An in-depth study reviewing 22 years of effort-related harbour porpoise data from around the UKfound the coastal waters of west Scotland, including the eastern sectors of The Minches and theSea of the Hebrides (around the Isles of Skye, Mull and the Small Isles), to be one of four sites ofkey importance to harbour porpoises in UK waters 82 . This analysis found that porpoises are presentin every month of the year, with concentrations in several months, including during the April-Septemberperiod when calving takes place. They also found a higher than average proportion of calves toadults in the area which suggests its importance for calving and/or breeding harbour porpoises.The long time series of data showed that this situation has been consistent over many years.Numerous studies in the region over recent years have reported high encounter rates for harbourporpoises. Encounter rates are generally highest in the spring and summer months, particularly inthe Gairloch region 83 , around the Isle of Skye 84 , and the Firth of Lorne and Sound of Jura 85 . Outsideof the summer months data are scarcer, but relatively high encounter rates have been recorded inthe northern part of the Minch in autumn 86 .Data on harbour porpoise calf presence are problematic due to difficulties in sighting such smallanimals and they are likely underreported. However, studies have reported relatively high numbersof groups with calves in The Minches and the Sea of the Hebrides 87 , and an aerial survey of theInner Hebrides found a high calf to adult ratio with calves making up 10% of all animals sighted 88 .The presence of a higher proportion of mother and calf pairs is an indication that the area is usedby harbour porpoises for calving 89 . This is supported by observations of researchers in the Gairlochregion suggesting behaviour typical of porpoises calving 90 .Spatial modelling of harbour porpoise data has been used to look at distribution in the region on afiner scale. One model, created using four years of survey data from the month of August, predictedfour areas of high relative abundance: 1) the region between Ardnamurchan, Coll and the SmallIsles, which the authors highlight is also important sandeel habitat (harbour porpoise prey),2) south east of Barra, an area they speculate may be preferentially used in seasonal movementsto enter the Sea of the Hebrides from offshore areas to the west, where other surveys have madesightings 91 , 3) north east of Skye to Gairloch, and 4) west of Pairc Peninsula (Isle of Lewis) tothe Shiant Islands 92 . The area south of Barra has been found to have relatively high porpoiseabundance by previous studies 93 and as discussed above, Gairloch has been identified as animportant area for porpoise calving, and other studies have noted it to be a highly significantlocation for harbour porpoises 94 .A second habitat modelling study using summer data from further south in the Hebrides predictedhighest densities close to the mainland – in the Sound of Jura, the Firth of Lorne, between Mull andthe Treshnish Islands, in patches around the Small Isles and the Sound of Sleat 95 . Although harbourporpoises clearly use the whole area, the same core, high-use areas were predicted over threeyears. Some inshore areas – the Sound of Mull, upper Firth of Lorne and upper Sound of Jura – hadsightings rates almost as high as the high density areas identified, but could not be included in theanalysis because of a lack of environmental data. The authors suggest that these are also important toharbour porpoises and should be considered alongside the areas identified by the model.Spatial habitat preference modelling such as this allows us to build on the knowledge gained fromsurvey data and observations, and can help to further define areas of critical habitat for cetaceans 96 .West and north of the Outer Hebrides, data are much sparser. Harbour porpoises are generallyonly recorded in low numbers although some studies have reported high relative encounter rateswest of the Uists and north east of Lewis 97 . Higher sea states and poorer weather offshore willinhibit the sighting of harbour porpoises and may mean they are underestimated in these areas.Far West ScotlandThe predominant view of harbour porpoises is of a species largely confined to nearshore watersless than 200m deep but despite this, several studies report their presence in deep offshore waters,albeit in relatively small numbers. The Wyville-Thomson Ridge and surrounding channels is an areanoted for porpoise presence 98 .2482Evans and Wang, 200283MacLeod, 200184Pollock et al, 2000; Goodwin and Speedie, 2008; Embling et al, 2010; Marubini et al, 200985Pollock et al, 2000; Embling et al, 201086MacLeod, 200187Weir, 2008; Evans and Wang, 200288SCANS II, unpublished data (in Embling et al, 2010)89Baines and Earl, 199890Dolman and Hodgins, 200991Macleod et al, 2003; Macleod et al, 200792Marubini et al, 200993MacLeod, 200194Dolman and Hodgins, 2009; Shrimpton and Parsons, 200095Embling et al, 201096Hoyt, 200997Pollock et al, 2000; Lewis et al, 199898Pollock et al, 2000; Evans and Wang, 2002; MacLeod, 2001
North ScotlandThe first SCANS survey (in July, 1994) recorded the highest densities in UK waters for the areaaround the Northern Isles – 0.783 animals/km 2 , and an abundance of 24,335 animals. The adjacentsurvey area, covering the waters to the east, west and north of the Northern Isles recorded amoderate density of 0.363 animals/km 2 (see Figure 4.3, p.21). Spatial modelling of this data predicteda large area of high density spanning much of the waters of east, north east and north Scotland(see Figure 4.4, p.21).A multi-year study during the 1990s covering porpoise distribution and habitat use in Shetland'scoastal waters found them to be widespread but with concentrations on the east and south coastsof mainland Shetland 99 . The most important areas were Sumburgh Head/Quendale Bay, MousaSound to Helliness, south Noss Sound, and east Whalsay/Out Skerries. These areas tend to becharacterised by strong tidal currents and as in other areas, harbour porpoises seem to takeadvantage of these conditions to forage 100 .Analysis of over 20 years of harbour porpoise data from around the UK found Shetland to be oneof four sites rated most important for this species 101 . Porpoises were shown to be present here inevery month, with concentrations during the April-September key calving period. They also foundthat a higher than average proportion of calves to adults was recorded here in August andSeptember, suggesting its importance as a calving and/or nursery area 102 .However, distributional changes appear to have occurred in recent years and porpoise sightingshave become much less common 103 . This observation was supported by the SCANS II survey thattook place in 2005 and found densities to be less than half what they were during the mid-1990s,at 0.274 animals/ km 2 around the Northern Isles 104 (see Figure 4.5, p.22). Spatial modelling of thisdataset suggested that at a large scale, the area of high density observed off south east Scotlandin 1994 had largely dispersed by 2005 105 (see Figures 4.4, p.21 and 4.6, p.22). To the north of theScottish mainland, high densities had persisted and intensified 106 . High densities of harbourporpoises during the summer, the time of year when these animals breed and produceyoung, suggests these waters may be important for these functions. This should beinvestigated further.The reasons for the apparent changes in distribution are unknown but may be related to preydistribution and abundance 107 . Shortages of sandeels, a staple food source for harbour porpoisesas well as other cetaceans and seabirds, have been observed in the region over recent years andlinked to starvation and breeding failures in seabirds 108 . The observed decrease in harbour porpoisesightings may be a result of reduced prey availability.Relatively high sightings rates have also been recorded near the Fair Isle and in Orkney's coastalwaters, during spring, summer and winter 109 . There is very little information about this area however.East ScotlandThis region is most famous for its bottlenose dolphin population but it is also home to significantnumbers of harbour porpoises. The SCANS survey (in July, 1994) recorded the highest density ofporpoises in UK waters off the Northern Isles and the Inner Moray Firth (see Figure 4.3, p.21) at0.783 animals/km 2 and an abundance of 24,335 animals 110 . The adjacent survey block coveringthe waters of north Scotland and the Outer Moray Firth recorded a moderate density of 0.363animals/km 2 . Spatial modelling of this data predicted a large area of high density covering thewaters of east, north east and north Scotland 111 (see Figure 4.4, p.21).Ten years later when this survey was repeated (SCANS II), this high density area had changed,and only moderately low densities of harbour porpoises were recorded for north east and eastScottish waters 112 (see Figures 4.5 and 4.6, p.22). The reasons for this are unknown but may berelated to prey distribution and abundance. (See East England section below for further discussion.)99Evans, 1997100Ibid101Evans and Wang, 2002102Ibid103Evans and Wang, 2008104SCANS II, 2006105Ibid106Ibid107Evans and Wang, 2008108JNCC press release, 2005 http://www.jncc.gov.uk/page3628109Pollock et al, 2000; Evans and Wang, 2002110Hammond et al, 1995111SCANS II, 2006112Ibid25
- Page 1 and 2: WDCS, the Whale and Dolphin Conserv
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6.1.1 HARBOUR PORPOISEWest and sout
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Assessment: Critical HabitatHigh re
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11. Area: Off Land's End, Cornwall
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Threats, status and relative import
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East Scotland3. Area: North east Sc
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Threats, status and relative import
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6.1.4 COMMON MINKE WHALEWest and so
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6.1.5 WHITEBEAKED DOLPHINWest and s
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Coastal Wales2. Area: Bardsey Islan
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6.1.8 ATLANTIC WHITESIDED DOLPHINNo
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Other areasOne other area was asses
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6.1.11 BEAKED WHALESThe northern Ro
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7. PROTECTING CRITICAL HABITAT - RE
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considered in the area, each of whi
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7.4 EAST SCOTLANDWithin the east Sc
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7.5 SOUTH WEST ENGLANDWithin the so
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An SAC is already in place for bott
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7.9 SUMMARYFour areas of identified
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iv. Licensing processes for industr
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9. IDENTIFICATION AND PRIORITISATIO
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Recommendations on particular data
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Coastal Wales northern Pembrokeshir
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ANNEX II SPECIES ACCOUNTS SUMMARY
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HARBOUR PORPOISEEast ScotlandStudyH
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HARBOUR PORPOISESouth West EnglandS
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HARBOUR PORPOISECoastal WalesStudyT
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BOTTLENOSE DOLPHINStudyTemporalReso
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SHORTBEAKED COMMON DOLPHINStudyTemp
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SHORTBEAKED COMMON DOLPHINStudyTemp
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COMMON MINKE WHALEWest ScotlandStud
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WHITEBEAKED DOLPHINStudyTemporalRes
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KILLER WHALEStudyTemporalResolution
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ATLANTIC WHITESIDED DOLPHINStudyTem
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SPERM WHALEStudyTemporalResolutionS
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BALEEN WHALESStudyTemporalResolutio
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ANNEX III THREATS TO CETACEANS - RE
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Actual orPotential ThreatActivitySp
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North ScotlandSpecies of most relev
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East EnglandSpecies of most relevan
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South East EnglandSpecies of most r
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South West EnglandSpecies of most r
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Irish SeaSpecies of most relevance
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Actual orPotential ThreatActivitySp
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Bravington, M., Borchers, D. and No
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Evans, P.G.H. and Anderwald, P. 200
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ICES WGMME. 2006. Report of the Wor
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ECS/ASCOBANS/ACCOBAMS Workshop held
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marine protected areas effective to
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ISBN: 9781901386233