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Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

Part 2: Nematocera and Aschiza not dealt with by Falk (1991) - JNCC

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Published sources Ch<strong>and</strong>ler (1992c); Cole & Ch<strong>and</strong>ler<br />

(1979); Gagné (1981).<br />

DIAZOSMA HIRTIPENNE<br />

A winter gnat LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family TRICHOCERIDAE<br />

Diazosma hirtipenne (Siebke, 1863)<br />

Identification Keyed <strong>by</strong> Edwards (1938), Freeman (1950a).<br />

Distribution Records thinly scattered in Engl<strong>and</strong><br />

(Wiltshire, Hertfordshire, Buckinghamshire, Herefordshire,<br />

Lincolnshire, Leicestershire, Yorkshire, Durham), Wales<br />

(Caernarvonshire) <strong>and</strong> Scotl<strong>and</strong> (Roxburghshire, Elgin,<br />

Easterness).<br />

Habitat Mainly broad-leaved woodl<strong>and</strong>.<br />

Ecology A female was found flying in the cavity of an old<br />

hollow Elm (Ulmus) at Grimsthorpe Park, Lincolnshire,<br />

suggesting a potential breeding site. Basden reared a male<br />

Diazosma from the nest of a blue tit or robin in<br />

Buckinghamshire (1931) but the nest was situated on damp<br />

decomposing vegetable matter 2 feet from the ground in a<br />

small hollow Elm (Ulmus) stump. Adults are on the wing<br />

from June to August.<br />

Status Little information <strong>and</strong> it has probably been<br />

overlooked to some extent. There are twelve post 1960<br />

records: Plucking Grove Wood, Lackham Park, Wiltshire<br />

(2002); Rotherfield Park, Hampshire (2002); Aldbury<br />

Common, Hertfordshire (1997); The Coombe,<br />

Buckinghamshire (1997); Grimsthorpe Park, Lincolnshire<br />

(1972); Hatfield Moor (<strong>1991</strong>) <strong>and</strong> Duncombe Park NNR<br />

(1996), Yorkshire (<strong>1991</strong>); Nesbitt Dene, Durham (1992);<br />

Coed Tremadog NNR, Caernarvonshire (1976); Newtown<br />

St Boswells, Roxburghshire (1988); Doune of Relugas,<br />

Elgin (1965); Butterstone Loch, Easterness (<strong>1991</strong>).<br />

Threats Woodl<strong>and</strong> clearance for agriculture or intensive<br />

forestry. Removal of old or diseased trees, which may<br />

support breeding sites.<br />

Management <strong>and</strong> conservation Maintain habitat diversity<br />

in woodl<strong>and</strong>, retaining any old or diseased trees or stumps,<br />

ensuring continuity of habitat in the future.<br />

Published sources Ch<strong>and</strong>ler (1977d, 2003); Edwards<br />

(1938); Payne (1967).<br />

TRICHOCERA MACULIPENNIS<br />

A winter gnat LOWER RISK (Nationally Scarce)<br />

Order DIPTERA<br />

Family TRICHOCERIDAE<br />

Trichocera maculipennis Meigen, 1818<br />

Identification Keyed <strong>by</strong> Freeman (1950a) <strong>and</strong> Laurence<br />

(1957).<br />

Distribution There are isolated records for southern<br />

Engl<strong>and</strong> (Somerset, Kent) <strong>and</strong> scattered records for the<br />

Midl<strong>and</strong>s <strong>and</strong> north of Engl<strong>and</strong> (Staffordshire, Der<strong>by</strong>shire,<br />

Cheshire, Lancashire, Yorkshire, Durham, Westmorl<strong>and</strong>),<br />

Wales (Breconshire, Denbighshire) <strong>and</strong> Scotl<strong>and</strong><br />

(Midlothian).<br />

Habitat Strongly associated <strong>with</strong> caves, although possibly<br />

utilising other shaded damp environments including old<br />

mines.<br />

Ecology Larvae probably develop in decaying matter, such<br />

as animal dung in caves or rotting vegetation <strong>and</strong> fungi<br />

elsewhere. Adults recorded throughout the year.<br />

Status Four known post 1960 localities: Castle Eden Dene<br />

NNR, Durham (1975); Moor House NNR, Westmorl<strong>and</strong><br />

(1963 to 1967); Ogof Ffynnon, Breconshire (1979); Ogof<br />

Dydd Byraf, Denbighshire (1972). Possibly overlooked due<br />

to its secretive nature. Status revised from RDB 3 (Shirt<br />

1987).<br />

Threat Quarrying as well as the degrading of cave systems<br />

<strong>by</strong> caving activities <strong>and</strong> general tidying up for tourism.<br />

Management <strong>and</strong> conservation Maintain caves in a natural<br />

state, placing limits on the amount of disturbance.<br />

Published sources Edwards (1938); Kidd & Brindle<br />

(1959).<br />

MYCETOBIA GEMELLA<br />

A wood gnat<br />

Order DIPTERA<br />

Mycetobia gemella Mamaev, 1968<br />

DATA DEFICIENT<br />

Family MYCETOBIIDAE<br />

Identification The species of Mycetobia are keyed <strong>by</strong><br />

Mamaev (1968). The females are keyed <strong>by</strong> Hancock et al.<br />

(1996).<br />

Distribution Three sites in the Scottish Highl<strong>and</strong>s are<br />

known for this species: Abernethy Forest NNR (1994,<br />

1997), Glen Affric (1996) <strong>and</strong> Rothiemurchus (<strong>1991</strong>),<br />

Easterness (rearings <strong>by</strong> the Malloch Society).<br />

Habitat All sites are Caledonian pine forest.<br />

Ecology All rearings were from Scots Pine (Pinus<br />

sylvestris). The Rothiemurchus record was a rearing from a<br />

rot hole (emergence of adults in late June to early July) <strong>and</strong><br />

the earlier Abernethy Forest NNR record was from a water<br />

run (resembling a sap run) on a trunk <strong>and</strong> from under bark<br />

of a dead st<strong>and</strong>ing pine (adults emerged late May to mid<br />

June).<br />

Status More than 40 individuals were reared from the above<br />

sources. The species was identified only in 1994 <strong>and</strong> was<br />

added to the British list <strong>by</strong> Hancock et al. (1996). Its<br />

distribution will continue to be investigated <strong>by</strong> the Malloch<br />

Society, <strong>and</strong> it is possible that it will be found to occur more<br />

widely in the remaining Scottish pine forests. As <strong>with</strong> M.<br />

pallipes Meigen, finding larvae is probably easier than<br />

finding adults. Currently there is inadequate information to<br />

assess the risk of extinction. Not included in Shirt (1987)<br />

<strong>and</strong> <strong>Falk</strong> (<strong>1991</strong>). Rotheray et al. (2001) recommended RDB<br />

3 status.<br />

Threats Clearance of native pine forest for agriculture or<br />

commercial forestry. Prevention of regeneration of forest <strong>by</strong><br />

deer grazing.<br />

Management <strong>and</strong> conservation Maintain remaining<br />

Caledonian pine forest <strong>with</strong> old <strong>and</strong> decayed trees.<br />

121

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