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Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

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Ecology There is a Russian record of association <strong>with</strong> the<br />

fungus Gyromitra esculenta. The larvae may develop in<br />

other soft fungi in the Scottish sites. Adults recorded in<br />

June.<br />

Status Only two males have been found in Britain, one of<br />

them recently but it may have been overlooked elsewhere<br />

<strong>with</strong>in this range. In Europe it is a boreo-alpine species<br />

known from northern Sc<strong>and</strong>inavia <strong>and</strong> the Alps, but also<br />

extends across Siberia to Alaska <strong>and</strong> Canada. Not listed in<br />

Shirt (1987) or <strong>Falk</strong> (<strong>1991</strong>).<br />

Threats Clearance of native forests for intensive forestry or<br />

agriculture. Overgrazing <strong>by</strong> deer preventing natural<br />

regeneration. Removal of old trees or dead wood, which<br />

may support host fungi.<br />

Management <strong>and</strong> conservation Maintain forest cover,<br />

retaining any old trees <strong>and</strong> dead wood <strong>and</strong> ensure<br />

succession of such habitats <strong>by</strong> restricting grazing <strong>and</strong> other<br />

damaging activities.<br />

Published sources Ch<strong>and</strong>ler (1992b).<br />

PSEUDEXECHIA PARALLELA<br />

A fungus gnat LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family MYCETOPHILIDAE<br />

Pseudexechia parallela (Edwards, 1925)<br />

Identification Keyed <strong>by</strong> Edwards (1925) <strong>and</strong> Ch<strong>and</strong>ler<br />

(1978d). The male genitalia were figured <strong>by</strong> Stackelberg<br />

(1948).<br />

Distribution This was known as British only from the<br />

female holotype found at Newmarket, Suffolk (23<br />

September 1888, G.H. Verrall) until it was found to be<br />

frequent in wetl<strong>and</strong> sites surveyed <strong>by</strong> the NCC from 1987 to<br />

1989 in East Anglia (Suffolk, Norfolk, Cambridgeshire) <strong>and</strong><br />

Wales (Glamorgan, Pembrokeshire, Merionethshire,<br />

Caernarvonshire, Denbighshire, Anglesey). It was also<br />

found in Irel<strong>and</strong> in 1983 (Ch<strong>and</strong>ler et al. 2000).<br />

Habitat Marshes <strong>and</strong> fens, wet heaths <strong>and</strong> bogs.<br />

Ecology Biology unknown. Other species of the genus<br />

develop in soft gill fungi, including both terrestrial <strong>and</strong><br />

lignicolous species. Adults recorded from June to October.<br />

Status This was unrecorded in Britain for almost a century<br />

after its original discovery but there are now records for<br />

three sites in Suffolk, 21 sites in Norfolk, Chippenham Fen<br />

NNR in Cambridgeshire <strong>and</strong> fourteen sites in Wales. Status<br />

revised from RDB 1 (Shirt 1987) <strong>and</strong> from Extinct (<strong>Falk</strong><br />

<strong>1991</strong>).<br />

Threats Drainage of wetl<strong>and</strong>s. Overgrazing or nonrotational<br />

cutting on some sites.<br />

Management <strong>and</strong> conservation Maintain stable water<br />

level <strong>and</strong> existing vegetation structure <strong>and</strong> mosaic of<br />

habitats. Grazing should be at a low level <strong>and</strong> where<br />

appropriate grazing or cutting should be carried out on<br />

rotation. Removal of the litter layer on which host fungi<br />

may develop should be avoided.<br />

Published sources Ch<strong>and</strong>ler et al. (2000); Edwards (1925).<br />

PSEUDORYMOSIA FOVEA<br />

A fungus gnat LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family MYCETOPHILIDAE<br />

Pseudorymosia fovea (Dziedzicki, 1910)<br />

Identification The male genitalia were figured <strong>by</strong><br />

Dziedzicki (1910).<br />

Distribution A northern <strong>and</strong> western species: Forest of<br />

Dean, Gloucestershire; several sites in Yorkshire; Helvellyn<br />

Gill, Cumberl<strong>and</strong>; Glamorgan, Denbighshire, Flintshire;<br />

several sites in Perthshire; the Mar Lodge Estate in<br />

Aberdeenshire; the Spey Valley in Elgin <strong>and</strong> Easterness;<br />

other sites in Easterness, East Ross <strong>and</strong> East Sutherl<strong>and</strong>.<br />

Habitat Damp woodl<strong>and</strong>, mostly broad-leaved but mixed at<br />

some Scottish sites.<br />

Ecology This has been reared in Germany from a terrestrial<br />

agaric, Tricholoma flavobrunneum. Adults recorded in June<br />

to July, September <strong>and</strong> October.<br />

Status A distinctive species but poorly known before 1980<br />

as of 30 recorded sites, 27 are more recent. It seems<br />

particularly widespread in the Scottish Highl<strong>and</strong>s <strong>and</strong><br />

Ch<strong>and</strong>ler (1987a) also recorded it from Irel<strong>and</strong>. Status<br />

revised from RDB 1 (Shirt 1987) <strong>and</strong> from RDB 3 (<strong>Falk</strong><br />

<strong>1991</strong>).<br />

Threats Clearance of native woodl<strong>and</strong> for intensive forestry<br />

or agriculture. Removal of old trees or damage to soil<br />

structure which could affect survival of mycorrhizal fungi.<br />

Management <strong>and</strong> conservation Maintain woodl<strong>and</strong>s in a<br />

natural state, <strong>with</strong> all successional stages, ensuring its<br />

continuity in the future. Avoid damage to litter layer or soil<br />

structure to safeguard terrestrial fungi.<br />

Published sources Ch<strong>and</strong>ler (1987a); Edwards (1923,<br />

1925); Kidd & Ackl<strong>and</strong> (1970).<br />

RYMOSIA ACTA<br />

A fungus gnat<br />

Order DIPTERA<br />

Rymosia acta Dziedzicki, 1910<br />

DATA DEFICIENT<br />

Family MYCETOPHILIDAE<br />

Identification Ch<strong>and</strong>ler (1994b) keyed this species <strong>and</strong><br />

figured the male genitalia, which had also been figured <strong>by</strong><br />

Laštovka & Matile (1974).<br />

Distribution Only known as British from single males<br />

found at six sites in Scotl<strong>and</strong>: Kilvrecht (1987) <strong>and</strong> Allt nan<br />

Bogair (2000), Rannoch, Perthshire; Luibeg, Mar Lodge<br />

Estate, Aberdeenshire (2000); Dulsie, Elgin (1999);<br />

Craigellachie NNR, Easterness (2002); Migdale Wood, East<br />

Sutherl<strong>and</strong> (1995).<br />

Habitat The known localities are broad-leaved or mixed<br />

woodl<strong>and</strong>.<br />

Ecology Biology unknown. The larvae probably develop in<br />

soft fungi. Adults recorded in June <strong>and</strong> August to October.<br />

Status A recent discovery in Britain which is widespread in<br />

northern Europe <strong>and</strong> may prove more widespread in<br />

105

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