eastern Channel does not seem to have had a similar increase in porpoise numbers. DuringSCANS (in July 1994) none were sighted off south east England. During SCANS II, a moderatedensity of 0.331 animals/km 2 was recorded for the block that included this area but spatial modellingof the results suggests higher densities of animals were in the western Channel, and only the verynorthern part of the eastern Channel stretching down from the region of high density off eastEngland (see Figures 4.5 and 4.6, p.22).South west EnglandThe waters off south west England are a further area where the SCANS surveys showedsignificant changes in distribution of harbour porpoises in 10 years. SCANS (in July 1994)recorded no porpoises in the <strong>English</strong> Channel region and a relatively low density for the CelticSea and Bristol Channel 144 (see Figure 4.3, p.21). Ten years later during SCANS II, moderatedensities were recorded in the <strong>English</strong> Channel (0.331 animals/km 2 ) and high densities in theCeltic Sea and Bristol Channel (0.408 animals/km 2 ) (see Figure 4.5, p.22). Spatial modelling ofthe SCANS II dataset predicted moderate densities for a large area of the Celtic Sea and the verywestern edge of the <strong>English</strong> Channel 145 (see Figure 4.6, p.22).An increase in the summer occurrence of harbour porpoises in the <strong>English</strong> Channel has also beensuggested by an 11-year study (1996-2006) using data collected from ferries travelling acrossthe <strong>English</strong> Channel and the Bay of Biscay 146 . Findings indicate that this increase has occurredsince 2002 and may only be the result of an increase in summer abundance, not a consistentyear-round increase 147 .Analysis of over 20 years of harbour porpoise data for UK waters highlighted several hotspotsfor harbour porpoises in this region 148 . North Devon was determined to be important, with highconcentrations of porpoises in several months of the year, particularly during the April to Septemberperiod. An area in the Outer Bristol Channel, south to north west Cornwall, was rated as animportant site, but with fewer months of high concentrations than north Devon. High densities ofharbour porpoises during the summer months, the time of year when these animals breedand produce young, suggest these waters may be important for these functions and thisshould be investigated further.Data were lacking for the western <strong>English</strong> Channel region but significant concentrations ofporpoises were still noted for the first quarter of the year 149 .Little other data were found for the north Devon coast. One study investigated the fine-scale habitatuse by harbour porpoises of two sites in this area and determined that one, Morte Point, was animportant feeding ground for harbour porpoises 150 . Morte Point is a headland with an area of tidalrapids nearby and porpoises were observed to spend 60% of their time in the area foraging. Thisactivity predominantly occurred within the tidal race 151 which is similar to results from other locationswhere porpoises are seen to take advantage of the aggregating effect of high tidal flow. At thesecond site, Lee Bay, porpoises were also observed feeding but spent more time travelling in thisarea and it seems to be used mainly as a corridor between more productive feeding sites 152 .Some survey work has taken place in the western <strong>English</strong> Channel and close to the Scilly Islesduring the winter 153 . Although overall relative abundance was fairly low, porpoises were not evenlydistributed, with large groups of apparently foraging porpoises seen at times 154 . A land-basedsurvey conducted at the same time of year reported relatively high densities of porpoises off Land'sEnd and frequently observed them to be foraging here 155 . These surveys would seem to supportother analyses of data from the region that found there to be concentrations of porpoises in thewestern <strong>English</strong> Channel during the first few months of the year 156 . Analysis of multiple years ofcetacean sighting data from the coastal waters of Cornwall, investigating spatial and temporal patterns,also found harbour porpoises were predominantly sighted off Land's End, with a peak in winter 157 .28144Hammond et al, 1995145SCANS II, 2006146Macleod et al, 2009147Ibid148Evans and Wang, 2002149Ibid150Goodwin, 2008151Ibid152Ibid153De Boer and Simmonds, 2004; WDCS, 2005b; De Boer and Saulino, 2007154De Boer and Simmonds, 2004155De Boer and Saulino, 2007156Evans and Wang, 2002157Pikesley et al, IN PRESS
Irish SeaOutside of coastal Welsh waters (see following section), the area of the Irish Sea consistentlyhighlighted for high porpoise numbers is the St. George's Channel in the south 158 . The waters offPembrokeshire appear to be the centre of high porpoise densities but further west into the St.George's Channel, several studies report high concentrations in some months, including during thesummer 159 . High densities of harbour porpoises during the summer, the time of year whenthese animals breed and produce young, suggest these waters may be important for thesefunctions and this should be investigated further.The central Irish Sea has also been noted by some studies for possible concentrations of porpoises,although there are fewer data 160 . There is a potential association here between harbour porpoisesand the western Irish Sea front which appears to create an area of increased productivity 161 . Thisfront develops seasonally, south from the Isle of Man, as a body of deep, thermally stratified watermeets a shallower body of mixed, nutrient rich water. There is some movement of this front but itremains relatively static throughout the season 162 . At this interface, an enhanced level of productivitydevelops, attracting top predators such as seabirds and harbour porpoises. A study here found thatporpoises occurred more frequently in the frontal area, compared to other areas of the central IrishSea included in the survey, and were predominantly found on the mixed side of the front 163 . The studysuggests that the aggregation of harbour porpoises may be the result of this increased productivity.Coastal WalesSeveral areas in coastal Welsh waters have been highlighted for their high sightings rates andapparent importance to harbour porpoises. Northern Pembrokeshire, particularly Strumble Headand around Ramsey and Skomer islands, is an area shown to have high concentrations of porpoisesin most months of the year, including the April to September calving period 164 . Several studies lookingat data over a long time period have highlighted the waters west of Pembrokeshire as a harbourporpoise hotspot 165 . Peaks in abundance have been noted for late summer and early autumn, andagain in April 166 , but surveys at other times of year such as late autumn have also found the highestnumber of porpoises in the region to be off Pembrokeshire 167 indicating the area’s relative importanceat other times of year.Foraging is the most frequently observed behaviour here — again, the area is characterised bystrong tidal currents and races — and animals are observed holding their position in the current,often for extended periods, and regularly joined by feeding seabirds 168 .More detailed investigations into the foraging strategy and behaviour of harbour porpoises atRamsey Sound found that foraging was almost entirely restricted to the ebb tidal phase 169 . At thissite, aggregations of up to 20 porpoises were recorded at times and it was not uncommon to seegroups repeatedly and closely surfacing together, appearing to pursue prey in a coordinatedmanner 170 . At other times, usually after extended periods of foraging, contact between groups ofindividuals seemed to take on a social context, with slow milling, intermingling, leaps and tailslapping observed 171 .These observations, combined with the presence of females with calves, throughout the year butparticularly through the summer, indicate that breeding takes place in this area 172 . This assertion issupported by investigations into the strandings records in the region which have shown that in Juneand July (peak harbour porpoise birthing months), high proportions of freshly stranded harbourporpoises were neonates (> 50%) 173 . Review of effort- and non effort-related sightings data fromthe region has indicated that harbour porpoise mothers with calves prefer the Skomer-Ramseyregion over Strumble Head — the area with the highest overall porpoise densities 174 .In a study looking at the comparative importance of sites around the UK, the area of Pembrokeshireand southern Cardigan Bay was rated as one of the four most important known sites for harbourporpoises in the UK 175 . Southern Cardigan Bay was also highlighted as an area with higher thanaverage proportions of calves throughout the summer months indicating its importance as a calvingand breeding area 176 .158Northridge et al, 1995; Bravington et al, 2002; Evans and Wang, 2002159Evans and Wang, 2002; Northridge et al, 1995160Bravington et al, 2002161Weir and O'Brien, 2000162WWF163Weir and O'Brien, 2000164Baines and Evans, 2009; Evans and Wang, 2002165Evans and Wang, 2002; Baines and Evans, 2009; Bravington et al, 2002; Northridge et al, 1995166Pierpoint et al, 1998167De Boer and Simmonds, 2003168Ibid169Pierpoint, 2008170Ibid171Ibid172Pierpoint et al, 1998; Baines and Earl, 1999173Penrose and Pierpoint, 1999174Baines and Earl, 1999175Evans and Wang, 2002176Evans and Wang, 2002; Baines and Earl, 199929
- Page 1 and 2: WDCS, the Whale and Dolphin Conserv
- Page 3 and 4: Towards Marine ProtectedAreas for C
- Page 5 and 6: South west EnglandCeltic Deep - com
- Page 7 and 8: CONTENTSExecutive Summary 2Acknowle
- Page 9 and 10: 1. INTRODUCTIONWe enter an exciting
- Page 11 and 12: Note: This schematic map shows thev
- Page 13 and 14: 2. IDENTIFYING MARINE PROTECTED ARE
- Page 15 and 16: 3. CRITICAL HABITATInterest in the
- Page 17 and 18: Foraging, travel, socialising and p
- Page 19 and 20: 4. DATA AND SPECIES ACCOUNTSIn orde
- Page 21 and 22: Killer whale (Orcinus orca)Atlantic
- Page 23 and 24: 4.4 SPECIES ACCOUNTS4.4.1 HARBOUR P
- Page 25 and 26: Although there is some variation in
- Page 27 and 28: North ScotlandThe first SCANS surve
- Page 29: East EnglandThe seasonal pattern in
- Page 33 and 34: 4.4.2 BOTTLENOSE DOLPHIN (Tursiops
- Page 35 and 36: This expansion in range means this
- Page 37 and 38: 4.4.3 SHORTBEAKED COMMON DOLPHIN (D
- Page 39 and 40: South west EnglandBased on observat
- Page 41 and 42: 4.4.4 COMMON MINKE WHALE (Balaenopt
- Page 43 and 44: Sightings for this area of the Oute
- Page 45 and 46: The JNCC Atlas of cetacean distribu
- Page 47 and 48: 4.4.5 WHITEBEAKED DOLPHIN (Lagenorh
- Page 49 and 50: 4.4.6 RISSO'S DOLPHIN (Grampus gris
- Page 51 and 52: 4.4.7 KILLER WHALE OR ORCA (Orcinus
- Page 53 and 54: 4.4.8 ATLANTIC WHITESIDED DOLPHIN (
- Page 55 and 56: 4.4.9 LONGFINNED PILOT WHALE (Globi
- Page 57 and 58: 4.4.10 SPERM WHALE (Physeter macroc
- Page 59 and 60: 4.4.11 BEAKED WHALESNORTHERN BOTTLE
- Page 61 and 62: which looked at worldwide beaked wh
- Page 63 and 64: There is no current population esti
- Page 65 and 66: 4.5 SCOTTISH MPA GUIDELINES - STAGE
- Page 67 and 68: Northern bottlenose whale and Sower
- Page 69 and 70: Fisheries BycatchFisheries bycatch
- Page 71 and 72: 5.2 AN OVERVIEW OF REGIONAL THREATS
- Page 73 and 74: North ScotlandAs in the west, threa
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- Page 77 and 78: 6.1.1 HARBOUR PORPOISEWest and sout
- Page 79 and 80: Assessment: Critical HabitatHigh re
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11. Area: Off Land's End, Cornwall
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Threats, status and relative import
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East Scotland3. Area: North east Sc
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Threats, status and relative import
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6.1.4 COMMON MINKE WHALEWest and so
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6.1.5 WHITEBEAKED DOLPHINWest and s
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Coastal Wales2. Area: Bardsey Islan
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6.1.8 ATLANTIC WHITESIDED DOLPHINNo
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Other areasOne other area was asses
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6.1.11 BEAKED WHALESThe northern Ro
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7. PROTECTING CRITICAL HABITAT - RE
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considered in the area, each of whi
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7.4 EAST SCOTLANDWithin the east Sc
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7.5 SOUTH WEST ENGLANDWithin the so
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An SAC is already in place for bott
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7.9 SUMMARYFour areas of identified
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iv. Licensing processes for industr
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9. IDENTIFICATION AND PRIORITISATIO
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Recommendations on particular data
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Coastal Wales northern Pembrokeshir
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ANNEX II SPECIES ACCOUNTS SUMMARY
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HARBOUR PORPOISEEast ScotlandStudyH
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HARBOUR PORPOISESouth West EnglandS
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HARBOUR PORPOISECoastal WalesStudyT
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BOTTLENOSE DOLPHINStudyTemporalReso
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SHORTBEAKED COMMON DOLPHINStudyTemp
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SHORTBEAKED COMMON DOLPHINStudyTemp
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COMMON MINKE WHALEWest ScotlandStud
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WHITEBEAKED DOLPHINStudyTemporalRes
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KILLER WHALEStudyTemporalResolution
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ATLANTIC WHITESIDED DOLPHINStudyTem
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SPERM WHALEStudyTemporalResolutionS
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BALEEN WHALESStudyTemporalResolutio
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ANNEX III THREATS TO CETACEANS - RE
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Actual orPotential ThreatActivitySp
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North ScotlandSpecies of most relev
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East EnglandSpecies of most relevan
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South East EnglandSpecies of most r
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South West EnglandSpecies of most r
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Irish SeaSpecies of most relevance
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Actual orPotential ThreatActivitySp
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Bravington, M., Borchers, D. and No
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Evans, P.G.H. and Anderwald, P. 200
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ICES WGMME. 2006. Report of the Wor
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ECS/ASCOBANS/ACCOBAMS Workshop held
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marine protected areas effective to
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ISBN: 9781901386233