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Bushland Weeds Manual - Environmental Weeds Action Network

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• Fire also offers a valuable window of opportunity<br />

for control.<br />

• Understanding the life-cycles of these weeds is an<br />

integral part of effective chemical and physical<br />

control and preventing spread into intact<br />

bushland.<br />

Dealing with weedy geophytes invading bushland can<br />

be a complex task. The information in the following<br />

case studies illustrates that with a consistent and<br />

carefully targeted effort over time, effective control and<br />

containment of populations can be achieved. The case<br />

Box 4.4 Timing herbicide treatment<br />

Annually renewed corms and bulbs<br />

Chapter 4 Corms, Bulbs and Tubers<br />

For those species with annually renewed corms and bulbs, the<br />

most effective time for herbicide application is when the parent<br />

corm or bulb has shrivelled and the new daughter corm/bulb has<br />

only partly developed. This usually occurs just before or just at<br />

flowering. Apply the herbicide too early and the parent corm/bulb<br />

could have enough reserves to survive; too late and the daughter<br />

corm/bulb will not be affected.<br />

In addition herbicide application at this time prevents the<br />

development of bulbils in many species. This is particularly<br />

important with Oxalis pes-caprae, O. glabra and O. purpurea where<br />

new bulbils are formed on the nodes of underground stems each<br />

year (Du Plessis and Duncan 1989, Peirce 1990, 1998). If herbicide<br />

application takes place after bulbils have begun to develop they<br />

will probably survive and go on to produce new plants the<br />

following season.<br />

Perennial corms<br />

studies cover managing and controlling the spread of<br />

the Yellow Soldier, two species of Watsonia, Harlequin<br />

Flower and Taro where they are invading wetlands and<br />

woodlands around Perth. The final table provides<br />

detailed, species-specific information on control<br />

techniques where they are known. The information in<br />

the table also highlights the many species of weedy<br />

geophytes that have similar life-cycles and reproductive<br />

biologies. The next chapter looks at a much more<br />

diverse group of weeds, the broadleaf herbs that are<br />

not geophytes and the sedges and the succulents.<br />

Only two species dealt with here arise from perennial corms, Black<br />

Flag (Ferraria crispa) and Taro (Colocasia esculenta). Timing for<br />

control of Black Flag is quite problematic as the parent corm does not shrivel and die as the new corm<br />

develops, so effectively there is no corm exhaustion. In addition there is a series of previous years’ corms<br />

below the parent which appear to be joined by a fine thread of living tissue and it is not known if herbicides<br />

can be translocated between these corms (Box 4.1)<br />

Effective control of Taro was obtained by cutting and painting the bases in early to mid summer with<br />

glyphosate and spraying regrowth in late summer with glyphosate and metsulfuron methyl (Case Study 4.3).<br />

Some workers have noted that excessive sap movement when cutting bases early in the season prevents cut<br />

stumps from taking up herbicide.<br />

Perennial bulbs<br />

The lowest dry weight of a perennial bulb generally occurs just when all the leaves are up but before flowering<br />

(Dafni et al.1981). This is probably the time plants would be most vulnerable to herbicide application. An<br />

interesting exception is the Easter Lily (Amaryllis belladonna) which belongs to a group of bulbs that flower as<br />

temperatures drop with the first rains – before the leaves come up. Dry weight in the perennial bulbs with this<br />

kind of life-cycle is generally lowest as the new leaves emerge, a couple of months after flowering (Dafni et al.<br />

1981) and this is probably the best time to apply herbicide.<br />

Perennial tubers<br />

developing daughter<br />

corm<br />

exhausted<br />

parent corm<br />

remains of<br />

previous years’<br />

corms<br />

Corm exhaustion in Watsonia meriana – late winter.<br />

Herbicide application to Arum Lily is effective throughout its period of active growth (Moore and Hoskins 1997,<br />

Brown unpublished data). Early herbicide application prevents the majority of the population flowering and<br />

setting seed but misses tubers that begin active growth later in the season.<br />

Studies on Bridal Creeper (Asparagus asparagoides) have shown herbicide application to have an impact from<br />

flowering through to the green berry stage, although is most effective at flowering (Pritchard 2002).<br />

43

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