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A comparative study of models for predation and parasitism

A comparative study of models for predation and parasitism

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(1)<br />

assumption <strong>of</strong> a PoxssoN distribution,<br />

~E [ {z (r, + Jr)'Aq s~ j!] e - ~'(' '+ dr)2X ,<br />

j~l<br />

(2) probability that all <strong>of</strong> these j points fall within a locus (ri, ri+dr), i.e.<br />

[ {z (r~ +dr) 2_ r, ri~}/~r (ri +dr) 2],,.<br />

With a similar calculation as <strong>for</strong> 7] in Appendix 1, we have<br />

Thus,<br />

Similarly,<br />

probability that at least one point <strong>of</strong> A's falls within radius ri+dr; this is, under the<br />

l. - wr~.X<br />

~m ta~=2~rXe dr.<br />

d r--*O<br />

co PR - *rr2X<br />

lira i=<br />

J r--~0 '=<br />

a~lFi=|o 2r, rXe dr<br />

=l_e-~rR~X<br />

oo PR ~ -wrO-X<br />

lira ]E rq2~=l 2.'rr Xe dr<br />

zIr~O i=1 dO<br />

<strong>and</strong> integrating by parts,<br />

(iii).<br />

fR2~,r,e-,rrO~X, ar=Jo ['R-wr~<br />

e ' dr-Re -'R~'x (iv).<br />

Let O(t) be the normal probability function, i.e.<br />

1 t - t 2/2<br />

$(t) =. 7,-- f e dt,<br />

V~Tr JO<br />

<strong>and</strong> setting t equal to l/2zXr,<br />

9 (1/~R)/V'X.<br />

the integral in the right-h<strong>and</strong> side <strong>of</strong> eq. (iv) will be written as<br />

Thus, from eqs. (ii), (iii), <strong>and</strong> (iv) <strong>and</strong> using symbol ~, we have<br />

L.=[, )<br />

89<br />

Appendix 3. The pro<strong>of</strong> <strong>of</strong> eq. (4i. 6)<br />

Let Y be the density <strong>of</strong> particles distributed at r<strong>and</strong>om over an area A, each one <strong>of</strong> the<br />

particles having a circle <strong>of</strong> area 6 around it. Then the density <strong>of</strong> the particles, Y", within the<br />

total area covered with these circles, i.e. A', is<br />

Y"-=-AY/A'<br />

Suppose that a large number <strong>of</strong> points, i.e. U, with circles <strong>of</strong>, also, area ~ are placed to<br />

cover the whole area <strong>of</strong> A independently <strong>of</strong> the distribution <strong>of</strong> the particles. Then, the number<br />

U' <strong>of</strong> these points that include at least one <strong>of</strong> the particles will he, on the assumption <strong>of</strong> the<br />

Poissos distribution,<br />

U'= U(1-e -~r)<br />

(vi),<br />

<strong>and</strong> the proportion U/U ~ must be equal to the proportion A/A'. Thus from eqs. (v <strong>and</strong> vi), we<br />

find<br />

Y"=Y/(1-e-~Y).<br />

Thus a circle <strong>of</strong> area c~ around each particle contains on the average 6Y~P number <strong>of</strong> particles.<br />

However, since ~Y~ is, as defined in w 4i, the mean number <strong>of</strong> particles in each circle less<br />

the one at the centre, we have<br />

6Y'=6Y"--I<br />

=~Y/(1-e -~r) -1.

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