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114 FIELDIANA: BOTANY, VOLUME 28 Carptotepala Moldenke, gen. nov. ERIOCAULACEAE l Flores trimeri androgyni. Bracteae involucrati paucae imbricatae squarrosae. Receptaculum dense longeque villosum. Bracteae receptaculi nullae. Flores masculi central!; sepala 3 libera glabra; receptaculum ultra calycem in stipitem angustum production corollam staminaque gerens; petala 3 libera firma adscendentes glabra; stamina 3 libera; filamenta elongata. Flores femini peripherati; sepala 3 libera glabra; petala 3 libera elongata erecta in medio constricta, ad apicem saepe appendiculata; stylus brevissimus, appendicibus 3 ornatis; stigmata 3 brevissima recurvata. Flowers trimerous, androgynous; involucral bracelets rather few, imbricate, squarrose; receptacle densely long-villous; receptacular bracelets absent; staminate florets central in the head, easily detached, their sepals 3, separate, glabrous; the remainder of the floret inserted at the apex of a prolongation of the receptacle; their petals 3, separate, firm, ascending, glabrous; stamens 3, inserted alternate to the petals, free; filaments elongate; anthers sagittate, versatile, composed of 2 thecae, each dehiscing by means of a longitudinal slit; rudimentary ovary present at the base of the stamens; pistillate florets peripheral, firmly attached to the receptacle, with all the floral parts inserted at one level; their sepals 3, separate, glabrous; their petals 3, separate, elongate, erect, narrowed in the middle, often appendaged at the apex; style very short; stigmas 3, very short, recurved; style-appendages 3, issuing from the same level as the stigmas, elongate, erect, fimbriate; ovary large, 3-angulate or 3- alate, 3-celled, 3-ovulate. This genus is known thus far only from the type species C. insolita Moldenke, which exhibits most remarkable floral characters. Among its distinctive characters are the definitely central and loosely attached staminate florets and the distinctly peripheral firmly attached pistillate florets; the prolongation of the floral receptacle (regarded as such in the absence of detailed morphological examination) above the calyx in the staminate florets; the free elongate stamens; the completely separate sepals and petals in flowers of both sexes; the elongated exserted petals of the pistillate florets, with their narrowed mid-portion and caudate appendage; and the 3- alate ovary. The generic name is compounded from the Latin word "carptim", meaning in separate parts, and the latinized French word "tepal", meaning a division of the perianth. Carptotepala insolita Moldenke, sp. nov. Herba parva caulescens; caulibus gracilibus 4-6 cm. longis rectis simplicibus ad apicem longe villosis; foliis numerosissimis rectis 1 By Harold N. Moldenke
BOTANICAL EXPLORATION IN VENEZUELA - I 115 linearibus 4.5-8 cm. longis raultistriatis non fenestratis utrinque glabris vel ad basin plusminusve longe villosis; vaginis cylindricis angustis arete adpressis glabris ca. 3.5 cm. longis; pedunculis stramineis vel flavis 15-25 cm. longis 3-costatis tortis glabris. Small caulescent herb; stems slender, 4-6 cm. long, completely hidden by the sheathing leaf-bases, erect, apparently unbranched, long-villous at the summit; leaves very numerous, erect, linear, grass-green, thin-membranous, 4.5-8 cm. long, rather regularly attenuate from the acute apex to the ampliate and sheathing base, about 1.5 mm. wide at the mid-point, many-striate, not fenestrate, glabrous on both surfaces or more or less long-villous on the subhyaline margins of the top of the sheathing base; sheaths cylindric, narrow, closely appressed to the peduncles, shorter than the leaves, about 3.5 cm. long, glabrous, many-striate, slightly twisted, obliquely split at the apex, the blade ovate-lanceolate, appressed to the peduncle, about 7 mm. long, acute, glabrous; peduncles about 3 per plant, slender, stramineous or yellow, 15-25 cm. long, 3-costate, twisted, glabrous, shiny; heads solitary, turbinate, 4-5 mm. long, 3-4 mm. wide; involucral bractlets rather few, closely imbricate in several series, oblanceolate-spatulate, about 1.9 mm. long and 0.8 mm. wide, firm, stramineous or light-brown, rounded at the apex, more or less squarrose, glabrous on both surfaces, shiny, concave on the inner, convex on the outer surface; receptacle long-villous; receptacular bractlets absent; staminate florets central in the head, loosely attached; sepals 3, separate to the base, inserted about 0.4 mm. below the remainder of the floret, narrowly elliptic, yellowish, about 1.9 mm. long and 0.4 mm. wide, subacute at the apex, glabrous throughout, not bearded, not glanduliferous, firm, erect or spreading; petals 3, separate to the base, elliptic, about 2.1 mm. long and 0.8 mm. wide, obtuse at the apex, yellowish, thin-textured, firm, ascending, glabrous throughout, not bearded, not glanduliferous, not involute; stamens 3, separate and free to the base, alternate with the petals; filaments flattened, about 1.8 mm. long, glabrous; anthers sagittate, versatile, about 0.4 mm. long, 0.2-0.3 mm. wide, the 2 thecae separate and opening by longitudinal slits; ovary rudimentary, at the base of the stamens and petals, dark-brown, about 0.2 mm. long and wide, 3-angled, 3-celled, sometimes surmounted by a very abbreviated 3-parted style; pistillate florets peripheral in the head, all its parts borne at one level: sepals 3, separate to the base, oblong, firm, brownish, ascending, about 1.6 mm. long and 0.3 mm. wide, acute at the apex, glabrous on both surfaces, not bearded, not glanduliferous; petals 3, separate to the base, elongate-linear, erect, narrowed in the mid-portion, to 4.2 mm. long, about 0.2 mm. wide for the lowest 1.2 mm. above the base and there hyaline, then narrowed to about half that diameter for about 0.4 mm., then expanded to 0.15 mm. and somewhat involute and brownish for about 0.8 mm. or more, acute at the apex, the apex often surmounted by a terminal caudate erect of spreading appendage to 0.8 mm. in length, the lowest expanded hyaline portion of the petal long-hirsute on the back, the
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o 2 CM XI E> RAR.Y OF THL UNIVERSIT
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FIELDIANA: BOTANY A Continuation of
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NUMBER 1 LIST OF ILLUSTRATIONS TEXT
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PAGE 67. Emmotum ptarianum 342 68.
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CONTRIBUTIONS TO THE FLORA OF VENEZ
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BOTANICAL EXPLORATION IN VENEZUELA
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