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Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

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CEPHALOSPHAERA GERMANICA<br />

A big-headed fly LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family PIPUNCULIDAE<br />

Cephalosphaera germanica Aczél, 1940<br />

Identification Keyed <strong>by</strong> Coe (1966) <strong>and</strong> De Meyer (1989).<br />

Distribution Widely distributed in Engl<strong>and</strong> <strong>and</strong> also present<br />

in South Wales: Cornwall, Devon, Somerset, Wiltshire,<br />

Hampshire, Sussex, Kent, Surrey, Berkshire, Oxfordshire,<br />

Suffolk, Cambridgeshire, Gloucestershire, Herefordshire,<br />

Worcestershire, Westmorl<strong>and</strong> <strong>and</strong> Glamorgan.<br />

Habitat Broad-leaved woodl<strong>and</strong>. Most recent records are<br />

from ancient forest areas but the Cornish record was from<br />

s<strong>and</strong> dunes <strong>and</strong> the Caharton Bay site in Devon is a cliff<br />

<strong>with</strong> scrub backed <strong>by</strong> woodl<strong>and</strong>. The Welsh record was<br />

from scrub at the edge of an Oak (Quercus) wood.<br />

Ecology Biology unknown. Probably as in Cephalops a<br />

parasitoid of leaf hoppers of the family Delphacidae. Adults<br />

recorded from May to early August.<br />

Status This was recorded from fourteen counties <strong>by</strong> Coe<br />

(1966) but there have been comparatively few recent<br />

records, although three further counties have been added.<br />

Ten post 1960 records have been confirmed: Penhale S<strong>and</strong>s,<br />

Cornwall (2002); Charlton Bay, Devon (2003); Vallis Vale<br />

<strong>and</strong> Goblin Combe, Somerset (1985); Savernake Forest,<br />

Wiltshire (1990); Alice Holt Forest, Hampshire (1970);<br />

Orlestone Forest, Kent (1980); Staffhurst Wood, Surrey<br />

(1972); King’s Forest, Suffolk (1995); Pwlldu Bay,<br />

Glamorgan (1994). Not listed in Shirt (1987) or <strong>Falk</strong> (<strong>1991</strong>).<br />

Threats Unclear, other than clearance of native woodl<strong>and</strong><br />

for agriculture or commercial forestry.<br />

Management <strong>and</strong> conservation Maintain woodl<strong>and</strong>s in<br />

natural state <strong>with</strong> all successional stages including rides <strong>and</strong><br />

clearings <strong>with</strong> mosaic of herbaceous vegetation.<br />

Published sources Ch<strong>and</strong>ler (1969); Coe (1966).<br />

CLARAEOLA HALTERATA<br />

A big-headed fly LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family PIPUNCULIDAE<br />

Claraeola halterata (Meigen, 1838)<br />

Identification Keyed <strong>by</strong> Coe (1966) <strong>and</strong> von der Dunk<br />

(1997) as Eudorylas halteratus <strong>and</strong> <strong>by</strong> Ackl<strong>and</strong> (2002). This<br />

species was transferred to Claraeola <strong>by</strong> Skevington &<br />

Yeates (2001).<br />

Distribution Records scattered widely in southern Engl<strong>and</strong>,<br />

north to Lancashire <strong>and</strong> one record from South Wales:<br />

Somerset, Wiltshire, Hampshire, Kent, Berkshire,<br />

Oxfordshire, Suffolk, Cambridgeshire, Bedfordshire,<br />

Lancashire; Glamorgan.<br />

Habitat Most records relate to grassl<strong>and</strong> in chalk or<br />

limestone areas, <strong>with</strong> a preference for species-rich short<br />

turf.<br />

Ecology Biology unknown; members of the tribe Eudorylini<br />

are parasitoids of leaf hoppers of the family Cicadellidae as<br />

larvae. Adults recorded from late May to early July, but<br />

mostly in June so a rather short flight period is evident.<br />

Status There are fifteen known post 1970 sites <strong>and</strong> it is may<br />

be more widespread in calcareous areas than present records<br />

indicate. There is a long history at Aston Rowant NNR<br />

(records in 1934, 1968 <strong>and</strong> 1990) but no more recent record.<br />

In Cambridgeshire it has apparently disappeared from<br />

Devil’s Ditch, where it occurred in 1988 <strong>and</strong> 1993 but it<br />

was found at a new site, Fleam Dyke, in 2004 (I. Perry pers.<br />

comm.). Not listed in Shirt (1987) <strong>and</strong> status revised from<br />

Notable in <strong>Falk</strong> <strong>1991</strong>, where it was included as Eudorylas<br />

halteratus.<br />

Threats Habitat loss to agriculture or afforestation;<br />

especially changes in grazing management of chalk or<br />

limestone grassl<strong>and</strong> or scrub encroachment due to a<br />

reduction in grazing. Overgrazing may be a problem at<br />

some sites including Aston Rowant NNR. One of the<br />

Hampshire sites, a SSSI adjoining Stockbridge Down,<br />

where this species was recorded in 1970 subsequently<br />

suffered scrub encroachment <strong>and</strong> was manually cleared to<br />

restore it in 1992, when there was a temporary loss of<br />

vegetation.<br />

Management <strong>and</strong> conservation Maintain a mosaic of<br />

vegetation types <strong>and</strong> existing grazing patterns on grassl<strong>and</strong><br />

sites, employing rotational grazing where necessary.<br />

Published sources Coe (1966); Skevington & Yeates<br />

(2001); Verrall (1901).<br />

CLARAEOLA MELANOSTOLA<br />

A big-headed fly LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family PIPUNCULIDAE<br />

Claraeola melanostola (Becker, 1898)<br />

Identification Keyed <strong>by</strong> Coe (1966) <strong>and</strong> von der Dunk<br />

(1997) as Eudorylas melanostolus <strong>and</strong> <strong>by</strong> Ackl<strong>and</strong> (2002).<br />

This species was transferred to Claraeola <strong>by</strong> Skevington &<br />

Yeates (2001)<br />

Distribution Nineteen scattered sites in southern Engl<strong>and</strong>,<br />

north to Yorkshire: Cogley Wood (1979), Somerset; Wylye<br />

(1977), Wiltshire; Lyndhurst (1894) <strong>and</strong> Shortheath<br />

Common (1990), Hampshire; Tunbridge Wells (1920) <strong>and</strong><br />

Woolwich Wood (1956, 1957), Kent; Weybridge (1909),<br />

Sydenham Hill Wood (2004) <strong>and</strong> Bransl<strong>and</strong> Wood,<br />

Bletchingley (2004), Surrey; Epping Forest (1907), Essex;<br />

Knebworth (1927), Hertfordshire; Wytham Wood (1949)<br />

<strong>and</strong> Windsor Great Park (1989), Berkshire; Worlington<br />

(1950), Barton Mills (1958) <strong>and</strong> King’s Forest (1989),<br />

Suffolk; Woodditton Wood (1996), Cambridgeshire;<br />

Bishop’s Wood (1954), Staffordshire; Slings<strong>by</strong> (1947),<br />

Yorkshire.<br />

Habitat Old broad-leaved woodl<strong>and</strong>, including marshy<br />

areas; some sheltered chalk grassl<strong>and</strong> sites <strong>and</strong> a limestone<br />

gorge in the Mendips.<br />

Ecology Biology unknown; members of the tribe Eudorylini<br />

are parasitoids of leaf hoppers of the family Cicadellidae as<br />

larvae. The Epping Forest individual was reared from a<br />

pupa found in rotten wood although it seems likely that this<br />

was merely a pupation site. Adults recorded in July.<br />

142

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