Porpoises are present throughout the year in southern Cardigan Bay but with slight peaks duringthe winter months 177 . They are widespread throughout the inshore and offshore areas but havebeen found to have some preferred areas — densities on the whole are higher in the southern partof Cardigan Bay than further north 178 , and presence is greatest around New Quay head, Aberporth,Ynys Lochtyn, and between Cemaes Head and Ceibwr Bay 179. Abundance has been estimated atbetween 167 and 214 for harbour porpoises in the southern Cardigan Bay SAC 180 .Along the south coast of Wales, the Gower Peninsula has also been highlighted as an area ofpossible importance to harbour porpoises as it was found to have concentrations in some monthsof the year 181 . There are few data available for this region but one study looking at fine scaledistribution found from acoustic and visual surveys that Mumbles Head and Burry Holms on theGower Peninsula were high-use areas 182 .To the north coast of Wales, analysis of multiple-year data has found the Isle of Anglesey and theLleyn Peninsula to have high relative densities of harbour porpoises, with concentrations for severalmonths of the year 183 . A higher than average proportion of calves to adults was found to occur offAnglesey in July, suggesting the area’s importance for calving.Studies of the waters around Anglesey have found them to support high densities of harbourporpoises, particularly Point Lynas and South Stacks on the north and west of the island 184 .A three-year study (May-September) recorded the highest densities of porpoises at Point Lynas, anarea of strong tidal currents and races, along with a high degree of aggregation of animals at thissite 185 . Based on this, and that other studies have observed large aggregations of porpoises feedingin the area, researchers suggest it is likely an important foraging and feeding site for harbourporpoises. At South Stacks, porpoises were not found to be so highly aggregated and behaviouraldata for the area is not currently available, so conclusions about site use are harder to reach.However, high densities of porpoises are found here and the area contains the oceanographicfeatures (tidal races and eddies for example) that seem to facilitate foraging for harbour porpoisesso it may also be a site used for feeding 186 .The Lleyn Peninsula and Bardsey Island just off the coast have been the subject of study for anumber of years in recognition of the high numbers of porpoises found in these waters 187 . Mostsurvey work has taken place in spring and summer, with the highest levels of sightings in April andlate July-September, and the lowest sighting levels in early July 188 . April seems to be a particularlyfavoured time of year for porpoises to aggregate and feed in the Sound that separates BardseyIsland from the mainland 189 . The waters around Bardsey Island are influenced strongly by the fastand complex tidal currents that occur and porpoise activity was found to be related to tidal state,with peaks observed during the low water period, and following the neap tide 190 . As at otherlocations, porpoises were observed holding their position into the current, apparently using theconditions to aid feeding. Lower numbers recorded during spring tides are possibly due to the tidesbecoming too strong and energetically demanding for porpoises to forage.In addition to feeding, the area is also believed to be important as a porpoise breeding and nursingsite 191 . A high proportion of sightings (10-24%) are females with young, particularly in August butalso September and July 192 . They appear to favour the calmer, inshore waters, an observation thathas been made at other sites (e.g. Ramsey Sound, south Wales).Analysis of five years of stranding data from around England and Wales found that strandings ofneonate porpoises occurred more often in Wales than anywhere else, suggesting that CardiganBay and west Wales is an important breeding and calving ground for harbour porpoises 193 .Little information for this area is available from outside the spring and summer months. One surveythat took place in the autumn of 2002 and included the waters of northern Cardigan Bay noted acluster of harbour porpoise sightings in the northern Cardigan Bay-Bardsey Island area, indicatingthat this area may still be important to harbour porpoises later in the year 194 .30177Pesante et al, 2008178Although there has been much more survey effort in southern Cardigan Bay179Pesante et al, 2008180Ibid181Meaning it was found to have porpoises present in at least three months of the year, with concentrations in atleast two months although not necessarily during the AprilSeptember period (when calving takes place), andwith records over several years182Watkins and Colley, 2004183Evans and Wang, 2002; Baines and Evans, 2009184Shucksmith et al, 2008185Ibid186Ibid187WDCS, 2002, 2003, 2004, 2005a, 2006; De Boer and Simmonds, 2003188WDCS, 2005a189Ibid190Ibid191Ibid192Ibid193Bennett et al, 2002194De Boer and Simmonds, 2003
4.4.2 BOTTLENOSE DOLPHIN (Tursiops truncatus)Bottlenose dolphins in the North Atlantic appear to consist of two forms, a coastal and an offshoreform. The better known coastal form is locally common in the Irish Sea and off north east Scotland,and in smaller numbers in the Hebrides (west Scotland), and in the western half of the Channel, offsouth west England. It is present in coastal UK waters year-round but numbers peak in most placesbetween July and October. Coastal bottlenose dolphins have a varied diet that includes several fishspecies, including cod (Gadus morhua), salmon (Salmo salar) and whiting, and they will also preyon invertebrates 195 .No overall population estimate exists for bottlenose dolphins in UK coastal waters but due to theirrelatively predictable and coastal habitat preferences, and suitability for photo-ID studies, goodpopulation data are available for the best known populations. The east coast of Scotland bottlenosedolphin population is estimated to number 193 animals (95% Probability Interval = 162-245) 196while the Cardigan Bay (Irish Sea) population is estimated to vary between 154 and 248 197 (seeFigure 4.8, p.31). The less well studied population of dolphins inhabiting the west coast of Scotland(and consisting of two communities) apparently numbers about 45 animals (95% Probability Interval =33-66) 198 . A poorly-known group of bottlenose dolphins appears to be resident to the waters ofsouth west England and photo-ID studies conducted during the 1990s documented at least 50animals 199 . Genetic testing of these coastal populations has indicated that a level of populationstructure exists and they do not form one single randomly mating population. However, there is notenough genetic divergence to suggest these populations are completely isolated from each other;there is at least a low level of gene flow occurring 200 .Little is known about the offshore form of bottlenose dolphins, including the relationship betweenthe offshore and coastal forms. On the other side of the Atlantic, off the US east coast, whereoffshore and coastal forms of bottlenose dolphins are also observed, they appear to be ecologicallyand genetically discrete from each other 201 . There are indications that this situation is repeatedacross the North Atlantic 202 but further research is needed to confirm that this is the case in UKwaters. Offshore bottlenose dolphins are thought to prey on mesopelagic fish and cephalopods.They are frequently found in mixed pods with pilot whales, and this is thought to be because of ashared preference for squid 203 . No population estimate exists for the offshore form of bottlenosedolphins in UK waters.195Culik, 2004196Thompson et al, IN PREP197Pesante et al, 2008198Thompson et al, IN PREP199Wood, 1998200Thompson et al, IN PREP201Hoelzel et al, 1998202Quérouil et al, 2007203Skov et al, 199531
- Page 1 and 2: WDCS, the Whale and Dolphin Conserv
- Page 3 and 4: Towards Marine ProtectedAreas for C
- Page 5 and 6: South west EnglandCeltic Deep - com
- Page 7 and 8: CONTENTSExecutive Summary 2Acknowle
- Page 9 and 10: 1. INTRODUCTIONWe enter an exciting
- Page 11 and 12: Note: This schematic map shows thev
- Page 13 and 14: 2. IDENTIFYING MARINE PROTECTED ARE
- Page 15 and 16: 3. CRITICAL HABITATInterest in the
- Page 17 and 18: Foraging, travel, socialising and p
- Page 19 and 20: 4. DATA AND SPECIES ACCOUNTSIn orde
- Page 21 and 22: Killer whale (Orcinus orca)Atlantic
- Page 23 and 24: 4.4 SPECIES ACCOUNTS4.4.1 HARBOUR P
- Page 25 and 26: Although there is some variation in
- Page 27 and 28: North ScotlandThe first SCANS surve
- Page 29 and 30: East EnglandThe seasonal pattern in
- Page 31: Irish SeaOutside of coastal Welsh w
- Page 35 and 36: This expansion in range means this
- Page 37 and 38: 4.4.3 SHORTBEAKED COMMON DOLPHIN (D
- Page 39 and 40: South west EnglandBased on observat
- Page 41 and 42: 4.4.4 COMMON MINKE WHALE (Balaenopt
- Page 43 and 44: Sightings for this area of the Oute
- Page 45 and 46: The JNCC Atlas of cetacean distribu
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- Page 51 and 52: 4.4.7 KILLER WHALE OR ORCA (Orcinus
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- Page 55 and 56: 4.4.9 LONGFINNED PILOT WHALE (Globi
- Page 57 and 58: 4.4.10 SPERM WHALE (Physeter macroc
- Page 59 and 60: 4.4.11 BEAKED WHALESNORTHERN BOTTLE
- Page 61 and 62: which looked at worldwide beaked wh
- Page 63 and 64: There is no current population esti
- Page 65 and 66: 4.5 SCOTTISH MPA GUIDELINES - STAGE
- Page 67 and 68: Northern bottlenose whale and Sower
- Page 69 and 70: Fisheries BycatchFisheries bycatch
- Page 71 and 72: 5.2 AN OVERVIEW OF REGIONAL THREATS
- Page 73 and 74: North ScotlandAs in the west, threa
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- Page 77 and 78: 6.1.1 HARBOUR PORPOISEWest and sout
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Threats, status and relative import
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East Scotland3. Area: North east Sc
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Threats, status and relative import
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6.1.4 COMMON MINKE WHALEWest and so
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6.1.5 WHITEBEAKED DOLPHINWest and s
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Coastal Wales2. Area: Bardsey Islan
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6.1.8 ATLANTIC WHITESIDED DOLPHINNo
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Other areasOne other area was asses
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6.1.11 BEAKED WHALESThe northern Ro
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7. PROTECTING CRITICAL HABITAT - RE
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considered in the area, each of whi
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7.4 EAST SCOTLANDWithin the east Sc
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7.5 SOUTH WEST ENGLANDWithin the so
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An SAC is already in place for bott
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7.9 SUMMARYFour areas of identified
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iv. Licensing processes for industr
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9. IDENTIFICATION AND PRIORITISATIO
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Recommendations on particular data
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Coastal Wales northern Pembrokeshir
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ANNEX II SPECIES ACCOUNTS SUMMARY
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HARBOUR PORPOISEEast ScotlandStudyH
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HARBOUR PORPOISESouth West EnglandS
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HARBOUR PORPOISECoastal WalesStudyT
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BOTTLENOSE DOLPHINStudyTemporalReso
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SHORTBEAKED COMMON DOLPHINStudyTemp
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SHORTBEAKED COMMON DOLPHINStudyTemp
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COMMON MINKE WHALEWest ScotlandStud
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WHITEBEAKED DOLPHINStudyTemporalRes
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KILLER WHALEStudyTemporalResolution
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ATLANTIC WHITESIDED DOLPHINStudyTem
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SPERM WHALEStudyTemporalResolutionS
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BALEEN WHALESStudyTemporalResolutio
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ANNEX III THREATS TO CETACEANS - RE
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Actual orPotential ThreatActivitySp
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North ScotlandSpecies of most relev
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East EnglandSpecies of most relevan
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South East EnglandSpecies of most r
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South West EnglandSpecies of most r
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Irish SeaSpecies of most relevance
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Actual orPotential ThreatActivitySp
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Bravington, M., Borchers, D. and No
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Evans, P.G.H. and Anderwald, P. 200
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ICES WGMME. 2006. Report of the Wor
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ECS/ASCOBANS/ACCOBAMS Workshop held
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marine protected areas effective to
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ISBN: 9781901386233