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156 FIELDIANA: BOTANY, VOLUME 28
The dimorphous leaves of the pistillate and staminate plants are
noteworthy, those of the staminate plants being broader. The older
mature leaves of both sexes are non-reticulate and show immersed
nerves, whereas the younger leaves of both sexes manifest a reticulate
conspicuously nerved and veined appearance. In age, apparently,
the leaves become much more coriaceous, and the veins become
immersed on the upper leaf surface. On these older leaves usually
just one main nerve stands out, the other lateral ones being only
slightly evident.
This species is related to S. immersa A. C. Smith of British
Guiana, the principal likeness being found in the nerves of the upper
leaf surface becoming immersed in the leaf tissue of the older
leaves. From this species it differs in its shorter petioles, smooth
rather than minutely tuberculate branches, more narrowly lanceolate
or oblong-lanceolate leaves, longer pedicels of the staminate flower,
and staminate inflorescence composed of only one umbel.
Smilax staminea Griseb., f.
obtusata Steyermark, forma nova.
A typo differt apice foliorum obtuso vel subobtuso; pedunculis
foemineis axillaribus solitariis 2-4 mm. longis, 1.5 ram. latis; pedicellis
bacciferis 5-6 mm. longis.
Stems striatulate, glabrous, unarmed; petioles thickened, 6-15 mm.
long; leaf-blades deep green above, dull paler green below, ovateoblong
(older leaves) to narrowly oblong (those on lateral shoots),
6.5-H cm. long. 2.5 cm. wide, shining above, conspicuously reticulate-veined
on both sides, obtuse to rounded at base, obtuse or
subobtuse at apex, 3-5-nerved from the base; fruiting peduncles
short, solitary, 2-4 mm. long; fruiting pedicels 5-6 mm. long; berries
pale green, 7-8 mm. in diameter.
Type in herb. Chi. Nat. Hist. Mus., collected on forested sandstone
south-facing slopes between pleteau portion and "Cave Camp,"
Ptari-tepuf, state of Bolfvar, alt. 1700-1800 m., November 1, 1944,
Julian A. Steyermark 59695.
Tofieldia duidae Steyermark, sp. nov.
Herba caespitosa, foliis elongato-linearibus acicularibus, 5.5-14
cm. longis, 2-3 mm. latis, glabris, longitudinaliter 6-10-striatis, basi
distiche vaginantibus; scapo erecto tereti, 13-25 cm. longo, glabro
foliis longioribus, 2-6-floro; bracteis minutis; floribus strictis racemosis;
pedicellis erectis, 8-12 mm. longis; involucello bracteolis
late ovatis acutis 1.5-2 ram. longis; perianthii segmentis oblongolanceolatis
acutis, 8 ram. longis, 1.75-2 mm. latis, 5-7-striatis;
filamentis linearibus planis 2.7 mm. longis; antheris lineari-oblongis,
1.2 ram. longis, 0.5 ram. latis; stylis 3 coalitis, 1.5 ram. longis;
stigmatibus 3 subcapitatis; ovario 3-lobato conico-oblongo 3.5 mm.
longo apicera versus acuto, triloculari; ovulis 10 in quoque loculo.
Type in herb. Chi. Nat. Hist. Mus., collected along stream bank