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Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

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(1977); Monk Soham, Suffolk (frequent in orchard, June to<br />

August 1917); Kirtling (1896, 1897), Whittlesford (1904),<br />

Chippenham Fen NNR (1943) <strong>and</strong> Cambridge (frequent in<br />

garden, 1901 to 1906), Cambridgeshire; Blaise Woods<br />

(1952) <strong>and</strong> Newent (1979), Gloucestershire; Stoke Edith,<br />

Herefordshire (in garden, 1912).<br />

Habitat Associations are unclear; records include fens,<br />

woods, gardens <strong>and</strong> orchards. The presence of trees of the<br />

family Rosaceae seems to be the over riding factor.<br />

Ecology Biology unknown, but the larvae probably develop<br />

in bracket fungi such as Phellinus pomaceus which grows<br />

on old trees of the family Rosaceae. Adults recorded from<br />

April to October <strong>and</strong> males have been observed swarming<br />

about 0.6m above grass in an orchard (Morley 1918).<br />

Status Apparently formerly widespread in the south, but<br />

<strong>with</strong> only two post 1960 records. It is <strong>not</strong> known whether it<br />

may still occur in garden or orchard situations but is being<br />

overlooked due to insufficient recording.<br />

Threats The destruction of sites containing old rosaceous<br />

trees; the loss of old orchards (which has increased in recent<br />

years) or the removal of old trees from gardens could both<br />

be significant as threats. The shading out of rides <strong>and</strong><br />

clearings in woods which may be required for adult<br />

swarming.<br />

Management <strong>and</strong> conservation Retain any old or diseased<br />

rosaceous trees. Maintain open rides <strong>and</strong> clearings <strong>with</strong>in<br />

woods.<br />

Published sources Aston (1957); Collin (1904, 1938);<br />

Jenkinson (1903); Morley (1918); Verrall (1901).<br />

AGATHOMYIA ELEGANTULA<br />

A flat-footed fly LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family PLATYPEZIDAE<br />

Agathomyia elegantula (Fallén, 1815)<br />

Identification Keyed <strong>by</strong> Ch<strong>and</strong>ler (2001b).<br />

Distribution Scattered localities, mainly in southern<br />

Engl<strong>and</strong> (Hampshire, Surrey, Hertfordshire, Middlesex,<br />

Berkshire, Oxfordshire, Buckinghamshire, Suffolk, Norfolk,<br />

Cambridgeshire, Herefordshire, Warwickshire) but also one<br />

Welsh record (Monmouthshire).<br />

Habitat Usually found in old broad-leaved woodl<strong>and</strong> <strong>with</strong> a<br />

good amount of dead wood present.<br />

Ecology Biology unknown; related species develop in tough<br />

bracket fungi <strong>and</strong> most are fairly specific in their choice of<br />

host. Adults recorded from May to October.<br />

Status There are 22 known post 1960 sites. This species is<br />

usually found in small numbers <strong>and</strong> is probably underrecorded.<br />

The concept of A. elegantula recognised here<br />

represents both sexes of Agathomyia boreella (Zetterstedt),<br />

but females only referred to A. elegantula <strong>by</strong> Ch<strong>and</strong>ler<br />

(1974); the males placed under A. elegantula in that paper<br />

are now referred to A. woodella Ch<strong>and</strong>ler.<br />

Threats The loss of woodl<strong>and</strong> to agriculture or intensive<br />

forestry, <strong>and</strong> the removal of dead wood <strong>and</strong> old or diseased<br />

trees which may support suitable fungi. Also the shading<br />

out of rides or clearings which may be required for adult<br />

swarming.<br />

Management <strong>and</strong> conservation Retain any old or diseased<br />

trees <strong>and</strong> dead wood <strong>with</strong>in a site together <strong>with</strong> their<br />

associated fungi, ensuring the continuity of these habitats in<br />

the future. Maintain open rides <strong>and</strong> clearings.<br />

Published sources Ch<strong>and</strong>ler (1974, 1995); Miles (1985);<br />

Perry & Langton (2000); Wood (1905).<br />

AGATHOMYIA FALLENI<br />

A flat-footed fly LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family PLATYPEZIDAE<br />

Agathomyia falleni (Zetterstedt, 1819)<br />

Identification Keyed <strong>by</strong> Ch<strong>and</strong>ler (2001b).<br />

Distribution Scattered localities in south-east Engl<strong>and</strong>:<br />

Everleigh Ashes, Wiltshire (2003); Harry’s Wood,<br />

Ashdown Forest (1976) <strong>and</strong> Marley & Blackdown Common<br />

(1989), Sussex; Knole Park (1966, 1967, 1972, 1977), Pond<br />

Wood, Chislehurst (1967), Walk Wood, Chislehurst (1972,<br />

1973), Scadbury Park (1983, 1984), Horniman Nature Trail<br />

<strong>and</strong> Sydenham Hill Wood (1987), Maryon-Wilson Park,<br />

Charlton (1980, 1981), Aylesford Old Pit near Maidstone<br />

(1992), Kiln Wood, Lenham (1994) <strong>and</strong> Tudely Wood<br />

RSPB Reserve (2004), Kent; Box Hill (1952), Mickleham<br />

(1956), RHS Garden, Wisley (1999), Wisley Common<br />

(1971), Chobham Common (1976), <strong>and</strong> West End<br />

Common, Esher (2001) Surrey; Buckingham Palace<br />

Garden, Middlesex (1996, 1998); Highst<strong>and</strong>ing Hill,<br />

Windsor Forest (1988), California Country Park (1997),<br />

S<strong>and</strong>ford Copse, Dinton Pastures Country Park (1998) <strong>and</strong><br />

Kennet & Avon Canal, Reading (2003), Berkshire;<br />

Burnham Beeches NNR, Buckinghamshire (1995);<br />

Ickworth, Suffolk (2004); Wayl<strong>and</strong> Wood, Norfolk (2004);<br />

W<strong>and</strong>lebury, Cambridgeshire (2004).<br />

Habitat Sites include broad-leaved woodl<strong>and</strong>s <strong>and</strong><br />

heathl<strong>and</strong> <strong>with</strong> old trees.<br />

Ecology The larvae develop in the bracket fungus<br />

Bjerk<strong>and</strong>era adusta (a common species growing on a range<br />

of mainly broad-leaved trees). Adults recorded from<br />

September to November <strong>and</strong> may be found running about or<br />

feeding on surface deposits on broad leaves such as<br />

Sycamore (Acer pseudoplatanus). Male swarms were<br />

observed at the Horniman Nature Trail (Godfrey 1989);<br />

swarming took place at a height of 3-4m in a small<br />

woodl<strong>and</strong> clearing.<br />

Status Of the 28 sites listed above, 24 are post 1960<br />

records. Status revised from RDB 2 (Shirt 1987). Until<br />

recently it appeared to be restricted to the eastern counties,<br />

<strong>with</strong> no records north of the Thames valley. However, there<br />

have been recent extensions to its range to Wiltshire (2003)<br />

<strong>and</strong> East Anglia (2004), which may suggest a population<br />

increase. It is particularly well established in West Kent<br />

(especially that part now included in Greater London) <strong>and</strong><br />

has no doubt been overlooked in many other woods in this<br />

region because of its late flight period. The common host<br />

fungus is shared <strong>by</strong> Agathomyia unicolor (Zetterstedt),<br />

which although a less conspicuous species, <strong>with</strong> the same<br />

flight period, is nevertheless known to occur frequently<br />

129

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