>в</strong>. Ру<strong>с</strong><strong>с</strong><strong>к</strong>.<br />

энт. <strong>о</strong>б<strong>о</strong>зр., т. 26, № 1—4, <strong>с</strong>тр. 25-^58.-1947. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые<br />

зл<strong>а</strong>т<strong>к</strong>и из бин<strong>а</strong>г<strong>а</strong>дин<strong>с</strong><strong>к</strong>их мир<strong>о</strong><strong>в</strong>ых <strong>с</strong>л<strong>о</strong>е<strong>в</strong>. Д<strong>о</strong><strong>к</strong>л.<br />

АН Арм. ССР, т. 4, <strong>в</strong>ып. 5, <strong>с</strong>тр. 147—150. Р<strong>о</strong>денд<strong>о</strong>рф<br />

Б. Б. (Rohdendorf В. В.) 1938. Д<strong>в</strong>у<strong>к</strong>рылые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

мез<strong>о</strong>з<strong>о</strong>я К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР,<br />

т. 7, <strong>в</strong>ып. 3, <strong>с</strong>тр. 29—67.—1939<strong>а</strong>. О ми<strong>о</strong>цен<strong>о</strong><strong>в</strong><strong>о</strong>й ф<strong>а</strong>уне н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

<strong>о</strong><strong>к</strong>ре<strong>с</strong>тн<strong>о</strong><strong>с</strong>тей г. В<strong>о</strong>р<strong>о</strong>шил<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>. «Прир<strong>о</strong>д<strong>а</strong>», № 2,<br />

<strong>с</strong>тр. 85—88.— 19396. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель <strong>о</strong>тряд<strong>а</strong> Protelytropteг<strong>а</strong><br />

из Ур<strong>а</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>й перми. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 23, № 5,<br />

<strong>с</strong>тр. 505—507.— 1940<strong>а</strong>. Н<strong>а</strong>х<strong>о</strong>ждение <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Scytohymenidae <strong>в</strong> Ур<strong>а</strong>ль<strong>с</strong><strong>к</strong><strong>о</strong>й перми. Д<strong>о</strong><strong>к</strong>л. АН<br />

СССР, т. 26, № 1, <strong>с</strong>тр. 106—107.— 19406. Стр<strong>о</strong>ение тел<strong>а</strong><br />

Archodonata и п<strong>о</strong>л<strong>о</strong>жение эт<strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>а</strong> <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме Palaeoptera.<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 26, № 1, Стр. 108—ПО.—1943.<br />

Об э<strong>в</strong><strong>о</strong>люции п<strong>о</strong>лет<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 40,<br />

№ 4, <strong>с</strong>тр. 187—189.—1944. Н<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> жу<strong>к</strong><strong>о</strong><strong>в</strong> из перми<br />

Ур<strong>а</strong>л<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 44, № 6, <strong>с</strong>тр. 277—279,—<br />

1946. Э<strong>в</strong><strong>о</strong>люция <strong>к</strong>рыл<strong>а</strong> и фил<strong>о</strong>генез длинн<strong>о</strong>у<strong>с</strong>ых д<strong>в</strong>у<strong>к</strong>рылых<br />

Oligoneura. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 13,<br />

<strong>в</strong>ып. 2, <strong>с</strong>тр. 5—108.—1947<strong>а</strong>. Об и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых из<br />

В<strong>о</strong>р<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> уг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong> б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 57,<br />

№ 4, <strong>с</strong>тр. 391—393.—19476. Ф<strong>а</strong>ун<strong>а</strong> д<strong>в</strong>у<strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

К<strong>а</strong>р<strong>а</strong>т<strong>а</strong>у и зн<strong>а</strong>чение ее для п<strong>о</strong>ним<strong>а</strong>ния э<strong>в</strong><strong>о</strong>люции<br />

<strong>о</strong>тряд<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 55, № 8, <strong>с</strong>тр. 757—760.—<br />

1947<strong>в</strong>. О ме<strong>с</strong>т<strong>о</strong>н<strong>а</strong>х<strong>о</strong>ждении лейя<strong>с</strong><strong>о</strong><strong>в</strong>ых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых С<strong>о</strong>гюты.<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 57, № 6, <strong>с</strong>тр. 607—608.—<br />

1948. Жил<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>рылье<strong>в</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и ег<strong>о</strong> зн<strong>а</strong>чение<br />

для <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер. би<strong>о</strong>л., № 2,<br />

<strong>с</strong>тр. 199—203.— 1949. Э<strong>в</strong><strong>о</strong>люция и <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция лет<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong><br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН<br />

СССР, т. 16, <strong>с</strong>тр. 1—176.— 1950<strong>а</strong>. Об изучении <strong>к</strong><strong>о</strong>н<strong>к</strong>ретных<br />

путей э<strong>в</strong><strong>о</strong>люции н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Из<strong>в</strong>. АН СССР, <strong>с</strong>ер.<br />

би<strong>о</strong>л., № 1, <strong>с</strong>тр. 78—97.— 19506. Пр<strong>о</strong>гре<strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие<br />

э<strong>в</strong><strong>о</strong>люции. Ве<strong>с</strong>тн. АН СССР, № 6, <strong>с</strong>тр. 53—58.—1951. Орг<strong>а</strong>ны<br />

д<strong>в</strong>ижения д<strong>в</strong>у<strong>к</strong>рылых н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и их пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение.<br />

Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 35, <strong>с</strong>тр. 1—180.—<br />

1952. Р<strong>а</strong>з<strong>в</strong>итие и зн<strong>а</strong>чение п<strong>а</strong>ле<strong>о</strong>энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их р<strong>а</strong>б<strong>о</strong>т <strong>в</strong><br />

Кузнец<strong>к</strong><strong>о</strong>м б<strong>а</strong><strong>с</strong><strong>с</strong>ейне. Бюлл. М<strong>о</strong><strong>с</strong><strong>к</strong>. <strong>о</strong>б-<strong>в</strong><strong>а</strong> и<strong>с</strong>пыт. прир<strong>о</strong>ды,<br />

<strong>о</strong>тд. ге<strong>о</strong>л., т. 27, <strong>в</strong>ып. 3, <strong>с</strong>тр. 1.— 1954. Р<strong>а</strong>з<strong>в</strong>итие п<strong>а</strong>ле<strong>о</strong>энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний <strong>в</strong> Кузнец<strong>к</strong><strong>о</strong>м б<strong>а</strong><strong>с</strong><strong>с</strong>ейне. Тр.<br />

Т<strong>о</strong>м<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> г<strong>о</strong><strong>с</strong>. уни<strong>в</strong>ер<strong>с</strong>итет<strong>а</strong>, т. 132, <strong>с</strong>тр. 126—128.— 1956.<br />

П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Южн<strong>о</strong>й Сибири. Энт<strong>о</strong>м<strong>о</strong>л.<br />

<strong>о</strong>б<strong>о</strong>зр., т. 35, № 3, <strong>с</strong>тр. 611—619.— 1957. П<strong>а</strong>ле<strong>о</strong>энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>в</strong> СССР. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong>,<br />

т. 66, <strong>с</strong>тр. 1—102.— 1958<strong>а</strong>. Les. Insectes Paleozoiques du<br />

Sud de la Siberie. Proc. Tenth Int. Congress entomol,<br />

vol. 1, pp. 853—859.—1958b. Paleontologicky vyzkum v<br />

SSSR. Vesmir, 37, 10, pp. 341—343.— 1959. В<strong>о</strong>пр<strong>о</strong><strong>с</strong>ы п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. журн<strong>а</strong>л, № 3,<br />

<strong>с</strong>тр. 15—26.— 1961<strong>а</strong>. О<strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>о</strong>нт<strong>о</strong>генез<strong>а</strong> и их зн<strong>а</strong>чение<br />

<strong>в</strong> э<strong>в</strong><strong>о</strong>люции н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. The Onthogeny of Insects.<br />

Acta symposii de evolutione insectorum, Praha, 1959,<br />

pp. 56—60.—19616. П<strong>а</strong>ле<strong>о</strong>энт<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>в</strong> СССР. С<strong>о</strong>р<strong>о</strong><strong>к</strong> лет <strong>с</strong><strong>о</strong><strong>в</strong>ет<strong>с</strong><strong>к</strong><strong>о</strong>й п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Тр. Чет<strong>в</strong>ерт<strong>о</strong>й<br />

<strong>с</strong>е<strong>с</strong><strong>с</strong>ии <strong>в</strong><strong>с</strong>е<strong>с</strong>. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. <strong>о</strong>б-<strong>в</strong><strong>а</strong>, <strong>с</strong>тр. 23—30.— 1961<strong>в</strong>.<br />

Н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>я южн<strong>о</strong>й Сибири и их зн<strong>а</strong>чение для<br />

п<strong>а</strong>ле<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фии и <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фии. Congreso Geologico International.<br />

ХХ<strong>а</strong> Sesion. Section VII — Paleontologia, Taxonomia<br />

у Evolution. Mexico, D. F., pags 37—54.— 1961r.<br />

Дре<strong>в</strong>нейшие инфр<strong>а</strong><strong>о</strong>тряды д<strong>в</strong>у<strong>к</strong>рылых из три<strong>а</strong><strong>с</strong><strong>а</strong> Средней<br />

Азии. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. журн., № 3, <strong>с</strong>тр. 90—100.— 1961 д. Опи<strong>с</strong><strong>а</strong>ние<br />

пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>т<strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> из де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений<br />

Тим<strong>а</strong>н<strong>а</strong> (Insecta, Pterygota). Энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>бз<strong>о</strong>р., т. 40,<br />

'№ 3, <strong>с</strong>тр. 485—489.— 1961е. Neue Angeben fiber das System<br />

der Dipteren. XI. Intern. Kongrefi f. Entomol, Verhandlungen,<br />

В. I, SS. 153—158.— 1961ж. Die Palaoentomologie<br />

in der UdSSR. Т<strong>а</strong>м же, <strong>с</strong>тр. 313—318.— 1962. Тип<br />

Arthropoda. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие. Chelicerata, Picnogonides, Tracheata<br />

В. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гия бе<strong>с</strong>п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных. П<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией<br />

Ю. А. Орл<strong>о</strong><strong>в</strong><strong>а</strong>, <strong>с</strong>тр. 266, 306—328. Р<strong>о</strong>денд<strong>о</strong>рф Б. Б.,<br />

Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong> Е. Э., М<strong>а</strong>ртын<strong>о</strong><strong>в</strong><strong>а</strong> О. М.,<br />

Ш<strong>а</strong>р<strong>о</strong><strong>в</strong> А. Г. 1961. П<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Тр. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 85,<br />

<strong>с</strong>тр. 1—705.<br />

Тер-Мин <strong>а</strong> <strong>с</strong> ян М. Э. 1947. Н<strong>о</strong><strong>в</strong>ый и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емый<br />

<strong>в</strong>ид р<strong>о</strong>д<strong>а</strong> Rhynohites из бин<strong>а</strong>г<strong>а</strong>дин<strong>с</strong><strong>к</strong>их Кир<strong>о</strong><strong>в</strong>ых <strong>с</strong>л<strong>о</strong>е<strong>в</strong>.<br />

Д<strong>о</strong><strong>к</strong>л. АН Арм. ССР, т. 7, № 5, <strong>с</strong>тр. 227^229.<br />

Черн<strong>о</strong><strong>в</strong><strong>а</strong> О. А. 1961. О <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>м п<strong>о</strong>л<strong>о</strong>жении<br />

и ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>м <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>те п<strong>о</strong>ден<strong>о</strong><strong>к</strong> р<strong>о</strong>д<strong>а</strong> Ephemeropsis<br />

Eichwald (Ephemeroptera, Hexagenitidae). Энт<strong>о</strong>м<strong>о</strong>л. <strong>о</strong>б<strong>о</strong>зрен.,<br />

т. 40, № 4, <strong>с</strong>тр. 858—869.— 1962. Личин<strong>к</strong>и п<strong>о</strong>ден<strong>к</strong>и<br />

из не<strong>о</strong>ген<strong>а</strong> З<strong>а</strong>п<strong>а</strong>дн<strong>о</strong>й Сибири (Ephemeroptera, Heptageniidae).<br />

З<strong>о</strong><strong>о</strong>л<strong>о</strong>г, журн., т. 41, № 6, <strong>с</strong>тр. 943—945.<br />

Ш<strong>а</strong>п<strong>о</strong>шни<strong>к</strong><strong>о</strong><strong>в</strong> Г. X. 1953. Жи<strong>в</strong><strong>о</strong>тный мир СССР,<br />

т. IV, ч. 8, <strong>с</strong>тр. 505—521. Ш <strong>а</strong> р <strong>о</strong> <strong>в</strong> А. Г. 1948. Три<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ые<br />

Thysanura из Приур<strong>а</strong>лья. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 61, № 3,<br />

<strong>с</strong>тр. 517—619.—1963. Пер<strong>в</strong><strong>а</strong>я н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong> перм<strong>с</strong><strong>к</strong><strong>о</strong>й личин<strong>к</strong>и<br />

<strong>в</strong>и<strong>с</strong>л<strong>о</strong><strong>к</strong>рыл<strong>о</strong>г<strong>о</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>г<strong>о</strong> (Megaloptera) из К<strong>а</strong>рг<strong>а</strong>лы.<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 89, № 4, <strong>с</strong>тр. 731—732.—1957<strong>а</strong>.<br />

Нимфы ми<strong>о</strong>м<strong>о</strong>птер из перм<strong>с</strong><strong>к</strong>их <strong>о</strong>тл<strong>о</strong>жений Кузнец<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<strong>а</strong>. Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 142, № 6, <strong>с</strong>тр. 1106—<br />

1108.—19676. С<strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зные л<strong>а</strong>лез<strong>о</strong>й<strong>с</strong><strong>к</strong>ие бе<strong>с</strong><strong>к</strong>рылые н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мые<br />

н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>а</strong> Monura (Insecta, Apterygota).<br />

Д<strong>о</strong><strong>к</strong>л. АН СССР, т. 115, № 4, <strong>с</strong>тр. 795—798.—1967<strong>в</strong>.<br />

Пер<strong>в</strong><strong>а</strong>я н<strong>а</strong>х<strong>о</strong>д<strong>к</strong><strong>а</strong> мел<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> ж<strong>а</strong>л<strong>о</strong>н<strong>о</strong><strong>с</strong>н<strong>о</strong>г<strong>о</strong> переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong><strong>к</strong>рыл<strong>о</strong>г<strong>о</strong><br />

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372<br />

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ПОДТИП CHELICEROPHORA<br />

ХЕЛИЦЕРОНОСНЫЕ ЧЛЕНИСТОНОГИЕ<br />

http://jurassic.ru/

http://jurassic.ru/

ПОДТИП CHELICEROPHORA.<br />

ХЕЛИЦЕРОНОСНЫЕ ЧЛЕНИСТОНОГИЕ<br />

(В. Б. Дубинин])<br />

Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующие<strong>с</strong>я <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>г<strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> <strong>о</strong>тдел<strong>а</strong>, реду<strong>к</strong>цией<br />

тергит<strong>а</strong> переднег<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> и ег<strong>о</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

— <strong>а</strong>нтеннул. Вт<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

пре<strong>в</strong>р<strong>а</strong>тил<strong>а</strong><strong>с</strong>ь <strong>в</strong> х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные <strong>о</strong>рг<strong>а</strong>ны —<br />

хелицеры. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти III—VI <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

приближены <strong>к</strong> р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>му <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тию, и их <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ные<br />

члени<strong>к</strong>и пер<strong>в</strong>ичн<strong>о</strong> игр<strong>а</strong>ют р<strong>о</strong>ль р<strong>о</strong>т<strong>о</strong>­<br />

<strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> или гн<strong>а</strong>т<strong>о</strong>б<strong>а</strong>з. Передние <strong>с</strong>егменты<br />

тел<strong>а</strong> (I—VI), <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong><strong>с</strong><strong>о</strong>бый<br />

<strong>о</strong>тдел тел<strong>а</strong>, или т<strong>а</strong>гму (пр<strong>о</strong><strong>с</strong><strong>о</strong>му), не<strong>с</strong>ущую<br />

х<strong>о</strong>дильные н<strong>о</strong>ги.<br />

К эт<strong>о</strong>му п<strong>о</strong>дтипу члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я<br />

д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: Chelicerata или хелицер<strong>о</strong><strong>в</strong>ые, и<br />

Pycnogonides, или м<strong>о</strong>р<strong>с</strong><strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и.<br />

http://jurassic.ru/

НАДКЛАСС CHELICERATA. ХЕЛИЦЕРОВЫЕ<br />

ОБЩАЯ ХАРАКТЕРИСТИКА ХЕЛИЦЕРОВЫХ<br />

( В. Б. Дубинин )<br />

В э<strong>в</strong><strong>о</strong>люции хелицер<strong>о</strong><strong>в</strong>ых члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

<strong>в</strong><strong>а</strong>жным м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>им пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>о</strong>м был<strong>о</strong><br />

изгиб<strong>а</strong>ние пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>; эт<strong>о</strong> при<strong>в</strong>ел<strong>о</strong> <strong>к</strong><br />

перемещению г<strong>о</strong>л<strong>о</strong><strong>в</strong>ных <strong>с</strong>егмент<strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>с</strong>пинную<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну и реду<strong>к</strong>ции передней п<strong>а</strong>ры г<strong>о</strong>л<strong>о</strong><strong>в</strong>ных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, именн<strong>о</strong> <strong>а</strong>нтеннул. В<strong>с</strong>е хелицер<strong>о</strong><strong>в</strong>ые,<br />

<strong>в</strong> <strong>о</strong>тличие <strong>о</strong>т других члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием у<strong>с</strong>и<strong>к</strong><strong>о</strong><strong>в</strong> (<strong>а</strong>нтенн и <strong>а</strong>нтеннул)<br />

и н<strong>а</strong>личием у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых <strong>о</strong><strong>с</strong><strong>о</strong>бей ше<strong>с</strong>ти<br />

п<strong>а</strong>р х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итых <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

д<strong>в</strong>е передние п<strong>а</strong>ры (хелицеры и педип<strong>а</strong>льпы)<br />

игр<strong>а</strong>ют р<strong>о</strong>ль х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельных, р<strong>а</strong>змельч<strong>а</strong>ющих<br />

пищу или чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>, <strong>а</strong> четыре<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льные <strong>в</strong>ып<strong>о</strong>лняют фун<strong>к</strong>цию перед<strong>в</strong>ижения<br />

(х<strong>о</strong>дильные н<strong>о</strong>ги). К<strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, тел<strong>о</strong> хелицер<strong>о</strong><strong>в</strong>ых<br />

р<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь (пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>)<br />

и брюш<strong>к</strong><strong>о</strong> (<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>). В <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди<br />

<strong>в</strong>х<strong>о</strong>дят <strong>а</strong><strong>к</strong>р<strong>о</strong>н (<strong>а</strong><strong>с</strong>г<strong>о</strong>п), <strong>о</strong>т<strong>в</strong>еч<strong>а</strong>ющий г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти (prostomium) <strong>а</strong>ннелид, и ше<strong>с</strong>ть тул<strong>о</strong><strong>в</strong>ищных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующих ше<strong>с</strong>ти п<strong>а</strong>р<strong>а</strong>м<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Четыре передних<br />

<strong>с</strong>егмент<strong>а</strong> л<strong>а</strong>р<strong>в</strong><strong>а</strong>льные; <strong>о</strong>ни <strong>о</strong>т<strong>в</strong>еч<strong>а</strong>ют<br />

четырем <strong>с</strong>егмент<strong>а</strong>м пр<strong>о</strong>т<strong>а</strong><strong>с</strong>пи<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong>й личин<strong>к</strong>и<br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>, <strong>о</strong>бр<strong>а</strong>зующих <strong>в</strong> д<strong>а</strong>льнейшем г<strong>о</strong>л<strong>о</strong><strong>в</strong>у<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лых трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>. Г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гия <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

и <strong>с</strong>егмент<strong>а</strong>ции тел<strong>а</strong> п<strong>о</strong>нятны из д<strong>а</strong>нных<br />

т<strong>а</strong>бл. 1 (<strong>с</strong>тр. 377).<br />

Стр<strong>о</strong>ение тел<strong>а</strong> и р<strong>а</strong>з<strong>в</strong>итие. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь,<br />

или пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> хелицер<strong>о</strong><strong>в</strong>ых<br />

(Merostomata, Arachnida, б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> Scorpionomorpha)<br />

п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> щит<strong>о</strong>м, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из <strong>с</strong>ли<strong>в</strong>ших<strong>с</strong>я тергит<strong>о</strong><strong>в</strong> ше<strong>с</strong>ти передних <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й щит бы<strong>в</strong><strong>а</strong>ет п<strong>о</strong>др<strong>а</strong>зделен<br />

н<strong>а</strong> б<strong>о</strong>лее <strong>к</strong>рупную переднюю ч<strong>а</strong><strong>с</strong>ть —-<br />

378<br />

http://jurassic.ru/<br />

пр<strong>о</strong>пельтидий, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичный г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>му щиту<br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>, и д<strong>в</strong>е леж<strong>а</strong>щие п<strong>о</strong>з<strong>а</strong>ди нег<strong>о</strong> ч<strong>а</strong><strong>с</strong>ти<br />

— мез<strong>о</strong>- и мет<strong>а</strong>пельтидий (Palpigradi, Solifugae<br />

и др.). У <strong>в</strong><strong>с</strong>ех <strong>к</strong>лещей (Acaromorpha)<br />

н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я <strong>с</strong>лияние <strong>с</strong>егмент<strong>о</strong><strong>в</strong> хелицер и педип<strong>а</strong>льп<br />

<strong>в</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленную т<strong>а</strong>гму — гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му,<br />

или г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>к</strong>у, х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>: <strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, <strong>с</strong>пинн<strong>о</strong>й,<br />

или пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льный, щит <strong>в</strong><strong>с</strong>ех <strong>к</strong>лещей не<br />

г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичен <strong>с</strong>пинн<strong>о</strong>му щиту других хелицер<strong>о</strong><strong>в</strong>ых.<br />

Сегмент<strong>а</strong>рный <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы (брюш<strong>к</strong><strong>а</strong>)<br />

н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лный у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> (Scorpionida)<br />

и э<strong>в</strong>риптер<strong>о</strong>ид<strong>о</strong><strong>в</strong> (Eurypterida) (12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

и <strong>а</strong>н<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть, или тель<strong>с</strong><strong>о</strong>н). Кр<strong>о</strong>ме<br />

т<strong>о</strong>г<strong>о</strong>, для мн<strong>о</strong>гих Merostomata и Scorpionomorpha<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделение <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

н<strong>а</strong> д<strong>в</strong>е <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленные ч<strong>а</strong><strong>с</strong>ти — шир<strong>о</strong><strong>к</strong>ую мез<strong>о</strong><strong>с</strong><strong>о</strong>му<br />

(переднебрюшье) и уз<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му<br />

(з<strong>а</strong>днебрюшье) • Сегменты мет<strong>а</strong><strong>с</strong><strong>о</strong>мы у Merostomata<br />

ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м и <strong>о</strong>бр<strong>а</strong>зуют<br />

длинную х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ую иглу, у Eurypterida<br />

u Scorpionida мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>е<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> (пять), <strong>а</strong> у<br />

других <strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Scorpionomorpha п<strong>о</strong>д<strong>в</strong>ерг<strong>а</strong>ют<strong>с</strong>я<br />

реду<strong>к</strong>ции. У пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

Solifugomorpha и Arachnida мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong><br />

пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Среди <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

хелицер<strong>о</strong><strong>в</strong>ых <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Arachnida и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Acaromorpha н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я д<strong>а</strong>льнейш<strong>а</strong>я<br />

реду<strong>к</strong>ция з<strong>а</strong>дних <strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong>, при<strong>в</strong><strong>о</strong>дящ<strong>а</strong>я<br />

<strong>к</strong> п<strong>о</strong>лн<strong>о</strong>му и<strong>с</strong>чезн<strong>о</strong><strong>в</strong>ению ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мет<strong>а</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>а</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong><strong>а</strong>р<strong>к</strong><strong>о</strong>пт<strong>о</strong>идных <strong>к</strong>лещей<br />

<strong>о</strong>тряд<strong>а</strong> Acariformes т<strong>а</strong><strong>к</strong>же <strong>к</strong> реду<strong>к</strong>ции <strong>о</strong>дн<strong>о</strong>г<strong>о</strong><br />

— трех <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы.

Сегменты<br />

тел<strong>а</strong><br />

I<br />

II<br />

III<br />

IV<br />

V<br />

VI<br />

VII<br />

VIII<br />

IX<br />

X<br />

XI<br />

XII<br />

XIII<br />

XIV<br />

XV<br />

XVI<br />

XVII<br />

IXVIII<br />

XIX<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Trilobitomorpha<br />

Пред<strong>а</strong>нтенн<strong>а</strong>льный<strong>с</strong>егментАнтенн<strong>а</strong>льный<br />

<strong>с</strong>егмент<br />

П<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>а</strong>льный<br />

1-й<br />

П<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>а</strong>льный<br />

2-й<br />

Предз<strong>а</strong>тыл<strong>о</strong>чный<br />

З<strong>а</strong>тыл<strong>о</strong>чный<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

Н<strong>о</strong>ги<br />

<strong>о</strong><br />

ч<br />

<strong>о</strong><br />

СХЕМА РАСПРЕДЕЛЕНИЯ КОНЕЧНОСТЕЙ И ИХ ДЕРИВАТОВ ПО СЕГМЕНТАМ И ТАГМАМ ТЕЛА<br />

ОСНОВНЫХ НАДКЛАССОВ И КЛАССОВ ЧЛЕНИСТОНОГИХ<br />

<strong>а</strong><br />

К<br />

ю<br />

<strong>о</strong><br />

ч<br />

>><br />

н<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Merostomata<br />

Хелицеры<br />

Педип<strong>а</strong>льпы<br />

Н<strong>о</strong>ги I<br />

Н<strong>о</strong>ги II<br />

<strong>о</strong><br />

Н<strong>о</strong>ги III <strong>о</strong><br />

Н<strong>о</strong>ги IV<br />

Генит<strong>а</strong>льные<br />

<strong>к</strong>рыш<strong>к</strong>и<br />

Ж<strong>а</strong>бры<br />

Ж<strong>а</strong>бры<br />

г<strong>а</strong><br />

Ж<strong>а</strong>бры' I<br />

a<br />

<strong>о</strong><br />

Ж<strong>а</strong>бры Jj<br />

Ж<strong>а</strong>бры<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Scorpionomorpha<br />

Хелицеры |<br />

<strong>о</strong><br />

и<br />

Педип<strong>а</strong>льпы <strong>о</strong>.<br />

Н<strong>о</strong>ги I<br />

Н<strong>о</strong>ги II <strong>о</strong><br />

CQ<br />

О<br />

Н<strong>о</strong>ги III <strong>о</strong><br />

1—I<br />

Н<strong>о</strong>ги IV<br />

Генит<strong>а</strong>льные ^<br />

<strong>к</strong>рыш<strong>к</strong>и %<br />

<strong>а</strong><br />

» 2<br />

Гребенч<strong>а</strong>тый <strong>о</strong>.<br />

<strong>о</strong>рг<strong>а</strong>н ^<br />

Лег<strong>к</strong>ие g<br />

Лег<strong>к</strong>ие<br />

Лег<strong>к</strong>ие<br />

Лег<strong>к</strong>ие<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Chelicerata<br />

<strong>а</strong> <strong>а</strong><br />

<strong>о</strong> 2<br />

Си<br />

2 ю<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Arachnida<br />

Хелицеры s<br />

<strong>о</strong><br />

и<br />

_ <strong>о</strong><br />

Педип<strong>а</strong>льпы р.<br />

Н<strong>о</strong>ги I<br />

Н<strong>о</strong>ги II 9<br />

Н<strong>о</strong>ги III<br />

Н<strong>о</strong>ги IV<br />

Лег<strong>к</strong>ие или<br />

генит<strong>а</strong>льные<br />

<strong>к</strong>рыш<strong>к</strong>и<br />

Лег<strong>к</strong>ие<br />

П<strong>а</strong>утинные<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и<br />

П<strong>а</strong>утинные<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и<br />

<strong>о</strong><br />

<strong>а</strong><br />

s<br />

Си<br />

Ш<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Solifugomorpha<br />

г<br />

Хелицеры S<br />

<strong>с</strong>и<br />

Педип<strong>а</strong>льпы g<br />

Си<br />

С<br />

НОГИ I<br />

НОГИ II<br />

Н<strong>о</strong>ги III<br />

Н<strong>о</strong>ги IV g<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

Стигмы<br />

Стигмы<br />

Стигмы<br />

http://jurassic.ru/<br />

Хелицеры<br />

Педип<strong>а</strong>льпы<br />

Н<strong>о</strong>ги I<br />

|Н<strong>о</strong>ги II<br />

S<br />

<strong>о</strong><br />

Н<strong>о</strong>ги III <strong>о</strong><br />

<strong>о</strong><br />

С<br />

Н<strong>о</strong>ги IV<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Acaromorpha<br />

<strong>о</strong><br />

С=[ г<strong>а</strong><br />

С С,<br />

О О<br />

Си<br />

I °<br />

S<br />

4<br />

я<br />

<strong>о</strong><br />

<strong>о</strong><br />

<strong>о</strong><br />

н<br />

и<br />

К<br />

<strong>с</strong><br />

<strong>о</strong><br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Pycnogonides<br />

Хелиф<strong>о</strong>ры <strong>о</strong><br />

Яйцен<strong>о</strong><strong>с</strong>цы<br />

Н<strong>о</strong>ги I<br />

Н<strong>о</strong>ги II г<strong>а</strong><br />

S<br />

<strong>о</strong><br />

Н<strong>о</strong>ги III <strong>о</strong><br />

Н<strong>о</strong>ги IV "<br />

(Н<strong>о</strong>ги V) S<br />

(Н<strong>о</strong>ги IV)<br />

<strong>о</strong><br />

<strong>к</strong><br />

<strong>а</strong><br />

2<br />

Си<br />

Т<strong>а</strong>блиц<strong>а</strong> 1<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Tracheata,<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

Insecta<br />

Антенны<br />

М<strong>а</strong>ндибулы [<br />

М<strong>а</strong><strong>к</strong><strong>с</strong>иллы I<br />

М<strong>а</strong><strong>к</strong><strong>с</strong>иллы II<br />

Н<strong>о</strong>ги I<br />

Н<strong>о</strong>ги II<br />

Н<strong>о</strong>ги III<br />

<strong>к</strong> <strong>о</strong> Ь;<br />

a g|<br />

г<strong>а</strong><br />

-*-»<br />

<strong>о</strong> .<br />

<strong>а</strong>д х г<strong>а</strong><br />

E?at5<br />

(У Ю Й)<br />

•+-» О W<br />

О. Я С<br />

< г<strong>а</strong>­<br />

г<strong>а</strong> 3 х<br />

м <strong>о</strong>, S<br />

ч « и<br />

<strong>к</strong> н <strong>о</strong><br />

ш <strong>к</strong>

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер <strong>с</strong><strong>о</strong>членения <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

р<strong>а</strong>зличен. У б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> ф<strong>о</strong>рм эти д<strong>в</strong><strong>а</strong><br />

<strong>о</strong>тдел<strong>а</strong> тел<strong>а</strong> <strong>с</strong><strong>о</strong>членяют<strong>с</strong>я п<strong>о</strong> <strong>в</strong><strong>с</strong>ей ширине VI<br />

и VII <strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong>, и лишь у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>ни<br />

<strong>с</strong><strong>о</strong>единены п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м уз<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>тебель<strong>к</strong><strong>а</strong>. Сегмент<strong>а</strong>ция<br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong>енн<strong>а</strong> <strong>в</strong><strong>с</strong>ем пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителям<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> Merostomata u Scorpionomorpha.<br />

Среди Arachnida у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>с</strong>егменты брюш<strong>к</strong><strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м.<br />

И<strong>с</strong>чезн<strong>о</strong><strong>в</strong>ение <strong>с</strong>егмент<strong>а</strong>ции тел<strong>а</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

т<strong>а</strong><strong>к</strong>же у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Acaromorpha.<br />

С<strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>с</strong> изменением х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>а</strong><br />

мет<strong>а</strong>мерии пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дит <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>ция <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

и ч<strong>а</strong><strong>с</strong>тичн<strong>а</strong>я их реду<strong>к</strong>ция. В <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е<br />

<strong>о</strong>бщей черты <strong>о</strong>тметим, чт<strong>о</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

у <strong>в</strong><strong>с</strong>ех хелицер<strong>о</strong><strong>в</strong>ых д<strong>а</strong>же при <strong>в</strong>ых<strong>о</strong>де н<strong>а</strong> <strong>с</strong>ушу<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я р<strong>о</strong>т<strong>о</strong><strong>в</strong>ыми и л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>т<strong>о</strong>рными, <strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы — дых<strong>а</strong>тельными; <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>мы, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итые у Trilobitomorpha,<br />

<strong>с</strong>реди Chelicerata <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Вт<strong>о</strong>р<strong>о</strong>й <strong>о</strong>бщей<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тью изменений <strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении'и фун<strong>к</strong>ции<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей у Chelicerata <strong>с</strong>лужит пре<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

пер<strong>в</strong>ичн<strong>о</strong>-<strong>в</strong><strong>о</strong>дных ж<strong>а</strong>берных н<strong>о</strong>же<strong>к</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы у Merostomata u Eurypterida<br />

<strong>в</strong> <strong>о</strong>рг<strong>а</strong>ны <strong>в</strong><strong>о</strong>здушн<strong>о</strong>г<strong>о</strong> дых<strong>а</strong>ния у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Scorpionomorpha и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Arachnida, Solifugomorpha.<br />

Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, <strong>о</strong>бщей тенденцией я<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>о</strong>гр<strong>а</strong>ничение уч<strong>а</strong><strong>с</strong>тия пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>в</strong> приеме пищи. У Merostomata<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>о</strong><strong>к</strong>ружен<strong>о</strong> м<strong>а</strong><strong>к</strong><strong>с</strong>иллярными<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми (эндит<strong>а</strong>ми) <strong>в</strong><strong>с</strong>ех <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>к</strong>р<strong>о</strong>ме хелицер. У Eurypterida и <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нии предр<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти приним<strong>а</strong>ют<br />

уч<strong>а</strong><strong>с</strong>тие т<strong>о</strong>ль<strong>к</strong><strong>о</strong> эндиты- педип<strong>а</strong>льп и<br />

пер<strong>в</strong>ых д<strong>в</strong>ух п<strong>а</strong>р х<strong>о</strong>дильных н<strong>о</strong>г. Д<strong>а</strong>лее, у Opiliones<br />

и Pedipalpides п<strong>о</strong>д<strong>о</strong>бную фун<strong>к</strong>цию <strong>в</strong>ып<strong>о</strong>лняют<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> эндиты педип<strong>а</strong>льп и т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>о</strong>г пер<strong>в</strong>ых (реже и н<strong>о</strong>г <strong>в</strong>т<strong>о</strong>рых), <strong>а</strong> у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

других хелицер<strong>о</strong><strong>в</strong>ых — т<strong>о</strong>ль<strong>к</strong><strong>о</strong> эндиты т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

педип<strong>а</strong>льп. Фун<strong>к</strong>цию же<strong>в</strong><strong>а</strong>ния, <strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> и<br />

з<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>а</strong> пищи <strong>в</strong>ып<strong>о</strong>лняют д<strong>в</strong>ух-четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

хелицеры, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Chelicerata <strong>к</strong>лешне<strong>в</strong>идные <strong>с</strong> з<strong>а</strong>зубренными<br />

<strong>к</strong>р<strong>а</strong>ями их п<strong>а</strong>льце<strong>в</strong>. Одн<strong>а</strong><strong>к</strong><strong>о</strong> у Аг<strong>а</strong>пе<strong>а</strong>е и Anthracomarti<br />

хелицеры <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идные, <strong>а</strong> <strong>с</strong><strong>а</strong>м пр<strong>о</strong>це<strong>с</strong><strong>с</strong><br />

пище<strong>в</strong><strong>а</strong>рения у этих хелицер<strong>о</strong><strong>в</strong>ых н<strong>а</strong>ружный;<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ии <strong>с</strong> этим н<strong>а</strong> эндит<strong>а</strong>х т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

педип<strong>а</strong>льп <strong>о</strong>бр<strong>а</strong>зует<strong>с</strong>я <strong>с</strong>пеци<strong>а</strong>льн<strong>а</strong>я фильтр<strong>а</strong>ци<strong>о</strong>нн<strong>а</strong>я<br />

<strong>с</strong>ет<strong>к</strong><strong>а</strong>, через <strong>к</strong><strong>о</strong>т<strong>о</strong>рую пр<strong>о</strong>цежи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

р<strong>а</strong>зжиженн<strong>а</strong>я пищ<strong>а</strong>. П<strong>о</strong>чти у <strong>в</strong><strong>с</strong>ех Merostomata<br />

и Scorpionomorpha н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я <strong>с</strong>ильн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие<br />

педип<strong>а</strong>льп, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я <strong>к</strong>лешне<strong>в</strong>идными<br />

и и<strong>с</strong>п<strong>о</strong>льзуют<strong>с</strong>я для з<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>а</strong>, <strong>а</strong> ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

и для р<strong>а</strong>змельчения пищи. У других<br />

хелицер<strong>о</strong><strong>в</strong>ых педип<strong>а</strong>льпы н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные и <strong>в</strong>ып<strong>о</strong>лняют<br />

фун<strong>к</strong>цию <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> <strong>о</strong><strong>с</strong>яз<strong>а</strong>ния. Педип<strong>а</strong>льпы,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>ги м<strong>а</strong>ндибул р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>з­<br />

380<br />

http://jurassic.ru/<br />

ных и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, и у примити<strong>в</strong>ных Chelicerata<br />

ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют ' <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующую фун<strong>к</strong>цию.<br />

У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>-п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льпы <strong>в</strong>ып<strong>о</strong>лняют<br />

р<strong>о</strong>ль <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>, перен<strong>о</strong><strong>с</strong>ящих <strong>с</strong>перму <strong>в</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> и бы<strong>в</strong><strong>а</strong>ют <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong>ид<strong>о</strong>изменены.<br />

Другие <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы — х<strong>о</strong>дильные<br />

н<strong>о</strong>ги. Очень ч<strong>а</strong><strong>с</strong>т<strong>о</strong> н<strong>о</strong>ги I п<strong>а</strong>ры у<strong>с</strong>тр<strong>о</strong>ены <strong>о</strong>тличн<strong>о</strong><br />

<strong>о</strong>т других н<strong>о</strong>г и фун<strong>к</strong>ци<strong>о</strong>нируют <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>рг<strong>а</strong>ны<br />

<strong>о</strong><strong>с</strong>яз<strong>а</strong>ния: у мн<strong>о</strong>гих Pedipalpides, н<strong>а</strong>пример,<br />

<strong>о</strong>ни <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я биче<strong>в</strong>идными. Х<strong>о</strong>дильные<br />

н<strong>о</strong>ги у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong>лещей м<strong>о</strong>гут п<strong>о</strong>д<strong>в</strong>ерг<strong>а</strong>ть<strong>с</strong>я<br />

реду<strong>к</strong>ции, <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> чем <strong>о</strong>бщее <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей м<strong>о</strong>жет <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>тить<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong>дну (н<strong>а</strong>пример, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые Tarsonemini)<br />

или д<strong>в</strong>е п<strong>а</strong>ры (н<strong>а</strong>пример, Eriophyidae). Интере<strong>с</strong>н<strong>о</strong>,<br />

чт<strong>о</strong> реду<strong>к</strong>ции п<strong>о</strong>д<strong>в</strong>ерг<strong>а</strong>ют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> з<strong>а</strong>дние<br />

п<strong>а</strong>ры н<strong>о</strong>г, т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти л<strong>а</strong>р<strong>в</strong><strong>а</strong>льных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я у <strong>в</strong><strong>с</strong>ех Chelicerata.<br />

Р<strong>а</strong><strong>с</strong>членение <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> п<strong>о</strong><strong>с</strong>т<strong>о</strong>янн<strong>о</strong>.<br />

Т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong> <strong>о</strong>дних групп<strong>а</strong>х неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><br />

<strong>с</strong>литы <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми тел<strong>а</strong>, <strong>в</strong> других <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

п<strong>о</strong>д<strong>в</strong>ижными. Д<strong>а</strong>лее <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть<br />

(энд<strong>о</strong>п<strong>о</strong>дит) р<strong>а</strong><strong>с</strong>членяет<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ертлуг, бедр<strong>о</strong>,<br />

<strong>к</strong><strong>о</strong>лен<strong>о</strong>, г<strong>о</strong>лень и л<strong>а</strong>п<strong>к</strong>у; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бедр<strong>о</strong> и <strong>в</strong>ертлуг<br />

п<strong>о</strong>др<strong>а</strong>зделяют<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельных члени<strong>к</strong><strong>а</strong><br />

(basi- и telofemur), <strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>членяет<strong>с</strong>я<br />

д<strong>о</strong>п<strong>о</strong>лнительн<strong>о</strong> н<strong>а</strong> д<strong>в</strong><strong>а</strong> (metatarsus и tarsus)<br />

или пять члени<strong>к</strong><strong>о</strong><strong>в</strong>. Н<strong>а</strong> <strong>в</strong>ершине л<strong>а</strong>п<strong>к</strong>и при<strong>к</strong>репляют<strong>с</strong>я<br />

д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong> и прет<strong>а</strong>рз, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ющий<strong>с</strong>я<br />

<strong>в</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идный прид<strong>а</strong>т<strong>о</strong><strong>к</strong>, или<br />

пери<strong>с</strong>тый эмп<strong>о</strong>дий. Н<strong>а</strong> члени<strong>к</strong><strong>а</strong>х н<strong>о</strong>г м<strong>о</strong>гут<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ть<strong>с</strong>я р<strong>а</strong>зличные <strong>о</strong>рг<strong>а</strong>ны чу<strong>в</strong><strong>с</strong>т<strong>в</strong>. Между<br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>бр<strong>а</strong>зует<strong>с</strong>я <strong>с</strong>терн<strong>а</strong>льный щит.<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти мез<strong>о</strong><strong>с</strong><strong>о</strong>мы у пер<strong>в</strong>ичн<strong>о</strong>-<strong>в</strong><strong>о</strong>дных<br />

хелицер<strong>о</strong><strong>в</strong>ых (Merostomata, Eurypterida, <strong>с</strong>илурий<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны Apoxypodes) пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

четырьмя п<strong>а</strong>р<strong>а</strong>ми ж<strong>а</strong>берных н<strong>о</strong>же<strong>к</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

н<strong>а</strong> IX—XIII <strong>с</strong>егмент<strong>а</strong>х, и п<strong>а</strong>р<strong>о</strong>й<br />

генит<strong>а</strong>льных <strong>к</strong>рышече<strong>к</strong> н<strong>а</strong> VIII <strong>с</strong>терните. Среди<br />

н<strong>а</strong>земных хелицер<strong>о</strong><strong>в</strong>ых н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong>лный<br />

н<strong>а</strong>б<strong>о</strong>р мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей имеет<strong>с</strong>я<br />

у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, где <strong>о</strong>ни пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены п<strong>а</strong>р<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых<br />

<strong>к</strong>рыше<strong>к</strong> н<strong>а</strong> VIII <strong>с</strong>егменте, четырьмя п<strong>а</strong>р<strong>а</strong>ми<br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х X—XIII<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> и п<strong>а</strong>р<strong>о</strong>й гребне<strong>в</strong>идных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong><br />

чу<strong>в</strong><strong>с</strong>т<strong>в</strong> н<strong>а</strong> IX <strong>с</strong>егменте; лег<strong>о</strong>чные меш<strong>к</strong>и я<strong>в</strong>ляют<strong>с</strong>я<br />

п<strong>о</strong>груженными п<strong>о</strong>д <strong>к</strong><strong>о</strong>жу ж<strong>а</strong>берными<br />

н<strong>о</strong>ж<strong>к</strong><strong>а</strong>ми их пред<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>о</strong>бит<strong>а</strong><strong>в</strong>ших <strong>в</strong> <strong>в</strong><strong>о</strong>де.<br />

У других хелицер<strong>о</strong><strong>в</strong>ых н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я уменьшение<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. У Uropygi,<br />

Amblypygi, Palpigradi, Ricinulei и др. <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> д<strong>в</strong>е п<strong>а</strong>ры лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><br />

IX и X <strong>с</strong>егмент<strong>а</strong>х, у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> их <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>дн<strong>а</strong><br />

или д<strong>в</strong>е п<strong>а</strong>ры. У п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Аг<strong>а</strong>пе<strong>а</strong>е<br />

з<strong>а</strong>дние п<strong>а</strong>ры мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

пре<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ли<strong>с</strong>ь <strong>в</strong> д<strong>в</strong>е п<strong>а</strong>ры п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>-

д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>; третья п<strong>а</strong>р<strong>а</strong> п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ем пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении не <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>а</strong> <strong>с</strong> <strong>с</strong>егмент<strong>а</strong>ми<br />

тел<strong>а</strong>.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>й <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тью <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции<br />

мн<strong>о</strong>гих хелицер<strong>о</strong><strong>в</strong>ых я<strong>в</strong>ляет<strong>с</strong>я <strong>с</strong>терн<strong>а</strong>льный<br />

<strong>с</strong><strong>к</strong>елет мез<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ующий у трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>,<br />

Merostomata и мн<strong>о</strong>гих примити<strong>в</strong>ных Scorpionomorpha,<br />

Solifugomorpha и Arachnida. Эт<strong>о</strong><br />

<strong>о</strong>бу<strong>с</strong>л<strong>о</strong><strong>в</strong>лен<strong>о</strong> у<strong>с</strong>илением фун<strong>к</strong>ции н<strong>о</strong>г при н<strong>а</strong>земн<strong>о</strong>м<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>нии и не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>тью у<strong>к</strong>репления<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны груди.<br />

Т<strong>а</strong>зи<strong>к</strong>и <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми тел<strong>а</strong> и<br />

<strong>о</strong>бр<strong>а</strong>зуют зде<strong>с</strong>ь <strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льные щиты, или<br />

же р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong>пер<strong>в</strong><strong>а</strong> мет<strong>а</strong>мерные, <strong>а</strong> з<strong>а</strong>тем<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>с</strong>терн<strong>а</strong>льные щиты.:<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие у <strong>в</strong><strong>с</strong>ех Chelicerata р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>терните VIII тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

(<strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>);<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>н<strong>о</strong> лежит н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не или<br />

п<strong>о</strong>з<strong>а</strong>ди т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV. У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong>лещей,<br />

<strong>а</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у Opiliones, н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я <strong>с</strong>мещение<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия <strong>в</strong>перед и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение ег<strong>о</strong><br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г I.<br />

В э<strong>в</strong><strong>о</strong>люции нер<strong>в</strong>н<strong>о</strong>й <strong>с</strong>и<strong>с</strong>темы у Chelicerata<br />

<strong>о</strong>тмеч<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong><strong>с</strong>тепенн<strong>а</strong>я <strong>к</strong><strong>о</strong>нцентр<strong>а</strong>ция брюшн<strong>о</strong>й<br />

цеп<strong>о</strong>ч<strong>к</strong>и. У <strong>с</strong><strong>о</strong><strong>в</strong>ременных Merostomata<br />

г<strong>а</strong>нглии пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong>литы <strong>в</strong>близи пище<strong>в</strong><strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

<strong>о</strong>дну нер<strong>в</strong>ную м<strong>а</strong><strong>с</strong><strong>с</strong>у, <strong>а</strong> г<strong>а</strong>нглии мез<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельными, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, имеет<strong>с</strong>я<br />

<strong>о</strong>дин г<strong>а</strong>нглий <strong>в</strong> мет<strong>а</strong><strong>с</strong><strong>о</strong>ме. Д<strong>а</strong>лее, у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

<strong>в</strong><strong>о</strong>един<strong>о</strong> <strong>с</strong>литы <strong>в</strong><strong>с</strong>е г<strong>а</strong>нглии пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и четыре<br />

г<strong>а</strong>нглия мез<strong>о</strong><strong>с</strong><strong>о</strong>мы; <strong>о</strong><strong>с</strong>т<strong>а</strong>льные д<strong>в</strong><strong>а</strong> г<strong>а</strong>нглия мез<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и ше<strong>с</strong>ть г<strong>а</strong>нглие<strong>в</strong> мет<strong>а</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными. У Uropygi и Solifugae <strong>в</strong><strong>с</strong>е г<strong>а</strong>нглии<br />

брюшн<strong>о</strong>й нер<strong>в</strong>н<strong>о</strong>й цеп<strong>о</strong>ч<strong>к</strong>и <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

<strong>о</strong>дну п<strong>о</strong>дгл<strong>о</strong>т<strong>о</strong>чную м<strong>а</strong><strong>с</strong><strong>с</strong>у и т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> з<strong>а</strong>д<strong>а</strong>ем<br />

<strong>к</strong><strong>о</strong>нце тел<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я тр<strong>о</strong>йн<strong>о</strong>й <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленный<br />

г<strong>а</strong>нглий, иннер<strong>в</strong>ирующий три п<strong>о</strong><strong>с</strong>ледние <strong>с</strong>егмент<strong>а</strong><br />

тел<strong>а</strong>. У <strong>в</strong><strong>с</strong>ех <strong>о</strong><strong>с</strong>т<strong>а</strong>льных хелицер<strong>о</strong><strong>в</strong>ых <strong>в</strong><strong>с</strong>е<br />

г<strong>а</strong>нглии <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>с</strong>пл<strong>о</strong>шную нер<strong>в</strong>ную м<strong>а</strong><strong>с</strong><strong>с</strong>у<br />

(«м<strong>о</strong>зг»), пр<strong>о</strong>низ<strong>а</strong>нную пище<strong>в</strong><strong>о</strong>д<strong>о</strong>м.<br />

Пище<strong>в</strong><strong>а</strong>рительн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет <strong>в</strong> цел<strong>о</strong>м<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные черты <strong>к</strong>ишечни<strong>к</strong><strong>а</strong> члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

и х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я п<strong>о</strong><strong>с</strong>т<strong>о</strong>янным н<strong>а</strong>личием<br />

н<strong>а</strong> <strong>с</strong>редней <strong>к</strong>иш<strong>к</strong>е длинных б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>в</strong>ыпячи<strong>в</strong><strong>а</strong>ний<br />

(ди<strong>в</strong>ерти<strong>к</strong>ул<strong>о</strong><strong>в</strong>). Н<strong>а</strong> гр<strong>а</strong>нице <strong>с</strong>редней<br />

и з<strong>а</strong>дней <strong>к</strong>ише<strong>к</strong> <strong>в</strong> пище<strong>в</strong><strong>а</strong>рительный <strong>к</strong><strong>а</strong>н<strong>а</strong>л <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>рг<strong>а</strong>ны <strong>в</strong>ыделения — <strong>с</strong>леп<strong>о</strong> з<strong>а</strong>м<strong>к</strong>нутые<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>х труб<strong>к</strong>и м<strong>а</strong>льпигие<strong>в</strong>ых <strong>с</strong><strong>о</strong><strong>с</strong>уд<strong>о</strong><strong>в</strong>.<br />

Из других <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> <strong>в</strong>ыделения у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

Chelicerata имеют<strong>с</strong>я <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зные <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льные<br />

железы; <strong>о</strong>ни б<strong>о</strong>лее р<strong>а</strong>з<strong>в</strong>иты у з<strong>а</strong>р<strong>о</strong>дышей и <strong>в</strong><br />

м<strong>о</strong>л<strong>о</strong>д<strong>о</strong>м <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>те, у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых <strong>о</strong><strong>с</strong><strong>о</strong>бей ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

редуцируют<strong>с</strong>я.<br />

В<strong>с</strong>е Chelicerata р<strong>а</strong>здельн<strong>о</strong>п<strong>о</strong>лы. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

дим<strong>о</strong>рфизм пр<strong>о</strong>я<strong>в</strong>ляет<strong>с</strong>я, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, нерез<strong>к</strong><strong>о</strong><br />

и <strong>о</strong>бычн<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>ж<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> р<strong>а</strong>зличиях р<strong>а</strong>змер<strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> и <strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении <strong>о</strong>тдельных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. П<strong>о</strong>чти<br />

<strong>в</strong><strong>с</strong>е Chelicerata <strong>о</strong>т<strong>к</strong>л<strong>а</strong>ды<strong>в</strong><strong>а</strong>ют яйц<strong>а</strong>, зн<strong>а</strong>чительн<strong>о</strong><br />

реже <strong>с</strong>реди них н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я жи<strong>в</strong><strong>о</strong>р<strong>о</strong>дящие<br />

ф<strong>о</strong>рмы (<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>к</strong>лещи). Ин<strong>о</strong>гд<strong>а</strong><br />

яйц<strong>а</strong> <strong>с</strong><strong>о</strong>бир<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>о</strong><strong>к</strong><strong>о</strong>ны, <strong>о</strong>плетенные п<strong>а</strong>утин<strong>о</strong>й<br />

(п<strong>а</strong>у<strong>к</strong>и), <strong>в</strong>ын<strong>а</strong>ши<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюш<strong>к</strong>е <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

(л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны) или при<strong>к</strong>репляют<strong>с</strong>я н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>к</strong>лещи,<br />

Oribatei). Яйц<strong>а</strong> хелицер<strong>о</strong><strong>в</strong>ых <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

<strong>к</strong>рупные, б<strong>о</strong>г<strong>а</strong>тые желт<strong>к</strong><strong>о</strong>м, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong><br />

др<strong>о</strong>бление яйце<strong>к</strong>лет<strong>к</strong>и ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong>е, п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тн<strong>о</strong>е.<br />

У з<strong>а</strong>р<strong>о</strong>дышей, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, бы<strong>в</strong><strong>а</strong>ет <strong>о</strong>чень<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> мет<strong>а</strong>мерия, <strong>а</strong> н<strong>а</strong> брюш<strong>к</strong>е<br />

имеют<strong>с</strong>я <strong>с</strong>леды <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей (у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> пяти<br />

<strong>с</strong>егмент<strong>а</strong>х, <strong>а</strong> у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> — н<strong>а</strong> <strong>с</strong>еми <strong>с</strong>егмент<strong>а</strong>х<br />

брюш<strong>к</strong><strong>а</strong>). Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, у эмбри<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

прет<strong>а</strong>рзу<strong>с</strong>ы пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong> бы<strong>в</strong><strong>а</strong>ют <strong>о</strong>дн<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>ые:<br />

эт<strong>о</strong> <strong>о</strong>блегч<strong>а</strong>ет <strong>с</strong>уждение <strong>о</strong>б их<br />

фил<strong>о</strong>генезе- П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие прям<strong>о</strong>е,<br />

н<strong>о</strong> у <strong>к</strong>лещей д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>с</strong>л<strong>о</strong>жный мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з.<br />

Интере<strong>с</strong>н<strong>о</strong>, чт<strong>о</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых хелицер<strong>о</strong><strong>в</strong>ых<br />

(<strong>к</strong>лещи, Ricinulei) <strong>в</strong>ылупляющ<strong>а</strong>я<strong>с</strong>я из яйц<strong>а</strong><br />

личин<strong>к</strong><strong>а</strong> имеет т<strong>о</strong>ль<strong>к</strong><strong>о</strong> ше<strong>с</strong>ть н<strong>о</strong>г, т<strong>о</strong>гд<strong>а</strong><br />

<strong>к</strong><strong>а</strong><strong>к</strong> у других Chelicerata личии<strong>к</strong><strong>а</strong> <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью<br />

д<strong>о</strong>г<strong>а</strong>ми.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

В э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>м <strong>о</strong>тн<strong>о</strong>шении<br />

хелицер<strong>о</strong><strong>в</strong>ые <strong>к</strong>р<strong>а</strong>йне р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны. Пер<strong>в</strong>ичн<strong>о</strong><strong>в</strong><strong>о</strong>дными<br />

<strong>с</strong>реди них я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong><strong>с</strong>е<br />

Merostomata, пр<strong>о</strong>из<strong>о</strong>шедшие <strong>о</strong>т них Eurypterida<br />

и <strong>с</strong>илурий<strong>с</strong><strong>к</strong>ие <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Ар<strong>о</strong>xypodes.<br />

Уже <strong>в</strong> де<strong>в</strong><strong>о</strong>не пр<strong>о</strong>из<strong>о</strong>шел <strong>в</strong>ых<strong>о</strong>д хелицер<strong>о</strong><strong>в</strong>ых<br />

н<strong>а</strong> <strong>с</strong>ушу, чт<strong>о</strong> <strong>о</strong>пределил<strong>о</strong> <strong>в</strong><strong>с</strong>ё<br />

д<strong>а</strong>льнейшее р<strong>а</strong>з<strong>в</strong>итие этих жи<strong>в</strong><strong>о</strong>тных. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

<strong>о</strong>тряд<strong>а</strong> Palpigradi, <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong> н<strong>а</strong><strong>с</strong>ыщенн<strong>о</strong>й<br />

<strong>в</strong><strong>о</strong>д<strong>о</strong>й п<strong>о</strong>ч<strong>в</strong>е, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили н<strong>а</strong>иб<strong>о</strong>лее<br />

примити<strong>в</strong>ные черты <strong>с</strong>тр<strong>о</strong>ения <strong>с</strong>реди <strong>в</strong><strong>с</strong>ех н<strong>а</strong>земных<br />

хелицер<strong>о</strong><strong>в</strong>ых. В д<strong>а</strong>льнейшем не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители или целые <strong>к</strong>рупные <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие<br />

<strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>ы ф<strong>о</strong>рм <strong>в</strong>н<strong>о</strong><strong>в</strong>ь перех<strong>о</strong>дили<br />

<strong>к</strong> <strong>в</strong><strong>о</strong>дн<strong>о</strong>му <strong>о</strong>бр<strong>а</strong>зу жизни (<strong>в</strong><strong>о</strong>дяные п<strong>а</strong>у<strong>к</strong>и <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Argyronetidae, пре<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>дные и м<strong>о</strong>р<strong>с</strong><strong>к</strong>ие<br />

<strong>к</strong>лещи Hydrachnellae и Halacarae), <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong><br />

при эт<strong>о</strong>м <strong>в</strong> <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции их не пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<strong>о</strong><br />

<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енных изменений. Н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> п<strong>а</strong>нцирные и <strong>с</strong><strong>а</strong>р<strong>к</strong><strong>о</strong>пт<strong>о</strong>идные<br />

<strong>к</strong>лещи и п<strong>а</strong>у<strong>к</strong>и, ныне <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><strong>в</strong><strong>с</strong>еме<strong>с</strong>тн<strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>уше. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> групп <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременную<br />

эп<strong>о</strong>ху н<strong>а</strong><strong>с</strong>еляет преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>ие <strong>с</strong>тр<strong>а</strong>ны. П<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющее<br />

б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> Chelicerata — хищни<strong>к</strong>и,<br />

н<strong>о</strong> <strong>с</strong>реди них имеют<strong>с</strong>я и р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные<br />

ф<strong>о</strong>рмы, <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги и п<strong>а</strong>р<strong>а</strong>зиты жи<strong>в</strong><strong>о</strong>тных.<br />

Гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>е б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> хелицер<strong>о</strong><strong>в</strong>ых —<br />

<strong>с</strong>умеречные или н<strong>о</strong>чные жи<strong>в</strong><strong>о</strong>тные, чт<strong>о</strong>, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>,<br />

<strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong> з<strong>а</strong>труднениями терм<strong>о</strong>регуляции<br />

и <strong>в</strong><strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бмен<strong>а</strong>. Одн<strong>а</strong><strong>к</strong><strong>о</strong> имеют<strong>с</strong>я<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>иды, <strong>о</strong>х<strong>о</strong>тящие<strong>с</strong>я при яр<strong>к</strong><strong>о</strong>м <strong>с</strong><strong>о</strong>лн-<br />

381<br />

http://jurassic.ru/

це; т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы, н<strong>а</strong>пример, п<strong>а</strong>у<strong>к</strong>и-<strong>с</strong><strong>к</strong><strong>а</strong><strong>к</strong>уны, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цы и др.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме и фил<strong>о</strong>гения.<br />

В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее <strong>в</strong>ремя <strong>в</strong><strong>с</strong>еми призн<strong>а</strong>н<strong>о</strong>, чт<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременные н<strong>а</strong>земные Chelicerata пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят<br />

<strong>о</strong>т примити<strong>в</strong>ных <strong>в</strong><strong>о</strong>дных хелицер<strong>о</strong><strong>в</strong>ых, <strong>а</strong> те, <strong>в</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>ю <strong>о</strong>чередь,— <strong>о</strong>т "Trilobitomorpha. Одн<strong>а</strong><strong>к</strong><strong>о</strong><br />

<strong>к</strong><strong>о</strong>н<strong>к</strong>ретные пути э<strong>в</strong><strong>о</strong>люции <strong>о</strong>тдельных групп<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я не<strong>в</strong>ыя<strong>с</strong>ненными и <strong>с</strong>п<strong>о</strong>рными. Вы<strong>с</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong><strong>о</strong>мнение <strong>в</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении <strong>в</strong><strong>с</strong>ех из<strong>в</strong>е<strong>с</strong>тных<br />

Chelicerata т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>т Merostomata,<br />

Eurypterida и Apoxypodes и предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

нез<strong>а</strong><strong>в</strong>и<strong>с</strong>им<strong>о</strong>е пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение <strong>о</strong>тдельных групп<br />

н<strong>а</strong>земных хелицер<strong>о</strong><strong>в</strong>ых <strong>о</strong>т р<strong>а</strong>зличных и не из<strong>в</strong>е<strong>с</strong>тных<br />

п<strong>о</strong><strong>к</strong><strong>а</strong> н<strong>а</strong>м групп <strong>в</strong><strong>о</strong>дных хелицер<strong>о</strong><strong>в</strong>ых<br />

(Бе<strong>к</strong>лемише<strong>в</strong>, 1952)- Т<strong>а</strong><strong>к</strong><strong>о</strong>е пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ление <strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ет,<br />

не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>, п<strong>о</strong>т<strong>о</strong>му, чт<strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция<br />

Chelicerata <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>л<strong>а</strong><strong>с</strong>ь н<strong>а</strong> <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong>м<br />

принципе р<strong>а</strong>зделения хелицер<strong>о</strong><strong>в</strong>ых н<strong>а</strong> д<strong>в</strong>е<br />

<strong>с</strong>б<strong>о</strong>рные и, п<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>у, э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие группы:<br />

<strong>в</strong><strong>о</strong>дные (Merostomata) и н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые<br />

(Arachnoidea). Предп<strong>о</strong>л<strong>о</strong>жению <strong>о</strong> п<strong>о</strong>лифилетиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении <strong>в</strong><strong>с</strong>ех <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

<strong>о</strong>тряд<strong>о</strong><strong>в</strong> пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>речит н<strong>а</strong>личие у <strong>в</strong><strong>с</strong>ех Chelicerata<br />

<strong>о</strong>бщих призн<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>, <strong>к</strong><strong>а</strong><strong>к</strong>-т<strong>о</strong>: п<strong>о</strong>лн<strong>а</strong>я реду<strong>к</strong>ция<br />

<strong>а</strong>нтеннул, з<strong>а</strong>мен<strong>а</strong> <strong>а</strong>нтенн хелицер<strong>а</strong>ми, п<strong>о</strong><strong>с</strong>т<strong>о</strong>ян<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

чи<strong>с</strong>л<strong>а</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и мез<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

н<strong>а</strong>личие рудимент<strong>а</strong>рн<strong>о</strong>г<strong>о</strong> VII <strong>с</strong>егмент<strong>а</strong><br />

тел<strong>а</strong>, един<strong>с</strong>т<strong>в</strong><strong>о</strong> пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> изгиб<strong>а</strong>ния пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й<br />

<strong>о</strong><strong>с</strong>и тел<strong>а</strong> <strong>с</strong>пер<strong>в</strong><strong>а</strong> <strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти, <strong>а</strong> з<strong>а</strong>тем<br />

<strong>в</strong> х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й и т. п. При п<strong>о</strong>лифилетиче<strong>с</strong><strong>к</strong><strong>о</strong>м р<strong>а</strong>з<strong>в</strong>итии<br />

группы <strong>в</strong>ряд ли м<strong>о</strong>жн<strong>о</strong> <strong>о</strong>жид<strong>а</strong>ть п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>п<strong>а</strong>дения изменений <strong>с</strong>тр<strong>о</strong>ения. Перечи<strong>с</strong>ленные<br />

призн<strong>а</strong><strong>к</strong>и <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>уют т<strong>а</strong><strong>к</strong>ж^е <strong>о</strong><br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении н<strong>а</strong>земных групп Chelicerata не<br />

неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>т трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>, <strong>а</strong> <strong>о</strong>т <strong>к</strong><strong>а</strong><strong>к</strong>их-т<strong>о</strong><br />

<strong>в</strong><strong>о</strong>дных хелицер<strong>о</strong><strong>в</strong>ых.<br />

Неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енные р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные <strong>о</strong>тн<strong>о</strong>шения<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> хелицер<strong>о</strong><strong>в</strong>ых<br />

жи<strong>в</strong><strong>о</strong>тных — мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> (Merostomata) — <strong>с</strong><br />

Trilobitomorpha <strong>с</strong><strong>о</strong>мнений не <strong>в</strong>ызы<strong>в</strong><strong>а</strong>ет. Они<br />

д<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>ими<br />

и эмбри<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ими д<strong>а</strong>нными п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению и<br />

фун<strong>к</strong>ции д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тых брюшных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>с</strong> ж<strong>а</strong>берными прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми, п<strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждению<br />

<strong>с</strong>литн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди <strong>с</strong> ее л<strong>а</strong>р<strong>в</strong><strong>а</strong>льн<strong>о</strong>й и п<strong>о</strong><strong>с</strong>тл<strong>а</strong>р<strong>в</strong><strong>а</strong>льн<strong>о</strong>й<br />

ч<strong>а</strong><strong>с</strong>тями, п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> <strong>в</strong>ыделения,<br />

<strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>щения и т. п. Не <strong>в</strong>ызы<strong>в</strong><strong>а</strong>ет<br />

т<strong>а</strong><strong>к</strong>же <strong>с</strong><strong>о</strong>мнений и неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>а</strong>я фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>язь Merostomata и Eurypterida,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых не<strong>о</strong>дн<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong> д<strong>а</strong>же <strong>о</strong>бъединяли <strong>в</strong><br />

<strong>о</strong>дин <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Palaeostraca.<br />

Зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>льшие трудн<strong>о</strong><strong>с</strong>ти и р<strong>а</strong>зн<strong>о</strong>гл<strong>а</strong><strong>с</strong>ия<br />

<strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ли при у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лении р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

н<strong>а</strong>земных Chelicerata <strong>с</strong> перечи<strong>с</strong>ленными <strong>в</strong>ыше<br />

групп<strong>а</strong>ми. Сближение н<strong>а</strong>земных п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

«<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>» Arachnoidea <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>ж<strong>к</strong><strong>а</strong>ми и<br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мыми п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Tracheata н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е<br />

382<br />

http://jurassic.ru/<br />

н<strong>а</strong>личия <strong>в</strong> <strong>о</strong>беих этих групп<strong>а</strong>х п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

<strong>в</strong><strong>о</strong>зни<strong>к</strong>ших тр<strong>а</strong>хей и м<strong>а</strong>льпигие<strong>в</strong>ых <strong>с</strong><strong>о</strong><strong>с</strong>уд<strong>о</strong><strong>в</strong><br />

я<strong>в</strong>н<strong>о</strong> <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong>.<br />

Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> (преем<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><strong>с</strong>ть)<br />

мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> <strong>с</strong> другими Chelicerata п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong>бщн<strong>о</strong><strong>с</strong>тью <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния гл<strong>а</strong><strong>в</strong>ных<br />

р<strong>а</strong>здел<strong>о</strong><strong>в</strong> тел<strong>а</strong> и х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>о</strong>м мет<strong>а</strong>мерии, <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди,<br />

г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гией ж<strong>а</strong>берных брюшных н<strong>о</strong>же<strong>к</strong> мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><br />

и э<strong>в</strong>риптер<strong>о</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong> лег<strong>о</strong>чными меш<strong>к</strong><strong>а</strong>ми<br />

н<strong>а</strong>земных хелицер<strong>о</strong><strong>в</strong>ых, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием <strong>в</strong><br />

<strong>о</strong>беих групп<strong>а</strong>х <strong>а</strong>нтеннул и <strong>а</strong>нтенн, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гией<br />

<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льных желез, <strong>с</strong>лепых <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> <strong>к</strong>ишечни<strong>к</strong><strong>а</strong>,<br />

<strong>с</strong>тр<strong>о</strong>ением и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жением гл<strong>а</strong>з и т. п.<br />

П<strong>о</strong>эт<strong>о</strong>му имеет<strong>с</strong>я б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> <strong>в</strong>е<strong>с</strong><strong>к</strong>их д<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>тель<strong>с</strong>т<strong>в</strong><br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждения н<strong>а</strong>земных Chelicerata<br />

<strong>о</strong>т <strong>в</strong><strong>о</strong>дных хелицер<strong>о</strong><strong>в</strong>ых и, <strong>в</strong>идим<strong>о</strong>, <strong>о</strong>т <strong>к</strong><strong>а</strong><strong>к</strong>ихт<strong>о</strong><br />

э<strong>в</strong>риптер<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong>, <strong>о</strong>бщий <strong>в</strong>ид<br />

и <strong>с</strong>тр<strong>о</strong>ение <strong>о</strong>тдельных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых не <strong>о</strong>тличимы<br />

пр<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong>и д<strong>а</strong>же <strong>о</strong>т <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>.<br />

При р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трении и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых и <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

хелицер<strong>о</strong><strong>в</strong>ых прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong>бр<strong>а</strong>щ<strong>а</strong>ет н<strong>а</strong><br />

<strong>с</strong>ебя <strong>в</strong>ним<strong>а</strong>ние н<strong>а</strong>личие <strong>с</strong>реди них <strong>о</strong>бширн<strong>о</strong>й<br />

группир<strong>о</strong><strong>в</strong><strong>к</strong>и <strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>с</strong> прям<strong>о</strong>й пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й<br />

<strong>о</strong><strong>с</strong>ью тел<strong>а</strong>,- Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители этих <strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, длинным тел<strong>о</strong>м, брюш<strong>к</strong><strong>о</strong><br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделен<strong>о</strong> н<strong>а</strong> ут<strong>о</strong>лщенный<br />

передний уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong> — переднебрюшье (мез<strong>о</strong><strong>с</strong><strong>о</strong>му)<br />

и б<strong>о</strong>лее т<strong>о</strong>н<strong>к</strong><strong>о</strong>е «х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>идн<strong>о</strong>е» з<strong>а</strong>днебрюшье<br />

(мет<strong>а</strong><strong>с</strong><strong>о</strong>му), <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е м<strong>о</strong>жет п<strong>о</strong>д<strong>в</strong>ерг<strong>а</strong>ть<strong>с</strong>я<br />

зн<strong>а</strong>чительн<strong>о</strong>й реду<strong>к</strong>ции. Эти жи<strong>в</strong><strong>о</strong>тные<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я н<strong>а</strong>личием, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>к</strong>рупных, <strong>к</strong>лешне<strong>в</strong>идных х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельных педип<strong>а</strong>льп.<br />

П<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>е <strong>с</strong>тр<strong>о</strong>ение н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я у Eurypterida,<br />

Scorpiones, Palpigradi, Uropygu<br />

Amblypygi, Ricinulei, Pseudoscorpionoidea и Kustarachnida.<br />

В<strong>с</strong>е эти <strong>о</strong>тряды <strong>о</strong>бъединяют<strong>с</strong>я<br />

н<strong>а</strong>ми <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Scorpionomorpha, <strong>в</strong>едущий <strong>с</strong><strong>в</strong><strong>о</strong>е<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>т примити<strong>в</strong>ных<br />

Merostomata, <strong>с</strong>х<strong>о</strong>дных <strong>с</strong> <strong>к</strong>ембрий<strong>с</strong><strong>к</strong>ими<br />

Aglaspida. Среди <strong>в</strong><strong>с</strong>ех Chelicerata пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

Scorpionomorpha н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ны.<br />

Другие <strong>о</strong>тряды хелицер<strong>о</strong><strong>в</strong>ых я<strong>с</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>а</strong>д<strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> три группы, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующие<strong>с</strong>я изменениями<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> пл<strong>а</strong>н<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ения, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тями<br />

<strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>г<strong>о</strong> изгиб<strong>а</strong>ния пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и<br />

тел<strong>а</strong>, <strong>с</strong>тр<strong>о</strong>ения педип<strong>а</strong>льп и т. п.<br />

В <strong>о</strong>дну группу — <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha —<br />

<strong>в</strong>х<strong>о</strong>дит <strong>о</strong>тряд Solifugae, у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

три <strong>с</strong>егмент<strong>а</strong> передней ч<strong>а</strong><strong>с</strong>ти тел<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

и <strong>о</strong>бр<strong>а</strong>зуют пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляя<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными <strong>с</strong>егменты III и IV п<strong>а</strong>р н<strong>о</strong>г. Пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я<br />

<strong>о</strong><strong>с</strong>ь тел<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я прям<strong>о</strong>й, <strong>а</strong> педип<strong>а</strong>льпы<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я <strong>к</strong>рупными, <strong>в</strong> <strong>в</strong>иде н<strong>о</strong>ги,<br />

н<strong>о</strong> лишенными <strong>к</strong>лешне<strong>в</strong>идных г<strong>о</strong>лени и л<strong>а</strong>п<strong>к</strong>и.<br />

В<strong>о</strong> <strong>в</strong>т<strong>о</strong>рую группу—<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Acaromorpha —<br />

<strong>о</strong>бъединяют<strong>с</strong>я три <strong>о</strong>тряд<strong>а</strong> <strong>к</strong>лещей (Acarifor-

mes, Parasitiformes, Opilioacarina), р<strong>а</strong>нее непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пределенных <strong>в</strong> д<strong>в</strong>ух н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>х<br />

(З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин, 1952). У <strong>к</strong>лещей <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>а</strong>я<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, н<strong>о</strong> <strong>в</strong> <strong>о</strong>тличие<br />

<strong>о</strong>т <strong>в</strong><strong>с</strong>ех других хелицер<strong>о</strong><strong>в</strong>ых, пер<strong>в</strong>ые д<strong>в</strong><strong>а</strong> <strong>с</strong>егмент<strong>а</strong><br />

п<strong>о</strong><strong>с</strong>т<strong>о</strong>янн<strong>о</strong> ф<strong>о</strong>рмируют <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зный<br />

р<strong>а</strong>здел — гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленный <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

тел<strong>а</strong>. П<strong>о</strong><strong>с</strong>леднее у <strong>к</strong>лещей либ<strong>о</strong> бы<strong>в</strong><strong>а</strong>ет<br />

р<strong>а</strong>зделен<strong>о</strong> н<strong>а</strong> три (пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му, мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му<br />

и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му — не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые Endeostigmata) или<br />

д<strong>в</strong><strong>а</strong> р<strong>а</strong>здел<strong>а</strong> (п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му или пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му<br />

и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му — б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> Acariformes),<br />

либ<strong>о</strong> <strong>в</strong><strong>с</strong>е <strong>с</strong>егменты тел<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong><strong>о</strong>един<strong>о</strong>, <strong>о</strong>бр<strong>а</strong>зуя нер<strong>а</strong><strong>с</strong>члененную иди<strong>о</strong><strong>с</strong><strong>о</strong>му<br />

(б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> Parasitiformes) (<strong>с</strong>м. т<strong>а</strong>бл. 1).<br />

Дых<strong>а</strong>ние этих хелицер<strong>о</strong><strong>в</strong>ых <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляет<strong>с</strong>я <strong>с</strong><br />

п<strong>о</strong>м<strong>о</strong>щью тр<strong>а</strong>хей, либ<strong>о</strong> <strong>о</strong>рг<strong>а</strong>ны дых<strong>а</strong>ния <strong>в</strong><strong>о</strong><strong>о</strong>бще<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; пр<strong>о</strong>д<strong>о</strong>льн<strong>а</strong>я <strong>о</strong><strong>с</strong>ь тел<strong>а</strong> изгиб<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти, чт<strong>о</strong> при<strong>в</strong><strong>о</strong>дит <strong>к</strong> зн<strong>а</strong>чительн<strong>о</strong>й<br />

реду<strong>к</strong>ции <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong>ы Solifugomorpha (б<strong>о</strong>лее примити<strong>в</strong>ные)<br />

и Acaromorpha (б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нные)<br />

<strong>в</strong>едут <strong>с</strong><strong>в</strong><strong>о</strong>е пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение <strong>о</strong>т <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> примити<strong>в</strong>ных<br />

ф<strong>о</strong>рм п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Pedipalpides, <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

Scorpionomorpha, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти <strong>о</strong>т <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong>,<br />

<strong>с</strong>х<strong>о</strong>дных <strong>с</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременными Palpigradi и Schizopeltidia.<br />

Об<strong>о</strong><strong>с</strong><strong>о</strong>бление этих <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> пр<strong>о</strong>из<strong>о</strong>шл<strong>о</strong><br />

не п<strong>о</strong>зднее <strong>в</strong>ерхнег<strong>о</strong> <strong>с</strong>илур<strong>а</strong>; <strong>в</strong> де<strong>в</strong><strong>о</strong>не уже<br />

<strong>в</strong><strong>с</strong>тречены <strong>к</strong>лещи, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящие<strong>с</strong>я <strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>м.<br />

Перечи<strong>с</strong>ленным д<strong>в</strong>ум <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>м н<strong>а</strong>земных<br />

хелицер<strong>а</strong>т пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<strong>с</strong>я третья rpyn- t<br />

пир<strong>о</strong><strong>в</strong><strong>к</strong><strong>а</strong> <strong>о</strong>тряд<strong>о</strong><strong>в</strong>, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>к</strong><strong>о</strong>т<strong>о</strong>рых х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я<br />

н<strong>а</strong>личием х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>ф<strong>о</strong>рмленн<strong>о</strong>й<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нением лег<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> дых<strong>а</strong>ния,<br />

н<strong>а</strong>личием н<strong>а</strong> брюш<strong>к</strong>е <strong>в</strong>ид<strong>о</strong>измененных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>в</strong> <strong>в</strong>иде п<strong>а</strong>рных генит<strong>а</strong>льных <strong>к</strong>рыше<strong>к</strong>,<br />

п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> и т. и, <strong>а</strong> т<strong>а</strong><strong>к</strong>же<br />

рез<strong>к</strong>им изгиб<strong>а</strong>нием пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong> <strong>в</strong> з<strong>а</strong>дней<br />

ч<strong>а</strong><strong>с</strong>ти, чт<strong>о</strong> при<strong>в</strong><strong>о</strong>дит <strong>к</strong> перемещению не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну и <strong>к</strong> реду<strong>к</strong>ции<br />

их <strong>с</strong>тернит<strong>о</strong><strong>в</strong>. Этими призн<strong>а</strong><strong>к</strong><strong>а</strong>ми х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я<br />

четыре <strong>в</strong>ымерших <strong>о</strong>тряд<strong>а</strong><br />

(Haptopoda, Phalangiotarbi, AnthracomaHi и<br />

Trigonotarbi) и д<strong>в</strong><strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>о</strong>тряд<strong>а</strong> (Opiliones<br />

и Araneae), <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong>бъединяют<strong>с</strong>я <strong>в</strong><br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельный <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida. С<strong>в</strong><strong>о</strong>е пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida <strong>в</strong>едет <strong>о</strong>т <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>а</strong><br />

б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нных Scorpionomorpha,<br />

<strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти, <strong>о</strong>т примити<strong>в</strong>ных ф<strong>о</strong>рм<br />

<strong>о</strong>тряд<strong>а</strong> Amblypygi и Ricinulei. Об<strong>о</strong><strong>с</strong><strong>о</strong>бление<br />

этих групп <strong>о</strong>т Scorpionomorpha пр<strong>о</strong>из<strong>о</strong>шл<strong>о</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>в</strong>ерхнем де<strong>в</strong><strong>о</strong>не и <strong>к</strong><strong>а</strong>рб<strong>о</strong>не.<br />

Предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>емые фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения<br />

<strong>о</strong>тдельных <strong>о</strong>тряд<strong>о</strong><strong>в</strong> и <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> Chelicerata<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены н<strong>а</strong> ри<strong>с</strong>. 1152. От<strong>с</strong>юд<strong>а</strong> <strong>в</strong>ыте<strong>к</strong><strong>а</strong>ют<br />

и н<strong>а</strong>ши пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ления <strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Chelicerata, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые принци­<br />

пи<strong>а</strong>льн<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующей <strong>с</strong>и<strong>с</strong>темы.<br />

Д<strong>о</strong> п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>в</strong>ремени <strong>в</strong><strong>с</strong>ех хелицер<strong>о</strong><strong>в</strong>ых<br />

жи<strong>в</strong><strong>о</strong>тных п<strong>о</strong>дтип<strong>а</strong> Chelicerata р<strong>а</strong>зделяют н<strong>а</strong><br />

три <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельных <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: Eurypterodea,<br />

Xiphosura (эти д<strong>в</strong><strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> мн<strong>о</strong>гие и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тели<br />

<strong>о</strong>бъединяли <strong>в</strong> <strong>о</strong>дин <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Merostomata) и<br />

Arachnoidea (или Arachnida). П<strong>о</strong>д<strong>о</strong>бн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>,<br />

н<strong>а</strong>пример, принят<strong>а</strong> <strong>в</strong> мн<strong>о</strong>г<strong>о</strong>т<strong>о</strong>мн<strong>о</strong>м ру<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>д<strong>с</strong>т<strong>в</strong>е<br />

«Traite de zoologie» (т. VI, <strong>с</strong>тр. 217),<br />

изд<strong>а</strong><strong>в</strong><strong>а</strong>ем<strong>о</strong>м п<strong>о</strong>д ред<strong>а</strong><strong>к</strong>цией Гр<strong>а</strong><strong>с</strong><strong>с</strong>е (Grasse,<br />

1949) и <strong>в</strong> «Treatise on invertebrate paleontology*<br />

(Petrunkevitch, 1955).<br />

Мн<strong>о</strong>гие и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тели не<strong>о</strong>дн<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong> п<strong>о</strong>дним<strong>а</strong>ли<br />

<strong>в</strong><strong>о</strong>пр<strong>о</strong><strong>с</strong> <strong>о</strong> не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>ти пере<strong>с</strong>м<strong>о</strong>треть<br />

<strong>с</strong>и<strong>с</strong>тему Chelicerata, <strong>о</strong>тмеч<strong>а</strong>я <strong>с</strong>б<strong>о</strong>рный х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<br />

эт<strong>о</strong>й группир<strong>о</strong><strong>в</strong><strong>к</strong>и. Т<strong>а</strong><strong>к</strong>, <strong>в</strong> 1949 г. <strong>а</strong>мери<strong>к</strong><strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>г Петрун<strong>к</strong>е<strong>в</strong>ич (Petrunkevitch)<br />

предпринял пер<strong>в</strong>ую п<strong>о</strong>пыт<strong>к</strong>у улучшить<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующую <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию Chelicerata.<br />

Он <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил р<strong>а</strong>зделение п<strong>о</strong>дтип<strong>а</strong> Chelicerata<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> «<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>» — <strong>в</strong><strong>о</strong>дных Merostomata и н<strong>а</strong>земных<br />

Arachnoidea — и предл<strong>о</strong>жил лишь<br />

р<strong>а</strong>зделить <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Arachnoidea н<strong>а</strong> четыре п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>:<br />

Latigastra (<strong>с</strong> <strong>о</strong>тряд<strong>а</strong>ми Scorpiones,<br />

Pseudoscorpiones, Opiliones, Architarbi u Acari),<br />

Stethostomata (<strong>с</strong> <strong>о</strong>тряд<strong>а</strong>ми Haptopoda,<br />

Anthracomarti), Soluta (<strong>с</strong> <strong>о</strong>дним <strong>о</strong>тряд<strong>о</strong>м Trigonotarbi)<br />

и Caulogastra, р<strong>а</strong><strong>с</strong>п<strong>а</strong>д<strong>а</strong>ющий<strong>с</strong>я н<strong>а</strong><br />

пять <strong>в</strong>ет<strong>в</strong>ей: Latisterna (Palpigradi), Camarostomata<br />

(Uropygi, Kustarachnae), Labellata<br />

(Phrynichida, Araneae),. Rostrata (Solifugae)<br />

u Cucullifera (Ricinulei).<br />

Эту <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ци<strong>о</strong>нную <strong>с</strong>хему <strong>а</strong><strong>в</strong>т<strong>о</strong>р принял<br />

и <strong>в</strong> п<strong>о</strong><strong>с</strong>ледующих <strong>с</strong><strong>в</strong><strong>о</strong>д<strong>к</strong><strong>а</strong>х (Petrunkevitch,<br />

1950, 1953, 1955), ее придержи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я и В<strong>а</strong>терл<strong>о</strong><br />

и (Waterlot, 1953). Н<strong>а</strong>иб<strong>о</strong>лее з<strong>а</strong>метным нед<strong>о</strong>чет<strong>о</strong>м<br />

д<strong>а</strong>нн<strong>о</strong>й <strong>с</strong>и<strong>с</strong>темы я<strong>в</strong>ляет<strong>с</strong>я р<strong>а</strong>з<strong>о</strong>бщение<br />

мн<strong>о</strong>гих близ<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енных <strong>о</strong>тряд<strong>о</strong><strong>в</strong>, н<strong>а</strong>пример,<br />

Opiliones и Haptopoda, или Anthracomarti<br />

и Trigonotarbi, или Acarina и Solifugae и<br />

т. д. <strong>в</strong> р<strong>а</strong>зные п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы. Н<strong>о</strong> г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong> б<strong>о</strong>лее<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енным нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong>м .эт<strong>о</strong>й <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции<br />

Петрун<strong>к</strong>е<strong>в</strong>ич<strong>а</strong>, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й ф<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong>и<br />

лишь н<strong>а</strong> учете ф<strong>о</strong>рмы брюш<strong>к</strong><strong>а</strong> и <strong>о</strong>бщег<strong>о</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<strong>а</strong><br />

<strong>с</strong>тр<strong>о</strong>ения р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

игн<strong>о</strong>рир<strong>о</strong><strong>в</strong><strong>а</strong>ние фил<strong>о</strong>гении, именн<strong>о</strong> —<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>о</strong>бширн<strong>о</strong>г<strong>о</strong> <strong>а</strong>р<strong>с</strong>ен<strong>а</strong>л<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>их<br />

и эмбри<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их д<strong>а</strong>нных п<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременным й и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емым.<br />

Вт<strong>о</strong>р<strong>а</strong>я п<strong>о</strong>пыт<strong>к</strong><strong>а</strong> уп<strong>о</strong>ряд<strong>о</strong>чить <strong>с</strong>и<strong>с</strong>тему<br />

Chelicerata был<strong>а</strong> <strong>с</strong>дел<strong>а</strong>н<strong>а</strong> А- А. З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>а</strong>нным<br />

(1952 <strong>а</strong>, б), <strong>к</strong><strong>о</strong>т<strong>о</strong>рый н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии изучения<br />

л<strong>о</strong><strong>к</strong><strong>а</strong>лиз<strong>а</strong>ции <strong>о</strong>птиче<strong>с</strong><strong>к</strong>и <strong>а</strong>низ<strong>о</strong>тр<strong>о</strong>пн<strong>о</strong>г<strong>о</strong> <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>в</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>х р<strong>а</strong>зличных хелицер<strong>о</strong><strong>в</strong>ых и<br />

и<strong>с</strong>п<strong>о</strong>льз<strong>о</strong><strong>в</strong><strong>а</strong>ния не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых м<strong>а</strong>тери<strong>а</strong>л<strong>о</strong><strong>в</strong> п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>й<br />

<strong>а</strong>н<strong>а</strong>т<strong>о</strong>мии (гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м <strong>к</strong>лещей)<br />

р<strong>а</strong>зделил <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Chelicerata н<strong>а</strong> четыре<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>. Н<strong>а</strong>иб<strong>о</strong>лее <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енным улучше-<br />

383<br />

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ARACHNIDA SCORPIONOMORPHA<br />

-Л<br />

-J <strong>о</strong><br />

CO »*<br />

TRILOBITOMORPHA<br />

ACAROMORPHA<br />

A<br />

Ри<strong>с</strong>. 1152. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>, п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> и <strong>о</strong>тряд<strong>о</strong><strong>в</strong> хелицер<strong>о</strong><strong>в</strong>ых (Дубинин, 1959) Н<strong>а</strong><br />

ф<strong>о</strong>не ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>л<strong>о</strong>н<strong>к</strong>и п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>о</strong> <strong>в</strong>ремя п<strong>о</strong>я<strong>в</strong>ления, р<strong>а</strong>з<strong>в</strong>ития и р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нения <strong>о</strong>тдельных <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>,<br />

п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> и <strong>о</strong>тряд<strong>о</strong><strong>в</strong>.<br />

384<br />

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нием <strong>с</strong>и<strong>с</strong>темы я<strong>в</strong>ил<strong>о</strong><strong>с</strong>ь <strong>о</strong>бъединение <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

Eurypterida (= Gigantostraca), Scorpiones,<br />

Telyphones (= Holopeltidia) <strong>в</strong> <strong>о</strong>дн<strong>о</strong>м н<strong>а</strong>д<strong>о</strong>тряде<br />

Holactinochitinosi Zachvatkin. Эт<strong>о</strong>т н<strong>а</strong>д<strong>о</strong>тряд<br />

(п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> <strong>в</strong> н<strong>а</strong>шем п<strong>о</strong>ним<strong>а</strong>нии) <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> (н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>) Chelicerata р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong><strong>с</strong>лед з<strong>а</strong> н<strong>а</strong>д<strong>о</strong>тряд<strong>о</strong>м (<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong>м) Merostomata.<br />

Д<strong>в</strong><strong>а</strong> других н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong> <strong>с</strong>и<strong>с</strong>темы Chelicerata, п<strong>о</strong><br />

З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ину (1952) —Actinochaeta и Actinoderma,<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>ли<strong>с</strong>ь <strong>с</strong>б<strong>о</strong>рными, т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>тряды<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных <strong>в</strong> них р<strong>а</strong><strong>с</strong>пределены н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> лишь ф<strong>о</strong>рм<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> призн<strong>а</strong><strong>к</strong><strong>а</strong> —<br />

н<strong>а</strong>личия или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ия <strong>о</strong>птиче<strong>с</strong><strong>к</strong>и <strong>а</strong>низ<strong>о</strong>тр<strong>о</strong>пн<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>х или т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong> <strong>о</strong><strong>с</strong>е<strong>в</strong><strong>о</strong>м <strong>с</strong>тержне щетин<strong>о</strong><strong>к</strong>. В<strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие эт<strong>о</strong>г<strong>о</strong>,<br />

н<strong>а</strong>пример, <strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нные З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>иным три<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельные <strong>о</strong>тряд<strong>а</strong> <strong>к</strong>лещей «<strong>о</strong>тряд<strong>а</strong>»<br />

Acarina—Acariformes, Parasitiformes и Opilioacarina<br />

— <strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ли<strong>с</strong>ь р<strong>а</strong>з<strong>о</strong>бщенными <strong>в</strong> р<strong>а</strong>зных<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>х.<br />

В пр<strong>о</strong><strong>в</strong>еденных и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниях <strong>а</strong><strong>в</strong>т<strong>о</strong>р<strong>о</strong>м не<br />

были д<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> <strong>о</strong>ценены принципи<strong>а</strong>льные <strong>о</strong>тличия<br />

я<strong>с</strong>н<strong>о</strong> <strong>о</strong>ф<strong>о</strong>рмленн<strong>о</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> Arachnida и Scorpionomorpha<br />

<strong>о</strong>т л<strong>о</strong>жн<strong>о</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы (пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> з<strong>а</strong> <strong>в</strong>ычет<strong>о</strong>м<br />

гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы) <strong>к</strong>лещей или пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

ф<strong>а</strong>л<strong>а</strong>нг.<br />

Си<strong>с</strong>тему З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин<strong>а</strong> принял В. Н. Бе<strong>к</strong>лемише<strong>в</strong><br />

(1952), <strong>в</strong>ы<strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей интере<strong>с</strong>н<strong>о</strong>й<br />

<strong>к</strong>ниге ряд з<strong>а</strong>меч<strong>а</strong>ний п<strong>о</strong> п<strong>о</strong><strong>в</strong><strong>о</strong>ду нее.<br />

Предл<strong>а</strong>г<strong>а</strong>ем<strong>а</strong>я ниже н<strong>о</strong><strong>в</strong><strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> хелицер<strong>о</strong><strong>в</strong>ых<br />

жи<strong>в</strong><strong>о</strong>тных н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Chelicerata у<strong>с</strong>тр<strong>а</strong>няет<br />

нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и предыдущих <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ций<br />

и, <strong>в</strong>идим<strong>о</strong>, б<strong>о</strong>лее п<strong>о</strong>лн<strong>о</strong> <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ет н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ления<br />

и эт<strong>а</strong>пы фил<strong>о</strong>генез<strong>а</strong> (Дубинин, 1959).<br />

П<strong>о</strong>дтип 'Chelicerophora<br />

Н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Chelicerata<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Merostomata<br />

Отряд Aglaspidida<br />

Отряд Chasmataspidida<br />

Отряд Synziphosura<br />

Отряд Limulida<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Scorpionomorpha<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Holactinochitinosi<br />

Отряд Eurypterida<br />

П<strong>о</strong>д<strong>о</strong>тряд Eurypterina<br />

П<strong>о</strong>д<strong>о</strong>тряд Stylonurina<br />

Отряд Scorpionida<br />

25'<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

http://jurassic.ru/<br />

П<strong>о</strong>д<strong>о</strong>тряд Apoxypodes<br />

П<strong>о</strong>д<strong>о</strong>тряд Dionychopodes<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Pedipalpides<br />

Отряд Uropygi<br />

П<strong>о</strong>д<strong>о</strong>тряд Schizopeltidia<br />

П<strong>о</strong>д<strong>о</strong>тряд Holopeltidia<br />

Отряд AmbJypygi<br />

Отряд Kustarachnida<br />

Отряд Palpigradi<br />

Отряд Pseudoscorpionodea<br />

П<strong>о</strong>д<strong>о</strong>тряд Chthoniinea<br />

П<strong>о</strong>д<strong>о</strong>тряд Neobisiinea<br />

П<strong>о</strong>д<strong>о</strong>тряд Cheliferinea<br />

Отряд Ricinulei<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha<br />

Отряд Solifugae<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Acaromorpha<br />

Отряд Acariformes<br />

П<strong>о</strong>д<strong>о</strong>тряд Palaeacariformes<br />

П<strong>о</strong>д<strong>о</strong>тряд Oribatei<br />

П<strong>о</strong>д<strong>о</strong>тр яд Acar i d i <strong>а</strong>е<br />

П<strong>о</strong>д<strong>о</strong>тряд Trombidiiformes<br />

П<strong>о</strong>д<strong>о</strong>тряд Tetrapodili<br />

Отряд Parasitiformes<br />

П<strong>о</strong>д<strong>о</strong>тряд Holothyroidea<br />

П<strong>о</strong>д<strong>о</strong>тряд Mesostigmata<br />

П<strong>о</strong>д<strong>о</strong>тряд Ixodides<br />

Отряд Opilioacarina<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Opiliomorphae<br />

Отряд Haptopoda<br />

Отряд Phalangiotarbi<br />

Отряд Opiliones<br />

П<strong>о</strong>д<strong>о</strong>тряд Cyphophthalmi<br />

П<strong>о</strong>д<strong>о</strong>тряд Laniatores<br />

П<strong>о</strong>д<strong>о</strong>тряд Palpatores<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Soluta<br />

Отряд Trigonotarbi<br />

Отряд Anthracomarti<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Araneae<br />

Отряд Liphistiomorphae<br />

Отряд Arachnomorphae<br />

Отряд Mygalomorphae<br />

П<strong>о</strong>д<strong>о</strong>тряд Nelipoda<br />

П<strong>о</strong>д<strong>о</strong>тряд Hypodemata<br />

П<strong>о</strong>д<strong>о</strong>тряд Pycnotheloides<br />

Отряд Araneomorphae<br />

П<strong>о</strong>д<strong>о</strong>тряд Ecribellatae<br />

П<strong>о</strong>д<strong>о</strong>тряд Cribellatae

КЛАСС MEROSTOMATA. МЕЧЕХВОСТЫ<br />

В<strong>о</strong>дные хелицер<strong>о</strong><strong>в</strong>ые. Р<strong>а</strong>змеры тел<strong>а</strong> у <strong>с</strong><strong>а</strong>мых<br />

мел<strong>к</strong>их п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм <strong>о</strong>т 1—3 <strong>с</strong>м, у<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных д<strong>о</strong> 60 <strong>с</strong>м. Тел<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong><br />

переднюю, <strong>о</strong>бычн<strong>о</strong> <strong>о</strong>чень <strong>к</strong>рупную ч<strong>а</strong><strong>с</strong>ть —<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>му (или г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь) и з<strong>а</strong>днюю — <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му<br />

(или <strong>а</strong>бд<strong>о</strong>мен, брюш<strong>к</strong><strong>о</strong>), <strong>с</strong> м<strong>о</strong>щным<br />

<strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>зным тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

ин<strong>о</strong>гд<strong>а</strong> делит<strong>с</strong>я н<strong>а</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>му (передняя ч<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями) и мет<strong>а</strong><strong>с</strong><strong>о</strong>му (з<strong>а</strong>дняя ч<strong>а</strong><strong>с</strong>ть,<br />

без <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее или менее<br />

<strong>о</strong><strong>к</strong>ругл<strong>о</strong>г<strong>о</strong> или шлем<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> з<strong>а</strong>днеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми (ген<strong>а</strong>льные<br />

или щечные углы), н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

и н<strong>а</strong>з<strong>а</strong>д. Ред<strong>к</strong><strong>о</strong> эти <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

(Liomesaspididae). Н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме р<strong>а</strong>з<strong>в</strong>иты <strong>с</strong>рединные<br />

гл<strong>а</strong>з<strong>к</strong>и и <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> (реже <strong>о</strong>ни <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют—<br />

Synziphosura) и ше<strong>с</strong>ть п<strong>а</strong>р <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пер<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> — хелицеры,<br />

не<strong>с</strong>ущие <strong>к</strong>лешни, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> тремя или<br />

четырьмя члени<strong>к</strong><strong>а</strong>ми. Другие <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>в</strong><strong>к</strong>люч<strong>а</strong>я педип<strong>а</strong>льпы (<strong>в</strong>т<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong>),<br />

<strong>с</strong>луж<strong>а</strong>т х<strong>о</strong>дильными н<strong>о</strong>г<strong>а</strong>ми; их пр<strong>о</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льные<br />

члени<strong>к</strong>и (<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ы или т<strong>а</strong>зи<strong>к</strong>и) ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> и<strong>с</strong>п<strong>о</strong>лняют<br />

фун<strong>к</strong>цию челю<strong>с</strong>тей, р<strong>а</strong>змельч<strong>а</strong>я<br />

пищу. Сегменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы изменяют<strong>с</strong>я <strong>в</strong><br />

чи<strong>с</strong>ле и <strong>в</strong> т<strong>о</strong>й или ин<strong>о</strong>й <strong>с</strong>тепени редуцируют<strong>с</strong>я<br />

или <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я. Ср<strong>а</strong><strong>с</strong>т<strong>а</strong>ние <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong>иб<strong>о</strong>лее<br />

п<strong>о</strong>лн<strong>о</strong> у Limulida. Прид<strong>а</strong>т<strong>к</strong>и ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong> пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong>д пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; <strong>о</strong>ни <strong>о</strong>бр<strong>а</strong>зуют<br />

п<strong>а</strong>ру пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong> — т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емые хил<strong>а</strong>рии<br />

(или мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong>). Пер<strong>в</strong><strong>о</strong>й п<strong>а</strong>р<strong>о</strong>й <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>а</strong> именн<strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> тел<strong>а</strong>,<br />

я<strong>в</strong>ляет<strong>с</strong>я <strong>о</strong>пер<strong>к</strong>улум, не<strong>с</strong>ущий п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия.<br />

Другие <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные<br />

и д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тые; <strong>к</strong><strong>а</strong>жд<strong>а</strong>я <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из мн<strong>о</strong>гих редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нных <strong>с</strong>рединных и б<strong>о</strong><strong>к</strong><strong>о</strong>­<br />

(Не<strong>с</strong>т<strong>о</strong>р Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>)<br />

386<br />

http://jurassic.ru/<br />

<strong>в</strong>ых, <strong>с</strong>ильн<strong>о</strong> упл<strong>о</strong>щенных <strong>в</strong>ет<strong>в</strong>ей, не<strong>с</strong>ущих<br />

ж<strong>а</strong>бры.<br />

М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия. Тел<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><br />

п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> пр<strong>о</strong>чным, б<strong>о</strong>лее или менее минер<strong>а</strong>лиз<strong>о</strong><strong>в</strong><strong>а</strong>нным<br />

хитин<strong>о</strong><strong>в</strong>ым п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong>м (э<strong>к</strong>з<strong>о</strong><strong>с</strong><strong>к</strong>елет).<br />

Пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>ть п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> других, <strong>в</strong>ымерших<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>, т<strong>а</strong><strong>к</strong>же был<strong>а</strong> <strong>в</strong>ели<strong>к</strong><strong>а</strong>.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее или менее п<strong>о</strong>лу<strong>к</strong>ругл<strong>а</strong>я или<br />

шлем<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния, упл<strong>о</strong>щенн<strong>а</strong>я или<br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я. Н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

(ри<strong>с</strong>. 1153) имеет<strong>с</strong>я <strong>с</strong>рединн<strong>а</strong>я <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>а</strong>я<br />

д<strong>о</strong>ля (или гл<strong>а</strong>беллярн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть, <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong>) <strong>с</strong><strong>о</strong> <strong>с</strong>рединным<br />

гребнем, н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м у м<strong>о</strong>л<strong>о</strong>дых <strong>о</strong><strong>с</strong><strong>о</strong>бей<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>г<strong>о</strong> мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>а</strong> р<strong>а</strong>з<strong>в</strong>иты шипы.<br />

Впереди <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й д<strong>о</strong>ли р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>с</strong>рединные<br />

гл<strong>а</strong>з<strong>к</strong>и. Сл<strong>о</strong>жные б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>мещ<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>бычн<strong>о</strong> близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>к</strong>р<strong>а</strong>ю пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. Вперед<br />

и н<strong>а</strong>з<strong>а</strong>д <strong>о</strong>т <strong>к</strong><strong>а</strong>жд<strong>о</strong>г<strong>о</strong> <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> гл<strong>а</strong>з<strong>а</strong> ин<strong>о</strong>гд<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ет<strong>с</strong>я гребень или ребр<strong>о</strong> (гл<strong>а</strong>зн<strong>о</strong>й гребень<br />

или гл<strong>а</strong>зн<strong>о</strong>е ребр<strong>о</strong>), не<strong>с</strong>ущие у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

ф<strong>о</strong>рм (н<strong>а</strong>пример, у Euproopidae) или м<strong>о</strong>л<strong>о</strong>дых<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бей шипы, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленные н<strong>а</strong>з<strong>а</strong>д. Пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

между гл<strong>а</strong>зными ребр<strong>а</strong>ми н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>к</strong><strong>а</strong>рди<strong>о</strong>фт<strong>а</strong>льмиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью, <strong>а</strong> пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

между <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й д<strong>о</strong>лей и <strong>к</strong><strong>а</strong>ждым гл<strong>а</strong>зным<br />

ребр<strong>о</strong>м — межгл<strong>а</strong>зн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью (<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

<strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует гл<strong>а</strong>беллярн<strong>а</strong>я пл<strong>о</strong>щ<strong>а</strong>д<strong>к</strong><strong>а</strong>). Р<strong>а</strong>змещение<br />

му<strong>с</strong><strong>к</strong>ул<strong>о</strong><strong>в</strong> <strong>в</strong> межгл<strong>а</strong>зн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти и<br />

<strong>о</strong>тч<strong>а</strong><strong>с</strong>ти <strong>в</strong> <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й д<strong>о</strong>ле <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ет <strong>с</strong>егмент<strong>а</strong>цию<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>тделены н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не п<strong>о</strong>перечным ш<strong>в</strong><strong>о</strong>м. Эт<strong>о</strong>т ш<strong>о</strong><strong>в</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и<br />

пере<strong>с</strong>е<strong>к</strong><strong>а</strong>ет <strong>с</strong>едьм<strong>о</strong>й <strong>с</strong>егмент, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

зн<strong>а</strong>чительн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н. Шир<strong>о</strong><strong>к</strong>ий<br />

<strong>о</strong>б<strong>о</strong>д, или лимб пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong><strong>к</strong>руж<strong>а</strong>ющий <strong>к</strong><strong>а</strong>рди<strong>о</strong>фт<strong>а</strong>льмиче<strong>с</strong><strong>к</strong>ую<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, прин<strong>а</strong>длежит, п<strong>о</strong><br />

<strong>в</strong><strong>с</strong>ей <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти, <strong>к</strong> пер<strong>в</strong><strong>о</strong>му <strong>с</strong>егменту.

С.ID<br />

Ри<strong>с</strong> 1153—1154. М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>с</strong><strong>о</strong><strong>в</strong>ременных мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> (St0rmer, 1955)<br />

*<br />

1153. Tachypleus gigas (Muller), д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть м<strong>о</strong>л<strong>о</strong>д<strong>о</strong>г<strong>о</strong><br />

инди<strong>в</strong>ид<strong>а</strong> <strong>с</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>нн<strong>о</strong>й <strong>с</strong>егмент<strong>а</strong>цией. 1154. Limulus polyphemus<br />

(Linnaeus); <strong>в</strong>ентр<strong>а</strong>льн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>. I VI <strong>с</strong>егменты<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; VII — XII — <strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы; м. <strong>о</strong>. — му<strong>с</strong><strong>к</strong>ульные<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong>и; <strong>с</strong>. г.— <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>; <strong>с</strong>. ш. — <strong>с</strong><strong>о</strong>член<strong>о</strong><strong>в</strong>ный ш<strong>о</strong><strong>в</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы; т.— _„ тель<strong>с</strong><strong>о</strong>н; II т т — V Л7 — - пер<strong>в</strong><strong>а</strong>я чет<strong>в</strong>ерт<strong>а</strong>я<br />

х<strong>о</strong>дильные н<strong>о</strong>ги; VI - пят<strong>а</strong>я п<strong>а</strong>р<strong>а</strong> х<strong>о</strong>дильных н<strong>о</strong>г; х —<br />

хил<strong>а</strong>рий (прин<strong>а</strong>длежит VI <strong>с</strong>егменту пр<strong>о</strong><strong>с</strong><strong>о</strong>мы); <strong>о</strong> - <strong>о</strong>пер<strong>к</strong>улум;<br />

р - р<strong>о</strong>т; л - л<strong>а</strong>брум; <strong>в</strong> - <strong>в</strong>ентр<strong>а</strong>льный <strong>о</strong>б<strong>о</strong>нятельный<br />

<strong>о</strong>рг<strong>а</strong>н; <strong>о</strong>пи<strong>с</strong>.— <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>. Сильн<strong>о</strong> уменьшен<strong>о</strong><br />

С <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны пр<strong>о</strong><strong>с</strong><strong>о</strong>мы (ри<strong>с</strong>.<br />

1154) п<strong>о</strong> ее <strong>к</strong>р<strong>а</strong>ям имеет<strong>с</strong>я дублюр<strong>а</strong>, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляющ<strong>а</strong>я<br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>й з<strong>а</strong>гнутую <strong>в</strong>низ и <strong>в</strong>нутрь <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ую<br />

ч<strong>а</strong><strong>с</strong>ть д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> э<strong>к</strong>з<strong>о</strong><strong>с</strong><strong>к</strong>елет<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Уз<strong>к</strong><strong>а</strong>я <strong>в</strong>ерхняя губ<strong>а</strong> (labrum) <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет<br />

р<strong>о</strong>т <strong>с</strong>переди. У <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния губы р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен неб<strong>о</strong>льш<strong>о</strong>й<br />

<strong>о</strong><strong>к</strong>руглый <strong>о</strong>б<strong>о</strong>нятельный <strong>о</strong>рг<strong>а</strong>н. Сз<strong>а</strong>ди<br />

р<strong>о</strong>т <strong>о</strong><strong>к</strong><strong>а</strong>ймлен неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>нутренней пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>о</strong>й<br />

(энд<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>й). К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

(ри<strong>с</strong>. 1155) т<strong>о</strong>н<strong>к</strong>ие, члени<strong>с</strong>тые. Хелицеры неб<strong>о</strong>льшие.<br />

Педип<strong>а</strong>льпы и четыре п<strong>а</strong>ры х<strong>о</strong>дильных<br />

н<strong>о</strong>г <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м <strong>с</strong>х<strong>о</strong>дны между <strong>с</strong><strong>о</strong>б<strong>о</strong>й.<br />

Крупные пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тые т<strong>а</strong>зи<strong>к</strong>и этих <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>с</strong>н<strong>а</strong>бжены п<strong>о</strong> <strong>с</strong>рединн<strong>о</strong>й линии<br />

шип<strong>а</strong>ми, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong>луж<strong>а</strong>т для перед<strong>в</strong>ижения<br />

ч<strong>а</strong><strong>с</strong>тиц пищи <strong>в</strong> р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие.<br />

Гребень или буг<strong>о</strong>р н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х н<strong>о</strong>ги<br />

IV <strong>с</strong>лужит для р<strong>а</strong>змельчения пищи. У Limulidae<br />

педип<strong>а</strong>льпы и н<strong>о</strong>ги III—V <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>к</strong>лешнями, н<strong>о</strong>ги VI не<strong>с</strong>ут упл<strong>о</strong>щенные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленные для <strong>к</strong><strong>о</strong>п<strong>а</strong>ния. К б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>му<br />

<strong>к</strong>р<strong>а</strong>ю <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ы н<strong>о</strong>ги VI при<strong>к</strong>реплен нечлени<strong>с</strong>тый<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й прид<strong>а</strong>т<strong>о</strong><strong>к</strong> (фл<strong>а</strong>беллум).<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> у р<strong>а</strong>зных Merostomata не<strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ения, у Limulidae <strong>о</strong>н<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зует<br />

<strong>с</strong>пл<strong>о</strong>шн<strong>о</strong>й, умеренн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый щит. Ин<strong>о</strong>гд<strong>а</strong> н<strong>а</strong><br />

387<br />

http://jurassic.ru/<br />

II-V<br />

<strong>о</strong>пи<strong>с</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я трехд<strong>о</strong>льн<strong>о</strong><strong>с</strong>ть, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

<strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чен<strong>а</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми. У Limulidae<br />

эти б<strong>о</strong>р<strong>о</strong>зды з<strong>а</strong>менены ше<strong>с</strong>тью п<strong>а</strong>р<strong>а</strong>ми<br />

<strong>в</strong>ыпячи<strong>в</strong><strong>а</strong>ний, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>уют<br />

ме<strong>с</strong>т<strong>а</strong>м при<strong>к</strong>репления мышц. Длинный <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

тель<strong>с</strong><strong>о</strong>н Limulidae причленяет<strong>с</strong>я<br />

<strong>к</strong> <strong>а</strong>бд<strong>о</strong>мену п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>с</strong>пеци<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> Т-<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>, <strong>о</strong>че<strong>в</strong>идн<strong>о</strong>, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляющег<strong>о</strong><br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>й терг<strong>а</strong>льный <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т з<strong>а</strong>днег<strong>о</strong> рудимент<strong>а</strong>рн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong>. Тель<strong>с</strong><strong>о</strong>н п<strong>о</strong>д<strong>в</strong>ижен; у <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Limulidae <strong>о</strong>н игр<strong>а</strong>ет <strong>о</strong>пределенную р<strong>о</strong>ль при<br />

перед<strong>в</strong>ижении и при з<strong>а</strong><strong>к</strong><strong>а</strong>пы<strong>в</strong><strong>а</strong>нии яиц <strong>в</strong> пе<strong>с</strong><strong>о</strong><strong>к</strong>.<br />

Р<strong>а</strong>з<strong>в</strong>итие. Онт<strong>о</strong>генез <strong>с</strong><strong>о</strong><strong>в</strong>ременных мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет б<strong>о</strong>льш<strong>о</strong>й интере<strong>с</strong> для<br />

<strong>в</strong>ыя<strong>с</strong>нения их фил<strong>о</strong>гении. Н<strong>а</strong> р<strong>а</strong>нних <strong>с</strong>т<strong>а</strong>диях<br />

р<strong>а</strong>з<strong>в</strong>ития у лимулид н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> четыре<br />

пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льные <strong>с</strong>егмент<strong>а</strong> (<strong>к</strong><strong>а</strong><strong>к</strong> у пр<strong>о</strong>т<strong>а</strong><strong>с</strong>пи<strong>с</strong><strong>а</strong><br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>). В эт<strong>о</strong>м пр<strong>о</strong>я<strong>в</strong>ляет<strong>с</strong>я фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

близ<strong>о</strong><strong>с</strong>ть хелицер<strong>о</strong><strong>в</strong>ых и трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>.<br />

В д<strong>а</strong>льнейшем р<strong>а</strong>з<strong>в</strong>итии мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты<br />

пр<strong>о</strong>х<strong>о</strong>дят <strong>с</strong>т<strong>а</strong>дию, у <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й тель<strong>с</strong><strong>о</strong>н не удлинен<br />

и <strong>о</strong>бщий <strong>о</strong>бли<strong>к</strong> жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет<br />

трил<strong>о</strong>бит<strong>а</strong>: п<strong>о</strong>т<strong>о</strong>му эту <strong>с</strong>т<strong>а</strong>дию ч<strong>а</strong><strong>с</strong>т<strong>о</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<br />

«трил<strong>о</strong>бит<strong>о</strong><strong>в</strong><strong>о</strong>й», х<strong>о</strong>тя <strong>в</strong> дей<strong>с</strong>т<strong>в</strong>ительн<strong>о</strong><strong>с</strong>ти г<strong>о</strong>р<strong>а</strong>зд<strong>о</strong><br />

б<strong>о</strong>льшее <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> имеет<strong>с</strong>я <strong>с</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ими<br />

мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>ми Euproopidae.<br />

26*

<strong>в</strong><br />

Ри<strong>с</strong>. 1155. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong> (III, VII и VIII) пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>г<strong>о</strong> мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>а</strong> Limutus<br />

polyphemus (Linnaeus) (Sttfrmer, 1955).<br />

Об<strong>о</strong>зн<strong>а</strong>чения: хе — хелицеры; <strong>в</strong> — <strong>в</strong>ентр<strong>а</strong>льный <strong>о</strong>б<strong>о</strong>нятельный<br />

<strong>о</strong>рг<strong>а</strong>н; э — энд<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong>; р — р<strong>о</strong>т; <strong>к</strong> — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ы (т<strong>а</strong>зи<strong>к</strong>и); л — л<strong>а</strong>бруи;<br />

ф — фл<strong>а</strong>беллум; х — хил<strong>а</strong>рий; п. <strong>к</strong>. — <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых<br />

<strong>к</strong><strong>а</strong>н<strong>а</strong>л<strong>о</strong><strong>в</strong>; <strong>а</strong> — <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong>, д — дублюр<strong>а</strong>; <strong>о</strong> — <strong>о</strong>пер<strong>к</strong>улюм; an —<br />

<strong>а</strong>п<strong>о</strong>дем<strong>а</strong>; ж — ж<strong>а</strong>бры; <strong>с</strong> — <strong>с</strong><strong>о</strong>членение б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и <strong>а</strong>бд<strong>о</strong>мин<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

прид<strong>а</strong>т<strong>к</strong><strong>а</strong>; б и <strong>с</strong>р — б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я и <strong>с</strong>рединн<strong>а</strong>я <strong>в</strong>ет<strong>в</strong>и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льн<strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти. Сильн<strong>о</strong> уменьшен<strong>о</strong><br />

В<strong>о</strong> <strong>в</strong>ремя линь<strong>к</strong>и н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>в</strong><strong>с</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>г<strong>о</strong> ш<strong>в</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

С<strong>о</strong><strong>в</strong>ременные мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты<br />

— м<strong>о</strong>р<strong>с</strong><strong>к</strong>ие ф<strong>о</strong>рмы. Взр<strong>о</strong><strong>с</strong>лые жи<strong>в</strong><strong>о</strong>тные<br />

п<strong>о</strong>лз<strong>а</strong>ют п<strong>о</strong> дну и з<strong>а</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong>. ил и пе<strong>с</strong><strong>о</strong><strong>к</strong>;<br />

для р<strong>а</strong>змн<strong>о</strong>жения <strong>о</strong>ни <strong>в</strong>ып<strong>о</strong>лз<strong>а</strong>ют н<strong>а</strong> берег.<br />

М<strong>о</strong>л<strong>о</strong>дые личин<strong>к</strong>и (и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>зр<strong>о</strong><strong>с</strong>лые)<br />

<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бны пл<strong>а</strong><strong>в</strong><strong>а</strong>ть н<strong>а</strong> <strong>с</strong>пине при п<strong>о</strong>м<strong>о</strong>щи <strong>в</strong>зм<strong>а</strong>х<strong>о</strong><strong>в</strong><br />

брюшных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> дре<strong>в</strong>них<br />

мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> не был<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong> м<strong>о</strong>рем, п<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>у<br />

м<strong>о</strong>жн<strong>о</strong> <strong>с</strong>удить п<strong>о</strong> их и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емым<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м, н<strong>а</strong>х<strong>о</strong>димым гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м <strong>в</strong><br />

пре<strong>с</strong>н<strong>о</strong>- и <strong>с</strong><strong>о</strong>л<strong>о</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>дных <strong>о</strong><strong>с</strong><strong>а</strong>д<strong>к</strong><strong>а</strong>х. Н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> ред<strong>к</strong>и, п<strong>о</strong>эт<strong>о</strong>му <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нению <strong>с</strong> из<strong>в</strong>е<strong>с</strong>тными р<strong>о</strong>д<strong>а</strong>ми<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> не<strong>в</strong>ели<strong>к</strong><strong>о</strong>. В предел<strong>а</strong>х СССР<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и дре<strong>в</strong>них мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> д<strong>о</strong> <strong>с</strong>ег<strong>о</strong> <strong>в</strong>ремени<br />

н<strong>а</strong>йдены <strong>в</strong> <strong>о</strong>чень <strong>о</strong>гр<strong>а</strong>ниченн<strong>о</strong>м <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е: из<strong>в</strong>е<strong>с</strong>тны<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>в</strong><strong>с</strong>ег<strong>о</strong> четырех <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Paleomeridae и Aglaspididae из <strong>о</strong>тряд<strong>а</strong> <strong>а</strong>гл<strong>а</strong><strong>с</strong>-<br />

388<br />

http://jurassic.ru/<br />

пидид, н<strong>а</strong>йденные <strong>в</strong> <strong>с</strong>реднем <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>е В. Сибири<br />

(ре<strong>к</strong>и Анг<strong>а</strong>р<strong>а</strong> и Лен<strong>а</strong>), Bunodidae из<br />

<strong>о</strong>тряд<strong>а</strong> <strong>с</strong>инзиф<strong>о</strong>зур из <strong>в</strong>. <strong>с</strong>илур<strong>а</strong> Э<strong>с</strong>т<strong>о</strong>нии и<br />

Belinuridae из <strong>о</strong>тряд<strong>а</strong> лимулид из <strong>в</strong>. де<strong>в</strong><strong>о</strong>н<strong>а</strong><br />

Липец<strong>к</strong><strong>о</strong>й <strong>о</strong>бл.<br />

Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> из<strong>в</strong>е<strong>с</strong>тных <strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нен<strong>о</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>е (т<strong>а</strong>бл. 2). Мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ие ф<strong>о</strong>рмы<br />

<strong>в</strong> неб<strong>о</strong>льш<strong>о</strong>м <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е из<strong>в</strong>е<strong>с</strong>тны из три<strong>а</strong><strong>с</strong><strong>а</strong>,<br />

юры и мел<strong>а</strong>. Из не<strong>о</strong>ген<strong>а</strong> н<strong>а</strong>йдены <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителя<br />

р<strong>о</strong>д<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны (Tachypleus<br />

Leach). Мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты, <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующие <strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее<br />

<strong>в</strong>ремя, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены тремя р<strong>о</strong>д<strong>а</strong>ми<br />

<strong>с</strong> пятью <strong>в</strong>ид<strong>а</strong>ми, <strong>о</strong>бит<strong>а</strong>ющими <strong>в</strong> мел<strong>к</strong>их <strong>в</strong><strong>о</strong>д<strong>а</strong>х<br />

<strong>в</strong>д<strong>о</strong>ль <strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>чных п<strong>о</strong>бережий Се<strong>в</strong>ерн<strong>о</strong>й Амери<strong>к</strong>и<br />

и Азии.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>иий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Общ<strong>а</strong>я<br />

ф<strong>о</strong>рм<strong>а</strong> тел<strong>а</strong>, <strong>с</strong>тр<strong>о</strong>ение пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и гл<strong>а</strong>з, <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы (<strong>о</strong>тч<strong>а</strong><strong>с</strong>ти и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы), <strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>с</strong>тр<strong>о</strong>ение нер<strong>в</strong>н<strong>о</strong>й <strong>с</strong>и<strong>с</strong>темы,<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых желез и другие м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие при-

зн<strong>а</strong><strong>к</strong>и мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> близ<strong>к</strong>и <strong>к</strong> примити<strong>в</strong>ным<br />

Trilobitomorpha. Одн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>в</strong> э<strong>в</strong><strong>о</strong>люции мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><br />

н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я <strong>с</strong>пецифиче<strong>с</strong><strong>к</strong>ие черты: <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

хелицер, реду<strong>к</strong>ция <strong>а</strong>нтеннул, <strong>в</strong>ид<strong>о</strong>изменение<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>в</strong>ыз<strong>в</strong><strong>а</strong>нн<strong>о</strong>е изгиб<strong>а</strong>нием<br />

передней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong> <strong>в</strong><strong>в</strong>ерх и<br />

н<strong>а</strong>з<strong>а</strong>д, при<strong>с</strong><strong>о</strong>единение пят<strong>о</strong>г<strong>о</strong> и ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>к</strong> пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льным четырем<br />

личин<strong>о</strong>чным <strong>с</strong>егмент<strong>а</strong>м примити<strong>в</strong>ных трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие пигидия. Оче<strong>в</strong>идн<strong>о</strong>, чт<strong>о</strong> мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят <strong>о</strong>т н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ных<br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>, <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ция <strong>к</strong><strong>о</strong>т<strong>о</strong>рых был<strong>а</strong> н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не р<strong>а</strong>з<strong>в</strong>ития н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Olenelloidea.<br />

Мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты были пер<strong>в</strong>ичными хелицер<br />

<strong>о</strong> <strong>в</strong> ы м и. В пр<strong>о</strong>це<strong>с</strong><strong>с</strong>е их р<strong>а</strong>з<strong>в</strong>ития пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<strong>о</strong><br />

и<strong>с</strong><strong>к</strong>ри<strong>в</strong>ление передней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й<br />

<strong>о</strong><strong>с</strong>и тел<strong>а</strong>, чт<strong>о</strong> при<strong>в</strong>ел<strong>о</strong> <strong>к</strong> изменению р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, р<strong>а</strong>з<strong>в</strong>итие эт<strong>о</strong>г<strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> <strong>с</strong><strong>о</strong>пр<strong>о</strong><strong>в</strong><strong>о</strong>жд<strong>а</strong>л<strong>о</strong><strong>с</strong>ь<br />

пр<strong>о</strong>це<strong>с</strong><strong>с</strong><strong>о</strong>м цеф<strong>а</strong>лиз<strong>а</strong>ции, <strong>в</strong> д<strong>а</strong>льнейшем<br />

д<strong>о</strong><strong>с</strong>тигшим <strong>с</strong><strong>в</strong><strong>о</strong>их <strong>к</strong>р<strong>а</strong>йних <strong>с</strong>т<strong>а</strong>дий.<br />

Примити<strong>в</strong>ные мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты (<strong>о</strong>тряд Aglaspidida)<br />

<strong>с</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и<br />

м<strong>а</strong>л<strong>о</strong><strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленными мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й были,<br />

<strong>о</strong>че<strong>в</strong>идн<strong>о</strong>, неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енными пред<strong>к</strong><strong>а</strong>ми т<strong>а</strong><strong>к</strong><br />

н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емых «р<strong>а</strong><strong>к</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>р<strong>о</strong>пи<strong>о</strong>н<strong>о</strong><strong>в</strong>» (Eurypterida),<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые, п<strong>о</strong> <strong>в</strong><strong>с</strong>ей <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляли<br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>й р<strong>о</strong>д<strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льную группу <strong>в</strong><strong>с</strong>ех н<strong>а</strong>земных<br />

Chelicerata. К<strong>а</strong><strong>к</strong> <strong>о</strong>тметил Ст<strong>о</strong>рмер (Stormer,<br />

1955), Paleomerus Stormer, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<br />

перех<strong>о</strong>дную ф<strong>о</strong>рму <strong>о</strong>т Merostomata,<br />

<strong>а</strong> именн<strong>о</strong> <strong>о</strong>т Aglaspididae, <strong>к</strong> Eurypterida. Призн<strong>а</strong><strong>к</strong>и<br />

перех<strong>о</strong>дн<strong>о</strong><strong>с</strong>ти между у<strong>к</strong><strong>а</strong>з<strong>а</strong>нными групп<strong>а</strong>ми<br />

еще б<strong>о</strong>лее <strong>в</strong>ыр<strong>а</strong>жены у Angarocaris Chernyshev.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция Merostomata <strong>о</strong>бычн<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>л<strong>а</strong><strong>с</strong>ь<br />

не т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их призн<strong>а</strong><strong>к</strong><strong>а</strong>х,<br />

н<strong>о</strong> и н<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зе жизни. Ст<strong>о</strong>рмер (Starmer,<br />

1944, 1955) <strong>в</strong><strong>к</strong>люч<strong>а</strong>ет <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Merostomata<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: Xiphosura {<strong>с</strong><strong>о</strong>б<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты)<br />

и Eurypterida. З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин (1952) и Дубинин<br />

(1959), <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>я<strong>с</strong>ь гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м<br />

н<strong>а</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниях, <strong>о</strong>бъединяют<br />

Eurypterida (р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>я их т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>тряд) <strong>с</strong> <strong>о</strong>тряд<strong>о</strong>м Scorpionida <strong>в</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><br />

<strong>с</strong><strong>к</strong><strong>о</strong>р<strong>о</strong>пи<strong>о</strong>н<strong>о</strong><strong>в</strong>ых — Holactinochitinosi. Эти д<strong>в</strong><strong>а</strong><br />

<strong>о</strong>тряд<strong>а</strong>, <strong>о</strong>бъединенные <strong>в</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>ых,<br />

Дубинин п<strong>о</strong>мещ<strong>а</strong>ет <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong>м<br />

жгут<strong>о</strong>н<strong>о</strong>гих Pedipalpida <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бый <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Scorpionomorpha.<br />

Дубининым зн<strong>а</strong>чительн<strong>о</strong> перер<strong>а</strong>б<strong>о</strong>т<strong>а</strong>н<strong>а</strong><br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция <strong>в</strong><strong>с</strong>ех Chelicerata, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> предл<strong>а</strong>г<strong>а</strong>ем<strong>о</strong>е<br />

им <strong>о</strong>тделение Eurypterida <strong>о</strong>т Xiphosura,<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, неуд<strong>а</strong>чн<strong>о</strong>.<br />

С<strong>о</strong>гл<strong>а</strong><strong>с</strong>н<strong>о</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции, принят<strong>о</strong>й для н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong><br />

изд<strong>а</strong>ния, Merostomata {<strong>в</strong> уз<strong>к</strong><strong>о</strong>м<br />

<strong>с</strong>мы<strong>с</strong>ле) и Eurypterida п<strong>о</strong>мещены <strong>в</strong> р<strong>а</strong>зличные<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы. Кл<strong>а</strong><strong>с</strong><strong>с</strong> Merostomata <strong>в</strong><strong>к</strong>люч<strong>а</strong>ет четыре<br />

<strong>о</strong>тряд<strong>а</strong>: Aglaspidida, Chasmataspidida, Synziphosura,<br />

Limulida.<br />

Т<strong>а</strong>блиц<strong>а</strong> 2<br />

Си<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция и <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение Merostomata<br />

Merostomata<br />

Отряд Aglaspidida<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Paleomeridae;<br />

У<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ленный ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий<br />

<strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<br />

Paleomerus H. и <strong>с</strong>р. <strong>к</strong>ембрий<br />

Angarocaris Ср. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Aglaspididae:<br />

Aglaspis В. <strong>к</strong>ембрий<br />

Girardevia Ср. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

Aglaspella В. <strong>к</strong>ембрий<br />

Aglaspoides В. <strong>к</strong>ембрий<br />

Glypharthrus В. <strong>к</strong>ембрий<br />

Uarthrus В. <strong>к</strong>ембрий<br />

Cyclopites В. <strong>к</strong>ембрий<br />

Craspedops В. <strong>к</strong>ембрий<br />

Set as pis В. <strong>к</strong>ембрий<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Strabopidae:<br />

Strabops В. <strong>к</strong>ембрий<br />

Neostrabops В. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Beckwithiidae :<br />

Beckwithia Ср. <strong>к</strong>ембрий<br />

Отряд<br />

Chasmataspidida:<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Chasmataspidae:<br />

Chasmataspis Н. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

Отряд Synziphosura<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Limuloididae:<br />

Limuloides В. <strong>с</strong>илур<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Bunodidae:<br />

Bunodes В.<strong>с</strong>илур—д<strong>а</strong>унт<strong>о</strong>н<br />

Bunaia Д<strong>а</strong>унт<strong>о</strong>н<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Neolirrrulidae:<br />

Neolimulus В.<strong>с</strong>илур—н.де<strong>в</strong><strong>о</strong>н<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pseudoniscidae:<br />

Pseudoniscus В. <strong>с</strong>илур<br />

Cyamocephatus В. <strong>с</strong>илур<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Weinberginidae:<br />

389<br />

http://jurassic.ru/<br />

Weinbergina Н. де<strong>в</strong><strong>о</strong>н

Merostomata<br />

У<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ленный ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий<br />

<strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<br />

Merostomata<br />

Т<strong>а</strong>блиц<strong>а</strong> 2 (<strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ние)<br />

У<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ленный ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий<br />

<strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<br />

Отряд Limulida<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Liomesaspididea<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Belinuridea<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomesaspididae:<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Belinuridae:<br />

Belinurus<br />

Protolimulus<br />

В.де<strong>в</strong><strong>о</strong>н—<strong>в</strong>.<strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Ср. де<strong>в</strong><strong>о</strong>н<br />

Liotnesaspis<br />

Prolimutus<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Limulidea<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Н. пермь<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Austrolimulidae:<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Paleolimulidae<br />

Austrolimulus Ср. три<strong>а</strong><strong>с</strong> Paleolimulus К<strong>а</strong>рб<strong>о</strong>н—пермь<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Euproopidae:<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesolimulidae:<br />

Euproops В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н—н. пермь<br />

Psammolimulus: Н. три<strong>а</strong><strong>с</strong><br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Elleriidae:<br />

Limulitella<br />

Mesolimulus<br />

Н.—В. три<strong>а</strong><strong>с</strong><br />

Юр<strong>а</strong>—? мел<br />

Elleria<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Kiaeriidae:<br />

В.де<strong>в</strong><strong>о</strong>н—<strong>в</strong>.<strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Limulidae:<br />

Tachypleus Ми<strong>о</strong>цен—ныне<br />

Kiaerta Д<strong>а</strong>унт<strong>о</strong>н<br />

Мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тым, б<strong>о</strong>лее или<br />

менее <strong>в</strong>ыпу<strong>к</strong>лым тел<strong>о</strong>м. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>с</strong> б<strong>о</strong>лее или менее<br />

<strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лыми <strong>с</strong>л<strong>о</strong>жными гл<strong>а</strong>з<strong>а</strong>ми. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> 11 и 12 <strong>с</strong>егмент<strong>а</strong>ми, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые п<strong>о</strong>чти<br />

<strong>в</strong><strong>с</strong>е п<strong>о</strong>д<strong>в</strong>ижны, ред<strong>к</strong><strong>о</strong> три з<strong>а</strong>дние <strong>с</strong>егмент<strong>а</strong><br />

<strong>с</strong>р<strong>о</strong><strong>с</strong>шие<strong>с</strong>я (Beckwithiidae). Трехд<strong>о</strong>льн<strong>о</strong><strong>с</strong>ть<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы не <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> или <strong>о</strong>чень нея<strong>с</strong>н<strong>а</strong>.<br />

П<strong>о</strong>д з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>тью <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы имеют<strong>с</strong>я п<strong>о</strong><strong>с</strong>т<strong>в</strong>ентр<strong>а</strong>льные<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и. Тель<strong>с</strong><strong>о</strong>н <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

б<strong>а</strong>з<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью. Хелицеры четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые;<br />

<strong>в</strong><strong>с</strong>е <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong>з<strong>в</strong>иты <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их х<strong>о</strong>дильных<br />

н<strong>о</strong>г. Н<strong>а</strong>ружн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> <strong>с</strong><br />

р<strong>а</strong>зличн<strong>о</strong>й зерни<strong>с</strong>т<strong>о</strong><strong>с</strong>тью.<br />

М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия. Агл<strong>а</strong><strong>с</strong>пидиды — <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м<br />

неб<strong>о</strong>льшие ф<strong>о</strong>рмы: <strong>в</strong> <strong>с</strong>реднем <strong>о</strong>т 2 д<strong>о</strong> 6 <strong>с</strong>м<br />

длины, х<strong>о</strong>тя из<strong>в</strong>е<strong>с</strong>тны <strong>о</strong>тдельные э<strong>к</strong>земпляры<br />

длиннее 21 <strong>с</strong>м. Тел<strong>о</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>е, б<strong>о</strong>лее или<br />

менее <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, без н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих <strong>о</strong><strong>с</strong>е<strong>в</strong>ых б<strong>о</strong>р<strong>о</strong>зд,<br />

н<strong>о</strong> б<strong>о</strong>лее или менее <strong>о</strong>тчетли<strong>в</strong><strong>а</strong>я трехд<strong>о</strong>льн<strong>о</strong><strong>с</strong>ть<br />

н<strong>а</strong>меч<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> <strong>в</strong>иде р<strong>а</strong>зличн<strong>о</strong>й <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ти <strong>о</strong><strong>с</strong>е<strong>в</strong><strong>о</strong>й<br />

и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>о</strong>бл<strong>а</strong><strong>с</strong>тей. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

<strong>к</strong>рупн<strong>а</strong>я и шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>р<strong>а</strong>з<strong>в</strong>итыми<br />

ген<strong>а</strong>льными угл<strong>а</strong>ми. Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены близ<strong>к</strong><strong>о</strong> <strong>к</strong> <strong>с</strong>рединн<strong>о</strong>й<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й линии и <strong>к</strong> переднему <strong>к</strong>р<strong>а</strong>ю. Гл<strong>а</strong>з­<br />

ОТРЯД AGLASP1DIDA<br />

(Aglaspida)<br />

390<br />

http://jurassic.ru/<br />

<strong>к</strong>и м<strong>о</strong>гут быть р<strong>а</strong>з<strong>в</strong>иты, н<strong>о</strong> не <strong>в</strong><strong>с</strong>егд<strong>а</strong> <strong>в</strong>идимы.<br />

Гл<strong>а</strong>беллярн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть между гл<strong>а</strong>з<strong>а</strong>ми и з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>уют межгл<strong>а</strong>зн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

б<strong>о</strong>лее п<strong>о</strong>здних ф<strong>о</strong>рм; эт<strong>а</strong> <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>в</strong><strong>о</strong>з<strong>в</strong>ыш<strong>а</strong>ет<strong>с</strong>я<br />

н<strong>а</strong>д б<strong>о</strong>лее упл<strong>о</strong>щенн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Гл<strong>а</strong>беллярн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть м<strong>о</strong>жет быть д<strong>о</strong>льч<strong>а</strong>т<strong>о</strong>й<br />

(ри<strong>с</strong>. 1156), б<strong>о</strong>лее или менее треуг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>черт<strong>а</strong>ния <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>й з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>тью. Пле<strong>в</strong>ры<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> пле<strong>в</strong>р<strong>а</strong>льными <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> пере<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют друг друг<strong>а</strong>. Из<strong>о</strong>гнутые<br />

пле<strong>в</strong>р<strong>а</strong>льные ребр<strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют п<strong>о</strong>перечн<strong>о</strong>й<br />

<strong>с</strong><strong>о</strong>член<strong>о</strong><strong>в</strong>н<strong>о</strong>й линии, чт<strong>о</strong> <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует<br />

<strong>о</strong> примити<strong>в</strong>н<strong>о</strong>м <strong>с</strong><strong>о</strong>членении <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, тергиты<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых м<strong>о</strong>гут <strong>в</strong>д<strong>в</strong>иг<strong>а</strong>ть<strong>с</strong>я друг <strong>в</strong> друг<strong>а</strong><br />

(ри<strong>с</strong>. 1157). П<strong>о</strong>др<strong>а</strong>зделения <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong><br />

мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>му не н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я, з<strong>а</strong> и<strong>с</strong><strong>к</strong>лючением<br />

Beckwithiidae, у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бляет<strong>с</strong>я<br />

з<strong>а</strong>дний п<strong>о</strong>лу<strong>о</strong><strong>к</strong>руглый щит, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующий<br />

пигидию трил<strong>о</strong>бит<strong>о</strong><strong>в</strong> или мет<strong>а</strong><strong>с</strong><strong>о</strong>ме.<br />

Н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong>з<strong>а</strong>ди<br />

дублюры р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> четырех<strong>с</strong>т<strong>о</strong>р<strong>о</strong>нняя пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует<br />

эпи<strong>с</strong>т<strong>о</strong>ме. П<strong>о</strong>з<strong>а</strong>ди п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> и <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния тель<strong>с</strong><strong>о</strong>н<strong>а</strong> имеют<strong>с</strong>я<br />

<strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong><strong>с</strong>т<strong>в</strong>ентр<strong>а</strong>льные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и<br />

(ри<strong>с</strong>. 1158); м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>а</strong>я прир<strong>о</strong>д<strong>а</strong> этих<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ний нея<strong>с</strong>н<strong>а</strong>.

1158<br />

Ри<strong>с</strong>. 1156—1160. М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>о</strong>тряд<strong>а</strong><br />

1156. Craspedops modest a Raasch, пр<strong>о</strong><strong>с</strong><strong>о</strong>и<strong>а</strong>: п. д.—переднее п<strong>о</strong>нижение;<br />

<strong>с</strong>.г. — <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>; г — гл<strong>а</strong>беллярн<strong>а</strong>я д<strong>о</strong>ля; х2 (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция).<br />

1157. Aglaspis splnlfer (Raasch); <strong>с</strong><strong>о</strong>члененные<br />

•<strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> пле<strong>в</strong>р<strong>а</strong>ми н<strong>а</strong> <strong>к</strong>р<strong>а</strong>ях. 1158. Uarthrus Instabi-<br />

Us Raasch: n.n.— п<strong>о</strong><strong>с</strong>т<strong>в</strong>ентр<strong>а</strong>льные пл<strong>а</strong><strong>с</strong>тнн<strong>к</strong>н; <strong>с</strong> — 11-й <strong>с</strong>егмент!<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> незн<strong>а</strong>чительн<strong>о</strong>й <strong>с</strong>тепени<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у р<strong>о</strong>д<strong>а</strong> Aglaspis Hall (ри<strong>с</strong>. 1159—<br />

1160). Четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые хелицеры <strong>с</strong>н<strong>а</strong>бжены<br />

<strong>к</strong>лешней; <strong>в</strong><strong>с</strong>е другие <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленные для х<strong>о</strong>ждения, пр<strong>о</strong><strong>с</strong>тые, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Пер<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы х<strong>о</strong>дильн<strong>а</strong>я, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я; эт<strong>о</strong><br />

у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет н<strong>а</strong> примити<strong>в</strong>ный х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер Aglaspididae<br />

п<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нению <strong>с</strong> б<strong>о</strong>лее п<strong>о</strong>здними Merostomata,<br />

у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>о</strong>ги <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы не при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены<br />

для х<strong>о</strong>ждения, <strong>а</strong> пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идные<br />

прид<strong>а</strong>т<strong>к</strong>и, ж<strong>а</strong>бры. С<strong>к</strong>ульптур<strong>а</strong>,<br />

р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я н<strong>а</strong> э<strong>к</strong>з<strong>о</strong><strong>с</strong><strong>к</strong>елете <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидид, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong><br />

буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми р<strong>а</strong>зличн<strong>о</strong>й <strong>в</strong>еличины, ин<strong>о</strong>гд<strong>а</strong><br />

имеющими <strong>в</strong>ид пузырь<strong>к</strong><strong>о</strong><strong>в</strong>, или м<strong>о</strong>рщин<strong>к</strong><strong>а</strong>ми,<br />

п<strong>о</strong>х<strong>о</strong>жими н<strong>а</strong> <strong>в</strong>етр<strong>о</strong><strong>в</strong>ые зн<strong>а</strong><strong>к</strong>и н<strong>а</strong> пе<strong>с</strong><strong>к</strong>е.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. П<strong>о</strong> ф<strong>о</strong>рме<br />

тел<strong>а</strong> <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидид н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е<br />

<strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> между ними и трил<strong>о</strong>бит<strong>а</strong>ми и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

<strong>с</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми мер<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бными трил<strong>о</strong>бит<strong>о</strong>м<strong>о</strong>рф<strong>а</strong>ми,<br />

<strong>а</strong> именн<strong>о</strong> <strong>с</strong> Leanchoillidae и<br />

Emeraldellidae. Одн<strong>а</strong><strong>к</strong><strong>о</strong> у этих ф<strong>о</strong>рм г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

<strong>о</strong>тдел т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> четырьмя п<strong>а</strong>р<strong>а</strong>ми <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

п<strong>о</strong>з<strong>а</strong>ди рт<strong>а</strong>, <strong>в</strong> т<strong>о</strong> <strong>в</strong>ремя <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидид их<br />

имеет<strong>с</strong>я ше<strong>с</strong>ть. От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие у <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидид <strong>а</strong>нтенн,<br />

<strong>к</strong><strong>а</strong><strong>к</strong>ие р<strong>а</strong>з<strong>в</strong>иты у трил<strong>о</strong>бит<strong>о</strong><strong>в</strong> и мер<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных,<br />

т<strong>а</strong><strong>к</strong>же <strong>о</strong>тлич<strong>а</strong>ют Aglaspidida <strong>о</strong>т<br />

тех и других. Среди мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> <strong>с</strong> Aglaspidida<br />

имеют мн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бщег<strong>о</strong> Synziphosura, т<strong>а</strong><strong>к</strong>ие, <strong>к</strong><strong>а</strong><strong>к</strong><br />

U60G<br />

Aglaspidida, <strong>в</strong>. <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong> (St0rmer, 1955).<br />

т—тель<strong>с</strong><strong>о</strong>н, х2,0. 1159. Aglaspis splnlfer (Raasch); передняя<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, xl,3; э — эпи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>; хе — хелицер<strong>а</strong>.<br />

1160. Aglaspis splnlfer (Raasch); А —п<strong>о</strong><strong>с</strong>ледняя <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; Б — пер<strong>в</strong><strong>а</strong>я <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

Pseudoniscidae, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я б<strong>о</strong>лее<br />

<strong>к</strong>рупн<strong>о</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, лишенн<strong>о</strong>й гл<strong>а</strong>з, и ин<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й.<br />

П<strong>о</strong> ф<strong>о</strong>рме тел<strong>а</strong> <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидиды, <strong>а</strong> именн<strong>о</strong><br />

Strabopidae (<strong>в</strong>ерхний <strong>к</strong>ембрий) и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

Paleomeridae (нижний <strong>к</strong>ембрий), имеют мн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>с</strong> Eurypterida, т<strong>а</strong><strong>к</strong>ими, <strong>к</strong><strong>а</strong><strong>к</strong> Hughmilleriidae<br />

(<strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> — н. де<strong>в</strong><strong>о</strong>н). Н<strong>а</strong>личие<br />

12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидид <strong>о</strong>бъединяет<br />

их <strong>с</strong> Eurypterida. Еще б<strong>о</strong>льше <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>с</strong><br />

Hughmilleriidae имеет б<strong>о</strong>лее п<strong>о</strong>здний пред<strong>с</strong>т<strong>а</strong><strong>в</strong>итель<br />

Paleomeridae — р<strong>о</strong>д Angarocaris из<br />

<strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong>. У эт<strong>о</strong>й ф<strong>о</strong>рмы <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее<br />

уз<strong>к</strong><strong>а</strong>я, <strong>в</strong>еретен<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я, б<strong>о</strong>льше н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющ<strong>а</strong>я<br />

Eurypterida, чем Merostomata.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

О<strong>с</strong>т<strong>а</strong>т<strong>к</strong>и из<strong>в</strong>е<strong>с</strong>тных <strong>а</strong>гл<strong>а</strong><strong>с</strong>пидид<br />

были н<strong>а</strong>йдены <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> трил<strong>о</strong>бит<strong>а</strong>ми<br />

и плечен<strong>о</strong>гими м<strong>о</strong>ллю<strong>с</strong><strong>к</strong><strong>а</strong>ми. П<strong>о</strong> <strong>в</strong><strong>с</strong>ей <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти,<br />

<strong>а</strong>гл<strong>а</strong><strong>с</strong>пидиды <strong>о</strong>бит<strong>а</strong>ли <strong>в</strong> б<strong>о</strong>лее или менее<br />

<strong>с</strong>п<strong>о</strong><strong>к</strong><strong>о</strong>йн<strong>о</strong>й мел<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>дн<strong>о</strong>й м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>реде. Н. <strong>к</strong>ембрий<br />

— <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Paleomeridae, Aglaspididae,<br />

Strabopidae, Beckwithiidae.<br />

391<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PALEOMERIDAE ST0RMER, 1955<br />

Ф<strong>о</strong>рмы <strong>с</strong>редней <strong>в</strong>еличины (6—10 <strong>с</strong>м). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

из 12 <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>тый — тр<strong>а</strong>пец<strong>о</strong>ид<strong>а</strong>льный. Тель<strong>с</strong><strong>о</strong>н<br />

и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти неиз<strong>в</strong>е<strong>с</strong>тны. Н. <strong>к</strong>ембрий —<br />

<strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.

'.! 1161Б \ ,' 1161А<br />

Ри<strong>с</strong>. 1161—1162. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Paleomeridae<br />

1161. Paleomerus hamiltonl St; ;rmer; A — б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>;<br />

Б— <strong>с</strong>пинн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>. х0,8, н. <strong>к</strong>ембрий, Ш<strong>в</strong>еция (Sttfrmer,<br />

955). 1162. Angarocaris tschekanowskll (Schmidt); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>,<br />

Xl,2, <strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>, В. Сибирь (<strong>о</strong>риг. ри<strong>с</strong>.)<br />

Paleomerus St0rmer, 1955. Тип р<strong>о</strong>д<strong>а</strong> — P. hamiltonl<br />

Stormer, 1955; н. <strong>к</strong>ембрий, Ш<strong>в</strong>еция.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>' и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong>ме<strong>с</strong>те <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния,<br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я,<br />

<strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми <strong>с</strong>л<strong>о</strong>жными гл<strong>а</strong>з<strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

близ<strong>к</strong><strong>о</strong> <strong>к</strong> переднему и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым <strong>к</strong>р<strong>а</strong>ям;<br />

д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>тый <strong>с</strong>егмент <strong>а</strong>бд<strong>о</strong>мен<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тый,<br />

тр<strong>а</strong>пеци<strong>о</strong>ид<strong>а</strong>льный (ри<strong>с</strong>. 1161). Один <strong>в</strong>ид.<br />

Н. <strong>к</strong>ембрий Ш<strong>в</strong>еции.<br />

Angarocaris Chernyshev, 1953. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Eurypterus? tschekanowskll Schmidt,<br />

1886 (г<strong>о</strong>м<strong>о</strong>тип, <strong>о</strong>пи<strong>с</strong><strong>а</strong>нный Черныше<strong>в</strong>ым);<br />

<strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>, В. Сибирь (<strong>к</strong>ри<strong>в</strong><strong>о</strong>луц<strong>к</strong>ий яру<strong>с</strong>, интей<strong>с</strong><strong>к</strong>ий<br />

г<strong>о</strong>риз<strong>о</strong>нт). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее или менее<br />

упл<strong>о</strong>щенн<strong>а</strong>я, п<strong>а</strong>р<strong>а</strong>б<strong>о</strong>личе<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния, <strong>с</strong><br />

длинными ген<strong>а</strong>льными <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми или т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

392<br />

http://jurassic.ru/<br />

<strong>с</strong> ген<strong>а</strong>льными угл<strong>а</strong>ми; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>ильн<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лые,<br />

<strong>о</strong><strong>к</strong>руглые, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены близ<strong>к</strong><strong>о</strong> <strong>к</strong><br />

переднему и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым <strong>к</strong>р<strong>а</strong>ям пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> м<strong>о</strong>рщин<strong>к</strong><strong>а</strong>ми, п<strong>о</strong>х<strong>о</strong>жими н<strong>а</strong><br />

<strong>в</strong>етр<strong>о</strong><strong>в</strong>ые зн<strong>а</strong><strong>к</strong>и н<strong>а</strong> пе<strong>с</strong><strong>к</strong>е, ме<strong>с</strong>т<strong>а</strong>ми имеют<strong>с</strong>я<br />

мел<strong>к</strong>ие ям<strong>к</strong>и. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я, уз<strong>к</strong><strong>а</strong>я,<br />

из 12 <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>тый<br />

<strong>с</strong>егмент пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тый; тель<strong>с</strong><strong>о</strong>н <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>в</strong>идный<br />

(ри<strong>с</strong>. 1162). В<strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Ср. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

В. Сибири.<br />

СЕМЕЙСТВО AGLASPIDIDAE MILLER, 1877<br />

Ф<strong>о</strong>рмы <strong>с</strong>редней <strong>в</strong>еличины (4—16 <strong>с</strong>м), <strong>с</strong> 11<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными <strong>с</strong>егмент<strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими умеренн<strong>о</strong><br />

пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тую <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му. В<strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

В. Кембрий — <strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>.<br />

1164<br />

Ри<strong>с</strong>. Г163—1165. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Aglaspididae<br />

1163. Aglaspis splnlfer Raasch; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, X 2<br />

- 0<br />

- <strong>в</strong><br />

- <strong>к</strong>ембрий, С.<br />

Амери<strong>к</strong><strong>а</strong> (St0rmer, 1955). 1164. Aglaspis simplex Raasch; x0,7,<br />

<strong>в</strong>", <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong> (Stermer, 1955). 1165. Glrardevla тиsculus<br />

Andreeva; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, xl,5, <strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>, В. Сибирь<br />

(<strong>о</strong>риг. ри<strong>с</strong>.)<br />

Aglaspis Hall, 1862. Тип р<strong>о</strong>д<strong>а</strong> —Л. barrandi<br />

Hall, 1862; <strong>в</strong>. <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong>. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

п<strong>а</strong>р<strong>а</strong>б<strong>о</strong>личе<strong>с</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong> длинными ген<strong>а</strong>льными<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми или т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> ген<strong>а</strong>льными угл<strong>а</strong>ми;<br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong>чти прям<strong>о</strong>й, <strong>в</strong>ыпу<strong>к</strong>лый;<br />

межгл<strong>а</strong>зн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>-<strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я;<br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ть д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет з<strong>а</strong>тыл<strong>о</strong>чн<strong>о</strong>г<strong>о</strong>

<strong>к</strong>р<strong>а</strong>я; тергиты <strong>с</strong> п<strong>а</strong>рными п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными<br />

н<strong>а</strong>р<strong>о</strong><strong>с</strong>т<strong>а</strong>ми (ри<strong>с</strong>. 1163—1164). Б<strong>о</strong>лее трех <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

В. <strong>к</strong>ембрий С. Амери<strong>к</strong>и.<br />

Girardevia Andreeva, 1957. Тип р<strong>о</strong>д<strong>а</strong> —<br />

G. musculus Andreeva, 1957; <strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>,<br />

B. Сибирь (<strong>к</strong>ри<strong>в</strong><strong>о</strong>луц<strong>к</strong>ий яру<strong>с</strong>, <strong>к</strong>ирен<strong>с</strong><strong>к</strong>ий г<strong>о</strong>риз<strong>о</strong>нт,<br />

р. Лен<strong>а</strong>). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я;<br />

ее передняя п<strong>о</strong>л<strong>о</strong><strong>в</strong>ин<strong>а</strong> треуг<strong>о</strong>льн<strong>а</strong>я, <strong>с</strong> з<strong>а</strong><strong>к</strong>ругленным<br />

передним угл<strong>о</strong>м, з<strong>а</strong>дняя — прям<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я;<br />

з<strong>а</strong>тыл<strong>о</strong>чный <strong>к</strong>р<strong>а</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы прям<strong>о</strong>й.<br />

Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лые, <strong>о</strong><strong>в</strong><strong>а</strong>льные,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные немн<strong>о</strong>г<strong>о</strong> <strong>в</strong>переди <strong>о</strong>т <strong>с</strong>ередины<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. С<strong>к</strong>ульптур<strong>а</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> м<strong>о</strong>рщин<strong>к</strong><strong>а</strong>ми,<br />

п<strong>о</strong>х<strong>о</strong>жими н<strong>а</strong> <strong>в</strong>етр<strong>о</strong><strong>в</strong>ые зн<strong>а</strong><strong>к</strong>и н<strong>а</strong> пе<strong>с</strong><strong>к</strong>е.<br />

Длин<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы 9—13 мм (ри<strong>с</strong>. 1165). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. Ср. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> В. Сибири.<br />

Вне СССР: <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong> из <strong>в</strong>. <strong>к</strong>ембрия<br />

C. Амери<strong>к</strong>и: Aglaspella Raasch, 1939; Aglaspoides<br />

Raasch, 1939; Glypharthrus Raasch,- 1939;<br />

Uarthrus Raasch, 1939; Cyclopites Raasch,<br />

1939; Craspedops Raasch, 1939; Setaspis<br />

Raasch, 1939.<br />

СЕМЕЙСТВО STRABOPIDAE GERHARDT, 1932<br />

Ф<strong>о</strong>рмы <strong>с</strong>редней <strong>в</strong>еличины (12—15 <strong>с</strong>м) <strong>с</strong> 11<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными <strong>с</strong>егмент<strong>а</strong>ми <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, без эпимер<strong>о</strong><strong>в</strong>;<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, эллиптиче<strong>с</strong><strong>к</strong><strong>а</strong>я.<br />

Сл<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>в</strong> <strong>с</strong>ередине пр<strong>о</strong><strong>с</strong><strong>о</strong>мы или близ<br />

переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я,<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я. Тель<strong>с</strong><strong>о</strong>н шир<strong>о</strong><strong>к</strong>ий. В. <strong>к</strong>ембрий —<br />

<strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Strabops Beecher, 1901. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. thacheri Beecher, 1901; <strong>в</strong>. <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong>.<br />

Тергиты <strong>с</strong> нея<strong>с</strong>ными чешуй<strong>к</strong><strong>а</strong>ми близ<br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я; тел<strong>о</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>е; гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

близ переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

(ри<strong>с</strong>. 1166). Один <strong>в</strong>ид. В. <strong>к</strong>ембрий С. Амери<strong>к</strong>и.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из<strong>в</strong>е<strong>с</strong>тен Neostrabops Caster<br />

et Meek, 1952 из <strong>в</strong>. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong> С. Амери<strong>к</strong>и; <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жением<br />

гл<strong>а</strong>з <strong>в</strong> <strong>с</strong>ередине пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

СЕМЕЙСТВО BECKWITHIIDAE RAASCH, 1939<br />

Три з<strong>а</strong>дние <strong>с</strong>егмент<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>с</strong>пл<strong>о</strong>шн<strong>о</strong>й щит. Ср. <strong>к</strong>ембрий. Один р<strong>о</strong>д.<br />

Beckwithia Res ser, 1931. Тип р<strong>о</strong>д<strong>а</strong> —<br />

В. typa Resser, 1931; <strong>с</strong>р. <strong>к</strong>ембрий, С Амери<strong>к</strong><strong>а</strong>.<br />

Опи<strong>с</strong><strong>а</strong>ны Б. И. Черныше<strong>в</strong>ым три р<strong>о</strong>д<strong>а</strong> из<br />

<strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong> В. Сибири: Obrutschewia Chernyshev,<br />

1953; Intejocaris Chernyshev, 1953;<br />

Chacharejocaris Chernyshev, 1953. П<strong>о</strong> ф<strong>о</strong>рме<br />

Ри<strong>с</strong>. 1166—1167. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Strabopidae, Beckwithiidae<br />

1166. Strabops thacheri Moysey; x0,7, <strong>в</strong>. <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong><br />

(St0rmer, 1955). 1167. Beckwithia typa Resser; xl,0, <strong>с</strong>р. <strong>к</strong>ембрий,<br />

С. Амери<strong>к</strong><strong>а</strong> (St0rmer, 1955)<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>чти п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>с</strong> упл<strong>о</strong>щенным<br />

<strong>о</strong>б<strong>о</strong>д<strong>о</strong>м; гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>мещ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й; з<strong>а</strong>дняя<br />

б<strong>о</strong>лее или менее п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я <strong>с</strong>р<strong>о</strong><strong>с</strong>шими<strong>с</strong>я <strong>с</strong>егмент<strong>а</strong>ми, <strong>с</strong><br />

д<strong>о</strong>р<strong>с</strong><strong>а</strong>льным шип<strong>о</strong>м; тель<strong>с</strong><strong>о</strong>н предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

уз<strong>к</strong>ий, <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>в</strong>идный (ри<strong>с</strong>. 1167). Один <strong>в</strong>ид.<br />

Ср. <strong>к</strong>ембрий С. Амери<strong>к</strong>и.<br />

AGLASPIDIDA INCERTAE SEDJS<br />

393<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и, <strong>о</strong>пи<strong>с</strong><strong>а</strong>нные п<strong>о</strong>д этими н<strong>а</strong>з<strong>в</strong><strong>а</strong>ниями,<br />

н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют Aglaspidida, н<strong>о</strong> <strong>в</strong><strong>с</strong>е-т<strong>а</strong><strong>к</strong>и<br />

<strong>с</strong>тр<strong>о</strong>ение тел<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я неиз<strong>в</strong>е<strong>с</strong>тным.<br />

http://jurassic.ru/

ОТРЯД CHASMATASPIDIDA<br />

[nom. corr. Novojilov, hie (pro Chasmataspida Caster et Brooks, 1956)]<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>лу<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, упл<strong>о</strong>щенн<strong>а</strong>я,<br />

<strong>с</strong> неб<strong>о</strong>льшими ген<strong>а</strong>льными<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>к</strong>рупные,<br />

<strong>в</strong>ыпу<strong>к</strong>лые, <strong>о</strong><strong>к</strong>руглые; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

чет<strong>к</strong><strong>о</strong> р<strong>а</strong>зделен<strong>а</strong> н<strong>а</strong> м<strong>о</strong>з<strong>о</strong><strong>с</strong><strong>о</strong>му и<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>му. Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> упл<strong>о</strong>щенн<strong>а</strong>я,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из трех <strong>с</strong>р<strong>о</strong><strong>с</strong>ших<strong>с</strong>я <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>бр<strong>а</strong>зующих целый щит, зубч<strong>а</strong>тый<br />

п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым <strong>к</strong>р<strong>а</strong>ям; трехд<strong>о</strong>льн<strong>о</strong><strong>с</strong>ть<br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> д<strong>в</strong>умя б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляющими<br />

эпимеры <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> и третьег<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы; <strong>в</strong>ентр<strong>а</strong>льн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong><br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong>пл<strong>о</strong>шн<strong>а</strong>я, не<strong>с</strong>ущ<strong>а</strong>я н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х<br />

щеле<strong>в</strong>идн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие ж<strong>а</strong>берн<strong>о</strong>й<br />

<strong>к</strong><strong>а</strong>меры. Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> из де<strong>в</strong>яти <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>с</strong> эпимер<strong>а</strong>ми, уз<strong>к</strong><strong>а</strong>я и длинн<strong>а</strong>я.<br />

Тель<strong>с</strong><strong>о</strong>н <strong>с</strong>тержне<strong>в</strong>идный, неп<strong>о</strong>д<strong>в</strong>ижный.<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й <strong>в</strong> <strong>в</strong>иде буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong><br />

или пузырь<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й и<br />

<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>х. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Chasmataspididae Caster et<br />

Brooks, 1956 <strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м Chasmataspis<br />

Caster et Brooks, 1956 (ри<strong>с</strong>.<br />

1168). H. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> К<strong>а</strong>н<strong>а</strong>ды.<br />

Ри<strong>с</strong>. 1168. Отряд Chasmataspidida<br />

1168. Chasmataspis laurenci Caster et Brooks;<br />

A — <strong>с</strong><strong>в</strong>ерху, x 1,5; Б — <strong>с</strong>низу, X 1,5; <strong>о</strong> — <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть,<br />

Xl,5, <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>, К<strong>а</strong>н<strong>а</strong>д<strong>а</strong> (Caster et<br />

Brooks, 1956)<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> без <strong>в</strong><strong>с</strong>я<strong>к</strong>их <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з,<br />

•<strong>с</strong> <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью или без нее; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> де<strong>в</strong>ятью или де<strong>с</strong>ятью <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными <strong>с</strong>егмент<strong>а</strong>ми,<br />

ин<strong>о</strong>гд<strong>а</strong> ше<strong>с</strong>т<strong>о</strong>й и <strong>с</strong>едьм<strong>о</strong>й <strong>с</strong>егменты<br />

•<strong>с</strong>р<strong>о</strong><strong>с</strong>шие<strong>с</strong>я; <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong> упл<strong>о</strong>щенными ди<strong>с</strong>т<strong>а</strong>льными<br />

<strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми <strong>в</strong>ме<strong>с</strong>т<strong>о</strong> <strong>к</strong>лешней.<br />

М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия. Синзиф<strong>о</strong>зуры неб<strong>о</strong>льш<strong>о</strong>й<br />

<strong>в</strong>еличины — <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 5 <strong>с</strong>м длин<strong>о</strong>й (<strong>в</strong><strong>к</strong>люч<strong>а</strong>я<br />

тель<strong>с</strong><strong>о</strong>н). Тел<strong>о</strong> эллиптиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния.<br />

Стр<strong>о</strong>ение тел<strong>а</strong> у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей р<strong>о</strong>д<strong>а</strong> Pseudoniscus<br />

Nieszkowski п<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет, чт<strong>о</strong> <strong>с</strong>инзиф<strong>о</strong>зуры<br />

м<strong>о</strong>гли <strong>с</strong><strong>в</strong>ерты<strong>в</strong><strong>а</strong>ть<strong>с</strong>я. Крупн<strong>а</strong>я, п<strong>о</strong>чти<br />

п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я или шлем<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ф<strong>о</strong>рм <strong>с</strong> ген<strong>а</strong>льными уг­<br />

ОТРЯД SYNZIPHOSURA<br />

394<br />

http://jurassic.ru/<br />

л<strong>а</strong>ми, <strong>с</strong>лег<strong>к</strong><strong>а</strong> пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тыми (Pseudoniscus<br />

Nieszkowski, Weinbergina Richter et Richter),<br />

у других р<strong>о</strong>д<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> меньше, <strong>в</strong> р<strong>а</strong>зличн<strong>о</strong>й<br />

<strong>с</strong>тепени <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я и <strong>с</strong><strong>к</strong>ульптир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я. К<strong>а</strong>рди<strong>а</strong>льн<strong>а</strong>я<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть <strong>о</strong>тчетли<strong>в</strong><strong>а</strong>я у Bunodidae и<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у Neolimulidae. Гл<strong>а</strong>зные ребр<strong>а</strong> <strong>о</strong>бычн<strong>о</strong><br />

<strong>о</strong><strong>к</strong>руж<strong>а</strong>ют гл<strong>а</strong>беллярную <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть (Limuloididae,<br />

Neolimulidae), чт<strong>о</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм. Сегмент<strong>а</strong>ция н<strong>а</strong>ружн<strong>о</strong>й<br />

з<strong>о</strong>ны <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти ин<strong>о</strong>гд<strong>а</strong><br />

<strong>о</strong>тмечен<strong>а</strong> р<strong>а</strong>ди<strong>а</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми (Bunodidae)<br />

и п<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong><strong>о</strong>м р<strong>а</strong>ди<strong>а</strong>льных ребер (Limuloides).<br />

Обычн<strong>о</strong> нет ни<strong>к</strong><strong>а</strong><strong>к</strong>их <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>с</strong>л<strong>о</strong>жных<br />

гл<strong>а</strong>з, н<strong>о</strong>, м<strong>о</strong>жет быть, <strong>о</strong>ни имели<strong>с</strong>ь у Neolimulus

и Pseudoniscus. Н<strong>а</strong> прежних ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>циях<br />

б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong> Pseudoniscus из<strong>о</strong>бр<strong>а</strong>ж<strong>а</strong>ли<strong>с</strong>ь<br />

б<strong>о</strong>льшие <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> у <strong>а</strong>мери<strong>к</strong><strong>а</strong>н<strong>с</strong><strong>к</strong>их<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> эт<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> не был<strong>о</strong> <strong>о</strong>бн<strong>а</strong>ружен<strong>о</strong><br />

ни<strong>к</strong><strong>а</strong><strong>к</strong>их <strong>с</strong>лед<strong>о</strong><strong>в</strong> гл<strong>а</strong>з, з<strong>а</strong> и<strong>с</strong><strong>к</strong>лючением не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>с</strong>тру<strong>к</strong>тур, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые Рудем<strong>а</strong>н и<strong>с</strong>т<strong>о</strong>л<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>л<br />

<strong>к</strong><strong>а</strong><strong>к</strong> неб<strong>о</strong>льшие гл<strong>а</strong>з<strong>а</strong> и ф<strong>а</strong>ци<strong>а</strong>льные ш<strong>в</strong>ы.<br />

Ст<strong>о</strong>рмер (Stermer, 1956) <strong>о</strong>тмеч<strong>а</strong>ет, чт<strong>о</strong> эти <strong>о</strong>бр<strong>а</strong>зцы<br />

были <strong>с</strong>ильн<strong>о</strong> деф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ны <strong>в</strong> глини<strong>с</strong>т<strong>о</strong>м<br />

<strong>с</strong>л<strong>а</strong>нце. Неб<strong>о</strong>льшие з<strong>в</strong>ездч<strong>а</strong>тые <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

н<strong>а</strong> гл<strong>а</strong>зных ребр<strong>а</strong>х Limuloides, <strong>к</strong><strong>а</strong><strong>к</strong><br />

<strong>о</strong>тмеч<strong>а</strong>ет Ст<strong>о</strong>рмер, м<strong>о</strong>гли быть чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительными<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичными<br />

д<strong>о</strong>р<strong>с</strong><strong>а</strong>льным <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>м у з<strong>а</strong>р<strong>о</strong>дышей <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>.<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> имеет де<strong>в</strong>ять или де<strong>с</strong>ять <strong>о</strong>тчетли<strong>в</strong>ых<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> чет<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>о</strong><br />

деление <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong> шир<strong>о</strong><strong>к</strong>ую мез<strong>о</strong><strong>с</strong><strong>о</strong>му и<br />

уз<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му. В эт<strong>о</strong>м <strong>с</strong>луч<strong>а</strong>е мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из <strong>с</strong>еми или из ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, причем<br />

ше<strong>с</strong>т<strong>о</strong>й и <strong>с</strong>едьм<strong>о</strong>й тергиты ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я.<br />

Реже деление <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong> д<strong>в</strong><strong>а</strong> р<strong>а</strong>здел<strong>а</strong> нея<strong>с</strong>н<strong>о</strong>е<br />

(Neolimulidae, Pseudoniscidae). Трехд<strong>о</strong>льн<strong>о</strong><strong>с</strong>ть<br />

<strong>о</strong>тчетли<strong>в</strong><strong>а</strong>я у Pseudoniscidae и Neolimulidae<br />

и менее <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> у других ф<strong>о</strong>рм,<br />

не имеющих д<strong>о</strong>р<strong>с</strong><strong>а</strong>льных б<strong>о</strong>р<strong>о</strong>зд. Пле<strong>в</strong>ры м<strong>о</strong>гут<br />

быть <strong>с</strong> пр<strong>о</strong>резями и пр<strong>о</strong>д<strong>о</strong>лж<strong>а</strong>ть<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

пле<strong>в</strong>р<strong>а</strong>льные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и.<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти из<strong>в</strong>е<strong>с</strong>тны т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме<br />

у Weinberginidae — <strong>в</strong> <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е пяти п<strong>а</strong>р х<strong>о</strong>дильных<br />

н<strong>о</strong>г; хелицеры неиз<strong>в</strong>е<strong>с</strong>тны. Стр<strong>о</strong>ение<br />

н<strong>о</strong>г <strong>о</strong>чень <strong>с</strong>х<strong>о</strong>дн<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>тр<strong>о</strong>ением н<strong>о</strong>ги IV <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Limulida; <strong>к</strong>лешни <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, имеют<strong>с</strong>я<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льных<br />

члени<strong>к</strong><strong>а</strong>х. От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие з<strong>а</strong>метных <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

н<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме у х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>ших<strong>с</strong>я<br />

<strong>о</strong>бр<strong>а</strong>зц<strong>о</strong><strong>в</strong> не и<strong>с</strong><strong>к</strong>люч<strong>а</strong>ет <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ти н<strong>а</strong>личия<br />

н<strong>а</strong> них пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тых <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, т<strong>а</strong><strong>к</strong>их, <strong>к</strong><strong>а</strong><strong>к</strong><br />

у Limulida и Eurypterida. Н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><br />

<strong>о</strong>бычн<strong>о</strong> гл<strong>а</strong>д<strong>к</strong>ий, реже б<strong>о</strong>лее или менее <strong>с</strong>ильн<strong>о</strong><br />

<strong>о</strong>рн<strong>а</strong>ментир<strong>о</strong><strong>в</strong><strong>а</strong>нный.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения,<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Р<strong>а</strong>зличие<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>терных <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

Pseudoniscidae и Weinberginidae, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

и <strong>о</strong><strong>с</strong>т<strong>а</strong>льных из<strong>в</strong>е<strong>с</strong>тных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>с</strong> друг<strong>о</strong>й,<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет н<strong>а</strong>метить д<strong>в</strong>е группы (? н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>),<br />

н<strong>о</strong> т<strong>а</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>др<strong>а</strong>зделение будет<br />

<strong>в</strong><strong>о</strong>зм<strong>о</strong>жным при н<strong>а</strong>личии б<strong>о</strong>лее <strong>о</strong>пределенных<br />

д<strong>а</strong>нных. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие <strong>о</strong>тн<strong>о</strong>шения Synziphosura<br />

еще м<strong>а</strong>л<strong>о</strong> и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ны. Н<strong>а</strong>личие <strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

групп<strong>а</strong>х <strong>о</strong>тряд<strong>а</strong> чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>мы и пл<strong>о</strong>тных <strong>о</strong>рн<strong>а</strong>ментир<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> <strong>с</strong>ближ<strong>а</strong>ет этих мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> <strong>с</strong> Chasmataspididae.<br />

С друг<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, <strong>к</strong>рупн<strong>а</strong>я шир<strong>о</strong><strong>к</strong><strong>а</strong>я<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и м<strong>а</strong>л<strong>о</strong>дифференцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я<br />

<strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щ<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют н<strong>а</strong> я<strong>в</strong>ные<br />

<strong>с</strong><strong>в</strong>язи <strong>с</strong> Limulida, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, я<strong>в</strong>­<br />

ляют<strong>с</strong>я п<strong>о</strong>т<strong>о</strong>м<strong>к</strong><strong>а</strong>ми не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong>инзиф<strong>о</strong>зур,<br />

п<strong>о</strong><strong>к</strong><strong>а</strong> еще не из<strong>в</strong>е<strong>с</strong>тных.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

Синзиф<strong>о</strong>зуры, п<strong>о</strong> <strong>в</strong><strong>с</strong>ей<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти, прин<strong>а</strong>длеж<strong>а</strong>ли <strong>к</strong> бент<strong>о</strong><strong>с</strong>у и не<br />

<strong>в</strong>х<strong>о</strong>дили <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> типичн<strong>о</strong> м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>а</strong>уны. Из<strong>в</strong>е<strong>с</strong>тные<br />

<strong>с</strong>илурий<strong>с</strong><strong>к</strong>ие и д<strong>а</strong>унт<strong>о</strong>н<strong>с</strong><strong>к</strong>ие <strong>в</strong>иды <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляли<br />

ч<strong>а</strong><strong>с</strong>ть э<strong>в</strong>риптерид<strong>о</strong><strong>в</strong><strong>о</strong>й ф<strong>а</strong>уны, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

н<strong>а</strong><strong>с</strong>еля<strong>в</strong>шей <strong>с</strong><strong>о</strong>л<strong>о</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>тые или пре<strong>с</strong>ные<br />

<strong>в</strong><strong>о</strong>дные б<strong>а</strong><strong>с</strong><strong>с</strong>ейны. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> нижнеде<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

Weinbergina, н<strong>а</strong>йденн<strong>а</strong>я <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> трил<strong>о</strong>бит<strong>а</strong>ми,<br />

был<strong>а</strong>, <strong>о</strong>че<strong>в</strong>идн<strong>о</strong>, м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й ф<strong>о</strong>рм<strong>о</strong>й. В. <strong>с</strong>илур —<br />

н. де<strong>в</strong><strong>о</strong>н. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Limuloididae, Bunodidae,<br />

Neolimulidae, Pseudoniscidae, Weinberginidae.<br />

395<br />

СЕМЕЙСТВО LIMULOIDIDAE STORMER, 1952<br />

(HEMIASPIDIDAE ZITTEL, 1881)<br />

Ф<strong>о</strong>рмы неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>еличины. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><br />

<strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью, гл<strong>а</strong>зными ребр<strong>а</strong>ми, р<strong>а</strong>ди<strong>а</strong>льными<br />

ребр<strong>а</strong>ми и <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ыми шип<strong>а</strong>ми; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

р<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> тр<strong>а</strong>пецие<strong>в</strong>идную<br />

ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ую мез<strong>о</strong><strong>с</strong><strong>о</strong>му и уз<strong>к</strong>ую трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ую<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>му. Ше<strong>с</strong>т<strong>о</strong>й <strong>с</strong>егмент мез<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

д<strong>в</strong><strong>о</strong>йн<strong>о</strong>й, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из <strong>с</strong>р<strong>о</strong><strong>с</strong>ших<strong>с</strong>я ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong> и<br />

<strong>с</strong>едьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> шир<strong>о</strong><strong>к</strong>им <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><strong>о</strong>м<br />

(<strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью); <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><br />

ряд<strong>а</strong>ми <strong>к</strong>рупных буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong>е и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

ч<strong>а</strong><strong>с</strong>тях пер<strong>в</strong>ых четырех тергит<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Один р<strong>о</strong>д: Limuloides Stormer, 1952<br />

[= Hemiaspis Woodward, 1864 (non Fitzinger,<br />

1861)] (ри<strong>с</strong>. 1169). В. <strong>с</strong>илур 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО BUNODIDAE PACKARD, 1886<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>лу<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>а</strong>я, р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>д<strong>о</strong>льч<strong>а</strong>т<strong>а</strong>я;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>др<strong>а</strong>зделен<strong>а</strong> н<strong>а</strong> шир<strong>о</strong><strong>к</strong>ую<br />

ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ую мез<strong>о</strong><strong>с</strong><strong>о</strong>му, не<strong>с</strong>ущую<br />

<strong>с</strong><strong>о</strong>члененные пле<strong>в</strong>р<strong>а</strong>льные шипы, и четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ую<br />

уз<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му, лишенную пле<strong>в</strong>р<strong>а</strong>льных<br />

шип<strong>о</strong><strong>в</strong>. Пер<strong>в</strong>ый <strong>с</strong>егмент мет<strong>а</strong><strong>с</strong><strong>о</strong>мы уз<strong>к</strong>ий;<br />

тергиты мез<strong>о</strong><strong>с</strong><strong>о</strong>мы не<strong>с</strong>ут <strong>с</strong>рединный ряд<br />

<strong>к</strong>рупных буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> без буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong> (<strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть) н<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме не<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен. В. <strong>с</strong>илур — ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) '.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

1<br />

Д<strong>а</strong>унт<strong>о</strong>н пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>с</strong><strong>а</strong>мый <strong>в</strong>ерхний (чет<strong>в</strong>ертый<br />

п<strong>о</strong> <strong>с</strong>чету) яру<strong>с</strong> <strong>в</strong>ерхнег<strong>о</strong> <strong>о</strong>тдел<strong>а</strong> <strong>с</strong>илурий<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>и<strong>с</strong>темы,<br />

неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> н<strong>а</strong>д лудл<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>им яру<strong>с</strong><strong>о</strong>м (Сл<strong>о</strong><strong>в</strong><strong>а</strong>рь п<strong>о</strong><br />

ге<strong>о</strong>л<strong>о</strong>гии нефти, Г<strong>о</strong><strong>с</strong>т<strong>о</strong>птехизд<strong>а</strong>т, 1962). С нед<strong>а</strong><strong>в</strong>нег<strong>о</strong> <strong>в</strong>ремени<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong><strong>с</strong>ть эт<strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>а</strong> <strong>о</strong><strong>с</strong>п<strong>а</strong>ри<strong>в</strong><strong>а</strong>ет<strong>с</strong>я не<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми<br />

<strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>ф<strong>а</strong>ми. Оче<strong>в</strong>идн<strong>о</strong>, п<strong>о</strong> эт<strong>о</strong>й причине<br />

д<strong>а</strong>унт<strong>о</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong> не п<strong>о</strong>мещен <strong>в</strong> <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>с</strong>хеме,<br />

принят<strong>о</strong>й для «О<strong>с</strong>н<strong>о</strong><strong>в</strong> п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии» (<strong>с</strong>м. <strong>с</strong>тр. 6),<br />

и <strong>в</strong><strong>о</strong><strong>о</strong>бще не н<strong>а</strong>шел ни<strong>к</strong><strong>а</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тр<strong>а</strong>жения <strong>в</strong> эт<strong>о</strong>й <strong>с</strong>хеме.<br />

В «Treatise on Invertebrate Paleontology*, Pt P, Arthropoda<br />

2, д<strong>а</strong>унт<strong>о</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong> цитирует<strong>с</strong>я <strong>с</strong><strong>о</strong> зн<strong>а</strong><strong>к</strong><strong>о</strong>м 1<br />

«?» <strong>в</strong><br />

нижнем де<strong>в</strong><strong>о</strong>не: «? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н)». Мн<strong>о</strong>гие Eurypterida<br />

и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые мечех<strong>в</strong><strong>о</strong><strong>с</strong>ты, р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>тренные <strong>в</strong> «О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>х<br />

п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии», шр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят <strong>к</strong><strong>а</strong><strong>к</strong> из лудл<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>,<br />

т<strong>а</strong><strong>к</strong> и из д<strong>а</strong>унт<strong>о</strong>н<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> яру<strong>с</strong><strong>о</strong><strong>в</strong> и, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>с</strong><strong>о</strong>б-<br />

http://jurassic.ru/

1174 А<br />

Ри<strong>с</strong>. 1169—1174. Отряд Synziphosura (<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Limuloididae, Bunodidae, Neolimulidae,<br />

Pseudoniscidae, Weinberginidae)<br />

1169. Limuloldes Umuloidts (Woodward); xl,0, <strong>в</strong>. <strong>с</strong>илур, Англия<br />

<strong>в</strong>переди, и <strong>о</strong><strong>с</strong>трыми ген<strong>а</strong>льными угл<strong>а</strong>ми;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, б<strong>о</strong>лее или менее п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я,<br />

<strong>с</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют <strong>к</strong><strong>а</strong>рди<strong>а</strong>льную <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>. Тель<strong>с</strong><strong>о</strong>н, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий.<br />

Тел<strong>о</strong> гл<strong>а</strong>д<strong>к</strong><strong>о</strong>е, без <strong>с</strong><strong>к</strong>ульптуры (ри<strong>с</strong>.<br />

1172). Один р<strong>о</strong>д — Neolimulus Woodward,<br />

1863. В. <strong>с</strong>илур — н. де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО PSEUDONISCIDAE PACKARD, 1886<br />

Ф<strong>о</strong>рмы неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>еличины; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я — <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины <strong>в</strong><strong>с</strong>ей<br />

длины тел<strong>а</strong>, гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я и упл<strong>о</strong>щенн<strong>а</strong>я, <strong>с</strong> <strong>о</strong><strong>с</strong>трыми<br />

или <strong>о</strong><strong>к</strong>руглыми ген<strong>а</strong>льными угл<strong>а</strong>ми; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> рез<strong>к</strong>ими <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми, <strong>с</strong><br />

де<strong>в</strong>ятью — де<strong>с</strong>ятью <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными <strong>с</strong>егмент<strong>а</strong>ми,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые не<strong>с</strong>ут <strong>о</strong><strong>с</strong>трые пле<strong>в</strong>р<strong>а</strong>льные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и;<br />

SYNZIPHOSURA INCERTAE SEDIS<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> не <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и: Pseudoniscus Nieszkowski,<br />

1859 (ри<strong>с</strong>. 1173),? Cyamocephalus Currie, 1927.<br />

СЕМЕЙСТВО WEINBERGINIDAE RICHTER<br />

ET RICHTER, 1929<br />

Ф<strong>о</strong>рмы <strong>с</strong>редней <strong>в</strong>еличины; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я,<br />

гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я, <strong>к</strong>рупн<strong>а</strong>я, р<strong>а</strong><strong>в</strong>н<strong>а</strong>я п<strong>о</strong> <strong>в</strong>еличине<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>с</strong>яти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>, из трех <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льн<strong>а</strong>я (<strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>а</strong>я) <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

н<strong>а</strong>мечен<strong>а</strong> н<strong>а</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>ме ряд<strong>а</strong>ми буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>; хелицеры<br />

<strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, пять п<strong>а</strong>р х<strong>о</strong>дильных<br />

н<strong>о</strong>г <strong>с</strong> шип<strong>а</strong>ми. Один р<strong>о</strong>д — Weinbergina Richter<br />

et Richter, 1929 из н. де<strong>в</strong><strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

(ри<strong>с</strong>. 1174А.Б).<br />

П<strong>о</strong> фр<strong>а</strong>гмент<strong>а</strong>рным <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м <strong>о</strong>чень неп<strong>о</strong>л- (<strong>с</strong>илур Ш<strong>о</strong>тл<strong>а</strong>ндии), Bunodella Matthew, 1889<br />

н<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>ны р<strong>о</strong>ды Bembicosoma Laurie, 1899 (н. <strong>с</strong>илур С. Амери<strong>к</strong>и).<br />

ОТРЯД LIMULIDA<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong> <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью,<br />

гл<strong>а</strong>зными ребр<strong>а</strong>ми, гл<strong>а</strong>з<strong>к</strong><strong>а</strong>ми и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>с</strong>л<strong>о</strong>жными гл<strong>а</strong>з<strong>а</strong>ми; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> де<strong>в</strong>ятью<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми <strong>с</strong>егмент<strong>а</strong>ми или п<strong>о</strong>д<strong>в</strong>ижными,<br />

или же <strong>в</strong> т<strong>о</strong>й или ин<strong>о</strong>й <strong>с</strong>тепени <strong>с</strong>р<strong>о</strong><strong>с</strong>шими<strong>с</strong>я;<br />

тель<strong>с</strong><strong>о</strong>н <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>в</strong>идный, <strong>в</strong><strong>с</strong>егд<strong>а</strong> длиннее<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия. Крупн<strong>а</strong>я, б<strong>о</strong>лее или менее<br />

п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> не<strong>с</strong>ет ген<strong>а</strong>льные<br />

шипы, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong>чень ред<strong>к</strong><strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют (Liomesaspididae);<br />

<strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть <strong>к</strong>лин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>к</strong>иле<strong>в</strong><strong>а</strong>т<strong>а</strong>я, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми<br />

или узл<strong>а</strong>ми н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е,<br />

<strong>о</strong><strong>к</strong>руженн<strong>а</strong>я <strong>в</strong>переди <strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я гл<strong>а</strong>зными<br />

ребр<strong>а</strong>ми. Эти ребр<strong>а</strong> пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ют<strong>с</strong>я <strong>в</strong>д<strong>о</strong>ль б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти, через б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

гл<strong>а</strong>з<strong>а</strong>, и <strong>с</strong>мы<strong>к</strong><strong>а</strong>ют<strong>с</strong>я у переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. В <strong>к</strong><strong>а</strong>рди<strong>о</strong>фт<strong>а</strong>льмиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

м<strong>о</strong>гут быть р<strong>а</strong>з<strong>в</strong>иты межгл<strong>а</strong>зные ребр<strong>а</strong>, <strong>о</strong><strong>к</strong><strong>а</strong>ймляющие<br />

прям<strong>о</strong>уг<strong>о</strong>льную пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

<strong>в</strong><strong>к</strong>люч<strong>а</strong>ет <strong>к</strong><strong>а</strong>рди<strong>а</strong>льную л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть и уз<strong>к</strong>ие п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и<br />

у ее <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н. Е<strong>с</strong>ли межгл<strong>а</strong>зные ребр<strong>а</strong><br />

имеют<strong>с</strong>я, <strong>к</strong><strong>а</strong>рди<strong>о</strong>фт<strong>а</strong>льмиче<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я<br />

н<strong>а</strong> пять ч<strong>а</strong><strong>с</strong>тей <strong>в</strong>ме<strong>с</strong>т<strong>о</strong> трех. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, б<strong>о</strong>лее или менее треуг<strong>о</strong>льн<strong>а</strong>я<br />

или непр<strong>а</strong><strong>в</strong>ильн<strong>а</strong>я ше<strong>с</strong>тиуг<strong>о</strong>льн<strong>а</strong>я, не<br />

р<strong>а</strong>зделяющ<strong>а</strong>я<strong>с</strong>я н<strong>а</strong> мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>му. Сегмент<strong>а</strong>ция<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы у п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их ф<strong>о</strong>рм <strong>в</strong>ыр<strong>а</strong>­<br />

(Limulina)<br />

жен<strong>а</strong> <strong>о</strong>чень чет<strong>к</strong><strong>о</strong> пле<strong>в</strong>р<strong>а</strong>льными ребр<strong>а</strong>ми.<br />

Реже <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из цельн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>,<br />

<strong>с</strong>егмент<strong>а</strong>ция <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> н<strong>а</strong>меч<strong>а</strong>ет<strong>с</strong>я лишь ряд<strong>а</strong>ми<br />

ям<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льных б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>х и пле<strong>в</strong>р<strong>а</strong>льными<br />

шип<strong>а</strong>ми (Limulidea). Ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

без <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>с</strong>егмент<strong>а</strong>ции (Liomesaspididae)<br />

или т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> <strong>о</strong>дним <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дным <strong>с</strong>егмент<strong>о</strong>м<br />

перед тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м (Paleolimulidae). У р<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Euproopidea <strong>к</strong><strong>о</strong>льч<strong>а</strong>тый <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><br />

<strong>с</strong> <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ими шип<strong>а</strong>ми. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

из<strong>в</strong>е<strong>с</strong>тны лишь у немн<strong>о</strong>гих ф<strong>о</strong>рм и <strong>о</strong>чень п<strong>о</strong>х<strong>о</strong>жи<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong><strong>о</strong><strong>в</strong>ременных Limulidae.<br />

С<strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зны <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти у Liomesaspididae,<br />

у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых х<strong>о</strong>дильные н<strong>о</strong>ги р<strong>а</strong>зличн<strong>о</strong>й <strong>в</strong>еличины.<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть щит<strong>а</strong> гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или <strong>с</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми<br />

р<strong>а</strong>зличных <strong>в</strong>еличины и ф<strong>о</strong>рмы.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Фил<strong>о</strong>генез Limulida пр<strong>о</strong><strong>с</strong>лежи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т <strong>с</strong>илурий<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>инзиф<strong>о</strong>зур Neolimulidae, з<strong>а</strong>тем Belinuridae<br />

и д<strong>а</strong>лее <strong>к</strong> мез<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>им и <strong>с</strong><strong>о</strong><strong>в</strong>ременным<br />

Limulidea. Эт<strong>о</strong>т пр<strong>о</strong>це<strong>с</strong><strong>с</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я<br />

<strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щением длины <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>нием<br />

ее <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, <strong>а</strong> т<strong>а</strong><strong>к</strong>же п<strong>о</strong><strong>с</strong>тепенным <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>нием<br />

<strong>в</strong>еличины. В цел<strong>о</strong>м пр<strong>о</strong>х<strong>о</strong>дит х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женн<strong>а</strong>я <strong>о</strong>лиг<strong>о</strong>мериз<strong>а</strong>ция <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>с</strong><strong>к</strong>елет<strong>а</strong>.<br />

?Н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) — ныне. Три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Belinuridea, Liomesaspididea, Limulidea.<br />

397<br />

http://jurassic.ru/

117Б<br />

Ри<strong>с</strong>. 1175—1177. Отряд Limulida, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

1175. Neobellnurus rossicus (Chernyshev); X0,5, <strong>в</strong>. де<strong>в</strong><strong>о</strong>н, Липец<strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. 1176. Belinurus Baldwini Woodward; X2.0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

НАДСЕМЕЙСТВО BELINURIDEA<br />

[nom. transl. Novojilov, hie (ex Be'.inuracea Stormer,<br />

1955; Belimiridae Zittel et Eastman, 1913)]<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й члени<strong>с</strong>т<strong>о</strong><strong>с</strong>тью<br />

и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>тью.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong> длинными<br />

ген<strong>а</strong>льными угл<strong>а</strong>ми. Ср. де<strong>в</strong>ен — пермь. Пять<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Belimiridae, Austrolimulidae, Euproopidae,<br />

Elleriidae, Kiaeriidae.<br />

СЕМЕЙСТВО BELINURIDAE ZITTEL ET EASTMAN,<br />

1913<br />

(Bellinuridae Packard, 1886)<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> тр<strong>а</strong>пецие<strong>в</strong>идн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния,<br />

из <strong>в</strong><strong>о</strong><strong>с</strong>ьми чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

<strong>с</strong>н<strong>а</strong>бженных п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям п<strong>о</strong>д<strong>в</strong>ижными пле<strong>в</strong>р<strong>а</strong>льными<br />

шип<strong>а</strong>ми. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong> длине р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>ме или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>о</strong>льше ее. Ср. де<strong>в</strong><strong>о</strong>н —<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Три р<strong>о</strong>д<strong>а</strong>.<br />

Neobelinurus Е 11 er, 1938. Тип р<strong>о</strong>д<strong>а</strong> — Bel-<br />

Unuroopsis rossicus Chernyshev, 1933; <strong>в</strong> де<strong>в</strong><strong>о</strong>н,<br />

Липец<strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (близ Лебедяни). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><br />

<strong>к</strong>лин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>а</strong>я, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>а</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>, <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью п<strong>о</strong>д<strong>в</strong>ижными <strong>с</strong>егмент<strong>а</strong>ми,<br />

п<strong>о</strong><strong>с</strong>ледний из них, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>с</strong>д<strong>в</strong><strong>о</strong>енный.<br />

Сегменты <strong>с</strong> пле<strong>в</strong>р<strong>а</strong>льными <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 1175). Один <strong>в</strong>ид из <strong>в</strong>. де<strong>в</strong><strong>о</strong>н<strong>а</strong> СССР.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, Belinurus Konig, 1820, <strong>в</strong> де<strong>в</strong><strong>о</strong>н<br />

— <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и;<br />

Protollmulus Packard, 1886, <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н С. Амери<strong>к</strong>и<br />

(ри<strong>с</strong>. 1176, 1177).<br />

СЕМЕЙСТВО AUSTROLIMULIDAE RIEK, 1955<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>че пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>а</strong>я ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из трех<br />

<strong>с</strong>р<strong>о</strong><strong>с</strong>ших<strong>с</strong>я передних и трех п<strong>о</strong>д<strong>в</strong>ижных з<strong>а</strong>дних<br />

1176<br />

398<br />

http://jurassic.ru/<br />

| 1177<br />

Belinuridea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Belimiridae (Stermer, 1955)<br />

Англия. 1177. Belinurus reglna Bally; x2,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Ирл<strong>а</strong>ндия<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Ген<strong>а</strong>льные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>чень<br />

<strong>в</strong>ели<strong>к</strong>и и шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны. Три<strong>а</strong><strong>с</strong>.<br />

Один р<strong>о</strong>д.<br />

Austrollmulus Riek, 1955. Тип р<strong>о</strong>д<strong>а</strong> — А.<br />

fleetcheri Riek, 1955; <strong>с</strong>р. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>к</strong>ругл<strong>а</strong>я <strong>с</strong>переди,<br />

<strong>с</strong> длинными ген<strong>а</strong>льными <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми, <strong>к</strong><strong>о</strong>нцы<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют ур<strong>о</strong><strong>в</strong>ня п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены,,<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> непр<strong>а</strong><strong>в</strong>ильными буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми,<br />

<strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ме<strong>с</strong>т<strong>а</strong>х — ребрыш<strong>к</strong><strong>а</strong>ми;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>о</strong>тчетли<strong>в</strong>ым уз<strong>к</strong>им <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><strong>о</strong>м,.<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>: передние три<strong>с</strong>р<strong>о</strong><strong>с</strong>шие<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зующие тр<strong>а</strong>пецие<strong>в</strong>идный щит,,<br />

<strong>с</strong>уж<strong>а</strong>ющий<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д, три з<strong>а</strong>дние <strong>с</strong>егмент<strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижные,<br />

уз<strong>к</strong>ие; тель<strong>с</strong><strong>о</strong>н трехгр<strong>а</strong>нный, <strong>в</strong> 1,5 р<strong>а</strong>з<strong>а</strong><br />

длиннее пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>ме<strong>с</strong>те<strong>в</strong>зятых<br />

(ри<strong>с</strong>. 1178; т<strong>а</strong>бл. XVII). Один <strong>в</strong>ид. Ср.<br />

три<strong>а</strong><strong>с</strong> А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии.<br />

СЕМЕЙСТВО EUPROOPIDAE ELLER, 1938<br />

Неб<strong>о</strong>льшие ф<strong>о</strong>рмы <strong>с</strong> <strong>к</strong>лин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью, <strong>о</strong><strong>к</strong><strong>а</strong>ймленн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми;<br />

<strong>к</strong><strong>а</strong>рди<strong>о</strong>фт<strong>а</strong>льмиче<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

гл<strong>а</strong>зные ребр<strong>а</strong> <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я п<strong>о</strong>з<strong>а</strong>ди гл<strong>а</strong>з;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я и <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящ<strong>а</strong>я<br />

из ше<strong>с</strong>ти или <strong>с</strong>еми чет<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женных, пл<strong>о</strong>тн<strong>о</strong><strong>с</strong><strong>о</strong>единенных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, <strong>с</strong> <strong>о</strong><strong>с</strong>трыми б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми; имеет<strong>с</strong>я <strong>к</strong><strong>о</strong>льч<strong>а</strong>тый <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong>, н<strong>а</strong> п<strong>о</strong><strong>с</strong>ледних<br />

<strong>с</strong>егмент<strong>а</strong>х <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>ие<br />

<strong>в</strong>ы<strong>с</strong>тупы. Один р<strong>о</strong>д Euproops Meek, 1867 (ри<strong>с</strong>.<br />

1179, 1180). <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н —<br />

пермь С. Амери<strong>к</strong>и; н. пермь В. Азии.<br />

СЕМЕЙСТВО ELLERIIDAE RAYMOND, 1944<br />

Нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>я групп<strong>а</strong>. Сегмент<strong>а</strong>ция<br />

<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы и пле<strong>в</strong>р<strong>а</strong>льн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и<br />

<strong>о</strong>тчетли<strong>в</strong>ые; <strong>к</strong><strong>а</strong>ждый <strong>с</strong>егмент <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><strong>а</strong> <strong>с</strong>

Ри<strong>с</strong>. 1178—1180. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Austrolimulidae и Euproopidae<br />

1178. Austrolimulus fleetchert Riek; ХЬ5, <strong>с</strong>р. три<strong>а</strong><strong>с</strong>, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>- <strong>с</strong>тяни. Х0,7, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия (St0rmer, 1945). 1180. Euproops.<br />

лня (ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> Н. Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong><strong>а</strong>, п<strong>о</strong> ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии). 1179. Euproops thompsonl Raymond, Xl,0, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (St0rmer, 1955<br />

anthrax Woodward; д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong> <strong>с</strong> <strong>о</strong>т<strong>к</strong>рытыми <strong>к</strong><strong>о</strong>нечн<strong>о</strong>-<br />

тремя буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми; з<strong>а</strong>дняя ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

глуб<strong>о</strong><strong>к</strong><strong>о</strong> <strong>в</strong>ырез<strong>а</strong>н<strong>а</strong>; тель<strong>с</strong><strong>о</strong>н и пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тны.<br />

Один р<strong>о</strong>д Elleria Raymond, 1944. В. де<strong>в</strong><strong>о</strong>н—<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО KIAER11DAE ST0RMER, 1952<br />

Нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>я групп<strong>а</strong>. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong>м<strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я, <strong>с</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми и гл<strong>а</strong>д<strong>к</strong>им<br />

<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><strong>о</strong>м; <strong>с</strong>егмент<strong>а</strong>ция и шир<strong>о</strong><strong>к</strong>ий з<strong>а</strong>дний <strong>в</strong>ырез<br />

у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют н<strong>а</strong> <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ть п<strong>о</strong>д<strong>в</strong>ижных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

тель<strong>с</strong><strong>о</strong>н и пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тны. Один<br />

р<strong>о</strong>д Kiaeria Stormer, 1934. ? Н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н)<br />

С<strong>к</strong><strong>а</strong>ндин<strong>а</strong><strong>в</strong>ии.<br />

BELINURIDEA INCERTAE SEDIS<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>пи<strong>с</strong><strong>а</strong>ны п<strong>о</strong> пл<strong>о</strong>х<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шим<strong>с</strong>я<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м р<strong>о</strong>ды Pringlia Raymond, 1944<br />

(<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н З.Е<strong>в</strong>р<strong>о</strong>пы и С.Амери<strong>к</strong>и),Anaconti-<br />

ит Raymond, 1944 (пермь С. Амери<strong>к</strong>и). Pin.combella<br />

Chapman, 1932 (<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы),Belinuropsis<br />

Matthew, 1909 (<strong>в</strong>. <strong>с</strong>илур С. Амери<strong>к</strong>и).<br />

НАДСЕМЕЙСТВО LIOMESASPIDIDEA<br />

[nom. transl. Novojilov, his<br />

(ex Liomesaspididae Raymond, 1944)]<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> цельн<strong>а</strong>я, без <strong>с</strong>лед<strong>о</strong><strong>в</strong> <strong>с</strong>егмент<strong>а</strong>ции<br />

п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong><strong>к</strong>руглые,<br />

<strong>с</strong> пл<strong>о</strong>тн<strong>о</strong>й дублюр<strong>о</strong>й п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям, без шип<strong>о</strong><strong>в</strong>,<br />

<strong>с</strong>х<strong>о</strong>дные п<strong>о</strong> ф<strong>о</strong>рме и <strong>в</strong>еличине. Ген<strong>а</strong>льные углы<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — н. пермь. Одн<strong>а</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomesaspididae.<br />

СЕМЕЙСТВО LIOMESASPIDIDAE RAYMOND, 1944<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льн<strong>а</strong>я <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я;<br />

ген<strong>а</strong>льные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и <strong>о</strong>т<strong>с</strong>тут<strong>с</strong>т<strong>в</strong>уют; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

без <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых шип<strong>о</strong><strong>в</strong> и пле<strong>в</strong>р<strong>а</strong>льных<br />

399<br />

http://jurassic.ru/

Ри<strong>с</strong>. 1181—1182. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Liomesaspididea, <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Liomesaspididae<br />

1181. Liomesaspis laevis Raymond; xl,7, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

(St0rmer, 1955). 1182. Prolimulus woodward! Fritsch; <strong>с</strong>низу,<br />

X2,0, н. пермь, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия (St0rmer, 1955)<br />

ребер; з<strong>а</strong>дний <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льный <strong>с</strong>егмент без <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong><strong>в</strong>.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Liomesaspis Raymond, 1944<br />

(<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и), Prolimulus Fritsch,<br />

1899 (пермь 3. Е<strong>в</strong>р<strong>о</strong>пы) (ри<strong>с</strong>. 1181, 1182).<br />

1183<br />

НАДСЕМЕЙСТВО LIMULIDEA<br />

[nom. transl. Novojilov, hie<br />

(ex Limulidae Zittel, 1885;<br />

Limulacea Raymond, 1944)]<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong> <strong>в</strong>иде цельн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>; ин<strong>о</strong>гд<strong>а</strong><br />

п<strong>о</strong>д<strong>в</strong>ижен лишь <strong>о</strong>дин з<strong>а</strong>дний <strong>с</strong>егмент перед<br />

тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м. Сегмент<strong>а</strong>ция <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong>злич<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жению пле<strong>в</strong>р<strong>а</strong>льных шип<strong>о</strong><strong>в</strong> или<br />

зубц<strong>о</strong><strong>в</strong> и при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ию ям<strong>о</strong><strong>к</strong> <strong>в</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льных б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>х.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными з<strong>а</strong>дними<br />

<strong>в</strong>ет<strong>в</strong>ями гл<strong>а</strong>зных ребер и умеренн<strong>о</strong> удлиненными<br />

ген<strong>а</strong>льными <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми. (Ин<strong>о</strong>гд<strong>а</strong>, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

<strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>нения, <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>тмеч<strong>а</strong>ет<br />

Ст<strong>о</strong>рмер, <strong>в</strong>ентр<strong>а</strong>льн<strong>а</strong>я <strong>с</strong>тру<strong>к</strong>тур<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<br />

<strong>с</strong>леды н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти, чт<strong>о</strong> <strong>с</strong><strong>о</strong>зд<strong>а</strong>ет<br />

непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>е пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ление <strong>о</strong> <strong>с</strong>тру<strong>к</strong>туре пр<strong>о</strong><strong>с</strong><strong>о</strong>мы).<br />

Пермь — ныне. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Paleolimulidae,<br />

Mesolimulidae, Limulidae.<br />

СЕМЕЙСТВО PALEOLIMULIDAE<br />

RAYMOND, 1944<br />

Ф<strong>о</strong>рмы неб<strong>о</strong>льш<strong>о</strong>й <strong>в</strong>еличины. Гл<strong>а</strong>зные ребр<strong>а</strong><br />

<strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong>переди <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти;<br />

<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы я<strong>в</strong>н<strong>о</strong> <strong>к</strong><strong>о</strong>льч<strong>а</strong>тый; имеет<strong>с</strong>я<br />

Ри<strong>с</strong>. 1183—1188. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Paleolimulidae, Mesolimulidae и Limulidae (St^rmer, 1955)<br />

1183. Paleolimulus avitus Dunbar, xl,0, пермь, К<strong>а</strong>н<strong>а</strong>д<strong>а</strong>. 1184. Mesolimulus walchi (Demarest); x0,5, юр<strong>а</strong>, Герм<strong>а</strong>ния. 1187.<br />

Psammolimulus goitingensis Lange; XO,5, н. три<strong>а</strong><strong>с</strong>, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. 1185. Mesolimulus syriacus (Woodward); X0,2, Ср. Азия, <strong>с</strong>р. мел.<br />

Limuliiella bronni (Schimper); x0,5, н. три<strong>а</strong><strong>с</strong>, Фр<strong>а</strong>нция. 1186. 1188. Tachypleus decheni (Zinken); x0,13, ми<strong>о</strong>цен, Герм<strong>а</strong>ния<br />

400<br />

1184<br />

http://jurassic.ru/

п<strong>о</strong>д<strong>в</strong>ижный <strong>с</strong>егмент <strong>в</strong>переди тель<strong>с</strong><strong>о</strong>н<strong>а</strong>. Один<br />

poR — Paleolimulus Dunbar, 1923 (ри<strong>с</strong>. 1183).<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО MESOLIMUL1DAE ST0RMER, 1952<br />

Ф<strong>о</strong>рмы <strong>о</strong>т неб<strong>о</strong>льш<strong>о</strong>й д<strong>о</strong> <strong>с</strong>редней <strong>в</strong>еличины.<br />

Гл<strong>а</strong>зные ребр<strong>а</strong> не <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong>переди <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти; ген<strong>а</strong>льные углы умеренн<strong>о</strong> пр<strong>о</strong>д<strong>о</strong>лжены<br />

н<strong>а</strong>з<strong>а</strong>д; <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льные б<strong>о</strong>р<strong>о</strong>зды <strong>о</strong>тчетли<strong>в</strong>ые;<br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>т<strong>о</strong><strong>с</strong>ть <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>с</strong><strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы не<strong>о</strong>тчетли<strong>в</strong><strong>а</strong>я;<br />

пер<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых шип<strong>о</strong><strong>в</strong> <strong>в</strong> предел<strong>а</strong>х<br />

передней трети <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Три р<strong>о</strong>д<strong>а</strong>: Psammolitnulus<br />

Lange, 1923 (н. три<strong>а</strong><strong>с</strong> Е<strong>в</strong>р<strong>о</strong>пы),<br />

Limulitella Stormer, 1952 (н.— <strong>в</strong>. три<strong>а</strong><strong>с</strong> Е<strong>в</strong>р<strong>о</strong>пы,<br />

С. Амери<strong>к</strong>и), Mesolimulus Stormer, 1952<br />

26 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

(юр<strong>а</strong>, ?мел 3. Е<strong>в</strong>р<strong>о</strong>пы и ?Азии) (ри<strong>с</strong>. 1184—<br />

1187).<br />

СЕМЕЙСТВО LIMULIDAE Z1TTEL, 1885<br />

(Xiphosuridae Pocock, 1902)<br />

Ф<strong>о</strong>рмы <strong>о</strong>т <strong>с</strong>редней д<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>й <strong>в</strong>еличины.<br />

Гл<strong>а</strong>зные ребр<strong>а</strong> не <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong>переди <strong>к</strong><strong>а</strong>рди<strong>а</strong>льн<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти, ген<strong>а</strong>льные углы зн<strong>а</strong>чительн<strong>о</strong><br />

пр<strong>о</strong>д<strong>о</strong>лжены н<strong>а</strong>з<strong>а</strong>д; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее или<br />

менее ше<strong>с</strong>тиуг<strong>о</strong>льн<strong>а</strong>я, <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льные б<strong>о</strong>р<strong>о</strong>зды<br />

не<strong>о</strong>тчетли<strong>в</strong>ые; пер<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых шип<strong>о</strong><strong>в</strong><br />

приблизительн<strong>о</strong> н<strong>а</strong> <strong>с</strong>ередине между передним и<br />

з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ями <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Не<strong>о</strong>ген — ныне.<br />

Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них Tachypleus<br />

Leach, 1819 (ри<strong>с</strong>. 1188) из<strong>в</strong>е<strong>с</strong>тен и <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии (ми<strong>о</strong>цен, 3- Е<strong>в</strong>р<strong>о</strong>пы).<br />

http://jurassic.ru/

КЛАСС SCORPIONOMORPHA. СКОРПИОНОМОРФЫ<br />

Тел<strong>о</strong> чет<strong>к</strong><strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> передний<br />

<strong>о</strong>тдел, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь, или пр<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую<br />

из пер<strong>в</strong>ичных ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и не<strong>с</strong>ущую<br />

члени<strong>с</strong>тые <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, и з<strong>а</strong>дний <strong>о</strong>тдел, или<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую из 12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong> и<br />

тель<strong>с</strong><strong>о</strong>н<strong>а</strong>, или <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти, лишенную<br />

л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>т<strong>о</strong>рных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, <strong>в</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>ю <strong>о</strong>чередь, ин<strong>о</strong>гд<strong>а</strong> п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>му<br />

и мет<strong>а</strong><strong>с</strong><strong>о</strong>му. В<strong>с</strong>е тергиты пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong>литы<br />

между <strong>с</strong><strong>о</strong>б<strong>о</strong>й и <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>дин г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный<br />

щит, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичный г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>му щиту<br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы утр<strong>а</strong>тили дых<strong>а</strong>тельную<br />

фун<strong>к</strong>цию, чт<strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и<br />

<strong>в</strong>ыр<strong>а</strong>зил<strong>о</strong><strong>с</strong>ь <strong>в</strong> реду<strong>к</strong>ции преэпип<strong>о</strong>дит<strong>о</strong><strong>в</strong>; <strong>о</strong>ни<br />

<strong>в</strong>ып<strong>о</strong>лняют лишь л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>т<strong>о</strong>рную и же<strong>в</strong><strong>а</strong>тельную<br />

фун<strong>к</strong>ции. Д<strong>а</strong>же у типичн<strong>о</strong> <strong>в</strong><strong>о</strong>дных Scorpionomorpha<br />

— п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их э<strong>в</strong>риптерид и у <strong>с</strong>илурий<strong>с</strong><strong>к</strong>их<br />

<strong>в</strong><strong>о</strong>дных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong><br />

Apoxypodes н<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тях пр<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли ж<strong>а</strong>берные прид<strong>а</strong>т<strong>к</strong>и.<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти мез<strong>о</strong><strong>с</strong><strong>о</strong>мы, н<strong>а</strong><strong>о</strong>б<strong>о</strong>р<strong>о</strong>т, утр<strong>а</strong>тили<br />

тел<strong>о</strong>п<strong>о</strong>диты и же<strong>в</strong><strong>а</strong>тельные прид<strong>а</strong>т<strong>к</strong>и (б<strong>а</strong>зэндиты)<br />

примити<strong>в</strong>н<strong>о</strong>й <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> преэпип<strong>о</strong>диты, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>ып<strong>о</strong>лняют<br />

зде<strong>с</strong>ь фун<strong>к</strong>ции дых<strong>а</strong>ния (ж<strong>а</strong>бры у Eurypterida,<br />

лег<strong>к</strong>ие у Scorpiones и др.) и при<strong>о</strong>брели еще<br />

фун<strong>к</strong>ции, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нные <strong>с</strong> р<strong>а</strong>з<strong>в</strong>итием н<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия (п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые, <strong>к</strong>рышеч<strong>к</strong>и).<br />

Б<strong>о</strong>лее примити<strong>в</strong>ные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

(п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Holactinochitinosi) <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> чет<strong>к</strong><strong>о</strong>е р<strong>а</strong>зделение <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленных <strong>о</strong>тдел<strong>а</strong> — мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>му,<br />

т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>в</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е Pedipalpides эти<br />

т<strong>а</strong>глы <strong>с</strong>литы <strong>в</strong>ме<strong>с</strong>те, и т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Uropygi п<strong>о</strong><strong>с</strong>ледние<br />

<strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены.<br />

Одн<strong>а</strong><strong>к</strong><strong>о</strong> <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е Scorpionomorpha не<br />

н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я т<strong>о</strong>й м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>тепени <strong>с</strong>лия­<br />

(I В. Б. Дубинин \)<br />

402<br />

http://jurassic.ru/<br />

ния <strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я т<strong>а</strong><strong>к</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>а</strong><br />

для Arachnida.<br />

Орг<strong>а</strong>ны дых<strong>а</strong>ния пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены тремя тип<strong>а</strong>ми<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ний. У <strong>в</strong><strong>о</strong>дных пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> (Eurypterida, Apoxypodes) имел<strong>о</strong><strong>с</strong>ь<br />

ж<strong>а</strong>берн<strong>о</strong>е дых<strong>а</strong>ние, <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ля<strong>в</strong>шее<strong>с</strong>я <strong>с</strong> п<strong>о</strong>м<strong>о</strong>щью<br />

ж<strong>а</strong>берных н<strong>о</strong>же<strong>к</strong> III—VI <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

У н<strong>а</strong>земных ф<strong>о</strong>рм (<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong><br />

Dionychopodes, и б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> жгут<strong>о</strong>н<strong>о</strong>гих п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

Pedipalpides) пер<strong>в</strong>ичные ж<strong>а</strong>берные<br />

н<strong>о</strong>ж<strong>к</strong>и пре<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ли<strong>с</strong>ь <strong>в</strong> лег<strong>о</strong>чные меш<strong>к</strong>и,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющие<strong>с</strong>я н<strong>а</strong> тех же <strong>с</strong>егмент<strong>а</strong>х мез<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я н<strong>а</strong>ружу четырьмя (<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны)<br />

или д<strong>в</strong>умя (жгут<strong>о</strong>н<strong>о</strong>гие) п<strong>а</strong>р<strong>а</strong>ми уз<strong>к</strong>их<br />

дых<strong>а</strong>лец. Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, у л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> (Pseudoscorpionidea)<br />

р<strong>а</strong>з<strong>в</strong>ит тр<strong>а</strong>хейный тип дых<strong>а</strong>ния<br />

и н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я утр<strong>а</strong>т<strong>а</strong> лег<strong>к</strong>их. Для <strong>в</strong><strong>с</strong>ех<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> типичн<strong>о</strong> н<strong>а</strong>личие д<strong>в</strong>ух<br />

<strong>к</strong>рупных меди<strong>а</strong>льных гл<strong>а</strong>з, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющих<strong>с</strong>я<br />

н<strong>а</strong> <strong>с</strong>ередине <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>,<br />

и трех — ше<strong>с</strong>ти п<strong>а</strong>р мел<strong>к</strong>их пр<strong>о</strong><strong>с</strong>тых гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

леж<strong>а</strong>щих <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых угл<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме и фил<strong>о</strong>гения.<br />

Ряд <strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>их <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует<br />

<strong>о</strong> б<strong>о</strong>льш<strong>о</strong>й фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>й дре<strong>в</strong>н<strong>о</strong><strong>с</strong>ти<br />

<strong>в</strong><strong>с</strong>ех пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Scorpionomorpha,<br />

я<strong>в</strong>и<strong>в</strong>ших<strong>с</strong>я пер<strong>в</strong>ыми н<strong>а</strong>земными<br />

жи<strong>в</strong><strong>о</strong>тными п<strong>о</strong>дтип<strong>а</strong> Chelicerata, <strong>а</strong> быть м<strong>о</strong>жет,<br />

и Arthropoda. Арх<strong>а</strong>ичн<strong>о</strong><strong>с</strong>ть их <strong>в</strong>ыр<strong>а</strong>ж<strong>а</strong>ет<strong>с</strong>я<br />

прежде <strong>в</strong><strong>с</strong>ег<strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нении <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мн<strong>о</strong>й <strong>с</strong>егмент<strong>а</strong>ции тел<strong>а</strong> при н<strong>а</strong>личии м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

чи<strong>с</strong>л<strong>а</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>—19 (Scorpiones),<br />

18 (Uropygi, Amblypygi, Pseudoscorpionidea)<br />

или 16 (Palpigradi), <strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нении<br />

пер<strong>в</strong>ичн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> <strong>в</strong><strong>о</strong>здушн<strong>о</strong>г<strong>о</strong> дых<strong>а</strong>ния <strong>с</strong> п<strong>о</strong>м<strong>о</strong>щью<br />

п<strong>о</strong>груженных п<strong>о</strong>д п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> ж<strong>а</strong>берных<br />

н<strong>о</strong>же<strong>к</strong> — лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> (тр<strong>а</strong>хеи

<strong>в</strong><strong>о</strong>зни<strong>к</strong>ли т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у н<strong>а</strong>иб<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

ф<strong>о</strong>рм — Pseudoscorpionoidea) и, н<strong>а</strong><strong>к</strong><strong>о</strong>нец,<br />

<strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>нении <strong>в</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>л<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии у<br />

мн<strong>о</strong>гих групп <strong>с</strong>едьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>, леж<strong>а</strong>щег<strong>о</strong> н<strong>а</strong><br />

гр<strong>а</strong>нице между пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>г<strong>о</strong> у мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>. Седьм<strong>о</strong>й <strong>с</strong>егмент<br />

лучше <strong>в</strong><strong>с</strong>ег<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит у Palpigradi, где <strong>о</strong>н <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong><strong>с</strong>ть и <strong>с</strong>лужит <strong>с</strong><strong>о</strong>единительным<br />

<strong>с</strong>тебель<strong>к</strong><strong>о</strong>м между пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й;<br />

у Uropygi <strong>о</strong>н <strong>с</strong>лит <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьмым <strong>с</strong>егмент<strong>о</strong>м,<br />

н<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет тергит и <strong>с</strong>тернит и группы<br />

д<strong>о</strong>р<strong>с</strong><strong>о</strong><strong>в</strong>ентр<strong>а</strong>льных мышц; у Amblypygi <strong>о</strong>н р<strong>а</strong>з<strong>в</strong>ит<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>, у л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я ег<strong>о</strong><br />

тергит; у других Scorpionomorpha и <strong>в</strong><strong>с</strong>ех<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных хелицер<strong>о</strong><strong>в</strong>ых реду<strong>к</strong>ция <strong>с</strong>едьм<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> (Petrunkevitch, 1922) з<strong>а</strong>х<strong>о</strong>дит <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong><br />

д<strong>а</strong>ле<strong>к</strong><strong>о</strong>, и ин<strong>о</strong>гд<strong>а</strong> ег<strong>о</strong> н<strong>а</strong>личие уд<strong>а</strong>ет<strong>с</strong>я з<strong>а</strong>метить<br />

лишь <strong>в</strong> течение эмбри<strong>о</strong>генез<strong>а</strong>. Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, х<strong>а</strong>р<strong>а</strong><strong>к</strong>терны<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нение у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Scorpionomorpha<br />

(н<strong>а</strong>пример, у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Scorpiones и у<br />

<strong>в</strong><strong>с</strong>ех Palpigradi) р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>а</strong>, н<strong>а</strong> <strong>в</strong>ершине<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я р<strong>о</strong>т, и при<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие у<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> м<strong>а</strong><strong>к</strong><strong>с</strong>иллярных <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х <strong>в</strong><strong>с</strong>ех <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

уч<strong>а</strong><strong>с</strong>т<strong>в</strong>ующих <strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нии предр<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти<br />

(педип<strong>а</strong>льпы и д<strong>в</strong>е передние п<strong>а</strong>ры<br />

х<strong>о</strong>дильных н<strong>о</strong>г). В<strong>с</strong>ё <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ющ<strong>а</strong>я у Chelicerata<br />

гетер<strong>о</strong>н<strong>о</strong>мн<strong>о</strong><strong>с</strong>ть <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы при<strong>в</strong><strong>о</strong>дит<br />

<strong>к</strong> т<strong>о</strong>му, чт<strong>о</strong> <strong>в</strong> принятии пищи у Arachnida<br />

приним<strong>а</strong>ют уч<strong>а</strong><strong>с</strong>тие т<strong>о</strong>ль<strong>к</strong><strong>о</strong> хелицеры, т<strong>о</strong>гд<strong>а</strong><br />

<strong>к</strong><strong>а</strong><strong>к</strong> у Scorpionomorpha зн<strong>а</strong>чительную р<strong>о</strong>ль <strong>в</strong><br />

эт<strong>о</strong>м игр<strong>а</strong>ют и педип<strong>а</strong>льпы, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичные<br />

м<strong>а</strong>ндибул<strong>а</strong>м р<strong>а</strong><strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных, н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых и мн<strong>о</strong>г<strong>о</strong>н<strong>о</strong>же<strong>к</strong>;<br />

<strong>с</strong>реди <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Scorpionomorpha т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Palpigradi педип<strong>а</strong>льпы<br />

не уч<strong>а</strong><strong>с</strong>т<strong>в</strong>уют <strong>в</strong> з<strong>а</strong>х<strong>в</strong><strong>а</strong>те пищи.<br />

Д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> <strong>в</strong>ремени <strong>о</strong>тряды хелицер<strong>о</strong><strong>в</strong>ых,<br />

<strong>о</strong>бъединяемые зде<strong>с</strong>ь <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Scorpionomorpha,<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>или<strong>с</strong>ь <strong>о</strong>дними и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телями<br />

<strong>к</strong> р<strong>а</strong>зличным <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>м (Merostomata и Arachnoidea;<br />

Grasse, 1949), другими — <strong>к</strong> р<strong>а</strong>зличным<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong>м (З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин, 1952; Бе<strong>к</strong>лемише<strong>в</strong>,<br />

1952) или <strong>к</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>у Eurypterida и д<strong>в</strong>ум п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>м<br />

Arachnida (Petrunkevitsh, 1949, 1953).<br />

П<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение был<strong>о</strong> <strong>в</strong>ыз<strong>в</strong><strong>а</strong>н<strong>о</strong> тем, чт<strong>о</strong><br />

<strong>с</strong><strong>о</strong> <strong>в</strong>ремени Л<strong>а</strong>н<strong>к</strong><strong>а</strong><strong>с</strong>тер<strong>а</strong> (Lankester, 1911) и<br />

П<strong>о</strong><strong>к</strong><strong>о</strong><strong>к</strong><strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1910) <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е Arachnida и<br />

п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е Eurachnida были <strong>в</strong>ыделены д<strong>в</strong>е и<strong>с</strong><strong>к</strong>у<strong>с</strong><strong>с</strong>т<strong>в</strong>енные<br />

группы хелицер<strong>о</strong><strong>в</strong>ых: <strong>в</strong><strong>о</strong>дные и<br />

н<strong>а</strong>земные. В д<strong>а</strong>льнейшем з<strong>а</strong> п<strong>о</strong><strong>с</strong>ледней <strong>с</strong>б<strong>о</strong>рн<strong>о</strong>й,<br />

и<strong>с</strong><strong>к</strong>у<strong>с</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й групп<strong>о</strong>й н<strong>а</strong>земных хелицер<strong>о</strong><strong>в</strong>ых<br />

з<strong>а</strong><strong>к</strong>репил<strong>о</strong><strong>с</strong>ь н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние «<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>» п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных—<br />

Arachnoidea. Т<strong>а</strong><strong>к</strong><strong>о</strong>е <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бление<br />

н<strong>а</strong>земных хелицер<strong>о</strong><strong>в</strong>ых <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льных групп,<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> чи<strong>с</strong>т<strong>о</strong> э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

принципе, при<strong>в</strong>ел<strong>о</strong> <strong>к</strong> п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ению <strong>в</strong>нутри эт<strong>о</strong>г<strong>о</strong><br />

«<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>» <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> з<strong>а</strong>пут<strong>а</strong>н<strong>о</strong>й и <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>й <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> не <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ющей е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енных<br />

<strong>в</strong>з<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шений этих члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

(Petrunkevitsh, 1949, 1953). Эти<br />

нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и не уд<strong>а</strong>л<strong>о</strong><strong>с</strong>ь пре<strong>о</strong>д<strong>о</strong>леть и З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ину<br />

(1952), предл<strong>о</strong>жи<strong>в</strong>шему н<strong>о</strong><strong>в</strong>ую <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Chelicerata, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong><strong>в</strong>шему<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные <strong>о</strong>тн<strong>о</strong>шения <strong>в</strong><strong>о</strong>дных Eurypterida<br />

<strong>с</strong> н<strong>а</strong>земными Scorpiones и Thelyphonida, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

были <strong>о</strong>бъединены <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельный<br />

н<strong>а</strong>д<strong>о</strong>тряд Holactinochitinosi. Си<strong>с</strong>тему З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин<strong>а</strong><br />

<strong>с</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми <strong>о</strong>г<strong>о</strong><strong>в</strong><strong>о</strong>р<strong>к</strong><strong>а</strong>ми принял Бе<strong>к</strong>лемише<strong>в</strong><br />

(1952).<br />

В дей<strong>с</strong>т<strong>в</strong>ительн<strong>о</strong><strong>с</strong>ти же Arachnoidea р<strong>а</strong>зделяют<strong>с</strong>я<br />

н<strong>а</strong> три <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>яльные <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>.<br />

Пер<strong>в</strong>ый из этих <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong> — <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Scorpionomorpha,<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных, н<strong>о</strong>г<strong>о</strong>р<strong>о</strong>тых или<br />

лег<strong>о</strong>чн<strong>о</strong>дыш<strong>а</strong>щих хелицер<strong>о</strong><strong>в</strong>ых пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<br />

д<strong>в</strong>умя п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>ми: Holactinochitinosi (<strong>о</strong>тряды<br />

Eurypterida и Scorpiones) и- Pedipalpides (<strong>о</strong>тряды:<br />

Uropygi, Amblypygi, Palpigradi, Kustarachnida,<br />

Pseudoscorpioidea).<br />

ПОДКЛАСС HOLACTINOCHITINOSI.<br />

СКОРПИОНООБРАЗНЫЕ<br />

Сильн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е <strong>в</strong> длину тел<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>о</strong>тдел<strong>а</strong>: пр<strong>о</strong><strong>с</strong><strong>о</strong>му, п<strong>о</strong><strong>к</strong>рытую г<strong>о</strong>л<strong>о</strong><strong>в</strong>ным<br />

щит<strong>о</strong>м и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую из <strong>с</strong>еми пер<strong>в</strong>ичных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, и длинную 13-члени<strong>к</strong><strong>о</strong><strong>в</strong>ую<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделенную н<strong>а</strong> шир<strong>о</strong><strong>к</strong>ую<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую из ше<strong>с</strong>ти или <strong>с</strong>еми<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, и уз<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящую из<br />

ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; п<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент мет<strong>а</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>с</strong><strong>о</strong>членен <strong>с</strong> тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м, имеющим ф<strong>о</strong>рму<br />

из<strong>о</strong>гнут<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>гтя, прям<strong>о</strong>г<strong>о</strong> шип<strong>а</strong> или пл<strong>а</strong><strong>в</strong>ни<strong>к</strong><strong>а</strong>.<br />

Хелицеры трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, ред<strong>к</strong><strong>о</strong><br />

удлиненные (Pterygotidae), <strong>к</strong>лешне<strong>в</strong>идные.<br />

Педип<strong>а</strong>льпы р<strong>а</strong>з<strong>в</strong>иты р<strong>а</strong>зличн<strong>о</strong>: у Eurypterida<br />

<strong>о</strong>ни либ<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, либ<strong>о</strong> длинные, ните<strong>в</strong>идные<br />

(Slimonia) или х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные (н<strong>а</strong>пример<br />

у Mixopterus); у <strong>в</strong><strong>с</strong>ех Scorpiones <strong>о</strong>ни х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные—<br />

<strong>в</strong> <strong>в</strong>иде м<strong>о</strong>щных <strong>к</strong>рупных <strong>к</strong>лешней.<br />

В<strong>с</strong>е <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы (<strong>к</strong>р<strong>о</strong>ме хелицер)<br />

имеют н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х же<strong>в</strong><strong>а</strong>тельные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и;<br />

ж<strong>а</strong>берные прид<strong>а</strong>т<strong>к</strong>и н<strong>а</strong> них п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти мез<strong>о</strong><strong>с</strong><strong>о</strong>мы пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ыми <strong>к</strong>рышеч<strong>к</strong><strong>а</strong>ми н<strong>а</strong> <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>м <strong>с</strong>егменте<br />

и ж<strong>а</strong>берными н<strong>о</strong>ж<strong>к</strong><strong>а</strong>ми н<strong>а</strong> де<strong>в</strong>ят<strong>о</strong>м —<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>м <strong>с</strong>егмент<strong>а</strong>х у Eurypterida и <strong>в</strong><strong>о</strong>дных<br />

403 26»<br />

http://jurassic.ru/

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>. Эти <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> <strong>в</strong>ых<strong>о</strong>д<strong>о</strong>м<br />

н<strong>а</strong> <strong>с</strong>ушу пре<strong>в</strong>р<strong>а</strong>тили<strong>с</strong>ь <strong>в</strong> лег<strong>о</strong>чные меш<strong>к</strong>и н<strong>а</strong>земных<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>.<br />

В н<strong>а</strong>д<strong>о</strong>тряде Holactinochitinosi, не<strong>с</strong>м<strong>о</strong>тря<br />

н<strong>а</strong> пр<strong>о</strong>из<strong>о</strong>шедшие рез<strong>к</strong>ие изменения <strong>в</strong> э<strong>к</strong><strong>о</strong>л<strong>о</strong>гии<br />

(перех<strong>о</strong>д <strong>о</strong>т <strong>в</strong><strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>к</strong> н<strong>а</strong>земн<strong>о</strong>му <strong>о</strong>бр<strong>а</strong>зу жизни),<br />

<strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех групп<strong>а</strong>х <strong>с</strong>т<strong>о</strong>й<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<br />

р<strong>а</strong><strong>с</strong>членения тел<strong>а</strong> и <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей,<br />

<strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>о</strong>бщий м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бный <strong>о</strong>бли<strong>к</strong>. Н<strong>а</strong>земных и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

и <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> п<strong>о</strong> пр<strong>а</strong><strong>в</strong>у мн<strong>о</strong>гие<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тели н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ют н<strong>а</strong>земными<br />

э<strong>в</strong>риптерид<strong>а</strong>ми.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Из<strong>в</strong>е<strong>с</strong>т­<br />

Хелицер<strong>о</strong><strong>в</strong>ые, п<strong>о</strong>х<strong>о</strong>жие н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых дре<strong>в</strong>них<br />

Merostomata, именн<strong>о</strong> Aglaspidida. Тел<strong>о</strong><br />

пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>е, б<strong>о</strong>лее или менее л<strong>а</strong>нцет<strong>о</strong><strong>в</strong>идн<strong>о</strong>е,<br />

ред<strong>к</strong><strong>о</strong> трехд<strong>о</strong>льн<strong>о</strong>е (у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных<br />

и Mycteropidae: Borchgrevinkium).<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й ф<strong>о</strong>рмы: п<strong>о</strong>чти<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я, <strong>в</strong>ырез<strong>а</strong>нн<strong>а</strong>я <strong>с</strong>з<strong>а</strong>ди, п<strong>о</strong>лу<strong>к</strong>ругл<strong>а</strong>я, п<strong>о</strong>лу<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я,<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я, тр<strong>а</strong>пец<strong>о</strong>ид<strong>а</strong>льн<strong>а</strong>я,<br />

пятиуг<strong>о</strong>льн<strong>а</strong>я, шлем<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния.<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>тью п<strong>о</strong>д<strong>в</strong>ижными<br />

<strong>с</strong>егмент<strong>а</strong>ми, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передние <strong>с</strong>емь<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют мез<strong>о</strong><strong>с</strong><strong>о</strong>му (или пре<strong>а</strong>бд<strong>о</strong>мен), <strong>а</strong><br />

з<strong>а</strong>дние пять — мет<strong>а</strong><strong>с</strong><strong>о</strong>му (или п<strong>о</strong><strong>с</strong>т<strong>а</strong>бд<strong>о</strong>мен).<br />

К п<strong>о</strong><strong>с</strong>леднему <strong>с</strong>егменту мет<strong>а</strong><strong>с</strong><strong>о</strong>мы причленен<br />

тель<strong>с</strong><strong>о</strong>н, <strong>о</strong>бычн<strong>о</strong> <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>в</strong>идный или друг<strong>о</strong>й<br />

•ф<strong>о</strong>рмы, <strong>к</strong><strong>о</strong>пье- или л<strong>о</strong>п<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зный, н<strong>а</strong><strong>к</strong><strong>о</strong>нец,<br />

<strong>к</strong><strong>о</strong>гте<strong>в</strong>идный, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>. Хелицеры<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>лешне<strong>в</strong>идные. Педип<strong>а</strong>льпы и<br />

четыре з<strong>а</strong>дние п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены<br />

для х<strong>о</strong>ждения; ин<strong>о</strong>гд<strong>а</strong> педип<strong>а</strong>льпы и третья<br />

п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные. З<strong>а</strong>дняя<br />

(VI) п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong>ид<strong>о</strong>изменен<strong>а</strong><br />

<strong>в</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные н<strong>о</strong>ги. Р<strong>о</strong>т центр<strong>а</strong>льный, <strong>о</strong>гр<strong>а</strong>ниченный<br />

<strong>с</strong>з<strong>а</strong>ди д<strong>в</strong>умя пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми: энд<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>й<br />

и мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>й. Генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong><br />

<strong>с</strong> п<strong>о</strong><strong>с</strong>тгенит<strong>а</strong>льн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>о</strong>й (<strong>о</strong>пер<strong>к</strong>улум);<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> <strong>в</strong>иде пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong> <strong>с</strong> нер<strong>а</strong><strong>с</strong>щепленными<br />

ж<strong>а</strong>бр<strong>а</strong>ми.<br />

М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия. В б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong>е из<strong>в</strong>е<strong>с</strong>тные<br />

Eurypterida <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> неб<strong>о</strong>льшие—10—<br />

20 <strong>с</strong>м длины, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые ф<strong>о</strong>рмы были<br />

<strong>о</strong>чень <strong>к</strong>рупными — д<strong>о</strong> 100—180 <strong>с</strong>м. Т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы<br />

Megalograptidae <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong>, Slimonia Page и<br />

Pterygotus Agassiz <strong>с</strong>илур<strong>а</strong> и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые Stylonuridae<br />

де<strong>в</strong><strong>о</strong>н<strong>а</strong> (TarsoptereUa Stormer, Ctenopterus<br />

Clarke et Ruedemann). П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong><br />

"р<strong>а</strong>зличн<strong>о</strong>й пл<strong>о</strong>тн<strong>о</strong><strong>с</strong>ти. Ч<strong>а</strong><strong>с</strong>т<strong>о</strong> имеют<strong>с</strong>я мн<strong>о</strong>г<strong>о</strong>­<br />

ОТРЯД EURYPTERIDA<br />

(Не<strong>с</strong>т<strong>о</strong>р Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>)<br />

404<br />

http://jurassic.ru/<br />

ны <strong>с</strong> <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong> д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> <strong>в</strong>ремени. Пер<strong>в</strong>ичн<strong>о</strong><strong>в</strong><strong>о</strong>дные,<br />

<strong>в</strong>едущие <strong>с</strong><strong>в</strong><strong>о</strong>е пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение <strong>о</strong>т<br />

Merostomata, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>о</strong>тряд<strong>а</strong> Eurypterida,<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли <strong>в</strong> течение п<strong>о</strong>чти <strong>в</strong><strong>с</strong>ег<strong>о</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>я.<br />

В <strong>с</strong>илуре н<strong>а</strong>метил<strong>о</strong><strong>с</strong>ь <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бление <strong>о</strong>т<br />

них <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong> т<strong>а</strong><strong>к</strong>же<br />

<strong>в</strong>ели <strong>в</strong><strong>о</strong>дный <strong>о</strong>бр<strong>а</strong>з жизни (п<strong>о</strong>д<strong>о</strong>тряд<br />

Apoxypodes), <strong>а</strong> н<strong>а</strong>чин<strong>а</strong>я <strong>с</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> <strong>с</strong>т<strong>а</strong>ли типичн<strong>о</strong><br />

н<strong>а</strong>земными жи<strong>в</strong><strong>о</strong>тными (п<strong>о</strong>д<strong>о</strong>тряд<br />

Dionychopodes). В <strong>с</strong><strong>о</strong><strong>в</strong>ременную эп<strong>о</strong>ху пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>тряд Scorpiones, <strong>в</strong>иды <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н<strong>а</strong>х (<strong>с</strong>м.<br />

<strong>с</strong>тр. 423).<br />

чи<strong>с</strong>ленные <strong>о</strong><strong>с</strong>яз<strong>а</strong>тельные <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и или <strong>с</strong><strong>о</strong>хр<strong>а</strong>няющие<strong>с</strong>я<br />

<strong>о</strong>т них п<strong>о</strong>ры, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> н<strong>а</strong> пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тях <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Н<strong>а</strong><br />

тергит<strong>а</strong>х мез<strong>о</strong><strong>с</strong><strong>о</strong>мы ин<strong>о</strong>гд<strong>а</strong> были р<strong>а</strong>з<strong>в</strong>иты <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

цилиндриче<strong>с</strong><strong>к</strong>ие, <strong>в</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные друг <strong>в</strong><br />

друг<strong>а</strong> труб<strong>о</strong>ч<strong>к</strong>и неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>зн<strong>а</strong>чения, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>в</strong>д<strong>о</strong>ль <strong>с</strong>рединн<strong>о</strong>й линии (у Tylopterella<br />

Stormer). Орн<strong>а</strong>мент<strong>а</strong>ция хитин<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong> Eurypterida р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong> и бы<strong>в</strong><strong>а</strong>ет<br />

<strong>с</strong>х<strong>о</strong>дн<strong>о</strong>й у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей р<strong>а</strong>зных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, и<br />

н<strong>а</strong><strong>о</strong>б<strong>о</strong>р<strong>о</strong>т, р<strong>а</strong>зличн<strong>о</strong>й у близ<strong>к</strong>их ф<strong>о</strong>рм. Н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong>ледующие ф<strong>о</strong>рмы <strong>о</strong>рн<strong>а</strong>мент<strong>а</strong>ции:<br />

1) буг<strong>о</strong>рч<strong>а</strong>т<strong>а</strong>я и ш<strong>а</strong>грене<strong>в</strong><strong>а</strong>я, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляющие<br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>й мел<strong>к</strong>ие или <strong>к</strong>рупные буг<strong>о</strong>р<strong>к</strong>и, нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итые н<strong>а</strong> р<strong>а</strong>зличных ч<strong>а</strong><strong>с</strong>тях тел<strong>а</strong>, или<br />

упл<strong>о</strong>щенные буг<strong>о</strong>р<strong>к</strong>и р<strong>а</strong>зличных <strong>в</strong>еличины и<br />

ф<strong>о</strong>рмы, н<strong>о</strong> <strong>в</strong> <strong>в</strong>иде <strong>с</strong>пл<strong>о</strong>шн<strong>о</strong>й ш<strong>а</strong>грене<strong>в</strong>идн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти;<br />

2) чешуйч<strong>а</strong>т<strong>а</strong>я, <strong>в</strong> <strong>в</strong>иде <strong>с</strong>ближенных<br />

или р<strong>а</strong>зреженных чешуеп<strong>о</strong>д<strong>о</strong>бных ут<strong>о</strong>лщений:<br />

шир<strong>о</strong><strong>к</strong>их б<strong>о</strong>лее или менее п<strong>о</strong>лу<strong>к</strong>рулых (т<strong>а</strong><strong>к</strong><br />

н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емых «п<strong>о</strong>лулунных»), язы<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных,<br />

перье<strong>о</strong>бр<strong>а</strong>зных <strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>с</strong>тр<strong>о</strong>уг<strong>о</strong>льных ут<strong>о</strong>лщений,<br />

н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющих перье<strong>в</strong><strong>о</strong>й п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> птиц;<br />

3) ямч<strong>а</strong>т<strong>а</strong>я, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я углублениями р<strong>а</strong>зличных<br />

<strong>в</strong>еличины и ф<strong>о</strong>рмы. Н<strong>а</strong> <strong>о</strong>тдельных уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>х<br />

тел<strong>а</strong> (н<strong>а</strong>пример, н<strong>а</strong> <strong>с</strong>егмент<strong>а</strong>х <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

Megalograptus ohioensis Caster et Kjellesvig-Waering),<br />

<strong>в</strong>ме<strong>с</strong>те <strong>с</strong> чешуй<strong>к</strong><strong>а</strong>ми р<strong>а</strong>з<strong>в</strong>иты<br />

еще и штрих<strong>о</strong><strong>в</strong><strong>а</strong>тые ребрыш<strong>к</strong>и.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> (ри<strong>с</strong>. 1189 А) не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я:<br />

ее ф<strong>о</strong>рм<strong>а</strong> <strong>о</strong>чень р<strong>а</strong>зличн<strong>а</strong> у р<strong>а</strong>зных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

В передней, <strong>с</strong>редней, реже <strong>в</strong> з<strong>а</strong>дней (у<br />

Melbournopterus Caster et Kjellesvig-Waering)<br />

ч<strong>а</strong><strong>с</strong>тях пр<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong>з<strong>в</strong>иты <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>г<strong>о</strong>,<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> или п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> <strong>о</strong>черт<strong>а</strong>ния,<br />

р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные п<strong>о</strong> <strong>в</strong>еличине (ри<strong>с</strong>. 1189Б,<br />

В). Кр<strong>о</strong>ме <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з, н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме имеют<strong>с</strong>я

Ри<strong>с</strong>. 1189.М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы у<br />

А — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> Hughmitleria <strong>с</strong>низу;'' Б'— 7личин<strong>к</strong><strong>а</strong> Stylonurus<br />

myops Clarke; Х'ЬО, <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>; В — <strong>с</strong>л<strong>о</strong>жный гл<strong>а</strong>з<br />

(<strong>с</strong>. г.) и ф<strong>а</strong><strong>с</strong>ет<strong>к</strong>и \ф) Pterygotus sp., <strong>с</strong>илур,» Э<strong>с</strong>т<strong>о</strong>ния; Г — <strong>с</strong>рединные<br />

гл<strong>а</strong>з<strong>к</strong>и ^Eurypterus fischeri Eichwald,^ Х4,0; эп — эпи<strong>с</strong>т<strong>о</strong>м<strong>а</strong>;<br />

пр<strong>о</strong><strong>с</strong>тые гл<strong>а</strong>з<strong>а</strong> — гл<strong>а</strong>з<strong>к</strong>и, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные н<strong>а</strong><br />

<strong>с</strong>рединн<strong>о</strong>й пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й линии пр<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>е (ри<strong>с</strong>. 1189 Г). Гл<strong>а</strong>з<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

близ<strong>к</strong><strong>о</strong> друг <strong>к</strong> другу, ин<strong>о</strong>гд<strong>а</strong> неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>я<strong>с</strong>ь <strong>о</strong>дин друг<strong>о</strong>г<strong>о</strong>.<br />

Дублюр<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ет ме<strong>с</strong>т<strong>а</strong><br />

при<strong>к</strong>репления ше<strong>с</strong>ти п<strong>а</strong>р <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>о</strong><strong>к</strong>руж<strong>а</strong>ющих р<strong>о</strong>т (ри<strong>с</strong>. 1189 А). Пер<strong>в</strong><strong>а</strong>я<br />

п<strong>а</strong>р<strong>а</strong> — <strong>к</strong>лешне<strong>в</strong>идные трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые хелицеры,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>ып<strong>о</strong>лняли фун<strong>к</strong>ции <strong>к</strong><strong>о</strong>лющих или<br />

х<strong>в</strong><strong>а</strong>т<strong>а</strong>ющих <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1190). У Pterygotidae<br />

<strong>о</strong>ни <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, и <strong>о</strong>б<strong>а</strong> члени<strong>к</strong><strong>а</strong><br />

<strong>к</strong>лешней <strong>с</strong> <strong>о</strong><strong>с</strong>трыми, д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> м<strong>о</strong>щными зубц<strong>а</strong>ми.<br />

Следующие пять п<strong>а</strong>р (II—VI) <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>о</strong>бычн<strong>о</strong> л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>т<strong>о</strong>рные: или <strong>в</strong><strong>с</strong>е х<strong>о</strong>дильные<br />

(Stylonurina), или же <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти п<strong>о</strong><strong>с</strong>ледней<br />

п<strong>а</strong>ры (VI п<strong>а</strong>р<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей)<br />

<strong>в</strong>ид<strong>о</strong>изменены <strong>в</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные гребные н<strong>о</strong>ги<br />

(Eurypterina). Обычн<strong>о</strong> пер<strong>в</strong><strong>о</strong>й п<strong>а</strong>р<strong>о</strong>й л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>т<strong>о</strong>рных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей я<strong>в</strong>ляет<strong>с</strong>я II п<strong>а</strong>р<strong>а</strong>, или педи<strong>а</strong>льпы.<br />

Ин<strong>о</strong>гд<strong>а</strong> педи<strong>а</strong>льпы и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти III<br />

п<strong>а</strong>ры пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> <strong>с</strong>л<strong>о</strong>жн<strong>о</strong> п<strong>о</strong><strong>с</strong>тр<strong>о</strong>енные х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные<br />

<strong>о</strong>рг<strong>а</strong>ны (Megalograptidae, Mixopteridae).<br />

Х<strong>о</strong>дильные н<strong>о</strong>ги б<strong>о</strong>лее или менее <strong>о</strong>дн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ения, из цилиндриче<strong>с</strong><strong>к</strong>их члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>с</strong> шип<strong>а</strong>ми или без шип<strong>о</strong><strong>в</strong>. Пер<strong>в</strong>ый, члени<strong>к</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>й<br />

т<strong>а</strong><strong>к</strong><strong>о</strong>й н<strong>о</strong>ги я<strong>в</strong>ляет<strong>с</strong>я б<strong>о</strong>лее или менее шир<strong>о</strong><strong>к</strong>им<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong>м (<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>, <strong>с</strong><strong>о</strong>х<strong>а</strong>) <strong>с</strong> неб<strong>о</strong>льшими<br />

эпи<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ит<strong>а</strong>ми у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния зубч<strong>а</strong>т<strong>о</strong>г<strong>о</strong> р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я т<strong>а</strong>зи<strong>к</strong><strong>а</strong> (ри<strong>с</strong>. 1190, Б—Г). У Eurypterus<br />

т<strong>а</strong>зи<strong>к</strong> V н<strong>о</strong>ги <strong>с</strong> неб<strong>о</strong>льшими <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляющими,<br />

п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, <strong>с</strong>лух<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>о</strong>рг<strong>а</strong>н<br />

(ри<strong>с</strong>. 1190, В). Кр<strong>о</strong>ме т<strong>а</strong>зи<strong>к</strong><strong>а</strong>, х<strong>о</strong>дильные н<strong>о</strong>ги<br />

А<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Eurypterida (Stormer, 1955)<br />

р — р<strong>о</strong>т; эн — энд<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong>; м.<strong>к</strong>. — ме<strong>с</strong>т<strong>а</strong> при<strong>к</strong>репления <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей;<br />

м.м. — ме<strong>с</strong>т<strong>о</strong> при<strong>к</strong>репления мет<strong>а</strong><strong>с</strong>т<strong>о</strong>мы; <strong>с</strong>е. — <strong>с</strong>л<strong>о</strong>жные<br />

гл<strong>а</strong>з<strong>а</strong>; ф — ф<strong>а</strong><strong>с</strong>ет<strong>к</strong>и <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з<br />

Ри<strong>с</strong>. 1190. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы (/, IV, V, VI) Eurypterus.<br />

fischeri Eichwald, <strong>с</strong>илур, Э<strong>с</strong>т<strong>о</strong>ния<br />

А — трехчлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я хелицер<strong>а</strong>; Б — х<strong>о</strong>дильн<strong>а</strong>я н<strong>о</strong>г<strong>а</strong> чет<strong>в</strong>ерт<strong>о</strong>й<br />

п<strong>а</strong>ры; В — х<strong>о</strong>дильн<strong>а</strong>я н<strong>о</strong>г<strong>а</strong> пят<strong>о</strong>й п<strong>а</strong>ры; Г — пл<strong>а</strong><strong>в</strong><strong>а</strong>тельн<strong>а</strong>я н<strong>о</strong>г<strong>а</strong>;<br />

э<strong>к</strong> — эпи<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>иты, <strong>с</strong>.<strong>о</strong>.?<strong>с</strong>лух<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>о</strong>рг<strong>а</strong>н, <strong>к</strong> — т<strong>а</strong>зи<strong>к</strong> (<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>).<br />

л. <strong>о</strong>.—гребн<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть, xl (St0rmer, 1955)<br />

405<br />

http://jurassic.ru/

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из <strong>с</strong>еми — <strong>в</strong><strong>о</strong><strong>с</strong>ьми члени<strong>к</strong><strong>о</strong><strong>в</strong>: <strong>в</strong>ертлуги<br />

(тр<strong>о</strong>х<strong>а</strong>нтер<strong>а</strong>), <strong>к</strong><strong>о</strong>т<strong>о</strong>рый бы<strong>в</strong><strong>а</strong>ет д<strong>в</strong><strong>о</strong>йным <strong>в</strong> V<br />

н<strong>о</strong>ге), д<strong>в</strong><strong>о</strong>йн<strong>о</strong>г<strong>о</strong> бедр<strong>а</strong> (praefemur и femur),<br />

ч<strong>а</strong>шеч<strong>к</strong>и (patella), г<strong>о</strong>лени (tibia), л<strong>а</strong>п<strong>к</strong>и (tarsus)<br />

<strong>с</strong> ди<strong>с</strong>т<strong>а</strong>льными шип<strong>а</strong>ми, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующими<br />

прет<strong>а</strong>рзу (praetarsus) н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х<br />

имеют<strong>с</strong>я неп<strong>о</strong>д<strong>в</strong>ижные или <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> п<strong>о</strong>д<strong>в</strong>ижные<br />

шипы н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти.<br />

От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие шип<strong>о</strong><strong>в</strong> н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ф<strong>о</strong>рм<br />

<strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует <strong>о</strong> примити<strong>в</strong>н<strong>о</strong><strong>с</strong>ти <strong>с</strong>тр<strong>о</strong>ения.<br />

У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>о</strong>д<strong>о</strong><strong>в</strong> шипы <strong>в</strong>ид<strong>о</strong>изменены <strong>в</strong> <strong>с</strong>пеци<strong>а</strong>льные<br />

длинные иглы, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>о</strong>бл<strong>а</strong>д<strong>а</strong><strong>в</strong>шие<br />

яд<strong>о</strong><strong>в</strong>итым <strong>с</strong>е<strong>к</strong>рет<strong>о</strong>м (у Megalograptidae и<br />

Mixopteridae).<br />

У Eurypterina ше<strong>с</strong>т<strong>а</strong>я п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> для пл<strong>а</strong><strong>в</strong><strong>а</strong>ния. Д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong><strong>с</strong>ледние члени<strong>к</strong><strong>а</strong> этих н<strong>о</strong>г, б<strong>о</strong>лее или менее<br />

р<strong>а</strong><strong>с</strong>ширенные, <strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong>е<strong>с</strong>л<strong>о</strong><strong>в</strong>ые л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти или<br />

греб<strong>к</strong>и (ри<strong>с</strong>. 1190, Г). У Dolichopteridae греб<strong>к</strong>и<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты и имеют я<strong>в</strong>н<strong>о</strong> перех<strong>о</strong>дный<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер <strong>о</strong>т х<strong>о</strong>дильных <strong>к</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong>тельным н<strong>о</strong>г<strong>а</strong>м<br />

(или, н<strong>а</strong><strong>о</strong>б<strong>о</strong>р<strong>о</strong>т, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют п<strong>о</strong><strong>с</strong>тепенную<br />

утр<strong>а</strong>ту пл<strong>а</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>й фун<strong>к</strong>ции?).<br />

406<br />

http://jurassic.ru/<br />

П<strong>о</strong>з<strong>а</strong>ди рт<strong>а</strong> д<strong>в</strong>е п<strong>о</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и — энд<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> и мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong>;<br />

п<strong>о</strong><strong>с</strong>ледняя, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичн<strong>а</strong> хил<strong>а</strong>риям<br />

Xiphosura и прин<strong>а</strong>длежит пер<strong>в</strong><strong>о</strong>му <strong>с</strong>егменту<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Опер<strong>к</strong>улум (ри<strong>с</strong>. 1191) пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет прид<strong>а</strong>т<strong>к</strong>и<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> или генит<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

тел<strong>а</strong> и я<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong>т<strong>о</strong>рым <strong>с</strong>егмент<strong>о</strong>м <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

и длинные, ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>щепленные у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>;<br />

<strong>с</strong>ледует <strong>о</strong>тметить, чт<strong>о</strong> д<strong>о</strong> <strong>с</strong>их п<strong>о</strong>р <strong>о</strong>пределение<br />

прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ти этих <strong>с</strong>тру<strong>к</strong>тур <strong>к</strong> т<strong>о</strong>му или<br />

друг<strong>о</strong>му п<strong>о</strong>лу еще не я<strong>с</strong>н<strong>о</strong>.<br />

Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> н<strong>а</strong>иб<strong>о</strong>лее х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> у <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

Eurypteridae, Hughmilleriidae, Dolichopteridae,<br />

и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у Mixopteridae, <strong>в</strong> т<strong>о</strong><br />

<strong>в</strong>ремя <strong>к</strong><strong>а</strong><strong>к</strong> у Stylonuridae (<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у р<strong>о</strong>д<strong>а</strong><br />

Stylonurus Page) <strong>о</strong>н<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, и <strong>в</strong><strong>с</strong>я <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

имеет <strong>в</strong>ид <strong>к</strong>лин<strong>а</strong>. П<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент мет<strong>а</strong><strong>с</strong><strong>о</strong>мы<br />

(претель<strong>с</strong><strong>о</strong>н) бы<strong>в</strong><strong>а</strong>ет <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми<br />

(эпимер<strong>а</strong>ми). Эпимеры м<strong>о</strong>гут быть<br />

р<strong>а</strong>з<strong>в</strong>иты и н<strong>а</strong> других <strong>с</strong>егмент<strong>а</strong>х мет<strong>а</strong>- и мез<strong>о</strong><strong>с</strong><strong>о</strong>мы.

Тель<strong>с</strong><strong>о</strong>н б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный,<br />

уз<strong>к</strong>ий или б<strong>о</strong>лее или менее шир<strong>о</strong><strong>к</strong>ий,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>уженный <strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>ей <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти (у<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей р<strong>а</strong>зных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Stylonuridae—Drepanopterus<br />

Laurie, Eurypteridae —<br />

Onychopterella Stormer, Hughmilleriidae —<br />

Salteropterus Kjellesvig-Waering). Реже (у<br />

Pterygotidae) ф<strong>о</strong>рм<strong>а</strong> тель<strong>с</strong><strong>о</strong>н<strong>а</strong> б<strong>о</strong>лее или менее<br />

л<strong>о</strong>п<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я <strong>с</strong> <strong>к</strong>лин<strong>о</strong><strong>в</strong>идным <strong>к</strong>илем н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и. П<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

<strong>к</strong>иль пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>с</strong><strong>о</strong>б<strong>о</strong>й <strong>о</strong>бычный <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный<br />

тель<strong>с</strong><strong>о</strong>н, <strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> — <strong>с</strong>ильн<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итые эпимеры тель<strong>с</strong><strong>о</strong>н<strong>а</strong>. У Slimonia эт<strong>а</strong><br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> тель<strong>с</strong><strong>о</strong>н<strong>а</strong> н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет н<strong>а</strong><strong>к</strong><strong>о</strong>нечни<strong>к</strong><br />

<strong>к</strong><strong>о</strong>пья. Тель<strong>с</strong><strong>о</strong>н <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных Carcinosomatidae<br />

и Mixopteridae имеет <strong>в</strong>ид уз<strong>к</strong><strong>о</strong>г<strong>о</strong>, длинн<strong>о</strong>г<strong>о</strong>,<br />

из<strong>о</strong>гнут<strong>о</strong>г<strong>о</strong> шип<strong>а</strong> <strong>с</strong> ут<strong>о</strong>лщенным <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

(<strong>к</strong><strong>о</strong>гте<strong>о</strong>бр<strong>а</strong>зный).<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

Среди типичн<strong>о</strong>й м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ф<strong>а</strong>уны Eurypterida <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>а</strong>х<strong>о</strong>ждение<br />

их <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> <strong>в</strong> <strong>к</strong><strong>о</strong>нтинент<strong>а</strong>льных <strong>о</strong>тл<strong>о</strong>жениях<br />

у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет н<strong>а</strong> т<strong>о</strong>, чт<strong>о</strong> эти хелицер<strong>о</strong><strong>в</strong>ые <strong>о</strong>бит<strong>а</strong>ли<br />

<strong>в</strong> пре<strong>с</strong>н<strong>о</strong>й или <strong>с</strong><strong>о</strong>л<strong>о</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>й <strong>в</strong><strong>о</strong>де. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

был<strong>о</strong>, <strong>о</strong>че<strong>в</strong>идн<strong>о</strong>, д<strong>о</strong>нными <strong>о</strong>бит<strong>а</strong>телями, х<strong>о</strong>тя<br />

<strong>о</strong>бл<strong>а</strong>д<strong>а</strong><strong>в</strong>шие пл<strong>а</strong><strong>в</strong><strong>а</strong>тельными н<strong>о</strong>г<strong>а</strong>ми м<strong>о</strong>гли <strong>а</strong><strong>к</strong>ти<strong>в</strong>н<strong>о</strong><br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>ть. Птериг<strong>о</strong>тиды, <strong>с</strong><strong>к</strong><strong>о</strong>рее <strong>в</strong><strong>с</strong>ег<strong>о</strong>,<br />

были пл<strong>о</strong>х<strong>о</strong>п<strong>о</strong>д<strong>в</strong>ижными ф<strong>о</strong>рм<strong>а</strong>ми не<strong>к</strong>т<strong>о</strong>н<strong>а</strong>;<br />

<strong>с</strong>л<strong>а</strong>б<strong>а</strong>я <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ть <strong>к</strong> перед<strong>в</strong>ижению <strong>в</strong><strong>о</strong><strong>с</strong>п<strong>о</strong>лнял<strong>а</strong><strong>с</strong>ь<br />

у них м<strong>о</strong>щными п<strong>о</strong>д<strong>в</strong>ижными <strong>к</strong>лешнями,<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ля<strong>в</strong>шими н<strong>а</strong><strong>с</strong>тиг<strong>а</strong>ть д<strong>о</strong>бычу. Б<strong>о</strong>лее<br />

<strong>а</strong><strong>к</strong>ти<strong>в</strong>ными хищни<strong>к</strong><strong>а</strong>ми, <strong>к</strong> т<strong>о</strong>му же х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong><strong>в</strong>шими,<br />

были, <strong>о</strong>че<strong>в</strong>идн<strong>о</strong>, Mixopteridea <strong>с</strong> передним<br />

п<strong>о</strong>л<strong>о</strong>жением <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з и <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итыми передними н<strong>о</strong>г<strong>а</strong>ми, м<strong>о</strong>щные<br />

иглы <strong>к</strong><strong>о</strong>т<strong>о</strong>рых я<strong>в</strong>н<strong>о</strong> были при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блением<br />

хищни<strong>к</strong><strong>а</strong>.<br />

Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — пермь. Р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены были<br />

п<strong>о</strong>чти <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех ч<strong>а</strong><strong>с</strong>тях С<strong>в</strong>ет<strong>а</strong>, н<strong>о</strong> немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

(т<strong>а</strong>бл. 3).<br />

П<strong>о</strong>л<strong>о</strong>жение<strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Eurypterida<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, другим <strong>в</strong><strong>о</strong>дным<br />

хелицер<strong>о</strong><strong>в</strong>ым— мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>м, <strong>с</strong> друг<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

н<strong>а</strong>земным хелицер<strong>о</strong><strong>в</strong>ым — <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>нид<strong>а</strong>м.<br />

Eurypterida, <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бные не т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong><br />

х<strong>о</strong>ждению, н<strong>о</strong> и <strong>к</strong> пл<strong>а</strong><strong>в</strong><strong>а</strong>нию (п<strong>о</strong>д<strong>о</strong>тряд Eurypterina),<br />

<strong>в</strong>ер<strong>о</strong>ятнее <strong>в</strong><strong>с</strong>ег<strong>о</strong>, были п<strong>о</strong>т<strong>о</strong>м<strong>к</strong><strong>а</strong>ми мечех<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><br />

Aglaspidida. П<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong> <strong>с</strong><br />

ними <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли и Eurypterida, <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бные<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong> х<strong>о</strong>ждению (п<strong>о</strong>д<strong>о</strong>тряд Stylonurina).<br />

С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бные Mixopteridea п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong><br />

Eurypterina, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, были близ<strong>к</strong>ими<br />

пред<strong>к</strong><strong>а</strong>ми н<strong>а</strong>земных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>. Т<strong>а</strong><strong>к</strong>им <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м,<br />

р<strong>а</strong>з<strong>в</strong>итие <strong>о</strong>беих фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их <strong>в</strong>ет<strong>в</strong>ей:<br />

х<strong>о</strong>дяще-пл<strong>а</strong><strong>в</strong><strong>а</strong>ющей и т<strong>о</strong>ль<strong>к</strong><strong>о</strong> х<strong>о</strong>дящей — пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дил<strong>о</strong><br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>, н<strong>о</strong> х<strong>о</strong>дящие фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>и<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> м<strong>о</strong>л<strong>о</strong>же. П<strong>о</strong>д<strong>о</strong>тряды:<br />

Eurypterina, Stylonurina.<br />

Т<strong>а</strong>блиц<strong>а</strong> 3<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция и <strong>с</strong>тр<strong>а</strong>тигр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>а</strong> Eurypterida<br />

Eurypterida<br />

П<strong>о</strong>д<strong>о</strong>тряд Eurypterina<br />

У<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ленный<br />

ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridea<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hughmilleriidae:<br />

Hughmilleria В .<strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong>—<strong>с</strong>р.де<strong>в</strong><strong>о</strong>н<br />

Salteropterus Н. де<strong>в</strong><strong>о</strong>н<br />

Grossopterus Н. и <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н<br />

Lepidoderma В. де<strong>в</strong><strong>о</strong>н — <strong>в</strong>. пермь<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Slimoniidae:<br />

Slimonia В. <strong>с</strong>илур<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridae:<br />

Eurypterus Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — н. де<strong>в</strong><strong>о</strong>н<br />

Erieopterus Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — де<strong>в</strong><strong>о</strong>н<br />

Tylopterella В. <strong>с</strong>илур — н. де<strong>в</strong><strong>о</strong>н<br />

Onychopterella В. <strong>с</strong>илур<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dolichopteridae:<br />

Dolichopterus Силур<br />

Strobllopterus Н. де<strong>в</strong><strong>о</strong>н<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pterygotidea<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pterygotidae:<br />

Pterygotus Силур — <strong>в</strong>. де<strong>в</strong><strong>о</strong>н<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mixopteridea<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mixopteridae:<br />

Mixopterus В. <strong>с</strong>илур — д<strong>а</strong>унт<strong>о</strong>н<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Megalograptidae:<br />

407<br />

http://jurassic.ru/<br />

Megalograptus Орд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

Echinognatus Орд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Carcinosomatidae;<br />

Carcinosoma Орд<strong>о</strong><strong>в</strong>и<strong>к</strong>-<strong>с</strong>илур<br />

Rhinocarcinosoma Силур

Eurypterida<br />

П<strong>о</strong>д<strong>о</strong>тряд Stylonurina<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Stylonuri<br />

dea<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Stylonuridae<br />

П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Stylonurinae:<br />

Stytonurus<br />

Drepanopterus<br />

Brachyopterus<br />

Ctenopterus<br />

Tarsopterella<br />

П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mar<br />

supipterinae:<br />

Marsupipterus<br />

Polystomurum<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hibbertopteridae:<br />

Hibbertopterus<br />

Campylocephalus<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhenopteridae:<br />

Rhenopterus<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mycteropidea<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mycteropidae:<br />

Mycterops<br />

Borchgrevinkium<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Woodwardopteridae:<br />

Woodwardopterus<br />

Т<strong>а</strong>блиц<strong>а</strong> 3 (<strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>ние)<br />

У<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ленный<br />

ге<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ий <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<br />

Силур — <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н<br />

В. <strong>с</strong>илур — <strong>в</strong>. де<strong>в</strong><strong>о</strong>н<br />

Ср. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> - д<strong>а</strong>унт<strong>о</strong>н<br />

Силур — <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н<br />

Н. и <strong>в</strong>. де<strong>в</strong><strong>о</strong>н<br />

Д<strong>а</strong>унт<strong>о</strong>н<br />

В. де<strong>в</strong><strong>о</strong>н<br />

В. де<strong>в</strong><strong>о</strong>н — н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н — <strong>в</strong>. пермь<br />

Н. де<strong>в</strong><strong>о</strong>н<br />

В. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Н. де<strong>в</strong><strong>о</strong>н<br />

Н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

ПОДОТРЯД EURYPTERINA<br />

[nom. transl. Novojilov, hie (ex<br />

Eurypteridae Burmeiter, 1845)]<br />

П<strong>о</strong><strong>с</strong>ледняя п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> <strong>в</strong>иде<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных гребных н<strong>о</strong>г: п<strong>о</strong><strong>с</strong>ледние их<br />

408<br />

http://jurassic.ru/<br />

члени<strong>к</strong>и р<strong>а</strong><strong>с</strong>ширены и <strong>о</strong>бр<strong>а</strong>зуют гребные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти.<br />

Орд<strong>о</strong><strong>в</strong>и<strong>к</strong>—пермь. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Eurypteridea,<br />

Pterygotidea, Mixopteridea.<br />

НАДСЕМЕЙСТВО<br />

EURYPTERIDEA BURMEISTER, 1845<br />

[nom. corr. Novojilov, hie (ex Eurypteraceae<br />

Stormer, 1951 )1<br />

Хелицеры <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итые; мез<strong>о</strong>и<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> р<strong>а</strong>згр<strong>а</strong>ничены д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong>. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong>—пермь.<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Hughmilleriidae,<br />

Slimoniidae, Eurypteridae, Dolichopteridae.<br />

СЕМЕЙСТВО HUGHMILLERIIDAE<br />

KJELLESVIG-WAERING, 1951<br />

Длин<strong>а</strong> ф<strong>о</strong>рм 7—30 <strong>с</strong>м; н<strong>а</strong>ружн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

<strong>с</strong> п<strong>о</strong>перечными <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong>ми, или п<strong>о</strong>лу<strong>к</strong>руглыми<br />

«чешуй<strong>к</strong><strong>а</strong>ми», <strong>к</strong>рупными или мел<strong>к</strong>ими;<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены у передней<br />

ч<strong>а</strong><strong>с</strong>ти б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н или близ <strong>с</strong>ередины<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; дублюр<strong>а</strong> <strong>с</strong> эпи<strong>с</strong>т<strong>о</strong>м<strong>о</strong>й; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я: генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> уз<strong>к</strong>ий,<br />

без б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>к</strong><strong>о</strong>нечий; хелицеры без<br />

зубч<strong>а</strong>т<strong>о</strong><strong>с</strong>ти, х<strong>о</strong>дильные н<strong>о</strong>ги <strong>с</strong> шип<strong>а</strong>ми или зубчи<strong>к</strong><strong>а</strong>ми;<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти уз<strong>к</strong>ие; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>членяет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширенную<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>му и <strong>к</strong>лин<strong>о</strong><strong>в</strong>идную мет<strong>а</strong><strong>с</strong><strong>о</strong>му; тель<strong>с</strong><strong>о</strong>н<br />

л<strong>а</strong>нцет<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный или р<strong>а</strong><strong>с</strong>ширенный <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии.<br />

Четыре р<strong>о</strong>д<strong>а</strong>. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — пермь.<br />

Hughtnilleria Sarle, 1903. Тип р<strong>о</strong>д<strong>а</strong> — Я.<br />

socialis Sarle, 1902; н. <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

п<strong>о</strong>луэллиптиче<strong>с</strong><strong>к</strong><strong>а</strong>я д<strong>о</strong> п<strong>о</strong>лу<strong>к</strong>ругл<strong>о</strong>й; <strong>с</strong>л<strong>о</strong>жные<br />

гл<strong>а</strong>з<strong>а</strong> близ <strong>к</strong>р<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

ч<strong>а</strong><strong>с</strong>тях; дублюр<strong>а</strong> <strong>с</strong> <strong>о</strong>тчетли<strong>в</strong>ыми эпи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льными<br />

ш<strong>в</strong><strong>а</strong>ми и менее <strong>о</strong>тчетли<strong>в</strong>ыми<br />

переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми ш<strong>в</strong><strong>а</strong>ми; генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong><br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и шир<strong>о</strong><strong>к</strong>ий; мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>ч<strong>о</strong>н<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>черт<strong>а</strong>ния, н<strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лыми<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>к</strong>р<strong>а</strong>ями, п<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> з<strong>а</strong>днеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми угл<strong>а</strong>ми; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong>уж<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я <strong>к</strong>лин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>; тель<strong>с</strong><strong>о</strong>н<br />

<strong>к</strong>лин<strong>о</strong><strong>в</strong>идный; <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong><br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> ряд<strong>а</strong>ми пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>, п<strong>о</strong>х<strong>о</strong>жих<br />

н<strong>а</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые у Pterygotus, н<strong>о</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>н<strong>а</strong><br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> «терр<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>ыми» линиями (ри<strong>с</strong>.<br />

1192; т<strong>а</strong>бл. XVIII, фиг. 1—4). 15 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

К<strong>а</strong>н<strong>а</strong>ды; н. и <strong>в</strong>. <strong>с</strong>илур С. Амери<strong>к</strong>и, Ш<strong>о</strong>тл<strong>а</strong>ндии,<br />

Англии; ?н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) С<strong>к</strong><strong>а</strong>ндин<strong>а</strong><strong>в</strong>ии,<br />

?н. и <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н Азии (Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>яр<strong>с</strong><strong>к</strong>ий<br />

<strong>к</strong>р<strong>а</strong>й, Ту<strong>в</strong><strong>а</strong>).<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong>не СССР: Salteropterus Kjellesvig-Waering,<br />

1951 (н. де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы);

1194Б<br />

Ри<strong>с</strong>. 1192—1196. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Hughmilleriidae и Slimoniidae (St0rmer, 1955)<br />

1192. Hughmllleria norvegica (Kjaer)?; н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н), Н<strong>о</strong>р<strong>в</strong>егия;<br />

А — <strong>с</strong><strong>в</strong>ерху; Б — <strong>с</strong>низу, <strong>с</strong> ме<strong>с</strong>т<strong>а</strong>ми ж<strong>а</strong>берных прид<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы (пун<strong>к</strong>тир), Х0,5; В — генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и <strong>с</strong><strong>а</strong>мц<strong>а</strong>;<br />

Г — т<strong>о</strong> же у <strong>с</strong><strong>а</strong>м<strong>к</strong>и/Xl. 1193. Тель<strong>с</strong><strong>о</strong>н Salteropterus abbtevlatus<br />

(Salter), X 1,0? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н), Англия. 1194. Grossopterus<br />

overatht (Gross); А — <strong>с</strong><strong>в</strong>ерху, x0,25, Б — генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong>?^<strong>с</strong><strong>а</strong>мц<strong>а</strong>,<br />

Х0,3; н. де<strong>в</strong><strong>о</strong>н, Герм<strong>а</strong>ния. 1195. Lepldoderma Reuss:<br />

4Q9<br />

1194А<br />

А — L. mansfteldt (С. Е. Hall), <strong>с</strong><strong>в</strong>ерху, Х1,3, <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С.-Амери<strong>к</strong><strong>а</strong>;<br />

Б, В — L. masonense (Meek et Worthen), Xl; Б — генит<strong>а</strong>льный<br />

прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и, В — мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> и пл<strong>а</strong><strong>в</strong><strong>а</strong>тельн<strong>а</strong>я н<strong>о</strong>г<strong>а</strong>,<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> . 1196. Slimonia acuminata (Salter); А —<br />

<strong>с</strong><strong>в</strong>ерху, Х0,1; Б — генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и <strong>с</strong><strong>а</strong>м<strong>к</strong>и (?), В — т<strong>о</strong><br />

же у <strong>с</strong><strong>а</strong>мц<strong>а</strong> (?), х0,13; <strong>с</strong>илур, Ш<strong>о</strong>тл<strong>а</strong>ндия<br />

http://jurassic.ru/

Grossopterus Stormer, 1934 (н. и <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н 3.<br />

Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и); Lepidoderma Reuss, 1855<br />

(<strong>в</strong>. де<strong>в</strong><strong>о</strong>н — <strong>в</strong>. пермь 3. Е<strong>в</strong>р<strong>о</strong>пы, С. Амери<strong>к</strong>и,<br />

Азии) (ри<strong>с</strong>. 1193—1195).<br />

СЕМЕЙСТВО SLIMONIIDAE NOVOJILOV,<br />

FAM. NOV.<br />

Длин<strong>а</strong> ф<strong>о</strong>рм <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 <strong>с</strong>м; п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тел<strong>а</strong><br />

гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> н<strong>а</strong> угл<strong>а</strong>х переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я рез<strong>к</strong><strong>о</strong> притуплённ<strong>о</strong>й, б<strong>о</strong>лее или менее<br />

прям<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> уз<strong>к</strong><strong>о</strong><strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>й<br />

ф<strong>о</strong>рмы; генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong><br />

<strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>о</strong><strong>с</strong>тр<strong>о</strong><strong>к</strong><strong>о</strong>нечиями; педип<strong>а</strong>льпы<br />

т<strong>о</strong>н<strong>к</strong>ие, 'без шип<strong>о</strong><strong>в</strong>; тель<strong>с</strong><strong>о</strong>н р<strong>а</strong><strong>с</strong>ширенный, <strong>в</strong><br />

<strong>в</strong>иде пл<strong>а</strong><strong>в</strong>ни<strong>к</strong><strong>а</strong>. Один р<strong>о</strong>д — Slimonia Page,<br />

1856. В. <strong>с</strong>илур 3. Е<strong>в</strong>р<strong>о</strong>пы (ри<strong>с</strong>. 1196).<br />

СЕМЕЙСТВО EURYPTERIDAE<br />

BURMEISTER, 1845<br />

Ф<strong>о</strong>рмы 10—30 <strong>с</strong>м длины; <strong>с</strong> <strong>о</strong>тчетли<strong>в</strong><strong>о</strong>й чешуе-<br />

или перьебр<strong>а</strong>зн<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й или т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ребри<strong>с</strong>т<strong>о</strong>й; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>чти п<strong>о</strong>лу<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я<br />

или п<strong>о</strong>чти прям<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> б<strong>о</strong>лее<br />

или менее п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

близ <strong>с</strong>ередины пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; тель<strong>с</strong><strong>о</strong>н <strong>к</strong>лин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зный,<br />

реже <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>бым пережим<strong>о</strong>м; дублюр<strong>а</strong> без<br />

эпи<strong>с</strong>т<strong>о</strong>мы, мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я или <strong>с</strong>ердце<strong>в</strong>идн<strong>а</strong>я<br />

<strong>о</strong>переди; х<strong>о</strong>дильные н<strong>о</strong>ги у<strong>в</strong>еличи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> длине <strong>с</strong>переди н<strong>а</strong>з<strong>а</strong>д, <strong>с</strong> шип<strong>а</strong>ми или без шип<strong>о</strong><strong>в</strong>;<br />

гребные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й шп<strong>о</strong>р<strong>о</strong>й или без нее; генит<strong>а</strong>льный<br />

прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

д<strong>о</strong>лями у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, длинный, <strong>с</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я<br />

ди<strong>с</strong>т<strong>а</strong>льными шип<strong>а</strong>ми у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — <strong>в</strong>.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Четыре р<strong>о</strong>д<strong>а</strong>.<br />

Eurypterus De Kay, 1825. Тип р<strong>о</strong>д<strong>а</strong> — Е.<br />

remipes De Kay, 1825; <strong>в</strong>. <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> уз<strong>к</strong>им <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ым <strong>о</strong>б<strong>о</strong>д<strong>к</strong><strong>о</strong>м; дублюр<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong><strong>о</strong> <strong>с</strong>рединным ш<strong>в</strong><strong>о</strong>м; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я,<br />

<strong>с</strong> глуб<strong>о</strong><strong>к</strong>им зубч<strong>а</strong>тым <strong>в</strong>ырез<strong>о</strong>м <strong>о</strong>переди;<br />

передние три н<strong>а</strong>ры х<strong>о</strong>дильных н<strong>о</strong>г <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> шип<strong>а</strong>ми н<strong>а</strong> <strong>в</strong><strong>с</strong>ех члени<strong>к</strong><strong>а</strong>х, чет<strong>в</strong>ерт<strong>а</strong>я<br />

п<strong>а</strong>р<strong>а</strong> х<strong>о</strong>дильных н<strong>о</strong>г длиннее и <strong>с</strong> шип<strong>а</strong>ми<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м члени<strong>к</strong>е; гребные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширены,<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й шп<strong>о</strong>р<strong>о</strong>й. Н<strong>а</strong>ружный<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью чешуе<strong>в</strong>идный (ри<strong>с</strong>,<br />

1197). В<strong>о</strong><strong>с</strong>емь <strong>в</strong>ид<strong>о</strong><strong>в</strong> и <strong>с</strong>емь п<strong>о</strong>д<strong>в</strong>ид<strong>о</strong><strong>в</strong>. В. <strong>с</strong>илур<br />

(лудл<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий яру<strong>с</strong>) Э<strong>с</strong>т<strong>о</strong>нии, Н<strong>о</strong>р<strong>в</strong>егии,<br />

С. Амери<strong>к</strong>и; ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) Ш<strong>о</strong>тл<strong>а</strong>ндии;<br />

н. де<strong>в</strong><strong>о</strong>н США (В<strong>а</strong>й<strong>о</strong>минг), У<strong>к</strong>р<strong>а</strong>ины, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>,<br />

Сибири, Ш<strong>о</strong>тл<strong>а</strong>ндии, Фр<strong>а</strong>нции, Бельгии,<br />

Герм<strong>а</strong>нии, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии, пермь П<strong>о</strong>ртуг<strong>а</strong>лии,<br />

Кит<strong>а</strong>я, С. Амери<strong>к</strong>и.<br />

Erieopterus Kjellesvig-Waering,<br />

1958. Тип р<strong>о</strong>д<strong>а</strong> — Eurypterus microphthalmus<br />

410<br />

http://jurassic.ru/<br />

Hall, 1859; <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> уз*<br />

<strong>к</strong>им или шир<strong>о</strong><strong>к</strong>им <strong>о</strong>б<strong>о</strong>д<strong>о</strong>м; дублюр<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

без <strong>с</strong>рединн<strong>о</strong>г<strong>о</strong> ш<strong>в</strong><strong>а</strong>, жел<strong>о</strong>бч<strong>а</strong>т<strong>а</strong>я н<strong>а</strong> <strong>в</strong>нутреннем<br />

<strong>к</strong>р<strong>а</strong>е; х<strong>о</strong>дильные н<strong>о</strong>ги <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения,<br />

<strong>в</strong><strong>с</strong>е без шип<strong>о</strong><strong>в</strong> или <strong>с</strong> неп<strong>о</strong>д<strong>в</strong>ижными шип<strong>а</strong>ми<br />

н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льных члени<strong>к</strong><strong>а</strong>х: гребные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей шир<strong>о</strong><strong>к</strong>ие, <strong>с</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

шп<strong>о</strong>р<strong>о</strong>й <strong>в</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м <strong>в</strong>ырезе; гр<strong>а</strong>ниц<strong>а</strong><br />

между мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й б<strong>о</strong>лее рез<strong>к</strong><strong>а</strong>я,<br />

чем у Eurypterus; тель<strong>с</strong><strong>о</strong>н <strong>к</strong>лин<strong>о</strong><strong>в</strong>идный. Н<strong>а</strong>ружный<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> гл<strong>а</strong>д<strong>к</strong>ий или пузырьч<strong>а</strong>тый<br />

(т<strong>а</strong>бл. XX, фиг. 2). Де<strong>с</strong>ять <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong><br />

С. Амери<strong>к</strong>и; <strong>с</strong>илур Э<strong>с</strong>т<strong>о</strong>нии и С. Амери<strong>к</strong>и; де<strong>в</strong><strong>о</strong>н<br />

Ш<strong>о</strong>тл<strong>а</strong>ндии и С. Амери<strong>к</strong>и.<br />

Tylopterella Stormer, 1951 ( = Tylopterus<br />

Clarke et Ruedemann, 1912). Тип р<strong>о</strong>д<strong>а</strong> — Eurypterus<br />

boyli Whiteaves, 1884; <strong>в</strong>. <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong><br />

(Онт<strong>а</strong>ри<strong>о</strong>). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее или менее <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я,<br />

немн<strong>о</strong>г<strong>о</strong> <strong>с</strong>рез<strong>а</strong>нн<strong>а</strong>я з<strong>а</strong>тыл<strong>о</strong>чным <strong>к</strong>р<strong>а</strong>ем<br />

или <strong>о</strong><strong>к</strong>ругл<strong>о</strong>-<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные<br />

или <strong>о</strong><strong>в</strong><strong>а</strong>льные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные б<strong>о</strong>лее<br />

или менее близ<strong>к</strong><strong>о</strong> <strong>к</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ым <strong>к</strong>р<strong>а</strong>ям и н<strong>а</strong><br />

ур<strong>о</strong><strong>в</strong>не <strong>с</strong>ередины длины пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; гл<strong>а</strong>з<strong>к</strong>и между<br />

<strong>с</strong>л<strong>о</strong>жными гл<strong>а</strong>з<strong>а</strong>ми или немн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>з<strong>а</strong>ди;<br />

х<strong>о</strong>дильные н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, уз<strong>к</strong>ие, без шип<strong>о</strong><strong>в</strong>;<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong> умеренн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й<br />

гребн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> длинн<strong>а</strong>я,<br />

<strong>к</strong>лин<strong>о</strong><strong>в</strong>идн<strong>а</strong>я или <strong>с</strong><strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; н<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых тергит<strong>а</strong>х мез<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>в</strong>д<strong>о</strong>ль <strong>с</strong>рединн<strong>о</strong>й линии р<strong>а</strong>з<strong>в</strong>иты п<strong>а</strong>рные цилиндриче<strong>с</strong><strong>к</strong>ие<br />

не<strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong>ие труб<strong>о</strong>ч<strong>к</strong>и или <strong>о</strong><strong>к</strong>руглые<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и; тель<strong>с</strong><strong>о</strong>н <strong>к</strong>лин<strong>о</strong><strong>в</strong>идный (ри<strong>с</strong>. 1198,<br />

1199; т<strong>а</strong>бл. XVIII, фиг. 5). Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. В. <strong>с</strong>илур<br />

С. Амери<strong>к</strong>и; н. де<strong>в</strong><strong>о</strong>н С. Сибири (юг<strong>о</strong>-з<strong>а</strong>п<strong>а</strong>дный<br />

Т<strong>а</strong>ймыр).<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, Onychopterella Stormer, 1951<br />

(<strong>в</strong>. <strong>с</strong>илур С. Амери<strong>к</strong>и).<br />

СЕМЕЙСТВО DOLICHOPTERIDAE<br />

KJELLESVIG-WAERING ET ST0RMER, 1952<br />

Длин<strong>а</strong> ф<strong>о</strong>рм <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 <strong>с</strong>м; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> нея<strong>с</strong>н<strong>о</strong><br />

<strong>о</strong>тгр<strong>а</strong>ниченными шир<strong>о</strong><strong>к</strong><strong>о</strong>й мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и<br />

уз<strong>к</strong><strong>о</strong>й мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; тель<strong>с</strong><strong>о</strong>н <strong>к</strong>лин<strong>о</strong><strong>в</strong>идный; <strong>с</strong>л<strong>о</strong>жные<br />

гл<strong>а</strong>з<strong>а</strong> <strong>к</strong>рупные, <strong>в</strong>ыпу<strong>к</strong>лые, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

близ переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ленные;<br />

пер<strong>в</strong>ые три п<strong>а</strong>ры х<strong>о</strong>дильных н<strong>о</strong>г (<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

II—IV <strong>с</strong>егмент<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы) <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

т<strong>о</strong>л<strong>с</strong>тые и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> <strong>к</strong>рупными шип<strong>а</strong>ми;<br />

д<strong>в</strong>е п<strong>о</strong><strong>с</strong>ледние п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

б<strong>о</strong>лее длинные, без шип<strong>о</strong><strong>в</strong>, н<strong>о</strong> <strong>с</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце; з<strong>а</strong>дняя п<strong>а</strong>р<strong>а</strong> не<strong>с</strong>ет р<strong>а</strong><strong>с</strong>ширенные греб<strong>к</strong>и,<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нные у<strong>в</strong>еличенными тремя п<strong>о</strong><strong>с</strong>ледними<br />

члени<strong>к</strong><strong>а</strong>ми; эти гребные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ены<br />

ин<strong>а</strong>че, чем у других <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong><br />

уз<strong>к</strong><strong>а</strong>я, <strong>о</strong>переди <strong>в</strong><strong>о</strong>гнут<strong>а</strong>я; генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> длинные; н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> м<strong>о</strong>рщини<strong>с</strong>тый<br />

или <strong>с</strong> пузыре<strong>в</strong>идными буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми и

Ри<strong>с</strong>. 1197. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridae.<br />

Eurypterus fischeri Eichwald; А —<strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); Х0,5, <strong>с</strong>р. <strong>с</strong>илур, Э<strong>с</strong>т<strong>о</strong>ния (St0rmer, 1955)<br />

п<strong>о</strong>лу<strong>к</strong>руглыми «чешуй<strong>к</strong><strong>а</strong>ми». Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> Dolichopterus<br />

Hall, 1859 (ри<strong>с</strong>. 1200А, Б); <strong>с</strong>илур<br />

С. Амери<strong>к</strong>и; Strobilopterus Ruedemann, 1935<br />

(ри<strong>с</strong>. 1201А, Б), н. де<strong>в</strong><strong>о</strong>н С. Амери<strong>к</strong>и.<br />

НАДСЕМЕЙСТВО<br />

PTERYGOTIDEA CLARKE ET RUEDEMANN,<br />

1912<br />

[nom. transl. Novojilov, hie<br />

(ex Pterygotidae Clarke et Ruedemann, 1912)]<br />

Хелицеры <strong>о</strong>чень <strong>к</strong>рупные, зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее <strong>в</strong><strong>с</strong>ей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми<br />

<strong>к</strong>лешнями, н<strong>а</strong> <strong>о</strong>беих <strong>в</strong>ет<strong>в</strong>ях <strong>к</strong><strong>о</strong>т<strong>о</strong>рых имеют<strong>с</strong>я<br />

длинные уз<strong>к</strong>ие или шир<strong>о</strong><strong>к</strong>ие зубцы; п<strong>о</strong><strong>с</strong>ледняя<br />

п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> <strong>в</strong>иде гребных<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных н<strong>о</strong>г, <strong>с</strong>тр<strong>о</strong>ение <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong>х<strong>о</strong>дн<strong>о</strong> <strong>с</strong><strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ением их у <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Eurypteridae и Hugh-<br />

411<br />

milleriidae; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>к</strong>рупные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>к</strong>р<strong>а</strong>ях пр<strong>о</strong><strong>с</strong><strong>о</strong>мы;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> нея<strong>с</strong>н<strong>о</strong> р<strong>а</strong>згр<strong>а</strong>ничен<strong>а</strong> н<strong>а</strong> мез<strong>о</strong>и<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>му. Р<strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> — де<strong>в</strong><strong>о</strong>н. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО PTERYGOTIDAE CLARKE<br />

ET RUEDEMANN, 1912<br />

Ф<strong>о</strong>рмы <strong>о</strong>т неб<strong>о</strong>льших д<strong>о</strong> <strong>о</strong>чень <strong>к</strong>рупных<br />

(<strong>к</strong>рупные д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют 80—200 <strong>с</strong>м длины); <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

б<strong>о</strong>лее или менее <strong>в</strong>еретен<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я, без<br />

рез<strong>к</strong><strong>о</strong>й гр<strong>а</strong>ницы между мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й;<br />

тель<strong>с</strong><strong>о</strong>н <strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>к</strong>руглых или <strong>о</strong><strong>в</strong><strong>а</strong>льных пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>,<br />

<strong>с</strong> <strong>к</strong>илем н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти;<br />

х<strong>о</strong>дильные н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, из цилиндриче<strong>с</strong><strong>к</strong>их<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> без шип<strong>о</strong><strong>в</strong> или шп<strong>о</strong>р; генит<strong>а</strong>льный<br />

прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, груше<strong>в</strong>идный у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, уз-<br />

http://jurassic.ru/

Ри<strong>с</strong>. 1198—1199. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridae<br />

1198. Tylopterella boyli (Whiteaves); XO, 17, <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong><br />

(St0rmer, 1955). 1199. Tylopterella mennrl (Novojilov); x4,2,<br />

н. де<strong>в</strong><strong>о</strong>н, С. CH6Hpb^(Novjilov, 1958)<br />

<strong>к</strong>ий и туп<strong>о</strong><strong>к</strong><strong>о</strong>нечный у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>; н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><br />

из чешуе<strong>в</strong>идн<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптуры. Один р<strong>о</strong>д. ?<br />

Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — де<strong>в</strong><strong>о</strong>н.<br />

Pterygotus A g a s s i z, 1844. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. anglicus Agassiz, 1844; н. де<strong>в</strong><strong>о</strong>н,<br />

Ш<strong>о</strong>тл<strong>а</strong>ндия. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>лу<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, б<strong>о</strong>лее или<br />

менее тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>а</strong>я или б<strong>о</strong>лее или менее<br />

прям<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я <strong>с</strong><strong>о</strong> <strong>с</strong>гл<strong>а</strong>женными передними<br />

угл<strong>а</strong>ми, н<strong>а</strong> ме<strong>с</strong>те <strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>о</strong>чень<br />

<strong>к</strong>рупные <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong>; дублюр<strong>а</strong> <strong>с</strong> эпи<strong>с</strong>т<strong>о</strong>м<strong>о</strong>й;<br />

мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> б<strong>о</strong>лее или менее <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я д<strong>о</strong> <strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>й<br />

(ри<strong>с</strong>. 1202—1204; т<strong>а</strong>бл. XVIII, фиг.<br />

6—7). О<strong>к</strong><strong>о</strong>л<strong>о</strong> 30 <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Силур Э<strong>с</strong>т<strong>о</strong>нии, Ш<strong>о</strong>тл<strong>а</strong>ндии,<br />

С. Амери<strong>к</strong>и, ? А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лии; ? н. де<strong>в</strong><strong>о</strong>н<br />

(д<strong>а</strong>унт<strong>о</strong>н) Ш<strong>о</strong>тл<strong>а</strong>ндии и Н<strong>о</strong>р<strong>в</strong>егии; н. де<strong>в</strong><strong>о</strong>н<br />

С. Сибири (юг<strong>о</strong>-з<strong>а</strong>п<strong>а</strong>дный Т<strong>а</strong>ймыр), Герм<strong>а</strong>нии,<br />

С. Амери<strong>к</strong>и; <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н К<strong>а</strong>н<strong>а</strong>ды; <strong>в</strong>. де<strong>в</strong><strong>о</strong>н<br />

Ш<strong>о</strong>тл<strong>а</strong>ндии, С. Амери<strong>к</strong>и; де<strong>в</strong><strong>о</strong>н Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

412<br />

http://jurassic.ru/<br />

НАДСЕМЕЙСТВО<br />

MIXOPTERIDEA CASTER<br />

ET KJELLESVIG-WAERING, 1955<br />

[nom. transl. Novojilov, hie (ex Mixopteridae Caster e;<br />

Kjellesvig-Waering, 1955)]<br />

П<strong>о</strong>х<strong>о</strong>жие н<strong>а</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> Eurypterida, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му^<br />

п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>й мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, не<strong>с</strong>ущей из<strong>о</strong>гнутый<br />

<strong>к</strong><strong>о</strong>гте<strong>в</strong>идный или прям<strong>о</strong>й тель<strong>с</strong><strong>о</strong>н; хелицеры<br />

<strong>к</strong>лешне<strong>в</strong>идные, <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

чет<strong>к</strong><strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> р<strong>а</strong><strong>с</strong>ширенную<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>му и уз<strong>к</strong>ую цилиндриче<strong>с</strong><strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Эт<strong>а</strong> групп<strong>а</strong> э<strong>в</strong>риптерид х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я яр<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>женным при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блением для <strong>а</strong><strong>к</strong>ти<strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

хищниче<strong>с</strong>т<strong>в</strong><strong>а</strong>: <strong>к</strong><strong>о</strong>лющим <strong>о</strong>рг<strong>а</strong>н<strong>о</strong>м, тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м и<br />

х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельными <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями. Фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>в</strong>язи этих ф<strong>о</strong>рм <strong>с</strong> другими э<strong>в</strong>риптерид<strong>а</strong>ми<br />

еще не я<strong>с</strong>ны: н<strong>а</strong>личие пл<strong>а</strong><strong>в</strong><strong>а</strong>тельных н<strong>о</strong>г <strong>с</strong>ближ<strong>а</strong>ет<br />

их <strong>с</strong> Eurypteridea, примити<strong>в</strong>ные ф<strong>о</strong>рмы<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, были близ<strong>к</strong>и <strong>к</strong> пред<strong>к</strong><strong>а</strong>м<br />

Mixopteridea. Третья п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

Megalograptidae т<strong>а</strong><strong>к</strong> же <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и у Mixopteridae. Длинные п<strong>а</strong>рные иглы<br />

н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льных четырех (у Mixopterus) или пяти<br />

(у Megalograptus) члени<strong>к</strong><strong>а</strong>х н<strong>о</strong>г этих ф<strong>о</strong>рм<br />

<strong>о</strong>чень <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зны и, п<strong>о</strong> <strong>в</strong><strong>с</strong>ей <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти, <strong>в</strong>ып<strong>о</strong>лняли<br />

фун<strong>к</strong>цию яд<strong>о</strong><strong>в</strong>итых, <strong>к</strong><strong>о</strong>лющих и х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельных<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. Дифференци<strong>а</strong>ция <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

и <strong>о</strong>бщ<strong>а</strong>я ф<strong>о</strong>рм<strong>а</strong> тел<strong>а</strong> у <strong>о</strong>б<strong>о</strong>их н<strong>а</strong>з<strong>в</strong><strong>а</strong>нных<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>о</strong>бщи. Б<strong>о</strong>лее дре<strong>в</strong>ние Megalograptidae<br />

<strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т Mixopteridae т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ф<strong>о</strong>рм<strong>о</strong>й уз<strong>к</strong><strong>о</strong>г<strong>о</strong>, прям<strong>о</strong>г<strong>о</strong>, игл<strong>о</strong><strong>в</strong>идн<strong>о</strong>г<strong>о</strong> тель<strong>с</strong><strong>о</strong>н<strong>а</strong>.<br />

С друг<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Carcinosomatidae, имеющие<br />

т<strong>а</strong><strong>к</strong><strong>о</strong>й же тель<strong>с</strong><strong>о</strong>н, <strong>к</strong><strong>а</strong><strong>к</strong> Mixopteridae, <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т них (т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>о</strong>т Megalograptidae!)<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием у<strong>в</strong>еличенных х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей и б<strong>о</strong>лее р<strong>а</strong>з<strong>в</strong>итыми пл<strong>а</strong><strong>в</strong><strong>а</strong>тельными<br />

н<strong>о</strong>г<strong>а</strong>ми. Б<strong>о</strong>льш<strong>о</strong>е <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> третьей п<strong>а</strong>ры<br />

(р<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й — педип<strong>а</strong>льп) <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы у Megalograptidae и Mixopteridae,<br />

<strong>о</strong>че<strong>в</strong>идн<strong>о</strong>, у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет н<strong>а</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

этих <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

Вер<strong>о</strong>ятн<strong>о</strong>, были д<strong>о</strong>нными<br />

<strong>о</strong>бит<strong>а</strong>телями <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>, где <strong>в</strong>ели <strong>о</strong>бр<strong>а</strong>з жизни<br />

п<strong>о</strong>д<strong>с</strong>терег<strong>а</strong>ющих <strong>с</strong><strong>в</strong><strong>о</strong>ю д<strong>о</strong>бычу хищни<strong>к</strong><strong>о</strong><strong>в</strong>. Х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>, <strong>с</strong>лужили <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти<br />

л<strong>о</strong><strong>в</strong>чими <strong>с</strong>етями, имитирующими з<strong>а</strong>р<strong>о</strong><strong>с</strong>ли<br />

р<strong>а</strong><strong>с</strong>тений. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н).<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Carcinosomatidae, Mixopteridae, Megalograptidae.<br />

СЕМЕЙСТВО CARCINOSOMATIDAE ST0RMER, 1934<br />

Неб<strong>о</strong>льшие д<strong>о</strong> <strong>к</strong>рупных ф<strong>о</strong>рмы (4—60 <strong>с</strong>м<br />

длины); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>а</strong>я, <strong>с</strong>уж<strong>а</strong>ющ<strong>а</strong>я<strong>с</strong>я<br />

<strong>к</strong>переди; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие, <strong>о</strong><strong>в</strong><strong>а</strong>ль-

Ри<strong>с</strong>. 1200—1201. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Dolichopteridae<br />

1200. Dolichopterus macrocheirus Hall; А— <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу Б — пл<strong>а</strong><strong>в</strong><strong>а</strong>тельн<strong>а</strong>я н<strong>о</strong>г<strong>а</strong>, Х0.75; н. де<strong>в</strong><strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (St0r-<br />

(ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); х0,3, <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>. 1201. Strobilop- mer, 1955)<br />

ierus princetoni Ruedemann; A — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>о</strong>дним тергит<strong>о</strong>м;<br />

ные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong> 'передней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> рез<strong>к</strong><strong>о</strong> р<strong>а</strong>згр<strong>а</strong>ничен<strong>а</strong> н<strong>а</strong> <strong>о</strong>чень<br />

шир<strong>о</strong><strong>к</strong>ую, <strong>о</strong><strong>к</strong>руглую, <strong>в</strong>ыпу<strong>к</strong>лую мез<strong>о</strong><strong>с</strong><strong>о</strong>му и<br />

уз<strong>к</strong>ую, цилиндриче<strong>с</strong><strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му; тель<strong>с</strong><strong>о</strong>н <strong>в</strong><br />

<strong>в</strong>иде из<strong>о</strong>гнут<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>гте<strong>в</strong>идн<strong>о</strong>г<strong>о</strong> длинн<strong>о</strong>г<strong>о</strong> шип<strong>а</strong>;<br />

дублюр<strong>а</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й уз<strong>к</strong><strong>о</strong>й эпи<strong>с</strong>т<strong>о</strong>м<strong>о</strong>й; хелицеры<br />

т<strong>о</strong>н<strong>к</strong>ие и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> <strong>к</strong>реп<strong>к</strong>ими шип<strong>а</strong>ми,<br />

т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong>ледующие три л<strong>а</strong>ры х<strong>о</strong>дильных<br />

н<strong>о</strong>г; <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и <strong>к</strong>реп<strong>к</strong>ие педип<strong>а</strong>льпы немн<strong>о</strong>гим<br />

длиннее хелицер; пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные н<strong>о</strong>ги м<strong>о</strong>щные;<br />

мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>а</strong>я <strong>с</strong><strong>о</strong> <strong>с</strong>гл<strong>а</strong>женными<br />

угл<strong>а</strong>ми, уз<strong>к</strong><strong>а</strong>я <strong>с</strong>з<strong>а</strong>ди; генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и шир<strong>о</strong><strong>к</strong>ий (? у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>), длинный <strong>с</strong><br />

ди<strong>с</strong>т<strong>а</strong>льными шип<strong>а</strong>ми (? у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>); н<strong>а</strong>ружный<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>с</strong> р<strong>а</strong>зреженными неб<strong>о</strong>льшими буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми<br />

или <strong>с</strong> язы<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зными «чешуй<strong>к</strong><strong>а</strong>ми-». Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>: Carcinosoma Claypole, 1890, <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> —<br />

413<br />

<strong>с</strong>илур С. Амери<strong>к</strong>и (ри<strong>с</strong>. 1205) и Rhinocarcinosotna<br />

Novojilov, gen. nov., <strong>с</strong>илур С. Амери<strong>к</strong>и.<br />

Rhinocarcinosoma Novojilov, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Carcinosoma vaningeni Clark et Ruedemann,<br />

1912; <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>. Шлем<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми<br />

<strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти, <strong>с</strong> <strong>в</strong><strong>о</strong>гнутыми <strong>в</strong> з<strong>а</strong>дней<br />

ч<strong>а</strong><strong>с</strong>ти и <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им тупым <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong>м <strong>в</strong>переди;<br />

неб<strong>о</strong>льшие <strong>о</strong><strong>в</strong><strong>а</strong>льные, <strong>с</strong>уженные <strong>к</strong>переди<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>мещ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>;<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> у<strong>с</strong>еченн<strong>о</strong>-<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я <strong>с</strong> б<strong>о</strong>лее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им<br />

передним <strong>с</strong>егмент<strong>о</strong>м; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>а</strong>я;<br />

н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> буг<strong>о</strong>рч<strong>а</strong>тый. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти,<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, тель<strong>с</strong><strong>о</strong>н и генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и<br />

неиз<strong>в</strong>е<strong>с</strong>тны (ри<strong>с</strong>. 1206). Один <strong>в</strong>ид из <strong>с</strong>илур<strong>а</strong><br />

С. Амери<strong>к</strong>и.<br />

http://jurassic.ru/

1202А<br />

JL_<br />

1202Б<br />

1202Д<br />

1202В 1202 г<br />

1202. Pterygotus 'Pterygotus) rhenaniae Jaekel; A — <strong>с</strong><strong>в</strong>ерху,<br />

Б — п<strong>о</strong>перечн<strong>о</strong>е <strong>с</strong>ечение тель<strong>с</strong><strong>о</strong>н<strong>а</strong>, Х0,14; В — генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong>, Г — т<strong>о</strong> же у <strong>с</strong><strong>а</strong>м<strong>к</strong>и, Х0,2; н. де<strong>в</strong><strong>о</strong>н, Герм<strong>а</strong>ния;<br />

Д — мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> Р.(Р.) anglicus Agassiz; Х0,3, н. де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия.<br />

1203. Pterygotus (Erettopterus) osiliensis Schmidt; A — <strong>с</strong><strong>в</strong>ер­<br />

СЕМЕЙСТВО MIXOPTERIDAE CASTER<br />

ET KJELLESVIG-WAERING, 1955<br />

Длин<strong>а</strong> ф<strong>о</strong>рм 20—70 <strong>с</strong>м; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>чти<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>а</strong>я <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей <strong>с</strong>ередин<strong>о</strong>й переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие, п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены у переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я;<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>а</strong>я, <strong>с</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, <strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong>ими<br />

<strong>с</strong>егмент<strong>а</strong>ми; тель<strong>с</strong><strong>о</strong>н <strong>к</strong><strong>о</strong>гте<strong>о</strong>бр<strong>а</strong>зный; педип<strong>а</strong>льпы<br />

и третья п<strong>а</strong>р<strong>а</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

удлиненные, <strong>о</strong>бе <strong>с</strong> длинными п<strong>а</strong>рными шип<strong>а</strong>ми;<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>в</strong>ели<strong>к</strong>и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти третьей п<strong>а</strong>ры;<br />

чет<strong>в</strong>ерт<strong>а</strong>я и пят<strong>а</strong>я п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные н<strong>о</strong>ги х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, <strong>с</strong> длинным<br />

шир<strong>о</strong><strong>к</strong>им <strong>с</strong>едьмым и менее р<strong>а</strong>з<strong>в</strong>итым<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьмым члени<strong>к</strong><strong>а</strong>ми; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> <strong>с</strong>ердце<strong>в</strong>идн<strong>а</strong>я,<br />

<strong>с</strong>уженн<strong>а</strong>я <strong>с</strong>з<strong>а</strong>ди; генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

длинные; н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>с</strong> р<strong>а</strong>зреженными<br />

буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми или п<strong>о</strong>лу<strong>к</strong>руглыми «чешуй<strong>к</strong><strong>а</strong>ми».<br />

Один р<strong>о</strong>д — Mixopterus Ruedemann, 1921. Четыре<br />

<strong>в</strong>ид<strong>а</strong> (ри<strong>с</strong>. 1207). В. <strong>с</strong>илур (<strong>в</strong>енл<strong>о</strong><strong>к</strong><strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>) Ш<strong>о</strong>тл<strong>а</strong>ндии; ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) С<strong>к</strong><strong>а</strong>ндин<strong>а</strong><strong>в</strong>ии.<br />

1203Д<br />

1203 Б<br />

1203Е<br />

1203 Г<br />

1203В<br />

Ри<strong>с</strong>. 1202—1203. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pterygotidae (St/armer, 1955)<br />

414<br />

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1203А<br />

ху, x0,1; Б — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льн<strong>а</strong>я дублюр<strong>а</strong> <strong>с</strong> эпи<strong>с</strong>т<strong>о</strong>м<strong>о</strong>й, х0,3; В —<br />

мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong>, х 0,2; Г — генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> (?) <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х 0,3;<br />

Д — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, Х0,7; <strong>с</strong>илур, Э<strong>с</strong>т<strong>о</strong>ния; £ — P. (P.) bilobus (Salter);<br />

х<strong>о</strong>дильн<strong>а</strong>я н<strong>о</strong>г<strong>а</strong>, Х1,0, <strong>с</strong>илур, Ш<strong>о</strong>тл<strong>а</strong>ндия<br />

СЕМЕЙСТВО MEGALOGRAPTIDAE CASTER<br />

ET KJELLESVIG-WAERING, 1955<br />

Ф<strong>о</strong>рмы 60—70 <strong>с</strong>м длины; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> шлем<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>черт<strong>а</strong>ния, р<strong>а</strong><strong>с</strong>ширенн<strong>а</strong>я <strong>с</strong>переди;<br />

ме<strong>с</strong>т<strong>о</strong>п<strong>о</strong>л<strong>о</strong>жение <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е;<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>а</strong>я, <strong>с</strong> <strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льными<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми, п<strong>о</strong><strong>с</strong>ледний ее <strong>с</strong>егмент<br />

без пле<strong>в</strong>р; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, <strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong>ими<br />

<strong>с</strong>егмент<strong>а</strong>ми, <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими пл<strong>о</strong><strong>с</strong><strong>к</strong>ими <strong>к</strong><strong>о</strong>гте<strong>о</strong>бр<strong>а</strong>зными<br />

прид<strong>а</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>егменте; тель<strong>с</strong><strong>о</strong>н<br />

т<strong>о</strong>н<strong>к</strong>ий, <strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>с</strong>тр<strong>о</strong>й иглы; дублюр<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й, шир<strong>о</strong><strong>к</strong><strong>о</strong>й эпи<strong>с</strong>т<strong>о</strong>м<strong>о</strong>й; <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я<br />

и третья п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные,<br />

<strong>с</strong> длинными п<strong>а</strong>рными игл<strong>а</strong>ми, причем<br />

третья п<strong>а</strong>р<strong>а</strong> б<strong>о</strong>лее м<strong>о</strong>щн<strong>а</strong>я, р<strong>а</strong>з<strong>а</strong> <strong>в</strong> 3 б<strong>о</strong>льше<br />

педип<strong>а</strong>льп (II п<strong>а</strong>ры); чет<strong>в</strong>ерт<strong>а</strong>я и пят<strong>а</strong>я п<strong>а</strong>ры<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> т<strong>о</strong>н<strong>к</strong>ие и <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щими<br />

шип<strong>а</strong>ми; пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные н<strong>о</strong>ги <strong>с</strong> <strong>о</strong>чень <strong>к</strong>рупными<br />

чет<strong>в</strong>ертым и <strong>с</strong>едьмым члени<strong>к</strong><strong>а</strong>ми; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong><br />

<strong>о</strong>т <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й д<strong>о</strong> <strong>с</strong>ердце<strong>в</strong>идн<strong>о</strong>й. Н<strong>а</strong>ружный<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>с</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льными или треуг<strong>о</strong>льными «чешуй<strong>к</strong><strong>а</strong>ми»<br />

и пр<strong>о</strong>д<strong>о</strong>льными ребрыш<strong>к</strong><strong>а</strong>ми н<strong>а</strong> з<strong>а</strong>дней

Pterygotus (Acuttramus) buffatoensis (Pohlman) A — <strong>с</strong><strong>в</strong>ерху;<br />

£—<strong>с</strong>низу, x0,045; В — мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, X 0,3; Г — пл<strong>а</strong><strong>в</strong><strong>а</strong>тельн<strong>а</strong>я<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ть, Х0,3; Д — генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> (?) <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х0,17;<br />

<strong>с</strong>илур, С.Амери<strong>к</strong><strong>а</strong> (St0rmer, 1955). Е.Ж—Р. (Acutiramus)<br />

waylandsmithi Kjellesvig-Waering et Caster: п<strong>о</strong>д<strong>в</strong>ижные <strong>в</strong>ет<strong>в</strong>и<br />

ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Генит<strong>а</strong>льные<br />

прид<strong>а</strong>т<strong>к</strong>и <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие дубин<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, у<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> длинные, уз<strong>к</strong>ие. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong><strong>а</strong><br />

С. Амери<strong>к</strong>и: Megatograptus Miller, 1874<br />

(ри<strong>с</strong>. 1208) и Echinognatus Walcott, 1882.<br />

Ри<strong>с</strong>. 1204. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pterygotidae.<br />

<strong>к</strong>лешни <strong>с</strong> др<strong>о</strong>бящими зубц<strong>а</strong>ми (1—7). 3— P. (Acutiramus) floveri<br />

Kjellesvig-Waering et Caster; 3 — <strong>к</strong>лешня; И — п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я<br />

<strong>в</strong>ет<strong>в</strong>ь <strong>к</strong>лешни; х2, <strong>с</strong>илур (<strong>с</strong>л<strong>а</strong>нцы <strong>в</strong>ерн<strong>о</strong>й), С. Амери<strong>к</strong><strong>а</strong> (Kjellesvig-Waering<br />

et Caster, 1955)<br />

ПОДОТРЯД STYLONURINA<br />

[nom. transl. Novojilov, hie (ex Stylonuridae Diener,<br />

1924; Stylonuracea Stormer, 1951)]<br />

В<strong>с</strong>е <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong>чин<strong>а</strong>я <strong>с</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й<br />

п<strong>а</strong>ры <strong>в</strong> <strong>в</strong>иде длинных х<strong>о</strong>дильных н<strong>о</strong>г. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong>—<br />

н. пермь. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Stylonuridea,<br />

Mycteropidea.<br />

415<br />

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1205 В<br />

Ри<strong>с</strong>. 1205—1206. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Carcinosomatidae<br />

1205. Carctnosoma seorplonls (Grote et Pitt); <strong>с</strong><strong>а</strong>мец (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>- 1206. Rhlnocarclnosoma vanlngent (Clark, et Ruederaann); пр<strong>о</strong>ция);<br />

A - <strong>с</strong><strong>в</strong>ерху, x 0,2, Б - <strong>с</strong>низу, X0,2; В - <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> <strong>с</strong><strong>о</strong>м<strong>а</strong> и пять тергит<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, х 2,0 <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong><br />

прид<strong>а</strong>т<strong>к</strong><strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы; x2,0 f <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong> (Stermer, 1955).

Ри<strong>с</strong>. 1207. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mixopteridae.<br />

Mixopterus kiaert St0rmer; A — <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция), Х0,2,? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н), Н<strong>о</strong>р<strong>в</strong>егия (Starrner, 1955)<br />

27 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

417<br />

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Ри<strong>с</strong>. 1208. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Megalograptidae. Megatograptus 0hioensis<br />

Caster et Kjellesvig-Waering; <strong>с</strong><strong>в</strong>ерху (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру <strong>к</strong>ция)<br />

X 0,2, <strong>в</strong>.рд<strong>о</strong><strong>в</strong>и<strong>к</strong>, С. Амери<strong>к</strong><strong>а</strong> (St0rmer, 1955)<br />

н<strong>о</strong><strong>с</strong>ти длинные, х<strong>о</strong>дильные, <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ющие <strong>в</strong><br />

<strong>в</strong>еличине <strong>с</strong>переди н<strong>а</strong>з<strong>а</strong>д, <strong>с</strong> шип<strong>а</strong>ми или без<br />

шип<strong>о</strong><strong>в</strong>; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> <strong>в</strong> <strong>в</strong>иде пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>т<strong>о</strong>й уз<strong>к</strong><strong>о</strong>й<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и; генит<strong>а</strong>льные прид<strong>а</strong>т<strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

418<br />

http://jurassic.ru/<br />

шир<strong>о</strong><strong>к</strong>ие у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> и длинные у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>; н<strong>а</strong>ружный<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>с</strong> «чешуй<strong>к</strong><strong>а</strong>ми», перех<strong>о</strong>дящими <strong>в</strong><br />

буг<strong>о</strong>р<strong>к</strong>и или шипы. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — н. пермь. П<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Stylonurinae, Marsupipterinae.<br />

ПОДСЕМЕЙСТВО STYLONURINAE DIENER, 1924<br />

[nom. transl. Novojilov, 1958<br />

(ex Stylonuridae Diner, 1924)]<br />

Тель<strong>с</strong><strong>о</strong>н без <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

и без п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й ег<strong>о</strong><br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не. Орд<strong>о</strong><strong>в</strong>и<strong>к</strong> — н. пермь. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Stylonurus Page, 1856. Тип р<strong>о</strong>д<strong>а</strong> — S. р<strong>о</strong>wrieri<br />

Page, 1856; де<strong>в</strong><strong>о</strong>н; Ш<strong>о</strong>тл<strong>а</strong>ндия. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

п<strong>о</strong>лу<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я или <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, <strong>с</strong>рез<strong>а</strong>нн<strong>а</strong>я <strong>с</strong>з<strong>а</strong>ди;<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, близ б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> передней ее п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине; гл<strong>а</strong>з<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены между <strong>с</strong>л<strong>о</strong>жными гл<strong>а</strong>з<strong>а</strong>ми;<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти длинные, <strong>с</strong> уз<strong>к</strong>ими, пр<strong>о</strong>д<strong>о</strong>лг<strong>о</strong><strong>в</strong><strong>а</strong>тыми<br />

члени<strong>к</strong><strong>а</strong>ми; пер<strong>в</strong>ые три п<strong>а</strong>ры н<strong>о</strong>г <strong>с</strong> шип<strong>а</strong>ми,<br />

п<strong>о</strong><strong>с</strong>ледняя п<strong>а</strong>р<strong>а</strong> н<strong>о</strong>г р<strong>а</strong><strong>в</strong>н<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме или<br />

длиннее ее; генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong><br />

м<strong>о</strong>щный, длинный, <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями у<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния и р<strong>а</strong>зд<strong>в</strong><strong>о</strong>енный н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м <strong>к</strong><strong>о</strong>нце,<br />

п<strong>о</strong> <strong>о</strong>черт<strong>а</strong>нию н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет д<strong>в</strong>ухл<strong>о</strong>п<strong>а</strong><strong>с</strong>тный<br />

я<strong>к</strong><strong>о</strong>рь; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>згр<strong>а</strong>ниченными<br />

мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; тель<strong>с</strong><strong>о</strong>н <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>в</strong>идный;<br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть м<strong>о</strong>рщини<strong>с</strong>т<strong>а</strong>я (т<strong>а</strong>бл. XIX,<br />

фиг. 1—4). Д<strong>о</strong> де<strong>с</strong>яти <strong>в</strong>ид<strong>о</strong><strong>в</strong>. Силур Англии,<br />

Ш<strong>о</strong>тл<strong>а</strong>ндии; ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) С<strong>к</strong><strong>а</strong>ндин<strong>а</strong><strong>в</strong>ии;<br />

де<strong>в</strong><strong>о</strong>н Ш<strong>о</strong>тл<strong>а</strong>ндии; ? <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н Ю. Сибири<br />

и С. Амери<strong>к</strong>и.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>: Drepanopterus Laurie, 1892<br />

(ри<strong>с</strong>. 1210), <strong>в</strong>. <strong>с</strong>илур Ш<strong>о</strong>тл<strong>а</strong>ндии и С. Амери<strong>к</strong>и<br />

(Инди<strong>а</strong>н<strong>а</strong>); <strong>в</strong>. де<strong>в</strong><strong>о</strong>н Ирл<strong>а</strong>ндии; Brachyopterus<br />

Stormer, 1951 (ри<strong>с</strong>. 1211), <strong>с</strong>р. <strong>о</strong>рд<strong>о</strong><strong>в</strong>и<strong>к</strong> Англии,<br />

? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) Н<strong>о</strong>р<strong>в</strong>егии; Ctenopterus<br />

Clarke et Ruedemann, 1912 (ри<strong>с</strong>. 1212),<br />

<strong>с</strong>илур и <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н США (шт<strong>а</strong>т Нью-Й<strong>о</strong>р<strong>к</strong>);<br />

Tarsopterella Stormer, 1951, н. и <strong>в</strong>. де<strong>в</strong><strong>о</strong>н Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

ПОДСЕМЕЙСТВО MARSUPIPTERINAE NOVOJILOV,<br />

1958<br />

Тель<strong>с</strong><strong>о</strong>н <strong>с</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

и <strong>с</strong> дупл<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>тью н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не. ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н) — <strong>в</strong>.<br />

де<strong>в</strong><strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Potystomurum Novojilov, 1958. Тип р<strong>о</strong>д<strong>а</strong>—<br />

P. stermeri Novojilov, 1958; <strong>в</strong>. де<strong>в</strong><strong>о</strong>н, В<strong>о</strong>р<strong>о</strong>неж<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. Тель<strong>с</strong><strong>о</strong>н г<strong>в</strong><strong>о</strong>зде<strong>о</strong>бр<strong>а</strong>зный н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>с</strong> <strong>к</strong>илем и п<strong>а</strong>рными <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями.<br />

Брюшн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong> тель<strong>с</strong><strong>о</strong>н<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тн<strong>а</strong>;<br />

предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong>личие <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> у р<strong>о</strong>д<strong>а</strong> Marsupipterus. П<strong>о</strong>л<strong>о</strong><strong>с</strong>ть тель<strong>с</strong><strong>о</strong>н<strong>а</strong>,<br />

п<strong>о</strong> <strong>в</strong><strong>с</strong>ей <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong><strong>с</strong>ти, з<strong>а</strong><strong>к</strong>люч<strong>а</strong>л<strong>а</strong> яд<strong>о</strong><strong>в</strong>итую железу,<br />

<strong>с</strong>е<strong>к</strong>рет <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>в</strong>ыделял<strong>с</strong>я через д<strong>о</strong>р<strong>с</strong><strong>а</strong>льные<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия (ри<strong>с</strong>. 1213).

Ри<strong>с</strong>. 1209—1212. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> !<br />

1209. Hibbertopterus scouleri (Hibbert); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> д<strong>в</strong>умя тергит<strong>а</strong>ми,<br />

X 0,25, и. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия (St0r«ier, 1955). 1210.<br />

Drepanopterus abonensis Simpson (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); X0.23, <strong>в</strong>.<br />

де<strong>в</strong><strong>о</strong>н, Англия (Simpson, 1951). 1211. Brachyopterus? pentago-<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, Marsupipterus Kjellesvig-Waering,<br />

1955 (ри<strong>с</strong>. 1214), ? н. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н)<br />

Англии.<br />

СЕМЕЙСТВО HIBBERTOPTERIDAE<br />

KJELLESVIG-WAERING, 1959<br />

Крупные ф<strong>о</strong>рмы (<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 50 <strong>с</strong>м); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> <strong>о</strong><strong>к</strong>руглые или п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идные,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

между ними п<strong>о</strong>мещ<strong>а</strong>ют<strong>с</strong>я гл<strong>а</strong>з<strong>к</strong>и; <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ы<br />

з<strong>а</strong>дних <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы без зубч<strong>а</strong>тых <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>о</strong>-<strong>с</strong>ердце<strong>в</strong>идн<strong>а</strong>я,<br />

<strong>с</strong>х<strong>о</strong>дящ<strong>а</strong>я<strong>с</strong>я <strong>в</strong>переди, <strong>с</strong><strong>о</strong> <strong>с</strong>рединным<br />

ш<strong>в</strong><strong>о</strong>м <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти; <strong>о</strong>пер<strong>к</strong>улум <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> неиз<strong>в</strong>е<strong>с</strong>тен,<br />

у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им дубин<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зным<br />

прид<strong>а</strong>т<strong>к</strong><strong>о</strong>м и передней яйце<strong>в</strong>идн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин­<br />

'lonuridae и Hibbertopteridae<br />

nalis (StiBrmer); A — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, Б — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями<br />

<strong>с</strong>низу, X 0,4,? и. де<strong>в</strong><strong>о</strong>н (д<strong>а</strong>унт<strong>о</strong>н), Н<strong>о</strong>р<strong>в</strong>егия. 1212.<br />

Ctenopterus cestrotus (Clarke); <strong>с</strong><strong>в</strong>ерху, Х0,5, <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong><br />

(St0rmer, 1955) j<br />

<strong>к</strong><strong>о</strong>й. Сегменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы уз<strong>к</strong>ие; тель<strong>с</strong><strong>о</strong>н неиз<strong>в</strong>е<strong>с</strong>тен.<br />

Н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из мел<strong>к</strong>их перье<strong>в</strong>идных «чешуе<strong>к</strong>»; з<strong>а</strong>дние<br />

<strong>к</strong>р<strong>а</strong>я тергит<strong>о</strong><strong>в</strong> <strong>с</strong> т<strong>а</strong><strong>к</strong>ими же «чешуй<strong>к</strong><strong>а</strong>ми» <strong>в</strong><br />

<strong>в</strong>иде <strong>к</strong><strong>а</strong>йм,ы. В. де<strong>в</strong><strong>о</strong>н (?) — н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. "Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>. !<br />

Campylocephalus Е i <strong>с</strong> h wa 1 d, 1860 [Eidothea<br />

Scouler, 1831 (non Risso, 1826, nec Chambers,<br />

1873)]. Тип р<strong>о</strong>д<strong>а</strong> — Limulus oculatus Kutorga,<br />

1838; <strong>в</strong>. пермь, Б<strong>а</strong>ш<strong>к</strong>ирия (б<strong>а</strong><strong>с</strong><strong>с</strong>ейн p.<br />

Бел<strong>о</strong>й, Дур<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий рудни<strong>к</strong>). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я,<br />

<strong>с</strong> <strong>в</strong>ырез<strong>о</strong>м н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е и уз<strong>к</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>йм<strong>о</strong>й;<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идные, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; гл<strong>а</strong>з<strong>к</strong>и <strong>в</strong>переди<br />

<strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з; тергиты прямые, уз<strong>к</strong>ие. О<strong>с</strong>т<strong>а</strong>льные<br />

ч<strong>а</strong><strong>с</strong>ти тел<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тны (т<strong>а</strong>бл. XXI). Один<br />

<strong>в</strong>ид.<br />

419 27*<br />

http://jurassic.ru/

* ill<br />

(213<br />

I214A<br />

Ри<strong>с</strong>. 1213—1214. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Stylonuridae<br />

1213. Potystomurum starmeri Novojilov; тель<strong>с</strong><strong>о</strong>н, X12.0, <strong>в</strong>. де<strong>в</strong><strong>о</strong>н,<br />

СССР (В<strong>о</strong>р<strong>о</strong>неж<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.) (Н<strong>о</strong><strong>в</strong><strong>о</strong>жил<strong>о</strong><strong>в</strong>, 1958). 1214. М<strong>а</strong>гsuplpterus<br />

sculpturatus Kjellesvlg-Waering; тель<strong>с</strong><strong>о</strong>н; А — <strong>с</strong><strong>в</strong>ерху,<br />

Б — <strong>с</strong>низу, х0,5, <strong>в</strong>. <strong>с</strong>илур, Англия (Kjellesvig-Waerlng, 1955)<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, Hibbertopterus Kjellesvig-Waering,<br />

1959 (ри<strong>с</strong>. 1209), <strong>в</strong>. де<strong>в</strong><strong>о</strong>н (?) Ирл<strong>а</strong>ндии,<br />

н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

СЕМЕЙСТВО RHENOPTERIDAE ST0RMER, 1951<br />

Неб<strong>о</strong>льшие ф<strong>о</strong>рмы (5—10 <strong>с</strong>м длины); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

п<strong>о</strong>лу<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я, <strong>с</strong> п<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зными <strong>с</strong>л<strong>о</strong>жными<br />

гл<strong>а</strong>з<strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными у ее б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

420<br />

http://jurassic.ru/<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong>, примерн<strong>о</strong> н<strong>а</strong> <strong>с</strong>ередине; дублюр<strong>а</strong><br />

<strong>с</strong><strong>о</strong> <strong>с</strong>рединным ш<strong>в</strong><strong>о</strong>м, н<strong>о</strong> без<br />

эпи<strong>с</strong>т<strong>о</strong>мы; пер<strong>в</strong>ые д<strong>в</strong>е (или три?)<br />

п<strong>а</strong>ры н<strong>о</strong>г <strong>с</strong> шип<strong>а</strong>ми, п<strong>о</strong><strong>с</strong>ледние д<strong>в</strong>е<br />

п<strong>а</strong>ры без шип<strong>о</strong><strong>в</strong>; мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<strong>а</strong> груше<strong>в</strong>идн<strong>а</strong>я<br />

<strong>с</strong> <strong>в</strong><strong>о</strong>гнутым передним <strong>к</strong>р<strong>а</strong>ем,<br />

генит<strong>а</strong>льный прид<strong>а</strong>т<strong>о</strong><strong>к</strong> <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий,<br />

<strong>с</strong> д<strong>в</strong>умя ди<strong>с</strong>т<strong>а</strong>льными шип<strong>а</strong>ми;<br />

н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>с</strong> р<strong>а</strong>зличными<br />

буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми (т<strong>а</strong>бл. XIX, фиг. 5).<br />

Один р<strong>о</strong>д — Rhenopterus Strmer,<br />

1936 (ри<strong>с</strong>. 1215). н. де<strong>в</strong><strong>о</strong>н Герм<strong>а</strong>нии.<br />

НАДСЕМЕЙСТВО<br />

MYCTEROPIDEA СОРЕ, 1886<br />

(nom. transl. Novojilov, hie (ex Mycteropidae<br />

Cope, 1886)]<br />

Пер<strong>в</strong>ый <strong>с</strong>егмент <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>о</strong>чень <strong>к</strong>рупный, немн<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че пр<strong>о</strong><strong>с</strong><strong>о</strong>мы;<br />

п<strong>о</strong><strong>с</strong>ледующие <strong>с</strong>егменты <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> уменьш<strong>а</strong>ющие<strong>с</strong>я <strong>в</strong><br />

ширине и <strong>с</strong>х<strong>о</strong>дящие<strong>с</strong>я <strong>к</strong>лин<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>к</strong> тель<strong>с</strong><strong>о</strong>ну; <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти уз<strong>к</strong>ие, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

или длинные, <strong>к</strong><strong>а</strong><strong>к</strong> у Stylonuridae.<br />

Eurypterida <strong>с</strong> <strong>к</strong>рупным пер<strong>в</strong>ым<br />

<strong>с</strong>егмент<strong>о</strong>м <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы из<strong>в</strong>е<strong>с</strong>тны п<strong>о</strong><br />

неп<strong>о</strong>лным э<strong>к</strong>земпляр<strong>а</strong>м четырех <strong>в</strong>ид<strong>о</strong><strong>в</strong>:<br />

Mycterops ordinatus С<strong>о</strong>ре, 1886<br />

(<strong>к</strong><strong>а</strong>рб<strong>о</strong>н Пен<strong>с</strong>иль<strong>в</strong><strong>а</strong>нии), Eurypterus<br />

scabrosus Woodward, 1887 (н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Ш<strong>о</strong>тл<strong>а</strong>ндии), Glaucodes mathieui<br />

Pruvost, 1923 (<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н Бельгии),<br />

Borchgrevinkium taimyrense Novojilov,1958<br />

(н. де<strong>в</strong><strong>о</strong>н С. Сибири). Х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер<br />

т<strong>а</strong>гм<strong>а</strong>тиз<strong>а</strong>ции этих ф<strong>о</strong>рм,<br />

именн<strong>о</strong> н<strong>а</strong>личие <strong>к</strong>рупн<strong>о</strong>г<strong>о</strong> переднег<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> (или д<strong>в</strong>ух <strong>с</strong>егмент<strong>о</strong><strong>в</strong>?)<br />

1214Б<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ет их <strong>о</strong>т<br />

Stylonurina д<strong>а</strong> и <strong>в</strong><strong>о</strong><strong>о</strong>бще <strong>о</strong>т <strong>в</strong><strong>с</strong>ех других<br />

Eurypterida. Т<strong>а</strong><strong>к</strong><strong>а</strong>я м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ть з<strong>а</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ть<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> этих ф<strong>о</strong>рм <strong>с</strong> другими<br />

хелицер<strong>о</strong><strong>в</strong>ыми. От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие д<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>ерных<br />

д<strong>а</strong>нных <strong>о</strong> <strong>с</strong>тр<strong>о</strong>ении <strong>в</strong><strong>с</strong>ег<strong>о</strong><br />

тел<strong>а</strong> и ег<strong>о</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей не п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет<br />

<strong>с</strong>дел<strong>а</strong>ть <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>тельных <strong>в</strong>ы<strong>в</strong><strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>о</strong><br />

п<strong>о</strong>л<strong>о</strong>жении Mycteropidea <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме.<br />

Выделение эт<strong>о</strong>й группы <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е<br />

<strong>о</strong>тдельн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> пред<strong>в</strong><strong>а</strong>рительн<strong>о</strong>;<br />

<strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>, чт<strong>о</strong> <strong>в</strong> д<strong>а</strong>льнейшем<br />

эти ф<strong>о</strong>рмы н<strong>а</strong>д<strong>о</strong> будет п<strong>о</strong>ме-<br />

<strong>с</strong>тить <strong>в</strong><br />

т<strong>а</strong><strong>к</strong><strong>с</strong><strong>о</strong>н б<strong>о</strong>лее <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong>нг<strong>а</strong>.<br />

Н<strong>а</strong>д<strong>о</strong> <strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть, чт<strong>о</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и, п<strong>о</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым изучены<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители Eurypterida <strong>с</strong> <strong>к</strong>рупным<br />

пер<strong>в</strong>ым <strong>с</strong>егмент<strong>о</strong>м <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>чень <strong>с</strong><strong>к</strong>удны.<br />

У Mycterops ordinatus (т<strong>а</strong>бл. XXII), п<strong>о</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>му<br />

у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лены р<strong>о</strong>д и <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь

Ри<strong>с</strong>. 1215. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Rhenopteridae. Rhenopterus diensti<br />

St0rmer ; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>;<br />

Л—<strong>с</strong><strong>в</strong>ерху, Б—<strong>с</strong>низу, XI,0, н. де<strong>в</strong><strong>о</strong>н, Герм<strong>а</strong>ния (St0rmer, 1955)<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> уз<strong>к</strong><strong>а</strong>я <strong>с</strong>переди ' тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

и пер<strong>в</strong>ый тергит <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>бл<strong>о</strong>м<strong>а</strong>нные<br />

<strong>с</strong> пр<strong>а</strong><strong>в</strong><strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны (пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong> эти <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и примити<strong>в</strong>ных п<strong>о</strong>­<br />

Ри<strong>с</strong>. 1216—1218. Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mycteropidea,<br />

1216. Woodwardopterus scabrosus (Н. Woodward); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> ше<strong>с</strong>тью<br />

тергит<strong>а</strong>ми мез<strong>о</strong><strong>с</strong><strong>о</strong>мы и тремя пр<strong>а</strong><strong>в</strong>ыми н<strong>о</strong>г<strong>а</strong>ми. Х0.5, и.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия, (St0rmer, 1955). 1217. Mycterops mathieui<br />

(Pruvost); A — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> пер<strong>в</strong>ым <strong>к</strong>рупным <strong>с</strong>егмент<strong>о</strong>м мез<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

Х0.38, £ — <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме и тергите, х3,0;<br />

421<br />

з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных). С<strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я непр<strong>а</strong><strong>в</strong>ильн<strong>а</strong>я <strong>с</strong>етч<strong>а</strong>т<strong>о</strong>-ячеи<strong>с</strong>т<strong>а</strong>я<br />

<strong>с</strong><strong>к</strong>ульптур<strong>а</strong> п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

<strong>в</strong><strong>с</strong>ей пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и пер<strong>в</strong><strong>о</strong>г<strong>о</strong> тергит<strong>а</strong>. П<strong>о</strong>д<strong>о</strong>бные<br />

же <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и (пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и пер<strong>в</strong>ый тергит), н<strong>о</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> лучшей <strong>с</strong><strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ти н<strong>а</strong>йдены <strong>в</strong> <strong>в</strong>.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>не Бельгии. Они <strong>о</strong>пи<strong>с</strong><strong>а</strong>ны <strong>к</strong><strong>а</strong><strong>к</strong> Glaucodes<br />

mathieui (т<strong>а</strong>бл. XIX, фиг. 6), н<strong>о</strong>, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му,<br />

д<strong>о</strong>лжны быть причи<strong>с</strong>лены <strong>к</strong> т<strong>о</strong>му же р<strong>о</strong>ду<br />

Mycterops.<br />

П<strong>о</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м из н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Ш<strong>о</strong>тл<strong>а</strong>ндии,<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>нным пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong> <strong>к</strong><strong>а</strong><strong>к</strong> Eurypterus scabrosus,<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие р<strong>а</strong>зры<strong>в</strong><strong>о</strong><strong>в</strong> и <strong>с</strong>мещений тергит<strong>о</strong><strong>в</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я нея<strong>с</strong>ным, <strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong>рупных <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong>бр<strong>а</strong>зуют переднюю ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы —<br />

<strong>о</strong>дин или д<strong>в</strong><strong>а</strong>. О п<strong>о</strong><strong>с</strong>ледующих <strong>с</strong>егмент<strong>а</strong>х м<strong>о</strong>жн<strong>о</strong><br />

<strong>с</strong>удить т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>м четырех передних:<br />

<strong>о</strong>ни <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong> уз<strong>к</strong>ие и <strong>с</strong> пле<strong>в</strong>р<strong>а</strong>льными<br />

угл<strong>а</strong>ми. С<strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шие<strong>с</strong>я три пр<strong>а</strong><strong>в</strong>ые <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

уз<strong>к</strong>ие и длинные, <strong>к</strong><strong>а</strong><strong>к</strong> у Stylonuridea.<br />

С<strong>к</strong>ульптур<strong>а</strong> из мел<strong>к</strong>их перье<strong>в</strong>идных «чешуе<strong>к</strong>»<br />

п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет пр<strong>о</strong><strong>с</strong><strong>о</strong>му и <strong>к</strong>рупный тергит. П<strong>о</strong> <strong>к</strong>руп-<br />

1217Б<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Mycteropidae и Woodwardopteridae<br />

<strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Бельгия (Pruvost, 1923). 1218. Borchgrevinklum tatmyrense<br />

Novojilov; A — <strong>с</strong><strong>в</strong>ерху (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); Б — <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

н<strong>а</strong> <strong>с</strong><strong>к</strong><strong>о</strong>л<strong>о</strong>т<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>с</strong>лед генит<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong>,<br />

Х3,4; В — чешуе<strong>в</strong>идн<strong>а</strong>я <strong>с</strong><strong>к</strong>ульптур<strong>а</strong> н<strong>а</strong> тергит<strong>а</strong>х <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

Х10; и. де<strong>в</strong><strong>о</strong>н, С. Сибирь. (Novojilov, 1959)<br />

http://jurassic.ru/

н<strong>о</strong>му (или д<strong>в</strong>ум?) <strong>с</strong>егменту пр<strong>о</strong><strong>с</strong><strong>о</strong>мы Ст<strong>о</strong>рмер<br />

(St0rmer, 1951, 1955) причи<strong>с</strong>лил эт<strong>о</strong>т <strong>в</strong>ид <strong>к</strong><br />

р<strong>о</strong>ду Mycterops. Челле<strong>с</strong><strong>в</strong>иг-Веринг (Kjellesvig-<br />

Waering, 1959) п<strong>о</strong> эт<strong>о</strong>му же э<strong>к</strong>земпляру у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ил<br />

н<strong>о</strong><strong>в</strong>ый р<strong>о</strong>д Woodwardopterus и н<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Woodwardopteridae, п<strong>о</strong>ме<strong>с</strong>ти<strong>в</strong> ег<strong>о</strong> <strong>в</strong><br />

н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridea п<strong>о</strong> <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong>у <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей.<br />

Б<strong>о</strong>лее п<strong>о</strong>лн<strong>о</strong>е пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ление <strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гии<br />

<strong>к</strong>рупн<strong>о</strong><strong>с</strong>егментных Eurypterida д<strong>а</strong>ет п<strong>о</strong>чти п<strong>о</strong>лный<br />

э<strong>к</strong>земпляр р<strong>о</strong>д<strong>а</strong> Borchgrevinkium, х<strong>о</strong>тя и<br />

пл<strong>о</strong>х<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>ший<strong>с</strong>я (ри<strong>с</strong>. 1218). Ег<strong>о</strong> н<strong>а</strong>ружный<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>, т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> у Mycterops, <strong>с</strong>етч<strong>а</strong>т<strong>о</strong>ячеи<strong>с</strong>тый,<br />

н<strong>о</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н пр<strong>а</strong><strong>в</strong>ильными ячей<strong>к</strong><strong>а</strong>ми,<br />

пр<strong>а</strong><strong>в</strong>д<strong>а</strong>, чешуе<strong>в</strong>идными. Н<strong>а</strong> <strong>с</strong><strong>к</strong><strong>о</strong>л<strong>о</strong>т<strong>о</strong>й <strong>с</strong>редней<br />

ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>идны уз<strong>к</strong>ие <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> у Stylonuridae, н<strong>о</strong> <strong>в</strong><strong>с</strong>е <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие (ри<strong>с</strong>. 1218<br />

Б, т<strong>а</strong>бл. XIX фиг. 7).<br />

С учет<strong>о</strong>м н<strong>а</strong>личия <strong>к</strong>рупн<strong>о</strong>г<strong>о</strong> (или д<strong>в</strong>ух <strong>к</strong>рупных)<br />

<strong>с</strong>егмент<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы у <strong>в</strong><strong>с</strong>ех четырех уп<strong>о</strong>мянутых<br />

ф<strong>о</strong>рм, <strong>а</strong> т<strong>а</strong><strong>к</strong>же р<strong>а</strong>зличия <strong>в</strong> <strong>с</strong><strong>к</strong>ульптуре<br />

н<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong> <strong>о</strong>ни <strong>о</strong>бъединены зде<strong>с</strong>ь<br />

<strong>в</strong> <strong>о</strong>дн<strong>о</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mycteropidea <strong>с</strong> д<strong>в</strong>умя <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>ми:<br />

Mycteropidae и Woodwardopteridae.<br />

Н. де<strong>в</strong><strong>о</strong>н — <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н.<br />

СЕМЕЙСТВО MYCTEROPIDAE<br />

ST0RMER, 1951<br />

Ф<strong>о</strong>рмы <strong>о</strong>т неб<strong>о</strong>льших д<strong>о</strong> б<strong>о</strong>лее или менее<br />

<strong>к</strong>рупных; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>а</strong>р<strong>а</strong>б<strong>о</strong>личе<strong>с</strong><strong>к</strong><strong>а</strong>я или тр<strong>а</strong>пецеид<strong>а</strong>льн<strong>а</strong>я;<br />

<strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> неб<strong>о</strong>льшие, зерн<strong>о</strong><strong>в</strong>идные<br />

или <strong>о</strong><strong>к</strong>руглые, <strong>в</strong> передней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы ин<strong>о</strong>гд<strong>а</strong> не<strong>о</strong>тчетли<strong>в</strong>ые; пер<strong>в</strong>ый <strong>с</strong>егмент<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>чень <strong>в</strong>ели<strong>к</strong>, рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т<br />

других и <strong>о</strong>бр<strong>а</strong>зует ее переднюю ч<strong>а</strong><strong>с</strong>ть; б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

<strong>к</strong>р<strong>а</strong>я б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> переднег<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> <strong>с</strong>луж<strong>а</strong>т <strong>к</strong><strong>а</strong><strong>к</strong><br />

бы пр<strong>о</strong>д<strong>о</strong>лжением б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н пр<strong>о</strong><strong>с</strong><strong>о</strong>мы;<br />

тель<strong>с</strong><strong>о</strong>н уз<strong>к</strong>ий, <strong>к</strong>лин<strong>о</strong><strong>в</strong>идный; <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти уз<strong>к</strong>ие,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие или длинные; н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><br />

чешуе<strong>в</strong>идный или непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>-ячеи<strong>с</strong>тый. Н.<br />

де<strong>в</strong><strong>о</strong>н — <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Borchgrevinkium Novojilov, 1958. Тип<br />

р<strong>о</strong>д<strong>а</strong> — В. taimyrense Novojilov, 1958; н. де<strong>в</strong><strong>о</strong>н<br />

(<strong>с</strong><strong>а</strong>мый <strong>в</strong>ерхний г<strong>о</strong>риз<strong>о</strong>нт), юг<strong>о</strong>-з<strong>а</strong>п<strong>а</strong>дный Т<strong>а</strong>ймыр<br />

(р. Им<strong>а</strong>игд<strong>а</strong>). П<strong>а</strong>р<strong>а</strong>б<strong>о</strong>личе<strong>с</strong><strong>к</strong><strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>о</strong>дну треть (или б<strong>о</strong>лее) <strong>в</strong><strong>с</strong>ей длины;<br />

передняя ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я <strong>о</strong>дним<br />

б<strong>о</strong>льшим <strong>с</strong>егмент<strong>о</strong>м (или д<strong>в</strong>умя?), <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<br />

немн<strong>о</strong>г<strong>о</strong> меньше трети <strong>в</strong><strong>с</strong>ей <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы;<br />

EURYPTERIDA INCERTAE SEDIS<br />

• Melbournopterus Caster et Kjellesvig-Waering,<br />

1953 (ри<strong>с</strong>. 1219); <strong>в</strong>. <strong>с</strong>илур, А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия<br />

(Ви<strong>к</strong>т<strong>о</strong>рия). Из<strong>в</strong>е<strong>с</strong>тен т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я<br />

<strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т пр<strong>о</strong><strong>с</strong><strong>о</strong>м <strong>в</strong><strong>с</strong>ех из<strong>в</strong>е<strong>с</strong>тных<br />

422<br />

http://jurassic.ru/<br />

п<strong>о</strong><strong>с</strong>ледующие <strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>бр<strong>а</strong>зуют<br />

ее з<strong>а</strong>днюю треуг<strong>о</strong>льную ч<strong>а</strong><strong>с</strong>ть; п<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> д<strong>в</strong>ур<strong>а</strong>здельным <strong>с</strong>пинным<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м; тель<strong>с</strong><strong>о</strong>н уз<strong>к</strong>ий, <strong>к</strong>лин<strong>о</strong><strong>в</strong>идный; н<strong>а</strong><br />

тергит<strong>а</strong>х з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы имеют<strong>с</strong>я р<strong>а</strong>ди<strong>а</strong>льные<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и, <strong>с</strong>уж<strong>а</strong>ющие<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д и д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющие<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>ередины де<strong>с</strong>ят<strong>о</strong>г<strong>о</strong> тергит<strong>а</strong>;<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти т<strong>о</strong>н<strong>к</strong>ие, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, <strong>с</strong> шип<strong>а</strong>ми н<strong>а</strong><br />

ди<strong>с</strong>т<strong>а</strong>льных члени<strong>к</strong><strong>а</strong>х; н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными <strong>с</strong>етч<strong>а</strong>тыми<br />

ячей<strong>к</strong><strong>а</strong>ми <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>лу<strong>к</strong>руглых «чешуе<strong>к</strong>». Гл<strong>а</strong>з<strong>а</strong><br />

нея<strong>с</strong>ны (ри<strong>с</strong>. 1218; т<strong>а</strong>бл. XIX, фиг. 7). Один<br />

<strong>в</strong>ид. Н. де<strong>в</strong><strong>о</strong>н С. Сибири.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, Mycterops С<strong>о</strong>ре, 1886 (= Glaucodes<br />

Pruvost, 1923) (ри<strong>с</strong>. 1217; т<strong>а</strong>бл. XIX,<br />

фиг. 6; т<strong>а</strong>бл. XXII), <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н США (Пен<strong>с</strong>иль<strong>в</strong><strong>а</strong>ния)<br />

и Бельгии.<br />

СЕМЕЙСТВО WOODWARDOPTERIDAE<br />

KJELLESVIG-WAERING, 1959<br />

Длин<strong>а</strong> ф<strong>о</strong>рм 20—30 <strong>с</strong>м; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>а</strong>р<strong>а</strong>б<strong>о</strong>личе<strong>с</strong><strong>к</strong><strong>а</strong>я;<br />

неб<strong>о</strong>льшие б<strong>о</strong>лее или менее п<strong>о</strong>ч<strong>к</strong><strong>о</strong>- или<br />

зерн<strong>о</strong><strong>в</strong>идные <strong>с</strong>л<strong>о</strong>жные гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong><br />

передней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>тделяя <strong>с</strong>реднюю<br />

треть ее ширины <strong>в</strong> эт<strong>о</strong>м ме<strong>с</strong>те; гл<strong>а</strong>з<strong>к</strong>и неиз<strong>в</strong>е<strong>с</strong>тны;<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы уз<strong>к</strong>ие, <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ющие<br />

<strong>в</strong> длине <strong>с</strong>переди н<strong>а</strong>з<strong>а</strong>д, без шип<strong>о</strong><strong>в</strong> н<strong>а</strong> члени<strong>к</strong><strong>а</strong>х.<br />

Один (или д<strong>в</strong><strong>а</strong>?) передний <strong>с</strong>егмент <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>к</strong>рупный, <strong>о</strong><strong>с</strong>т<strong>а</strong>льные <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> б<strong>о</strong>-<br />

Ри<strong>с</strong>. 1219. Отряд Eurypterida. Ф<strong>о</strong>рм<strong>а</strong> не<strong>о</strong>пределенн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> п<strong>о</strong>л<strong>о</strong>жения, пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> Melbournopterus<br />

crossotus Caster et Kjellesvig-Waering; Xl,5, <strong>в</strong>. <strong>с</strong>илур,<br />

А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия (шт. Ви<strong>к</strong>т<strong>о</strong>рия) (St0rmer, 1955)<br />

<strong>к</strong><strong>о</strong><strong>в</strong>ыми л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями. Н<strong>а</strong>ружный п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и тергит<strong>о</strong><strong>в</strong> перье<strong>в</strong>идный или бугри<strong>с</strong>тый (?).<br />

Один р<strong>о</strong>д — Woodwardopterus Kjellesvig-Waering,<br />

1959 (ри<strong>с</strong>. 1216) из н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

Eurypterida <strong>в</strong><strong>о</strong>гнутым передним и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

<strong>к</strong>р<strong>а</strong>ями и зубч<strong>а</strong>т<strong>о</strong>й б<strong>а</strong>хр<strong>о</strong>м<strong>о</strong>й н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е.<br />

О<strong>с</strong><strong>о</strong>бенн<strong>о</strong>е же <strong>о</strong>тличие з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ет<strong>с</strong>я <strong>в</strong> ф<strong>о</strong>рме<br />

и п<strong>о</strong>л<strong>о</strong>жении <strong>с</strong>л<strong>о</strong>жных гл<strong>а</strong>з; п<strong>о</strong> ф<strong>о</strong>рме <strong>о</strong>ни н<strong>а</strong>-

п<strong>о</strong>мин<strong>а</strong>ют р<strong>о</strong>мбиче<strong>с</strong><strong>к</strong>ие <strong>с</strong>те<strong>к</strong>л<strong>а</strong> <strong>а</strong><strong>в</strong>т<strong>о</strong>м<strong>о</strong>бильных<br />

<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong> и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Эти <strong>о</strong>тличия и неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>е <strong>с</strong>тр<strong>о</strong>ение <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

не п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляют у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ить <strong>о</strong>пределенн<strong>о</strong>е<br />

т<strong>а</strong><strong>к</strong><strong>с</strong><strong>о</strong>н<strong>о</strong>миче<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение <strong>а</strong><strong>в</strong><strong>с</strong>тр<strong>а</strong>лий<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

ф<strong>о</strong>рмы <strong>в</strong> <strong>о</strong>дн<strong>о</strong>м из д<strong>в</strong>ух п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> Eurypterida.<br />

Unionopterus Chernyshev, 1948; н. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>.<br />

П<strong>о</strong>чти <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>о</strong>-<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> неб<strong>о</strong>льшими,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными у переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

<strong>с</strong>л<strong>о</strong>жными гл<strong>а</strong>з<strong>а</strong>ми и центр<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

гл<strong>а</strong>з<strong>к</strong><strong>а</strong>ми; х<strong>о</strong>дильные н<strong>о</strong>ги (<strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь<br />

В нед<strong>а</strong><strong>в</strong>н<strong>о</strong> <strong>в</strong>ышедшей р<strong>а</strong>б<strong>о</strong>те Челле<strong>с</strong><strong>в</strong>иг<strong>а</strong>-<br />

Веринг<strong>а</strong> (Е. N. Kjellesvig-Waering «The Silurian<br />

Eurypterida of the Welsh Borderland» — Jour.<br />

Paleont., v. 35, No 4, p. 789—835) у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лен<br />

н<strong>о</strong><strong>в</strong>ый р<strong>о</strong>д Parahughmilleria <strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е Hugh-<br />

ОТРЯД SCORPION1DA. СКОРПИОНЫ<br />

С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны, дре<strong>в</strong>нейшие н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые,<br />

из<strong>в</strong>е<strong>с</strong>тные уже <strong>с</strong> <strong>с</strong>илур<strong>а</strong>. С<strong>о</strong><strong>в</strong>ременные<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны — <strong>с</strong><strong>а</strong>мые <strong>к</strong>рупные н<strong>а</strong>земные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>, длин<strong>о</strong>й д<strong>о</strong> 20 <strong>с</strong>м. Кр<strong>о</strong>ме<br />

т<strong>о</strong>г<strong>о</strong>, <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны — един<strong>с</strong>т<strong>в</strong>енные <strong>с</strong><strong>о</strong><strong>в</strong>ременные<br />

хелицер<strong>о</strong><strong>в</strong>ые, <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шие м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong><br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong> — 19.<br />

Тел<strong>о</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>в</strong>ытянут<strong>о</strong>е, чет<strong>к</strong><strong>о</strong> р<strong>а</strong>зделяет<strong>с</strong>я<br />

н<strong>а</strong> три <strong>о</strong>тдел<strong>а</strong>: неб<strong>о</strong>льшую г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

(пр<strong>о</strong><strong>с</strong><strong>о</strong>му), шир<strong>о</strong><strong>к</strong>ую переднюю ч<strong>а</strong><strong>с</strong>ть брюш<strong>к</strong><strong>а</strong><br />

— мез<strong>о</strong><strong>с</strong><strong>о</strong>му (<strong>с</strong>емь <strong>с</strong>егмент<strong>о</strong><strong>в</strong>) и <strong>с</strong>уженый<br />

з<strong>а</strong>дний <strong>о</strong>тдел брюш<strong>к</strong><strong>а</strong> — мет<strong>а</strong><strong>с</strong><strong>о</strong>му (пять <strong>с</strong>егмент<strong>о</strong><strong>в</strong>),<br />

з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ющую<strong>с</strong>я тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м <strong>с</strong> <strong>к</strong><strong>о</strong>гте<strong>в</strong>идным<br />

ж<strong>а</strong>лящим <strong>о</strong>рг<strong>а</strong>н<strong>о</strong>м н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не.<br />

В <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди, или пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и других Chelicerata (<strong>с</strong>м. <strong>с</strong>тр. 00),<br />

<strong>в</strong>х<strong>о</strong>дят <strong>а</strong><strong>к</strong>р<strong>о</strong>н и ше<strong>с</strong>ть <strong>с</strong>егмент<strong>о</strong><strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей: хелицер, педип<strong>а</strong>льп и четырех<br />

п<strong>а</strong>р х<strong>о</strong>дильных н<strong>о</strong>г. Седьм<strong>о</strong>й <strong>с</strong>егмент, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящий<strong>с</strong>я<br />

уже <strong>к</strong> брюш<strong>к</strong>у, у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых жи<strong>в</strong><strong>о</strong>тных<br />

рудимент<strong>а</strong>рен; <strong>с</strong>леды ег<strong>о</strong> пр<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я<br />

лишь <strong>в</strong> н<strong>а</strong>личии <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующей ему п<strong>а</strong>ры<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>а</strong>ртерий и лиг<strong>а</strong>мент<strong>о</strong><strong>в</strong> <strong>с</strong>ердц<strong>а</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> з<strong>а</strong>ним<strong>а</strong>ет<br />

<strong>в</strong><strong>с</strong>ег<strong>о</strong> '/<strong>в</strong> ч<strong>а</strong><strong>с</strong>ть тел<strong>а</strong> (ри<strong>с</strong>. 1220, 1257).<br />

Спинн<strong>о</strong>й щит г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди (<strong>с</strong><strong>а</strong>г<strong>а</strong>р<strong>а</strong>х)—<strong>с</strong>ли<strong>в</strong>шие<strong>с</strong>я<br />

<strong>а</strong><strong>к</strong>р<strong>о</strong>н и тергиты ше<strong>с</strong>ти <strong>с</strong>ледующих <strong>с</strong>егмент<strong>о</strong><strong>в</strong>;<br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть ег<strong>о</strong> нер<strong>о</strong><strong>в</strong>н<strong>а</strong>я, <strong>с</strong><strong>к</strong>ульптир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong><br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми, ям<strong>к</strong><strong>а</strong>ми и ряд<strong>а</strong>ми буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong><br />

и не<strong>с</strong>ет <strong>о</strong>тдельные <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и. В <strong>с</strong>редней<br />

ч<strong>а</strong><strong>с</strong>ти щит<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>к</strong>рупные меди<strong>а</strong>льные<br />

гл<strong>а</strong>з<strong>а</strong>, п<strong>о</strong>л<strong>о</strong>жение <strong>к</strong><strong>о</strong>т<strong>о</strong>рых шир<strong>о</strong><strong>к</strong><strong>о</strong> и<strong>с</strong>п<strong>о</strong>льзует<strong>с</strong>я<br />

<strong>в</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>е (ри<strong>с</strong>. 1221 А). О<strong>к</strong><strong>о</strong>л<strong>о</strong> перед­<br />

<strong>в</strong>т<strong>о</strong>р<strong>а</strong>я пр<strong>а</strong><strong>в</strong><strong>а</strong>я, третья ле<strong>в</strong><strong>а</strong>я неп<strong>о</strong>лн<strong>а</strong>я и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> чет<strong>в</strong>ерт<strong>о</strong>й) <strong>с</strong> шип<strong>а</strong>ми, н<strong>о</strong> не<br />

у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лен<strong>о</strong>, <strong>с</strong> п<strong>о</strong>д<strong>в</strong>ижными или неп<strong>о</strong>д<strong>в</strong>ижными;<br />

пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong> уз<strong>к</strong>ими л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями<br />

и <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й шп<strong>о</strong>р<strong>о</strong>й н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м<br />

<strong>к</strong>р<strong>а</strong>е л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

и уз<strong>к</strong><strong>о</strong>й мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; тель<strong>с</strong><strong>о</strong>н неиз<strong>в</strong>е<strong>с</strong>тен.<br />

Пл<strong>о</strong>хие иллю<strong>с</strong>тр<strong>а</strong>ции не д<strong>а</strong>ют <strong>о</strong>тчетли<strong>в</strong><strong>о</strong>г<strong>о</strong><br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ления <strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тях<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>нн<strong>о</strong>й Черныше<strong>в</strong>ым ф<strong>о</strong>рмы из <strong>к</strong><strong>а</strong>р<strong>а</strong>г<strong>а</strong>ндин<strong>с</strong><strong>к</strong><strong>о</strong>й<br />

<strong>с</strong><strong>в</strong>иты К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н<strong>а</strong>. К<strong>о</strong>лле<strong>к</strong>ци<strong>о</strong>нный<br />

н<strong>о</strong>мер и ме<strong>с</strong>т<strong>о</strong> хр<strong>а</strong>нения <strong>о</strong>бр<strong>а</strong>зц<strong>а</strong> неиз<strong>в</strong>е<strong>с</strong>тны.<br />

(| В. Б. Дубинин \)<br />

milleriidae и <strong>в</strong><strong>о</strong>з<strong>в</strong>еден <strong>в</strong> р<strong>а</strong>нг р<strong>о</strong>д<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> т<strong>а</strong><strong>к</strong><strong>с</strong><strong>о</strong>н<strong>а</strong><br />

п<strong>о</strong>др<strong>о</strong>д Erettopterus Salter, 1859 (р<strong>о</strong>д Pterygotus,<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pterygotidae). У<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лен<strong>о</strong><br />

11 н<strong>о</strong><strong>в</strong>ых <strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong>зличных р<strong>о</strong>д<strong>о</strong><strong>в</strong>, гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м<br />

п<strong>о</strong> прежде <strong>о</strong>пи<strong>с</strong><strong>а</strong>нным ф<strong>о</strong>рм<strong>а</strong>м.<br />

не-б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых угл<strong>о</strong><strong>в</strong> щит<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я <strong>о</strong>т д<strong>в</strong>ух<br />

д<strong>о</strong> пяти м<strong>а</strong>лень<strong>к</strong>их б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых гл<strong>а</strong>з (ри<strong>с</strong>. 1228,<br />

1235). Брюшн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди п<strong>о</strong>чти<br />

цели<strong>к</strong><strong>о</strong>м п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми педип<strong>а</strong>льп и х<strong>о</strong>дильных<br />

н<strong>о</strong>г (ри<strong>с</strong>. 1223, 1226А, Е). П<strong>о</strong>з<strong>а</strong>ди<br />

между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г III и IV имеет<strong>с</strong>я неб<strong>о</strong>льш<strong>а</strong>я<br />

треуг<strong>о</strong>льн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> — мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м, или<br />

<strong>с</strong>тернум (ри<strong>с</strong>. 1226А, Е),— пр<strong>о</strong>из<strong>о</strong>шедш<strong>а</strong>я <strong>о</strong>т<br />

<strong>с</strong>лияния <strong>с</strong>тернит<strong>о</strong><strong>в</strong> пят<strong>о</strong>г<strong>о</strong> и ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

П<strong>о</strong>з<strong>а</strong>ди <strong>с</strong>тернум<strong>а</strong> леж<strong>а</strong>т п<strong>а</strong>рные п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>к</strong>рышеч<strong>к</strong>и<br />

(ри<strong>с</strong>. 1223, 1226А, Е) п<strong>о</strong>л<strong>о</strong>жение <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

и<strong>с</strong>п<strong>о</strong>льзует<strong>с</strong>я <strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции. Т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп<br />

и н<strong>о</strong>г I и II н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

имеют л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти — т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емые челю<strong>с</strong>тные<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и, т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х н<strong>о</strong>г III и IV<br />

<strong>о</strong>ни <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют (ри<strong>с</strong>. 1226А, Е). В р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти<br />

имеет<strong>с</strong>я д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> <strong>к</strong>рупный р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>ну<strong>с</strong>,<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> лежит р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие;<br />

м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>о</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

между <strong>с</strong>егмент<strong>а</strong>ми педип<strong>а</strong>льп и н<strong>о</strong>г I. Клешне<strong>в</strong>идные<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые хелицеры леж<strong>а</strong>т <strong>в</strong>переди<br />

и н<strong>а</strong>д р<strong>о</strong>т<strong>о</strong><strong>в</strong>ым <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тием (ри<strong>с</strong>. 1232,<br />

1244); п<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец хелицер у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong>зд<strong>в</strong><strong>о</strong>ен н<strong>а</strong> <strong>к</strong><strong>о</strong>нце. Педип<strong>а</strong>льпы у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

зн<strong>а</strong>чительн<strong>о</strong> т<strong>о</strong>лще и длиннее других<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей; <strong>о</strong>ни <strong>к</strong>лешне<strong>в</strong>идны и <strong>с</strong>луж<strong>а</strong>т для<br />

л<strong>о</strong><strong>в</strong>ли, удерж<strong>а</strong>ния и п<strong>о</strong>дне<strong>с</strong>ения <strong>к</strong> хелицер<strong>а</strong>м<br />

д<strong>о</strong>бычи. Педип<strong>а</strong>льпы <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>:<br />

п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>, <strong>в</strong>ертлуг<strong>а</strong>, бедр<strong>а</strong>, <strong>к</strong><strong>о</strong>лен<strong>а</strong>,<br />

г<strong>о</strong>лени и <strong>к</strong><strong>о</strong>нечн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> л<strong>а</strong>п<strong>к</strong>и. Х<strong>о</strong>дильные<br />

н<strong>о</strong>ги <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие у <strong>в</strong><strong>о</strong>дных — <strong>с</strong>илурий<strong>с</strong><strong>к</strong>их—<br />

ф<strong>о</strong>рм, п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> у<strong>в</strong>еличи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> р<strong>а</strong>змер<strong>а</strong>х <strong>о</strong>т пер<strong>в</strong><strong>о</strong>й д<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>й з<strong>а</strong>дней п<strong>а</strong>ры,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных. В<strong>с</strong>е н<strong>о</strong>ги <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из<br />

423<br />

http://jurassic.ru/

<strong>в</strong><strong>о</strong><strong>с</strong>ьми члени<strong>к</strong><strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пер<strong>в</strong>ые пять т<strong>а</strong><strong>к</strong>ие<br />

же, <strong>к</strong><strong>а</strong><strong>к</strong> у педип<strong>а</strong>льп, ше<strong>с</strong>т<strong>о</strong>й и <strong>с</strong>едьм<strong>о</strong>й н<strong>о</strong><strong>с</strong>ят<br />

н<strong>а</strong>з<strong>в</strong><strong>а</strong>ния пер<strong>в</strong><strong>о</strong>г<strong>о</strong> и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> л<strong>а</strong>п<strong>к</strong>и<br />

(metatarsus), <strong>а</strong> -п<strong>о</strong><strong>с</strong>ледний — <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>й —<br />

члени<strong>к</strong> не<strong>с</strong>ет <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и. Т<strong>а</strong>зи<strong>к</strong>и х<strong>о</strong>дильных н<strong>о</strong>г<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong>литы <strong>с</strong> тел<strong>о</strong>м. Н<strong>о</strong>ги<br />

<strong>с</strong> ред<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных <strong>о</strong>тдел<strong>а</strong>х и<br />

<strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными щетин<strong>к</strong><strong>а</strong>ми и шп<strong>о</strong>р<strong>а</strong>ми н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце<strong>в</strong>ых члени<strong>к</strong><strong>а</strong>х. В<strong>с</strong>е <strong>в</strong>зр<strong>о</strong><strong>с</strong>лые <strong>о</strong><strong>с</strong><strong>о</strong>би <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> имеют н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х н<strong>о</strong>г<br />

п<strong>о</strong> д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong> (Dionychopodes), т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>с</strong>илурий<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны были «<strong>о</strong>дн<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>ые»<br />

(Apoxypodes) (ри<strong>с</strong>. 1220, 1222), т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и<br />

пер<strong>в</strong>ые, <strong>с</strong><strong>а</strong>мые м<strong>о</strong>л<strong>о</strong>дые <strong>с</strong>т<strong>а</strong>дии р<strong>а</strong>з<strong>в</strong>ития у <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

ф<strong>о</strong>рм (П<strong>а</strong><strong>в</strong>л<strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong>ий, 1917).<br />

Пер<strong>в</strong>ые ше<strong>с</strong>ть <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы (VIII—<br />

XIII) <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ид<strong>о</strong>измененными <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями<br />

(п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>к</strong>рыш<strong>к</strong>и, гребне<strong>в</strong>идные прид<strong>а</strong>т<strong>к</strong>и,<br />

лег<strong>к</strong>ие). У <strong>с</strong>илурий<strong>с</strong><strong>к</strong>их <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> (Apoxypodes)<br />

при гребне<strong>в</strong>идных <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>х были еще и<br />

рудименты тел<strong>о</strong>п<strong>о</strong>дит<strong>о</strong><strong>в</strong> н<strong>о</strong>г (ри<strong>с</strong>. 1222). П<strong>о</strong><strong>с</strong>ледний<br />

<strong>с</strong>егмент р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>г<strong>о</strong> переднебрюшья<br />

(XIV), т<strong>а</strong><strong>к</strong> же <strong>к</strong><strong>а</strong><strong>к</strong> и <strong>в</strong><strong>с</strong>е <strong>с</strong>егменты мет<strong>а</strong><strong>с</strong><strong>о</strong>мы,<br />

лишены <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей или их рудимент<strong>о</strong><strong>в</strong>. Шир<strong>о</strong><strong>к</strong>ие<br />

и упл<strong>о</strong>щенные <strong>с</strong>егменты мез<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong><strong>о</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р'<strong>о</strong>ны п<strong>о</strong><strong>к</strong>рыты тергит<strong>а</strong>ми (ри<strong>с</strong>.<br />

1220), <strong>с</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>тчленяет<strong>с</strong>я<br />

п<strong>о</strong> м<strong>а</strong>лень<strong>к</strong><strong>о</strong>му б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>му щит<strong>к</strong>у. С брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>с</strong>егменты мез<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong><strong>к</strong>рыты <strong>с</strong>тернит<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1244). Стернит IX <strong>с</strong>егмент<strong>а</strong> (<strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы) пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен неб<strong>о</strong>льш<strong>о</strong>й<br />

треуг<strong>о</strong>льн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>о</strong>й — мет<strong>а</strong><strong>с</strong>тернум<strong>о</strong>м,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женным между гребне<strong>в</strong>идными<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми (ри<strong>с</strong>. 1244). Стерниты <strong>о</strong><strong>с</strong>т<strong>а</strong>льных<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы шир<strong>о</strong><strong>к</strong>ие. Н<strong>а</strong> VIII <strong>с</strong>егменте<br />

у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> и у <strong>в</strong><strong>с</strong>ех других хелицер<strong>о</strong><strong>в</strong>ых,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong> д<strong>в</strong>умя<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ыми <strong>к</strong>рышеч<strong>к</strong><strong>а</strong>ми. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>к</strong>рышеч<strong>к</strong>и<br />

г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичны <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тям пер<strong>в</strong><strong>о</strong>г<strong>о</strong> брюшн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong>. Рудимент<strong>а</strong>ми <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

брюшн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> я<strong>в</strong>ляют<strong>с</strong>я гребне<strong>в</strong>идные<br />

прид<strong>а</strong>т<strong>к</strong>и (ри<strong>с</strong>. 1243) — <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зные <strong>о</strong>рг<strong>а</strong>ны<br />

чу<strong>в</strong><strong>с</strong>т<strong>в</strong>, <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong>енные т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong>м. Н<strong>а</strong><br />

<strong>с</strong>ледующих четырех <strong>с</strong>егмент<strong>а</strong>х (X—XIII) р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong><br />

п<strong>о</strong> п<strong>а</strong>ре лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>; у з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я <strong>к</strong><strong>а</strong>жд<strong>о</strong>г<strong>о</strong> из них <strong>в</strong>идн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я<br />

щель — дых<strong>а</strong>льце. Лег<strong>к</strong>ие <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичны<br />

ж<strong>а</strong>бр<strong>о</strong>н<strong>о</strong><strong>с</strong>ным <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тям Eurypterodea.<br />

Интере<strong>с</strong>н<strong>о</strong>, чт<strong>о</strong> у <strong>с</strong>илурий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>в</strong><strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong><br />

лег<strong>к</strong>ие <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли (ри<strong>с</strong>. 1222), <strong>а</strong> у<br />

Isobuthus Fritsch и Microlabis Corda н<strong>а</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не имели<strong>с</strong>ь еще <strong>к</strong>рупные п<strong>о</strong>лу<strong>к</strong>руглые<br />

«ж<strong>а</strong>берные» щит<strong>к</strong>и (ри<strong>с</strong>. 1245). П<strong>о</strong><strong>с</strong>ледний<br />

<strong>с</strong>егмент мез<strong>о</strong><strong>с</strong><strong>о</strong>мы (XIV или VII тул<strong>о</strong><strong>в</strong>ищный)<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я уже <strong>к</strong> мет<strong>а</strong><strong>с</strong><strong>о</strong>ме и я<strong>в</strong>ляет<strong>с</strong>я, п<strong>о</strong><br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>у, ее пер<strong>в</strong>ым <strong>с</strong>егмент<strong>о</strong>м; н<strong>а</strong> нем <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, н<strong>о</strong> имеют<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льные<br />

<strong>в</strong><strong>а</strong>ли<strong>к</strong>и, х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные для <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мет<strong>а</strong><strong>с</strong><strong>о</strong>мы<br />

424<br />

http://jurassic.ru/<br />

(ри<strong>с</strong>. 1222, 1258Б). Н<strong>а</strong> пяти члени<strong>к</strong><strong>а</strong>х мет<strong>а</strong><strong>с</strong><strong>о</strong>мы<br />

н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>лн<strong>о</strong>е <strong>с</strong>лияние тергит<strong>о</strong><strong>в</strong> и<br />

<strong>с</strong>тернит<strong>о</strong><strong>в</strong>; <strong>к</strong><strong>а</strong>ждый члени<strong>к</strong> зде<strong>с</strong>ь имеет <strong>в</strong>ид хит<strong>о</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>льц<strong>а</strong>. Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>чень п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong> и<br />

м<strong>о</strong>жет з<strong>а</strong>гиб<strong>а</strong>ть<strong>с</strong>я дуг<strong>о</strong>й н<strong>а</strong>д <strong>с</strong>пин<strong>о</strong>й жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong>.<br />

Ан<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не между п<strong>о</strong><strong>с</strong>ледним <strong>с</strong>егмент<strong>о</strong>м з<strong>а</strong>днебрюшья<br />

и тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м. Тель<strong>с</strong><strong>о</strong>н не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong>здут <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии и, изгиб<strong>а</strong>я<strong>с</strong>ь н<strong>а</strong> <strong>с</strong>пинную<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну, з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ет<strong>с</strong>я и<strong>с</strong><strong>к</strong>ри<strong>в</strong>ленн<strong>о</strong>й игл<strong>о</strong>й —<br />

ж<strong>а</strong>л<strong>о</strong>м (ри<strong>с</strong>. 1222, 1243, 1254). В тель<strong>с</strong><strong>о</strong>не р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

д<strong>в</strong>е яд<strong>о</strong><strong>в</strong>итые железы, пр<strong>о</strong>т<strong>о</strong><strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ершине ж<strong>а</strong>л<strong>а</strong>.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и б и <strong>о</strong> г е <strong>о</strong> г р <strong>а</strong> ф и я. В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящее<br />

<strong>в</strong>ремя <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены <strong>в</strong><strong>о</strong><br />

<strong>в</strong><strong>с</strong>ех ч<strong>а</strong><strong>с</strong>тях <strong>с</strong><strong>в</strong>ет<strong>а</strong>, преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>м п<strong>о</strong>я<strong>с</strong><strong>а</strong>х. Обит<strong>а</strong>ют <strong>в</strong><br />

<strong>с</strong>ухих пу<strong>с</strong>тынных, <strong>с</strong>тепных и г<strong>о</strong>рных у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях,<br />

<strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>в</strong> ле<strong>с</strong><strong>а</strong>х. Гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м н<strong>о</strong>чные жи<strong>в</strong><strong>о</strong>тные,<br />

хищни<strong>к</strong>и. В <strong>с</strong>илурий<strong>с</strong><strong>к</strong><strong>о</strong>м пери<strong>о</strong>де<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong>ели <strong>в</strong><strong>о</strong>дный <strong>о</strong>бр<strong>а</strong>з жизни,<br />

н<strong>а</strong>п<strong>о</strong>д<strong>о</strong>бие э<strong>в</strong>риптерид. С <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я<br />

н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие н<strong>а</strong>земные ф<strong>о</strong>рмы, близ<strong>к</strong>ие <strong>к</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременным <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong>м.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме и фил<strong>о</strong>гения.<br />

С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны близ<strong>к</strong>и <strong>к</strong> э<strong>в</strong>риптерид<strong>а</strong>м<br />

(Eurypterida — <strong>с</strong>м. <strong>с</strong>тр. 407), п<strong>о</strong>т<strong>о</strong>м<strong>к</strong><strong>а</strong>ми <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>ни я<strong>в</strong>ляют<strong>с</strong>я. Ан<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong><strong>а</strong>я близ<strong>о</strong><strong>с</strong>ть<br />

этих д<strong>в</strong>ух <strong>о</strong>тряд<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>, чт<strong>о</strong><br />

д<strong>а</strong>ет пр<strong>а</strong><strong>в</strong><strong>о</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ть <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> н<strong>а</strong>земными<br />

э<strong>в</strong>риптерид<strong>а</strong>ми (Бе<strong>к</strong>лемише<strong>в</strong>, 1952).Близ<strong>о</strong><strong>с</strong>ть<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>к</strong> э<strong>в</strong>риптерид<strong>а</strong>м зн<strong>а</strong>чительн<strong>о</strong> б<strong>о</strong>льш<strong>а</strong>я,<br />

чем <strong>к</strong> другим <strong>о</strong>тряд<strong>а</strong>м н<strong>а</strong>земных п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> и<br />

их <strong>в</strong>з<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шение <strong>с</strong> рецентными ф<strong>о</strong>рм<strong>а</strong>ми<br />

<strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>тельн<strong>о</strong> не у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лены. Пер<strong>в</strong>ым Т<strong>о</strong>релл<br />

(Thorell et Lindstrom 1885) <strong>в</strong>ыделил <strong>с</strong>илурий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong> Palaeophonus nuncius Thoroll<br />

et Lindstrom, 1885 <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бый п<strong>о</strong>д<strong>о</strong>тряд Apoxypodes,<br />

пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>и<strong>в</strong> ему <strong>в</strong><strong>с</strong>ех других<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, имеющих н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>,<br />

— п<strong>о</strong>д<strong>о</strong>тряд Dionychopodes. Д<strong>а</strong>льнейшие и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ли, чт<strong>о</strong> Apoxypodes жили,<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong>илуре, <strong>а</strong> <strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не уже <strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли<br />

типичн<strong>о</strong> н<strong>а</strong>земные Dionychopodes. Эт<strong>о</strong> деление<br />

приним<strong>а</strong>ют Вернер (Werner, 1935), В<strong>а</strong>терл<strong>о</strong><br />

(Waterlot 1949), Бе<strong>к</strong>лемише<strong>в</strong> (1952) и др.<br />

К<strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> Т<strong>о</strong>релл (1885)<br />

<strong>о</strong>бъединил <strong>в</strong> группу Anthracoscorpii, пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>и<strong>в</strong><br />

им <strong>в</strong><strong>с</strong>ех <strong>с</strong><strong>о</strong><strong>в</strong>ременных Neoscorpii, нез<strong>а</strong><strong>в</strong>и<strong>с</strong>им<strong>о</strong><br />

<strong>о</strong>т м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>й близ<strong>о</strong><strong>с</strong>ти п<strong>о</strong><strong>с</strong>ледних<br />

<strong>к</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льным ф<strong>о</strong>рм<strong>а</strong>м. П<strong>о</strong>зднее<br />

П<strong>о</strong><strong>к</strong><strong>о</strong><strong>к</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911), <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>я<strong>с</strong>ь н<strong>а</strong> р<strong>а</strong>зличиях<br />

<strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди,<br />

р<strong>а</strong>зделил <strong>в</strong><strong>с</strong>ех <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> н<strong>а</strong> группу б<strong>о</strong>лее<br />

примити<strong>в</strong>ных <strong>в</strong>ид<strong>о</strong><strong>в</strong> — Lobosterni и н<strong>а</strong> группу<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, п<strong>о</strong>х<strong>о</strong>жих н<strong>а</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных,— Ortho-

sterni. Д<strong>а</strong>лее Бялыниц<strong>к</strong>ий-Бируля (1917) <strong>в</strong><strong>с</strong>ех<br />

<strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>о</strong>бъединял <strong>в</strong> д<strong>в</strong>е<br />

группы — Palacoscorpiones и Neoscorpiones,<br />

у<strong>с</strong><strong>о</strong><strong>в</strong>ершен<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong><strong>в</strong> этим <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию П<strong>о</strong><strong>к</strong><strong>о</strong><strong>к</strong><strong>а</strong>.<br />

Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, Петрун<strong>к</strong>е<strong>в</strong>ич (Petrunkewitch,<br />

1949, 1953) предл<strong>о</strong>жил н<strong>о</strong><strong>в</strong>ую <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, р<strong>а</strong>здели<strong>в</strong> их н<strong>а</strong> д<strong>в</strong><strong>а</strong><br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Protoscorpiones, имеющих <strong>в</strong><strong>о</strong><strong>с</strong>емь<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, и Euscorpiones <strong>с</strong><br />

<strong>с</strong>емью мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ими <strong>с</strong>егмент<strong>а</strong>ми и. д<strong>в</strong>умя<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х. В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящем изд<strong>а</strong>нии<br />

приним<strong>а</strong>ет<strong>с</strong>я эт<strong>о</strong> деление, н<strong>о</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>м Петрун<strong>к</strong>е<strong>в</strong>ич<strong>а</strong><br />

прид<strong>а</strong>ет<strong>с</strong>я зн<strong>а</strong>чение д<strong>в</strong>ух н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Dionychopodes Thorell.<br />

Р<strong>а</strong><strong>с</strong>пределение <strong>в</strong><strong>о</strong> <strong>в</strong> р е м е н и. С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны<br />

из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong> <strong>с</strong>илур<strong>а</strong> (Apoxypodes). В <strong>к</strong><strong>а</strong>рб<strong>о</strong>не<br />

<strong>о</strong>ни пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены уже мн<strong>о</strong>гими р<strong>о</strong>д<strong>а</strong>ми<br />

Dionychopodes, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых т<strong>о</strong>ль<strong>к</strong><strong>о</strong> Palaeoscorpiidae<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ли<strong>с</strong>ь <strong>о</strong>т <strong>с</strong><strong>о</strong><strong>в</strong>ременных;<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льные р<strong>о</strong>ды я<strong>в</strong>ляют<strong>с</strong>я типичными Neoscorpiones,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и <strong>с</strong><strong>о</strong><strong>в</strong>ременные <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны (Бялыниц<strong>к</strong>ий<br />

— Бируля, 1917; Бе<strong>к</strong>лемише<strong>в</strong>,<br />

1905).<br />

ПОДОТРЯД APOXYPODES.<br />

ЗАОСТРЕННОНОГИЕ СКОРПИОНЫ<br />

Тел<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> н<strong>а</strong> неб<strong>о</strong>льшую г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь,<br />

<strong>с</strong>емь <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мез<strong>а</strong><strong>с</strong><strong>о</strong>мы и мет<strong>а</strong><strong>с</strong><strong>о</strong>му.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>рединными гл<strong>а</strong>з<strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными<br />

н<strong>а</strong> меди<strong>а</strong>льн<strong>о</strong>м буг<strong>о</strong>р<strong>к</strong>е, реже без них<br />

(Dolichophonidae). Вд<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

е<strong>с</strong>ть п<strong>о</strong>перечн<strong>о</strong>е <strong>в</strong>д<strong>а</strong><strong>в</strong>ление, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>е г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гизирует<strong>с</strong>я<br />

<strong>с</strong><strong>о</strong> ш<strong>в</strong><strong>о</strong>м между VI и VII тергит<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1221 А). Хелицеры <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong>лешне<strong>в</strong>идные.<br />

Педип<strong>а</strong>льпы длинные, пяти-члени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>с</strong> <strong>к</strong>рупн<strong>о</strong>й <strong>к</strong>лешней. Х<strong>о</strong>дильные н<strong>о</strong>ги<br />

<strong>с</strong> <strong>с</strong>емью г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мными члени<strong>к</strong><strong>а</strong>ми; <strong>о</strong>ни<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пр<strong>о</strong><strong>с</strong>тым з<strong>а</strong><strong>о</strong><strong>с</strong>тренным<br />

члени<strong>к</strong><strong>о</strong>м (ри<strong>с</strong>. 1221Б). Мет<strong>а</strong><strong>с</strong>т<strong>о</strong>м<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен п<strong>о</strong>з<strong>а</strong>ди н<strong>о</strong>г IV, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong><br />

<strong>в</strong><strong>с</strong>е <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, <strong>к</strong><strong>а</strong><strong>к</strong> и у Eurypterodea, группируют<strong>с</strong>я<br />

<strong>в</strong><strong>о</strong><strong>к</strong>руг рт<strong>а</strong>. Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из<br />

<strong>о</strong>бычных <strong>с</strong>еми <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

н<strong>а</strong> пер<strong>в</strong><strong>о</strong>м <strong>с</strong>егменте. Гребне<strong>в</strong>идные <strong>о</strong>рг<strong>а</strong>ны<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> имеют <strong>с</strong>леды р<strong>а</strong><strong>с</strong>членения<br />

преэпип<strong>о</strong>дит<strong>о</strong><strong>в</strong> (тел<strong>о</strong>п<strong>о</strong>дит<strong>о</strong><strong>в</strong>) брюшн<strong>о</strong>й <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1222). Дых<strong>а</strong>лец н<strong>а</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных четырех <strong>с</strong>егмент<strong>а</strong>х нет,— <strong>в</strong>идим<strong>о</strong>,<br />

зде<strong>с</strong>ь р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ли<strong>с</strong>ь ж<strong>а</strong>берные н<strong>о</strong>ж<strong>к</strong>и, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

не <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь. Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из пяти<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> и тель<strong>с</strong><strong>о</strong>н<strong>а</strong> (ри<strong>с</strong>. 1222).<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Силурий<strong>с</strong><strong>к</strong>ие<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны Apoxypodes <strong>в</strong>ели, п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong><br />

э<strong>в</strong>риптерид<strong>а</strong>м, <strong>в</strong><strong>о</strong>дный <strong>о</strong>бр<strong>а</strong>з жизни, <strong>о</strong>бит<strong>а</strong>я,<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong> неглуб<strong>о</strong><strong>к</strong>их уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>х <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>.<br />

С <strong>в</strong><strong>о</strong>дным, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м жизни <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны и <strong>в</strong><strong>с</strong>е <strong>о</strong>тме­<br />

425<br />

ченные <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>с</strong>тр<strong>о</strong>ения. Обн<strong>а</strong>ружены <strong>в</strong><br />

<strong>о</strong>тл<strong>о</strong>жениях <strong>в</strong>. <strong>с</strong>илур<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (Ш<strong>в</strong>еция, Англия)<br />

и С. Амери<strong>к</strong>и.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер р<strong>а</strong><strong>с</strong>членения тел<strong>а</strong> и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие<br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>с</strong>т<strong>о</strong>ль х<strong>а</strong>р<strong>а</strong><strong>к</strong>терных<br />

для <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют н<strong>а</strong><br />

близ<strong>о</strong><strong>с</strong>ть <strong>с</strong>илурий<strong>с</strong><strong>к</strong>их <strong>в</strong><strong>о</strong>дных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>к</strong><br />

э<strong>в</strong>риптерид<strong>а</strong>м. Силур. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Palaeophonidae<br />

и Dolichophonidae.<br />

СЕМЕЙСТВО PALAEOPHONIDAE THORELL<br />

ET LINDSTROM, 1885<br />

В<strong>с</strong>е члени<strong>к</strong>и н<strong>о</strong>г <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

длины; л<strong>а</strong>п<strong>к</strong>и з<strong>а</strong><strong>о</strong><strong>с</strong>тренные (ри<strong>с</strong>. 1221Б).<br />

Педип<strong>а</strong>льпы <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими члени<strong>к</strong><strong>а</strong>ми. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>.<br />

Palaeophonus Thorell et Lindstrom,<br />

1885. Тип р<strong>о</strong>д<strong>а</strong> — P. nuncius Thorell et Lindstrom,<br />

1885; <strong>с</strong>илур, Ш<strong>в</strong>еция. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный<br />

щит п<strong>о</strong>чти <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тный, <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong><strong>о</strong>гнутый <strong>с</strong>переди.<br />

Клешня педип<strong>а</strong>льп <strong>о</strong>чень м<strong>о</strong>щн<strong>а</strong>я, <strong>с</strong><strong>о</strong> <strong>в</strong>здутым<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием и <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

п<strong>а</strong>льц<strong>а</strong>ми; длин<strong>а</strong> их р<strong>а</strong><strong>в</strong>н<strong>а</strong> длине <strong>в</strong>здут<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния.<br />

Бедр<strong>о</strong> педип<strong>а</strong>льп примерн<strong>о</strong> т<strong>а</strong><strong>к</strong><strong>о</strong>й же<br />

длины, <strong>к</strong><strong>а</strong><strong>к</strong> г<strong>о</strong>лень. Крупные <strong>в</strong>иды; <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><br />

длин<strong>о</strong>й 7,5—8 мм; длин<strong>а</strong> тел<strong>а</strong> д<strong>о</strong> 35,5 мм<br />

(ри<strong>с</strong>. 1221 — 1223). Три <strong>в</strong>ид<strong>а</strong>. Силур 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Proscorpius Whitfield, 1885. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. osborni Whitfield, 1885; <strong>с</strong>илур, С. Амери<strong>к</strong><strong>а</strong>.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>в</strong>ытянут <strong>в</strong> ширину, <strong>в</strong>ыпу<strong>к</strong>лый<br />

<strong>с</strong>переди. Клешня педип<strong>а</strong>льп <strong>в</strong>ытянут<strong>а</strong>я,<br />

<strong>с</strong> уз<strong>к</strong>ими, из<strong>о</strong>гнутыми п<strong>а</strong>льц<strong>а</strong>ми, длин<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

б<strong>о</strong>льше длины-<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти. Бедр<strong>о</strong> педип<strong>а</strong>льп<br />

<strong>в</strong> 2 р<strong>а</strong>з<strong>а</strong> длиннее г<strong>о</strong>лени. Мел<strong>к</strong>ий <strong>в</strong>ид;<br />

длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 4 мм (ри<strong>с</strong>. 1220). Один <strong>в</strong>ид.<br />

Силур С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО DOLICHOPHONIDAE<br />

PETRUNKEVITCH, 1953<br />

Члени<strong>к</strong>и н<strong>о</strong>г, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>,<br />

р<strong>а</strong>зличн<strong>о</strong>й длины. Педип<strong>а</strong>льпы длинные,<br />

<strong>с</strong> <strong>в</strong>ытянутыми т<strong>о</strong>н<strong>к</strong>ими члени<strong>к</strong><strong>а</strong>ми. Один р<strong>о</strong>д.<br />

Силур 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Dolichophonus Petrunkevitch, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Palaeophonus loudonensis Laurie,<br />

1889; <strong>с</strong>илур, Ш<strong>о</strong>тл<strong>а</strong>ндия. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит<br />

з<strong>а</strong>метн<strong>о</strong> <strong>в</strong>ытянут, <strong>в</strong><strong>о</strong>гнут н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>ю.<br />

П<strong>а</strong>льцы педип<strong>а</strong>льп <strong>с</strong> <strong>в</strong><strong>о</strong>гнутыми <strong>в</strong>нутренними<br />

<strong>к</strong>р<strong>а</strong>ями; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong>лешни длиннее п<strong>а</strong>льце<strong>в</strong>.<br />

К<strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong> 9,25 мм длин<strong>о</strong>й и 6,4 мм ширин<strong>о</strong>й <strong>в</strong><br />

з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти (ри<strong>с</strong>. 1224). Один <strong>в</strong>ид. Силур<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

http://jurassic.ru/

Ри<strong>с</strong>. 1220—1225. CeMeftcTBaJPalaeophonidae, Dolichophonidae и Mazoniidae<br />

1220. Proscorpius osborni Whitfield; <strong>о</strong>бщий <strong>в</strong>ид, x2,5, <strong>с</strong>илур, (Petrunkevitch, 1949). 1223. Palaeophonus nunclus Thorell et<br />

С. Амери<strong>к</strong><strong>а</strong> (Werner, 1934). 1221. Palaeophonus nunclus Thorell Lindstrom; <strong>с</strong>хем<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ения <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, Х3,5,<br />

et Lindestrom; А — передний <strong>к</strong><strong>о</strong>нец тел<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, <strong>с</strong>илур, Ш<strong>в</strong>еция (Petrunkevitch, 1953). 1224. Dollchophonus lou-<br />

Х3,5; Б — л<strong>а</strong>п<strong>к</strong><strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й н<strong>о</strong>ги <strong>с</strong> шип<strong>о</strong>м н<strong>а</strong> <strong>в</strong>ершине г<strong>о</strong>лени, donensis (Laurie): <strong>о</strong>бщий <strong>в</strong>ид, Х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>, Ш<strong>о</strong>тл<strong>а</strong>н-<br />

Х30.0; <strong>с</strong>илур, Ш<strong>в</strong>еция (Petrunkevitch, 1949, 1953). 1222. Р<strong>а</strong>- дня (Petrunkevitch, 1953). 1225. Мazonla woodlana Meek et Wortlaeophonus<br />

caledontcus Hunter; <strong>о</strong>бщий <strong>в</strong>ид, x3,5, <strong>с</strong>илур, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> hen; <strong>о</strong>бщий <strong>в</strong>ид, Х3,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1953)<br />

ПОДОТРЯД DIONYCHOPODES.<br />

ДВУКОГОТКОВЫЕ СКОРПИОНЫ<br />

Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>с</strong>еми, реже из <strong>в</strong><strong>о</strong><strong>с</strong>ьми<br />

•<strong>с</strong>егмент<strong>о</strong><strong>в</strong> (т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у Mazoniidae их <strong>в</strong><strong>о</strong><strong>с</strong>емь).<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>к</strong>рупными, <strong>с</strong>рединными гл<strong>а</strong>з<strong>а</strong>ми,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными <strong>в</strong> центре или <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я щит<strong>а</strong>, и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими мел<strong>к</strong>ими гл<strong>а</strong>з<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бычн<strong>о</strong> леж<strong>а</strong>щими <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> ег<strong>о</strong> передне-б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

угл<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1227). Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г з<strong>а</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1241). Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong> п<strong>о</strong>з<strong>а</strong>ди н<strong>о</strong>г II (ри<strong>с</strong>. 1226<strong>а</strong>—е),<br />

426<br />

http://jurassic.ru/<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> з<strong>а</strong>дние п<strong>а</strong>ры н<strong>о</strong>г <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т<strong>о</strong>д<strong>в</strong>инутыми <strong>о</strong>т р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия. Гребне<strong>в</strong>идные<br />

<strong>о</strong>рг<strong>а</strong>ны х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, тел<strong>о</strong>п<strong>о</strong>диты<br />

их нер<strong>а</strong><strong>с</strong>члененные (ри<strong>с</strong>. 1243). Н<strong>а</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не четырех <strong>с</strong>редних <strong>с</strong>егмент<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

(X—XIII) имеют<strong>с</strong>я щеле<strong>в</strong>идные дых<strong>а</strong>тельные<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия, <strong>в</strong>едущие <strong>в</strong> лег<strong>к</strong>ие (ри<strong>с</strong>. 1243).<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Из<strong>в</strong>е<strong>с</strong>тны<br />

<strong>с</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> <strong>в</strong>ремени. Эт<strong>о</strong><br />

типичн<strong>о</strong> <strong>с</strong>ух<strong>о</strong>путные <strong>о</strong>рг<strong>а</strong>низмы, дыш<strong>а</strong>щие <strong>с</strong><br />

п<strong>о</strong>м<strong>о</strong>щью лег<strong>к</strong>их; их <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и н<strong>а</strong>йдены <strong>в</strong> <strong>к</strong><strong>о</strong>нтинент<strong>а</strong>льных<br />

<strong>о</strong>тл<strong>о</strong>жениях. С<strong>о</strong><strong>в</strong>ременн<strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет их приур<strong>о</strong>-

1226 А<br />

1234<br />

1239<br />

1226. Схем<strong>а</strong> <strong>с</strong>тр<strong>о</strong>ения <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>о</strong>тдельных<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Dionychopodes. А — Archaeoctonidae,<br />

Б — Centromachldae, В — Mesophonldae, Г — Isobuthidae,<br />

Д — Cyclophthalmidae, Е — Eoscorpionidae (Petrunkevitch,<br />

1953). 1227. Mazonia woodiana Meek et Worthen; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху,<br />

X9,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1953). 1228. Isobuthus<br />

kralupensis Thorell et Lindstrom; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, x7,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1229. Eoctonus miniatus<br />

Petrunkevitch; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, x8,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch,<br />

1913). 1230. Alloscorpius tuberculatum (Peach); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong><strong>в</strong>ерху, x7,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953).<br />

1231 .Palaeoptsthacanthus schuchertl Petrunkevitch;пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху,<br />

Х8,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1232. Trigonoscorpio<br />

americanus Petrunkevitch; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, x7,0, <strong>к</strong><strong>а</strong>р<strong>о</strong>й,<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1233. Buthioscorpius<br />

ценн<strong>о</strong><strong>с</strong>ть <strong>к</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>им и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>им <strong>о</strong>бл<strong>а</strong><strong>с</strong>тям<br />

(<strong>с</strong>м.<strong>с</strong>тр. 424).<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Д<strong>в</strong>у<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют при­<br />

Ри<strong>с</strong>. 1226—1240. Стр<strong>о</strong>ение пр<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong>зличных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

427<br />

http://jurassic.ru/<br />

buthljormis (Pocock); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, X8,6, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Petrunkevitch, 1953). 1234. Cyclophthalmus senior (Corda); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong><strong>в</strong>ерху, X10,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953).<br />

1235. Compsoscorplus elegans Petrunkevitch; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху,<br />

ХЮ.О, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1236. Europhthalmus<br />

longimanus Petrunkevitch; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху; X3,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1237. Typhloscorplus distinctus<br />

Petrunkevitch; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, x 6,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Petrunkevitch, 1953). 1238. Lichnophthalmus pulcher Petrunkevitch,<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, x2,6, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1949). 1239. Mesophonus perornatus Wills; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, X3,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1240. Garnettlus hungerfordi<br />

(Elias); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, X 3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

(Petrunkevitch, 1949)<br />

мер д<strong>а</strong>льнейшег<strong>о</strong> при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления п<strong>о</strong>т<strong>о</strong>м<strong>к</strong><strong>о</strong><strong>в</strong><br />

пер<strong>в</strong>ичн<strong>о</strong><strong>в</strong><strong>о</strong>дных Eurypterida и Apoxypodes <strong>к</strong><br />

н<strong>а</strong>земн<strong>о</strong>му <strong>о</strong>бр<strong>а</strong>зу жизни (<strong>с</strong>м. <strong>с</strong>тр. 382). Среди<br />

из<strong>в</strong>е<strong>с</strong>тных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>не США (шт<strong>а</strong>т Пен<strong>с</strong>иль<strong>в</strong><strong>а</strong>ния) н<strong>а</strong>йден<strong>а</strong>

Mazonia woodiana Meek et Worthen, 1868<br />

(ри<strong>с</strong>. 1225, 1227), <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>с</strong><strong>о</strong>чет<strong>а</strong>ет призн<strong>а</strong><strong>к</strong>и<br />

<strong>о</strong>б<strong>о</strong>их п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong>. Кл<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льных,<br />

перм<strong>с</strong><strong>к</strong>их и других и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> и их <strong>в</strong>з<strong>а</strong>им<strong>о</strong><strong>о</strong>тн<strong>о</strong>шение <strong>с</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременными<br />

ф<strong>о</strong>рм<strong>а</strong>ми <strong>о</strong><strong>к</strong><strong>о</strong>нч<strong>а</strong>тельн<strong>о</strong> не <strong>в</strong>ыя<strong>с</strong>нены.<br />

В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящем изд<strong>а</strong>нии мы приним<strong>а</strong>ем п<strong>о</strong><strong>с</strong>леднюю<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ци<strong>о</strong>нную <strong>с</strong>хему, предл<strong>о</strong>женную<br />

Петрун<strong>к</strong>е<strong>в</strong>ичем (Petrunkevitch, 1949), причем<br />

п<strong>о</strong>д<strong>о</strong>тряды р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е<br />

н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>.<br />

НАДСЕМЕЙСТВО MAZONIIDEA<br />

Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>в</strong><strong>о</strong><strong>с</strong>ьми <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>й<br />

щит м<strong>а</strong>лень<strong>к</strong>ий, <strong>с</strong> передним <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идным<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>трением; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й ег<strong>о</strong> п<strong>о</strong>лу<strong>к</strong><strong>о</strong>льц<strong>о</strong>м<br />

<strong>о</strong>х<strong>в</strong><strong>а</strong>ты<strong>в</strong><strong>а</strong>ет уз<strong>к</strong><strong>а</strong>я п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>а</strong>, <strong>о</strong>тделенн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й <strong>о</strong>т <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й передней ч<strong>а</strong><strong>с</strong>ти щит<strong>а</strong>;<br />

эт<strong>а</strong> <strong>к</strong><strong>а</strong>йм<strong>а</strong>, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>с</strong><strong>о</strong>б<strong>о</strong>й рудимент<br />

ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong> тергит<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. О<strong>к</strong><strong>о</strong>л<strong>о</strong> переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

угл<strong>о</strong><strong>в</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> имеют<strong>с</strong>я п<strong>о</strong>лулунные<br />

щеле<strong>в</strong>идные <strong>в</strong>д<strong>а</strong><strong>в</strong>ления. Н<strong>а</strong>ряду <strong>с</strong> этими<br />

примити<strong>в</strong>ными призн<strong>а</strong><strong>к</strong><strong>а</strong>ми <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льные<br />

Mazoniidae <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ли ряд<strong>о</strong>м <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей<br />

<strong>с</strong>тр<strong>о</strong>ения, <strong>с</strong>ближ<strong>а</strong>ющих их <strong>с</strong> <strong>в</strong>ымершими и <strong>с</strong><strong>о</strong><strong>в</strong>ременными<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong>ми п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Dionychopodes.<br />

Педип<strong>а</strong>льпы у них были т<strong>о</strong>н<strong>к</strong>ие и длинные,<br />

<strong>с</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong>ертлуг<strong>а</strong>ми и бедр<strong>а</strong>ми,<br />

<strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>с</strong> <strong>о</strong>чень длинным трубч<strong>а</strong>тым <strong>к</strong><strong>о</strong>ленным<br />

члени<strong>к</strong><strong>о</strong>м и г<strong>о</strong>ленью. Стр<strong>о</strong>ение н<strong>о</strong>г, дых<strong>а</strong>тельн<strong>о</strong>й<br />

<strong>с</strong>и<strong>с</strong>темы и других <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я неиз<strong>в</strong>е<strong>с</strong>тным.<br />

Длин<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 11 мм, ширин<strong>а</strong><br />

ее <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 10 мм. К<strong>а</strong>рб<strong>о</strong>н. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Mazoniidae <strong>с</strong> р<strong>о</strong>д<strong>о</strong>м Mazonia Meek et Worthen,<br />

1868 из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и (ри<strong>с</strong>. 1225,<br />

1227).<br />

НАДСЕМЕЙСТВО SCORPIONIDEA<br />

НАСТОЯЩИЕ СКОРПИОНЫ<br />

Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>с</strong>еми <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Н<strong>о</strong>ги<br />

длинные, <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью члени<strong>к</strong><strong>а</strong>ми р<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>змер<strong>а</strong>.<br />

Педип<strong>а</strong>льпы длинные, <strong>с</strong> пятью члени<strong>к</strong><strong>а</strong>ми<br />

и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>о</strong>й <strong>к</strong>лешней. К<strong>а</strong>рб<strong>о</strong>н —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Buthidae, Scorpionidae, Diplocentridae, Chactidae,<br />

Vejovidae, Bothriuridae, из них д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых<br />

из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong>. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

де<strong>в</strong>ять <strong>в</strong>ымерших <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> из де<strong>в</strong><strong>о</strong>н<strong>а</strong>, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

и три<strong>а</strong><strong>с</strong><strong>а</strong>: Palaeoscorpiidae, Archaeoctonidae,<br />

Centromachidae, Isobuthidae, Cyclophthalmidae,<br />

Eoscorpiidae, Trigonoscorpionidae, Garnettiidae<br />

и Mesophonidae.<br />

СЕМЕЙСТВО PALAEOSCORPIIDAE<br />

LEHMANN, 1944<br />

Члени<strong>к</strong>и мет<strong>а</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong><strong>о</strong> <strong>с</strong>редним пр<strong>о</strong>д<strong>о</strong>льным<br />

ут<strong>о</strong>лщением, или <strong>в</strong><strong>а</strong>ли<strong>к</strong><strong>о</strong>м. Срединные гл<strong>а</strong>з<strong>а</strong><br />

428<br />

http://jurassic.ru/<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong>.<br />

Стр<strong>о</strong>ение <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong>-<strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>.<br />

Один р<strong>о</strong>д Palaeoscorpius Lehmann, 1944<br />

из н. де<strong>в</strong><strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ARCHAEOCTONIDAE<br />

PETRUNKEVITCH, 1949<br />

Члени<strong>к</strong>и мет<strong>а</strong><strong>с</strong><strong>о</strong>мы без <strong>с</strong>рединн<strong>о</strong>г<strong>о</strong> <strong>в</strong><strong>а</strong>ли<strong>к</strong><strong>а</strong>,<br />

у<strong>с</strong>тр<strong>о</strong>ены, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1241). Стернум <strong>к</strong>рупный, прям<strong>о</strong>уг<strong>о</strong>льный,<br />

з<strong>а</strong>ним<strong>а</strong>ет <strong>в</strong><strong>с</strong>ю <strong>с</strong>реднюю ч<strong>а</strong><strong>с</strong>ть груди; т<strong>а</strong>зи<strong>к</strong>и<br />

трех з<strong>а</strong>дних п<strong>а</strong>р н<strong>о</strong>г р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>о</strong>т нег<strong>о</strong>. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I <strong>в</strong>нутренними<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми, н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>бр<strong>а</strong>зуют<strong>с</strong>я м<strong>а</strong><strong>к</strong><strong>с</strong>иллярные<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти, д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong> <strong>с</strong>редней линии тел<strong>а</strong>;<br />

<strong>о</strong>ни р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong>переди <strong>с</strong>тернум<strong>а</strong> (ри<strong>с</strong>.<br />

1226А). П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие лежит п<strong>о</strong>з<strong>а</strong>ди ур<strong>о</strong><strong>в</strong>ня<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV. Н<strong>о</strong>ги <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

<strong>с</strong> шип<strong>а</strong>ми н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>в</strong>ершин<br />

г<strong>о</strong>леней и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong>рупными <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. Педип<strong>а</strong>льпы<br />

п<strong>о</strong>чти <strong>в</strong>д<strong>в</strong><strong>о</strong>е т<strong>о</strong>лще и длиннее х<strong>о</strong>дильных<br />

н<strong>о</strong>г; <strong>к</strong>лешня б<strong>о</strong>льш<strong>а</strong>я, <strong>с</strong> длинными<br />

п<strong>а</strong>льц<strong>а</strong>ми (ри<strong>с</strong>. 1241). Длин<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

9 мм. К<strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Archaeoctonus Pocock,<br />

1911 из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и Eoctonus Petrunkevitch,<br />

1913 из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.<br />

СЕМЕЙСТВО CENTROMACHIDAE<br />

PETRUNKEVITCH, 1953<br />

Ст<strong>в</strong><strong>о</strong>р<strong>к</strong>и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

между <strong>в</strong>нутренними <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г<br />

III и IV. Впереди них лежит треуг<strong>о</strong>льный<br />

<strong>с</strong>тернум, д<strong>о</strong>х<strong>о</strong>дящий д<strong>о</strong> т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г I; п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

ег<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г II (ри<strong>с</strong>.<br />

1226Б). Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими м<strong>а</strong><strong>к</strong><strong>с</strong>иллярными<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м<br />

н<strong>а</strong> <strong>с</strong>редней линии тел<strong>а</strong>. Педип<strong>а</strong>льпы <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими<br />

и т<strong>о</strong>л<strong>с</strong>тыми члени<strong>к</strong><strong>а</strong>ми, п<strong>о</strong><strong>к</strong>рытыми <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>ими<br />

буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми; длин<strong>а</strong> педип<strong>а</strong>льп <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

9 мм; п<strong>а</strong>льпы <strong>к</strong>лешни длинные, т<strong>о</strong>н<strong>к</strong>ие, з<strong>а</strong><strong>о</strong><strong>с</strong>тренные<br />

(ри<strong>с</strong>. 1242). Один р<strong>о</strong>д Centromachus<br />

Thorell, 1885 из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ISOBUTHIDAE<br />

PETRUNKEVITCH, 1913<br />

Ст<strong>в</strong><strong>о</strong>р<strong>к</strong>и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г IV. Впереди них м<strong>а</strong>лень<strong>к</strong>ий<br />

треуг<strong>о</strong>льный <strong>с</strong>тернум, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый р<strong>а</strong>зделяет<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г III. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I и II<br />

<strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м н<strong>а</strong> <strong>с</strong>редней линии<br />

тел<strong>а</strong> (ри<strong>с</strong>. 1226Г). К<strong>а</strong>рб<strong>о</strong>н. Три р<strong>о</strong>д<strong>а</strong>: Isobuthus<br />

Fritsch, 1904 (ри<strong>с</strong>. 1243, 1244), Microlabis<br />

Corda, 1839 (ри<strong>с</strong>. 1245) из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и Palaeobuthus Petrunkevitch, 1913, из<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Амери<strong>к</strong>и.

СЕМЕЙСТВО CYCLOPHTHALMIDAE<br />

PETRUNKEVITCH, 1913<br />

Ст<strong>в</strong><strong>о</strong>р<strong>к</strong>и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

п<strong>о</strong>з<strong>а</strong>ди т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV. Т<strong>а</strong>зи<strong>к</strong>и пер<strong>в</strong>ых и <strong>в</strong>т<strong>о</strong>рых<br />

н<strong>о</strong>г имеют <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянутые м<strong>а</strong><strong>к</strong><strong>с</strong>иллярные<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющие<strong>с</strong>я <strong>в</strong>д<strong>о</strong>ль <strong>с</strong>редней<br />

линии тел<strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> друг другу. Стернум<br />

м<strong>а</strong>лень<strong>к</strong>ий, треуг<strong>о</strong>льный, лежит между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

н<strong>о</strong>г IV (ри<strong>с</strong>. 1226Д). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит<br />

<strong>в</strong> ф<strong>о</strong>рме <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong> прям<strong>о</strong>уг<strong>о</strong>льни<strong>к</strong><strong>а</strong>,<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>уженн<strong>о</strong>г<strong>о</strong> <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти; <strong>с</strong>рединные<br />

гл<strong>а</strong>з<strong>а</strong> н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м р<strong>о</strong>мб<strong>о</strong><strong>в</strong>идн<strong>о</strong>м буг<strong>о</strong>р<strong>к</strong>е<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> передней трети щит<strong>а</strong><br />

(ри<strong>с</strong>. 1233). Один р<strong>о</strong>д Cyclophthalmus Corda,<br />

1835 из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО EOSCORPIIDAE SCUDDER, 1884<br />

Ст<strong>в</strong><strong>о</strong>р<strong>к</strong>и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия <strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong>ие,<br />

леж<strong>а</strong>т п<strong>о</strong>з<strong>а</strong>ди т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV. Стернум<br />

шир<strong>о</strong><strong>к</strong>ий, пятиуг<strong>о</strong>льный, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен между<br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г III и IV, шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong>зд<strong>в</strong>иг<strong>а</strong>я их <strong>в</strong><br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны. М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г<br />

I и II длинные, прямые, пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ют<strong>с</strong>я <strong>в</strong>перед<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> друг другу (ри<strong>с</strong>. 1226Е). Педип<strong>а</strong>льпы<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> т<strong>о</strong>н<strong>к</strong>ие, длинные (ри<strong>с</strong>.<br />

1247). Члени<strong>к</strong>и н<strong>о</strong>г гл<strong>а</strong>д<strong>к</strong>ие, без шил<strong>о</strong><strong>в</strong>идных<br />

р<strong>а</strong>зр<strong>а</strong>щений. Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я и т<strong>о</strong>л<strong>с</strong>т<strong>а</strong>я, не<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> тел<strong>а</strong>. Передний <strong>к</strong>р<strong>а</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

шир<strong>о</strong><strong>к</strong>ий и туп<strong>о</strong>й. К<strong>а</strong>рб<strong>о</strong>н. Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Eoscorpius Meek et Worthen, 1868. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Е. carbonarius Meek et Worthen, 1868;<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Передне- и з<strong>а</strong>днебрюшье<br />

примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й длины. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный<br />

щит <strong>с</strong> б<strong>о</strong>лее или менее прямыми и п<strong>а</strong>р<strong>а</strong>ллельными<br />

друг другу <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми. Гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong><br />

<strong>с</strong>мещен <strong>в</strong> переднюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину щит<strong>а</strong>; мел<strong>к</strong>ие<br />

гл<strong>а</strong>з<strong>а</strong> н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е ег<strong>о</strong> (ри<strong>с</strong>. 1247). Н<strong>о</strong>ги<br />

т<strong>о</strong>н<strong>к</strong>ие, длинные. Педип<strong>а</strong>льпы длинные, т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong> 1,5 р<strong>а</strong>з<strong>а</strong> т<strong>о</strong>лще х<strong>о</strong>дильных н<strong>о</strong>г; <strong>к</strong><strong>о</strong>ленный<br />

члени<strong>к</strong> их примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>ен длине <strong>к</strong>лешни. Длин<strong>а</strong><br />

тел<strong>а</strong> д<strong>о</strong> 130 мм. Четыре <strong>в</strong>ид<strong>а</strong>, из них д<strong>в</strong><strong>а</strong> <strong>в</strong><br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>не Е<strong>в</strong>р<strong>о</strong>пы.<br />

Alloscorpius Petrunkevitch, 1949. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Eoscorpius granulosus Petrunkevitch,<br />

1913; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет предыдущий<br />

р<strong>о</strong>д, н<strong>о</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>лу<strong>к</strong>ругл<strong>о</strong>й<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й, <strong>о</strong>тделяющей переднюю ч<strong>а</strong><strong>с</strong>ть<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> <strong>с</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женным зде<strong>с</strong>ь<br />

<strong>к</strong>рупным груше<strong>в</strong>идным гл<strong>а</strong>зным буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м,<br />

Ри<strong>с</strong>. 1241—1246. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archaeoctonidae, Centromachidae,<br />

Isobuthidae и Cyclophthalmidae<br />

1241. Archaeoctonus glaber (Peach); <strong>о</strong>бщий <strong>в</strong>ид, Xl,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Werner, 1934). 1242. Centromachus euglyptus (Peach);<br />

педип<strong>а</strong>льпы, Хб,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953).<br />

1243. Isobuthus kralupensls Thorell et Lindstrom; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу,<br />

V3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия (Werner, 1934). 1244. Isobuthus holti<br />

Pocock; <strong>о</strong>бщий <strong>в</strong>ид и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>г<strong>о</strong>, X5,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия<br />

(Petrunkevitch, 1953). 1245. Microlabls sternbergii (Corda); <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong>низу, X3,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия (Pocock, 1911). 1246.<br />

Cyclophthalmus senior (Corda); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, Xl,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия (Werner, 1934)<br />

429<br />

http://jurassic.ru/<br />

п<strong>о</strong>чти д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ющим переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong><br />

(ри<strong>с</strong>. 1236). Гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong>лу<strong>с</strong>фериче<strong>с</strong><strong>к</strong>ие. Клешня<br />

р<strong>а</strong><strong>в</strong>н<strong>а</strong> или длиннее <strong>к</strong><strong>о</strong>ленн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> педип<strong>а</strong>льп<br />

(ри<strong>с</strong>. 1248). Длин<strong>а</strong> тел<strong>а</strong> д<strong>о</strong> 130 мм; дли-

1265Б<br />

http://jurassic.ru/

н<strong>а</strong> <strong>к</strong>лешни педип<strong>а</strong>льп 29 мм.. Четыре <strong>в</strong>ид<strong>а</strong>: д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и и д<strong>в</strong><strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

Anthracoscorpius Rust <strong>а</strong>, 1888 (= Lichnoscorpius<br />

Petrunkevitch, 1949). Тип р<strong>о</strong>д<strong>а</strong> — Cyclophthalmus<br />

(Anthracoscorpius) juvenis Kusta,<br />

1885 (= C. senior Kusta, 1884, pars); <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. З<strong>а</strong>днебрюшье <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

т<strong>о</strong>л<strong>с</strong>т<strong>о</strong>е, примерн<strong>о</strong> т<strong>а</strong><strong>к</strong><strong>о</strong>й же длины, <strong>к</strong><strong>а</strong><strong>к</strong> переднебрюшье<br />

(ри<strong>с</strong>. 1249, 1250); г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный<br />

щит <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми и <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тью,<br />

<strong>с</strong>переди шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен, п<strong>о</strong>луди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идный,<br />

длин<strong>а</strong> ег<strong>о</strong> примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> ширине<br />

(ри<strong>с</strong>. 1249); гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong>близи переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я; гребенч<strong>а</strong>тые <strong>о</strong>рг<strong>а</strong>ны длинные<br />

(ри<strong>с</strong>. 1250). Длин<strong>а</strong> тел<strong>а</strong> 9—11 мм; длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong><br />

0,95—1,41 мм, ширин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти<br />

1,50—1,71 мм. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы.<br />

Europhthalmus Petrunkevitch, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Е. longlmanus Petrunkevitch, 1949;<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит эллиптиче<strong>с</strong><strong>к</strong>ий,<br />

у<strong>с</strong>еченный н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце; <strong>с</strong>рединные<br />

<strong>к</strong>рупные гл<strong>а</strong>з<strong>а</strong> леж<strong>а</strong>т <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

щит<strong>а</strong>; <strong>о</strong>ни р<strong>а</strong>зделены неб<strong>о</strong>льшим треуг<strong>о</strong>льным<br />

<strong>в</strong><strong>о</strong>з<strong>в</strong>ышением (ри<strong>с</strong>. 1236); п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть щит<strong>а</strong> <strong>с</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными мел<strong>к</strong>ими буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми; педип<strong>а</strong>льпы<br />

<strong>о</strong>чень длинные; <strong>к</strong><strong>о</strong>нцы п<strong>а</strong>льце<strong>в</strong> при<br />

<strong>о</strong>т<strong>о</strong>гнут<strong>о</strong>й н<strong>а</strong>з<strong>а</strong>д педип<strong>а</strong>льпе д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют третьег<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> з<strong>а</strong>днебрюшья; <strong>к</strong>лешня <strong>с</strong> длинными,<br />

прямыми п<strong>а</strong>льц<strong>а</strong>ми (ри<strong>с</strong>. 1254). Длин<strong>а</strong><br />

тел<strong>а</strong> 60 мм ;длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 10,5 мм; длин<strong>а</strong><br />

<strong>к</strong>лешни педип<strong>а</strong>льп 18,7 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы: Typhloscorpius<br />

Petrunkevitch, 1949 (ри<strong>с</strong>. 1237);<br />

Lichnophthalmus Petrunkevitch, 1949 (ри<strong>с</strong>.<br />

1238); Buthlscorpius Petrunkevitch, 1953<br />

(ри<strong>с</strong>. 1252).<br />

СЕМЕЙСТВО TRIGONOSCORPIONIDAE<br />

W. DUBININ, FAM. NOV.<br />

Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> и р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

Уз—Уг <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> тел<strong>а</strong>; <strong>с</strong>егменты мет<strong>а</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие (ри<strong>с</strong>. 1232, 1251, 1256). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>й<br />

щит <strong>с</strong>ужен <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти, не притуплён<br />

и не р<strong>а</strong><strong>с</strong>ширен <strong>с</strong>переди. Четыре р<strong>о</strong>д<strong>а</strong> из<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

Trigonoscorpio Petrunkevitch, 1913.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Т. americanus Petrunkevitch, 1913;<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>с</strong><strong>о</strong><br />

<strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я <strong>с</strong>переди <strong>к</strong>р<strong>а</strong>ями, треуг<strong>о</strong>льный,<br />

у<strong>с</strong>еченный или з<strong>а</strong><strong>к</strong>ругленный <strong>с</strong>переди, зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины, <strong>с</strong> п<strong>о</strong>перечн<strong>о</strong>й<br />

прям<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й, р<strong>а</strong>зделяющей п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

щит<strong>а</strong> н<strong>а</strong> д<strong>в</strong>е ч<strong>а</strong><strong>с</strong>ти (ри<strong>с</strong>. 1232, 1251); гл<strong>а</strong>з<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> уз<strong>к</strong><strong>о</strong>м треуг<strong>о</strong>льн<strong>о</strong>м п<strong>о</strong>ле<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я щит<strong>а</strong>, з<strong>а</strong>днебрюшье<br />

<strong>о</strong>чень т<strong>о</strong>н<strong>к</strong><strong>о</strong>е, примерн<strong>о</strong> т<strong>а</strong><strong>к</strong><strong>о</strong>й же т<strong>о</strong>лщины, <strong>к</strong><strong>а</strong><strong>к</strong><br />

х<strong>о</strong>дильные н<strong>о</strong>ги (ри<strong>с</strong>. 1251); педип<strong>а</strong>льпы длинные,<br />

<strong>в</strong>д<strong>в</strong><strong>о</strong>е т<strong>о</strong>лще х<strong>о</strong>дильных н<strong>о</strong>г, <strong>к</strong><strong>о</strong>ленный<br />

члени<strong>к</strong> их <strong>к</strong><strong>о</strong>р<strong>о</strong>че длины <strong>к</strong>лешни. Длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong><br />

1,7 мм, ширин<strong>а</strong> ег<strong>о</strong> <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти<br />

3,2 мм. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и<br />

(ри<strong>с</strong>. 1251) и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Compsoscorpius Petrunkevitch, 1949.<br />

Тип р<strong>о</strong>д<strong>а</strong> — С. elegans Petrunkevitch, 1949;<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит п<strong>о</strong>лутреуг<strong>о</strong>льный,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>ругленный н<strong>а</strong> переднем<br />

<strong>к</strong><strong>о</strong>нце, прям<strong>о</strong> <strong>о</strong>брез<strong>а</strong>нный <strong>с</strong>з<strong>а</strong>ди (ри<strong>с</strong>. 1235);<br />

<strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>к</strong>р<strong>а</strong>е<strong>в</strong> пр<strong>о</strong>х<strong>о</strong>дит т<strong>о</strong>н<strong>к</strong><strong>а</strong>я<br />

длинн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>, <strong>о</strong>тделяющ<strong>а</strong>я уз<strong>к</strong>ий лент<strong>о</strong><strong>в</strong>идный<br />

<strong>к</strong>р<strong>а</strong>й; из других трех б<strong>о</strong>р<strong>о</strong>зд д<strong>в</strong>е з<strong>а</strong>дние<br />

пр<strong>о</strong>х<strong>о</strong>дят п<strong>о</strong>пере<strong>к</strong> тел<strong>а</strong>, <strong>а</strong> передняя — п<strong>о</strong>лу<strong>к</strong>ругл<strong>а</strong>я<br />

— <strong>в</strong><strong>о</strong><strong>к</strong>руг <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> гл<strong>а</strong>д<strong>к</strong><strong>о</strong>г<strong>о</strong> бугр<strong>а</strong><br />

и мел<strong>к</strong>их <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1235, 1253);<br />

педип<strong>а</strong>льпы б<strong>о</strong>льшие, <strong>в</strong> п<strong>о</strong>лт<strong>о</strong>р<strong>а</strong> р<strong>а</strong>з<strong>а</strong> т<strong>о</strong>лще<br />

х<strong>о</strong>дильных н<strong>о</strong>г, <strong>в</strong>нутренние <strong>к</strong>р<strong>а</strong>я их члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

бугри<strong>с</strong>тые; <strong>к</strong>лешня уз<strong>к</strong><strong>а</strong>я, <strong>с</strong> длинными, т<strong>о</strong>н<strong>к</strong>ими<br />

п<strong>а</strong>льц<strong>а</strong>ми; длин<strong>а</strong> <strong>к</strong>лешни <strong>в</strong> 1,5—2 р<strong>а</strong>з<strong>а</strong> пре<strong>в</strong>ыш<strong>а</strong>ет<br />

длину <strong>к</strong><strong>о</strong>ленн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

б<strong>о</strong>льше бедр<strong>а</strong>. Длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 2,57 мм. Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Typhlopisthacanthus Petrunkevitch,<br />

1949. Тип р<strong>о</strong>д<strong>а</strong> — Palaeopisthacanthus mazonensis<br />

Petrunkevitch, 1913; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>.<br />

Мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>чень <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я и т<strong>о</strong>н<strong>к</strong><strong>а</strong>я; з<strong>а</strong>гиб<strong>а</strong>я<strong>с</strong>ь<br />

<strong>в</strong>перед, <strong>о</strong>н<strong>а</strong> тель<strong>с</strong><strong>о</strong>н<strong>о</strong>м не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

з<strong>а</strong>дней гр<strong>а</strong>ницы ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> (ри<strong>с</strong>. 1256А),<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит, <strong>в</strong>идим<strong>о</strong>, гл<strong>а</strong>д<strong>к</strong>ий, тр<strong>а</strong>пецие<strong>в</strong>идный,<br />

<strong>в</strong>ытянутый, <strong>с</strong><strong>о</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong>ыпу<strong>к</strong>лыми<br />

б<strong>о</strong><strong>к</strong><strong>а</strong>ми; гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; пецип<strong>а</strong>льпы <strong>о</strong>чень<br />

т<strong>о</strong>л<strong>с</strong>тые и длинные (ри<strong>с</strong>. 1256Б); р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г, <strong>с</strong>тернум<strong>а</strong> и <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н<strong>о</strong><strong>в</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> н<strong>а</strong> ри<strong>с</strong>. 1256Б. Длин<strong>а</strong><br />

тел<strong>а</strong> 23 мм, <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong> 6,6 мм и 6,3 мм <strong>в</strong><br />

ширину н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Кр<strong>о</strong>ме тип<strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>, <strong>о</strong>дин <strong>в</strong>ид из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и Palaeopisthacanthus<br />

Petrunkevitch, 1913 (ри<strong>с</strong>. 1231).<br />

Ри<strong>с</strong>. 1247—1257. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Eoscorpiidae, Trigonoscorpionidae, Garnettiidae<br />

1247 Eoscorpius dunlopi Pocock; <strong>о</strong>бщий <strong>в</strong>ид, Xl,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3 Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Pocock, 1911). 1248. Alloscorpius granulosus (Petrunkevitch);<br />

<strong>о</strong>бщий <strong>в</strong>ид, хЗ/4, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С Амери<strong>к</strong><strong>а</strong> (Petrunkevitch,<br />

1913) 1249. Anthracoscorpius minutus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид,<br />

х 3/4 '<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия (Petrunkevitch, 1953). 1250. Anthracoscorpius<br />

iuvenls Kusta; <strong>о</strong>бщий <strong>в</strong>ид, хЗ/4, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия<br />

(Petrunkevitch, 1953). 1251. Trigonoscorpio americanus<br />

Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, Xl,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch<br />

1913). 1252. Buthlscorpius buthiformis Pocock; <strong>о</strong>бщий <strong>в</strong>ид,<br />

xl <strong>о</strong>' <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Pocock, 1911). 1253. Compsoscorpius<br />

elegans Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>г<strong>о</strong>, х5,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

431<br />

http://jurassic.ru/<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1949). 1254. Europhthalmus longlmanus<br />

Petrunkevitch; А — <strong>о</strong>бщий <strong>в</strong>ид, xl,7; Б — педип<strong>а</strong>льпы<br />

Х2,5; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1940). 1255. Palaeopisthacanthus<br />

schucherti Petrunkevitch; A — <strong>о</strong>бщий <strong>в</strong>ид, X4,0; Б —<br />

<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть; Х4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch,<br />

1913). 1256. Typhlopisthacanthus mazonensts (Petrunkevitch),<br />

A — <strong>о</strong>бщий <strong>в</strong>ид, X3,0; Б — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть,<br />

Х3,0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1257. Garnettius<br />

hungerfordi Elias); A — <strong>о</strong>бщий <strong>в</strong>ид, X2,5; Б — н<strong>о</strong>г<strong>а</strong> III,<br />

X18.0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913)

СЕМЕЙСТВО GARNETTIIDAE W. DUBININ,<br />

FAM. NOV.<br />

К<strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong>-<strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, <strong>к</strong><strong>а</strong><strong>к</strong> у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

Eoscorpiidae и <strong>в</strong><strong>с</strong>ех <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>.<br />

Мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong>,<br />

<strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>му <strong>с</strong>переди причленяет<strong>с</strong>я неб<strong>о</strong>льш<strong>а</strong>я<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>. Тергиты мез<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>с</strong>переди уз<strong>к</strong>ие, п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширяют<strong>с</strong>я <strong>к</strong> з<strong>а</strong>днему <strong>к</strong><strong>о</strong>нцу; тергит<br />

<strong>с</strong>едьм<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> <strong>с</strong><strong>а</strong>мый длинный и уз<strong>к</strong>ий,<br />

п<strong>о</strong>чти <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тный; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

т<strong>о</strong>л<strong>с</strong>т<strong>а</strong>я, тель<strong>с</strong><strong>о</strong>н уз<strong>к</strong>ий, <strong>с</strong> прямым <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м<br />

(ри<strong>с</strong>. 1257); г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит, п<strong>о</strong>чти <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тный<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии, <strong>с</strong>переди имеет треуг<strong>о</strong>льный<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т, <strong>с</strong><strong>а</strong>мый передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>тделен<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й; гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> <strong>о</strong>чень<br />

м<strong>а</strong>лень<strong>к</strong>ий, <strong>о</strong><strong>в</strong><strong>а</strong>льный р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я тел<strong>а</strong> (ри<strong>с</strong>. 1240). Г<strong>о</strong>лень и мет<strong>а</strong>т<strong>а</strong>рз<br />

н<strong>о</strong>г III <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными д<strong>о</strong>рз<strong>а</strong>льными<br />

шил<strong>о</strong><strong>в</strong>идными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми (ри<strong>с</strong>. 1257 А). Н<strong>о</strong>ги<br />

<strong>в</strong><strong>с</strong>ех п<strong>а</strong>р т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Педип<strong>а</strong>льпы<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>с</strong> м<strong>о</strong>щными<br />

<strong>к</strong>лешнями (ри<strong>с</strong>. 1257 А, Б). Один р<strong>о</strong>д Garnettius<br />

Petrunkevitch, 1953 из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.<br />

Ри<strong>с</strong>. 1258—1260. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mesophonidae и Scorpionidae<br />

incertae sedis<br />

1258. Mesophonus sp. (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); A — <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

Xl,5; Б — <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, XI,5; три<strong>а</strong><strong>с</strong>. 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Werner,<br />

1934). 1259. Palacomachus anglicus (Woodward); A — <strong>с</strong>редние <strong>с</strong>егменты<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>мы, Xl,0; Б— мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и п<strong>о</strong><strong>с</strong>ледний <strong>с</strong>егмент мез<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

XI,0; В — педип<strong>а</strong>льпы, Х*>"; Г — педип<strong>а</strong>льпы <strong>с</strong> р<strong>а</strong><strong>с</strong><strong>к</strong>рыт<strong>о</strong>й<br />

<strong>к</strong>лешней, Х1,0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>. 1911). 1260.<br />

Palaeomachus sp.; <strong>к</strong>лешня педип<strong>а</strong>льпы, Х',0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>,<br />

(Petrunkevitch, 1949)<br />

432<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО MESOPHONIDAE WILLS, 1910<br />

Крупные <strong>к</strong>л<strong>а</strong>п<strong>а</strong>ны п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

п<strong>о</strong>з<strong>а</strong>ди ур<strong>о</strong><strong>в</strong>ня т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV.<br />

Стернум <strong>к</strong>рупный, треуг<strong>о</strong>льный или пятиуг<strong>о</strong>льный,<br />

<strong>в</strong>д<strong>а</strong>ет<strong>с</strong>я угл<strong>о</strong>м между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г III<br />

и IV. М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и имеют т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г II прямые, д<strong>о</strong>х<strong>о</strong>дят<br />

д<strong>о</strong> <strong>с</strong>редней линии тел<strong>а</strong>, <strong>в</strong>переди <strong>с</strong>тернум<strong>а</strong><br />

(ри<strong>с</strong>. 1226 В). Сегменты мез<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>мы рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены друг <strong>о</strong>т друг<strong>а</strong>;<br />

<strong>с</strong>егменты з<strong>а</strong>днебрюшья имеют пр<strong>о</strong>д<strong>о</strong>льные<br />

ряды буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1258 А, В). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит прям<strong>о</strong>уг<strong>о</strong>льный,<br />

<strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим <strong>в</strong>перед <strong>с</strong>редним <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м,<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я гл<strong>а</strong>з<strong>а</strong><br />

(ри<strong>с</strong>. 1239). Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные; педип<strong>а</strong>льпы<br />

<strong>к</strong>рупные, <strong>с</strong> м<strong>о</strong>щными <strong>к</strong>лешнями (ри<strong>с</strong>. 1258<br />

А, Б). Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из три<strong>а</strong><strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Mesophonus Wills, 1910. Тип р<strong>о</strong>д<strong>а</strong> — М.<br />

perornatus Wills, 1910; три<strong>а</strong><strong>с</strong>, Англия. Стернум<br />

пятиуг<strong>о</strong>льный; п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть хитин<strong>а</strong> н<strong>а</strong> <strong>с</strong>егмент<strong>а</strong>х<br />

гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или <strong>с</strong> р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й т<strong>о</strong>чечн<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й;<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>в</strong> ф<strong>о</strong>рме п<strong>о</strong>перечн<strong>о</strong>г<strong>о</strong><br />

прям<strong>о</strong>уг<strong>о</strong>льни<strong>к</strong><strong>а</strong>, <strong>с</strong> <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong> <strong>с</strong>рез<strong>а</strong>нными<br />

передними угл<strong>а</strong>ми и <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющей <strong>с</strong>редней<br />

ч<strong>а</strong><strong>с</strong>тью, н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я гл<strong>а</strong>з<strong>а</strong>; з<strong>а</strong>дний<br />

и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые <strong>к</strong>р<strong>а</strong>я щит<strong>а</strong> <strong>о</strong>гр<strong>а</strong>ничены б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й<br />

(ри<strong>с</strong>. 1239). Ше<strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong> из три<strong>а</strong><strong>с</strong><strong>а</strong> Англии.<br />

Spongiotarsus Wills, 1947. Тип р<strong>о</strong>д<strong>а</strong> —<br />

S. pustulosus Wills, 1947; три<strong>а</strong><strong>с</strong>, Англия. От предыдущег<strong>о</strong><br />

р<strong>о</strong>д<strong>а</strong> <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я треуг<strong>о</strong>льным <strong>с</strong>тернум<strong>о</strong>м<br />

и <strong>с</strong>етч<strong>а</strong>тым <strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

хитин<strong>а</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong> (ри<strong>с</strong>. 1226 В). Один <strong>в</strong>ид<br />

из три<strong>а</strong><strong>с</strong><strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО BUTHIDAE<br />

С. L. КОСН, 1837<br />

Стернум треуг<strong>о</strong>льный. Д<strong>в</strong><strong>а</strong> шип<strong>а</strong> н<strong>а</strong> меж<strong>с</strong>егмент<strong>а</strong>льн<strong>о</strong>й<br />

переп<strong>о</strong>н<strong>к</strong>е между мет<strong>а</strong>т<strong>а</strong>рз<strong>о</strong>м<br />

и л<strong>а</strong>п<strong>к</strong><strong>о</strong>й. Три или пять п<strong>а</strong>р б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых гл<strong>а</strong>з. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 34 р<strong>о</strong>д<strong>а</strong>,<br />

из них Tityus С. L. Koch, 1837 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО SCORPIONIDAE LEACH, 1815<br />

Стернум пятиуг<strong>о</strong>льный. Един<strong>с</strong>т<strong>в</strong>енный шип<br />

н<strong>а</strong> меж<strong>с</strong>егментн<strong>о</strong>й переп<strong>о</strong>н<strong>к</strong>е между мет<strong>а</strong>т<strong>а</strong>рз<strong>о</strong>м<br />

и л<strong>а</strong>п<strong>к</strong><strong>о</strong>й I и II н<strong>о</strong>г; III и IV н<strong>о</strong>ги без эт<strong>о</strong>г<strong>о</strong><br />

шип<strong>а</strong>. Три п<strong>а</strong>ры б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых гл<strong>а</strong>з. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из<br />

них Scorpio Linnaeus, 1758 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) и ми<strong>о</strong>цен<strong>а</strong> Герм<strong>а</strong>нии.

SC0RP10NIDA INCERTAE SEDIS<br />

Palaeomachus Pocock, 1911. Тип р<strong>о</strong>д<strong>а</strong>—<br />

Eoscorpius angticus Woodward, 1876, nec Petrunkevitch,<br />

1949; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия. Опи<strong>с</strong><strong>а</strong>ны д<strong>в</strong>е<br />

педип<strong>а</strong>льпы (ри<strong>с</strong>. 1259, 1260) и д<strong>в</strong><strong>а</strong> фр<strong>а</strong>гмент<strong>а</strong><br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>мы (ри<strong>с</strong>. 1259 А, Б).<br />

Liassoscorpionides Bode, 1951. Тип р<strong>о</strong>д<strong>а</strong> —•<br />

L. schmidti Bode, 1951; н. юр<strong>а</strong>, Герм<strong>а</strong>ния.<br />

Ве<strong>с</strong>ьм<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тн<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>нный <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н пл<strong>о</strong>х<strong>о</strong>й<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нн<strong>о</strong><strong>с</strong>ти. Длин<strong>а</strong> тел<strong>а</strong> 14,4 мм, т<strong>о</strong>лщин<strong>а</strong><br />

4,8 мм. Один <strong>в</strong>ид. Юр<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ПОДКЛАСС PEDIPALPIDES. ЖГУТОНОГИЕ<br />

Тел<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е или <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>о</strong>тдел<strong>а</strong>: пр<strong>о</strong><strong>с</strong><strong>о</strong>му, п<strong>о</strong><strong>к</strong>рытую<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит<strong>о</strong>м, и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му <strong>с</strong><br />

я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женн<strong>о</strong>й <strong>с</strong>егмент<strong>а</strong>цией. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

не п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> <strong>к</strong>рупные переднюю шир<strong>о</strong><strong>к</strong>ую<br />

и з<strong>а</strong>днюю уз<strong>к</strong>ую ч<strong>а</strong><strong>с</strong>ти, <strong>к</strong><strong>а</strong><strong>к</strong> у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Holactinochitinosi. З<strong>а</strong>дняя<br />

ч<strong>а</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы рез<strong>к</strong><strong>о</strong> <strong>с</strong>ужен<strong>а</strong>, <strong>а</strong> три ее п<strong>о</strong><strong>с</strong>ледних<br />

<strong>с</strong>егмент<strong>а</strong> (XVI—XVIII) <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую<br />

и т<strong>о</strong>н<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му, <strong>о</strong><strong>к</strong><strong>а</strong>нчи<strong>в</strong><strong>а</strong>ющую<strong>с</strong>я<br />

у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> длинн<strong>о</strong>й, члени<strong>с</strong>т<strong>о</strong>й х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

нитью (ри<strong>с</strong>. 1262, 1263, 1265). Лишь у<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>з<strong>а</strong>ди<br />

туп<strong>а</strong>я, без <strong>с</strong>лед<strong>о</strong><strong>в</strong> мет<strong>а</strong><strong>с</strong><strong>о</strong>мы. Педип<strong>а</strong>льпы <strong>о</strong>чень<br />

<strong>к</strong>рупные (не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые Uropygi), <strong>к</strong>лешне<strong>в</strong>идные<br />

(Pseudoscorpionodea), х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные (Amblypygi,<br />

Uropygi) или н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные (Palpigradi,<br />

Ricinulei). Же<strong>в</strong><strong>а</strong>тельные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и имеют<strong>с</strong>я<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х педип<strong>а</strong>льп и н<strong>о</strong>г I у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1263). Х<strong>о</strong>дильные н<strong>о</strong>ги<br />

т<strong>о</strong>н<strong>к</strong>ие и длинные, <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> ут<strong>о</strong>лщенными<br />

бедр<strong>а</strong>ми и т<strong>о</strong>н<strong>к</strong>ими, длинными ди<strong>с</strong>т<strong>а</strong>льными<br />

члени<strong>к</strong><strong>а</strong>ми; л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 1262, 1270). Н<strong>о</strong>ги I <strong>о</strong>бычн<strong>о</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> б<strong>о</strong>льш<strong>о</strong>й длины, <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я биче<strong>в</strong>идными;<br />

<strong>о</strong>ни не <strong>в</strong>ып<strong>о</strong>лняют л<strong>о</strong><strong>к</strong><strong>о</strong>м<strong>о</strong>т<strong>о</strong>рн<strong>о</strong>й фун<strong>к</strong>ции<br />

и я<strong>в</strong>ляют<strong>с</strong>я <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми <strong>о</strong><strong>с</strong>яз<strong>а</strong>ния (ри<strong>с</strong>. 1262,<br />

1270). Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> и рицинулей<br />

н<strong>о</strong>ги I п<strong>о</strong>чти не <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т других.<br />

Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не мез<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>ид<strong>о</strong>измененные<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены <strong>в</strong> <strong>в</strong>иде <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>о</strong><strong>к</strong><br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия н<strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>м брюшн<strong>о</strong>м<br />

<strong>с</strong>терните (ри<strong>с</strong>. 1264) и д<strong>в</strong>ух п<strong>а</strong>р лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong><br />

или <strong>с</strong>тигм (Pseudoscorpionodea) н<strong>а</strong> III и<br />

IV <strong>с</strong>егмент<strong>а</strong>х <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Стр<strong>о</strong>ение <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ти <strong>в</strong> цел<strong>о</strong>м <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<br />

«<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>ый» тип, <strong>с</strong> т<strong>о</strong>й лишь р<strong>а</strong>зницей, чт<strong>о</strong><br />

у Amblypygi и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у Kustarachnida т<strong>а</strong>зи<strong>к</strong>и<br />

<strong>в</strong><strong>с</strong>ех пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> и <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я <strong>в</strong>ершин<strong>а</strong>ми <strong>в</strong> центре<br />

груди (ри<strong>с</strong>. 1266 Б, 1271, 1273). Среди не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

Pedipalpides, <strong>в</strong>пер<strong>в</strong>ые у хелицер<strong>о</strong><strong>в</strong>ых,<br />

н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я р<strong>а</strong>з<strong>в</strong>итие н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> грудн<strong>о</strong>г<strong>о</strong><br />

(\ В. Б. Дубинин \)<br />

28 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 433<br />

http://jurassic.ru/<br />

щит<strong>а</strong> — <strong>с</strong>тернум<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я з<strong>а</strong><br />

<strong>с</strong>чет <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>а</strong>ции брюшных п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> груди,<br />

<strong>о</strong>х<strong>в</strong><strong>а</strong>ты<strong>в</strong><strong>а</strong>ющих <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>в</strong><strong>с</strong>ех <strong>с</strong>егмент<strong>о</strong><strong>в</strong> х<strong>о</strong>дильных<br />

н<strong>о</strong>г. У Uropygi и Amblypygi н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

р<strong>а</strong>з<strong>в</strong>итие м<strong>а</strong>лень<strong>к</strong>их р<strong>о</strong>мб<strong>о</strong><strong>в</strong>идных уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>о</strong><strong>в</strong><br />

упл<strong>о</strong>тненн<strong>о</strong>г<strong>о</strong> хитин<strong>а</strong> <strong>в</strong>д<strong>о</strong>ль <strong>с</strong>редней линии<br />

тел<strong>а</strong>, между <strong>в</strong>нутренними <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>о</strong>г. Интере<strong>с</strong>н<strong>о</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>тдельных грудных<br />

<strong>с</strong>тернит<strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>в</strong><strong>с</strong>ех <strong>с</strong>егмент<strong>а</strong>х пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы у и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>г<strong>о</strong> Sternarthron zitteli<br />

Haase (ри<strong>с</strong>. 1276). П<strong>о</strong>д<strong>о</strong>бные <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельные<br />

<strong>с</strong>терниты е<strong>с</strong>ть н<strong>а</strong> <strong>с</strong>егмент<strong>а</strong>х хелицер, трех з<strong>а</strong>дних<br />

п<strong>а</strong>р н<strong>о</strong>г у <strong>с</strong><strong>о</strong><strong>в</strong>ременных п<strong>а</strong>льпигр<strong>а</strong>д (К<strong>о</strong>еneniidae):<br />

<strong>с</strong>терниты педип<strong>а</strong>льп и н<strong>о</strong>г I у них<br />

<strong>с</strong>литы <strong>в</strong> <strong>о</strong>бщую пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

имеет мел<strong>к</strong>ие р<strong>а</strong>змеры: длин<strong>а</strong> тел<strong>а</strong> <strong>в</strong><strong>а</strong>рьирует<br />

<strong>о</strong>т 0,8—2,8 мм (Palpigradi) д<strong>о</strong> 75 мм (Uro-<br />

Pygi)-<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Н<strong>а</strong>земные<br />

хелицер<strong>о</strong><strong>в</strong>ые р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н<strong>а</strong>х;<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны м<strong>о</strong>гут д<strong>а</strong>ле<strong>к</strong><strong>о</strong><br />

пр<strong>о</strong>ни<strong>к</strong><strong>а</strong>ть н<strong>а</strong> <strong>с</strong>е<strong>в</strong>ер, где <strong>о</strong>ни <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> гнезд<strong>а</strong>х<br />

и н<strong>о</strong>р<strong>а</strong>х жи<strong>в</strong><strong>о</strong>тных и <strong>в</strong> жилище чел<strong>о</strong><strong>в</strong>е<strong>к</strong><strong>а</strong>. Мн<strong>о</strong>гие<br />

<strong>в</strong>иды жи<strong>в</strong>ут <strong>в</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тных <strong>с</strong>л<strong>о</strong>ях п<strong>о</strong>ч<strong>в</strong>ы,<br />

<strong>в</strong> <strong>с</strong><strong>к</strong><strong>о</strong>плениях <strong>в</strong>л<strong>а</strong>жных р<strong>а</strong><strong>с</strong>тительных <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>о</strong><strong>в</strong><br />

и т. п. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> — хищни<strong>к</strong>и.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Н<strong>а</strong>д<strong>о</strong>тряд<br />

Pedipalpides я<strong>в</strong>ляет<strong>с</strong>я группир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

хелицер<strong>о</strong><strong>в</strong>ых, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дящих<br />

<strong>о</strong>т <strong>о</strong>бщих <strong>с</strong><strong>о</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong>ми пред<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

О был<strong>о</strong>м <strong>в</strong><strong>о</strong>дн<strong>о</strong>м <strong>о</strong>бр<strong>а</strong>зе жизни их пред<strong>к</strong><strong>о</strong><strong>в</strong><br />

г<strong>о</strong><strong>в</strong><strong>о</strong>рит н<strong>а</strong>личие у Pedipalpides лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

пр<strong>о</strong>из<strong>в</strong><strong>о</strong>дных пер<strong>в</strong>ичных ж<strong>а</strong>берных н<strong>о</strong>же<strong>к</strong><br />

<strong>в</strong><strong>о</strong>дных Chelicerata. У других, б<strong>о</strong>лее<br />

<strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нных Pedipalpides, н<strong>а</strong>пример,<br />

у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Pseudoscorpionodea,<br />

н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я реду<strong>к</strong>ция лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> и<br />

р<strong>а</strong>з<strong>в</strong>итие н<strong>а</strong> их <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>е тр<strong>а</strong>хей. К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Ше<strong>с</strong>ть <strong>о</strong>тряд<strong>о</strong><strong>в</strong>: Uropygi, Amblypygi, Kustarachnida<br />

(<strong>в</strong>ымершие), Palpigradi, Pseudoscorpionodea,<br />

Ricinulei.

Ри<strong>с</strong>. 1261—1265. Отряд Uropygi<br />

1261. Schizomus tenutcaudatus (Cambridge); <strong>о</strong>бщий <strong>в</strong>ид, x 5,0,<br />

р<strong>о</strong>ны, х 2,5 (Pocock, 1911); <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. 1264. Prothelg.<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Roewer, 1934). 1262. Calcitro fisheri Petrunkevich;<br />

phonus giganteus (Petrunkevitch); <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, хЗ.О, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,.<br />

<strong>о</strong>бщий <strong>в</strong>ид, x? не<strong>о</strong>ген, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1955). 1263.'<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1265. Thelyphonus caudatul<br />

Prothelyphonus brttannlcus (Pocock); А — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>­ Linnaeus; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>, <strong>о</strong>бщий <strong>в</strong>ид, х2,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Millot, 1949)<br />

р<strong>о</strong>ны, Х3,5 (Petrunkevitch, 1949); Б — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>-<br />

ОТРЯД UROPYGI. ЖГУТОНОГИЕ СКОРПИОНЫ<br />

(Thelyphonida)<br />

Тел<strong>о</strong> удлиненн<strong>о</strong>е, <strong>с</strong> уз<strong>к</strong><strong>о</strong>й, <strong>в</strong>ытянут<strong>о</strong>й пр<strong>о</strong>- му, <strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й при<strong>к</strong>реплен<strong>а</strong> длинн<strong>а</strong>я х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я<br />

<strong>с</strong><strong>о</strong>м<strong>о</strong>й и <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>й нить (ри<strong>с</strong>. 1263 Б, 1265). Н<strong>о</strong>ги длинные, т<strong>о</strong>нн<strong>а</strong><br />

12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; три з<strong>а</strong>дних <strong>с</strong>егмент<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>к</strong>ие; н<strong>о</strong>ги I <strong>с</strong> <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ими, <strong>в</strong>ытянутыми чле<strong>с</strong>ужены<br />

и <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую, т<strong>о</strong>н<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>- ни<strong>к</strong><strong>а</strong>ми <strong>о</strong>бр<strong>а</strong>зуют длинные ните<strong>в</strong>идные <strong>о</strong><strong>с</strong>яз<strong>а</strong>-<br />

434<br />

http://jurassic.ru/

тельные <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, пр<strong>о</strong>тянутые <strong>в</strong>перед<br />

ОРИС. 1265). О<strong>с</strong>т<strong>а</strong>льные три п<strong>а</strong>ры х<strong>о</strong>дильных<br />

н<strong>о</strong>г <strong>с</strong> ут<strong>о</strong>лщенными бедр<strong>а</strong>ми, <strong>в</strong><strong>о</strong><strong>с</strong>ьмичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>е<strong>в</strong><br />

грудн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тдел<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. Между ними у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> бы<strong>в</strong><strong>а</strong>ют р<strong>а</strong>з<strong>в</strong>иты <strong>о</strong>тдельные мел<strong>к</strong>ие<br />

<strong>с</strong>терн<strong>а</strong>льные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и, не <strong>о</strong>т<strong>о</strong>д<strong>в</strong>иг<strong>а</strong>ющие<br />

т<strong>а</strong>зи<strong>к</strong>и <strong>к</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди. Хелицеры <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>к</strong>лешне<strong>в</strong>идные, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Педип<strong>а</strong>льпы<br />

<strong>о</strong>чень <strong>к</strong>рупные, т<strong>о</strong>л<strong>с</strong>тые, <strong>о</strong>бр<strong>а</strong>зуют <strong>в</strong>переди<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы <strong>к</strong>лещи (ри<strong>с</strong>. 1264). Н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х<br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> и третьег<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

д<strong>в</strong>е п<strong>а</strong>ры <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>а</strong> <strong>в</strong>переди них, <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>,—<strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong>и<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия. Н<strong>а</strong> переднем<br />

<strong>к</strong>р<strong>а</strong>е г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> — д<strong>в</strong><strong>а</strong> <strong>к</strong>рупных гл<strong>а</strong>-<br />

Ри<strong>с</strong>. 1266—1270.<br />

Х266. Protophrynus carbonartus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид:<br />

А — <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х3,0; Б — <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х 3<br />

>";<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1267. Thelyphrynus<br />

tlongatus Petrunkevitch; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

13,0; Б — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, ХЗ.О; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunke-<br />

з<strong>а</strong>, прип<strong>о</strong>днятых н<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>м буг<strong>о</strong>р<strong>к</strong>е, <strong>а</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых угл<strong>о</strong><strong>в</strong> — п<strong>о</strong> три м<strong>а</strong>лень<strong>к</strong>их<br />

гл<strong>а</strong>з<strong>к</strong><strong>а</strong>. Р<strong>а</strong>змеры тел<strong>а</strong> <strong>о</strong>т 3 д<strong>о</strong> 75 мм.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Жгут<strong>о</strong>н<strong>о</strong>гие<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны, или телиф<strong>о</strong>ны, <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong><strong>о</strong><br />

<strong>в</strong>л<strong>а</strong>жных ле<strong>с</strong><strong>а</strong>х тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тей Амери<strong>к</strong>и, Афри<strong>к</strong>и и юг<strong>о</strong>-<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>чн<strong>о</strong>й<br />

Азии. В г<strong>о</strong>ры п<strong>о</strong>дним<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong>ту д<strong>о</strong><br />

1200 м н<strong>а</strong>д ур<strong>о</strong><strong>в</strong>нем м<strong>о</strong>ря. К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 100 <strong>в</strong>ид<strong>о</strong><strong>в</strong>.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Отряд<br />

Uropygi близ<strong>о</strong><strong>к</strong> <strong>к</strong> <strong>о</strong>тряду Amblypygi (<strong>с</strong>м.<br />

<strong>с</strong>тр. 436), <strong>с</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым ег<strong>о</strong> ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бъединяют<br />

<strong>в</strong> <strong>о</strong>бщий <strong>о</strong>тряд Pedipalpi. Одн<strong>а</strong><strong>к</strong><strong>о</strong> Милл<strong>о</strong> (Millot,<br />

1942, 1949) п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>л не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>ть р<strong>а</strong>зде-<br />

Отряд Amblypygi<br />

vitch, 1913). 1268. Graeophonus anglicus Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий<strong>в</strong>ид <strong>с</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х2,5; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911). 1269.<br />

Graeophonus anglicus Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

Х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911).' 1270. Stygophrynus<br />

dammermani Roewer; <strong>о</strong>бщий <strong>в</strong>ид, Х3,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Werner, 1935)<br />

435 28*<br />

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ления телиф<strong>о</strong>н<strong>о</strong><strong>в</strong> и жгут<strong>о</strong>н<strong>о</strong>гих п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельные<br />

<strong>о</strong>тряды. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Schizopeltidia<br />

и Holopeltidia.<br />

ПОДОТРЯД SCHIZOPELTIDIA<br />

(Tartarides, Schizomida)<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит р<strong>а</strong><strong>с</strong>членен н<strong>а</strong> <strong>к</strong>рупный<br />

пр<strong>о</strong>пельтидий и д<strong>в</strong><strong>а</strong> м<strong>а</strong>лень<strong>к</strong>их тергит<strong>а</strong>, леж<strong>а</strong>щих<br />

п<strong>о</strong>з<strong>а</strong>ди нег<strong>о</strong>. Х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я нить т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> <strong>о</strong>дним<br />

или тремя чдени<strong>к</strong><strong>а</strong>ми. Педип<strong>а</strong>льпы т<strong>о</strong>н<strong>к</strong>ие,<br />

н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные; члени<strong>к</strong>и их без шип<strong>о</strong><strong>в</strong> н<strong>а</strong><br />

<strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не. Бедр<strong>а</strong> з<strong>а</strong>дних н<strong>о</strong>г б<strong>о</strong>ч<strong>о</strong>и<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong><br />

<strong>в</strong>здуты; н<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льных н<strong>о</strong>г<strong>а</strong>х бедр<strong>а</strong><br />

т<strong>о</strong>н<strong>к</strong>ие (ри<strong>с</strong>. 1275). Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>т 2,5 д<strong>о</strong> 6 мм.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>е<br />

Schizomidae и третичн<strong>о</strong>е Calcitronidae.<br />

ПОДОТРЯД HOLOPELTIDIA<br />

(Oxypoel, Urotricha, Thelyphonida)<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит не р<strong>а</strong><strong>с</strong>членен. Х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я<br />

нить длинн<strong>а</strong>я, мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я. Педип<strong>а</strong>льпы<br />

<strong>о</strong>чень т<strong>о</strong>л<strong>с</strong>тые; члени<strong>к</strong>и их <strong>с</strong> <strong>к</strong>рупными<br />

зуб<strong>о</strong><strong>в</strong>идными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

(ри<strong>с</strong>. 1265), реже без них, н<strong>о</strong> т<strong>о</strong>гд<strong>а</strong> педи­<br />

п<strong>а</strong>льпы <strong>к</strong><strong>о</strong>р<strong>о</strong>че <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> (ри<strong>с</strong>. 1264). Н<strong>о</strong>ги<br />

т<strong>о</strong>н<strong>к</strong>ие и длинные, бедр<strong>а</strong> <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> ут<strong>о</strong>лщенные, н<strong>о</strong> не <strong>в</strong>здутые, <strong>к</strong><strong>а</strong><strong>к</strong> у<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> предыдущег<strong>о</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>. Б<strong>о</strong>лее <strong>к</strong>рупные<br />

<strong>в</strong>иды: длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>т 18 д<strong>о</strong> 75 мм. К<strong>а</strong>рб<strong>о</strong>н —<br />

ныне. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО THELYPHONIDAE LUCAS, 1882<br />

(Geralinuridae Scudder, 1884)<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я призн<strong>а</strong><strong>к</strong><strong>а</strong>ми п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне. Де<strong>с</strong>ять <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong> и<br />

д<strong>в</strong><strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong>.<br />

Prothelyphonus Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Thetyphonus bohemicus Kusta, 1884); <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>в</strong>ытянутый,<br />

ше<strong>с</strong>тиуг<strong>о</strong>льный. Гл<strong>а</strong>з<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты. П<strong>о</strong><br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м гл<strong>а</strong>з <strong>о</strong>т переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я тел<strong>а</strong> н<strong>а</strong>з<strong>а</strong>д<br />

пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ют<strong>с</strong>я д<strong>в</strong>е <strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнутые б<strong>о</strong>р<strong>о</strong>зды, н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong>х <strong>к</strong><strong>о</strong>т<strong>о</strong>рых неб<strong>о</strong>льшие <strong>о</strong><strong>в</strong><strong>а</strong>льные ут<strong>о</strong>лщения.<br />

Длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 10 мм, ширин<strong>а</strong> 6 мм,<br />

длин<strong>а</strong> брюш<strong>к</strong><strong>а</strong> 20 мм (ри<strong>с</strong>. 1263, 1264). Д<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>а</strong> из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы и <strong>о</strong>дин <strong>в</strong>ид из <strong>с</strong>р.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, р<strong>о</strong>д Gerallnura Scudder, 1884<br />

из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и и <strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д<br />

Thelyphonus Latreille, 1802 из не<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ОТРЯД AMBLYPYGI. ЖГУТОНОГИЕ ПАУКИ<br />

Тел<strong>о</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong>, <strong>с</strong> п<strong>о</strong>чти ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и неб<strong>о</strong>льшим <strong>о</strong><strong>в</strong><strong>а</strong>льным<br />

брюш<strong>к</strong><strong>о</strong>м, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящим из 12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; п<strong>о</strong><strong>с</strong>ледние<br />

<strong>с</strong>егменты не <strong>о</strong>бр<strong>а</strong>зуют уз<strong>к</strong><strong>о</strong>г<strong>о</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

прид<strong>а</strong>т<strong>к</strong><strong>а</strong>, т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце тел<strong>а</strong> имеет<strong>с</strong>я <strong>о</strong>дин м<strong>а</strong>лень<strong>к</strong>ий <strong>с</strong>егмент<br />

(ри<strong>с</strong>. 1270). П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

углублениями, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющими<strong>с</strong>я<br />

р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong>: ее передняя ч<strong>а</strong><strong>с</strong>ть не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>з<strong>в</strong>ышен<strong>а</strong>;<br />

н<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>м переднем <strong>к</strong>р<strong>а</strong>е р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

д<strong>в</strong><strong>а</strong> <strong>к</strong>рупных <strong>с</strong>рединных гл<strong>а</strong>з<strong>а</strong>, <strong>а</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

<strong>о</strong>т них д<strong>в</strong>е группы гл<strong>а</strong>з<strong>к</strong><strong>о</strong><strong>в</strong>, п<strong>о</strong> три <strong>в</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>й<br />

(ри<strong>с</strong>. 1270). Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены т<strong>а</strong>зи<strong>к</strong>и<br />

х<strong>о</strong>дильных н<strong>о</strong>г и педип<strong>а</strong>льп. Хелицеры <strong>к</strong>рупные,<br />

<strong>к</strong>лешне<strong>в</strong>идные, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Педип<strong>а</strong>льпы<br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>й <strong>в</strong>еличины —<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> 4 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше длины тел<strong>а</strong> (Phrynichus<br />

Karsch) — и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>к</strong><strong>о</strong>т<strong>о</strong>рых мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

т<strong>о</strong>н<strong>к</strong>ие игл<strong>о</strong><strong>в</strong>идные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты (ри<strong>с</strong>. 1268, 1270).<br />

Н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong> длинными бедр<strong>а</strong>ми и<br />

г<strong>о</strong>ленями. Члени<strong>к</strong>и н<strong>о</strong>г I <strong>о</strong>чень удлиненные,<br />

(Phrynida, Phrynichida)<br />

436<br />

http://jurassic.ru/<br />

п<strong>а</strong>л<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идные, <strong>с</strong> длинн<strong>о</strong>й, т<strong>о</strong>н<strong>к</strong><strong>о</strong>й, биче<strong>в</strong>идн<strong>о</strong>й,<br />

мн<strong>о</strong>г<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>о</strong><strong>с</strong>яз<strong>а</strong>тельн<strong>о</strong>й л<strong>а</strong>п<strong>к</strong><strong>о</strong>й<br />

(ри<strong>с</strong>. 1270). Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

и мяг<strong>к</strong><strong>о</strong>й, п<strong>о</strong>душ<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й, пуль<strong>в</strong>илл<strong>о</strong>й <strong>в</strong> их <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии.<br />

Орг<strong>а</strong>ны дых<strong>а</strong>ния пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены д<strong>в</strong>умя<br />

п<strong>а</strong>р<strong>а</strong>ми лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я<br />

щеле<strong>в</strong>идными дых<strong>а</strong>льц<strong>а</strong>ми н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х II и<br />

III <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>т 7 д<strong>о</strong><br />

35 мм.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Обит<strong>а</strong>ют<br />

<strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жных ле<strong>с</strong><strong>а</strong>х тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>тр<strong>а</strong>н Амери<strong>к</strong>и, Афри<strong>к</strong>и и юг<strong>о</strong>-<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>к</strong><strong>а</strong><br />

Азии. К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Отряд<br />

Amblypygi близ<strong>о</strong><strong>к</strong> <strong>к</strong> <strong>о</strong>тряду Uropygi, <strong>с</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым<br />

ег<strong>о</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>бъединяют <strong>в</strong> <strong>с</strong>б<strong>о</strong>рный «<strong>о</strong>тряд»<br />

Pedipalpi. Н<strong>а</strong>ряду <strong>с</strong> б<strong>о</strong>льшим г<strong>а</strong>биту<strong>а</strong>льным<br />

<strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong>м этих д<strong>в</strong>ух групп, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong> имеющих<br />

<strong>о</strong>бщее пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение, Amblypygi х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т Uropygi принципи<strong>а</strong>льн<strong>о</strong> иным<br />

<strong>с</strong>тр<strong>о</strong>ением <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong>-<strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти, утр<strong>а</strong>т<strong>о</strong>й<br />

тенденции <strong>к</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нию «<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>идн<strong>о</strong>й»

уз<strong>к</strong><strong>о</strong>й мет<strong>а</strong><strong>с</strong><strong>о</strong>мы, <strong>а</strong> тем б<strong>о</strong>лее х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й нити<br />

и т. п. И<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния Милл<strong>о</strong> (Millot, 1942,<br />

1949) п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ли, чт<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енные р<strong>а</strong>зличия<br />

между этими д<strong>в</strong>умя групп<strong>а</strong>ми имеют<strong>с</strong>я <strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й, <strong>к</strong>р<strong>о</strong><strong>в</strong>ен<strong>о</strong><strong>с</strong>н<strong>о</strong>й, нер<strong>в</strong>н<strong>о</strong>й и пище<strong>в</strong><strong>а</strong>рительн<strong>о</strong>й<br />

<strong>с</strong>и<strong>с</strong>тем, <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льных и <strong>а</strong>н<strong>а</strong>льных желез<br />

и т. п. Интере<strong>с</strong>н<strong>о</strong>, чт<strong>о</strong> н<strong>а</strong>дгл<strong>о</strong>т<strong>о</strong>чн<strong>а</strong>я<br />

нер<strong>в</strong>н<strong>а</strong>я м<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> (пр<strong>о</strong>цеф<strong>а</strong>л<strong>о</strong>н) <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Amblypygi имеет <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong> т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й у телиф<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

из <strong>о</strong>тряд<strong>а</strong> Uropygi, т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> п<strong>о</strong>дгл<strong>о</strong>т<strong>о</strong>чн<strong>о</strong>е<br />

<strong>с</strong><strong>к</strong><strong>о</strong>пление г<strong>а</strong>нглие<strong>в</strong> м<strong>о</strong>зг<strong>а</strong> н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет <strong>с</strong>тр<strong>о</strong>ение<br />

члени<strong>с</strong>тых п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Liphistiomorphae<br />

(<strong>с</strong>м. <strong>с</strong>тр. 494). Близ<strong>о</strong><strong>с</strong>ть Amblypygi <strong>к</strong><br />

примити<strong>в</strong>ным Аг<strong>а</strong>пе<strong>а</strong>е п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я т<strong>а</strong><strong>к</strong>же<br />

<strong>с</strong>тр<strong>о</strong>ением <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong>-<strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти и других<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>. След<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, <strong>о</strong>тряды Amblepygi<br />

и Uropygi <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>уют нез<strong>а</strong><strong>в</strong>и<strong>с</strong>им<strong>о</strong> <strong>о</strong>дин <strong>о</strong>т<br />

ОТРЯД KUST;<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и<br />

б<strong>о</strong>лее уз<strong>к</strong>им яйце<strong>в</strong>идным или <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идным<br />

брюш<strong>к</strong><strong>о</strong>м. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идный<br />

(ри<strong>с</strong>. 1273 А) или п<strong>о</strong>лу<strong>к</strong>руглый (ри<strong>с</strong>. 1272).<br />

В центре ег<strong>о</strong> или немн<strong>о</strong>г<strong>о</strong> ближе <strong>к</strong> переднему<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>му <strong>к</strong>р<strong>а</strong>ю р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен м<strong>а</strong>лень<strong>к</strong>ий<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льный буг<strong>о</strong>р<strong>о</strong><strong>к</strong> <strong>с</strong> д<strong>в</strong>умя гл<strong>а</strong>з<strong>а</strong>ми;<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые гл<strong>а</strong>з<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют (ри<strong>с</strong>. 1272,<br />

1273 А). Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянутых треуг<strong>о</strong>льни<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

пл<strong>о</strong>тн<strong>о</strong> приж<strong>а</strong>тых друг <strong>к</strong> другу,<br />

<strong>с</strong>х<strong>о</strong>дящих<strong>с</strong>я р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> <strong>к</strong> центру брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

груди; т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп <strong>с</strong>литы друг <strong>с</strong><br />

друг<strong>о</strong>м и <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>дну треуг<strong>о</strong>льную пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у<br />

(ри<strong>с</strong>. 1271, 1273 Б). Педип<strong>а</strong>льпы <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

четырех-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>лешне<strong>в</strong>идные. Н<strong>о</strong>ги<br />

<strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие и длинные (ри<strong>с</strong>. 1271, 1273 Б).<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong> 12 <strong>с</strong>егмент<strong>а</strong>ми, п<strong>о</strong><strong>с</strong>тепенн<strong>о</strong> ут<strong>о</strong>няющее<strong>с</strong>я<br />

<strong>к</strong> з<strong>а</strong>днему <strong>к</strong><strong>о</strong>нцу. П<strong>о</strong><strong>с</strong>ледние члени<strong>к</strong>и ег<strong>о</strong><br />

(<strong>о</strong>дин или д<strong>в</strong><strong>а</strong>) <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие, <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е<br />

з<strong>а</strong>днебрюшье (ри<strong>с</strong>. 1273 Б). Хелицеры и<br />

<strong>о</strong>рг<strong>а</strong>ны дых<strong>а</strong>ния неиз<strong>в</strong>е<strong>с</strong>тны; <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, Kustarachnida,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные им Amblypygi,<br />

имели д<strong>в</strong>е п<strong>а</strong>ры лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я<br />

н<strong>а</strong> третьем и чет<strong>в</strong>ерт<strong>о</strong>м <strong>с</strong>тернит<strong>а</strong>х<br />

брюш<strong>к</strong><strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> 11—15 мм.<br />

друг<strong>о</strong>г<strong>о</strong>, х<strong>о</strong>тя я имеют <strong>о</strong>бщее пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение.<br />

П<strong>о</strong>пыт<strong>к</strong>и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей (Werner,<br />

1935) н<strong>а</strong>йти и<strong>с</strong>х<strong>о</strong>дных Protopalpigradi, д<strong>а</strong>бы<br />

д<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>ть не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>ть <strong>о</strong>бъединения этих<br />

д<strong>в</strong>ух <strong>о</strong>тряд<strong>о</strong><strong>в</strong>, не м<strong>о</strong>гут быть призн<strong>а</strong>ны уд<strong>а</strong>чными.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Tarantulidae.<br />

СЕМЕЙСТВО TARANTULIDAE KARSCH, 1897<br />

(Phrynidae Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1902;<br />

Cnarontidae Simon, 1936)<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> р<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> четыре п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и три р<strong>о</strong>д<strong>а</strong>: Graeophonus<br />

Scudder, 1890 (ри<strong>с</strong>. 1268, 1269), Protophrynus<br />

Petrunkevitch, 1913 (ри<strong>с</strong>. 1266), Thelyphrynus<br />

Petrunkevitch, 1913 (ри<strong>с</strong>. 1267).<br />

ОТРЯД PALPIGRADI. ПАЛЬПИГРАДЫ<br />

Очень мел<strong>к</strong>ие хелицер<strong>о</strong><strong>в</strong>ые (длин<strong>а</strong> тел<strong>а</strong><br />

0,8—2,8 мм) <strong>с</strong> удлиненным тел<strong>о</strong>м, ут<strong>о</strong>няющим<strong>с</strong>я<br />

н<strong>а</strong> гр<strong>а</strong>нице пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, и <strong>с</strong> длинн<strong>о</strong>й<br />

члени<strong>с</strong>т<strong>о</strong>й х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й нитью (ри<strong>с</strong>. 1274).<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong>. П<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющий<br />

ее г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит р<strong>а</strong><strong>с</strong>членен н<strong>а</strong><br />

<strong>к</strong>рупный г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й щит — пр<strong>о</strong>пельтидий, леж<strong>а</strong>щий<br />

п<strong>о</strong>з<strong>а</strong>ди нег<strong>о</strong> м<strong>а</strong>лень<strong>к</strong>ий мет<strong>а</strong>пельтидий и<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Эти<br />

жгут<strong>о</strong>н<strong>о</strong>гие <strong>о</strong>бит<strong>а</strong>ли, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong> п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е<br />

<strong>в</strong>л<strong>а</strong>жных ле<strong>с</strong><strong>о</strong><strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>я<strong>с</strong><strong>о</strong><strong>в</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong> пери<strong>о</strong>д<strong>а</strong> <strong>в</strong><br />

Амери<strong>к</strong>е.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

<strong>о</strong>тряд<strong>а</strong> Kustarachnida р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны <strong>о</strong>тряду<br />

Amblypygi, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, и пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят.<br />

Предп<strong>о</strong>л<strong>о</strong>жение Мел<strong>а</strong>нжер<strong>а</strong> (Melanger,<br />

1903) и П<strong>о</strong><strong>к</strong><strong>о</strong><strong>к</strong><strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1910) <strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>е Kustarachnida<br />

<strong>с</strong> Ricinulei <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong> и не п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>ими<br />

д<strong>а</strong>нными. Отряд Kustarachnida, <strong>в</strong>ер<strong>о</strong>ятнее <strong>в</strong><strong>с</strong>ег<strong>о</strong>,<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>с</strong><strong>о</strong>б<strong>о</strong>й неб<strong>о</strong>льшую б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ую<br />

<strong>с</strong>лепую <strong>в</strong>ет<strong>в</strong>ь жгут<strong>о</strong>н<strong>о</strong>гих п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>в</strong><strong>о</strong>тряд<strong>а</strong> Amblypygi,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бил<strong>а</strong><strong>с</strong>ь и <strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>ре <strong>в</strong>ымерл<strong>а</strong><br />

<strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не С. Амери<strong>к</strong>и. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Kustarachnidae.<br />

СЕМЕЙСТВО KUSTARACHNIDAE PETRUNKEVITCH,<br />

1913<br />

Один р<strong>о</strong>д Kustarachne Scudder, 1890 из<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и (ри<strong>с</strong>. 1271, 1273).<br />

(Microthelyphonida, Palpigradida)<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> м<strong>а</strong>лень<strong>к</strong>их щит<strong>к</strong><strong>а</strong> (мез<strong>о</strong>пельтидии), леж<strong>а</strong>щих<br />

н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди, н<strong>а</strong>д <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями<br />

н<strong>о</strong>г III (ри<strong>с</strong>. 1275А). Меди<strong>а</strong>льные гл<strong>а</strong>з<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> передне-б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых угл<strong>о</strong><strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

щит<strong>а</strong> <strong>с</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны имеет<strong>с</strong>я п<strong>о</strong><br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>му гл<strong>а</strong>з<strong>к</strong>у. Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не груди<br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г I и педип<strong>а</strong>льп у <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Palpigradi р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я <strong>к</strong>рупный<br />

437<br />

http://jurassic.ru/

438<br />

http://jurassic.ru/<br />

<strong>с</strong>тернит (дейт<strong>о</strong><strong>с</strong>тернум) <strong>в</strong> <strong>в</strong>иде б<strong>о</strong>льш<strong>о</strong>й тр<strong>а</strong>пецие<strong>в</strong>идн<strong>о</strong>й<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и; п<strong>о</strong>з<strong>а</strong>ди нег<strong>о</strong>, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>в</strong><strong>с</strong>ех з<strong>а</strong>дних п<strong>а</strong>р н<strong>о</strong>г, леж<strong>а</strong>т м<strong>а</strong>лень<strong>к</strong>ие<br />

<strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тные <strong>с</strong>терниты (трит<strong>о</strong><strong>с</strong>тернум, пент<strong>а</strong><strong>с</strong>тернум<br />

и мет<strong>а</strong><strong>с</strong>тернум). У <strong>в</strong>ерхнеюр<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

Sternarthron zitteli Haase н<strong>а</strong> <strong>в</strong><strong>с</strong>ех грудных<br />

<strong>с</strong>егмент<strong>а</strong>х были <strong>о</strong>тдельные ше<strong>с</strong>ть м<strong>а</strong>лень<strong>к</strong>их<br />

<strong>с</strong>терн<strong>а</strong>льных щит<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1276); эт<strong>о</strong> п<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет,<br />

чт<strong>о</strong> дейт<strong>о</strong><strong>с</strong>тернум <strong>с</strong><strong>о</strong><strong>в</strong>ременных ф<strong>о</strong>рм <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>л<strong>с</strong>я<br />

<strong>в</strong> результ<strong>а</strong>те <strong>с</strong>лияния трех <strong>с</strong>терн<strong>а</strong>льных<br />

щит<strong>к</strong><strong>о</strong><strong>в</strong> (хелицер<strong>а</strong>льный, педип<strong>а</strong>льпный и<br />

н<strong>о</strong>жн<strong>о</strong>й I). Н<strong>а</strong>личие этих р<strong>а</strong>зр<strong>о</strong>зненных щит<strong>к</strong><strong>о</strong><strong>в</strong><br />

длительн<strong>о</strong>е <strong>в</strong>ремя <strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong>л<strong>о</strong><strong>с</strong>ь не<strong>о</strong>бъя<strong>с</strong>ненным,<br />

<strong>а</strong> юр<strong>с</strong><strong>к</strong>ие Palpigradi (р<strong>о</strong>д Sternarthron<br />

Haase) <strong>о</strong>тн<strong>о</strong><strong>с</strong>или<strong>с</strong>ь <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong> <strong>к</strong> «<strong>о</strong>тряду» Pedipalpi;<br />

лишь Вер<strong>с</strong>луи<strong>с</strong> и Дэм<strong>о</strong>ль (Versluys und<br />

Demoll, 1922) у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>или пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>е <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е<br />

п<strong>о</strong>л<strong>о</strong>жение эт<strong>о</strong>г<strong>о</strong> жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong>. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> <strong>о</strong>динн<strong>а</strong>дц<strong>а</strong>тью я<strong>с</strong>н<strong>о</strong> р<strong>а</strong><strong>с</strong>члененными <strong>с</strong>егмент<strong>а</strong>ми<br />

<strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т пр<strong>о</strong><strong>с</strong><strong>о</strong>мы зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>с</strong>уженным <strong>с</strong>едьмым <strong>с</strong>егмент<strong>о</strong>м, тергиты и <strong>с</strong>терниты<br />

н<strong>а</strong> <strong>с</strong>егмент<strong>а</strong>х брюш<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. З<strong>а</strong>дние<br />

три <strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong> (XV—XVII) т<strong>о</strong>н<strong>к</strong>ие,<br />

цилиндриче<strong>с</strong><strong>к</strong>ие, <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую и т<strong>о</strong>н<strong>к</strong>ую<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>му, н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем <strong>с</strong>егменте <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й причленяет<strong>с</strong>я<br />

длинн<strong>а</strong>я члени<strong>с</strong>т<strong>а</strong>я и п<strong>о</strong><strong>к</strong>рыт<strong>а</strong>я <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми<br />

х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я нить (flagellum) (ри<strong>с</strong>.<br />

1274—1276). Хелицеры <strong>к</strong>рупные, <strong>к</strong>лешне<strong>в</strong>идные,<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, рез<strong>к</strong><strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ют <strong>в</strong>перед,<br />

<strong>с</strong> из<strong>о</strong>гнут<strong>о</strong>й н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну <strong>к</strong>лешней<br />

(ри<strong>с</strong>. 1275А). Педип<strong>а</strong>льпы длинные, т<strong>о</strong>н<strong>к</strong>ие<br />

де<strong>в</strong>ятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные (ри<strong>с</strong>. 1274); н<strong>а</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии эт<strong>о</strong>й <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти <strong>с</strong>тр<strong>о</strong>ения педип<strong>а</strong>льп<br />

и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых других <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> Palpigradi<br />

<strong>с</strong>ближ<strong>а</strong>ли <strong>с</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong>м Schizopeltidia из <strong>о</strong>тряд<strong>а</strong><br />

Uropygi. Н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong> б<strong>о</strong>льшим<br />

чи<strong>с</strong>л<strong>о</strong>м г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мных мел<strong>к</strong>их члени<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

У юр<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> Sternasthron н<strong>о</strong>ги н<strong>а</strong> <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong>е<br />

имеют <strong>в</strong>ид т<strong>о</strong>н<strong>к</strong>их, прямых б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 1276). Дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых других Scorpionomorpha,<br />

четырьмя п<strong>а</strong>р<strong>а</strong>ми упл<strong>о</strong>щенных лег<strong>о</strong>чных<br />

меш<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

третьег<strong>о</strong> — ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; <strong>в</strong> глубине этих<br />

меш<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>реди мышц имеют<strong>с</strong>я примити<strong>в</strong>ные<br />

тр<strong>а</strong>хеи. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Зде<strong>с</strong>ь имеет<strong>с</strong>я трехл<strong>о</strong>п<strong>а</strong><strong>с</strong>тн<strong>о</strong>й генит<strong>а</strong>льный<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong>, пр<strong>о</strong>из<strong>о</strong>шедший из г<strong>о</strong>н<strong>а</strong>п<strong>о</strong>физ II и III<br />

Ри<strong>с</strong>. 1271—1276. Отряды Kustarachnida и Palpigradi<br />

1271. Kustarachne conica Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>. брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1272.<br />

Kustarachne tenulpes Scudder; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, Х4.0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1273. Kustarachne extincta<br />

Melander; A — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, X^.O; Б — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х3,0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913),<br />

1274. Koenenia mlrabilis Qrassi; <strong>о</strong>бщий <strong>в</strong>ид, Х3,5, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (В<strong>о</strong>еwer,<br />

1934). 1275. Koenenia mlrabilis Grassi; A — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>а</strong>м<strong>к</strong>и<br />

<strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х7.0; Б —брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х7.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Millot,<br />

1949). 1276. Sternarthron zitteli Haase; <strong>о</strong>бщий <strong>в</strong>ид, X 5<br />

>°.<br />

юр<strong>а</strong>, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Werner, 1935).

брюшных <strong>с</strong>егмент<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1275 Б). П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong> у <strong>о</strong>б<strong>о</strong>их п<strong>о</strong>л<strong>о</strong><strong>в</strong> р<strong>а</strong>з<strong>в</strong>ит примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. С<strong>о</strong><strong>в</strong>ременные<br />

Palpigradi <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> н<strong>а</strong><strong>с</strong>ыщенн<strong>о</strong>й<br />

<strong>в</strong>л<strong>а</strong>г<strong>о</strong>й п<strong>о</strong>ч<strong>в</strong>е тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>тей С. и Ю. Амери<strong>к</strong>и, Афри<strong>к</strong>и, Е<strong>в</strong>р<strong>о</strong>пы и<br />

юг<strong>о</strong>-<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>чн<strong>о</strong>й Азии; б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> н<strong>а</strong>х<strong>о</strong>д<strong>о</strong><strong>к</strong><br />

был<strong>о</strong> <strong>с</strong>дел<strong>а</strong>н<strong>о</strong> н<strong>а</strong> берег<strong>а</strong>х м<strong>о</strong>рей и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

пре<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>дных <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>о</strong><strong>в</strong>. В <strong>в</strong>. Афри<strong>к</strong>е (Kijabe)<br />

<strong>к</strong>енении были н<strong>а</strong>йдены н<strong>а</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong>те 2100 м н<strong>а</strong>д<br />

ур<strong>о</strong><strong>в</strong>нем м<strong>о</strong>ря. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны<br />

из <strong>в</strong>. юры Б<strong>а</strong><strong>в</strong><strong>а</strong>рии.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Palpigradi<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют <strong>с</strong><strong>о</strong>б<strong>о</strong>й <strong>о</strong>тдельный <strong>о</strong>тряд <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>м<strong>о</strong>рф,<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енный примити<strong>в</strong>ным жгут<strong>о</strong>н<strong>о</strong>гим<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>м<strong>о</strong>рф<strong>а</strong>м <strong>о</strong>тряд<strong>а</strong> Uropygi (п<strong>о</strong>д<strong>о</strong>тряд<br />

Schizopeltidia). Объединение этих д<strong>в</strong>ух<br />

<strong>о</strong>тряд<strong>о</strong><strong>в</strong> не<strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>. Palpigradi, <strong>в</strong>идим<strong>о</strong>,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и Schizopeltidia, <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>били<strong>с</strong>ь <strong>о</strong>т <strong>о</strong>бщег<strong>о</strong><br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong> Scorpionomorpha <strong>в</strong> <strong>с</strong><strong>а</strong>мый н<strong>а</strong>ч<strong>а</strong>льный<br />

пери<strong>о</strong>д ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ния н<strong>а</strong>земных хелицер<strong>о</strong><strong>в</strong>ых;<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременные Palpigradi, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<br />

еще д<strong>о</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>с</strong>тепени пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льный<br />

<strong>в</strong><strong>о</strong>дный <strong>о</strong>бр<strong>а</strong>з жизни. Общие пред<strong>к</strong><strong>о</strong><strong>в</strong>ые ф<strong>о</strong>рмы,<br />

д<strong>а</strong><strong>в</strong>шие н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителям<br />

н<strong>а</strong>д<strong>о</strong>тряд<strong>а</strong> Pedipalpides, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, я<strong>в</strong>или<strong>с</strong>ь и<br />

и<strong>с</strong>т<strong>о</strong>чни<strong>к</strong><strong>о</strong>м р<strong>а</strong>з<strong>в</strong>ития не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых групп Arachnida,<br />

<strong>в</strong> пер<strong>в</strong>ую <strong>о</strong>чередь <strong>о</strong>тряд<strong>а</strong> Solifugae и<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Acaromorpha (<strong>с</strong>м. <strong>с</strong>тр. 446, 447).<br />

Отряд Palpigradi пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен д<strong>в</strong>умя <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>ми:<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременными Eukoeneniidae и юр<strong>с</strong><strong>к</strong>ими<br />

Sternarthronidae.<br />

СЕМЕЙСТВО STERNARTHRONIDAE HAASE, 1890<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>с</strong> ут<strong>о</strong>нченными<br />

<strong>к</strong><strong>о</strong>нце<strong>в</strong>ыми <strong>с</strong>егмент<strong>а</strong>ми и х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

нитью. Сегменты, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные н<strong>а</strong> гр<strong>а</strong>нице<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, не <strong>с</strong>ужены. Н<strong>а</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди ше<strong>с</strong>ть <strong>о</strong>тдельных<br />

<strong>с</strong>терн<strong>а</strong>льных щит<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных мет<strong>а</strong>мерн<strong>о</strong>,<br />

<strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> ше<strong>с</strong>ти п<strong>а</strong>р<strong>а</strong>м <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей.<br />

Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные (ри<strong>с</strong>. 1276). Длин<strong>а</strong><br />

тел<strong>а</strong> 15 мм. Юр<strong>а</strong>. Один р<strong>о</strong>д Sternarthron<br />

Haase, 1890 (= Halometra Oppenheim, 1887)<br />

из <strong>в</strong>. юры Герм<strong>а</strong>нии.<br />

ОТРЯД PSEUDOSCORPIONODEA. ЛОЖНОСКОРПИОНЫ<br />

(Pseudoscorpion.es, Chelonethi, Chelonethida)<br />

Тел<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е или <strong>в</strong>ытянут<strong>о</strong>е, упл<strong>о</strong>щенн<strong>о</strong>е<br />

<strong>в</strong> д<strong>о</strong>р<strong>с</strong><strong>о</strong><strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>м н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лении, шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>е или <strong>с</strong> <strong>о</strong>дним м<strong>а</strong>лень<strong>к</strong>им<br />

цилиндриче<strong>с</strong><strong>к</strong>им члени<strong>к</strong><strong>о</strong>м (п<strong>о</strong>д<strong>о</strong>тряд Neobisiiпе<strong>а</strong>)<br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце. С<strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

тел<strong>о</strong> чет<strong>к</strong><strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделен<strong>о</strong> н<strong>а</strong> уз<strong>к</strong>ую и <strong>о</strong>бычн<strong>о</strong><br />

треуг<strong>о</strong>льную пр<strong>о</strong><strong>с</strong><strong>о</strong>му и н<strong>а</strong> б<strong>о</strong>лее <strong>к</strong>рупн<strong>о</strong>е<br />

брюш<strong>к</strong><strong>о</strong>. Об<strong>а</strong> эти р<strong>а</strong>здел<strong>а</strong> неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong><strong>о</strong>членены<br />

друг <strong>с</strong> друг<strong>о</strong>м. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> пл<strong>о</strong>тным<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит<strong>о</strong>м, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый имеет или<br />

д<strong>в</strong>е п<strong>о</strong>перечные б<strong>о</strong>р<strong>о</strong>зды, р<strong>а</strong>зделяющие ег<strong>о</strong> н<strong>а</strong><br />

три <strong>о</strong>тдел<strong>а</strong> (пр<strong>о</strong>пельтидий, мез<strong>о</strong>- и мет<strong>а</strong>пельтидий),<br />

или т<strong>о</strong>ль<strong>к</strong><strong>о</strong> д<strong>в</strong>е п<strong>о</strong>перечные <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong>и,<br />

<strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>ющие гр<strong>а</strong>ницы <strong>с</strong>лияния этих пер<strong>в</strong>ичных<br />

щит<strong>к</strong><strong>о</strong><strong>в</strong>. Н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е щит<strong>а</strong> или н<strong>а</strong><br />

ег<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х д<strong>в</strong><strong>а</strong> или четыре <strong>к</strong>рупных гл<strong>а</strong>з<strong>а</strong>; реже<br />

гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют или <strong>в</strong> т<strong>о</strong>й или ин<strong>о</strong>й <strong>с</strong>тепени<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Вд<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я щит<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<br />

уз<strong>к</strong>ий прям<strong>о</strong>уг<strong>о</strong>льный тергит пер<strong>в</strong><strong>о</strong>г<strong>о</strong><br />

брюшн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый не р<strong>а</strong>зделен н<strong>а</strong><br />

д<strong>в</strong>е п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины. В<strong>с</strong>е <strong>о</strong><strong>с</strong>т<strong>а</strong>льные тергиты<br />

(II—X) и <strong>с</strong>терниты (III—X) <strong>о</strong>бычн<strong>о</strong> р<strong>а</strong>зделены<br />

<strong>в</strong>д<strong>о</strong>ль <strong>с</strong>редней линии пр<strong>о</strong><strong>с</strong><strong>в</strong>ет<strong>а</strong>ми<br />

мяг<strong>к</strong><strong>о</strong>й мембр<strong>а</strong>ны н<strong>а</strong> ле<strong>в</strong>ую и пр<strong>а</strong><strong>в</strong>ую<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины. Тергиты и <strong>с</strong>терниты XI <strong>с</strong>егмент<strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> цельные, нер<strong>а</strong>зделенные (ри<strong>с</strong>. 1277—<br />

1283). У примити<strong>в</strong>ных л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

Chthoniinea и мн<strong>о</strong>гих Neobisiinea тер­<br />

гиты и <strong>с</strong>терниты цельные, не р<strong>а</strong>зделенные пр<strong>о</strong>д<strong>о</strong>льными<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми мяг<strong>к</strong><strong>о</strong>й мембр<strong>а</strong>ны<br />

(ри<strong>с</strong>. 1278, 1279). Н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong><br />

у мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> имеет<strong>с</strong>я рудимент<strong>а</strong>рный <strong>а</strong>н<strong>а</strong>льный<br />

(XII) <strong>с</strong>егмент, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый ин<strong>о</strong>гд<strong>а</strong> имеет <strong>в</strong>ид<br />

уз<strong>к</strong><strong>о</strong>й, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й труб<strong>к</strong>и, н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>я этим <strong>о</strong><br />

бы<strong>в</strong>шей мет<strong>а</strong><strong>с</strong><strong>о</strong>ме пред<strong>к</strong><strong>о</strong><strong>в</strong>ых ф<strong>о</strong>рм (Amblypygi).<br />

Стерн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди цели<strong>к</strong><strong>о</strong>м<br />

п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> <strong>с</strong><strong>о</strong>м<strong>к</strong>нутыми друг <strong>с</strong> друг<strong>о</strong>м т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

педип<strong>а</strong>льп и х<strong>о</strong>дильных н<strong>о</strong>г. Хелицеры<br />

неб<strong>о</strong>льшие, <strong>к</strong>лешне<strong>в</strong>идные, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Педип<strong>а</strong>льпы <strong>о</strong>чень <strong>к</strong>рупные, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>с</strong> <strong>к</strong>рупными <strong>к</strong>лешнями, н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющие педип<strong>а</strong>льпы<br />

<strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, н<strong>о</strong> менее <strong>с</strong>ильн<strong>о</strong> хитинизир<strong>о</strong><strong>в</strong><strong>а</strong>нные,<br />

и <strong>с</strong> длинными чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительными<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. Н<strong>а</strong> <strong>к</strong><strong>о</strong>нце п<strong>а</strong>льц<strong>а</strong> <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

пр<strong>о</strong>т<strong>о</strong><strong>к</strong>и п<strong>а</strong>утинных желез, <strong>с</strong>е<strong>к</strong>рет<strong>о</strong>м <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>о</strong>б<strong>в</strong><strong>о</strong>л<strong>а</strong><strong>к</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я д<strong>о</strong>быч<strong>а</strong> и <strong>к</strong>л<strong>а</strong>д<strong>к</strong>и яиц. Х<strong>о</strong>дильные<br />

н<strong>о</strong>ги <strong>с</strong>редней длины, <strong>с</strong> пятью члени<strong>к</strong><strong>а</strong>ми<br />

(л<strong>а</strong>п<strong>к</strong><strong>а</strong> не р<strong>а</strong><strong>с</strong>членен<strong>а</strong>); бедр<strong>а</strong> н<strong>о</strong>г не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ут<strong>о</strong>лщены; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и <strong>с</strong>тебельч<strong>а</strong>тым<br />

прет<strong>а</strong>рз<strong>о</strong>м (arolium). В <strong>с</strong>ередине<br />

<strong>с</strong>тернит<strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие, при<strong>к</strong>рыт<strong>о</strong>е <strong>с</strong>переди<br />

<strong>к</strong>рыш<strong>к</strong><strong>о</strong>й (operculum). П<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м III и IV <strong>с</strong>тернит<strong>о</strong><strong>в</strong>,<br />

н<strong>а</strong> ме<strong>с</strong>т<strong>а</strong>х р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жения лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong><br />

других Scorpionomorpha, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

д<strong>в</strong>е п<strong>а</strong>ры <strong>с</strong>тигм, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>нутрь тел<strong>а</strong> <strong>о</strong>тх<strong>о</strong>-<br />

439<br />

http://jurassic.ru/

1277. Heterolophus kochil (Menge); <strong>о</strong>бщий <strong>в</strong>ид, y- 1 - Oligcihernes weigandi (Menge); <strong>о</strong>бщий <strong>в</strong>ид<br />

o n<br />

l o v n F ? 8 i «б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,'<br />

1955). 1282. Oltgochelifer berendtl (Menge); <strong>о</strong>бщий <strong>в</strong>ид, Х37.0, п<strong>а</strong>!<br />

7§2» ен<br />

Л, ? <strong>в</strong><br />

,8? п<strong>а</strong><br />

http://jurassic.ru/<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий «янт<strong>а</strong>рь) (Petrunkevitch, 1955)<br />

1.283. Cheiridiumhartmannl (Menge); <strong>о</strong>бщий <strong>в</strong>ид, ХЗЗ.О, п<strong>а</strong>ле<strong>о</strong>ген<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1955)<br />

ременных <strong>в</strong>ид<strong>о</strong><strong>в</strong>. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны<br />

из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. В <strong>с</strong><strong>в</strong>язи <strong>с</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> примити<strong>в</strong>ных<br />

л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные <strong>о</strong>тн<strong>о</strong>шения<br />

<strong>о</strong>тряд<strong>а</strong> Pseudoscorpionodea <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

не<strong>в</strong>ыя<strong>с</strong>ненными. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей<br />

<strong>с</strong>ближ<strong>а</strong>ет <strong>о</strong>тряд Pseudoscorpionodea <strong>с</strong> групп<strong>о</strong>й<br />

Pedipalpi (Amblypygi + Uropygi) (Щел<strong>к</strong><strong>а</strong>н<strong>о</strong><strong>в</strong>це<strong>в</strong>,<br />

1898—1910; Kastner, 1928; Beier, 1932—<br />

1939) и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>а</strong> Araneae (Chamberlin,<br />

1924). Ср<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>ие и<br />

эмбри<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ие м<strong>а</strong>тери<strong>а</strong>лы п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ют<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><strong>с</strong>ть <strong>с</strong>ближения <strong>о</strong>тряд<strong>а</strong> Pseudoscor-

pionodea <strong>с</strong> групп<strong>о</strong>й «Pedipalpi» (<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>с</strong><br />

Amblypygi), <strong>к</strong><strong>о</strong>т<strong>о</strong>рые я<strong>в</strong>или<strong>с</strong>ь <strong>в</strong>идим<strong>о</strong>, и<strong>с</strong>х<strong>о</strong>дн<strong>о</strong>й<br />

не т<strong>о</strong>ль<strong>к</strong><strong>о</strong> для л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, н<strong>о</strong> и для<br />

<strong>о</strong>тряд<strong>а</strong> Аг<strong>а</strong>пе<strong>а</strong>е (<strong>с</strong>м. <strong>с</strong>тр. 382).<br />

Отряд Pseudoscorpionodea из<strong>в</strong>е<strong>с</strong>тен <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь). С<strong>о</strong><strong>в</strong>ременные Pseudoscorpionodea<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены 20 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>ми и 200 р<strong>о</strong>д<strong>а</strong>ми.<br />

Три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Chthoniinea, Neobisiinea,<br />

Cheliferinea.<br />

ПОДОТРЯД CHTHONIINEA<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>с</strong> п<strong>а</strong>р<strong>а</strong>ллельными <strong>к</strong>р<strong>а</strong>ями<br />

или же <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>с</strong>ужен <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти. Четыре<br />

гл<strong>а</strong>з<strong>а</strong>; реже <strong>о</strong>ни бы<strong>в</strong><strong>а</strong>ют редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны.<br />

Хелицеры <strong>к</strong>рупные, м<strong>о</strong>щные; <strong>о</strong>ни <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

<strong>в</strong>ыд<strong>а</strong>ют<strong>с</strong>я з<strong>а</strong> передний <strong>к</strong>р<strong>а</strong>й щит<strong>а</strong>; з<strong>а</strong>зубренный<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т н<strong>а</strong> <strong>в</strong>ершине п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> п<strong>а</strong>льц<strong>а</strong><br />

(galea) <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; гребенч<strong>а</strong>т<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong><br />

(serrula) н<strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>м п<strong>а</strong>льце <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>,<br />

при<strong>к</strong>реплен<strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии; пуч<strong>о</strong><strong>к</strong> щетин<strong>о</strong><strong>к</strong><br />

н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния хелицеры<br />

(flagellum) длинный, <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> <strong>о</strong>дн<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>о</strong>нне<br />

з<strong>а</strong>зубренными щетин<strong>к</strong><strong>а</strong>ми. Педип<strong>а</strong>льпы<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, г<strong>о</strong>лень <strong>в</strong>здут<strong>а</strong>я, ч<strong>а</strong>ше<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я.<br />

Обе передние п<strong>а</strong>ры н<strong>о</strong>г <strong>с</strong> нер<strong>а</strong><strong>с</strong>члененными<br />

(<strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ыми) л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми, <strong>а</strong> л<strong>а</strong>п<strong>к</strong>и<br />

<strong>о</strong>беих з<strong>а</strong>дних п<strong>а</strong>р н<strong>о</strong>г д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

(ри<strong>с</strong>. 1279). Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I, II и III <strong>с</strong> шип<strong>а</strong>ми.<br />

Бедр<strong>а</strong> н<strong>о</strong>г IV ут<strong>о</strong>лщены. К<strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> пр<strong>о</strong><strong>с</strong>тые,<br />

длинные. Вт<strong>о</strong>ричн<strong>о</strong>п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые призн<strong>а</strong><strong>к</strong>и <strong>в</strong>ыр<strong>а</strong>жены<br />

нерез<strong>к</strong><strong>о</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Dithidae и Chthoniidae.<br />

СЕМЕЙСТВО DITHIDAE CHAMBERLIN, 1931<br />

Стигмы р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>к</strong><strong>о</strong><strong>с</strong><strong>о</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е п<strong>о</strong>ле<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и упл<strong>о</strong>тненн<strong>о</strong>е. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>: из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Heterolophus<br />

Tomosvary, 1884 (ри<strong>с</strong>. 1277).<br />

СЕМЕЙСТВО CHTHONIIDAE HANSEN, 1894<br />

Стигмы р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong>перечн<strong>о</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

п<strong>о</strong>ле <strong>с</strong><strong>а</strong>м<strong>к</strong>и мяг<strong>к</strong><strong>о</strong>е (ри<strong>с</strong>. 1278). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>: из них <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Chthonius<br />

С. L. Koch, 1843.<br />

ПОДОТРЯД NEOBISIINEA<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>с</strong> б<strong>о</strong>лее или менее п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>лег<strong>к</strong><strong>а</strong> <strong>в</strong><strong>о</strong>гнутыми.<br />

Гл<strong>а</strong>з д<strong>в</strong><strong>а</strong> или четыре; у пещерных <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

их нет. Хелицеры <strong>к</strong>рупные и м<strong>о</strong>щные, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

<strong>в</strong>ыд<strong>а</strong>ют<strong>с</strong>я з<strong>а</strong> <strong>к</strong>р<strong>а</strong>й щит<strong>а</strong>; galea имеет<strong>с</strong>я или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

serrula п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> п<strong>а</strong>льц<strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong> н<strong>а</strong> ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>м <strong>к</strong>р<strong>а</strong>е. Г<strong>о</strong>лень<br />

441<br />

http://jurassic.ru/<br />

педип<strong>а</strong>льп <strong>в</strong>ытянут<strong>а</strong>я. Л<strong>а</strong>п<strong>к</strong>и четырех п<strong>а</strong>р х<strong>о</strong>дильных<br />

н<strong>о</strong>г <strong>о</strong>бычн<strong>о</strong> д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Бедр<strong>а</strong><br />

<strong>в</strong><strong>с</strong>ех н<strong>о</strong>г р<strong>а</strong>з<strong>в</strong>иты примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>. Вт<strong>о</strong>ричн<strong>о</strong>п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые<br />

призн<strong>а</strong><strong>к</strong>и <strong>в</strong>ыр<strong>а</strong>жены рез<strong>к</strong><strong>о</strong>. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Neobisiidea,<br />

Garypidea и Feaellidea.<br />

НАДСЕМЕЙСТВО NEOBISIIDEA<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. П<strong>о</strong>д<strong>в</strong>ижный<br />

п<strong>а</strong>лец <strong>к</strong>лешни хелицеры з<strong>а</strong>зубренный. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Neobisiidae, Syarinidae, Ideoroncidae;<br />

пер<strong>в</strong><strong>о</strong>е из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО NEOBISIIDAE CHAMBERLIN^ 1930<br />

Basi- и telofemur н<strong>о</strong>ги IV <strong>о</strong>тгр<strong>а</strong>ничены б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й.<br />

Яд<strong>о</strong><strong>в</strong>итые железы т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>м<br />

п<strong>а</strong>льце педип<strong>а</strong>льп. Пле<strong>в</strong>р<strong>а</strong>льные <strong>о</strong>бл<strong>а</strong><strong>с</strong>ти<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 11 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Neobisiutn<br />

Chamberlin, 1930 (ри<strong>с</strong>. 1279).<br />

НАДСЕМЕЙСТВО GARYPIDEA<br />

Л<strong>а</strong>п<strong>к</strong>и д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. П<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец<br />

хелицер гл<strong>а</strong>д<strong>к</strong>ий. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Menthidae, Olpiidae,-<br />

Garypidae, из них <strong>о</strong>б<strong>а</strong> п<strong>о</strong><strong>с</strong>ледних из<strong>в</strong>е<strong>с</strong>тны<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО OLPI1DAE CHAMBERLIN, 1930<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> незн<strong>а</strong>чительн<strong>о</strong> шире пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Об<strong>а</strong> п<strong>а</strong>льц<strong>а</strong> <strong>к</strong>лешни педип<strong>а</strong>льп <strong>с</strong> яд<strong>о</strong><strong>в</strong>итыми<br />

желез<strong>а</strong>ми. Пле<strong>в</strong>ры <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы гл<strong>а</strong>д<strong>к</strong>ие. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 18 р<strong>о</strong>д<strong>о</strong><strong>в</strong>,,<br />

из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

Garypinus Daday, 1888.<br />

СЕМЕЙСТВО GARYPIDAE HANSEN, 1894<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> шире пр<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Об<strong>а</strong> п<strong>а</strong>льц<strong>а</strong> педип<strong>а</strong>льп <strong>с</strong> яд<strong>о</strong><strong>в</strong>итыми желез<strong>а</strong>ми.<br />

Пле<strong>в</strong>ры <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> буг<strong>о</strong>рч<strong>а</strong>т<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тью.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них Garypus С. L. Koch, 1873 и<br />

Geogarypus Chamberlin, 1930 <strong>в</strong> ми<strong>о</strong>цене<br />

Ю. Азии и п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

НАДСЕМЕЙСТВО FEAELLIDEA<br />

В<strong>с</strong>е л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми или л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Synsphyronidae, Feaellidae, Pseudogarypidae;.<br />

п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.

СЕМЕЙСТВО PSEUDOGARYPIDAE CHAMBERLIN,<br />

1923<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их р<strong>о</strong>г<strong>о</strong><strong>в</strong>.<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> без пле<strong>в</strong>р<strong>а</strong>льных пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong>.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong>дин<br />

р<strong>о</strong>д Pseudogarypus Ellingsen, 1909, из<strong>в</strong>е<strong>с</strong>тный<br />

т<strong>а</strong><strong>к</strong>же из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(ри<strong>с</strong>. 1280).<br />

ПОДОТРЯД CHELIFERINEA<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й щит <strong>с</strong> <strong>в</strong>ыпу<strong>к</strong>лыми б<strong>о</strong><strong>к</strong><strong>а</strong>ми, шир<strong>о</strong><strong>к</strong>ий<br />

или треуг<strong>о</strong>льный и <strong>с</strong> прямыми <strong>к</strong>р<strong>а</strong>ями,<br />

<strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я <strong>в</strong>переди. Д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>а</strong>, или их нет<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть щит<strong>а</strong> гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>,<br />

<strong>с</strong> шип<strong>о</strong><strong>в</strong><strong>а</strong>тыми щетин<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>.<br />

1282). Тергиты и <strong>с</strong>терниты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong>зделены<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong> п<strong>о</strong> меди<strong>а</strong>льн<strong>о</strong>й линии<br />

(ри<strong>с</strong>. 1280, 1281, 1283). Хелицеры м<strong>а</strong>лень<strong>к</strong>ие;<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> п<strong>а</strong>льц<strong>а</strong> имеет<strong>с</strong>я б<strong>о</strong>лее<br />

или менее х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я galea, serrula п<strong>о</strong><br />

<strong>в</strong><strong>с</strong>ей длине прир<strong>а</strong><strong>с</strong>т<strong>а</strong>ет <strong>к</strong> п<strong>а</strong>льцу хелицеры. Педип<strong>а</strong>льпы<br />

длинные, <strong>с</strong> <strong>о</strong>чень <strong>к</strong>рупн<strong>о</strong>й и м<strong>о</strong>щн<strong>о</strong>й<br />

<strong>к</strong>лешней. В<strong>с</strong>е л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>о</strong>дн<strong>о</strong>члени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Третичные<br />

— ныне. Д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Cheliferidea<br />

u Cheiridiidea.<br />

НАДСЕМЕЙСТВО CHELIFERIDEA<br />

Н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>м щите пр<strong>о</strong>х<strong>о</strong>дят <strong>о</strong>дн<strong>а</strong> или<br />

д<strong>в</strong>е п<strong>о</strong>перечные б<strong>о</strong>р<strong>о</strong>зды (ри<strong>с</strong>. 1281, 1282), или<br />

<strong>о</strong>ни <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Неп<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец <strong>к</strong>лешни<br />

<strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью, <strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец <strong>с</strong> четырьмя<br />

чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. Бедр<strong>а</strong> н<strong>о</strong>г I и II<br />

п<strong>о</strong>др<strong>а</strong>зделены б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й ин<strong>а</strong>че, чем у н<strong>о</strong>г III и<br />

IV. Между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г хитин<strong>о</strong><strong>в</strong><strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong><br />

п<strong>с</strong>е<strong>в</strong>д<strong>о</strong><strong>с</strong>тер ну м <strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Atemnidae,<br />

Chernetidae, Myrmochernetidae, Cheliferidae,<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их и <strong>с</strong>убтр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их<br />

л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые из<strong>в</strong>е<strong>с</strong>тны<br />

лишь из <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Chernetidae и Cheliferidae.<br />

СЕМЕЙСТВО CHERNETIDAE MENGE, 1855<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>в</strong>ытянут; длин<strong>а</strong> ег<strong>о</strong><br />

пре<strong>в</strong>ыш<strong>а</strong>ет ширину; передний <strong>к</strong>р<strong>а</strong>й ег<strong>о</strong> б<strong>о</strong>лее<br />

или менее з<strong>а</strong><strong>к</strong>руглен; <strong>о</strong>дн<strong>а</strong> или д<strong>в</strong>е п<strong>о</strong>перечные<br />

линии <strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине щит<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жены<br />

(ри<strong>с</strong>. 1281). Гл<strong>а</strong>з<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты; <strong>о</strong>чень<br />

ред<strong>к</strong><strong>о</strong> гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Щетин<strong>к</strong>и тел<strong>а</strong> и педип<strong>а</strong>льп<br />

р<strong>а</strong>з<strong>в</strong>иты н<strong>о</strong>рм<strong>а</strong>льн<strong>о</strong> и не р<strong>а</strong><strong>с</strong>ширены<br />

л<strong>о</strong>ж<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>. Пуч<strong>о</strong><strong>к</strong> щетин<strong>о</strong><strong>к</strong> flagellum хелицер<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен из д<strong>в</strong>ух — четырех щетин<strong>о</strong><strong>к</strong>.<br />

Пр<strong>о</strong>т<strong>о</strong><strong>к</strong>и яд<strong>о</strong><strong>в</strong>ит<strong>о</strong>й желез<strong>к</strong>и имеют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong><br />

неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>м п<strong>а</strong>льце <strong>к</strong>лешни педип<strong>а</strong>льп: н<strong>а</strong><br />

•<strong>о</strong>б<strong>о</strong>их п<strong>а</strong>льц<strong>а</strong>х <strong>к</strong>лешни педип<strong>а</strong>льп, <strong>к</strong>р<strong>о</strong>ме ряд<strong>а</strong><br />

442<br />

http://jurassic.ru/<br />

зуб<strong>о</strong><strong>в</strong> н<strong>а</strong> режущем <strong>к</strong>р<strong>а</strong>е, имеет<strong>с</strong>я <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й, б<strong>о</strong>лее<br />

ред<strong>к</strong>ий ряд зуб<strong>о</strong><strong>в</strong>, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>т<strong>с</strong>туп<strong>а</strong>ющий <strong>о</strong>т<br />

<strong>к</strong>р<strong>а</strong>я. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г IV <strong>с</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительн<strong>о</strong>й<br />

щетин<strong>к</strong><strong>о</strong>й. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 43 р<strong>о</strong>д<strong>а</strong>. В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) <strong>о</strong>дин <strong>в</strong>ид р<strong>о</strong>д<strong>а</strong> Oligochernes<br />

Beier, 1937 (ри<strong>с</strong>. 1281).<br />

СЕМЕЙСТВО CHELIFERIDAE HAGEN, 1879<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит удлиненный, треуг<strong>о</strong>льный,<br />

либ<strong>о</strong> длин<strong>а</strong> и ширин<strong>а</strong> ег<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ы. П<strong>о</strong>перечные<br />

б<strong>о</strong>р<strong>о</strong>зды и гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong>з<strong>в</strong>иты х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong>; у<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> гл<strong>а</strong>з<strong>а</strong> бы<strong>в</strong><strong>а</strong>ют редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны,<br />

и т<strong>о</strong>гд<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> гл<strong>а</strong>зные пятн<strong>а</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> и тергит<strong>о</strong><strong>в</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><br />

груб<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й. Щетин<strong>к</strong>и н<strong>а</strong> теле и н<strong>а</strong><br />

педип<strong>а</strong>льп<strong>а</strong>х з<strong>а</strong>зубрены или ут<strong>о</strong>лщены <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии.<br />

Flagellum хелицер <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен из трех —<br />

пяти щетин<strong>о</strong><strong>к</strong>. Н<strong>а</strong> <strong>о</strong>б<strong>о</strong>их п<strong>а</strong>льц<strong>а</strong>х педип<strong>а</strong>льп<br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я пр<strong>о</strong>т<strong>о</strong><strong>к</strong> яд<strong>о</strong><strong>в</strong>итых желез<strong>о</strong><strong>к</strong>. Б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й<br />

ряд зуб<strong>о</strong><strong>в</strong> н<strong>а</strong> п<strong>а</strong>льц<strong>а</strong>х <strong>к</strong>лешни педип<strong>а</strong>льп<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х н<strong>о</strong>г IV имеет<strong>с</strong>я чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительн<strong>а</strong>я<br />

щетин<strong>к</strong><strong>а</strong>, реже <strong>о</strong>н<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует<br />

(ри<strong>с</strong>. 1282). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong>. Из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тны четыре <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong>: Electrochelifer Beier, 1937; Oligochelifer<br />

Beier, 1937; Oligowithius Beier, 1937; Pycnochelifer<br />

Beier, 1937.<br />

НАДСЕМЕЙСТВО CHEIRIDIIDEA<br />

Н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>м щите пр<strong>о</strong>х<strong>о</strong>дят д<strong>в</strong>е п<strong>о</strong>перечные<br />

б<strong>о</strong>р<strong>о</strong>зды (ри<strong>с</strong>. 1283); п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть щит<strong>а</strong><br />

пл<strong>о</strong>тн<strong>а</strong>я, груб<strong>о</strong> <strong>с</strong><strong>к</strong>ульптир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я. Н<strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>м<br />

п<strong>а</strong>льце т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дин — д<strong>в</strong><strong>а</strong> чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительных<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>. Бедр<strong>а</strong> <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>с</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чным, не<br />

фун<strong>к</strong>ци<strong>о</strong>нирующим <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е <strong>с</strong><strong>о</strong>членения<br />

штрих<strong>о</strong>м или б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й. Между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

н<strong>о</strong>г — п<strong>с</strong>е<strong>в</strong>д<strong>о</strong><strong>с</strong>тернум. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Cheiridiidae и<br />

Sternophoridae, из них пер<strong>в</strong><strong>о</strong>е пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО CHEIRIDIIDAE CHAMBERLIN, 1931<br />

Передний <strong>к</strong>р<strong>а</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> <strong>с</strong> углублением;<br />

<strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти щит<strong>а</strong> д<strong>в</strong>е п<strong>о</strong>перечные<br />

б<strong>о</strong>р<strong>о</strong>зды. Имеют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> д<strong>в</strong><strong>а</strong> м<strong>а</strong>лень<strong>к</strong>их<br />

гл<strong>а</strong>з<strong>а</strong>, или же и <strong>о</strong>ни <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы щит<strong>а</strong><br />

т<strong>о</strong>н<strong>к</strong><strong>о</strong> или груб<strong>о</strong> 'пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Flagellum<br />

хелицер <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лен из д<strong>в</strong>ух или трех щетин<strong>о</strong><strong>к</strong>.<br />

Пр<strong>о</strong>т<strong>о</strong><strong>к</strong>и яд<strong>о</strong><strong>в</strong>ит<strong>о</strong>й желез<strong>к</strong>и <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>нце неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> п<strong>а</strong>льц<strong>а</strong>. Х<strong>о</strong>дильные<br />

н<strong>о</strong>ги пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые (ри<strong>с</strong>. 1283). П<strong>а</strong>ле<strong>о</strong>ген •—<br />

ныне. В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий ян«<br />

т<strong>а</strong>рь) р<strong>о</strong>д Cheiridium Menge, 1855.

Ри<strong>с</strong>. 1284—1289. Огря!<br />

1284. Polyochera alticeps Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид, Х7.0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911). 1285. Curculioid.es ansticii Buckland;<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, Х4,0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911).<br />

1286. Curculioides sulcatus (Melander); <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху,<br />

Хб,0; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1287. Ricinoi-<br />

ОТРЯД RICINULEI.<br />

Мел<strong>к</strong>ие члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие <strong>с</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им, т<strong>о</strong>л<strong>с</strong>тым,<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льным тел<strong>о</strong>м, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящим из 17 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

Тел<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>му и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му.<br />

Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я,<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я или призм<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>а</strong>я; п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющий<br />

ее щит <strong>о</strong>бычн<strong>о</strong> <strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льным <strong>с</strong>рединным<br />

<strong>в</strong>д<strong>а</strong><strong>в</strong>лением и п<strong>о</strong>перечн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й, идущей<br />

<strong>в</strong>д<strong>о</strong>ль з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я и <strong>о</strong>тделяющей уз<strong>к</strong>ий уч<strong>а</strong><strong>с</strong>т<strong>о</strong><strong>к</strong><br />

щит<strong>а</strong> (<strong>в</strong>идим<strong>о</strong>, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичный мет<strong>а</strong>пельтидию).<br />

К переднему <strong>к</strong>р<strong>а</strong>ю щит<strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> при-<br />

443<br />

Ricinulei и Solifugae<br />

des feae Hansen; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, x5,0; <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Millot, 1949).<br />

1288. Protosolpuga carbonarla Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, X2,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1289. Oylippus judaicus<br />

Kraepelin; <strong>о</strong>бщий <strong>в</strong>ид: A —<strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу, Х2,0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Бялыниц<strong>к</strong>ий-Бируля, 1913)<br />

РИЦИНУЛЕЙ<br />

http://jurassic.ru/<br />

членен<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> (cucullus), <strong>к</strong><strong>о</strong>т<strong>о</strong>-«<br />

р<strong>а</strong>я, <strong>с</strong>пу<strong>с</strong><strong>к</strong><strong>а</strong>я<strong>с</strong>ь <strong>в</strong>низ, при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>р*<br />

г<strong>а</strong>ны. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>в</strong>яти — де<strong>с</strong>яти<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых т<strong>о</strong>ль<strong>к</strong><strong>о</strong> четыре <strong>о</strong>тчетли<strong>в</strong><strong>о</strong><br />

<strong>в</strong>идны бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>ря н<strong>а</strong>личию пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тых<br />

тергит<strong>о</strong><strong>в</strong> и <strong>с</strong>тернит<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1284—1287). Н<strong>а</strong><br />

з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong> п<strong>о</strong><strong>с</strong>ледние д<strong>в</strong><strong>а</strong> — три <strong>с</strong>егмент<strong>а</strong><br />

<strong>с</strong>ильн<strong>о</strong> ут<strong>о</strong>нены, труб<strong>к</strong><strong>о</strong><strong>в</strong>идны и <strong>о</strong>бр<strong>а</strong>зуют<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>терную для п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Pedipalpides<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ую мет<strong>а</strong><strong>с</strong><strong>о</strong>му (ри<strong>с</strong>. 1284, 1287). Р<strong>о</strong>т<strong>о</strong>-

<strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны из <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>к</strong>лешне<strong>в</strong>идных<br />

хелицер и <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> длинных<br />

педип<strong>а</strong>льп, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящих из ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>: <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>в</strong>ертлуг<strong>а</strong> и бедр<strong>а</strong>, т<strong>о</strong>л<strong>с</strong>т<strong>о</strong>г<strong>о</strong>, длинн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>лен<strong>а</strong>, трубч<strong>а</strong>т<strong>о</strong>й длинн<strong>о</strong>й г<strong>о</strong>лени и <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong> <strong>с</strong>переди. Субтермин<strong>а</strong>льн<strong>о</strong> н<strong>а</strong><br />

ней причленяет<strong>с</strong>я м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я <strong>к</strong><strong>о</strong>гте<strong>в</strong>идн<strong>а</strong>я п<strong>о</strong>д<strong>в</strong>ижн<strong>а</strong>я<br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зующ<strong>а</strong>я н<strong>а</strong> <strong>в</strong>ершине г<strong>о</strong>лени<br />

<strong>к</strong>лешню (ри<strong>с</strong>. 1287). Б<strong>а</strong>з<strong>а</strong>льный члени<strong>к</strong><br />

педип<strong>а</strong>льп — т<strong>а</strong>зи<strong>к</strong> — <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий; <strong>о</strong>б<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong><br />

<strong>с</strong>литы, <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льную пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у, <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ющую<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong>ую п<strong>о</strong>л<strong>о</strong><strong>с</strong>ть <strong>с</strong>з<strong>а</strong>ди. Н<strong>о</strong>ги<br />

длинные, <strong>с</strong><strong>о</strong> мн<strong>о</strong>гими <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими члени<strong>к</strong><strong>а</strong>ми;<br />

н<strong>о</strong>ги I <strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, н<strong>о</strong>ги II <strong>с</strong> ше<strong>с</strong>тью <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ными<br />

члени<strong>к</strong><strong>а</strong>ми и л<strong>а</strong>п<strong>к</strong><strong>о</strong>й, р<strong>а</strong>зделенн<strong>о</strong>й н<strong>а</strong><br />

пять мел<strong>к</strong>их члени<strong>к</strong><strong>о</strong><strong>в</strong>; л<strong>а</strong>п<strong>к</strong><strong>а</strong> н<strong>о</strong>ги III р<strong>а</strong>зделен<strong>а</strong><br />

н<strong>а</strong> четыре члени<strong>к</strong><strong>а</strong>, <strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong> — н<strong>о</strong>ги IV —<br />

н<strong>а</strong> пять члени<strong>к</strong><strong>о</strong><strong>в</strong>. Н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х <strong>в</strong><strong>с</strong>ех л<strong>а</strong>п<strong>о</strong><strong>к</strong> п<strong>о</strong><br />

д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г <strong>к</strong>рупные, и, <strong>к</strong>р<strong>о</strong>ме т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

I и II п<strong>а</strong>р, <strong>с</strong>ближены друг <strong>с</strong> друг<strong>о</strong>м. Н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не брюш<strong>к</strong><strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, имеет<strong>с</strong>я<br />

<strong>с</strong>рединный ряд <strong>к</strong>рупных тергит<strong>о</strong><strong>в</strong>, п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> м<strong>а</strong>лень<strong>к</strong>их<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых тергит<strong>о</strong><strong>в</strong>, <strong>о</strong>тделенных <strong>о</strong>т <strong>с</strong>рединных<br />

пр<strong>о</strong>д<strong>о</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми мяг<strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>жи<br />

(Polyocheridae и Ricinoididae). В эт<strong>о</strong>м <strong>о</strong>тн<strong>о</strong>шении<br />

Ricinulei н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных<br />

Anthracomarti и Trigonotarbi, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые,<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong>едут <strong>с</strong><strong>в</strong><strong>о</strong>е пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение <strong>о</strong>т<br />

рицинулеип<strong>о</strong>д<strong>о</strong>бных пред<strong>к</strong><strong>о</strong><strong>в</strong> (<strong>с</strong>м. <strong>с</strong>хему н<strong>а</strong><br />

<strong>с</strong>тр. 3841. Ин<strong>о</strong>гд<strong>а</strong> имеет<strong>с</strong>я <strong>о</strong>бщий <strong>с</strong>пинн<strong>о</strong>й щит,<br />

р<strong>а</strong>зделенный <strong>с</strong>рединн<strong>о</strong>й линией (Curculioididae).<br />

Орг<strong>а</strong>ны дых<strong>а</strong>ния пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены п<strong>а</strong>р<strong>о</strong>й<br />

<strong>с</strong>тигм, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>о</strong>г III,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>едут <strong>в</strong> <strong>о</strong>бщий меш<strong>к</strong><strong>о</strong><strong>в</strong>идный <strong>а</strong>триум<br />

<strong>с</strong> р<strong>а</strong>з<strong>в</strong>ет<strong>в</strong>ляющими<strong>с</strong>я тр<strong>а</strong>хеями. Эти <strong>о</strong>рг<strong>а</strong>ны<br />

п<strong>о</strong>х<strong>о</strong>жи н<strong>а</strong> <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гичные <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния у л<strong>о</strong>жн<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong><br />

(Pseudoscorpionodea) и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

Araneae. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> у <strong>в</strong><strong>с</strong>ех Pedipalpides, между пер<strong>в</strong>ым и <strong>в</strong>т<strong>о</strong>рым<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ими <strong>с</strong>тернит<strong>а</strong>ми. У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong><br />

<strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я з<strong>а</strong><br />

<strong>с</strong>чет ди<strong>с</strong>т<strong>а</strong>льных члени<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г III. Длин<strong>а</strong> тел<strong>а</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых рицинулей 15,0—16,5 мм, <strong>с</strong><strong>о</strong><strong>в</strong>ременных—<br />

4,5—10 мм.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. С<strong>о</strong><strong>в</strong>ременные<br />

Ricinulei <strong>о</strong>бит<strong>а</strong>ют н<strong>а</strong> п<strong>о</strong>ч<strong>в</strong>е и <strong>с</strong>реди<br />

ли<strong>с</strong>тье<strong>в</strong> р<strong>а</strong><strong>с</strong>тений, <strong>о</strong>бр<strong>а</strong>зующих п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>у н<strong>а</strong><br />

ее п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти, <strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>й Афри<strong>к</strong>е, Центр,<br />

и Ю. Амери<strong>к</strong>е. И<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые рицинулей из<strong>в</strong>е<strong>с</strong>тны<br />

из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Отряд<br />

Ricinulei д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> <strong>в</strong>ремени <strong>с</strong>ближ<strong>а</strong>ли <strong>с</strong><br />

<strong>о</strong>тряд<strong>а</strong>ми Anthracomarti, Opiliones и Uropygi<br />

(или <strong>в</strong><strong>с</strong>еми Pedipalpi). Одн<strong>а</strong><strong>к</strong><strong>о</strong> бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>ря н<strong>а</strong>личию<br />

п<strong>а</strong>ры <strong>с</strong>тигм н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии н<strong>о</strong>г<br />

III и ше<strong>с</strong>тин<strong>о</strong>г<strong>о</strong>й личин<strong>к</strong>и <strong>в</strong>ы<strong>с</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ли<strong>с</strong>ь предп<strong>о</strong>л<strong>о</strong>жения<br />

<strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>е Ricinulei <strong>с</strong> Solifugae и-<br />

Acarina. Сближение Ricinulei <strong>с</strong> этими д<strong>в</strong>умя<br />

<strong>о</strong>тряд<strong>а</strong>ми ничем не <strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, <strong>а</strong> п<strong>о</strong>л<strong>о</strong>жение<br />

<strong>с</strong>тигм н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>ме и нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>итие<br />

чет<strong>в</strong>ерт<strong>о</strong>й п<strong>а</strong>ры н<strong>о</strong>г у личин<strong>к</strong>и я<strong>в</strong>ляют<strong>с</strong>я<br />

<strong>к</strong><strong>о</strong>н<strong>в</strong>ергентными. Сближение <strong>о</strong>тряд<strong>а</strong><br />

Ricinulei <strong>с</strong> <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><strong>о</strong>м Pedipalpi, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти<br />

<strong>с</strong> Amblypygi, <strong>в</strong>п<strong>о</strong>лне <strong>о</strong>пр<strong>а</strong><strong>в</strong>д<strong>а</strong>н<strong>о</strong> и п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я<br />

д<strong>а</strong>нными <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>й <strong>а</strong>н<strong>а</strong>т<strong>о</strong>мии. Ricinulei,<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят <strong>о</strong>т <strong>а</strong>мблипиг<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных<br />

пред<strong>к</strong><strong>о</strong><strong>в</strong> и <strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не п<strong>о</strong>лучили пышн<strong>о</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие, д<strong>а</strong><strong>в</strong> н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> д<strong>в</strong>ум <strong>к</strong>рупным <strong>о</strong>тряд<strong>а</strong>м—<br />

Anthracomarti и Trigonotarbi. Р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> Ricinulei<br />

<strong>с</strong> Opiliones не <strong>в</strong>п<strong>о</strong>лне <strong>в</strong>ыя<strong>с</strong>нен<strong>о</strong>. К<strong>а</strong>рб<strong>о</strong>н —<br />

ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Polyocheridae, Curculioididae,<br />

Ricinoididae, причем п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> из <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уны.<br />

СЕМЕЙСТВО POLYOCHERIDAE SCUDDER, 1884<br />

Н<strong>а</strong> брюш<strong>к</strong>е х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женные четыре<br />

тергит<strong>а</strong> и <strong>с</strong>тернит<strong>а</strong> (ри<strong>с</strong>. 1284). Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I,<br />

III и IV <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м, т<strong>а</strong>зи<strong>к</strong>и<br />

II шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены. Один р<strong>о</strong>д из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

С. Амери<strong>к</strong>и и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Polyochera Scudder, 1884 (=Poliochera<br />

Petrunkevitch, 1953). Тип р<strong>о</strong>д<strong>а</strong> —P. punctulata<br />

Scudder, 1884; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Mazon<br />

Creek). Длин<strong>а</strong> тел<strong>а</strong> 12,0—15,5 мм (ри<strong>с</strong>. 1284).<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и и <strong>о</strong>дин из<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Англии.<br />

СЕМЕЙСТВО CURCULIOIDIDAE COCKERELL, 190&<br />

(Holotergidae Petrunkevitch, 1913)<br />

Тергиты брюш<strong>к</strong><strong>а</strong> <strong>с</strong>литы <strong>в</strong> <strong>о</strong>бщий щит, <strong>о</strong>бычн<strong>о</strong><br />

р<strong>а</strong>зделенный п<strong>о</strong> <strong>с</strong>редней линии н<strong>а</strong> ле<strong>в</strong>уюи<br />

пр<strong>а</strong><strong>в</strong>ую п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины, реже <strong>о</strong>н цельный, нер<strong>а</strong>зделенный.<br />

Стернит третьег<strong>о</strong> члени<strong>к</strong><strong>а</strong> брюш<strong>к</strong><strong>а</strong><br />

р<strong>а</strong>зделен н<strong>а</strong> три ч<strong>а</strong><strong>с</strong>ти; ин<strong>о</strong>гд<strong>а</strong> н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не брюш<strong>к</strong><strong>а</strong> не <strong>в</strong>идн<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>ции. Т<strong>а</strong>зи<strong>к</strong>и<br />

н<strong>о</strong>г IV <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я или <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я. К<strong>а</strong>рб<strong>о</strong>н<br />

С. Амери<strong>к</strong>и и Англии.<br />

Curculioides Buckland, 1837. Тип р<strong>о</strong>д<strong>а</strong><br />

— С. ansticii Buckland, 1837; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия<br />

(Coseley). Х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я призн<strong>а</strong><strong>к</strong><strong>а</strong>ми <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 15,0—16,5 мм (ри<strong>с</strong>. 1285,<br />

1286). Три <strong>в</strong>ид<strong>а</strong> из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и <strong>к</strong><br />

три из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Англии.<br />

http://jurassic.ru/

КЛАСС SOLIFUGOMORPHA. СОЛЬПУГОПОДОБНЫЕ,<br />

ИЛИ ПРОТЕРОСОМНЫЕ ПАУКООБРАЗНЫЕ<br />

Тел<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> п<strong>о</strong> измененн<strong>о</strong>му трил<strong>о</strong>битн<strong>о</strong>му<br />

(тритире<strong>о</strong>идн<strong>о</strong>му) типу н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел<br />

— пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му и тул<strong>о</strong><strong>в</strong>ищный <strong>о</strong>тдел — ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му.<br />

В <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>х<strong>о</strong>дят<br />

три пер<strong>в</strong>ых п<strong>о</strong><strong>с</strong>т<strong>о</strong>р<strong>а</strong>льных <strong>с</strong>егмент<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зующих<br />

пер<strong>в</strong>ичн<strong>о</strong>-г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й — л<strong>а</strong>р<strong>в</strong><strong>а</strong>льный — <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong>,<br />

и <strong>о</strong>дин п<strong>о</strong><strong>с</strong>тл<strong>а</strong>р<strong>в</strong><strong>а</strong>льный— з<strong>а</strong>тыл<strong>о</strong>чный.<br />

Тул<strong>о</strong><strong>в</strong>ищные, ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие, <strong>с</strong>егменты<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленными <strong>с</strong>егмент<strong>а</strong>ми н<strong>о</strong>г<br />

III и IV (мез<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>) и <strong>в</strong><strong>с</strong>еми брюшными( <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ими)<br />

<strong>с</strong>егмент<strong>а</strong>ми; тел<strong>о</strong> Solifugomorpha<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит <strong>в</strong> <strong>о</strong>бщем из 16 <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1288—1289). Пр<strong>о</strong>пельтидий <strong>к</strong>рупный, я<strong>с</strong>н<strong>о</strong><br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>к</strong>рупных <strong>к</strong>лешне<strong>в</strong>идных хелицер,<br />

уч<strong>а</strong><strong>с</strong>т<strong>в</strong>ующих <strong>в</strong> <strong>а</strong><strong>к</strong>те р<strong>а</strong>змельчения (же<strong>в</strong><strong>а</strong>ния)<br />

пищи. Между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями хелицер<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я <strong>к</strong>рупн<strong>а</strong>я р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я труб<strong>к</strong><strong>а</strong> — rostrum,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящ<strong>а</strong>я из <strong>с</strong>ли<strong>в</strong>ших<strong>с</strong>я <strong>в</strong>ерхней губы <strong>с</strong><br />

пери<strong>с</strong>тыми щетин<strong>к</strong><strong>а</strong>ми н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце, <strong>о</strong>бр<strong>а</strong>зующими<br />

п<strong>о</strong>д<strong>о</strong>бие <strong>с</strong>ет<strong>к</strong>и, д<strong>в</strong>ух б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых л<strong>о</strong>п<strong>а</strong><strong>с</strong>тей,<br />

т<strong>а</strong><strong>к</strong>же <strong>с</strong><strong>о</strong> щетин<strong>к</strong><strong>а</strong>ми, и <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

дейт<strong>о</strong><strong>с</strong>тернум<strong>а</strong>. Педип<strong>а</strong>льпы <strong>к</strong>рупные, н<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные.<br />

Н<strong>о</strong>ги длинные, х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>; н<strong>о</strong>ги<br />

I <strong>в</strong>ып<strong>о</strong>лняют фун<strong>к</strong>цию <strong>о</strong><strong>с</strong>яз<strong>а</strong>ния и т<strong>о</strong>ньше<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных п<strong>а</strong>р. Н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х II—IV <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

брюш<strong>к</strong><strong>а</strong> имеют<strong>с</strong>я дых<strong>а</strong>тельные <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия<br />

тр<strong>а</strong>хейных <strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>в</strong>. Д<strong>о</strong>п<strong>о</strong>лнительн<strong>а</strong>я п<strong>а</strong>р<strong>а</strong><br />

<strong>с</strong>тигм между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г II и III я<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>о</strong><strong>в</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нием. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>с</strong>яти<br />

я<strong>в</strong><strong>с</strong>т<strong>в</strong>енных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; <strong>с</strong>едьм<strong>о</strong>й <strong>с</strong>егмент тел<strong>а</strong><br />

(рудимент<strong>а</strong>рный пер<strong>в</strong>ый <strong>с</strong>егмент брюш<strong>к</strong><strong>а</strong>)<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен <strong>в</strong> <strong>в</strong>иде неб<strong>о</strong>льш<strong>о</strong>й треуг<strong>о</strong>льн<strong>о</strong>й<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г IV; у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> ей <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует и м<strong>а</strong>лень<strong>к</strong>ий тергит,<br />

леж<strong>а</strong>щий п<strong>о</strong>з<strong>а</strong>ди тергит<strong>о</strong><strong>в</strong> д<strong>в</strong>ух грудных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичных мез<strong>о</strong>- и мет<strong>а</strong>пельтидиям<br />

(ри<strong>с</strong>. 1289). П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

(\ В. Б. Дубинин \)<br />

445 http://jurassic.ru/<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong> трехл<strong>о</strong>п<strong>а</strong><strong>с</strong>тн<strong>о</strong>м генит<strong>а</strong>льн<strong>о</strong>м<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong>е.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

един<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>г<strong>о</strong> <strong>о</strong>тряд<strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Solifugomorpha<br />

<strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> <strong>с</strong>ухих пу<strong>с</strong>тынных и<br />

<strong>с</strong>тепных, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>в</strong> г<strong>о</strong>рных ме<strong>с</strong>тн<strong>о</strong><strong>с</strong>тях тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти умеренн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>я<strong>с</strong><strong>о</strong><strong>в</strong> земн<strong>о</strong>г<strong>о</strong><br />

ш<strong>а</strong>р<strong>а</strong>. Преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> н<strong>о</strong>чные жи<strong>в</strong><strong>о</strong>тные;<br />

хищни<strong>к</strong>и. К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме и фил<strong>о</strong>гения.<br />

Д<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>тель<strong>с</strong>т<strong>в</strong><strong>о</strong>м <strong>с</strong><strong>а</strong>м<strong>о</strong>бытн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

я <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>г<strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждения <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Solifugomorpha<br />

<strong>о</strong>т примити<strong>в</strong>ных Pedipalpides я<strong>в</strong>ляет<strong>с</strong>я<br />

н<strong>а</strong>личие у <strong>с</strong><strong>о</strong>льпуг п<strong>а</strong>рных <strong>с</strong>тигм дых<strong>а</strong>тельных<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>с</strong>тернит<strong>а</strong>х IX—XI <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> (<strong>в</strong>т<strong>о</strong>р<strong>о</strong>й — чет<strong>в</strong>ертый <strong>с</strong>егменты брюш<strong>к</strong><strong>а</strong>).<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha <strong>с</strong> един<strong>с</strong>т<strong>в</strong>енным <strong>о</strong>тряд<strong>о</strong>м<br />

Solifugae пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>с</strong>реди других<br />

хелицер<strong>о</strong><strong>в</strong>ых <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленную группу, р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енную<br />

примити<strong>в</strong>ным Pedipalpides, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти<br />

<strong>о</strong>тряду Palpigrades и п<strong>о</strong>д<strong>о</strong>тряду Schizopeltidia<br />

(Uropygi), <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>ни и <strong>в</strong>едут <strong>с</strong><strong>в</strong><strong>о</strong>е н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong>.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha не р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енен другим<br />

н<strong>а</strong>земным п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зным, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые имеют<br />

б<strong>о</strong>лее п<strong>о</strong>зднее пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение и <strong>в</strong>едут <strong>с</strong><strong>в</strong><strong>о</strong>е н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong><br />

<strong>о</strong>т <strong>а</strong>мблипиг<strong>о</strong>м<strong>о</strong>рфных <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong> и <strong>о</strong>бъединяют<strong>с</strong>я<br />

<strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida — п<strong>а</strong>у<strong>к</strong><strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных,<br />

или пр<strong>о</strong><strong>с</strong><strong>о</strong>мных п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных (<strong>с</strong>м.<br />

<strong>с</strong>тр. 474 и <strong>с</strong>хему н<strong>а</strong> <strong>с</strong>тр. 384).<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha <strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>ет не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

черты р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>с</strong> <strong>к</strong>лещ<strong>а</strong>ми <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Acaromorpha,<br />

<strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong>, <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я ряд<strong>о</strong>м<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>енных <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тей <strong>с</strong>тр<strong>о</strong>ения и би<strong>о</strong>л<strong>о</strong>гии,<br />

<strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле <strong>с</strong><strong>о</strong>хр<strong>а</strong>нением нер<strong>а</strong><strong>с</strong>члененн<strong>о</strong>й<br />

л<strong>а</strong>р<strong>в</strong><strong>а</strong>льн<strong>о</strong>й т<strong>а</strong>гмы тел<strong>а</strong> (пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы), р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>й<br />

у <strong>к</strong>лещей н<strong>а</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му и пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Solifugomorpha пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен <strong>о</strong>дним<br />

<strong>о</strong>тряд<strong>о</strong>м Solifugae.

ОТРЯД SOLIFUGAE. ФАЛАНГИ, ИЛИ СОЛЬПУГИ<br />

Крупные хелицер<strong>о</strong><strong>в</strong>ые (длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>т 10 д<strong>о</strong><br />

70 мм) <strong>с</strong> чет<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным тритире<strong>о</strong>идным<br />

р<strong>а</strong><strong>с</strong>членением тел<strong>а</strong>. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>лит<strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> <strong>о</strong>дним<br />

грудным <strong>с</strong>егмент<strong>о</strong>м (пр<strong>о</strong>пельтидий); д<strong>а</strong>лее<br />

<strong>с</strong>ледуют три я<strong>в</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>тделенных друг <strong>о</strong>т<br />

друг<strong>а</strong> грудных <strong>с</strong>егмент<strong>а</strong> (т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Hexisopodidae Pocock <strong>о</strong>ни <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong>ме<strong>с</strong>те) и з<strong>а</strong>тем де<strong>с</strong>ять <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong><br />

(ри<strong>с</strong>. 1289). Спереди г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> у<strong>с</strong>ечен<strong>а</strong> и не<strong>с</strong>ет<br />

гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> <strong>с</strong> п<strong>а</strong>р<strong>о</strong>й гл<strong>а</strong>з или <strong>с</strong> пуч<strong>к</strong><strong>о</strong>м<br />

щетин<strong>о</strong><strong>к</strong> и шип<strong>о</strong><strong>в</strong>. Хелицеры <strong>о</strong>чень <strong>к</strong>рупные, <strong>с</strong><br />

т<strong>о</strong>л<strong>с</strong>тыми <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями и длинными п<strong>а</strong>льц<strong>а</strong>ми<br />

<strong>к</strong>лешни, <strong>с</strong> з<strong>а</strong>зубренными режущими <strong>к</strong>р<strong>а</strong>ями.<br />

Внутренняя п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть хелицер гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или<br />

<strong>с</strong> пр<strong>о</strong>д<strong>о</strong>льными <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong>ми <strong>о</strong>рг<strong>а</strong>н<strong>а</strong> <strong>с</strong>тре<strong>к</strong><strong>о</strong>т<strong>а</strong>ния.<br />

Снизу между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями хелицер п<strong>о</strong>мещ<strong>а</strong>ет<strong>с</strong>я<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>а</strong>я труб<strong>к</strong><strong>а</strong>. Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>к</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>му<br />

члени<strong>к</strong>у неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> прир<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<br />

<strong>к</strong>рупные треуг<strong>о</strong>льные т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп и б<strong>о</strong>лее<br />

мел<strong>к</strong>ие т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I (ри<strong>с</strong>. 1289). Педип<strong>а</strong>льпы<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены для <strong>с</strong>х<strong>в</strong><strong>а</strong>ты<strong>в</strong><strong>а</strong>ния и <strong>о</strong>щупы<strong>в</strong><strong>а</strong>ния<br />

д<strong>о</strong>бычи. Н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

п<strong>а</strong>льп р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я р<strong>а</strong>зличные шипы, <strong>а</strong><br />

у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, для удержи<strong>в</strong><strong>а</strong>ния <strong>с</strong><strong>а</strong>м<strong>к</strong>и<br />

при <strong>к</strong><strong>о</strong>пуляции — р<strong>а</strong>зличные <strong>с</strong>пеци<strong>а</strong>льные шипы<br />

и желези<strong>с</strong>тые <strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong>ч<strong>к</strong>и.<br />

Сегменты груди, н<strong>а</strong>именьшие п<strong>о</strong> <strong>о</strong>бъему,<br />

п<strong>о</strong><strong>к</strong>рыты мяг<strong>к</strong>ими хитин<strong>о</strong><strong>в</strong>ыми <strong>о</strong>б<strong>о</strong>л<strong>о</strong>ч<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong> тремя тергит<strong>а</strong>ми н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не.<br />

Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не им <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>уют три<br />

п<strong>а</strong>ры т<strong>в</strong>ердых пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong> — т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> х<strong>о</strong>дильных<br />

н<strong>о</strong>г (ри<strong>с</strong>. 1289). Четыре п<strong>а</strong>ры х<strong>о</strong>дильных н<strong>о</strong>г,<br />

причленяющие<strong>с</strong>я <strong>к</strong> <strong>с</strong>егмент<strong>а</strong>м груди, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

длинные и т<strong>о</strong>н<strong>к</strong>ие, реже <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Пер<strong>в</strong><strong>а</strong>я<br />

п<strong>а</strong>р<strong>а</strong> н<strong>о</strong>г <strong>о</strong>бычн<strong>о</strong> бы<strong>в</strong><strong>а</strong>ет <strong>с</strong><strong>а</strong>м<strong>о</strong>й т<strong>о</strong>н<strong>к</strong><strong>о</strong>й<br />

и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>й, <strong>с</strong> ше<strong>с</strong>тью члени<strong>к</strong><strong>а</strong>ми (<strong>в</strong>ертлуг р<strong>а</strong>зделен<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> члени<strong>к</strong><strong>а</strong>); эти н<strong>о</strong>ги игр<strong>а</strong>ют <strong>в</strong><strong>с</strong>п<strong>о</strong>м<strong>о</strong>г<strong>а</strong>тельную<br />

р<strong>о</strong>ль <strong>к</strong><strong>а</strong><strong>к</strong> <strong>о</strong>рг<strong>а</strong>ны <strong>о</strong><strong>с</strong>яз<strong>а</strong>ния. Н<strong>о</strong>ги<br />

II — <strong>с</strong> ше<strong>с</strong>тью члени<strong>к</strong><strong>а</strong>ми, <strong>а</strong> н<strong>о</strong>ги III и IV <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят<br />

из <strong>с</strong>еми члени<strong>к</strong><strong>о</strong><strong>в</strong>, т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> их <strong>в</strong>ертлуги<br />

р<strong>а</strong>зделены н<strong>а</strong> три ч<strong>а</strong><strong>с</strong>ти. Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех з<strong>а</strong>дних<br />

п<strong>а</strong>р н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>гтями: <strong>к</strong><strong>а</strong>ждый <strong>к</strong><strong>о</strong>г<strong>о</strong>ть н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>нце <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>тчлененным неб<strong>о</strong>льшим уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>о</strong>м<br />

— <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м (unguiculus). К<strong>о</strong>гти причленяют<strong>с</strong>я<br />

<strong>к</strong> <strong>о</strong>бщему прет<strong>а</strong>рзу, <strong>о</strong>бр<strong>а</strong>зующему <strong>с</strong>низу<br />

http://jurassic.ru/<br />

неб<strong>о</strong>льшую л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть (pulvillum). Члени<strong>к</strong>и н<strong>о</strong>г<br />

п<strong>о</strong><strong>к</strong>рыты б<strong>о</strong>льшим чи<strong>с</strong>л<strong>о</strong>м <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>, длинных<br />

щетин<strong>о</strong><strong>к</strong> и б<strong>о</strong>лее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их шип<strong>о</strong><strong>в</strong>. Н<strong>а</strong> бедр<strong>а</strong>х<br />

и н<strong>а</strong> пер<strong>в</strong><strong>о</strong>м и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>м <strong>в</strong>ертлуг<strong>а</strong>х н<strong>о</strong>г IV имеет<strong>с</strong>я<br />

<strong>о</strong>т д<strong>в</strong>ух д<strong>о</strong> пяти <strong>о</strong><strong>с</strong><strong>о</strong>бых треуг<strong>о</strong>льных при<strong>в</strong>е<strong>с</strong><strong>о</strong><strong>к</strong><br />

(malleoli) — <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>зн<strong>а</strong>чения<br />

(ри<strong>с</strong>. 1289).<br />

Брюш<strong>к</strong><strong>о</strong> <strong>к</strong>рупн<strong>о</strong>е, меш<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>е, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из<br />

де<strong>с</strong>яти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г IV и<br />

пер<strong>в</strong>ым <strong>с</strong>егмент<strong>о</strong>м брюш<strong>к</strong><strong>а</strong> — х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метн<strong>а</strong>я<br />

н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не треуг<strong>о</strong>льн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я я<strong>в</strong>ляет<strong>с</strong>я, <strong>в</strong>идим<strong>о</strong>, рудимент<strong>о</strong>м пер<strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong>. С<strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

<strong>к</strong><strong>а</strong>ждый <strong>с</strong>егмент <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чен тергит<strong>о</strong>м, п<strong>о</strong><strong>к</strong>рытым<br />

щетин<strong>к</strong><strong>а</strong>ми или шип<strong>а</strong>ми, <strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й —<br />

п<strong>а</strong>рными <strong>с</strong>тернит<strong>а</strong>ми, р<strong>а</strong>зделенными уз<strong>к</strong><strong>о</strong>й п<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong>й<br />

мяг<strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>жи <strong>в</strong>д<strong>о</strong>ль <strong>с</strong>редней линии тел<strong>а</strong>.<br />

И<strong>с</strong><strong>к</strong>лючение <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет <strong>с</strong>тернит пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

брюш<strong>к</strong><strong>а</strong>, н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие, при<strong>к</strong>рыт<strong>о</strong>е п<strong>о</strong>лулунными<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1289). П<strong>о</strong>д з<strong>а</strong>дними <strong>к</strong>р<strong>а</strong>ями<br />

<strong>в</strong>т<strong>о</strong>рых и третьих <strong>с</strong>тернит<strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>с</strong>редней линии<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>а</strong>рные дых<strong>а</strong>льц<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong><br />

при<strong>к</strong>рытые гребеш<strong>к</strong><strong>а</strong>ми или пери<strong>с</strong>тыми щетин<strong>к</strong><strong>а</strong>ми.<br />

П<strong>о</strong>д <strong>к</strong>р<strong>а</strong>ем чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> е<strong>с</strong>ть<br />

неп<strong>а</strong>рн<strong>о</strong>е рудимент<strong>а</strong>рн<strong>о</strong>е дых<strong>а</strong>льце. Н<strong>а</strong> п<strong>о</strong><strong>с</strong>леднем<br />

<strong>с</strong>егменте брюш<strong>к</strong><strong>а</strong> <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я з<strong>а</strong>днепр<strong>о</strong>х<strong>о</strong>дн<strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й дим<strong>о</strong>рфизм <strong>в</strong>ыр<strong>а</strong>жен<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong>. С<strong>а</strong>м<strong>к</strong>и яйце<strong>к</strong>л<strong>а</strong>дущие. Опл<strong>о</strong>д<strong>о</strong>т<strong>в</strong><strong>о</strong>рение<br />

<strong>с</strong>перм<strong>а</strong>т<strong>о</strong>ф<strong>о</strong>рн<strong>о</strong>е. К<strong>а</strong>рб<strong>о</strong>н — ныне. Де<strong>с</strong>ять<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне и <strong>о</strong>дн<strong>о</strong> (Protosolpugidae)<br />

<strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не.<br />

СЕМЕЙСТВО PROTOSOLPUGIDAE<br />

PETRUNKEVITCH, 1953<br />

Три <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных грудных <strong>с</strong>егмент<strong>а</strong> и т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>с</strong>емь <strong>с</strong>егмент<strong>о</strong><strong>в</strong> брюш<strong>к</strong><strong>а</strong>. Хелицеры <strong>с</strong> <strong>о</strong>чень т<strong>о</strong>л<strong>с</strong>тыми<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими, з<strong>а</strong><strong>о</strong><strong>с</strong>тренными<br />

п<strong>а</strong>льц<strong>а</strong>ми. Педип<strong>а</strong>льпы н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные, <strong>к</strong>рупные.<br />

Н<strong>о</strong>ги I зн<strong>а</strong>чительн<strong>о</strong> т<strong>о</strong>ньше <strong>о</strong><strong>с</strong>т<strong>а</strong>льных п<strong>а</strong>р<br />

н<strong>о</strong>г. Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong> тел<strong>а</strong> <strong>с</strong> хелицер<strong>а</strong>ми<br />

24 мм, длин<strong>а</strong> брюш<strong>к</strong><strong>а</strong> 12 мм. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

С. Амери<strong>к</strong>и. Один р<strong>о</strong>д — Protosolpuga Petrunkevitch,<br />

1913 (ри<strong>с</strong>. 1288).

КЛАСС ACAROMORPHA.КЛЕЩИ,<br />

ИЛИ ГНАТОСОМНЫЕ ХЕЛИЦЕРОВЫЕ<br />

В <strong>о</strong>тличие <strong>о</strong>т <strong>в</strong><strong>с</strong>ех других Chelicerata, р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые<br />

<strong>о</strong>рг<strong>а</strong>ны 1<br />

<strong>к</strong>лещей <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

тел<strong>а</strong> <strong>в</strong> <strong>в</strong>иде л<strong>о</strong>жн<strong>о</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>к</strong>и, или гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>о</strong>й т<strong>а</strong><strong>к</strong>же <strong>к</strong><strong>а</strong>питулум<strong>о</strong>м, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й или<br />

х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> тремя передними<br />

<strong>с</strong>егмент<strong>а</strong>ми: пре<strong>о</strong>р<strong>а</strong>льными (<strong>а</strong><strong>с</strong>г<strong>о</strong>п), <strong>в</strong>т<strong>о</strong>рым<br />

<strong>с</strong>егмент<strong>о</strong>м, не<strong>с</strong>ущим хелицеры, и третьим<br />

— педип<strong>а</strong>льпы.<br />

Сегмент<strong>а</strong>ция <strong>о</strong><strong>с</strong>т<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> тел<strong>а</strong>, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няющ<strong>а</strong>я<strong>с</strong>я<br />

у немн<strong>о</strong>гих, н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ньгх,<br />

ф<strong>о</strong>рм (п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Palaeacariformes, н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

Pediculoididea, Endeostigmata и др.),<br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует у п<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющег<strong>о</strong><br />

б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong>. О н<strong>а</strong>личии прежней<br />

<strong>с</strong>егмент<strong>а</strong>ции м<strong>о</strong>жн<strong>о</strong> <strong>с</strong>удить п<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жению<br />

щетин<strong>о</strong><strong>к</strong>, <strong>о</strong>бр<strong>а</strong>зующих н<strong>а</strong> теле б<strong>о</strong>лее или менее<br />

чет<strong>к</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женные п<strong>о</strong>перечные ряды, ин<strong>о</strong>гд<strong>а</strong><br />

п<strong>о</strong> <strong>с</strong>егмент<strong>а</strong>рн<strong>о</strong>му р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жению б<strong>о</strong>р<strong>о</strong>зд, щит<strong>о</strong><strong>в</strong><br />

и т. п. В предел<strong>а</strong>х <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Acaromorpha н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

тенденция <strong>к</strong> ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong>й реду<strong>к</strong>ции<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong>, <strong>а</strong> <strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, <strong>к</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чению<br />

тел<strong>а</strong> з<strong>а</strong> <strong>с</strong>чет и<strong>с</strong>чезн<strong>о</strong><strong>в</strong>ения мет<strong>а</strong><strong>с</strong><strong>о</strong>мы, <strong>а</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

Acariformes — <strong>о</strong>дн<strong>о</strong>г<strong>о</strong>-трех <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы. Н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ный — р<strong>а</strong>т<strong>о</strong>идный<br />

— тип р<strong>а</strong><strong>с</strong>членения <strong>с</strong>реди Acaromorpha<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>няют не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>к</strong>лещи Palaeacariformes,<br />

у <strong>к</strong><strong>о</strong>т<strong>о</strong>рых г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>а</strong>я л<strong>о</strong>п<strong>а</strong><strong>с</strong>ть и д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых <strong>с</strong>егмент<strong>а</strong><br />

(пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>) п<strong>о</strong><strong>к</strong>рыты щит<strong>о</strong>м (пр<strong>о</strong>пельтидий,<br />

<strong>с</strong>хиз<strong>о</strong>пельтидий), <strong>в</strong> т<strong>о</strong> <strong>в</strong>ремя <strong>к</strong><strong>а</strong><strong>к</strong><br />

<strong>с</strong>егменты н<strong>о</strong>г III—IV <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены 1<br />

. Вид<strong>о</strong>изменением<br />

эт<strong>о</strong>г<strong>о</strong> пер<strong>в</strong>ичн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> р<strong>а</strong><strong>с</strong>членения я<strong>в</strong>ляет<strong>с</strong>я<br />

тритире<strong>о</strong>идн<strong>о</strong>е р<strong>а</strong><strong>с</strong>членение тел<strong>а</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

других примити<strong>в</strong>ных <strong>к</strong>лещей (Nanorhestidae;<br />

ри<strong>с</strong>. 1339), при <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бляет<strong>с</strong>я<br />

(\ В. Б. Дубинин \)<br />

447<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бый щит, мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> для н<strong>о</strong>г III—IV.<br />

При д<strong>а</strong>льнейшем пр<strong>о</strong>це<strong>с</strong><strong>с</strong>е <strong>с</strong>лияния <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> <strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ет <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>идный (х<strong>а</strong>р<strong>а</strong><strong>к</strong>терный для<br />

<strong>о</strong>тряд<strong>а</strong> Acariformes) и <strong>а</strong>р<strong>а</strong>хн<strong>о</strong>идный типы 1<br />

р<strong>а</strong><strong>с</strong>­<br />

членения (х<strong>а</strong>р<strong>а</strong><strong>к</strong>терен для <strong>о</strong>тряд<strong>а</strong> Parasitiformes).<br />

Пер<strong>в</strong>ый тип <strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ет при <strong>с</strong>лиянии мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>с</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й <strong>в</strong> единую ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му,.<br />

<strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й — при <strong>с</strong>лиянии мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

и <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>м, <strong>к</strong><strong>о</strong>н<strong>в</strong>ергентн<strong>о</strong>м, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

«пр<strong>о</strong><strong>с</strong><strong>о</strong>мы» (Parasitiformes), н<strong>о</strong> при<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нении <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

Тел<strong>о</strong> <strong>к</strong>лещей <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из пер<strong>в</strong>ичны'х 13 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых три <strong>о</strong>бр<strong>а</strong>зуют гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му,<br />

д<strong>в</strong><strong>а</strong> (IV—V) — пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong>-<strong>с</strong><strong>о</strong>му, не<strong>с</strong>ущую д<strong>в</strong>е передние<br />

п<strong>а</strong>ры н<strong>о</strong>г, д<strong>в</strong><strong>а</strong> (VI—VII) — мет<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му,<br />

не<strong>с</strong>ущую д<strong>в</strong>е п<strong>а</strong>ры з<strong>а</strong>дних н<strong>о</strong>г, <strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льные<br />

ше<strong>с</strong>ть <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емую «брюш<strong>к</strong><strong>о</strong>м»; <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льных <strong>с</strong>егмент<strong>о</strong><strong>в</strong> м<strong>о</strong>жет <strong>в</strong><strong>а</strong>рьир<strong>о</strong><strong>в</strong><strong>а</strong>ть<br />

<strong>о</strong>т ше<strong>с</strong>ти (у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>к</strong>лещей) д<strong>о</strong><br />

<strong>с</strong>еми — де<strong>в</strong>яти (у Parhypochthonius и Pachygnathus)<br />

и д<strong>а</strong>же д<strong>о</strong> де<strong>с</strong>яти — <strong>о</strong>динн<strong>а</strong>дц<strong>а</strong>ти<br />

(Opitiocarina или Notostigmata). Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> ф<strong>о</strong>рм бы<strong>в</strong><strong>а</strong>ет <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т мет<strong>а</strong>­<br />

п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы 1<br />

http://jurassic.ru/<br />

п<strong>о</strong>перечн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие<br />

чег<strong>о</strong> тел<strong>о</strong> <strong>к</strong>лещей принят<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделять н<strong>а</strong> д<strong>в</strong><strong>а</strong><br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных <strong>о</strong>тдел<strong>а</strong>: передний — пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му,<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы и пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы, и<br />

<strong>в</strong><strong>а</strong>дний — ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и <strong>с</strong>лит<strong>о</strong>й <strong>с</strong> нею <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

у немн<strong>о</strong>гих ф<strong>о</strong>рм (н<strong>а</strong>пример, у Speleorchestes;<br />

ри<strong>с</strong>. 1339) гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> бы<strong>в</strong><strong>а</strong>ют <strong>в</strong>нешне <strong>о</strong>тделены<br />

друг <strong>о</strong>т друг<strong>а</strong>.

Приняты <strong>с</strong>ледующие <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чения ч<strong>а</strong><strong>с</strong>тей (т<strong>а</strong>г<strong>а</strong>) тел<strong>а</strong> "<strong>к</strong>лещей:<br />

I. Ч<strong>а</strong><strong>с</strong>ть тел<strong>а</strong>, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящ<strong>а</strong>я из р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> (хелидеры<br />

и педип<strong>а</strong>льпы) Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> I<br />

II. Ч<strong>а</strong><strong>с</strong>ть тел<strong>а</strong>, <strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й причленяют<strong>с</strong>я н<strong>о</strong>ги I и | Пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

II Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> J . |<br />

III. Ч<strong>а</strong><strong>с</strong>ть тел<strong>а</strong>, <strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й причленяют<strong>с</strong>я н<strong>о</strong>ги , | П<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> i<br />

III и IV Мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> I<br />

^ i Ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

IV. З<strong>а</strong>дний <strong>о</strong>тдел («брюш<strong>к</strong><strong>о</strong>») тел<strong>а</strong> Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> J /<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

Иди<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong>лещей п<strong>о</strong> н<strong>а</strong>б<strong>о</strong>ру <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ует<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди других п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных,<br />

н<strong>о</strong> <strong>с</strong>егменты, <strong>в</strong>х<strong>о</strong>дящие <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> ее, не <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

и не <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>тдельную ч<strong>а</strong><strong>с</strong>ть — т<strong>а</strong>гму<br />

— тел<strong>а</strong>, н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емую г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудью, и пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляемую<br />

з<strong>а</strong>дней брюшн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>ти —<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы других хелицер<strong>о</strong><strong>в</strong>ых.<br />

Хитин<strong>о</strong><strong>в</strong>ые п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы 1<br />

гидие<strong>в</strong><strong>а</strong> <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х <strong>к</strong>лещей Ixodidae и<br />

Rhagidiidae и т. п. У мн<strong>о</strong>гих <strong>а</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущих<br />

<strong>к</strong>лещей имеют<strong>с</strong>я пр<strong>о</strong><strong>с</strong>тые <strong>о</strong>дин<strong>о</strong>чные или д<strong>в</strong><strong>о</strong>йные<br />

гл<strong>а</strong>з<strong>а</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющие<strong>с</strong>я н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы. Хелицеры <strong>о</strong>бычн<strong>о</strong> д<strong>в</strong>ухтрех,<br />

реже четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>лешне<strong>в</strong>идные;<br />

<strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых групп<strong>а</strong>х <strong>о</strong>ни <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я <strong>с</strong>тилет<strong>о</strong>-<br />

тел<strong>а</strong> р<strong>а</strong>з<strong>в</strong>иты <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>в</strong>идными, при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленными для пр<strong>о</strong><strong>к</strong><strong>а</strong>лы<strong>в</strong><strong>а</strong>­<br />

р<strong>а</strong>зличн<strong>о</strong>; у Notostigmata <strong>о</strong>ни <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые, тип<strong>а</strong> ния жи<strong>в</strong><strong>о</strong>тных или р<strong>а</strong><strong>с</strong>тительных т<strong>к</strong><strong>а</strong>ней. У <strong>с</strong><strong>а</strong>м­<br />

т<strong>о</strong>н<strong>к</strong><strong>о</strong>й гр<strong>а</strong>нулир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й <strong>к</strong>ути<strong>к</strong>улы, у б<strong>о</strong>льшинц<strong>о</strong><strong>в</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> Mesostigmata п<strong>о</strong>д<strong>в</strong>ижный<br />

<strong>с</strong>т<strong>в</strong><strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Acariformes п<strong>о</strong><strong>к</strong>­ п<strong>а</strong>лец хелицер пре<strong>в</strong>р<strong>а</strong>щен <strong>в</strong> д<strong>о</strong>п<strong>о</strong>лнительный<br />

р<strong>о</strong><strong>в</strong>ы т<strong>о</strong>н<strong>к</strong>ие, эл<strong>а</strong><strong>с</strong>тичные и пр<strong>о</strong>зр<strong>а</strong>чные, ме<strong>с</strong>т<strong>а</strong>­ ' <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный <strong>о</strong>рг<strong>а</strong>н. Педип<strong>а</strong>льпы р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ми<br />

<strong>о</strong>бр<strong>а</strong>зующие пл<strong>о</strong>тные щит<strong>к</strong>и; н<strong>а</strong><strong>о</strong>б<strong>о</strong>р<strong>о</strong>т, у ют<strong>с</strong>я п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> хелицер; у де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

Oribatei,. Holothyroidea и мн<strong>о</strong>гих Mesostigmata Protacarus crani Hirst (ри<strong>с</strong>. 1341) <strong>о</strong>ни <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят<br />

тел<strong>о</strong> п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> т<strong>в</strong>ердыми з<strong>а</strong>щитными щит<strong>а</strong>ми, из <strong>с</strong>еми члени<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>а</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>к</strong>лещей —<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м. П<strong>о</strong><strong>в</strong>ерх­ не б<strong>о</strong>лее чем из ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>, чт<strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>н<strong>о</strong><strong>с</strong>ть<br />

<strong>к</strong>ути<strong>к</strong>улы м<strong>о</strong>жет быть гл<strong>а</strong>д<strong>к</strong><strong>о</strong>й или п<strong>о</strong><strong>к</strong>дит <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>с</strong>лияния basi- и telofemur. Перрыт<strong>о</strong>й<br />

б<strong>о</strong>лее или менее <strong>с</strong>л<strong>о</strong>жным <strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м; <strong>в</strong>ичн<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые, п<strong>о</strong>х<strong>о</strong>жие н<strong>а</strong> н<strong>о</strong>ги, педип<strong>а</strong>льпы<br />

мн<strong>о</strong>гие <strong>с</strong>тру<strong>к</strong>турные <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния — пр<strong>о</strong>из<strong>в</strong><strong>о</strong>д­ у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong>лещей бы<strong>в</strong><strong>а</strong>ют <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ид<strong>о</strong>изные<br />

<strong>к</strong>лет<strong>о</strong><strong>к</strong> гип<strong>о</strong>дермы. Н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти тел<strong>а</strong> менены и пре<strong>в</strong>р<strong>а</strong>щены <strong>в</strong> <strong>к</strong>лешне<strong>в</strong>идные х<strong>в</strong><strong>а</strong>т<strong>а</strong>­<br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong>пеци<strong>а</strong>льные железы пип<strong>о</strong>дертельные <strong>о</strong>рг<strong>а</strong>ны хищных Cheyletidae или, н<strong>а</strong>м<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждения. Щетин<strong>к</strong>и н<strong>а</strong> теле <strong>о</strong>б<strong>о</strong>р<strong>о</strong>т, бы<strong>в</strong><strong>а</strong>ют <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны д<strong>о</strong> не­<br />

и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тях у <strong>к</strong>лещей имеют р<strong>а</strong>зличную <strong>в</strong>еб<strong>о</strong>льших п<strong>а</strong>пилл.<br />

личину и ф<strong>о</strong>рму; <strong>о</strong>ни <strong>в</strong>ып<strong>о</strong>лняют фун<strong>к</strong>цию <strong>о</strong>р­ Личин<strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, имеют три п<strong>а</strong>ры<br />

г<strong>а</strong>н<strong>о</strong><strong>в</strong> <strong>о</strong><strong>с</strong>яз<strong>а</strong>ния, хем<strong>о</strong>рецепт<strong>о</strong>р<strong>о</strong><strong>в</strong>, мех<strong>а</strong>ниче­ х<strong>о</strong>дильных н<strong>о</strong>г, <strong>а</strong> нимфы и <strong>в</strong>зр<strong>о</strong><strong>с</strong>лые <strong>к</strong>лещи —<br />

<strong>с</strong><strong>к</strong><strong>о</strong>й з<strong>а</strong>щиты, <strong>о</strong>пределения <strong>с</strong>илы д<strong>а</strong><strong>в</strong>ления и четыре п<strong>а</strong>ры, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> г<strong>а</strong>лл<strong>о</strong><strong>в</strong>ые <strong>к</strong>лещи п<strong>о</strong>д<strong>о</strong>т­<br />

д<strong>в</strong>ижения <strong>с</strong>реды, з<strong>а</strong>щиты <strong>о</strong>т чрезмерн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыдеряд<strong>а</strong> Eriophyiformes (ри<strong>с</strong>. 1361) н<strong>а</strong> <strong>в</strong><strong>с</strong>ех ф<strong>а</strong>з<strong>а</strong>х<br />

ления <strong>в</strong><strong>о</strong>ды и т. п. Ф<strong>о</strong>рм<strong>а</strong> и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>же­ р<strong>а</strong>з<strong>в</strong>ития имеют т<strong>о</strong>ль<strong>к</strong><strong>о</strong> д<strong>в</strong>е п<strong>а</strong>ры н<strong>о</strong>г, <strong>а</strong> не<strong>к</strong><strong>о</strong>ния<br />

щетин<strong>о</strong><strong>к</strong> (х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер хет<strong>о</strong>т<strong>а</strong><strong>к</strong><strong>с</strong>ии и <strong>с</strong><strong>о</strong>т<strong>о</strong>рые Podapolipodidae — <strong>в</strong><strong>с</strong>ег<strong>о</strong> лишь <strong>о</strong>дну п<strong>а</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><br />

хет<strong>о</strong>м<strong>а</strong>) имеют б<strong>о</strong>льш<strong>о</strong>е зн<strong>а</strong>чение <strong>в</strong> ру. У личин<strong>о</strong><strong>к</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ции. Химиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong> <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong> (н<strong>а</strong>пример, Demodex bovis Stiles) н<strong>а</strong>блюд<strong>а</strong>ет­<br />

щетин<strong>о</strong><strong>к</strong> и ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> тел<strong>а</strong> у <strong>о</strong>т<strong>с</strong>я п<strong>о</strong>лн<strong>о</strong>е нед<strong>о</strong>р<strong>а</strong>з<strong>в</strong>итие н<strong>о</strong>г. Н<strong>о</strong>ги, <strong>к</strong><strong>а</strong><strong>к</strong> и педельных<br />

групп <strong>к</strong>лещей р<strong>а</strong>зличен: щетиндип<strong>а</strong>льпы, р<strong>а</strong><strong>с</strong>членяют<strong>с</strong>я н<strong>а</strong> ше<strong>с</strong>ть — <strong>с</strong>емь чле<strong>к</strong>и,<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>с</strong>тен<strong>к</strong>и их <strong>в</strong>нутреннег<strong>о</strong> <strong>к</strong><strong>а</strong>н<strong>а</strong>л<strong>а</strong>, ни<strong>к</strong><strong>о</strong><strong>в</strong>; у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> чи<strong>с</strong>л<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> м<strong>о</strong>­<br />

у <strong>к</strong>лещей <strong>о</strong>тряд<strong>а</strong> Acariformes (Sarcoptiformes жет уменьшить<strong>с</strong>я д<strong>а</strong>же д<strong>о</strong> <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> з<strong>а</strong><br />

и Trombidiiformes) <strong>в</strong> п<strong>о</strong>ляриз<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>м <strong>с</strong><strong>в</strong>ете <strong>с</strong>чет <strong>с</strong>лияния пер<strong>в</strong>ичяы'х члени<strong>к</strong><strong>о</strong><strong>в</strong> друг <strong>с</strong> дру­<br />

<strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>ют <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong><strong>о</strong> ц<strong>в</strong>етн<strong>о</strong>г<strong>о</strong> лучепрег<strong>о</strong>м: реже имеет ме<strong>с</strong>т<strong>о</strong> <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>е п<strong>о</strong>др<strong>а</strong>зделел<strong>о</strong>мления<br />

(<strong>с</strong><strong>в</strong>ечение белым ц<strong>в</strong>ет<strong>о</strong>м) («<strong>а</strong><strong>к</strong>тиние н<strong>а</strong> б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> члени<strong>к</strong><strong>о</strong><strong>в</strong> (д<strong>о</strong> 18). Л<strong>а</strong>п<strong>к</strong>и<br />

н<strong>о</strong>хитин») и х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong><strong>к</strong>р<strong>а</strong>ши<strong>в</strong><strong>а</strong>ют<strong>с</strong>я й<strong>о</strong>д<strong>о</strong>м, н<strong>о</strong>г <strong>в</strong> типичных <strong>с</strong>луч<strong>а</strong>ях не<strong>с</strong>ут <strong>о</strong>дин — д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>­<br />

т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>к</strong>лещей <strong>о</strong>тряд<strong>о</strong><strong>в</strong> Parasitiformes г<strong>о</strong>т<strong>к</strong><strong>а</strong>.<br />

и Opilioacarina <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>о</strong> щетин<strong>о</strong><strong>к</strong> <strong>о</strong>птиче<strong>с</strong><strong>к</strong>и<br />

Для <strong>в</strong><strong>с</strong>ех Acaromorpha, <strong>в</strong> пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong>л<strong>о</strong>ж­<br />

не<strong>а</strong><strong>к</strong>ти<strong>в</strong>н<strong>о</strong> и не <strong>о</strong><strong>к</strong>р<strong>а</strong>ши<strong>в</strong><strong>а</strong>ет<strong>с</strong>я й<strong>о</strong>д<strong>о</strong>м. Кр<strong>о</strong>ме<br />

н<strong>о</strong><strong>с</strong>ть пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителям <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Arachnida, х<strong>а</strong>р<strong>а</strong><strong>к</strong>­<br />

щетин<strong>о</strong><strong>к</strong>, <strong>в</strong> р<strong>а</strong>зных ме<strong>с</strong>т<strong>а</strong>х тел<strong>а</strong> и н<strong>о</strong>г имеют<strong>с</strong>я<br />

терн<strong>о</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие цельн<strong>о</strong>й грудины. Мет<strong>а</strong>мерн<strong>о</strong><br />

<strong>с</strong>пеци<strong>а</strong>льные <strong>о</strong>рг<strong>а</strong>ны чу<strong>в</strong><strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>в</strong>иде п<strong>с</strong>е<strong>в</strong>д<strong>о</strong><strong>с</strong>тиг-<br />

<strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нные <strong>с</strong>терниты груди <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я<br />

м<strong>а</strong>льных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>, <strong>с</strong>ен<strong>с</strong>илл или трих<strong>о</strong>б<strong>о</strong>трий,<br />

у <strong>к</strong>лещей Parasitiformes, т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>к</strong>лещей<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong><strong>о</strong>т<strong>о</strong>рых р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> <strong>с</strong>л<strong>о</strong>жн<strong>о</strong><br />

Acariformes грудные <strong>с</strong>терниты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют и<br />

у<strong>с</strong>тр<strong>о</strong>енные ч<strong>а</strong>ш<strong>к</strong><strong>о</strong><strong>в</strong>идных углублениях (б<strong>о</strong>три-<br />

з<strong>а</strong>менены <strong>к</strong>рупными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми х<strong>о</strong>дильных н<strong>о</strong>г,<br />

диях, или п<strong>с</strong>е<strong>в</strong>д<strong>о</strong><strong>с</strong>тигм<strong>а</strong>х), г<strong>а</strong>ллер<strong>о</strong><strong>в</strong><strong>а</strong> или р<strong>а</strong>-<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми тел<strong>а</strong> и <strong>о</strong>б-<br />

448<br />

http://jurassic.ru/

р<strong>а</strong>зующими <strong>о</strong><strong>с</strong><strong>о</strong>бые <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льные щиты. У р<strong>а</strong><strong>с</strong>члененных<br />

<strong>к</strong>лещей н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pachygnathidea,<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Rhodacaridae и др. <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я<br />

пер<strong>в</strong>ый (прегенит<strong>а</strong>льный) брюшн<strong>о</strong>й <strong>с</strong>егмент<br />

<strong>с</strong> -<strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующим н<strong>а</strong>б<strong>о</strong>р<strong>о</strong>м чу<strong>в</strong><strong>с</strong>т<strong>в</strong>ительных<br />

щетин<strong>о</strong><strong>к</strong>.<br />

Н<strong>а</strong>ружный п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>, <strong>а</strong> ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

и <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н трехл<strong>о</strong>п<strong>а</strong><strong>с</strong>тным п<strong>о</strong>л<strong>о</strong><strong>в</strong>ым<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong>м. У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>к</strong>лещей н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

реду<strong>к</strong>ция эт<strong>о</strong>г<strong>о</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>а</strong> и пре<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

ег<strong>о</strong> <strong>в</strong> пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>е щеле<strong>в</strong>идн<strong>о</strong>е генит<strong>а</strong>льн<strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> мн<strong>о</strong>гих <strong>к</strong>лещей<br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти тел<strong>а</strong> бы<strong>в</strong><strong>а</strong>ет <strong>в</strong>идн<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я<br />

щель, при<strong>к</strong>рыт<strong>а</strong>я <strong>к</strong><strong>о</strong>жными <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong>ми и <strong>о</strong>бычн<strong>о</strong><br />

неп<strong>а</strong>рным п<strong>о</strong>л<strong>о</strong><strong>в</strong>ым <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н<strong>о</strong>м (эпигинием).<br />

Дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>, имеющ<strong>а</strong>я <strong>в</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>е<br />

Acarina б<strong>о</strong>льш<strong>о</strong>е т<strong>а</strong><strong>к</strong><strong>с</strong><strong>о</strong>н<strong>о</strong>миче<strong>с</strong><strong>к</strong><strong>о</strong>е зн<strong>а</strong>чение,<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> <strong>в</strong> <strong>о</strong>дних групп<strong>а</strong>х (п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>х)<br />

тр<strong>а</strong>хеями и дых<strong>а</strong>тельными <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями (<strong>с</strong>тигм<strong>а</strong>ми),<br />

<strong>а</strong> у других <strong>о</strong>н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong><br />

или <strong>с</strong><strong>о</strong><strong>в</strong><strong>с</strong>ем <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; дых<strong>а</strong>ние у этих<br />

<strong>к</strong>лещей <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляет<strong>с</strong>я через т<strong>о</strong>н<strong>к</strong>ие п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

тел<strong>а</strong>. П<strong>о</strong>л<strong>о</strong>жение дых<strong>а</strong>тельных <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий и их<br />

<strong>с</strong>тр<strong>о</strong>ение <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны. У <strong>к</strong>лещей-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>це<strong>в</strong><br />

<strong>о</strong>тряд<strong>а</strong> Opilioacarina (Notostigmata)<br />

четыре п<strong>а</strong>ры <strong>с</strong>тигм р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> тергит<strong>а</strong>х<br />

пер<strong>в</strong>ых четырех <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. У <strong>к</strong>лещей<br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Holothyroidea (Parasitiformes)<br />

д<strong>в</strong>е п<strong>а</strong>ры <strong>с</strong>тигм, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передняя п<strong>а</strong>р<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>а</strong><br />

п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м тел<strong>а</strong> н<strong>а</strong>д т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г III.<br />

В других групп<strong>а</strong>х Parasitiformes (п<strong>о</strong>д<strong>о</strong>тряды<br />

Mesostigmata и Ixodides) имеет<strong>с</strong>я п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>й<br />

п<strong>а</strong>ре <strong>с</strong>тигм, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>о</strong>бширных<br />

перитрем<strong>а</strong>х п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м тел<strong>а</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не или<br />

<strong>в</strong>переди т<strong>а</strong>зи<strong>к</strong><strong>а</strong><strong>в</strong> н<strong>о</strong>г III (Mesostigmata), либ<strong>о</strong><br />

п<strong>о</strong>з<strong>а</strong>ди т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV (Ixodides). У <strong>а</strong><strong>к</strong><strong>а</strong>риф<strong>о</strong>рмных<br />

<strong>к</strong>лещей п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Trombidiiformes<br />

<strong>с</strong>тигмы <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

или между гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

(у Tarsonemidea <strong>о</strong>дн<strong>а</strong>-д<strong>в</strong>е п<strong>а</strong>ры <strong>с</strong>тигм <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>ме). У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>к</strong>лещей дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

У п<strong>а</strong>нцирных <strong>к</strong>лещей п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Oribatei<br />

(<strong>в</strong>зр<strong>о</strong><strong>с</strong>лые <strong>к</strong>лещи) <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> р<strong>а</strong>з<strong>в</strong>итием пл<strong>о</strong>тных<br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> тел<strong>а</strong> р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я тр<strong>а</strong>хеи, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я<br />

<strong>с</strong>тигм<strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г, и п<strong>о</strong>ри<strong>с</strong>тыми п<strong>о</strong>лями н<strong>а</strong> мн<strong>о</strong>гих<br />

уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>х тел<strong>а</strong>.<br />

В<strong>с</strong>е <strong>к</strong>лещи р<strong>а</strong>здельн<strong>о</strong>п<strong>о</strong>лы. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й дим<strong>о</strong>рфизм<br />

<strong>в</strong>ыр<strong>а</strong>жен р<strong>а</strong>зличн<strong>о</strong>. Опл<strong>о</strong>д<strong>о</strong>т<strong>в</strong><strong>о</strong>рение<br />

<strong>в</strong>нутреннее, <strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляет<strong>с</strong>я путем <strong>к</strong><strong>о</strong>пуляции<br />

или <strong>в</strong><strong>в</strong>едения <strong>с</strong>перм<strong>а</strong>т<strong>о</strong>ф<strong>о</strong>р<strong>о</strong><strong>в</strong>. Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>к</strong>лещи<br />

<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бны <strong>к</strong> п<strong>а</strong>ртен<strong>о</strong>генезу. С<strong>а</strong>м<strong>к</strong>и яйце<strong>к</strong>л<strong>а</strong>дущие<br />

или жи<strong>в</strong><strong>о</strong>р<strong>о</strong>дящие.<br />

Типичный и б<strong>о</strong>лее примити<strong>в</strong>ный жизненный<br />

ци<strong>к</strong>л х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я <strong>с</strong>мен<strong>о</strong>й ше<strong>с</strong>ти<br />

п<strong>о</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я ф<strong>а</strong>з: яйц<strong>а</strong><br />

(у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> из<strong>в</strong>е<strong>с</strong>тны <strong>о</strong>дн<strong>а</strong>-д<strong>в</strong>е<br />

эмбри<strong>о</strong>н<strong>а</strong>льные линь<strong>к</strong>и) — Личин<strong>к</strong>и — пр<strong>о</strong>т<strong>о</strong>нимфы<br />

— дейт<strong>о</strong>нимфы — трит<strong>о</strong>- или теле<strong>о</strong>нимфы<br />

и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> и <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong>. В<strong>о</strong><br />

мн<strong>о</strong>гих групп<strong>а</strong>х <strong>к</strong>лещей жизненные ци<strong>к</strong>лы<br />

м<strong>о</strong>гут быть б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>ны, м<strong>о</strong>гут<br />

<strong>в</strong>ып<strong>а</strong>д<strong>а</strong>ть не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые ф<strong>а</strong>зы мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>, <strong>а</strong> <strong>в</strong><strong>с</strong>е<br />

р<strong>а</strong>з<strong>в</strong>итие н<strong>о</strong><strong>с</strong>ит черты 1<br />

29 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 449<br />

http://jurassic.ru/<br />

глуб<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong>шедшей<br />

эмбри<strong>о</strong>низ<strong>а</strong>ции <strong>с</strong> я<strong>в</strong>лениями <strong>а</strong>н<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. Клещи<br />

<strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong><strong>в</strong><strong>с</strong>еме<strong>с</strong>тн<strong>о</strong>. Одни из них <strong>в</strong>едут<br />

н<strong>а</strong>земный или п<strong>о</strong>дземный, <strong>с</strong><strong>а</strong>пр<strong>о</strong>фитный, <strong>о</strong>бр<strong>а</strong>з<br />

жизни, или я<strong>в</strong>ляют<strong>с</strong>я хищни<strong>к</strong><strong>а</strong>ми, т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong><br />

другие жи<strong>в</strong>ут <strong>в</strong> пре<strong>с</strong>н<strong>о</strong>й или м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>в</strong><strong>о</strong>де,<br />

либ<strong>о</strong> п<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong> теле или д<strong>а</strong>же <strong>в</strong><strong>о</strong> <strong>в</strong>нутренних<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>х п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных жи<strong>в</strong><strong>о</strong>тных и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых;<br />

между этими <strong>к</strong>р<strong>а</strong>йними э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>ими<br />

тип<strong>а</strong>ми имеют<strong>с</strong>я <strong>в</strong><strong>с</strong>е<strong>в</strong><strong>о</strong>зм<strong>о</strong>жные перех<strong>о</strong>ды. Огр<strong>о</strong>мн<strong>о</strong>е<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> и <strong>о</strong><strong>с</strong><strong>о</strong>бей <strong>к</strong>лещей<br />

<strong>в</strong><strong>с</strong>треч<strong>а</strong>ет<strong>с</strong>я <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е ле<strong>с</strong><strong>о</strong><strong>в</strong> и <strong>о</strong>тч<strong>а</strong><strong>с</strong>ти<br />

н<strong>а</strong> дере<strong>в</strong>ьях, чем, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, и <strong>о</strong>бъя<strong>с</strong>няет<strong>с</strong>я<br />

н<strong>а</strong>х<strong>о</strong>ждение б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> чи<strong>с</strong>л<strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> (95)<br />

<strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре. Мел<strong>к</strong>ие р<strong>а</strong>змеры <strong>к</strong>лещей<br />

з<strong>а</strong>трудняют н<strong>а</strong>х<strong>о</strong>ждение их <strong>в</strong> б<strong>о</strong>лее дре<strong>в</strong>них<br />

<strong>о</strong>тл<strong>о</strong>жениях, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> из э<strong>о</strong>цен<strong>а</strong> Пендж<strong>а</strong>б<strong>а</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<br />

тир<strong>о</strong>глиф<strong>о</strong>идный <strong>к</strong>лещ Palaeotyroglyphus<br />

fossilis (Mani, 1945) (= Gamasus fossilis), из<br />

э<strong>о</strong>цен<strong>а</strong> У<strong>а</strong>й<strong>о</strong>минг<strong>а</strong> (Green River) — п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ий<br />

<strong>к</strong>лещ р<strong>о</strong>д<strong>а</strong> Ixodes, <strong>а</strong> <strong>в</strong> де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>их <strong>к</strong>р<strong>а</strong><strong>с</strong>ных<br />

пе<strong>с</strong>ч<strong>а</strong>ни<strong>к</strong><strong>а</strong>х Ш<strong>о</strong>тл<strong>а</strong>ндии Хир<strong>с</strong>т<strong>о</strong>м (Hirst,<br />

1923) <strong>о</strong>бн<strong>а</strong>ружен<strong>а</strong> цел<strong>а</strong>я групп<strong>а</strong> примити<strong>в</strong>ных<br />

<strong>к</strong>лещей четырех <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> (Nanorchestidae, Р<strong>а</strong>chygnathidae,<br />

Alicorhagiidae и Tydeidae), <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong> были <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong> <strong>о</strong>тне<strong>с</strong>ены<br />

<strong>к</strong> <strong>о</strong>дн<strong>о</strong>му <strong>в</strong>иду Protacarus crani Hirst (<strong>с</strong>м.<br />

<strong>с</strong>тр. 462). Кр<strong>о</strong>ме <strong>с</strong><strong>а</strong>мих <strong>к</strong>лещей, <strong>в</strong>" бурых углях<br />

Герм<strong>а</strong>нии (Heyden, 1860, 1862; Thomas, 1877;<br />

Trotter, 1899; Kiister, 1911) и <strong>в</strong> з<strong>а</strong>х<strong>о</strong>р<strong>о</strong>ненн<strong>о</strong>м<br />

<strong>с</strong>ф<strong>а</strong>гн<strong>о</strong><strong>в</strong><strong>о</strong>м б<strong>о</strong>л<strong>о</strong>те де<strong>с</strong>яти — тридц<strong>а</strong>титы<strong>с</strong>ячелетней<br />

д<strong>а</strong><strong>в</strong>н<strong>о</strong><strong>с</strong>ти <strong>в</strong> Англии (Harrison, 1926)<br />

н<strong>а</strong>йдены ли<strong>с</strong>тья <strong>с</strong><strong>в</strong>ыше 15 <strong>в</strong>ид<strong>о</strong><strong>в</strong> дере<strong>в</strong>ье<strong>в</strong> и <strong>к</strong>у<strong>с</strong>т<strong>а</strong>рни<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>с</strong> <strong>в</strong><strong>о</strong>йл<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>ыми или р<strong>о</strong>ж<strong>к</strong><strong>о</strong><strong>в</strong>идными<br />

г<strong>а</strong>лл<strong>а</strong>ми <strong>к</strong>лещей п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Eriophyiformes;<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и г<strong>а</strong>лл<strong>о</strong><strong>в</strong> г<strong>а</strong>ллиц, <strong>а</strong> <strong>в</strong>идим<strong>о</strong>, т<strong>а</strong><strong>к</strong>же и г<strong>а</strong>лл<strong>о</strong><strong>в</strong>идных<br />

<strong>к</strong>лещей из<strong>в</strong>е<strong>с</strong>тны из <strong>с</strong><strong>а</strong>рм<strong>а</strong>т<strong>а</strong> Ам<strong>в</strong>р<strong>о</strong><strong>с</strong>ие<strong>в</strong><strong>к</strong>и<br />

(юг У<strong>к</strong>р<strong>а</strong>ин<strong>с</strong><strong>к</strong><strong>о</strong>й ССР) (Пимен<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1954). Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>к</strong>лещей из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я <strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным<br />

р<strong>о</strong>д<strong>а</strong>м и <strong>в</strong>ид<strong>а</strong>м; т<strong>о</strong>ль<strong>к</strong><strong>о</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые из<br />

них пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют <strong>о</strong><strong>с</strong><strong>о</strong>бые и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емые ф<strong>о</strong>рмы.<br />

Интере<strong>с</strong>н<strong>о</strong>, чт<strong>о</strong> и <strong>к</strong>лещи из де<strong>в</strong><strong>о</strong>н<strong>а</strong> я<strong>в</strong>ляют<strong>с</strong>я<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителями <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> (<strong>с</strong>м.<br />

<strong>с</strong>тр. 462).<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и-<strong>с</strong> теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей <strong>с</strong><strong>к</strong>л<strong>о</strong>нн<strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть<br />

Acarina <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> из <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Arachnida, р<strong>а</strong><strong>с</strong>п<strong>а</strong>д<strong>а</strong>ющег<strong>о</strong><strong>с</strong>я н<strong>а</strong> <strong>с</strong>емь —<br />

де<strong>с</strong>ять п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> и мн<strong>о</strong>же<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>. Пет-

рун<strong>к</strong>е<strong>в</strong>ич (Petrunkevitch, 1949) предл<strong>о</strong>жил н<strong>о</strong><strong>в</strong>ую<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Arachnida, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

13 <strong>о</strong>тряд<strong>о</strong><strong>в</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных группируют<strong>с</strong>я<br />

<strong>в</strong> четыре п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>; <strong>о</strong>тряд Acari н<strong>а</strong>ряду <strong>с</strong> <strong>о</strong>тряд<strong>а</strong>ми<br />

Architarbi, Opiliones, Pseudoscorpiones,<br />

Scorpiones <strong>о</strong>тн<strong>о</strong><strong>с</strong>ит<strong>с</strong>я <strong>к</strong> п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>у Latigastra.<br />

Одн<strong>а</strong><strong>к</strong><strong>о</strong> уже Бэ<strong>к</strong>ер и У<strong>а</strong>рт<strong>о</strong>н (Baker and Wharton,<br />

1952) <strong>о</strong>тметили, чт<strong>о</strong> е<strong>с</strong>ли призн<strong>а</strong>ть <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию<br />

Петрун<strong>к</strong>е<strong>в</strong>ич<strong>а</strong>, т<strong>о</strong> пр<strong>а</strong><strong>в</strong>ильнее р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть<br />

Acarina <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>. Интере<strong>с</strong>ную <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>цию <strong>в</strong><strong>с</strong>ег<strong>о</strong><br />

н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Chelicerata предл<strong>о</strong>жил З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин<br />

(1952), <strong>к</strong><strong>о</strong>т<strong>о</strong>рый, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>я<strong>с</strong>ь н<strong>а</strong> и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниях<br />

Гр<strong>а</strong>ндж<strong>а</strong>н<strong>а</strong> и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>шири<strong>в</strong> их, п<strong>о</strong><br />

н<strong>а</strong>личию <strong>а</strong>низ<strong>о</strong>тр<strong>о</strong>пн<strong>о</strong>г<strong>о</strong> <strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong> («хитин<strong>а</strong>»)<br />

<strong>в</strong> щетин<strong>к</strong><strong>а</strong>х <strong>к</strong>лещей предл<strong>о</strong>жил р<strong>а</strong>зделить <strong>в</strong><strong>с</strong>ех<br />

<strong>к</strong>лещей Acarina н<strong>а</strong> три <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельных <strong>о</strong>тряд<strong>а</strong>:<br />

1-й <strong>о</strong>тряд н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих <strong>к</strong>лещей, Acariformes,<br />

<strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующий <strong>с</strong>умме п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> Sarcoptiformes,<br />

Trombidiiformes, Palaeacariformes, Oribatei<br />

и Tetrapodili (Eriophyiformes); 2-й <strong>о</strong>тряд<br />

г<strong>а</strong>м<strong>а</strong>з<strong>о</strong>идных <strong>к</strong>лещей, Parasitiformes, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующий<br />

<strong>о</strong>дн<strong>о</strong>именн<strong>о</strong>му п<strong>о</strong>д<strong>о</strong>тряду <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ующей<br />

<strong>с</strong>и<strong>с</strong>темы <strong>с</strong> <strong>в</strong><strong>к</strong>лючением <strong>в</strong> нег<strong>о</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong><br />

<strong>к</strong>лещей Holothyroidea; 3-й <strong>о</strong>тряд <strong>к</strong>лещей-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>це<strong>в</strong>,<br />

Opilioacarina, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из <strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Opilioacaridae. Эти три <strong>о</strong>тряд<strong>а</strong> д<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

чет<strong>к</strong><strong>о</strong> р<strong>а</strong>злич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тями пл<strong>а</strong>н<strong>а</strong><br />

<strong>с</strong>тр<strong>о</strong>ения (т<strong>а</strong>гм<strong>о</strong>зи<strong>с</strong><strong>а</strong> и р<strong>а</strong>з<strong>в</strong>ития, <strong>с</strong>тр<strong>о</strong>ения дых<strong>а</strong>тельн<strong>о</strong>й<br />

и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>с</strong>и<strong>с</strong>тем и т. п.). Одн<strong>а</strong><strong>к</strong><strong>о</strong><br />

у<strong>с</strong>т<strong>а</strong>н<strong>а</strong><strong>в</strong>ли<strong>в</strong><strong>а</strong>емые А. А. З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>а</strong>нным (1952)<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные <strong>о</strong>тн<strong>о</strong>шения <strong>о</strong>тдельных <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

<strong>к</strong>лещей <strong>с</strong> другими <strong>о</strong>тряд<strong>а</strong>ми п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных,<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нные п<strong>о</strong>чти преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> н<strong>а</strong> н<strong>а</strong>личии<br />

или н<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ии <strong>а</strong>низ<strong>о</strong>тр<strong>о</strong>пн<strong>о</strong>г<strong>о</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

<strong>в</strong>еще<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> щетин<strong>к</strong><strong>а</strong>х тел<strong>а</strong> и <strong>о</strong>пи<strong>с</strong><strong>а</strong>нные<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии эт<strong>о</strong>г<strong>о</strong> н<strong>о</strong><strong>в</strong>ые п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ы Chelicerata,<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют<strong>с</strong>я нед<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нными.<br />

В н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящей р<strong>а</strong>б<strong>о</strong>те мы <strong>в</strong>ыделяем <strong>в</strong><strong>с</strong>ех<br />

<strong>к</strong>лещей <strong>в</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельный <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Acaromorpha,<br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ющий три <strong>о</strong>тряд<strong>а</strong>, <strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>а</strong>нным (1952). Для <strong>о</strong>тряд<strong>а</strong> Acariformes,<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>о</strong>г<strong>о</strong> н<strong>а</strong>иб<strong>о</strong>льшим <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>м<br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых <strong>в</strong>ид<strong>о</strong><strong>в</strong> и групп н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих <strong>к</strong>лещей,<br />

приним<strong>а</strong>ет<strong>с</strong>я <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция, предл<strong>о</strong>женн<strong>а</strong>я<br />

В. Дубининым (1956).<br />

Ср<strong>а</strong><strong>в</strong>нение из<strong>в</strong>е<strong>с</strong>тных и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емых и <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

<strong>к</strong>лещей п<strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет, чт<strong>о</strong> <strong>в</strong> течение <strong>в</strong><strong>с</strong>ег<strong>о</strong><br />

длительн<strong>о</strong>г<strong>о</strong> и<strong>с</strong>т<strong>о</strong>риче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> пери<strong>о</strong>д<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ития<br />

н<strong>а</strong>земные ф<strong>о</strong>рмы <strong>с</strong>т<strong>о</strong>й<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няли примити<strong>в</strong>ные<br />

черты <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции и р<strong>а</strong>з<strong>в</strong>ития. Б<strong>о</strong>лее<br />

пр<strong>о</strong>гре<strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие имели <strong>в</strong>иды, перешедшие<br />

<strong>к</strong> хищниче<strong>с</strong>т<strong>в</strong>у и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> <strong>к</strong> п<strong>а</strong>р<strong>а</strong>зитизму<br />

н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях и жи<strong>в</strong><strong>о</strong>тных.<br />

Р<strong>а</strong><strong>с</strong>пределение <strong>в</strong><strong>о</strong> <strong>в</strong>ремени. Кле­<br />

щи из<strong>в</strong>е<strong>с</strong>тны 1<br />

<strong>с</strong> де<strong>в</strong><strong>о</strong>н<strong>а</strong>. Вер<strong>о</strong>ятн<strong>о</strong>, <strong>о</strong>ни были<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены п<strong>о</strong>чти <strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех ч<strong>а</strong><strong>с</strong>тях<br />

<strong>с</strong><strong>в</strong>ет<strong>а</strong> (ныне неиз<strong>в</strong>е<strong>с</strong>тны <strong>к</strong>лещи т<strong>о</strong>ль<strong>к</strong><strong>о</strong> из Ант<strong>а</strong>р<strong>к</strong>тиды).<br />

О шир<strong>о</strong><strong>к</strong><strong>о</strong>м их р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нении<br />

г<strong>о</strong><strong>в</strong><strong>о</strong>рит, н<strong>а</strong>пример, н<strong>а</strong>х<strong>о</strong>ждение <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

янт<strong>а</strong>ре <strong>в</strong>ид<strong>о</strong><strong>в</strong>, близ<strong>к</strong>их <strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным индий<strong>с</strong><strong>к</strong>им<br />

и я<strong>в</strong><strong>а</strong>н<strong>с</strong><strong>к</strong>им (<strong>о</strong>дин р<strong>о</strong>д). Об эт<strong>о</strong>м же <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует<br />

цир<strong>к</strong>умп<strong>о</strong>лярн<strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Oribatei.<br />

ОТРЯД ACAR1DIFORMES. НАСТОЯЩИЕ КЛЕЩИ<br />

В <strong>о</strong>тряде Acariformes н<strong>а</strong>блюд<strong>а</strong>ют<strong>с</strong>я три тип<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>членения тел<strong>а</strong>: <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>идн<strong>о</strong>е [тел<strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из<br />

пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы, не<strong>с</strong>ущей р<strong>о</strong>т<strong>о</strong><strong>в</strong>ые <strong>о</strong>рг<strong>а</strong>ны и д<strong>в</strong>е<br />

передние п<strong>а</strong>ры н<strong>о</strong>г, и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы •— тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>тдел<strong>а</strong> <strong>с</strong> д<strong>в</strong>умя з<strong>а</strong>дними п<strong>а</strong>р<strong>а</strong>ми н<strong>о</strong>г<br />

(б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> групп <strong>с</strong>м. н<strong>а</strong> ри<strong>с</strong>. 1290—1338)];<br />

тритире<strong>о</strong>идн<strong>о</strong>е {<strong>о</strong>т <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы б<strong>о</strong>лее или менее<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>егмент<strong>а</strong>, т. е. V и VI, <strong>о</strong>бр<strong>а</strong>зующих мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му:<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые Pachygnathidea, н<strong>а</strong>пример,<br />

Nanorchestidae (ри<strong>с</strong>. 1339) и др.], р<strong>а</strong>г<strong>о</strong>идн<strong>о</strong>е<br />

[<strong>с</strong>егмент н<strong>о</strong>г II п<strong>а</strong>ры (з<strong>а</strong>тыл<strong>о</strong>чный или<br />

<strong>о</strong><strong>к</strong>ципит<strong>а</strong>льный) ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен <strong>о</strong>т <strong>о</strong><strong>с</strong>т<strong>а</strong>льных<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong>ных; <strong>с</strong>егменты н<strong>о</strong>г III и IV п<strong>а</strong>р б<strong>о</strong>лее<br />

или менее я<strong>в</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong> р<strong>а</strong>згр<strong>а</strong>ничены друг <strong>о</strong>т друг<strong>а</strong><br />

и <strong>о</strong>т <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы: н<strong>а</strong>пример, Palaeacariformes].<br />

У .не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ф<strong>о</strong>рм н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>е<br />

п<strong>о</strong>лн<strong>о</strong>е <strong>с</strong>лияние <strong>в</strong><strong>с</strong>ех р<strong>а</strong>здел<strong>о</strong><strong>в</strong> (<strong>к</strong>р<strong>о</strong>ме гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы)<br />

, при <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м тел<strong>о</strong> <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ит<strong>с</strong>я м<strong>о</strong>н<strong>о</strong>литным<br />

и п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью теряет <strong>с</strong>леды <strong>с</strong>егмент<strong>а</strong>ции<br />

(Trombidii-Sarcoptiformes; Actinochitinosi)<br />

450<br />

http://jurassic.ru/<br />

(н<strong>а</strong>пример, <strong>в</strong>ы<strong>с</strong>шие Trombidiiformes; ри<strong>с</strong>. 1346<br />

и 1350—1356).<br />

Н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители Acariformes<br />

имеют мяг<strong>к</strong>ие п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>н<strong>с</strong>и<strong>с</strong>тенции <strong>с</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>й рельефн<strong>о</strong>й ми<strong>к</strong>р<strong>о</strong><strong>с</strong>тру<strong>к</strong>тур<strong>о</strong>й<br />

из ребрыше<strong>к</strong> или зерен, <strong>о</strong>бычн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющих<strong>с</strong>я<br />

ряд<strong>а</strong>ми. С<strong>к</strong>лер<strong>о</strong>тиз<strong>а</strong>ция п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong>я<strong>в</strong>ляет<strong>с</strong>я <strong>в</strong> п<strong>о</strong>лн<strong>о</strong>м <strong>о</strong>бъеме лишь при<br />

п<strong>о</strong><strong>с</strong>ледней линь<strong>к</strong>е, имея типичн<strong>о</strong> им<strong>а</strong>гин<strong>а</strong>льный<br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер. М<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ития <strong>с</strong>л<strong>о</strong>жный<br />

и т<strong>в</strong>ердый п<strong>а</strong>нцирь д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у Oribatei,<br />

х<strong>о</strong>тя <strong>о</strong>н н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> з<strong>а</strong>ч<strong>а</strong>т<strong>о</strong>чн<strong>о</strong>м<br />

<strong>в</strong>иде д<strong>а</strong>же у п<strong>о</strong><strong>с</strong>ледних нимф<strong>а</strong>льных ф<strong>а</strong>з<br />

этих <strong>к</strong>лещей. У <strong>в</strong><strong>с</strong>ех других групп Acariformes<br />

пл<strong>о</strong>тный хитин<strong>о</strong><strong>в</strong>ый п<strong>а</strong>нцирь у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм<br />

не р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я; п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелые <strong>к</strong>лещи эт<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>уют теле<strong>о</strong>нимф<strong>а</strong>льн<strong>о</strong>му<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янию Oribatei. Видим<strong>о</strong>,<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у <strong>в</strong>ы<strong>с</strong>ших п<strong>а</strong>нцирных <strong>к</strong>лещей н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>лный мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з д<strong>о</strong> им<strong>а</strong>гин<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>яния.

Щетин<strong>к</strong>и тел<strong>а</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризуют<strong>с</strong>я и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong>й<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><strong>с</strong>тью р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жения и п<strong>о</strong><strong>с</strong>т<strong>о</strong>ян<strong>с</strong>т<strong>в</strong><strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong>, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы и мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы. В течение мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong><br />

н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я ритмичн<strong>о</strong>е (п<strong>о</strong>ф<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>о</strong>е)<br />

приб<strong>а</strong><strong>в</strong>ление н<strong>о</strong><strong>в</strong>ых <strong>с</strong>егментных групп щетин<strong>о</strong><strong>к</strong><br />

(хет<strong>о</strong>мер<strong>о</strong><strong>в</strong>) н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ющее<br />

<strong>а</strong>н<strong>а</strong>м<strong>о</strong>рфный х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер инди<strong>в</strong>иду<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>ития. В<strong>с</strong>е щетин<strong>к</strong>и тел<strong>а</strong> и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей,<br />

<strong>а</strong> т<strong>а</strong><strong>к</strong>же их пр<strong>о</strong>из<strong>в</strong><strong>о</strong>дные (трих<strong>о</strong>б<strong>о</strong>трии, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и,<br />

зубцы хелицер, при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и и т. п.) <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют<br />

<strong>а</strong><strong>к</strong>тин<strong>о</strong>хитин<strong>о</strong><strong>в</strong>ым <strong>с</strong>тержнем, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующим<strong>с</strong>я<br />

<strong>о</strong>птиче<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>а</strong>низ<strong>о</strong>тр<strong>о</strong>пн<strong>о</strong><strong>с</strong>тью.<br />

Н<strong>о</strong>ги, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мны, умеренн<strong>о</strong>й<br />

длины и т<strong>о</strong>лщины 1<br />

. Они <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из <strong>с</strong>еми пер<strong>в</strong>ичных<br />

члени<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые м<strong>о</strong>гут <strong>с</strong>ли<strong>в</strong><strong>а</strong>ть<strong>с</strong>я<br />

друг <strong>с</strong> друг<strong>о</strong>м, и т<strong>о</strong>гд<strong>а</strong> н<strong>о</strong>г<strong>а</strong> <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ит<strong>с</strong>я четырех-ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й;<br />

у <strong>в</strong><strong>с</strong>ех Oribatei и Sarcoptiformes<br />

(s. lato) и б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> Trombididae<br />

зде<strong>с</strong>ь <strong>в</strong>иды <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я <strong>к</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>м, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленным<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, н<strong>о</strong> <strong>о</strong>бл<strong>а</strong>д<strong>а</strong>ют<br />

ряд<strong>о</strong>м <strong>о</strong>чень примити<strong>в</strong>ных черт <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции:<br />

н<strong>а</strong>личие <strong>с</strong>егмент<strong>а</strong>ции тел<strong>а</strong>, <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>членения<br />

н<strong>о</strong>г и п<strong>а</strong>льп, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нение мн<strong>о</strong>г<strong>о</strong>члени<strong>с</strong>тых<br />

хелицер и т. п. (<strong>с</strong>м. <strong>с</strong>тр. 462—466).<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Отряд<br />

Acariformes <strong>о</strong>бъединяет н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ные<br />

группы <strong>к</strong>лещей, <strong>о</strong>бн<strong>а</strong>ружи<strong>в</strong><strong>а</strong>ющих не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные черты <strong>с</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>ми Solifugomorpha<br />

и Scorpionomorpha (<strong>о</strong>тряд<strong>а</strong>ми Uropygi,<br />

Palpigradi и PseudOscorpionodea).<br />

Де<strong>в</strong><strong>о</strong>н — ныне. Пять п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong>: Palaeacariformes,<br />

Acaridiae, Oribatei, Trombidiiformes<br />

и Tetrapodili (В. Дубинин, 1954), из них пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители<br />

п<strong>о</strong><strong>с</strong>ледних четырех н<strong>а</strong>йдены<br />

т<strong>а</strong>зи<strong>к</strong>и неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми<br />

брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зуя <strong>с</strong>пл<strong>о</strong>шн<strong>о</strong>й <strong>к</strong><strong>о</strong><strong>к</strong>-<br />

С<strong>о</strong>-<strong>с</strong>терн<strong>а</strong>льный <strong>с</strong><strong>к</strong>елет: н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие <strong>с</strong>терниты<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х л<strong>а</strong>п<strong>о</strong><strong>к</strong> имеет<strong>с</strong>я <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>о</strong><strong>к</strong><br />

(Oribatei, Sarcoptiformes), <strong>к</strong><strong>о</strong>т<strong>о</strong>рый у Trombidiiformes<br />

имеет <strong>в</strong>ид эмп<strong>о</strong>дия, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> пери<strong>с</strong>т<strong>о</strong>г<strong>о</strong><br />

или <strong>с</strong>л<strong>о</strong>жн<strong>о</strong> р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>г<strong>о</strong>; ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong>ершинные<br />

п<strong>а</strong>рные щетин<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong><br />

п<strong>а</strong>рные <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и (ри<strong>с</strong>. 1297).<br />

Тр<strong>а</strong>хейн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> у низших Acariformes,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у Oribatei <strong>о</strong>н<strong>а</strong><br />

<strong>в</strong><strong>о</strong>зни<strong>к</strong><strong>а</strong>ет <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong> <strong>с</strong> р<strong>а</strong>з<strong>в</strong>итием пл<strong>о</strong>тн<strong>о</strong>г<strong>о</strong><br />

хитин<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> п<strong>а</strong>нциря, т. е. т<strong>о</strong>ль<strong>к</strong><strong>о</strong> при пре<strong>в</strong>р<strong>а</strong>щении<br />

<strong>в</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лую «п<strong>а</strong>нцирную» ф<strong>а</strong>зу. У Trombidiiformes<br />

тр<strong>а</strong>хейн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я<br />

н<strong>а</strong>личием д<strong>в</strong>ух п<strong>а</strong>р г<strong>о</strong>л<strong>о</strong><strong>в</strong>ных тр<strong>а</strong>хей, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я<br />

<strong>с</strong>тигм<strong>а</strong>ми н<strong>а</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии хелицер.<br />

П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>те<strong>к</strong><strong>а</strong>ет п<strong>о</strong><br />

типу тринимф<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>; у мн<strong>о</strong>гих<br />

Sarcoptiformes (s. lato), <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong>, дейт<strong>о</strong>нимф<strong>а</strong><br />

либ<strong>о</strong> теряет <strong>с</strong><strong>в</strong><strong>о</strong>и <strong>в</strong>егет<strong>а</strong>ти<strong>в</strong>ные фун<strong>к</strong>ции,<br />

пре<strong>в</strong>р<strong>а</strong>щ<strong>а</strong>я<strong>с</strong>ь <strong>в</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>елительн<strong>о</strong>г<strong>о</strong> или п<strong>о</strong><strong>к</strong><strong>о</strong>ющег<strong>о</strong><strong>с</strong>я<br />

гип<strong>о</strong>пу<strong>с</strong><strong>а</strong>, либ<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> и<strong>с</strong>чез<strong>а</strong>ет из<br />

ци<strong>к</strong>л<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ития. При р<strong>а</strong>з<strong>в</strong>итии чет<strong>к</strong><strong>о</strong> пр<strong>о</strong><strong>с</strong>лежи<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

я<strong>в</strong>ления <strong>а</strong>н<strong>о</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. В э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>м<br />

и би<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>м <strong>о</strong>тн<strong>о</strong>шениях <strong>в</strong>иды<br />

Acariformes чрез<strong>в</strong>ыч<strong>а</strong>йн<strong>о</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны. Н<strong>а</strong>иб<strong>о</strong>лее<br />

примити<strong>в</strong>ные ф<strong>о</strong>рмы жи<strong>в</strong>ут <strong>в</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>тных<br />

<strong>с</strong>л<strong>о</strong>ях п<strong>о</strong>ч<strong>в</strong>ы и <strong>в</strong> п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е из гниющих<br />

р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>, б<strong>о</strong>лее же <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

и измененные <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> пре<strong>с</strong>ных<br />

и м<strong>о</strong>р<strong>с</strong><strong>к</strong>их <strong>в</strong><strong>о</strong>д<strong>а</strong>х, <strong>а</strong> т<strong>а</strong><strong>к</strong>же п<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>тениях и жи<strong>в</strong><strong>о</strong>тных. Н<strong>а</strong>иб<strong>о</strong>льшее <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> Acariformes из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

янт<strong>а</strong>ря; н<strong>а</strong>йденные зде<strong>с</strong>ь ф<strong>о</strong>рмы <strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong>м<br />

чи<strong>с</strong>ле <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят<strong>с</strong>я <strong>к</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременным р<strong>о</strong>д<strong>а</strong>м<br />

и д<strong>а</strong>же <strong>в</strong>ид<strong>а</strong>м. С<strong>а</strong>мые дре<strong>в</strong>ние Acariformes из<strong>в</strong>е<strong>с</strong>тны<br />

из де<strong>в</strong><strong>о</strong>н<strong>а</strong> Ш<strong>о</strong>тл<strong>а</strong>ндии; <strong>в</strong><strong>с</strong>треченные<br />

1<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

ПОДОТРЯД ACARIDIAE.<br />

САРКОПТОИДН'ЫЕ КЛЕЩИ<br />

(Sarcoptiformes)<br />

Р<strong>а</strong><strong>с</strong>членение тел<strong>а</strong> <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>идн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

нежные, гл<strong>а</strong>д<strong>к</strong>ие, т<strong>о</strong>н<strong>к</strong><strong>о</strong>штрих<strong>о</strong><strong>в</strong><strong>а</strong>нные или<br />

<strong>с</strong> зерни<strong>с</strong>т<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й. С<strong>к</strong>лер<strong>о</strong>тизир<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

щиты, е<strong>с</strong>ли <strong>о</strong>ни имеют<strong>с</strong>я, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong>; <strong>о</strong>дин пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий,<br />

н<strong>а</strong>иб<strong>о</strong>лее п<strong>о</strong><strong>с</strong>т<strong>о</strong>янный, <strong>о</strong>дин <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий<br />

и <strong>о</strong>дн<strong>а</strong> — д<strong>в</strong>е п<strong>а</strong>ры <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых; <strong>о</strong>чень<br />

ред<strong>к</strong><strong>о</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> цели<strong>к</strong><strong>о</strong>м <strong>с</strong><strong>к</strong>лер<strong>о</strong>тизируют<strong>с</strong>я<br />

(р<strong>о</strong>д Gohieria Oudemans из <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Glycyphagidae).<br />

Трих<strong>о</strong>б<strong>о</strong>трии <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я, причленяет<strong>с</strong>я н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце<br />

тел<strong>а</strong>, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>идн<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; <strong>к</strong><strong>а</strong>мер<strong>о</strong><strong>с</strong>т<strong>о</strong>м,<br />

е<strong>с</strong>ли <strong>о</strong>н е<strong>с</strong>ть, при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет <strong>о</strong>бычн<strong>о</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы 1<br />

. Хелицеры м<strong>а</strong>лень<strong>к</strong>ие,<br />

<strong>к</strong>лешне<strong>в</strong>идные, д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Педип<strong>а</strong>льпы<br />

пр<strong>о</strong><strong>с</strong>тые, д<strong>в</strong>ух-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Н<strong>о</strong>ги<br />

х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong> <strong>с</strong> пятью <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными члени<strong>к</strong><strong>а</strong>ми.<br />

К<strong>о</strong><strong>к</strong>-<strong>с</strong>ы <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны 1<br />

тел<strong>а</strong> и бы<strong>в</strong><strong>а</strong>ют пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены лишь<br />

уз<strong>к</strong>ими <strong>с</strong><strong>к</strong>лерит<strong>а</strong>ми — эпимер<strong>а</strong>ми и эпимерит<strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>зующими <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>терный <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong>-<strong>с</strong>терн<strong>а</strong>льный<br />

<strong>с</strong><strong>к</strong>елет. Кл<strong>а</strong>п<strong>а</strong>ны п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий<br />

т<strong>о</strong>н<strong>к</strong>ие, нежные, не имеют щетин<strong>о</strong><strong>к</strong>.<br />

Оф<strong>о</strong>рмленных дых<strong>а</strong>тельных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> нет; дых<strong>а</strong>ние<br />

<strong>о</strong><strong>с</strong>уще<strong>с</strong>т<strong>в</strong>ляет<strong>с</strong>я неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> через<br />

т<strong>о</strong>н<strong>к</strong>ие п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong>.<br />

Мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные <strong>в</strong>иды <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е,<br />

<strong>в</strong> <strong>с</strong><strong>к</strong><strong>о</strong>плениях р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> и п<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> жи<strong>в</strong><strong>о</strong>тных или <strong>в</strong> их <strong>в</strong>нутренних<br />

<strong>о</strong>рг<strong>а</strong>н<strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Семь н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>:<br />

Pediculoiodidea, Anoetidea, Canestrinidea, Sarcoptidea,<br />

Analgesidea, Listrophoridea и Acaridea;<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

п<strong>о</strong><strong>с</strong>леднее.<br />

451 29*<br />

http://jurassic.ru/

НАДСЕМЕЙСТВО ACARIDEA.<br />

ХЛЕБНЫЕ (АМБАРНЫЕ) КЛЕЩИ<br />

(Tyroglyphoidea Auctorum) 1<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы т<strong>о</strong>н<strong>к</strong>ие, нежные, гл<strong>а</strong>д<strong>к</strong>ие, зерни<strong>с</strong>тые<br />

или <strong>с</strong>л<strong>а</strong>б<strong>о</strong>шип<strong>о</strong><strong>в</strong><strong>а</strong>тые, <strong>к</strong>р<strong>а</strong>йне ред<strong>к</strong><strong>о</strong> <strong>с</strong> т<strong>о</strong>н<strong>к</strong><strong>о</strong>й<br />

б<strong>о</strong>лее или менее п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>й штрих<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й.<br />

Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong><strong>о</strong>мп<strong>а</strong><strong>к</strong>тн<strong>а</strong>я, м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я, <strong>о</strong>т<strong>к</strong>рыт<strong>а</strong>я,<br />

причленяет<strong>с</strong>я н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong>.<br />

Н<strong>о</strong>ги г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мные, у <strong>о</strong>б<strong>о</strong>их п<strong>о</strong>л<strong>о</strong><strong>в</strong> р<strong>а</strong>з<strong>в</strong>иты примерн<strong>о</strong><br />

<strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>днржи<strong>в</strong>ущие, <strong>с</strong><strong>а</strong>пр<strong>о</strong>би<strong>о</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>в</strong>иды 1<br />

,<br />

<strong>о</strong>бычн<strong>о</strong> <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong> <strong>с</strong><strong>к</strong><strong>о</strong>плениях д<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>о</strong>чн<strong>о</strong><br />

<strong>в</strong>л<strong>а</strong>жных или гниющих р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>иды жи<strong>в</strong>ут н<strong>а</strong> ли<strong>с</strong>тьях р<strong>а</strong><strong>с</strong>тений<br />

и <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е. Р<strong>а</strong><strong>с</strong><strong>с</strong>елительны'е гип<strong>о</strong>пу<strong>с</strong>ы шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

и<strong>с</strong>п<strong>о</strong>льзуют эпиз<strong>о</strong><strong>о</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бы р<strong>а</strong><strong>с</strong><strong>с</strong>еления<br />

<strong>с</strong> п<strong>о</strong>м<strong>о</strong>щью н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых или п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных<br />

жи<strong>в</strong><strong>о</strong>тных.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>о</strong>дн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО ACARIDAE LATREILLE, 1802<br />

(Tyroglyphidae Donnadieu, 1868; Parasitidae<br />

Mani, 1945, 1946)<br />

Щетин<strong>к</strong>и иди<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong><strong>с</strong>егд<strong>а</strong> гл<strong>а</strong>д<strong>к</strong>ие, <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong>идные<br />

или иг<strong>о</strong>льч<strong>а</strong>тые. С<strong>а</strong>мцы 1<br />

<strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми<br />

<strong>а</strong>н<strong>а</strong>льными и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong> д<strong>в</strong>умя<br />

п<strong>а</strong>р<strong>а</strong>ми т<strong>а</strong>рз<strong>а</strong>льных <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ных при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong>.<br />

С<strong>а</strong>м<strong>к</strong>и без эпигиния; генит<strong>а</strong>льные щуп<strong>а</strong>льцы 1<br />

п<strong>а</strong>льце<strong>в</strong>идные (ри<strong>с</strong>. 1290). Не<strong>о</strong>ген — ныне.<br />

Б<strong>о</strong>льш<strong>о</strong>е чи<strong>с</strong>л<strong>о</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны р<strong>о</strong>ды Ac&rus<br />

Linnaeus, 1758 (Tyroglyphus Latreille, 1796) из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), Туroglyphites<br />

Pampaloni, 1902 из ми<strong>о</strong>цен<strong>а</strong> Сицилии<br />

и Palaeotyroglyphus W. Dubinin, nom. nov.<br />

(= Gamasus Mani, 1945, 1946). Тип р<strong>о</strong>д<strong>а</strong>—<br />

Gamasus fossilis Mani, 1945; личин<strong>к</strong><strong>а</strong> (<strong>а</strong> не<br />

пр<strong>о</strong>т<strong>о</strong>нимф<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> пишут Mani, 1945 и George,<br />

1952) (ри<strong>с</strong>. 1290). Длин<strong>а</strong> тел<strong>а</strong> 0,24 мм. Не<strong>о</strong>ген<br />

(?) Пендж<strong>а</strong>б<strong>а</strong>.<br />

ПОДОТРЯД ORIBATEI.<br />

ПАНЦИРНЫЕ КЛЕЩИ<br />

Р<strong>а</strong><strong>с</strong>членение тел<strong>а</strong> <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>идн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>, н<strong>о</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong><strong>к</strong>рыт<strong>а</strong> п<strong>о</strong>д <strong>о</strong>бширным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы и причленяет<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>м<br />

углублении <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

— <strong>к</strong><strong>а</strong>мер<strong>о</strong><strong>с</strong>т<strong>о</strong>ме.<br />

Ри<strong>с</strong>. 1290—1297. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Acaridae,<br />

1290. Palaeotyroglyphus fossilis (Mani); <strong>о</strong>бщий <strong>в</strong>ид, X 100,0,<br />

не<strong>о</strong>ген, Индия (George, 1952). 1291. Trhqpochthonius tectorum<br />

(Berlese); <strong>о</strong>бщий <strong>в</strong>ид, X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1882). 1292.<br />

Brachychthonius brevls (Michael); <strong>о</strong>бщий <strong>в</strong>ид, x 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Hanmer, 1952). 1293. Camisia spinifer (Koch); <strong>о</strong>бщий <strong>в</strong>ид, X110.0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1887). 1294. Nothrus palustris Koch; <strong>о</strong>бщий <strong>в</strong>ид,<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых ф<strong>о</strong>рм, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

чрез<strong>в</strong>ыч<strong>а</strong>йн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны и темн<strong>о</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>шены,<br />

<strong>о</strong>бр<strong>а</strong>зуя пл<strong>о</strong>тный п<strong>а</strong>нцирь (ри<strong>с</strong>.<br />

1298—1338); у низших <strong>о</strong>риб<strong>а</strong>тид (ри<strong>с</strong>. 1292)<br />

т<strong>а</strong><strong>к</strong><strong>о</strong>й п<strong>а</strong>нцирь р<strong>а</strong>з<strong>в</strong>ит менее <strong>с</strong>ильн<strong>о</strong>, и т<strong>о</strong>гд<strong>а</strong><br />

п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> бы<strong>в</strong><strong>а</strong>ют <strong>к</strong><strong>о</strong>жи<strong>с</strong>тыми. У неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых<br />

ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>нными. Н<strong>а</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>ме<br />

у <strong>в</strong><strong>с</strong>ех ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития имеют<strong>с</strong>я трих<strong>о</strong>б<strong>о</strong>трии, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>о</strong>гружены 1<br />

<strong>в</strong> <strong>с</strong>л<strong>о</strong>жн<strong>о</strong> у<strong>с</strong>тр<strong>о</strong>енные<br />

ч<strong>а</strong>шеч<strong>к</strong>и — б<strong>о</strong>тридии (ри<strong>с</strong>. 1297); <strong>в</strong>еличин<strong>а</strong><br />

и ф<strong>о</strong>рм<strong>а</strong> трих<strong>о</strong>б<strong>о</strong>трии <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> изменчи<strong>в</strong>ы. Педип<strong>а</strong>льпы<br />

пр<strong>о</strong><strong>с</strong>тые, четырех-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы* у мн<strong>о</strong>гих<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пр<strong>о</strong>д<strong>о</strong>льные <strong>к</strong>или (ри<strong>с</strong>.<br />

1303, 1307), ребр<strong>а</strong> (ри<strong>с</strong>. 1316) или <strong>к</strong>рупные<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и. Н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы м<strong>о</strong>гут р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ть<strong>с</strong>я<br />

р<strong>а</strong>зличные пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тые <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты —<br />

те<strong>к</strong>т<strong>о</strong>педии (Damaeidae; ри<strong>с</strong>. 1301), <strong>а</strong> н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х<br />

н<strong>о</strong>т<strong>о</strong>г<strong>а</strong><strong>с</strong>тр<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> у <strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Notaspididae (ри<strong>с</strong>. 1324—1336) р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идные неп<strong>о</strong>д<strong>в</strong>ижные (ри<strong>с</strong>. 1331) или п<strong>о</strong>д<strong>в</strong>ижные<br />

(ри<strong>с</strong>. 1335) пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и — птер<strong>о</strong>м<strong>о</strong>рфы.<br />

В<strong>с</strong>е эти <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>в</strong>ып<strong>о</strong>лняют фун<strong>к</strong>ции з<strong>а</strong>щиты<br />

н<strong>о</strong>г и <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> их <strong>с</strong><strong>о</strong>членений <strong>о</strong>т з<strong>а</strong>грязнения<br />

или п<strong>о</strong><strong>в</strong>реждения. Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>иды <strong>в</strong> течение<br />

<strong>в</strong><strong>с</strong>ей жизни <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>дин<strong>о</strong>чные ш<strong>к</strong>ур<strong>к</strong>и предыдущих<br />

ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития (ри<strong>с</strong>. 1295), <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong> чем<br />

у них р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong>пеци<strong>а</strong>льные при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бления<br />

для з<strong>а</strong><strong>к</strong>репления этих ш<strong>к</strong>ур<strong>о</strong><strong>к</strong>.<br />

Н<strong>о</strong>ги х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. К<strong>о</strong><strong>к</strong><strong>с</strong>ы <strong>с</strong>литы <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми<br />

брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong>; эпимеры их<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены зде<strong>с</strong>ь грубыми <strong>в</strong><strong>а</strong>ли<strong>к</strong><strong>а</strong>ми, леж<strong>а</strong>щими<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> друг другу п<strong>о</strong>пере<strong>к</strong> тел<strong>а</strong><br />

и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я <strong>с</strong> хитин<strong>о</strong><strong>в</strong>ым <strong>к</strong><strong>о</strong>льц<strong>о</strong>м,<br />

<strong>о</strong><strong>к</strong>руж<strong>а</strong>ющим <strong>к</strong><strong>а</strong>мер<strong>о</strong><strong>с</strong>т<strong>о</strong>м. Н<strong>а</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>х л<strong>а</strong>п<strong>о</strong><strong>к</strong> у<br />

<strong>в</strong><strong>с</strong>ех неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития имеет<strong>с</strong>я п<strong>о</strong><br />

<strong>о</strong>дн<strong>о</strong>му <strong>к</strong>рупн<strong>о</strong>му <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>у, <strong>а</strong> у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых <strong>к</strong>лещей<br />

п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>му-трем «<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>м» (ри<strong>с</strong>. 1293,<br />

1297).<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й дим<strong>о</strong>рфизм не <strong>в</strong>ыр<strong>а</strong>жен. Генит<strong>а</strong>льных<br />

при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong> три п<strong>а</strong>ры. Кл<strong>а</strong>п<strong>а</strong>не п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий прям<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы<br />

<strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> хитиниз<strong>о</strong><strong>в</strong><strong>а</strong>ны, <strong>о</strong><strong>к</strong>ружены хитин<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

р<strong>а</strong>м<strong>к</strong><strong>о</strong>й (ри<strong>с</strong>. 1305). С<strong>а</strong>м<strong>к</strong>и имеют <strong>о</strong>чень<br />

длинный, теле<strong>с</strong><strong>к</strong><strong>о</strong>пиче<strong>с</strong><strong>к</strong>ий яйце<strong>к</strong>л<strong>а</strong>д, <strong>с</strong> тремя<br />

п<strong>а</strong>льце<strong>в</strong>идными <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми н<strong>а</strong> <strong>в</strong>ершине.<br />

Дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> тр<strong>а</strong>хеями<br />

и <strong>с</strong>тигм<strong>а</strong>ми, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

н<strong>о</strong>г. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, у мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong> пл<strong>о</strong>тн<strong>о</strong>м<br />

п<strong>а</strong>нцире имеют<strong>с</strong>я п<strong>о</strong>ри<strong>с</strong>тые уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и — т<strong>а</strong><strong>к</strong> н<strong>а</strong>-<br />

Hypochthoniidae, Camisiidae и Neoliodidae<br />

452<br />

http://jurassic.ru/<br />

X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1896). 1295. Neollodes theleproctus<br />

(Hermann); <strong>о</strong>бщий <strong>в</strong>ид, х 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1887). 1296.<br />

Embolacarus pergratus Sellnick; <strong>о</strong>бщий <strong>в</strong>ид: A — <strong>с</strong>низу, Б —<br />

<strong>с</strong><strong>в</strong>ерху; х 100,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий HHTapb)(Sellnick,<br />

1918). 1297. Platyliodes doderleinii (Berlese); <strong>о</strong>бщий <strong>в</strong>ид, х 100.0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1896)

453<br />

http://jurassic.ru/

зы<strong>в</strong><strong>а</strong>емые п<strong>о</strong>р<strong>о</strong><strong>в</strong>ые п<strong>о</strong>ля, <strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>ры'м п<strong>о</strong>дх<strong>о</strong>дят<br />

тр<strong>а</strong>хеи. Тр<strong>а</strong>хейн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> Oribatei р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>в</strong>т<strong>о</strong>ричн<strong>о</strong>; у <strong>в</strong><strong>с</strong>ех неп<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>зрелых ф<strong>а</strong>з, <strong>а</strong> т<strong>а</strong><strong>к</strong>же<br />

у примити<strong>в</strong>ных п<strong>а</strong>нцирных <strong>к</strong>лещей, не имеющих<br />

пл<strong>о</strong>тн<strong>о</strong>г<strong>о</strong> хитин<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> п<strong>а</strong>нциря, тр<strong>а</strong>хеи и<br />

<strong>с</strong>тигмы <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>те<strong>к</strong><strong>а</strong>ет п<strong>о</strong><br />

типу тринимф<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>. Личин<strong>к</strong>и<br />

и <strong>в</strong><strong>с</strong>е нимфы 1<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я <strong>о</strong>т <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

<strong>к</strong>лещей.<br />

П<strong>а</strong>нцирные <strong>к</strong>лещи <strong>в</strong> <strong>к</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>с</strong><strong>а</strong>льных <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>а</strong>х<br />

(д<strong>о</strong> 10—20 ты<strong>с</strong>. н<strong>а</strong> 1 <strong>к</strong><strong>в</strong>. м) н<strong>а</strong><strong>с</strong>еляют<br />

<strong>в</strong>ерхние г<strong>о</strong>риз<strong>о</strong>нты п<strong>о</strong>ч<strong>в</strong>ы, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong>й р<strong>а</strong><strong>с</strong>тительн<strong>о</strong><strong>с</strong>тью;<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>о</strong>ни <strong>в</strong> ле<strong>с</strong><strong>а</strong>х<br />

и н<strong>а</strong> луг<strong>а</strong>х, где н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти п<strong>о</strong>ч<strong>в</strong>ы 1<br />

имеет<strong>с</strong>я<br />

<strong>о</strong>бильн<strong>а</strong>я <strong>в</strong>л<strong>а</strong>жн<strong>а</strong>я п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong><strong>а</strong>. Клещи з<strong>а</strong>п<strong>о</strong>лз<strong>а</strong>ют<br />

н<strong>а</strong> дере<strong>в</strong>ья н<strong>а</strong> <strong>в</strong>ы<strong>с</strong><strong>о</strong>ту д<strong>о</strong> 20 м, з<strong>а</strong><strong>с</strong>еляют<br />

дупл<strong>а</strong> дере<strong>в</strong>ье<strong>в</strong>, н<strong>о</strong>ры грызун<strong>о</strong><strong>в</strong> и другие убежищ<strong>а</strong>.<br />

Из<strong>о</strong>билием <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong> ф<strong>а</strong>уны п<strong>а</strong>нцирных<br />

<strong>к</strong>лещей ле<strong>с</strong><strong>а</strong> <strong>с</strong>ледует <strong>о</strong>бъя<strong>с</strong>нить их <strong>в</strong><strong>с</strong>треч<strong>а</strong>ем<strong>о</strong><strong>с</strong>ть<br />

преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong> янт<strong>а</strong>ре. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Д<strong>в</strong><strong>а</strong> инфр<strong>а</strong><strong>о</strong>тряд<strong>а</strong>: Aptyctima и<br />

Ptyctima.<br />

ИНФРАОТРЯД APTYCTIMA<br />

Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong><strong>о</strong>единен<strong>а</strong> <strong>с</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й;<br />

н<strong>а</strong> ме<strong>с</strong>те их <strong>с</strong>лияния у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong><br />

(ри<strong>с</strong>. 1291—1297); т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у <strong>о</strong>тдельны'х <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

(р<strong>о</strong>д Amerus Berlese) п<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> м<strong>о</strong>жет<br />

и<strong>с</strong>чез<strong>а</strong>ть <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>с</strong>лияния п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>с</strong> н<strong>о</strong>т<strong>о</strong>г<strong>а</strong><strong>с</strong>тр<strong>а</strong>льным щит<strong>о</strong>м. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Де<strong>с</strong>ять н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Hypochthoniidea,<br />

Camisiidea, Hermanniidea, Damaeidea,<br />

Carabodidea, Hermanniellidea, Notaspididea;<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льные три н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> — Epilohmanniidea,<br />

Eulohmanniidea, Nanhermanniidea — из<strong>в</strong>е<strong>с</strong>тны<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

НАДСЕМЕЙСТВО HYPOCHTHON1IDEA<br />

(Circummarginatae)<br />

Спинн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong>л<strong>а</strong>б<strong>о</strong>­<br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я или упл<strong>о</strong>щенн<strong>а</strong>я. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы 1<br />

тел<strong>а</strong> <strong>о</strong>т­<br />

н<strong>о</strong><strong>с</strong>ительн<strong>о</strong> нежные, т<strong>о</strong>н<strong>к</strong>ие и мяг<strong>к</strong>ие; п<strong>а</strong>нлгирь,<br />

<strong>с</strong>т<strong>о</strong>ль х<strong>а</strong>р<strong>а</strong><strong>к</strong>терный для <strong>в</strong><strong>с</strong>ех <strong>о</strong>риб<strong>а</strong>тид,<br />

зде<strong>с</strong>ь <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; н<strong>а</strong> теле м<strong>о</strong>гут быть лишь<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные мел<strong>к</strong>ие щит<strong>к</strong>и (ри<strong>с</strong>. 1292).<br />

Кл<strong>а</strong>п<strong>а</strong>ны п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий<br />

<strong>о</strong>чень <strong>к</strong>рупные, <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м,<br />

з<strong>а</strong>ним<strong>а</strong>ют <strong>в</strong><strong>с</strong>ю длину брюшн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы и <strong>о</strong><strong>к</strong>ружены <strong>о</strong>бщей уз<strong>к</strong><strong>о</strong>й хитин<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

р<strong>а</strong>м<strong>к</strong><strong>о</strong>й. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Malaconothridae, Lohmanniidae, Trhypochthoniidae,<br />

Hypochthoniidae; п<strong>о</strong><strong>с</strong>леднее<br />

из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО HYPOCHTHONIIDAE BERLESE, 1910<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы б<strong>о</strong>лее<br />

упл<strong>о</strong>тненные уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я пятн<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1292). Спинн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

р<strong>а</strong>зделен<strong>а</strong>, <strong>о</strong>дним — четырьмя п<strong>о</strong>перечными<br />

ш<strong>в</strong><strong>а</strong>ми (б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми), <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> зде<strong>с</strong>ь<br />

имеет<strong>с</strong>я д<strong>в</strong><strong>а</strong> — пять <strong>о</strong>тдельных щит<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>.<br />

1291). Генит<strong>а</strong>льные щит<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong> <strong>к</strong>рупнее<br />

<strong>а</strong>н<strong>а</strong>льных. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 18 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Trhypochthonius<br />

Berlese, 1904 (ри<strong>с</strong>. 1291) (д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>) и Вг<strong>а</strong>chychthonius<br />

Berlese, 1910 (ри<strong>с</strong>. 1292) (<strong>о</strong>дин<br />

<strong>в</strong>ид).<br />

НАДСЕМЕЙСТВО CAMISIIDEA<br />

Спинн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы пл<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>я,<br />

<strong>с</strong> <strong>в</strong><strong>о</strong>гнут<strong>о</strong>й <strong>с</strong>рединн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью и прип<strong>о</strong>днятыми<br />

<strong>к</strong>р<strong>а</strong>ями. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы мяг<strong>к</strong>ие, н<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> хитиниз<strong>о</strong><strong>в</strong><strong>а</strong>нны'е.<br />

З<strong>а</strong>дний <strong>к</strong><strong>о</strong>нец тел<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен,<br />

у<strong>с</strong>ечен или <strong>с</strong> <strong>в</strong>ыем<strong>к</strong><strong>о</strong>й и <strong>с</strong> <strong>к</strong>рупными щетин<strong>к</strong><strong>а</strong>ми<br />

р<strong>а</strong>зличных ф<strong>о</strong>рм и <strong>в</strong>еличины, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

н<strong>а</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>х или <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>х щит<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Camisiidae.<br />

СЕМЕЙСТВО CAMISIIDAE SELLNICK, 1928<br />

(Oribatidae Karsch, 1884)<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е-и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я<br />

друг <strong>с</strong> друг<strong>о</strong>м и ряд<strong>о</strong>м <strong>с</strong> ними р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

неб<strong>о</strong>льшие щит<strong>к</strong>и (ри<strong>с</strong>. 1293, 1294).<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. Ше<strong>с</strong>ть <strong>с</strong><strong>о</strong><strong>в</strong>ременных р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

причем д<strong>в</strong><strong>а</strong> из них из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь): Catnisia Heyden, 1826 (ри<strong>с</strong>.<br />

1293) и Nothrus Koch, 1836 (ри<strong>с</strong>. 1294).<br />

НАДСЕМЕЙСТВО HERMANNIIDEA<br />

(Immarginatae)<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> хитиниз<strong>о</strong><strong>в</strong><strong>а</strong>ны; н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы 1<br />

пл<strong>о</strong>тный н<strong>о</strong>т<strong>о</strong>г<strong>а</strong><strong>с</strong>тр<strong>а</strong>льный<br />

щит (п<strong>а</strong>нцирь); з<strong>а</strong>дний и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

Ри<strong>с</strong>. 1298—1310. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Cymbaeremaeidae, Damaeidae, Oribatidae<br />

1298. Cymbaeremaeus (Scapheremaeus) marginalis (Banks); <strong>о</strong>бщий<br />

<strong>в</strong>ид, хбО.О, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Baker and Wharton, 1952). 1299. Micreremus<br />

brevipes (Michael); <strong>о</strong>бщий <strong>в</strong>ид, X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese,<br />

1910). 1300. Tectocymba rara Sellnick; <strong>о</strong>бщий <strong>в</strong>ид, X 100,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Sellnick, 1918). 1301. Damaeus<br />

auritus (Koch); <strong>о</strong>бщий <strong>в</strong>ид, X35.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1887).<br />

1302 Gymnodamaeus retusus (Berlese); <strong>о</strong>бщий <strong>в</strong>ид, x40,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Berlese, 1883). 1303. Eremaeus oblongus (Koch); <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1910). 1304. Caleremaeus moniltpes<br />

(Michael)'; <strong>о</strong>бщий <strong>в</strong>ид, X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1887). 1305.<br />

454<br />

http://jurassic.ru/<br />

Gradidorsum asper Sellnick; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

Б — <strong>с</strong>т<strong>в</strong><strong>о</strong>р<strong>к</strong>и генит<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий; х 110,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Sellnick, 1918). 1306. Licneremaeus<br />

caesareus Berlese; <strong>о</strong>бщий <strong>в</strong>ид, x 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese,<br />

1910). 1307. Lucoppia lucorum (Koch); <strong>о</strong>бщий <strong>в</strong>ид, х 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Berlese, 1910). 1308. Oppia longilamellata (Michael); <strong>о</strong>бщий <strong>в</strong>ид.<br />

X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1887). 1309. Ceratoppla bipllis (Hermann);<br />

<strong>о</strong>бщий <strong>в</strong>ид, x 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1887). 1310. Strieremaeus<br />

illlbatus Sellnick; <strong>о</strong>бщий <strong>в</strong>ид, x85,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Sellnick, 1918)

1305Б<br />

http://jurassic.ru/

<strong>к</strong>р<strong>а</strong>я щит<strong>а</strong> з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я<br />

друг <strong>с</strong> друг<strong>о</strong>м, н<strong>о</strong> не <strong>о</strong><strong>к</strong>ружены <strong>о</strong>бщей хитин<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

р<strong>а</strong>м<strong>к</strong><strong>о</strong>й. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Neoliodidae и Hermanniidae, из них <strong>в</strong>т<strong>о</strong>р<strong>о</strong>е из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

СЕМЕЙСТВО NEOLIODIDAE WILLMANN, 1913<br />

Кр<strong>а</strong>й <strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>, з<strong>а</strong>гиб<strong>а</strong>я<strong>с</strong>ь н<strong>а</strong> брюшную<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну, <strong>о</strong>бр<strong>а</strong>зует н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong><br />

уз<strong>к</strong>ий <strong>в</strong>ы<strong>с</strong>туп (х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong>). К<strong>а</strong>ждый <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong>зделен п<strong>о</strong>перечн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й<br />

н<strong>а</strong> д<strong>в</strong>е ч<strong>а</strong><strong>с</strong>ти. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я личин<strong>о</strong>чные ш<strong>к</strong>ур<strong>к</strong>и<br />

<strong>в</strong><strong>с</strong>ех предыдущих ф<strong>а</strong>з р<strong>а</strong>з<strong>в</strong>ития (ри<strong>с</strong>. 1295,<br />

1297). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

четыре р<strong>о</strong>д<strong>а</strong> и <strong>о</strong>дин р<strong>о</strong>д <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь): Embolacarus Sellnick,<br />

1919 (ри<strong>с</strong>. 1296). Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре<br />

<strong>о</strong>бн<strong>а</strong>ружены <strong>с</strong><strong>о</strong><strong>в</strong>ременные р<strong>о</strong>ды Neoliod.es<br />

Berlese, 1888 (ри<strong>с</strong>. 1295) и Platyliodes Berlese,<br />

1916 (ри<strong>с</strong>. 1297).<br />

НАДСЕМЕЙСТВО DAMAEIDEA<br />

Кр<strong>а</strong>я <strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

<strong>к</strong>р<strong>а</strong>ях тел<strong>а</strong> или з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну<br />

тел<strong>а</strong>. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть н<strong>о</strong>т<strong>о</strong>г<strong>а</strong><strong>с</strong>тр<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong><br />

гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или <strong>с</strong> мел<strong>к</strong><strong>о</strong>т<strong>о</strong>чечн<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й.<br />

Спинн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я,<br />

или н<strong>а</strong> ней имеют<strong>с</strong>я темн<strong>о</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>шенные уч<strong>а</strong><strong>с</strong>т<strong>к</strong>и<br />

(ри<strong>с</strong>. 1298—1302). Н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы 1<br />

,<br />

н<strong>а</strong>д <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями н<strong>о</strong>г I и II, имеют<strong>с</strong>я шир<strong>о</strong><strong>к</strong>ие<br />

треуг<strong>о</strong>льные или р<strong>о</strong>г<strong>о</strong><strong>в</strong>ые <strong>в</strong>ы<strong>с</strong>тупы. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Cymbaeremaeidae,<br />

Damaeidae, Oribatidae.<br />

СЕМЕЙСТВО CYMBAEREMAEIDAE WILLMANN, 1931<br />

Кр<strong>а</strong>я <strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

тел<strong>а</strong>, з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

<strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> и пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мэтиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

щит<strong>о</strong><strong>в</strong> <strong>с</strong> гу<strong>с</strong>тым <strong>с</strong>ете<strong>в</strong>идны'м ри<strong>с</strong>ун<strong>к</strong><strong>о</strong>м (ри<strong>с</strong>.<br />

1298—1300). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один <strong>в</strong>ымерший<br />

р<strong>о</strong>д Tectocymba Sellnick. 1919 из<strong>в</strong>е<strong>с</strong>тен из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь); три<br />

р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из них Cymbaeremaeus<br />

Berlese, 1896 и Micreremus Berlese,<br />

1908, т<strong>а</strong><strong>к</strong>же <strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре.<br />

СЕМЕЙСТВО DAMAEIDAE BERLESE, 1896<br />

(Belbidae Willmann, 1931)<br />

Спинн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы 1<br />

гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я,<br />

без <strong>к</strong><strong>а</strong><strong>к</strong>их-либ<strong>о</strong> ут<strong>о</strong>лщений <strong>в</strong> передней ее<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине. Кр<strong>а</strong>я <strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> не з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну тел<strong>а</strong>. П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть <strong>с</strong>пинн<strong>о</strong>г<strong>о</strong><br />

щит<strong>а</strong> гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я или <strong>с</strong> мел<strong>к</strong><strong>о</strong>т<strong>о</strong>чечн<strong>о</strong>й<br />

456<br />

http://jurassic.ru/<br />

пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й. Н<strong>о</strong>ги <strong>о</strong>бычн<strong>о</strong> длинные, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

длиннее тел<strong>а</strong>; члени<strong>к</strong>и их длинн<strong>о</strong><strong>с</strong>тебельч<strong>а</strong>тые,<br />

чет<strong>к</strong><strong>о</strong><strong>в</strong>идные или бутыл<strong>к</strong><strong>о</strong><strong>в</strong>идные. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>в</strong><strong>о</strong><strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>; из<br />

них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пь( (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

р<strong>о</strong>ды Damaeus С. L. Koch, 1836 и Gymnodamaeus<br />

Kulczynski, 1902 (ри<strong>с</strong>. 1301, 1302). Кр<strong>о</strong>ме<br />

т<strong>о</strong>г<strong>о</strong>, из ми<strong>о</strong>цен<strong>а</strong> 'Ю. Е<strong>в</strong>р<strong>о</strong>пы <strong>о</strong>пи<strong>с</strong><strong>а</strong>н р<strong>о</strong>д Belbites<br />

Pampaloni, 1902.<br />

СЕМЕЙСТВО ORIBATIDAE KRAMER, 1877<br />

(Eremaeidae Willmann, 1931)<br />

В передней ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>пинн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м и <strong>в</strong>переди б<strong>о</strong>тридий (трих<strong>о</strong>б<strong>о</strong>трии),<br />

имеют<strong>с</strong>я темн<strong>о</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>шенные <strong>в</strong><strong>а</strong>ли<strong>к</strong>и.<br />

Н<strong>о</strong>ги <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, чет<strong>к</strong><strong>о</strong><strong>в</strong>идные, <strong>с</strong><br />

ут<strong>о</strong>лщенными члени<strong>к</strong><strong>а</strong>ми, н<strong>о</strong> бедр<strong>а</strong> н<strong>о</strong>г I и II<br />

не<strong>с</strong>тебельч<strong>а</strong>тые (ри<strong>с</strong>. 1303—1310). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong>;<br />

<strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, три р<strong>о</strong>д<strong>а</strong> <strong>в</strong>ымершие: Gradidorsum<br />

Sellnick, 1919 (ри<strong>с</strong>. 1305); Strieremaeus Sellnick,<br />

1919 (ри<strong>с</strong>. 1310); Oppites Pampaloni, 1902,<br />

причем пер<strong>в</strong>ые д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь), п<strong>о</strong><strong>с</strong>ледний — из ми<strong>о</strong>цен<strong>а</strong><br />

Сицилии. П<strong>о</strong>мим<strong>о</strong> т<strong>о</strong>г<strong>о</strong>, из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий ян<strong>а</strong>т<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тны <strong>с</strong><strong>о</strong><strong>в</strong>ременные<br />

р<strong>о</strong>ды Caleremaeus Berlese, 1910 (ри<strong>с</strong>. 1304);<br />

Licneremaeus Paoli, 1908 (ри<strong>с</strong>. 1306); Lucoppia<br />

Berlese, 1908 (ри<strong>с</strong>. 1307); Oppia С. L. Koch,<br />

1836 (ри<strong>с</strong>. 1308); Ceratoppia Berlese, 1908<br />

(ри<strong>с</strong>. 1309); Suctobelba Paoli, 1908; Eremaeus<br />

C. L. Koch, 1836.<br />

НАДСЕМЕЙСТВО<br />

CARABODIDEA W. DUBININ, 1954<br />

Тел<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы, умеренн<strong>о</strong><br />

<strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

имеют<strong>с</strong>я пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тые р<strong>а</strong>зр<strong>а</strong>щения<br />

<strong>в</strong> <strong>в</strong>иде <strong>к</strong>илей, ребер и гребней (ри<strong>с</strong>. 1313);<br />

у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> (у <strong>с</strong><strong>о</strong><strong>в</strong>ременных Ameronothridae)<br />

<strong>о</strong>ни <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жены или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют,<br />

н<strong>о</strong> т<strong>о</strong>гд<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> между пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й бы<strong>в</strong><strong>а</strong>ет п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>а</strong>ми<br />

<strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> и г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> щит<strong>о</strong><strong>в</strong>. Кр<strong>а</strong>я пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> з<strong>а</strong><strong>к</strong>руглены. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong> <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ньг,<br />

<strong>с</strong> груб<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й, <strong>о</strong>бычн<strong>о</strong> темн<strong>о</strong>г<strong>о</strong><br />

ц<strong>в</strong>ет<strong>а</strong>. Н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Пять<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Plateremaeidae, Zetorchestidae, Ameronothridae,<br />

Carabodidae, Liacaridae, н<strong>о</strong> лишь<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО CARABODIDAE WILLMANN, 1931<br />

Н<strong>о</strong>ги III и IV причленяют<strong>с</strong>я н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х тел<strong>а</strong>.<br />

Бедр<strong>а</strong> н<strong>о</strong>г I и II <strong>с</strong>тебельч<strong>а</strong>тые, бутыл<strong>к</strong><strong>о</strong><strong>в</strong>идные.<br />

Н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы грубый <strong>с</strong><strong>к</strong>уль-

Ри<strong>с</strong>. 1311—1320. П<strong>а</strong>нцирные <strong>к</strong>лещи. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Carabodidae, Liacaridae и Hermanniellidae<br />

1311. Carabodes femoratus Berlese; <strong>о</strong>бщий <strong>в</strong>ид, x 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Berlese, 1913). 1312. Otocepheus damaeoides Berlese; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1913). 1313. Cepheus latus (Koch);<br />

<strong>о</strong>бщий <strong>в</strong>ид, x90,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1887). 1314. Plategeocranus<br />

suicatus (Karsch); <strong>о</strong>бщий <strong>в</strong>ид, x85,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Sellnick, 1918). 1315. Scutoribates perornatus Sel-<br />

Inlck; <strong>о</strong>бщий <strong>в</strong>ид, x90,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

птурный ри<strong>с</strong>ун<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 1311—1318). Очень <strong>с</strong>ух<strong>о</strong>люби<strong>в</strong>ые<br />

<strong>в</strong>иды. П<strong>а</strong>ле<strong>о</strong>ген — ныне. 21 р<strong>о</strong>д. Из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из-<br />

457<br />

http://jurassic.ru/<br />

1312<br />

1320<br />

1315<br />

(Sellnick, 1918). 1316. Xenlllus tegeocranus (Hermann); <strong>о</strong>бщий <strong>в</strong>ид,<br />

Х 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1882). 1317. Tectocepheus velatus (Michael);<br />

<strong>о</strong>бщий <strong>в</strong>ид, х80,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1887). 1318. Ommatocepheus<br />

elongatus (Michael); <strong>о</strong>бщий <strong>в</strong>ид, Х80.0. <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael,<br />

1879). 1319. Cultroribula juncta (Michael); <strong>о</strong>бщий <strong>в</strong>ид, X 100,0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1882). 1320. Hermanniella granulata (Nicolet);<br />

<strong>о</strong>бщий <strong>в</strong>ид, х 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1882)<br />

<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>: <strong>в</strong>ымершие Plategeocranus<br />

Sellnick, 1918 (ри<strong>с</strong>. 1314) и Scutoribates Sellnick,<br />

1919 (ри<strong>с</strong>. 1315) и жи<strong>в</strong>ущие ныне С<strong>а</strong>г<strong>а</strong>-

Ри<strong>с</strong>. 1321—1330. П<strong>а</strong>нцирные <strong>к</strong>лещи. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Oripodidae, Oribatulidae, Ceratozetidae и Oribatellidae<br />

1321. Oripoda elongata Banks; <strong>о</strong>бщий <strong>в</strong>ид, XllO.O, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Baker<br />

and Wharton, 1952). 1322. Eporibatula rauschenensis (Sellnick);<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Sellnick, 1908). 1323. Liebstadia<br />

slmilis (Michael); <strong>о</strong>бщий <strong>в</strong>ид, X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1895).<br />

1324. Scheloribates latipes (Koch); <strong>о</strong>бщий <strong>в</strong>ид, хЮ0,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Michael, 1888). 1325. Chamobates pusillus (Berlese); <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1895). 1326. Melanozetes mollicomus<br />

bodes Koch, 1836 (ри<strong>с</strong>. 1311); Otocepheus Berlese,<br />

1910 (ри<strong>с</strong>. 1312); Cepheus Koch, 1836<br />

(ри<strong>с</strong>. 1313); Xenillus Robineau-Desvoidy, 1839<br />

(ри<strong>с</strong>. 1316); Tectocepheus Berlese, 1913 (ри<strong>с</strong>.<br />

1317); Ommatocepheus Berlese,1913 (ри<strong>с</strong>. 1318).<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из ми<strong>о</strong>цен<strong>а</strong> Сицилии <strong>о</strong>пи<strong>с</strong><strong>а</strong>н р<strong>о</strong>д<br />

Carabodites Pampaloni, 1902.<br />

458<br />

http://jurassic.ru/<br />

(K.och); <strong>о</strong>бщий <strong>в</strong>ид, х60,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Michael, 1888). 1327. Sphaerozetes<br />

prudens Berlese; <strong>о</strong>бщий <strong>в</strong>ид, 80,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1895).<br />

1328. Oribatella calcarata (Koch); <strong>о</strong>бщий <strong>в</strong>ид, x 100,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем.<br />

(Berlese, 1895). 1329. Tectoribates piriformis (Nicolet); <strong>о</strong>бщий <strong>в</strong>ид,<br />

X60,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese, 1895). 1330. Tectoribates parvus Sellnick;<br />

<strong>о</strong>бщий <strong>в</strong>ид, x90,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь,<br />

Sellnick, 1918)<br />

СЕМЕЙСТВО LIACARIDAE WILLMANN, 1931<br />

Н<strong>о</strong>ги III и IV причленяют<strong>с</strong>я н<strong>а</strong> брюшн<strong>о</strong>й<br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии <strong>о</strong>т<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> тел<strong>а</strong>. Спинн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я,<br />

<strong>с</strong> т<strong>о</strong>чечн<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й. В<strong>с</strong>е члени<strong>к</strong>и<br />

н<strong>о</strong>г б<strong>о</strong>лее или менее цилиндриче<strong>с</strong><strong>к</strong>ие. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й

Ри<strong>с</strong>. 1331—1338. П<strong>а</strong>нцирные <strong>к</strong>лещи. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Notaspididae, Haplozetidae, Phenopelopidae, Galumnidae<br />

и Phthiracaridae<br />

1337А<br />

1331. Notaspls nicoletli (Berlese); <strong>о</strong>бщий <strong>в</strong>ид, X100.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем, ronata (Canestrini Q. et R.); <strong>о</strong>бщий <strong>в</strong>ид. ХбО.О, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Berlese,<br />

ширн<strong>а</strong>я групп<strong>а</strong> п<strong>а</strong>нцирных <strong>к</strong>лещей, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленн<strong>а</strong>я<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 11 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>ми:<br />

Microzetidae, Epactozetidae, Tenuialidae, Oripodidae,<br />

Oribatulidae, Ceratozetidae, OribatelHdae,<br />

Notaspididae, Haplozetidae, Pelopsidae,<br />

Galumnidae; пер<strong>в</strong>ые три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тны.<br />

СЕМЕЙСТВО ORIPODIDAE JACOT, 1925<br />

Вд<strong>о</strong>ль б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ют<strong>с</strong>я<br />

лент<strong>о</strong><strong>в</strong>идные р<strong>а</strong>зр<strong>а</strong>щения хитин<strong>а</strong> (неп<strong>о</strong>д<strong>в</strong>ижные<br />

птер<strong>о</strong>м<strong>о</strong>рфы); <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>ни не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>ширены, пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ют<strong>с</strong>я <strong>в</strong>перед д<strong>о</strong><br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>и<strong>к</strong>ь! длины пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы, где <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я,<br />

<strong>о</strong>бр<strong>а</strong>зуя шир<strong>о</strong><strong>к</strong>ую, т<strong>о</strong>н<strong>к</strong>ую п<strong>о</strong>перечную пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у,<br />

н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющую н<strong>а</strong>д пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющей<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния трих<strong>о</strong>б<strong>о</strong>трий и интерл<strong>а</strong>меллярных<br />

щетин<strong>о</strong><strong>к</strong>, <strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> и <strong>с</strong><strong>а</strong>ми щетин<strong>к</strong>и.<br />

У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых ф<strong>о</strong>рм эт<strong>о</strong>т <strong>к</strong><strong>о</strong>зыре<strong>к</strong> <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы. Генит<strong>а</strong>льных щетин<strong>о</strong><strong>к</strong><br />

д<strong>в</strong>е — три п<strong>а</strong>ры (ри<strong>с</strong>. 1321). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре р<strong>о</strong>д<strong>а</strong>; из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тен<br />

р<strong>о</strong>д Oripoda Banks, 1904.<br />

СЕМЕЙСТВО ORIBATULIDAE JACOT, 1929<br />

(Scheloribatidae Grandjean, 1933)<br />

В передней ч<strong>а</strong><strong>с</strong>ти ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы имеют<strong>с</strong>я<br />

<strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong>ие плече<strong>в</strong>ые хитин<strong>о</strong><strong>в</strong>ые <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты,<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнутые н<strong>а</strong>з<strong>а</strong>д, х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метные<br />

при р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>нии <strong>к</strong>лещей <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны. Н<strong>а</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>ме <strong>с</strong>л<strong>а</strong>б<strong>о</strong>р<strong>а</strong>з<strong>в</strong>итые <strong>к</strong>или<br />

или ребр<strong>а</strong>. Щетин<strong>к</strong>и н<strong>а</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>ме р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> четыре пр<strong>о</strong>д<strong>о</strong>льных ряд<strong>а</strong>. Н<strong>а</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м<br />

<strong>к</strong>л<strong>а</strong>п<strong>а</strong>не п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong> четыре щетин<strong>к</strong>и<br />

(ри<strong>с</strong>. 1322—1324). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 13 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) р<strong>о</strong>ды 1<br />

Eporibatula<br />

Sellnick, 1928 (ри<strong>с</strong>. 1322); Liebstadia Oudemans,<br />

1906 (ри<strong>с</strong>. 1323); Scheloribates Berlese, 1908<br />

(ри<strong>с</strong>. 1324).<br />

СЕМЕЙСТВО CERATOZETIDAE JACOT, 1925<br />

Н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы имеют<strong>с</strong>я я<strong>в</strong><strong>с</strong>т<strong>в</strong>енные<br />

шир<strong>о</strong><strong>к</strong>ие неп<strong>о</strong>д<strong>в</strong>ижные пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тые <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты<br />

(птер<strong>о</strong>м<strong>о</strong>рфы). Вд<strong>о</strong>ль переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

птер<strong>о</strong>м<strong>о</strong>рфы <strong>с</strong><strong>о</strong>единены друг <strong>с</strong> друг<strong>о</strong>м<br />

уз<strong>к</strong>им хитин<strong>о</strong><strong>в</strong>ым м<strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>о</strong>м (ри<strong>с</strong>. 1327), <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

не при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет <strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong>-нибудь зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>с</strong>пинную п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы; передний<br />

<strong>к</strong>р<strong>а</strong>й м<strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong> д<strong>а</strong>ле<strong>к</strong><strong>о</strong> не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет ур<strong>о</strong><strong>в</strong>ня<br />

б<strong>о</strong>тридий (трих<strong>о</strong>б<strong>о</strong>трий). Н<strong>а</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м <strong>к</strong>л<strong>а</strong>п<strong>а</strong>не<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong> ше<strong>с</strong>ти щетин<strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 1325—1327). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 29 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) р<strong>о</strong>ды Chamobates<br />

460<br />

http://jurassic.ru/<br />

Hull, 1916 (ри<strong>с</strong>. 1325); Melanozetes Hull, 1916<br />

(ри<strong>с</strong>. 1326); Sphaerozetes Berlese, 1885 (= Euzetes<br />

Berlese, 1908) (ри<strong>с</strong>. 1327).<br />

СЕМЕЙСТВО ORIBATELLIDAE JACOT, 1925<br />

Неп<strong>о</strong>д<strong>в</strong>ижные птер<strong>о</strong>м<strong>о</strong>рфы <strong>к</strong>рупные, <strong>с</strong> <strong>о</strong><strong>с</strong>триями<br />

н<strong>а</strong> переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых угл<strong>а</strong>х. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>чень <strong>к</strong>рупные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и,<br />

при<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющие п<strong>о</strong>чти <strong>в</strong><strong>с</strong>ю <strong>с</strong>пинную п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы; передние <strong>к</strong>р<strong>а</strong>я их <strong>с</strong> т<strong>о</strong>рч<strong>а</strong>щими<br />

<strong>о</strong><strong>с</strong>триями (ри<strong>с</strong>. 1328). Трих<strong>о</strong>б<strong>о</strong>трий<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ют з<strong>а</strong> <strong>к</strong>р<strong>а</strong>й уз<strong>к</strong><strong>о</strong>г<strong>о</strong> хитин<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> м<strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong><br />

между птер<strong>о</strong>м<strong>о</strong>рф<strong>а</strong>ми (ри<strong>с</strong>. 1328—1330). П<strong>а</strong>ле<strong>о</strong>ген—-ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре'<br />

р<strong>о</strong>д<strong>а</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) р<strong>о</strong>ды: Oribatella Banks, 1885 (ри<strong>с</strong>.<br />

1328) и Tectoribates Berlese, 1910 (ри<strong>с</strong>. 1329—<br />

1330).<br />

СЕМЕЙСТВО NOTASPIDIDAE OUDEMANS, 1900<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и<br />

<strong>о</strong>чень <strong>к</strong>рупные. Т<strong>о</strong>н<strong>к</strong>ие, з<strong>а</strong><strong>о</strong><strong>с</strong>тренные передние<br />

углы птер<strong>о</strong>м<strong>о</strong>рф <strong>о</strong><strong>с</strong>триями п<strong>о</strong>чти д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют<br />

ур<strong>о</strong><strong>в</strong>ня р<strong>о</strong><strong>с</strong>трум<strong>а</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Передние<br />

<strong>к</strong>р<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>о</strong><strong>к</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы у<strong>с</strong>ечены, не <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>в</strong>ытянутых пл<strong>а</strong><strong>с</strong>тинч<strong>а</strong>тых з<strong>а</strong><strong>о</strong><strong>с</strong>трений.<br />

Пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я между<br />

<strong>с</strong><strong>о</strong>б<strong>о</strong>й <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти (ри<strong>с</strong>. 1331, 1332). П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре<br />

р<strong>о</strong>д<strong>а</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) р<strong>о</strong>ды: Notaspis Hermann, 1804 (ри<strong>с</strong>.<br />

1331) и Punctoribates Berlese, 1908 (ри<strong>с</strong>.1332).<br />

СЕМЕЙСТВО HAPLOZETIDAE GRANDJEAN, 1936<br />

Птер<strong>о</strong>м<strong>о</strong>рфы м<strong>а</strong>лень<strong>к</strong>ие, треуг<strong>о</strong>льные. Те<strong>к</strong>т<strong>о</strong>педии<br />

н<strong>о</strong>г IV <strong>к</strong>рупные, ли<strong>с</strong>т<strong>о</strong><strong>в</strong>идные. Н<strong>а</strong><br />

<strong>к</strong><strong>а</strong>жд<strong>о</strong>м <strong>к</strong>л<strong>а</strong>п<strong>а</strong>не п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong> пяти<br />

щетин<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 1333). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) р<strong>о</strong>д Рг<strong>о</strong>toribates<br />

Berlese, 1908.<br />

СЕМЕЙСТВО PHENOPELOPIDAE<br />

PETRUNKEVITCH, 1955<br />

(Pelopsidae Ewing, 1917)<br />

Птер<strong>о</strong>м<strong>о</strong>рфы п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong><strong>о</strong>членены <strong>с</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й;<br />

хитин<strong>о</strong><strong>в</strong>ый м<strong>о</strong><strong>с</strong>ти<strong>к</strong> между ними уз<strong>к</strong>ий;<br />

<strong>о</strong>н р<strong>а</strong><strong>с</strong>ширен т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти,<br />

имеющей <strong>в</strong>ид шир<strong>о</strong><strong>к</strong><strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти, н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющей<br />

н<strong>а</strong> з<strong>а</strong>днюю ч<strong>а</strong><strong>с</strong>ть пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы; передний <strong>к</strong>р<strong>а</strong>й<br />

<strong>с</strong>рединн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти д<strong>о</strong>х<strong>о</strong>дит ин<strong>о</strong>гд<strong>а</strong> п<strong>о</strong>чти д<strong>о</strong><br />

<strong>с</strong>ередины пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы (ри<strong>с</strong>. 1334). Хелицеры<br />

<strong>с</strong> шир<strong>о</strong><strong>к</strong>ими <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями и рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженными<br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong>ми; <strong>в</strong>ершинн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть их <strong>о</strong>бычн<strong>о</strong> длинн<strong>а</strong>я,<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я <strong>с</strong> м<strong>а</strong>лень<strong>к</strong><strong>о</strong>й <strong>к</strong>лешней. Н<strong>а</strong> <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н<strong>а</strong>х<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong> ше<strong>с</strong>ти щетин<strong>о</strong><strong>к</strong>.

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне де<strong>с</strong>ять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>; из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

из<strong>в</strong>е<strong>с</strong>тен р<strong>о</strong>д Phaenopelops Petrunkevitch,<br />

1955 ( = Pelops Koch, 1836) (ри<strong>с</strong>. 1334)<br />

СЕМЕЙСТВО GALUMNIDAE GRANDJEAN, 1936<br />

(Parakalummidae Grandjean, 1936; З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин, 1953)<br />

Птер<strong>о</strong>м<strong>о</strong>рфы <strong>к</strong>рыл<strong>о</strong><strong>в</strong>идные, п<strong>о</strong>д<strong>в</strong>ижные; хитин<strong>о</strong><strong>в</strong>ый<br />

м<strong>о</strong><strong>с</strong>ти<strong>к</strong> между ними <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Хелицеры<br />

н<strong>о</strong>рм<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения; передняя ч<strong>а</strong><strong>с</strong>ть<br />

их не <strong>в</strong>ытянут<strong>а</strong>. П<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> между<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й не <strong>в</strong><strong>с</strong>егд<strong>а</strong> я<strong>с</strong>н<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жен<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Н<strong>а</strong> <strong>к</strong><strong>а</strong>жд<strong>о</strong>м<br />

<strong>к</strong>л<strong>а</strong>п<strong>а</strong>не п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong> пяти или ше<strong>с</strong>ти<br />

щетин<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 1335). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Galumna<br />

Heyden, 1826 и Neoribates Berlese, 1914<br />

(ри<strong>с</strong>. 1336).<br />

ИНФРАОТРЯД PTYCTIMA<br />

Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong><strong>о</strong>членен<strong>а</strong> <strong>с</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й,<br />

бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>ря чему <strong>о</strong>н<strong>а</strong> м<strong>о</strong>жет пригиб<strong>а</strong>ть<strong>с</strong>я<br />

<strong>к</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong>, при<strong>к</strong>ры<strong>в</strong><strong>а</strong>я<br />

н<strong>о</strong>ги и гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му (ри<strong>с</strong>. 1337Б). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Одн<strong>о</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phthiracaridea.<br />

НАДСЕМЕЙСТВО PHTHIRACARIDEA<br />

(PTYCTIMA ANDRE, 1949)<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я призн<strong>а</strong><strong>к</strong><strong>а</strong>ми инфр<strong>а</strong><strong>о</strong>тряд<strong>а</strong><br />

(ри<strong>с</strong>. 1337, 1338). П<strong>а</strong>ле<strong>о</strong>ген — ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне: Protoplophoridae,<br />

Mesoplophoridae и Phthiracaridae, из них<br />

п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО PHTHIRACARIDAE PERTY, 1841<br />

Ги<strong>с</strong>тер<strong>о</strong>е<strong>о</strong>м<strong>а</strong> не р<strong>а</strong>зделен<strong>а</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми н<strong>а</strong><br />

«<strong>с</strong>егменты»; <strong>с</strong>пинн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong> ее п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> пл<strong>о</strong>тным,<br />

цельным щит<strong>о</strong>м. Кл<strong>а</strong>п<strong>а</strong>ны п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> и<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий уз<strong>к</strong>ие, у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>в</strong>ытянуты <strong>в</strong> длину, <strong>с</strong>мы<strong>к</strong><strong>а</strong>ют<strong>с</strong>я или д<strong>а</strong>же <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

друг <strong>с</strong> друг<strong>о</strong>м, <strong>о</strong>бр<strong>а</strong>зуя т<strong>о</strong>гд<strong>а</strong> чрез<strong>в</strong>ыч<strong>а</strong>йн<strong>о</strong><br />

длинные <strong>а</strong>н<strong>о</strong>-генит<strong>а</strong>льные пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и.<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong> мяг<strong>к</strong>ие; брюшн<strong>о</strong>й<br />

щит не <strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong><strong>о</strong> <strong>с</strong>пинным щит<strong>о</strong>м<br />

(ри<strong>с</strong>. 1337—1338). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 13 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>: Н<strong>о</strong>ploderma<br />

Michael, 1898 и Oriboritia Jacot, 1924.<br />

ПОДОТРЯД TROMBIDIIFORMES.<br />

КРАСНОТЕЛКОВЫЕ КЛЕЩИ<br />

Р<strong>а</strong><strong>с</strong>членение тел<strong>а</strong> <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>идн<strong>о</strong>г<strong>о</strong> или тритире<strong>о</strong>идн<strong>о</strong>г<strong>о</strong><br />

(ри<strong>с</strong>. 1339) тип<strong>о</strong><strong>в</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы т<strong>о</strong>н<strong>к</strong>ие,<br />

переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>й <strong>к</strong><strong>о</strong>н<strong>с</strong>и<strong>с</strong>тенции; щиты р<strong>а</strong>з<strong>в</strong>иты<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти хитин<strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> имеет<strong>с</strong>я нежн<strong>а</strong>я пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>а</strong> или<br />

т<strong>о</strong>н<strong>к</strong><strong>а</strong>я и<strong>с</strong>черченн<strong>о</strong><strong>с</strong>ть, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я <strong>о</strong>бр<strong>а</strong>зует х<strong>а</strong>р<strong>а</strong><strong>к</strong>терный<br />

<strong>с</strong><strong>к</strong>ульптурный ри<strong>с</strong>ун<strong>о</strong><strong>к</strong>, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий<br />

из <strong>с</strong>и<strong>с</strong>темы п<strong>а</strong>р<strong>а</strong>ллельных линий. Н<strong>а</strong> теле<br />

<strong>о</strong>бычны д<strong>о</strong>п<strong>о</strong>лнительные щетин<strong>к</strong>и (не<strong>о</strong>трихия;<br />

ри<strong>с</strong>. 1357), м<strong>а</strong><strong>с</strong><strong>к</strong>ирующие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение пер<strong>в</strong>ичных<br />

щетин<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 1344); д<strong>о</strong>п<strong>о</strong>лнительные<br />

щетин<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у немн<strong>о</strong>гих групп,<br />

н<strong>а</strong>пример, у Tarsonemidea, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Tydeidea<br />

(ри<strong>с</strong>. 1346—1350), Bdellidae (ри<strong>с</strong>. 1352),<br />

Tetranychidae (ри<strong>с</strong>. 1358) и т. п. Н<strong>а</strong>ряду <strong>с</strong><br />

гл<strong>а</strong>д<strong>к</strong>ими щетин<strong>к</strong><strong>а</strong>ми шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены<br />

р<strong>а</strong>зличн<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>а</strong> <strong>о</strong>пуш<strong>о</strong>нные, пер<strong>о</strong><strong>в</strong>идные или<br />

гребенч<strong>а</strong>тые щетин<strong>к</strong>и (ри<strong>с</strong>. 1340, 1342). Н<strong>а</strong><br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>ме трих<strong>о</strong>б<strong>о</strong>трии имеют<strong>с</strong>я или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>бычн<strong>о</strong> неб<strong>о</strong>льш<strong>а</strong>я и причленяет<strong>с</strong>я<br />

н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>ю тел<strong>а</strong> (Cheyletidae,<br />

ри<strong>с</strong>. 1353; Cunaxidae, Bdellidae, ри<strong>с</strong>.<br />

1352 и др.). Сли<strong>в</strong>шие<strong>с</strong>я <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния хелицер и<br />

гип<strong>о</strong><strong>с</strong>т<strong>о</strong>м <strong>о</strong>бр<strong>а</strong>зуют <strong>к</strong>рупную <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идную<br />

<strong>с</strong><strong>о</strong><strong>с</strong>ущую труб<strong>к</strong>у, п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

б<strong>о</strong>льшие х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные педип<strong>а</strong>льпы,<br />

ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong> <strong>в</strong>иде м<strong>о</strong>щных щипц<strong>о</strong><strong>в</strong> (Cheyletidae и<br />

др.; ри<strong>с</strong>. 1351, 1353, 1356). Хелицеры р<strong>а</strong>з<strong>в</strong>иты<br />

р<strong>а</strong>зличн<strong>о</strong>: <strong>к</strong>лешне<strong>в</strong>идные у <strong>о</strong>дних <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>с</strong>тилет<strong>о</strong><strong>в</strong>идные<br />

у других (Tetranychidea, Myobiidae,<br />

Cheyletidae, Trichadenidea) или т<strong>о</strong>л<strong>с</strong>тые<br />

п<strong>а</strong>льце<strong>в</strong>идные <strong>с</strong> м<strong>а</strong>лень<strong>к</strong>им, <strong>в</strong> <strong>в</strong>иде шип<strong>а</strong> или<br />

<strong>к</strong><strong>о</strong>люч<strong>к</strong>и, п<strong>о</strong>д<strong>в</strong>ижным п<strong>а</strong>льцем (Tydeidea,<br />

ри<strong>с</strong>. 1350; Speleognathidae). Н<strong>о</strong>ги, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>, н<strong>о</strong> у п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

м<strong>о</strong>гут быть при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены для з<strong>а</strong><strong>к</strong>репления<br />

н<strong>а</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>а</strong>х жи<strong>в</strong><strong>о</strong>тных-х<strong>о</strong>зяе<strong>в</strong> (н<strong>о</strong>ги I у<br />

Myobiidae) или быть <strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нными<br />

(Demodicidae, Psorergatidae); у <strong>в</strong><strong>о</strong>дяных<br />

<strong>к</strong>лещей (Hydrachnellae) не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые п<strong>а</strong>ры<br />

н<strong>о</strong>г при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены для пл<strong>а</strong><strong>в</strong><strong>а</strong>ния, члени<strong>к</strong>и<br />

их <strong>с</strong>н<strong>а</strong>бжены длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г<br />

<strong>с</strong>литы <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми тел<strong>а</strong>. В не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых групп<strong>а</strong>х,<br />

<strong>в</strong>едущих п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ий <strong>о</strong>бр<strong>а</strong>з жизни н<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

(Tarsonemidea) или <strong>в</strong> р<strong>а</strong><strong>с</strong>тительных<br />

г<strong>а</strong>лл<strong>а</strong>х (Trichodenidea), н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я реду<strong>к</strong>ция<br />

<strong>о</strong>тдельных н<strong>о</strong>ет. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong>рупными<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и пери<strong>с</strong>тым эмп<strong>о</strong>дием между ними;<br />

ин<strong>о</strong>гд<strong>а</strong> эмп<strong>о</strong>дий и <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и бы<strong>в</strong><strong>а</strong>ют редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны.<br />

Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong>з<strong>в</strong>иты или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Тр<strong>а</strong>хейн<strong>а</strong>я<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong> п<strong>о</strong>чти у <strong>в</strong><strong>с</strong>ех <strong>в</strong>ид<strong>о</strong><strong>в</strong>;<br />

<strong>о</strong>дн<strong>а</strong> п<strong>а</strong>р<strong>а</strong> <strong>с</strong>тигм <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы или <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> нее; <strong>с</strong>тигмы <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>о</strong>жн<strong>о</strong> у<strong>с</strong>тр<strong>о</strong>енными перитрем<strong>а</strong>ми. У<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>-<strong>к</strong>лещей <strong>с</strong>тигмы <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

дим<strong>о</strong>рфизм <strong>в</strong>ыр<strong>а</strong>жен нерез<strong>к</strong><strong>о</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е и<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong>з<strong>о</strong>бщены; <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>о</strong>б<strong>а</strong> <strong>о</strong>ни р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не,<br />

н<strong>о</strong> у Eupodidae Koch з<strong>а</strong>днепр<strong>о</strong>х<strong>о</strong>дн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

<strong>с</strong>мещен<strong>о</strong> н<strong>а</strong> <strong>с</strong>пинную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну, <strong>а</strong> у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

н<strong>а</strong>д <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Cheyletidea и Demodicidea<br />

н<strong>а</strong> <strong>с</strong>пинную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> перех<strong>о</strong>дит <strong>к</strong><strong>о</strong>пу-<br />

461<br />

http://jurassic.ru/

ляти<strong>в</strong>н<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие. Ан<strong>а</strong>льные при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и <strong>в</strong><strong>с</strong>егд<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; 'п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и у <strong>о</strong>тдельных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> бы<strong>в</strong><strong>а</strong>ют р<strong>а</strong>з<strong>в</strong>иты.<br />

П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие пр<strong>о</strong>те<strong>к</strong><strong>а</strong>ет п<strong>о</strong><br />

типу три- или бинимф<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>.<br />

Би<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и Trombidiiformes <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зны.<br />

Н<strong>а</strong>ряду <strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущими <strong>с</strong><strong>а</strong>пр<strong>о</strong>фитными<br />

<strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> примити<strong>в</strong>ными ф<strong>о</strong>рм<strong>а</strong>ми<br />

(Pachygnathidea) <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

хищни<strong>к</strong>и (Cheyletidea, Rhaphignatidea<br />

и др.), п<strong>а</strong>р<strong>а</strong>зиты р<strong>а</strong><strong>с</strong>тений, н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых (Tarsonemidea,<br />

Tetranychidea, Trichadenidea и др.) и<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных (Demodicidae, Pterygosomatidae,<br />

Myobiidae, Trombiculidae и др.). Б<strong>о</strong>льш<strong>а</strong>я<br />

групп<strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емых <strong>в</strong><strong>о</strong>дяных <strong>к</strong>лещей<br />

пр<strong>о</strong><strong>в</strong>из<strong>о</strong>рн<strong>о</strong>й группы Hydrachnellae и н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Halacaridea <strong>о</strong>бит<strong>а</strong>ет <strong>в</strong> пре<strong>с</strong>ных или<br />

м<strong>о</strong>р<strong>с</strong><strong>к</strong>их <strong>в</strong><strong>о</strong>д<strong>а</strong>х.<br />

П<strong>о</strong>д<strong>о</strong>тряд Trombidiiformes р<strong>а</strong><strong>с</strong>п<strong>а</strong>д<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

ряд пр<strong>о</strong><strong>в</strong>из<strong>о</strong>рных группир<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>: Tarsonemini,<br />

Prostigmata и Hydrachnellae (Hydracarina),<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рым ч<strong>а</strong><strong>с</strong>т<strong>о</strong> прид<strong>а</strong>ют зн<strong>а</strong>чение е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енных<br />

<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их групп (Baker and Wharton,<br />

1952). И<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниями ряд<strong>а</strong> <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<br />

Trombidiiformes р<strong>а</strong>зделен н<strong>а</strong> 26 н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>,<br />

<strong>о</strong>бъединяющих 86 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>. Де<strong>в</strong><strong>о</strong>н —<br />

ныне. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны де<strong>в</strong>ять<br />

н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Pachygnathidea, Rhaphignathidea,<br />

Tydeidea, Anystiidea, Bdellidea, Cheyletidea,<br />

Smaridiidea, Trombidiidea и Tetranychidea.<br />

З<strong>а</strong>меч<strong>а</strong>ние. Пять <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>тн<strong>о</strong><strong>с</strong>ящих<strong>с</strong>я<br />

<strong>к</strong> д<strong>в</strong>ум н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>м (Pachygnathidea и Tydeidea),<br />

из<strong>в</strong>е<strong>с</strong>тны из де<strong>в</strong><strong>о</strong>н<strong>а</strong> Ш<strong>о</strong>тл<strong>а</strong>ндии. Эти<br />

ф<strong>о</strong>рмы д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> <strong>в</strong>ремени <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>или<strong>с</strong>ь<br />

<strong>к</strong> <strong>о</strong>дн<strong>о</strong>му <strong>в</strong>иду Protacarus crani Hirst,<br />

1923. Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тели, <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong>, у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ли,<br />

чт<strong>о</strong> п<strong>о</strong>д н<strong>а</strong>з<strong>в</strong><strong>а</strong>нием «Protacarus crani»<br />

<strong>о</strong>бъединен<strong>о</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> из р<strong>а</strong>зличных <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><br />

примити<strong>в</strong>ных Trombidiformes (З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин,<br />

1952, <strong>с</strong>тр. 24).<br />

НАДСЕМЕЙСТВО PACHYGNATHIDEA<br />

(Endeostigmata)<br />

Мел<strong>к</strong>ие <strong>к</strong>лещи, <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, <strong>с</strong> чет<strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>й ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, щетин<strong>к</strong>и н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong>перечными<br />

ряд<strong>а</strong>ми. Н<strong>а</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>ме имеют<strong>с</strong>я<br />

трих<strong>о</strong>б<strong>о</strong>трий. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> <strong>к</strong>рупн<strong>а</strong>я.<br />

Педип<strong>а</strong>льпы пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Хелицеры<br />

четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>рупные, <strong>к</strong>лешне<strong>в</strong>идные.<br />

В <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии их, н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я тр<strong>а</strong>хеи. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong><br />

<strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие, штрих<strong>о</strong><strong>в</strong><strong>а</strong>нные или <strong>с</strong> нежными<br />

буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми; щиты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>о</strong>ги х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>, <strong>с</strong> р<strong>а</strong><strong>с</strong>члененными н<strong>а</strong> basi- и telofemur<br />

бедр<strong>а</strong>ми. Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х имеют<strong>с</strong>я <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и и пери<strong>с</strong>тый<br />

эмп<strong>о</strong>дий. Кл<strong>а</strong>п<strong>а</strong>ны п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых и <strong>а</strong>н<strong>а</strong>льных<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий <strong>с</strong><strong>о</strong> щетин<strong>к</strong><strong>а</strong>ми. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ых при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong><br />

д<strong>в</strong>е или три п<strong>а</strong>ры. Де<strong>в</strong><strong>о</strong>н — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Terpnacaridae, Lordalychidae,<br />

Sphaerolichidae, Pachygnathidae,.<br />

Alicorhagiidae, Nanorchestidae, из них три п<strong>о</strong><strong>с</strong>ледних<br />

из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО NANORCHESTIDAE<br />

GRANDJEAN, 1937<br />

Тел<strong>о</strong> я<strong>в</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> перетяж<strong>к</strong><strong>а</strong>ми<br />

н<strong>а</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му, мет<strong>а</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му и<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му (тритире<strong>о</strong>идный тип р<strong>а</strong><strong>с</strong>членения).<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы мяг<strong>к</strong>ие, <strong>с</strong> т<strong>о</strong>н<strong>к</strong>ими б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми. Н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены 64 х<strong>а</strong>р<strong>а</strong><strong>к</strong>терные<br />

бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>-<strong>в</strong><strong>о</strong>р<strong>о</strong>н<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные щетин<strong>к</strong>и <strong>с</strong> шип<strong>а</strong>ми<br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти (ри<strong>с</strong>. 1340). Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

м<strong>а</strong><strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>а</strong>я. Педип<strong>а</strong>льпы <strong>с</strong> четырьмя — пятью<br />

п<strong>о</strong>д<strong>в</strong>ижными члени<strong>к</strong><strong>а</strong>ми, пр<strong>о</strong><strong>с</strong>тые. Н<strong>о</strong>ги <strong>с</strong>мещены<br />

<strong>в</strong> переднюю п<strong>о</strong>л<strong>о</strong><strong>в</strong>ину тел<strong>а</strong> <strong>о</strong>т меш<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы. Н<strong>о</strong>ги IV <strong>с</strong> прыг<strong>а</strong>тельным<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блением. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong> <strong>о</strong>дним<br />

эмп<strong>о</strong>ди<strong>а</strong>льным <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м. Имеет<strong>с</strong>я три п<strong>а</strong>ры<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong>. Очень мел<strong>к</strong>ие (0,165—<br />

0,350 мм <strong>в</strong> длину) бе<strong>с</strong>ц<strong>в</strong>етные или <strong>к</strong>р<strong>а</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>тые<br />

<strong>к</strong>лещи, <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong><strong>о</strong> мху и <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е. Де<strong>в</strong><strong>о</strong>н<br />

— ныне. Пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых четыре<br />

из<strong>в</strong>е<strong>с</strong>тны т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

Protospeleorchestes W. Dubinin, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. pseudoprotacarus W. Dubinin,<br />

n. n. (Protacarus crani Hirst, 1923; ч<strong>а</strong><strong>с</strong>тью, <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и<br />

№ 6. Не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>к</strong>у<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> <strong>к</strong>лещей<br />

<strong>с</strong><strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зными «щетин<strong>к</strong><strong>а</strong>ми» Hirst, 1923,<br />

<strong>с</strong>тр. 459, ри<strong>с</strong>. 1<strong>в</strong>); де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия (Aberdeenshire,<br />

<strong>к</strong>р<strong>а</strong><strong>с</strong>ный пе<strong>с</strong>ч<strong>а</strong>ни<strong>к</strong>). С<strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

щетин<strong>к</strong>и <strong>с</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й. Щетин<strong>к</strong>и имеют<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идную ф<strong>о</strong>рму, у<strong>с</strong>ечены н<strong>а</strong> <strong>в</strong>ершине, <strong>с</strong><br />

мел<strong>к</strong>ими треуг<strong>о</strong>льными шип<strong>а</strong>ми н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти;<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я б<strong>о</strong>лее или менее п<strong>о</strong>п<strong>а</strong>рн<strong>о</strong>;<br />

н<strong>а</strong>ряду <strong>с</strong> <strong>к</strong>рупными щетин<strong>к</strong><strong>а</strong>ми н<strong>а</strong> теле <strong>к</strong>лещ<strong>а</strong><br />

имели<strong>с</strong>ь <strong>в</strong>д<strong>в</strong><strong>о</strong>е б<strong>о</strong>лее мел<strong>к</strong>ие <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и п<strong>о</strong>д<strong>о</strong>бных<br />

ф<strong>о</strong>рмы и <strong>с</strong>тр<strong>о</strong>ения. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> <strong>к</strong>лещ<strong>а</strong> т<strong>о</strong>н<strong>к</strong>ие,<br />

<strong>с</strong> нежн<strong>о</strong>й п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>й штрих<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й<br />

(ри<strong>с</strong>. 1340). Один <strong>в</strong>ид.<br />

Ри<strong>с</strong>. 1339—1344. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Nanorchestidae, Pachygnathidae, Alicorhagiidae<br />

1339. Speleorchestes poduroides Hirst; <strong>о</strong>бщий <strong>в</strong>ид, X 130,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем<br />

(Hirst, 1917). 1340. Protospeleorchestes pseudoprotacarus W. Dubinin,<br />

gen. nov., sp. nov., щетин<strong>к</strong>и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, х 120,0, де<strong>в</strong><strong>о</strong>н<br />

3. Е<strong>в</strong>р<strong>о</strong>пы (Hirst, 1923). 1341. Protacarus crani Hirst; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X400.0, де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы (Hirst, 1923). 1342. Petralycus unicor­<br />

462<br />

http://jurassic.ru/<br />

nis Grandjean; <strong>о</strong>бщий <strong>в</strong>ид, X600.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Grandjean, 1943).<br />

1343. Pseudoprotacarus scoticus W. Dubinin, gen. nov , sp nov •<br />

<strong>о</strong>бщий <strong>в</strong>ид, X450.0, де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы (Hirst, 1923). 1344. Alicorhagia<br />

fragilis Berlese; A — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х450.0;<br />

Б — н<strong>о</strong>г<strong>а</strong> пер<strong>в</strong><strong>а</strong>я, Х 1500, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Grandjean, 1939)

1342<br />

http://jurassic.ru/

СЕМЕЙСТВО PACHYGNATHIDAE<br />

KRAMER, 1877<br />

Тел<strong>о</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong>, р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>е<br />

н<strong>а</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му и<br />

ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му (<strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>идный тип). Ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong><strong>с</strong>членен<strong>а</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1341, 1342). П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы мяг<strong>к</strong>ие,<br />

<strong>с</strong> б<strong>о</strong>р<strong>о</strong>здч<strong>а</strong>тыми, <strong>с</strong>етч<strong>а</strong>тыми или иными<br />

нежными уз<strong>о</strong>р<strong>а</strong>ми. Передний <strong>к</strong>р<strong>а</strong>й пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>о</strong>бр<strong>а</strong>зует <strong>в</strong>ы<strong>с</strong>туп (те<strong>к</strong>тум, р<strong>о</strong><strong>с</strong>трум),<br />

н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющий ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> н<strong>а</strong>д <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нием<br />

гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы д<strong>в</strong>е п<strong>а</strong>ры <strong>с</strong>ен<strong>с</strong><strong>о</strong>рных щетин<strong>о</strong><strong>к</strong>.<br />

Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong>рупн<strong>а</strong>я.<br />

Педип<strong>а</strong>льпы у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>в</strong>ид<strong>о</strong><strong>в</strong> пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>а</strong> у и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>г<strong>о</strong> Protacarus<br />

crani Hirst <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ят из ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и пери<strong>с</strong>тым эмп<strong>о</strong>дием.<br />

Н<strong>о</strong>ги I—III у <strong>с</strong><strong>о</strong><strong>в</strong>ременных <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>а</strong> н<strong>о</strong>ги IV <strong>с</strong> бедр<strong>о</strong>м, р<strong>а</strong>зделенным<br />

н<strong>а</strong> basi- и telofemur. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ых<br />

при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong> три п<strong>а</strong>ры. Мел<strong>к</strong>ие бел<strong>о</strong><strong>в</strong><strong>а</strong>тые<br />

<strong>к</strong>лещи (длин<strong>а</strong> тел<strong>а</strong> 0,3—0,4 мм), <strong>о</strong>бит<strong>а</strong>ющие<br />

<strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е и <strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й ле<strong>с</strong>н<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е.<br />

Де<strong>в</strong><strong>о</strong>н — ныне. Четыре р<strong>о</strong>д<strong>а</strong>, три из них<br />

<strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

Protacarus Hirst, 1923. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. crani Hirst, 1923 (ч<strong>а</strong><strong>с</strong>тью, э<strong>к</strong>з. № 2;<br />

т<strong>а</strong>бл. XI, ри<strong>с</strong>. 6); де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия<br />

(Aberdeenshire, <strong>к</strong>р<strong>а</strong><strong>с</strong>ный пе<strong>с</strong>ч<strong>а</strong>ни<strong>к</strong>). Н<strong>а</strong><br />

ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>ме п<strong>о</strong>перечные б<strong>о</strong>р<strong>о</strong>зды; гл<strong>а</strong>з<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют: н<strong>о</strong>ги ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; щетин<strong>к</strong>и<br />

н<strong>а</strong> теле и трих<strong>о</strong>б<strong>о</strong>трий н<strong>а</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>ме<br />

<strong>с</strong>лег<strong>к</strong><strong>а</strong> бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идные, из<strong>о</strong>гнутые,<br />

педип<strong>а</strong>льпы <strong>с</strong> р<strong>а</strong><strong>с</strong>члененными бедр<strong>а</strong>ми.<br />

Р<strong>а</strong>змеры мел<strong>к</strong>ие (ри<strong>с</strong>. 1341). Один <strong>в</strong>ид.<br />

СЕМЕЙСТВО ALICORHAGIIDAE<br />

GRANDJEAN, 1939<br />

Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным ш<strong>в</strong><strong>о</strong>м<br />

(ри<strong>с</strong>. 1343, 1344). П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы мяг<strong>к</strong>ие, <strong>с</strong> буг<strong>о</strong>рч<strong>а</strong>тыми<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми. Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й<br />

п<strong>а</strong>р<strong>о</strong>й трих<strong>о</strong>б<strong>о</strong>трий; п<strong>а</strong>льце<strong>в</strong>идный<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т переднег<strong>о</strong> ее <strong>к</strong>р<strong>а</strong>я, н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющий<br />

н<strong>а</strong>д гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, <strong>с</strong> д<strong>в</strong>умя щетин<strong>к</strong><strong>а</strong>ми.<br />

Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Педип<strong>а</strong>льпы пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

пр<strong>о</strong><strong>с</strong>тые. Н<strong>о</strong>ги <strong>с</strong> пятью п<strong>о</strong>д<strong>в</strong>ижными<br />

члени<strong>к</strong><strong>а</strong>ми; бедр<strong>а</strong> их р<strong>а</strong>зделены н<strong>а</strong><br />

basi- и telofemur; т<strong>а</strong><strong>к</strong> же р<strong>а</strong>зделены <strong>в</strong>ертлуги<br />

н<strong>о</strong>г I и II (ри<strong>с</strong>. 1344Б). П<strong>о</strong>л<strong>о</strong><strong>в</strong>ых<br />

при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong> д<strong>в</strong>е п<strong>а</strong>ры. Мел<strong>к</strong>ие <strong>к</strong>лещи (длин<strong>а</strong><br />

тел<strong>а</strong> 0,22—0,30 мм), <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong><strong>о</strong><br />

<strong>в</strong>л<strong>а</strong>жн<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е ле<strong>с</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1344).<br />

Де<strong>в</strong><strong>о</strong>н — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

<strong>о</strong>дин р<strong>о</strong>д, <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>о</strong>д — <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

1348<br />

Ри<strong>с</strong>. 1345—1350. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Cryptognathidae, Tydeidae и<br />

Penthalodidae<br />

1345. Cryptognathus rhombeus (Koch et Berendt); A — <strong>о</strong>бщий <strong>в</strong>ид<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, xlOO.O; Б —<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и н<strong>а</strong> л<strong>а</strong>п<strong>к</strong>е пер<strong>в</strong><strong>о</strong>й<br />

н<strong>о</strong>ги, хбОО.О, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

et Berendt, 1854). 1346. Palaeotydeus devonicus W. Dubinin, gen.<br />

nov., sp. nov.; A — л<strong>а</strong>п<strong>к</strong><strong>а</strong> х<strong>о</strong>дильн<strong>о</strong>й н<strong>о</strong>ги, X600.0; Б — <strong>в</strong>ид<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х300.0; де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы (Hirst, 1923). 1347. Р<strong>а</strong>raprotacarus<br />

hirstl W. Dubinin, gen. nov., sp. nov.; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, хЗОО.О, де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы (Hirst, 1923). 1348.<br />

Lorryia superba Oudemans; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

300,0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Thor, 1933). 1349. Tydeus (Triophtydeus) pinicolus<br />

(Oudemans); <strong>о</strong>бщий <strong>в</strong>ид, X200.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Thor, 1933).<br />

1350. Penthalodes tristiculus(K.och et Berandt); <strong>о</strong>бщий <strong>в</strong>ид, X 100,0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854)<br />

464<br />

http://jurassic.ru/

Р<strong>о</strong>д Pseudoprotacarus W. Dubinin, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. scoticus W. Dubinin, nom. nov.<br />

(Protacarus crani Hirst, 1923; ч<strong>а</strong><strong>с</strong>тью: э<strong>к</strong>земпляр<br />

№ 1. Клещ <strong>с</strong>б<strong>о</strong><strong>к</strong>у — pi. XI, fig. <strong>а</strong>); де<strong>в</strong><strong>о</strong>н,<br />

Ш<strong>о</strong>тл<strong>а</strong>ндия. Тел<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е ,<strong>с</strong> меш<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong>рупн<strong>а</strong>я;<br />

гип<strong>о</strong><strong>с</strong>т<strong>о</strong>м <strong>с</strong> шил<strong>о</strong><strong>в</strong>идным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м н<strong>а</strong><br />

переднем <strong>к</strong><strong>о</strong>нце. Хелицеры <strong>к</strong>рупные, <strong>к</strong>лешне<strong>в</strong>идные.<br />

Н<strong>о</strong>ги б<strong>о</strong>льшие, <strong>с</strong> т<strong>о</strong>л<strong>с</strong>тыми л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми,<br />

пяти-ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими,<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> из<strong>о</strong>гнутыми хет<strong>а</strong>ми. Н<strong>а</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не з<strong>а</strong>днег<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> тел<strong>а</strong>, <strong>в</strong>идим<strong>о</strong>, п<strong>о</strong><br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й щели, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я четырьмя<br />

ряд<strong>а</strong>ми <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие из<strong>о</strong>гнутые щетин<strong>к</strong>и,<br />

п<strong>о</strong> три и четыре щетин<strong>к</strong>и <strong>в</strong> <strong>о</strong>дн<strong>о</strong>м ряду. Щетин<strong>к</strong>и<br />

тел<strong>а</strong> и н<strong>о</strong>г, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, т<strong>о</strong>н<strong>к</strong><strong>о</strong><strong>о</strong>пуш<strong>о</strong>нные,<br />

чт<strong>о</strong> не <strong>о</strong>тмечен<strong>о</strong> Хир<strong>с</strong>т<strong>о</strong>м (Hirst, 1923) ни <strong>в</strong><br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>нии, ни н<strong>а</strong> ри<strong>с</strong>ун<strong>к</strong>е. Длин<strong>а</strong> тел<strong>а</strong> 0,29 мм<br />

(ри<strong>с</strong>. 1343). Один <strong>в</strong>ид. Де<strong>в</strong><strong>о</strong>н Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

НАДСЕМЕЙСТВО<br />

RHAPHIGNATHIDEA GRANDJEAN, 1944<br />

Тел<strong>о</strong> <strong>с</strong>литн<strong>о</strong>е; п<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> между<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

Щит<strong>к</strong>и н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> имеют<strong>с</strong>я или<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong> длину,<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идн<strong>а</strong>я: педип<strong>а</strong>льпы зн<strong>а</strong>чительн<strong>о</strong><br />

длиннее р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>а</strong>. У <strong>о</strong>дних <strong>в</strong>ид<strong>о</strong><strong>в</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, <strong>а</strong> у других (Cryptognathidae)<br />

з<strong>а</strong><strong>к</strong>лючен<strong>а</strong> <strong>в</strong> труб<strong>к</strong><strong>о</strong><strong>в</strong>идный <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы (ри<strong>с</strong>. 1345А). Хелицеры<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>тилет<strong>о</strong><strong>в</strong>идными п<strong>а</strong>льц<strong>а</strong>ми. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> <strong>о</strong>бычный н<strong>а</strong>б<strong>о</strong>р щетин<strong>о</strong><strong>к</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

тел<strong>а</strong> т<strong>о</strong>н<strong>к</strong>ие, и<strong>с</strong>черченные или <strong>с</strong> <strong>с</strong>етч<strong>а</strong>тым<br />

<strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м. Н<strong>о</strong>ги пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>, <strong>с</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и эмп<strong>о</strong>дием; у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> бедр<strong>о</strong> р<strong>а</strong>зделен<strong>о</strong> н<strong>а</strong> д<strong>в</strong>е ч<strong>а</strong><strong>с</strong>ти. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые<br />

при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Rhaphignathidae,<br />

Cunaxidae, Pomerantziidae, Cryptognathidae;<br />

п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО CRYPTOGNATH1DAE OUDEMANS, 1902<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и ги<strong>с</strong>тер<strong>о</strong>е<strong>о</strong>м<strong>а</strong><br />

не р<strong>а</strong>зделены п<strong>о</strong>перечн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

<strong>с</strong>л<strong>а</strong>б<strong>о</strong> хитиниз<strong>о</strong><strong>в</strong><strong>а</strong>ны, <strong>с</strong> т<strong>о</strong>чечным <strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м,<br />

н<strong>а</strong> <strong>с</strong>пине — <strong>с</strong> <strong>с</strong>етч<strong>а</strong>тым уз<strong>о</strong>р<strong>о</strong>м. Передний<br />

<strong>к</strong>р<strong>а</strong>й пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я <strong>в</strong>перед, <strong>о</strong>бр<strong>а</strong>зуя<br />

труб<strong>к</strong>у, <strong>о</strong>т<strong>к</strong>рытую <strong>с</strong>низу; гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, <strong>с</strong><strong>к</strong>рыт<strong>а</strong>я<br />

<strong>в</strong> эт<strong>о</strong>й труб<strong>к</strong>е, м<strong>о</strong>жет <strong>в</strong>ыд<strong>в</strong>иг<strong>а</strong>ть<strong>с</strong>я или<br />

<strong>в</strong>тяги<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong>нутрь. Хелицеры <strong>к</strong>лешне<strong>в</strong>идные,<br />

<strong>с</strong> м<strong>а</strong>лень<strong>к</strong>ими п<strong>а</strong>льц<strong>а</strong>ми, без зуб<strong>о</strong><strong>в</strong>; <strong>о</strong>ни з<strong>а</strong><strong>о</strong><strong>с</strong>трены<br />

н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х и при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены для пр<strong>о</strong><strong>к</strong><strong>а</strong>лы<strong>в</strong><strong>а</strong>ния.<br />

Педип<strong>а</strong>льпы т<strong>о</strong>н<strong>к</strong>ие, прямые, <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ют <strong>в</strong> <strong>в</strong>иде м<strong>а</strong>лень<strong>к</strong>их «р<strong>о</strong>же<strong>к</strong>» н<strong>а</strong><br />

переднем <strong>к</strong><strong>о</strong>нце пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льн<strong>о</strong>й труб<strong>к</strong>и<br />

30 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 465<br />

http://jurassic.ru/<br />

(ри<strong>с</strong>. 1345). Н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong><br />

д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. Мел<strong>к</strong>ие <strong>к</strong>лещи (длин<strong>а</strong> тел<strong>а</strong><br />

0,30—0,40 мм), <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong><strong>о</strong> мху, п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями,<br />

п<strong>о</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>й и <strong>в</strong> гнезд<strong>а</strong>х птиц и бел<strong>о</strong><strong>к</strong>. Вер<strong>о</strong>ятн<strong>о</strong>,<br />

хищни<strong>к</strong>и. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Один р<strong>о</strong>д<br />

Cryptognathus Kramer, 1878, из<strong>в</strong>е<strong>с</strong>тный т<strong>а</strong><strong>к</strong>же<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО TYDEIDEA.<br />

КЛЕЩИ ТИДЕИДЕИ<br />

(Eupodidea, pars)<br />

Мел<strong>к</strong>ие <strong>к</strong>лещи <strong>с</strong> мяг<strong>к</strong>ими п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми, и<strong>с</strong>черченными<br />

пун<strong>к</strong>тирными линиями. Иди<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> не<br />

<strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong>; имеет<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>перечн<strong>а</strong>я<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> между пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

(ри<strong>с</strong>. 1346). Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, <strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong><strong>а</strong>я;<br />

п<strong>а</strong>льпы пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые у <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущих<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> и трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые у Speleognathidae<br />

Womersley, п<strong>а</strong>р<strong>а</strong>зитирующие <strong>в</strong> н<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong>ых п<strong>о</strong>л<strong>о</strong><strong>с</strong>тях<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных. Хелицеры <strong>с</strong> т<strong>о</strong>л<strong>с</strong>тыми <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>литыми друг <strong>с</strong> друг<strong>о</strong>м, и<br />

п<strong>а</strong>льце<strong>в</strong>идн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й. П<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец<br />

хелицер <strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong>ий и имеет <strong>в</strong>ид <strong>о</strong><strong>с</strong>тр<strong>о</strong>г<strong>о</strong><br />

шип<strong>а</strong> или <strong>к</strong><strong>о</strong>люч<strong>к</strong>и; п<strong>а</strong>льцы хелицер не пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

друг другу. Передние и з<strong>а</strong>дние<br />

п<strong>а</strong>ры н<strong>о</strong>г уд<strong>а</strong>лены друг <strong>о</strong>т друг<strong>а</strong>; н<strong>о</strong>ги пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые;<br />

л<strong>а</strong>п<strong>к</strong>и их <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и<br />

пери<strong>с</strong>тым эмп<strong>о</strong>дием. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

тел<strong>а</strong> <strong>о</strong>т <strong>с</strong>еми д<strong>о</strong> д<strong>в</strong>ен<strong>а</strong>дц<strong>а</strong>ти п<strong>а</strong>р щетин<strong>к</strong><strong>о</strong><strong>в</strong>идных<br />

хет.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> п<strong>а</strong>р<strong>а</strong>зиты н<strong>а</strong>земных<br />

м<strong>о</strong>ллю<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>, земн<strong>о</strong><strong>в</strong><strong>о</strong>дных и птиц. Де<strong>в</strong><strong>о</strong>н<br />

— ныне. Семь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне: Speleognathidae, Ereynatidae, Paratydeidae,<br />

Eupodidae, Penthaleidae, Tydeidae и<br />

Penthalodidae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО TYDEIDAE KRAMER, 1877<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е <strong>с</strong> нея<strong>с</strong>ными, мяг<strong>к</strong>ими п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми,<br />

и<strong>с</strong>черченными т<strong>о</strong>н<strong>к</strong>ими пун<strong>к</strong>тирными б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>ми.<br />

П<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> тел<strong>а</strong> имеет<strong>с</strong>я<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Щиты р<strong>а</strong>з<strong>в</strong>иты <strong>с</strong>л<strong>а</strong>б<strong>о</strong>. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я. Педип<strong>а</strong>льпы <strong>с</strong><br />

пятью •— ше<strong>с</strong>тью члени<strong>к</strong><strong>а</strong>ми. Хелицеры <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие;<br />

п<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец их <strong>к</strong>инж<strong>а</strong>л<strong>о</strong><strong>в</strong>идный,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленный для пр<strong>о</strong><strong>к</strong><strong>а</strong>лы<strong>в</strong><strong>а</strong>ния, зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че и т<strong>о</strong>ньше т<strong>о</strong>л<strong>с</strong>т<strong>о</strong>г<strong>о</strong> неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong><br />

п<strong>а</strong>льц<strong>а</strong> и не пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет<strong>с</strong>я ему (ри<strong>с</strong>.<br />

1346, 1347). Н<strong>о</strong>ги х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong><br />

д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и пери<strong>с</strong>тым эмп<strong>о</strong>дием, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый<br />

м<strong>о</strong>жет быть <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идным. Очень <strong>к</strong>рупн<strong>о</strong>е<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е и щеле<strong>в</strong>идн<strong>о</strong>е <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти брюшн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

тел<strong>а</strong>. Де<strong>в</strong><strong>о</strong>н — ныне. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне и д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.

Palaeotydeus W. Dubinin, gen. nov. Тип<br />

р<strong>о</strong>д<strong>а</strong> — P. devonicus W. Dubinin, nom. nov.<br />

(Protacarus crani Hirst, 1923, ч<strong>а</strong><strong>с</strong>тью: э<strong>к</strong>земпляр<br />

«№ 3. Клещ <strong>с</strong>б<strong>о</strong><strong>к</strong>у. Н<strong>о</strong>ги II и IV б<strong>о</strong>лее<br />

длинные». Hirst, 1923, р. 459, textfig. 1); де<strong>в</strong><strong>о</strong>н,<br />

Ш<strong>о</strong>тл<strong>а</strong>ндия. (Aberdeenshire, <strong>к</strong>р<strong>а</strong><strong>с</strong>ный пе<strong>с</strong>ч<strong>а</strong>ни<strong>к</strong>).<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е; пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> з<strong>а</strong>ним<strong>а</strong>ет<br />

треть длины, <strong>о</strong>тделен<strong>а</strong> <strong>о</strong>т ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й; гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я, шил<strong>о</strong><strong>в</strong>идный<br />

п<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец хелицер <strong>к</strong><strong>о</strong>нц<strong>о</strong>м не д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет<br />

<strong>в</strong>ершины б<strong>о</strong>лее т<strong>о</strong>л<strong>с</strong>т<strong>о</strong>г<strong>о</strong> неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong><br />

п<strong>а</strong>льц<strong>а</strong>. Педип<strong>а</strong>льпы пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Н<strong>о</strong>ги<br />

ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>о</strong>бычн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ения; н<strong>о</strong>ги II<br />

и IV зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>о</strong><strong>с</strong>т<strong>а</strong>льных п<strong>а</strong>р.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя длинными <strong>с</strong><strong>о</strong>гнутыми <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

и т<strong>а</strong><strong>к</strong><strong>о</strong>й же длины эмп<strong>о</strong>дием между ними<br />

(ри<strong>с</strong>. 1346). Генит<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong><br />

н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний н<strong>о</strong>г IV; <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

термин<strong>а</strong>льн<strong>о</strong>е, н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 0,31 мм. Один <strong>в</strong>ид. Де<strong>в</strong><strong>о</strong>н Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

Paraprotacarus W. Dubinin, gen. nov.<br />

Тип р<strong>о</strong>д<strong>а</strong> — P. hirsti W. Dubinin, nom. nov.<br />

(Protacarus crani (?) Hirst, 1923, ч<strong>а</strong><strong>с</strong>тью: э<strong>к</strong>земпляр<br />

«№ 4. Вид <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны», Hirst,<br />

1923, р. 458—459; textfig. 2, la); де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия<br />

(Aberdeenshire, <strong>к</strong>р<strong>а</strong><strong>с</strong>ный пе<strong>с</strong>ч<strong>а</strong>ни<strong>к</strong>). Тел<strong>о</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е; пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет меньше<br />

трети длины тел<strong>а</strong>; гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong> длину,<br />

треуг<strong>о</strong>льн<strong>а</strong>я; хелицеры четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

(?); п<strong>а</strong>л<strong>в</strong>пы длинные, ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

В<strong>с</strong>е н<strong>о</strong>ги <strong>с</strong> ше<strong>с</strong>тью члени<strong>к</strong><strong>а</strong>ми, р<strong>а</strong>з<strong>в</strong>иты н<strong>о</strong>рм<strong>а</strong>льн<strong>о</strong>,<br />

примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й длины. Крупн<strong>о</strong>е<br />

генит<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>в</strong> <strong>в</strong>иде б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> п<strong>о</strong>з<strong>а</strong>ди ур<strong>о</strong><strong>в</strong>ня <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний<br />

н<strong>о</strong>г IV; <strong>о</strong>н<strong>о</strong> приближен<strong>о</strong> <strong>к</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й щели,<br />

леж<strong>а</strong>щей <strong>с</strong>уб<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>. Длин<strong>а</strong> тел<strong>а</strong> 0,31 мм<br />

(ри<strong>с</strong>. 1347). Один <strong>в</strong>ид. Де<strong>в</strong><strong>о</strong>н Ш<strong>о</strong>тл<strong>а</strong>ндии.<br />

СЕМЕЙСТВО PENTHALODIDAE THOR, 1933<br />

Тел<strong>о</strong> груше<strong>в</strong>идн<strong>о</strong>е, <strong>с</strong>уж<strong>а</strong>ющее<strong>с</strong>я <strong>в</strong> з<strong>а</strong>дней<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине. П<strong>о</strong>перечн<strong>а</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong> между пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

<strong>о</strong>бычн<strong>о</strong> бы<strong>в</strong><strong>а</strong>ют <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>ны,<br />

<strong>с</strong> <strong>с</strong>етч<strong>а</strong>тым или т<strong>о</strong>чечным <strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м,<br />

у жи<strong>в</strong>ых <strong>к</strong>лещей черн<strong>о</strong>г<strong>о</strong> ц<strong>в</strong>ет<strong>а</strong> <strong>с</strong> <strong>к</strong>р<strong>а</strong><strong>с</strong>ными<br />

пятн<strong>а</strong>ми. Н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>му<br />

гл<strong>а</strong>зу. Щетин<strong>к</strong>и н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х и теле немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие. Н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong><br />

ше<strong>с</strong>тью п<strong>о</strong>д<strong>в</strong>ижными члени<strong>к</strong><strong>а</strong>ми; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и <strong>о</strong>пуш<strong>о</strong>нным эмп<strong>о</strong>дием. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

неб<strong>о</strong>льш<strong>а</strong>я, треуг<strong>о</strong>льн<strong>а</strong>я. Педип<strong>а</strong>льпы<br />

т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong> четырьмя члени<strong>к</strong><strong>а</strong>ми. Хелицеры м<strong>а</strong>лень<strong>к</strong>ие,<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> и<strong>с</strong><strong>к</strong>ри<strong>в</strong>ленные, н<strong>о</strong>жнице<strong>о</strong>бр<strong>а</strong>зные.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong>ых при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong> д<strong>в</strong>е п<strong>а</strong>ры. Длин<strong>а</strong><br />

гел<strong>а</strong> <strong>в</strong><strong>а</strong>рьирует <strong>о</strong>т 0,33 д<strong>о</strong> 1,50 мм. С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие<br />

<strong>в</strong>иды, <strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong><strong>о</strong> мху и гниющих<br />

466<br />

http://jurassic.ru/<br />

р<strong>а</strong><strong>с</strong>тительных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>а</strong>х (ри<strong>с</strong>. 1350). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Три р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне, из<br />

них Penthalodes Murray, 1877 <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО ANYSTIIDEA<br />

(Anystoidea)<br />

Мел<strong>к</strong>ие <strong>к</strong>р<strong>а</strong><strong>с</strong>ные <strong>к</strong>лещи <strong>с</strong> <strong>о</strong><strong>к</strong>руглым тел<strong>о</strong>м.<br />

Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и ги<strong>с</strong>тер<strong>о</strong>е<strong>о</strong>м<strong>а</strong> <strong>с</strong>литы или р<strong>а</strong>зделены<br />

ед<strong>в</strong><strong>а</strong> н<strong>а</strong>меченным ш<strong>в</strong><strong>о</strong>м. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы т<strong>о</strong>н<strong>к</strong>ие,<br />

гу<strong>с</strong>т<strong>о</strong> и<strong>с</strong>черченные штрих<strong>а</strong>ми. Имеют<strong>с</strong>я пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий,<br />

<strong>а</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий<br />

щиты. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>е<br />

<strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> щетин<strong>о</strong><strong>к</strong>; трих<strong>о</strong>б<strong>о</strong>трии<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong>рупн<strong>а</strong>я.<br />

П<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец хелицер <strong>с</strong>тилет<strong>о</strong><strong>в</strong>идный,<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блен для пр<strong>о</strong><strong>к</strong><strong>а</strong>лы<strong>в</strong><strong>а</strong>ния. Педип<strong>а</strong>льпы<br />

длинные, н<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные или <strong>с</strong> <strong>к</strong>рю<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

г<strong>о</strong>ленью и м<strong>а</strong>лень<strong>к</strong><strong>о</strong>й л<strong>а</strong>п<strong>к</strong><strong>о</strong>й, причленяющей<strong>с</strong>я<br />

н<strong>а</strong> ее <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не. Н<strong>о</strong>ги<br />

длинные, р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г<br />

I и II имеют п<strong>о</strong> <strong>о</strong>дн<strong>о</strong>му <strong>с</strong>ен<strong>с</strong><strong>о</strong>рн<strong>о</strong>му <strong>о</strong>рг<strong>а</strong>ну.<br />

З<strong>а</strong>днепр<strong>о</strong>х<strong>о</strong>дн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие термин<strong>а</strong>льн<strong>о</strong>е или<br />

<strong>с</strong>убд<strong>о</strong>рз<strong>а</strong>льн<strong>о</strong>е. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие хищные <strong>к</strong>лещи, <strong>о</strong>бит<strong>а</strong>ющие<br />

<strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, <strong>в</strong><strong>о</strong> мху и н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях. Длин<strong>а</strong><br />

тел<strong>а</strong> <strong>о</strong>т 0,55 д<strong>о</strong> 1,50 мм. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО ANYST1DAE OUDEMANS, 1902<br />

Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближены <strong>в</strong> передней трети<br />

тел<strong>а</strong>; н<strong>о</strong>ги <strong>о</strong>чень длинные, <strong>о</strong>тх<strong>о</strong>дят <strong>о</strong>т тел<strong>а</strong><br />

<strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, луче<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> (ри<strong>с</strong>. 1351). Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий<br />

щит<strong>о</strong><strong>к</strong> имеет<strong>с</strong>я не <strong>в</strong><strong>с</strong>егд<strong>а</strong>, н<strong>о</strong> д<strong>а</strong>же<br />

<strong>к</strong><strong>о</strong>гд<strong>а</strong> <strong>о</strong>н е<strong>с</strong>ть, ф<strong>а</strong><strong>к</strong>тиче<strong>с</strong><strong>к</strong>и имеет ме<strong>с</strong>т<strong>о</strong> не<br />

<strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>а</strong>ция п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong>, <strong>а</strong> р<strong>а</strong>з<strong>в</strong>итие пл<strong>о</strong>тн<strong>о</strong>г<strong>о</strong><br />

б<strong>о</strong>р<strong>о</strong>здч<strong>а</strong>т<strong>о</strong>г<strong>о</strong> <strong>о</strong>рн<strong>а</strong>мент<strong>а</strong> н<strong>а</strong> <strong>к</strong><strong>о</strong>же. Перитремы<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии хелицер. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идн<strong>а</strong>я.<br />

П<strong>о</strong>д<strong>в</strong>ижный п<strong>а</strong>лец хелицер <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зный.<br />

Педип<strong>а</strong>льпы <strong>к</strong>рупные, н<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные;<br />

г<strong>о</strong>лень их <strong>с</strong> <strong>о</strong>дним — тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми;<br />

л<strong>а</strong>п<strong>к</strong>и длинные, «<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>а</strong>тые», причленяют<strong>с</strong>я <strong>к</strong><br />

г<strong>о</strong>лени <strong>с</strong>уб<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>. Длин<strong>а</strong> тел<strong>а</strong> 0,55—<br />

1,30 мм.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие хищные <strong>к</strong>лещи, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные н<strong>а</strong> ли<strong>с</strong>тьях р<strong>а</strong><strong>с</strong>тений. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 17 р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

из них Anystis Heyden, 1826 из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО BDELLIDEA.<br />

БДЕЛЛИДЫ, ИЛИ НОСАТЫЕ КЛЕЩИ<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, нежн<strong>о</strong>е, р<strong>а</strong>зделен<strong>о</strong> глуб<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й н<strong>а</strong> пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>о</strong><strong>о</strong>му и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му.<br />

Хелицеры и гип<strong>о</strong><strong>с</strong>т<strong>о</strong>мы <strong>в</strong>ытянуты <strong>в</strong> длинный<br />

<strong>к</strong>лю<strong>в</strong>, <strong>о</strong>т <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дят <strong>к</strong><strong>о</strong>ленч<strong>а</strong>-

1352<br />

Ри<strong>с</strong>. 1351—1356. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Anystidae, Bdellidae, Cheyletidiae, Erythraeidae<br />

1351. Anystis venusiula (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид, X17.0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1352. Bdellodes lata (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид, X50.0;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1353. Cheyletus portentosus Koch et Berendt, X40.0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854).<br />

тые т<strong>о</strong>н<strong>к</strong>ие п<strong>а</strong>льтгы <strong>с</strong> пятью члени<strong>к</strong><strong>а</strong>ми; н<strong>а</strong><br />

<strong>в</strong>ершине л<strong>а</strong>п<strong>о</strong><strong>к</strong> их п<strong>о</strong> д<strong>в</strong>е т<strong>о</strong>рч<strong>а</strong>щих <strong>в</strong>перед<br />

щетин<strong>к</strong>и (ри<strong>с</strong>. 1352). Хелицеры удлиненные, <strong>с</strong><br />

<strong>о</strong>дн<strong>о</strong>й или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими д<strong>о</strong>р<strong>с</strong><strong>а</strong>льными щетин<strong>к</strong><strong>а</strong>ми<br />

и <strong>с</strong> <strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong>ими <strong>к</strong>лешнями. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

т<strong>о</strong>н<strong>к</strong>ие, б<strong>о</strong>р<strong>о</strong>здч<strong>а</strong>тые; щиты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены д<strong>в</strong>е п<strong>а</strong>ры трих<strong>о</strong>б<strong>о</strong>трий и д<strong>в</strong>е п<strong>а</strong>ры<br />

р<strong>а</strong>зделенных гл<strong>а</strong>з; не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые р<strong>о</strong>ды имеют<br />

неп<strong>а</strong>рный <strong>с</strong>рединный гл<strong>а</strong>з н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы. Н<strong>о</strong>ги <strong>с</strong> ше<strong>с</strong>тью п<strong>о</strong>д<strong>в</strong>ижными<br />

члени<strong>к</strong><strong>а</strong>ми. Три п<strong>а</strong>ры п<strong>о</strong>л<strong>о</strong><strong>в</strong>ых при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong>. Длин<strong>а</strong><br />

тел<strong>а</strong> 0,45—1,50 мм. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие хищни<strong>к</strong>и, <strong>о</strong>бит<strong>а</strong>ющие<br />

<strong>в</strong><strong>с</strong>юду <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>жных <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>а</strong>х, где е<strong>с</strong>ть другие<br />

<strong>к</strong>лещи или <strong>с</strong><strong>к</strong><strong>о</strong>пления мел<strong>к</strong>их н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых.<br />

Bdellidae <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>в</strong><strong>о</strong> мху и<br />

ли<strong>с</strong>т<strong>в</strong>енн<strong>о</strong>й п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е, ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

ли<strong>с</strong>тьях р<strong>а</strong><strong>с</strong>тений. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО BDELLIDAE DUGES, 1834<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я призн<strong>а</strong><strong>к</strong><strong>а</strong>ми, перечи<strong>с</strong>ленными<br />

<strong>в</strong> ди<strong>а</strong>гн<strong>о</strong>зе н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> — Bdellodes Oudemans,1937 и Bdella<br />

Latreille, 1795 — из<strong>в</strong>е<strong>с</strong>тны из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

1356<br />

1353<br />

1354. Leptus incertus (Koch et Berendt); X 4<br />

- 0<br />

- п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854), 1355. Balaustium<br />

illustrls (Koch et Berendt), x40,0, п<strong>а</strong>ле<strong>о</strong>ген. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1356. Erythraeus foveolatus<br />

(Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид, X4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1854)<br />

НАДСЕМЕЙСТВО CHEYLETIDEA.<br />

ХИЩНЫЕ КЛЕЩИ<br />

Мел<strong>к</strong>ие и <strong>с</strong>редней <strong>в</strong>еличины <strong>к</strong>лещи <strong>с</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льным<br />

или <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ытянутым тел<strong>о</strong>м. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

треуг<strong>о</strong>льн<strong>а</strong>я, <strong>с</strong> <strong>к</strong>рупным <strong>с</strong>рединным хелицер<strong>а</strong>льным<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong>м — «<strong>к</strong>лю<strong>в</strong><strong>о</strong>м», <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м з<strong>а</strong><strong>к</strong>лючены<br />

<strong>с</strong>тилет<strong>о</strong><strong>в</strong>идные хелицеры. Педип<strong>а</strong>льпы<br />

трех-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; <strong>в</strong>ершины г<strong>о</strong>леней<br />

п<strong>а</strong>льп <strong>в</strong>ытянуты <strong>в</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>ть, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

причленяет<strong>с</strong>я м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я л<strong>а</strong>п<strong>к</strong><strong>а</strong>; ин<strong>о</strong>гд<strong>а</strong> педип<strong>а</strong>льпы<br />

<strong>в</strong>ид<strong>о</strong>изменены <strong>в</strong> м<strong>о</strong>щные х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные<br />

<strong>о</strong>рг<strong>а</strong>ны (ри<strong>с</strong>. 1353). Х<strong>о</strong>дильные н<strong>о</strong>ги р<strong>а</strong>з<strong>в</strong>иты<br />

р<strong>а</strong>зличн<strong>о</strong>; у <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущих <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>ни<br />

длинные, <strong>о</strong>бычн<strong>о</strong>г<strong>о</strong> р<strong>а</strong>змер<strong>а</strong> и <strong>с</strong>тр<strong>о</strong>ения, <strong>а</strong> у<br />

п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их м<strong>о</strong>гут быть у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены или<br />

при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>блены для з<strong>а</strong>щемления <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong> жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong>-х<strong>о</strong>зяин<strong>а</strong>.<br />

Перитремы н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>к</strong>рупные, чет<strong>к</strong><strong>о</strong><strong>в</strong>идные. Ведут <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дный<br />

или п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ий <strong>о</strong>бр<strong>а</strong>з жизни. Хищни<strong>к</strong>и.<br />

Длин<strong>а</strong> тел<strong>а</strong> 0,2—0,8 мм. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>:<br />

Myobiidae, Syringophilidae, Harpirhynchidae,<br />

Cheyletidae; п<strong>о</strong><strong>с</strong>леднее из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

467 30*<br />

http://jurassic.ru/

СЕМЕЙСТВО CHEYLETIDAE LEACH, 1814.<br />

ХИЩНЫЕ КЛЕЩИ ИЛИ ХЕЙЛЕТИДЫ<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>здут<strong>о</strong>е, реже удлиненн<strong>о</strong>е.<br />

Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и ги<strong>с</strong>тер<strong>о</strong>е<strong>о</strong>м<strong>а</strong> чет<strong>к</strong><strong>о</strong><br />

<strong>в</strong>ыделяют<strong>с</strong>я бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>ря <strong>к</strong><strong>о</strong>нфигур<strong>а</strong>ции <strong>к</strong>р<strong>а</strong>е<strong>в</strong><br />

тел<strong>а</strong> и н<strong>а</strong>личию д<strong>в</strong>ух или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>их <strong>с</strong>пинных<br />

щит<strong>о</strong><strong>в</strong>. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> <strong>к</strong>рупным <strong>с</strong>рединным «<strong>к</strong>лю<strong>в</strong><strong>о</strong>м»,<br />

п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>о</strong>чень <strong>к</strong>рупные п<strong>а</strong>льпы, пре<strong>в</strong>р<strong>а</strong>щенные <strong>в</strong> м<strong>о</strong>щные<br />

щипцы <strong>с</strong> т<strong>о</strong>л<strong>с</strong>тыми бедр<strong>а</strong>ми и н<strong>о</strong>гте<strong>в</strong>идными<br />

г<strong>о</strong>ленями; н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х м<strong>а</strong>лень<strong>к</strong>их л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

имеют<strong>с</strong>я пр<strong>о</strong><strong>с</strong>тые, гл<strong>а</strong>д<strong>к</strong>ие и гребенч<strong>а</strong>тые щетин<strong>к</strong>и.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие хищни<strong>к</strong>и; не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

<strong>в</strong>иды регулярн<strong>о</strong> <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> теле птиц и<br />

мле<strong>к</strong><strong>о</strong>пит<strong>а</strong>ющих, где п<strong>о</strong>жир<strong>а</strong>ют п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong>лещей и н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Длин<strong>а</strong> тел<strong>а</strong> 0,20—<br />

0,80 мм (ри<strong>с</strong>. 1353). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 37 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Cheyletus Latreille,<br />

1796. Определение эт<strong>о</strong>й ф<strong>о</strong>рмы требует<br />

ут<strong>о</strong>чнения.<br />

НАДСЕМЕЙСТВО SMARIDIIDEA<br />

(Apobolostigmata)<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> удлиненн<strong>о</strong>е, <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>е,<br />

п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых <strong>к</strong>лещей мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными,<br />

гу<strong>с</strong>т<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными щетин<strong>к</strong><strong>а</strong>ми.<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> т<strong>о</strong>н<strong>к</strong>ие, штрих<strong>о</strong><strong>в</strong><strong>а</strong>нные. П<strong>о</strong><strong>с</strong>редине<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы лежит пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий<br />

щит<strong>о</strong><strong>к</strong>, имеющий ф<strong>о</strong>рму уз<strong>к</strong><strong>о</strong>г<strong>о</strong> п<strong>а</strong>л<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong>иля <strong>с</strong> р<strong>а</strong><strong>с</strong>ширениями н<strong>а</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>х. Имеет<strong>с</strong>я<br />

<strong>о</strong>дн<strong>а</strong> или д<strong>в</strong>е п<strong>а</strong>ры линз<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных гл<strong>а</strong>з. Т<strong>а</strong>зи<strong>к</strong>и<br />

н<strong>о</strong>г р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я д<strong>в</strong>умя групп<strong>а</strong>ми. Н<strong>о</strong>ги<br />

длинные, бег<strong>а</strong>тельные, <strong>с</strong> ше<strong>с</strong>тью п<strong>о</strong>д<strong>в</strong>ижными<br />

члени<strong>к</strong><strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и передних н<strong>о</strong>г <strong>о</strong>бычн<strong>о</strong> бы<strong>в</strong><strong>а</strong>ют<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>здуты; <strong>о</strong>ни <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

и пери<strong>с</strong>тым эмп<strong>о</strong>дием. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong> ф<strong>о</strong>рме<br />

<strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong> <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>а</strong>. Хелицеры <strong>с</strong>тилет<strong>о</strong><strong>в</strong>идные,<br />

длинные, м<strong>о</strong>гут <strong>в</strong>тяги<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong>нутрь тел<strong>а</strong>. Педип<strong>а</strong>льпы<br />

<strong>к</strong>рупные, <strong>с</strong> пятью члени<strong>к</strong><strong>а</strong>ми; г<strong>о</strong>лени<br />

их н<strong>а</strong> <strong>в</strong>ершине <strong>с</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м; л<strong>а</strong>п<strong>к</strong>и педип<strong>а</strong>льп<br />

причленяют<strong>с</strong>я <strong>с</strong>уб<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>, <strong>о</strong>тчег<strong>о</strong> п<strong>а</strong>льпы<br />

<strong>к</strong><strong>а</strong>жут<strong>с</strong>я д<strong>в</strong>у<strong>в</strong>ершинными. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие личин<strong>к</strong>и п<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Длин<strong>а</strong> тел<strong>а</strong> 0,5—2,0 мм (ри<strong>с</strong>.<br />

1354—1356). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Smaridiidae и Erythraeidae;<br />

п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ERYTHRAEIDAE OUDEMANS, 1902<br />

(Balaustiidae Grandjean, 1947)<br />

Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> не <strong>в</strong>ытянут<strong>а</strong> <strong>в</strong> длинный х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>.<br />

Щетин<strong>к</strong>и тел<strong>а</strong> и н<strong>о</strong>г гл<strong>а</strong>д<strong>к</strong>ие или т<strong>о</strong>н<strong>к</strong><strong>о</strong>пери<strong>с</strong>тые.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

468<br />

http://jurassic.ru/<br />

18 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре Leptus<br />

Latreille, 1795 (ри<strong>с</strong>. 1354), Erythraeus Latreille,<br />

1806 (ри<strong>с</strong>. 1356) и Balaustium Heyden, 1826<br />

(ри<strong>с</strong>. 1355). Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>пи<strong>с</strong><strong>а</strong>н <strong>о</strong>дин <strong>в</strong>ымерший<br />

р<strong>о</strong>д из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) —Arythaena Menge, 1954.<br />

НАДСЕМЕЙСТВО TROMBIDIIDEA.<br />

КЛЕЩИ-КРАСНОТЕЛКИ<br />

(Trombeae)<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е или перетянут<strong>о</strong>е п<strong>о</strong><strong>с</strong>редине<br />

(ри<strong>с</strong>. 1357); п<strong>о</strong><strong>к</strong>рыт<strong>о</strong> <strong>о</strong>чень гу<strong>с</strong>тым п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong>м<br />

щетин<strong>о</strong><strong>к</strong>, прид<strong>а</strong>ющих <strong>к</strong>лещ<strong>а</strong>м б<strong>а</strong>рх<strong>а</strong>тный <strong>в</strong>ид:<br />

у личин<strong>о</strong><strong>к</strong> щетин<strong>о</strong><strong>к</strong> н<strong>а</strong> теле неб<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

Тел<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>о</strong>тчетли<strong>в</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong><br />

н<strong>а</strong> гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>му и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>му. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

тел<strong>а</strong> б<strong>о</strong>р<strong>о</strong>здч<strong>а</strong>тые, <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> щетин<strong>о</strong><strong>к</strong> —<br />

гл<strong>а</strong>д<strong>к</strong>ие. Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий щит прям<strong>о</strong>уг<strong>о</strong>льный<br />

у личин<strong>о</strong><strong>к</strong>, н<strong>о</strong> <strong>к</strong>иле<strong>в</strong>идный у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бей. Н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти щит<strong>к</strong><strong>а</strong> <strong>о</strong>дн<strong>а</strong> п<strong>а</strong>р<strong>а</strong> трих<strong>о</strong>б<strong>о</strong>трии,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бы<strong>в</strong><strong>а</strong>ют ут<strong>о</strong>лщенными,<br />

бул<strong>а</strong><strong>в</strong><strong>о</strong><strong>в</strong>идными, и не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>а</strong>р <strong>о</strong>пуш<strong>о</strong>нных<br />

щетин<strong>о</strong><strong>к</strong>. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> неб<strong>о</strong>льш<strong>а</strong>я, треуг<strong>о</strong>льн<strong>а</strong>я;<br />

педип<strong>а</strong>льпы х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты, <strong>с</strong> пятью члени<strong>к</strong><strong>а</strong>ми;<br />

м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я л<strong>а</strong>п<strong>к</strong><strong>а</strong> педип<strong>а</strong>льп причленяет<strong>с</strong>я<br />

н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не г<strong>о</strong>лени. Н<strong>о</strong>ги <strong>с</strong><br />

ше<strong>с</strong>тью п<strong>о</strong>д<strong>в</strong>ижными члени<strong>к</strong><strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие н<strong>а</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong>, н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г IV. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ых<br />

при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong> три п<strong>а</strong>ры. Стигмы <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м члени<strong>к</strong>е хелицер. С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие<br />

<strong>в</strong>иды, хищни<strong>к</strong>и. Личин<strong>к</strong>и п<strong>а</strong>р<strong>а</strong>зитируют<br />

н<strong>а</strong> члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих (Trombidiidae) и н<strong>а</strong> п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных<br />

(Trombiculidae). Длин<strong>а</strong> тел<strong>а</strong> 0,5—<br />

2,0 мм. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Trombidiidae и Trombiculidae;<br />

пер<strong>в</strong><strong>о</strong>е из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО TROMBIDIIDAE LEACH, 1815<br />

Тел<strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых <strong>о</strong><strong>с</strong><strong>о</strong>бей не перетянут<strong>о</strong> п<strong>о</strong><strong>с</strong>редине.<br />

Личин<strong>к</strong>и <strong>о</strong>бычн<strong>о</strong> имеют б<strong>о</strong>лее <strong>о</strong>дн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>пинн<strong>о</strong>г<strong>о</strong> щит<strong>к</strong><strong>а</strong>. Те<strong>к</strong>тум (гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху) <strong>с</strong><br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными щетин<strong>к</strong><strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 60 р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

из них Trombidium Fabricius, 1775 и Allotrom-<br />

Ыит Berlese, 1903 (ри<strong>с</strong>. 1357) из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО TETRANYCHIDEA.<br />

ПАУТИННЫЕ,<br />

ИЛИ ТЕТРАНИХОВЫЕ, КЛЕЩИ<br />

Р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные <strong>к</strong>лещи <strong>с</strong>редней <strong>в</strong>еличины<br />

(длин<strong>а</strong> тел<strong>а</strong> <strong>в</strong> <strong>с</strong>реднем 0,8 мм). Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е,<br />

ин<strong>о</strong>гд<strong>а</strong> груше<strong>в</strong>идн<strong>о</strong>е, <strong>с</strong> б<strong>о</strong>лее или менее<br />

<strong>о</strong>тчетли<strong>в</strong><strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женными плече<strong>в</strong>ыми <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong>-

<strong>с</strong>тями. Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>литы, либ<strong>о</strong><br />

р<strong>а</strong>зделены <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>женным ш<strong>в</strong><strong>о</strong>м. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

мяг<strong>к</strong>ие, щит<strong>к</strong>и не р<strong>а</strong>з<strong>в</strong>иты. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющ<strong>а</strong>я, <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong>рупн<strong>а</strong>я, <strong>с</strong><strong>о</strong> <strong>с</strong>рединным<br />

хелицер<strong>о</strong><strong>в</strong>ым <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong>м (ри<strong>с</strong>. 1358).<br />

Хелицеры <strong>с</strong>тилет<strong>о</strong><strong>в</strong>идные. Педип<strong>а</strong>льпы <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>гте<strong>в</strong>идн<strong>о</strong>й г<strong>о</strong>ленью и м<strong>а</strong>лень<strong>к</strong><strong>о</strong>й л<strong>а</strong>п<strong>к</strong><strong>о</strong>й<br />

н<strong>а</strong> ее <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не. Н<strong>о</strong>ги длинные.<br />

Амбул<strong>а</strong><strong>к</strong>ры и эмп<strong>о</strong>дий н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х м<strong>о</strong>гут быть<br />

<strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Жи<strong>в</strong>ут н<strong>а</strong> зеленых <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>х р<strong>а</strong><strong>с</strong>тений,<br />

пит<strong>а</strong>я<strong>с</strong>ь их <strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>ми и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> причиняя б<strong>о</strong>льш<strong>о</strong>й<br />

<strong>в</strong>ред. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Bryobiidae и Tetranychidae;<br />

п<strong>о</strong><strong>с</strong>леднее из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО TETRANYCHIDAE DONNADIEU, 1875<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> 12—13 п<strong>а</strong>р щетин<strong>о</strong><strong>к</strong>;<br />

эмп<strong>о</strong>дий без желези<strong>с</strong>тых <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>, ин<strong>о</strong>гд<strong>а</strong><br />

<strong>с</strong><strong>о</strong><strong>в</strong>ершенн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н; перитремы при<strong>к</strong>реплены<br />

н<strong>а</strong> <strong>с</strong>тен<strong>к</strong><strong>а</strong>х хелицер<strong>а</strong>льн<strong>о</strong>й <strong>в</strong><strong>о</strong>р<strong>о</strong>н<strong>к</strong>и;н<strong>а</strong><br />

<strong>к</strong>л<strong>а</strong>п<strong>а</strong>н<strong>а</strong>х <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия п<strong>о</strong> д<strong>в</strong>е щетин<strong>к</strong>и.<br />

Н<strong>а</strong> ли<strong>с</strong>тьях р<strong>а</strong><strong>с</strong>тений жи<strong>в</strong>ут <strong>к</strong><strong>о</strong>л<strong>о</strong>ниями, <strong>в</strong>ыделяя<br />

<strong>о</strong>бильную п<strong>а</strong>утину. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь):<br />

Metatranychus Oudemans, 1931 (ри<strong>с</strong>.<br />

1358) и Schizotetranychus Tragardt, 1915.<br />

НАДСЕМЕЙСТВО LIMNOCHARIDEA.<br />

ЛИМНОХАРИДЫ<br />

(Limnocharae)<br />

Тел<strong>о</strong> мяг<strong>к</strong><strong>о</strong>е, меш<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>е, <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>к</strong>р<strong>а</strong><strong>с</strong>н<strong>о</strong>г<strong>о</strong><br />

ц<strong>в</strong>ет<strong>а</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы мяг<strong>к</strong>ие, переп<strong>о</strong>нч<strong>а</strong>т<strong>о</strong>й<br />

<strong>к</strong><strong>о</strong>н<strong>с</strong>и<strong>с</strong>тенции. Б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые гл<strong>а</strong>з<strong>а</strong> <strong>в</strong> <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итых<br />

<strong>к</strong><strong>а</strong>п<strong>с</strong>ул<strong>а</strong>х, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м <strong>с</strong>рединн<strong>о</strong>г<strong>о</strong><br />

п<strong>а</strong>л<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>г<strong>о</strong> щит<strong>к</strong><strong>а</strong>. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

цилиндриче<strong>с</strong><strong>к</strong><strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>а</strong>я, <strong>с</strong> р<strong>о</strong>г<strong>о</strong><strong>в</strong>ым ди<strong>с</strong><strong>к</strong><strong>о</strong>м,<br />

<strong>о</strong><strong>к</strong><strong>а</strong>ймленным б<strong>а</strong>хр<strong>о</strong>м<strong>о</strong>й. Хелицеры <strong>к</strong>рупные<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м. Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные,<br />

<strong>с</strong> ше<strong>с</strong>тью члени<strong>к</strong><strong>а</strong>ми; т<strong>а</strong>зи<strong>к</strong>и их (эпимеры)<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены четырьмя групп<strong>а</strong>ми. Пл<strong>а</strong><strong>в</strong><strong>а</strong>тельные<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> члени<strong>к</strong><strong>а</strong>х н<strong>о</strong>г IV <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong>ых при<strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>к</strong> нет,. Жи<strong>в</strong>ут <strong>в</strong> пре<strong>с</strong>-<br />

Ри<strong>с</strong>. 1357—1361. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Trombidiidae, Tetranychidae,<br />

Eriophyidae<br />

1357. Allothrombium clavipes (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид,<br />

X40.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1358. Metatranychus gibbus (Koch et Berendt);<br />

<strong>о</strong>бщий <strong>в</strong>ид, X35.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

und Berendt, 1854). 1359. Phytoptus antiquus (Heyden); <strong>в</strong><strong>о</strong>йл<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

г<strong>а</strong>ллы н<strong>а</strong> ли<strong>с</strong>те Passiflora; Х1.0, п<strong>а</strong>ле<strong>о</strong>ген, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Heyden, 1862). 1360. Aceria ceanothi Keifer; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong>б<strong>о</strong><strong>к</strong>у, x600,0,<br />

<strong>с</strong><strong>о</strong><strong>в</strong>рем. (КеИег, 1939). 1361. Aceria diversicoloris Roivainen;<br />

А — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; Б — т<strong>о</strong> же <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

Х700.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Liro und Roivainen, 1951)<br />

469<br />

http://jurassic.ru/

Ри<strong>с</strong>. 1362—1365. Отряд Parasitiformes<br />

1S62. Seius vlduus K.och; <strong>о</strong>бщий <strong>в</strong>ид, x25,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> center pictus Hermann; A — <strong>с</strong><strong>а</strong>мец <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854), 1363. Hypoaspls Б — т<strong>о</strong> же <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х15.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (П<strong>о</strong>мер<strong>а</strong>нце<strong>в</strong>,<br />

aculelfer (Canestrini); А — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; Б — т<strong>о</strong> же 1950).1365. Ixodes rlclnus (Linnaeus); <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х65.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Л<strong>а</strong>нге, 1955). 1364. Derma- X20.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (П<strong>о</strong>мер<strong>а</strong>нце<strong>в</strong>, 1950)<br />

470<br />

http://jurassic.ru/

ных <strong>в</strong><strong>о</strong>д<strong>о</strong>ем<strong>а</strong>х. Длин<strong>а</strong> тел<strong>а</strong> д<strong>о</strong> 4—5 мм. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Limnocharidae,<br />

Eylaidae, Protziidae. У<strong>к</strong><strong>а</strong>з<strong>а</strong>ние н<strong>а</strong> н<strong>а</strong>х<strong>о</strong>ждение<br />

р<strong>о</strong>д<strong>а</strong> Limnochares Latreille, 1796 <strong>в</strong> третичных<br />

<strong>о</strong>тл<strong>о</strong>жениях Герм<strong>а</strong>нии (Heyden, 1862; Oudemans,<br />

1937) <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong>.<br />

ПОДОТРЯД TETRAPODILI.<br />

ЧЕТЫРЕХНОГИЕ ГАЛЛОВЫЕ<br />

КЛЕЩИ<br />

(Eriophyiformes)<br />

Очень мел<strong>к</strong>ие чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зные р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядные<br />

<strong>к</strong>лещи (длин<strong>а</strong> тел<strong>а</strong> 0,10—0,25 мм) <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

ут<strong>о</strong>лщенным передним <strong>к</strong><strong>о</strong>нц<strong>о</strong>м и<br />

длинным, <strong>с</strong>уж<strong>а</strong>ющим<strong>с</strong>я н<strong>а</strong>з<strong>а</strong>д <strong>к</strong><strong>о</strong>льч<strong>а</strong>тым тел<strong>о</strong>м.<br />

Н<strong>а</strong> <strong>в</strong><strong>с</strong>ех ф<strong>а</strong>з<strong>а</strong>х р<strong>а</strong>з<strong>в</strong>ития т<strong>о</strong>ль<strong>к</strong><strong>о</strong> д<strong>в</strong>е передние<br />

п<strong>а</strong>ры н<strong>о</strong>г <strong>с</strong> пятью <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими члени<strong>к</strong><strong>а</strong>ми.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> шил<strong>о</strong><strong>в</strong>идными и <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong>идными щетин<strong>к</strong><strong>а</strong>ми,<br />

длинным, из<strong>о</strong>гнутым п<strong>а</strong>л<strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong>идным<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м и пери<strong>с</strong>тым эмп<strong>о</strong>дием; н<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льных<br />

члени<strong>к</strong><strong>а</strong>х н<strong>о</strong>г единичные щетин<strong>к</strong><strong>о</strong><strong>в</strong>идные хеты<br />

(ри<strong>с</strong>. 1360, 1361 А, Б). Передний — р<strong>а</strong><strong>с</strong>ширенный<br />

— <strong>к</strong><strong>о</strong>нец тел<strong>а</strong> н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не имеет<br />

щит<strong>о</strong><strong>к</strong> <strong>с</strong> п<strong>а</strong>р<strong>о</strong>й щетин<strong>о</strong><strong>к</strong> н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е, ин<strong>о</strong>гд<strong>а</strong><br />

и <strong>с</strong><strong>о</strong> щетин<strong>к</strong><strong>а</strong>ми <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти щит<strong>к</strong><strong>а</strong>. О<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>е<br />

тел<strong>о</strong> у <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>бит<strong>а</strong>ющих <strong>в</strong> г<strong>а</strong>лл<strong>а</strong>х<br />

;(ри<strong>с</strong>. 1359), <strong>с</strong> нежными п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми и р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й<br />

т<strong>о</strong>н<strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>льце<strong>в</strong>идн<strong>о</strong>й и<strong>с</strong>черченн<strong>о</strong><strong>с</strong>тью; у<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>о</strong>бит<strong>а</strong>ющих н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти ли<strong>с</strong>тье<strong>в</strong>, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

б<strong>о</strong>лее <strong>с</strong><strong>к</strong>лер<strong>о</strong>тиз<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>, причем <strong>к</strong><strong>о</strong>льд<strong>а</strong><br />

н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не р<strong>а</strong><strong>с</strong>ширены. Н<strong>а</strong> з<strong>а</strong>днем<br />

<strong>к</strong><strong>о</strong>нце тел<strong>а</strong> имеют<strong>с</strong>я при<strong>с</strong><strong>а</strong><strong>с</strong>ы<strong>в</strong><strong>а</strong>тельные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

(ри<strong>с</strong>. 1361 А, Б). Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>ильн<strong>о</strong> <strong>в</strong>ид<strong>о</strong>изменен<strong>а</strong>;<br />

<strong>с</strong>реднюю ч<strong>а</strong><strong>с</strong>ть ее з<strong>а</strong>ним<strong>а</strong>ет <strong>к</strong>лю<strong>в</strong><strong>о</strong><strong>в</strong>идный<br />

хелицер<strong>а</strong>льный <strong>к</strong><strong>о</strong>ну<strong>с</strong>, п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены т<strong>о</strong>л<strong>с</strong>тые, у<strong>к</strong><strong>о</strong>р<strong>о</strong>ченные п<strong>а</strong>ль-<br />

ОТРЯД PARASITIFORMES.<br />

Тел<strong>о</strong> <strong>о</strong>бычн<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong>е <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

и брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н р<strong>а</strong>зличными щит<strong>к</strong><strong>а</strong>ми. Р<strong>а</strong><strong>с</strong>членения<br />

иди<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>бычн<strong>о</strong> нет, и т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у примити<strong>в</strong>ных<br />

<strong>к</strong>лещей <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Rhodacaridae н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я<br />

типичный для <strong>в</strong><strong>с</strong>ех хелицер<strong>о</strong><strong>в</strong>ых <strong>а</strong>р<strong>а</strong>хн<strong>о</strong>идный<br />

тип р<strong>а</strong><strong>с</strong>членения н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь или<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>му, не<strong>с</strong>ущую <strong>в</strong><strong>с</strong>е <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му;<br />

или брюш<strong>к</strong><strong>о</strong>; х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>а</strong>я для <strong>в</strong><strong>с</strong>ех <strong>к</strong>лещей<br />

гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> у Parasitiformes <strong>о</strong>чень чет<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong><br />

<strong>в</strong> <strong>в</strong>иде <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>г<strong>о</strong> р<strong>а</strong>здел<strong>а</strong> тел<strong>а</strong> (ри<strong>с</strong>. 1362,<br />

1363А). У гр<strong>о</strong>м<strong>а</strong>дн<strong>о</strong>г<strong>о</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> других г<strong>а</strong>м<strong>а</strong>з<strong>о</strong>идных<br />

<strong>к</strong>лещей г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь и брюш<strong>к</strong><strong>о</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью<br />

<strong>с</strong>литы, <strong>а</strong> гр<strong>а</strong>ницу между ними уд<strong>а</strong>ет<strong>с</strong>я<br />

п<strong>о</strong>дмет<strong>а</strong>ть лишь п<strong>о</strong> <strong>с</strong>тр<strong>о</strong>ению элемент<strong>о</strong><strong>в</strong> <strong>с</strong><strong>к</strong>елет<strong>а</strong><br />

и п<strong>о</strong>. р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жению щетин<strong>о</strong><strong>к</strong>. С<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упн<strong>о</strong><strong>с</strong>ть<br />

пы <strong>с</strong> при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й л<strong>а</strong>п<strong>к</strong><strong>о</strong>й. Хелицеры т<strong>о</strong>н<strong>к</strong>ие,<br />

<strong>с</strong>тилет<strong>о</strong><strong>в</strong>идные; дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong><br />

брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> п<strong>о</strong>з<strong>а</strong>ди <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льных п<strong>о</strong>лей<br />

н<strong>о</strong>г II. Ан<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я н<strong>а</strong><br />

з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong>.<br />

Четырехн<strong>о</strong>гие <strong>к</strong>лещи п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Tetrapodili<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны другим р<strong>а</strong><strong>с</strong>тительн<strong>о</strong>ядным <strong>к</strong>лещ<strong>а</strong>м,<br />

<strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти <strong>к</strong>лещ<strong>а</strong>м н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Trichadenidea.<br />

Четырехн<strong>о</strong>гие <strong>к</strong>лещи п<strong>а</strong>р<strong>а</strong>зитируют н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях,<br />

пит<strong>а</strong>я<strong>с</strong>ь их <strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>ми; ч<strong>а</strong><strong>с</strong>ть <strong>в</strong>ид<strong>о</strong><strong>в</strong> (Eriophyidae)<br />

<strong>в</strong>ызы<strong>в</strong><strong>а</strong>ет <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ние г<strong>а</strong>лл<strong>о</strong><strong>в</strong> (ри<strong>с</strong>.<br />

1359), т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> другие <strong>о</strong>бит<strong>а</strong>ют т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

ли<strong>с</strong>тье<strong>в</strong> (Phyllocoptidae). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Eriophyidae и Phyllocoptidae.<br />

СЕМЕЙСТВО ERIOPHYIDAE NALEPA, 1898.<br />

ЧЕТЫРЕХНОГИЕ ГАЛЛООБРАЗУЮЩИЕ КЛЕЩИ<br />

Спинн<strong>о</strong>й щит ни<strong>к</strong><strong>о</strong>гд<strong>а</strong> не имеет <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>,<br />

н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющег<strong>о</strong> н<strong>а</strong>д р<strong>о</strong><strong>с</strong>трум<strong>о</strong>м. Н<strong>а</strong> ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>ме<br />

<strong>в</strong><strong>о</strong> <strong>в</strong><strong>с</strong>ех ф<strong>а</strong>з<strong>а</strong>х р<strong>а</strong>з<strong>в</strong>ития е<strong>с</strong>ть г<strong>о</strong>м<strong>о</strong>н<strong>о</strong>мн<strong>а</strong>я нежн<strong>а</strong>я<br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>т<strong>о</strong><strong>с</strong>ть; ширин<strong>а</strong> <strong>о</strong>тдельных <strong>к</strong><strong>о</strong>лец н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й и брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>х тел<strong>а</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я<br />

.(ри<strong>с</strong>. 1361). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них д<strong>в</strong><strong>а</strong> — Phytoptus Dujardin,<br />

1851 и Eriophyes Siebold, 1850 — <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО PHYLLOCOPTIDAE NALEPA, 1898<br />

Спинн<strong>о</strong>й щит <strong>о</strong>бычн<strong>о</strong> имеет передний <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т,<br />

н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющий н<strong>а</strong>д р<strong>о</strong><strong>с</strong>трум<strong>о</strong>м. Хитин<strong>о</strong><strong>в</strong>ые<br />

<strong>к</strong><strong>о</strong>льц<strong>а</strong> н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong><br />

шире, чем н<strong>а</strong> брюшн<strong>о</strong>й, и б<strong>о</strong>лее упл<strong>о</strong>тненные.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. О<strong>к</strong><strong>о</strong>л<strong>о</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне и <strong>о</strong>дн<strong>а</strong> (м<strong>а</strong>л<strong>о</strong>изученн<strong>а</strong>я) ф<strong>о</strong>рм<strong>а</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong>, <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<strong>а</strong> Геер<strong>о</strong>м (Thomas, 1877) п<strong>о</strong><br />

г<strong>а</strong>лл<strong>а</strong>м н<strong>а</strong> ли<strong>с</strong>тьях т<strong>о</strong>п<strong>о</strong>ля.<br />

471<br />

ГАМАЗОИДНЫЕ КЛЕЩИ<br />

имеющих<strong>с</strong>я <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>-<strong>а</strong>н<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong>их и эмбри<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>их<br />

д<strong>а</strong>нных <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует <strong>о</strong> т<strong>о</strong>м,<br />

чт<strong>о</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> Parasitiformes <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>в</strong>яти<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти. Передний<br />

<strong>к</strong>р<strong>а</strong>й <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>ытянут <strong>в</strong> эпи<strong>с</strong>т<strong>о</strong>м,<br />

п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> у Mesostigmata р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

же<strong>в</strong><strong>а</strong>тельные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти гн<strong>а</strong>т<strong>о</strong><strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>, <strong>о</strong>бычн<strong>о</strong><br />

<strong>с</strong>мы<strong>к</strong><strong>а</strong>ющие<strong>с</strong>я п<strong>о</strong> <strong>с</strong>редней линии. Уз<strong>к</strong><strong>о</strong>е щеле<strong>в</strong>идн<strong>о</strong>е<br />

р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong><strong>в</strong>ерху при<strong>к</strong>рыт<strong>о</strong><br />

<strong>в</strong>ерхней губ<strong>о</strong>й, <strong>а</strong> <strong>с</strong>низу <strong>в</strong>нутренним <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м<br />

гн<strong>а</strong>т<strong>о</strong>б<strong>а</strong>зы — гип<strong>о</strong><strong>с</strong>т<strong>о</strong>м<strong>о</strong>м. Д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong> <strong>о</strong>т р<strong>с</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>й<br />

щели р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>д<strong>в</strong>ижные <strong>к</strong>рупные<br />

хелицеры, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

К<strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>ни <strong>к</strong>лешне<strong>в</strong>идные,<br />

причем у хищных г<strong>а</strong>м<strong>а</strong>зид (Rhodacaridae. Р<strong>а</strong>-<br />

http://jurassic.ru/

asitidae) <strong>о</strong>ни <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельными, <strong>с</strong><br />

длинными и груб<strong>о</strong>зубч<strong>а</strong>тыми п<strong>а</strong>льц<strong>а</strong>ми; у <strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong>ущих<br />

Laelaptidae, Haemogamasidae, Dermanyssidae<br />

и др., н<strong>а</strong><strong>о</strong>б<strong>о</strong>р<strong>о</strong>т, хелицеры <strong>в</strong>ытяги<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>в</strong>иде гиб<strong>к</strong>их <strong>с</strong>тилет<strong>о</strong><strong>в</strong>, <strong>с</strong>луж<strong>а</strong>щих для<br />

пр<strong>о</strong><strong>к</strong><strong>а</strong>лы<strong>в</strong><strong>а</strong>ния п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> и т<strong>к</strong><strong>а</strong>ней жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>г<strong>о</strong>х<strong>о</strong>зяин<strong>а</strong>;<br />

н<strong>а</strong><strong>к</strong><strong>о</strong>нец, у Ixodides хелицеры <strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я<br />

я<strong>к</strong><strong>о</strong>ре<strong>в</strong>идными, <strong>в</strong>ып<strong>о</strong>лняющими фун<strong>к</strong>цию<br />

пр<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> з<strong>а</strong><strong>к</strong>репления <strong>в</strong> т<strong>о</strong>лще <strong>к</strong><strong>о</strong>жи х<strong>о</strong>зяин<strong>а</strong>.<br />

П<strong>о</strong>з<strong>а</strong>ди <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны гн<strong>а</strong>т<strong>о</strong>б<strong>а</strong>зы<br />

у Mesostigmata п<strong>о</strong>мещ<strong>а</strong>ет<strong>с</strong>я <strong>в</strong>ильч<strong>а</strong>тый трит<strong>о</strong><strong>с</strong>тернум.<br />

Оф<strong>о</strong>рмленный <strong>с</strong><strong>к</strong>елет п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> иди<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

имеет<strong>с</strong>я н<strong>а</strong> <strong>в</strong><strong>с</strong>ех ф<strong>а</strong>з<strong>а</strong>х р<strong>а</strong>з<strong>в</strong>ития и <strong>о</strong>тлич<strong>а</strong>ет<strong>с</strong>я<br />

б<strong>о</strong>льш<strong>о</strong>й <strong>с</strong>л<strong>о</strong>жн<strong>о</strong><strong>с</strong>тью. Обычн<strong>о</strong> у <strong>в</strong><strong>с</strong>ех пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

Parasitiformes е<strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>й щит<br />

(<strong>с</strong><strong>а</strong>г<strong>а</strong>р<strong>а</strong>х) (ри<strong>с</strong>. 1364, 1365), грудн<strong>о</strong>й щит, или<br />

грудин<strong>а</strong> (ри<strong>с</strong>. 1363А), <strong>а</strong>н<strong>а</strong>льный щит и у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

еще генит<strong>а</strong>льный щит, или эпигин<strong>а</strong> (ри<strong>с</strong>.<br />

1313А). Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, у мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong> з<strong>а</strong>дней<br />

ч<strong>а</strong><strong>с</strong>ти <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong> р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я н<strong>о</strong>т<strong>о</strong>г<strong>а</strong><strong>с</strong>тр<strong>а</strong>льный,<br />

или пигиди<strong>а</strong>льный, щит; ин<strong>о</strong>гд<strong>а</strong><br />

эт<strong>о</strong>т щит <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный и <strong>о</strong>бр<strong>а</strong>зует<br />

единый <strong>с</strong>пинн<strong>о</strong>й щит, з<strong>а</strong>ним<strong>а</strong>ющий п<strong>о</strong>чти<br />

<strong>в</strong><strong>с</strong>ю <strong>с</strong>пинную п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тел<strong>а</strong> (ри<strong>с</strong>. 1363А).<br />

брюшн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти тел<strong>а</strong>, <strong>к</strong>р<strong>о</strong>ме х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> щит<strong>к</strong><strong>а</strong>, м<strong>о</strong>гут быть р<strong>а</strong>з<strong>в</strong>иты переднегрудные<br />

(югулярные) щит<strong>к</strong>и, леж<strong>а</strong>щие п<strong>о</strong><br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>о</strong>т трит<strong>о</strong><strong>с</strong>тернум<strong>а</strong> у г<strong>а</strong>м<strong>а</strong>зид и п<strong>о</strong>з<strong>а</strong>ди<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия — л<strong>а</strong>тер<strong>а</strong>льные, эпигин<strong>а</strong>льные,<br />

мет<strong>а</strong>п<strong>о</strong>ди<strong>а</strong>льные, <strong>в</strong>ентр<strong>а</strong>льный и <strong>а</strong>н<strong>а</strong>льный<br />

щит<strong>к</strong>и. Стр<strong>о</strong>ение брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

и<strong>с</strong>п<strong>о</strong>льзует<strong>с</strong>я для целей <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>и.<br />

В<strong>с</strong>е щетин<strong>к</strong>и тел<strong>а</strong> и <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти лишены <strong>а</strong><strong>к</strong>тин<strong>о</strong>хитин<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тержня, <strong>о</strong>птиче<strong>с</strong><strong>к</strong>и из<strong>о</strong>тр<strong>о</strong>пны.<br />

Н<strong>а</strong> м<strong>о</strong>л<strong>о</strong>дых ф<strong>а</strong>з<strong>а</strong>х р<strong>а</strong>з<strong>в</strong>ития щетин<strong>к</strong>и немн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны<br />

и пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены, <strong>а</strong> з<strong>а</strong>тем<br />

р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> н<strong>о</strong><strong>в</strong>ых, д<strong>о</strong>п<strong>о</strong>лнительных<br />

щетин<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 1363Б, 1365).<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терны для <strong>в</strong><strong>с</strong>ех Parasitiformes н<strong>а</strong>личие<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дных цилиндриче<strong>с</strong><strong>к</strong>их т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> и р<strong>а</strong>зделение<br />

телеп<strong>о</strong>дит<strong>а</strong> н<strong>о</strong>ги н<strong>а</strong> ше<strong>с</strong>ть члени<strong>к</strong><strong>о</strong><strong>в</strong>:<br />

<strong>в</strong>ертлуг, нер<strong>а</strong><strong>с</strong>члененн<strong>о</strong>е бедр<strong>о</strong>, п<strong>а</strong>теллю, г<strong>о</strong>лень<br />

и л<strong>а</strong>п<strong>к</strong>у. Н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х л<strong>а</strong>п<strong>о</strong><strong>к</strong> имеют<strong>с</strong>я переп<strong>о</strong>нч<strong>а</strong>тый<br />

прет<strong>а</strong>рз и д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>; эмя<strong>о</strong>дий бы<strong>в</strong><strong>а</strong>ет<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идным и <strong>о</strong>бычн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из м<strong>а</strong>лень<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong><strong>к</strong>лерит<strong>а</strong> и шир<strong>о</strong><strong>к</strong><strong>о</strong>й трехл<strong>о</strong>п<strong>а</strong><strong>с</strong>тн<strong>о</strong>й<br />

при<strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>и (pulvillus) (ри<strong>с</strong>. 1363Б). Очень ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

н<strong>о</strong>ги пер<strong>в</strong><strong>о</strong>й п<strong>а</strong>ры не уч<strong>а</strong><strong>с</strong>т<strong>в</strong>уют <strong>в</strong> х<strong>о</strong>ждении, <strong>а</strong><br />

<strong>с</strong>луж<strong>а</strong>т <strong>о</strong>рг<strong>а</strong>н<strong>а</strong>ми <strong>о</strong><strong>с</strong>яз<strong>а</strong>ния. У Ixodides т<strong>а</strong>зи<strong>к</strong>и<br />

н<strong>о</strong>г <strong>в</strong>т<strong>о</strong>ричн<strong>о</strong> <strong>с</strong>р<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я <strong>с</strong> п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми тел<strong>а</strong> и<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>бширные <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>льные щиты<br />

(ри<strong>с</strong>. 1364Б).<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> между <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>а</strong>ми<br />

н<strong>о</strong>г. Дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong><br />

тр<strong>а</strong>хеями, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я п<strong>а</strong>р<strong>о</strong>й <strong>с</strong>тигм н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х<br />

тел<strong>а</strong> <strong>в</strong>переди (<strong>к</strong><strong>а</strong><strong>к</strong> у Mesostigmata) или п<strong>о</strong>з<strong>а</strong>ди<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний н<strong>о</strong>г IV (у Ixodides).<br />

472<br />

http://jurassic.ru/<br />

П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие у г<strong>а</strong>м<strong>а</strong>з<strong>о</strong><strong>в</strong>ых<br />

<strong>к</strong>лещей (Mesostigmata) пр<strong>о</strong>те<strong>к</strong><strong>а</strong>ет п<strong>о</strong> типу бинимф<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>а</strong>, <strong>а</strong> у Ixodides <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дн<strong>а</strong> нимф<strong>а</strong>льн<strong>а</strong>я ф<strong>а</strong>з<strong>а</strong>. Пре<strong>в</strong>р<strong>а</strong>щение<br />

<strong>в</strong><strong>о</strong> <strong>в</strong>зр<strong>о</strong><strong>с</strong>лую ф<strong>а</strong>зу, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, не <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong><br />

<strong>с</strong> <strong>к</strong><strong>а</strong><strong>к</strong>ими-либ<strong>о</strong> рез<strong>к</strong>ими пре<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ниями,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> эт<strong>о</strong> н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я у Acariformes. Опл<strong>о</strong>д<strong>о</strong>т<strong>в</strong><strong>о</strong>рение<br />

<strong>в</strong><strong>с</strong>ех <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong>перм<strong>а</strong>т<strong>о</strong>ф<strong>о</strong>рн<strong>о</strong>е. Жизненные<br />

ци<strong>к</strong>лы, <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong> у п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>их <strong>в</strong>ид<strong>о</strong><strong>в</strong>,<br />

м<strong>о</strong>гут быть <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>с</strong>л<strong>о</strong>жными.<br />

Среди Parasitiformes имеют<strong>с</strong>я <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие<br />

<strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги и хищни<strong>к</strong>и и б<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

п<strong>а</strong>р<strong>а</strong>зит<strong>о</strong><strong>в</strong>; п<strong>о</strong>д<strong>о</strong>тряд Ixodides пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ими, <strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong>ущими<br />

<strong>в</strong>ид<strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Mesostigmata, Ixodides,<br />

Holothyroidea, из них д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

ПОДОТРЯД MESOSTIGMATA.<br />

ГАМАЗОИДНЫЕ КЛЕЩИ<br />

Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>й гн<strong>а</strong>т<strong>о</strong>б<strong>а</strong>з<strong>о</strong>й,<br />

<strong>в</strong>нутри <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>к</strong>рупные хелицеры.<br />

Педип<strong>а</strong>льпы длинные, н<strong>о</strong>г<strong>о</strong><strong>о</strong>бр<strong>а</strong>зные. Н<strong>а</strong><br />

брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не н<strong>о</strong>г I имеет<strong>с</strong>я<br />

<strong>в</strong>ильч<strong>а</strong>тый трит<strong>о</strong><strong>с</strong>тернум, <strong>а</strong> <strong>в</strong>д<strong>о</strong>ль <strong>с</strong>редней линии<br />

тел<strong>а</strong> <strong>о</strong>дин или не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>с</strong>терн<strong>а</strong>льных щит<strong>о</strong><strong>в</strong>.<br />

Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г цилиндриче<strong>с</strong><strong>к</strong>ие, п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>-<strong>с</strong><strong>о</strong>члененные<br />

<strong>с</strong> тел<strong>о</strong>м. Нпг-.и длинные, бег<strong>а</strong>тельн<strong>о</strong>г<strong>о</strong>тип<strong>а</strong><br />

(ри<strong>с</strong>. 1362, 1363А). Дых<strong>а</strong>тельные <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м тел<strong>а</strong> н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний н<strong>о</strong>г III; <strong>о</strong><strong>к</strong>руж<strong>а</strong>ющие их перитремы<br />

<strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>чень длинный лент<strong>о</strong><strong>в</strong>идный<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т, пр<strong>о</strong><strong>с</strong>тир<strong>а</strong>ющий<strong>с</strong>я <strong>в</strong>д<strong>о</strong>ль б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> д<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний н<strong>о</strong>г II или I (ри<strong>с</strong>. 1363А).Н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> — пл<strong>о</strong>тный г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>й<br />

щит или д<strong>в</strong><strong>а</strong> щит<strong>а</strong>, ин<strong>о</strong>гд<strong>а</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я<br />

друг <strong>с</strong> друг<strong>о</strong>м и т<strong>о</strong>гд<strong>а</strong> п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих п<strong>о</strong>чти <strong>в</strong><strong>с</strong>ю<br />

<strong>с</strong>пинную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну тел<strong>а</strong>. С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие и п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ие<br />

<strong>в</strong>иды <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ения и би<strong>о</strong>л<strong>о</strong>гии. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 11 н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Megisthanidea,<br />

Liroaspididea, Microgyniidea, Camasidea.<br />

Thinozerconidea, Diarthrophalidea, Zerconidea,<br />

Trachytidea, Uropodidea, Celaenopseidea, Fedrizziidea,<br />

из них <strong>о</strong>дн<strong>о</strong> •— Gamasidea — из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong><br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

НАДСЕМЕЙСТВО GAMASIDEA<br />

(Gamasides)<br />

Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> и <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong><br />

эпигин<strong>а</strong>льный щит<strong>о</strong><strong>к</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит; б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

щит<strong>к</strong>и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong><strong>а</strong>мц<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> <strong>в</strong>переди <strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> щит<strong>к</strong><strong>а</strong>. С<strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>о</strong>жи<strong>в</strong>ущие<br />

или э<strong>к</strong>т<strong>о</strong>п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ие <strong>к</strong>лещи;

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>иды (Rhinonyssidae, Halarachnidae,<br />

Entonyssidae) <strong>в</strong>едут энд<strong>о</strong>п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ий <strong>о</strong>бр<strong>а</strong>з<br />

жизни <strong>в</strong> <strong>в</strong><strong>о</strong>здушных путях тел<strong>а</strong> мле<strong>к</strong><strong>о</strong>пит<strong>а</strong>ющих<br />

и птиц. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 20 <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии Phytoseiidae и Laelaptidae.<br />

СЕМЕЙСТВО PHYTOSEIIDAE BERLESE, 1916<br />

Эпигин<strong>а</strong>льный щит<strong>о</strong><strong>к</strong> у<strong>с</strong>ечен или <strong>в</strong><strong>о</strong>гнут <strong>с</strong>з<strong>а</strong>ди.<br />

Н<strong>а</strong> теле имеет<strong>с</strong>я неб<strong>о</strong>льш<strong>о</strong>е <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> щетин<strong>о</strong><strong>к</strong>.<br />

Те<strong>к</strong>тум <strong>с</strong> зубчи<strong>к</strong><strong>а</strong>ми п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям. Трит<strong>о</strong><strong>с</strong>тернум<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит, у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>о</strong>н <strong>с</strong> ц<strong>а</strong>циниями.<br />

В<strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> р<strong>а</strong><strong>с</strong>тениях, где <strong>о</strong>ни <strong>о</strong>х<strong>о</strong>тят<strong>с</strong>я<br />

з<strong>а</strong> другими <strong>к</strong>лещ<strong>а</strong>ми и мел<strong>к</strong>ими н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мыми;<br />

<strong>о</strong>чень <strong>а</strong><strong>к</strong>ти<strong>в</strong>ные хищни<strong>к</strong>и. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 20 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

Sejus С. L. Koch, 1836, (ри<strong>с</strong>. 1362) <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО LAELAPTIDAE BERLESE, 1892<br />

Эпигин<strong>а</strong>льный щит<strong>о</strong><strong>к</strong> имеет <strong>к</strong><strong>а</strong>пле<strong>в</strong>идную<br />

ф<strong>о</strong>рму или <strong>в</strong>ырез<strong>а</strong>н п<strong>о</strong> з<strong>а</strong>днему <strong>к</strong>р<strong>а</strong>ю. Брюшные<br />

щиты у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>бычн<strong>о</strong> <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>о</strong>дин<br />

<strong>о</strong>бщий щит; <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дным, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong><strong>с</strong>т<strong>а</strong>ет<strong>с</strong>я<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>а</strong>н<strong>а</strong>льный щит<strong>о</strong><strong>к</strong>. Щетин<strong>о</strong><strong>к</strong> н<strong>а</strong> <strong>о</strong>пине<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> немн<strong>о</strong>г<strong>о</strong> — <strong>о</strong>бычн<strong>о</strong> <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 п<strong>а</strong>р.<br />

Н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х II у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют, <strong>а</strong><br />

н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г II имеет<strong>с</strong>я шил<strong>о</strong><strong>в</strong>идный<br />

<strong>в</strong>ы<strong>с</strong>туп. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> их — п<strong>а</strong>р<strong>а</strong>зитиче<strong>с</strong><strong>к</strong>ие<br />

ф<strong>о</strong>рмы. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тен Hypoaspis<br />

G. Canestrini, 1885 (ри<strong>с</strong>. 1363).<br />

ПОДОТРЯД. IXODES.<br />

ИКСОДОВЫЕ КЛЕЩИ<br />

Н<strong>а</strong>иб<strong>о</strong>лее <strong>к</strong>рупные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

Acaromorpha:- п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью н<strong>а</strong>пит<strong>а</strong><strong>в</strong>шие<strong>с</strong>я <strong>к</strong>р<strong>о</strong><strong>в</strong>ью<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>о</strong>д<strong>а</strong> Hyalomma д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют<br />

2 <strong>с</strong>м <strong>в</strong> длину.<br />

Тел<strong>о</strong> и<strong>к</strong><strong>с</strong><strong>о</strong>д<strong>о</strong><strong>в</strong>ых <strong>к</strong>лещей <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

упл<strong>о</strong>щенн<strong>о</strong>е — у г<strong>о</strong>л<strong>о</strong>дных и меш<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>е —<br />

у н<strong>а</strong>пит<strong>а</strong><strong>в</strong>ших<strong>с</strong>я <strong>к</strong>р<strong>о</strong><strong>в</strong>ью <strong>о</strong><strong>с</strong><strong>о</strong>бей. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> имеет<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>й<br />

щит (Ixodidae), или же щиты <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют (Argasidae).<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы <strong>к</strong><strong>о</strong>жи<strong>с</strong>тые. Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тр<strong>о</strong>ения. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние ее <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из тр<strong>а</strong>пецие<strong>в</strong>идн<strong>о</strong>й<br />

хитин<strong>о</strong><strong>в</strong><strong>о</strong>й <strong>к</strong><strong>а</strong>п<strong>с</strong>улы <strong>с</strong> <strong>в</strong>ытянутым<br />

язы<strong>к</strong><strong>о</strong><strong>в</strong>идным эпи<strong>с</strong>т<strong>о</strong>м<strong>о</strong>м. Педип<strong>а</strong>льпы т<strong>о</strong>л<strong>с</strong>тые<br />

и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Хелицеры т<strong>о</strong>н<strong>к</strong>ие,<br />

<strong>к</strong><strong>о</strong>лющие у Argasidae, я<strong>к</strong><strong>о</strong>ре<strong>в</strong>идные у Ixodidae.<br />

Трит<strong>о</strong><strong>с</strong>тернум <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><br />

<strong>с</strong>р<strong>а</strong>щены <strong>с</strong> тел<strong>о</strong>м, имеют <strong>в</strong>ид упл<strong>о</strong>тненных<br />

щит<strong>о</strong><strong>в</strong>. Дых<strong>а</strong>тельные <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их и шир<strong>о</strong><strong>к</strong>их <strong>о</strong><strong>в</strong><strong>а</strong>льных перитрем<strong>а</strong>х,<br />

не имеющих длинн<strong>о</strong>г<strong>о</strong> уз<strong>к</strong><strong>о</strong>г<strong>о</strong> переднег<strong>о</strong><br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>; <strong>о</strong>ни р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не или п<strong>о</strong>з<strong>а</strong>ди<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний н<strong>о</strong>г IV. Н<strong>о</strong>ги длинные, х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>. Ан<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие близ з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я тел<strong>а</strong>; <strong>к</strong>л<strong>а</strong>п<strong>а</strong>ны ег<strong>о</strong> <strong>с</strong><strong>о</strong> щетин<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>.<br />

1364Б). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Argasidae и Ixodidae. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны т<strong>о</strong>ль<strong>к</strong><strong>о</strong> и<strong>к</strong><strong>с</strong><strong>о</strong>д<strong>о</strong><strong>в</strong>ые<br />

<strong>к</strong>лещи.<br />

СЕМЕЙСТВО IXODIDAE MURRAY, 1877<br />

Гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ет рез<strong>к</strong><strong>о</strong> <strong>в</strong>перед <strong>в</strong> <strong>в</strong>иде<br />

«г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>к</strong>и, х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>а</strong> или <strong>к</strong><strong>а</strong>питулюм<strong>а</strong>» и не <strong>с</strong><strong>к</strong>рыт<strong>а</strong><br />

н<strong>а</strong><strong>в</strong>и<strong>с</strong><strong>а</strong>ющим <strong>к</strong>р<strong>а</strong>ем пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>й<br />

щит<strong>о</strong><strong>к</strong> у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я тел<strong>а</strong> (ри<strong>с</strong>. 1364 А), <strong>а</strong> у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>о</strong>н, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

бы<strong>в</strong><strong>а</strong>ет <strong>о</strong>гр<strong>а</strong>ничен лишь з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем<br />

пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы (ри<strong>с</strong>. 1365). Жизненный ци<strong>к</strong>л<br />

Ixodidae <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> <strong>с</strong>л<strong>о</strong>жен и <strong>о</strong>бычн<strong>о</strong> пр<strong>о</strong>те<strong>к</strong><strong>а</strong>ет<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>мен<strong>о</strong>й х<strong>о</strong>зяе<strong>в</strong>, пери<strong>о</strong>диче<strong>с</strong><strong>к</strong>им н<strong>а</strong><strong>с</strong><strong>а</strong><strong>с</strong>ы<strong>в</strong><strong>а</strong>нием<br />

<strong>о</strong>гр<strong>о</strong>мных п<strong>о</strong>рций <strong>к</strong>р<strong>о</strong><strong>в</strong>и и т. п. Мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<br />

<strong>с</strong>ильн<strong>о</strong> упр<strong>о</strong>щен; <strong>с</strong><strong>о</strong>хр<strong>а</strong>няет<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дн<strong>а</strong> нимф<strong>а</strong>льн<strong>а</strong>я<br />

ф<strong>а</strong>з<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне 16 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них Ixodes Latreille,<br />

1795 и Dermacentor С. L. Koch, 1844, из<strong>в</strong>е<strong>с</strong>тны<br />

из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong> С. Амери<strong>к</strong>и.<br />

ОТРЯД OPILIOACARINA. КЛЕЩИ-СЕНОКОСЦЫ<br />

Клещи <strong>с</strong>редней <strong>в</strong>еличины — длин<strong>о</strong>й <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><br />

1 мм. Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми <strong>с</strong>егмент<strong>а</strong>ции.<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> нежные, и<strong>с</strong>черчены б<strong>о</strong>р<strong>о</strong>зд<strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нными мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными п<strong>о</strong>р<strong>а</strong>ми.<br />

Имеет<strong>с</strong>я р<strong>а</strong><strong>с</strong>члененный трит<strong>о</strong><strong>с</strong>тернум. П<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м<br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong> д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>а</strong>.<br />

Хелицеры <strong>к</strong>лешне<strong>в</strong>идные. Н<strong>о</strong>ги <strong>о</strong>чень длинные,<br />

т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong> длинными цилиндриче<strong>с</strong><strong>к</strong>ими члени<strong>к</strong><strong>а</strong>ми.<br />

Т<strong>а</strong>зи<strong>к</strong>и п<strong>о</strong>д<strong>в</strong>ижны. Вертлуги н<strong>о</strong>г III и IV<br />

р<strong>а</strong><strong>с</strong>членены. Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х н<strong>о</strong>г п<strong>о</strong> д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>. Н<strong>а</strong><br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>в</strong> передней ч<strong>а</strong><strong>с</strong>ти ги<strong>с</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

(Notostigmata; Onychopalpida, pars)<br />

473<br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я четыре п<strong>а</strong>ры дых<strong>а</strong>тельных <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> между<br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г III. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>в</strong>ыд<strong>в</strong>ижн<strong>о</strong>й яйце<strong>к</strong>л<strong>а</strong>д.<br />

Клещи-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цы пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong><br />

примити<strong>в</strong>ных <strong>к</strong>лещей. Ведут <strong>с</strong><strong>к</strong>рытный <strong>о</strong>бр<strong>а</strong>з<br />

жизни. Обит<strong>а</strong>ют <strong>в</strong> п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е п<strong>о</strong>ч<strong>в</strong>ы, п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями.<br />

Вер<strong>о</strong>ятн<strong>о</strong>, хищни<strong>к</strong>и. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

<strong>о</strong>дн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Opilioacaridae <strong>с</strong> тремя р<strong>о</strong>д<strong>а</strong>ми;<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тны.<br />

http://jurassic.ru/

КЛАСС ARACHNIDA. ПАУКООБРАЗНЫЕ<br />

Н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые. Тел<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> н<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>му и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му. О<strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тью пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Arachnida, х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризующей<br />

т<strong>а</strong><strong>к</strong>же мн<strong>о</strong>гих <strong>к</strong>лещей <strong>о</strong>тряд<strong>а</strong> Acariformes (Oribatei,<br />

Sarcoptiformes) и де<strong>с</strong>ятин<strong>о</strong>гих р<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>,<br />

н<strong>а</strong>д<strong>о</strong> <strong>с</strong>чит<strong>а</strong>ть изгиб<strong>а</strong>ние пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>,<br />

при <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м IX и X <strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы перемещ<strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну. Н<strong>а</strong>пример,<br />

у <strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>це<strong>в</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Cyphophthalmi X тергит<br />

при эт<strong>о</strong>м <strong>о</strong><strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>в</strong>переди де<strong>в</strong>ят<strong>о</strong>г<strong>о</strong><br />

(<strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>с</strong>д<strong>в</strong>иг<strong>а</strong>ет<strong>с</strong>я <strong>к</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>му <strong>к</strong><strong>о</strong>нцу<br />

и <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие), <strong>а</strong> <strong>с</strong>терниты VI—IX <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> изгиб<strong>а</strong>ют<strong>с</strong>я,<br />

<strong>о</strong><strong>к</strong>руж<strong>а</strong>я переме<strong>с</strong>ти<strong>в</strong>шие<strong>с</strong>я н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну<br />

тергиты. У<strong>к</strong><strong>о</strong>р<strong>о</strong>чение тел<strong>а</strong> путем з<strong>а</strong>гиб<strong>а</strong>ния<br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> ег<strong>о</strong> н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну при<strong>в</strong><strong>о</strong>дит<br />

у Arachnida <strong>к</strong> зн<strong>а</strong>чительн<strong>о</strong>й реду<strong>к</strong>ции<br />

<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны з<strong>а</strong>гнутых <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, чт<strong>о</strong><br />

х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует <strong>в</strong>ы<strong>с</strong>шую <strong>с</strong>тепень интегр<strong>а</strong>ции<br />

тул<strong>о</strong><strong>в</strong>ищ<strong>а</strong> <strong>с</strong>реди хелицер<strong>о</strong><strong>в</strong>ых. Общее <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong> Arachnida <strong>в</strong><strong>а</strong>рьирует <strong>о</strong>т<br />

16 (Opiliones) д<strong>о</strong> 18 (Araneae). У <strong>в</strong>ы<strong>с</strong>ших<br />

Araneae <strong>в</strong> результ<strong>а</strong>те изгиб<strong>а</strong>ния з<strong>а</strong>днег<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong><br />

тел<strong>а</strong> н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дит ч<strong>а</strong><strong>с</strong>тичн<strong>а</strong>я<br />

реду<strong>к</strong>ция ше<strong>с</strong>ти з<strong>а</strong>дних <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; мет<strong>а</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

при эт<strong>о</strong>м м<strong>о</strong>жет п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью и<strong>с</strong>чез<strong>а</strong>ть.<br />

Следы п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>й реду<strong>к</strong>ции яр<strong>к</strong><strong>о</strong> пр<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong><br />

н<strong>а</strong>личии у эмбри<strong>о</strong>н<strong>о</strong><strong>в</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> (р<strong>о</strong>д<br />

Pholcus Walcken) д<strong>в</strong>ух<strong>с</strong>егментн<strong>о</strong>г<strong>о</strong> п<strong>о</strong><strong>с</strong>т<strong>а</strong>бд<strong>о</strong>мен<strong>а</strong><br />

<strong>с</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью (Holm, 1940). Суще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>о</strong>ф<strong>о</strong>рмленн<strong>о</strong>г<strong>о</strong><br />

з<strong>а</strong>днебрюшья <strong>в</strong> эмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><strong>с</strong>ть предп<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ем<strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Arachnida <strong>с</strong> Amblypygi и другими примити<strong>в</strong>ными<br />

Scorpionomorpha.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> <strong>о</strong>бщим<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит<strong>о</strong>м, н<strong>о</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Opiliones и у не<strong>к</strong><strong>о</strong>т<strong>о</strong>­<br />

(] В. Б. Дубинин |)<br />

474<br />

http://jurassic.ru/<br />

рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> Trigonotarbi и Anthracomarti н<strong>а</strong><br />

щите имеют<strong>с</strong>я п<strong>о</strong>перечные ш<strong>в</strong>ы, <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>ющие<br />

гр<strong>а</strong>ницы пер<strong>в</strong><strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>членения н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

щит (пр<strong>о</strong>пельтидий), мез<strong>о</strong>- и мет<strong>а</strong>пельтидий<br />

(ри<strong>с</strong>. 1393, 1401).<br />

Ин<strong>о</strong>гд<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий щит <strong>с</strong>ли<strong>в</strong><strong>а</strong>ет<strong>с</strong>я <strong>с</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими тергит<strong>а</strong>ми <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, н<strong>о</strong> гр<strong>а</strong>ницы<br />

<strong>с</strong>ли<strong>в</strong>ших<strong>с</strong>я <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>в</strong> <strong>в</strong>иде ш<strong>в</strong><strong>о</strong><strong>в</strong> бы<strong>в</strong><strong>а</strong>ют<br />

х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> з<strong>а</strong>метны. Тергиты мез<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельными или п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью редуцируют<strong>с</strong>я<br />

(у <strong>в</strong><strong>с</strong>ех п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>к</strong>р<strong>о</strong>ме пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Liphistiomorphae).<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> н<strong>а</strong>личие (<strong>о</strong>бычн<strong>о</strong> <strong>к</strong>рупн<strong>о</strong>й)<br />

<strong>с</strong>терн<strong>а</strong>льн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и н<strong>а</strong> брюшн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong> т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г, н<strong>а</strong>пример<br />

у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>, бы<strong>в</strong><strong>а</strong>ют р<strong>а</strong>зд<strong>в</strong>инуты <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

(ри<strong>с</strong>. 1413, 1422, 1434).<br />

Стернит <strong>с</strong>егмент<strong>а</strong> педип<strong>а</strong>льп у Araneae игр<strong>а</strong>ет<br />

р<strong>о</strong>ль нижней губы, <strong>в</strong> других групп<strong>а</strong>х <strong>о</strong>н<br />

з<strong>а</strong>метн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н. Брюшн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> (<strong>с</strong>егменты <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>). К<strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

имеют<strong>с</strong>я и <strong>с</strong>терниты (<strong>к</strong>р<strong>о</strong>ме <strong>в</strong><strong>с</strong>ех пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

Araneae).<br />

Гетер<strong>о</strong>н<strong>о</strong>мн<strong>о</strong><strong>с</strong>ть <strong>в</strong> р<strong>а</strong>з<strong>в</strong>итии <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей у<br />

Arachnida д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ет м<strong>а</strong><strong>к</strong><strong>с</strong>им<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жения<br />

<strong>с</strong>реди <strong>в</strong><strong>с</strong>ех Chelicerata. Хелицеры <strong>в</strong> <strong>о</strong>дних<br />

групп<strong>а</strong>х трех-пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>лешне<strong>в</strong>идные<br />

(Opiliomorphae), т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>в</strong> других (Araneae,<br />

Anthracomarti) <strong>о</strong>ни <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идные (ри<strong>с</strong>. 1383,<br />

1447, 1451); п<strong>о</strong><strong>с</strong>леднее, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong><br />

р<strong>а</strong>з<strong>в</strong>итием <strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бн<strong>о</strong><strong>с</strong>ти н<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong> пище<strong>в</strong><strong>а</strong>рения<br />

и прием<strong>а</strong> уже р<strong>а</strong>зжиженн<strong>о</strong>й пищи. У п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

и Anthracomarti эндиты т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп<br />

п<strong>о</strong><strong>к</strong>рыты <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зующими фильтр,<br />

через <strong>к</strong><strong>о</strong>т<strong>о</strong>рый пр<strong>о</strong>фильтр<strong>о</strong><strong>в</strong>ы<strong>в</strong><strong>а</strong>ет<strong>с</strong>я р<strong>а</strong>зжиженн<strong>а</strong>я<br />

пищ<strong>а</strong> (ри<strong>с</strong>. 1383 Б). Педип<strong>а</strong>льпы у <strong>в</strong><strong>с</strong>ех<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>тые, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные,

<strong>с</strong> з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>й л<strong>а</strong>п<strong>к</strong><strong>о</strong>й, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>ни бы<strong>в</strong><strong>а</strong>ют<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими, х<strong>о</strong>тя м<strong>о</strong>гут д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ть и б<strong>о</strong>лее<br />

<strong>к</strong>рупных р<strong>а</strong>змер<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1378, 1426), н<strong>о</strong> ни<strong>к</strong><strong>о</strong>гд<strong>а</strong><br />

при эт<strong>о</strong>м не <strong>в</strong>ып<strong>о</strong>лняют х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельн<strong>о</strong>й фун<strong>к</strong>ции.<br />

Х<strong>о</strong>дильные н<strong>о</strong>ги длинные, р<strong>а</strong><strong>с</strong>членены н<strong>а</strong><br />

ше<strong>с</strong>ть <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>ных члени<strong>к</strong><strong>о</strong><strong>в</strong> (т<strong>а</strong>зи<strong>к</strong>, <strong>в</strong>ертлуг,<br />

бедр<strong>о</strong>, patella, г<strong>о</strong>лень и л<strong>а</strong>п<strong>к</strong><strong>а</strong>); у мн<strong>о</strong>гих <strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

л<strong>а</strong>п<strong>к</strong>и м<strong>о</strong>гут быть д<strong>о</strong>п<strong>о</strong>лнительн<strong>о</strong> р<strong>а</strong><strong>с</strong>членены<br />

н<strong>а</strong> д<strong>в</strong><strong>а</strong> — пять члени<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>а</strong> <strong>в</strong>ертлуги и<br />

реже бедр<strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> р<strong>а</strong>зделяют<strong>с</strong>я <strong>к</strong><strong>а</strong>жд<strong>о</strong>е н<strong>а</strong><br />

д<strong>в</strong><strong>а</strong> члени<strong>к</strong><strong>а</strong>; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы зн<strong>а</strong>чительн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>к</strong>рыш<strong>к</strong>и н<strong>а</strong> VIII <strong>с</strong>егменте <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Opiliones и Haptopoda.<br />

У Аг<strong>а</strong>пе<strong>а</strong>е н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е эт<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я п<strong>а</strong>р<strong>а</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong>ид<strong>о</strong>измененными<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ыми <strong>к</strong>рыш<strong>к</strong><strong>а</strong>ми, п<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>у у <strong>с</strong>тернит<strong>о</strong><strong>в</strong><br />

IX и X <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>и п<strong>а</strong>рные лег<strong>о</strong>чные<br />

меш<strong>к</strong>и. Н<strong>а</strong> X <strong>с</strong>егменте у Аг<strong>а</strong>пе<strong>а</strong>е е<strong>с</strong>ть <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

<strong>в</strong> <strong>в</strong>иде пер<strong>в</strong><strong>о</strong>й п<strong>а</strong>ры п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>.<br />

Н<strong>а</strong>личие лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющих<strong>с</strong>я<br />

н<strong>а</strong> тех же <strong>с</strong>егмент<strong>а</strong>х, чт<strong>о</strong> и у <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных<br />

хелицер<strong>а</strong>т, у<strong>к</strong><strong>а</strong>зы<strong>в</strong><strong>а</strong>ет н<strong>а</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

этих групп. Интере<strong>с</strong>н<strong>о</strong>, чт<strong>о</strong> <strong>в</strong>ме<strong>с</strong>т<strong>о</strong> гребне<strong>в</strong>идных<br />

<strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> IX <strong>с</strong>егмент<strong>а</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>о</strong><strong>в</strong>, т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

у Amblypygi и Uropygi, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, и у<br />

Аг<strong>а</strong>пе<strong>а</strong>е, <strong>с</strong> друг<strong>о</strong>й, имеют<strong>с</strong>я либ<strong>о</strong> <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong><br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, либ<strong>о</strong> п<strong>а</strong>р<strong>а</strong> тр<strong>а</strong>хей. У п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>а</strong> XI <strong>с</strong>егменте <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong> п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>.<br />

О<strong>с</strong>т<strong>а</strong>льные <strong>с</strong>егменты мез<strong>о</strong>- и мет<strong>а</strong><strong>с</strong><strong>о</strong>мы у<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зных (Arachnida) лишены брюшных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Н<strong>а</strong>ружные ч<strong>а</strong><strong>с</strong>ти п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong><br />

Arachnida имеют <strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й генит<strong>а</strong>льн<strong>о</strong>й<br />

щели, при<strong>к</strong>рыт<strong>о</strong>й <strong>с</strong>переди неп<strong>а</strong>рн<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>к</strong><strong>о</strong>й, <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й <strong>с</strong>тернит<strong>о</strong>м VIII <strong>с</strong>егмент<strong>а</strong>.<br />

От<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует у Arachnida и трехл<strong>о</strong>п<strong>а</strong><strong>с</strong>тн<strong>о</strong>й<br />

генит<strong>а</strong>льный <strong>к</strong><strong>о</strong>ну<strong>с</strong>, <strong>с</strong><strong>в</strong><strong>о</strong>й<strong>с</strong>т<strong>в</strong>енный <strong>в</strong><strong>с</strong>ем<br />

Solifugomorpha и н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ным<br />

Scorpionomorpha (Palpigradi).<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida <strong>в</strong>едет <strong>с</strong><strong>в</strong><strong>о</strong>е пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение <strong>о</strong>т<br />

<strong>а</strong>мблипиг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зных члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих, д<strong>а</strong><strong>в</strong>ших <strong>в</strong><br />

<strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>м пери<strong>о</strong>де целый <strong>к</strong><strong>о</strong>мпле<strong>к</strong><strong>с</strong><br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных хелицер<strong>о</strong><strong>в</strong>ых, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых д<strong>в</strong>е<br />

группир<strong>о</strong><strong>в</strong><strong>к</strong>и (Opiliones и Аг<strong>а</strong>пе<strong>а</strong>е) <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили<strong>с</strong>ь<br />

д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> <strong>в</strong>ремени. От<strong>в</strong>ет<strong>в</strong>ление и<br />

д<strong>а</strong>льнейшее ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>о</strong>тдельных группир<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong><br />

(п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>) пр<strong>о</strong>те<strong>к</strong><strong>а</strong>л<strong>о</strong> не <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong>.<br />

Н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ными Arachnida н<strong>а</strong>д<strong>о</strong><br />

<strong>с</strong>чит<strong>а</strong>ть пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Opiliomorphae,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили п<strong>о</strong>лную <strong>с</strong>егмент<strong>а</strong>цию<br />

тел<strong>а</strong>, п<strong>а</strong>рные п<strong>о</strong>л<strong>о</strong><strong>в</strong>ые <strong>к</strong>рышеч<strong>к</strong>и н<strong>а</strong> VIII <strong>с</strong>тер-<br />

ните, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение <strong>с</strong>рединных гл<strong>а</strong>з н<strong>а</strong> буг<strong>о</strong>р<strong>к</strong>е,<br />

<strong>в</strong> центре г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong>. Вер<strong>о</strong>ятн<strong>о</strong>,<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бление Opiliomorphae <strong>о</strong>т и<strong>с</strong>х<strong>о</strong>дных ф<strong>о</strong>рм<br />

Amblypygi пр<strong>о</strong>из<strong>о</strong>шл<strong>о</strong> д<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бления <strong>о</strong>тряд<strong>а</strong><br />

Ricinulei. Р<strong>а</strong>з<strong>в</strong>итие п<strong>о</strong><strong>с</strong>ледних был<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>н<strong>о</strong> <strong>с</strong><br />

упл<strong>о</strong>тнением п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong> тел<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong> <strong>с</strong>илу<br />

не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong><strong>с</strong>ти <strong>с</strong><strong>о</strong>хр<strong>а</strong>нить п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong><strong>с</strong>ть ее р<strong>а</strong><strong>с</strong>ширения при<br />

п<strong>о</strong>треблении пищи и р<strong>а</strong>з<strong>в</strong>итии яиц, <strong>с</strong><strong>к</strong>леритиз<strong>о</strong><strong>в</strong><strong>а</strong>ли<strong>с</strong>ь<br />

<strong>о</strong>тдельными уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми, <strong>о</strong>бр<strong>а</strong>зуя <strong>с</strong>рединную<br />

и д<strong>в</strong>е <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ые з<strong>о</strong>ны тергит<strong>о</strong><strong>в</strong> и <strong>с</strong>тернит<strong>о</strong><strong>в</strong>.<br />

При эт<strong>о</strong>м Ricinulei <strong>с</strong><strong>о</strong>хр<strong>а</strong>нили б<strong>о</strong>лее<br />

примити<strong>в</strong>ные черты <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции дре<strong>в</strong>них<br />

Scorpionomorpha, <strong>а</strong> пр<strong>о</strong>и<strong>с</strong>шедшие <strong>о</strong>т их пред<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong>тряды п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Soluta, <strong>в</strong>идим<strong>о</strong>, <strong>в</strong> результ<strong>а</strong>те<br />

перех<strong>о</strong>д<strong>а</strong> <strong>к</strong> <strong>о</strong>бит<strong>а</strong>нию <strong>в</strong> н<strong>а</strong>земн<strong>о</strong><strong>в</strong><strong>о</strong>здушных<br />

з<strong>а</strong><strong>с</strong>ушли<strong>в</strong>ых у<strong>с</strong>л<strong>о</strong><strong>в</strong>иях, при<strong>о</strong>брели <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong><br />

пл<strong>о</strong>тный п<strong>а</strong>нцирь. Н<strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong>е гл<strong>а</strong><strong>в</strong>н<strong>о</strong>е изменение<br />

з<strong>а</strong><strong>к</strong>люч<strong>а</strong>л<strong>о</strong><strong>с</strong>ь у них, <strong>к</strong><strong>а</strong><strong>к</strong> и у других Arachnida,<br />

<strong>в</strong> р<strong>а</strong>з<strong>в</strong>итии п<strong>о</strong>дгиб<strong>а</strong>ния з<strong>а</strong>дних <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну. Н<strong>а</strong><strong>к</strong><strong>о</strong>нец, <strong>о</strong>тряд<br />

Аг<strong>а</strong>пе<strong>а</strong>е, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong><strong>о</strong>зни<strong>к</strong> п<strong>о</strong>чти <strong>о</strong>дн<strong>о</strong><strong>в</strong>ременн<strong>о</strong><br />

<strong>с</strong> <strong>о</strong>тряд<strong>а</strong>ми п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Soluta, прит<strong>о</strong>м не<br />

<strong>о</strong>т типичных Amblypygi, <strong>а</strong> <strong>о</strong>т н<strong>а</strong>ч<strong>а</strong><strong>в</strong>ших т<strong>о</strong>гд<strong>а</strong><br />

р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ть<strong>с</strong>я Ricinulei. Р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>о</strong>тряд<strong>а</strong> Anthracomarti<br />

и пр<strong>о</strong>из<strong>о</strong>шедшег<strong>о</strong> <strong>о</strong>т нег<strong>о</strong> Trigonotarbi<br />

<strong>с</strong> п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми <strong>о</strong>тряд<strong>а</strong> Аг<strong>а</strong>пе<strong>а</strong>е п<strong>о</strong>дчер<strong>к</strong>и<strong>в</strong><strong>а</strong>ет<strong>с</strong>я<br />

<strong>о</strong>дн<strong>о</strong><strong>о</strong>бр<strong>а</strong>зием <strong>с</strong>тр<strong>о</strong>ения р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> (р<strong>а</strong>з<strong>в</strong>итие<br />

<strong>к</strong><strong>о</strong>гте<strong>в</strong>идных хелицер и фильтр<strong>а</strong>ци<strong>о</strong>нн<strong>о</strong>й<br />

<strong>с</strong>ет<strong>к</strong>и н<strong>а</strong> эндит<strong>а</strong>х т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп<br />

(ри<strong>с</strong>. 1383 Б), <strong>к</strong>рупн<strong>о</strong>г<strong>о</strong> <strong>с</strong>тернум<strong>а</strong> и т. п. П<strong>а</strong>у<strong>к</strong>и<br />

<strong>о</strong>тряд<strong>а</strong> Аг<strong>а</strong>пе<strong>а</strong>е <strong>в</strong>едут <strong>с</strong><strong>в</strong><strong>о</strong>е н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong> <strong>о</strong>т б<strong>о</strong>лее<br />

примити<strong>в</strong>ных Amblypygi, чт<strong>о</strong> п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нением <strong>в</strong> эмбри<strong>о</strong>генезе не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

рудимент<strong>а</strong> з<strong>а</strong>днебрюшья, <strong>с</strong><strong>о</strong>хр<strong>а</strong>нением<br />

у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей примити<strong>в</strong>ных групп д<strong>в</strong>ух п<strong>а</strong>р<br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> и других брюшных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей,<br />

VII <strong>с</strong>егмент<strong>а</strong> тел<strong>а</strong> и т. п. и <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ием<br />

н<strong>а</strong> их <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме пр<strong>о</strong>д<strong>о</strong>льных б<strong>о</strong>р<strong>о</strong>зд, р<strong>а</strong>зделяющих<br />

тергиты и <strong>с</strong>терниты н<strong>а</strong> три з<strong>о</strong>ны. След<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>,<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida д<strong>о</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й<br />

<strong>с</strong>тепени гетер<strong>о</strong>генен п<strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждению, чт<strong>о</strong> и<br />

н<strong>а</strong>шл<strong>о</strong> <strong>о</strong>тр<strong>а</strong>жение <strong>в</strong> р<strong>а</strong>зделении <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> н<strong>а</strong> три<br />

п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>. Не<strong>с</strong>м<strong>о</strong>тря н<strong>а</strong> эт<strong>о</strong>, цельн<strong>о</strong><strong>с</strong>ть <strong>в</strong><strong>с</strong>ей<br />

<strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упн<strong>о</strong><strong>с</strong>ти <strong>о</strong>тряд<strong>о</strong><strong>в</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Arachnida п<strong>о</strong>дт<strong>в</strong>ержд<strong>а</strong>ет<strong>с</strong>я<br />

единым н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лением э<strong>в</strong><strong>о</strong>люции,<br />

н<strong>а</strong>шедшим <strong>в</strong>ыр<strong>а</strong>жение <strong>в</strong> пр<strong>о</strong>це<strong>с</strong><strong>с</strong>е и<strong>с</strong><strong>к</strong>ри<strong>в</strong>ления<br />

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong> и <strong>с</strong><strong>в</strong>яз<strong>а</strong>нн<strong>о</strong>й <strong>с</strong> этим <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>й<br />

интегр<strong>а</strong>цией <strong>с</strong>егмент<strong>а</strong>рн<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong><strong>а</strong> тел<strong>а</strong>.<br />

Кл<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida п<strong>о</strong>лучил р<strong>а</strong>знитие <strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не,<br />

<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>о</strong>тряд<strong>о</strong><strong>в</strong> Opiliones и Аг<strong>а</strong>пе<strong>а</strong>е<br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong>уют и ныне. Кл<strong>а</strong><strong>с</strong><strong>с</strong> Arachnida<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен) тремя п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>ми: Opiliomorphae,<br />

Soluta и Аг<strong>а</strong>пе<strong>а</strong>е.<br />

475<br />

http://jurassic.ru/

ПОДКЛАСС OPILIOMORPHAE.<br />

СЕНОКОСЦЕПОДОБНЫЕ ИЛИ ЛОЖНОПАУКИ<br />

Тел<strong>о</strong> <strong>к</strong>ругл<strong>о</strong>е или <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. Пр<strong>о</strong>п<strong>о</strong>д<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong> брюш<strong>к</strong><strong>о</strong>м. Н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>м<br />

щите имеют<strong>с</strong>я б<strong>о</strong>р<strong>о</strong>зды — <strong>с</strong>леды пр<strong>о</strong>шл<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>членения<br />

н<strong>а</strong> пр<strong>о</strong>пельтидий, мет<strong>а</strong>- и мез<strong>о</strong>пельтидий.<br />

Гл<strong>а</strong>з<strong>а</strong> м<strong>о</strong>гут быть или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ть;<br />

у <strong>о</strong>дних <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>ни р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти щит<strong>а</strong> <strong>в</strong> <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е д<strong>в</strong>ух<br />

или ше<strong>с</strong>ти, <strong>а</strong> у других <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>ни леж<strong>а</strong>т н<strong>а</strong><br />

треуг<strong>о</strong>льн<strong>о</strong>м гл<strong>а</strong>зн<strong>о</strong>м буг<strong>о</strong>р<strong>к</strong>е. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е,<br />

<strong>с</strong> 9—12 <strong>с</strong>егмент<strong>а</strong>ми, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

п<strong>о</strong><strong>с</strong>ледние р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти<br />

брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны тел<strong>а</strong>; X или XII тергиты <strong>о</strong>бр<strong>а</strong>зуют<br />

<strong>а</strong>н<strong>а</strong>льную <strong>к</strong>рышеч<strong>к</strong>у. Стернум имеет<strong>с</strong>я<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует; у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>н<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

между <strong>в</strong>ершин<strong>а</strong>ми т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г м<strong>о</strong>жет<br />

быть <strong>с</strong>ильн<strong>о</strong> хитинизир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>. Стерниты х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

<strong>в</strong>ыр<strong>а</strong>жены. Стернит I <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> <strong>в</strong> <strong>о</strong>дних групп<strong>а</strong>х треуг<strong>о</strong>льный, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<br />

между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г IV и п<strong>о</strong>з<strong>а</strong>ди них,<br />

<strong>а</strong> у других передние <strong>с</strong>терниты зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>с</strong>мещ<strong>а</strong>ют<strong>с</strong>я <strong>в</strong>перед и <strong>в</strong><strong>к</strong>лини<strong>в</strong><strong>а</strong>ют<strong>с</strong>я между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

н<strong>о</strong>г (<strong>в</strong>ы<strong>с</strong>шие Opiliones). Хелицеры<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>лешне<strong>в</strong>идные. П<strong>а</strong>льпы н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные.<br />

Н<strong>о</strong>ги <strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong> <strong>с</strong>терните VIII <strong>с</strong>егмент<strong>а</strong><br />

тел<strong>а</strong>; у мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>н<strong>о</strong> <strong>с</strong>н<strong>а</strong>бжен<strong>о</strong> п<strong>а</strong>рными<br />

генит<strong>а</strong>льными <strong>к</strong>рышеч<strong>к</strong><strong>а</strong>ми. Орг<strong>а</strong>ны дых<strong>а</strong>ния<br />

у низших Opiliomorphae (<strong>о</strong>тряды Haptopoda и<br />

Phalangiotarbi) пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены д<strong>в</strong>умя п<strong>а</strong>р<strong>а</strong>ми<br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных н<strong>а</strong> IX и X<br />

<strong>с</strong>тернит<strong>а</strong>х, <strong>а</strong> у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> Opiliones н<strong>а</strong> <strong>с</strong>тер­<br />

ОТРЯД HAPTOPODA<br />

Мел<strong>к</strong>ие н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые <strong>с</strong> <strong>к</strong>рупн<strong>о</strong>й<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й;<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь шир<strong>о</strong><strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong>членяет<strong>с</strong>я <strong>с</strong> брюш<strong>к</strong><strong>о</strong>м.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong>рупн<strong>а</strong>я, з<strong>а</strong>ним<strong>а</strong>ет<br />

треть длины тел<strong>а</strong>, п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы ее пл<strong>о</strong>тные, <strong>с</strong> груб<strong>о</strong>й<br />

т<strong>о</strong>чечн<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й; <strong>в</strong>д<strong>о</strong>ль пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й<br />

<strong>о</strong><strong>с</strong>и щит<strong>а</strong> пр<strong>о</strong>х<strong>о</strong>дят четыре б<strong>о</strong>р<strong>о</strong>зды, <strong>с</strong>рединные<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых чет<strong>к</strong><strong>о</strong> <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют уз<strong>к</strong>ую<br />

р<strong>а</strong>хиди<strong>а</strong>льную з<strong>о</strong>ну пр<strong>о</strong>пельтидия, <strong>а</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые<br />

— ег<strong>о</strong> пле<strong>в</strong>р<strong>а</strong>льные ч<strong>а</strong><strong>с</strong>ти. Близ переднег<strong>о</strong><br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> щит<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я п<strong>а</strong>р<strong>а</strong> <strong>к</strong>рупных гл<strong>а</strong>з.<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из 11 <strong>с</strong>егмент<strong>о</strong><strong>в</strong> (Petrunkevitch,<br />

1949); тергиты и <strong>с</strong>терниты <strong>с</strong> т<strong>о</strong>чечн<strong>о</strong>й<br />

пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й. Пле<strong>в</strong>ры мяг<strong>к</strong>ие, без <strong>с</strong><strong>к</strong>лерит<strong>о</strong><strong>в</strong>.<br />

П<strong>о</strong><strong>с</strong>ледние <strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я<br />

н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну (ри<strong>с</strong>. 1366), <strong>о</strong>дн<strong>а</strong><strong>к</strong><strong>о</strong> эт<strong>о</strong>т<br />

пр<strong>о</strong>це<strong>с</strong><strong>с</strong> <strong>в</strong> <strong>о</strong>тряде Haptopoda т<strong>о</strong>ль<strong>к</strong><strong>о</strong> еще н<strong>а</strong>ме­<br />

(Haptopodida)<br />

476<br />

http://jurassic.ru/<br />

ните IX <strong>с</strong>егмент<strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>о</strong>дн<strong>а</strong> п<strong>а</strong>р<strong>а</strong> тр<strong>а</strong>хей.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

Н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые,<br />

<strong>о</strong>бит<strong>а</strong>ющие <strong>в</strong> п<strong>о</strong>ч<strong>в</strong>е, <strong>в</strong><strong>о</strong> мху, п<strong>о</strong>д <strong>к</strong><strong>о</strong>рнями дере<strong>в</strong>ье<strong>в</strong>,<br />

<strong>в</strong> р<strong>а</strong>зличных п<strong>о</strong>л<strong>о</strong><strong>с</strong>тях и углублениях<br />

п<strong>о</strong>ч<strong>в</strong>ы и т. п. Хищни<strong>к</strong>и, ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременную эп<strong>о</strong>ху р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены п<strong>о</strong><strong>в</strong><strong>с</strong>еме<strong>с</strong>тн<strong>о</strong>,<br />

<strong>к</strong>р<strong>о</strong>ме х<strong>о</strong>л<strong>о</strong>дных <strong>о</strong>бл<strong>а</strong><strong>с</strong>тей. В <strong>к</strong><strong>а</strong>рб<strong>о</strong>не<br />

были р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены н<strong>а</strong> террит<strong>о</strong>рии Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Сен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цеп<strong>о</strong>д<strong>о</strong>бные<br />

хелицер<strong>о</strong><strong>в</strong>ые р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны примити<strong>в</strong>ным<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>м Scorpionomorpha, <strong>в</strong> ч<strong>а</strong><strong>с</strong>тн<strong>о</strong><strong>с</strong>ти<br />

Amblypygi, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых и <strong>в</strong>едут <strong>с</strong><strong>в</strong><strong>о</strong>е н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong><br />

(<strong>с</strong>м. <strong>с</strong>хему н<strong>а</strong> <strong>с</strong>тр. 384). Н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>ные<br />

Haptopoda, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, я<strong>в</strong>или<strong>с</strong>ь и<strong>с</strong>х<strong>о</strong>дн<strong>о</strong>й<br />

групп<strong>о</strong>й для <strong>в</strong>ы<strong>с</strong>ших Opiliomorphae, <strong>с</strong>реди <strong>к</strong><strong>о</strong>т<strong>о</strong>рых,<br />

н<strong>а</strong>пример, п<strong>о</strong>д<strong>о</strong>тряд Cyphophthalmi имеет<br />

мн<strong>о</strong>г<strong>о</strong> <strong>о</strong>бщег<strong>о</strong> <strong>с</strong> Haptopoda. Р<strong>а</strong>з<strong>в</strong>итие <strong>в</strong><strong>с</strong>ех<br />

Opiliomorphae пр<strong>о</strong>те<strong>к</strong><strong>а</strong>л<strong>о</strong> <strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong>й <strong>с</strong>тепени<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> <strong>с</strong> д<strong>в</strong>умя другими п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>ми<br />

Arachnida. В<strong>с</strong>е эти группы <strong>с</strong><strong>в</strong>язы<strong>в</strong><strong>а</strong>ют пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение<br />

<strong>о</strong>т <strong>о</strong>бщих <strong>а</strong>мблипиг<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных<br />

пред<strong>к</strong><strong>о</strong><strong>в</strong> и <strong>о</strong>бщие тенденции м<strong>о</strong>рф<strong>о</strong>генез<strong>а</strong>, <strong>с</strong><strong>в</strong>яз<strong>а</strong>нные<br />

<strong>с</strong> и<strong>с</strong><strong>к</strong>ри<strong>в</strong>лением пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>.<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Opiliomorphae р<strong>а</strong><strong>с</strong>п<strong>а</strong>д<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong><br />

три <strong>о</strong>тряд<strong>а</strong>: Haptopoda, Phalangiotarbi, Opiliones;<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>с</strong>уще<strong>с</strong>т<strong>в</strong>уют<br />

ныне.<br />

чен. Н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не пр<strong>о</strong><strong>с</strong><strong>о</strong>мы трех<strong>с</strong>егментный<br />

<strong>с</strong>тернум. Хелицеры трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>к</strong>лешне<strong>в</strong>идные. Педип<strong>а</strong>льпы<br />

пр<strong>о</strong><strong>с</strong>тые, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные, ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые;<br />

т<strong>а</strong>зи<strong>к</strong>и хелицер <strong>с</strong>литы <strong>с</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми педип<strong>а</strong>льп<br />

<strong>в</strong> <strong>о</strong>дну пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у. Эндиты т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп<br />

<strong>в</strong>ытянуты и <strong>о</strong>бр<strong>а</strong>зуют м<strong>а</strong><strong>к</strong><strong>с</strong>иллярные<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты. В<strong>с</strong>е н<strong>о</strong>ги длинные, т<strong>о</strong>н<strong>к</strong>ие, х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>, <strong>с</strong> п<strong>о</strong>д<strong>в</strong>ижными цилиндриче<strong>с</strong><strong>к</strong>ими<br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми, <strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Л<strong>а</strong>п<strong>к</strong>и т<strong>о</strong>н<strong>к</strong>ие, пятичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

П<strong>а</strong>телл<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че или р<strong>а</strong><strong>в</strong>н<strong>а</strong> п<strong>о</strong><br />

длине г<strong>о</strong>лени. Пер<strong>в</strong>ый <strong>с</strong>тернит <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

треуг<strong>о</strong>льный, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г<br />

IV и п<strong>о</strong>з<strong>а</strong>ди них. Следующий <strong>с</strong>тернит VIII <strong>с</strong>егмент<strong>а</strong><br />

<strong>о</strong>чень <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширен; <strong>в</strong>близи з<strong>а</strong>днег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я ег<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие,<br />

имеющее <strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й щели. П<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м

пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й <strong>в</strong>д<strong>о</strong>ль <strong>с</strong>редней линии<br />

тел<strong>а</strong>.<br />

Haptopoda были н<strong>а</strong>земными, дыш<strong>а</strong><strong>в</strong>шими <strong>с</strong><br />

п<strong>о</strong>м<strong>о</strong>щью лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, хищными<br />

хелицер<strong>о</strong><strong>в</strong>ыми. К<strong>а</strong>рб<strong>о</strong>н.<br />

С<strong>а</strong>мые примити<strong>в</strong>ные пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong><br />

Opiliomorphae, пр<strong>о</strong>из<strong>о</strong>шедшие <strong>о</strong>т <strong>а</strong>м-<br />

Ри<strong>с</strong>. 1366. Отряд Haptopoda.<br />

1366. Plesiosiro madeleyi Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид; А — <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, X6,0;<br />

Х4,5; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1910)<br />

<strong>о</strong>т нег<strong>о</strong> имеют<strong>с</strong>я <strong>о</strong><strong>к</strong>руглые п<strong>а</strong>рные <strong>к</strong>л<strong>а</strong>п<strong>а</strong>ны,<br />

рудименты брюшных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей VIII, п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong>,<br />

<strong>с</strong>егмент<strong>а</strong> .(ри<strong>с</strong>. 1366 А). П<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям IX и X<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>с</strong><strong>к</strong>л<strong>а</strong>дч<strong>а</strong>тые лег<strong>о</strong>чные<br />

меш<strong>к</strong>и. Стерниты X—XVIII <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong><br />

Вымершие н<strong>а</strong>земные хелицер<strong>о</strong><strong>в</strong>ые <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льным<br />

или <strong>в</strong>ытянутым тел<strong>о</strong>м (ри<strong>с</strong>. 1372,<br />

1373). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong>членен<strong>а</strong> <strong>с</strong> брюш<strong>к</strong><strong>о</strong>м п<strong>о</strong> <strong>в</strong><strong>с</strong>ей<br />

ширине тел<strong>а</strong>. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий щит <strong>к</strong>рупный,<br />

<strong>в</strong>ытянутый (ри<strong>с</strong>. 1373), треуг<strong>о</strong>льный (ри<strong>с</strong>.<br />

1372) или <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тный (ри<strong>с</strong>. 1375 Б). З<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й ег<strong>о</strong> прям<strong>о</strong>линейный (ри<strong>с</strong>. 1374 Б), <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>лый<br />

(ри<strong>с</strong>. 1373) или <strong>с</strong> <strong>с</strong>ильн<strong>о</strong> <strong>о</strong>ттянут<strong>о</strong>й<br />

н<strong>а</strong>з<strong>а</strong>д <strong>с</strong>рединн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью (ри<strong>с</strong>. 1369). Передний<br />

<strong>к</strong><strong>о</strong>нец щит<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен, либ<strong>о</strong> <strong>с</strong> уз<strong>к</strong>им<br />

п<strong>а</strong>льце<strong>в</strong>идным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м (ри<strong>с</strong>. 1368, 1369).<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть щит<strong>а</strong> <strong>с</strong> <strong>к</strong>рупн<strong>о</strong>т<strong>о</strong>чечн<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й,<br />

без пр<strong>о</strong>д<strong>о</strong>льных б<strong>о</strong>р<strong>о</strong>зд; <strong>в</strong> передней<br />

•трети ег<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены д<strong>в</strong><strong>а</strong> <strong>к</strong>рупных гл<strong>а</strong>з<strong>а</strong>.<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>а</strong>я или яйце<strong>в</strong>идн<strong>а</strong>я, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит<br />

из 10—12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, п<strong>о</strong><strong>с</strong>ледние из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong>; тергит<br />

XII <strong>с</strong>егмент<strong>а</strong> брюш<strong>к</strong><strong>а</strong> у мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> пред­<br />

ОТРЯД PHALANGIOTARBI<br />

(Architarbida)<br />

4<br />

• <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

блипиг<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных <strong>о</strong>рг<strong>а</strong>низм<strong>о</strong><strong>в</strong> и д<strong>а</strong><strong>в</strong>шие н<strong>а</strong>ч<strong>а</strong>л<strong>о</strong><br />

другим п<strong>а</strong>у<strong>к</strong><strong>а</strong>м-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>ц<strong>а</strong>м <strong>о</strong>тряд<strong>о</strong><strong>в</strong> Phalangiotarbi<br />

и Opiliones. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Plesiosironidae<br />

<strong>с</strong> <strong>о</strong>дним р<strong>о</strong>д<strong>о</strong>м и <strong>в</strong>ид<strong>о</strong>м (Plesiosiro<br />

Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911) из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> Англии (ри<strong>с</strong>. 1366).<br />

<strong>с</strong>т<strong>а</strong><strong>в</strong>лен <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н<strong>о</strong>м <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия. Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>й<br />

<strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тью р<strong>а</strong><strong>с</strong>членения <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы я<strong>в</strong>ляет<strong>с</strong>я зн<strong>а</strong>чительн<strong>о</strong>е<br />

у<strong>к</strong><strong>о</strong>р<strong>о</strong>чение тергит<strong>о</strong><strong>в</strong> пер<strong>в</strong>ых <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>; рез<strong>к</strong><strong>о</strong> <strong>с</strong>уженные тергиты <strong>о</strong>бычн<strong>о</strong><br />

<strong>о</strong>гиб<strong>а</strong>ют з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong><br />

(ри<strong>с</strong>. 1368, 1372 А). Стернит<strong>о</strong><strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong><br />

меньше, чем тергит<strong>о</strong><strong>в</strong>; <strong>о</strong>бычн<strong>о</strong> их бы<strong>в</strong><strong>а</strong>ет <strong>с</strong>емь;<br />

реже <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong><strong>о</strong> их р<strong>а</strong><strong>в</strong>н<strong>о</strong> чи<strong>с</strong>лу тергит<strong>о</strong><strong>в</strong>.<br />

Стерниты имеют д<strong>в</strong>е пр<strong>о</strong>д<strong>о</strong>льные б<strong>о</strong>р<strong>о</strong>зды, идущие<br />

<strong>в</strong>д<strong>о</strong>ль <strong>к</strong>р<strong>а</strong>е<strong>в</strong> тел<strong>а</strong> (ри<strong>с</strong>. 1372 Б). Стернит<br />

пер<strong>в</strong><strong>о</strong>г<strong>о</strong> брюшн<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> треуг<strong>о</strong>льный, н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я<br />

между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г IV и п<strong>о</strong>з<strong>а</strong>ди них.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>в</strong> <strong>в</strong>иде п<strong>о</strong>перечн<strong>о</strong>й щели<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong> VIII <strong>с</strong>терните (ри<strong>с</strong>. 1370);<br />

имеют<strong>с</strong>я п<strong>а</strong>рные <strong>к</strong>рыш<strong>к</strong>и п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия.<br />

Дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> лег<strong>о</strong>чными<br />

меш<strong>к</strong><strong>а</strong>ми, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женными п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м<br />

477<br />

http://jurassic.ru/

1376А<br />

http://jurassic.ru/

IX—XI <strong>с</strong>тернит<strong>о</strong><strong>в</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Хелицеры<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>лешне<strong>в</strong>идные. Педип<strong>а</strong>льпы<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, т<strong>о</strong>н<strong>к</strong>ие, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные, <strong>с</strong> ше<strong>с</strong>тью<br />

члени<strong>к</strong><strong>а</strong>ми. Т<strong>а</strong>зи<strong>к</strong>и педин<strong>а</strong>льп <strong>с</strong> м<strong>а</strong><strong>к</strong><strong>с</strong>иллярными<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г треуг<strong>о</strong>льные,<br />

пл<strong>о</strong>тн<strong>о</strong> приж<strong>а</strong>ты друг <strong>к</strong> другу, неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><br />

<strong>с</strong>литы <strong>с</strong> грудью; у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong>, <strong>в</strong><strong>о</strong>зм<strong>о</strong>жн<strong>о</strong>,<br />

были п<strong>о</strong>д<strong>в</strong>ижными т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г IV. Н<strong>о</strong>ги длинные,<br />

<strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н.<br />

Н<strong>а</strong>земные <strong>с</strong><strong>а</strong>пр<strong>о</strong>ф<strong>а</strong>ги или хищни<strong>к</strong>и, <strong>о</strong>бит<strong>а</strong><strong>в</strong>шие,<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>в</strong> п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е ле<strong>с</strong><strong>о</strong><strong>в</strong>.<br />

Эт<strong>о</strong>т <strong>о</strong>тряд <strong>о</strong>чень близ<strong>о</strong><strong>к</strong> <strong>к</strong> <strong>о</strong>тряду Haptopoda,<br />

<strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong>, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, и пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дит. Близ<strong>о</strong><strong>к</strong><br />

<strong>к</strong> р<strong>о</strong>д<strong>о</strong>н<strong>а</strong>ч<strong>а</strong>льн<strong>о</strong>й группе <strong>о</strong>тряд<strong>а</strong> Opiliones,<br />

пер<strong>в</strong>ые пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> т<strong>а</strong><strong>к</strong>же п<strong>о</strong>я<strong>в</strong>или<strong>с</strong>ь<br />

<strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не, н<strong>о</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рый имеет и <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Heterotarbidae,<br />

Opiliotarbidae и Architarbidae.<br />

СЕМЕЙСТВО HETEROTARBIDAE PETRUNKEVITCH,<br />

1913<br />

Н<strong>о</strong>ги пер<strong>в</strong><strong>о</strong>й п<strong>а</strong>ры <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие и длинные,<br />

<strong>с</strong> трубч<strong>а</strong>тыми, <strong>в</strong>ытянутыми члени<strong>к</strong><strong>а</strong>ми. Н<strong>о</strong>ги<br />

II <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>а</strong> н<strong>о</strong>ги III и IV б<strong>о</strong>лее длинные, <strong>с</strong><br />

т<strong>о</strong>л<strong>с</strong>тыми члени<strong>к</strong><strong>а</strong>ми. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г I р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

п<strong>о</strong>д <strong>о</strong><strong>с</strong>трым угл<strong>о</strong>м <strong>к</strong> пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>,<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены друг <strong>о</strong>т друг<strong>а</strong> н<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong>е<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>яние. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г II м<strong>а</strong>лень<strong>к</strong>ие, леж<strong>а</strong>т<br />

п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г III и IV<br />

имеют ф<strong>о</strong>рму уз<strong>к</strong>их прям<strong>о</strong>уг<strong>о</strong>льни<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>в</strong>ытянутых<br />

п<strong>о</strong>пере<strong>к</strong> тел<strong>а</strong> и <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ющих<strong>с</strong>я друг <strong>с</strong><br />

друг<strong>о</strong>м н<strong>а</strong> <strong>с</strong>редней линии (ри<strong>с</strong>. 1367). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> де<strong>с</strong>ятью тергит<strong>а</strong>ми, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пер<strong>в</strong>ые<br />

пять <strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены. Длин<strong>а</strong> тел<strong>а</strong> 14 мм.<br />

К<strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и. Один р<strong>о</strong>д Heterotarbus<br />

Petrunkevitch, 1913.<br />

СЕМЕЙСТВО OPILIOTARBIDAE PETRUNKEVITCH,<br />

1945<br />

Н<strong>о</strong>ги <strong>в</strong><strong>с</strong>ех п<strong>а</strong>р р<strong>а</strong>з<strong>в</strong>иты <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>; <strong>о</strong>ни удлиненные,<br />

умеренн<strong>о</strong> т<strong>о</strong>л<strong>с</strong>тые. Т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г<br />

треуг<strong>о</strong>льные, р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены пл<strong>о</strong>тн<strong>о</strong><br />

друг <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> друг<strong>а</strong>; <strong>в</strong>нутренними <strong>к</strong><strong>о</strong>нц<strong>а</strong>ми <strong>о</strong>ни<br />

п<strong>о</strong>чти <strong>с</strong>мы<strong>к</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong> центре груди. Вертлуги н<strong>о</strong>г<br />

III и IV р<strong>а</strong>зделены н<strong>а</strong> д<strong>в</strong><strong>а</strong> члени<strong>к</strong><strong>а</strong>. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> 11 <strong>с</strong>егмент<strong>а</strong>ми, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пер<strong>в</strong>ые ше<strong>с</strong>ть<br />

Ри<strong>с</strong>. 1367—1375.<br />

1367. Heterotarbus ovatus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, Х4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1368. Opiliotarbus elongatus<br />

(Scudder); A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу; Б — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху,<br />

х4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1369. Architarbus<br />

minor Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1370. Architarbus rotundatus<br />

Scudder; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, X 2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch,<br />

1913). 1371. Phalangiotarbus subovalis (Woodward);<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, Х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911).<br />

<strong>с</strong>ильн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены (ри<strong>с</strong>. 1368). Длин<strong>а</strong> тел<strong>а</strong> 14—<br />

15 мм. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и. Один р<strong>о</strong>д<br />

Opiliotarbus Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1900.<br />

СЕМЕЙСТВО ARCHITARBIDAE KARSCH, 1882<br />

Н<strong>о</strong>ги <strong>в</strong><strong>с</strong>ех п<strong>а</strong>р р<strong>а</strong>з<strong>в</strong>иты примерн<strong>о</strong> <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>;<br />

<strong>о</strong>ни т<strong>о</strong>н<strong>к</strong>ие, <strong>с</strong>редней <strong>в</strong>еличины, х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong><br />

тип<strong>а</strong>. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г треуг<strong>о</strong>льные, пл<strong>о</strong>тн<strong>о</strong> приж<strong>а</strong>ты<br />

друг <strong>к</strong> другу, н<strong>о</strong> <strong>в</strong>ершин<strong>а</strong>ми не д<strong>о</strong>х<strong>о</strong>дят д<strong>о</strong><br />

центр<strong>а</strong> груди. Т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп <strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>с</strong><br />

т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г I или <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дными и<br />

т<strong>о</strong>гд<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я между ними. Н<strong>а</strong> брюш<strong>к</strong>е<br />

де<strong>с</strong>ять тергит<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых пер<strong>в</strong>ые пять —<br />

ше<strong>с</strong>ть з<strong>а</strong>метн<strong>о</strong> у<strong>к</strong><strong>о</strong>р<strong>о</strong>чены, и де<strong>в</strong>ять <strong>с</strong>тернит<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1369, 1375). К<strong>а</strong>рб<strong>о</strong>н. 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Architarbus Scudder, 1868 (Geraphrynus<br />

Scudder, 1884). Тип р<strong>о</strong>д<strong>а</strong> — A. rotundatus Scudder,<br />

1868; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> без<br />

гл<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>; гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце щит<strong>а</strong>. Ф<strong>о</strong>рм<strong>а</strong><br />

ег<strong>о</strong> р<strong>о</strong>мб<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, <strong>с</strong> п<strong>а</strong>льце<strong>в</strong>идным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м<br />

н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце и <strong>с</strong> <strong>о</strong>ттянут<strong>о</strong>й <strong>с</strong>редней<br />

ч<strong>а</strong><strong>с</strong>тью з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я. Длин<strong>а</strong> тел<strong>а</strong> 15—22 мм<br />

(ри<strong>с</strong>. 1369, 1370). Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Кр<strong>о</strong>ме тип<strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>, д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из С. Амери<strong>к</strong>и и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Phalangiotarbus Н a a s е, 1890. Тип р<strong>о</strong>д<strong>а</strong> —-<br />

Architarbus subovalis Woodward, 1872; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Англия. Гл<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong> и гл<strong>а</strong>з нет. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

шире <strong>с</strong><strong>в</strong><strong>о</strong>ей длины, <strong>с</strong>переди шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен<strong>а</strong>,<br />

п<strong>о</strong>лу<strong>к</strong>ругл<strong>а</strong>я; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й.<br />

Брюш<strong>к</strong><strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> б<strong>о</strong>лее шир<strong>о</strong><strong>к</strong><strong>о</strong>е<br />

<strong>с</strong>переди. Четыре пер<strong>в</strong>ых тергит<strong>а</strong> <strong>с</strong>ужены. Длин<strong>а</strong><br />

<strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 5,8 мм (ри<strong>с</strong>. 1371). Один <strong>в</strong>ид.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Goniotarbus Petrunkevitch, 1945. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Geraphrynus tuberculatus Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия. Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. О<strong>к</strong><strong>о</strong>л<strong>о</strong> переднег<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>мы гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> <strong>с</strong><br />

ше<strong>с</strong>тью гл<strong>а</strong>з<strong>а</strong>ми п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong> ф<strong>о</strong>рме<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>мб<strong>а</strong>, треуг<strong>о</strong>льн<strong>а</strong>я <strong>с</strong>переди и <strong>с</strong><br />

<strong>о</strong>ттянут<strong>о</strong>й <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>тью з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я<br />

(ри<strong>с</strong>. 1372А, Б). Пять пер<strong>в</strong>ых тергит<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>с</strong>ж<strong>а</strong>ты, <strong>о</strong>ни ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> <strong>о</strong>гиб<strong>а</strong>ют з<strong>а</strong>дний<br />

<strong>к</strong>р<strong>а</strong>й щит<strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> 16—18 мм. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Mesotarbus Petrunkevitch, 1949. Тип<br />

р<strong>о</strong>д<strong>а</strong> — М. intermedius Petrunkevitch, 1949;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия. Н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет предыдущий<br />

р<strong>о</strong>д, н<strong>о</strong> гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> треуг<strong>о</strong>льный, <strong>с</strong> тремя<br />

п<strong>а</strong>р<strong>а</strong>ми гл<strong>а</strong>з п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м ег<strong>о</strong>. Передний <strong>к</strong>р<strong>а</strong>й<br />

ряд Phalangiotarbi<br />

479<br />

http://jurassic.ru/<br />

1372. Goniotarbus angulatus (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>); <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong><strong>в</strong>ерху,<br />

Б — <strong>с</strong>низу; Х4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911). 1373. Leptotarbus<br />

torpedo (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>); <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong>низу, Б — <strong>с</strong><strong>в</strong>ерху;<br />

Х4.0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911). 1374. Orthotarbus mtnutus<br />

(Petrunkevitch); <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong>низу, £ — <strong>с</strong>рерху; х5,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1375. Geratarbus<br />

lacoet Scudder; <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong>низу, F <strong>с</strong><strong>в</strong>ерху; Х4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913)

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренным <strong>с</strong>рединным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м;<br />

з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен. Пять передних<br />

тергит<strong>о</strong><strong>в</strong> уз<strong>к</strong>ие, <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти <strong>с</strong> д<strong>в</strong>умя<br />

ряд<strong>а</strong>ми буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>. Длин<strong>а</strong> тел<strong>а</strong> 10—18 мм. Три<br />

<strong>в</strong>ид<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Leptotarbus Petrunkevitch, 1945. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Geraphrynus torpedo Pocock, 1911;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия. Тел<strong>о</strong> уз<strong>к</strong><strong>о</strong>е, <strong>с</strong>иг<strong>а</strong>р<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>е.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины,<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>трен<strong>а</strong> <strong>с</strong>переди и шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен<strong>а</strong><br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце. Гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> треуг<strong>о</strong>льный,<br />

<strong>с</strong> ше<strong>с</strong>тью гл<strong>а</strong>з<strong>а</strong>ми. Четыре передних тергит<strong>а</strong><br />

<strong>с</strong>ужены, <strong>а</strong> длин<strong>а</strong> пят<strong>о</strong>г<strong>о</strong> тергит<strong>а</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине длины ше<strong>с</strong>т<strong>о</strong>г<strong>о</strong>. Длин<strong>а</strong> брюш<strong>к</strong><strong>а</strong> <strong>в</strong><br />

2,5 р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>льше ег<strong>о</strong> ширины. Длин<strong>а</strong> тел<strong>а</strong><br />

13,5 мм, ширин<strong>а</strong> 2,86 мм (ри<strong>с</strong>. 1373). Один <strong>в</strong>ид.<br />

Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Orthotarbus Petrunkevitch, 1945. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Geratarbus minutus Petrunkevitch,<br />

1913; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>. Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> прямым з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ем, <strong>к</strong>уп<strong>о</strong>л<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я<br />

<strong>с</strong>переди, <strong>с</strong> м<strong>а</strong>лень<strong>к</strong>им з<strong>а</strong><strong>о</strong><strong>с</strong>тренным<br />

<strong>с</strong>рединным <strong>в</strong>ы<strong>с</strong>туп<strong>о</strong>м. Брюш<strong>к</strong><strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>в</strong><br />

1,5 р<strong>а</strong>з<strong>а</strong> длиннее <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong>. Передние пять<br />

тергит<strong>о</strong><strong>в</strong> <strong>с</strong>ужены, <strong>с</strong><strong>о</strong> <strong>с</strong>рединн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й н<strong>а</strong><br />

п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г IV з<strong>а</strong>метн<strong>о</strong> длиннее<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных. Вершины <strong>в</strong><strong>с</strong>ех т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> з<strong>а</strong><strong>к</strong>руглены;<br />

пл<strong>о</strong>щ<strong>а</strong>д<strong>к</strong><strong>а</strong> <strong>к</strong><strong>о</strong>жи между ними д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я.<br />

Пер<strong>в</strong>ый <strong>с</strong>тернит брюш<strong>к</strong><strong>а</strong> м<strong>а</strong>лень<strong>к</strong>ий,<br />

треуг<strong>о</strong>льный (ри<strong>с</strong>. 1374). Н<strong>о</strong>ги I и II т<strong>о</strong>ньше<br />

н<strong>о</strong>г з<strong>а</strong>дних п<strong>а</strong>р. Длин<strong>а</strong> тел<strong>а</strong> 9,0—11,5 мм. Ср.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из С. Амери<strong>к</strong>и и <strong>о</strong>дин из Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

Geratarbus Scudder, 1890. Тип р<strong>о</strong>д<strong>а</strong> —<br />

G, tacoei Scudder, 1890; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong>.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> прям<strong>о</strong>уг<strong>о</strong>льн<strong>а</strong>я, <strong>с</strong> прямыми п<strong>а</strong>р<strong>а</strong>ллельными<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми и з<strong>а</strong><strong>к</strong>ругленными переднеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми<br />

угл<strong>а</strong>ми и <strong>с</strong>редним л<strong>о</strong>бным <strong>в</strong>ы<strong>с</strong>туп<br />

п<strong>о</strong>м (ри<strong>с</strong>. 1375). З<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

прям<strong>о</strong>й; н<strong>а</strong> ее переднем <strong>к</strong>р<strong>а</strong>е <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

л<strong>о</strong>бн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong> имеет<strong>с</strong>я треуг<strong>о</strong>льный гл<strong>а</strong>зн<strong>о</strong>й<br />

буг<strong>о</strong>р<strong>о</strong><strong>к</strong>. Брюш<strong>к</strong><strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. Пер<strong>в</strong>ые пять тергит<strong>о</strong><strong>в</strong><br />

уз<strong>к</strong>ие. Пер<strong>в</strong>ый <strong>с</strong>тернит <strong>о</strong>чень <strong>к</strong>рупный,<br />

треуг<strong>о</strong>льный. Н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не члени<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>о</strong>г и педип<strong>а</strong>льп имеют<strong>с</strong>я притуплённые<br />

шил<strong>о</strong><strong>в</strong>идные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты (ри<strong>с</strong>. 1375 А, Б). Длин<strong>а</strong><br />

тел<strong>а</strong> 10—12 мм. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из<br />

С. Амери<strong>к</strong>и и Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и р<strong>о</strong>ды<br />

Hadrachne Melander, 1903; Metatarbus<br />

Petrunkevitch, 1913; Paratarbus Petrunkevitch,<br />

1945; Discotarbus Petrunkevitch, 1913; Ootarbus<br />

Petrunkevitch, 1945.<br />

ОТРЯД OPILIONES. ПАУКИ-СЕНОКОСЦЫ,<br />

л<strong>о</strong>жн<strong>о</strong> ПАУКИ, ИЛИ <strong>о</strong>пили<strong>о</strong>ны<br />

Тел<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>е, <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е или <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong><br />

р<strong>а</strong><strong>с</strong>члененн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

<strong>с</strong><strong>о</strong>членен<strong>а</strong> <strong>с</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й. У б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> ф<strong>о</strong>рм<br />

н<strong>о</strong>ги д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>о</strong>гр<strong>о</strong>мн<strong>о</strong>й <strong>в</strong>еличины (ри<strong>с</strong>. 1382,<br />

1379). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>лит<strong>а</strong> из ше<strong>с</strong>ти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>,<br />

п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> щит<strong>о</strong>м, н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> у <strong>о</strong>тдельных<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>няют<strong>с</strong>я б<strong>о</strong>р<strong>о</strong>зды, <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>ющие<br />

гр<strong>а</strong>ницы <strong>с</strong>егмент<strong>о</strong><strong>в</strong> (Gonyleptidae,<br />

Gagrellinae). Н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

имеет<strong>с</strong>я гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> <strong>с</strong> д<strong>в</strong>умя (ин<strong>о</strong>гд<strong>а</strong><br />

<strong>о</strong>чень <strong>к</strong>рупными) или <strong>с</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong>ими гл<strong>а</strong>з<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1382). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>а</strong>я, <strong>с</strong> де<strong>с</strong>ятью<br />

члени<strong>к</strong><strong>а</strong>ми, де<strong>в</strong>ять из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых я<strong>с</strong>н<strong>о</strong> <strong>в</strong>ыр<strong>а</strong>жены,<br />

<strong>а</strong> тергит де<strong>с</strong>ят<strong>о</strong>г<strong>о</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>й <strong>к</strong>рышеч<strong>к</strong><strong>о</strong>й<br />

(operculum). У не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong><br />

1376. Nemastomoides elaveris Thevenin; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, Х5,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1377. totrogulus jayoll<br />

Thevenin; A — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>низу, <strong>с</strong> т<strong>а</strong>зи<strong>к</strong><strong>о</strong>м н<strong>о</strong>г; Б — <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

передних н<strong>о</strong>г и педип<strong>а</strong>льп; v 10,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1953). 1378. Gonyleptes nemastomoides Koch et Berendt;<br />

<strong>о</strong>бщий <strong>в</strong>ид, xl,5, п<strong>а</strong>ле<strong>о</strong>ген. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

und Berendt, 1854). 1379. Nemastoma tuberculatum Koch et Berendt;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X 2,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

(Phalangioidea)<br />

Ри<strong>с</strong>. 1376—1382. Отряд Opilones<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>ме р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я длинные шипы (Triaenonychidae,<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые Gonyleptidae). К<strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

II и III <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ие <strong>с</strong>терниты<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я <strong>в</strong>ме<strong>с</strong>те и <strong>о</strong>бр<strong>а</strong>зуют <strong>о</strong>бширную <strong>о</strong>бщую<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>у, п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

д<strong>в</strong><strong>а</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия тр<strong>а</strong>хей. Тергиты IX и X<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы перемещены н<strong>а</strong> брюшную<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну з<strong>а</strong>днег<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong> тел<strong>а</strong>. Хелицеры<br />

трехчлени<strong>с</strong>тые, <strong>к</strong>лешне<strong>в</strong>идные; ин<strong>о</strong>гд<strong>а</strong> (Ischyropsalidae)<br />

<strong>о</strong>чень <strong>к</strong>рупные. Педип<strong>а</strong>льпы р<strong>а</strong>з<strong>в</strong>иты<br />

р<strong>а</strong>зличн<strong>о</strong>; <strong>о</strong>ни либ<strong>о</strong> пр<strong>о</strong><strong>с</strong>тые, т<strong>о</strong>н<strong>к</strong>ие,<br />

н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные (ри<strong>с</strong>. 1377 б, 1382), либ<strong>о</strong> бы<strong>в</strong><strong>а</strong>ют<br />

<strong>о</strong>чень длинные (ри<strong>с</strong>. 1379), либ<strong>о</strong> н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х<br />

г<strong>о</strong>леней или п<strong>а</strong>телл их п<strong>о</strong>я<strong>в</strong>ляют<strong>с</strong>я <strong>к</strong>рупные<br />

п<strong>а</strong>льце<strong>в</strong>идные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты, либ<strong>о</strong>, н<strong>а</strong><strong>к</strong><strong>о</strong>нец, педи-<br />

480<br />

http://jurassic.ru/<br />

(Koch und Berendt, 1854). 1380. Opilio ovalis Koch et Berendt<br />

<strong>о</strong>бщий <strong>в</strong>ид, x 2,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

und Berendt, 1854). 1381. Opillo ramiger Koch et Berendt; п<strong>а</strong>льпы<br />

и передний <strong>к</strong><strong>о</strong>нец тел<strong>а</strong>, X 4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt, 1854). 1382. Platybunus dentipalpus<br />

Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид, Х4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1854)

31 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

http://jurassic.ru/

п<strong>а</strong>льпы д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>о</strong>гр<strong>о</strong>мн<strong>о</strong>й длины <strong>с</strong> <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

л<strong>а</strong>п<strong>к</strong><strong>о</strong>й и <strong>в</strong>ид<strong>о</strong>измененными <strong>о</strong><strong>с</strong>т<strong>а</strong>льными<br />

члени<strong>к</strong><strong>а</strong>ми. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г <strong>к</strong>рупные, неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong><br />

<strong>с</strong>р<strong>а</strong>щены <strong>с</strong> тел<strong>о</strong>м, пл<strong>о</strong>тн<strong>о</strong> прилег<strong>а</strong>ют друг <strong>к</strong><br />

другу, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>в</strong>ыд<strong>в</strong>инутых<br />

<strong>в</strong>перед <strong>с</strong>тернит<strong>о</strong><strong>в</strong>. Эндиты т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

педип<strong>а</strong>льп и н<strong>о</strong>г I, <strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> т<strong>а</strong><strong>к</strong>же и т<strong>а</strong>зи<strong>к</strong>и<br />

н<strong>о</strong>г II и III <strong>о</strong>бр<strong>а</strong>зуют м<strong>а</strong><strong>к</strong><strong>с</strong>иллярные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты.<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти н<strong>а</strong> <strong>с</strong>егмент<strong>а</strong>х мез<strong>о</strong><strong>с</strong><strong>о</strong>мы пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Opiliones п<strong>а</strong>рными <strong>к</strong>рыш<strong>к</strong><strong>а</strong>ми<br />

п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия и п<strong>а</strong>р<strong>о</strong>й тр<strong>а</strong>хейных<br />

дых<strong>а</strong>лец, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичны пер<strong>в</strong><strong>о</strong>й<br />

п<strong>а</strong>ре лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> других хелицер<strong>о</strong><strong>в</strong>ых.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong>мещен<strong>о</strong> д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>перед;<br />

у <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> имеет<strong>с</strong>я длинный яйце<strong>к</strong>л<strong>а</strong>д, <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющий<br />

между п<strong>а</strong>льп<strong>а</strong>ми. Н<strong>о</strong>ги т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> Cyphophthalmi<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, у других Opiliones,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>о</strong>чень т<strong>о</strong>н<strong>к</strong>ие и длинные, <strong>с</strong> трубч<strong>а</strong>тыми<br />

члени<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1380). Н<strong>о</strong>г<strong>а</strong> р<strong>а</strong><strong>с</strong>членен<strong>а</strong><br />

н<strong>а</strong> <strong>с</strong>емь члени<strong>к</strong><strong>о</strong><strong>в</strong>; л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Длин<strong>а</strong> тел<strong>а</strong> 1—22 мм.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия, ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

Обит<strong>а</strong>ют <strong>в</strong> земле, <strong>в</strong><strong>о</strong> мху,<br />

п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями, <strong>с</strong>реди гниющих р<strong>а</strong><strong>с</strong>тительных<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>и. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> немн<strong>о</strong>гие <strong>в</strong>ы<strong>с</strong>шие<br />

Opiliones (н<strong>а</strong>пример, Phalangiidae) п<strong>о</strong><strong>к</strong>ид<strong>а</strong>ют<br />

у<strong>в</strong>л<strong>а</strong>жненные <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>ты и <strong>о</strong>бит<strong>а</strong>ют н<strong>а</strong> <strong>с</strong><strong>к</strong><strong>а</strong>л<strong>а</strong>х,<br />

<strong>с</strong>т<strong>в</strong><strong>о</strong>л<strong>а</strong>х дере<strong>в</strong>ье<strong>в</strong>, <strong>с</strong>тен<strong>а</strong>х д<strong>о</strong>м<strong>о</strong><strong>в</strong> и т. п. Хищни<strong>к</strong>и.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременную эп<strong>о</strong>ху Opiliones р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены<br />

п<strong>о</strong> <strong>в</strong><strong>с</strong>ему земн<strong>о</strong>му ш<strong>а</strong>ру, и<strong>с</strong><strong>к</strong>люч<strong>а</strong>я<br />

Ант<strong>а</strong>р<strong>к</strong>тиду. В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны<br />

из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> и юры С. Амери<strong>к</strong>и и 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Ряд <strong>в</strong>ид<strong>о</strong><strong>в</strong> из<strong>в</strong>е<strong>с</strong>тен из б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> янт<strong>а</strong>ря.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Си<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение Opiliones д<strong>о</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong><br />

<strong>в</strong>ремени т<strong>о</strong>чн<strong>о</strong> не <strong>в</strong>ыя<strong>с</strong>нен<strong>о</strong>. Ке<strong>с</strong>тнер<br />

(Kastner, 1935) <strong>с</strong>ближ<strong>а</strong>ет их, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

<strong>с</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>емыми Phalangiotarbi и Haptopoda,<br />

<strong>а</strong> <strong>с</strong> друг<strong>о</strong>й — <strong>с</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рыми Scorpionomorpha и<br />

<strong>к</strong>лещ<strong>а</strong>ми. Б<strong>о</strong>лее дет<strong>а</strong>льн<strong>о</strong>е р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>о</strong>трение <strong>о</strong>бщих<br />

черт <strong>в</strong> <strong>с</strong>тр<strong>о</strong>ении Opiliones и других Chelicerata<br />

при<strong>в</strong>ели К<strong>а</strong><strong>с</strong>тнер<strong>а</strong>. <strong>к</strong> <strong>в</strong>ы<strong>в</strong><strong>о</strong>ду <strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>е<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>це<strong>в</strong> <strong>с</strong> <strong>о</strong>тряд<strong>а</strong>ми п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Pedipalpides,<br />

<strong>а</strong> з<strong>а</strong>тем уже <strong>с</strong><strong>о</strong> <strong>с</strong><strong>к</strong><strong>о</strong>рпи<strong>о</strong>н<strong>а</strong>ми и <strong>к</strong>лещ<strong>а</strong>ми.<br />

П<strong>о</strong> н<strong>а</strong>шим <strong>в</strong><strong>о</strong>ззрениям, <strong>о</strong>тряд Opiliones<br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енен <strong>о</strong>тряд<strong>а</strong>м Phalangiotarbi и Haptopoda,<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляет единую группу<br />

(п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>) Opiliomorphae (<strong>с</strong>м. <strong>с</strong>тр. 384). С<strong>в</strong><strong>о</strong>е<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение п<strong>а</strong>у<strong>к</strong>и-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>цы не<strong>с</strong><strong>о</strong>мненн<strong>о</strong><br />

<strong>в</strong>едут <strong>о</strong>т примити<strong>в</strong>ных Scorpionomorpha, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>,<br />

<strong>о</strong>т <strong>а</strong>мблипиги<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных хелицер<strong>а</strong>т.<br />

Об<strong>о</strong><strong>с</strong><strong>о</strong>бление Opiliones пр<strong>о</strong>из<strong>о</strong>шл<strong>о</strong> не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>а</strong>ньше р<strong>а</strong>з<strong>в</strong>ития Ricinulei, Soluta и Araneae<br />

(<strong>с</strong>м. <strong>с</strong>тр. 384 и <strong>с</strong>хему н<strong>а</strong> <strong>с</strong>тр. 383).<br />

Отряд Opiliones р<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>:<br />

Cyphophthalmi, Laniatores, Palpatores.<br />

482<br />

http://jurassic.ru/<br />

ПОДОТРЯД CYPHOPHTHALMI<br />

(Cyphophthalmina)<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, <strong>с</strong>л<strong>а</strong>б<strong>о</strong> дифференцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е.<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> тергиты пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьми <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> <strong>о</strong>б<strong>о</strong>зн<strong>а</strong>чены:<br />

IX и X тергиты р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>с</strong>уб<strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие <strong>с</strong>мещен<strong>о</strong> <strong>в</strong>перед,<br />

н<strong>о</strong> не при<strong>к</strong>рыт<strong>о</strong> <strong>с</strong>тернит<strong>а</strong>ми VIII и IX <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

Педип<strong>а</strong>льпы <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, т<strong>о</strong>н<strong>к</strong>ие, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные.<br />

Длин<strong>а</strong> тел<strong>а</strong> 2—3 мм. Предп<strong>о</strong>л<strong>о</strong>жение<br />

Петрун<strong>к</strong>е<strong>в</strong>ич<strong>а</strong> (Petrunkevitch, 1949, 1953) <strong>о</strong><br />

прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ти р<strong>о</strong>д<strong>о</strong><strong>в</strong> Nemastomoides Thevenin<br />

и Eotrogulus Thevenin <strong>к</strong> <strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>ц<strong>а</strong>м п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong><br />

Palpatores <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong>, т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>в</strong><strong>с</strong>ех пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

эт<strong>о</strong>г<strong>о</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

<strong>с</strong>мещен<strong>о</strong> д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>перед и при<strong>к</strong>рыт<strong>о</strong> <strong>с</strong>тернит<strong>а</strong>ми<br />

переднебрюшных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. К<strong>а</strong>рб<strong>о</strong>н —ныне.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне (Sironidae)<br />

и <strong>о</strong>дн<strong>о</strong> <strong>в</strong> <strong>к</strong><strong>а</strong>рб<strong>о</strong>не (Nemastomoididae><br />

СЕМЕЙСТВО NEMASTOMOIDIDAE<br />

PETRUNKEVITCH, 1955<br />

(Eotrogulidae Petrunkevitch, 1965)<br />

Тел<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е или <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> з<strong>а</strong>ним<strong>а</strong>ет<br />

1<br />

/з— l<br />

h ч<strong>а</strong><strong>с</strong>ть тел<strong>а</strong>. Имеют<strong>с</strong>я д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>а</strong>.<br />

Т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блены. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г<br />

<strong>к</strong>рупные, <strong>в</strong>еличин<strong>а</strong> их <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ет <strong>о</strong>т передних<br />

н<strong>о</strong>г <strong>к</strong> з<strong>а</strong>дним. Внутренние <strong>к</strong>р<strong>а</strong>я т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г<br />

IV н<strong>а</strong>иб<strong>о</strong>лее длинные, прям<strong>о</strong>уг<strong>о</strong>льные, п<strong>о</strong>чти<br />

д<strong>о</strong>х<strong>о</strong>дят друг д<strong>о</strong> друг<strong>а</strong> или <strong>с</strong>мы<strong>к</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong><br />

<strong>с</strong>редней линии тел<strong>а</strong> (ри<strong>с</strong>. 1376). Н<strong>о</strong>ги длинные<br />

и т<strong>о</strong>н<strong>к</strong>ие. Внутренние <strong>к</strong>р<strong>а</strong>я т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>бр<strong>а</strong>мляют<br />

неб<strong>о</strong>льшую <strong>о</strong><strong>в</strong><strong>а</strong>льную пл<strong>о</strong>щ<strong>а</strong>д<strong>к</strong>у н<strong>а</strong> груди.<br />

П<strong>а</strong>льпы т<strong>о</strong>н<strong>к</strong>ие, пр<strong>о</strong><strong>с</strong>тые, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные (ри<strong>с</strong>.<br />

1377Б). Длин<strong>а</strong> тел<strong>а</strong> 2,7—12,0 мм. Ср. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

С. Амери<strong>к</strong>и и Е<strong>в</strong>р<strong>о</strong>пы. Эт<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>люч<strong>а</strong>ет<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые Петрун<strong>к</strong>е<strong>в</strong>ич (Petrunkevitch,<br />

1955) п<strong>о</strong>мещ<strong>а</strong>ет <strong>в</strong> д<strong>в</strong><strong>а</strong> <strong>о</strong><strong>с</strong><strong>о</strong>бых<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>.<br />

Nemastomoides Thevenin, 1901 (Protopilio<br />

Petrunkevitch, 1913). Тип р<strong>о</strong>д<strong>а</strong> — N. etaveris<br />

Thevenin, 1901; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

Т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г без м<strong>а</strong><strong>к</strong><strong>с</strong>иллярных л<strong>о</strong>п<strong>а</strong><strong>с</strong>тей;<br />

<strong>в</strong>еличин<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> <strong>в</strong><strong>о</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ет <strong>о</strong>т пер<strong>в</strong>ых н<strong>о</strong>г <strong>к</strong><br />

п<strong>о</strong><strong>с</strong>ледним; т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г IV прям<strong>о</strong>уг<strong>о</strong>льные,<br />

<strong>с</strong>мы<strong>к</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> <strong>с</strong>редней линии тел<strong>а</strong>; н<strong>о</strong>ги <strong>о</strong>чень<br />

длинные и т<strong>о</strong>н<strong>к</strong>ие, <strong>в</strong>ертлуги их не р<strong>а</strong><strong>с</strong>членены.<br />

О<strong>к</strong><strong>о</strong>л<strong>о</strong> переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я пр<strong>о</strong><strong>с</strong><strong>о</strong>мы имеют<strong>с</strong>я д<strong>в</strong><strong>а</strong><br />

неб<strong>о</strong>льших гл<strong>а</strong>з<strong>а</strong> (ри<strong>с</strong>. 1376). Ср. и <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н.<br />

Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong> из 3. Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

Eotrogulus Thevenin, 1901. Тип р<strong>о</strong>д<strong>а</strong>.—<br />

Е. fayoll Thevenin, 1901; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> треуг<strong>о</strong>льн<strong>а</strong>я, зн<strong>а</strong>чительн<strong>о</strong> длиннее<br />

<strong>с</strong><strong>в</strong><strong>о</strong>ей ширины, <strong>с</strong> д<strong>в</strong>умя неб<strong>о</strong>льшими гл<strong>а</strong>з<strong>а</strong>ми<br />

н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце. Т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>в</strong> д<strong>в</strong><strong>а</strong> п<strong>а</strong>р<strong>а</strong>ллельных ряд<strong>а</strong>, <strong>о</strong><strong>к</strong>руглые; педип<strong>а</strong>ль-

пы м<strong>а</strong>лень<strong>к</strong>ие, пр<strong>о</strong><strong>с</strong>тые, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные (ри<strong>с</strong>.<br />

1377 А; <strong>в</strong>ертлуги н<strong>о</strong>г трубч<strong>а</strong>тые. Длин<strong>а</strong> тел<strong>а</strong><br />

12 мм, длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>к</strong><strong>с</strong><strong>а</strong> 5,5 мм. Один <strong>в</strong>ид. В.<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

ПОДОТРЯД LANIOTORES<br />

Тел<strong>о</strong> б<strong>о</strong>лее у<strong>к</strong><strong>о</strong>р<strong>о</strong>чен<strong>о</strong>, з<strong>а</strong><strong>к</strong>руглен<strong>о</strong>, рез<strong>к</strong><strong>о</strong><br />

п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>му и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му;<br />

пер<strong>в</strong><strong>а</strong>я лишь <strong>с</strong> нея<strong>с</strong>ными <strong>с</strong>лед<strong>а</strong>ми <strong>с</strong>егмент<strong>а</strong>ции.<br />

П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие при<strong>к</strong>рыт<strong>о</strong> <strong>с</strong>д<strong>в</strong>инутыми <strong>в</strong>перед<br />

<strong>с</strong>тернит<strong>а</strong>ми брюш<strong>к</strong><strong>а</strong>. Педип<strong>а</strong>льпы <strong>к</strong>рупные,<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>гте<strong>в</strong>идными л<strong>а</strong>п<strong>к</strong><strong>а</strong>ми и р<strong>а</strong>зличн<strong>о</strong><br />

<strong>в</strong>ид<strong>о</strong>измененными другими члени<strong>к</strong><strong>а</strong>ми, <strong>с</strong>н<strong>а</strong>бженными<br />

шип<strong>а</strong>ми , игл<strong>а</strong>ми и т. п. (ри<strong>с</strong>. 1378).<br />

Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г III и IV <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми или<br />

<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>ят<strong>с</strong>я трех<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>ыми з<strong>а</strong> <strong>с</strong>чет <strong>в</strong>ид<strong>о</strong>изменения<br />

предл<strong>а</strong>п<strong>к</strong>и (arolium) или <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>г<strong>о</strong><br />

р<strong>а</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ния д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>к</strong>р<strong>а</strong>я <strong>в</strong>ершины<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong>й л<strong>а</strong>п<strong>к</strong>и. Б<strong>о</strong>лее <strong>к</strong>рупные <strong>в</strong>иды; длин<strong>а</strong><br />

тел<strong>а</strong> д<strong>о</strong> 10 мм. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне ше<strong>с</strong>ть <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Oncopodidae, Phalangodidae,<br />

Assamiidae, Cosmetidae, Triaenonychidae<br />

и Gonyleptidae; п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

СЕМЕЙСТВО GONYLEPTIDAE SIMON, 1879<br />

Педип<strong>а</strong>льпы т<strong>о</strong>л<strong>с</strong>тые, длинные; <strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>о</strong>й<br />

члени<strong>к</strong> их <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идный, м<strong>о</strong>гущий прижим<strong>а</strong>ть<strong>с</strong>я<br />

<strong>к</strong> г<strong>о</strong>лени; н<strong>а</strong> п<strong>о</strong><strong>с</strong>ледней, <strong>а</strong> т<strong>а</strong><strong>к</strong>же н<strong>а</strong><br />

<strong>к</strong><strong>о</strong>ленн<strong>о</strong>м члени<strong>к</strong>е и ин<strong>о</strong>гд<strong>а</strong> н<strong>а</strong> других члени<strong>к</strong><strong>а</strong>х—<br />

шипы (ри<strong>с</strong>. 1378). Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г III и IV<br />

<strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми, <strong>с</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й <strong>в</strong>ершин<strong>о</strong>й<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong>й л<strong>а</strong>п<strong>к</strong>и (pseuderychium). Тергиты VI и<br />

VIII <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные. Х<strong>о</strong>дильные<br />

н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные. Длин<strong>а</strong> тел<strong>а</strong><br />

<strong>о</strong>т 5 мм (б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong>) д<strong>о</strong> 18 мм. П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 300<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них Gonyleptes Kjrby, 1819 (ри<strong>с</strong>. 1378)<br />

н<strong>а</strong>йден <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

.<br />

ПОДОТРЯД PALPATORES<br />

Тел<strong>о</strong> <strong>о</strong><strong>к</strong>ругл<strong>о</strong>е или <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине ин<strong>о</strong>гд<strong>а</strong> имеет<br />

п<strong>о</strong>перечные б<strong>о</strong>р<strong>о</strong>зды. Гл<strong>а</strong>з<strong>а</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>ют <strong>о</strong>гр<strong>о</strong>мн<strong>о</strong>й <strong>в</strong>еличины (ри<strong>с</strong>. 1382).<br />

Педип<strong>а</strong>льпы пр<strong>о</strong><strong>с</strong>тые, <strong>о</strong>бычн<strong>о</strong> неб<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> р<strong>а</strong>змер<strong>а</strong>;<br />

л<strong>а</strong>п<strong>к</strong>и их з<strong>а</strong><strong>к</strong>руглены н<strong>а</strong> <strong>в</strong>ершине. У<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не г<strong>о</strong>леней<br />

или <strong>к</strong><strong>о</strong>ленных члени<strong>к</strong><strong>о</strong><strong>в</strong> имеют<strong>с</strong>я п<strong>а</strong>льце<strong>в</strong>идные<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты (ри<strong>с</strong>. 1381). Н<strong>о</strong>ги <strong>о</strong>чень<br />

длинные и т<strong>о</strong>н<strong>к</strong>ие. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г III и IV т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>с</strong> <strong>о</strong>дним <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м. К<strong>а</strong>рб<strong>о</strong>н — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Trogulidea и<br />

Phalangiidea.<br />

483<br />

НАДСЕМЕЙСТВО TROGULIDEA<br />

(Dyspnoi)<br />

Л<strong>а</strong>п<strong>к</strong><strong>а</strong> педип<strong>а</strong>льп <strong>к</strong><strong>о</strong>р<strong>о</strong>че г<strong>о</strong>лени. П<strong>а</strong>льце<strong>в</strong>идный<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong>е н<strong>о</strong>г II <strong>о</strong>т<strong>с</strong>у<strong>с</strong>т<strong>в</strong>ует<br />

или <strong>о</strong>чень м<strong>а</strong>лень<strong>к</strong>ий. Н<strong>о</strong>ги без <strong>с</strong>тигм. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Trogulidae,<br />

Acropsopilionidae, Ischyropsalidae, Nemastomatidae;<br />

д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ISCHYROPSALIDAE<br />

SIMON, 1879<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> без <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>. Хелицеры <strong>о</strong>чень<br />

<strong>к</strong>рупные, б<strong>о</strong>льше тел<strong>а</strong>. Мет<strong>а</strong>пельтидий <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленный.<br />

Педип<strong>а</strong>льпы т<strong>о</strong>н<strong>к</strong>ие, <strong>к</strong><strong>о</strong>р<strong>о</strong>че н<strong>о</strong>г и<br />

хелицер. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне четыре р<strong>о</strong>д<strong>а</strong>, из них Sabacon Simon,<br />

1879, <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО NEMASTOMATIDAE<br />

SIMON, 1872<br />

Передний <strong>к</strong>р<strong>а</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> з<strong>а</strong><strong>к</strong>руглен,<br />

без <strong>с</strong>рединн<strong>о</strong>г<strong>о</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>. Мез<strong>о</strong>- и мет<strong>а</strong>пельтидий<br />

<strong>с</strong>ли<strong>в</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м (ри<strong>с</strong>.<br />

1379). Педип<strong>а</strong>льпы длинные, н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>,<br />

из них Nemastoma С. L. Koch, 1836 <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО PHALANGIIDEA<br />

(Eupnoi)<br />

Л<strong>а</strong>п<strong>к</strong><strong>а</strong> педип<strong>а</strong>льп длиннее г<strong>о</strong>лени. Выр<strong>о</strong><strong>с</strong>т<br />

н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х н<strong>о</strong>г II я<strong>в</strong><strong>с</strong>т<strong>в</strong>енный. Н<strong>о</strong>г<strong>а</strong> <strong>с</strong> <strong>а</strong><strong>к</strong>це<strong>с</strong><strong>с</strong><strong>о</strong>рными<br />

<strong>с</strong>тигм<strong>а</strong>ми н<strong>а</strong> бедр<strong>а</strong>х. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>о</strong>дн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Phalangiidae, из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>е<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО PHALANGIIDAE<br />

SIMON, 1879<br />

Тел<strong>о</strong> мяг<strong>к</strong><strong>о</strong>е. Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> буг<strong>о</strong>р<strong>к</strong>е<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленными<br />

члени<strong>к</strong><strong>а</strong>ми. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 120 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Opilio Herbst,<br />

1798 (ри<strong>с</strong>. 1380, 1381); Platybunus С. L. Koch,<br />

1854 (ри<strong>с</strong>. 1382); Caddo Banks, 1892; Cheiromachus<br />

Menge, 1854; Dicranopalpus Doleschall,<br />

1852; Liobunum C. L. Koch, 1854; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и из<strong>в</strong>е<strong>с</strong>тен р<strong>о</strong>д Phalan~<br />

gium Linnaeus, 1758.<br />

http://jurassic.ru/<br />

31*

Ри<strong>с</strong>. 1383—1386. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pal<br />

1383. Pa.laeocharinoid.es hornei Hirst; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х40.0; Б — хелицеры и <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ы педип<strong>а</strong>льп и передних<br />

н<strong>о</strong>г, Х120.0; В — <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong> <strong>с</strong>б<strong>о</strong><strong>к</strong>у, X 40,0; де<strong>в</strong><strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Hirst, 1923). 1384. Palaeocharinus sp.; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, Х30.0,<br />

484<br />

DCharinidae и Anthracosironidae<br />

http://jurassic.ru/<br />

де<strong>в</strong><strong>о</strong>н,» 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hirst, 1923). 1385. Palaecharinus rhyniensis<br />

Hirst; <strong>о</strong>бщий <strong>в</strong>идХ<strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х40.0, де<strong>в</strong><strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hirst, J1923).<br />

1386. Anthracosiro woodwardi Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху; Х5,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Англия (Petrunkevitch, 1949)

ПОДКЛАСС SOLUTA. ПАНЦИРНЫЕ ПАУКИ<br />

Тел<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>е, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>с</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong><strong>о</strong>членен<strong>а</strong> <strong>с</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; <strong>с</strong><strong>о</strong>член<strong>о</strong><strong>в</strong>н<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

ширине тел<strong>а</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы <strong>в</strong><strong>с</strong>ег<strong>о</strong> тел<strong>а</strong> <strong>о</strong>чень пл<strong>о</strong>тные,<br />

<strong>с</strong> буг<strong>о</strong>рч<strong>а</strong>тым (ри<strong>с</strong>. 1393, 1401), <strong>с</strong>етч<strong>а</strong>тым<br />

(ри<strong>с</strong>. 1399Б), т<strong>о</strong>чечным (ри<strong>с</strong>. 1386) или иным<br />

рез<strong>к</strong>им <strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м (ри<strong>с</strong>. 1407, 1413А). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий<br />

щит р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong> <strong>в</strong>ыпу<strong>к</strong>лый (б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Anthracomarti) или <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й<br />

<strong>с</strong>рединн<strong>о</strong>й з<strong>о</strong>н<strong>о</strong>й и <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ыми п<strong>о</strong>лями<br />

(ри<strong>с</strong>. 1387, 1393). В з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти щит<strong>а</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> имеют<strong>с</strong>я п<strong>о</strong>перечные б<strong>о</strong>р<strong>о</strong>зды,<br />

<strong>о</strong>б<strong>о</strong>зн<strong>а</strong>ч<strong>а</strong>ющие гр<strong>а</strong>ницы <strong>с</strong>лияния пр<strong>о</strong>-, мез<strong>о</strong>- и<br />

мет<strong>а</strong>пельтидия. Обычн<strong>о</strong> имеет<strong>с</strong>я д<strong>в</strong><strong>а</strong> <strong>к</strong>рупных<br />

гл<strong>а</strong>з<strong>а</strong> (ри<strong>с</strong>. 1385). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> р<strong>а</strong><strong>с</strong>членен<strong>а</strong> н<strong>а</strong><br />

11 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые из них <strong>с</strong>литны и нея<strong>с</strong>ны.<br />

Д<strong>в</strong><strong>а</strong> <strong>с</strong>егмент<strong>а</strong> з<strong>а</strong>гиб<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> брюшную<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну; <strong>с</strong>терниты этих <strong>с</strong>егмент<strong>о</strong><strong>в</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны,<br />

<strong>а</strong> тергит XI <strong>с</strong>егмент<strong>а</strong> пре<strong>в</strong>р<strong>а</strong>щен <strong>в</strong><br />

<strong>а</strong>н<strong>а</strong>льный <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н (ри<strong>с</strong>. 1383А, 1409). Тергиты<br />

д<strong>в</strong>умя пр<strong>о</strong>д<strong>о</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми р<strong>а</strong>зделены н<strong>а</strong><br />

<strong>с</strong>рединную и д<strong>в</strong>е <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ые ч<strong>а</strong><strong>с</strong>ти. У Anthracomarti,<br />

<strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и тергит<strong>о</strong><strong>в</strong><br />

<strong>с</strong>ильн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширяют<strong>с</strong>я и <strong>о</strong>бр<strong>а</strong>зуют б<strong>о</strong>лее или<br />

менее шир<strong>о</strong><strong>к</strong>ие <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ые з<strong>о</strong>ны (ри<strong>с</strong>. 1408А,<br />

1413А). П<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>е <strong>с</strong>тр<strong>о</strong>ение тергит<strong>о</strong><strong>в</strong> п<strong>а</strong>нцирные<br />

п<strong>а</strong>у<strong>к</strong>и не<strong>с</strong><strong>о</strong>мненн<strong>о</strong> ун<strong>а</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ли <strong>о</strong>т пред<strong>к</strong><strong>о</strong><strong>в</strong><br />

Ricinulei, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>ни пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят.<br />

Брюшн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы п<strong>о</strong><strong>к</strong>рыт<strong>а</strong> шир<strong>о</strong><strong>к</strong>ими<br />

и, <strong>в</strong>идим<strong>о</strong>, не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong>д<strong>в</strong>ижными т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

педип<strong>а</strong>льп и х<strong>о</strong>дильных н<strong>о</strong>г, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у<br />

единичных <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong>при<strong>к</strong><strong>а</strong><strong>с</strong><strong>а</strong>ют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м<br />

н<strong>а</strong> <strong>с</strong>редней линии тел<strong>а</strong> (ри<strong>с</strong>. 1402Б), т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong><br />

у п<strong>о</strong>д<strong>а</strong><strong>в</strong>ляющег<strong>о</strong> б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong>ни <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют<br />

д<strong>о</strong><strong>в</strong><strong>о</strong>льн<strong>о</strong> шир<strong>о</strong><strong>к</strong>ую <strong>о</strong><strong>в</strong><strong>а</strong>льную пл<strong>о</strong>щ<strong>а</strong>д<strong>к</strong>у,<br />

н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен <strong>с</strong>тернум<br />

(ри<strong>с</strong>. 1383А). Т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп и ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

т<strong>а</strong><strong>к</strong>же н<strong>о</strong>г I н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>с</strong>н<strong>а</strong>бжены<br />

длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми (ин<strong>о</strong>гд<strong>а</strong> з<strong>а</strong>зубрены), <strong>о</strong>бр<strong>а</strong>зующими<br />

фильтр<strong>а</strong>ци<strong>о</strong>нную <strong>с</strong>ет<strong>к</strong>у, <strong>к</strong><strong>а</strong><strong>к</strong> у<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>о</strong>тряд<strong>а</strong> Araneae (ри<strong>с</strong>. 1383В).<br />

Стерниты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>иты: з<strong>а</strong>дние<br />

<strong>с</strong>терниты дуг<strong>о</strong><strong>о</strong>бр<strong>а</strong>зн<strong>о</strong> из<strong>о</strong>гнуты, <strong>о</strong><strong>к</strong>руж<strong>а</strong>я<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие и тергиты п<strong>о</strong><strong>с</strong>ледних <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

тел<strong>а</strong> (ри<strong>с</strong>. 1392Б, 1408Б). Хелицеры <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие,<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идные, <strong>к</strong><strong>а</strong><strong>к</strong> у<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1383); <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, п<strong>а</strong>нцирные п<strong>а</strong>у<strong>к</strong>и<br />

имели н<strong>а</strong>ружн<strong>о</strong>е пище<strong>в</strong><strong>а</strong>рение. Педип<strong>а</strong>льпы<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, пр<strong>о</strong><strong>с</strong>тые (ри<strong>с</strong>. I413A). Х<strong>о</strong>дильные<br />

н<strong>о</strong>ги <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, ут<strong>о</strong>лщенные, <strong>с</strong>е-<br />

мичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я н<strong>а</strong> <strong>с</strong>терните<br />

VIII <strong>с</strong>егмент<strong>а</strong>; генит<strong>а</strong>льные <strong>к</strong>л<strong>а</strong>п<strong>а</strong>ны п<strong>а</strong>рные,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>в</strong> <strong>в</strong>иде д<strong>в</strong>ух <strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1402Б). П<strong>о</strong><br />

<strong>к</strong>р<strong>а</strong>ям II, III и IV <strong>с</strong>тернит<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

лег<strong>о</strong>чные меш<strong>к</strong>и. Де<strong>в</strong><strong>о</strong>н — пермь.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение.<br />

Сух<strong>о</strong>путные хелицер<strong>о</strong><strong>в</strong>ые,<br />

<strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, хищни<strong>к</strong>и. Н<strong>а</strong>личие у Anthracomarti<br />

шир<strong>о</strong><strong>к</strong>их <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ых з<strong>о</strong>н, м<strong>о</strong>жет быть <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует<br />

<strong>о</strong>б <strong>о</strong>бит<strong>а</strong>нии н<strong>а</strong> <strong>с</strong>ыпучих <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>а</strong>х.<br />

В де<strong>в</strong><strong>о</strong>не, <strong>к</strong><strong>а</strong>рб<strong>о</strong>не и перми были шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нены н<strong>а</strong> м<strong>а</strong>тери<strong>к</strong><strong>а</strong>х Е<strong>в</strong>р<strong>а</strong>зии и<br />

Амери<strong>к</strong>и.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. П<strong>а</strong>нцирные<br />

п<strong>а</strong>у<strong>к</strong>и п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Soluta пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят <strong>о</strong>т примити<strong>в</strong>ных<br />

Scorpionomorpha, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>о</strong>т<br />

пред<strong>к</strong><strong>о</strong><strong>в</strong> Amblypygi. П<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>телями п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>г<strong>о</strong><br />

фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong> я<strong>в</strong>ляет<strong>с</strong>я н<strong>а</strong>личие<br />

<strong>с</strong>тернум<strong>а</strong>, п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>г<strong>о</strong> <strong>с</strong><strong>о</strong>членения г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди<br />

и брюш<strong>к</strong><strong>а</strong>, лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> пер<strong>в</strong>ых брюшных<br />

<strong>с</strong>тернит<strong>а</strong>х, п<strong>а</strong>рных <strong>к</strong>рыше<strong>к</strong> у п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong><br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия, <strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>с</strong><strong>о</strong>хр<strong>а</strong>нение у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>их Trigonotarbi (Palaeocharinoid.es<br />

hornei Hirst), рудимент<strong>о</strong><strong>в</strong> XII и XIII <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> <strong>в</strong>иде м<strong>а</strong>лень<strong>к</strong><strong>о</strong>г<strong>о</strong> цилиндриче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

«х<strong>в</strong><strong>о</strong><strong>с</strong>ти<strong>к</strong><strong>а</strong>» п<strong>о</strong>з<strong>а</strong>ди <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия<br />

(ри<strong>с</strong>. 1383А); у других Trigonotarbi и <strong>в</strong><strong>с</strong>ех<br />

Anthracomarti этих рудимент<strong>о</strong><strong>в</strong> уже нет, н<strong>о</strong><br />

<strong>о</strong>ни, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

п<strong>о</strong>я<strong>в</strong>ляли<strong>с</strong>ь при эмбри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>м р<strong>а</strong>з<strong>в</strong>итии.<br />

Предп<strong>о</strong>л<strong>о</strong>жение не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей <strong>о</strong><br />

р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>е и д<strong>а</strong>же <strong>о</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении <strong>о</strong>т Anthracomarti<br />

<strong>к</strong>лещей Ixodides, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е н<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нении<br />

<strong>с</strong>тр<strong>о</strong>ения <strong>с</strong>пинных щит<strong>о</strong><strong>в</strong>, <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong>. В<strong>с</strong>е<br />

Soluta, <strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле и Anthracomarti, имеют<br />

н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящую пр<strong>о</strong><strong>с</strong><strong>о</strong>му, т<strong>о</strong>гд<strong>а</strong> <strong>к</strong><strong>а</strong><strong>к</strong> <strong>в</strong><strong>с</strong>е Acarina, <strong>а</strong><br />

<strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, и Ixodides имеют <strong>к</strong><strong>о</strong>жную пр<strong>о</strong><strong>с</strong><strong>о</strong>му,<br />

<strong>в</strong> ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>ние <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й не <strong>в</strong>х<strong>о</strong>дят <strong>а</strong><strong>к</strong>р<strong>о</strong>н<br />

и д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых р<strong>о</strong>т<strong>о</strong><strong>в</strong>ых <strong>с</strong>егмент<strong>а</strong>, <strong>о</strong>бр<strong>а</strong>зующих<br />

у <strong>к</strong>лещей <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленную гн<strong>а</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му. Т<strong>а</strong><strong>к</strong>им<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м, г<strong>о</strong>л<strong>о</strong><strong>в</strong>ные щиты Anthracomarti и Ixodides<br />

не я<strong>в</strong>ляют<strong>с</strong>я г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичными <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ниями.<br />

П<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Soluta р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енен п<strong>а</strong>у<strong>к</strong><strong>а</strong>м<br />

п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong> Araneae, <strong>с</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым имеет <strong>о</strong>бщее<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждение. Отряд Trigonotarbi б<strong>о</strong>лее примити<strong>в</strong>ен,<br />

a Anthracomarti <strong>с</strong> их <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ыми р<strong>а</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ниями<br />

тергит<strong>о</strong><strong>в</strong> б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>н.<br />

Д<strong>в</strong><strong>а</strong> <strong>о</strong>тряд<strong>а</strong>: Trigonotarbi и Anthracomarti.<br />

485<br />

http://jurassic.ru/

Тел<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>му (г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь)<br />

и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му (брюш<strong>к</strong><strong>о</strong>), <strong>с</strong><strong>о</strong>члененные п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong><strong>с</strong>егд<strong>а</strong> меньше брюш<strong>к</strong><strong>а</strong>; <strong>о</strong>н<strong>а</strong><br />

имеет <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>зличную ф<strong>о</strong>рму—<strong>о</strong>т п<strong>о</strong>луш<strong>а</strong>р<strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

(ри<strong>с</strong>. 1386) д<strong>о</strong> треуг<strong>о</strong>льн<strong>о</strong>й или <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й<br />

(ри<strong>с</strong>. 1395). П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>чень<br />

пл<strong>о</strong>тны, <strong>о</strong>бр<strong>а</strong>зуют <strong>с</strong>пл<strong>о</strong>шн<strong>о</strong>й п<strong>а</strong>нцирь (ри<strong>с</strong>.<br />

1383Б). К<strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>с</strong>рединн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong> <strong>о</strong>т б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых ч<strong>а</strong><strong>с</strong>тей, <strong>а</strong> у<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong> б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми <strong>о</strong>тмечены гр<strong>а</strong>ницы<br />

<strong>с</strong>лияния пер<strong>в</strong>ичных пр<strong>о</strong>,- мет<strong>а</strong>- и мез<strong>о</strong>пельтидия<br />

(ри<strong>с</strong>. 1387). Обычн<strong>о</strong> имеют<strong>с</strong>я д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>а</strong>,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупных (ри<strong>с</strong>. 1385, 1401). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

р<strong>а</strong><strong>с</strong>членен<strong>а</strong> н<strong>а</strong> 11 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>: т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Palaeocharinidae имеют<strong>с</strong>я рудименты<br />

еще д<strong>в</strong>ух (XII и XIII) <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>в</strong> <strong>в</strong>иде «х<strong>в</strong><strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>идн<strong>о</strong>г<strong>о</strong>» прид<strong>а</strong>т<strong>к</strong><strong>а</strong> п<strong>о</strong>з<strong>а</strong>ди<br />

з<strong>а</strong>днепр<strong>о</strong>х<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия (ри<strong>с</strong>. 1383А).<br />

С д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны з<strong>а</strong>метны лишь <strong>в</strong><strong>о</strong><strong>с</strong>емь<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Тергиты р<strong>а</strong><strong>с</strong>членены д<strong>в</strong>умя пр<strong>о</strong>д<strong>о</strong>льными<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми н<strong>а</strong> <strong>с</strong>рединную и <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ые<br />

ч<strong>а</strong><strong>с</strong>ти; з<strong>а</strong>днеб<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые углы тергит<strong>о</strong><strong>в</strong> ин<strong>о</strong>гд<strong>а</strong><br />

<strong>о</strong>бр<strong>а</strong>зуют п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям или н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong><br />

шил<strong>о</strong><strong>в</strong>идные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты (ри<strong>с</strong>. 1390, 1403). Хелицеры<br />

трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идные. Стр<strong>о</strong>ение<br />

их, при н<strong>а</strong>личии фильтр<strong>а</strong>ци<strong>о</strong>нн<strong>о</strong>г<strong>о</strong> <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т<strong>а</strong><br />

н<strong>а</strong> т<strong>а</strong>зи<strong>к</strong><strong>а</strong>х педип<strong>а</strong>льп и н<strong>о</strong>г I (ри<strong>с</strong>. 1383Б),<br />

п<strong>о</strong>з<strong>в</strong><strong>о</strong>ляет п<strong>о</strong> <strong>а</strong>н<strong>а</strong>л<strong>о</strong>гии <strong>с</strong> п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми г<strong>о</strong><strong>в</strong><strong>о</strong>рить <strong>о</strong><br />

<strong>с</strong>уще<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>нии у Trigonotarbi н<strong>а</strong>ружн<strong>о</strong>г<strong>о</strong> пище<strong>в</strong><strong>а</strong>рения.<br />

Среднюю ч<strong>а</strong><strong>с</strong>ть груди з<strong>а</strong>ним<strong>а</strong>ет<br />

<strong>с</strong>тернум, п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

п<strong>о</strong>д<strong>в</strong>ижные т<strong>а</strong>зи<strong>к</strong>и х<strong>о</strong>дильных н<strong>о</strong>г (ри<strong>с</strong>.<br />

1383А, 1392Б). Н<strong>о</strong>ги умеренн<strong>о</strong>й длины, <strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые.<br />

Не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые члени<strong>к</strong>и их н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не имеют зубцы (ри<strong>с</strong>. 1383А,<br />

1386). Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> н<strong>а</strong> VIII <strong>с</strong>терните тел<strong>а</strong><br />

(<strong>в</strong>т<strong>о</strong>р<strong>о</strong>м брюшн<strong>о</strong>м!); <strong>о</strong>н<strong>о</strong> имеет <strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й<br />

щели <strong>с</strong> д<strong>в</strong>умя <strong>к</strong>рышеч<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1392Б,<br />

1402Б). Дых<strong>а</strong>тельн<strong>а</strong>я <strong>с</strong>и<strong>с</strong>тем<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong><br />

тремя п<strong>а</strong>р<strong>а</strong>ми лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>с</strong>тернит<strong>о</strong><strong>в</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong>, третьег<strong>о</strong><br />

и чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

Де<strong>в</strong><strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы и <strong>к</strong><strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>генния<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Д<strong>о</strong><br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ний Петрун<strong>к</strong>е<strong>в</strong>ич<strong>а</strong> (Petrunkevitch,<br />

1949) <strong>о</strong>тряд Trigonotarbi <strong>о</strong>бъединял<strong>с</strong>я <strong>с</strong> Anthracomarti.<br />

Отмеч<strong>а</strong>л<strong>о</strong><strong>с</strong>ь р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> ег<strong>о</strong>, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, <strong>с</strong> п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми-<strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>ц<strong>а</strong>ми <strong>о</strong>тряд<strong>а</strong> Opiliones,<br />

<strong>а</strong> <strong>с</strong> друг<strong>о</strong>й — <strong>с</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящими п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми<br />

<strong>о</strong>тряд<strong>а</strong> Аг<strong>а</strong>пе<strong>а</strong>е. П<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>е пр<strong>о</strong>межут<strong>о</strong>чн<strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение<br />

пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong> <strong>о</strong>тр<strong>а</strong>ж<strong>а</strong>ет дей<strong>с</strong>т<strong>в</strong>ительн<strong>о</strong>е<br />

фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> Trigonotarbi. Они<br />

пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят <strong>о</strong>т <strong>о</strong>бщих <strong>с</strong> п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми и <strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>ц<strong>а</strong>ми<br />

ОТРЯД TRIGONOTARBI<br />

486<br />

http://jurassic.ru/<br />

<strong>а</strong>мблипиг<strong>о</strong>п<strong>о</strong>д<strong>о</strong>бных хелицер<strong>о</strong><strong>в</strong>ых, <strong>с</strong><strong>о</strong>чет<strong>а</strong>я <strong>в</strong><br />

<strong>с</strong>ебе призн<strong>а</strong><strong>к</strong>и <strong>о</strong>беих этих групп, <strong>а</strong> т<strong>а</strong><strong>к</strong>же не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

призн<strong>а</strong><strong>к</strong>и примити<strong>в</strong>ных Scorpionomorpha<br />

(Amblypygi, Ricinulei): <strong>с</strong><strong>о</strong>хр<strong>а</strong>нение рудимент<strong>а</strong>рн<strong>о</strong>й<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>мы, р<strong>а</strong>зделение тергит<strong>о</strong><strong>в</strong><br />

пр<strong>о</strong>д<strong>о</strong>льными б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми н<strong>а</strong> три ч<strong>а</strong><strong>с</strong>ти и т. п.<br />

Пять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Palaeocharinidae, Anthracosironidae,<br />

Eophrynidae, Trigonotarbidae, Trigonomartidae.<br />

СЕМЕЙСТВО PALAEOCHARINIDAE HIRST, 1923<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> из 13 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

де<strong>в</strong>ять пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми тергит<strong>а</strong>ми<br />

и <strong>с</strong>тернит<strong>а</strong>ми. Тергиты X и XI <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> з<strong>а</strong>гнуты н<strong>а</strong> брюшную<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну; тергит <strong>о</strong>динн<strong>а</strong>дц<strong>а</strong>т<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> ф<strong>о</strong>рмирует<br />

<strong>а</strong>н<strong>а</strong>льный <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н. П<strong>о</strong>з<strong>а</strong>ди нег<strong>о</strong> д<strong>в</strong><strong>а</strong><br />

рудимент<strong>а</strong>рных трубч<strong>а</strong>тых <strong>с</strong>егмент<strong>а</strong>, г<strong>о</strong>м<strong>о</strong>л<strong>о</strong>гичных<br />

мет<strong>а</strong><strong>с</strong><strong>о</strong>ме Amblypygi и других Scorpionomorpha<br />

(ри<strong>с</strong>. 1383). Де<strong>в</strong><strong>о</strong>н. Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Palaeocharinoides Hirst, 1923. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. hornet Hirst, 1923; де<strong>в</strong><strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>, Ш<strong>о</strong>тл<strong>а</strong>ндия.<br />

Стернум «з<strong>в</strong>ездч<strong>а</strong>тый» <strong>с</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емью з<strong>а</strong><strong>о</strong><strong>с</strong>тренными<br />

угл<strong>а</strong>ми; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ыпу<strong>к</strong>лый.<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди р<strong>о</strong><strong>в</strong>н<strong>а</strong>я, пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я<br />

(ри<strong>с</strong>. 1383А, Б). Длин<strong>а</strong> тел<strong>а</strong> 2,2 мм.<br />

Один <strong>в</strong>ид. Де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Palaeocharinus Hirst, 1923. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. rhyniensis Hirst, 1923; де<strong>в</strong><strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>,<br />

Ш<strong>о</strong>тл<strong>а</strong>ндия. Стернум <strong>о</strong><strong>в</strong><strong>а</strong>льный <strong>с</strong> з<strong>а</strong><strong>к</strong>ругленными<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими угл<strong>а</strong>ми; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й<br />

щит<strong>а</strong> прям<strong>о</strong>й. Имеют<strong>с</strong>я д<strong>в</strong><strong>а</strong> <strong>к</strong>рупных гл<strong>а</strong>з<strong>а</strong><br />

(ри<strong>с</strong>. 1384, 1385). Длин<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы 1,5—2,0 мм,<br />

длин<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы 2 мм. Четыре <strong>в</strong>ид<strong>а</strong>. Де<strong>в</strong><strong>о</strong>н<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО ANTHRACOSIRONIDAE РОСОСК, 1903<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>с</strong>яти <strong>в</strong>идимых <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

(пер<strong>в</strong>ый тергит редуцир<strong>о</strong><strong>в</strong><strong>а</strong>н). Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

п<strong>о</strong>луш<strong>а</strong>р<strong>о</strong><strong>в</strong>идн<strong>а</strong>я, шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен<strong>а</strong> <strong>с</strong><br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> и <strong>с</strong>переди. Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы тел<strong>а</strong> <strong>с</strong> рез<strong>к</strong><strong>о</strong>й ямч<strong>а</strong>т<strong>о</strong>й<br />

<strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й. Н<strong>о</strong>ги длинные; бедр<strong>о</strong>, <strong>к</strong><strong>о</strong>лен<strong>а</strong> и<br />

г<strong>о</strong>лени пер<strong>в</strong>ых д<strong>в</strong>ух п<strong>а</strong>р н<strong>о</strong>г <strong>с</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны буг<strong>о</strong>рч<strong>а</strong>тые (ри<strong>с</strong>. 1386). К<strong>а</strong>рб<strong>о</strong>н<br />

3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д Anthracosiro Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>,<br />

1903.<br />

СЕМЕЙСТВО EOPHRYNIDAE KARSCH, 1882<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>в</strong>яти <strong>в</strong>идимых <strong>с</strong><strong>в</strong>ерху<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>; IX тергит з<strong>а</strong><strong>к</strong>лючен между пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми<br />

<strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>г<strong>о</strong> тергит<strong>а</strong>. Ди<strong>с</strong>т<strong>а</strong>льные <strong>с</strong>егменты<br />

редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны и <strong>с</strong>мещены н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льную <strong>с</strong>т<strong>о</strong>-

1390<br />

1387. Kreischeria wiedel Geinitz; <strong>о</strong>бщий <strong>в</strong>ид, X3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Petrunkevitch, 1953). 1388. Stenotrogulus salmii (Stur); <strong>о</strong>бщий<br />

<strong>в</strong>ид, x 2,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1389.<br />

Cyclotrogulus sturmii (Haase); <strong>о</strong>бщий <strong>в</strong>ид, x 2,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Petrunkevitch, 1953). 1390. Petrovicia proditoria Fritsch; <strong>о</strong>бщий<br />

<strong>в</strong>ид, x2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1391. Hemiphrynus<br />

longipes Fritsch; <strong>о</strong>бщий <strong>в</strong>ид, x,2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1953). 1392. Eophrynus prestvici (Buchland); A — <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу, х 2,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>,<br />

1392Б<br />

Ри<strong>с</strong>. 1387—1397. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eophrynidae<br />

1911). 1393. Acrokreischeria verrucosa Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху •<br />

Х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911). 1394. Hemikreischeria<br />

gelnitzi Thevenin; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>низу, Х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1953). 1395. Pleophrynus ensifer Petrunkevitch;<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х 3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch,<br />

1949). 1396. Pocononia whltei (Ewing); <strong>о</strong>бщий <strong>в</strong>ид, X 2,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н.<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1949). 1397. Areomartus ovatus<br />

Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong><br />

(Petrunkevitch, 1913)<br />

487<br />

http://jurassic.ru/

р<strong>о</strong>ну, <strong>о</strong>бр<strong>а</strong>зуя <strong>а</strong>н<strong>а</strong>льный <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н (ри<strong>с</strong>. 1392).<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий щит треуг<strong>о</strong>льный, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>с</strong> уз<strong>к</strong>им р<strong>о</strong><strong>с</strong>трум<strong>о</strong>м и <strong>в</strong><strong>о</strong>гнутым з<strong>а</strong>дним<br />

<strong>к</strong>р<strong>а</strong>ем. Гл<strong>а</strong>з<strong>а</strong> имеют<strong>с</strong>я или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Четыре п<strong>а</strong>ры лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных<br />

н<strong>а</strong> II—V <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мэтиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тернит<strong>а</strong>х.<br />

К<strong>а</strong>рб<strong>о</strong>н С. Амери<strong>к</strong>и и Е<strong>в</strong>р<strong>о</strong>пы. 14 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Kfeischeria Geinitz, 1882. Тип р<strong>о</strong>д<strong>а</strong> —<br />

К- wiedei Geinitz, 1882; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Герм<strong>а</strong>ния.<br />

П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы <strong>в</strong><strong>с</strong>ег<strong>о</strong> тел<strong>а</strong> буг<strong>о</strong>рч<strong>а</strong>тые. Срединн<strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> рез<strong>к</strong><strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>блен<strong>а</strong>;<br />

з<strong>а</strong>дняя ч<strong>а</strong><strong>с</strong>ть щит<strong>а</strong> <strong>о</strong>тгр<strong>а</strong>ничен<strong>а</strong> п<strong>о</strong>перечн<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й<br />

(ри<strong>с</strong>. 1387); <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>й тергит <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>с</strong>лит <strong>с</strong> де<strong>в</strong>ятым; з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й тел<strong>а</strong> гл<strong>а</strong>д<strong>к</strong>ий, без<br />

шил<strong>о</strong><strong>в</strong>идных <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>. Длин<strong>а</strong> тел<strong>а</strong> 50 мм, длин<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы 15 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Stenotrogulus Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Eophrynus salmii Stur, 1887; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

З<strong>а</strong>дний <strong>к</strong><strong>о</strong>нец тел<strong>а</strong> <strong>с</strong> четырьмя м<strong>а</strong>лень<strong>к</strong>ими<br />

з<strong>а</strong><strong>о</strong><strong>с</strong>трениями; тергиты VIII и IX <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>льных<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong>литы, тергиты <strong>с</strong>рединн<strong>о</strong>й<br />

з<strong>о</strong>ны <strong>с</strong> д<strong>в</strong>умя пр<strong>о</strong>д<strong>о</strong>льными <strong>в</strong>д<strong>а</strong><strong>в</strong>лениями<br />

и четырьмя ряд<strong>а</strong>ми из <strong>к</strong>рупных и б<strong>о</strong>лее мел<strong>к</strong>их<br />

буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1388); <strong>в</strong> <strong>с</strong>редней ч<strong>а</strong><strong>с</strong>ти<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> щит<strong>а</strong> имеет<strong>с</strong>я групп<strong>а</strong> из<br />

д<strong>в</strong>ух <strong>к</strong>рупных з<strong>а</strong>дних и трех б<strong>о</strong>лее мел<strong>к</strong>их<br />

<strong>о</strong><strong>в</strong><strong>а</strong>льных передних буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>; ф<strong>о</strong>рм<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

треуг<strong>о</strong>льн<strong>а</strong>я, <strong>с</strong> удлиненн<strong>о</strong>й р<strong>о</strong><strong>с</strong>тр<strong>а</strong>льн<strong>о</strong>й ч<strong>а</strong><strong>с</strong>тью.<br />

Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Длин<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

10 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Cyclotrogulus Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Eophrynus sturii Haase, 1890; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Тел<strong>о</strong> груше<strong>в</strong>идн<strong>о</strong>й ф<strong>о</strong>рмы <strong>с</strong> неб<strong>о</strong>льш<strong>о</strong>й<br />

треуг<strong>о</strong>льн<strong>о</strong>й, рез<strong>к</strong><strong>о</strong> з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>й <strong>с</strong>переди пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й<br />

и п<strong>о</strong>чти ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>й <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й, шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>й н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце; тергиты<br />

<strong>с</strong>рединн<strong>о</strong>й з<strong>о</strong>ны <strong>с</strong> д<strong>в</strong>умя пр<strong>о</strong>д<strong>о</strong>льными <strong>в</strong>д<strong>а</strong><strong>в</strong>лениями;<br />

<strong>в</strong>д<strong>о</strong>ль этих <strong>в</strong>д<strong>а</strong><strong>в</strong>лений и б<strong>о</strong>р<strong>о</strong>зд н<strong>а</strong> тергит<strong>а</strong>х<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я ше<strong>с</strong>ть ряд<strong>о</strong><strong>в</strong> буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Тергит IX <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> без<br />

буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1389). Длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 6,5 мм,<br />

длин<strong>а</strong> брюш<strong>к</strong><strong>а</strong> 14 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Petrovicia Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. proditoria Fritsch, 1904; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Тел<strong>о</strong> яйце<strong>в</strong>идн<strong>о</strong>е; н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце <strong>в</strong> <strong>о</strong>дин<br />

ряд р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены четыре <strong>к</strong>рупных шип<strong>а</strong>; тергиты<br />

<strong>с</strong> четырьмя пр<strong>о</strong>д<strong>о</strong>льными ряд<strong>а</strong>ми буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> треуг<strong>о</strong>льн<strong>а</strong>я <strong>с</strong> <strong>в</strong>ытянутым <strong>о</strong><strong>с</strong>трым<br />

р<strong>о</strong><strong>с</strong>трум<strong>о</strong>м. Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют (ри<strong>с</strong>. 1390).<br />

Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Hemiphrynus Fritsch, 1899. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Н. longipes Fritsch, 1899; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Тел<strong>о</strong> яйце<strong>в</strong>идн<strong>о</strong>е, зн<strong>а</strong>чительн<strong>о</strong> р<strong>а</strong><strong>с</strong>ширенн<strong>о</strong>е<br />

<strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине, <strong>с</strong> четырьмя неб<strong>о</strong>льшими<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ми н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>ю; тергиты без<br />

488<br />

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пр<strong>о</strong>д<strong>о</strong>льных <strong>в</strong>д<strong>а</strong><strong>в</strong>лений и буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

треуг<strong>о</strong>льн<strong>а</strong>я <strong>с</strong> длинным уз<strong>к</strong>им р<strong>о</strong><strong>с</strong>трум<strong>о</strong>м. Гл<strong>а</strong>з<strong>а</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Н<strong>о</strong>ги длинные и т<strong>о</strong>н<strong>к</strong>ие<br />

(ри<strong>с</strong>. 1391). Длин<strong>а</strong> тел<strong>а</strong> 28 мм. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Vratislavia Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Architarbus silesiacus Roemer, 1878; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

П<strong>о</strong>льш<strong>а</strong>. Тергиты <strong>с</strong>рединн<strong>о</strong>й з<strong>о</strong>ны не р<strong>а</strong>зделены<br />

пр<strong>о</strong>д<strong>о</strong>льными <strong>в</strong>д<strong>а</strong><strong>в</strong>лениями, без буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>;<br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong> четыре треуг<strong>о</strong>льных<br />

<strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>; тергиты VIII и IX <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> р<strong>а</strong>зделены б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й; н<strong>о</strong>ги<br />

<strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> т<strong>о</strong>л<strong>с</strong>тые и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; <strong>с</strong>тр<strong>о</strong>ение<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы: Eophrynus<br />

Woodward, 1871 (ри<strong>с</strong>. 1392); Acrokreischeria<br />

Petrunkevitch, 1953 (ри<strong>с</strong>. 1393); Pseudokreischeria<br />

Petrunkevitch, 1953; Anzinia Petrunkevitch,<br />

1953; Hemikreischeria Fritsch, 1904<br />

(ри<strong>с</strong>. 1394), <strong>а</strong> т<strong>а</strong><strong>к</strong>же из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> С. Амери<strong>к</strong>и —<br />

Pleophrynus Petrunkevitch, 1945 (ри<strong>с</strong>. 1395);<br />

Pocononia Petrunkevitch, 1953 (ри<strong>с</strong>. 1396);<br />

Areomartus Petrunkevitch, 1913 (ри<strong>с</strong>. 1397).<br />

СЕМЕЙСТВО TRIGONOTARBIDAE<br />

PETRUNKEVITCH, 1949<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>в</strong><strong>о</strong><strong>с</strong>ьми (<strong>в</strong>идимых<br />

<strong>с</strong><strong>в</strong>ерху) тергит<strong>о</strong><strong>в</strong>. Кр<strong>а</strong>е<strong>в</strong>ые з<strong>о</strong>ны тергит<strong>о</strong><strong>в</strong> имеют<strong>с</strong>я<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> н<strong>а</strong> <strong>с</strong>еми <strong>с</strong>егмент<strong>а</strong>х: VIII тергит<br />

<strong>о</strong><strong>к</strong>ружен пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>ми VII тергит<strong>а</strong>. Н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не тел<strong>а</strong> <strong>с</strong>емь <strong>с</strong>тернит<strong>о</strong><strong>в</strong> и д<strong>в</strong><strong>а</strong><br />

тергит<strong>а</strong> переме<strong>с</strong>ти<strong>в</strong>ших<strong>с</strong>я <strong>с</strong>юд<strong>а</strong> <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Четыре<br />

з<strong>а</strong>дних <strong>с</strong>тернит<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>лу<strong>к</strong>руг<strong>а</strong>ми<br />

<strong>в</strong>переди з<strong>а</strong>днепр<strong>о</strong>х<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный<br />

щит треуг<strong>о</strong>льный. Д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>а</strong>. Н<strong>а</strong><br />

<strong>в</strong>т<strong>о</strong>р<strong>о</strong>м и третьем <strong>с</strong>тернит<strong>а</strong>х (м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и<br />

третьем и чет<strong>в</strong>ерт<strong>о</strong>м!) имеют<strong>с</strong>я лег<strong>о</strong>чные меш<strong>к</strong>и.<br />

Длин<strong>а</strong> тел<strong>а</strong> 4,7 мм (ри<strong>с</strong>. 1398А, Б). К<strong>а</strong>рб<strong>о</strong>н.<br />

Один р<strong>о</strong>д Trigonotarbus Pocock, 1911 из<br />

3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

СЕМЕЙСТВО TRIGONOMARTIDAE<br />

PETRUNKEVITCH, 1949<br />

Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>в</strong><strong>о</strong><strong>с</strong>ьми <strong>в</strong>идимых<br />

<strong>с</strong><strong>в</strong>ерху <strong>с</strong>егмент<strong>о</strong><strong>в</strong>; <strong>в</strong><strong>о</strong><strong>с</strong>ьм<strong>о</strong>й тергит <strong>с</strong> треуг<strong>о</strong>льными<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong>ыми уч<strong>а</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми. Меж<strong>с</strong>егментные<br />

гр<strong>а</strong>ницы тергит<strong>о</strong><strong>в</strong> прямые, п<strong>а</strong>р<strong>а</strong>ллельны друг<br />

другу. Пр<strong>о</strong>д<strong>о</strong>льные б<strong>о</strong>р<strong>о</strong>зды н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

брюш<strong>к</strong><strong>а</strong> <strong>с</strong>уж<strong>а</strong>ют<strong>с</strong>я <strong>к</strong> з<strong>а</strong>днему <strong>к</strong><strong>о</strong>нцу тел<strong>а</strong>.<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong><strong>с</strong><strong>о</strong>мы и тергит<strong>о</strong><strong>в</strong> <strong>с</strong>етч<strong>а</strong>т<strong>а</strong>я<br />

(ри<strong>с</strong>. 1399 Б) или буг<strong>о</strong>рч<strong>а</strong>т<strong>а</strong>я (ри<strong>с</strong>. 1401). Гл<strong>а</strong>з<strong>а</strong><br />

имеют<strong>с</strong>я. К<strong>а</strong>рб<strong>о</strong>н. Ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Trigonomartus Petrunkevitch, 1913.<br />

Тип р<strong>о</strong>д<strong>а</strong> — Anthracomartus pustulatus Scudder,<br />

1884; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> и 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> уз<strong>к</strong><strong>а</strong>я, треуг<strong>о</strong>льн<strong>а</strong>я, з<strong>а</strong><strong>к</strong>ругленн<strong>а</strong>я н<strong>а</strong>

Ри<strong>с</strong>. 1398—1404. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Tri]<br />

1398. Trigonotarbus johnsoni Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong><strong>в</strong>ерху,<br />

Б — <strong>с</strong>низу, Х7,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1910). 1399. Trigonomartus<br />

pustulatus (Scudder); <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong>низу, Б —<br />

<strong>с</strong><strong>в</strong>ерху, Х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1913). 1400.<br />

Planomartus krejcii Kusta; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, Х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1913). 1401. Aphantomartus areolatus<br />

Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, X 6,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>,<br />

<strong>в</strong>ершине; п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тел<strong>а</strong> <strong>с</strong><strong>в</strong>ерху <strong>с</strong> <strong>с</strong>етч<strong>а</strong>тым<br />

<strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> р<strong>а</strong>зделен<strong>а</strong> п<strong>о</strong>перечн<strong>о</strong>й<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й н<strong>а</strong> д<strong>в</strong>е ч<strong>а</strong><strong>с</strong>ти; гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; т<strong>а</strong>зи<strong>к</strong>и<br />

н<strong>о</strong>г. IV зн<strong>а</strong>чительн<strong>о</strong> <strong>к</strong>рупнее т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных н<strong>о</strong>г (ри<strong>с</strong>. 1399А, Б). Длин<strong>а</strong> тел<strong>а</strong><br />

16—18 мм. Четыре <strong>в</strong>ид<strong>а</strong>. Ср. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

и С. Амери<strong>к</strong>и.<br />

Planomartus Petrunkevitch, 1953. Тип<br />

р<strong>о</strong>д<strong>а</strong>—Anthracomartus krejcii Kusta, 1883;<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>к</strong>рупн<strong>а</strong>я, треуг<strong>о</strong>льн<strong>а</strong>я;<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, яйце<strong>в</strong>идн<strong>а</strong>я,<br />

;onotarbidae и Trigonomartidae<br />

1911). 1402. Elaverimartus pococki (Petrunkevitch); <strong>о</strong>бщий <strong>в</strong>ид:<br />

А — <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу, Х6,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>,<br />

1911). 1403. Phrynomartus waechteri Guthorl; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху,<br />

Х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1404. Lissomartus<br />

carbonarius (Petrunkevitch); <strong>о</strong>бщий <strong>в</strong>ид, <strong>с</strong>низу, х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

С. Амери<strong>к</strong><strong>а</strong> (Petrunkevitch, 1949)<br />

з<strong>а</strong><strong>к</strong>ругленн<strong>а</strong>я н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце; тергит VIII<br />

<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>к</strong><strong>в</strong><strong>а</strong>др<strong>а</strong>тн<strong>о</strong>й ф<strong>о</strong>рмы; <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ые ч<strong>а</strong><strong>с</strong>ти<br />

тергит<strong>о</strong><strong>в</strong> н<strong>а</strong><strong>к</strong>л<strong>о</strong>нены <strong>к</strong>з<strong>а</strong>ди (ри<strong>с</strong>. 1400).<br />

Длин<strong>а</strong> брюш<strong>к</strong><strong>а</strong> 11,5 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н<br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы — Aphantomartus<br />

Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911 (ри<strong>с</strong>. 1401); Elaverimartus<br />

Petrunkevitch, 1953 (ри<strong>с</strong>. 1402); Phrynomartus<br />

Petrunkevitch, 1945 (ри<strong>с</strong>. 1403); из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

С. Амери<strong>к</strong>и — Lissomartus Petrunkevitch, 1949<br />

(ри<strong>с</strong>. 1404).<br />

489<br />

http://jurassic.ru/

ОТРЯД ANT<br />

Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> у б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>о</strong><strong>к</strong>ругл<strong>а</strong>я или<br />

треуг<strong>о</strong>льн<strong>а</strong>я (ри<strong>с</strong>. 1405 А, 1410); <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

<strong>с</strong>пинн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong><strong>с</strong><strong>о</strong>мы без рез<strong>к</strong><strong>о</strong> <strong>в</strong>ыделяющей<strong>с</strong>я<br />

<strong>с</strong>рединн<strong>о</strong>й з<strong>о</strong>ны 1<br />

, р<strong>о</strong><strong>в</strong>н<strong>а</strong>я или <strong>с</strong>л<strong>а</strong>б<strong>о</strong><strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я,<br />

<strong>с</strong> т<strong>о</strong>чечн<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й (ри<strong>с</strong>.<br />

1413 А). Гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong> (Maiocercus Pocock) <strong>с</strong><strong>к</strong>ульптур<strong>а</strong><br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ет т<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong>ую у Trigonotarbi<br />

(ри<strong>с</strong>. 1408А). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong> <strong>с</strong><strong>о</strong>членен<strong>а</strong><br />

<strong>с</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й; ме<strong>с</strong>т<strong>о</strong> <strong>с</strong><strong>о</strong>членения шир<strong>о</strong><strong>к</strong><strong>о</strong>е.<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из де<strong>с</strong>яти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

д<strong>в</strong><strong>а</strong> з<strong>а</strong>дних з<strong>а</strong>гнуты н<strong>а</strong> брюшную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну,<br />

и тергиты их ф<strong>о</strong>рмируют п<strong>о</strong>д<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong>идный <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong><strong>а</strong>н<strong>а</strong>льны'й<br />

<strong>с</strong>егмент и <strong>а</strong>н<strong>а</strong>льный <strong>к</strong>л<strong>а</strong>п<strong>а</strong>н (ри<strong>с</strong>.<br />

1409 Б). Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не брюш<strong>к</strong><strong>а</strong> тергиты<br />

р<strong>а</strong>зделены пр<strong>о</strong>д<strong>о</strong>льными линиями н<strong>а</strong> <strong>с</strong>рединную<br />

и д<strong>в</strong>е <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ые з<strong>о</strong>ны, <strong>к</strong><strong>а</strong><strong>к</strong> у Trigonomarti.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>к</strong>р<strong>а</strong>я тергит<strong>о</strong><strong>в</strong> р<strong>а</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны<br />

и ф<strong>о</strong>рмируют шир<strong>о</strong><strong>к</strong>ую <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ую з<strong>о</strong>ну,<br />

зн<strong>а</strong>чительн<strong>о</strong> у<strong>в</strong>еличи<strong>в</strong><strong>а</strong>ющую пл<strong>о</strong>щ<strong>а</strong>дь тел<strong>а</strong><br />

(ри<strong>с</strong>. 1408, 1413); <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых р<strong>а</strong>зр<strong>а</strong>щений<br />

не н<strong>а</strong>блюд<strong>а</strong>ет<strong>с</strong>я у пер<strong>в</strong><strong>о</strong>г<strong>о</strong> тергит<strong>а</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женн<strong>о</strong>г<strong>о</strong><br />

н<strong>а</strong> гр<strong>а</strong>нице <strong>с</strong><strong>о</strong>членения пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы. Брюшн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

р<strong>а</strong><strong>с</strong>членен<strong>а</strong> н<strong>а</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емь — де<strong>в</strong>ять <strong>с</strong>тернит<strong>о</strong><strong>в</strong>,<br />

з<strong>а</strong>дние из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых дуг<strong>о</strong><strong>в</strong>идн<strong>о</strong> из<strong>о</strong>гнуты и<br />

<strong>о</strong><strong>к</strong>руж<strong>а</strong>ют <strong>с</strong>переди <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие (ри<strong>с</strong>.<br />

1408, 1413 Б). Стернит пер<strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> м<strong>а</strong>лень<strong>к</strong>ий, треуг<strong>о</strong>льный,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми н<strong>о</strong>г IV и п<strong>о</strong>з<strong>а</strong>ди<br />

них. Брюшн<strong>а</strong>я п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

б<strong>о</strong>лее <strong>в</strong>ыпу<strong>к</strong>л<strong>а</strong>я, чем <strong>с</strong>пинн<strong>а</strong>я (ри<strong>с</strong>. 1413 В).<br />

П<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тергит<strong>о</strong><strong>в</strong> <strong>с</strong> т<strong>о</strong>чечн<strong>о</strong>й пун<strong>к</strong>тир<strong>о</strong><strong>в</strong><strong>к</strong><strong>о</strong>й,<br />

<strong>о</strong>чень ред<strong>к</strong><strong>о</strong> <strong>с</strong> буг<strong>о</strong>р<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1407) или<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>о</strong>жным <strong>о</strong>рн<strong>а</strong>мент<strong>о</strong>м (ри<strong>с</strong>. 1413А).<br />

Хелицеры трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идные;<br />

из<strong>о</strong>гнуты н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну. Педип<strong>а</strong>льпы 1<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, пр<strong>о</strong><strong>с</strong>тые (ри<strong>с</strong>.<br />

1413); их п<strong>о</strong>д<strong>в</strong>ижные т<strong>а</strong>зи<strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong>переди<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г (ри<strong>с</strong>. 1413 Б). Т<strong>а</strong>зи<strong>к</strong>и <strong>в</strong><strong>с</strong>ех<br />

н<strong>о</strong>г п<strong>о</strong>д<strong>в</strong>ижные, уз<strong>к</strong>ие, треуг<strong>о</strong>льн<strong>о</strong>-цилиндриче<strong>с</strong><strong>к</strong>ие,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ю <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны пр<strong>о</strong><strong>с</strong><strong>о</strong>мы (ри<strong>с</strong>. 1413 Б). Стернум <strong>о</strong>бычн<strong>о</strong><br />

<strong>в</strong>ытянутый, реже <strong>о</strong><strong>к</strong>руглый. Х<strong>о</strong>дильные н<strong>о</strong>ги<br />

<strong>с</strong>емичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, <strong>с</strong> р<strong>а</strong><strong>с</strong>члененными <strong>в</strong>ертлуг<strong>а</strong>ми:<br />

т<strong>а</strong>зи<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые, <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идные, <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми.<br />

П<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong>, третьег<strong>о</strong> и чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тернит<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены три п<strong>а</strong>ры<br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>. П<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие н<strong>а</strong> II<br />

<strong>с</strong>егменте <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы имеет <strong>в</strong>ид п<strong>о</strong>перечн<strong>о</strong>й<br />

щели, при<strong>к</strong>рыт<strong>о</strong>й неп<strong>а</strong>рн<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>о</strong>й (ри<strong>с</strong>.<br />

1409). В п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>м <strong>а</strong>триуме <strong>с</strong><strong>а</strong>м<strong>к</strong>и имеют<strong>с</strong>я п<strong>а</strong>рные<br />

<strong>с</strong>еменные меш<strong>о</strong>ч<strong>к</strong>и, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ющие<strong>с</strong>я н<strong>а</strong><br />

третьем <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>м <strong>с</strong>терните.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и б и <strong>о</strong> г е <strong>о</strong> г р <strong>а</strong> ф и я. Н<strong>а</strong>земные<br />

хелицер<strong>о</strong><strong>в</strong>ые, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, хищни<strong>к</strong>и <strong>с</strong> н<strong>а</strong>ружным<br />

пище<strong>в</strong><strong>а</strong>рением, <strong>о</strong>бит<strong>а</strong>ли <strong>в</strong> п<strong>о</strong>д<strong>с</strong>тил<strong>к</strong>е ле<strong>с</strong><strong>о</strong><strong>в</strong><br />

н<strong>а</strong> р<strong>а</strong>змельченных <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>а</strong>х. Р<strong>а</strong><strong>с</strong>ширенн<strong>а</strong>я<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>а</strong>я з<strong>о</strong>н<strong>а</strong> тергит<strong>о</strong><strong>в</strong> зн<strong>а</strong>чительн<strong>о</strong> у<strong>в</strong>еличи­<br />

<strong>в</strong><strong>а</strong>л<strong>а</strong> удельную п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть тел<strong>а</strong> и препят<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong><br />

п<strong>о</strong>гружению жи<strong>в</strong><strong>о</strong>тных <strong>в</strong> <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т, <strong>а</strong> упл<strong>о</strong>щенн<strong>а</strong>я<br />

ф<strong>о</strong>рм<strong>а</strong> тел<strong>а</strong> <strong>о</strong>блегч<strong>а</strong>л<strong>а</strong> перед<strong>в</strong>ижение<br />

<strong>в</strong> т<strong>о</strong>лще <strong>с</strong>уб<strong>с</strong>тр<strong>а</strong>т<strong>а</strong> (ри<strong>с</strong>. 1413В). Anthracomarti<br />

были мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленны <strong>в</strong> эп<strong>о</strong>хи <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> —<br />

перми. Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>о</strong>ни, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ли<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременным п<strong>о</strong>ч<strong>в</strong>енным п<strong>а</strong>нцирным<br />

<strong>к</strong>лещ<strong>а</strong>м (Oribatei) и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым <strong>с</strong>ен<strong>о</strong><strong>к</strong><strong>о</strong><strong>с</strong>ц<strong>а</strong>м<br />

(Nemastomoididae). Н<strong>а</strong><strong>с</strong>еляли террит<strong>о</strong>рии<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременных Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и.<br />

П<strong>о</strong>лжение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>генияи<strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

Отряд<br />

Anthracomarti <strong>о</strong>чень близ<strong>о</strong><strong>к</strong> <strong>к</strong> <strong>о</strong>тряду Trigonotarbi,<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым длительн<strong>о</strong>е <strong>в</strong>ремя <strong>о</strong>бъединял<strong>с</strong>я<br />

<strong>в</strong> <strong>о</strong>дну группу. Antharcomarti пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>дят <strong>о</strong>т<br />

примити<strong>в</strong>ных Trigonotarbi (<strong>с</strong>м. <strong>с</strong>хему н<strong>а</strong> <strong>с</strong>тр.<br />

384), н<strong>о</strong> я<strong>в</strong>ляют<strong>с</strong>я б<strong>о</strong>лее <strong>с</strong>пеци<strong>а</strong>лизир<strong>о</strong><strong>в</strong><strong>а</strong>нными<br />

Soluta. Пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители Anthracomarti не<strong>о</strong>дн<strong>о</strong><strong>к</strong>р<strong>а</strong>тн<strong>о</strong><br />

и<strong>с</strong>п<strong>о</strong>льз<strong>о</strong><strong>в</strong><strong>а</strong>ли<strong>с</strong>ь для п<strong>о</strong><strong>с</strong>тр<strong>о</strong>ения фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их<br />

<strong>с</strong>хем Chelicerata. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Anthracomartidae.<br />

СЕМЕЙСТВО ANTHRACOMARTIDAE HAASE, 1890<br />

(Brachypygidae Pocock, 1911; Pleomartidae<br />

Petrunkevitch, 1945; Promygalidae Fritsch,<br />

1899; Coryphomartidae Petrunkevitch, 1945)<br />

Х<strong>а</strong>р<strong>а</strong><strong>к</strong>теризует<strong>с</strong>я призн<strong>а</strong><strong>к</strong><strong>а</strong>ми <strong>о</strong>тряд<strong>а</strong>.<br />

К<strong>а</strong>рб<strong>о</strong>н — пермь. Де<strong>с</strong>ять р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>ют<strong>с</strong>я <strong>с</strong> не<strong>в</strong>ыя<strong>с</strong>ненным <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>им<br />

п<strong>о</strong>л<strong>о</strong>жением.<br />

Brachylycosa Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Arthrolycosa carcinoides Fritsch, 1899; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> неб<strong>о</strong>льш<strong>а</strong>я,<br />

<strong>о</strong><strong>к</strong>ругл<strong>а</strong>я; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>а</strong>я, <strong>с</strong> р<strong>о</strong><strong>в</strong>ными<br />

<strong>к</strong>р<strong>а</strong>ями; <strong>в</strong>ерхняя п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ть пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я, передний <strong>к</strong>р<strong>а</strong>й ее шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен;<br />

ширин<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>ширения тергит<strong>о</strong><strong>в</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

ширине б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых ч<strong>а</strong><strong>с</strong>тей тергит<strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1405).<br />

Длин<strong>а</strong> тел<strong>а</strong> 12,0 — 20,3 мм. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

Promygale Fritsch, 1899. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Kreischeria bohemica Fritsch, 1899; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины;<br />

передние <strong>к</strong>р<strong>а</strong>я ее <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я п<strong>о</strong>д тупым<br />

угл<strong>о</strong>м; <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>а</strong>я, ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>а</strong>я,<br />

<strong>с</strong> р<strong>о</strong><strong>в</strong>ными <strong>к</strong>р<strong>а</strong>ями. Ширин<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>шире­<br />

ния меньше ширины 1<br />

490<br />

http://jurassic.ru/<br />

б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых ч<strong>а</strong><strong>с</strong>тей тергит<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1406А, Б). Длин<strong>а</strong> тел<strong>а</strong> 9,8—20,5 мм.<br />

Три <strong>в</strong>ид<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии.<br />

Oomartus Petrunkevitch, 1953. Тип<br />

р<strong>о</strong>д<strong>а</strong> — О. nyranensis Petrunkevitch, 1953; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Тел<strong>о</strong> яйце<strong>в</strong>идн<strong>о</strong>е, р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong><br />

з<strong>а</strong><strong>к</strong>ругленн<strong>о</strong>е н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

и н<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong>м з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

(ри<strong>с</strong>. 1411); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> п<strong>о</strong>чти п<strong>о</strong>луш<strong>а</strong>р<strong>о</strong><strong>в</strong>идн<strong>а</strong>я:<br />

длин<strong>а</strong> ее б<strong>о</strong>льше ширины, з<strong>а</strong>дний <strong>к</strong>р<strong>а</strong>й прям<strong>о</strong>й.<br />

Длин<strong>а</strong> тел<strong>а</strong> 17,5 мм. Один <strong>в</strong>ид.

405. Brachylycosa carcinoides (Fritsch); <strong>о</strong>бщий <strong>в</strong>ид: A — <strong>с</strong><strong>в</strong>ерху,<br />

Б — <strong>с</strong>низу; х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1406.<br />

Promygale bohemica (Fritsch); <strong>о</strong>бщий<strong>в</strong>ид: A— <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу,<br />

Х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1407. Brachypyge<br />

carbonis Woodward; брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, х5,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911). 1408. Maiocercus carbonis<br />

Ри<strong>с</strong>. 1405—1410. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Anthracomartidae<br />

(Howard et Thomas); <strong>о</strong>бщий <strong>в</strong>ид: A — <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу, х 4.0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1409. Cleptomartus<br />

plantus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу,<br />

Х8,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1410. Coryphomartus<br />

triangularis (Petrunkevitch); <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, x4,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1913'<br />

491<br />

http://jurassic.ru/

1411. Oomartus nyranensls Petrunkevitch, <strong>о</strong>бщий <strong>в</strong>ид, X3,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1412. Anthracophrynus<br />

tubercutatus Andree; брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Anthracomartus Karsch, 1882. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. voelkelianus Karsch, 1882; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

П<strong>о</strong>льш<strong>а</strong>. Виды р<strong>о</strong>д<strong>а</strong> п<strong>о</strong>х<strong>о</strong>жи н<strong>а</strong> Pleomartus, н<strong>о</strong><br />

н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong>ыделяет<strong>с</strong>я б<strong>о</strong>лее <strong>в</strong>ы­<br />

пу<strong>к</strong>л<strong>а</strong>я <strong>с</strong>рединн<strong>а</strong>я з<strong>о</strong>н<strong>а</strong>; <strong>о</strong>тн<strong>о</strong>шение длины 1<br />

Ри<strong>с</strong>. 1411—1413. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Anthracomartidae<br />

<strong>с</strong>ре­<br />

динн<strong>о</strong>й з<strong>о</strong>ны тергит<strong>о</strong><strong>в</strong>^<strong>к</strong> ее ширине р<strong>а</strong><strong>в</strong>н<strong>о</strong> 1 : 0,8;<br />

б<strong>о</strong>р<strong>о</strong>зды зде<strong>с</strong>ь р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

друг другу; передний <strong>к</strong>р<strong>а</strong>й пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>руглен, <strong>а</strong> з<strong>а</strong>дний прям<strong>о</strong>й. Из<strong>в</strong>е<strong>с</strong>тны<br />

четыре пл<strong>о</strong>х<strong>о</strong> <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных <strong>в</strong>ид<strong>а</strong>, <strong>в</strong><strong>с</strong>е из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии и П<strong>о</strong>льши.<br />

Anthracophrynus Andree, 1913. Тип<br />

р<strong>о</strong>д<strong>а</strong> — A. tubercutatus Andree, 1913; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Dudweiler, Saar). Опи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

з<strong>а</strong><strong>к</strong>руглен<strong>а</strong> н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых и з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>ях,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953)<strong>Л1</strong>413. Cryptomartus hindi (Pocock);<br />

<strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу, В — <strong>с</strong>б<strong>о</strong><strong>к</strong>у; х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Pocock, 1911).<br />

п<strong>о</strong>чти прям<strong>о</strong> у<strong>с</strong>ечен<strong>а</strong> <strong>с</strong>переди; ширин<strong>а</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong><strong>о</strong>й<br />

з<strong>о</strong>ны примерн<strong>о</strong> р<strong>а</strong><strong>в</strong>н<strong>а</strong> ширине б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых<br />

ч<strong>а</strong><strong>с</strong>тей тергит<strong>о</strong><strong>в</strong>, <strong>о</strong>тделенных <strong>о</strong>т <strong>с</strong>рединн<strong>о</strong>й шир<strong>о</strong><strong>к</strong><strong>о</strong>й<br />

з<strong>о</strong>ны их прямыми, <strong>с</strong>х<strong>о</strong>дящими<strong>с</strong>я <strong>с</strong>з<strong>а</strong>ди<br />

б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми; <strong>в</strong>д<strong>о</strong>ль них п<strong>о</strong> <strong>к</strong>р<strong>а</strong>ям тергит<strong>о</strong><strong>в</strong> <strong>с</strong>рединн<strong>о</strong>й<br />

з<strong>о</strong>ны р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> из пяти<br />

<strong>о</strong><strong>к</strong>руглы'х буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong> (ри<strong>с</strong>. 1412). Один <strong>в</strong>ид.<br />

К<strong>а</strong>рб<strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы: Brachypyge<br />

Woodward, 1878 (ри<strong>с</strong>. (1407); Maiocercus<br />

Pocock, 1911 (ри<strong>с</strong>. 1408A, Б); Cleptomartus Petrunkevitch,<br />

1949 (ри<strong>с</strong>. 1409A, Б); Cryptomartus<br />

Petrunkevitch, 1949 (ри<strong>с</strong>. 1413A, Б); из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong><br />

С. Амери<strong>к</strong>и — Coryphomartus Petrunkevitch,<br />

1945 (ри<strong>с</strong>. 1410); Pleomartus Petrunkevitch,<br />

1945.<br />

ПОДКЛАСС ARANEAE. ПАУКИ<br />

Тел<strong>о</strong> р<strong>а</strong><strong>с</strong>членен<strong>о</strong> н<strong>а</strong> <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> неб<strong>о</strong>льшую<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>му, или г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь, не<strong>с</strong>ущую <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти<br />

и <strong>к</strong>рупную <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>му, или брюш<strong>к</strong><strong>о</strong>;<br />

п<strong>о</strong><strong>с</strong>леднее, <strong>в</strong> <strong>о</strong>тличие <strong>о</strong>т <strong>в</strong><strong>с</strong>ех других Chelicerata,<br />

<strong>с</strong><strong>о</strong>единяет<strong>с</strong>я <strong>с</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й уз<strong>к</strong>им, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>им,<br />

492<br />

http://jurassic.ru/<br />

трубч<strong>а</strong>тым <strong>с</strong>тебель<strong>к</strong><strong>о</strong>м (ри<strong>с</strong>. 1427). В <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>,<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>к</strong><strong>а</strong><strong>к</strong> и у других хелицер<strong>о</strong><strong>в</strong>ых, <strong>в</strong>х<strong>о</strong>дят<br />

<strong>а</strong><strong>к</strong>р<strong>о</strong>н и ше<strong>с</strong>ть <strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong> <strong>с</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями.<br />

Стебеле<strong>к</strong> <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н VII тул<strong>о</strong><strong>в</strong>ищным <strong>с</strong>егмент<strong>о</strong>м.<br />

Брюш<strong>к</strong><strong>о</strong> (<strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>) у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>о</strong><strong>в</strong>.

Liphistiomorphae и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых Arachnomorphae<br />

(<strong>с</strong>м. <strong>с</strong>тр. 494 и 497) р<strong>а</strong><strong>с</strong>членен<strong>о</strong> н<strong>а</strong> 12 <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1419 А). У <strong>в</strong>ы<strong>с</strong>ших п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>о</strong>бъединяемых<br />

<strong>в</strong> <strong>о</strong>тряды Mygalomorphae и Araneomorphae,<br />

брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>с</strong>ли<strong>в</strong>ших<strong>с</strong>я пяти <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

мез<strong>о</strong><strong>с</strong><strong>о</strong>мы (VII—XI и п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> XVIII); <strong>о</strong><strong>с</strong>т<strong>а</strong>льные брюшные<br />

<strong>с</strong>егменты, р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ющие<strong>с</strong>я т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у з<strong>а</strong>р<strong>о</strong>дышей,<br />

<strong>в</strong> течение д<strong>а</strong>льнейшег<strong>о</strong> <strong>о</strong>нт<strong>о</strong>генез<strong>а</strong> редуцируют<strong>с</strong>я,<br />

и у <strong>в</strong>зр<strong>о</strong><strong>с</strong>лых жи<strong>в</strong><strong>о</strong>тных бы<strong>в</strong><strong>а</strong>ют пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лены<br />

(XII и XIII <strong>с</strong>егменты) не<strong>в</strong>р<strong>о</strong>мер<strong>а</strong>ми<br />

п<strong>о</strong>дгл<strong>о</strong>т<strong>о</strong>чн<strong>о</strong>й нер<strong>в</strong>н<strong>о</strong>й м<strong>а</strong><strong>с</strong><strong>с</strong>ы.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит цели<strong>к</strong><strong>о</strong>м <strong>с</strong><strong>в</strong>ерху п<strong>о</strong><strong>к</strong>ры<strong>в</strong><strong>а</strong>ет<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>му, <strong>а</strong> <strong>к</strong>р<strong>а</strong>я ег<strong>о</strong> <strong>о</strong>пу<strong>с</strong><strong>к</strong><strong>а</strong>ют<strong>с</strong>я и <strong>о</strong>бр<strong>а</strong>зуют<br />

л<strong>о</strong>б и б<strong>о</strong><strong>к</strong><strong>а</strong>. В <strong>с</strong>ередине щит<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>е<br />

<strong>в</strong>д<strong>а</strong><strong>в</strong>ление — <strong>с</strong>рединн<strong>а</strong>я ям<strong>к</strong><strong>а</strong>, <strong>о</strong>т <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong><br />

<strong>о</strong>тх<strong>о</strong>дят б<strong>о</strong>р<strong>о</strong>зды (ри<strong>с</strong>. 1419 Б). Н<strong>а</strong> переднем<br />

<strong>к</strong>р<strong>а</strong>ю щит<strong>а</strong> или н<strong>а</strong> лбу имеет<strong>с</strong>я <strong>в</strong><strong>о</strong><strong>с</strong>емь<br />

гл<strong>а</strong>з, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных <strong>о</strong>бычн<strong>о</strong> <strong>в</strong> д<strong>в</strong><strong>а</strong>-три п<strong>о</strong>перечных<br />

ряд<strong>а</strong> (ри<strong>с</strong>. 1426Б, 1433Б). Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>не брюш<strong>к</strong><strong>а</strong> т<strong>о</strong>ль<strong>к</strong><strong>о</strong> у члени<strong>с</strong>тых п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>о</strong>тряд<strong>а</strong> Liphistiomorphae е<strong>с</strong>ть тергиты; у<br />

<strong>в</strong><strong>с</strong>ех <strong>о</strong><strong>с</strong>т<strong>а</strong>льных п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> — не<strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е<br />

брюш<strong>к</strong><strong>о</strong>, п<strong>о</strong><strong>к</strong>рыт<strong>о</strong>е мяг<strong>к</strong>ими хитин<strong>о</strong><strong>в</strong>ыми п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>а</strong>ми;<br />

т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> ме<strong>с</strong>т<strong>а</strong>х при<strong>к</strong>репления пуч<strong>к</strong><strong>о</strong><strong>в</strong><br />

д<strong>о</strong>рз<strong>о</strong><strong>в</strong>ентр<strong>а</strong>льных мышц и н<strong>а</strong> т<strong>а</strong><strong>к</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>емых<br />

лег<strong>о</strong>чных <strong>к</strong>рыш<strong>к</strong><strong>а</strong>х имеют<strong>с</strong>я неб<strong>о</strong>льшие хитиниз<strong>о</strong><strong>в</strong><strong>а</strong>нные<br />

пл<strong>о</strong>щ<strong>а</strong>ди. Брюшн<strong>а</strong>я <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы<br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> <strong>к</strong>рупным <strong>с</strong>тернум<strong>о</strong>м, <strong>в</strong><strong>о</strong><strong>к</strong>руг<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>к</strong>рупные т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп<br />

и х<strong>о</strong>дильных н<strong>о</strong>г (ри<strong>с</strong>. 1428, 1434Б,<br />

1438 А). Впереди <strong>с</strong>тернум<strong>а</strong> у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен неб<strong>о</strong>льш<strong>о</strong>й <strong>с</strong>тернит педип<strong>а</strong>льп<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

(<strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>) <strong>с</strong>егмент<strong>а</strong>.<br />

Стернум <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>н <strong>с</strong>ли<strong>в</strong>шими<strong>с</strong>я <strong>с</strong>тернит<strong>а</strong>ми<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong> х<strong>о</strong>дильных н<strong>о</strong>г (III—VI); у мн<strong>о</strong>гих<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>с</strong>леды эт<strong>о</strong>г<strong>о</strong> <strong>с</strong>лияния <strong>в</strong>идны бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>ря<br />

н<strong>а</strong>личию п<strong>о</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>м <strong>с</strong>тернум<strong>а</strong> <strong>в</strong>д<strong>а</strong><strong>в</strong>лений<br />

и <strong>в</strong>ыпячи<strong>в</strong><strong>а</strong>ний (ри<strong>с</strong>. 1456А); з<strong>а</strong>дняя ч<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong>тернум<strong>а</strong> мн<strong>о</strong>гих п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>рединным <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рый, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>о</strong>т<strong>в</strong>еч<strong>а</strong>ет <strong>с</strong>терниту<br />

п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> пр<strong>о</strong><strong>с</strong><strong>о</strong>мы. Хелицеры <strong>к</strong>рупные,<br />

д<strong>в</strong>учлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; имеют м<strong>а</strong><strong>с</strong><strong>с</strong>и<strong>в</strong>н<strong>о</strong>е <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

и <strong>к</strong><strong>о</strong>гте<strong>в</strong>идную л<strong>а</strong>п<strong>к</strong>у (ри<strong>с</strong>. 1451, 1490 Б,<br />

1497 Б), н<strong>а</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>х <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong>ид<strong>о</strong><strong>в</strong><br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пр<strong>о</strong>т<strong>о</strong><strong>к</strong>и яд<strong>о</strong><strong>в</strong>итых желез. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния<br />

хелицер <strong>о</strong>гр<strong>а</strong>ничи<strong>в</strong><strong>а</strong>ют <strong>с</strong>переди предр<strong>о</strong>т<strong>о</strong><strong>в</strong>ую<br />

п<strong>о</strong>л<strong>о</strong><strong>с</strong>ть. С б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>н<strong>а</strong> <strong>о</strong>гр<strong>а</strong>ничен<strong>а</strong> м<strong>а</strong><strong>к</strong><strong>с</strong>иллярными<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп,<br />

<strong>а</strong> <strong>с</strong>низу — передним <strong>к</strong>р<strong>а</strong>ем педип<strong>а</strong>льп<strong>а</strong>льн<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>тернит<strong>а</strong>, н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ем<strong>о</strong>г<strong>о</strong> ин<strong>о</strong>гд<strong>а</strong> нижней губ<strong>о</strong>й<br />

(ри<strong>с</strong>. 1490Б). М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти педип<strong>а</strong>льп<br />

я<strong>в</strong>ляют<strong>с</strong>я <strong>в</strong>ид<strong>о</strong>измененными же<strong>в</strong><strong>а</strong>тельными<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>ми пер<strong>в</strong>ичн<strong>о</strong>й <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих;<br />

н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е <strong>о</strong>ни имеют гу<strong>с</strong>тую<br />

<strong>с</strong>еть <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые, пере<strong>к</strong>рещи<strong>в</strong><strong>а</strong>я<strong>с</strong>ь,<br />

<strong>с</strong><strong>о</strong>зд<strong>а</strong>ют <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зную <strong>с</strong>ет<strong>к</strong>у, <strong>с</strong>луж<strong>а</strong>щую для<br />

фильтр<strong>а</strong>ции пищи, п<strong>о</strong>гл<strong>о</strong>щ<strong>а</strong>ем<strong>о</strong>й уже <strong>в</strong> жид<strong>к</strong><strong>о</strong>м<br />

<strong>в</strong>иде. Педип<strong>а</strong>льпы у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong>тые, <strong>к</strong><strong>о</strong>­<br />

493<br />

http://jurassic.ru/<br />

р<strong>о</strong>т<strong>к</strong>ие, <strong>о</strong>бычн<strong>о</strong> <strong>в</strong> 3—4 р<strong>а</strong>з<strong>а</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>че х<strong>о</strong>дильных<br />

н<strong>о</strong>г. У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>нце<strong>в</strong>ые члени<strong>к</strong>и педип<strong>а</strong>льп<br />

<strong>с</strong>ильн<strong>о</strong> <strong>в</strong>здуты и <strong>в</strong>ид<strong>о</strong>изменены; <strong>о</strong>ни фун<strong>к</strong>ци<strong>о</strong>нируют<br />

<strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>, перен<strong>о</strong><strong>с</strong>ящих <strong>с</strong>перму<br />

<strong>в</strong> жен<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие (ри<strong>с</strong>. 1439 Б,<br />

1493, 1516). У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> педип<strong>а</strong>льпы р<strong>а</strong>з<strong>в</strong>иты<br />

н<strong>о</strong>рм<strong>а</strong>льн<strong>о</strong>, ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Х<strong>о</strong>дильные н<strong>о</strong>ги<br />

имеют <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е <strong>с</strong>тр<strong>о</strong>ение, н<strong>о</strong> р<strong>а</strong>злич<strong>а</strong>ют<strong>с</strong>я<br />

длин<strong>о</strong>й; <strong>о</strong>бычн<strong>о</strong> н<strong>о</strong>ги III бы<strong>в</strong><strong>а</strong>ют <strong>к</strong><strong>о</strong>р<strong>о</strong>че<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>льных. К<strong>а</strong>жд<strong>а</strong>я н<strong>о</strong>г<strong>а</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>с</strong>еми члени<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

л<strong>а</strong>п<strong>к</strong>и их р<strong>а</strong><strong>с</strong>членены н<strong>а</strong> пят<strong>к</strong>у (metatarsus)<br />

и <strong>с</strong><strong>о</strong>б<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> л<strong>а</strong>п<strong>к</strong>у (tarsus). Н<strong>а</strong> <strong>в</strong>ершине<br />

л<strong>а</strong>п<strong>о</strong><strong>к</strong> д<strong>в</strong><strong>а</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong> и пуч<strong>о</strong><strong>к</strong> щетин<strong>о</strong><strong>к</strong>;<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и ч<strong>а</strong><strong>с</strong>т<strong>о</strong> бы<strong>в</strong><strong>а</strong>ют гребенч<strong>а</strong>тыми, <strong>а</strong> <strong>с</strong>реди<br />

<strong>в</strong>ершинных щетин<strong>о</strong><strong>к</strong> не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые р<strong>а</strong>з<strong>в</strong>и<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> <strong>в</strong>иде <strong>к</strong>рупных з<strong>а</strong>зубренных шип<strong>о</strong><strong>в</strong>, при<strong>с</strong>п<strong>о</strong><strong>с</strong><strong>о</strong>бленных<br />

для перед<strong>в</strong>ижения п<strong>о</strong> п<strong>а</strong>утинным <strong>с</strong><strong>о</strong><strong>о</strong>ружениям<br />

(тенет<strong>а</strong>м). У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> мн<strong>о</strong>гих<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х л<strong>а</strong>п<strong>о</strong><strong>к</strong> н<strong>о</strong>г IV е<strong>с</strong>ть<br />

<strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>е <strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>ние — calamistrum, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящее<br />

из длинн<strong>о</strong>г<strong>о</strong> ряд<strong>а</strong> из<strong>о</strong>гнутых <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женные<br />

<strong>с</strong> б<strong>о</strong>льш<strong>о</strong>й пр<strong>а</strong><strong>в</strong>ильн<strong>о</strong><strong>с</strong>тью <strong>в</strong>д<strong>о</strong>ль<br />

предл<strong>а</strong>п<strong>к</strong>и.<br />

Н<strong>а</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не брюш<strong>к</strong><strong>а</strong>, неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

п<strong>о</strong>з<strong>а</strong>ди ме<strong>с</strong>т<strong>а</strong> при<strong>к</strong>репления <strong>с</strong>тебель<strong>к</strong><strong>а</strong>,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> п<strong>о</strong>перечн<strong>о</strong>е п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие,<br />

п<strong>о</strong>л<strong>о</strong>жение <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>еч<strong>а</strong>ет VIII тул<strong>о</strong><strong>в</strong>ищн<strong>о</strong>му<br />

<strong>с</strong>егменту (или <strong>в</strong>т<strong>о</strong>р<strong>о</strong>му <strong>о</strong>пи<strong>с</strong>т<strong>о</strong>м<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>му).<br />

У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>с</strong>тр<strong>о</strong>ение п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия у<strong>с</strong>л<strong>о</strong>жнен<strong>о</strong><br />

<strong>о</strong>бр<strong>а</strong>з<strong>о</strong><strong>в</strong><strong>а</strong>нием эпигины, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящей из<br />

неп<strong>а</strong>рн<strong>о</strong>г<strong>о</strong> чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>г<strong>о</strong> хитин<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> прид<strong>а</strong>т<strong>к</strong><strong>а</strong><br />

и п<strong>а</strong>рных ут<strong>о</strong>лщений <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и.<br />

Н<strong>а</strong> ур<strong>о</strong><strong>в</strong>не п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я<br />

пер<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> (<strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е<br />

Caponiidae <strong>о</strong>ни з<strong>а</strong>мещ<strong>а</strong>ют<strong>с</strong>я тр<strong>а</strong>хеями). Н<strong>а</strong> IX<br />

<strong>с</strong>егменте у низших п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> (<strong>о</strong>тряды Liphistiomorphae,<br />

Arachnomorphae, Mygalomorphae)<br />

н<strong>а</strong>х<strong>о</strong>дит<strong>с</strong>я <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong> лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> —<br />

<strong>в</strong>ид<strong>о</strong>измененных брюшных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей мез<strong>о</strong><strong>с</strong><strong>о</strong>мы.<br />

У <strong>в</strong>ы<strong>с</strong>ших п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>а</strong> Araneomorphae<br />

н<strong>а</strong> эт<strong>о</strong>м ме<strong>с</strong>те, <strong>в</strong>переди п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>,<br />

лежит неп<strong>а</strong>рн<strong>а</strong>я <strong>с</strong>тигм<strong>а</strong> тр<strong>а</strong>хей, <strong>в</strong>едущ<strong>а</strong>я <strong>в</strong><br />

<strong>а</strong>триум, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я д<strong>в</strong>е п<strong>а</strong>ры тр<strong>а</strong>хей.<br />

Н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> (н<strong>а</strong> ме<strong>с</strong>те<br />

и<strong>с</strong>чезну<strong>в</strong>ших X и XI тул<strong>о</strong><strong>в</strong>ищных <strong>с</strong>егмент<strong>о</strong><strong>в</strong>)<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены три п<strong>а</strong>ры п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 1435 Д, 1473 Б, 1513 В, 1516). Передние<br />

и з<strong>а</strong>дние п<strong>а</strong>ры этих б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>, д<strong>в</strong>ух-трехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>чень длинные (ри<strong>с</strong>. 1428 Б,<br />

1436), пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ляют <strong>с</strong><strong>о</strong>б<strong>о</strong>й <strong>в</strong>ид<strong>о</strong>измененные<br />

брюшные н<strong>о</strong>ж<strong>к</strong>и X и XI <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, <strong>а</strong> <strong>в</strong>нутренняя<br />

п<strong>а</strong>р<strong>а</strong> м<strong>а</strong>лень<strong>к</strong>их б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> при р<strong>а</strong>з<strong>в</strong>итии<br />

<strong>о</strong>тделяет<strong>с</strong>я <strong>о</strong>т з<strong>а</strong>ч<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> з<strong>а</strong>дней п<strong>а</strong>ры и, <strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>,<br />

<strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельными <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями не<br />

я<strong>в</strong>ляет<strong>с</strong>я. Щель между <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями з<strong>а</strong>дних<br />

п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> з<strong>а</strong>п<strong>о</strong>лнен<strong>а</strong> м<strong>а</strong>лень<strong>к</strong>им<br />

<strong>к</strong><strong>о</strong>ниче<strong>с</strong><strong>к</strong>им прид<strong>а</strong>т<strong>к</strong><strong>о</strong>м cohilus. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

имеющих calamistrum, <strong>в</strong>переди з<strong>а</strong>дних<br />

п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> (<strong>в</strong>ме<strong>с</strong>т<strong>о</strong> colulus) р<strong>а</strong><strong>с</strong>п<strong>о</strong>-

л<strong>о</strong>жен<strong>а</strong> п<strong>о</strong>перечн<strong>а</strong>я, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> д<strong>в</strong><strong>о</strong>йн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>а</strong>я и<strong>с</strong>пещрен<strong>а</strong> т<strong>о</strong>ч<strong>к</strong><strong>а</strong>ми, прид<strong>а</strong>ющими ей<br />

<strong>в</strong>ид <strong>с</strong>ит<strong>а</strong> — cribellum. Н<strong>а</strong>личие эт<strong>о</strong>й пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и<br />

игр<strong>а</strong>ет <strong>в</strong><strong>а</strong>жную р<strong>о</strong>ль <strong>в</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>ти<strong>к</strong>е. П<strong>а</strong>утинные<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и имеют <strong>с</strong>пеци<strong>а</strong>льные труб<strong>о</strong>ч<strong>к</strong>и,<br />

через <strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>ых<strong>о</strong>дят нити <strong>с</strong>е<strong>к</strong>рет<strong>а</strong> желез,<br />

з<strong>а</strong><strong>с</strong>ты<strong>в</strong><strong>а</strong>ющие н<strong>а</strong> <strong>в</strong><strong>о</strong>здухе <strong>в</strong> <strong>в</strong>иде п<strong>а</strong>утины<br />

(ри<strong>с</strong>. 1473 Б). П<strong>о</strong>з<strong>а</strong>ди п<strong>а</strong>утинн<strong>о</strong>г<strong>о</strong> п<strong>о</strong>ля, н<strong>а</strong><br />

з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong>, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ет<strong>с</strong>я <strong>а</strong>н<strong>а</strong>льн<strong>о</strong>е<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие, ин<strong>о</strong>гд<strong>а</strong> леж<strong>а</strong>щее н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>м <strong>в</strong><strong>о</strong>з<strong>в</strong>ышении. К<strong>а</strong>рб<strong>о</strong>н —<br />

ныне.<br />

Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и би<strong>о</strong>ге<strong>о</strong>гр<strong>а</strong>фия. П<strong>а</strong>у<strong>к</strong>и,<br />

з<strong>а</strong> единичными и<strong>с</strong><strong>к</strong>лючениями (р<strong>о</strong>д Argyroneta),<br />

<strong>в</strong>едут н<strong>а</strong>земный <strong>о</strong>бр<strong>а</strong>з жизни, <strong>в</strong><strong>с</strong>треч<strong>а</strong>я<strong>с</strong>ь<br />

бу<strong>к</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong> п<strong>о</strong><strong>в</strong><strong>с</strong>юду. Хищни<strong>к</strong>и; п<strong>о</strong>жир<strong>а</strong>ют члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих,<br />

гл<strong>а</strong><strong>в</strong>ным <strong>о</strong>бр<strong>а</strong>з<strong>о</strong>м н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых, <strong>в</strong>ы<strong>с</strong><strong>а</strong><strong>с</strong>ы<strong>в</strong><strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong> р<strong>а</strong>змельченн<strong>о</strong>е хелицер<strong>а</strong>ми<br />

и р<strong>а</strong><strong>с</strong>т<strong>в</strong><strong>о</strong>ренн<strong>о</strong>е желуд<strong>о</strong>чным <strong>с</strong><strong>о</strong><strong>к</strong><strong>о</strong>м тел<strong>о</strong><br />

н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>с</strong>тр<strong>о</strong>ит<br />

л<strong>о</strong><strong>в</strong>чие <strong>с</strong>ети из п<strong>а</strong>утины, или <strong>о</strong>ни жи<strong>в</strong>ут <strong>в</strong><br />

н<strong>о</strong>р<strong>к</strong><strong>а</strong>х.<br />

П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>гения<br />

и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>. Б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>телей <strong>с</strong>пр<strong>а</strong><strong>в</strong>едли<strong>в</strong><strong>о</strong> <strong>с</strong>ближ<strong>а</strong>ет<br />

Аг<strong>а</strong>пе<strong>а</strong>е <strong>с</strong> Amblypygi, <strong>с</strong>чит<strong>а</strong>я Liphistiomorphae<br />

н<strong>а</strong>иб<strong>о</strong>лее примити<strong>в</strong>н<strong>о</strong>й групп<strong>о</strong>й. Р<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>о</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

<strong>с</strong> другими п<strong>о</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>ми Arachnida <strong>о</strong>тмечен<strong>о</strong><br />

<strong>в</strong>ыше. Шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненн<strong>о</strong>е р<strong>а</strong>зделение<br />

Аг<strong>а</strong>пе<strong>а</strong>е н<strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряды (= <strong>о</strong>тряды!)<br />

четырехлег<strong>о</strong>чных (Tetrapneumones) и д<strong>в</strong>улег<strong>о</strong>чных<br />

(Dipneumones) п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>, или, <strong>к</strong><strong>а</strong><strong>к</strong> их<br />

предл<strong>о</strong>жил н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ть Берл<strong>а</strong>н (Berlan, 1932),<br />

н<strong>а</strong> «п<strong>о</strong>д<strong>о</strong>тряды» Orthognatha и Labidognatha,<br />

неуд<strong>а</strong>чн<strong>о</strong>, т<strong>а</strong><strong>к</strong> <strong>к</strong><strong>а</strong><strong>к</strong> при эт<strong>о</strong>м т<strong>а</strong><strong>к</strong><strong>а</strong>я е<strong>с</strong>те<strong>с</strong>т<strong>в</strong>енн<strong>а</strong>я<br />

групп<strong>а</strong>, <strong>к</strong><strong>а</strong><strong>к</strong> члени<strong>с</strong>тые п<strong>а</strong>у<strong>к</strong>и <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Liphistiidae,<br />

и<strong>с</strong><strong>к</strong>у<strong>с</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>о</strong>бъединяет<strong>с</strong>я <strong>с</strong> четырех-<br />

лег<strong>о</strong>чными п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми, <strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

Hypochilidae, имеющие четыре лег<strong>к</strong>их, тем<br />

не менее п<strong>о</strong> мн<strong>о</strong>гим другим призн<strong>а</strong><strong>к</strong><strong>а</strong>м <strong>о</strong>бъединяют<strong>с</strong>я<br />

<strong>с</strong> д<strong>в</strong>улег<strong>о</strong>чными п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми. Эти нед<strong>о</strong>четы<br />

и<strong>с</strong>пр<strong>а</strong><strong>в</strong>ил Ке<strong>с</strong>тнер (Kastner, 1938), р<strong>а</strong>здели<strong>в</strong><br />

<strong>в</strong><strong>с</strong>ех п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong> — Mesothelae,<br />

Orthognatha и Labidognatha, <strong>о</strong>тр<strong>а</strong>зи<strong>в</strong> <strong>в</strong><br />

н<strong>а</strong>з<strong>в</strong><strong>а</strong>ниях <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти п<strong>о</strong>л<strong>о</strong>жения хелицер<br />

<strong>о</strong>тн<strong>о</strong><strong>с</strong>ительн<strong>о</strong> пр<strong>о</strong>д<strong>о</strong>льн<strong>о</strong>й <strong>о</strong><strong>с</strong>и тел<strong>а</strong>. Одн<strong>а</strong><strong>к</strong><strong>о</strong> еще<br />

р<strong>а</strong>ньше П<strong>о</strong><strong>к</strong><strong>о</strong><strong>к</strong> (Pocock, 1892) и Милл<strong>о</strong> (Millot,<br />

1933) предл<strong>о</strong>жили для этих трех п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong><br />

б<strong>о</strong>лее нейтр<strong>а</strong>льные н<strong>а</strong>имен<strong>о</strong><strong>в</strong><strong>а</strong>ния: Liphistiomorphae<br />

и Mygalomorphae и Araneomorphae.<br />

Из <strong>о</strong>пи<strong>с</strong><strong>а</strong>нных Петрун<strong>к</strong>е<strong>в</strong>ичем (Petrunkevitch,<br />

1933) трех н<strong>о</strong><strong>в</strong>ых п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> — Hypochilomorphae,<br />

Apneumomorphae и Arachnomorphae —<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ители т<strong>о</strong>ль<strong>к</strong><strong>о</strong> п<strong>о</strong><strong>с</strong>леднег<strong>о</strong> из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong><br />

и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии (<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archaeometidae<br />

и Pyritaraneidae). С<strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельн<strong>о</strong><strong>с</strong>ть<br />

других д<strong>в</strong>ух п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> п<strong>о</strong>д<strong>в</strong>ерг<strong>а</strong>ет<strong>с</strong>я <strong>с</strong><strong>о</strong>мнению,<br />

и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>а</strong><strong>в</strong>т<strong>о</strong>ры <strong>о</strong>тн<strong>о</strong><strong>с</strong>ят <strong>в</strong>х<strong>о</strong>дящие <strong>в</strong><br />

эти п<strong>о</strong>д<strong>о</strong>тряды <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>к</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>м Megalomorphae<br />

и Araneomorphae (Millot, 1949).<br />

Т<strong>а</strong><strong>к</strong> же <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>а</strong> предл<strong>а</strong>г<strong>а</strong>ем<strong>а</strong>я<br />

Петрун<strong>к</strong>е<strong>в</strong>ичем (1933) <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция <strong>о</strong>тдельных<br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>о</strong><strong>в</strong> и б<strong>о</strong>лее мел<strong>к</strong>их т<strong>а</strong><strong>к</strong><strong>с</strong><strong>о</strong>н<strong>о</strong>миче<strong>с</strong><strong>к</strong>их<br />

<strong>к</strong><strong>а</strong>тег<strong>о</strong>рий, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>а</strong>я преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

н<strong>а</strong> р<strong>а</strong>зличиях <strong>в</strong> чи<strong>с</strong>ле <strong>о</strong><strong>с</strong>тий <strong>с</strong>ердц<strong>а</strong>. Б<strong>о</strong>лее<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>й я<strong>в</strong>ляет<strong>с</strong>я <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция Ке<strong>с</strong>тнер<br />

<strong>а</strong> (1938) и Милл<strong>о</strong> (1949), <strong>к</strong><strong>о</strong>т<strong>о</strong>рую мы и<br />

приним<strong>а</strong>ем для н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящег<strong>о</strong> изд<strong>а</strong>ния, <strong>с</strong>чит<strong>а</strong>я<br />

лишь п<strong>о</strong>д<strong>о</strong>тряды, предл<strong>о</strong>женные этими <strong>а</strong><strong>в</strong>т<strong>о</strong>р<strong>а</strong>ми,<br />

з<strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>с</strong>т<strong>о</strong>ятельные <strong>о</strong>тряды. Де<strong>в</strong><strong>о</strong>н — ныне.<br />

Четыре <strong>о</strong>тряд<strong>а</strong>: Liphistiomorphae, Arachnomorphae,<br />

Mygalomorphae и Araneomorphae.<br />

ОТРЯД LIPHISTIOMORPHAE. Л ИФИСТОМОРФНЫЕ,<br />

ИЛИ ЧЛЕНИСТОБРЮХИЕ ПАУКИ<br />

Брюш<strong>к</strong><strong>о</strong> я<strong>в</strong><strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> и имеет<br />

11 тергит<strong>о</strong><strong>в</strong>. П<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> <strong>с</strong>емь или<br />

<strong>в</strong><strong>о</strong><strong>с</strong>емь; <strong>о</strong>ни р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>переди <strong>о</strong>т<br />

<strong>а</strong>н<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> бугр<strong>а</strong>; между п<strong>о</strong><strong>с</strong>ледним и п<strong>а</strong>утинными<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми леж<strong>а</strong>т <strong>с</strong>терниты з<strong>а</strong>дних<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти педип<strong>а</strong>льп не<br />

р<strong>а</strong>з<strong>в</strong>иты. Хелицеры п<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льные. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong><br />

тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. Гл<strong>а</strong>з<strong>а</strong> <strong>в</strong> чи<strong>с</strong>ле <strong>в</strong><strong>о</strong><strong>с</strong>ьми р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> <strong>о</strong>бщем буг<strong>о</strong>р<strong>к</strong>е. Эпигин<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

К<strong>а</strong>рб<strong>о</strong>н — ныне. Три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>; из них<br />

д<strong>в</strong><strong>а</strong> п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их (Arthromygalidae и Arthrolycosidae)<br />

и <strong>о</strong>дн<strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>е (Liphistiidae).<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО ARTHROMYGALIDAE<br />

PETRUNKEVITCH, 1923<br />

Н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>м щите гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует: гл<strong>а</strong>з, п<strong>о</strong>-<strong>в</strong>идим<strong>о</strong>му, нет. К<strong>а</strong>рб<strong>о</strong>н.<br />

Семь р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Protolycosa Roemer, 1866. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. anthracophila Roemer, 1866; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, П<strong>о</strong>льш<strong>а</strong>.<br />

Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не брюш<strong>к</strong><strong>а</strong> п<strong>о</strong>перечные<br />

ряды буг<strong>о</strong>р<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ующих н<strong>а</strong> <strong>о</strong>динн<strong>а</strong>дц<strong>а</strong>т<strong>о</strong>м<br />

<strong>с</strong>егменте. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Arthromygale Petrunkevitch, 1923.<br />

Тип р<strong>о</strong>д<strong>а</strong>—Arthrolycosa fortis Fritsch, 1904;

1419Б<br />

1415Б<br />

Ри<strong>с</strong>. 1414—1420. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Arthromygalidae и Arthrolycosidae<br />

1414. Arthromygala fortis (Fritsch); пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, x5,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1415. Geralycosa fritschi<br />

Kusta; А — <strong>о</strong>бщий <strong>в</strong>ид, x5,0, Б — Г<strong>о</strong>лень и л<strong>а</strong>п<strong>к</strong><strong>а</strong> н<strong>о</strong>ги III,<br />

Х15.0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1953). 1416. Kustaria<br />

carbonaria (Kusta); <strong>о</strong>бщий <strong>в</strong>ид, х3,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1953). 1417. Racovnicia antiqua Kusta; <strong>о</strong>бщий <strong>в</strong>ид, X 2,0,<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит<br />

п<strong>о</strong>чти ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идный, <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong> з<strong>а</strong><strong>к</strong>ругленными<br />

передней и б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми и <strong>с</strong> ше<strong>с</strong>тью<br />

чет<strong>к</strong>ими б<strong>о</strong>р<strong>о</strong>зд<strong>а</strong>ми, р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> <strong>о</strong>тх<strong>о</strong>дящими <strong>о</strong>т<br />

<strong>с</strong>рединн<strong>о</strong>й ям<strong>к</strong>и; з<strong>а</strong>дний <strong>к</strong><strong>о</strong>нец <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>г<strong>о</strong><br />

брюш<strong>к</strong><strong>а</strong> з<strong>а</strong><strong>к</strong>руглен; н<strong>о</strong>ги <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

т<strong>о</strong>л<strong>с</strong>тые; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь р<strong>а</strong><strong>в</strong>н<strong>а</strong> 2<br />

/ 3 длины брюш<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> 15 мм, длин<strong>а</strong> <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong> 6,57 мм<br />

(ри<strong>с</strong>. 1414). Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Palaranea Fritsch, 1873. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. borassifoliae Fritsch, 1873; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит з<strong>а</strong><strong>к</strong>руглен <strong>с</strong>переди,<br />

ширин<strong>а</strong> ег<strong>о</strong> пре<strong>в</strong>ыш<strong>а</strong>ет длину; л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г без<br />

пуль<strong>в</strong>иллы; н<strong>о</strong>ги <strong>о</strong>чень длинные; н<strong>о</strong>ги IV длиннее<br />

тел<strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong> 23 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н<br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

Geralycosa Kusta, 1888. Тип р<strong>о</strong>д<strong>а</strong> — G.<br />

fritschi Kusta, 1888; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Длин<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди пре<strong>в</strong>ыш<strong>а</strong>ет ее ширину; <strong>о</strong>н<strong>а</strong><br />

з<strong>а</strong><strong>к</strong>руглен<strong>а</strong> н<strong>а</strong> переднем <strong>к</strong><strong>о</strong>нце и <strong>в</strong><strong>о</strong>гнут<strong>а</strong> н<strong>а</strong><br />

з<strong>а</strong>днем, гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я, без гл<strong>а</strong>з, <strong>с</strong> пятью п<strong>а</strong>р<strong>а</strong>ми р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящих<strong>с</strong>я б<strong>о</strong>р<strong>о</strong>зд; н<strong>о</strong>ги длинные<br />

495<br />

1419А<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1933). 1418. Protocteniza britanт<strong>с</strong><strong>а</strong>:<br />

Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1953). 1419. Arthrolycosa danielsi Petrunkevitch; <strong>о</strong>бщий<br />

? ид<br />

:* Л. ~;„<br />

<strong>с</strong> н и з у<br />

' Б<br />

— <strong>с</strong><strong>в</strong><br />

ерху; Х2,5, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, С. Амери<strong>к</strong><strong>а</strong> (Petrun­<br />

kevitch, 1913). 1420. Eocteniza silvicola Pocock; <strong>о</strong>бщий <strong>в</strong>ид, X 3,0<br />

<strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Pocock, 1911)<br />

и т<strong>о</strong>н<strong>к</strong>ие (ри<strong>с</strong>. 1415); л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

и пуль<strong>в</strong>илл<strong>о</strong>й (ри<strong>с</strong>. 1415 Б). Длин<strong>а</strong> тел<strong>а</strong><br />

14,5 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Kustaria Petrunkevitch, 1953. Тип<br />

р<strong>о</strong>д<strong>а</strong> — Scudderia carbonaria Kusta, 1888; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ии. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>о</strong><strong>к</strong>руглый<br />

<strong>с</strong> <strong>в</strong><strong>о</strong>гнутым передним <strong>к</strong>р<strong>а</strong>ем и <strong>с</strong> шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

з<strong>а</strong><strong>к</strong>ругленными б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми и з<strong>а</strong>дним <strong>к</strong>р<strong>а</strong>ями;<br />

гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; педип<strong>а</strong>льпы <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, т<strong>о</strong>н<strong>к</strong>ие;<br />

н<strong>о</strong>ги длинные (ри<strong>с</strong>. 1416). Длин<strong>а</strong> тел<strong>а</strong><br />

5,7 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Rakovnicia Kusta, 1884. Тип р<strong>о</strong>д<strong>а</strong> — R.<br />

antiqua Kusta, 1884: <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный<br />

щит <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong><strong>о</strong><strong>в</strong><strong>а</strong>льн<strong>о</strong>й ф<strong>о</strong>рмы, немн<strong>о</strong>г<strong>о</strong><br />

длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей ширины, гл<strong>а</strong>д<strong>к</strong>ий, <strong>с</strong> тремя<br />

п<strong>а</strong>р<strong>а</strong>ми р<strong>а</strong>ди<strong>а</strong>льн<strong>о</strong> р<strong>а</strong><strong>с</strong>х<strong>о</strong>дящих<strong>с</strong>я б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong><strong>к</strong>;<br />

брюш<strong>к</strong><strong>о</strong> <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong> <strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong>, п<strong>о</strong>чти <strong>в</strong> 2<br />

р<strong>а</strong>з<strong>а</strong> б<strong>о</strong>лее длинн<strong>о</strong>г<strong>о</strong>, чем ег<strong>о</strong> ширин<strong>а</strong><br />

(ри<strong>с</strong>. 1417); н<strong>о</strong>ги <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие и т<strong>о</strong>н<strong>к</strong>ие. Длин<strong>а</strong><br />

тел<strong>а</strong> 6,6 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы Proto-.<br />

cteniza Petrunkevitch, 1949 (ри<strong>с</strong>. 1418).<br />

http://jurassic.ru/

СЕМЕЙСТВО ARTHROLYCOS1DAE<br />

PETRUNKEVITCH, 1923<br />

Члени<strong>с</strong>т<strong>о</strong>брюхие п<strong>а</strong>у<strong>к</strong>и, н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди <strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

имеет<strong>с</strong>я гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> (ри<strong>с</strong>. 1420,<br />

1419А). К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы и С. Амери<strong>к</strong>и. Д<strong>в</strong><strong>а</strong><br />

р<strong>о</strong>д<strong>а</strong>: Arthrolycosa Harger, 1874 (С. Амери<strong>к</strong><strong>а</strong>)<br />

и Eocteniza Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911 (3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>).<br />

Palaeocteniza Hirst, 1923. 1ип р<strong>о</strong>д<strong>а</strong> — Р.<br />

crassipes Hirst, 1923; де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия. Н<strong>а</strong>йден<br />

<strong>в</strong> <strong>к</strong>р<strong>а</strong><strong>с</strong>н<strong>о</strong>м пе<strong>с</strong>ч<strong>а</strong>ни<strong>к</strong>е Ш<strong>о</strong>тл<strong>а</strong>ндии <strong>с</strong>реди<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленных <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> Trigonotarbi и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>к</strong>лещей; п<strong>о</strong> <strong>о</strong>дним призн<strong>а</strong><strong>к</strong><strong>а</strong>м п<strong>о</strong>х<strong>о</strong>дит<br />

н<strong>а</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>, <strong>а</strong> п<strong>о</strong> другим — н<strong>а</strong> пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

1422А<br />

Ри<strong>с</strong>. 1421—1426. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archaeometidae, Pyritaraneidae и Dipluridae<br />

1421. Eopholcus pedatus Fritsch: <strong>о</strong>бщий <strong>в</strong>ид, x4,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Petrunkevitsch, 1946). 1422. Archaeometa nephilina Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>;<br />

<strong>о</strong>бщий <strong>в</strong>ид: А — <strong>с</strong><strong>в</strong>ерху, Б — <strong>с</strong>низу, х5,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(Р<strong>о</strong><strong>с</strong><strong>о</strong><strong>с</strong><strong>к</strong>, 1911; Petrunkevitsch, 1953). 1423. Arachnometa tuberculoid<br />

Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, х7,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1953). 1424. Pyritaranea tubifera Fritsch; <strong>о</strong>бщий <strong>в</strong>ид,<br />

ARTHROLYCOSIDAE INCERTAE SEDIS<br />

Eolycosa Kusta, 1885. Тип р<strong>о</strong>д<strong>а</strong> — E. lorenzi<br />

Kusta, 1885; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Пл<strong>о</strong>х<strong>о</strong><br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>ший<strong>с</strong>я <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong> п<strong>а</strong>у<strong>к</strong><strong>а</strong>, близ<strong>к</strong><strong>о</strong>г<strong>о</strong><br />

<strong>к</strong> р<strong>о</strong>ду Arthrolycosa Harger. Один <strong>в</strong>ид.<br />

496<br />

http://jurassic.ru/<br />

14225<br />

ХЗ.О, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch, 1946). 1425. Dinopillogi'gas<br />

Fritsch; <strong>о</strong>бщий <strong>в</strong>ид, х2,0, <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Petrunkevitch,<br />

1949). 1426. Clostes priscus Menge; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, х 5,5, Б — гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong>, х60,0; В — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и<br />

н<strong>о</strong>ги IV, X 250,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Petrunkevitch, 1946)<br />

<strong>о</strong>тряд<strong>а</strong> Trigonotarbi. Не<strong>о</strong>бх<strong>о</strong>дим<strong>о</strong> д<strong>о</strong>п<strong>о</strong>лнительн<strong>о</strong>е<br />

изучение.<br />

Pleurolycosa Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> —<br />

Arthrolycosa prolifera Fritsch, 1904; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия. Си<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>е п<strong>о</strong>л<strong>о</strong>жение<br />

нея<strong>с</strong>н<strong>о</strong>.

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong> цели<strong>к</strong><strong>о</strong>м или т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце; тергит<strong>о</strong><strong>в</strong> <strong>о</strong>т четырех д<strong>о</strong><br />

<strong>с</strong>еми. Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти педип<strong>а</strong>льп <strong>с</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми,<br />

<strong>о</strong>бр<strong>а</strong>зующими фильтр<strong>а</strong>ци<strong>о</strong>нную <strong>с</strong>ет<strong>к</strong>у<br />

(ри<strong>с</strong>. 1450). Хелицеры, н<strong>о</strong>ги и п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и<br />

р<strong>а</strong>з<strong>в</strong>иты т<strong>а</strong><strong>к</strong> же, <strong>к</strong><strong>а</strong><strong>к</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных<br />

Araneomorphae. Гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>ближены, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди. С<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упительные<br />

<strong>о</strong>рг<strong>а</strong>ны <strong>с</strong><strong>а</strong>м<strong>к</strong>и у<strong>с</strong>тр<strong>о</strong>ены <strong>с</strong>л<strong>о</strong>жн<strong>о</strong>. Орг<strong>а</strong>ны<br />

дых<strong>а</strong>ния <strong>к</strong><strong>а</strong><strong>к</strong> у н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих, <strong>а</strong>р<strong>а</strong>не<strong>о</strong>м<strong>о</strong>рфных,<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>.<br />

Неб<strong>о</strong>льш<strong>а</strong>я групп<strong>а</strong> п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>к</strong><strong>а</strong>менн<strong>о</strong>уг<strong>о</strong>льн<strong>о</strong>г<strong>о</strong><br />

пери<strong>о</strong>д<strong>а</strong>, <strong>с</strong><strong>о</strong>чет<strong>а</strong>ющ<strong>а</strong>я призн<strong>а</strong><strong>к</strong>и члени<strong>с</strong>т<strong>о</strong>брюхих<br />

п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>а</strong> Liphistiomorphae и<br />

н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong> <strong>о</strong>тряд<strong>а</strong> Araneomorphae.<br />

Петрун<strong>к</strong>е<strong>в</strong>ич (Petrunkevitch, 1933, 1949, 1953)<br />

<strong>о</strong>тмеч<strong>а</strong>ет р<strong>а</strong><strong>с</strong>члененн<strong>о</strong><strong>с</strong>ть брюш<strong>к</strong><strong>а</strong> этих п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong><br />

н<strong>а</strong>ряду <strong>с</strong> н<strong>а</strong>личием призн<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих Araneomorphae.<br />

Он не <strong>о</strong>пределяет т<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>л<strong>о</strong>жения Archaeometidae и Pyritaraneidae,<br />

<strong>а</strong> н<strong>а</strong>зы<strong>в</strong><strong>а</strong>ет их п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>ими <strong>а</strong>р<strong>а</strong>хн<strong>о</strong>м<strong>о</strong>рфными<br />

п<strong>а</strong>у<strong>к</strong><strong>а</strong>ми, близ<strong>к</strong>ими <strong>к</strong> рецентным.<br />

К<strong>а</strong>рб<strong>о</strong>н. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Archaeometidae и Руritaraneidae.<br />

СЕМЕЙСТВО ARCHAEOMETIDAE<br />

PETRUNKEVITSH, 1949<br />

П<strong>а</strong>у<strong>к</strong>и <strong>с</strong> члени<strong>с</strong>тым брюш<strong>к</strong><strong>о</strong>м и пр<strong>о</strong>гр<strong>а</strong>дными<br />

н<strong>о</strong>г<strong>а</strong>ми (ри<strong>с</strong>. 1422, 1423). К<strong>а</strong>рб<strong>о</strong>н. Три р<strong>о</strong>д<strong>а</strong>.<br />

Eopholcus Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> — Е.<br />

pedatus Fritsch, 1904; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>о</strong><strong>в</strong><strong>а</strong>льный, з<strong>а</strong><strong>к</strong>ругленный<br />

ОТРЯД ARACHNOMORPHAE<br />

<strong>с</strong>переди; брюш<strong>к</strong><strong>о</strong> ди<strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>е <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми <strong>с</strong>егмент<strong>а</strong>ции;<br />

д<strong>в</strong><strong>а</strong> гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> неб<strong>о</strong>льш<strong>о</strong>м<br />

буг<strong>о</strong>р<strong>к</strong>е; н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные; <strong>в</strong>ертлуги<br />

длиннее бедер (ри<strong>с</strong>. 1421). Длин<strong>а</strong> тел<strong>а</strong><br />

27,16 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н Е<strong>в</strong>р<strong>о</strong>пы.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<strong>а</strong> 3. Е<strong>в</strong>р<strong>о</strong>пы: Archaeometa<br />

Pocock, 1911 (ри<strong>с</strong>. 1422 А, Б) и Arachnometa<br />

Petrunkevitch, 1949 (ри<strong>с</strong>. 1423).<br />

СЕМЕЙСТВО PYRITARANEIDAE<br />

PETRUNKEVITCH, 1953<br />

Брюш<strong>к</strong><strong>о</strong> п<strong>о</strong>лн<strong>о</strong><strong>с</strong>тью <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>. Н<strong>о</strong>ги<br />

л<strong>а</strong>теригр<strong>а</strong>дные, т<strong>о</strong>н<strong>к</strong>ие и длинные. К<strong>а</strong>рб<strong>о</strong>н.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>.<br />

Pyritaranea Fritsch, 1899. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. tubifera Fritsch, 1899; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>о</strong><strong>в</strong><strong>а</strong>льный, длиннее <strong>с</strong><strong>в</strong><strong>о</strong>ей<br />

ширины, з<strong>а</strong><strong>к</strong>руглен <strong>с</strong>переди и прям<strong>о</strong> у<strong>с</strong>ечен<br />

<strong>с</strong>з<strong>а</strong>ди; гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют; н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие и длинные;<br />

длин<strong>а</strong> педип<strong>а</strong>льп б<strong>о</strong>льше ширины тел<strong>а</strong><br />

(ри<strong>с</strong>. 1424). Длин<strong>а</strong> тел<strong>а</strong> 11 мм. Один <strong>в</strong>ид. К<strong>а</strong>рб<strong>о</strong>н<br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

Dinopilio Fritsch, 1904. Тип р<strong>о</strong>д<strong>а</strong> — D.<br />

gigas Fritsch, 1904; <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Чех<strong>о</strong><strong>с</strong>л<strong>о</strong><strong>в</strong><strong>а</strong><strong>к</strong>ия.<br />

Н<strong>о</strong>ги <strong>о</strong>чень длинные, <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> т<strong>о</strong>л<strong>с</strong>тые,<br />

р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; т<strong>а</strong>зи<strong>к</strong>и н<strong>о</strong>г (ри<strong>с</strong>. 1425)<br />

леж<strong>а</strong>т п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> друг другу; т<strong>а</strong>зи<strong>к</strong>и<br />

педип<strong>а</strong>льп <strong>с</strong> уз<strong>к</strong>ими м<strong>а</strong><strong>к</strong><strong>с</strong>иллярными л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями<br />

и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1425). К<strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><br />

длин<strong>о</strong>й 7,5—8 мм; пр<strong>о</strong>межут<strong>о</strong><strong>к</strong> между т<strong>а</strong>зи<strong>к</strong><strong>а</strong>ми<br />

н<strong>о</strong>г р<strong>а</strong><strong>в</strong>ен 11,4 мм. Д<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>а</strong>. К<strong>а</strong>рб<strong>о</strong>н<br />

Е<strong>в</strong>р<strong>о</strong>пы.<br />

ОТРЯД MYGALOMORPHAE. ПАУКИ-ПТИЦЕЯДЫ<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong>н<strong>а</strong>ружи не<strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е.<br />

П<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>в</strong> <strong>к</strong><strong>о</strong>личе<strong>с</strong>т<strong>в</strong>е <strong>о</strong>т д<strong>в</strong>ух д<strong>о</strong><br />

ше<strong>с</strong>ти р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце брюш<strong>к</strong><strong>а</strong>.<br />

Хелицеры п<strong>а</strong>р<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льные. К<strong>о</strong><strong>к</strong><strong>с</strong>ы педип<strong>а</strong>льп<br />

<strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми челю<strong>с</strong>тными л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями,<br />

н<strong>о</strong> ч<strong>а</strong>ще без них. Лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong> д<strong>в</strong>е<br />

п<strong>а</strong>ры. Гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>ближены, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> переднем<br />

<strong>к</strong>р<strong>а</strong>е г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди. С<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упительные <strong>о</strong>рг<strong>а</strong>ны<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и пр<strong>о</strong><strong>с</strong>тые. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>: Nelipoda, Нур<strong>о</strong>demata<br />

и Pycnothelidea. Обит<strong>а</strong>ют преимуще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong><br />

<strong>в</strong> тр<strong>о</strong>пиче<strong>с</strong><strong>к</strong>их <strong>с</strong>тр<strong>а</strong>н<strong>а</strong>х; <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>м<br />

бр<strong>о</strong>дячие и дре<strong>в</strong>е<strong>с</strong>ные <strong>в</strong>иды, жи<strong>в</strong>ущие <strong>в</strong> н<strong>о</strong>р<strong>к</strong><strong>а</strong>х<br />

или <strong>в</strong> п<strong>а</strong>утин<strong>о</strong><strong>в</strong>ых труб<strong>к</strong><strong>а</strong>х-убежищ<strong>а</strong>х.<br />

В и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии из<strong>в</strong>е<strong>с</strong>тны <strong>о</strong>б<strong>а</strong> пер<strong>в</strong>ых<br />

п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>.<br />

32 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

(Tetrapneumones, Orthognatha)<br />

ПОДОТРЯД NELIPODA<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>в</strong>ершины<br />

л<strong>а</strong>п<strong>к</strong>и нет пуч<strong>к</strong><strong>о</strong><strong>в</strong> при<strong>к</strong>репительных щетин<strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 1426В). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Ctenizidae,<br />

Migidae, Atypidae, Dipluridae, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> п<strong>о</strong><strong>с</strong>леднее<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

497<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО DIPLURIDAE POCOCK, 1894<br />

П<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> <strong>о</strong>т д<strong>в</strong>ух д<strong>о</strong> трех п<strong>а</strong>р,<br />

из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых з<strong>а</strong>дняя <strong>о</strong>чень длинн<strong>а</strong>я. Хелицеры<br />

без «гр<strong>а</strong>бель». Прядут тенет<strong>а</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<br />

<strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ний дере<strong>в</strong>ье<strong>в</strong> или <strong>к</strong>у<strong>с</strong>т<strong>о</strong><strong>в</strong>,

<strong>а</strong> т<strong>а</strong><strong>к</strong>же между <strong>к</strong><strong>а</strong>мнями. Б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью мел<strong>к</strong>ие<br />

ф<strong>о</strong>рмы. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

<strong>о</strong>дин <strong>в</strong>ид р<strong>о</strong>д<strong>а</strong> Clostes Menge, 1869 (ри<strong>с</strong>. 1426).<br />

ПОДОТРЯД HYPODEMATA.<br />

НАСТОЯЩИЕ ПТИЦЕЯДЫ<br />

(Theraphosina)<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и пуч<strong>к</strong><strong>о</strong>м<br />

щетин<strong>о</strong><strong>к</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Barychelidae и Therapho-<br />

sidae; п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО THERAYHOS1DAE THORELL, 1869<br />

Д<strong>в</strong>е п<strong>а</strong>ры п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>: з<strong>а</strong>дняя из<br />

них <strong>о</strong>чень длинн<strong>а</strong>я, трехчлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>а</strong>я. П<strong>о</strong><strong>с</strong>ледний<br />

члени<strong>к</strong> з<strong>а</strong>дних б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> т<strong>а</strong><strong>к</strong><strong>о</strong>й же длины,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> и предп<strong>о</strong><strong>с</strong>ледний. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> де<strong>с</strong>ят<strong>к</strong><strong>о</strong><strong>в</strong> р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

Из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

<strong>о</strong>пи<strong>с</strong><strong>а</strong>н <strong>в</strong>ымерший р<strong>о</strong>д Eodiplurina Petrunkevitch,<br />

1922.<br />

ОТРЯД ARANEOMORPHAE. НАСТОЯЩИЕ ПАУКИ<br />

Брюш<strong>к</strong><strong>о</strong> <strong>в</strong>нешне не <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е. П<strong>а</strong>утинные<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong><strong>о</strong>нце тел<strong>а</strong>. Хелицеры,<br />

<strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, ди<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льные. Педип<strong>а</strong>льпы<br />

<strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итыми челю<strong>с</strong>тными л<strong>о</strong>п<strong>а</strong><strong>с</strong>тями.<br />

Передняя п<strong>а</strong>р<strong>а</strong> <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong> дых<strong>а</strong>ния<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>а</strong> лег<strong>к</strong>ими, з<strong>а</strong>дняя — тр<strong>а</strong>хеями;<br />

и<strong>с</strong><strong>к</strong>лючение <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют <strong>в</strong>иды <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Нур<strong>о</strong>chilidae<br />

<strong>с</strong> четырьмя лег<strong>к</strong>ими; <strong>в</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>е Telemidae<br />

лег<strong>к</strong>ие з<strong>а</strong>менены тр<strong>а</strong>хеями, <strong>а</strong> у пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> лег<strong>о</strong>чные меш<strong>к</strong>и<br />

м<strong>о</strong>гут <strong>в</strong><strong>о</strong><strong>о</strong>бще <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ть. Очень шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong>пр<strong>о</strong><strong>с</strong>тр<strong>а</strong>ненные п<strong>а</strong>у<strong>к</strong>и, дел<strong>а</strong>ющие п<strong>а</strong>утинные<br />

тенет<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне. Д<strong>в</strong><strong>а</strong> п<strong>о</strong>д<strong>о</strong>тряд<strong>а</strong>:<br />

Ecribellatae и Cribellatae.<br />

ПОДОТРЯД ECRIBELLATAE<br />

Между з<strong>а</strong>дними п<strong>а</strong>утинными б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми<br />

<strong>с</strong>ит<strong>о</strong><strong>в</strong>идн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> — cribellum — <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х н<strong>о</strong>г IV нет ряд<strong>а</strong> <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>бр<strong>а</strong>зующих calamistrum. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Д<strong>в</strong><strong>а</strong> инфр<strong>а</strong><strong>о</strong>тряд<strong>а</strong>: Haplogynae и Entelogynae.<br />

ИНФРАОТРЯД HAPLOGYNAE<br />

Ше<strong>с</strong>ть п<strong>а</strong>р гл<strong>а</strong>з. Семяприемни<strong>к</strong>и <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я<br />

неп<strong>о</strong><strong>с</strong>ред<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> <strong>в</strong><strong>о</strong> <strong>в</strong>л<strong>а</strong>г<strong>а</strong>лищ<strong>а</strong>.<br />

Муж<strong>с</strong><strong>к</strong><strong>о</strong>й <strong>к</strong><strong>о</strong>пуляти<strong>в</strong>ный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, без<br />

<strong>к</strong>р<strong>о</strong><strong>в</strong>еприемни<strong>к</strong><strong>а</strong> и л<strong>о</strong>д<strong>о</strong>ч<strong>к</strong>и. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

Д<strong>в</strong><strong>а</strong> н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Dysderidea и Caponiidea из<br />

них <strong>в</strong>т<strong>о</strong>р<strong>о</strong>е лишь <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне.<br />

НАДСЕМЕЙСТВО DYSDERIDEA<br />

(Dysderiformia)<br />

Н<strong>а</strong> <strong>к</strong>р<strong>а</strong>ю жел<strong>о</strong>б<strong>к</strong><strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> хелицер<br />

пл<strong>а</strong><strong>с</strong>тин<strong>к</strong>и (lamella) <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Пять<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне: Oonopidae,<br />

Dysderidae, Segestriidae, Leptonetidae, Telemidae,<br />

из них д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>ледних <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

неиз<strong>в</strong>е<strong>с</strong>тны.<br />

(Labidognatha, Dipneumonomorphae)<br />

498<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО OONOPIDAE SIMON, 1892<br />

От<strong>в</strong>ер<strong>с</strong>тия тр<strong>а</strong>хей <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>з<strong>а</strong>ди <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий<br />

лег<strong>к</strong>их. Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти <strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я<br />

<strong>в</strong>переди нижней губы. Н<strong>а</strong> <strong>в</strong>ершин<strong>а</strong>х л<strong>а</strong>п<strong>о</strong><strong>к</strong> п<strong>о</strong><br />

д<strong>в</strong><strong>а</strong> или п<strong>о</strong> три <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>. П<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены н<strong>а</strong> <strong>о</strong>бщем <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии. С<strong>о</strong><strong>в</strong>ременные<br />

<strong>в</strong>иды жи<strong>в</strong>ут п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями или <strong>с</strong>реди р<strong>а</strong><strong>с</strong>тительных<br />

<strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> <strong>в</strong><strong>с</strong>ех м<strong>а</strong>тери<strong>к</strong><strong>а</strong>х. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее де<strong>с</strong>яти<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) Orchestina Simon, 1892<br />

(ри<strong>с</strong>. 1427).<br />

СЕМЕЙСТВО DYSDERIDAE С. L. КОСН, 1837<br />

От<strong>в</strong>ер<strong>с</strong>тия тр<strong>а</strong>хей <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>з<strong>а</strong>ди лег<strong>о</strong>чных<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий. Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти длинные,<br />

п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельные. Нижняя губ<strong>а</strong> удлиненн<strong>а</strong>я.<br />

Имеет<strong>с</strong>я ше<strong>с</strong>ть гл<strong>а</strong>з, или гл<strong>а</strong>з<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

С<strong>о</strong><strong>в</strong>ременные <strong>в</strong>иды <strong>в</strong>едут п<strong>о</strong>лубр<strong>о</strong>дячий<br />

<strong>о</strong>бр<strong>а</strong>з жизни, ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>в</strong><strong>с</strong>треч<strong>а</strong>ют<strong>с</strong>я п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями<br />

и п<strong>о</strong>д <strong>к</strong><strong>о</strong>р<strong>о</strong>й дере<strong>в</strong>ье<strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 19 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Harpactes<br />

Templeton, 1834 (ри<strong>с</strong>. 1428А —Г); Thereola<br />

Petrunkevitch, 1955 (= Thera Koch u. Berend,<br />

1854) (ри<strong>с</strong>. 1429); Dysdera Latreille, 1804<br />

(ри<strong>с</strong>. 1430).<br />

СЕМЕЙСТВО SEGESTRIIDAE<br />

PETRUNKEVITCH, 1933<br />

Близ<strong>к</strong>и <strong>к</strong> предыдущему <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>у. Ше<strong>с</strong>ть<br />

гл<strong>а</strong>з. Стернум <strong>о</strong><strong>в</strong><strong>а</strong>льный, <strong>с</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong>м н<strong>а</strong> з<strong>а</strong>днем<br />

<strong>к</strong>р<strong>а</strong>е; <strong>о</strong>н <strong>о</strong>бычн<strong>о</strong> бы<strong>в</strong><strong>а</strong>ет чет<strong>к</strong><strong>о</strong> <strong>в</strong>иден бл<strong>а</strong>г<strong>о</strong>д<strong>а</strong>ря<br />

нежн<strong>о</strong><strong>с</strong>ти <strong>о</strong><strong>к</strong>руж<strong>а</strong>ющих п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong><strong>в</strong>. Л<strong>а</strong>п<strong>к</strong>и<br />

<strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1431). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>, из них<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Segestria<br />

Latreille, 1804.

Ри<strong>с</strong>. 1427—1431. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Oonopidae, Dysderidae и Segestriidae<br />

1427. Orchestina baltica Petrunkevitch; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у,<br />

X6,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>к</strong><strong>о</strong>нец <strong>с</strong>переди, Х10.0; В —н<strong>о</strong>г<strong>а</strong> IV, Х15;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942).<br />

1428. Harpactes extinctus Petrunkevitch; A — <strong>о</strong>бщий <strong>в</strong>ид, x7,0;<br />

Б — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, х20,0; В — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и<br />

н<strong>о</strong>ги III, X20.0; Г — п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и, X 20,0 п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1950). 1429. Thereo-<br />

ИНФРАОТРЯД ENTELOGYNAE<br />

У б<strong>о</strong>льшин<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong>ид<strong>о</strong><strong>в</strong> имеет<strong>с</strong>я <strong>в</strong><strong>о</strong><strong>с</strong>емь гл<strong>а</strong>з,<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong>. Семяприемни<strong>к</strong>и<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я н<strong>а</strong>ружу <strong>о</strong><strong>с</strong><strong>о</strong>быми <strong>с</strong><strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упительными<br />

<strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тиями (и<strong>с</strong><strong>к</strong>лючение <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют<br />

<strong>в</strong>иды н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Pholcidea и <strong>с</strong>ем. Теtragnathidae.<br />

Имеет<strong>с</strong>я х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит<strong>а</strong>я эпигин<strong>а</strong>.<br />

С<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упительный <strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong>л<strong>о</strong>жный.<br />

К<strong>о</strong>нце<strong>в</strong><strong>о</strong>й члени<strong>к</strong> педип<strong>а</strong>льп пре<strong>в</strong>р<strong>а</strong>щен <strong>в</strong> л<strong>о</strong>д<strong>о</strong>ч<strong>к</strong>у.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

11 н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, из них де<strong>в</strong>ять из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь): Pholcidea,<br />

Zodariidea, Hersiliidea, Argiopidea, Ly-<br />

499<br />

1428Г<br />

la petiolata (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид,. X4,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch et Berendt, 1854). 1430.,Dysderatersa<br />

Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид, x2,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch et Berendt, 1854). 1431. Segestria iomentoza<br />

Koch et Berendt;j <strong>о</strong>бщий <strong>в</strong>ид, X 3,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch et Berendt; 1854)<br />

cosidea, Drassodidea, Clubionidea, Thomisidea,<br />

Salticidea; т<strong>о</strong>ль<strong>к</strong><strong>о</strong> ныне — Ammoxenidea,' Homalonychidea.<br />

НАДСЕМЕЙСТВО PHOLCIDEA<br />

(Pholciformia)<br />

Трих<strong>о</strong>б<strong>о</strong>трий н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Хелицеры<br />

<strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>с</strong><strong>о</strong>единены <strong>о</strong>бщей мембр<strong>а</strong>н<strong>о</strong>й;<br />

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й их члени<strong>к</strong> <strong>с</strong> <strong>к</strong>рупным <strong>о</strong><strong>с</strong>трым<br />

зубц<strong>о</strong>м, <strong>к</strong><strong>о</strong>т<strong>о</strong>рый <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> п<strong>о</strong>д<strong>в</strong>ижным <strong>к</strong><strong>о</strong>гте<strong>о</strong>бр<strong>а</strong>зным<br />

<strong>к</strong><strong>о</strong>нце<strong>в</strong>ым члени<strong>к</strong><strong>о</strong>м <strong>о</strong>бр<strong>а</strong>зует <strong>к</strong>лешне<strong>в</strong>идный<br />

з<strong>а</strong>жим. Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти <strong>с</strong>ближены<br />

(ри<strong>с</strong>. 1432). Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Pholcidae.<br />

http://jurassic.ru/<br />

32*

СЕМЕЙСТВО PHOLCIDAE WALCKENAER,<br />

1837<br />

Обычн<strong>о</strong> <strong>в</strong><strong>о</strong><strong>с</strong>емь, реже ше<strong>с</strong>ть те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближенных<br />

гл<strong>а</strong>з. Нижняя губ<strong>а</strong> <strong>с</strong>лит<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>терн<strong>а</strong>льным<br />

щит<strong>к</strong><strong>о</strong>м. У мн<strong>о</strong>гих <strong>в</strong>ид<strong>о</strong><strong>в</strong> <strong>в</strong>переди п<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия<br />

имеет<strong>с</strong>я б<strong>о</strong>льш<strong>о</strong>й щит. Тр<strong>а</strong>хеи <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

С<strong>о</strong><strong>в</strong>ременные <strong>в</strong>иды плетут тенет<strong>а</strong> <strong>в</strong> пр<strong>о</strong><strong>с</strong><strong>в</strong>ет<strong>а</strong>х<br />

дупел, <strong>в</strong>ыб<strong>о</strong>ин<strong>а</strong>х <strong>с</strong><strong>к</strong><strong>а</strong>л и <strong>в</strong> других пу<strong>с</strong>т<strong>о</strong>т<strong>а</strong>х.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 33 р<strong>о</strong>д<strong>а</strong> и б<strong>о</strong>лее 220<br />

1435А<br />

1435Б<br />

1435Г<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong>. В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

<strong>о</strong>дин <strong>в</strong>ид р<strong>о</strong>д<strong>а</strong> Micropholcus Petrunkevitch<br />

(ри<strong>с</strong>. 1432).<br />

НАДСЕМЕЙСТВО ZODARIIDEA<br />

(Zodariiformia)<br />

З<strong>а</strong>дняя п<strong>а</strong>р<strong>а</strong> п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> н<strong>а</strong>мн<strong>о</strong>г<strong>о</strong><br />

меньше передних; ч<strong>а</strong><strong>с</strong>т<strong>о</strong> <strong>о</strong>ни <strong>с</strong>ильн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны<br />

или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>у-<br />

Ри<strong>с</strong>. 1432—1436. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Pholcidae, Zodariidae, Spatiatoridae, Urocteidae, Hersiliidae<br />

1432. Micropholcus heteropus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, X4,5;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942).<br />

1433. Annlculus balticus Petrunkevitch; A — <strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong>и<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong>ы<br />

педип<strong>а</strong>льп, Х25.0; Б — <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong>, Х1500; В — пер<strong>о</strong><strong>в</strong>идные<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и члени<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>о</strong>г, Х50,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1434. Spatlator praeceps<br />

Petrunkevitch; A — <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong>, x 35,0; Б — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть,<br />

X 20,0; В — брюш<strong>к</strong><strong>о</strong> <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х30,0; Г — н<strong>о</strong>г<strong>а</strong> II, Х12,0;<br />

Д — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, х15; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий ян­<br />

500<br />

http://jurassic.ru/<br />

Н34В<br />

1434Д<br />

т<strong>а</strong>рь) (Petrunkevitch, 1942). 1435. Paruroctea blauvelll Petrunkevitch;<br />

А — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>с</strong><strong>а</strong>мц<strong>а</strong>, Х15.0; Б — <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong>,<br />

Х15,0; В —<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х 15,0;<br />

Г — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги III, х20,0; Д — п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и,<br />

X25.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1436. Hersilla mlranda Koch und Berendt; <strong>о</strong>бщий <strong>в</strong>ид,<br />

ХЗ; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854)

не д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Palpimanidae и Zodariidae,<br />

из них п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> <strong>в</strong> б<strong>а</strong>лтий<strong>с</strong><strong>к</strong><strong>о</strong>м янт<strong>а</strong>ре,<br />

<strong>о</strong>т<strong>к</strong>уд<strong>а</strong>, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>о</strong>пи<strong>с</strong><strong>а</strong>н<strong>о</strong> <strong>в</strong>ымершее <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Spatiatoridae.<br />

СЕМЕЙСТВО ZODARIIDAE SIMON, 1892<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> тремя, реже <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми.<br />

Кр<strong>а</strong>й жел<strong>о</strong>б<strong>к</strong><strong>а</strong> н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии хелицер без зубц<strong>о</strong><strong>в</strong>,<br />

реже <strong>с</strong> <strong>о</strong>дним зубц<strong>о</strong>м. Colulus <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует.<br />

С<strong>о</strong><strong>в</strong>ременные <strong>в</strong>иды — п<strong>а</strong>у<strong>к</strong>и-<strong>о</strong>х<strong>о</strong>тни<strong>к</strong>и, плетущие<br />

меш<strong>к</strong><strong>о</strong><strong>в</strong>идные л<strong>о</strong>г<strong>о</strong><strong>в</strong>ищ<strong>а</strong> или не дел<strong>а</strong>ющие<br />

гнезд<strong>а</strong>; не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>в</strong>иды жи<strong>в</strong>ут <strong>в</strong> н<strong>о</strong>р<strong>к</strong><strong>а</strong>х<br />

<strong>с</strong> д<strong>в</strong>ерцей. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне <strong>с</strong>емь р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) <strong>в</strong>ымерший р<strong>о</strong>д Anniculus Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1433 А —Б).<br />

СЕМЕЙСТВО SPATIATORIDAE PETRUNKEVITCH,<br />

1942<br />

Срединн<strong>а</strong>я ч<strong>а</strong><strong>с</strong>ть лб<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>г<strong>о</strong> щит<strong>а</strong><br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>ыд<strong>а</strong>ет<strong>с</strong>я; <strong>с</strong><strong>а</strong>м щит <strong>в</strong> ф<strong>о</strong>рме <strong>в</strong>ытянут<strong>о</strong>г<strong>о</strong><br />

<strong>о</strong><strong>в</strong><strong>а</strong>л<strong>а</strong> <strong>с</strong> п<strong>о</strong>чти п<strong>а</strong>р<strong>а</strong>ллельными <strong>с</strong>т<strong>о</strong>р<strong>о</strong>н<strong>а</strong>ми.<br />

Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены тремя групп<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1434Д). О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния хелицер леж<strong>а</strong>т п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

друг другу, <strong>с</strong> уз<strong>к</strong>им <strong>к</strong>р<strong>а</strong>е<strong>в</strong>ым р<strong>а</strong><strong>с</strong>ширением,<br />

без <strong>в</strong>здутия. Scopula имеет<strong>с</strong>я. М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные<br />

<strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>а</strong>льп шир<strong>о</strong><strong>к</strong>ие, пятиуг<strong>о</strong>льные,<br />

<strong>о</strong>бле<strong>к</strong><strong>а</strong>ют <strong>с</strong> б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong> нижнюю губу<br />

(ри<strong>с</strong>. 1434). Н<strong>о</strong>ги <strong>с</strong> длинн<strong>о</strong>й п<strong>а</strong>телл<strong>о</strong>й; л<strong>а</strong>п<strong>к</strong>и<br />

их <strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми; имеют<strong>с</strong>я трих<strong>о</strong>б<strong>о</strong>трий.<br />

Colulus имеет<strong>с</strong>я или <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. П<strong>а</strong>ле<strong>о</strong>ген.<br />

Д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь): Spatiator Petrunkevitch, 1942 (ри<strong>с</strong>.<br />

1434 А —Д) и Adorator Petrunkevitch, 1942.<br />

НАДСЕМЕЙСТВО HERSILIIDEA<br />

(Hersiliaeformia)<br />

З<strong>а</strong>дние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и чрез<strong>в</strong>ыч<strong>а</strong>йн<strong>о</strong><br />

длинные и п<strong>о</strong>д<strong>в</strong>ижные. Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

<strong>с</strong>х<strong>о</strong>дят<strong>с</strong>я. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне:<br />

Urocteidae и Hersiliidae из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО UROCTEIDAE SIMON, 1875<br />

Хелицеры <strong>с</strong>л<strong>а</strong>бые, у <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны<br />

друг <strong>с</strong> друг<strong>о</strong>м мембр<strong>а</strong>н<strong>о</strong>й. Ан<strong>а</strong>льн<strong>а</strong>я <strong>к</strong>рышеч<strong>к</strong><strong>а</strong><br />

<strong>к</strong>рупн<strong>а</strong>я, делит<strong>с</strong>я н<strong>а</strong> д<strong>в</strong><strong>а</strong> <strong>о</strong>тдел<strong>а</strong> <strong>в</strong>енчи<strong>к</strong><strong>о</strong>м<br />

длинных желези<strong>с</strong>тых <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong>дин р<strong>о</strong>д; <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>в</strong>ымерший<br />

р<strong>о</strong>д Paruroctea Petrunkevitch, 1942 (ри<strong>с</strong>. 1435<br />

А-Д).<br />

501<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО HERSILIIDAE THORELL, 1869<br />

Передняя ч<strong>а</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

<strong>в</strong>здут<strong>а</strong>я, <strong>в</strong>ы<strong>с</strong><strong>о</strong><strong>к</strong><strong>а</strong>я (ри<strong>с</strong>. 1436). Длинные з<strong>а</strong>дние<br />

п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и н<strong>а</strong> <strong>в</strong>нутренней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

<strong>с</strong> д<strong>в</strong>умя непр<strong>а</strong><strong>в</strong>ильными ряд<strong>а</strong>ми п<strong>а</strong>утинных<br />

труб<strong>о</strong>че<strong>к</strong>. Colulus <strong>с</strong>ильн<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит. Бы<strong>с</strong>тр<strong>о</strong> бег<strong>а</strong>ют<br />

п<strong>о</strong> <strong>к</strong><strong>о</strong>ре дере<strong>в</strong>ье<strong>в</strong>, п<strong>о</strong>йм<strong>а</strong>нных н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>мых<br />

з<strong>а</strong>м<strong>а</strong>ты<strong>в</strong><strong>а</strong>ют пуч<strong>к</strong><strong>а</strong>ми п<strong>а</strong>утинных нитей. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Четыре р<strong>о</strong>д<strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне;<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

<strong>о</strong>дин <strong>с</strong><strong>о</strong><strong>в</strong>ременный р<strong>о</strong>д — Hersilia Audouin,<br />

1827 (ри<strong>с</strong>. 1436) и <strong>о</strong>дин <strong>в</strong>ымерший — Gerdia<br />

Menge, 1869.<br />

НАДСЕМЕЙСТВО ARANEIDEA<br />

(Argiopiformia)<br />

Трих<strong>о</strong>б<strong>о</strong>трий н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. П<strong>а</strong>утинные<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; <strong>о</strong>ни<br />

те<strong>с</strong>н<strong>о</strong> <strong>с</strong><strong>в</strong>яз<strong>а</strong>ны и <strong>в</strong>ме<strong>с</strong>те <strong>с</strong> colulus и <strong>а</strong>н<strong>а</strong>льным<br />

буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м <strong>о</strong>бр<strong>а</strong>зуют р<strong>о</strong>зет<strong>к</strong>у. Н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящие тенетные<br />

п<strong>а</strong>у<strong>к</strong>и. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне де<strong>с</strong>ять <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Nesticidae, Symphytognathidae,<br />

Theridiidae, Archaeidae, Mimetidae,<br />

Micryphantidae, Linyphiidae, Araneidae, Tetragnathidae,<br />

Ephalmatoridae, из них пер<strong>в</strong>ые<br />

д<strong>в</strong><strong>а</strong> <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тны.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Mithridae и<br />

Adjutoridae.<br />

СЕМЕЙСТВО THERIDIIDAE SUNDEVALL, 1833<br />

Г<strong>о</strong>лени и мет<strong>а</strong>т<strong>а</strong>рз<strong>а</strong>льные члени<strong>к</strong>и н<strong>о</strong>г без<br />

шип<strong>о</strong><strong>в</strong>. Н<strong>а</strong> нижней <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не л<strong>а</strong>п<strong>о</strong><strong>к</strong> з<strong>а</strong>дних н<strong>о</strong>г<br />

ше<strong>с</strong>ть — де<strong>с</strong>ять из<strong>о</strong>гнутых щетин<strong>о</strong><strong>к</strong>, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые<br />

и<strong>с</strong>п<strong>о</strong>льзуют<strong>с</strong>я при з<strong>а</strong>бр<strong>а</strong><strong>с</strong>ы<strong>в</strong><strong>а</strong>нии д<strong>о</strong>бычи <strong>к</strong>лей<strong>к</strong>ими<br />

нитями. Педип<strong>а</strong>льпы <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> без <strong>а</strong>п<strong>о</strong>физ<strong>о</strong><strong>в</strong><br />

н<strong>а</strong> г<strong>о</strong>лени и <strong>к</strong><strong>о</strong>лене (ри<strong>с</strong>. 1439 Б). В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>:<br />

Theridion Walcken-aer, 1805 (ри<strong>с</strong>. 1437) и Steatoda<br />

Sundevall, 1833 (ри<strong>с</strong>. 1438А, Б). Кр<strong>о</strong>ме<br />

т<strong>о</strong>г<strong>о</strong>, <strong>в</strong> т<strong>о</strong>й же ф<strong>а</strong>уне <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>в</strong>ымерших р<strong>о</strong>д<strong>о</strong><strong>в</strong>:<br />

Flegia Koch et Berendt, 1854 (ри<strong>с</strong>. 1439A, Б);<br />

Clya Koch et Berendt, 1854 (ри<strong>с</strong>. 1440); Mizalia<br />

Koch et Berendt, 1854 (ри<strong>с</strong>. 1441); Clythia<br />

Koch et Berend, 1954 (ри<strong>с</strong>. 1442); Eodipoena<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1443); Nactodipoena<br />

Petrunkevitch, 1942, (ри<strong>с</strong>. 1444); Municeps Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1445); Eomysmena Petrunkevitch<br />

1942 (ри<strong>с</strong>. 1446A— Г).<br />

СЕМЕЙСТВО ARCHAEIDAE KOCH, 1837<br />

Кр<strong>а</strong>й жел<strong>о</strong>б<strong>к</strong><strong>а</strong> хелицер <strong>с</strong> длинными шип<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1447 А, Б). Передняя ч<strong>а</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди<br />

<strong>с</strong>переди <strong>с</strong>ильн<strong>о</strong> прип<strong>о</strong>днят<strong>а</strong>, <strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие чег<strong>о</strong>

502<br />

http://jurassic.ru/

<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния хелицер и т<strong>а</strong>зи<strong>к</strong>и педип<strong>а</strong>льп зн<strong>а</strong>чительн<strong>о</strong><br />

уд<strong>а</strong>лены друг <strong>о</strong>т друг<strong>а</strong> (ри<strong>с</strong>. 1447). О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

члени<strong>к</strong> хелицер <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> длинный. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне южн<strong>о</strong>г<strong>о</strong><br />

п<strong>о</strong>луш<strong>а</strong>рия д<strong>в</strong><strong>а</strong> р<strong>о</strong>д<strong>а</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) р<strong>о</strong>д Archaea Koch<br />

et Berendt, 1854 (ри<strong>с</strong>. 1447А, Б).<br />

СЕМЕЙСТВО MIMETIDAE SIMON, 1895<br />

Хелицеры п<strong>о</strong>х<strong>о</strong>жи н<strong>а</strong> хелицеры предыдущег<strong>о</strong><br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>. Мет<strong>а</strong>т<strong>а</strong>рз<strong>а</strong>льные члени<strong>к</strong>и, <strong>а</strong> ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

и г<strong>о</strong>лени пер<strong>в</strong>ых д<strong>в</strong>ух п<strong>а</strong>р н<strong>о</strong>г имеют ряды<br />

длинных, <strong>с</strong>лег<strong>к</strong><strong>а</strong> из<strong>о</strong>гнутых шип<strong>о</strong><strong>в</strong>, между <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я б<strong>о</strong>лее мел<strong>к</strong>ие шипи<strong>к</strong>и,<br />

з<strong>а</strong>гнутые н<strong>а</strong> <strong>к</strong><strong>о</strong>нц<strong>а</strong>х. Л<strong>о</strong><strong>в</strong>чих <strong>с</strong>етей не дел<strong>а</strong>ют.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне четыре р<strong>о</strong>д<strong>а</strong>, из них <strong>в</strong><br />

п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) р<strong>о</strong>д<br />

Ег<strong>о</strong> С. L. Koch, 1837 (ри<strong>с</strong>. 1448).<br />

СЕМЕЙСТВО MICRYPHANT1DAE BERTKAU, 1885<br />

(Erigonidae Simon, 1926)<br />

Брюш<strong>к</strong><strong>о</strong> мяг<strong>к</strong><strong>о</strong>е, неб<strong>о</strong>льш<strong>о</strong>е. Н<strong>о</strong>ги без шип<strong>о</strong><strong>в</strong>.<br />

Б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ые гл<strong>а</strong>з<strong>а</strong> <strong>с</strong><strong>о</strong>единяют<strong>с</strong>я друг <strong>с</strong> друг<strong>о</strong>м.<br />

Л<strong>о</strong>б не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong>здут и прип<strong>о</strong>днят. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

<strong>в</strong> з<strong>а</strong>дней п<strong>о</strong>л<strong>о</strong><strong>в</strong>ине не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong>ширен<strong>а</strong>. Н<strong>а</strong><br />

пер<strong>в</strong><strong>о</strong>м и чет<strong>в</strong>ерт<strong>о</strong>м мет<strong>а</strong>т<strong>а</strong>рз<strong>а</strong>льных члени<strong>к</strong><strong>а</strong>х<br />

имеют<strong>с</strong>я трих<strong>о</strong>б<strong>о</strong>трий. Верхний <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>о</strong><strong>к</strong> л<strong>а</strong>п<strong>о</strong><strong>к</strong><br />

пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й, т<strong>о</strong>н<strong>к</strong>ий, <strong>с</strong> пятью нежными шипи<strong>к</strong><strong>а</strong>ми.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее де<strong>с</strong>яти р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из<br />

них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

три р<strong>о</strong>д<strong>а</strong>: Micryphantes Koch, 1875 (ри<strong>с</strong>. 1449);<br />

Erigone Savigny (ри<strong>с</strong>. 1450); Eogonatium Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1451).<br />

СЕМЕЙСТВО ADJ.UTORIDAE PETRUNKEVITCH, 1942<br />

Трих<strong>о</strong>б<strong>о</strong>трий н<strong>а</strong> г<strong>о</strong>ленях и мет<strong>а</strong>т<strong>а</strong>рз<strong>а</strong>льных<br />

члени<strong>к</strong><strong>а</strong>х л<strong>а</strong>п<strong>о</strong><strong>к</strong> <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>о</strong>чень длинные, <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>о</strong><strong>в</strong>идные<br />

(ри<strong>с</strong>. 1454). Colulus х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>ит.<br />

Ан<strong>а</strong>льный <strong>в</strong>ы<strong>с</strong>туп <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идный (ри<strong>с</strong>. 1452).<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит р<strong>а</strong><strong>с</strong>ширен <strong>в</strong> з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти,<br />

<strong>с</strong> п<strong>о</strong>лу<strong>к</strong>ругл<strong>о</strong>й б<strong>о</strong>р<strong>о</strong>зд<strong>о</strong>й <strong>в</strong> <strong>с</strong>ередине, <strong>о</strong>тделяющей<br />

не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong><strong>о</strong>з<strong>в</strong>ыш<strong>а</strong>ющую<strong>с</strong>я переднюю<br />

ч<strong>а</strong><strong>с</strong>ть. Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> (ри<strong>с</strong>.<br />

1452, 1454). Хелицеры <strong>с</strong> т<strong>о</strong>н<strong>к</strong>им <strong>к</strong>р<strong>а</strong>ем, н<strong>о</strong> без<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong>. Педип<strong>а</strong>льпы т<strong>о</strong>н<strong>к</strong>ие и <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong><br />

длинные (ри<strong>с</strong>. 1454). Н<strong>о</strong>ги <strong>с</strong> шип<strong>а</strong>ми, без scopu-<br />

1437. Theridion hirtum Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и,<br />

X2,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1438. Steatoda succini Petrunkevitch; A — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, X20.0; Б — п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и, Х25.0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1439. Flegia<br />

longimana Koch et Berendt; А — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x3,0; Б —<br />

т<strong>о</strong> же, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь и педип<strong>а</strong>льп<strong>а</strong>, х 20,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1950). 1440. Clya lugubrts<br />

Koch et Berendt; <strong>с</strong><strong>а</strong>мец (ряд<strong>о</strong>м из<strong>о</strong>бр<strong>а</strong>жен передний <strong>к</strong>р<strong>а</strong>й <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong><br />

<strong>с</strong> гл<strong>а</strong>з<strong>а</strong>ми), х2,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt, 1854). 1441. Mlzalla pilosula Koch et Berendt;<br />

<strong>о</strong>бщий <strong>в</strong>ид, X2,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

lae, <strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми, <strong>к</strong><strong>о</strong>т<strong>о</strong>рые ин<strong>о</strong>гд<strong>а</strong> м<strong>о</strong>гут<br />

быть зн<strong>а</strong>чительн<strong>о</strong> редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны. Длин<strong>а</strong> тел<strong>а</strong><br />

1—1,8 мм. П<strong>а</strong>ле<strong>о</strong>ген. В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) три р<strong>о</strong>д<strong>а</strong>: Adjutor Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1452); Admissor Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1453); Adjunctor Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1454).<br />

СЕМЕЙСТВО LINYPHIIDAE LATREILLE, 1804<br />

Трих<strong>о</strong>б<strong>о</strong>трий н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х и бедр<strong>а</strong>х н<strong>о</strong>г <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют.<br />

Гл<strong>а</strong>з<strong>а</strong> уд<strong>а</strong>лены <strong>о</strong>т переднег<strong>о</strong> <strong>к</strong>р<strong>а</strong>я г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди<br />

(ри<strong>с</strong>. 1455). Передний <strong>к</strong>р<strong>а</strong>й нижней губы<br />

<strong>в</strong>здут. Кр<strong>а</strong>й жел<strong>о</strong>б<strong>к</strong><strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния хелицер <strong>с</strong><strong>о</strong><br />

мн<strong>о</strong>гими зубц<strong>а</strong>ми. Педип<strong>а</strong>льпы <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>с</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong><br />

р<strong>а</strong>з<strong>в</strong>итым paracimbium (ри<strong>с</strong>. 1456). П<strong>а</strong>ле<strong>о</strong>ген—<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong><strong>к</strong><strong>о</strong>л<strong>о</strong> 40 р<strong>о</strong>д<strong>о</strong><strong>в</strong>;<br />

из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене С. Амери<strong>к</strong>и р<strong>о</strong>д Ыпуphia<br />

Latreille, 1804 (ри<strong>с</strong>. 1455). Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong><br />

т<strong>о</strong>й же ф<strong>а</strong>уне <strong>в</strong><strong>о</strong><strong>с</strong>емь <strong>в</strong>ымерших р<strong>о</strong>д<strong>о</strong><strong>в</strong>: Eopopiп<strong>о</strong><br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1456); Malleator<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1457А, Б); Custodela<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1458); Meditrina Petrunkevitsh,<br />

1942 (ри<strong>с</strong>. 1459 А, Б); Obnisus<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1460 А, Б); Liticen Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1461 А, Б); Mystagogus<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1462 А, Б); Impulsor<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1463 А, Б).<br />

Ри<strong>с</strong> 1437—1446. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Theridiidae<br />

СЕМЕЙСТВО MITHRAEIDAE KOCH ET BERENDT,<br />

1854<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь груше<strong>в</strong>идн<strong>а</strong>я <strong>с</strong><strong>о</strong> <strong>с</strong>л<strong>а</strong>б<strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим<br />

л<strong>о</strong>бным <strong>к</strong>р<strong>а</strong>ем, н<strong>а</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

д<strong>в</strong><strong>а</strong> <strong>к</strong>рупных гл<strong>а</strong>з<strong>а</strong>; <strong>о</strong><strong>с</strong>т<strong>а</strong>льные гл<strong>а</strong>з<strong>а</strong> <strong>с</strong>д<strong>в</strong>инуты<br />

н<strong>а</strong>з<strong>а</strong>д и леж<strong>а</strong>т н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не передней<br />

ч<strong>а</strong><strong>с</strong>ти <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong>. Н<strong>а</strong> г<strong>о</strong>ленях и <strong>к</strong><strong>о</strong>ленях<br />

единичные трих<strong>о</strong>б<strong>о</strong>трий. У <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>о</strong>чень <strong>к</strong>рупные<br />

<strong>к</strong><strong>о</strong>нце<strong>в</strong>ые члени<strong>к</strong>и педип<strong>а</strong>льп (ри<strong>с</strong>. 1464).<br />

В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong>дин<br />

р<strong>о</strong>д Androgeus Koch et Berendt, 1854.<br />

СЕМЕЙСТВО ARANEIDAE LEACH, 1819<br />

(Argiopidae Simon, 1892; Epeiridae<br />

Sundevall, 1833)<br />

П<strong>а</strong>у<strong>к</strong>и-<strong>к</strong>руг<strong>о</strong>пряды близ<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны предыдущим<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>м Theridiidae и Linyphiidae.<br />

Кр<strong>а</strong>я жел<strong>о</strong>б<strong>к</strong><strong>а</strong> н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниях их хелицер <strong>с</strong> зуб-<br />

503<br />

http://jurassic.ru/<br />

und Berendt, 1854). 1442. Clythla alma Koch et Berendt; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, x3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

und Berendt, 1854). 1443. Eodlpoena balttca Petrunkevitch; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong><br />

<strong>с</strong>б<strong>о</strong><strong>к</strong>у, x4,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1946). 1444. Nactodlpoena tnfulata (Koch et Berendt);<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x3,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt, 1864). 1445. Munlceps pulcher Petrunkevitch,<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги III, X20.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1955). 1446. Eomysmena morttura Petrunkevitch;<br />

A — <strong>с</strong><strong>а</strong>мец <strong>с</strong>б<strong>о</strong><strong>к</strong>у, x5,0; Б — т<strong>о</strong> же <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

Х5,0;В — педип<strong>а</strong>льп<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, Х15.0; Г — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, Х15.0п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1946)

14БЗ<br />

Ри<strong>с</strong>. 1447—1454. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archaeidae,<br />

1447. Archaea paradoxa Koch et Berendt; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и,<br />

X2,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, Х15.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1448. Ег<strong>о</strong> setulosa Koch;<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, Х3,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt; 1854).^ 1449. Micryphantes regularls Koch<br />

et Berendt; <strong>о</strong>бщий <strong>в</strong>ид, x2,5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt; 1854). 1450. Erigone stigmatosa Koch et<br />

Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1954). 1451. Eogonatium robustum<br />

504<br />

http://jurassic.ru/<br />

т<strong>а</strong><br />

Mimetidae, Micryphantidae, Adjutoridae<br />

Petrunkevitch; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, X20.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1946). 1452. Adjutor mirabilis<br />

Petrunkevitch; A — брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X7,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>а</strong>м<strong>к</strong>и,<br />

х7,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942), 1453. Admissor aculeatus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и<br />

<strong>с</strong>б<strong>о</strong><strong>к</strong>у, х9,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1454. Adjunctor similis Petrunkevitch; A — <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong><br />

<strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х8,0; Б — н<strong>о</strong>г<strong>а</strong> II <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х5,0, п<strong>а</strong>ле<strong>о</strong>ген»<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942)

ц<strong>а</strong>ми (ри<strong>с</strong>. 1470). Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми;<br />

п<strong>о</strong>д <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х п<strong>о</strong><strong>с</strong>ледней п<strong>а</strong>ры н<strong>о</strong>г,<br />

<strong>к</strong>р<strong>о</strong>ме зубч<strong>а</strong>тых щетин<strong>о</strong><strong>к</strong>, имеет<strong>с</strong>я еще т<strong>о</strong>л<strong>с</strong>тый<br />

шип (ри<strong>с</strong>. 1466 В). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) р<strong>о</strong>д Zilla<br />

С. L. Koch (ри<strong>с</strong>. 1465) и <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене С. Амери<strong>к</strong>и<br />

р<strong>о</strong>ды Nephila Leach, 1815 и Epeira Walckenaer,<br />

1805.<br />

Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong>ымершие р<strong>о</strong>ды из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong><br />

С. Амери<strong>к</strong>и: Tethneus Scudder, 1890, и 3. Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь); Eustaloides Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1466А —В); Geo. Koch et<br />

Berendt, 1854 (ри<strong>с</strong>. 1467); Acrometa Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1468); Theridiometa Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1469); Memoratrix Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1470A, Б); Elucus Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1471, 1472).<br />

СЕМЕЙСТВО TETRAGNATHIDAE SIMON, 1892<br />

Бедр<strong>а</strong> н<strong>о</strong>г <strong>с</strong> пр<strong>о</strong><strong>с</strong>тым и д<strong>в</strong><strong>о</strong>йным ряд<strong>о</strong>м трих<strong>о</strong>б<strong>о</strong>трий.<br />

Хелицеры длинные. Тел<strong>о</strong> удлиненн<strong>о</strong>е.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне пять<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене С. Амери<strong>к</strong>и р<strong>о</strong>д Tetragnatha<br />

Latreille, 1904; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, <strong>в</strong> т<strong>о</strong>й же<br />

ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших р<strong>о</strong>д<strong>а</strong>: Palaeometa Petrunkevitch,<br />

1922 и Palaeopachygnatha Petrunkevitch,<br />

1922.<br />

СЕМЕЙСТВО EPHALMATORIDAE<br />

PETRUNKEVITCH, 1950<br />

Н<strong>а</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не г<strong>о</strong>леней и мет<strong>а</strong>т<strong>а</strong>рз<strong>о</strong><strong>в</strong><br />

пер<strong>в</strong><strong>о</strong>й и <strong>в</strong>т<strong>о</strong>р<strong>о</strong>й п<strong>а</strong>р н<strong>о</strong>г имеют<strong>с</strong>я <strong>к</strong>рупные<br />

шил<strong>о</strong><strong>в</strong>идные щетин<strong>к</strong>и (ри<strong>с</strong>. 1473 А). Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х<br />

н<strong>о</strong>г три <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>, трих<strong>о</strong>б<strong>о</strong>трий единичны. Colulus<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует (ри<strong>с</strong>. 1473 Б). Хелицеры <strong>с</strong> <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми.<br />

Нижняя губ<strong>а</strong> <strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я, гл<strong>а</strong>д<strong>к</strong><strong>а</strong>я (ри<strong>с</strong>.<br />

1473 В); м<strong>а</strong><strong>к</strong><strong>с</strong>иллярные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> друг другу.<br />

Н<strong>о</strong>ги пр<strong>о</strong>гр<strong>а</strong>дные. Ше<strong>с</strong>ть п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong><br />

(ри<strong>с</strong>. 1473 Б). Н<strong>а</strong> п<strong>а</strong>льп<strong>а</strong>х <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> нет paracymlium<br />

и <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>о</strong><strong>в</strong> г<strong>о</strong>лени (ри<strong>с</strong>. 1473 А). П<strong>а</strong>ле<strong>о</strong>ген.<br />

Р<strong>о</strong>д Ephalmator Petrunkevitch, 1950, <strong>о</strong>дин<br />

<strong>в</strong>ид (ри<strong>с</strong>. 1473 А—В) <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

НАДСЕМЕЙСТВО LYCOSIDEA<br />

(Lycosformia)<br />

Н<strong>а</strong> н<strong>о</strong>г<strong>а</strong>х <strong>о</strong>дин-д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> трих<strong>о</strong>б<strong>о</strong>трий<br />

(ри<strong>с</strong>. 1480). П<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и т<strong>о</strong>л<strong>с</strong>тые, <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идные<br />

(ри<strong>с</strong>. 1476 Б). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>: Hahniidae,<br />

Oxyopidae, Senoculidae, Agelenidae, Argyronetidae,<br />

Lycosidae, Pisauridae, из них пер<strong>в</strong>ые три<br />

<strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тны, <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

<strong>о</strong>дн<strong>о</strong> <strong>в</strong>ымершее <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Insecutoridae.<br />

505<br />

СЕМЕЙСТВО AGELENIDAE WALCKENAER, 1805<br />

Н<strong>а</strong> <strong>к</strong>р<strong>а</strong>ях жел<strong>о</strong>б<strong>к</strong><strong>а</strong> <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии хелицер<br />

мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные зубцы. Л<strong>а</strong>п<strong>к</strong>и без scopulae.<br />

Г<strong>о</strong>лени педип<strong>а</strong>льп <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>с</strong> <strong>а</strong>п<strong>о</strong>физ<strong>а</strong>ми. В<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и<br />

н<strong>а</strong> теле б<strong>о</strong>льшей ч<strong>а</strong><strong>с</strong>тью пери<strong>с</strong>тые. Передние<br />

п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены, и т<strong>о</strong>гд<strong>а</strong><br />

<strong>о</strong>ни без colulus, <strong>а</strong> е<strong>с</strong>ли те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближены, т<strong>о</strong> colulus<br />

имеет<strong>с</strong>я. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Agelena Walckenaer,<br />

1805 (ри<strong>с</strong>. 1474) и Myro О. P. Cambridge,.<br />

1876 (ри<strong>с</strong>. 1475), <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из т<strong>о</strong>й же ф<strong>а</strong>уны<br />

из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong>ымершие р<strong>о</strong>ды Eocryphoeca Petrunkevitch,<br />

1946 ( = Tegenaria Koch et Berendt, 1854<br />

(ри<strong>с</strong>. 1476 А, Б); Thyelia Koch et Berendt, 1854<br />

(ри<strong>с</strong>. 1477); Textrix Sundevall, 1833 (ри<strong>с</strong>. 1478).<br />

СЕМЕЙСТВО ARGYRONETIDAE MENGE, 1869<br />

Близ<strong>к</strong>и <strong>к</strong> Agelenidae, н<strong>о</strong> неп<strong>а</strong>рн<strong>о</strong>е <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тие<br />

тр<strong>а</strong>хей р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен<strong>о</strong> д<strong>а</strong>ле<strong>к</strong><strong>о</strong> <strong>в</strong>переди <strong>о</strong>т п<strong>а</strong>утинных<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>. Н<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудн<strong>о</strong>м щите не<strong>с</strong><strong>к</strong><strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

пр<strong>а</strong><strong>в</strong>ильных ряд<strong>о</strong><strong>в</strong> <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1479). О<strong>с</strong>н<strong>о</strong><strong>в</strong>н<strong>о</strong>й члени<strong>к</strong> <strong>к</strong>рупных хелицер<br />

<strong>с</strong> четырьмя зубц<strong>а</strong>ми. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне <strong>о</strong>дин<br />

р<strong>о</strong>д, пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ленный и <strong>в</strong> ми<strong>о</strong>цене Е<strong>в</strong>р<strong>о</strong>пы: Агgyroneta<br />

Latreille, 1804.<br />

СЕМЕЙСТВО PISAURIDAE SIMON, 1897<br />

П<strong>а</strong>у<strong>к</strong>и, близ<strong>к</strong><strong>о</strong> р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енные <strong>к</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>м<br />

Agelenidae и Lycosidae. Н<strong>а</strong> д<strong>о</strong>р<strong>с</strong><strong>а</strong>льн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не<br />

л<strong>а</strong>п<strong>о</strong><strong>к</strong> мн<strong>о</strong>г<strong>о</strong>чи<strong>с</strong>ленные трих<strong>о</strong>б<strong>о</strong>трий, ч<strong>а</strong><strong>с</strong>т<strong>о</strong><br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> (ри<strong>с</strong>. 1480). В<strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и<br />

<strong>о</strong>бычн<strong>о</strong> пери<strong>с</strong>тые. Ди<strong>с</strong>т<strong>а</strong>льный <strong>к</strong>р<strong>а</strong>й <strong>в</strong>ертлуг<strong>о</strong><strong>в</strong><br />

<strong>с</strong> п<strong>о</strong>лулунн<strong>о</strong>й <strong>в</strong>ырез<strong>к</strong><strong>о</strong>й. Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

<strong>в</strong> три — четыре ряд<strong>а</strong> (ри<strong>с</strong>. 1481). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 12 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong>дин <strong>в</strong>ымерший<br />

р<strong>о</strong>д Esuritor Petrunkevitch, 1904 (ри<strong>с</strong>. 1480,<br />

1481).<br />

СЕМЕЙСТВО INSECUTORIDAE PETRUNKEVITCH,<br />

1942<br />

Хелицеры ди<strong>а</strong><strong>к</strong><strong>с</strong>и<strong>а</strong>льные (ри<strong>с</strong>. 1483). Д<strong>в</strong><strong>а</strong><br />

лег<strong>о</strong>чных меш<strong>к</strong><strong>а</strong>. Cribellum, calamistrum и colulus<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Имеет<strong>с</strong>я ше<strong>с</strong>ть п<strong>а</strong>утинных<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых передняя п<strong>а</strong>р<strong>а</strong> т<strong>о</strong>лще и<br />

длиннее з<strong>а</strong>дних. Ан<strong>а</strong>льный буг<strong>о</strong>р<strong>о</strong><strong>к</strong> д<strong>в</strong>учлени<strong>с</strong>тый,<br />

<strong>с</strong><strong>о</strong> щетин<strong>к</strong><strong>а</strong>ми. Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х scopulae <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует;<br />

три мел<strong>к</strong>их <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>. Н<strong>о</strong>ги длинные, <strong>с</strong><br />

шил<strong>о</strong><strong>в</strong>идными щетин<strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>. 1482 А). Трих<strong>о</strong>б<strong>о</strong>трий<br />

единичны — д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> н<strong>а</strong> г<strong>о</strong>лени и п<strong>о</strong><br />

<strong>о</strong>дн<strong>о</strong>му ряду н<strong>а</strong> т<strong>а</strong>рзе и мет<strong>а</strong>т<strong>а</strong>рзе. Хелицеры <strong>с</strong><br />

п<strong>а</strong>р<strong>а</strong>ллельными <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями и длинными <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 1483 В). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит <strong>с</strong><br />

http://jurassic.ru/

1463Б<br />

http://jurassic.ru/

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленн<strong>о</strong>й передней ч<strong>а</strong><strong>с</strong>тью (ри<strong>с</strong>. 1483). В<strong>о</strong><strong>с</strong>емь<br />

гл<strong>а</strong>з р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong>. Один р<strong>о</strong>д<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) —<br />

Insecutor Petrunkevitsh, 1942 (ри<strong>с</strong>. 1482 А, Б;<br />

1483 А, Б).<br />

НАДСЕМЕЙСТВО DRASSODIDEA<br />

(Drassiformia)<br />

Передние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>к</strong>рупные,<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены; имеет<strong>с</strong>я colulus. Хелицеры<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дные, их <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния не з<strong>а</strong><strong>к</strong>лючены <strong>в</strong><br />

<strong>о</strong>бщий футляр. В<strong>о</strong><strong>с</strong>емь гл<strong>а</strong>з <strong>в</strong> д<strong>в</strong>ух п<strong>о</strong>перечных<br />

ряд<strong>а</strong>х. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми, между <strong>к</strong><strong>о</strong>т<strong>о</strong>рыми<br />

пуч<strong>к</strong>и л<strong>о</strong>п<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идных <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong>. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Prodidomidae,<br />

Platoridae, Drassodidae; п<strong>о</strong><strong>с</strong>леднее из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО DRASSODIDAE PETRUNKEVITCH, 1942<br />

(Drassidae, Gnaphosidae)<br />

Передние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и зн<strong>а</strong>чительн<strong>о</strong><br />

<strong>к</strong>рупнее з<strong>а</strong>дних. Colulus <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х<br />

н<strong>о</strong>г д<strong>в</strong><strong>а</strong> з<strong>а</strong>зубренных <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>. Нижняя губ<strong>а</strong><br />

длинн<strong>а</strong>я. С<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упительные <strong>о</strong>рг<strong>а</strong>ны педип<strong>а</strong>льп<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> пр<strong>о</strong><strong>с</strong>тые. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 16 р<strong>о</strong>д<strong>о</strong><strong>в</strong>. В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong>бн<strong>а</strong>ружены д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших<br />

р<strong>о</strong>д<strong>а</strong> — Captrix Petrunkevitch, 1942 (ри<strong>с</strong>.<br />

1484) и Pytonyssa С. L. Koch, 1837 (ри<strong>с</strong>. 1485)<br />

и <strong>в</strong> <strong>о</strong>лиг<strong>о</strong>цене С. Амери<strong>к</strong>и <strong>в</strong>ымерший р<strong>о</strong>д Р<strong>а</strong>laeodrassus<br />

Petrunkevitch, 1942.<br />

НАДСЕМЕЙСТВО CLUBIONIDEA<br />

(Clubioniformia)<br />

Передние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>с</strong>ближены;<br />

colulus <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> пуч<strong>к</strong><strong>а</strong>ми л<strong>о</strong>п<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong>идных<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong> между <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми и <strong>с</strong><br />

д<strong>в</strong>умя ряд<strong>а</strong>ми трих<strong>о</strong>б<strong>о</strong>трие<strong>в</strong> и scopulae. Передние<br />

<strong>к</strong>рупные гл<strong>а</strong>з<strong>а</strong> п<strong>о</strong> <strong>в</strong>еличине не <strong>с</strong>ильн<strong>о</strong> <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>т других гл<strong>а</strong>з. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне четыре <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Ctenidae,<br />

Selenopidae, Clubionidae, Sparassidae; д<strong>в</strong><strong>а</strong> п<strong>о</strong><strong>с</strong>­<br />

Ри<strong>с</strong>. 1455—1464. Семей<strong>с</strong>п<br />

1455. Linyphla oblonga Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x2,5,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt;<br />

1854). 1456. Eopopino longipss Petrunkevitch; A — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, Xl5,0; Б — педип<strong>а</strong>льп<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, ХЮ,0; В — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и<br />

л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги II, х10,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь<br />

(Petrunkevitch, 1942). 1457. Matleator niger Petrunkevitch; A —<br />

г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>а</strong>мц<strong>а</strong>, X6,0; Б — педип<strong>а</strong>льп<strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong>, Х12,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch; 1942).<br />

1458. Custodela cheiracantha (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид,<br />

X2,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1459. Meditrina circumvallata Petrunkevitch; A — тел<strong>о</strong><br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, x5,5; Б —г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

Х10,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1460. Obnisus tenuipes Petrunkevitch; А — тел<strong>о</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong><br />

ледних <strong>о</strong>бн<strong>а</strong>ружены <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь); <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, д<strong>в</strong><strong>а</strong> <strong>в</strong>ымерших<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Inceptoridae и Parattidae,<br />

СЕМЕЙСТВО CLUBIONIDAE SIMON, 1895<br />

(Anyphaenidae Bertkau, 1878)<br />

Кр<strong>а</strong>й жел<strong>о</strong>б<strong>к</strong><strong>а</strong> н<strong>а</strong> хелицер<strong>а</strong>х зубч<strong>а</strong>тый (ри<strong>с</strong>.<br />

1490 Б). Передние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и длиннее<br />

з<strong>а</strong>дних (ри<strong>с</strong>. 1488). М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти<br />

т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп <strong>в</strong>ытянуты, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong> друг другу; длин<strong>а</strong> нижней губы<br />

б<strong>о</strong>льше ее ширины (ри<strong>с</strong>. 1490 Б). В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 35 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

С. Амери<strong>к</strong>и — Clubiona Latreille, 1804 (ри<strong>с</strong>.<br />

1486) и <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

— Anyphaena Sundevall, 1833 (ри<strong>с</strong>. 1487);<br />

Micarla L. Koch, 1875 (ри<strong>с</strong>. 1488); <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тны<br />

<strong>в</strong>ымершие р<strong>о</strong>ды Melaenophora С. L. Koch,<br />

1875 (ри<strong>с</strong>. 1489); Massula Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1490А, В); Ablator Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1491); Abliguritor Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1492А, Б); Desultor Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1493) и из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и — Eoversatrlx<br />

Petrunkevitch, 1922; Eobumbratrix Petrunkevitch,<br />

1922; Eostentatrix Petrunkevitch, 1922;<br />

507<br />

СЕМЕЙСТВО EUSPARASSIDAE PETRUNKEVITCH,<br />

1949<br />

(Sparassidae)<br />

Близ<strong>к</strong>и <strong>к</strong> Clubionidae. Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong>. Н<strong>а</strong> з<strong>а</strong>днем <strong>к</strong>р<strong>а</strong>е хелицер<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> жел<strong>о</strong>б<strong>к</strong><strong>а</strong><br />

не менее д<strong>в</strong>ух зубц<strong>о</strong><strong>в</strong>. Н<strong>о</strong>ги у мн<strong>о</strong>гих<br />

<strong>в</strong>ид<strong>о</strong><strong>в</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, <strong>к</strong><strong>а</strong><strong>к</strong> у п<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>в</strong>б<strong>о</strong><strong>к</strong><strong>о</strong>х<strong>о</strong>д<strong>о</strong><strong>в</strong><br />

(ри<strong>с</strong>. 1494—1496). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них<br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

р<strong>о</strong>ды Olios Walckenaer, 1805 (ри<strong>с</strong>. 1494); Heteropoda<br />

Latreille, 1804; Zachria С. L. Koch, 1875<br />

|(ри<strong>с</strong> 1495 А, Б) и <strong>в</strong>ымершие р<strong>о</strong>ды Sosybius<br />

Koch et Berenstein, 1854 (ри<strong>с</strong>. 1496); Collacteus<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1497A — В); Caduceator<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1498); Adulatrix<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1499); Eostaianus Pet-<br />

Linyphiidae и Mithraeidae<br />

<strong>с</strong>б<strong>о</strong><strong>к</strong>у, X2,5; Б — п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и, х10,0; В —<strong>к</strong><strong>о</strong>нце<strong>в</strong><strong>а</strong>я<br />

ч<strong>а</strong><strong>с</strong>ть ле<strong>в</strong><strong>о</strong>й педип<strong>а</strong>льпы <strong>с</strong><strong>а</strong>мц<strong>а</strong>, Х26.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1461. Liticen setosus Petrunkevitch;<br />

A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х5,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

<strong>с</strong><strong>в</strong>ерху, X 10,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1462. Mystagogus glaber Petrunkevitch; A — <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong><strong>в</strong>ерху, х 3,5; Б — педип<strong>а</strong>льп<strong>а</strong>, Х10.0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1463.<br />

Impulsor neglectus Petrunkevitch; <strong>с</strong><strong>а</strong>мец: A — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я<br />

<strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, х25,0; Б —педип<strong>а</strong>льпы <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

X15.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1464. Androgens mllitaris Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong>, x4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und<br />

Berendt, 1854)<br />

http://jurassic.ru/

1473А<br />

http://jurassic.ru/

Timkevitch, 1942 (ри<strong>с</strong>. 1500A, Б); Eostasina Petrunkevitch,<br />

1942 (рж. 1501 А, Б).<br />

СЕМЕЙСТВО INCEPTORIDAE PETRUNKEVITCH,<br />

1942<br />

Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудный щит яйце<strong>в</strong>идный, шир<strong>о</strong><strong>к</strong><strong>о</strong><br />

з<strong>а</strong><strong>к</strong>руглен н<strong>а</strong> <strong>о</strong>б<strong>о</strong>их <strong>к</strong><strong>о</strong>нц<strong>а</strong>х и н<strong>а</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>х (ри<strong>с</strong>.<br />

1503В). В<strong>о</strong><strong>с</strong>емь гл<strong>а</strong>з р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong><br />

<strong>в</strong>д<strong>о</strong>ль лб<strong>а</strong> (ри<strong>с</strong>. 1502). Хелицеры <strong>с</strong> шир<strong>о</strong><strong>к</strong>ими<br />

прям<strong>о</strong>уг<strong>о</strong>льными <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниями (ри<strong>с</strong>. 1503).<br />

М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп<br />

прям<strong>о</strong>уг<strong>о</strong>льные, уз<strong>к</strong>ие; нижняя губ<strong>а</strong> м<strong>а</strong>лень<strong>к</strong><strong>а</strong>я,<br />

<strong>с</strong><strong>в</strong><strong>о</strong>б<strong>о</strong>дн<strong>а</strong>я (ри<strong>с</strong>. 1503). Стернум <strong>о</strong><strong>в</strong><strong>а</strong>льный.<br />

Н<strong>о</strong>ги т<strong>о</strong>н<strong>к</strong>ие, длинные; л<strong>а</strong>п<strong>к</strong>и их <strong>с</strong> д<strong>в</strong>умя пр<strong>о</strong><strong>с</strong>тыми<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми; scopulae и р<strong>а</strong><strong>с</strong>ширенные щетин<strong>к</strong>и<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. У <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> педип<strong>а</strong>льпы <strong>с</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>о</strong>м.<br />

П<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> ше<strong>с</strong>ть, передняя<br />

п<strong>а</strong>р<strong>а</strong> их длиннее з<strong>а</strong>дних (ри<strong>с</strong>. 1503Г). Colulus<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы ('б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) р<strong>о</strong>д Inceptor Petrunkevitch, 1942 (ри<strong>с</strong>.<br />

1502, 1503 А —Г).<br />

СЕМЕЙСТВО PARATTIDAE<br />

PETRUNKEVITCH, 1922<br />

Гл<strong>а</strong>з<strong>а</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>г<strong>о</strong> ряд<strong>а</strong> зн<strong>а</strong>чительн<strong>о</strong> меньше<br />

гл<strong>а</strong>з переднег<strong>о</strong> ряд<strong>а</strong>. З<strong>а</strong>дний ряд рез<strong>к</strong><strong>о</strong> дуг<strong>о</strong><strong>в</strong>идный<br />

и б<strong>о</strong>лее <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий, чем передний. Гл<strong>а</strong>з<strong>а</strong><br />

переднег<strong>о</strong> ряд<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены примерн<strong>о</strong> н<strong>а</strong><br />

<strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>м р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>о</strong>янии. Д<strong>в</strong><strong>а</strong> <strong>с</strong>редних гл<strong>а</strong>з<strong>а</strong><br />

з<strong>а</strong>днег<strong>о</strong> ряд<strong>а</strong> <strong>с</strong>ближены. П<strong>а</strong>ле<strong>о</strong>ген. Олиг<strong>о</strong>цен<br />

С. Амери<strong>к</strong>и. Один р<strong>о</strong>д Parotitis Scudder, 1890.<br />

НАДСЕМЕЙСТВО THOMISIDEA<br />

(Thomisiformia)<br />

Передние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>с</strong>ближены.<br />

•Colulus <strong>в</strong><strong>с</strong>егд<strong>а</strong> имеет<strong>с</strong>я. Н<strong>а</strong> л<strong>а</strong>п<strong>к</strong><strong>а</strong>х <strong>о</strong>дин трих<strong>о</strong>б<strong>о</strong>трий<br />

или <strong>о</strong>дин ряд трих<strong>о</strong>б<strong>о</strong>трие<strong>в</strong>. Гл<strong>а</strong>з<strong>а</strong><br />

•б<strong>о</strong>лее или менее <strong>о</strong>дин<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>й <strong>в</strong>еличины (ри<strong>с</strong>.<br />

1506). Н<strong>о</strong>ги, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>лены <strong>в</strong> <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны,<br />

бегут б<strong>о</strong><strong>к</strong><strong>о</strong>м (ри<strong>с</strong>. 1506). П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. Д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне —<br />

Aphantochilidae и Thomisidae; п<strong>о</strong><strong>с</strong>леднее <strong>о</strong>бн<strong>а</strong>ружен<strong>о</strong><br />

<strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

Ри<strong>с</strong>. 1465—1473. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong><br />

1465. Zttla porrecta Koch et Berendt; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>, X3,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1466.<br />

Eustaloides succtni Petrunkevitch; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>: А — тел<strong>о</strong> <strong>с</strong>б<strong>о</strong><strong>к</strong>у, X7,0;<br />

Б — <strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, X 9,0; В — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги<br />

III, Х25.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1467. Geagpelroidea Koch et Berendt; <strong>с</strong><strong>а</strong>мец, x 3,0 (ряд<strong>о</strong>м<br />

из<strong>о</strong>бр<strong>а</strong>жены гл<strong>а</strong>з<strong>а</strong> н<strong>а</strong> переднем <strong>к</strong>р<strong>а</strong>е <strong>к</strong><strong>а</strong>р<strong>а</strong>п<strong>а</strong><strong>к</strong><strong>с</strong><strong>а</strong>); п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1468.<br />

Acrometa cristata Petrunkevitch, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й<br />

«т<strong>о</strong>р<strong>о</strong>ны, Х15.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1469. Therldlometa cdwardsi Petrunkevitch; м<strong>о</strong>л<strong>о</strong>д<strong>а</strong>я<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>, Хб,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Pet-<br />

СЕМЕЙСТВО THOMISIDAE SUNDEVALL, 1833<br />

К<strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и, <strong>к</strong><strong>а</strong><strong>к</strong> пр<strong>а</strong><strong>в</strong>ил<strong>о</strong>,<br />

те<strong>с</strong>н<strong>о</strong> <strong>с</strong>ближены между <strong>с</strong><strong>о</strong>б<strong>о</strong>й, <strong>а</strong> т<strong>а</strong><strong>к</strong>же <strong>с</strong><br />

colulus и <strong>а</strong>н<strong>а</strong>льным буг<strong>о</strong>р<strong>к</strong><strong>о</strong>м. Кр<strong>а</strong>й жел<strong>о</strong>б<strong>к</strong><strong>а</strong><br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии хелицер гл<strong>а</strong>д<strong>к</strong>ий или <strong>с</strong> <strong>о</strong>дним<br />

— тремя зубц<strong>а</strong>ми. Обе з<strong>а</strong>дние п<strong>а</strong>ры н<strong>о</strong>г<br />

<strong>о</strong>бычн<strong>о</strong> <strong>с</strong>л<strong>а</strong>бее и меньше передних н<strong>о</strong>г. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне б<strong>о</strong>лее 30 р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

<strong>в</strong> т<strong>о</strong>м чи<strong>с</strong>ле из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) Misumena Latreille, 1804 (ри<strong>с</strong>.<br />

1504); Philodromus Walckenaer, 1805 (ри<strong>с</strong>. 1505)<br />

и из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> С. Амери<strong>к</strong>и и не<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

р<strong>о</strong>д Thomisus Walckenaer, 1805. Кр<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>, из<br />

п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тны<br />

<strong>в</strong>ымершие р<strong>о</strong>ды: Siphax Koch et Berendt<br />

1854 (ри<strong>с</strong>. 1506): Fiducia Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1507); Eothanatus Petrunkevitch, 1950<br />

(ри<strong>с</strong>. 1508 А, Б); Facundia Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1509); Filiola Petrunkevitch, 1955 (ри<strong>с</strong>.<br />

1510 А, Б); Medela Petrunkevitch, 1942 (ри<strong>с</strong>.<br />

1511).<br />

НАДСЕМЕЙСТВО SALTICIDEA<br />

(Salticiformia)<br />

Передние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

ряд<strong>о</strong>м. Colulus <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> <strong>о</strong>дним<br />

— д<strong>в</strong>умя ряд<strong>а</strong>ми трих<strong>о</strong>б<strong>о</strong>трие<strong>в</strong>. Передние<br />

<strong>с</strong>рединные гл<strong>а</strong>з<strong>а</strong> не<strong>о</strong>быч<strong>а</strong>йн<strong>о</strong> <strong>в</strong>ели<strong>к</strong>и (ри<strong>с</strong>. 1512,<br />

1513). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

д<strong>в</strong><strong>а</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Lyssomanidae и Salticidae; п<strong>о</strong><strong>с</strong>леднее<br />

из<strong>в</strong>е<strong>с</strong>тн<strong>о</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО SALTICIDAE CAMBRIDGE, 1900<br />

ПАУКИ-СКАКУНЫ<br />

(Attidae Sundevall, 1833)<br />

В переднем ряду четыре гл<strong>а</strong>з<strong>а</strong>, <strong>а</strong> <strong>в</strong><strong>о</strong> <strong>в</strong>т<strong>о</strong>р<strong>о</strong>м<br />

и третьем ряд<strong>а</strong>х — п<strong>о</strong> д<strong>в</strong><strong>а</strong> (ри<strong>с</strong>. 1516 В). Кр<strong>а</strong>й<br />

хелицер<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> жел<strong>о</strong>б<strong>к</strong><strong>а</strong> <strong>с</strong> р<strong>а</strong>зличным <strong>в</strong><strong>о</strong><strong>о</strong>ружением<br />

(ри<strong>с</strong>. 1512А, 1513Б). В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне б<strong>о</strong>лее 50 р<strong>о</strong>д<strong>о</strong><strong>в</strong>; из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) из<strong>в</strong>е<strong>с</strong>тны <strong>в</strong>ымершие р<strong>о</strong>ды:<br />

Parevophrys Petrunkevitch 1942 (ри<strong>с</strong>.<br />

1512А — Д); Gorgopsina Petrunkevitch, 1955<br />

( = Gorgopsis Menge, 1854) (ри<strong>с</strong>. 1513A — Г);<br />

.raneidae и Ephalmatoridae<br />

runkevitch, 1942). 1470. Memoratrlx rydei Petrunkevitch; A — <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, х5,0; 15 —г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, Х12.0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942).<br />

1471. Elucus infelix Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид, х10,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1472. Elucus<br />

inermis Petrunkevitch; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>а</strong>мц<strong>а</strong>, х10,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1473. Ephalmator<br />

fossiles Petrunkevitch; <strong>с</strong><strong>а</strong>мец; А — <strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у, хЮ,0;<br />

Б — п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>с</strong> п<strong>а</strong>утинными нитями, Х7,0; В —<br />

<strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, х 10,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevich, 1950)<br />

509<br />

http://jurassic.ru/

1483Б<br />

Ри<strong>с</strong>. 1474—1483. П<strong>а</strong>у<strong>к</strong>и <strong>с</strong>емей<strong>с</strong>т<strong>в</strong> Agelenidae, Argyronetidae, Pisauridae, Insecutoridae<br />

1474. Agelena tabida Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, x2,5;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854).<br />

1475. Myro hirsutus Petrunkevitch; А — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, xl5,0;<br />

S — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, xl5,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) Petrunkevitch, 1942). 1476. Eocryphoeca distincta<br />

Petrunkevitch; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>; A — <strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, х 6,0;<br />

Б — п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и, X 20,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1950). 1477. Thyelia anomala Koch<br />

et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1478. Textrix funesta Koch et<br />

Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1479. Argyroneta aquatica<br />

510<br />

http://jurassic.ru/<br />

1483В<br />

Clerk; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, X15.0, <strong>с</strong><strong>о</strong><strong>в</strong>рем. (Crane, 1955). 1480. Esuritor<br />

spinipes Petrunkevitch; м<strong>о</strong>л<strong>о</strong>д<strong>а</strong>я <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>, н<strong>о</strong>г<strong>а</strong> IV, X6,0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942),<br />

1481. Esuritor spinipes Petrunkevitch; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>переди.<br />

Х7,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch.<br />

1942). 1482. Insecutor rufus Petrunkevitch; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>; A — <strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у,<br />

Х4,5; Б — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, Х20.0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1483. Insecutor<br />

mandibulatus Petrunkevitch; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>; A — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>в</strong>ерху,<br />

Хб,0; Б — т<strong>о</strong> же <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х6,0; В — т<strong>о</strong> же <strong>с</strong>переди, х 20,0;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942)

1484. Captrix lineata (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, x2,5,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1485. Pytonyssa affinis Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и,<br />

X3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt<br />

1854). 1486. Clublona sericea Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и<br />

X3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1487.Anyphaena fuscata Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>,<br />

X3.5, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1488. Mlcaria procera Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>[<br />

X3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt,<br />

1854). 1489. Melanophora mundulata Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong>, X4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und<br />

Berendt, 1854). 1490. Massula klebsi Petrunkevitch; <strong>с</strong><strong>а</strong>мец; A —<br />

14ЭЗА<br />

Ри<strong>с</strong>. 1484—1493. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Drassodidae и Clubionidae<br />

1493 Б<br />

<strong>в</strong>ид <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны <strong>с</strong> пр<strong>о</strong><strong>с</strong><strong>в</strong>ечи<strong>в</strong><strong>а</strong>ющими гр<strong>а</strong>ниц<strong>а</strong>ми <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong> и<br />

<strong>с</strong>тернум<strong>а</strong>, х12,0; Б —хелицеры, м<strong>а</strong><strong>к</strong><strong>с</strong>илярные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

педип<strong>а</strong>льп и нижняя губ<strong>а</strong> <strong>с</strong> брюшн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, хЗО.О, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1491. Ablator<br />

trlguttatus (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>,x3,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854) .1492.<br />

Abllgurltor nlger Petrunkevitch; <strong>с</strong><strong>а</strong>мец; A — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>в</strong>ерху<br />

X 15,0; Б — педип<strong>а</strong>льп<strong>а</strong>, Х25.0; В — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и и щетин<strong>к</strong>и н<strong>а</strong><br />

л<strong>а</strong>п<strong>к</strong>е н<strong>о</strong>ги III, х20,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Petrunkevitch, 1942). 1493. Desultor depressus Petrunkevitch,<br />

<strong>с</strong><strong>а</strong>мец; A — педип<strong>а</strong>льп<strong>а</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; Б — т<strong>о</strong> же, <strong>с</strong> брюшн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х35.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Petrunkevitch, 1942)<br />

511<br />

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1494. OUos crassipes (Koch et Berendt); <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X4,0;<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (K°ch und Berendt, 1854).<br />

1495. Zachria pecullata Petrunkevitch; A — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong><strong>в</strong>ерху,<br />

Х7,0; Б — л<strong>а</strong>п<strong>к</strong><strong>а</strong> н<strong>о</strong>ги IV, Х15,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1946). 1496. Sosybius minor Koch<br />

et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X,4,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1497. Collacteus aeptivus<br />

Petrunkevitch; А — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, x 15,0; Б — <strong>к</strong><strong>о</strong>г<strong>о</strong>ть хелицеры,<br />

Х35.0; В — передняя ч<strong>а</strong><strong>с</strong>ть <strong>с</strong>тернум<strong>а</strong>, нижняя губ<strong>а</strong> и<br />

м<strong>а</strong><strong>к</strong><strong>с</strong>илярные <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong>и т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп, X 25,0, п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ийянт<strong>а</strong>рь) (Petrunkevitch, 1942). 1498. Caducca-<br />

Ри<strong>с</strong>. 1494—1501. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eusparassidae<br />

512<br />

http://jurassic.ru/<br />

1500А<br />

tor quadrimaculatas Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х 6,0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,П1950).<br />

1499. Adulatrix fusca Petrunkevitch; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>переди,<br />

Х25.0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1500. Eostaianus succini Petrunkevitch; A — <strong>о</strong>бщий <strong>в</strong>ид<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong><strong>в</strong>ерху, Хб,5; Б — н<strong>о</strong>г<strong>а</strong> IV, X 15,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1950). 1501. Eostasina <strong>а</strong><strong>с</strong>иleata<br />

Petrunkevitch; А — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>низу, х 6,5; 5 —<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги IV, х20,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1942)

150?<br />

1608А<br />

1502. Inceptor dubius Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х 9,0,<br />

п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1946).<br />

1503. Inceptor aculeatus Petrunkevitch; A — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди,<br />

ХЮ.О; Б — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, 10,0; В — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

<strong>с</strong><strong>в</strong>ерху," х10,0; Г — <strong>а</strong>н<strong>а</strong>льный <strong>к</strong><strong>о</strong>ну<strong>с</strong> и п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и<br />

<strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х15.0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1504. Misumena samlandensis Petrunkevitch; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и,хб.О п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1505. Philodromus dubius Koch et Berendt; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>мц<strong>а</strong>, x3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch<br />

und Berendt, 1854). 1506. Syphax gracilis Koch et Berendt; <strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, x2,5; <strong>в</strong><strong>в</strong>ерху из<strong>о</strong>бр<strong>а</strong>жены гл<strong>а</strong>з<strong>а</strong> Х15.0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854). 1507.<br />

1503<br />

1510А<br />

1503В<br />

Ри<strong>с</strong>. 1502—1511. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Inceptoridae и Thomisidae<br />

33 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 513<br />

http://jurassic.ru/<br />

1503Б<br />

1511<br />

1508Б<br />

Fiducia tenuipes Petrunkevitch; <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть <strong>с</strong><strong>а</strong>м<strong>к</strong>и,<br />

Xl8,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1950). 1508. Eothanatus diritatis Petrunkevitch; A — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong>переди, х 15,0; Б — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и и <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и н<strong>а</strong> ч<strong>а</strong>п<strong>к</strong>е н<strong>о</strong>ги<br />

II <strong>с</strong><strong>а</strong>мц<strong>а</strong>, X 20,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1950). 1509. Facundia clara Petrunkevitch; <strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong> <strong>с</strong>переди,<br />

хб.О; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1510. Flliolaargentata Petrunkevitch; А — <strong>о</strong>бщий <strong>в</strong>ид.<strong>с</strong><strong>в</strong>ерху,<br />

X 5,0; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong>переди, х 10,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong><br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1511. Medela baltica<br />

Petrunkevitch; г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>переди, ХЮ,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942)

1б)6Б<br />

http://jurassic.ru/

Eolinus Petrunkevitch, 1942 (ри<strong>с</strong>. 1514, 1515);<br />

Paralinus Petrunkevitch, 1942 (ри<strong>с</strong>. 1516A —Г);<br />

Cenattus Petrunkevitch, 1942 (ри<strong>с</strong>. 1517); Pseudopropetes<br />

W. Dubinin, nom. nov.. (pro Propetes<br />

Menge, 1854, non Walker, 1851, Insecta); Phidippus<br />

Koch et Berendt, 1854 (ри<strong>с</strong>. 1518); Steneattus<br />

Bronn, 1856 (=Leda Koch et Berendt,<br />

1854, non Schumacher, 1817) (ри<strong>с</strong>. 1519);<br />

Eophrys Menge, 1854 (ри<strong>с</strong>. 1520) и из <strong>о</strong>лиг<strong>о</strong>цен<strong>а</strong><br />

Фр<strong>а</strong>нции — Eoattopsis Petrunkevitch, 1955<br />

(=Attopsis Gonrret, 1886, non Weer, 1849);<br />

Attoides Brongniart, 1901.<br />

ПОДОТРЯД CRIB ELL ATAE<br />

Между з<strong>а</strong>дними п<strong>а</strong>утинными б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong><strong>а</strong>ми<br />

имеет<strong>с</strong>я <strong>с</strong>ит<strong>о</strong><strong>в</strong>идн<strong>а</strong>я пл<strong>а</strong><strong>с</strong>тин<strong>к</strong><strong>а</strong> — cribellum. Н<strong>а</strong><br />

л<strong>а</strong>п<strong>к</strong><strong>а</strong>х н<strong>о</strong>г VI <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong>и, <strong>о</strong>бр<strong>а</strong>зующие calamistrum;<br />

<strong>о</strong>ни м<strong>о</strong>гут быть редуцир<strong>о</strong><strong>в</strong><strong>а</strong>ны у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых<br />

<strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>. Д<strong>в</strong><strong>а</strong> инфр<strong>а</strong><strong>о</strong>тряд<strong>а</strong>: Palaeocribellatae<br />

и Neocribellatae.<br />

ИНФРАОТРЯД<br />

PALAEOCRIBELLATAE<br />

Д<strong>в</strong>е п<strong>а</strong>ры <strong>в</strong>еер<strong>о</strong><strong>в</strong>идных лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющих<strong>с</strong>я н<strong>а</strong> <strong>с</strong>межных (VIII и IX) <strong>с</strong>егмент<strong>а</strong>х<br />

брюш<strong>к</strong><strong>а</strong>. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне: Hypochilidae.<br />

И НФРАОТРЯД<br />

NEOCRIBELLATAE<br />

Одн<strong>а</strong> п<strong>а</strong>р<strong>а</strong> лег<strong>о</strong>чных меш<strong>к</strong><strong>о</strong><strong>в</strong>; <strong>в</strong>т<strong>о</strong>р<strong>а</strong>я п<strong>а</strong>р<strong>а</strong><br />

дых<strong>а</strong>тельных <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тий <strong>с</strong>лит<strong>а</strong> <strong>в</strong> неп<strong>а</strong>рную<br />

щель, <strong>в</strong> глубине <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>й <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я тр<strong>а</strong>хеи.<br />

Ан<strong>а</strong>т<strong>о</strong>мия, <strong>к</strong><strong>а</strong><strong>к</strong> у типичных Araneomorphae. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. Ше<strong>с</strong>ть н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong> <strong>в</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне: Filistatidea, Oecobiidea, Zoropseidea,<br />

Eresidea, Dictynidea, Amaurobiidea,<br />

причем д<strong>в</strong><strong>а</strong> пер<strong>в</strong>ых неиз<strong>в</strong>е<strong>с</strong>тны <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии.<br />

НАДСЕМЕЙСТВО ZOROPSEIDEA<br />

(Zoropsidiformia)<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г <strong>с</strong> д<strong>в</strong>умя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми, пуч<strong>к</strong><strong>а</strong>ми<br />

при<strong>к</strong>репительных <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>о</strong><strong>в</strong> и <strong>с</strong><strong>о</strong> scopulae.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне д<strong>в</strong><strong>а</strong><br />

1512. Parevophrys succini Petrunkevitch; <strong>с</strong><strong>а</strong>мец; Д'—г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

<strong>с</strong>переди, х!5,0; Б — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, Х15,0;<br />

В — педип<strong>а</strong>льп<strong>а</strong>, х20,0; Г — передняя ч<strong>а</strong><strong>с</strong>ть г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди <strong>с</strong> гл<strong>а</strong>з<strong>а</strong>ми,<br />

Х7,0; Д — л<strong>а</strong>п<strong>к</strong><strong>а</strong> н<strong>о</strong>ги III, х 15,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1513. Gorgopsina frenata<br />

(Koch et Berendt); A — *бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X4,0; (Koch und<br />

Berendt, 1854); Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong><strong>а</strong>мц<strong>а</strong> <strong>с</strong>переди, Х15.0; В — <strong>а</strong>н<strong>а</strong>льный<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong> и п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, X 15,0; В — <strong>а</strong>н<strong>а</strong>льный<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong> и п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х15.0; Г — <strong>в</strong><strong>о</strong><strong>о</strong>ружение<br />

л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги III, х15,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Petrunkevitch, 1942). 1514. Eolinus succineus Petrunkevitch;<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong><strong>а</strong>; A —передний <strong>к</strong>р<strong>а</strong>й г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди, х10,0; Б —л<strong>а</strong>п<strong>к</strong><strong>а</strong><br />

н<strong>о</strong>ги IV, х15,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1515. Eolinus theryi Petrunkevitch; педип<strong>а</strong>льп<strong>а</strong><br />

Ри<strong>с</strong>. 1512—1520. Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Salticidae<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Zoropseidae и Acanthoctenidae, из<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых пер<strong>в</strong><strong>о</strong>е пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен<strong>о</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

СЕМЕЙСТВО ZOROPSEIDAE SIMON, 1892<br />

Передние н<strong>о</strong>ги <strong>с</strong> шип<strong>а</strong>ми и <strong>с</strong><strong>о</strong> scopulae н<strong>а</strong><br />

т<strong>а</strong>рз<strong>а</strong>льн<strong>о</strong>м, <strong>а</strong> ин<strong>о</strong>гд<strong>а</strong> и н<strong>а</strong> мет<strong>а</strong>т<strong>а</strong>рз<strong>а</strong>льн<strong>о</strong>м<br />

члени<strong>к</strong><strong>а</strong>х. Трих<strong>о</strong>б<strong>о</strong>трий е<strong>с</strong>ть. Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены<br />

<strong>в</strong> д<strong>в</strong><strong>а</strong>-три ряд<strong>а</strong>, причем <strong>в</strong> пер<strong>в</strong><strong>о</strong>м ряду<br />

<strong>в</strong><strong>с</strong>егд<strong>а</strong> четыре гл<strong>а</strong>з<strong>а</strong>. Cribellum м<strong>а</strong>лень<strong>к</strong>ий, р<strong>а</strong>зделен<br />

н<strong>а</strong> д<strong>в</strong>е п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины; calamistrum <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий.<br />

Крупные п<strong>а</strong>у<strong>к</strong>и, дел<strong>а</strong>ющие п<strong>о</strong>д <strong>к</strong><strong>а</strong>мнями шир<strong>о</strong><strong>к</strong>ие<br />

<strong>с</strong>ети <strong>с</strong> б<strong>о</strong>льшими непр<strong>а</strong><strong>в</strong>ильными петлями.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

три р<strong>о</strong>д<strong>а</strong>; <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

— <strong>в</strong>ымерший р<strong>о</strong>д Adamator Petrunkevitch,<br />

1942 (ри<strong>с</strong>. 1521).<br />

НАДСЕМЕЙСТВО ERESIDEA<br />

(Eresiformia)<br />

З<strong>а</strong>дние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие.<br />

П<strong>а</strong>рные <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>о</strong><strong>к</strong> нер<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>-зубч<strong>а</strong>тые.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong>.<br />

СЕМЕЙСТВО ERESIDAE С. L. КОСН, 1868<br />

Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> три ряд<strong>а</strong> (ри<strong>с</strong>. 1522).<br />

Хелицеры м<strong>о</strong>щные. Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

и <strong>с</strong> трих<strong>о</strong>б<strong>о</strong>триями. З<strong>а</strong>дние п<strong>а</strong>утинные б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>к</strong>и<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>че и т<strong>о</strong>ньше передних. С<strong>о</strong><strong>в</strong><strong>о</strong><strong>к</strong>упительный<br />

<strong>а</strong>пп<strong>а</strong>р<strong>а</strong>т <strong>с</strong><strong>а</strong>мц<strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й. П<strong>а</strong>ле<strong>о</strong>ген —<br />

ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>лен р<strong>о</strong>д Eresus Walckenaer,<br />

1805 (ри<strong>с</strong>. 1522).<br />

НАДСЕМЕЙСТВО DICTYNIDEA<br />

(Dictynaeformia)<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>в</strong><strong>с</strong>ех н<strong>о</strong>г без трих<strong>о</strong>б<strong>о</strong>трие<strong>в</strong>, <strong>с</strong> тремя<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния хелицер <strong>с</strong> зубц<strong>а</strong>ми<br />

(ри<strong>с</strong>. 1524 В). Мел<strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и, п<strong>о</strong> <strong>в</strong>нешн<strong>о</strong><strong>с</strong>ти<br />

н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ющие пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Theridiidae.<br />

П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне<br />

три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Dictynidae, Dinopidae, Uloboridae,<br />

пер<strong>в</strong><strong>о</strong>е из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>о</strong>бн<strong>а</strong>ружен<strong>о</strong> <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь).<br />

<strong>с</strong><strong>а</strong>мц<strong>а</strong>, хЮ,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1516. Paralinus crosbyl Petrunkevitch; <strong>с</strong><strong>а</strong>мец; A —<br />

<strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, х9,0; Б —п<strong>а</strong>льп<strong>а</strong> <strong>с</strong>б<strong>о</strong><strong>к</strong>у, хЗО.О; В —п<strong>а</strong>льп<strong>а</strong> <strong>с</strong>е<br />

<strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, Х30.0; Г — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>а</strong> <strong>с</strong>переди, Х8,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1517. Cenattus<br />

exophth almlcus Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong><strong>в</strong>ерху,<br />

х7,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1942). 1518. Phidippus paululus Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и,<br />

x2,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und<br />

Berendt, 1854). 1519. Steneattus promtssus (Koch et Berendt);<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X2,5; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt, 1854). 1520. Eophrys gibberulus Koch et Berendt;<br />

<strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, x3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Koch und Berendt, 1864)<br />

515 33*<br />

http://jurassic.ru/

1530В<br />

Ри<strong>с</strong>. 1521—1530. Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Zoropseidae, Eresidae, Dictynidae, Amaurobiidae, Arthrodictynidae и Psechridae<br />

1521. Adamator succienus Petrunkevitch; <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и, x 20,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

Petrunkevitch, 1942). 1522. Eresus monachus Koch et Berendt<strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, X3,0, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь)<br />

(Koch und Berendt, 1854). 1523. Ealathys succini Petrunkevitch<strong>о</strong>бщий<br />

<strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х9,0 п<strong>а</strong>ле<strong>о</strong>г<strong>о</strong>н, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1950). 1524. Eolathys debilis Petrunkevitch;<br />

A — <strong>к</strong><strong>о</strong><strong>к</strong><strong>с</strong><strong>о</strong><strong>с</strong>терн<strong>а</strong>льн<strong>а</strong>я <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть, xlO.O; Б — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

<strong>с</strong><strong>в</strong>ерху, х 10,0; В — г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong>переди, х 12,0; п<strong>а</strong>ле<strong>о</strong>ген,<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1950). 1525. Ат<strong>а</strong>иrobius<br />

rlmosus Koch et Berendt; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>а</strong>м<strong>к</strong>и, х4,0, п<strong>а</strong>ле<strong>о</strong>ген.<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Koch und Berendt, 1854).<br />

1526. Amaurobtus succini Petrunkevitch; A — передний <strong>к</strong><strong>о</strong>нец<br />

хелицеры, X 20,0; Б — <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги IV, х20,0 п<strong>а</strong>ле<strong>о</strong>ген<br />

516<br />

http://jurassic.ru/<br />

1530Г<br />

Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1527. Auximus<br />

succini Petrunkevitch; <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong> cri.<br />

bellum, X250, п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь (Petrunkevitch,<br />

1942). 1528. Auximus fossilis Petrunkevitch; <strong>о</strong>бщий <strong>в</strong>ид<br />

<strong>с</strong><strong>а</strong>м<strong>к</strong>и, х 15,0, п<strong>а</strong>л<strong>о</strong><strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) (Petrunkevitch,<br />

1950). 1529. Arthrodictyma segmentata Petrunkevitch;<br />

А — <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong>б<strong>о</strong><strong>к</strong>у, Х8,0; Б — брюш<strong>к</strong><strong>о</strong> <strong>с</strong>низу, X 8,0; В —<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong>и л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>ги III, х 10,0; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий<br />

янт<strong>а</strong>рь) (Petrunkevitch, 1942). 1530. Eomatachia latifrons Petrunkevitch;<br />

<strong>с</strong><strong>а</strong>мец; А— педип<strong>а</strong>льп<strong>а</strong>, x 15,0; Б — <strong>о</strong>бл<strong>а</strong><strong>с</strong>ть п<strong>а</strong>утинных<br />

б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>, х 20„0: В — р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жение гл<strong>а</strong>з н<strong>а</strong> лбу, х10,0;<br />

Г — нижняя губ<strong>а</strong> и м<strong>а</strong><strong>к</strong><strong>с</strong>илярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп,<br />

хЮ.О; п<strong>а</strong>ле<strong>о</strong>ген, Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь (Petrunkevitch,<br />

1942)

СЕМЕЙСТВО DICTYNIDAE SIMON, 1874<br />

Кр<strong>а</strong>й хелицер<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> жел<strong>о</strong>б<strong>к</strong><strong>а</strong> пр<strong>о</strong><strong>с</strong>т<strong>о</strong>й.<br />

В<strong>о</strong><strong>с</strong>емь гл<strong>а</strong>з р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> (ри<strong>с</strong>.<br />

1524 Б). Хелицеры у не<strong>к</strong><strong>о</strong>т<strong>о</strong>рых <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong><br />

<strong>в</strong>ид<strong>о</strong>изменены. Плетут тенет<strong>а</strong>. П<strong>а</strong>ле<strong>о</strong>ген<br />

— ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне 15 р<strong>о</strong>д<strong>о</strong><strong>в</strong>.<br />

В п<strong>а</strong>ле<strong>о</strong>гене Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>в</strong>ымерший<br />

р<strong>о</strong>д Eolathys Petrunkevitch, 1950 (ри<strong>с</strong>.<br />

1523, 1524А —В).<br />

НАДСЕМЕЙСТВО AMAUROBIIDEA<br />

(Amaurobiiformia)<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> <strong>о</strong>дним-д<strong>в</strong>умя ряд<strong>а</strong>ми трих<strong>о</strong>б<strong>о</strong>трие<strong>в</strong>.<br />

П<strong>о</strong> <strong>в</strong>нешнему <strong>в</strong>иду и не<strong>к</strong><strong>о</strong>т<strong>о</strong>рым <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тям<br />

<strong>с</strong>тр<strong>о</strong>ения н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей<br />

<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Agelenidae. П<strong>а</strong>ле<strong>о</strong>ген — ныне. В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й<br />

ф<strong>а</strong>уне три <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Amaurobiidae,<br />

Psechridae, Tengellidae; п<strong>о</strong><strong>с</strong>леднее <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии неиз<strong>в</strong>е<strong>с</strong>тн<strong>о</strong>; <strong>к</strong>р<strong>о</strong>ме т<strong>о</strong>г<strong>о</strong>,<br />

<strong>о</strong>дн<strong>о</strong> <strong>в</strong>ымершее <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Arthrodictynidae.<br />

СЕМЕЙСТВО AMAUROBIIDAE С. L. КОСН, 1868<br />

Гл<strong>а</strong>з<strong>а</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong>. Кр<strong>а</strong>я жел<strong>о</strong>б<strong>к</strong><strong>а</strong><br />

н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ниях хелицер <strong>с</strong> зубц<strong>а</strong>ми (ри<strong>с</strong>.<br />

1526). Челю<strong>с</strong>тные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти б<strong>о</strong>лее или менее<br />

п<strong>а</strong>р<strong>а</strong>ллельны. Cribellum р<strong>а</strong>зделен н<strong>а</strong> д<strong>в</strong>е п<strong>о</strong>л<strong>о</strong><strong>в</strong>ины.<br />

Л<strong>а</strong>п<strong>к</strong>и <strong>с</strong> <strong>о</strong>дним ряд<strong>о</strong>м трих<strong>о</strong>б<strong>о</strong>трие<strong>в</strong>,<br />

без scopulae (ри<strong>с</strong>. 1528). П<strong>а</strong>ле<strong>о</strong>ген — ныне. В<br />

<strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне пять р<strong>о</strong>д<strong>о</strong><strong>в</strong>, из них <strong>в</strong> п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) Ат<strong>а</strong>иг<strong>о</strong>bius<br />

С. L. Koch, 1837 (ри<strong>с</strong>. 1525, 1526А —Б)<br />

и Auximus Simon, 1892 (ри<strong>с</strong>. 1527, 1528).<br />

СЕМЕЙСТВО ARTHRODICTYNIDAE<br />

PETRUNKEVITCH, 1942<br />

Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong> <strong>с</strong>лед<strong>а</strong>ми <strong>с</strong>егмент<strong>а</strong>ции: <strong>в</strong> передней<br />

ч<strong>а</strong><strong>с</strong>ти ег<strong>о</strong> <strong>в</strong>ентр<strong>а</strong>льн<strong>о</strong>й п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти д<strong>в</strong><strong>а</strong><br />

<strong>с</strong>тернит<strong>а</strong>, п<strong>о</strong>з<strong>а</strong>ди них р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены шир<strong>о</strong><strong>к</strong>ий<br />

cribellum и ше<strong>с</strong>ть п<strong>а</strong>утинных б<strong>о</strong>р<strong>о</strong>д<strong>а</strong><strong>в</strong><strong>о</strong><strong>к</strong>, пер<strong>в</strong><strong>а</strong>я<br />

п<strong>а</strong>р<strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупн<strong>а</strong>я. Ан<strong>а</strong>льный <strong>к</strong><strong>о</strong>ну<strong>с</strong> шир<strong>о</strong><strong>к</strong>ий<br />

(ри<strong>с</strong>. 1529 Б). Хелицеры <strong>с</strong> неб<strong>о</strong>льшими<br />

р<strong>а</strong><strong>с</strong>ширениями. М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong><br />

педип<strong>а</strong>льп н<strong>а</strong><strong>к</strong>л<strong>о</strong>нены перед нижней губ<strong>о</strong>й.<br />

Стернум щит<strong>о</strong><strong>в</strong>идный (ри<strong>с</strong>. 1529 А). Л<strong>а</strong>п<strong>к</strong>и педип<strong>а</strong>льп<br />

<strong>с</strong> <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. Л<strong>а</strong>п<strong>к</strong>и н<strong>о</strong>г <strong>с</strong> тремя <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми<br />

(ри<strong>с</strong>. 1529 В). Calamistrum н<strong>а</strong> мет<strong>а</strong>т<strong>а</strong>рзе<br />

<strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из из<strong>о</strong>гнутых щетин<strong>о</strong><strong>к</strong>. П<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong>ы<br />

тел<strong>а</strong> <strong>с</strong> ред<strong>к</strong>ими, же<strong>с</strong>т<strong>к</strong>ими <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми (ри<strong>с</strong>.<br />

1529 А). Н<strong>а</strong> г<strong>о</strong>ленях, мет<strong>а</strong>т<strong>а</strong>рзу<strong>с</strong><strong>а</strong>х и т<strong>а</strong>рзу<strong>с</strong><strong>а</strong>х<br />

имеют<strong>с</strong>я длинные трих<strong>о</strong>б<strong>о</strong>трий. Мел<strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и;<br />

длин<strong>а</strong> брюш<strong>к</strong><strong>а</strong> 0,8 мм. П<strong>а</strong>ле<strong>о</strong>ген. В Е<strong>в</strong>р<strong>о</strong>пе<br />

(б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) <strong>о</strong>дин р<strong>о</strong>д — Arthrodictyna<br />

Petrunkevitch, 1942 (ри<strong>с</strong>. 1529А —В).<br />

СЕМЕЙСТВО PSECHRIDAE SIMON, 1892<br />

Крупные п<strong>а</strong>у<strong>к</strong>и <strong>с</strong> длинными передними н<strong>о</strong>г<strong>а</strong>ми.<br />

Тел<strong>о</strong> <strong>в</strong> длинных <strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>а</strong>х. Н<strong>а</strong>п<strong>о</strong>мин<strong>а</strong>ют<br />

пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Agelenidae. В<strong>о</strong><strong>с</strong>емь<br />

гл<strong>а</strong>з, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>женных <strong>в</strong> д<strong>в</strong><strong>а</strong> ряд<strong>а</strong> (ри<strong>с</strong>. 1530 В).<br />

Хелицеры м<strong>о</strong>щные, <strong>с</strong> зубц<strong>а</strong>ми н<strong>а</strong> <strong>к</strong>р<strong>а</strong>е жел<strong>о</strong>б<strong>к</strong><strong>а</strong><br />

и <strong>к</strong>рупным <strong>к</strong>рюч<strong>к</strong><strong>о</strong>м <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии ег<strong>о</strong>. Н<strong>а</strong><br />

мет<strong>а</strong>т<strong>а</strong>рзе и л<strong>а</strong>п<strong>к</strong>е имеют<strong>с</strong>я м<strong>а</strong>лень<strong>к</strong>ие scopu-<br />

1<strong>а</strong>. М<strong>а</strong><strong>к</strong><strong>с</strong>иллярные л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти т<strong>а</strong>зи<strong>к</strong><strong>о</strong><strong>в</strong> педип<strong>а</strong>льп<br />

з<strong>а</strong><strong>к</strong>руглены <strong>с</strong>переди, р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жены п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong><br />

друг другу (ри<strong>с</strong>. 1530 Г). П<strong>а</strong>ле<strong>о</strong>ген — ныне.<br />

В <strong>с</strong><strong>о</strong><strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне ше<strong>с</strong>ть р<strong>о</strong>д<strong>о</strong><strong>в</strong>. В п<strong>а</strong>ле<strong>о</strong>гене<br />

Е<strong>в</strong>р<strong>о</strong>пы (б<strong>а</strong>лтий<strong>с</strong><strong>к</strong>ий янт<strong>а</strong>рь) н<strong>а</strong>йден<br />

<strong>в</strong>ымерший р<strong>о</strong>д Eomatachia Petrunkevitch, 1942<br />

(ри<strong>с</strong>. 1530А — Г).<br />

http://jurassic.ru/

Андрее<strong>в</strong><strong>а</strong> О. Н. 1957. Н<strong>о</strong><strong>в</strong>ые н<strong>а</strong>х<strong>о</strong>д<strong>к</strong>и члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих<br />

<strong>в</strong> В<strong>о</strong><strong>с</strong>т<strong>о</strong>чн<strong>о</strong>й Сибири. Ежег<strong>о</strong>дн. В<strong>с</strong>е<strong>с</strong>. п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л. <strong>о</strong>б<strong>в</strong><strong>а</strong>,<br />

т. 16, <strong>с</strong>тр. 80—86.<br />

Бе<strong>к</strong>лемише<strong>в</strong> В. Н. 1952. О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>й<br />

<strong>а</strong>н<strong>а</strong>т<strong>о</strong>мии бе<strong>с</strong>п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных. Изд. «С<strong>о</strong><strong>в</strong>ет<strong>с</strong><strong>к</strong><strong>а</strong>я н<strong>а</strong>у<strong>к</strong><strong>а</strong>», М.,<br />

<strong>с</strong>тр. 1—698. Бе<strong>к</strong><strong>к</strong>ер Э. и У<strong>а</strong>рт<strong>о</strong>н Г. 1955. В<strong>в</strong>едение<br />

<strong>в</strong> <strong>а</strong><strong>к</strong><strong>а</strong>р<strong>о</strong>л<strong>о</strong>гию. Изд. ин<strong>о</strong><strong>с</strong>тр. лит-ры. М., <strong>с</strong>тр. 1—475.<br />

Б я л ы« и ц<strong>к</strong>'И й - Б я р у л я А. А. 1917. С<strong>к</strong><strong>о</strong>рпи<strong>о</strong>ны.<br />

Ф<strong>а</strong>ун<strong>а</strong> Р<strong>о</strong><strong>с</strong><strong>с</strong>ии. П<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные (Arachnida), т. I. Петр<strong>о</strong>гр<strong>а</strong>д,<br />

<strong>с</strong>тр. 1—227. Birula А. 1926. Zur ausseren Morphologie<br />

der fossilen und recenten Skorpione. Zool. Anz.,<br />

Bd. 67, S. 61—67.— 1938. Ф<strong>а</strong>л<strong>а</strong>нги (Solifuga). Ф<strong>а</strong>ун<strong>а</strong><br />

СССР. П<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, т. I, <strong>в</strong>ып. 3, <strong>с</strong>тр. 1—177.<br />

Дубинин В. Б. 1951. Перье<strong>в</strong>ые <strong>к</strong>лещи (Analgesoidea),<br />

ч. 1. Ф<strong>а</strong>ун<strong>а</strong>. СССР, т. 6, <strong>в</strong>. 5.—1954. Н<strong>о</strong><strong>в</strong><strong>а</strong>я<br />

<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong>ифи<strong>к</strong><strong>а</strong>ция перье<strong>в</strong>ых <strong>к</strong>лещей н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Analgesoidea<br />

и п<strong>о</strong>л<strong>о</strong>жение ег<strong>о</strong> <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме <strong>о</strong>тряд<strong>а</strong> Acariformes<br />

A. Zachv.— 1956. Отряды <strong>к</strong>лещей и п<strong>о</strong>л<strong>о</strong>жение их<br />

<strong>в</strong> <strong>с</strong>и<strong>с</strong>теме Chelicerata. Тр. II н<strong>а</strong>учн<strong>о</strong>й <strong>к</strong><strong>о</strong>нференции<br />

п<strong>а</strong>р<strong>а</strong>зит<strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong> УССР. Изд. З<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН УССР,<br />

Кие<strong>в</strong>, <strong>с</strong>тр. 6—51.— 1967. Об <strong>о</strong>риент<strong>а</strong>ции г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> у де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>их пи<strong>к</strong>н<strong>о</strong>г<strong>о</strong>н<strong>о</strong><strong>в</strong> р<strong>о</strong>д<strong>а</strong> Palaeoisopus<br />

и' п<strong>о</strong>л<strong>о</strong>жении их <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме Arthropoda. Д<strong>о</strong><strong>к</strong>л. АН<br />

СССР, т. 117, № 5, <strong>с</strong>тр. 881—884.— 1959<strong>а</strong>. Хелицер<strong>о</strong>н<strong>о</strong><strong>с</strong>ные<br />

жи<strong>в</strong><strong>о</strong>тные (п<strong>о</strong>дтип Chelicerophora W. Dubinin, nom.<br />

hov.) и п<strong>о</strong>л<strong>о</strong>жение их <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме. З<strong>о</strong><strong>о</strong>л. журя., т. 38,<br />

<strong>в</strong>ып. 8, <strong>с</strong>тр. 1163—1189.— 19596. О фил<strong>о</strong>генезе хелицер<strong>о</strong><strong>в</strong>ых<br />

жи<strong>в</strong><strong>о</strong>тных п<strong>о</strong>дтип<strong>а</strong> Chelicerophora W. Dub. и р<strong>о</strong>д-<br />

«т<strong>в</strong>е Chelicerata <strong>с</strong> Pycnogonides. Тр. Ин-т<strong>а</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия<br />

жи<strong>в</strong><strong>о</strong>тных АН СССР, <strong>в</strong>ып. 27, <strong>с</strong>тр. 134—150.<br />

З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин А. А. 1941. Тир<strong>о</strong>глиф<strong>о</strong>идные <strong>к</strong>лещи<br />

(Tyroglyphoidea). Ф<strong>а</strong>ун<strong>а</strong> СССР. П<strong>а</strong>у<strong>к</strong><strong>о</strong><strong>о</strong>бр<strong>а</strong>зные, т. 6,<br />

<strong>в</strong>ып. 2, <strong>с</strong>тр. 1—475.— 1952<strong>а</strong>. Р<strong>а</strong>зделение <strong>к</strong>лещей (Acarina)<br />

н<strong>а</strong> <strong>о</strong>тряды и их п<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме Chelicerata.<br />

П<strong>а</strong>р<strong>а</strong>зит<strong>о</strong>л<strong>о</strong>г, <strong>с</strong>б<strong>о</strong>'рн. З<strong>о</strong><strong>о</strong>л. ин-т<strong>а</strong> АН СССР, т. 14,<br />

<strong>с</strong>тр. 5—46.— 19526. Э<strong>в</strong><strong>о</strong>люция и м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>е <strong>о</strong>б<strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>с</strong>и<strong>с</strong>темы <strong>к</strong>лещей. Чтения п<strong>а</strong>мяти Н. А. Х<strong>о</strong>л<strong>о</strong>д<strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

з<strong>а</strong> 1951 г. Изд. АН СССР, <strong>с</strong>тр. 53—66,— 1953.<br />

И<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ния п<strong>о</strong> м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гии и п<strong>о</strong><strong>с</strong>тэм-бри<strong>о</strong>н<strong>а</strong>льн<strong>о</strong>му<br />

р<strong>а</strong>з<strong>в</strong>итию тир<strong>о</strong>глифид (Sarcoptiformes, Tyroglyphoidea).<br />

Сб. н<strong>а</strong>учн. р<strong>а</strong>б<strong>о</strong>т А. А. З<strong>а</strong>х<strong>в</strong><strong>а</strong>т<strong>к</strong>ин <strong>а</strong>. Изд. М<strong>о</strong><strong>с</strong><strong>к</strong>. ун-т<strong>а</strong>,<br />

М., <strong>с</strong>тр. 19—120.<br />

Китт<strong>а</strong>ры М. 1848. Ан<strong>а</strong>т<strong>о</strong>миче<strong>с</strong><strong>к</strong><strong>о</strong>е и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ние<br />

<strong>о</strong>бы<strong>к</strong>н<strong>о</strong><strong>в</strong>енн<strong>о</strong>й (Galeodes araneoides) и <strong>к</strong><strong>о</strong>лючей (G. dorsalis)<br />

<strong>с</strong><strong>о</strong>льпуги. Уч. з<strong>а</strong>п. К<strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>. ун-т<strong>а</strong>, № 2. К рейд<strong>о</strong><strong>в</strong><br />

<strong>с</strong><strong>к</strong>ий М. Е. 1885. Пре<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>о</strong>дные <strong>а</strong><strong>к</strong><strong>а</strong>риды Южн<strong>о</strong>й<br />

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НАДКЛАСС PYCNOGONIDES. МОРСКИЕ ПАУКИ<br />

(pantopoda)<br />

(\ В. Б. Дубинин \)<br />

С<strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>а</strong>я групп<strong>а</strong> м<strong>о</strong>р<strong>с</strong><strong>к</strong>их члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>- три ч<strong>а</strong><strong>с</strong>ти <strong>в</strong>ертлуг<strong>о</strong>м. О<strong>с</strong>т<strong>а</strong>льные три-четыре<br />

гих, <strong>с</strong><strong>о</strong>чет<strong>а</strong>ющ<strong>а</strong>я не<strong>к</strong><strong>о</strong>т<strong>о</strong>рые <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>ти р<strong>а</strong><strong>к</strong><strong>о</strong>­ п<strong>а</strong>ры х<strong>о</strong>дильных н<strong>о</strong>г (у Dodecalopoda имеет<strong>с</strong>я<br />

<strong>о</strong>бр<strong>а</strong>зных (х<strong>а</strong>р<strong>а</strong><strong>к</strong>тер н<strong>а</strong>ч<strong>а</strong>льных <strong>с</strong>т<strong>а</strong>дий др<strong>о</strong>бле­ д<strong>а</strong>же ше<strong>с</strong>ть п<strong>а</strong>р н<strong>о</strong>г) причленяют<strong>с</strong>я <strong>к</strong><strong>а</strong>жд<strong>а</strong>я н<strong>а</strong><br />

ния, н<strong>а</strong>личие н<strong>а</strong>упли<strong>а</strong>льн<strong>о</strong>г<strong>о</strong> гл<strong>а</strong>з<strong>а</strong> и <strong>о</strong><strong>с</strong><strong>о</strong>бых <strong>о</strong>тдельн<strong>о</strong>м <strong>с</strong>егменте, гр<strong>а</strong>ницы между <strong>к</strong><strong>о</strong>т<strong>о</strong>ры­<br />

л<strong>о</strong>бных <strong>о</strong>рг<strong>а</strong>н<strong>о</strong><strong>в</strong>) и н<strong>а</strong><strong>с</strong>т<strong>о</strong>ящих хелицер<strong>о</strong><strong>в</strong>ых ми ин<strong>о</strong>гд<strong>а</strong> и<strong>с</strong>чез<strong>а</strong>ют. Интере<strong>с</strong>н<strong>о</strong>, чт<strong>о</strong> <strong>в</strong> <strong>с</strong><strong>в</strong>язи <strong>с</strong><br />

(<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ие <strong>а</strong>нтеннул и deutocerebram <strong>в</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong>­ р<strong>а</strong>з<strong>в</strong>итием х<strong>о</strong>б<strong>о</strong>т<strong>а</strong> ни <strong>о</strong>дн<strong>а</strong> из <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей не<br />

<strong>в</strong>е н<strong>а</strong>дгл<strong>о</strong>т<strong>о</strong>чн<strong>о</strong>г<strong>о</strong> г<strong>а</strong>нглия, н<strong>а</strong>личие р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> имеет гн<strong>а</strong>т<strong>о</strong>б<strong>а</strong>з.<br />

<strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>а</strong>, <strong>в</strong>ид<strong>о</strong>изменение передних <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>в</strong>ременных п<strong>а</strong>нт<strong>о</strong>п<strong>о</strong>д рудимен­<br />

<strong>в</strong> хелицеры (хелиф<strong>о</strong>ры) и педип<strong>а</strong>льпы и т. п.). т<strong>а</strong>рн<strong>о</strong>е, нер<strong>а</strong><strong>с</strong>члененн<strong>о</strong>е, меш<strong>к</strong><strong>о</strong><strong>в</strong>идн<strong>о</strong>е, <strong>а</strong> у п<strong>а</strong>­<br />

М<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гия. Тел<strong>о</strong> п<strong>о</strong>др<strong>а</strong>зделяет<strong>с</strong>я н<strong>а</strong> ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их Palaeopantopoda брюш<strong>к</strong><strong>о</strong> длинн<strong>о</strong>е,<br />

х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>, г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь и брюш<strong>к</strong><strong>о</strong>. Одн<strong>а</strong><strong>к</strong><strong>о</strong> х<strong>о</strong>б<strong>о</strong>­ члени<strong>с</strong>т<strong>о</strong>е и <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>я<strong>в</strong>шее из пяти <strong>с</strong>егмент<strong>о</strong><strong>в</strong>.<br />

т<strong>о</strong><strong>к</strong> я<strong>в</strong>ляет<strong>с</strong>я лишь чрезмерным р<strong>а</strong>зр<strong>а</strong><strong>с</strong>т<strong>а</strong>нием В<strong>с</strong>ег<strong>о</strong>, <strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, тел<strong>о</strong> дре<strong>в</strong>них Palaeopan­<br />

<strong>к</strong>р<strong>а</strong>е<strong>в</strong> р<strong>о</strong>т<strong>о</strong><strong>в</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>т<strong>в</strong>ер<strong>с</strong>тия — р<strong>о</strong>т<strong>о</strong><strong>в</strong>ым <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong>м; topoda <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ял<strong>о</strong> из 14 <strong>с</strong>егмент<strong>о</strong><strong>в</strong>, из <strong>к</strong><strong>о</strong>т<strong>о</strong>рых у<br />

м<strong>о</strong>рф<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>о</strong>н <strong>в</strong>ентр<strong>а</strong>лен и не д<strong>о</strong>лжен <strong>с</strong><strong>о</strong><strong>в</strong>ременных Pantopoda <strong>с</strong><strong>о</strong>хр<strong>а</strong>нил<strong>о</strong><strong>с</strong>ь т<strong>о</strong>ль<strong>к</strong><strong>о</strong><br />

р<strong>а</strong><strong>с</strong><strong>с</strong>м<strong>а</strong>три<strong>в</strong><strong>а</strong>ть<strong>с</strong>я <strong>в</strong> <strong>к</strong><strong>а</strong>че<strong>с</strong>т<strong>в</strong>е <strong>о</strong>тдельн<strong>о</strong>й т<strong>а</strong>гмы <strong>с</strong>емь — де<strong>в</strong>ять. Эт<strong>о</strong>т пр<strong>о</strong>це<strong>с</strong><strong>с</strong> <strong>о</strong>лиг<strong>о</strong>мериз<strong>а</strong>ции<br />

тел<strong>а</strong>, не<strong>с</strong>м<strong>о</strong>тря н<strong>а</strong> т<strong>о</strong>, чт<strong>о</strong> т<strong>о</strong>п<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong>и у н<strong>а</strong>иб<strong>о</strong>лее <strong>с</strong>ильн<strong>о</strong> <strong>с</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>л<strong>с</strong>я <strong>в</strong> <strong>с</strong><strong>о</strong><strong>к</strong>р<strong>а</strong>щении брю­<br />

<strong>в</strong>зр<strong>о</strong><strong>с</strong>лых жи<strong>в</strong><strong>о</strong>тных <strong>о</strong>н <strong>о</strong>бр<strong>а</strong>зует передний <strong>к</strong><strong>о</strong>ш<strong>к</strong><strong>а</strong> (мет<strong>а</strong><strong>с</strong><strong>о</strong>мы) и <strong>с</strong>лиянии пер<strong>в</strong>ых четырех<br />

нец тел<strong>а</strong>. Ф<strong>о</strong>рм<strong>а</strong> и р<strong>а</strong>змеры х<strong>о</strong>б<strong>о</strong>т<strong>а</strong> <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> р<strong>а</strong>з­ <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>с</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й л<strong>о</strong>п<strong>а</strong><strong>с</strong>тью. Х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>й<br />

личны. Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из <strong>с</strong>еми — де<strong>в</strong>яти <strong>о</strong><strong>с</strong><strong>о</strong>бенн<strong>о</strong><strong>с</strong>тью <strong>с</strong>тр<strong>о</strong>ения пище<strong>в</strong><strong>а</strong>рительн<strong>о</strong>й <strong>с</strong>и­<br />

<strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Н<strong>а</strong> <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не переднег<strong>о</strong> <strong>с</strong>ег<strong>с</strong>темы <strong>с</strong>лужит з<strong>а</strong>х<strong>о</strong>ждение уз<strong>к</strong>их <strong>с</strong>лепых <strong>в</strong>ыр<strong>о</strong>мент<strong>а</strong><br />

<strong>о</strong>бычн<strong>о</strong> р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>о</strong>жен н<strong>а</strong>упли<strong>а</strong>льный гл<strong>а</strong>з. <strong>с</strong>т<strong>о</strong><strong>в</strong> <strong>с</strong>редней <strong>к</strong>иш<strong>к</strong>и <strong>в</strong> н<strong>о</strong>ги. Сюд<strong>а</strong> пр<strong>о</strong>ни<strong>к</strong><strong>а</strong>ют<br />

Сегменты г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди <strong>с</strong> рез<strong>к</strong><strong>о</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющими <strong>о</strong>тдельные <strong>в</strong>ыр<strong>о</strong><strong>с</strong>ты <strong>с</strong>еменни<strong>к</strong><strong>о</strong><strong>в</strong> у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong>, <strong>а</strong> у<br />

пень<strong>к</strong><strong>о</strong><strong>в</strong>идными б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ыми <strong>в</strong>ыр<strong>о</strong><strong>с</strong>т<strong>а</strong>ми, <strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>­ <strong>с</strong><strong>а</strong>м<strong>о</strong><strong>к</strong> <strong>в</strong> п<strong>о</strong>л<strong>о</strong><strong>с</strong>тях бедер н<strong>о</strong>г р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я<br />

рым причленяют<strong>с</strong>я <strong>с</strong>емь п<strong>а</strong>р <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Пе­ яични<strong>к</strong>и. Р<strong>а</strong>з<strong>в</strong>итие <strong>с</strong> мет<strong>а</strong>м<strong>о</strong>рф<strong>о</strong>з<strong>о</strong>м; из яйц<strong>а</strong><br />

редние четыре <strong>с</strong>егмент<strong>а</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди, <strong>с</strong>ли<strong>в</strong><strong>а</strong>я<strong>с</strong>ь, <strong>в</strong>ых<strong>о</strong>дит трехн<strong>о</strong>г<strong>а</strong>я личин<strong>к</strong><strong>а</strong>. П<strong>о</strong><strong>с</strong>тэмбри<strong>о</strong>н<strong>а</strong>ль­<br />

ф<strong>о</strong>рмируют г<strong>о</strong>л<strong>о</strong><strong>в</strong>у <strong>с</strong> <strong>в</strong>ы<strong>с</strong>туп<strong>а</strong>ющим <strong>в</strong>перед х<strong>о</strong>н<strong>о</strong>е р<strong>а</strong>з<strong>в</strong>итие у<strong>с</strong>л<strong>о</strong>жнен<strong>о</strong> я<strong>в</strong>лениями <strong>а</strong>н<strong>а</strong>м<strong>о</strong>рб<strong>о</strong>т<strong>к</strong><strong>о</strong>м<br />

и четырьмя п<strong>а</strong>р<strong>а</strong>ми <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей. Перф<strong>о</strong>з<strong>а</strong>.<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> (хелицеры) трех-четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые, Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гия и ге<strong>о</strong>гр<strong>а</strong>фиче<strong>с</strong><strong>к</strong><strong>о</strong>е р<strong>а</strong><strong>с</strong>­<br />

<strong>к</strong>лешне<strong>в</strong>идные; ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Следуюпр<strong>о</strong><strong>с</strong>тр<strong>а</strong>нение. М<strong>о</strong>р<strong>с</strong><strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и <strong>о</strong>бит<strong>а</strong>ют <strong>в</strong><strong>о</strong><br />

щие д<strong>в</strong>е п<strong>а</strong>ры <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong> <strong>к</strong><strong>о</strong>­ <strong>в</strong><strong>с</strong>ех <strong>о</strong><strong>к</strong>е<strong>а</strong>н<strong>а</strong>х и м<strong>о</strong>рях Земн<strong>о</strong>г<strong>о</strong> ш<strong>а</strong>р<strong>а</strong>. Хищнир<strong>о</strong>т<strong>к</strong>ие,<br />

н<strong>о</strong>г<strong>о</strong><strong>в</strong>идные, н<strong>о</strong><strong>с</strong>ят н<strong>а</strong>з<strong>в</strong><strong>а</strong>ние педип<strong>а</strong>льп <strong>к</strong>и; <strong>с</strong><strong>о</strong><strong>в</strong>ременные Pantopoda п<strong>о</strong>жир<strong>а</strong>ют преиму­<br />

и яйцен<strong>о</strong><strong>с</strong>це<strong>в</strong>; п<strong>о</strong><strong>с</strong>ледние ин<strong>о</strong>гд<strong>а</strong> имеют<strong>с</strong>я т<strong>о</strong>льще<strong>с</strong>т<strong>в</strong>енн<strong>о</strong> п<strong>о</strong>лип<strong>о</strong><strong>в</strong> <strong>к</strong>ишечн<strong>о</strong>п<strong>о</strong>л<strong>о</strong><strong>с</strong>тных. И<strong>с</strong><strong>к</strong><strong>о</strong><strong>к</strong><strong>о</strong><br />

у <strong>с</strong><strong>а</strong>мц<strong>о</strong><strong>в</strong> и <strong>с</strong>луж<strong>а</strong>т для з<strong>а</strong><strong>к</strong>репления яйцеп<strong>а</strong>емые <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong>и и<strong>с</strong><strong>к</strong>лючительн<strong>о</strong> ред<strong>к</strong>и.<br />

<strong>в</strong>ых <strong>к</strong><strong>о</strong>м<strong>к</strong><strong>о</strong><strong>в</strong>; ин<strong>о</strong>гд<strong>а</strong> и эти <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти м<strong>о</strong>гут П<strong>о</strong>л<strong>о</strong>жение <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме, фил<strong>о</strong>ге­<br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong><strong>о</strong><strong>в</strong><strong>а</strong>ть. К з<strong>а</strong>дней ч<strong>а</strong><strong>с</strong>ти б<strong>о</strong><strong>к</strong><strong>о</strong><strong>в</strong>ых <strong>в</strong>ы<strong>с</strong>туния и <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>ий <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>.<br />

п<strong>о</strong><strong>в</strong> <strong>к</strong>р<strong>а</strong>е<strong>в</strong> г<strong>о</strong>л<strong>о</strong><strong>в</strong>ы причленяет<strong>с</strong>я пер<strong>в</strong><strong>а</strong>я п<strong>а</strong>р<strong>а</strong> В<strong>к</strong>лючение Pycnogonides <strong>в</strong> н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> Chelicera­<br />

длинных х<strong>о</strong>дильных н<strong>о</strong>г <strong>с</strong> р<strong>а</strong><strong>с</strong>члененным н<strong>а</strong> ta, <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е н<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ии <strong>в</strong> <strong>о</strong>беих групп<strong>а</strong>х<br />

525<br />

http://jurassic.ru/

<strong>а</strong>нтенн I и н<strong>а</strong>личии <strong>к</strong>лешне<strong>в</strong>идных пер<strong>в</strong>ых п<strong>о</strong><strong>с</strong>т<strong>а</strong>нтеннулярных<br />

<strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, <strong>о</strong>шиб<strong>о</strong>чн<strong>о</strong>. Пр<strong>о</strong>ти<strong>в</strong><br />

эт<strong>о</strong>г<strong>о</strong> г<strong>о</strong><strong>в</strong><strong>о</strong>рит ин<strong>о</strong>й тип т<strong>а</strong>гм<strong>о</strong>зи<strong>с</strong><strong>а</strong> тел<strong>а</strong>;<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> у Pycnogonides <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, <strong>а</strong> п<strong>о</strong><strong>с</strong>ледние<br />

п<strong>а</strong>ры х<strong>о</strong>дильных н<strong>о</strong>г (VII—IX) причленяют<strong>с</strong>я<br />

н<strong>а</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>ме, и <strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тельн<strong>о</strong>, пр<strong>о</strong>ти<strong>в</strong><strong>о</strong>п<strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ление<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>ме, <strong>с</strong>т<strong>о</strong>ль х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong>е<br />

для Chelicerata, зде<strong>с</strong>ь ее <strong>в</strong>ыр<strong>а</strong>жен<strong>о</strong>.<br />

Слияние у Pycnogonides пер<strong>в</strong>ых четырех <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

<strong>в</strong> единый г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й <strong>о</strong>тдел, <strong>с</strong><strong>о</strong><strong>о</strong>т<strong>в</strong>ет<strong>с</strong>т<strong>в</strong>ующих<br />

пр<strong>о</strong>тер<strong>о</strong><strong>с</strong><strong>о</strong>ме Solifugae, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, я<strong>в</strong>ляет<strong>с</strong>я<br />

<strong>с</strong><strong>о</strong>хр<strong>а</strong>нением тип<strong>а</strong> <strong>с</strong>егмент<strong>а</strong>ции дре<strong>в</strong>них<br />

трил<strong>о</strong>бит<strong>о</strong><strong>в</strong>. О п<strong>о</strong>д<strong>о</strong>бн<strong>о</strong>м <strong>с</strong><strong>о</strong>хр<strong>а</strong>нении черт примити<strong>в</strong>н<strong>о</strong>й<br />

<strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гих <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует<br />

т<strong>а</strong><strong>к</strong>же н<strong>а</strong>личие де<strong>в</strong>яти п<strong>а</strong>р <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей.<br />

От трил<strong>о</strong>бит<strong>о</strong><strong>в</strong> Pycnogonides <strong>о</strong>тлич<strong>а</strong>ют<strong>с</strong>я<br />

<strong>о</strong>лиг<strong>о</strong>мериз<strong>а</strong>цией <strong>с</strong>егмент<strong>о</strong><strong>в</strong> тел<strong>а</strong>, п<strong>о</strong>терей<br />

пер<strong>в</strong>ых <strong>а</strong>нтенн, зн<strong>а</strong>чительным упр<strong>о</strong>щением <strong>в</strong><br />

<strong>с</strong>тр<strong>о</strong>ении пер<strong>в</strong>ичн<strong>о</strong> д<strong>в</strong>у<strong>в</strong>ет<strong>в</strong>и<strong>с</strong>тых н<strong>о</strong>г и <strong>с</strong>пеци<strong>а</strong>лиз<strong>а</strong>цией<br />

трех пер<strong>в</strong>ых п<strong>а</strong>р <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, <strong>в</strong>ып<strong>о</strong>лняющих<br />

фун<strong>к</strong>ции хелиф<strong>о</strong>р, педип<strong>а</strong>льп и<br />

яйцен<strong>о</strong><strong>с</strong>це<strong>в</strong>. Б<strong>о</strong>льш<strong>о</strong>г<strong>о</strong> <strong>в</strong>ним<strong>а</strong>ния з<strong>а</strong><strong>с</strong>лужи<strong>в</strong><strong>а</strong>ет<br />

э<strong>в</strong><strong>о</strong>люция <strong>а</strong>рхицеф<strong>а</strong>л<strong>о</strong>н<strong>а</strong> у Pycnogonides, <strong>с</strong> <strong>о</strong>дн<strong>о</strong>й<br />

<strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, и у Chelicerata — <strong>с</strong> друг<strong>о</strong>й. Для<br />

<strong>о</strong>беих этих групп х<strong>а</strong>р<strong>а</strong><strong>к</strong>терн<strong>о</strong> изгиб<strong>а</strong>ние передней<br />

ч<strong>а</strong><strong>с</strong>ти <strong>о</strong><strong>с</strong>и тел<strong>а</strong> <strong>в</strong><strong>в</strong>ерх и н<strong>а</strong>з<strong>а</strong>д, чт<strong>о</strong> при<strong>в</strong>ел<strong>о</strong><br />

<strong>к</strong> реду<strong>к</strong>ции <strong>а</strong>нтеннул и <strong>с</strong>мещению г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>й<br />

л<strong>о</strong>п<strong>а</strong><strong>с</strong>ти <strong>с</strong> гл<strong>а</strong>з<strong>а</strong>ми н<strong>а</strong> <strong>с</strong>пинную <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ну<br />

тел<strong>а</strong>. В<strong>с</strong>е эт<strong>о</strong> <strong>с</strong><strong>в</strong>идетель<strong>с</strong>т<strong>в</strong>ует <strong>о</strong> прин<strong>а</strong>длежн<strong>о</strong><strong>с</strong>ти<br />

<strong>о</strong>беих групп <strong>к</strong> <strong>о</strong><strong>с</strong><strong>о</strong>б<strong>о</strong>му п<strong>о</strong>дтипу Chelicerophora.<br />

http://jurassic.ru/<br />

Следы трил<strong>о</strong>бит<strong>о</strong>идн<strong>о</strong>й <strong>о</strong>рг<strong>а</strong>низ<strong>а</strong>ции не <strong>о</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>ляют<br />

<strong>с</strong><strong>о</strong>мнений <strong>в</strong> пр<strong>о</strong>и<strong>с</strong>х<strong>о</strong>ждении Pycnogonides<br />

<strong>о</strong>т примити<strong>в</strong>ных д<strong>о</strong>п<strong>а</strong>ле<strong>о</strong>з<strong>о</strong>й<strong>с</strong><strong>к</strong>их Trilobitomorpha.<br />

Вер<strong>о</strong>ятн<strong>о</strong>, пер<strong>в</strong>ые эт<strong>а</strong>пы р<strong>а</strong>з<strong>в</strong>ития;<br />

Pycnogonides и Chelicerata пр<strong>о</strong>х<strong>о</strong>дили п<strong>а</strong>р<strong>а</strong>ллельн<strong>о</strong>,<br />

чем и <strong>о</strong>бъя<strong>с</strong>няют<strong>с</strong>я <strong>о</strong>тмеченные <strong>в</strong>ыше<br />

черты <strong>с</strong>х<strong>о</strong>д<strong>с</strong>т<strong>в</strong><strong>а</strong> у <strong>с</strong><strong>о</strong><strong>в</strong>ременных п<strong>а</strong>нт<strong>о</strong>и<strong>о</strong>д и хелицер<strong>о</strong><strong>в</strong>ых.<br />

Одн<strong>а</strong><strong>к</strong><strong>о</strong> д<strong>а</strong>льнейшее р<strong>а</strong>з<strong>в</strong>итие <strong>в</strong><br />

<strong>о</strong>беих этих групп<strong>а</strong>х пр<strong>о</strong>те<strong>к</strong><strong>а</strong>л<strong>о</strong> <strong>с</strong><strong>в</strong><strong>о</strong>е<strong>о</strong>бр<strong>а</strong>зн<strong>о</strong>, чт<strong>о</strong>»<br />

и при<strong>в</strong>ел<strong>о</strong> <strong>к</strong> ф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>нию д<strong>в</strong>ух близ<strong>к</strong>их н<strong>а</strong>д<strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>о</strong><strong>в</strong>.<br />

Очень з<strong>а</strong>держ<strong>а</strong>л<strong>а</strong><strong>с</strong>ь пр<strong>а</strong><strong>в</strong>ильн<strong>а</strong>я <strong>о</strong>цен<strong>к</strong><strong>а</strong> фил<strong>о</strong>генетиче<strong>с</strong><strong>к</strong>их<br />

<strong>о</strong>тн<strong>о</strong>шений дре<strong>в</strong>них пи<strong>к</strong>н<strong>о</strong>г<strong>о</strong>н<strong>о</strong><strong>в</strong>;<br />

<strong>в</strong><strong>с</strong>лед<strong>с</strong>т<strong>в</strong>ие <strong>о</strong>шиб<strong>к</strong>и, д<strong>о</strong>пущенн<strong>о</strong>й <strong>в</strong> тр<strong>а</strong><strong>к</strong>т<strong>о</strong><strong>в</strong><strong>к</strong>е<br />

переднег<strong>о</strong> и з<strong>а</strong>днег<strong>о</strong> <strong>к</strong><strong>о</strong>нц<strong>о</strong><strong>в</strong> тел<strong>а</strong> де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>-<br />

Palaeoisopus probletnaticus Broili. Уз<strong>к</strong>ий и т<strong>о</strong>н<strong>к</strong>ий<br />

<strong>в</strong>ыр<strong>о</strong><strong>с</strong>т, <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ящий из ше<strong>с</strong>ти члени<strong>к</strong><strong>о</strong><strong>в</strong>, п<strong>о</strong><strong>к</strong>рытых<br />

<strong>в</strong><strong>о</strong>л<strong>о</strong><strong>с</strong><strong>к</strong><strong>а</strong>ми, н<strong>а</strong>пр<strong>а</strong><strong>в</strong>ленными ди<strong>с</strong>т<strong>а</strong>льн<strong>о</strong>,<br />

<strong>к</strong><strong>о</strong>т<strong>о</strong>рый Бр<strong>о</strong>йли (iBroili, 1928—1933) и другие<br />

<strong>а</strong><strong>в</strong>т<strong>о</strong>ры (Stormer, 1944; Hedgpeth, 1955) приним<strong>а</strong>ли<br />

з<strong>а</strong> х<strong>о</strong>б<strong>о</strong>т, <strong>в</strong> дей<strong>с</strong>т<strong>в</strong>ительн<strong>о</strong><strong>с</strong>ти я<strong>в</strong>ляет<strong>с</strong>я<br />

брюш<strong>к</strong><strong>о</strong>м. Эт<strong>о</strong> у<strong>с</strong>т<strong>а</strong>н<strong>о</strong><strong>в</strong>лен<strong>о</strong> н<strong>а</strong>ми при изучении<br />

из<strong>о</strong>бр<strong>а</strong>жений д<strong>а</strong>нн<strong>о</strong>й ф<strong>о</strong>рмы (Дубинин, 1957)<br />

и <strong>в</strong> д<strong>а</strong>льнейшем был<strong>о</strong> п<strong>о</strong>дт<strong>в</strong>ержден<strong>о</strong> <strong>с</strong>пеци<strong>а</strong>льными<br />

и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>ниями <strong>о</strong><strong>с</strong>т<strong>а</strong>т<strong>к</strong><strong>о</strong><strong>в</strong> н<strong>а</strong> рентген<strong>о</strong>гр<strong>а</strong>фиях<br />

(Lehmann, 1959). Изучение п<strong>о</strong><strong>к</strong><strong>а</strong>з<strong>а</strong>л<strong>о</strong>,<br />

чт<strong>о</strong> передний р<strong>а</strong>здутый <strong>к</strong><strong>о</strong>нец тел<strong>а</strong> — <strong>к</strong>рупные<strong>с</strong><strong>о</strong>м<strong>к</strong>нутые<br />

хелицеры, <strong>а</strong> <strong>в</strong><strong>о</strong><strong>в</strong><strong>с</strong>е не т<strong>о</strong>л<strong>с</strong>тый х<strong>о</strong>б<strong>о</strong>т<strong>о</strong><strong>к</strong>:<br />

п<strong>о</strong><strong>с</strong>ледний имеет <strong>в</strong>е<strong>с</strong>ьм<strong>а</strong> неб<strong>о</strong>льшие р<strong>а</strong>змеры<br />

и <strong>с</strong><strong>в</strong>ерху нер<strong>а</strong>зличим (ри<strong>с</strong>. 1531). Де<strong>в</strong><strong>о</strong>н—<br />

ныне. Д<strong>в</strong><strong>а</strong> <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong><strong>а</strong>: Protopantopoda »<br />

Pantopoda.

КЛАСС PROTOPANTOPODA.<br />

ПРОТОПАНТОПОДЫ, ИЛИ ЧЛЕНИСТОБРЮХИЕ<br />

МОРСКИЕ ПАУКИ<br />

Хелицеры <strong>к</strong>рупные, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые; х<strong>о</strong>б<strong>о</strong>т<br />

<strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ий и т<strong>о</strong>н<strong>к</strong>ий. Брюш<strong>к</strong><strong>о</strong> длинн<strong>о</strong>е,<br />

б<strong>о</strong>льше г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди; <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из пяти <strong>с</strong>егмент<strong>о</strong><strong>в</strong><br />

и з<strong>а</strong><strong>о</strong><strong>с</strong>тренн<strong>о</strong>г<strong>о</strong> тель<strong>с</strong><strong>о</strong>н<strong>а</strong>. Де<strong>в</strong><strong>о</strong>н. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong><br />

Palaeoisopodidae.<br />

СЕМЕЙСТВО PALAEOISOPODIDAE W. DUBININ,<br />

1957<br />

Имеет<strong>с</strong>я гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong> <strong>с</strong> гл<strong>а</strong>з<strong>а</strong>ми. Хелицеры<br />

<strong>в</strong> <strong>в</strong>иде м<strong>о</strong>щных, з<strong>а</strong>гнутых друг <strong>к</strong> другу<br />

<strong>к</strong>лешней <strong>с</strong> неб<strong>о</strong>льшим п<strong>о</strong>д<strong>в</strong>ижным чет<strong>в</strong>ертым<br />

члени<strong>к</strong><strong>о</strong>м. Педип<strong>а</strong>льпы т<strong>о</strong>н<strong>к</strong>ие и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, леж<strong>а</strong>т<br />

ряд<strong>о</strong>м <strong>с</strong> т<strong>о</strong>н<strong>к</strong>им х<strong>о</strong>б<strong>о</strong>т<strong>к</strong><strong>о</strong>м. Яйцен<strong>о</strong><strong>с</strong>цы<br />

длиннее хелицер, н<strong>а</strong> <strong>к</strong><strong>о</strong>нце <strong>к</strong>рюч<strong>к</strong><strong>о</strong><strong>в</strong>идные.<br />

Чет<strong>в</strong>ертый <strong>с</strong>егмент тел<strong>а</strong> д<strong>о</strong><strong>с</strong>тиг<strong>а</strong>л зн<strong>а</strong>чительных<br />

р<strong>а</strong>змер<strong>о</strong><strong>в</strong>: <strong>к</strong> нему при<strong>к</strong>реплял<strong>а</strong><strong>с</strong>ь передняя<br />

п<strong>а</strong>р<strong>а</strong> х<strong>о</strong>дильных н<strong>о</strong>г, р<strong>а</strong>з<strong>в</strong>итых <strong>с</strong>ильнее<br />

(\В. Б. Дубинин\)<br />

572<br />

http://jurassic.ru/<br />

других и, не<strong>с</strong><strong>о</strong>мненн<strong>о</strong>, <strong>в</strong>ып<strong>о</strong>лня<strong>в</strong>ших х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельную<br />

или фи<strong>к</strong><strong>с</strong><strong>а</strong>т<strong>о</strong>рную фун<strong>к</strong>цию (ри<strong>с</strong>. 1532).<br />

Де<strong>в</strong><strong>о</strong>н. Один р<strong>о</strong>д — Palaeoisopus Broilli 1928.<br />

Palaeoisopus Broil i, 1928. Тип р<strong>о</strong>д<strong>а</strong> —<br />

P. problematicus Broili, 1928; н. де<strong>в</strong><strong>о</strong>н, Герм<strong>а</strong>ния.<br />

Тел<strong>о</strong> <strong>в</strong>ытянут<strong>о</strong>е; <strong>в</strong><strong>с</strong>е <strong>с</strong>егменты г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди<br />

<strong>о</strong>б<strong>о</strong><strong>с</strong><strong>о</strong>бленные; три передние <strong>с</strong>ближены <strong>в</strong><br />

<strong>в</strong>иде <strong>с</strong><strong>к</strong>л<strong>а</strong>дч<strong>а</strong>т<strong>о</strong>й шее<strong>в</strong>идн<strong>о</strong>й перетяж<strong>к</strong>и между<br />

хелицер<strong>а</strong>ми и чет<strong>в</strong>ертым <strong>с</strong>егмент<strong>о</strong>м, не<strong>с</strong>ущим<br />

гр<strong>о</strong>м<strong>а</strong>дные х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельные н<strong>о</strong>ги. Брюш<strong>к</strong><strong>о</strong> длинн<strong>о</strong>е,<br />

ше<strong>с</strong>тичлени<strong>к</strong><strong>о</strong><strong>в</strong><strong>о</strong>е (<strong>в</strong><strong>к</strong>люч<strong>а</strong>я тель<strong>с</strong><strong>о</strong>н); длин<strong>а</strong><br />

ег<strong>о</strong> б<strong>о</strong>льше г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди. Н<strong>о</strong>ги чет<strong>в</strong>ерт<strong>о</strong>г<strong>о</strong><br />

<strong>с</strong>егмент<strong>а</strong> <strong>о</strong>чень <strong>к</strong>рупные, т<strong>о</strong>л<strong>с</strong>тые, <strong>а</strong> <strong>о</strong><strong>с</strong>т<strong>а</strong>льные<br />

три п<strong>а</strong>ры н<strong>о</strong>г — х<strong>о</strong>дильн<strong>о</strong>г<strong>о</strong> тип<strong>а</strong>. Длин<strong>а</strong> тел<strong>а</strong><br />

14,2 <strong>с</strong>м. Э<strong>к</strong><strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>и <strong>с</strong><strong>в</strong>яз<strong>а</strong>н <strong>с</strong> м<strong>о</strong>р<strong>с</strong><strong>к</strong>ими лилиями,<br />

<strong>к</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>рым жи<strong>в</strong><strong>о</strong>тн<strong>о</strong>е при<strong>к</strong>реплял<strong>о</strong><strong>с</strong>ь при<br />

п<strong>о</strong>м<strong>о</strong>щи х<strong>в</strong><strong>а</strong>т<strong>а</strong>тельных н<strong>о</strong>г (ри<strong>с</strong>. 1531). Один<br />

<strong>в</strong>ид. Де<strong>в</strong><strong>о</strong>н Герм<strong>а</strong>нии.

1632<br />

1631<br />

http://jurassic.ru/<br />

1633<br />

Ри<strong>с</strong>.41531—1533. HaAMacc^Pycnogonides<br />

1531. Palaeoisopus problematicus Broili (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция); (тел<strong>о</strong><strong>с</strong>низу,<br />

XI,0; де<strong>в</strong><strong>о</strong>н, Герм<strong>а</strong>ния (Lehmann, 1959). 1532. При<strong>к</strong>репленные<br />

Palaeoisopus problematicus Broili н<strong>а</strong> м<strong>о</strong>р<strong>с</strong><strong>к</strong><strong>о</strong>й лилии р<strong>о</strong>д<strong>а</strong><br />

Cyathocrtnus (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция) (Дубинин, 1957). 1533. Palaeopantopus<br />

mauchert Broili; <strong>о</strong>бщий <strong>в</strong>ид <strong>с</strong><strong>в</strong>ерху, X 0,7, и. де<strong>в</strong><strong>о</strong>н<br />

З.Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (Hedgpeth, 1955)

КЛАСС PANTOPODA<br />

В. Б. Дубинин |<br />

Хелицеры неб<strong>о</strong>льшие, трех- или д<strong>в</strong>учлени- ин<strong>о</strong>гд<strong>а</strong> члени<strong>с</strong>т<strong>о</strong>г<strong>о</strong> н<strong>а</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии (Palaeopanto<strong>к</strong><strong>о</strong><strong>в</strong>ые,<br />

ин<strong>о</strong>гд<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Х<strong>о</strong>б<strong>о</strong>т х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з- poda). Де<strong>в</strong><strong>о</strong>н — ныне. Д<strong>в</strong><strong>а</strong> <strong>о</strong>тряд<strong>а</strong>: Palaeopan<strong>в</strong>ит,<br />

ин<strong>о</strong>гд<strong>а</strong> длиннее г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>груди. Брюш<strong>к</strong><strong>о</strong> ре- topoda и Eupantopoda; J3 и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

дуцир<strong>о</strong><strong>в</strong><strong>а</strong>нн<strong>о</strong>е, <strong>в</strong> <strong>в</strong>иде <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong><strong>о</strong>г<strong>о</strong> <strong>о</strong>тр<strong>о</strong><strong>с</strong>т<strong>к</strong><strong>а</strong>, из<strong>в</strong>е<strong>с</strong>тен лишь пер<strong>в</strong>ый.<br />

ОТРЯД PA LA ЕОРА NTOPO DА<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния передней п<strong>а</strong>ры х<strong>о</strong>дильных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

рез<strong>к</strong><strong>о</strong> <strong>с</strong>ближены <strong>с</strong>переди. Гл<strong>а</strong>зн<strong>о</strong>й<br />

буг<strong>о</strong>р<strong>о</strong><strong>к</strong>, х<strong>о</strong>б<strong>о</strong>т, хелицеры и педип<strong>а</strong>льпы не<br />

<strong>в</strong>идны,— <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, <strong>о</strong>ни <strong>о</strong>чень м<strong>а</strong>лы или ч<strong>а</strong><strong>с</strong>тичн<strong>о</strong><br />

<strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>уют. Брюш<strong>к</strong><strong>о</strong> <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>ит из тель<strong>с</strong><strong>о</strong>н<strong>а</strong>,<br />

<strong>о</strong>дн<strong>о</strong>г<strong>о</strong> длинн<strong>о</strong>г<strong>о</strong> члени<strong>к</strong><strong>а</strong> и редуцир<strong>о</strong><strong>в</strong><strong>а</strong>нных<br />

<strong>в</strong> <strong>в</strong>иде <strong>с</strong><strong>к</strong>л<strong>а</strong>д<strong>о</strong><strong>к</strong> б<strong>а</strong>з<strong>а</strong>льных члени<strong>к</strong><strong>о</strong><strong>в</strong>. Де<strong>в</strong><strong>о</strong>н.<br />

Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Palaeopantopodidae.<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния передней п<strong>а</strong>ры х<strong>о</strong>дильных <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей<br />

шир<strong>о</strong><strong>к</strong><strong>о</strong> р<strong>а</strong><strong>с</strong><strong>с</strong>т<strong>а</strong><strong>в</strong>лены, и между ними<br />

р<strong>а</strong><strong>с</strong>п<strong>о</strong>л<strong>а</strong>г<strong>а</strong>ют<strong>с</strong>я х<strong>о</strong>б<strong>о</strong>т и <strong>о</strong>бычн<strong>о</strong> хелицеры, пе­<br />

ОТРЯД EUPANTOPODA<br />

34 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие 529<br />

http://jurassic.ru/<br />

СЕМЕЙСТВО PALAEOPANTOPODIDAE HEDGPETH,<br />

1965<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ния <strong>в</strong><strong>с</strong>ех <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей <strong>с</strong><strong>к</strong>л<strong>а</strong>дч<strong>а</strong>тые<br />

(рудимент<strong>а</strong>рн<strong>а</strong>я члеии<strong>с</strong>т<strong>о</strong><strong>с</strong>ть?). Г<strong>о</strong>л<strong>о</strong><strong>в</strong><strong>о</strong>грудь<br />

из трех чет<strong>к</strong>их <strong>с</strong>егмент<strong>о</strong><strong>в</strong>. Яйцен<strong>о</strong><strong>с</strong>цы т<strong>о</strong>н<strong>к</strong>ие<br />

и <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие, <strong>в</strong>ер<strong>о</strong>ятн<strong>о</strong>, четырехчлени<strong>к</strong><strong>о</strong><strong>в</strong>ые. Н.<br />

де<strong>в</strong><strong>о</strong>н 3. Е<strong>в</strong>р<strong>о</strong>пы. Один р<strong>о</strong>д Palaeopantopus<br />

Broili, 1929 <strong>с</strong> <strong>о</strong>дним <strong>в</strong>ид<strong>о</strong>м (ри<strong>с</strong>. 1533).<br />

дип<strong>а</strong>льпы и <strong>с</strong><strong>в</strong>ерху — гл<strong>а</strong>зн<strong>о</strong>й буг<strong>о</strong>р<strong>о</strong><strong>к</strong>. С<strong>о</strong><strong>в</strong>ременные.<br />

В<strong>о</strong><strong>с</strong>емь <strong>с</strong>емей<strong>с</strong>т<strong>в</strong>, <strong>в</strong> и<strong>с</strong><strong>к</strong><strong>о</strong>п<strong>а</strong>ем<strong>о</strong>м <strong>с</strong><strong>о</strong><strong>с</strong>т<strong>о</strong>янии<br />

неиз<strong>в</strong>е<strong>с</strong>тных.

Бе<strong>к</strong>лемише<strong>в</strong> В. Н. 1952. О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы <strong>с</strong>р<strong>а</strong><strong>в</strong>нительн<strong>о</strong>й<br />

<strong>а</strong>н<strong>а</strong>т<strong>о</strong>мии бе<strong>с</strong>п<strong>о</strong>з<strong>в</strong><strong>о</strong>н<strong>о</strong>чных. Изд. «С<strong>о</strong><strong>в</strong>ет<strong>с</strong><strong>к</strong><strong>а</strong>я н<strong>а</strong>у<strong>к</strong><strong>а</strong>», М.,<br />

<strong>с</strong>тр. 1—698.<br />

Д<strong>о</strong>гель В. А. 1913. Emibryologische Studien й. Р<strong>а</strong>пtopoden.<br />

Ztschr. f. wiss. Zoologie, Bd. 107; S.'4—210.<br />

Дубинин В. Б. 1957. Об <strong>о</strong>риент<strong>а</strong>ции г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong><br />

<strong>к</strong><strong>о</strong>нц<strong>а</strong> у де<strong>в</strong><strong>о</strong>н<strong>с</strong><strong>к</strong>их пи<strong>к</strong>н<strong>о</strong>г<strong>о</strong>н<strong>о</strong><strong>в</strong> р<strong>о</strong>д<strong>а</strong> Palaeoisopus и п<strong>о</strong>л<strong>о</strong>жении<br />

их <strong>в</strong> <strong>с</strong>и<strong>с</strong>теме Arthropoda. Д<strong>о</strong><strong>к</strong>л. АН СССР,<br />

т. 117, № 5, <strong>с</strong>тр. 881—«84.<br />

. Л<strong>о</strong>зин<strong>а</strong>-Л<strong>о</strong>зин<strong>с</strong><strong>к</strong>ий Л. 1923. Pantopoda, <strong>с</strong><strong>о</strong>бр<strong>а</strong>нные<br />

э<strong>к</strong><strong>с</strong>педицией' н<strong>а</strong>учн<strong>о</strong>-и<strong>с</strong><strong>с</strong>лед<strong>о</strong><strong>в</strong><strong>а</strong>тель<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> О<strong>к</strong>е<strong>а</strong>н<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

ин<strong>с</strong>титут<strong>а</strong> <strong>в</strong> 1921 г. Тр. Ин-т<strong>а</strong> <strong>о</strong><strong>к</strong>е<strong>а</strong>н<strong>о</strong>л<strong>о</strong>гии,<br />

т. IV (3), <strong>с</strong>тр. 63—87.<br />

Шим<strong>к</strong>е<strong>в</strong>ич В. 1930. М<strong>о</strong>р<strong>с</strong><strong>к</strong>ие п<strong>а</strong>у<strong>к</strong>и (Pantopoda).<br />

«Ф<strong>а</strong>ун<strong>а</strong> СССР и <strong>с</strong><strong>о</strong>предельных <strong>с</strong>тр<strong>а</strong>н», т. 5, <strong>в</strong>ып. 1,<br />

<strong>с</strong>тр. 1—532.<br />

Вбгпег С. 1903. Die Beingliederung der Arthropoden.<br />

Situngsber. Ges. nat. Frd. Berlin, S. 292—335.,В г <strong>о</strong> ili<br />

F. 1828. Crustacenfunde aus dem rhenischeft' Unterdevon.<br />

Situngsber. bayr. Akad. Wissensch. (Math.-Nat.),<br />

S. 197—201.— 1929. Beobachtugnen an neuen Arthropodenfunden<br />

aus den Hunsruckschiefern. Ein Pantopoda<br />

aus dem rheinischen Underdevon. Ibid., S. 272—280.—<br />

1930. Ueber ein neues Exemplar von Palaeopantopus.<br />

Iibid., 209—214.— 1932. Palaeoisopus ist ein Pantopode:<br />

ЛИТЕРАТУРА<br />

http://jurassic.ru/<br />

Ibid., S. 45—60.— 1933. Wietere Beobachtungen an Palaeoisopus.<br />

Ibid., S. 33—47.<br />

Caiman W. 1909. The problem of the Pycnogonus.<br />

Sci. Progress in the Twentieth Century. London, p. 687—<br />

693.— 1929. The- -Pycnogonida. Journ. Quekett. Microscop.<br />

Club., London, vol. 2, p. 16—106. Caiman W. et Gordon<br />

J. 1933. A Dodecapodous Pycnogonid. Proc. Roy.<br />

Soc, London, vol. 113, p. 107—115.<br />

F a g e L. 1949. Classe des Pycnogonides. Traite de<br />

Zoologie, v. VI, p. 906—941.<br />

Gordon J. 1932. Pycnogonida. Discovery reports.<br />

Cambridg. v. 6, p. 1—138.<br />

Hedgpeth I. W. 1955. Pycnogonida, Palaeoisopus.<br />

Treatise on Invertebrate Paleontology, pt. P, Arthropoda<br />

2, p. 163—173.<br />

Heifer H. 1932. Pantopoda. In: Kukenthel — Krumbach.<br />

Handbuch d. Zoolog., Bd. 3. S. 1—66. Heifer H.<br />

u. Schl.ottke E. 1935. Pantopoda. In: Bronn's Klassen<br />

a. Ordn. Tierreichs, Bd. 5, Teil 4, 1—160.<br />

Lehmann W. M. 1959. Neue Endeckungen an Palaeoisopus.<br />

Palaontol. Ztschr., Bd. 33, H. 1/2, S. 96—103.<br />

S t 0 r m e r L. 1944. On the relationships and phylogeny<br />

of fossil and- recent Arachnomorpha. Oslo, p. 1—158.<br />

Wieren W. 1918. Zur Morphologie und Phylogenie<br />

der Pantopoden. Zool. Bidrag fran Uppsala, Bd. 6, S.<br />

41—181.

ТИП MALACOPODA<br />

МЯГКО НО ГИЕ,<br />

ИЛИ ПЕРВИЧНОТРАХЕЙНЫЕ<br />

http://jurassic.ru/

http://jurassic.ru/

ТИП MALACOPODA.<br />

МЯГКОНОГИЕ, ИЛИ ПЕРВИЧНОТРАХЕЙНЫЕ<br />

Неб<strong>о</strong>льшие чер<strong>в</strong>е<strong>о</strong>бр<strong>а</strong>зные жи<strong>в</strong><strong>о</strong>тные (длин<strong>а</strong><br />

тел<strong>а</strong> 1,5—20 <strong>с</strong>м) <strong>с</strong> мяг<strong>к</strong>им хитин<strong>о</strong><strong>в</strong>ым п<strong>о</strong><strong>к</strong>р<strong>о</strong><strong>в</strong><strong>о</strong>м,<br />

п<strong>а</strong>р<strong>о</strong>й <strong>а</strong>нтенн, п<strong>а</strong>р<strong>о</strong>й челю<strong>с</strong>тей <strong>в</strong> п<strong>о</strong>л<strong>о</strong><strong>с</strong>ти<br />

рт<strong>а</strong>,п<strong>а</strong>р<strong>о</strong>й <strong>с</strong><strong>о</strong><strong>с</strong><strong>о</strong>ч<strong>к</strong><strong>о</strong><strong>в</strong> п<strong>о</strong>з<strong>а</strong>ди рт<strong>а</strong> и <strong>с</strong> р<strong>а</strong>зличным<br />

чи<strong>с</strong>л<strong>о</strong>м (не менее де<strong>с</strong>яти п<strong>а</strong>р) <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>их<br />

<strong>к</strong><strong>о</strong>льч<strong>а</strong>тых <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тей, <strong>в</strong> <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>нии <strong>к</strong><strong>о</strong>т<strong>о</strong>­<br />

(А. Г. Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>)<br />

КЛАСС ONYCHOPHORA<br />

рых <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ют<strong>с</strong>я пр<strong>о</strong>т<strong>о</strong><strong>к</strong>и нефридие<strong>в</strong>. С<strong>о</strong><strong>в</strong>ременные<br />

Malacopoda дыш<strong>а</strong>т тр<strong>а</strong>хеями, <strong>о</strong>т<strong>к</strong>ры<strong>в</strong><strong>а</strong>ющими<strong>с</strong>я<br />

<strong>в</strong> р<strong>а</strong>зличных ме<strong>с</strong>т<strong>а</strong>х н<strong>а</strong> п<strong>о</strong><strong>в</strong>ерхн<strong>о</strong><strong>с</strong>ти<br />

тел<strong>а</strong>. Р<strong>о</strong>д<strong>с</strong>т<strong>в</strong>енны <strong>к</strong><strong>о</strong>льч<strong>а</strong>тым чер<strong>в</strong>ям<br />

и не я<strong>в</strong>ляют<strong>с</strong>я прямыми пред<strong>к</strong><strong>а</strong>ми Arthropoda.<br />

Кембрий — ныне. Один <strong>к</strong>л<strong>а</strong><strong>с</strong><strong>с</strong> — Onychophora.<br />

Н<strong>а</strong>ружн<strong>а</strong>я <strong>с</strong>егмент<strong>а</strong>ция тел<strong>а</strong> <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует, phora, Euonychophora; п<strong>о</strong><strong>с</strong>ледний т<strong>о</strong>ль<strong>к</strong><strong>о</strong> <strong>в</strong> <strong>с</strong><strong>о</strong>-<br />

К<strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong> д<strong>в</strong>умя — ше<strong>с</strong>тью <strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми н<strong>а</strong> <strong>в</strong>ременн<strong>о</strong>й ф<strong>а</strong>уне,<br />

<strong>к</strong><strong>о</strong>нце. Кембрий — ныне. Отряды: Protonycho-<br />

Антенны <strong>к</strong><strong>о</strong>р<strong>о</strong>т<strong>к</strong>ие; п<strong>а</strong>р<strong>а</strong> х<strong>о</strong>р<strong>о</strong>ш<strong>о</strong> р<strong>а</strong>з<strong>в</strong>итых<br />

гл<strong>а</strong>з. Кембрий. Одн<strong>о</strong> <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> — Aysheaiidae.<br />

СЕМЕЙСТВО AYSHEAIIDAE<br />

WALCOTT, 1911<br />

Антенны ните<strong>в</strong>идные; <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong> ше<strong>с</strong>тью<br />

<strong>к</strong><strong>о</strong>г<strong>о</strong>т<strong>к</strong><strong>а</strong>ми. Один р<strong>о</strong>д. Кембрий.<br />

ОТРЯД PROTONYCHOPHORA<br />

533<br />

Aysheaia Walcott, 1911. Тип р<strong>о</strong>д<strong>а</strong> — А.<br />

peduncutata Walcott, 1911; <strong>с</strong>р. <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong>.<br />

Длин<strong>а</strong> тел<strong>а</strong> <strong>о</strong>т г<strong>о</strong>л<strong>о</strong><strong>в</strong>н<strong>о</strong>г<strong>о</strong> щит<strong>а</strong> д<strong>о</strong> де<strong>с</strong>ят<strong>о</strong>г<strong>о</strong>,<br />

не<strong>с</strong>ущег<strong>о</strong> <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти <strong>с</strong>егмент<strong>а</strong>, р<strong>а</strong><strong>в</strong>н<strong>а</strong><br />

33 мм (ри<strong>с</strong>. 1534). Один <strong>в</strong>ид. Ср. <strong>к</strong>ембрий С.<br />

Амери<strong>к</strong>и.<br />

http://jurassic.ru/

MALACOPODA INCERTAE SEDIS<br />

Xenusion Pompecky, 1927. Тип р<strong>о</strong>д<strong>а</strong> — X. <strong>а</strong>иerswaldae<br />

Pompecky, 1927; д<strong>о</strong><strong>к</strong>ембрий, 3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong>.<br />

Тел<strong>о</strong> я<strong>с</strong>н<strong>о</strong> <strong>с</strong>егментир<strong>о</strong><strong>в</strong><strong>а</strong>н<strong>о</strong>. Передние<br />

12 (?) <strong>с</strong>егмент<strong>о</strong><strong>в</strong> не<strong>с</strong>ут <strong>к</strong><strong>о</strong>ну<strong>с</strong><strong>о</strong><strong>в</strong>идные члеяи-<br />

<strong>с</strong>тые <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>ти, при<strong>к</strong>репленные п<strong>о</strong> б<strong>о</strong><strong>к</strong><strong>а</strong>м тел<strong>а</strong>.<br />

Н<strong>а</strong>" <strong>с</strong>пинн<strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>не <strong>к</strong><strong>а</strong>жд<strong>о</strong>г<strong>о</strong> <strong>с</strong>егмент<strong>а</strong> —<br />

п<strong>а</strong>рные <strong>в</strong>ыпу<strong>к</strong>л<strong>о</strong><strong>с</strong>ти. Длин<strong>а</strong> <strong>с</strong><strong>о</strong>хр<strong>а</strong>ни<strong>в</strong>шей<strong>с</strong>я ч<strong>а</strong><strong>с</strong>ти<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong> 8,5 <strong>с</strong>м (ри<strong>с</strong>. 1535). Один <strong>в</strong>ид.<br />

Д<strong>о</strong><strong>к</strong>ембрий 3. Е<strong>в</strong>р<strong>о</strong>пы.<br />

153В<br />

Ри<strong>с</strong>.. 1534—1535. Тип Malacopoda ' '<br />

!_ 34<br />

L-.d^£ m<br />

4. pedunculate:__WaJcott (ре<strong>к</strong><strong>о</strong>н<strong>с</strong>тру<strong>к</strong>ция), x 2,2, auerswaldoe Pompeckj; <strong>о</strong>бщий <strong>в</strong>ид <strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong><strong>а</strong>, x0,6, д<strong>о</strong><strong>к</strong>ембрий,<br />

<strong>с</strong>р. <strong>к</strong>ембрий, С. Амери<strong>к</strong><strong>а</strong> (Hutchinson, 1930). 1535. Xenusion<br />

3. Е<strong>в</strong>р<strong>о</strong>п<strong>а</strong> (<strong>о</strong>риг. ри<strong>с</strong>. п<strong>о</strong> ф<strong>о</strong>т<strong>о</strong>гр<strong>а</strong>фии г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>; Pompeckj, \1927)<br />

534<br />

http://jurassic.ru/

Dechaseaux С. 1953. Onychophora. In: J. Piveteau<br />

«Traite de paleontologies, v. Ill, p. 3—

http://jurassic.ru/

35. О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

ТАБЛИЦЫ I—XXII<br />

И ОБЪЯСНЕНИЯ К НИМ<br />

http://jurassic.ru/

ТАБЛИЦА I<br />

Отряд Ephemeroptera<br />

Фиг. 1. Ephemeropsis trisetalis Eichwald: a — <strong>о</strong>бщий <strong>в</strong>ид, п<strong>о</strong>зити<strong>в</strong>ный<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>, X 1,8; б — переднее <strong>к</strong>рыл<strong>о</strong>, п<strong>о</strong>зити<strong>в</strong>ный<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>; <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (Черн<strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 2. Ephemeropsis trisetalis Eichwald; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, нег<strong>а</strong>ти<strong>в</strong>ный<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>, X 3,0; <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (Черн<strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961)<br />

Фиг. 3. Ephemeropsis trisetalis Eichwald; личин<strong>к</strong>и, п<strong>о</strong>зити<strong>в</strong>ные<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>к</strong>и, X 3,6; <strong>в</strong>. юр<strong>а</strong>, З<strong>а</strong>б<strong>а</strong>й<strong>к</strong><strong>а</strong>лье (<strong>о</strong>риг. <strong>с</strong>р<strong>о</strong>т<strong>о</strong>)<br />

http://jurassic.ru/

http://jurassic.ru/<br />

ТАБЛИЦА <<br />

35

ТАБЛ ИЦА II<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacridae (п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacrinae)<br />

Фиг. 1. Kunguroblattina microdictya Becker-Migdisova: <strong>а</strong> — <strong>о</strong>бщий<br />

<strong>в</strong>ид, п<strong>о</strong>зити<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>, X 3; б — <strong>о</strong>бщий <strong>в</strong>ид,<br />

нег<strong>а</strong>ти<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>, X 1,7; <strong>в</strong> — яйце<strong>к</strong>л<strong>а</strong>д, п<strong>о</strong>зити<strong>в</strong>ный<br />

<strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>, X 4; г — яйце<strong>к</strong>л<strong>а</strong>д, нег<strong>а</strong>ти<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>,<br />

X 4; н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (<strong>о</strong>риг.<br />

ф<strong>о</strong>т<strong>о</strong>)<br />

http://jurassic.ru/

http://jurassic.ru/<br />

ТАБЛИЦА II

ТАБЛИЦА III<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacridae (п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Archimylacrinae)<br />

Фиг. 1. Phyloblatta picturata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье X3,2;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 2. Balachonoblatta zheltojarica Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 3,2; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 3. Grypoblattites tortuosus Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 4,9; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 4. Archaeotiphites petrablattinoides Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 3,7; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

http://jurassic.ru/

http://jurassic.ru/<br />

i л i. л и ц \ in

ТАБЛИЦА IV<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archimylacridae (п<strong>о</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Archimylacrinae<br />

и Spiloblattininae) и Poroblattinidae<br />

Фиг. 1. Phyloblatta poryvajkaensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 3,8; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, <strong>в</strong>ерхнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 2. Phyloblatta kuznetskiensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 3,7; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 3. Zavjaloblatta rotundata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 2,7; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 4. Ishanoblattina fodinensis (Becker-Migdisova); н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 4,0; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, <strong>в</strong>ерхнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1962)<br />

http://jurassic.ru/

http://jurassic.ru/<br />

ТА ЬЛ И U. А IV

ТАБЛИЦА V<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Euryptilonidae и Ideliidae<br />

Фиг. 1. Euryptilon blattoides Martynov; <strong>о</strong>бщий <strong>в</strong>ид, X 3,9;<br />

н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 2. Euryptilodes cascus Sharov. переднее <strong>к</strong>рыл<strong>о</strong>, X 5,0;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 3. Archidelia ovata Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, X 2,3;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 4. Aenigmidelia incredibilis Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, X 2,2;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

http://jurassic.ru/

http://jurassic.ru/<br />

ТАБЛИЦА V

ТАБЛИЦА VI<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> IdeSiidae, Narkemidae и Liomopteridae<br />

Фиг. 1. Rachimentomon reticulatum G. Zalessky; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 2,7; <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, н. пермь, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий,<br />

1939)<br />

Фиг. 2. Stenaropodites permiakovae (М. Zalessky); переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X 2,7; <strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Приур<strong>а</strong>лье<br />

(М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1929)<br />

Фиг. 3. Camptoneurites reticulatus Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 6,9; <strong>в</strong> пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1928)<br />

Фиг. 4. Narkemina angustiformis Sharov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 2,8; <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)<br />

Фиг. 5—6. Liomopteridae incertae sedis; передне<strong>с</strong>пин<strong>к</strong>и, X 1 1;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

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Г Л Б ЛИЦА VI

ТАБЛИЦА Vll<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Liomopteridae, Sylvaphlebidae и Lemmatophoridae<br />

Фиг. 1. Parapermula sibirica Sharov; переднее <strong>к</strong>рыл<strong>о</strong>. X 2,9;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 2. Climaconeurites asiaticus Sharov; переднее <strong>к</strong>рыл<strong>о</strong> X 4,6;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 3. Liomopterella vulgaris Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, Х4,5;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>. Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 4. Kaltanella lata Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, X 5,0; н. пермь,<br />

<strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)<br />

Фиг. 5. Kaltanympha thysanuriformis Sharov; нимф<strong>а</strong>, X 6,5;<br />

н. пермь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1961)<br />

Фиг 6. Sylviodes perloides Martynov; <strong>о</strong>бщий <strong>в</strong>ид, X 2,0;<br />

н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (<strong>о</strong>риг. ф<strong>о</strong>т<strong>о</strong> п<strong>а</strong>р<strong>а</strong>тип<strong>а</strong>)<br />

Фиг. 7. Lemmatophora typa Sellards; <strong>о</strong>бщий <strong>в</strong>ид, X 7,0; н. пермь;<br />

С. Амери<strong>к</strong><strong>а</strong> (Carpenter, 1935}<br />

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ТАБЛИЦА VII

ТАБЛИЦА VIII<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Atactophlebiidae, Megakhosaridae, Perlopseidae<br />

и Taeniopterygidae<br />

Фиг. 1. Atactophlebla termitoides Martynov; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 2,0; <strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Приур<strong>а</strong>лье (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1930)<br />

Фиг. 2. Megakhosara explicata Sharov, переднее <strong>к</strong>рыл<strong>о</strong>, X 4,3;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 3. Gurianovella blattoides (Martynov); нимф<strong>а</strong>, X 4,7;<br />

н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong> (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940)<br />

Фиг. 4. Perlopsis filicornis Martynov; <strong>о</strong>бщий <strong>в</strong>ид, Х2,6; н. пермь,<br />

<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (<strong>о</strong>риг. ф<strong>о</strong>т<strong>о</strong> плези<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 5. Mesonemura turanica Martynov; переднее <strong>к</strong>рыл<strong>о</strong>, X 0,4;<br />

н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

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I V I. Л И Ц V VIII

ТАБЛИЦА IX<br />

Plecoptera incertae sedis и <strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>о</strong> Sthenaropodidae<br />

Фиг. 1. Mesoleuctra gracilis Brauer, Redtenbacher, Ganglbauer;<br />

нимф<strong>а</strong>, X 5,0; н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ф<strong>о</strong>т<strong>о</strong> плези<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 2. Platyperla platypodia Brauer, Redtenbacher, Ganglbauer;<br />

нимф<strong>а</strong>, X 5,0; н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (<strong>о</strong>риг. ф<strong>о</strong>т<strong>о</strong> плези<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 3. Adumbratus extentus Sharov; переднее <strong>к</strong>рыл<strong>о</strong>, X 4,7;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>,<br />

1961)<br />

Фиг. 4. Kamia angustovenosa Martynov (Permacridites maximus<br />

Martynov); переднее <strong>к</strong>рыл<strong>о</strong>, X 2,5; <strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий<br />

яру<strong>с</strong>, Приур<strong>а</strong>лье (<strong>о</strong>риг. ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

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ТАБЛИЦА IX

ТАБЛИЦА X<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Oedischiidae и Tcholmanvissiidae<br />

Фиг. 1. Jasvia reticulata G. Zalessky; переднее <strong>к</strong>рыл<strong>о</strong>, X 4,0;<br />

н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (ф<strong>о</strong>т<strong>о</strong> Ю. М. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong>)<br />

Фиг. 2. Metoedischia magnifica Martynov; переднее <strong>к</strong>рыл<strong>о</strong> X 3,2;<br />

<strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1928)<br />

Фиг. 3. Pinegia longipes (Martynov); переднее <strong>к</strong>рыл<strong>о</strong>, X 2,4;<br />

н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 195П<br />

Фиг. 4. Pinegia longipes (Martynov); <strong>о</strong>бщий <strong>в</strong>ид, X 3,7; <strong>в</strong>. пермь,<br />

<strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Ур<strong>а</strong>л (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1940)<br />

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Г Л R Л и u \ \

ТАБЛИЦА XI<br />

Фиг. 1. Permelcana sojanensise Sharov: a — переднее <strong>к</strong>рыл<strong>о</strong>; б —<br />

з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 8,1; <strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я<br />

<strong>о</strong>бл. (Ш<strong>а</strong>р<strong>о</strong><strong>в</strong>, 1962)<br />

Фиг. 2. Abollus sp.; <strong>о</strong>бщий <strong>в</strong>ид, X 1,2; <strong>в</strong>. юр<strong>а</strong>, К<strong>а</strong>з<strong>а</strong>х<strong>с</strong>т<strong>а</strong>н (Ге<strong>к</strong><strong>к</strong>ер,<br />

1948)<br />

Фиг. 3. Pseudohagla pospelovi (О. Martynova); переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 3,7; юр<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 4. Archaboilus kisyl-kiensis Martynov; <strong>о</strong><strong>с</strong>н<strong>о</strong><strong>в</strong><strong>а</strong>ние переднег<strong>о</strong><br />

<strong>к</strong>рыл<strong>а</strong>, X 2,8; н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

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ТАБЛИЦА XI

ТАБЛИЦА XII<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Blattoprosbolidae и Prosbolidae<br />

Фиг. 1. Blattoprosbole tomiensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 5,5: a — п<strong>о</strong>зити<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>; б — нег<strong>а</strong>ти<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>;<br />

<strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н, нижнеб<strong>а</strong>л<strong>а</strong>х<strong>о</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1958)<br />

Фиг. 2. Evansia speciosa Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье, X 3;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961)<br />

Фиг. 3. Evansia rectimarginata Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>;<br />

X 3; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 4. Prosbole sojanensis Martynov; н<strong>а</strong>д<strong>к</strong>рылье, X 2,3;<br />

<strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>,<br />

1935)<br />

Фиг. 5. Kaltanetta nigra Becker-Migdisova: <strong>а</strong> — н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 9,1; б— з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 10,5; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 6. Pervestigia veteris Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

X 7,9; и. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 7. Permocicada umbrata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 5,7; <strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

(М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1928)<br />

Фиг. 8. Sojanoneura kazanensis М. Zalessky; н<strong>а</strong>д<strong>к</strong>рылье, X 4;<br />

<strong>в</strong>. пермь, <strong>к</strong><strong>а</strong>з<strong>а</strong>н<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Арх<strong>а</strong>нгель<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1940)<br />

Фиг. 9. Prosboloneura kondomensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 11,2; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

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ТА ЬЛ И Ц А XII

ТАБЛ ИЦА XIII<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Palaeontinidae, Tettigarctidae, Cicadidae<br />

и Prosbolidae<br />

Фиг. 1. Orthoscytina pallida Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 10,5; <strong>в</strong>. пермь, ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 2. Cicadoprosbole sogutensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 3,2: <strong>а</strong> — п<strong>о</strong>зити<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>; б — нег<strong>а</strong>ти<strong>в</strong>ный <strong>о</strong>тпеч<strong>а</strong>т<strong>о</strong><strong>к</strong>;<br />

рэт, И<strong>с</strong><strong>с</strong>ы<strong>к</strong>-Куль<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (ф<strong>о</strong>т<strong>о</strong> <strong>с</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 3. Soljuktocossus prosboloides Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 2,8; н. юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1949)<br />

Фиг. 4. Plachutella rotundata Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

X 3,7; н. юр<strong>а</strong>, Ср. Азия (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949)<br />

Фиг. 5. Pseudocossus zemcuznicovi Martynov; н<strong>а</strong>д<strong>к</strong>рылье, X 1,7;<br />

н. юр<strong>а</strong>, Ир<strong>к</strong>ут<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1931)<br />

Фиг. 6. Plachutella derupta Becker-Migdisova; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>,<br />

X 3; н. юр<strong>а</strong>, Ср. Азия. (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1949)<br />

Фиг. 7. Tomiocicada gorbunovi Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 2,5; ми<strong>о</strong>цен, 3. Сибирь (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1954)<br />

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ТАБЛИЦА XIII

ТАБЛИЦА XIV<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Scytinopteridae, Jassidae и Pereboriidae<br />

Фиг. 1. Scytinoptera picturata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье.<br />

X 9,3; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 2. Scytinoptera sp.; з<strong>а</strong>днее <strong>к</strong>рыл<strong>о</strong>, X 8,1; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я<br />

<strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>,<br />

1961)<br />

Фиг. 3. Permolamproptera grandis Becker-Migdisova; з<strong>а</strong>днее<br />

<strong>к</strong>рыл<strong>о</strong>, X 6,8; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 4. Kaltanospes kuznetskiensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 13,1; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 5. Permododa membracoides Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 10,3; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 6. Durgades miocoenica Becker-Migdisova; <strong>о</strong>бщий <strong>в</strong>ид,<br />

X 10,6; ми<strong>о</strong>цен, Ср. Азия (ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 7. Neuropibrocha ramisubcostalis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 3,4; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 8. Neuropibrocha paradunstanioides Becker-Migdisova;<br />

н<strong>а</strong>д<strong>к</strong>рылье, X 3,1; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 9. Borisrohdendorfia picturata Becker-Migdisova; ч<strong>а</strong><strong>с</strong>ть<br />

з<strong>а</strong>днег<strong>о</strong> <strong>к</strong>рыл<strong>а</strong>, X 3,2; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959)<br />

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ТАБЛИЦА XIV

ТАБЛИЦА XV<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong> Mesocixiidae, Ipsviciidae, Archescytinidae,<br />

Permaphidopseidae, Coleoscytidae, Cicadopsyllidae<br />

и Permaleurodidae<br />

Фиг. 1. Mesocixiella asiatica Martynov; н<strong>а</strong>д<strong>к</strong>рылые, X 12,5;<br />

н. юр<strong>а</strong>, Ср. Азия (М<strong>а</strong>ртын<strong>о</strong><strong>в</strong>, 1937)<br />

Фиг. 2. Reticulocicada brachyptera Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье,<br />

X 8,9; <strong>в</strong>. пермь, ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

•Фиг. 3. Tychtoscarta sokolovensis Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье<br />

и передне<strong>с</strong>пин<strong>к</strong><strong>а</strong>, X 12,2; <strong>в</strong>. пермь, ерун<strong>а</strong><strong>к</strong><strong>о</strong><strong>в</strong><strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>,<br />

Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 4. Permopsylla kuznetskiensis Becker-Migdisova; тел<strong>о</strong> <strong>с</strong><br />

<strong>к</strong>рыльями; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 5. Kaltanaphis permiensis Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>;<br />

.н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959)<br />

Фиг. 6. Kaltanoscyta reticulata Becker-Migdisova; н<strong>а</strong>д<strong>к</strong>рылье;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (Бе<strong>к</strong><strong>к</strong>ер-<br />

Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1959)<br />

Фиг. 7. Sojanopsylla kaltanica Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>;<br />

н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

"Фиг. 8. Cicadopsis rugosipenna Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

х9,3; н. пермь, <strong>к</strong>узнец<strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>, 1961)<br />

Фиг. 9. Permaleurodes rotundatus Becker-Migdisova; <strong>к</strong>у<strong>к</strong><strong>о</strong>л<strong>к</strong><strong>а</strong>,<br />

х8,3; <strong>в</strong>. пермь, ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(Бе<strong>к</strong><strong>к</strong>ер-Мигди<strong>с</strong><strong>о</strong><strong>в</strong><strong>а</strong>)<br />

У<strong>в</strong>еличения<br />

иг. 4 х 21; фиг. 5 X 8,4; фиг. 6 х 9,5; фиг. 7 х 10.<br />

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ТАЬЛИЦА XV

ТАБЛИЦА XVI<br />

Отряды Psocoptera и Perielytrodea<br />

Фиг. 1. Surijokopsocus radtchenkoi Becker-Migdisova; переднее<br />

<strong>к</strong>рыл<strong>о</strong>, X И; <strong>в</strong>. пермь, ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий<br />

б<strong>а</strong><strong>с</strong><strong>с</strong>ейн (ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 2. Zoropsocus tomiensis Becker-Migdisova; переднее <strong>к</strong>рыл<strong>о</strong>,<br />

X 18; <strong>в</strong>. пермь, ильин<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>с</strong><strong>в</strong>ит<strong>а</strong>, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн<br />

(ф<strong>о</strong>т<strong>о</strong> г<strong>о</strong>л<strong>о</strong>тип<strong>а</strong>)<br />

Фиг. 3. Perielytron mirabile G. Zalessky: a—цел<strong>о</strong>е н<strong>а</strong><strong>с</strong>е<strong>к</strong><strong>о</strong>м<strong>о</strong>е;<br />

б — переднее <strong>к</strong>рыл<strong>о</strong>; н. пермь, <strong>к</strong>унгур<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>, Че<strong>к</strong><strong>а</strong>рд<strong>а</strong><br />

(Ю. З<strong>а</strong>ле<strong>с</strong><strong>с</strong><strong>к</strong>ий, 1943)<br />

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О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

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ТАБЛИЦА XVI

ТАБЛИЦА XVII<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Austrolimulidae<br />

Austrolimulus fleetcheri Riek, 1955, X 2; <strong>с</strong>р. три<strong>а</strong><strong>с</strong>, Н<strong>о</strong><strong>в</strong>ый Южный<br />

Уэл<strong>с</strong> (А<strong>в</strong><strong>с</strong>тр<strong>а</strong>лия) (ф<strong>о</strong>т<strong>о</strong> при<strong>с</strong>л<strong>а</strong>н<strong>о</strong> Ри<strong>к</strong><strong>о</strong>м)<br />

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ТАБЛИЦА XVII

ТАБЛИЦА XVIII<br />

Н<strong>а</strong>д<strong>с</strong>емей<strong>с</strong>т<strong>в</strong><strong>а</strong> Eurypteridea и Pterygotidea<br />

Фиг. 1. Ни ь'.milleria зр. Чешуе- или терр<strong>а</strong><strong>с</strong><strong>о</strong><strong>в</strong>идные линии, <strong>о</strong>бр<strong>а</strong>зу!<br />

_>щие <strong>с</strong><strong>к</strong>ульптуру тергит<strong>о</strong><strong>в</strong>, X 10; н. де<strong>в</strong><strong>о</strong>н, С. Сибирь<br />

(юг<strong>о</strong>-з<strong>а</strong>п<strong>а</strong>дный Т<strong>а</strong>ймыр) (<strong>к</strong><strong>о</strong>лл. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong><br />

ин<strong>с</strong>титут<strong>а</strong>, № 1138/4)<br />

Фиг. 2. Hughmilleria sp. indet. Пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> д<strong>в</strong>умя ле<strong>в</strong>ыми <strong>к</strong><strong>о</strong>нечн<strong>о</strong><strong>с</strong>тями,<br />

X 1; <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н (н. жи<strong>в</strong>ет<strong>с</strong><strong>к</strong>ий яру<strong>с</strong>), Ту<strong>в</strong><strong>а</strong><br />

(Novojilov, 1958)<br />

Фиг. 3, 4. Hughmilleria lata? Stormer, 1933: 3 — тел<strong>о</strong> <strong>с</strong> <strong>о</strong>бл<strong>о</strong>м<strong>а</strong>нн<strong>о</strong>й<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>о</strong>й и без п<strong>о</strong><strong>с</strong>ледних <strong>с</strong>егмент<strong>о</strong><strong>в</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы;<br />

4 — п<strong>о</strong><strong>с</strong>ледние <strong>с</strong>егменты <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы т<strong>о</strong>г<strong>о</strong> же э<strong>к</strong>земпляр<strong>а</strong>,<br />

X 2; ? н. де<strong>в</strong><strong>о</strong>н, В. Сибирь (Кр<strong>а</strong><strong>с</strong>н<strong>о</strong>яр<strong>с</strong><strong>к</strong>ий <strong>к</strong>р<strong>а</strong>й)<br />

(Novojilov, 1958)<br />

Фиг. 5. Tylopterella menneri (Novojilov), 1958. Тел<strong>о</strong> без тель<strong>с</strong><strong>о</strong>н<strong>а</strong>,<br />

<strong>в</strong>идны пр<strong>а</strong><strong>в</strong>ые х<strong>о</strong>дильные и пл<strong>а</strong><strong>в</strong><strong>а</strong>тельн<strong>а</strong>я н<strong>о</strong>ги,<br />

X 3; н. де<strong>в</strong><strong>о</strong>н, С. Сибирь (юг<strong>о</strong>-з<strong>а</strong>п<strong>а</strong>дный Т<strong>а</strong>ймыр)<br />

(Novojilov, 1958)<br />

Фиг. 6, 7. Pterygotus (Pterygotus) Agassiz, 1839: 6 — фр<strong>а</strong>гмент<br />

неп<strong>о</strong>д<strong>в</strong>ижн<strong>о</strong>й <strong>в</strong>ет<strong>в</strong>и <strong>к</strong>лешни <strong>с</strong> м<strong>о</strong>щными зубц<strong>а</strong>ми; 7 —<br />

фр<strong>а</strong>гмент, тергит<strong>а</strong>, X 1; <strong>с</strong>р. де<strong>в</strong><strong>о</strong>н, Ш<strong>о</strong>тл<strong>а</strong>ндия (<strong>к</strong><strong>о</strong>лл.<br />

Г<strong>а</strong>м<strong>а</strong>лея, № 2084/36 и 2084/37)<br />

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ТАБЛИЦА XVTI1

ТАБЛ ИЦА XIX<br />

Семей<strong>с</strong>т<strong>в</strong><strong>а</strong>: Stylonuridae, Rhenopteridae, Mucteropidae<br />

Фиг. 1—4. Stylonurus ruedemanni? Stormer, 1933, X 2: 1 —<br />

пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>; 2 — три передние тергит<strong>а</strong> <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы; 3 —<br />

<strong>о</strong>пер<strong>к</strong>улум <strong>с</strong> генит<strong>а</strong>льный прид<strong>а</strong>т<strong>к</strong><strong>о</strong>м <strong>с</strong><strong>а</strong>мц<strong>а</strong>; 4 — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

и пять тергит<strong>о</strong><strong>в</strong> мез<strong>о</strong><strong>с</strong><strong>о</strong>мы, деф<strong>о</strong>рмир<strong>о</strong><strong>в</strong><strong>а</strong>нные и<br />

з<strong>а</strong><strong>к</strong>рытые п<strong>о</strong>р<strong>о</strong>д<strong>о</strong>й <strong>с</strong> ле<strong>в</strong><strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны; н. де<strong>в</strong><strong>о</strong>н, Х<strong>а</strong><strong>к</strong><strong>а</strong><strong>с</strong><strong>с</strong>ия<br />

(Novojilov, 1958).<br />

Фиг. 5. Rhenopteridae; фр<strong>а</strong>гмент из ше<strong>с</strong>ти тергит<strong>о</strong><strong>в</strong>? <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

X 2; 3. Сибирь, Кузнец<strong>к</strong>ий б<strong>а</strong><strong>с</strong><strong>с</strong>ейн, <strong>с</strong>р. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н<br />

К<strong>о</strong>лл. П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong><strong>о</strong>г<strong>о</strong> ин<strong>с</strong>титут<strong>а</strong>, № 1293/8.<br />

Фиг. 6. Glaucodes mathieui Pruvost, 1923; пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong> пер<strong>в</strong>ым<br />

<strong>к</strong>рупным <strong>с</strong>егмент<strong>о</strong>м <strong>о</strong>пи<strong>с</strong>т<strong>о</strong><strong>с</strong><strong>о</strong>мы, X 0,5; <strong>в</strong>. <strong>к</strong><strong>а</strong>рб<strong>о</strong>н,<br />

Бельгия (Pruvost, 1923)<br />

Фиг. 7. Borchgrevinkium taimyrense Novojilov, 1958, X 3, н. де­<br />

<strong>в</strong><strong>о</strong>н, С. Сибирь (юг<strong>о</strong>-з<strong>а</strong>п<strong>а</strong>дный Т<strong>а</strong>ймыр) (Novojilov,<br />

1958)<br />

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ТАБЛИЦА XIX

ТАБЛИЦА XX<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Eurypteridae<br />

Фиг. 1—2. Erleopterus microphthalmus microphthalmus (Hall)<br />

1859. 1 —пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>, X 1,2; 2 — фр<strong>а</strong>гмент пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>с</strong> <strong>в</strong>идим<strong>о</strong>й<br />

груб<strong>о</strong>й непр<strong>а</strong><strong>в</strong>ильн<strong>о</strong>й «пузырьч<strong>а</strong>т<strong>о</strong>й» <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й,<br />

X 3. Силур, С. Амери<strong>к</strong><strong>а</strong> (Нью-Й<strong>о</strong>р<strong>к</strong>). (Kjellesvig-<br />

Waering, 1958).<br />

Фиг. 3—-5. Е. microphthalmus eriensis (Whitfield) 1882. 3 — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong><br />

<strong>с</strong> 3-й х<strong>о</strong>дильн<strong>о</strong>й н<strong>о</strong>г<strong>о</strong>й и передняя ч<strong>а</strong><strong>с</strong>ть мез<strong>о</strong><strong>с</strong><strong>о</strong>мы,<br />

X 2,8; 4 — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> <strong>с</strong>низу, X 2; 5 — <strong>о</strong>пи<strong>с</strong>т<strong>с</strong><strong>о</strong>м<strong>а</strong>,<br />

X 1,3. Силур, С. Амери<strong>к</strong><strong>а</strong> (Ог<strong>а</strong>й<strong>о</strong>). (Kjellesvig-Waering,<br />

1958)<br />

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ТАЬЛ И Ц \ XX

ТАБЛИЦА XXI<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Hibbertopteridae<br />

Фиг. 1—3. Campylocephalus oculatus (Kutorga) 1838. 1 — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong>,<br />

7з -н<strong>а</strong>тур<strong>а</strong>льн<strong>о</strong>й <strong>в</strong>еличины; 2—ле<strong>в</strong>ый з<strong>а</strong>дний<br />

уг<strong>о</strong>л пр<strong>о</strong><strong>с</strong><strong>о</strong>мы <strong>в</strong> н<strong>а</strong>тур<strong>а</strong>льную <strong>в</strong>еличину; 3 —фр<strong>а</strong>гмент<br />

пр<strong>а</strong><strong>в</strong><strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны, <strong>в</strong> <strong>к</strong><strong>о</strong>т<strong>о</strong>р<strong>о</strong>м <strong>о</strong>т<strong>с</strong>ут<strong>с</strong>т<strong>в</strong>ует з<strong>а</strong>дний уг<strong>о</strong>л,<br />

н<strong>а</strong>тур<strong>а</strong>льн<strong>а</strong>я <strong>в</strong>еличин<strong>а</strong>. В. пермь, Б<strong>а</strong>ш<strong>к</strong>ирия. (Kutorga,<br />

1838).<br />

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I \ Ь Л И II V \XI

ТАБЛИЦА XXII<br />

Семей<strong>с</strong>т<strong>в</strong><strong>о</strong> Mycteropidae<br />

Фиг. 1—3. Mycterops ordinatus С<strong>о</strong>ре, 1886. 1 — пр<strong>о</strong><strong>с</strong><strong>о</strong>м<strong>а</strong> и пер<strong>в</strong>ый<br />

тергит мез<strong>о</strong><strong>с</strong><strong>о</strong>мы, <strong>о</strong>бл<strong>о</strong>м<strong>а</strong>нные <strong>с</strong> пр<strong>а</strong><strong>в</strong><strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны;<br />

2—фр<strong>а</strong>гмент ле<strong>в</strong><strong>о</strong>й <strong>с</strong>т<strong>о</strong>р<strong>о</strong>ны пр<strong>о</strong><strong>с</strong><strong>о</strong>мы; 3 — фр<strong>а</strong>гмент<br />

з<strong>а</strong>днег<strong>о</strong>? тергит<strong>а</strong> <strong>с</strong> чешуе<strong>в</strong>идн<strong>о</strong>й <strong>с</strong><strong>к</strong>ульптур<strong>о</strong>й. В<strong>с</strong>е <strong>о</strong>бр<strong>а</strong>зцы<br />

<strong>в</strong> н<strong>а</strong>тур<strong>а</strong>льную <strong>в</strong>еличину. Пен<strong>с</strong>иль<strong>в</strong><strong>а</strong>н<strong>с</strong><strong>к</strong>ий <strong>о</strong>тдел<br />

(<strong>а</strong>ллег<strong>а</strong>н<strong>с</strong><strong>к</strong><strong>а</strong>я ф<strong>о</strong>рм<strong>а</strong>ция); С. Амери<strong>к</strong><strong>а</strong> (Пен<strong>с</strong>иль<strong>в</strong><strong>а</strong>ния).<br />

(Kjellesvig-Waering, 1959).<br />

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41. О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

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2<br />

ТА Б.II И ЦА XXII

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АЛФАВИТНЫЙ УКАЗАТЕЛЬ<br />

СИСТЕМАТИЧЕСКИХ НАЗВАНИЙ<br />

Н<strong>о</strong>мер<strong>а</strong> <strong>с</strong>тр<strong>а</strong>ниц, где д<strong>а</strong>ны <strong>о</strong>пи<strong>с</strong><strong>а</strong>ния <strong>о</strong>пределенных <strong>с</strong>и<strong>с</strong>тем<strong>а</strong>тиче<strong>с</strong><strong>к</strong>их групп или<br />

р<strong>о</strong>д<strong>о</strong><strong>в</strong>, <strong>в</strong>ыделены <strong>к</strong>ур<strong>с</strong>и<strong>в</strong><strong>о</strong>м (жирным шрифт<strong>о</strong>м).<br />

Н<strong>о</strong>мер<strong>а</strong> <strong>с</strong>тр<strong>а</strong>ниц, где д<strong>а</strong>ны из<strong>о</strong>бр<strong>а</strong>жения пред<strong>с</strong>т<strong>а</strong><strong>в</strong>ителей тех или иных р<strong>о</strong>д<strong>о</strong><strong>в</strong>,<br />

<strong>о</strong>тмечены з<strong>в</strong>езд<strong>о</strong>ч<strong>к</strong><strong>о</strong>й.<br />

АЬ<strong>а</strong>рШ<strong>о</strong>п 52*, 53<br />

Abashevia 125*, 126<br />

АЫ<strong>а</strong> 273<br />

Abietinae 198<br />

Ablator 507, 511*<br />

Abliguritor 507, 511*<br />

Aboilidae 150<br />

Aboilus 151*<br />

Absoluta 176*<br />

Acalyptrata 342<br />

Acanaloniidae 181<br />

Acantherpestes 22*<br />

Acanthiidae 216<br />

Acanthocepkala 221*<br />

Acanhocephalina 221*<br />

Acanthoraeridae 336<br />

Acaridae 452*<br />

Acaridea 451, 452<br />

Acaridiae 384*, 385, 451<br />

Acariformes 378, 382, 383, 384*,<br />

385, 447, 448, 450, 451, 472,<br />

474<br />

Acari 383, 450<br />

Acarina 383, 385, 444, 449, 450,<br />

485<br />

Acaromorpha 378, 379, 380, 382,<br />

383, 384*, 385, 439, 445, 447,<br />

448, 479, 450<br />

Aceria 469*<br />

Achilidae 181, 185*. 186<br />

Achilixiidae 181<br />

Acmeoblatta 103<br />

Acoblatta 94<br />

Acocephalites 177<br />

Acosmoblatta 94<br />

Acrid idae 156*, 157<br />

Acrididea 155, 156<br />

Acridites 146, 157<br />

Acridoxeninae 155<br />

Acrocera 335*<br />

Acroceridae 334, 336<br />

41 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

Acrocerides 336<br />

Acrogiomeridae 23<br />

Acrokreischeria 487*, 488<br />

Acrometa 505, 509*<br />

Acromocoris 212<br />

Acropsopil ion idae 483<br />

Acttnoblatta 94<br />

Actinoblattula 103<br />

Actinochaeta 385<br />

Actinochitinosi 450<br />

Acfinoderma 385<br />

Actinomylacris 97<br />

Actinoscytina 225<br />

Actinoscytinidae 208, 210, 224*<br />

Actinophlebia 279<br />

Aculeata 349<br />

Acutiramus 415*<br />

Adamator 515, 516*<br />

Adelges 196*<br />

Adelgidae 196*, 197<br />

Adelginae 197<br />

Adelidae 304<br />

Adelinae 304<br />

Adeloblatta 94<br />

Adelocoris 212<br />

Adeloneuridae 122<br />

Ademosyne 252*, 253<br />

Ademosynoides 252*, 253<br />

Aderidae 263*<br />

Aderus 263*<br />

Adiphlebiidae 117<br />

Adjunctor 503, 504*<br />

Adjutor 503, 504*<br />

Adjutoridae 501, 503, 504*<br />

Admissor 503, 504*<br />

Adorator 501<br />

Adulatrix 507, 512*<br />

Adumbratus 145*, 146<br />

Aenigmatetla 116*, 117<br />

Aenigmatiasiidae 339<br />

Aenigmatidia 71*, 72<br />

539<br />

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Aenigmatidiidae 72<br />

Aenigmatodes 117<br />

Aenigmatodidae 117<br />

Aenigmidelia 120*. 121<br />

Aenimocoleus 251*, 253<br />

Aeolothripidae 237, 238<br />

Aeromyrma 357*<br />

Aerophasma 160*<br />

Aerophasmatidae 160<br />

Aeroplanidae 160<br />

Aeshnidae 83, 85<br />

Aeschnidiella 84*. 85<br />

Aeschnidiidae 83, 85<br />

Aeschnidiopsis 84*, 85<br />

Aeschnidium 84*, 85<br />

Aeschnopsis 85<br />

Aetaletidae 46<br />

Aetalion 178*<br />

Aetalionidae 173, 178*, 179<br />

Aetalionoideae 179<br />

Aetophlebiidae 117<br />

Aetophlebiopsis 131*, 132<br />

Agaontidae 352<br />

Agapetus 295<br />

Agelena 505, 510*<br />

Agelenidae 505, 510*, 517<br />

Agetochorista 287*<br />

Agetochoristella 287<br />

Agetopanorpa 288<br />

Agetopanorpidae 286<br />

Aglaspidida 384*, 385, 389<br />

390, 391*, 393, 404, 407<br />

Aglaspididae 388, 389, 391<br />

392*<br />

Aglaspella 389, 393<br />

Aglaspida 382, 390<br />

Aglaspis 389, 391*. 392*<br />

Aglaspoides 389, 393<br />

Agonoscena 207*<br />

Agraepterinae 155<br />

Agriolestes 79*

Agrion 79*<br />

Agrionidae 80, 81<br />

Agrionidea 76, 79*, 80<br />

Agromyzidae 342<br />

Aissoblatta 89, 90*<br />

Aldotrifidus 181<br />

Alena 272<br />

Aleurodes 203*<br />

Aleurodidae 203, 204<br />

Aleurodidea 203*<br />

Aleurolobus 203*<br />

Alicorhagia 463<br />

Alicorhagiidae 449 , 463*, 464<br />

Allactoneurites 323*<br />

Alleculidae 264*, 265<br />

Alloscorpius 427*, 429, 431*<br />

Allothrombidium 468, 469*<br />

Altajopanorpa 284*, 285<br />

Alysiinae 349<br />

Amaurobiidae 516*, 517<br />

Amaurobiidea 515, 5J7<br />

Amaurobiiformia 517<br />

Amaurobius 516*, 517<br />

Amatidae 307<br />

Amblyblatta 94<br />

Amblymylacris 97<br />

Amblypygi 380, 382, 383, 384*,<br />

385, 402, 403, 433 , 435*, 436,<br />

437, 439, 440, 441, 444, 474,<br />

475, 476, 485, 486, 494<br />

Amekeoblatta 99, 100*<br />

Amerus 454<br />

Ameronotridae 456<br />

Atnetroblatta 95<br />

Ametropodidae 64<br />

Ammoxenidea 499<br />

Amoeboblatta 94<br />

Amorphoblatta 94<br />

Amphientomidae 228, 231<br />

Amphientominae 231<br />

Amphientomum 232*<br />

Amphipterygidae 80<br />

Anacontium 399<br />

Anacoloptera 226<br />

Analgesidea 451<br />

Anamorpha 25, 26<br />

Anaplecta 108*<br />

Anaplectinae 108*, 110<br />

Anaplectidae 110<br />

Anaprosbole 174, 175*<br />

Anaspis 264*<br />

Anatopynia 317*<br />

A п<strong>а</strong>х 84*<br />

Anaxyela 346, 347*<br />

Anaxyelidae 346<br />

Ancylopsocus 228<br />

Andrena 358*<br />

Androgeus 503, 507*<br />

Anergetus 116*<br />

Angarapsychops 277*<br />

Angarocaris 389, 391, 392<br />

Anhomalophlebiidae 122<br />

Anisophlebia 82*<br />

Anisoptera 43, 73, 83, 84*, 85<br />

Anisozygoptera 43, 73, 80*, 81, 82*,<br />

83<br />

Anniculus 500*, 501<br />

Annulipalpia 295 , 296*<br />

Anobiidae 260*<br />

Anobium 260*<br />

Anoeciinae 197<br />

Anoetidea 451<br />

Anomalogramma 143*, 144<br />

Anomalogrammatidae 142, 144<br />

Anomaloscytina 170, 174<br />

Anomoblatta 94<br />

Anomomylacris 97<br />

Anomoscyta 174<br />

Anomothemis 83<br />

Anoplura 162*, 226<br />

Anormochoristidae 286<br />

Anosmus 210<br />

Antefungivora 326*, 325<br />

Anthicidae 263*, 264<br />

Anthicus 263*<br />

Anthocoridae 216*, 2/7<br />

Anthocorides 217<br />

Anthocoris 216*<br />

Anthomyiidae 343, 344<br />

Anthomyiinae 344<br />

Anthracoblattina 94<br />

Anthracoiul s 23<br />

Anthracomarti 380, 383, 384*,<br />

385, 444, 474, 475, 485, 486, 400<br />

Anthracomartidae 490, 491*, 492*<br />

Anthracomartus 488, 489, 492<br />

Anthracophrunus 492*<br />

Anthracoptilidae 117<br />

Anthracoscorpii 424<br />

Anthracoscorpius 431*<br />

Anthracosiro 484*, 486<br />

Anthracosironidae 484*. 486<br />

Anthracothremmidae 117<br />

Anthribidae 266, 267*<br />

Anthribus 267*<br />

Antiquamedia 326*, 328<br />

Anyphaena 507, 511*<br />

Anyphaenidae 507<br />

Anystiidae 466, 467*<br />

Anystiidea 462, 466<br />

Anystis 466, 467*<br />

Anystoidea 466<br />

Anzinia 488<br />

A pate 260*<br />

Apeirophlebia 278<br />

Apempherus 94<br />

Aphalarinae 206<br />

Aphaniptera 240*, 283, 284, 345*<br />

Aphantochillidae 509<br />

Aphantomartus 489*<br />

Aphelomylacris 97<br />

Apheloscyta 190<br />

Aphiden 198<br />

Aphididae 195, 196*. /07, 198<br />

Aphidii 197<br />

Aphidiinae 349<br />

Aphidinae 197, /06?*<br />

Aphidomorpha 180*, 190, 199<br />

Aphidulum 199<br />

Aphis 198*<br />

Aphlebocoridae 209*<br />

Aphlebocoris 209*<br />

Aphodius 256*<br />

Aphrophoroideae 179<br />

Aphryganeura 286<br />

Aphthoroblattina 92*, 04<br />

Apiariae 359<br />

Apidae 350<br />

Apidea 349, 358*, 359<br />

Apioceridae 337<br />

Apion 267*<br />

Apistoblattula 103<br />

Apneumonomorphae 494<br />

Apobolostigmata 468<br />

540<br />

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Apocrita 346, 340, 350*<br />

Apopnidae 209*<br />

Apopnus 209*<br />

Apotypoma 94<br />

Apoxypodes 380, 381, 382, 385,<br />

402, 404, 424, 425, 427<br />

Apsicoria 211*, 212<br />

Apterygota 36*, 37*, 45, 379<br />

Aptyctima 454<br />

Arachnida 378, 379, 380, 381, 383,<br />

384*, 385, 402, 403, 439, 445,<br />

448, 449, 450, 474, 475, 476, 494<br />

Arachnocampidae 330<br />

Arachnoidea 382, 383, 403<br />

Arachnometa 497<br />

Arachnomorphae 385, 493, 494, 407<br />

Aradidae 221*<br />

Aradiens 221<br />

Aradus 221*<br />

Araeopidae 187<br />

Araneae 380, 383, 384*, 385, 437,<br />

440, 441, 444, 474, 475, 482,<br />

485, 486, 402, 494<br />

Araneidae 501, 503, 509*<br />

Araneidea 501<br />

Araneomorphae 384*, 385, 493, 494,<br />

497, 498<br />

Archaboilus 151*, /52<br />

Archaea 503, 504*<br />

Archaeacridites 146<br />

Archaeidae 501, 504*<br />

Archaemetidae 494<br />

Archaemioptera 140, 141*<br />

Archaemiopteridae 140<br />

Archaeocronidae 427*<br />

Archaeocrunoecia 301*<br />

Archaeoctonidae 427*, 426?, 429*<br />

Archaeoctonus 428, 429*<br />

Archaeometa 496*, 497<br />

Archaeometidae 496*, 407<br />

Archaeoneoptera 35, 36*, 37, 85, 86<br />

Archaeoptera 37, 38, 86, 87*<br />

Archaeorrhynchus 267*<br />

A rchaeotiphe 94<br />

Archaeotiphites 91, 92*<br />

Archankothrips 237*<br />

Archebittacus 292<br />

Archedilaropsis 282*<br />

Archegetes 282<br />

Archeglyphis 168*, 170<br />

A rchegocimex 210 *<br />

Archegocimicidae 210*<br />

Archegoceris 210<br />

Archelytridae 115<br />

Archeosmylidae 274, 276<br />

A rcheosmylus 276<br />

A rchepsychops 278<br />

A rchescytina 192*<br />

Archescytinidae 190, 191, 192*, 226<br />

Archescytinidea 190<br />

Archescytinomorpha /00<br />

A rchescytinopsis 194*<br />

A rchexyela 346<br />

A rchicercopis 225<br />

Archidelia 119, 120*<br />

Archicupes 243, 244*<br />

Archidermatoptera 114<br />

Archidiptera 308, 309*<br />

Archihesperinus 321, 323*<br />

Archijassidae 181<br />

Archijassus 180*. 181<br />

Archijulidae 23

Archijulus 23<br />

Ascalaphides 274<br />

Archilycoria 326*. 328<br />

Asemoblatta 94<br />

Archimylacridae 89, 90*, 92*, 93*, Asiachorista 288, 289*<br />

95*,97, 110<br />

Asientomidae 231<br />

Archimylacrinae 89, 90*, 92*, 93*, Asientominae 231<br />

94<br />

Asientomum 232*<br />

Archimylacris 94<br />

Asilici 339<br />

Archipanorpidae 284<br />

Asilidae 339<br />

A rchiphora 340<br />

Asilidea 334, 337<br />

Archipleciofungivora 321, 323*<br />

Asilomorpha 308*, 310, 334 , 335*,<br />

Archipleciomima 321, 323*<br />

338*<br />

Archiprobnis 120*, 121<br />

Aslocixius 185*, 186<br />

Archiprobnisidae 119, 121<br />

Asiocoleidae 241, 242*. 243<br />

Archipsocidae 228, 234<br />

Asiocoleus 242*, 243<br />

Archipsocinae 234<br />

Asiocossus 164*, 166<br />

Archipsocus 232*<br />

Asiodictya 50, 51*<br />

Archipsyche 167<br />

Asionecta 214, 215*<br />

Archipsychops 277<br />

Asiopompidae 117<br />

Archipsylla 232*<br />

Asiopompus 116*, //7<br />

Archipsyllidae 228, 232<br />

Asiopsyllidium 199, 200*<br />

Archirhagio 335*, 337<br />

Asiphum 196*<br />

A rchirhyphus 331 *, 332<br />

Asphondylia 333*<br />

Archisargidae 334<br />

Aspidiotus 201*<br />

Archisargus 333*, 334<br />

Aspidohymen 67*, ft?<br />

Archiscudderia 22*<br />

Aspidohymenidae 67, 6S<br />

Archisialidae 141<br />

Aspidoneura 144<br />

Archisialis 142<br />

Aspidononina 155<br />

Architarbi 383, 450<br />

Aspidothoracidae 67<br />

Architarbida 477<br />

Aspidothorax 67*<br />

Architarbidae 479<br />

Aspongopus 223*<br />

Architarbus 479*, 488<br />

Assamiidae 483<br />

Architendipedidae 316, 317<br />

Astenohymen 71*, 72<br />

Architendipes 317*<br />

Astenohymenidae 72<br />

Archithemis 81<br />

Asyncritidae 117<br />

Archithemistidae 81<br />

Atactoblatta 95<br />

Architipula 312*, 313<br />

Atactophtebia 129*, /30<br />

Architipulidae 312<br />

Atactophlebiidae 124, 129*, /3<<br />

Archizelmira 324*, 326<br />

Atalophlebia 62<br />

Archizelmiridae 326<br />

Atemnidae 442<br />

Archizygoptera 78<br />

Atheroidinae 197<br />

Archoblattina 94<br />

Atimoblatta 94<br />

Archodonata 50, 54*, 55<br />

Atitizon 181<br />

Archoglossopteridae 157, 158* Attagenus 257*<br />

Archoglossopterum 157, 158* Attidae 509<br />

Arctiidae 307<br />

Attoides 515<br />

Arctocupes 243<br />

Attopsis 515<br />

Arctocoleus 268*<br />

Atypidae 497<br />

Arctolocusta 155<br />

Auchenorrhyncha 163, 189*, 190,<br />

Arctotypus 73*, 74<br />

191*<br />

Areomartus 487*, 488<br />

Aulacidae 349, 350<br />

Argasidae 473<br />

Austroblattula 103<br />

Argidae 348<br />

Austrocypha 230<br />

Argiopidae 503<br />

Austr omylacrites 103<br />

Argiopidea 499<br />

Austroidelia 121<br />

Argiopiformia 501<br />

Austrolestes 79*<br />

Argyroneta 494, 505, 510*<br />

Austrolimulidae 390, 39.S, 399*<br />

Argyronetidae 381, 505, 510* Austrolimulus 390, 39*, 399<br />

A rrhythmoblatta 95<br />

Arthrodictyna 516*, 517<br />

Arthrodictynidae 516*, 517<br />

Arthrolycosa 490, 494, 495*, 496<br />

Arthrolycosidae 494, 495*, 496<br />

Arthromygale 494, 495*<br />

Arthromygalidae 494, 495*<br />

Arthropleona 45, 46<br />

Arthropleura 22*, 25<br />

Arthropleuridae 25<br />

Arthropoda 402, 533<br />

Artinska 130<br />

Artitocoblatta 103<br />

Arythaena 468<br />

Ascalaphidae 273, 274<br />

:<br />

Austroperlidae 134<br />

Austroprosbole 170<br />

Austropsocidium 228<br />

Austroscytina 194<br />

Autoblattina 100<br />

Auxanoblatta 94<br />

Auximus 516*, 517<br />

Auxymyiidae 329<br />

Axobittacus 292<br />

Aysheaia 533, 534*<br />

Aysheai idae 533<br />

Baetidae 60, 61*, 62<br />

Baefts 61*, 62<br />

Baetiscidae 60<br />

Baisopteridae 272<br />

541<br />

http://jurassic.ru/<br />

Balachonoblatta 91, 92*<br />

Balaustiidae 468<br />

Balaustium 467*, 468<br />

Bardapteridae 52, 53<br />

Bardapteron 52*, 53<br />

Bardohymen 67*<br />

Bardohymenidae 67<br />

Barroisiblatta 94<br />

Barychetidae 498<br />

Bde/ta 467<br />

Bdellidae 461, 467*<br />

Bdellidea 463, 466<br />

Bdellodes 467*<br />

Bebiosis 232<br />

Beckwithia 389, 393*<br />

Beckwithiidae 389, 390, 391,<br />

393*<br />

Becquerelia 53<br />

Behningiidae 59<br />

Bekkerscytina 194<br />

Belbidae 456<br />

Belbididea 454<br />

Belbites 456<br />

Belinuracea 398<br />

Belimiridae 388, 390, 397, 39«*<br />

Belinuridea 390, 397, 39«*, 399<br />

Belinuropsis 399<br />

Belinurus 390, 398*<br />

Bellinuroopsis 398<br />

Belmontia 287*, 288<br />

Belmontiidae 286<br />

Beloptesis 167<br />

Belostomatidae 213, 2/4<br />

Belostomida 214<br />

Belpsylla 200<br />

Bembicosoma 397<br />

Berlichia 146<br />

Berothidae 277, 2

Blattinopsis 117<br />

Blattodea 37*. 86, 87*. 88, 108*,<br />

111, 241<br />

Blattoprosbole 38, 163*. 164<br />

Blattoprosbolidae 163*<br />

Blattoprosbolomorpha 163<br />

Blattopteroidea 36*, 88<br />

Blattula 100*. 103<br />

Blauchinae 349<br />

Blephariceratomorpha 308*<br />

Blitophaga 256*<br />

Bojoptera 53<br />

Bolitophila 331*<br />

Bolitophilidae 330<br />

Bolilophilidea 319, 330, 331*<br />

Bolitophilinae 330<br />

Boltonopruvostia 94<br />

Bombylarii 339<br />

Bombyliidae 339<br />

Bombyliidea 334, 339<br />

Borchgrevinkium 404, 408, 420,<br />

421*, 422<br />

Boreocixius 185*, 186<br />

Boreoscyta 194*<br />

Boreescitidae 190, 194*<br />

Borisrohdendorfia 190, 191*<br />

Bostrychidae 260*<br />

Bothrideres 261*<br />

Bothriuridae 428<br />

Brachycentrinae 302<br />

Brachycentrus 301*<br />

Brachychthonius 452*, 454<br />

Brachylycosa 490, 491*<br />

Brachymeria 352*<br />

Вrachymylacris 97<br />

Brachyopterus 408, 416, 418, 419*<br />

Brachypyge 491*, 492<br />

Brachypygidae 490<br />

Brachystoma 338*<br />

Braconidae 349<br />

Braconinae 349 •<br />

Braconoides 349<br />

Bradyblatta 94<br />

Bradyporinae 155<br />

Braulomorpha 308*<br />

Brechoblatta 102<br />

Breyeria 51*<br />

Breyeriidae 52<br />

Breyeriidea 50<br />

Brodia 65*<br />

Brodiana 156<br />

Brodiidae 64, 66<br />

Brongniartiella 279<br />

Brongniartiellidae 277, 278*. 279<br />

Bruchidae 266*<br />

Bruchomyia 316*<br />

Bruchus 266*<br />

Bryobiidae 469<br />

Bunaia 389, 396*<br />

Bunodella 397<br />

Bunodes 389, 396*<br />

Bunodidae 388 , 389, 394, 395, 396*<br />

Buprestidae 259*<br />

Buprestis 259*<br />

Burcha 272<br />

Buthidae 428, 432<br />

Buthiscorpius 427*, 431*<br />

Byrrhidae 257*, 258<br />

Byrrhus 257*<br />

Bythinus 254*<br />

Cachoplistinae 154<br />

Cacurgidae 122<br />

Cacurgidea 119, 122, 123*<br />

Caddo 483<br />

Caduceator 507, 512*<br />

Caeciliidae 234<br />

Caecilius 235*<br />

Caelifera 146, /55<br />

Caenidea 57<br />

Caenoblatta 89, 90*<br />

Caenoptilon 289<br />

Caenomyiidae 336, 377<br />

Cainoblattinidae 111<br />

Cainoblattinopsis 108*, III<br />

Calamoceratidae 299, 300<br />

Calamoceratinae 300<br />

Calophinidae 198<br />

Calaphis 196*<br />

Calcitrago 434*<br />

Calcitronidae 436<br />

Caleremaeus 454*, 456<br />

Callaphidinae 198<br />

Callicurgus 358*<br />

Callietheira 288, 289*<br />

Callimokaltania 75*, 76<br />

Callimokaltaniidae 74, 76<br />

Callimomidae 353<br />

Callipteriden 198<br />

Callipterinae 198<br />

Calocoris 218*<br />

Calohymen 67*, 68<br />

Caloneura 143*, 144<br />

Caloneuridae 142, 144<br />

Caloneurodea 37*, 118, 142<br />

Culotermitidae 112, 113<br />

Calvertiellidae 52<br />

Calvisia 160*<br />

Camarostomata 383<br />

Camisia 452*. 454<br />

Camisiidae 452*, 454<br />

Camisiidea 454<br />

Campodea 47*<br />

Campodeidae 47<br />

Campodeidea 47<br />

Camponotus 358<br />

Campterophlebia 83<br />

Camptoneurites 120*, /22<br />

Camptoneuritidae 119, /2/<br />

Campylocephalus 408, 416, 4/9<br />

Campyloptera 71*<br />

Campylopteridae 72<br />

Campylopteridea 64, 71, 72<br />

Canadaphis 199<br />

Canestrinidea 45/<br />

Cantharididae 257*. 258<br />

Cantharis 257*<br />

Canthyloscelididae 330<br />

Capniidae 134<br />

Caponiidae 493<br />

Caponiidea 498<br />

Capsidae 217<br />

Captrix 507, 511*<br />

Carabidae 254*<br />

Carabodes 457*<br />

Carabodidae 456, 457*<br />

Carabodidea 454 , 456<br />

Carabodites 458<br />

Carabopteron 254*<br />

Caraemmarginatae 454<br />

Carbonoptera 65*<br />

Carbonopteridae 64, 66<br />

Carcinosoma 407, 413, 416*<br />

Carcinosomatidae 407, 4/2, 416*<br />

542<br />

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Cardioblatta 97<br />

Carinicupes 247<br />

Carnettiidae 428<br />

Carsidarinae 206<br />

Cathalus 212<br />

Catocha 333*<br />

Cafops 34*<br />

Caulogestra 383<br />

Cebenniblatta 94<br />

Cecidomyiidae 332, 333<br />

Cecidomyiidea 319, 332, 333*<br />

Cecidomyiites 333<br />

Celyphidae 342<br />

Cenattus 515*<br />

Centromachidae 427*. 425, 429*<br />

Centromachus 428, 429*<br />

Centrotinae 177, 178<br />

Cepheus 457*. 458<br />

Cephidae 348<br />

Cephidea 346, 348<br />

Cephas 347*<br />

Cerambycidae 265, 266*<br />

Cerambyx 266*<br />

Ceratinoptera 105*<br />

Ceratocombidae 217<br />

Ceratocombus 216*<br />

Ceratopogonidae 316, 318<br />

Ceratopogoninae 318<br />

Ceratoppia 454*. 456<br />

Ceratoteleia 353*<br />

Ceratozetidae 458*. 460<br />

Cercopida 181<br />

Cercopidae 179, 181<br />

Cercopidea 173, 179, 181, 189<br />

Cercopidium 181<br />

Cercopinus 225<br />

Cercopis 181<br />

Cercoprisca 225<br />

Ceresopseinae /77<br />

Ceresopsis 178*<br />

Ceroplatidae 324<br />

Ceroplatina 324<br />

Cervaphidinae 197<br />

Ceton/a 256*<br />

Ceutorhynchus 267*<br />

Chacharejocaris 393<br />

Chaetidae 428<br />

Chaetoneurophora 340*<br />

Chaetopsocidium 227<br />

Chaitophorinae 197<br />

Chalcididae 352, 353<br />

Chalcididea 349, 352*<br />

Chalepomylacris 97<br />

Chamobates 458*, 460<br />

Chaoboridae 3/4<br />

Chaoborus 314<br />

Charipidae 352<br />

Charontidae 437<br />

Ghasmataspida 394<br />

Chamataspidida 384*, 385, 389,<br />

394*<br />

Chasmataspididae 389, 395<br />

Chasmataspis 389, 394*<br />

Chaulioditidae 124<br />

Cheiridiidae 442<br />

Cheiridiidea 442<br />

Cheiridium 440*, 442<br />

Cheiromachus 483<br />

Chelicerata 377, 375, 380, 381,<br />

382, 383, 384*, 385, 389,402, 403,<br />

423, 433, 447, 450, 474, 482,<br />

490, 492, 525, 526

Chelicerophora 375, 377, 385, 526<br />

Cheliferidae 442<br />

•Cheliferidiea 442<br />

Cheliferinea 384*, 385, 441, 442<br />

Cheliphlebiidae 117<br />

•Chelonethida 439<br />

Chelonethi 439<br />

Chelopteridae 119<br />

Cheloninae 349<br />

Chermesidae 197<br />

•Chernetidae 442<br />

Cheyletidae 448, 461, 467*, 468<br />

•Cheyletidea 462, 467<br />

Cheyletus 467*. 468<br />

Chiloblattuta 103<br />

Chilognatha 22, 23<br />

•Chilopoda 25, 26*<br />

•Chironomidee 43, 316, 317<br />

Chironomidea 310, 316, 317*<br />

Chironomides 3i7, 318<br />

•Chironominae 318<br />

Chironomus 317*<br />

•Chiromyzidae 334<br />

•Chloroperlidae 134<br />

Chloropidae 342, 343<br />

•Chloropinae 343<br />

•Chorisoneurini 107<br />

•Choristidae 286<br />

Choristopanorpa 291<br />

Choristopsyche 293*, 294<br />

•Choristopsychidae 292, 293<br />

Chresmodidae 160<br />

Chresmododea 159, 160<br />

•Chrysididae 354 , 355<br />

Chrysidides 355<br />

Chrysoleonites 273*. 274<br />

Chrysomela 266*<br />

Oirysomelidae 265 , 266*<br />

•Chrysopidae 273, 277, 282<br />

Chrysotoxum 341*<br />

Chthoniidae 441<br />

Chthoniinea 384, 385, 439, 441<br />

Chthonius 440*, 441<br />

•Cicadellidae 173, 176*, 356<br />

Cicadellidea 173<br />

Cicadellomorpha 163, 172, 181,<br />

190<br />

Cicadellium 177, 189<br />

Cicadellopsis 199, 200*<br />

•Cicadidae 167, 171, 172*<br />

Cicadidea 164, 167<br />

Cicadinae 172<br />

Cicadocoridae 208, 210, 224<br />

Cicadocoris 224*, 225<br />

•Cicadomorpha 163, 164, 172<br />

Cicadomorpha 164*<br />

Cicadomorphidae 164<br />

Cicadoprosbole 170, 171*<br />

Cicadopsis 205*, 206<br />

Cicadopsylla 205*, 206<br />

Cicadopsyllidae 204, 205*<br />

Cicadopsyllidea 203, 204<br />

Cicadopsyllidium 199, 200*<br />

Cicadopsyllinae 204<br />

Cicindelidae 254<br />

Cimbicidae 348<br />

•Cimbicinae 348<br />

Cimex 226<br />

Cimicidae 226<br />

Cimicidium 226<br />

Cinara 196*<br />

•Cinarinae 197<br />

Circummarginatae 454<br />

Ciriacreminae 206<br />

Cisidae 260*<br />

Cixiella 185*, 186<br />

Cixiidae 181, 184, 185*<br />

Cixioideae 184<br />

Cixiites 184<br />

Cixioides 186<br />

Cladochorista 296<br />

Cladochoristella 296<br />

Cladochoristidae 295<br />

Clastoptera 180*<br />

Clastopteridae 179, 180* 181<br />

Clavopsyllidium 200<br />

Cleptomartus 491*. 492<br />

Cleridae 258, 259*<br />

Climaconeuridae 124<br />

Climaconeura 128<br />

Climaconeurites 125*, 126<br />

Cloeon 62<br />

Clostes 496*. 498<br />

Clothodidae 139, 140<br />

Clubiona 507, 511*<br />

Clubionidae 507, 511*<br />

Clubionidea 499, 507<br />

Clubioniformia 507<br />

Clupeidae 41<br />

Clya 501, 503<br />

Clythia 341*. 501, 503*<br />

Clythiidae 341<br />

Cnemidolestes 123 *<br />

Cnemidolestidae 122, 124<br />

Cobaloblatta 94<br />

Coccidae 201, 202<br />

Coccidea 199, 200, 201*, 203<br />

Coccodes 202, 203<br />

Coccidomorpha 190, 199<br />

Coccinella 263*<br />

Coccinellidae 262, 263*<br />

Coccus 201*<br />

Cockerelliellidae 50<br />

Coenagrionidae 79, 80<br />

Coenagrionidea 76, 79*<br />

Coenomyidae 336, 337<br />

Coleoptera 42, 43, 240*, 241<br />

Coleopteroidea 36*, 239, 240*, 268<br />

Coleopteropsis 225*, 226<br />

Coleorrhyncha 163, 207, 208<br />

Coleoscyta 204, 205*<br />

Coleoscytidae 204, 205*<br />

Collacteus 507, 512*<br />

Collembola 37*, 45<br />

Colobognatha 23, 24<br />

Colydiidae 261*, 262<br />

Compsoblattula 94<br />

Compsoneura 53<br />

Compsoscorpius 427*, 431*<br />

Compylopterodea 64<br />

Conchostraca 41<br />

Confusio 144<br />

Coniopterygidae 269, 272, 282<br />

Coniopterygidea 273, 282*<br />

Conocephalella 156<br />

Conocephalinae 155<br />

Conopariae 342<br />

Conopidae 342<br />

Conopidea 341, 342<br />

Conostygmus 353*<br />

Constrictitermes 112*<br />

Copeognatha 226<br />

Copidopus 211<br />

Coptoclava 254*, 255<br />

543<br />

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Coptoclavidae 254, 255<br />

Coraloblatta 94<br />

Corduliidae 85<br />

Cordyluridae 343, 344<br />

Coreidae 221*. 222<br />

Coreides 222<br />

Corita 226<br />

Corlxa 213*<br />

Corixida 213<br />

Corixidae 213*, 225<br />

Corrodentia 226<br />

Corydalida 270<br />

Corydalidae 269, 270<br />

Corydaloides 65*<br />

Corydaloididae 64, 66<br />

Corydia 108*<br />

Corydiens 110<br />

Corydiidae 89, 108*. 110<br />

Corydiinae 108*. 110<br />

Corydiini 111<br />

Corydites 110<br />

Corynecoris 212<br />

Coryphomartidae 490<br />

Coryphomartus 491*, 492<br />

Coseliidae 122<br />

Cosmecidae 483<br />

Cossida 305<br />

Cossidae 304, 305<br />

Cramptonomyiidae 331<br />

Craspedops 389, 391*, 393<br />

Craspedosomatidae 24<br />

Creagroptera 282<br />

Cretavidae 356<br />

• Cretavidea 349, 356, 357*<br />

Cretavus 356, 357*<br />

Cribellatae 385, 498, 515<br />

Crockoblattina 94<br />

Cronicus 61*<br />

Crosbella 183<br />

Cryptinae 352<br />

Cryptocercinae 87*<br />

Cryptocercus 87*<br />

Cryptocerphus 353*<br />

Cryptocupes 248, 249*<br />

Cryptognathidae 464*, 465<br />

Cryptognathus 464*<br />

Cryptomartus 465, 492*<br />

Cryptopidae 25, 26<br />

Cryptostemma 216*<br />

Cryptostemmatidae 217<br />

Cryptoveniidae 52<br />

Ctenizidae 497<br />

Ctenidea 507<br />

Ctenoblattina 103<br />

Ctenopterus 404, 408, 416, 418, 419*<br />

Cucujidae 261*, 262<br />

Cucullifera 383<br />

Culex 315*<br />

Culicidae 314<br />

Culicidea 310, 314, 315*<br />

Culicides 314<br />

Cultriribula 457*. 459<br />

Cunaxidae 461, 465<br />

Cuneocoridae 211*, 212<br />

Cuneocoris 211*, 212<br />

Cupidae 241, 243, 244*<br />

Curculioides 443*, 444<br />

Curculioididae 444<br />

Curculionidae 267*<br />

Curculiopsis 250<br />

Custodela 503, 507*<br />

Cyathocrinus 528*

Cyamocephalus 389, 397<br />

Cyclocelidinae 66<br />

Cyclophtalmidae 427*, 428 , 429*<br />

D amphipnoidae 134<br />

£> aphanoptera 69, 70*<br />

D aphanopteridae 69<br />

Dltaxineurella 75*<br />

Ditaxineuridae 74, 75<br />

Dithidae 441<br />

Cyclophtalmus 427*, 429*, 431 D aphanipteritidae 69<br />

Dixa 315*<br />

Cyclopites 389, 393<br />

Cycloptera 284*, 285<br />

Cyclopteridae 284, 285<br />

Cyclorista 284*, 285<br />

Cycloristidae 284, 285<br />

Cycloscytina 185*<br />

Cyclotrogulus 487*, 488<br />

Cydnidae 223*, 224<br />

Cydnidea 224<br />

Cydnopsis 223*<br />

Cylindrotornidae 313<br />

Cylindrotomina 313<br />

Cyllonium 167<br />

Cymatophlebiopsis 85<br />

Cymbaetemaeidae 454*, 456<br />

Cymbaeremaeus 454*, 456<br />

Cymenophlebiidae 122<br />

Cymnodamaeus 456<br />

Cynipes 352<br />

Cynipidae 352<br />

Cynipidea 349, 350*, 352<br />

Cyphoderidae 46<br />

Cyphoderris 152<br />

Cyphomylacris 97<br />

Cyphoneura 230<br />

Cyphoneuridae 228<br />

Cyphoneurodes 230<br />

Cyphophthalmi 385, 474, 476,<br />

Cyphophthalmina 482<br />

Cyrtidae 334, 336<br />

Cyrtophyllites 152<br />

Cyrtophyllitinae 150, 152<br />

Cytocupes 244*<br />

Czekardia 134<br />

Damaeidae 452, 454*, 456<br />

Damaeidea 456<br />

Damaeus 454*, 456<br />

Darninae 177<br />

Darniopseinae 17-7<br />

Darniopsis 177, 178*<br />

Darthula 178*<br />

Dascillidae 257*<br />

Dasyleptidae 48<br />

Dasyleptus 48*<br />

Decticinae 155<br />

Deinacridinae 152<br />

Delopsocus 142<br />

Delopteridae 141<br />

Delopterum 141*, 142<br />

Delphacida 187<br />

Delphacidae 181, 187, 188*<br />

Demodex 448<br />

Demodicidae 461, 462<br />

Demopteridae 119<br />

Deracanthinae 155<br />

Deraeocoris 210<br />

Derbidae 181<br />

Dermacentor 473<br />

Dermanyssidae 472<br />

Dermaptera 37*, 88, 113,<br />

241<br />

Dermestidae 257*, 258<br />

Desuttor 507, 511*<br />

482<br />

114,<br />

D aphanopterodea 49, 64, 69<br />

D, aspides 202<br />

D aspididae 201, 202<br />

D astatommites 81<br />

D, atermes 112*<br />

D atilidae 211*, 212<br />

D atillus 211*. 212<br />

D chaspis 212<br />

D chentomum 227*<br />

D chronoblatta 103<br />

D dadoblatta 95<br />

D cranopalpus 483<br />

D cranoptila 282<br />

D ctynidae 515, 516*, 517<br />

D ctynidea 515<br />

D ctyniformia 515<br />

D, ctyoblatta 94<br />

D ctyodiptera 309*<br />

D ctyodipteridae 308<br />

D ctyodipteridea 308<br />

D<br />

ctyodipteromorpha 308*<br />

D<br />

ctyomylacridae 98<br />

D<br />

ctyomylacrinae 98<br />

ctyomylacris 98, 99*<br />

D<br />

ctyoneuridae 50<br />

D<br />

ctyoneuridea 50<br />

D;<br />

ctyophara 188*<br />

D<br />

ctyopha'ridae 181, 187*<br />

D<br />

'ctyopharoides 188<br />

D<br />

ctyoprosbole 167*, 168<br />

D<br />

ctyotipula 312*<br />

D cyrtomidae 47<br />

D echoblattina 102<br />

D, echoblattininae 100*<br />

D lar 273<br />

D larites 282*<br />

D meropterum 226<br />

D moblatta 94<br />

D nopanorpa 291<br />

D nopidae 515<br />

D nopilio 496*, 497<br />

D, nopsocidae 228<br />

D nopsocus 228<br />

D,<br />

D onychopodes 385, 402, 404, 424<br />

425, 426, 427*, 428<br />

D phtheropsis 171<br />

D plocentridae 428<br />

D plarchitipula 312*, 313<br />

D plopoda 22*<br />

D plopolyneura 310*<br />

D plopolyneuridae 310<br />

D plopolyneuromorpha 308*, 310*<br />

D plopteridae 89<br />

D plura 37*, 45, 47*<br />

D pluridae 496*, 497<br />

D ipluroblattina 102<br />

D pneumones 494<br />

D pneumorphae 498<br />

D prionidae 348, 349<br />

D psocoridae 216*, 217<br />

D ptera 43, 240*, 283, 284, 292<br />

307, 308*<br />

D pterodictya 309*<br />

D scoblatta 94<br />

Dixamima 314, 315*<br />

Dixamimidae 314<br />

Dixidae 314<br />

Dixidea 310, 314, 315*<br />

Dixinae 314<br />

Dobbertiniidae 292<br />

Dodecalopoda 525<br />

Dolichoderidae 358,<br />

Dolichoderinae 357, 358)<br />

Dolichoderus 357*, 358<br />

Dolichophonidae 384*, 425, 426*<br />

Dolichophonus 425, 426*<br />

Dolichopodes 339<br />

Dolichopodidae 339<br />

Dolichopteridae 406, 407, 408, 410,<br />

413*<br />

Dolichopterus 407, 411, 413*<br />

Dolichopus 338*<br />

Donacia 266*<br />

Doropteron 52*, 53"<br />

Doryblatta 95<br />

Doryctinae 349<br />

Dorylaidae 341<br />

Dory las 341*<br />

Doteridae 55<br />

Drassidae 507<br />

Drassiformia 507<br />

Drassodidae 507, 511*<br />

Dryssodidea 499, 507<br />

Drepanoblattina 99, 100<br />

Drepanopterus 407,408,416,418,419*-<br />

Drepanopteryx 280<br />

Drepanosiphiden 198<br />

Drepanosiphinae 197, 198<br />

Dromoblatta 94<br />

Drosicha 201*<br />

Drosophilidae 342<br />

Drosophilina 342<br />

Drybrookia 97<br />

Dryinidae 354 , 356<br />

Dunstania 164<br />

Dunstaniidae 164<br />

Dunstaniopsis 164<br />

Durdelstonia 103<br />

Durgades 176*<br />

Dyadozoarium 133*, 134<br />

Dysdera 498*<br />

Dysderidae 498<br />

Dysderidea 498<br />

Dysderiformia 498*<br />

Dysmenes 94<br />

Dysmorphoptila 189<br />

Dysmorphoptilidae 189<br />

Dysmorphoptiloides 189*<br />

Dyspnoi 483<br />

Dyspolyneura 309*, 310<br />

Dyspolyneuridae 310<br />

Dyspolyneuridea 308, 310-<br />

Dytiscidae 254*, 255<br />

Dytiscus 254*<br />

Echinognathus 407, 415<br />

•Echinomyia 344<br />

Echinopsocinae 231<br />

Ecnomidae 295, 298<br />

Ecnominae 298<br />

Deuterophlebiomorpha 308* D scotarbus 480<br />

Ecnomus 296*<br />

Diadocidia 324* D scomylacris 97<br />

Ecribellatae 385, 498<br />

Diadocidiidae 326 D statoblatta 94<br />

Ectinoblattula 103<br />

Diadocidiinae 326 Di taxineura 75*<br />

544<br />

http://jurassic.ru/

Ectobiens 107<br />

Ectobiinae 100, 107, 108*<br />

Ectobiini 107<br />

Ectobius 108*<br />

Eidothea 419<br />

Elaphroblatta 94<br />

Elasmoscelidium 188*<br />

Elateridae 258, 259*<br />

Elaveromartus 489*<br />

Elcana 149*<br />

Elcanidae 146, 149, 150<br />

Elcanopsis 149<br />

Electracanthinus 296*<br />

Electragapetus 296*<br />

Electrentominae 23/<br />

Electrentomum 232*<br />

Electresia 305*<br />

Electrocerum 301*<br />

Electrocherlifer 442<br />

Electrocrania 303*<br />

Electrogenia 63*, 64<br />

Electrostephanus 350*<br />

Electrotrichia 296*<br />

Elipsocidae 234<br />

Elipsocus 235*<br />

Elisama 103<br />

£//ешг 390, 399<br />

Elleriidae 390, 303<br />

Elliptiscarta 174<br />

£/moa 70*<br />

Elmoidae 60<br />

£/uc«s 505, 509*<br />

Elytroneuridae 115<br />

Embaneura 277*, 273<br />

Embiidae 139<br />

Embioptera 37*, 118, /30*<br />

Embolacarus 452*, 456<br />

Embolemidae 354<br />

Embolemoidae 354<br />

Embolemus 355*<br />

Emeraldellidae 391<br />

Emgerrophorus 212<br />

Empheria 232<br />

Empheriidae 232<br />

Empheriinae 232<br />

Emphylopteridae 122<br />

Empides 339<br />

Empididae 330<br />

Empididea 334, 330<br />

£mpis 338*<br />

Encyrtidae 352<br />

Endeostigmata 383, 384*, 447,<br />

449, 450, 463<br />

Endomychidae 262, 263*<br />

Eneopterinae 154<br />

Eneriblatta 94<br />

Engerophorus 212<br />

Engynabis 211<br />

Ensifera /46<br />

Ensphingocoris 212<br />

Entelegynae 498, 400<br />

Entomecoris 212<br />

Entomobryidae 46<br />

Entomobryomorpha 45, 46<br />

Entonyssidae 473<br />

Eottopsis 515<br />

Eoblattidae 117, //3<br />

Eoboletina 323*<br />

Eobumbratrix 507<br />

Eocercopis 225<br />

Eochiliocycla 208<br />

Eocicada 167<br />

Eocimex 210*, 211<br />

Eocimicidae 210*. 2//<br />

Eocryphaeca 505, 510*<br />

Eocteniza 495*, 496<br />

Eoctonus 427*, 428<br />

Eocupes 243, 244*<br />

Eodiplurina 498<br />

Eodipoena 501, 503*<br />

Eofulgoridium 183*, /34<br />

Eogerridiutn 211<br />

Eoglosselytridae 158<br />

Eoglosselytrum 158*<br />

Eogonatium 503, 504*<br />

Eohesperinus 323*<br />

Eohymen 52*<br />

Eohymenidae 52<br />

Eojassus 181<br />

Eolathys 516*, 517<br />

Eolimnobia 312*, 3/3*<br />

Eolimnobiidae 3/3<br />

Eolinus 515*<br />

Eolycosa 496<br />

Eomatachia 516*, 517<br />

Eomeganisoptera 73*<br />

Eomyia 333*, 334<br />

Eomyiidae 334<br />

Eomysmena 501, 503*<br />

Eonabidae 210*<br />

£ona6(s 210*, 211<br />

Eopachyneura 323*<br />

Eopholcus 496*, 407<br />

Eophrynidae 436, 487*<br />

Eophrynus 487, 488<br />

Eophrys 515*<br />

Eoplecia 329*<br />

Eopleciidae 320<br />

Eopodidae<br />

Eopolyneura 311*<br />

Eopolyneuridae 3//<br />

Eopolyneuridea 310, 3//*<br />

Eopopino 503, 507*<br />

Eopsychops 275*, 282*<br />

Eopsichopsis 282<br />

Eopsyllidium 200<br />

Eopteridae 36<br />

Eopterum 38, 36, 87*<br />

Eoptychoptera 314<br />

Eoptychopteridae 3/4<br />

Eoptychopteridea 310, 312*, 3/4<br />

Eoscarterella 179*, 180<br />

Eoscarterellidae /70*<br />

Eoscartoides 180<br />

Eoscorpiidae 428, 420, 431*, 432<br />

Eoscorpionidae 427*<br />

Eoscorpius 429, 431*, 433<br />

Eoscytina 194<br />

Eostaianus 507, 512*<br />

Eostaslna 500, 512*<br />

Eostentatrix 507<br />

Eostratiomyia 335*, 336<br />

Eostratiomyiidae 336<br />

Eotermes 112*<br />

Eotettigarcta 171<br />

Eothanatus 509, 513*<br />

Eotipula 313<br />

Eotrogulidae 482<br />

Eotrogutus 480*. 432<br />

Eoversatrix 507<br />

Epactinophelbia 278*, 270<br />

Epallage 79*<br />

Epallagidae 80, 3/<br />

Epeira 505<br />

545<br />

http://jurassic.ru/<br />

Epeiridae 503<br />

Ephalmator 505, 509*<br />

Ephalmatoridae 501, 505, 509*<br />

Epheboblatta 103<br />

Ephemera 59, 60*. 62<br />

Ephemerella 63*, 64<br />

Ephemerellidae 60, 62, 63*<br />

Ephemeridae 50, 60*<br />

Ephemeridea 57, 50 •<br />

Ephemeropsidae 50<br />

Ephemeropsis 58*, 59, 64<br />

Ephemeroptera 37*, 42, 50 , 55<br />

Ephialtitidae 348<br />

Ephippigerinae 155<br />

Ephoron 59, 60*<br />

Epideigmatidae 117<br />

Epididonotus 297<br />

Epigambria 274<br />

Epilampra 106*<br />

Epilampriens 107<br />

Epilamprinae 100, 106*, /67<br />

Epilamprini /07<br />

Epilestes 77*, 78<br />

Epimastacidae 117<br />

Epimastax 116*, 1/3<br />

Epimorpha 25<br />

Epiophlebia 80*<br />

Epiophlebiidae 81<br />

Epipsocidae 234<br />

Epipsocinae 234<br />

Epipsocus 235*<br />

Epitethe 53<br />

Eporibatula 458*, 460<br />

Erasipteridae 73<br />

Erasipteron 73*<br />

Ereischeria 490<br />

Eremaeidae 456<br />

Eremaeus 454*, 456<br />

Eresidae 5/5, 516*<br />

Eresidea 515<br />

Eresiformia 515<br />

£>esus 515, 516*<br />

Erettopterus 414*, 423<br />

Ereynetidae 465<br />

Erieopterus 407, 4/0<br />

Erigone 503, 504*<br />

Erigonidae 503<br />

Erinna 335*<br />

Erinnidae 336<br />

Eriocraniidae 303<br />

Eriophyes 471<br />

Eriophyidae 380, 469*, 47/<br />

Eriophyiformes 448, 449, 450, 471<br />

Eriosomatinae 197<br />

£rma 272<br />

£ro 503, 504*<br />

Erotesis 301*<br />

Erotylidae 261*, 262<br />

Erunakicupes 240, 250*<br />

Erythraeidae 467*, 463<br />

Erythraeus 467*, 468<br />

Esuritor 505, 510*<br />

Etoblattina 94<br />

Eublattula 103<br />

Eubleptidae 54<br />

Eubleptodea 50, 54*<br />

Eubleptus 54*<br />

Eucaenidae 117<br />

Eucnemis 259*<br />

Eucoenepidae 439<br />

Eucoilidae 352<br />

Eucosminae 306

Eudermaptera 114, 115<br />

Eudiaphanopterodea 69, 70*<br />

Eudiptera 308, 310<br />

Eudoter 56*<br />

Eudoteridae 56*<br />

Euembiaria 139<br />

Eugereon 51*<br />

Eugereonidae 50<br />

Euglyptoblatta 103<br />

Eukoeneniidae 439<br />

Eukuloja 71*. 72<br />

Eulahmaniidea 454<br />

Eulophidae 352<br />

Eumastacidae 156, 157<br />

Eumastacides 157<br />

Eumecoptera 283, 284, 286<br />

Eumegasecoptera 64, 65*<br />

Eumolpites 266<br />

Euonychophora 533<br />

Eumorphoblatta 94<br />

Eupalaeodictyoptera 50, 51*<br />

Eupantopoda 529<br />

Euphasmatodea 159, 160<br />

Euphoberia 22*<br />

Euphoberidae 23<br />

Euphorinae 349<br />

Eupincombea 195, 196<br />

Euplectoptera 56, 57<br />

Eupnoi 483<br />

Eupodidae 461, 465<br />

Eupodidea 465<br />

Euproopidae 386, 387, 390, 398,<br />

399*<br />

Euproopidea 397<br />

Euproops 390, 398, 399*<br />

Eupsocida 226, 234, 235*<br />

Eurachnida 403<br />

' Euraspidium 212<br />

Euremisca 128, 129*<br />

Euremiscidae 124, 128, 129*<br />

Europhthalmus 427*, 431*<br />

Euryblattula 100*, 103<br />

Eurybrachiidae 181<br />

Eurymelidae 173, 176*. 177<br />

Eurymelides 177<br />

Eurymelidium 177<br />

Eurynotis 212<br />

Eurypteracea 411<br />

Eurypterida 378, 380, 381, 382,<br />

384*, 385, 389, 391, 395, 402,<br />

403, 404, 405*, 406*, 407, 408,<br />

412, 420, 422*, 423, 424<br />

Eurypteridae 406, 407, 408, 410,<br />

411*, 412*<br />

Eurypteridea 407, 408, 412, 422<br />

Eurypterina 385, 405, 406, 407,<br />

408<br />

Eurypterodea 383, 424, 425, 427<br />

Eurypterus 392, 405*. 407, 410,<br />

411*, 420, 421<br />

Euryptilodes 116*, 118<br />

Euryptilon 116*. 118<br />

Euryptilonidae 117<br />

Eurysterna 23, 25<br />

Eury torn idae 352<br />

Euscorpiidea 428<br />

Euscorpiones 384*, 425<br />

Eusparassidae 507, 512*<br />

Euspilopteron 152<br />

Eustaloides 505, 509*<br />

Eustheniidae 134<br />

Euthygramma 143*, 144<br />

Euthygrammatidae 142, 144<br />

Euthyneura 144<br />

Euthyrrhapha 108*<br />

Euthyrrhaphinae 108*, 110, 111<br />

Euzetes 460<br />

Evansia 170<br />

Evansicicada 168*, 170<br />

Evania 350*<br />

Evaniades 350 '<br />

Evaniidae 349, 350<br />

Exapinurus 396<br />

Exochoblatta 94<br />

Exochomylacrls 97<br />

Eylaidae 471<br />

Facundia 509, 513*<br />

Fatjanoptera 270*, 271<br />

Fatjanopteridae 270*<br />

Fayoliellidae 117<br />

Feallidae 441<br />

Feallidea 441<br />

Ferganopsyche 293*, 294<br />

Fibla 271*. 272<br />

Fiducia 509, 513*<br />

Figitidae 352<br />

Filiola 509, 513*<br />

Filistatidea 515<br />

Flabellites 94<br />

Flatidae 181,187*<br />

Flatoides 187<br />

Flegia 501, 503<br />

Fletcheriana 167<br />

Forficula 114*<br />

Forficulidae 115<br />

Forficulidea 114, 115<br />

Forf iculina 114<br />

Foriria 65*<br />

Foririidae 64, 66<br />

Formicidae 358<br />

Formicidea 349, 357*<br />

Formicinae 357, 358<br />

Fouqueidae 50<br />

Frenata 304, 305*. 307*<br />

Friciella 94<br />

Frirenia 334<br />

Fulgorellae 183<br />

Fulgoridae 181, 183*, 356<br />

Fulgoridea 189<br />

Fulgoridiella 183*, 184<br />

Fulgoridiidae 181, 183*, 184<br />

Fulgoridiinae 184<br />

Fulgoridium 183*. 184<br />

Fulgoridulum 184<br />

Fulgorinae 188<br />

Fulgoromorpha 163, 181, 189<br />

Fulgoropsis 183*. 184<br />

Fungi vorae 324<br />

Fungivoridae 324<br />

Fungivorinae 324<br />

Fungivorites 323*<br />

Fungivoritidae 323<br />

Fungivoridea 319, 320*, 321*, 323*.<br />

324*, 326*.<br />

Gaesomyrmex 357*<br />

Gagrellinae 480<br />

Galerucites 226<br />

Galumna 459*, 461<br />

Galumnidae 459*, 460, 461<br />

Gamasidea 472<br />

Gamasides 472<br />

Gamasus 449, 452<br />

546<br />

http://jurassic.ru/<br />

Ganonema 301*<br />

Garnettiidae 428, 431*, 432<br />

Garnettius 427*, 431*. 432<br />

Garypidae 441<br />

Garypidea 441<br />

Garypinus 441<br />

Gary pus 441<br />

Gastrophilidea 341<br />

Gea 505, 509*<br />

Geinitzia 132<br />

Geinitziidae 124, 131*, 132<br />

Gelechiidae 306<br />

Gelechiidea 304, 306<br />

Genaphididae 195, 196*<br />

Genaphis 196*<br />

Genentomum 148<br />

Genopterygidae 142<br />

Geocorisae 208, 213<br />

Geogarypus 441<br />

Geophilellidae 21<br />

Geophilidae 25<br />

Geophilomorpha 25<br />

Geralinuridae 436<br />

Geralinura 436<br />

Geralycosa 495*<br />

Geraphrynus 479, 480<br />

Gerapomoidae 117<br />

Geraridae 122<br />

Geraridea 119, 122, 123*<br />

Gerarus 123*<br />

Geratarbus 479*, 480<br />

Gerdia 501<br />

Germanopriscidae 124<br />

Gerrida 219<br />

Gerridae 219, 220*<br />

Gerris 220*<br />

Gigantostraca 385<br />

Gigantotermes 282<br />

Girardevia 389, 392*, 393<br />

Giripopterygidae 134<br />

Glaphyrocoris 117<br />

Glaphyrophlebia 117<br />

Glaucodes 420, 421, 422<br />

Glendotricha _ 305*<br />

Glesseumeyrickia 305*<br />

Glomeridae 24<br />

Glomeris 22*<br />

Glosselytridae 157, 158, 159*<br />

Glosselytrodea 37*. 42, 145, 157Г<br />

Glosselytron 159*<br />

Glossina 343*, 344<br />

Glossinidae 344<br />

Glossosomatidae 295, 297<br />

Glycyphagidae 451<br />

Glypharthrus 389, 393<br />

Gnaphosidae 507<br />

Goera 301*<br />

Goerinae 302<br />

Gohieria 451<br />

Gojinae 234<br />

Gomphidae 83, 85<br />

Gonatocerus 352*<br />

Gondwanoblatta 94<br />

Gongyloblatta 94<br />

Goniomylacris 97<br />

Goniotarbus 479*<br />

Gonyleptes 480*, 483<br />

Gonyleptidae 480, 483<br />

Gorgopsina 509, 515*<br />

Gorgopsis 509<br />

Gradidorsum 454*, 456<br />

Graeophonus 435*, 437

Grammopsychops 277*, 278<br />

Grammositum 254<br />

Grandidorsum 454*<br />

Greenideinae 197<br />

Greenideini 197<br />

Grossopterus 407, 409*, 410<br />

Gryllacrididae 150, 151*, /52,153*<br />

Gryllacridinae 152<br />

Grylliae 154<br />

Gryllidae 150, 153*, 154<br />

Gryllidea 146, 150<br />

Gryllinae 154<br />

Gryllotalpa 153*<br />

Gryllotalpidae 150, 154<br />

Grypoblattina 94<br />

Grypoblattites 91, 92*<br />

Gurianovella 133*, 134<br />

Gylippus 443*<br />

Gymnodamaeus 454*, 456<br />

Gyrinidae 255<br />

Gyroblatta 94<br />

Hadentomidae 122<br />

Hadrachne 480<br />

Hadrocoridae 211*<br />

Hadrocoris 211*<br />

Haeberleoblatta 103<br />

Haemogamasidae 472<br />

Hagla 150, 151*Д<br />

Haglidae 150, 151*, 152<br />

Haglinae 150<br />

Hahniidae 505<br />

Halacarae 381<br />

Halacaridea 462<br />

Halarachnidae 473<br />

Haliplidae 254*, 255<br />

Haliplus 254*, 255<br />

Halometra 439<br />

Hapalopteridae 122<br />

Haplobittacus 292<br />

Haplogynae 498<br />

Haplopterum 131*, 132<br />

Haplozetidae 459*. 460<br />

Haptopoda 383, 384*, 385, 475,<br />

476, 477*, 479, 482<br />

Haptopodida 476<br />

Harpactes 498, 499*<br />

Harpirhynchidae 467<br />

Helea 317*<br />

Heleidae 43, 318<br />

Helicopsyche 301*<br />

Helicopsychidae 302<br />

Helicopsychinae 302<br />

Helicoptera 185*<br />

Heliothrips 237*<br />

Helodidae 257*<br />

Helophorus 256*<br />

Helops 264*<br />

Heloridae 353, 354<br />

Hemerobiidae 277, 280<br />

Hemerobiidea 273, 277, 281*. 282*<br />

Hemerobiini 280<br />

Hemerobius 281*<br />

Hemiaspididae 395<br />

Hemiaspis 395<br />

Hemikreischeria 487*, 488<br />

Hemimylacrella 29, 91, 92*<br />

Hemimylacridae 96, 97<br />

Hemimylacrides 96, 97<br />

Hemimylacrinae 96*. 97<br />

Hemimylacris 91, 96*, 97<br />

Hemiodoecus 207*<br />

42 О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие<br />

Hemiphrynus 487*, 488<br />

Hemiphlebia 79*<br />

Hemiphlebiidae 78<br />

Hemiphlebiidea 76, 78, 79*<br />

Hemipteron 226<br />

Hemizygopteron 75<br />

Henicinae 152<br />

Henicocephalida 219<br />

Henicocephalidae 219 , 220*<br />

Henicocephalus 220*<br />

Heolidae 52<br />

Heptagenia 63*<br />

Heptageniidae 63*, 64<br />

Heptageniidea 57, 64<br />

Hermanniella 457*, 459<br />

Hermanniellidae 457*, 459<br />

Hermanniellidea 454, 459<br />

Hermanniidae 456<br />

Hermanniidea 454<br />

Hermione 333*<br />

Hersilia 500*, 501<br />

Hersiliidae 500*, 501<br />

Hersiliidea 499, 501<br />

Hersilioformia 501<br />

Hesperinidae 329, 330<br />

Hesperinus 329*<br />

Hesperoblatta 94<br />

Heterologus 123*<br />

Heterolophus 440*, 441<br />

Heteronotinae 177<br />

Heteropeza 334<br />

Heteropezidae 332, 333<br />

Heteropezinae 333<br />

Heterophlebia 80*<br />

Heterophlebiidae 81<br />

Heterophlebiidea 80*, 81<br />

Heteropoda 507<br />

Heteroptera 42, 161, 162*, 189,<br />

208, 223, 224, 225, 241<br />

Heteroscytina 225<br />

Heterotarbidae 479<br />

Heterotarbus 479*<br />

Heterothemis 81<br />

Heterothripidae 237, 238<br />

Hetrodinae 155<br />

Hexagenia 59<br />

Hexagenites 58*, 59<br />

Hexagenitidae 58*, 59<br />

Hexagenitinae 59<br />

Hexisopodidae 446<br />

Hibbertopteridae 408, 416, 419<br />

Hibbertopterus 408, 419*. 420<br />

Hilarimorpha 338*, 339<br />

Hilarimorphidae 339<br />

Hippoboscidae 344<br />

Hippoboscidea 343*, 344<br />

Hister 256*<br />

Histeridae 256*<br />

Hodotermitidae 112, 113<br />

Hodotermitini 113<br />

Holactinochitinosi 384*, 385, 389,<br />

402, 403, 404, 433<br />

Holcorpa 291<br />

Holepyris 355*<br />

Holocompsa 108*<br />

Holocompsinae 108*, 110, ///<br />

Holocompsini 111<br />

Holometabola 35, 38, 85, 86, 239,<br />

240*. 241<br />

Holometopa 342<br />

Holopeltidia 384*. 385, 436<br />

Holotergidae 444<br />

547<br />

http://jurassic.ru/<br />

Holothyroidea 384*. 385, 448<br />

449, 450, 472<br />

Homaloneura 53<br />

Homaloneurina 53<br />

Homaloneurites 53<br />

Homalonychidae 499<br />

Homalophlebiidae 122<br />

Homaloptila 144<br />

Homaloscytina 174<br />

Homoeodictyidae 122<br />

Homoeodictyon 122, 123*<br />

Homoioptera 51*<br />

Homoiopteridae 52<br />

Homoiopteridea 50, 52<br />

Homoptera 42, 161, 162*, 163*, 208<br />

Homopterites 189<br />

Homopteron 190<br />

Homopterulum 177<br />

Homothetidae 53<br />

Hongaya 103<br />

Hoploderma 459*, 461<br />

Hughmilleria 405*, 407, 408, 409*<br />

Hughmilleriidae 391, 406, 407,<br />

408, 409*, 411, 423<br />

Humbertiella 152<br />

Hyalomma 473<br />

Hydracarina 462<br />

Hydrachnellae 381, 461, 462<br />

Hydrocorisae 209<br />

Hydrometra 220*, 226<br />

Hydrometraedes 220<br />

Hydrometridae 220*<br />

Hydrophilidae 256*<br />

Hydropsychidae 295, 298<br />

Hydropsychinae 299<br />

Hydroptilidae 295, 297<br />

Hylaeoneura 171<br />

Hylesinus 267*<br />

Hylicella 180*, 181<br />

Hylicellites 181<br />

Hylicellidae 180*, 181<br />

Hylicidae 173<br />

Hymenoptera 43, 240*, 284, 292,<br />

345<br />

Hymenopteroidea 36*, 239, 345<br />

Hypercompsa 108*<br />

Hyperpolyneura 309*<br />

Hyperpolyneuridae 309<br />

Hyperpolyneuroidea 308, 309<br />

Hyphinoinae 177<br />

Hypoaspis 470*, 473<br />

Hypoboscidae 344<br />

Hypoboscidea 341, 343, 344<br />

Hypochilidae 494, 498, 515<br />

Hypochilomorphae 494<br />

Hypocimex 212*. 213<br />

Hypochthoniidae 452*, 454<br />

Hypochthoniidea 454<br />

Hypocimicidae 212*, 213<br />

Hypodemata 385, 497, 498<br />

Hypodermatidae 343, 344<br />

Hypodermina 344<br />

Hypogastrura 46*<br />

Hypogastruridae 46<br />

Hypogastrurinae 46<br />

Hyponomeutidae 306<br />

Hypoperla 138*<br />

Hypoperlidae 138*<br />

Hypoperlopsis 138*<br />

Hypsoprorini 178<br />

Hystrichopsyllidae 345

Ibaliidae 352<br />

Ichneumonidae 349, 352<br />

Ichneumonidea 349, 350*<br />

Ichneutnonides 350<br />

Ichneumoninae 352<br />

Ichthyocupes 247*<br />

I delta 121<br />

Ideliella 54*, 55<br />

Ideliidae 119<br />

Ideliidea 119, 120*<br />

Ideliopsis 128<br />

Ideloblatta 98, 99*<br />

Idelopanorpa 288<br />

Idelopanorpidae 286<br />

Idelopsocus 228<br />

Ideoroncidae 441<br />

Idiomylacridae 97<br />

Idiomylacrinae 96*, 97<br />

Idiomylacris 96*, 97<br />

Ignarobtatta 92*. 93<br />

Ijacossus 164*, 165<br />

Ijapsyche 293<br />

Iliocoris 215*<br />

Immarginatae 454<br />

Impulsor 503, 507*<br />

Inceptor 509, 513*<br />

Inceptoridae 507, 509, 513*<br />

Incurvariidae 304<br />

Indusia 299*<br />

Ineptiae 282<br />

Ingruidae /75<br />

Ingruinae 173, /75*<br />

Ingruo 175*<br />

Inocellia 272<br />

Inocelliidae 270, 271*, 272<br />

Insecta 29, 379<br />

Insecutor 507, 510*<br />

Insecutoridae 505, 510*<br />

Integripalpia 295, 299, 301*<br />

Intejocaris 393<br />

Ipidae 267*<br />

Ipsvicia 189, 190, 191*<br />

Ipsviciella 190, 191*<br />

Ipsviciidae /*9, 190, 191*, 224<br />

Ipsviciopsis 189, 190<br />

Ischanoblattina 99, 100*<br />

Ischnocoris 212<br />

Ischnoneuridae 122<br />

Ischnopterini 104, /06<br />

Ischnoptilinae 66<br />

Ischnoptilus 65*<br />

Ischyopteron 226<br />

Ischyropsalidae 480, 4*3<br />

Isfaroptera 153*, /54<br />

Isfaropteridae 150, /53*<br />

Isobuthidae 427*, 428, 429*<br />

Isobuthus 424, 427*, 428, 429*<br />

Isoiulus 23<br />

Isonychia 61*<br />

Isonychiidae 61*<br />

Isonychiinae 6/<br />

Isophlebiidae 81, S3<br />

Isoptera 37*, 88, 112*<br />

Isotomidae 46<br />

Isotominae 46<br />

Issidae 181, 188*<br />

Issites 188<br />

Itonididae 333<br />

Itonididea 332<br />

Ivahymen 67*. 68<br />

Ivaia 174, 175*<br />

/vaiinae 173,174, 175*<br />

Ivascytina 192<br />

/jrades 449, 470*, 473<br />

Ixodidae 448, 473<br />

Ixodides 384*, 385, 449, 472, 473,<br />

485<br />

Japygidae 47<br />

Jassidea 189<br />

Jassidomorpha 172<br />

Jassites 177<br />

Jasvia 147*<br />

Jongmansiidae 52<br />

Jugata 303*<br />

Julidae 24<br />

Juliformia 23, 24<br />

Julus 22*<br />

Jurassobatea 152<br />

Jurina 158*, 159<br />

Jurinidae 157, 158*<br />

Kafar 94<br />

Kalligramma 278*<br />

Kalligrammatidae 277, 278*<br />

KalUgrammula 278*, 279<br />

Kalophryganeidae 299<br />

Kaltana 285<br />

Kaltanaphis 195*<br />

Kaltanella 125*, 126<br />

Kaltanelmoa 70*, 71<br />

Kaltanelmoidae 69, 71<br />

Kaltanetta 168*<br />

Kaltanicupes 243, 244*<br />

Kaltanidae 284*<br />

Kaltanochorista 285*, 286<br />

Kaltanocicada 163*, 172<br />

Kaltanocoleidae 241, 242*, 243<br />

Kaltanocoleus 242*. 243<br />

Kaltanoneura 75*<br />

Kaltanoneuridae 74, 75<br />

Kaltanopibrocha 182*, 183<br />

Kaltanopsis 168*, /70<br />

Kaltanopterodes 133*, /34<br />

Kaltanoseyta 204, 205*<br />

Kaltanoscytina 192*<br />

Kaltanospes 175*<br />

Kaltanympha 127*, /25<br />

Kamaites 148<br />

/Cam/a 145*, /46<br />

Kamopanorpa 287 , 288*<br />

Kansasia 55<br />

Karabasia 176*, /77<br />

Karajassus 176*, /77<br />

Karakanocoleodes 251*, 252<br />

Karakanocoleus 251*, 252<br />

Karanabis 218*, 2/9<br />

Karataviella 225*<br />

Karatavocoris 221*, 222<br />

Karatawia 82*, £3<br />

Karatawiidae 81, 52<br />

Kargarella 131*<br />

Kargalodes 131*, /32<br />

Kargalotypus 73*, 74<br />

Kazanella 127*, /2*<br />

Kebaona 100<br />

Kempyninae 276<br />

Kennedya 76, 77*<br />

Kennedyidae 76<br />

Kennedyidea 76, 77*<br />

Khosara 127*. /25<br />

Khosaridae 124, 132<br />

Khosarophlebia 131*, /32<br />

Kiaeria 390, 399<br />

548<br />

http://jurassic.ru/<br />

Kiaeriidae 390, 399<br />

/Cr'/ato? 439<br />

Kinklidoblatta 94<br />

Kinklidoptera 94<br />

Kirgisella 276<br />

Kirgisellodes 275*, 276<br />

Kirkorella 134<br />

Kisylblatta 90*, 9/<br />

К«г//ш /7/*<br />

Klebsiellidae 122<br />

Kleidocerys 226<br />

Kliveriidae 117, 122<br />

Koenenia 438*<br />

Koeneniidae 433<br />

Kokandoblattina 100*. /02<br />

Kondomoprosbole 168*. /70<br />

Kolvidelia 120*, /2/<br />

Kortshakolia 119, 120*<br />

Kreischeria 487*, 4*6', 490<br />

Kulmbachiellon 103<br />

/Cuto/a 71*, 72<br />

Kulojidae 72<br />

Kunguroblattina 89, 90*<br />

Kustarachnida 382, 384*, 385,<br />

403, 433,437,438*<br />

Kustarachnidae 437<br />

Kustarachnae 383<br />

Kustarachne 437, 438*<br />

Kustaria 495*<br />

Labellata 383<br />

Labidognatha 494, 498<br />

Labidura 114*<br />

Labiduridae 114<br />

Labiidae //5<br />

Labiinae //5<br />

Lachniden 197<br />

Lachninae /97<br />

Laelaptidae 472, 473<br />

Lagria 264*<br />

Lagriidae 264*, 265<br />

Laiyangia 110<br />

Lampropteridae 176<br />

Lamproptilidae 53<br />

Lampyridae 257*, 258<br />

Lampyris 257*<br />

Laniatores 482,4*3<br />

Lanthus 84*<br />

Laotoma 46*<br />

Largus 221*<br />

Lasius 357*, 358<br />

Laspeyresiellidae 122<br />

Laternariidae 183<br />

Lathridiidae 261*, 262<br />

Lathridius 261*<br />

Latigastra 383, 450<br />

Latindiinae 110<br />

Latisterna 383<br />

Lauxaniidae 342<br />

Leanchoillidae 39]<br />

Lecaniidae 202<br />

Lecanococcidae 202<br />

Lecorium 130<br />

Leda 515<br />

Lemmatophora 129*, 130<br />

Lemmatophoridae 124, 129*, /30<br />

Lepidillidae 231<br />

Lepidocyrtus 46*<br />

Lepidoderma 407, 409*, 410<br />

Lepidopsocidae 228, 231<br />

Lepidopsocinae 231

Lepidoptera 240*, 283, 284, 295<br />

299, 303<br />

Lepidostomatinae 302<br />

Lepisma 47*<br />

Lepismatidae 48<br />

Lepismida 48<br />

Leptoblattina 94<br />

Leptoceridae 299, 300<br />

Leptocerinae 300<br />

Leptonetidae 498<br />

Leptoneurula 142<br />

Leptophlebia 62, 63*<br />

Leptophlebiidae 60, 62, 63*<br />

Leptotarbus 479*, 480<br />

Leptus 467*, 468<br />

Lesna 272<br />

Lestes 79<br />

Lestidae 79<br />

Lestodryinus 355*<br />

Lestremiidae 332<br />

Lestremiinae 332<br />

Letopaloptera 270*, 271<br />

Letopalopteridae 270*, 271<br />

Leuctra 135*<br />

Leuctridae 134, 137<br />

Lezininae 152<br />

Liacaridae 456, 457*, 458<br />

Liadoblattina 103<br />

Liadopsylla 206, 207*<br />

Liadopsyllinae 206 •<br />

Liadotaulius 297<br />

Liadothemis 81<br />

Liadotypidae 73, 74<br />

Liadotypus 73*. 74<br />

Liadoxyela 346, 347*<br />

Liassocoris 211<br />

Liassochorista 289*<br />

Liassocicada 172<br />

Liassogomphidae 83<br />

Liassogomphus 84*<br />

Liassophila 297<br />

Liassophlebia 80*<br />

Liassophlebiidae 81<br />

Liassophyllutn 152<br />

Liassopsychops 278<br />

Liassoscorpionides 433<br />

Liassothripidae 237<br />

Liassothrips 237*, 233<br />

Liassotipula 313<br />

Libellulidae 85<br />

Libellulidea 83, 3,5<br />

Liburnia 188*<br />

Lichnophthalmus 427*, 431<br />

Lichnoscorpius 431<br />

Lichneremaeus 454*, 456<br />

Liebstadia 458*. 460<br />

Limacodites 167<br />

Limacomorpha 23<br />

Limmatoblatta 98, 99*<br />

Limnocharae 469<br />

Limnochares 471<br />

Limnocharidae 471<br />

Limnocharidea 469<br />

Limnogeton 213*<br />

Limnophilidae 299*, 300<br />

Limnophllldea 300<br />

Limnophllinae 300<br />

Limnophilus 299*<br />

Limoniidae 3/3<br />

Limulacea 400<br />

Limulida 384», 385, 386, 389,<br />

390, 395, 397, 398*<br />

Limulidae 390, 397, 400*, 401<br />

Limulidea 390, 397, 400<br />

Limulitella 390, 400*, 401<br />

Limuloides 389, 394, 395, 396*<br />

Limuloididae 387, 389, 394, 395,396*<br />

Limulus 387*, 388*, 419<br />

Lingulidae 41<br />

Linyphia 503, 507*<br />

Linyphiidae 501, 503, 507*<br />

Liobunum 483<br />

Liojassus 181<br />

Liomesaspididea 390, 397, 309,400*<br />

Liomesaspididae 386, 390, 397,<br />

399, 400*<br />

Liomesaspis 390, 400*<br />

Liomopterella 125*, /26<br />

Liomopteridae /24, 125*. 127*, 128<br />

Liomopteridea 119, /24, 125*,<br />

127*, 129*, 131*<br />

Liomopterites 126, 127*<br />

Liomopterum 127*, 128<br />

Liparoblatta 94<br />

Liphistiidae 494<br />

Liphistiomorphae 384*, 385, 437,<br />

474, 493, 494, 497<br />

Liposcelidae 228, 23/<br />

Liposcelis 229*<br />

Liriopeidae 3/3<br />

Liroaspididea 472<br />

Li'sca 130<br />

Lissomartus 489*<br />

Listrophoridea 451<br />

Lithentomidae 231<br />

Lithentomum 232, 236<br />

Lithobiidae 26, 27<br />

Lithobiomorpha 26<br />

Lithobius 26*<br />

Lithoblatta 103<br />

Lithomanteidae 50<br />

Lithomanteidea 50<br />

Lithomylacris 97<br />

Lithoneura 51*<br />

Lithopanorpa 289*<br />

Lithopsocidae 231<br />

Lithopsocidium 228<br />

Lithopsocus 232<br />

Lithoscytina 194<br />

Lithosmylus 275*<br />

Lithostoma 260*, 26/<br />

Lithymnetes 155<br />

Ш<strong>с</strong>еп 503, 507*<br />

Liviinae 206<br />

Livetiblatta 94<br />

Lobidognatha 498<br />

Lobosterni 424<br />

locm 180*<br />

Locustopseidae /55, 156*<br />

Locustopseidea /55<br />

Locustopsis 156*<br />

Lohmanniidae 454<br />

Lonchopteridae 340<br />

Lonchopterinae 340<br />

Lophiocypha 230<br />

Lophioneura 230<br />

Lophioneuridae 223<br />

Lophioneurodes 229*, 230<br />

Lophopida 184<br />

Lophopidae 181, /34<br />

Lordalycidae 462<br />

Lorryia 464*<br />

Loxoblatta 94, 97<br />

Loxophlebia 56<br />

549<br />

http://jurassic.ru/<br />

Loxophleps 282<br />

Lucanidae 257*<br />

Lucoppia 454*, 456<br />

Ludibrum 187*<br />

Lusiellidae 50<br />

Lusitanomylacris 97<br />

Lycidae 253, 259*<br />

Lycocercidae 50<br />

Lycoperdina 263*<br />

Lycoria 324*<br />

Lycoriidae 324<br />

Lycoriomima 324*, 326<br />

Lycoriomimella 326*, 323<br />

Lycoriomimodes 326*<br />

Lycorioplecia 324*, 326<br />

Lycos idae 505<br />

Lycosidea 499, 505<br />

Lycosiformia 505<br />

Lycostomus 259*<br />

Lyctidae 260*<br />

£ttc/us 260*<br />

Li/da 347*<br />

Lygaeidae 221*, 222<br />

Lugaeides 222<br />

Lygaenocoris 223*, 224<br />

Lygeosoma 221*<br />

Lygistorrhina 324*<br />

Lygistorrhinidae 326<br />

Lygobiidae 111<br />

Lygobius 111<br />

Lymexylidae 259*. 260<br />

Lymexylon 259*<br />

Lyonetiidae 306<br />

1д<strong>о</strong>е 296*<br />

Lyssomanidae 509<br />

Lystridae 183<br />

Lytta 264*<br />

Machaerotidae 179<br />

Machilidae 48<br />

Machitis 47*<br />

Macroblattina 103<br />

Macrocera 324*<br />

Macroceratidae 324<br />

Macroceratina 324<br />

Macrochile 312*<br />

Macrofrenata 304, 307*<br />

Macronematinae 295<br />

Macrophlebium 146<br />

Macropteridae 52<br />

Macropterocoris 212<br />

Maguviopsis 178*, /70<br />

Maiocereus 490, 491*, 492<br />

Malachius 259*<br />

Malaconothridae 454<br />

Malacopoda 531, 533, 534*<br />

Malleator 503, 507*<br />

Mallophaga 162*, 226<br />

Malmoblattina 103<br />

Mandibulata 17<br />

Manoblatta 94<br />

Manota 324*<br />

Manotidae 324<br />

Manteodea37*, 88, ///*<br />

Margarodidae 20/<br />

Margarodinae 201<br />

Margaroptilon 184<br />

Margaitea 105*<br />

Marietta 134<br />

Marimerobius 283*, 234<br />

Maripsocidae 191<br />

Maripsocus 192*, /04

Marsupipterinae 408, 418<br />

Marsupipterus 408, 418, 419, 420*<br />

Martynopsocidae 227, 228<br />

Martynopsocus 227*, 228<br />

Martynovia 70*<br />

Martynovietla 288<br />

Martynoviidae 69, 71<br />

Massula 507, 511*<br />

Mastotermes 112*, 113<br />

Mastotermitidae 112, 113<br />

Mastotermitinae 113<br />

Maueria 192*, 193<br />

Maueriidae 191<br />

Mazonia 426*, 427*, 428<br />

Mazoniidae 426*, 428<br />

Mazoniidea 384*, 428<br />

McLuckiepteron 53<br />

Meconeminae 155<br />

Mecopodinae 155<br />

Mecoptera 42, 240*, 283, 295<br />

Mecopteroidea 36*, 239, 240*, 283<br />

Macynopteridae 50<br />

Medela 509, 513*<br />

Meditrina 503, 507*<br />

Magacercopis 180*<br />

Megachorista 284*, 285<br />

Megachosara 132, 133*<br />

Megacoris 210<br />

Megakhosarella 132, 133*<br />

Megakhosaridae 132<br />

Megakhosaridea 119, 132, 133*<br />

Megaloblatta 108*<br />

Megalodontidae 346<br />

Megalodontidea 346<br />

Megalograptidae 404, 405, 406,<br />

407, 412, 414, 418*<br />

Megalograptus 404, 407, 412, 415,<br />

418*<br />

Megaloptera 240*, 269*<br />

Megalycoriomima 324*, 326<br />

Megalyridae 349<br />

Meganeura 73*<br />

Meganeuridae 73, 74<br />

Meganeurinae 74<br />

Meganeuruta 74<br />

Meganisoptera 42 , 73*<br />

Megapodagrionidae 79<br />

Megapolystoechus 282<br />

Megapsocidium 228<br />

Megaptilidae 50<br />

Megasecoptera 32, 37*, 64<br />

Megasecopteroidea 42, 49, 64<br />

Megaspilidae 353<br />

Megaspilini 353<br />

Megastigmus 352*<br />

Megatypus 74<br />

Megelytridae 115<br />

Megisthanidea 472<br />

Meinertillidae 48<br />

Meioneurites 279<br />

Melamnous 282<br />

Melaneimon 282<br />

Melaenophora 507, 511*<br />

Melanozetes 458*, 460<br />

Melasidae 259*<br />

Melbournopterus 404, 422<br />

Meloidae 264*, 265<br />

Melyridae 258, 259*<br />

Membracidae 173,177, 178*<br />

Membracides 177<br />

Membracinae 177, 178<br />

Memabrcini 178<br />

Memoratrix 505, 509*<br />

Mengea 268*<br />

Mengeidae 268<br />

Menthidae 441<br />

Merostomata 378, 379, 380, 381, 382<br />

383, 384*, 385, 386, 387, 389,<br />

391, 403, 404<br />

Merothripidae 237, 238<br />

Mesacridites 132<br />

Mesateuropsis 200 *<br />

Mesaulacinus 350*<br />

Mesephemera 57*<br />

Mesephemeridae 57*<br />

Mesepemeridea 56<br />

Meseumolpites 266*<br />

Mesitoblatta 94<br />

Mesoatracis 189*<br />

Mesobaetis 61*. 62<br />

Mesobelostomum 213*, 214<br />

Mesobittacus 292<br />

Mesoblattina 100*, 102, 103, 106<br />

Mesoblattininae 100*, 102<br />

Mesoblattopsis 103<br />

Mesoblattula 100*, 103<br />

Mesochorista 289*<br />

Mesochoristidae 286<br />

Mesochrysopa 282<br />

Mesochrysopidae 277, 281<br />

Mesocicada 168*, 170<br />

Mesocicadella 183<br />

Mesocixiella 184, 185*<br />

Mesocixiodes 181<br />

Mesocixioides 186<br />

Mesocixius 186<br />

Mesococcus 201*, 203<br />

Mesocupes 244*, 246<br />

Mesodascilla 257*<br />

Mesodiphthera 171<br />

Mesoforficula 115<br />

Mesogereon 163*, 164<br />

Mesogereonidae 163*, 164<br />

Mesogomphus 85<br />

Mesognjllacris 150<br />

Mesogryllus 152<br />

Mesohelorus 353*, 354<br />

Mesohemerobius 282<br />

Mesohemipteron 171<br />

Mesojassoides 177<br />

Mesojassula 181<br />

Mesojassus 176*, 177<br />

Mesojurina 159*<br />

Mesoledra 177<br />

Mesoleon 282<br />

Mesoleuctra 137*, /3S*<br />

Mesolimulidae 390, 400*, 40/<br />

Mesolimulus 390, 400*. 401<br />

Mesolygaeus 222<br />

Mesomphalocoris 212<br />

Mesonemura 135*, /36<br />

Mesonepa 214<br />

Mesoneta 62, 63*<br />

Mesonirvana 174<br />

Mesonotoperla 132<br />

Mesonymphes 274<br />

Mesopanorpa 290*, 29/<br />

Mesopanorpidae 286<br />

Mesopanorpodes 289<br />

Mesophantasma 319*<br />

Mesophantasmatidae 3/9<br />

Mesophantasmatidea 310, 3/S, 319*<br />

Mesophlebia 83<br />

Mesophonidae 427*, 428, 432<br />

550<br />

http://jurassic.ru/<br />

Mesophonus 427*, 432<br />

Mesoplecia 329*, 330<br />

Mesopleciella 329*, 330<br />

Mesoplectopteron 57*<br />

Mesopolphoridae 461<br />

Mesopolystoechotidae 274, 276<br />

Mesopolystoechus 275*, 276<br />

Mesopsyche 294<br />

Mesopsychidae 292, 294<br />

Mesopsychopsis 279<br />

Mesoraphidia 271*, 272<br />

Mesoraphidiidae 270, 271*, 272<br />

Mesorhypus 331*<br />

Mesorrhyphoides 331*<br />

Mesorthopteridae 124<br />

Mesorthopteron 123*<br />

Mesosagrites 266*<br />

Mesosciophila 324*<br />

Mesosciophilodes 324*<br />

Mesoscytina 181<br />

Mesosmylina 276<br />

Mesostigmata 384*, 385, 448, 449,<br />

471, 472<br />

Mesotaeniopteryx 135*, /36<br />

Mesotarbus 479<br />

Mesothelae 494<br />

Mesothoris 249<br />

Mesothrips 238<br />

Mesothytnburis 174<br />

Mesotipula 313<br />

Mesotitanidae 122<br />

Mesotrichopteridium 297<br />

Mesotubttustrium 188*<br />

Mesypochrysa 281*<br />

Metablattula 103<br />

Metachorus 94, 97<br />

Metadiaphanopterodea 69, 7/*<br />

Metafulgoridium 184<br />

Metapalaeodictyoptera 50, 52*, 53<br />

Metaphyloblatta 94<br />

Metapoblatta 94<br />

Metapsocidium 227<br />

Metarchitaulius 297<br />

Metatarbus 480<br />

Metatranychus 469*<br />

Metatrichopteridium 297<br />

Metaxys 94<br />

Metaxybittacus 292<br />

Metaxybtatta 96*, 97<br />

Meteorinae 349<br />

Metidelia 120*. /2/<br />

Metoedischia 147*<br />

Metropator 38, 239, 286, 287*<br />

Metropatridae 42, 2S6, 287*<br />

Miaroblatta 92*, 93<br />

Miastor 333*, 334<br />

Micaria 507, 511*<br />

Micraulacinus 350*<br />

Micreremus 454*, 456<br />

Microblattina 117<br />

Microdon 34*<br />

Microfrenata 304, 305*<br />

Microgastrinae 349<br />

Microlabius 424, 428, 429*<br />

Micropholcus 500*<br />

Micropterygidae 303<br />

Micropteryginae 303<br />

Micropteryx 303*<br />

Microptysma 295*<br />

Microptysmatidae 295<br />

Microptysmodes 295*<br />

Microthelyphonida 437

M<br />

icrotypus 350*<br />

Microzetidae 460<br />

Micryphantes 503, 504*<br />

Micryphantidae 501, 503, 504*<br />

Migalomorphae 384* 385<br />

Migidae 497<br />

Milichiidae 342<br />

Mimallactoneura 326*, 328<br />

Mimalycoria 323*<br />

Mimetidae 501, 503, 504*<br />

Mindarina 198<br />

Mindarinae 197, 198<br />

Mindarus 196*<br />

Minuta 178*, 179<br />

Miocenogenia 63*, 64<br />

Miomatoneura 141*<br />

Miomatoneurella 141*<br />

Miomoptera 118, 140, 141*<br />

Miridae 217, 218*<br />

Miridoides 217, 218*<br />

Mischoptera 65*<br />

Mischopteridae 64, 66<br />

Mischopterinae 66<br />

Misthodotes 56*<br />

Misthodotidae 56*<br />

Misumena 509, 513*<br />

Mitchelloneura 170<br />

Mithraeidae 501, 503, 507*<br />

Mitinovia 131*, 132<br />

Mixopteridae 405, 406, 407, 412,<br />

414, 417*<br />

Mixopteridea 407, 408, 412<br />

Mixopterus 403, 407, 412, 414, 417*<br />

Mixotermitidae 122<br />

Mizalia 501, 503<br />

Mochlonyx 315*<br />

Mogoplistinae 154<br />

Molanna 301*<br />

Molannidae 299, 302<br />

Molannodes 301*<br />

Monomorium 357*<br />

Mononeura 239*<br />

Monura 37*, 45, 48*<br />

Morbaeschna 85<br />

Mordellidae 264*<br />

Moristinae 155<br />

Mundidae 181, 182*<br />

Mundus 182*<br />

Municeps 501, 503*<br />

Musca 343*, 344<br />

Muscidae 34*, 343, 344<br />

Muscidea 341, 343*<br />

Muscides 344<br />

Muscidomorpha 308*<br />

Musidora 340*<br />

Musidoridae 340<br />

Musidoromima 311*<br />

Musidoromimidae 311<br />

Musidoromorpha 308*, 310, 340*<br />

Mutillariae 356<br />

Mutillidae 356<br />

Mycetobia 324*<br />

Mycetobiidae 326<br />

Mycetobiinae 326<br />

Mycetophagidae 261*, 262<br />

Mycteropidae 404, 408, 420, 421*,<br />

422<br />

Mycteropidea 408, 415, 420, 421*,422<br />

Mycterops 408, 420, 421*, 422<br />

Mycterus 263*<br />

Mydaidae 337<br />

Mydasii 337<br />

Mygalomorphae 384*, 385, 492,<br />

494 , 497<br />

Myiomorpha 308*, 310, 340<br />

Mylacridae 89, 96*<br />

Mylacrides 97<br />

Mylacridium 98, 99*<br />

Mylacrinae 96*, 97<br />

Mylacrinariae 97<br />

Mylacrini 97<br />

Mylacris 96*, 97<br />

Mymallactoneura 326*<br />

Mymarea 353<br />

Mymaridae 31, 352, 353<br />

Myobiidae 461, 462, 467<br />

Myodochidae 222<br />

Myriopoda 20<br />

Myrmecophilinae 154<br />

Myrmeleontidae 273, 274<br />

Myrmeleontoidea 273*<br />

Myrmicidae 357, 358<br />

Myrmicinae 357 , 358<br />

Myrmochernetidae 442<br />

Myro 505, 510*<br />

Mystagogus 503, 507*<br />

Nabidae 217, 218*<br />

Nabini 217<br />

Nacekomia 146<br />

Nactodipoena 501, 503*<br />

Nanhermanniidea 454<br />

Nannoblattina 103<br />

Nannochoristella 290<br />

Nannochoristidae 290<br />

Nannochoristinae 286<br />

Nanorchestidae 447, 449, 450, 462*<br />

Narkema 122<br />

Narkemidae 122<br />

Narkemina 122, 123*<br />

Naucoridae 209, 214, 215*<br />

Naucoridei 214<br />

Naucoris 215*<br />

Nearoblatta 103<br />

Nebrioides 254<br />

Necromera 263*<br />

Necrophasma 160*<br />

Necrophasmatidae 160<br />

Necrotaulidae 295, 297<br />

Necrotauliodes 297<br />

Necrotaulius 296*, 297<br />

Necymylacris 97<br />

Necymylacrodes 97<br />

Neelidae 47<br />

Neides 221*<br />

Nelipoda 385, 497<br />

Nemastoma 480*, 483<br />

Nemastomatidae 483, 490<br />

Nemastomoides 480*, 482<br />

Nemastomoididae 482, 490<br />

Nematophora 23, 24<br />

Nemestrinidae 336, 337<br />

Nemestrinus 338*<br />

Nemopalpidae 316<br />

Nemopalpinae 316<br />

Nemopteridae 272, 273, 274<br />

Nemoura 135*<br />

Nemouridae 134, 136<br />

Neoageta 287<br />

Neobelinurus 398*<br />

Neobisiidae 441<br />

Neobisiidea 441<br />

Neobisiinea 384*, 385, 439, 441<br />

Neobisium 440*, 441<br />

551<br />

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Neochoristella 290<br />

Neocribellatae 515<br />

Neoephemeridea 57<br />

Neohagla 151<br />

Neolimulidae 389, 394, 395, 396*,<br />

397<br />

Neolimulus 389, 394, 396*, 397<br />

Neoliodes 452*, 456<br />

Neoliodidae 452, 456<br />

Neomylacridae 98<br />

Neomylacrini 97, 98<br />

Neomylacrls 96*, 98<br />

Neoneurinae 349<br />

Neoparachorista 291<br />

Neopermopanorpa 286<br />

Neopetromantis 288<br />

Neoptera 35, 37, 49, 85<br />

Neoribates 459*, 461<br />

Neorthophlebia 291*, 292<br />

Neorthophlebiidae 286, 291*, 292<br />

Neorthroblattina 98, 99*<br />

Neorthroblattinidae 89, 98, 99*<br />

Neorthroblattininae 98<br />

Neoscorpii 424<br />

Neoscorpiones 425<br />

Neostrabops 389, 393<br />

Neosymplicius 97<br />

Nepa 213*<br />

Nepariae 214<br />

Nephila 505<br />

Nepidae 213*, 214, 226<br />

Nepioblatta 102<br />

Nepidium 214<br />

Neriidae 342<br />

Nereiides 342<br />

Nesticidae 501<br />

Neuburgia 52*, 53<br />

Neuburgiidae 53<br />

Neureclipsis 296*<br />

Neuropibrocha 182*, 183<br />

Neuropsyche 294<br />

Neuroptera 122 , 240*, 269, 272<br />

Neuropteroidea 36*, 239, 269<br />

Neurostigminae 234<br />

Neuroterus 350*<br />

Nicoletiidae 48<br />

Nitidula 261*<br />

Nitidulidae 261*<br />

Nitidulina 261*<br />

Noctuidae 307<br />

Noctuidea 307<br />

Nosodendridae 257*, 258<br />

Nosodendron 257*<br />

Notaspididae 452, 459*, 460<br />

Notaspididea 454, 459<br />

Notaspis 459*, 460<br />

Nothrus 452*, 454<br />

Notobtattites 103<br />

Notolampra 106*<br />

Notoligotomidae 139<br />

Notonecta 215*<br />

Notonectida 214<br />

Notonectidae 214, 215*<br />

Notonectites 215<br />

Notopamphagopsis 152<br />

Notostigmata 447, 448, 449, 473<br />

Nycteribiomorpha 308*<br />

Nyctiboridae 110<br />

Nyctiborides 110<br />

Nyctiborinae 100, 108*. 110<br />

Nyctophylax 183*<br />

Nymphites 27'4

Nymphitidae 273, 274<br />

Nymphomyia 309*<br />

Nymphomyiomorpha 308*<br />

Obnisus 503, 507*<br />

Obrutschewia 393<br />

Odonata 37*, 73<br />

Odonatoidea 49, 73<br />

Odontoceratidae 299, 301<br />

Odontopus 221*<br />

Oecanthinae 154<br />

Oecobiidea 515<br />

Oecophoridae 306<br />

Oecophorinae 306<br />

Oedemera 263*<br />

Oedemeridae 262, 263*<br />

Oedischia 147*<br />

Oedischiidae 146, 147*<br />

Oedischiidea 146, 150<br />

Oiratia 93*, 94<br />

Olbiogastridae 331, 332<br />

Olenelloidea 389<br />

Oletroblatta 95*<br />

Olgamartynovia 224*, 225<br />

Oligamatites 307*<br />

Oligochelifer 440*, 442<br />

Oligochern.es 440*, 442<br />

Oligoneoptera 35, 36*, 37*, 85, 86<br />

Oligoneuridae 60, 61<br />

Oligophryne 331*, 332<br />

Oligophryneidae 331,332<br />

Oligosaldina 216*<br />

OUgotoma 139*<br />

Oligotomidae 139, 140<br />

Oligotypus 73*<br />

Oligowithius 442<br />

Olios 507, 512*<br />

Olpiidae 441<br />

Omaliidae 122<br />

Ommatocepheus 457*, 458<br />

Omphralidae 337<br />

Oniscomorpha 23, 24<br />

Onychiuridae 46<br />

Onychopalpida 473<br />

Onychophora 533<br />

Onychopterella 407, 410<br />

Oochorista 287, 288*<br />

Oomartus 490, 492*<br />

Oonopidae 498*, 499*<br />

Ootarbus 480<br />

Ophion 350*<br />

Ophioninae 352<br />

Ophismoblatta 100*, 103<br />

Ophthalmocoris 210<br />

Opilio 480*. 483<br />

Opilioacaridae 450, 473<br />

Opilioacarina 383, 384*. 385, 448,<br />

449, 450, 473<br />

Opiliocarina 447<br />

Opiliomorphae 384*. 385, 474<br />

475, 476, 477, 482<br />

Opiliones 380, 381, 383, 384*. 385,<br />

444, 450, 474, 475, 476, 477, 479,<br />

480*, 481*<br />

Opiliotarbidae 479<br />

Opiliotarbus 479*<br />

Oppia 454*, 456<br />

Oppites 456<br />

Orchesia 264*<br />

Orchestina 498, 499*<br />

Oribatei 381, 384*. 385, 448, 449<br />

450 451 452, 454, 474, 490<br />

Oribatella 458*, 460<br />

Oribatellidae 458*. 460<br />

Oribatidae 454*, 456<br />

Oribatulidae 458*, 460<br />

Oribotritia 459*. 461<br />

Oripoda 458*, 460<br />

Oripodidae 458*, 460<br />

Orius 216*<br />

Ormenis 187*<br />

Ornaticosta 125*, 126<br />

Ortatides 342<br />

Ortalididae 342<br />

Orthezia 201*<br />

Orthezides 202<br />

Ortheziidae 201, 202<br />

Orthocostidae 52<br />

Orthognatha 494, 497<br />

Orthocladiinae 318<br />

Orthomylacris 97<br />

Orthophlebia 290*, 291, 292<br />

Orthophlebiidae 286, 290*<br />

Orthophlebiites 291<br />

Orthophlebioides 291<br />

Orthoprosbole 167*, 168<br />

Orthopsocus 228<br />

Orthoptea 145, 146<br />

Orthopteroidea 36*, 88, 144<br />

Orthoscytina 168*, 170<br />

Orthosterni 424<br />

Orthotarbus 479*, 480<br />

Orussidae 348<br />

Orussidea 346, 348<br />

Oryctothemis 81<br />

Osmylida 276<br />

Osmylidae 274, 276<br />

Osmylinae 276<br />

Osmylites 276<br />

Osmylitidae 274, 276<br />

Osmylopsis 282<br />

Osmylopsychopidae 277<br />

Ostoma 260*<br />

Ostomatidae 260*. 261<br />

Ostracoda 41<br />

Otitidae 342<br />

Otocepheus 457*, 458<br />

Otocryptops 26*<br />

Oulopterigidae 89<br />

Oxynoblatta 97<br />

Oxyopidae 505<br />

Oxypoei 436<br />

Oxyrhachinae 177<br />

Pachyblatta 94<br />

Pachygnathidae 449, 462*, 464<br />

Pachygnatidea 450, 462<br />

Pachygnathus 447<br />

Pachymeridiidae 211*<br />

Pachymeridium 211*<br />

Pachymerium 26*<br />

Pachymerus 226<br />

Pachyneuridea 310<br />

Pachyneuroblattina 103<br />

Pachypsyche 167<br />

Pachytroctidae 228, 230<br />

Pachytroctinae 230<br />

Pachytylopsidae 122<br />

Paedephemera 58*<br />

Paedephemeridae 58*<br />

Paedephemeridea 57, 58<br />

Palademosyne 252*, 253<br />

Palaeacariformes 450, 451<br />

Palaemerobiidae 277, 279, 280*<br />

552<br />

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Palaemerobius 279, 280*<br />

Palaeoacariformes 384*, 385, 447<br />

Palaeobuthus 428<br />

Palaeocharinidae 484*, 486<br />

Palaeocharinus 484*. 486<br />

Palaeocharinoides 484*, 485, 486<br />

Palaeochrysa 281*<br />

Palaeocixiidae 124<br />

Palaeocossus 164*, 166<br />

Palaeocoxopleura 23<br />

Palaeocribellatae 515<br />

Palaeocteniza 496<br />

Palaeocupes 243, 244*<br />

Palaeodictyoptera 32, 50, 161<br />

Palaeodictyopteroidea 49, 50<br />

Palaeodrassus 507<br />

Palaeoedischia 146<br />

Palaeogyrinidae 254*, 255<br />

Palaeogyrinus 254*, 255<br />

Palaeohesperinus 323*<br />

Palaeoheteroptera 214<br />

•Palaeoisopodidae 527<br />

Palaeoisopus 526, 527, 528*<br />

Palaeomachus 432*, 433<br />

Palaeomanteidae 140, 141<br />

Pataeomantis 141*, 142<br />

Palaeomantisca 142<br />

Palaeometa 505<br />

Palaeomorpha 23<br />

Palaeonemoura 135*, 136<br />

Palaeontina 167<br />

Palaeontinidae 43, 164*<br />

Palaeontinidea 164*<br />

Palaeontinodes 164*, 166<br />

Palaeontinopsis 164*, 166<br />

Palaeopachygnatha 505<br />

Palaeopantopoda 525,529<br />

Palaeopantopodidae 529<br />

Palaeopantopus 528*, 529<br />

Palaeoperla 134, 135*<br />

Palaeoperlidae 134, 136<br />

Palaeophonidae 384*, 425, 426*<br />

Palaeophonus 424, 425, 426*<br />

Palaeophora 340.*<br />

Palaeophoridae 340<br />

Palaeopisthacanthus 427*, 431 *<br />

Palaeoplecia 319, 320*<br />

Palaeopleciidae 3/9<br />

Palaeopsocus 235*<br />

Palaeopsylla 345*<br />

Palaeoptera 35, 36*, 37*. 49<br />

Palaeoptillus 53<br />

Palaeorehnia 153<br />

Palaeorehniinae 152, /53<br />

Palaeoscardiites 305*<br />

Palaeoscorpiidae 425, 423<br />

Palaeoscorpiones 425<br />

Palaeoscorpius 428<br />

Palaeoscytina 192*, /94<br />

Palaeostraca 382<br />

Palaeostratiomyia 333*. 334<br />

Palaeostratiomyiidae 334<br />

Palaeotaeniopteryx 135*, /36<br />

Palaeotaulius 297<br />

Palaeotendipes 317*<br />

Palaeotroctes 229*<br />

Palaeotyroglyphus 449, 452*<br />

Palaeotydeus 464*, 466"<br />

Palaeovieia 190<br />

Palaeoblattula 103<br />

Palaranea 495<br />

Paleobittacus 292

Paleolimulidae 390, 397, 400*<br />

Paleolimulus 390, 400*. 401<br />

Paleomeridae 388, 389, 39/, 392*<br />

Paleomerus 389, 392*<br />

Paleuthygramma 143*<br />

Paleuthygrammatidae 142<br />

Palingenia 59, 60*<br />

Palingeniidae 59, 60*<br />

Palingeniopsis 57*<br />

Palingeniopsidae 57<br />

Palmoblattina 103<br />

Palparites 282<br />

Palpatores 385,482, 483<br />

Palpigrades 445<br />

Palpigradi 378, 380, 381, 382, 383,<br />

384*, 385, 402, 403, 433, 437,<br />

438*, 439, 451, 475<br />

Palpigradida 437<br />

Palpimanidae 501<br />

Pamphagopsidae 150<br />

Pamphagopsis 152<br />

Pamphilidae 346, 347<br />

Pamphilinae 347<br />

Panchlora 106*<br />

Panchloriens 107<br />

Panchlorini 107<br />

Panestia 87*<br />

Panestiidae 87*. 89<br />

Panestiinae 87*<br />

Panorpa 32*, 291*<br />

Panorpidae 286, 290, 291*<br />

Panorpina 291<br />

Panorpodes 291<br />

Pantopoda 525, 526, 529<br />

Paoliidae 122<br />

Parabaetis 59<br />

Parabelmontia 288<br />

Parablattula 103<br />

Parabrodia 65*<br />

Parabrodiidae 64, 66<br />

Parachorista 287, 288*<br />

Paractinophlebia 279<br />

Paradictyodiptera 309*<br />

Paradunstania 164<br />

Parafulgoridium 184<br />

Parahughmilleria 423<br />

Paraiassus 190<br />

Parakalimmidae 461<br />

Paraknightia 189, 224*<br />

Paraknightiidae 189, 190, 224*<br />

Parakuloja 71*, 72<br />

Paralellophora 111<br />

Paralellophoridae 111<br />

Paralinus 515*<br />

Paraliomopterum 127*. 128<br />

Paralogidae 73, 74<br />

Paralycoriomima 324*, 326<br />

Paramecoptera 286<br />

Parandrexis 265, 266*<br />

Paraneoptera 35, 36*, 37*, 38, 85,<br />

86, 160, 162*, 238, 239*, 241<br />

Parapalaeodictyoptera 50, 52<br />

Parapanorpa 93*, 94<br />

Parapermula 125*. 126<br />

Paraplecoptera 32, 37*. 118, 119,<br />

133*<br />

Parapleurites 156*<br />

Paraprisca 129*, 130<br />

Paraprotocarus 464*, 466<br />

Parapsocida 226, 228, 229*. 232*<br />

Parapsocidium 227<br />

Parapsocus 236<br />

Parasisyra 280*<br />

Parasitica 349<br />

Parasitidae 452, 471<br />

Parasitiformes 383, 384*, 385, 447,<br />

448, 449, 450, 470*. 471, 472<br />

Parastylotermes 112*<br />

Parasylviodes 129*, 130<br />

Paratarbus 480<br />

Parataulius 297<br />

Parattidae 507, 509<br />

Parattus 509<br />

Paratenodera 111 *<br />

Paratrichoptera 283, 284, 292, 293*,<br />

294<br />

Paratrichosiphum 196*<br />

Paratydeidae 465<br />

Paraulacus 350<br />

Paraxymyia 328, 329*<br />

Paraxymyiidae 329<br />

Pareinoblatta 95<br />

Parelmoa 70*, 71<br />

Parelmoidae 69, 71<br />

Parelthoblatta 94<br />

Parelthotheniis 83<br />

Pareopolyneura 311*<br />

Parevophrys 509, 515*<br />

Parhypochthonius 447<br />

Paridelia 120*. /2/<br />

Paritonida 326*, 323<br />

Paromylacris 97<br />

Paroryssidae 343<br />

Paroryssus 347*, 343<br />

Paruroctea 500*, 501<br />

Patrodoron 89, 90*. 9/<br />

Pauropoda 20, 2/*<br />

Pauropsyllinae 206<br />

Pauropus 21*<br />

Paussidae254*. 255<br />

Paussus 254*<br />

Pectinariopsis 300<br />

Pediculoididea 447, 451<br />

Pedinoblatta 103<br />

Pedipalpi 435, 436, 438,440,441, 444<br />

Pedipalpida 389<br />

Pedipalpides 380, 383, 384*, 385,<br />

402, 403, 433, 439, 443, 444, 445,<br />

482<br />

Pelecinopteridae 349<br />

Pelecinopteron 350*<br />

Pelopiinae 318<br />

Pelops 461<br />

Pelopsidae 460<br />

Peloridiidae 207*, 208<br />

Peltoperlidae 134<br />

Pemphigiden 197<br />

Pendodelphax 189<br />

Penetoblatta 94<br />

Pentacentrinae 154<br />

Pentachrysis 355*<br />

Pentapsocidium 227<br />

Pentatomidae 223*. 226<br />

Pentatomidea 223<br />

Penthaleidae 465<br />

Penthalodes 464*. 466<br />

Penthalodidae 464*. 465, 466<br />

Penthetria 329*<br />

Penthetriidae 329, 330<br />

Penturoperlidae 134<br />

Pereboria 182*<br />

Pereboriidae 181,182*<br />

Periassophlebia 83<br />

Pergidae 348<br />

553<br />

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Perielytridae 236*<br />

Perielytrodea 226, 236<br />

Perielytron 236*<br />

Perientominae 231<br />

Perientomini 231<br />

Perientomum 232*<br />

Perilampidae 352<br />

Periplaneta 104*<br />

Peripsocidae 234<br />

Peripsocinae 234<br />

Peripsocus 235*<br />

Perisphaerini 106*, 107<br />

Perisphaerites 107<br />

Perla 32, 135*<br />

Perlariopsis 136<br />

Perlidae 134, 137<br />

Perlodidae 134<br />

Perlopseidae 134, 136<br />

Perlopsis 135*. 136<br />

Permacridites 146<br />

Permaeschna 75*, 76<br />

Permaeschnidae 74, 76<br />

Permagra 190<br />

Permagrion 77*<br />

Permagrionidae 77<br />

Permagrionidea 76, 77*<br />

Permaleurodes 203*, 204<br />

Permaleurodidae 203<br />

Permanisoptera 42, 75*<br />

Permaphidopseidae 195*<br />

Permaphidopsis 195*<br />

Permarrhaphidae 241, 242*, 243<br />

Permarrhaphus 242*. 243<br />

Permegalomidae 275<br />

Permegalomus 275*<br />

Permelcana 149*<br />

Permeleanidae 146, 149*<br />

Permella 134<br />

Permelytropsis 116<br />

Permembia 141*<br />

Permembiidae 140<br />

Permentomum 227<br />

Permepallage 77*, 78<br />

Permepallagidae 73<br />

Permepallagidea 76, 77*, 78<br />

Permiakovia 52*, 53<br />

Permithone 276<br />

Permithonidae 274, 275, 276<br />

Permithonopsis 275*, 276<br />

Permoberothidae 157<br />

Permobiella 143*<br />

Permobiellidae 142, /44<br />

Permoblattina 95<br />

Permobrachus 183<br />

Permocapnia 124<br />

Permocapniidae 124<br />

Permocentropidae 286, 290*<br />

Permocentropus 290*<br />

Permocentrus 190<br />

Permochorista 289<br />

Permochoristidae 236, 287*, 288*,<br />

289*<br />

Permochoristinae 286, 233<br />

Permocicada 168*<br />

Permocicadopsis 168<br />

Permocixiella 185*, /36<br />

Permocixius 174<br />

Permocrossos 252*. 253<br />

Permocupes 244*, 246<br />

Permocupoides 244*, 246<br />

Permodiphtera 170<br />

Permodiphtheroides 190

Permododa 175*, 176<br />

Permodunstania 163*, 172<br />

Permoedischia 148<br />

Permofulgor 189<br />

Permogtyphis 183, 190<br />

Permohymen 67*, 68<br />

Permojassinae 173, 174, 175*<br />

Permojassula 175<br />

Pernio jassus 175<br />

Permolamproptera 173*, /74<br />

Permolestes 77*<br />

Permolestidae 77<br />

Permoleuctropsis 137*, /35<br />

Permomerope 285*, 286<br />

Permomeropidae 284, 285*, 256<br />

Permoneuridae 55<br />

Permopanorpa 286, 287*<br />

Permopanorpidae 256<br />

Permopibrocha 182*, /53<br />

Permoplectoptera 56, 57*<br />

Permopsocida 226, 227*<br />

Permopsocidae 227, 225<br />

Permopsocus 227*, 228<br />

Permopsychops 276<br />

Permopsychopsidae 275<br />

Permopsylla 192*<br />

Permopsyllidae 199<br />

Permopsyllidium 200<br />

Permopsyllidops 200<br />

Permopsylloides 200<br />

Permopsyllopsis 192*, /94<br />

Permorhaphidiidae ; 146<br />

Permorapisma 276<br />

Permoscarta 190<br />

Permoscytina 194<br />

Permoscytinopsidae 191<br />

Permoscytinopsis 192<br />

Permosialidae 269<br />

Permosialis 269*, 270<br />

Permosisyra 280*<br />

Permosisyridae 279<br />

Permosmylus 276<br />

Permosyne 254<br />

Permosynidae 252*, 253<br />

Permotanyderidae 292<br />

Permotermopsis 120*, /2/<br />

Permothea 200<br />

Permotheella 200<br />

Permothemidia 54*, 55<br />

Permothemis 54*, 55<br />

Permothemistidae 55<br />

Permothripidae 237<br />

Permothrips 237*<br />

Permotipula 292, 293*<br />

Permotipulidae 292<br />

Permotrichoptera 295*<br />

Permovicia 190<br />

Permula 128<br />

Peromapteridae 50<br />

Pervestigia 168*, /70<br />

Petaluridae 53<br />

Petralycus 462*<br />

Petrochorista 288, 289*<br />

Petrablattina 94<br />

Petromantis 288, 289*<br />

Petrophlebia 83<br />

Petrothemis 83<br />

Petropypus 73*, 74<br />

Petrovicia 487*, 455<br />

Petrushevskia 275*, 276<br />

Phacelobranchus 59<br />

Phaenopteridae 124<br />

Phaenopelops 459*, 461<br />

Phalecocera 312*<br />

Phalacridae 261*, 262<br />

Phalacrus 261*<br />

Phalangiidae 482, 453<br />

Phalangiidea 480, 453<br />

Phalangiotarbi 383, 384*, 385, 476<br />

477, 479*, 482<br />

Phalangiotarbus 479*<br />

Phalangium 483<br />

Phaloniidae 306<br />

Phalangopsinae 154<br />

Phalangodidae 483<br />

Phaloniinae 306<br />

Phaneropterinae 155<br />

Phantoma 216*<br />

Phasmatidae /60<br />

Phasmatodes 31, 37*, 145, /59, 160*<br />

Phasmida 160<br />

Phasmodinae 155<br />

Phauloblatta 94<br />

Phenacolestes 79*<br />

Phenopelopidae 459*, 460<br />

Phenopelops 459*, 461<br />

Phenopteridae 124<br />

Phidippus 515*<br />

Philagra 180*<br />

Philiasptilon 69, 70*<br />

Philodromus 509, 513*<br />

Philonicus 338*<br />

Philonthus 254*<br />

Philopotamidae 295, 297<br />

Philotarsidae 234<br />

Philotarsus 235*<br />

Phipoides 287<br />

Phlebonotus 106*<br />

Phlebotomidae 3/6<br />

Phlebotominae 316<br />

Phiebotomus 316*<br />

Phloeomyzinae 197<br />

Phloeothripidae 235<br />

Phloeothrips 237*<br />

Phoberoblatta 94<br />

Pholcidae 499, 500*<br />

Pholcidea 459<br />

Pholciformia 499<br />

Pholcus 474<br />

Pholidoptilon 75*, 76<br />

Р/югЬш 343*<br />

Phoridae 339, 340<br />

Phoridea 339<br />

Phorites 340<br />

Phoromorpha 308*, 310, 339, 340*<br />

Phragmatoecicossus 164*, /66<br />

Phragmatoecites 164*, /66<br />

Phragmoligoneura 332, 333*<br />

Phragmoligoneuridae 332<br />

Phragmoligoneuridea 319, 332, 333*<br />

Phryganea 299*<br />

Phryganeidae 299*<br />

Phryganidium 184, 293, 332<br />

Phrynichida 383, 436<br />

Phrynichus 436<br />

Phrynida 436<br />

Phrynidae 437<br />

Phrynomartus 489*<br />

Phthartus 57*<br />

Phthinomytacris 97<br />

Phthiracaridae 459*. 46/<br />

Phthiracaridea 46/<br />

Phyllaphidinae 198<br />

Phyllidae 160<br />

554<br />

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Phyllocoptidae 47/<br />

Phyllophorinae 155<br />

Phylloxeridae 195, 196*, /97<br />

Phylloxerinae 197<br />

Phyloblatta89, 90*, 94<br />

Phylomylacris 97<br />

Phytoptus 469*, 471<br />

Phytoseiidae 473<br />

Pimplinae 352<br />

Pincombea 195*, 196<br />

Pincombeidae 195*, /96<br />

Pincombella 399<br />

P/negia /45 *<br />

Pinnachorista 284*, 255<br />

Pipunculidae 34/<br />

Pipunculini 341<br />

Pirontetia 83<br />

Pisauridae 505, 510*<br />

Plachutella 164*. /66<br />

Plagioblatta 94<br />

Planomartus 489*<br />

Plategeocranus 457*<br />

Plateremalidae 456<br />

Platoridae 507<br />

Platyblatta 94<br />

Platyblattina 94<br />

Platybunus 480*, 483<br />

Platycerus 257*<br />

Platychorista 283*, 284<br />

Platychoristidae 283*, 254<br />

Platycnemididae 79, 50<br />

Platycnemis 79*<br />

Platycrossos 254<br />

Platyliodes 452*, 456<br />

Platymeris 218*<br />

Platymylacris 97<br />

Platyperla 137*, /35<br />

Platypezidae 34/<br />

Platypezidea 34/<br />

Platypezinae 341<br />

Platypleurinae /72<br />

Platyseytinella 225<br />

Pleciodictya 319, 320*<br />

Pleciodictyidae 3/9<br />

Pleciodictyidea 3/9, 320*<br />

Pleciofungivora 323*<br />

Pleciofungivorella 323*<br />

Pleciofungivoridae 320<br />

Pleciomima 326*, 325<br />

Pleciomimella 326*, 325<br />

Pleciomimidae 326<br />

Plecoptera 37*, 118, /34, 135*, 137*,<br />

138<br />

Plecopteroidea 36*, 88, //5, 138<br />

Plectoptera 55, 56<br />

Plectopterinae 110<br />

Plectopterini 107, 110<br />

Pleisiogramma 142, 143*<br />

Pleisiogrammatidae /42<br />

Pleobyttacus 282<br />

Pleomartidae 490<br />

Pleomartus 492<br />

Pleophrynus 487*, 488<br />

Plesiadischia 148<br />

Plesiosiro 477*<br />

Plesiosironidae 477<br />

Pleurojulus 22*, 23<br />

Pleurolycosa 496<br />

Plyctoblattina 103<br />

Pneumoridae 156<br />

Pocononia 487*, 488<br />

Podapolipodidae 448

Poduridae 46<br />

Poduromorpha 45, 46<br />

Poecilogonalis 350*<br />

Poliochera 444<br />

Polycentropodidae 295, 298<br />

Polycentropinae 298<br />

Polycreagra 51*<br />

Polycreagridae 52<br />

Polycreagridea 50, 52<br />

Polyctoblatta 94<br />

Polycytella 159<br />

Polycytellidae 157,159*<br />

Polydesmodea 23, 24<br />

Polydesmus 22*<br />

Polydicrobittacus 292<br />

Polyglyptinae 177<br />

Polymitarcidae 59, 60*<br />

Polyneoptera 35, 36*, 37*, 85, 86,<br />

88, 241<br />

Polyneurisca 320, 321*<br />

Polyochera 443*, 444<br />

Polyocheridae 444<br />

Polyphaga 108*<br />

Polyphagidae 110<br />

Polyphaginae 110<br />

Polyphagini 111<br />

Polyphleboblatta 103<br />

Polysitum 252*, 253<br />

Polystoechotoidea 273, 274, 275*<br />

Potystomurum 408, 418, 420*<br />

Polystra 167<br />

Polytaxineura 75*, 76<br />

Polytaxineuridae 74, 76<br />

Polyxenidae 22<br />

Polyxenus 22*<br />

Polyzoniidae 24<br />

Pomerantziidae 465<br />

Pompilidae 358, 359<br />

Pompilidea 349, 358*<br />

Pompilides 359<br />

Pomponia 172*<br />

Ponera 357*<br />

Ponerinae 357<br />

Poroblattina 100<br />

Poroblattinidae 89, 98, 100*<br />

Potamanthidae 59, 60*<br />

Potamanthus 59, 60*<br />

Potamyia 296*<br />

Praemerothrips 237*<br />

Praemordella 264*<br />

Praemordellidae 264*<br />

Praemordellinae 264<br />

Prantlites 94<br />

Premnoblatta 100<br />

Pringlia 399<br />

Prisca 130<br />

Probascanion 209*, 210<br />

Probascanionidae 209*, 2/0<br />

Proberothidae 277<br />

Probittacus 291*, 292<br />

Probnisidae 124<br />

Procalosoma 254<br />

Procercopidae 179, /30*<br />

Procercopina 180*, /3/<br />

Procercopis 180*<br />

Prochilinae 155<br />

Prochoristella 298<br />

Prochoroptera 70*<br />

Prochoropteridae 69<br />

Procoptoblatta 94<br />

Proctotrupidae 353, 354<br />

Proctotrupidea 349, 353*<br />

Procyrtophyllites 152<br />

Prodidomidae 507<br />

Prodinapsis 350*<br />

Proelcana 149*<br />

Progenentomum 148<br />

Proglomeridae 23<br />

Progoneura 76, 77*<br />

Progonocimex 210*<br />

Progonocimicidae 2/0*<br />

Progonocoris 210*<br />

Progonophlebia 80*<br />

Progonophlebiidae 3/<br />

Progonopsocus 228<br />

Prohagla 151<br />

Prohaglinae 151<br />

Prohemerobiidae 277<br />

Prohesperinus 323*<br />

Proiulidae 23<br />

Projapygidae 47<br />

Prolimulus 390, 400*<br />

Prolyonetia 305*<br />

Promygale 490, 491*<br />

Promygalidae 490<br />

Promylacris 97<br />

Pronabis 211<br />

P roparagryllacris 150<br />

Propatrix 199, 200*<br />

Propetes 515<br />

Prophalangopseidae 150<br />

Prophalangopseinae 150, /5/<br />

Prophalangopsis 152<br />

Proptychoptera 312*, 314<br />

Proraphidia 271*. 272<br />

Prorhyacophilidae 295, 296<br />

Prosbole 167*, /63<br />

Prosbolidae 167*, /OS<br />

Prosbolina 168<br />

Prosboloneura 167*. /65<br />

Prosbolopseidae 203, 209*<br />

Prosbolopsis 208, 209*<br />

Prosbolopsites 179*, 180<br />

Proscopiidae 156<br />

Proscorpius 425, 426*<br />

Prosepididontidae 295, 297<br />

Prostenoneuridae 117<br />

Prostigmata 462<br />

Protacarus 448, 449, 462*, 464,465,<br />

466<br />

Protagriidae 50<br />

Protallactoneura 321*<br />

Protanisoptera 42, 73, 74, 75*<br />

Protelytridae 115, //0<br />

Protelytron 115*, 116<br />

Protelytropsis 116<br />

Protelytroptera 37*, 42, 88, //5*<br />

Protembiidae 124<br />

Protendipedidae 316, 3/7<br />

Protendipes 317*<br />

Protentomobrya 46<br />

Protentomobryidae 46<br />

Protentomobryomorpha 45, 46<br />

Protephemeroptera 55*<br />

Protereisma 56*<br />

Protereismatidae 56*<br />

Protereismatidea 56<br />

Protereismephemeridae 56<br />

Proterema 100*<br />

Protereminae /00*<br />

Proteroscarabeus 257<br />

Prothelyphonus 434*, 436<br />

Protipula 313<br />

Protobibio 329*. 330<br />

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Protobibionidae 330<br />

Protobittacus 291*, 292<br />

Protoblattodea 37*, 88, //6*, 118<br />

Protobrachyceridae 334<br />

Protobrachyceron 333*<br />

Protochorista 288<br />

Protochoristella 289<br />

Protochoristinae 236<br />

Protocoleidae 263*<br />

Protocoleus 268*<br />

Protocoridae 211*<br />

Protocoris 211*, 212, 226<br />

Protocteniza 494*<br />

Protocupes 244*, 246<br />

Protocupoides 244*, 246<br />

Protocyrtidae 334<br />

Protocyrtus 333*, 334<br />

Protodermaptera //4<br />

Protodiaphanopterodea 69, 70*<br />

Protodiplatydae //4<br />

Protodiplatys 114*<br />

Protogryllinae 154<br />

Protogryllus 153*<br />

Protohymen 67*, 63<br />

Protohymenidae 67, 68<br />

Protohymenoptera 42, 64, 66*, 67*<br />

Protojurina 158*<br />

Protokollariidae 122<br />

Protolbiogaster 331*, 332<br />

Protolbiogastridae 331/332<br />

Protoligoneura 319, 320*<br />

Protoligoneuria 62<br />

Protoligoneuridae 3/9<br />

Protoligoneuridea 3/9, 320*<br />

Protolimulus 390, 398<br />

Protolindenia 84*<br />

Protolycosa 494<br />

Protomecoptera 239, 284<br />

Protomeropidae 284<br />

Protomphrale 337, 338*<br />

Protomphralidae 337<br />

Protomutilla 357*<br />

Protomyrmeleon 78, 79 *<br />

Protomyrmeleontidae 73<br />

Protomyrmeleontoidea 76, 73, 79*<br />

Protonychophora 533<br />

Protopalpigradi, 437<br />

Protopanorpa 287, 288, 289 .<br />

Protopanorpoides 287, 288*<br />

Protopantopoda 526, 527<br />

Protophasmidae 117<br />

Protophrynus 435*. 437<br />

Protopilio 482<br />

Protopincombea 196<br />

Protoplecia 329*, 330<br />

Protopleciidae 329<br />

Protopolphoridae 461<br />

Vrotoprosbole 38, 163*, 164<br />

Protoprosbolidae 163, /64<br />

Protopsyche 167<br />

Protopsychopsis 278<br />

Protopsyllidiidae /99, 200*<br />

Protopsyllidiidea /99<br />

Protopsyllidium 200<br />

Protopsyllops 200<br />

Protorhagio 335*, 337<br />

Protorhyphidae 331, 332<br />

Protorhyphus 331*. 332<br />

Protoribates 459*, 460<br />

Protorthophlebia 290*, 29/<br />

Protorthoptera /45<br />

Protoscatopse 331*

Protoscatopsidae 330<br />

Protoscorpiones 425<br />

Protosmylinae 276<br />

Protosolpuga 443*, 446*<br />

Protosolpugidae 446<br />

Protospeleorchestes 462*<br />

Prototettigidae 122<br />

Protothrips 237*<br />

Protozygoptera 76<br />

Protura 37*, 45, 379<br />

Protziidae 471<br />

Pruvostites 147*<br />

Pruvostitidae 146<br />

Psammolimulus 390, 400*, 401<br />

Psechridae 516*, 517<br />

Pselaphidae 254*, 255<br />

Pselaphognatha 22<br />

Psephenidae 258<br />

Pseudagrion 79*<br />

Pseudelmoa 71<br />

Pseudipsvicia 190<br />

Pseudocistella 264*<br />

Pseudocossus 164*, 166<br />

Pseudodiptera 293*<br />

Pseudodipteridae 292 , 293<br />

Pseudogarypidae 441, 442<br />

Pseudogarypus 440*, 442<br />

Pseudogramma 143*<br />

Pseudohagla 151*, 152<br />

Pseudohumbertiella 151*, 152<br />

Pseudohymen 67*, 68<br />

Pseudokreischeria 488<br />

Pseudomantis 142<br />

Pseudomopidae 104<br />

Pseudomopinae 100, 104*, 105*,<br />

106<br />

Pseudomylacridae 97<br />

Pseudomylacrini 97<br />

Pseudomylacris 96*, 97, 98<br />

Pseudomyrmeleon 282<br />

Pseudomyrmicinae 357<br />

Pseudoniscidae 389, 391, 395, 396*,<br />

397<br />

Pseudoniscus 389, 394, 395, 396*,<br />

397<br />

Pseudooedischia 146<br />

Pseudoorthophlebia 297<br />

Pseudophanides 188<br />

Pseudophyllinae 155<br />

Pseudophytlodromia 106<br />

Pseudopolycentropidae 292<br />

Pseudopolycentropus 293*<br />

Pseudopropetes 515<br />

Pseudoprotacarus 462*. 465<br />

Pseudoscorpiones 383, 439, 450<br />

Pseudoscorpionoidea 382, 385, 402,<br />

403, 433, 439, 440* 441, 444, 451<br />

Pseudosirex 348<br />

Pseudosiricidae 348<br />

Pseudotettigonia 155*<br />

Pseudotettigoniinae 155<br />

Psocida 234<br />

Psocidae 234<br />

Psocidiidae 227<br />

Psocidium 227<br />

Psococicadallopsis 229*. 230<br />

Psocopsyllidium 200<br />

Psocoptera 162*, 226, 236<br />

Psocopteroidea 161, 226<br />

Psocoscytina 200<br />

Psocus 235*<br />

Psoquillidae 228<br />

Psorergatidae 461<br />

Psychidae 304<br />

Psychocoridae 211*, 212<br />

Psychocoris 211*, 212<br />

Psychoda 316*<br />

Psychodidae 316<br />

Psychodidea 310, 314, 316*<br />

Psychodites 316<br />

Psychomyidae 295, 298<br />

Psychopidae 43, 277*<br />

Psychopsidoidea 277<br />

Psychopsinae 277<br />

Psylla 207*<br />

Psyllidella 200<br />

Psyllidae 206, 207*<br />

Psyllidea 206<br />

Psyllidiana 200<br />

Psyllinae 206, 207<br />

Psyllipsocidae 228, 232<br />

Psyllipsocini 232<br />

Psyllidea 203<br />

Psyllipsocus 232*<br />

Psyllomorpha 190, 203<br />

Ptenidium 256*<br />

Pteridomylacridae 98<br />

Pteridomylacrinae 96*, 98<br />

Pteridomylacris 96*. 97, 98<br />

Pterodophyllites 203<br />

Pterinoblattina 279<br />

Pterocommatinae 197<br />

Pterogasterina 459<br />

Pteromalidae 352<br />

Pteronarcidae 134<br />

Pteronidea 347*<br />

Pteronidiidae 50<br />

Pterophyllinae 155<br />

Pteroplistinae 154<br />

Pteropteron 189<br />

Pterygosomatidae 462<br />

Pterygota 35, 36*. 38, 49, 86<br />

Pterygotidae 403, 405, 407, 411,<br />

414*. 415*<br />

Pterygotidea 407, 408, 411<br />

Pterygotus 404, 405*, 406*, 407,<br />

408, 412, 414*. 415*, 423<br />

Ptiliidae 31, 256*<br />

Ptilomylacris 97<br />

Ptinidae 260*<br />

Ptinus 260*<br />

Ptychoptera 312*<br />

Ptychopteropsis 293*, 294<br />

Ptyctima 454, 461<br />

Piccinites 203<br />

Punctoribates 459*, 460<br />

Purkynia 23<br />

Pursa 117<br />

Pycnochelifer 442<br />

Pycnogastrinae 155<br />

Pycnogonides 377, 379, 525, 526,<br />

528*<br />

Pycnophlebia 152<br />

Pycnoscelus 106*<br />

Pycnothelidea 497<br />

Pycnotheloides 385<br />

Pycnothemis 83<br />

Pyrojulus 23<br />

Pyralides 307<br />

Pyralididae 307<br />

Pyralidea 304, 307<br />

Pyrgotidae 342<br />

Pyritaranea 496*, 497<br />

Pyritaraneidae 494, 496*, 497<br />

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Pyrochroa 263*<br />

Pyrochroidae 263*<br />

Pyrrhocoridae 221*, 222<br />

Pyrrhoco rides 222<br />

Pythidae 263*<br />

Pythonissa 507, 511*<br />

Rachiceridae 334, 336<br />

Rachicerus 335*<br />

Rachimentomidae 119<br />

Rachimenotmon 119, 120*<br />

Rakovnicia 495*<br />

Rammeinae 155<br />

Ranatra 213*<br />

Raphidia 34*, 271*, 272<br />

Raphidiidae 270, 271*. 272<br />

Raphidiinae 272<br />

Raphldioptera 240*, 269, 270*, 271*<br />

Raphidiophorinae 152<br />

Rectineuridae 55<br />

Reculidae 124<br />

Reduviidae 218*, 219<br />

Reduviini 219<br />

Reeveana 254<br />

Reticulocicada 189*<br />

Rhabdoptilidae 52<br />

Rhabdothemis 83<br />

Rhaetiblattina 103<br />

Phaetofungivora 320*<br />

Rhaetofungivorella 320, 321*<br />

Rhaetofungivorodes 320, 321*<br />

Rhaetomyia 318*<br />

Rhaetomyiidae 318<br />

Rhaetomyiidea 310, 318*<br />

Rhagidiidae 448<br />

Rhagionempididae 336<br />

Rhagionempis 335*, 336<br />

Rhagionidae 336<br />

Rhagionides 336<br />

Rhagiophryne 335*. 337<br />

Rhaphidiopseidae 64, 66<br />

Rhaphidiopsis 65*<br />

Rhaphignathidae 463, 465<br />

Rhaphignathidea 462, 465<br />

Rhenopteridae 408, 416, 420, 421*<br />

Rhenopterus 408, 420, 421*<br />

Rhinocarcinosoma 407, 413, 416*<br />

Rhinocoris 218*<br />

Rhinonyssidae 473<br />

Rhinortharia 183<br />

Rhinotermitidae 112, 113<br />

Rhinotermitinae 113<br />

Rhipidioptera 117<br />

Rhipidoblattina 100*, 102<br />

Rhipiphoridae 265<br />

Rhizophagidae 261*, 262<br />

Rhizophagus 261*<br />

Rhodacaridae 449, 471<br />

Rhodesiomylacris 97<br />

Rhogadinae 349<br />

Rhombocoleidae 249, 250*<br />

Rhombocoleus 249, 250*<br />

Rhopalocera 307<br />

Rhopalosomatidae 358<br />

Rhyacophila 296*, 297<br />

Rhyacophilidae 295, 297<br />

Rhynchota 161<br />

Rhyncophthiraptera 226<br />

Rhyniella 46*<br />

Rhyniognatha 46<br />

Rhyphidae 331, 332<br />

Rhyphidea 319, 331*

Rhyphites 332<br />

Ricanides 186<br />

Ricaniidae 181, 186, 187*<br />

Ricaniites 187<br />

Ricinoides 443*<br />

Ricinoididae 444<br />

Ricinulei 380, 381, 382, T<br />

383,<br />

384*, 385, 433, 437, 443*,<br />

475, 482, 485, 486<br />

444,<br />

Rimnosentomon 147*<br />

Rithma 103<br />

Robinjohnia 292<br />

Robinjohniidae 292<br />

Rochlingiidae 50<br />

Roomeriidae 124<br />

Rossocoleus 250,<br />

Rostrata 383<br />

251*<br />

Saarlandiidae 50<br />

Sabacon 483<br />

Saginae 155<br />

Saldidae 216*<br />

Saldides 216<br />

Salomoninae 155<br />

Saltatoria 37* 144, 146<br />

Salteropterus 407, 408, 409*<br />

Salticidae 509, 515*<br />

Salticidea 499, 509<br />

Salticiformia 509<br />

Samaroblatta 100*, 103<br />

Samaroblattula 100*, 102<br />

Samarura 82*. 83<br />

Sarbaloptera 173*, 174<br />

Sarbalopterum 127*. 128<br />

Sarbaloscytina 192*<br />

Sarcophagidea 343, 344<br />

Sarcophagii 344<br />

Sarcoptidea 451<br />

Sarcoptiformes 448, 450, 451, 474<br />

Sardycblatta 93*, 94<br />

Scapanea 84*<br />

Scapheremaeus 454<br />

Scaphidiidae 256*<br />

Scaphidium 256*<br />

Scarabaeidae 256*, 257<br />

Scatopse 331*<br />

Scatopsidae 330, 331<br />

Scatopsidea 319, 330, 331<br />

Scatopsites, 331<br />

Scelionidae 353, 354<br />

Schambeloblattina 103<br />

Scheloribates 458*. 460<br />

Scheloribatidae 460<br />

Schistocerca 156*<br />

Schizoblatta 94<br />

Schizoblattina 94<br />

Schizocoleidae 249, 251*<br />

Schizocoteus 250, 251*<br />

Schizocupes 250, 251*<br />

Schizodactylidae 150<br />

Schizomida 436<br />

Schizomidae 436<br />

Schizomus 434*<br />

Schizoneuriden 197<br />

Schizoneurinae 197<br />

Schizopeltidia 383, 384*, 385,<br />

438, 439, 445<br />

Schizotaldycupes 246, 247*<br />

Schizotetranychus 469<br />

Schoutedeniinae 197<br />

Sciadocerldae 341<br />

Sciaraedes 324<br />

Sciaridae 324<br />

Sciomyzidae 342<br />

Sciomyzides 342<br />

Sciophilinae 324<br />

Scirtes 257*<br />

Scleropterinae 154<br />

Scoliidae 356<br />

Scoliidea 349, 356, 357*<br />

Scolopendrellidae 21<br />

Scolopendridae 25, 26<br />

Scolopendromorpha 25<br />

Scopeuma 343*<br />

Scopuridae 134<br />

Scorpio 432<br />

Scorpiones 382, 383, 385, 402, 403,<br />

404, 450<br />

Scorpionida 378, 384*.385, 389, 423,<br />

432*, 433<br />

Scorpionidae 428, 432<br />

Scorpionidea 428<br />

Scorpionomorpha 378, 379, 380, 381,<br />

382, 383, 384*, 385,389, 402, 403,<br />

438, 439, 451, 474, 475, 476, 482,<br />

485, 486<br />

Scraptia 264*<br />

Scraptiidae 264*, 265<br />

Scudderia 495<br />

Scudderulla 94<br />

Scutigera 26*<br />

Scutigerella 21*<br />

Scutigerellidae 21<br />

Scutigeridae 27<br />

Scutigeromorpha 26, 27<br />

Scutinoblattina 103<br />

Scutoribates 457*<br />

Scydmaenidae 255, 256*<br />

Scydmaenus 256*<br />

Scylacocoris 226<br />

Scytinoptera 173*, 174<br />

Scytinopteridae 168, 173*, 175<br />

Scytinopterinae 173*. 174<br />

Scytinopterula 17 A<br />

Scytocixius 185*, 186<br />

Scytohymen 67*. 68<br />

Scytohymenidae 67, 68<br />

Scytolestes 77*, 78<br />

Scytoneura 205*. 206 '<br />

Scytoneureila 205*. 206<br />

Scytophara 182*. 183<br />

Segestria 498, 499*<br />

Segestriidae 498, 499*<br />

Sejus 470*, 473<br />

Selenopidae 507<br />

Selenotemidae 81<br />

Selenothemis 80*, 81<br />

Sellardsiopsis 133*<br />

Sellardsula 94<br />

Semenoviola 114*, 115<br />

Semidalis 282*<br />

Semopterum 128<br />

Senoculidae 505<br />

Senzillidae 134<br />

Sepsidae 342<br />

Sequoja 299*<br />

Sericostomatidae 299,302<br />

Sericostomatinae 302<br />

Serphidae 354<br />

Serphites 353*. 354<br />

Serphitidae 353, 354<br />

Serropalpidae 264*, 265<br />

Setaspis 389, 393<br />

Severinopsis 53<br />

557<br />

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Severinula 53<br />

Sheimia 139*<br />

Sheimiidae 139<br />

Sheimioidea 139<br />

Shelfordella 104*<br />

Shepherdia 254<br />

Shurabella 225*<br />

Shurabia 131*, 132<br />

Shurabocossus 164*, 166<br />

Shuraboprosbole 171*<br />

Sialidae 269<br />

Sialidopseidae 277, 279, 280*<br />

Sialidopsis 280*<br />

Sialium 274<br />

Siberiohymen 65*, 66<br />

Sibirella 127*, 128<br />

Sibiroblatta 91, 92*<br />

Sieblosia 82*<br />

Sieblosiidae 81, 83<br />

Silpha 34*<br />

Silphidae 255, 256*<br />

Silvanus 261*<br />

Silvius 335*<br />

Simmondsia 249<br />

Simoderinae 155<br />

Simuliidae 316, 318<br />

Simulates 318<br />

Simulium 317*<br />

Sindon 116*. 117<br />

Sindonopsis 116*, 117<br />

Sinemedia 326*, 32*<br />

Sinemediidae 328<br />

Singulidae 41<br />

Sinoblatta 110<br />

Sinoperla 136<br />

Siphlonuridae 60, 6/*, 63*<br />

Siphlonuridea 57, 60<br />

Siphlonurinae 61<br />

Siphlonurites 61<br />

Siphlonurus 61*<br />

Siphonophorini 198<br />

Sirex 347*<br />

Siricidae 348<br />

Siricidea 346, 347<br />

Sironidae 482<br />

Sisyra 34*, 281*<br />

Sisyridae 277, 280<br />

Sisyrocoridae 211*. 212<br />

Sisyrocoris 211*, 212<br />

Slimonia 403, 404, 407, 409*, 410<br />

Slimoniidae 407, 408, 409*, 410<br />

Smaridiidae 468<br />

Smaridiidea 463, 468<br />

Smiliinae 177<br />

Sminthurelles 47<br />

Sminthuridae 47<br />

Sminthurus 47*<br />

Sogdoblatta 100*. 102<br />

Sogdopsyche 293*. 294<br />

Sogdothemis 80*, 81<br />

Sogjuta 273*, 274<br />

Sojanocoleidae 242*<br />

Sojanocoleus 242*<br />

Sojanoneura 167*, 168<br />

Sojanopsylla 204, 205*<br />

Sojanoraphidia 270*, 271<br />

Sojanoraphidiidae 270*, 271<br />

Sojanoscytina 192*, 194<br />

Solenoptilidae 273<br />

Solenoptilon 273*<br />

Solifugae 378, 381, 382, 383, 384*.<br />

385, 439, 443*, 444, 445, 446, 526

Solifugomorpha 378, 379, 380 381,<br />

382, 383, 384*, 385, 445, 451,475<br />

Solikamptilonidae 75<br />

Solikamptilon 77*, 78<br />

Soluta 383, 384*, 385, 475, 482,<br />

485, 490<br />

Solvidae 334<br />

Somatocoris 212<br />

Sooblatta 97<br />

Sooblattella 94<br />

Soomylacris 97<br />

Sosybius 507, 512*<br />

Spaniodera 123*<br />

Spanioderidae 122, 124<br />

Spaniodontella 299*<br />

Sparassidae 507<br />

Spathiinae 349<br />

Spatiator 500*, 501<br />

Spatiotoridae 500*, 501<br />

Speleognathidae 461, 465<br />

Speleorchestes 447, 462*<br />

Spermophorella 281*<br />

Sphaerherpestidae 23<br />

Sphaerolichidae 462<br />

Sphaeropsocidae 228, 230 '<br />

Sphaeropsocus 229*<br />

Sphaerozetes 458*, 460<br />

Sphaleroblattina 94<br />

Sphecoptera 65*<br />

Sphecopterinae 66<br />

Sphenomylacris 98<br />

Sphingidae 34*<br />

Sphingodea 307<br />

Spilaptera 53<br />

Spilapteridae 53<br />

Spiloblattina 95<br />

Spiloblattinidae 89, 94<br />

Spiloblattininae 89, 94, 95*<br />

Spondyliaspinae 206<br />

Spongiotarsus 432<br />

Spongoneura 146<br />

Spongophoriella 178*<br />

Spyharopteridae 69<br />

Stanleyana 190<br />

Stantoniella 104<br />

Staphilinidae 254*, 255<br />

Steatoda, 501, 503*<br />

Stedippus 515<br />

Stegaspinae 177<br />

Stegocupes 248*<br />

Stegopteridae 124, 128, 129*<br />

Stegopterum 128, 129*<br />

Stegosyne 252*, 253<br />

Steleopteron 79*, 81<br />

Stenaroceridae 122<br />

Stenaropoda 145*, 146<br />

Stenaropodidae 145*<br />

Stenaropodites 120*, /2/<br />

Steneattus 515*<br />

Stenodicranidae 58<br />

Stenodictyopteridae 50<br />

Stenoglyphis 181<br />

Stenomylacris 97<br />

Stenoneura 116*<br />

Stenoneuridae 117, //4?<br />

Stenoneurillidae 117<br />

Stenopelmatinae 152<br />

Stenophtebia 82*<br />

Stenophlebiidae 81, 53<br />

Stenopsocidium 228<br />

Stenopsyche 296*<br />

Stenopsychidae 295, 295<br />

Stenoscytina 174<br />

Stenotrogulus 487*, 455<br />

Stenovicia 190<br />

Stenoviciidae 189<br />

Stephanidia 350<br />

Stephanidae 349, 350<br />

Stephanoblatta 94<br />

Stephanomylacris 97<br />

Stereopteridae 124<br />

Sternarthron 433, 438*, 439<br />

Sternarthronidae 439<br />

Sternophoridae 442<br />

Sternorrhyncha 163, /90, 207<br />

Sterzelia 94<br />

Stethostomata 383<br />

Stigetoblatta 94<br />

Stigmorista 288*<br />

Stiphroschema 212<br />

Strabopidae 389, 391, 393*<br />

Strabops 389, 393*<br />

Stratiomididae 334, 336<br />

Stratiomididea 333*. 334<br />

Stratiomites 336<br />

Streblomorpha 308*<br />

Strebloblattula 103<br />

Strephocladidae 124<br />

Strephocladidea 119, 123*, /24<br />

Strephoneura 123*, /24<br />

Strephoneuridae /24<br />

Strepsiptera 240*, 241, 267, 268*<br />

Striatotegmen 103<br />

Strieremaeus 454*, 456<br />

Strobilocoris 212<br />

Strobilopterus 407, 411, 413*<br />

Stygneidae 122<br />

Stygophrynus 435*<br />

Stylonuracea 415, 416<br />

Stylonuridae 404, 406, 407, 408,<br />

415, 4/6, 418, 419*, 420, 422<br />

Stylonuridea 408, 415, 4/6, 421<br />

Stylonurina 385, 405, 407, 408,<br />

415, 420<br />

Stylonurinae 408, 4/5<br />

Stylonurus 405*, 406, 408, 416, 4/5<br />

Subilla 272<br />

Suctobelba 456<br />

Sulcoboltonia 94<br />

Suljuctocossus 164*, /65<br />

Suljuktaja 164*, /67<br />

Supellini 104, /05<br />

Surijoka 158*<br />

Surijokocixius 185*, /56<br />

Surijokocypha 229*, 230<br />

Surijokopsocidae 225<br />

Surijokopsocus 228, 229*<br />

Surijokovia 174, 175*<br />

Sushkinia 77*, 75<br />

Syarinidae 441<br />

Sycorax 316*<br />

Sylvaella 129*, /30<br />

Sylvaellidae 130<br />

Sylvaphlebia 129*, /30<br />

Sylvaphlebiidae 124, 129*, /30<br />

Sylvidelia 119, 120*<br />

Sylviodes 129*, /30<br />

Sylviodidae 130<br />

Sylvohymen 67*. 65<br />

Sylvopanorpa 286, 287*<br />

Symmocites 305*<br />

Symphyla 2/*, 36<br />

Symphyoblatta 94<br />

Symphypleona 45, 47*<br />

558<br />

http://jurassic.ru/<br />

Symphyta 346, 347*<br />

Symphytognathidae 501<br />

Symplicius 97<br />

Symploce 105*<br />

Symplocini /04<br />

Synarmogidae 52<br />

Syneoptoblatta 94<br />

Syndesmophylum 152<br />

Syndesmorpha 132<br />

Synneuridae 330<br />

Synomaloptila 142, 143*<br />

Synomaloptilidae /42<br />

Synonemoura 136<br />

Synorthophlebia 291<br />

Synsphyronidae 441<br />

Syntexidae 348<br />

Syntomidae 307<br />

Syntonopteridae 50<br />

Synziphosura 385, 386, 389. 391,<br />

394, 395, 397<br />

Sypharoptera 70*<br />

Sypharopteridae 69<br />

Syphax 513*<br />

Syringophilidae 467<br />

Syrphidae 34/<br />

Syrphidea 34/*<br />

Syrphinae 341<br />

Syscioblatta 95<br />

Sysciophtebla 95*<br />

Systellopthemis 83<br />

Tabanidae 336, 337<br />

Tabanidea 334, 336<br />

Tachina 344<br />

Tachinariae 344<br />

Tachinidae 343, 344<br />

Tachiniscidae 342<br />

Tachypeza 338*<br />

Tachypleus 387*, 388 , 390, 400*.<br />

401<br />

Taeniopterygidae, 117, 134, /36<br />

Taldycupes 248*<br />

Taldycupidae 241, 246, 247*, 248*,<br />

249*<br />

Taldycupidium 248, 249*<br />

Tanyderidae 3/3<br />

Tanyderina 313<br />

Tanyderophryne 314, 315*<br />

Tanyderophyneidae 3/4 .<br />

Tanyderophryneidea 310, 3/4, 315*<br />

Tanypina 318<br />

Tanypodinae 3/5<br />

Tapinellinae 230, 232<br />

Tapopterum 128<br />

Tarantulidae 437<br />

Tarsonemidea, 449, 461, 463<br />

Tarsonemini 380, 449, 462<br />

Tarsophlebia 82*<br />

Tarsophlebiidae 81, 52<br />

Tarsophlebiidea 81, 82*<br />

Tarsopterella 404, 408, 416, 418<br />

Tartarides 436, 451<br />

Taublatta 100*, 102<br />

Tchirkovaea 69, 70*<br />

Tcholmanvissia 148<br />

Tcholmanvissiidae 146, /45*, 150<br />

Tecticupes 248*<br />

Tectocepheus 457*, 458<br />

Tectocymba 454*, 456<br />

Tectoribates 458*, 460<br />

Tegenaria 505<br />

Telaxiidae 198

Telaxiinae 197,^198<br />

Telemidae- 498<br />

Telyphones 385<br />

Temnopteryx 105*<br />

Temnostigma 83<br />

Tendipedidae 317<br />

Tenebrionidae 264*, 265<br />

Teneopteridae 117<br />

Tengellidae 517<br />

Tenthredinidae 348<br />

Tenthredinidea 346, 348<br />

Tenualidae 460<br />

Terebrantia 237<br />

Terentiinae 177<br />

Termitidae 112, 113<br />

Termitidium 152<br />

Termitoides 133*<br />

Termitoxeniomorpha 308*<br />

Termopsis 112*<br />

Terpnacaridae 462<br />

Tersus 259*<br />

Tetanoptilon 276<br />

Tethneus 505<br />

Tetragnatha 505<br />

Tetragnathidae 499, 501, 505<br />

Tetranychidae 461, 469*<br />

Tetranychidea 461, 462, 468<br />

Tetrapneumones- 494, 497<br />

Tetrapodilii. 384*,"385, 450,451, 471<br />

Tetrigidae 156*, 157<br />

Tetrigides 157<br />

Tetrix 156*<br />

Tettigadinae 172<br />

Tettigarctidae 167, 170, 171*<br />

Tettigomotridae 181<br />

Tettigoniidae 155<br />

Tettigoniidea 146, 154<br />

Tettigoniinae 155<br />

Tettiniidae 181<br />

Textrix 505, 510*<br />

Thanatodictya 188*<br />

Thaumaleidea 310<br />

Thaumstoxeniidae 339<br />

Thelaxidae 198<br />

Thelaxinae 197, 198<br />

Thelyphonida 403, 434, 436<br />

Thelyphonidae 436<br />

Thelyphonus 434*, 436<br />

Thelyphrynus 435*, 437<br />

Theraphosidae 498<br />

Theraphosina 498<br />

Them 498<br />

Thereola 498, 499*<br />

Thereva 338*<br />

Therevidae 337 *<br />

Therevites 337<br />

Theridiidae 501, 503*<br />

theridiometa 505, 509*<br />

Theridion 501, 503*<br />

Thesoneuridae 50<br />

Thnetodes 148<br />

Thnetus 150*<br />

Thomisidae 509, 513*<br />

Thomisidea 499, 509<br />

Thomisiformia 509<br />

Thomisius 509<br />

Thoronysidae 122<br />

Thripidae 237, 238<br />

Throscidae 259*<br />

Throscus 259*<br />

Thuringoblatta 103<br />

Thyelia 505, 510*<br />

Thylacelia 232*<br />

Thylacinae 231<br />

Thyreocoridae 224<br />

Thysanoptera 162*, 236, 237*<br />

Thysanopteroidea 161, 236<br />

Thysanura 36, 37*, 45, 47*. 48<br />

Tibicininae 172<br />

Tillyardagrion 78<br />

Tillyardembia 123*. 124<br />

Tillyardembiidae 122, 124<br />

Tillyardiella 148<br />

Tinae 304<br />

Tineidae 304<br />

Tineidea 304<br />

Tineomorphinae 231<br />

Tingidae 215, 216*<br />

Tingidites 215<br />

Tingiopsis 215, 216*<br />

Tipula 312*<br />

Tipulidae 313<br />

Tipulidea 310, 312*<br />

Tipulodictya 311*<br />

Tipulodictyidae 310<br />

Tipulodictyidea 310, 311*<br />

Tipulomorpha 308*, 310<br />

Tipuloplecia 326*, 328<br />

Tipulopleciidae 328<br />

Titius 432<br />

Tiviinae 110<br />

Tolaniinae 177<br />

Tomacblatta 93*, 94<br />

Tomapsididae 181<br />

Tomia 131*<br />

Tomiidae 124, 131*<br />

Tomioblatta 89<br />

Tomiochorista 285*<br />

Tomiochoristidae 284, 285*<br />

Tomiocupes 244*, 246<br />

Tomiopsyllidium 199, 200*<br />

Tomioscarta 190, 191*<br />

Tomiulus 22*, 23<br />

Tomoceridae 46, 47<br />

Tomocerinae 47<br />

Tomocerus 46*<br />

Topinae 352<br />

Tortrices 306<br />

Tortricidae 305, 306<br />

Tortriciidea 304<br />

Torymidae 352, 353<br />

Tracheata 19, 20, 379, 382<br />

Trachycoris 212<br />

Transversiplecia 320, 321*<br />

Trepidariini 349<br />

Trhypochtoniidae 454<br />

Trhypochthonius 452*, 454<br />

Triadotypidae 73, 74<br />

Triadotypus 73*<br />

Triaenonychidae 480<br />

Triaspinae 349<br />

Triassagrion 78<br />

Triassoaphis 199<br />

Triassoblatta 103<br />

Triassocixius 186<br />

Triassocoridae 212*, 213<br />

Triassecoris 212*, 213<br />

TriassocSHe*J£6<br />

T riasso jassusuf<br />

Triassolestes 83<br />

Triassolocusta 156<br />

Triassomachilidae 48<br />

Triassomachilis 47*, 48<br />

Triassomanteidae 122<br />

559<br />

http://jurassic.ru/<br />

Triassophlebia 83<br />

Triassopsyche 294<br />

Triassopsychops 278<br />

Triassopsylla 194<br />

Triassoscarta 190<br />

Triassoscelis 174<br />

Triassoscytina 174<br />

T riassoscytinopsis 174<br />

Triassothea 200<br />

Trichadenidea 461, 463, 471<br />

Trichempheria 232*<br />

Trichocera 312*<br />

Trichoceridae 313<br />

Trichodes 259*<br />

Trichomyia 316*<br />

Trichomyiidae 316<br />

Trichomyiinae 316<br />

Trichoptera 240*, 283, 284, 294<br />

Tricorythidae 60<br />

Tricupes 244*, 246<br />

Tricyphona 312*<br />

Tridactilidae 155<br />

Trifidella 181<br />

Trigonalidae 349, 350<br />

Trigonidiinae 154<br />

Trigonomarti 490<br />

Trigonomartidae 486, 488, 489*<br />

Trigonomartus 488, 489*<br />

Trigonoscorpio 427*, 431*<br />

Trigonoscorpionidae 431<br />

Trigonotarbi 383, 384,* 385, 444,<br />

474, 475, 485, 486, 490, 496<br />

Trigonotarbidae 486, 488, 489*<br />

Trigonotarbus 488, 489*<br />

Trilobitomorpha 379, 380, 382,<br />

389, 526<br />

Trilophomylacris 97<br />

Triophtydeus 464<br />

Trioza 207*<br />

Triozinae 206, 207<br />

Triphyllus 261*<br />

Triplectides 301*<br />

Triplectidinae 300<br />

Triplosoba 55*<br />

Triplosobidae 56<br />

Tripsyllidium 207<br />

Trirhabdeloblatta 103<br />

Triscytina 225<br />

Tritoma 261*<br />

Triiophania 187*<br />

Troctidae 231<br />

Trogulidae 483<br />

Trogulidea 483<br />

Trombeae 468<br />

Trombiculidae 463, 468<br />

Trombidiformes 384*, 385, 448,<br />

449, 450, 451, 461, 463<br />

Trombidii 450<br />

Trombidiidae 451, 468, 469*<br />

Trombidiidea 463, 468<br />

Trombidiiformes 450, 451<br />

Trombidium 468<br />

Tropiduchidae 181<br />

Tryoniopsis 254<br />

Trypaneidae 342<br />

Trypetidae 342<br />

Tryphoninae 352<br />

Tschekardaella 195*<br />

Tschekardocoleidae 241, 242*<br />

Tschekardocoleus 242*<br />

Tschekardohymen 67*. 68<br />

Tshorkuphlebia 151*, 153

Tshorkuphlebiidae 150, 151*, 153<br />

Tubulifera 237, 238<br />

Tuqouseorpionidae 428<br />

Turfanelta 63*, 64<br />

Turanopsyche 293*, 294<br />

Turutanovis 171*<br />

Tychticola 190, 191*<br />

Tychticupes 248, 249*<br />

Tychticupoides 249*<br />

Tychtobius 279, 280*<br />

Tychtocoleus 252*, 253<br />

Tychtodelopterum 140, 141*<br />

Tychtopsyche 290*<br />

Tychtopsychidae 286, 290*<br />

Tychtoscarta 190, 191*<br />

Tychtoscytina 173*, 174<br />

Tydeidae 449, 464*, 465<br />

Tydeidea 461, 463, 465<br />

Tydeus 464*<br />

Tylidae 342<br />

Tylopterella 404, 407, 410, 412*<br />

Tylopterus 410<br />

Tymocicada 172*<br />

Tympanophoridae 155, 186<br />

Tympanophorinae 155<br />

Typhlopisthacanthus 431*<br />

Typhloscorpius 427*. 431<br />

Typinae 74<br />

Typus 74<br />

Tyroglyphidae 452<br />

Tyroglyphites 452<br />

Tyroglyphus 452<br />

Uarthrus 389, 391*, 393<br />

Ulmeriella 112*<br />

Uloboridae 515<br />

Ulomite^ 254<br />

Ungoneurites 93*, 94<br />

Uninervidae 23*<br />

Uninervus 238, 239*<br />

Unionopterus 423<br />

Uralelytron 115*. //6<br />

Uraloblatta 90*, 9/<br />

£/ ralocoleus 243<br />

Uralonympha 137*, /3*<br />

Uraloscytina 191, 192*<br />

Uraloscytinidae 191<br />

Uralotermes 133*<br />

Urocteidae 500*. 50/<br />

Urogomphus 85<br />

Uropodidea 472<br />

Uropygi 380, 381, 382, 383, 384*.<br />

385, 402, 403, 433, 434*, 435,<br />

436, 437, 438, 439, 440, 444, 445,<br />

475<br />

Urotricha 436<br />

Uskatelytridae 157, 159*<br />

Uskatelytrum 159*<br />

Uskaticupes 249*<br />

Uskatocoleus 251*. 252<br />

Variolina 203<br />

Vejovidae 428<br />

Velio 220*, 226<br />

Veliidae 219, 220*<br />

Vespidea 349<br />

Vilvia 143*<br />

Vilviopsis 143», 144<br />

Vitreacixius 185*, 186<br />

Vitriala 229*. 230<br />

Vorkutia 65*, 66<br />

Vorkutiidae 64, 66<br />

Vratislavia 488<br />

Weinbergina 389, 394, 395, 396*,<br />

397<br />

Weinberginidae 389, 395, 396*, 397<br />

Woodwardopteridae 408, 422<br />

Woodwardopterus 408, 421*, 422<br />

http://jurassic.ru/<br />

Xenillus 457*, 458<br />

Xenobtatta 94<br />

Xenochoristidae 286<br />

Xenogryllacris 132<br />

Xenopanorpa 286<br />

Xenopteridae 122<br />

Xenusion 534*<br />

Xiphydriidae 348, 350, 401<br />

Xyelidae 346<br />

Xyleutites 305*<br />

Xylobius 23<br />

Xylophagei 336<br />

Xylophagidae 334, 336<br />

Xyphosura 383, 389, 406<br />

Yezabura 198*<br />

Zacallitidae 80<br />

Zachria 507, 512*<br />

Zalmona 151<br />

Zalmonites 152<br />

Zavjaloblatta 95*<br />

Zelmira 324*<br />

Zeorchestidae 456<br />

Zeuneroptera 151*, 152<br />

Zilla 505, 509*<br />

7odariidae 500*, 501<br />

Zodariidea 499, 500*<br />

Zodariiform a 500<br />

Zoraptera 162*, 226<br />

Zoropseidea 5/5, 516*<br />

Zoropsidae 5/5, 516*<br />

Zoropsidiformia 515<br />

Zoropsocidae 228<br />

Zoropsocus 229*, 230<br />

Zygomyia 324*<br />

Zygopsocidae 228, 230<br />

Zygopsocus 229*<br />

Zygoptera 73, 76, 77*, 79*

ОСНОВЫ ПАЛЕОНТОЛОГИИ<br />

Спр<strong>а</strong><strong>в</strong><strong>о</strong>чни<strong>к</strong> для п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong> и ге<strong>о</strong>л<strong>о</strong>г<strong>о</strong><strong>в</strong> СССР<br />

Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие—тр<strong>а</strong>хейные, хелицер<strong>о</strong><strong>в</strong>ые<br />

Ут<strong>в</strong>ержден<strong>о</strong> <strong>к</strong> печ<strong>а</strong>ти<br />

П<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гиче<strong>с</strong><strong>к</strong>им ин<strong>с</strong>титут<strong>о</strong>м<br />

А<strong>к</strong><strong>а</strong>демии н<strong>а</strong>у<strong>к</strong> СССР<br />

Ред<strong>а</strong><strong>к</strong>т<strong>о</strong>р изд<strong>а</strong>тель<strong>с</strong>т<strong>в</strong><strong>а</strong> Л. С. О<strong>с</strong>ип<strong>о</strong><strong>в</strong><strong>а</strong><br />

Техниче<strong>с</strong><strong>к</strong>ий ред<strong>а</strong><strong>к</strong>т<strong>о</strong>р Е. В. М<strong>а</strong><strong>к</strong>уни<br />

К<strong>о</strong>рре<strong>к</strong>т<strong>о</strong>р А. С. См<strong>о</strong>гиле<strong>в</strong><strong>а</strong><br />

РИСО АН СССР № 93-56В. Сд<strong>а</strong>н<strong>о</strong> <strong>в</strong> н<strong>а</strong>б<strong>о</strong>р 15/111 1962 г.<br />

П<strong>о</strong>дпи<strong>с</strong><strong>а</strong>н<strong>о</strong> <strong>к</strong> печ<strong>а</strong>ти 5/XI 1962 г. Ф<strong>о</strong>рм<strong>а</strong>т 84 х ЮЭ 1<br />

/^.<br />

Печ. л. 35+22 <strong>в</strong><strong>к</strong>л. Уч.-изд. л. 62,9 + 2,6 <strong>в</strong><strong>к</strong>л.=65,5<br />

Тир<strong>а</strong>ж 3000 э<strong>к</strong>з. Т-12749. Изд. № 5091. Тип. з<strong>а</strong><strong>к</strong>. № 580<br />

Цен<strong>а</strong> 4 р. 92 <strong>к</strong>.<br />

Изд<strong>а</strong>тель<strong>с</strong>т<strong>в</strong><strong>о</strong> А<strong>к</strong><strong>а</strong>демии н<strong>а</strong>у<strong>к</strong> СССР.<br />

М<strong>о</strong><strong>с</strong><strong>к</strong><strong>в</strong><strong>а</strong>, Б-62, П<strong>о</strong>д<strong>с</strong><strong>о</strong><strong>с</strong>ен<strong>с</strong><strong>к</strong>ий пер., 21<br />

2-я тип<strong>о</strong>гр<strong>а</strong>фия Изд<strong>а</strong>тель<strong>с</strong>т<strong>в</strong><strong>а</strong> АН СССР.<br />

М<strong>о</strong><strong>с</strong><strong>к</strong><strong>в</strong><strong>а</strong>, Г-99, Шубин<strong>с</strong><strong>к</strong>ий пер., 10<br />

http://jurassic.ru/

ОПЕЧАТКИ И ИСПРАВЛЕНИЯ<br />

Стр<strong>а</strong>ниц<strong>а</strong> Стр<strong>о</strong><strong>к</strong><strong>а</strong> Н<strong>а</strong>печ<strong>а</strong>т<strong>а</strong>н<strong>о</strong> Д<strong>о</strong>лжн<strong>о</strong> быть<br />

89 9 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

91 17 <strong>с</strong><strong>в</strong>. <strong>с</strong>ле<strong>в</strong><strong>а</strong> Оренбург<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл. Перм<strong>с</strong><strong>к</strong><strong>а</strong>я <strong>о</strong>бл.<br />

153 24 <strong>с</strong><strong>в</strong>. <strong>с</strong>ле<strong>в</strong><strong>а</strong> Rhaphidiphoridae Rhaphidophorinae<br />

171 13 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> Wife White<br />

187<br />

П<strong>о</strong>дпи<strong>с</strong>ь п<strong>о</strong>д ри<strong>с</strong>.,<br />

2 <strong>с</strong><strong>в</strong>. <strong>с</strong>ле<strong>в</strong><strong>а</strong><br />

1 531<br />

198 7 <strong>с</strong>н. <strong>с</strong>ле<strong>в</strong><strong>а</strong> Alzenberg Aizenberg<br />

206' 14 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> 1—4 мм 1—4 мм (ри<strong>с</strong>. 604)<br />

207 11 <strong>с</strong>н. <strong>с</strong>ле<strong>в</strong><strong>а</strong> Psylla nigrita Psylla nigritopsis<br />

207 3 <strong>с</strong>н. <strong>с</strong>ле<strong>в</strong><strong>а</strong> (ри<strong>с</strong>. 604—606) (ри<strong>с</strong>. 606)<br />

219 20 <strong>с</strong><strong>в</strong>. <strong>с</strong>ле<strong>в</strong><strong>а</strong> р<strong>а</strong>зн<strong>о</strong>мерн<strong>о</strong>й р<strong>а</strong><strong>в</strong>н<strong>о</strong>мерн<strong>о</strong>й<br />

242 2 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> С Си<br />

266 30 <strong>с</strong><strong>в</strong>. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> Ср. юр<strong>а</strong> Н. юр<strong>а</strong><br />

316 16 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> CuA А<br />

329 П<strong>о</strong>дпи<strong>с</strong>ь п<strong>о</strong>д ри<strong>с</strong>.,<br />

2 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong><br />

1964 1064<br />

329 2 <strong>с</strong><strong>в</strong>. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> <strong>к</strong>реп<strong>к</strong>ие <strong>в</strong>ет<strong>в</strong>и и <strong>к</strong>реп<strong>к</strong>ие и<br />

332 9 <strong>с</strong><strong>в</strong>. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> Три Тип<br />

333 4 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> 1908 1902<br />

341 4 <strong>с</strong>н. <strong>с</strong>ле<strong>в</strong><strong>а</strong> Platypezidea Syrphidea<br />

354 18 <strong>с</strong><strong>в</strong>. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> из п<strong>а</strong>ле<strong>о</strong>ген<strong>а</strong> из <strong>в</strong>. мел<strong>а</strong><br />

357 4 <strong>с</strong>н. <strong>с</strong>пр<strong>а</strong><strong>в</strong><strong>а</strong> Myrmicidae Poneridae<br />

422<br />

27 <strong>с</strong><strong>в</strong>. <strong>с</strong>ле<strong>в</strong><strong>а</strong> Mycteropidae<br />

St0rmer, 1951<br />

Mycteropidae С<strong>о</strong>ре, 1886<br />

(= Mycteropidae<br />

St0rmer, 1951)<br />

450 22 <strong>с</strong>низу Acaridiformes Acariformes<br />

О<strong>с</strong>н<strong>о</strong><strong>в</strong>ы п<strong>а</strong>ле<strong>о</strong>нт<strong>о</strong>л<strong>о</strong>гии. Члени<strong>с</strong>т<strong>о</strong>н<strong>о</strong>гие.<br />

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