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american samoa - University of Hawaii at Manoa

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ipinn<strong>at</strong>e lamina, lanceol<strong>at</strong>e pinnules 10-20 cm long<br />

having an enlarged base, and sori <strong>of</strong> 7-12 sporangia<br />

free from each other, in a dense row 1-2 mm from<br />

the margins. Common in lowland to cloud forest,<br />

reported from near sea level to 1400 m elev<strong>at</strong>ion.<br />

Indigenous, ranging from Malaysia to Tahiti. Saf-<br />

Sortl 404 I; McMullin 17; Comstock s.n.; Sctchell<br />

38, 207; Eames 727; Garber 838; Long 1950; Whi-<br />

stler 8975!, 9125.<br />

Angiopteris opaca Copeland<br />

nase<br />

Large terrestrial fern similar to the preceding spe-<br />

cies, but with rachis and stipe densely scaly r<strong>at</strong>her<br />

than bearing sc<strong>at</strong>tered hairs. Occasional in lowland<br />

to montane forest, reported from 50 to 600 m eleva-<br />

tion. Indigenous, also found in Fiji. Christophersen<br />

99 1 ; Whistler 9047!, 9198.<br />

Mar<strong>at</strong>tia smithii Mett.<br />

nase<br />

Large terrestrial fern similar to the preceding two<br />

species, but with sporangia fused l<strong>at</strong>erally to form a<br />

single structure. Uncommon in montane forest,<br />

reported from 400 to 600 m elev<strong>at</strong>ion. Indigenous,<br />

ranging from Vanu<strong>at</strong>u to Samoa. Wilder s.n.;<br />

Sledge 1823; Whistler 8734!, 9090,9197.<br />

OPHIOGLOSSACEAE<br />

Ophioglossum pendulum L.<br />

Epiphytic fern with a small rhizome, pendulous,<br />

strap-shaped fronds 40-120 cm long, and sori<br />

embedded in a spike <strong>at</strong>tached to the center <strong>of</strong> the<br />

frond. Uncommon in coastal to montane forest,<br />

reported from near sea level to 550 m elev<strong>at</strong>ion.<br />

Indigenous, ranging from Madagascar to Hawai'i.<br />

Lamoureux 3001; Sledge 1822; Whistler 2818,<br />

2926, 300 1, 8746!.<br />

*Ophioglossum reticul<strong>at</strong>um L.<br />

Tiny terrestrial fern with a single ov<strong>at</strong>e lamina and<br />

sori embedded in a single erect spike. Uncommon<br />

in lowland to cloud forest, reported from 300 to<br />

1700 m elev<strong>at</strong>ion. Indigenous, ranging from tropi-<br />

cal Asia to Polynesia. Powell 127.<br />

POLYPODIACEAE<br />

*Dipteris conjug<strong>at</strong>a Reinwardt<br />

Large terrestrial fern with a creeping rhizome, re-<br />

mote stipes,veins dichotomously branched, lamina<br />

divided to the base into two spreading halves, these<br />

divided into four lobes which arc further divided,<br />

and numerous sori sc<strong>at</strong>tered on the lower surface.<br />

Locally abundant in montane scrub, reported from<br />

500 to 650 m elev<strong>at</strong>ion. Indigenous, ranging from<br />

Thailand to Samoa. Reinecke 173; Setchell 380;<br />

Garber 784; Christophersen 1034, 3564; Sledge<br />

1832; Whistler 2714,9162.<br />

*Lemmaphyllum accedens (Bl.) Donk<br />

Small epiphytic fern with a long-creeping rhizome,<br />

short stipes, dimorphic fronds, the sterile ones<br />

elliptic to ov<strong>at</strong>e, the fertile ones similar but with an<br />

<strong>at</strong>tenu<strong>at</strong>e tip, and sori in a single row on both sides<br />

<strong>of</strong> the midrib <strong>of</strong> the upper <strong>at</strong>tenu<strong>at</strong>e part. Occasional<br />

in lowland to montane forest, reported from 300 to<br />

900 m elev<strong>at</strong>ion. Indigenous, ranging from Malaysia<br />

to Samoa. Graeffe 459 (n.s.); Powell 93? ("Manua<br />

and Tutuila"); Whistler 9093, 9186.<br />

Microsorium sylv<strong>at</strong>icum (Bracken.) Copeland<br />

Large terrestrial fern with a creeping rhizome, sim-<br />

ple, deeply lobed, simply pinn<strong>at</strong>e fronds up to 2 m<br />

in height, and numerous tiny sori sc<strong>at</strong>tered<br />

irregularly over the lower surface <strong>of</strong> the lobes.<br />

Occasional in lowland to montane forest, reported<br />

from 25 to 850 m elev<strong>at</strong>ion. Indigenous, apparently<br />

ranging eastward to Tahiti. Kuruc 151; Whistler<br />

3842, 8501,8557!.<br />

Phym<strong>at</strong>osorus nigrescens (Bl.) Pichi Serm.<br />

Medium-sized terrestrial fern with a stout, creeping<br />

rhizome, frond deeply cut into 1-10 pairs <strong>of</strong> lobes<br />

(or sometimes simple and lanceol<strong>at</strong>e), and deeply<br />

embedded sori forming conspicuous tubercles on the<br />

upper surface. Occasional in lowland to montane<br />

forest, reported from 200 to 1000 m elev<strong>at</strong>ion. Indi-<br />

genous, ranging from Fiji to Tahiti. Christophersen<br />

997; Long 1996d; Whistler 8643!, 9132.<br />

Phym<strong>at</strong>osorus scolopendria (Burm, f.) Pichi Serm.<br />

lau auta; lau magamaga<br />

Large terrestrial or epiphytic fern with a long-<br />

creeping, scaly rhizome, simple, glossy-green

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