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Abschlußbericht zum Forschungsprojekt ST121 - Technische ...

Abschlußbericht zum Forschungsprojekt ST121 - Technische ...

Abschlußbericht zum Forschungsprojekt ST121 - Technische ...

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XII<br />

Abstract<br />

all examined trees, independent of the soil properties of the planting site. Moreover, the<br />

overall root lengths (diameters >5 mm) on the up-slope side of the trees were shorter than<br />

those of roots orientated to the down-slope side. In the long-run, this could endanger the<br />

stability of afforested redevelopment areas against snow movement.<br />

Both the seedlings on Ah horizons and the planted as well as the naturally regenerated spruces<br />

on “Mull”-rendzinas showed limited biomass development (for the seedlings) and/or needle<br />

weights and therefore a low overall vitality. Despite concentration effects in the small needles,<br />

the spruces on “Mull”-rendzinas showed heavy nutrient deficiencies especially for N, P, Mn,<br />

and rarely for K. On account of the low total needle masses, their internal nutrient stores were<br />

small and correspondingly the shoot increments were low. Higher N-concentrations and more<br />

easily decomposable N-forms in the organic layers would lead to an enhanced N-availability.<br />

In addition, organic layers were characterised by higher contents of extractable P and K than<br />

the mineral soils. Due to acid pH-values and a decreasing (Ca+Mg)/K ratio and therefore a<br />

decreasing ion-antagonism for K, P- and K-availability was higher in organic layers. In<br />

addition, the Mn-availability increased in organic layers. Thus, seedlings and naturally<br />

regenerated plants on thick, acid organic layers showed a higher vitality and possessed a<br />

higher total biomass (seedlings) and needle mass than spruces on “Mull”-rendzinas.<br />

Moreover, their larger needles indicated a less pronounced deficiency of N, P, and rarely of K<br />

and of Mn and their shoot increments were higher on account of the ability to translocate<br />

nutrients from the higher nutrient pool in the parenchyma. In addition, highly decayed<br />

deadwood was very favourable with regard to the nutrition and the biomass development of<br />

the seedlings. Manuring measures resulted only in a short-term increase of needle nutrient<br />

concentrations and growth rates. Moreover, the manuring effects were more marked on vital<br />

trees growing on “Moder” sites.<br />

The abundance of ectomycorrhizae (ECM) was highest in the organic soil layers of “Moder”rendzinas<br />

compared to the mineral soil horizons. The ordination of the ECM communities and<br />

of the exploration types revealed segregation in dependency on differences in soil horizon<br />

properties. While Cenococcum geophilum preferred the organic soil layers, Lactarius spp.,<br />

Tomentella spp., and Craterellus tubaeformis were generally most abundant in the mineral<br />

soil horizons. C. geophilum was the predominant species, possibly due to enhanced<br />

competitiveness under the prevailing site conditions. The short-distance exploration types<br />

preferentially colonised the organic soil layer, whereas the contact types together with<br />

medium-distance types were primarily associated with the underlying A-horizons. The spatial<br />

niche differentiation of ECM genera and exploration types was probably caused by the soil

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