For the Inner and Outer Moray Firth, harbour porpoises are the most commonly sighted cetacean,found throughout the area, in inshore and offshore waters 113 . For most or all of the year, harbourporpoises are widely distributed throughout the Inner Moray Firth 114 , an area designated a SpecialArea of Conservation (SAC) 115 for bottlenose dolphins. There does appear to be some segregationbetween the two species in this area, perhaps unsurprising given that bottlenose dolphins areknown to attack harbour porpoises 116 . Although porpoises can be sighted throughout the InnerMoray Firth, surveys and habitat modelling studies suggest higher relative densities away from thecoastal areas used by bottlenose dolphins, towards the centre of the area from the outer SACboundary towards the Inverness Firth, and also off Helmsdale on the northern coast 117 . High ratiosof young to adult harbour porpoises were also found for this region off the north coast, suggestingits importance for breeding and calving 118 .Fewer data are available overall for the Outer Moray Firth, particularly the northern and centralwaters, but recent visual and acoustic surveys have found porpoises to be widespread in this areafor much of the year 119 . Monitoring with PODs 120 found the highest level of detections for the OuterMoray Firth and recorded harbour porpoises at 97.5% of sites for an average of 6.7 hours a day 121 .Visual surveys carried out in conjunction with the acoustic monitoring supported these results, withharbour porpoises encountered on all surveys across the Outer Moray Firth, and with relativelyhigh sightings rates 122 . High densities of harbour porpoises during the summer, the time ofyear when these animals breed and produce young, suggest these waters may be importantfor these functions and this should be investigated further.A multi-year study of the southern coastal region of the Outer Moray Firth also reports high relativeabundance of harbour porpoises for this area 123 . Numbers apparently increase progressively fromMay through to October. During the summer months, females and calves moved inshore, which theauthors consider is due to sheltered inshore waters providing preferred calving habitat. Similar toother areas, neonatal calves were mostly observed between May and July. Increases in harbourporpoises over the summer are also likely due to sandeel availability in the area, providing goodforaging opportunities 124 .Nearby, along the Aberdeenshire coast, harbour porpoises are also present throughout the year,with abundance peaking in August and September. Researchers concluded that the seasonalincrease in harbour porpoises in the area was partly due to their preferential use of inshore watersfor calving, and also to take advantage of foraging opportunities 125 . Most calves and juveniles wererecorded between June and September, with June the peak month. This is the key period forharbour porpoise calving and indicates that the area is used for this purpose 126 . The timing of theincrease in harbour porpoise sightings matches when mackerel are known to move inshore, andthe particular area most favoured by porpoises is reportedly also where large numbers of mackerelare present 127 .In a study looking at several decades of harbour porpoise data from around the UK, the southerncoast of the Moray Firth and the north east Aberdeen coast came through strongly as an importantarea for harbour porpoises 128 . Records over many years show they are present for most of the yearand in significant concentrations during the summer calving months. Data on young are generallyproblematic because of difficulties with sighting calves, but several areas in this region were identifiedas having higher than average proportions of calves to adults – the Banff coast (June), the InnerMoray Firth and north Aberdeen coast (July), and the Aberdeen coast again in August 129 .For the remainder of this region – coastal south east Scotland, offshore east Scotland and thenorthern Moray Firth – there are insufficient data available to make an assessment of itsimportance for harbour porpoises.26113Thompson et al, 2010114Bailey and Thompson, 2009; Hastie et al, 2003; Evans and Wang, 2002115Designated under the EC Habitats Directive116Bailey and Thompson, 2009; Thompson et al, 2004117Bailey and Thompson, 2009; Hastie et al, 2003118Evans and Wang, 2002119Eisfeld et al, 2009; Thompson et al, 2010120Porpoise acoustic monitoring devices121Thompson et al, 2010122Ibid123Robinson et al, 2007124Ibid125Weir et al, 2007126Ibid127Ibid128Evans and Wang, 2002129Ibid
East EnglandThe seasonal pattern in the southern North Sea appears to be for an early spring peak in numbersin coastal waters to be followed by a northward migration towards more offshore waters. However,whilst some studies report that numbers of harbour porpoises in coastal waters around the southernNorth Sea then remains low for the summer 130 , others have found a second peak to take place offeast England in August and September, possibly highlighting a regional difference 131 .The major SCANS surveys undertaken in the summers of 1994 (SCANS) and 2005 (SCANS II)across the North Sea and European Atlantic continental shelf waters provided some interestinginsights into possible large-scale changes in harbour porpoise distribution in the North Sea overrecent years (see Figures 4.3-4.4, p.21 and 4.5-4.6, p.22). The 1994 survey 132 recorded moderatenumbers of harbour porpoises off east England and in the central/southern North Sea region (0.387animals/km 2 and 0.34 animals/km 2 ). Yet when SCANS II took place in 2005 133 , numbers in the southNorth Sea had increased greatly and densities of 0.562 animals/km 2 were recorded, while in thenorthern North Sea a corresponding decrease was detected. This may indicate a redistribution ofporpoises from north to south, which is thought to be most likely due to changes in the distributionor availability of prey 134 . Highest densities in 2005 stretched from The Wash and north Norfolkcoast, north and west to the Dogger Bank region 135 . Many sandbanks exist in this area, importanthabitat for sandeel and other prey species utilised by harbour porpoises, other marine mammals,seabirds and fish 136 .Other studies have highlighted the importance of the Dogger Bank area to harbour porpoises,other cetaceans and many species of seabirds, and multi-species feeding associations have beendocumented here 137 . Surveys over the German sector of the Bank found high densities of porpoises(1 - 1.5 animals/km 2 ) 138 . The German side of the bank has been designated as an SAC and theDutch and UK areas of the bank have also been proposed for an SAC, but not with harbour porpoisesas a qualifying feature 139 . An acoustic study based on offshore installations on the Dogger Bankrecorded porpoises regularly around the installations and considered these may be importantforaging areas for this species 140 . The authors also noted that if porpoises cluster around theseinstallations as their research suggests, these animals may be omitted by population surveys suchas SCANS as survey vessels would have to remain outside the 500m exclusion zone.Trying to understand these large-scale changes in distribution, and the driving forces behind them,will only be possible by repeating North Sea wide surveys such as SCANS on decadal or morefrequent basis, and also by investigating porpoise-prey dynamics. The SCANS surveys — designedto provide large-scale population estimates — provide no information on the relative importance ofhabitats at a fine scale. It may be that smaller areas within the North Sea are important to harbourporpoises. Smaller scale surveys are also necessary to determine if some areas remainimportant to harbour porpoises throughout these larger scale fluctuations.A review and analysis of over 20 years of harbour porpoise data from around the UK identifiedthree areas off the east coast of England that are potentially consistently important to harbourporpoises – east of Northumberland, east of Yorkshire, and particularly, east of the Wash near theNorfolk coastline. Porpoises have been recorded in these locations for some or most months of theyear, with concentrations in several months, and records over several years 141 . For east of theWash these concentrations occur during the April to September key calving period for harbourporpoises 142 . High densities of harbour porpoises during the summer, the time of year whenthese animals breed and produce young, suggest these waters may be important for thesefunctions and this should be investigated further.South east EnglandDeclines in harbour porpoise abundance have been observed in the southern North Sea regionsince the 1950s, for reasons not understood 143 . For the last few years, sightings and strandingsdata from Belgian and Dutch waters have indicated an increase in numbers for the southern NorthSea, and the SCANS II survey results (see East England section) support this. However, the130Haelters and Camphuysen, 2009131Evans and Wang, 2002132Hammond et al, 2002133SCANS II, 2006134Ibid135Ibid136JNCC, 2010137Camphuysen et al, 1995; Gubbay et al, 2002138Gilles et al, 2008139JNCC, 2010140Todd et al, 2009141Evans and Wang 2002142Ibid143Haelters and Camphuysen, 200927
- Page 1 and 2: WDCS, the Whale and Dolphin Conserv
- Page 3 and 4: Towards Marine ProtectedAreas for C
- Page 5 and 6: South west EnglandCeltic Deep - com
- Page 7 and 8: CONTENTSExecutive Summary 2Acknowle
- Page 9 and 10: 1. INTRODUCTIONWe enter an exciting
- Page 11 and 12: Note: This schematic map shows thev
- Page 13 and 14: 2. IDENTIFYING MARINE PROTECTED ARE
- Page 15 and 16: 3. CRITICAL HABITATInterest in the
- Page 17 and 18: Foraging, travel, socialising and p
- Page 19 and 20: 4. DATA AND SPECIES ACCOUNTSIn orde
- Page 21 and 22: Killer whale (Orcinus orca)Atlantic
- Page 23 and 24: 4.4 SPECIES ACCOUNTS4.4.1 HARBOUR P
- Page 25 and 26: Although there is some variation in
- Page 27: North ScotlandThe first SCANS surve
- Page 31 and 32: Irish SeaOutside of coastal Welsh w
- Page 33 and 34: 4.4.2 BOTTLENOSE DOLPHIN (Tursiops
- Page 35 and 36: This expansion in range means this
- Page 37 and 38: 4.4.3 SHORTBEAKED COMMON DOLPHIN (D
- Page 39 and 40: South west EnglandBased on observat
- Page 41 and 42: 4.4.4 COMMON MINKE WHALE (Balaenopt
- Page 43 and 44: Sightings for this area of the Oute
- Page 45 and 46: The JNCC Atlas of cetacean distribu
- Page 47 and 48: 4.4.5 WHITEBEAKED DOLPHIN (Lagenorh
- Page 49 and 50: 4.4.6 RISSO'S DOLPHIN (Grampus gris
- Page 51 and 52: 4.4.7 KILLER WHALE OR ORCA (Orcinus
- Page 53 and 54: 4.4.8 ATLANTIC WHITESIDED DOLPHIN (
- Page 55 and 56: 4.4.9 LONGFINNED PILOT WHALE (Globi
- Page 57 and 58: 4.4.10 SPERM WHALE (Physeter macroc
- Page 59 and 60: 4.4.11 BEAKED WHALESNORTHERN BOTTLE
- Page 61 and 62: which looked at worldwide beaked wh
- Page 63 and 64: There is no current population esti
- Page 65 and 66: 4.5 SCOTTISH MPA GUIDELINES - STAGE
- Page 67 and 68: Northern bottlenose whale and Sower
- Page 69 and 70: Fisheries BycatchFisheries bycatch
- Page 71 and 72: 5.2 AN OVERVIEW OF REGIONAL THREATS
- Page 73 and 74: North ScotlandAs in the west, threa
- Page 75 and 76: 6. IDENTIFYING CRITICAL HABITAT IN
- Page 77 and 78: 6.1.1 HARBOUR PORPOISEWest and sout
- Page 79 and 80:
Assessment: Critical HabitatHigh re
- Page 81 and 82:
11. Area: Off Land's End, Cornwall
- Page 83 and 84:
Threats, status and relative import
- Page 85 and 86:
East Scotland3. Area: North east Sc
- Page 87 and 88:
Threats, status and relative import
- Page 89 and 90:
6.1.4 COMMON MINKE WHALEWest and so
- Page 91 and 92:
6.1.5 WHITEBEAKED DOLPHINWest and s
- Page 93 and 94:
Coastal Wales2. Area: Bardsey Islan
- Page 95 and 96:
6.1.8 ATLANTIC WHITESIDED DOLPHINNo
- Page 97 and 98:
Other areasOne other area was asses
- Page 99 and 100:
6.1.11 BEAKED WHALESThe northern Ro
- Page 101 and 102:
7. PROTECTING CRITICAL HABITAT - RE
- Page 103 and 104:
considered in the area, each of whi
- Page 105 and 106:
7.4 EAST SCOTLANDWithin the east Sc
- Page 107 and 108:
7.5 SOUTH WEST ENGLANDWithin the so
- Page 109 and 110:
An SAC is already in place for bott
- Page 111 and 112:
7.9 SUMMARYFour areas of identified
- Page 113 and 114:
iv. Licensing processes for industr
- Page 115 and 116:
9. IDENTIFICATION AND PRIORITISATIO
- Page 117 and 118:
Recommendations on particular data
- Page 119 and 120:
Coastal Wales northern Pembrokeshir
- Page 121 and 122:
ANNEX II SPECIES ACCOUNTS SUMMARY
- Page 123 and 124:
HARBOUR PORPOISEEast ScotlandStudyH
- Page 125 and 126:
HARBOUR PORPOISESouth West EnglandS
- Page 127 and 128:
HARBOUR PORPOISECoastal WalesStudyT
- Page 129 and 130:
BOTTLENOSE DOLPHINStudyTemporalReso
- Page 131 and 132:
SHORTBEAKED COMMON DOLPHINStudyTemp
- Page 133 and 134:
SHORTBEAKED COMMON DOLPHINStudyTemp
- Page 135 and 136:
COMMON MINKE WHALEWest ScotlandStud
- Page 137 and 138:
WHITEBEAKED DOLPHINStudyTemporalRes
- Page 139 and 140:
KILLER WHALEStudyTemporalResolution
- Page 141 and 142:
ATLANTIC WHITESIDED DOLPHINStudyTem
- Page 143 and 144:
SPERM WHALEStudyTemporalResolutionS
- Page 145 and 146:
BALEEN WHALESStudyTemporalResolutio
- Page 147 and 148:
ANNEX III THREATS TO CETACEANS - RE
- Page 149 and 150:
Actual orPotential ThreatActivitySp
- Page 151:
North ScotlandSpecies of most relev
- Page 156 and 157:
East EnglandSpecies of most relevan
- Page 158 and 159:
South East EnglandSpecies of most r
- Page 160 and 161:
South West EnglandSpecies of most r
- Page 162 and 163:
Irish SeaSpecies of most relevance
- Page 164 and 165:
Actual orPotential ThreatActivitySp
- Page 166 and 167:
Bravington, M., Borchers, D. and No
- Page 168 and 169:
Evans, P.G.H. and Anderwald, P. 200
- Page 170 and 171:
ICES WGMME. 2006. Report of the Wor
- Page 172 and 173:
ECS/ASCOBANS/ACCOBAMS Workshop held
- Page 174 and 175:
marine protected areas effective to
- Page 176:
ISBN: 9781901386233