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The Greenland White-fronted Goose Anser albifrons flavirostris

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4 Spring staging in Iceland and the flight to <strong>Greenland</strong><br />

4.1 Introduction<br />

Until the 1980s, little was known about the status<br />

of staging <strong>Greenland</strong> <strong>White</strong>-<strong>fronted</strong> Geese in Iceland.<br />

It was known that large numbers passed<br />

through Iceland in autumn, but now it is known<br />

that the population also stops there in spring en<br />

route to the breeding grounds (MS4). It is clearly<br />

important to establish whether, where and for<br />

how long birds stage on spring migration since<br />

feeding at staging sites enables individuals to recoup<br />

depleted energy and nutrient stores and<br />

hence potentially affect subsequent breeding success<br />

(see review by Thomas 1983). Weather conditions<br />

in Iceland in spring had been shown to<br />

correlate with subsequent breeding success of the<br />

Pink-footed Geese <strong>Anser</strong> brachyrhynchus which<br />

nest there (Fox et al. 1989). Hence, it was reasonable<br />

to assume that weather conditions and the<br />

availability of forage were likely to affect the condition<br />

of migrating <strong>Greenland</strong> <strong>White</strong>-<strong>fronted</strong><br />

Geese departing for the breeding grounds from<br />

Iceland in spring. During the late 1980s and<br />

throughout the 1990s, a series of studies were carried<br />

out examining the spring staging of different<br />

goose species in Iceland, to assess the importance<br />

of feeding ecology of pre-nesting staging<br />

in that country. An important aim was to establish<br />

the importance of specific staging areas and<br />

to assess the distribution and abundance of the<br />

different goose populations there, examining the<br />

use of different habitats and the way in which<br />

the geese exploit these (e.g. Boyd & Fox 1992,<br />

1995, Fox et al. 1991, 1992, 1994a, 1994b, 1999). In<br />

very recent years, these studies have involved a<br />

Ph.D study (Kristiansen 2001) and Masters student<br />

study (Nyegaard 2001), but from which selected<br />

results are presented here.<br />

In conservation terms, the most important elements<br />

in the study of spring staging relate to the<br />

identification of staging areas, the habitats and<br />

food plants used by the geese, the level of energetic<br />

gain that can be achieved by geese utilising<br />

different habitat types and how they may derive<br />

the maximum nutritional benefit from them. In<br />

this way, it is possible to establish some basis for<br />

ranking the importance of sites and habitats in<br />

terms of the number of birds using different sites<br />

for site safeguard and nature conservation management<br />

purposes. Finally, studies have also con-<br />

sidered how much nutrient gain can be derived<br />

from the staging period in Iceland during the prelude<br />

to migration over the <strong>Greenland</strong> Ice Cap to<br />

the breeding grounds on the west coast.<br />

4.2Distribution of spring staging<br />

<strong>Greenland</strong> <strong>White</strong>-<strong>fronted</strong> Geese in<br />

Iceland<br />

Observations from many years confirm that a<br />

large proportion (if not all) <strong>Greenland</strong> <strong>White</strong><strong>fronted</strong><br />

Geese stage in Iceland from c. 10 April to<br />

c.12 May in most seasons (MS4, MS19). At this<br />

time, they use two main areas, namely the southern<br />

lowlands (Árnessýsla, Rangárvallassýsla and<br />

Vestur-Skaftafellssýsla) and the western lowlands<br />

(Kjósarsýsla, Borgarfjarðarsýsla, Mýrarsýsla and<br />

Snæfellsness- og Hnappadalssýsla). Spring migration<br />

phenology appears to differ between areas,<br />

with earlier arrivals but a slower build-up to<br />

maximum numbers in the southern lowlands.<br />

Here, numbers peaked during 24-26 April in 1990-<br />

1992, but dispersed earlier compared to the rapid<br />

build up in western staging areas to peak numbers<br />

during 18-22 April in 1997-1999, where substantial<br />

numbers of birds remained well into May<br />

each year (MS19). <strong>The</strong>se differences most likely<br />

relate to the timing of migration in the years concerned,<br />

but could also reflect different migration<br />

strategies of birds using the two areas (records of<br />

individuals using both staging areas are rare,<br />

MS27). <strong>The</strong> southern lowlands, especially along<br />

the coast, experience a slightly milder climate and<br />

hence earlier thaw than the west, which may initiate<br />

plant growth a little earlier in the spring. In<br />

addition, there are substantial areas of tillage in<br />

the southern lowlands that do not occur in the<br />

west (see below).<br />

At the most important staging site in the western<br />

lowlands, Hvanneyri, up to a maximum of 1,600<br />

birds have been counted (MS19). Based on observations<br />

of individually marked birds at Hvanneyri,<br />

more than half of the geese remained for less<br />

than a week, mostly early on in migration, whilst<br />

more than a third stayed for almost the entire staging<br />

period (c. 24 days).<br />

From observations of individually marked <strong>Greenland</strong><br />

<strong>White</strong>-<strong>fronted</strong> Geese, at least 90% of goslings<br />

35

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