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A comparative study of models for predation and parasitism

A comparative study of models for predation and parasitism

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73<br />

the curve must be decreasing <strong>for</strong> large values <strong>of</strong> Y. Although the pro<strong>of</strong> is<br />

curtailed here (because it can easily be confirmed by calculating the second<br />

order derivative), it should be mentioned that whether the rate <strong>of</strong> decrease is<br />

accelerated or decelerated depends on the rate <strong>of</strong> decrease in 2~(X) with increasing<br />

i; the curve is decreasing with an increasing rate if the value <strong>of</strong> 2<br />

decreases <strong>comparative</strong>ly fast as i increases, but the rate <strong>of</strong> decrease in the curve<br />

may become lower if the value <strong>of</strong> 2 decreases only slowly with increasing i.<br />

Some examples <strong>of</strong> curves generated by eq. (4i. 9) are shown in Fig. 12. These hypothetical<br />

curves cannot be compared directly with the observed curves <strong>and</strong> scattergram<br />

in Fig. 11, because the values <strong>of</strong> f(X), t, <strong>and</strong> ~ are not known in these observations.<br />

tOO<br />

0,50<br />

Xo(x} = 1.0<br />

e<br />

)h(x) = 0,5 I<br />

(i~<br />

1l<br />

0,10<br />

(1} o= 0<br />

(2) = 0,25<br />

0,0. ~<br />

13) = 0.50<br />

[z, ) = 0,75<br />

X<br />

:0<br />

l,f,-<br />

0<br />

=,<br />

~ 1.0C<br />

i ' ' ' I I<br />

0,1<br />

I I I L I I I i| I I I I I I I II I I<br />

1.0 10,0<br />

o~5o<br />

t<br />

{1)<br />

(2)<br />

0 , 2 0 , 5 , B,0,2,,,5 \ \,~,<br />

1.00 1,50 0,80 0.50 0.25 0,21 0.18 014 011 0.09 0.08 0.07<br />

1oo ,.,2o loo .... o.o oJo 0,50 o: 030 21; 0.27 ::: 0.25 ::; 0,23<br />

0,IC<br />

(1}<br />

..... 0:, . . . . . . . . ,'0 . . . . . . . . ,~0 ' '<br />

MEAN No. OF PARASITES PER EFFECTIVE AREA OF INTERACTION<br />

Fig. 12a. Hypothetical relationships between the values <strong>of</strong> ii/{f(X)t/X} <strong>and</strong> 6Y,<br />

(mean number <strong>of</strong> parasites per effective area) calculated from eqs. (4i. 6) <strong>and</strong><br />

(4i. 8), plotted in the natural logarithmic scale on both axes. The values <strong>of</strong><br />

2i (X) shown in the figure decreases as i increases, indicating that social<br />

interference only is considered here.<br />

Fig. 12b. The same as in Fig. 12a, but the value <strong>of</strong> 2i(X) increases from i=0<br />

to 1, indicating social facilitation, <strong>and</strong> then decreases towards higher values <strong>of</strong> i.

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