West ScotlandNumerous studies over a long time period have noted high relative abundance of white-beakeddolphins for the northern Hebrides, particularly the northern Minch 345 . There are fewer dataavailable for the winter months but it appears that white-beaked dolphins use this area throughoutthe year. The waters north of here, extending east to Orkney, along with the western Sea of theHebrides, also have reportedly high relative abundance of white-beaked dolphins although thereare fewer data available 346 . Studies in The Minch have reported calves to be present in 60% ofgroups in the summer months 347 , indicating the area provides calving and nursing habitat forwhite-beaked dolphins.Based on the reports of some sightings surveys, and on strandings records, it has been suggestedthat there has been a decrease in abundance of white-beaked dolphins for the west coast ofScotland 348 . A decrease has not been conclusively proven yet, and some recent surveys havecontinued to record high encounter rates and abundance for white-beaked dolphins in this area 349 .This suspected decrease has coincided with an apparent increase in recent years of commondolphin sightings in the region. Common dolphins are typically a warmer water species, andthese observations have led to the suggestion that these may be changes linked to risingsea temperatures 350 .Studies looking at the pattern of white-beaked dolphin strandings for the whole of the UK suggestchanges may be occurring over a wider area than the west of Scotland, with an apparent northwardshift in the location of strandings 351 . Decreases in the number of reported strandings have beennoted for all regions except the north east of Scotland 352 .Longer-term monitoring of distribution and abundance will be needed to confirm the statusof white-beaked dolphins and to determine the nature of any possible shifts in distribution.East ScotlandA seasonal inshore movement of white-beaked dolphins has been suggested for the summermonths 353 . There appears to be evidence of this for the east coast 354 , and particularly offAberdeenshire 355 where studies have shown strong seasonality with peaks in the summer months.Recorded white-beaked dolphin strandings also peak during the summer months 356 suggesting aninshore movement of animals at this time of year. More in-depth analysis of strandings data suggeststhat females move into inshore waters first, to be followed later by males 357 . This earlier peak infemale strandings coincides with a peak in calf strandings and this has led to the suggestion thatfemales make this movement to inshore waters to give birth 358 . The later increase in strandings ofmale white-beaked dolphins may signify their arrival into the area in order to breed with femalesthat have calved 359 .Sightings records of calves in UK waters support this timing, with studies reporting their presenceonly in the summer months 360 . As with most cetacean species, data on calf presence are generallyscarce and hindered by difficulties in sighting them. However, a study in Aberdeenshire coastalwaters reported calves to be present in 32% of groups in the summer months 361 . This furtherindicates that this area provides calving and nursing habitat for white-beaked dolphins.The observed summer peaks in sighting frequencies and possible inshore movement could also berelated to changes in prey distribution and abundance. White-beaked dolphins take a wide varietyof fish but cod, haddock, whiting and hake have been found to be the predominant species in thestomachs of stranded animals 362 . The North Sea provides habitat including spawning grounds formany of these fish species 363 and their distribution and abundance will in turn affect that ofwhite-beaked dolphins. Anecdotal evidence has been put forward suggesting white-beakeddolphins follow mackerel or herring into inshore waters 364 but so far, no particular movements orconcentrations of favoured prey species have been correlated with the observed distribution andincreased sighting frequencies of white-beaked dolphins 365 .46345Reid et al, 2003; Northridge et al, 1995; Evans et al, 2003; MacLeod, 2001; Pollock et al, 2000; Weir et al, 2009;Shrimpton and Parsons, 2000; Gill et al, 1997; Boran et al, 1999346Reid et al, 2003; Northridge et al, 1995; Evans et al, 2003; MacLeod, 2001; Pollock et al, 2000; Shrimpton andParsons, 2000; Boran et al, 1999347Weir, 2008348MacLeod et al, 2005; Evans, 1992349Weir, 2008; SCANS II, 2006350MacLeod et al, 2005; Weir et al, 2009351Canning et al, 2008; Jepson, 2006 (in JNCC, 2007)352Canning et al, 2008353Evans, 1992354Reid et al, 2003355Canning et al, 2008; Weir et al, 2007; Evans et al, 2008356Canning et al, 2008357Ibid358Ibid359Ibid360MacLeod, 2001; Weir et al, 2007; Pollock et al, 2000; Canning et al, 2008361Weir et al, 2007362Reid et al, 2003; Canning et al, 2008363CEFAS, 2001364Weir et al, 2007; Canning et al, 2008; Evans, 1980365Canning et al, 2008
4.4.6 RISSO'S DOLPHIN (Grampus griseus)Risso's dolphins have a wide distribution through temperate and tropical regions of the world 366 ; theUK's waters are at the northern limit of the species’ range. The status of Risso's dolphins in the UKis currently unknown, and there are no population estimates available. Large scale surveys such asSCANS and SCANS II made insufficient sightings to be able to estimate abundance for any areasof UK waters 367 .Preliminary results from genetic studies on this species have indicated that UK Risso'sdolphins are genetically different from the better-studied Mediterranean animals which have lessgenetic diversity 368 .In UK waters they tend to be seen in small- to medium-sized groups, most commonly 6-12individuals but between 2-50 could be regarded as typical 369 . In the North Atlantic they are at timesobserved swimming with other cetaceans including long-finned pilot whales, bottlenose dolphins,and white-beaked and Atlantic white-sided dolphins 370 .Life history information is generally scarce. A peak in calving has been noted for the wintermonths 371 in some locations but researchers suggest that in Scottish waters, births are likely tooccur between July and December 372 .Typical prey for Risso's dolphins are neritic and oceanic squid 373 ; they may also feed oncrustaceans 374 and octopus 375 .In most places in the world, Risso's dolphins show a strong preference for the deep waters of thecontinental slope 376 . It is believed they take advantage of the highly-productive frontal systems thatoceanographic mechanisms create near the shelf break 377 . In the UK, however, they are found incoastal areas, over the continental shelf and in some cases, close to the shore of some islands 378 .They are most common off the Outer Hebrides of west Scotland, and in parts of the Irish Sea 379 .Few sightings of Risso's dolphins are made on the east coast of the UK or in the Channel, exceptfor its westernmost parts 380 .This species is present in UK waters all year round but with some seasonal patterns of occurrence.They are seen in greater numbers in shelf waters from May to October, and during the remainder ofthe year sightings are greater in offshore areas 381 .West ScotlandThe Outer Hebrides in north west Scotland, and particularly the Isle of Lewis, have been shown tohave significant concentrations of Risso's dolphins. Sightings indicate possible year-round residencybut are most frequent over the summer and autumn months, with clear peaks in numbers inAugust and September 382 . The suggestion that at least a part of this population is resident hereis supported by studies off the north east of Lewis that have repeatedly resighted individuals 383 .A photo-ID study conducted over two years identified 142 individuals, with at least 52 animalsresighted between years 384 . In this area, Risso's dolphins seem generally to spend May and Julyfurther offshore in deeper waters and in larger groups, moving into nearshore habitat to forage insmaller groups for August and September. This movement is possibly linked to prey availability asthe lesser octopus (Eledone cirrhosa), thought to be a prey species for these Risso's dolphins 385 ,is abundant in Scottish coastal waters during August and September 386 .366Baird, 2002367Hammond et al, 1995; SCANS II, 2006368Gaspari et al, 2007369Reid et al, 2003370Ibid371Baird, 2002372Atkinson and Gill, 1996 (in Wharam and Simmonds, 2008)373Baird, 2002; Culik, 2004374Culik, 2004375Atkinson et6 al, 1998376Baumgartner, 1997; Mangion and Gannier, 2002; Casacci and Gannier, 2000377Baumgartner, 1997378Gaspari et al, 2007; Reid et al, 2003379Reid et al, 2003380Evans et al, 2003; Kiszka et al, 2007381Reid et al, 2003; Evans et al, 2003382Wharam and Simmonds, 2008; Pollock et al, 2000; Atkinson et al, 1998383Atkinson et al, 1998384Ibid385A stranded animal had lesser octopus beaks in its stomach Atkinson et al, 1998386Atkinson et al, 1998; Hastie et al, 200647
- Page 1 and 2: WDCS, the Whale and Dolphin Conserv
- Page 3 and 4: Towards Marine ProtectedAreas for C
- Page 5 and 6: South west EnglandCeltic Deep - com
- Page 7 and 8: CONTENTSExecutive Summary 2Acknowle
- Page 9 and 10: 1. INTRODUCTIONWe enter an exciting
- Page 11 and 12: Note: This schematic map shows thev
- Page 13 and 14: 2. IDENTIFYING MARINE PROTECTED ARE
- Page 15 and 16: 3. CRITICAL HABITATInterest in the
- Page 17 and 18: Foraging, travel, socialising and p
- Page 19 and 20: 4. DATA AND SPECIES ACCOUNTSIn orde
- Page 21 and 22: Killer whale (Orcinus orca)Atlantic
- Page 23 and 24: 4.4 SPECIES ACCOUNTS4.4.1 HARBOUR P
- Page 25 and 26: Although there is some variation in
- Page 27 and 28: North ScotlandThe first SCANS surve
- Page 29 and 30: East EnglandThe seasonal pattern in
- Page 31 and 32: Irish SeaOutside of coastal Welsh w
- Page 33 and 34: 4.4.2 BOTTLENOSE DOLPHIN (Tursiops
- Page 35 and 36: This expansion in range means this
- Page 37 and 38: 4.4.3 SHORTBEAKED COMMON DOLPHIN (D
- Page 39 and 40: South west EnglandBased on observat
- Page 41 and 42: 4.4.4 COMMON MINKE WHALE (Balaenopt
- Page 43 and 44: Sightings for this area of the Oute
- Page 45 and 46: The JNCC Atlas of cetacean distribu
- Page 47: 4.4.5 WHITEBEAKED DOLPHIN (Lagenorh
- Page 51 and 52: 4.4.7 KILLER WHALE OR ORCA (Orcinus
- Page 53 and 54: 4.4.8 ATLANTIC WHITESIDED DOLPHIN (
- Page 55 and 56: 4.4.9 LONGFINNED PILOT WHALE (Globi
- Page 57 and 58: 4.4.10 SPERM WHALE (Physeter macroc
- Page 59 and 60: 4.4.11 BEAKED WHALESNORTHERN BOTTLE
- Page 61 and 62: which looked at worldwide beaked wh
- Page 63 and 64: There is no current population esti
- Page 65 and 66: 4.5 SCOTTISH MPA GUIDELINES - STAGE
- Page 67 and 68: Northern bottlenose whale and Sower
- Page 69 and 70: Fisheries BycatchFisheries bycatch
- Page 71 and 72: 5.2 AN OVERVIEW OF REGIONAL THREATS
- Page 73 and 74: North ScotlandAs in the west, threa
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- Page 77 and 78: 6.1.1 HARBOUR PORPOISEWest and sout
- Page 79 and 80: Assessment: Critical HabitatHigh re
- Page 81 and 82: 11. Area: Off Land's End, Cornwall
- Page 83 and 84: Threats, status and relative import
- Page 85 and 86: East Scotland3. Area: North east Sc
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- Page 89 and 90: 6.1.4 COMMON MINKE WHALEWest and so
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- Page 97 and 98: Other areasOne other area was asses
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6.1.11 BEAKED WHALESThe northern Ro
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7. PROTECTING CRITICAL HABITAT - RE
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considered in the area, each of whi
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7.4 EAST SCOTLANDWithin the east Sc
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7.5 SOUTH WEST ENGLANDWithin the so
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An SAC is already in place for bott
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7.9 SUMMARYFour areas of identified
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iv. Licensing processes for industr
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9. IDENTIFICATION AND PRIORITISATIO
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Recommendations on particular data
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Coastal Wales northern Pembrokeshir
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ANNEX II SPECIES ACCOUNTS SUMMARY
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HARBOUR PORPOISEEast ScotlandStudyH
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HARBOUR PORPOISESouth West EnglandS
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HARBOUR PORPOISECoastal WalesStudyT
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BOTTLENOSE DOLPHINStudyTemporalReso
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SHORTBEAKED COMMON DOLPHINStudyTemp
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SHORTBEAKED COMMON DOLPHINStudyTemp
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COMMON MINKE WHALEWest ScotlandStud
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WHITEBEAKED DOLPHINStudyTemporalRes
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KILLER WHALEStudyTemporalResolution
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ATLANTIC WHITESIDED DOLPHINStudyTem
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SPERM WHALEStudyTemporalResolutionS
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BALEEN WHALESStudyTemporalResolutio
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ANNEX III THREATS TO CETACEANS - RE
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Actual orPotential ThreatActivitySp
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North ScotlandSpecies of most relev
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East EnglandSpecies of most relevan
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South East EnglandSpecies of most r
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South West EnglandSpecies of most r
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Irish SeaSpecies of most relevance
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Actual orPotential ThreatActivitySp
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Bravington, M., Borchers, D. and No
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Evans, P.G.H. and Anderwald, P. 200
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ICES WGMME. 2006. Report of the Wor
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ECS/ASCOBANS/ACCOBAMS Workshop held
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marine protected areas effective to
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ISBN: 9781901386233