Juveniles have been sighted in this region between March and November 387 . Off the north eastcoast of Lewis, whole groups comprised exclusively of sub-adults or juveniles were noted onseveral occasions, and a group consisting of eight females, each with a calf, has also beenobserved 388 . Sightings such as these strongly suggest this area has importance for breeding,nursing and/or raising young.North east Lewis is the focus of Risso's dolphins activity in this area but high sightings rates arereported for the rest of the western part of the Hebrides during the summer and there may befurther areas of importance in these less studied waters.There are generally few data available about Risso's dolphins in more offshore waters but sightingsoccur mostly during autumn and winter and are concentrated along the continental slope 389 .Sightings are too few to suggest particular areas of importance.Coastal Wales and the Irish SeaCertain locations in the Irish Sea – the Isle of Man, Anglesey, Bardsey Island and the LleynPeninsula, and Pembrokeshire – record Risso's dolphin sightings relatively regularly 390 . BardseyIsland records the highest sightings rates, followed by northern Pembrokeshire 391 .The waters off Bardsey Island in North Wales have been subject to a small-scale, multi-year studyof its cetacean fauna and this has confirmed the area’s importance for Risso's dolphins 392 . Despitebeing predominantly a land-based survey, 133 individual Risso's dolphins have been catalogued,with a few resightings between years 393 . The survey has been conducted in April, July, August andSeptember and Risso's have been observed in every month. Bardsey Islanders report seeingRisso's at other times also but it is unknown whether Risso's are present all year round in coastalwaters, present all year but further offshore or only seasonally present.As in the Isle of Lewis study, Bardsey Island researchers have observed groups containing onlysub-adults and 'nursery groups' consisting of multiple females, each with a calf. High numbers ofyoung animals are seen here; 10% of photographed Risso's dolphins have been calves andjuveniles and 29% were identified as sub-adults 394 . Some apparently very young calves have beenobserved, including one with foetal folds 395 . These observations indicate that the area is importantto Risso's dolphins for breeding, nursing and/or raising calves.Behaviour indicative of feeding (such as deep dives or remaining submerged for a long period) hasbeen observed often around Bardsey 396 and prey availability is likely a driver for them being in thearea. It has been suggested that the complex topography and deep ocean trenches around Bardseymay create areas of upwelling with increased productivity that Risso's dolphins exploit 397 . The lesseroctopus, known prey for Risso's dolphins from stomach studies of stranded animals found in Walesand Scotland 398 , is found in the trenches offshore and reportedly comes inshore to lay its eggs onthe horse mussel reefs in this area 399 . Risso's dolphins have been sighted feeding over thesereefs 400 , possibly following their prey inshore.There is much less information available for the other locations in the Irish Sea where Risso'sdolphins are sighted. Young have been observed off Pembrokeshire and Anglesey between Julyand September 401 but beyond this, the relative importance of these locations is unclear.Further research on Risso’s dolphins is needed in the waters of north east Lewis andBardsey Island to improve our knowledge of these important populations. Expanding thephoto-ID catalogues in the surrounding areas will help us understand the wider-scalemovements and possible connections between these populations.48387Pollock et al, 2000388Atkinson et al, 1998389Reid et al, 2003; Pollock et al, 2000390Evans et al, 2003391Baines and Evans, 2009392WDCS, 2002, 2003, 2004, 2005a and 2006; De Boer, 2009393De Boer, 2009394Ibid395WDCS, 2004396WDCS, 2005a397Wharam and Simmonds, 2008398Atkinson et al, 1998; Wharam and Simmonds, 2008399Wharam and Simmonds, 2008400Ibid401Baines and Evans, 2009
4.4.7 KILLER WHALE OR ORCA (Orcinus orca)Killer whales are widely distributed over the continental shelf and the deep offshore waters of thenorth east Atlantic. In UK waters they are primarily seen north and west of Scotland and key areasappear to be the continental slope to the north and north west of Shetland, the northern North Seato the east of Shetland, and coastal waters of Shetland and the Hebrides 402 . Killer whales are alsoreported around Orkney, Farne Islands (north east England), the Isle of Man, in the St. George'sChannel and off south west England 403 , but there are few data available about these animals, andtheir occurrence at these locations appears to be more sporadic. The presence of killer whales offCornwall has been linked to that of basking sharks (Cetorhinus maximus), a potential preyspecies 404 . Elsewhere around the UK sightings are considered rare 405 .No overall population estimate exists for the north east Atlantic. Only recently has research begunto shed light on the population structure of killer whales in this region. Comparisons of morphologicaltraits, DNA and ecology of killer whales throughout the area have suggested that two types exist,with differing physical and ecological characteristics 406 . The first type appear to be generalists, feedingmainly on lipid-rich pelagic fish such as herring or mackerel, but with a subset of the populationalso persistently feeding on marine mammals 407 . The second type appears to be highly specialised,and it is suggested that their main prey may be other cetaceans 408 . Type one animals were foundacross the North Atlantic, from Norway to Newfoundland, whereas type two specimens were onlysampled from Scotland and the Faroe Islands 409 .These two types overlap geographically and temporally. The existence of sympatric populations ofkiller whales has been seen elsewhere in the world, most notably in the north east Pacific. Howeverin contrast, north east Pacific killer whales all appear to be highly specialised, either predating onfish or marine mammals, whereas in the north east Atlantic there seems to exist a 'generalist' typethat will exploit fish, marine mammals and even seabirds at different times.Killer whales have been observed in UK waters in every month of the year 410 . A strong peak insightings occurs during June and July in Shetland coastal waters 411 . This timing is highly correlatedwith the harbour seal pupping season; killer whales have been observed hunting and feeding uponharbour seals, as well as grey seals and harbour porpoises in this area 412 .Photo-ID and connectivity analysis conducted between killer whales from Iceland, Norway, theNorthern and Western Isles of Scotland and the northern North Sea, has indicated that there is ahigh degree of inter-annual site fidelity in all locations 413 . This work also concluded that there ismovement between the Northern Isles, Scotland and East Iceland, and that killer whales predatingseals around the Northern Isles are linked to those that exploit Icelandic herring 414 . This connectionis supported by isotope analysis which has suggested that subsets of individuals within the Icelandicherring- and mackerel-eating killer whale population also persistently forage on marine mammals415 . These initial investigations suggest that at least some of the killer whales foundpredating on seals off Shetland in the summer are of the 'generalist' type, and move to Icelandicwaters for the winter to exploit fish resources. A high degree of inter-annual site fidelity was foundfor animals around the Northern Isles 416 .A peak in sightings occurs over the continental slope north and west of Shetland during May andJune 417 , slightly earlier than the peak in Shetland waters. This earlier peak may reveal themovement of killer whales into the area, as they travel from winter feeding grounds around Iceland.402Evans et al, 2003; Pollock et al, 2000; Stone, 1997, 1998, 2000, 2001, 2003, 2006; Evans et al, 1996; Luque et al, 2006403Evans et al, 2003; Weir, 2002404Weir, 2002405Reid et al, 2003; Evans et al, 2003406Foote et al, 2009407Ibid408Ibid409Ibid410Evans et al, 2003411Bolt et al, 2009412Weir, 2002; Bolt et al, 2009; Foote et al, 2010413Foote et al, 2010414Ibid415Foote et al, 2009416Foote et al, 2010417Pollock et al, 2000; Reid et al, 2003 (monthly data)49
- Page 1 and 2: WDCS, the Whale and Dolphin Conserv
- Page 3 and 4: Towards Marine ProtectedAreas for C
- Page 5 and 6: South west EnglandCeltic Deep - com
- Page 7 and 8: CONTENTSExecutive Summary 2Acknowle
- Page 9 and 10: 1. INTRODUCTIONWe enter an exciting
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- Page 13 and 14: 2. IDENTIFYING MARINE PROTECTED ARE
- Page 15 and 16: 3. CRITICAL HABITATInterest in the
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7. PROTECTING CRITICAL HABITAT - RE
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considered in the area, each of whi
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7.4 EAST SCOTLANDWithin the east Sc
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7.5 SOUTH WEST ENGLANDWithin the so
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An SAC is already in place for bott
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7.9 SUMMARYFour areas of identified
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iv. Licensing processes for industr
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9. IDENTIFICATION AND PRIORITISATIO
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Recommendations on particular data
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Coastal Wales northern Pembrokeshir
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ANNEX II SPECIES ACCOUNTS SUMMARY
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HARBOUR PORPOISEEast ScotlandStudyH
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HARBOUR PORPOISESouth West EnglandS
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HARBOUR PORPOISECoastal WalesStudyT
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BOTTLENOSE DOLPHINStudyTemporalReso
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SHORTBEAKED COMMON DOLPHINStudyTemp
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SHORTBEAKED COMMON DOLPHINStudyTemp
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COMMON MINKE WHALEWest ScotlandStud
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WHITEBEAKED DOLPHINStudyTemporalRes
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KILLER WHALEStudyTemporalResolution
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ATLANTIC WHITESIDED DOLPHINStudyTem
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SPERM WHALEStudyTemporalResolutionS
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BALEEN WHALESStudyTemporalResolutio
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ANNEX III THREATS TO CETACEANS - RE
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Actual orPotential ThreatActivitySp
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North ScotlandSpecies of most relev
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East EnglandSpecies of most relevan
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South East EnglandSpecies of most r
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South West EnglandSpecies of most r
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Irish SeaSpecies of most relevance
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Actual orPotential ThreatActivitySp
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Bravington, M., Borchers, D. and No
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Evans, P.G.H. and Anderwald, P. 200
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ICES WGMME. 2006. Report of the Wor
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ECS/ASCOBANS/ACCOBAMS Workshop held
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marine protected areas effective to
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ISBN: 9781901386233