SPECIES PLANTARUM — FLORA OF THE WORLD (2003) Embelia urophylla Wall., Numer. List no. 2309 (1830), nom. nud. Diemenia racemosa Korth., Ned. Kruidk. Arch. 3: 388 (1854); Licania diemenia Blume, Mélanges Bot. 2: 10 (1855), nom. superfl. illeg.; Trichocarya racemosa (Korth.) Miq., Fl. Ned. Ind. 1(1): 358 (1855). T: Sumatera, P.W.Korthals s.n.; holo: L. Parastemon spicatus Ridl., J. Straits Branch Roy. Asiat. Soc. 75: 29 (1917). T: Borneo: Sarawak, Kuching, G.D.Haviland 723; holo: K; iso: SAR. Illustrations: G.T.Prance & F.White, Philos. Trans. B 320: 97 (1988); G.T.Prance, Fl. Males. Ser. 1, 10(4): 648 (1989). Map: G.T.Prance, Fl. Males., Ser. 1, 10(4): 650 (1989). Tree to 40 m tall, or shrub. Trunk often buttressed. Branches glabrous when young. Leaves with stipules triangular, c. 1 mm long, caducous; petioles 4–5 mm long, canaliculate, glabrous; lamina thinly coriaceous, narrowly oblong, 2.5–8 × 1.4–2.5 cm, cuspidate acuminate at apex with tip 5–15 mm long, cuneate at base; midrib plane above, prominulous beneath; primary veins 8–11 pairs. Inflorescence of axillary and rarely terminal racemes or occasionally slightly branched, 4–14 cm long; rachis glabrous. Flowers polygamo-dioecious, c. 1.5 mm long. Receptacle broadly cupuliform to flattened saucer-shaped, glabrous on exterior, tomentose within; pedicels up to 2 mm long. Calyx lobes acute, glabrous on exterior. Petals 5. Stamens 2 fertile with 3 sterile staminodes opposite. Ovary pilose on exterior. Style pilose at base, glabrous above; stigma trifid. Fruit ellipsoid, 1–1.5 cm long; epicarp smooth, glabrous; mesocarp thin, hard; endocarp thin, hard, bony, glabrous within. Fig. 35. Widely distributed species from the Nicobar Islands and Thailand through the Malay Peninsula to Borneo and Sumatera. 41: THA. 42: BOR, MLY, SUM. 43: NCB. Lowland and coastal forest; commonest in swamp and secondary forest. Map 234. 41. NICOBAR IS.: Without detailed locality, Clarke 25960 (K). THAILAND: Tako, Langsuan, A.F.G.Kerr 1762 (BM). 42. BORNEO: Brunei, Tutong, Sungei Medit, D.A.Simpson 2582 (K); Sabah, Jesselton Distr., Mandahan, Kimanis F. R., W.Meijer SAN36251 (K, SAN); Sarawak, Binatang Distr., Sg. Nangar, Matu Daro P. F., J.A.R.Anderson 12306 (K, L, NB, S); Kalimantan, Loa Djanan, W of Saraminda, A.J.G.H.Kostermans 9975 (BO, K, L); E Kalimantan, Nunukan Is., A.J.G.H.Kostermans 8992 (BM, K, L, NY, P). MALAYA: Peninsular Malaysia, King’s Collector 10980 (A, BM, L, P, US, Z); Singapore, J.Sinclair SF39504 (K, L, M, NY). SUMATERA: Langga Pajoeng, Kota Pinang, R.S.Toroes 3319 (A, K, L, NY, US). 2. Parastemon versteeghii Merr. & L.M.Perry, J. Arnold Arbor. 21: 197 (1940) T: New Guinea: West Irian, Idenburg R., L.J.Brass & C.Versteegh 13544; holo: A; iso: BM, L. Illustration: G.T.Prance & F.White, Philos. Trans. B 320: 97 (1988). Map: G.T.Prance, Fl. Males. Ser. 1, 10(4): 650 (1989). Tree up to 40 m tall. Branches sparsely puberulous when young, soon glabrous. Leaves with stipules triangular, c. 1 mm long, caducous; petioles 3–7 mm long, terete, glabrescent; lamina thinly coriaceous, narrowly oblong, 5–9.5 × 1.8–3.7 cm, cuspidate acuminate at apex with tip 7–15 mm long, cuneate at base; midrib plane above, prominulous beneath; primary veins 8–12 pairs, inconspicuous, slightly prominulous beneath. Inflorescence of axillary and terminal racemes, 2–9 cm long; rachis sparsely villous. Flowers hermaphrodite, c. 1.5 mm long; pedicels 1.5–3 mm long, sparsely villous. Receptacle broadly cupuliform-flattened, sparsely villous on exterior, tomentose within. Calyx lobes acute, with hirsute margins. Petals 5, with hirsute margins. Stamens 5, all fertile, opposite the petals in a complete circle. Ovary glabrous on exterior except at base. Style pilose at base, glabrous above; stigma with 2 large apical lobes with the third reduced or missing. Fruit ellipsoid, c. 1.6 cm long; epicarp smooth, glabrous; mesocarp thin, hard; endocarp thin, hard, bony, glabrous within. Eastern Indonesia from the Moluccas through West Irian to Papua and the Admiralty Islands. 42: BIS, MOL, SUL. 43: NWG. Mostly in drier forest than P. urophyllus. Map 235. 42. MALUKU: Sangi & Talaud Is., Morotai, H.J.Lam 3529 (A, K, L). 43. BISMARCK ARCHIPELAGO: Admiralty Is., K.Mair 533 (L). NEW GUINEA: Irian Jaya, vicinity of Hollandia, Forest Serv. bb25062 (A, K, L, NY); Irian Jaya, Babo, Forest Serv. bb 22298 (K, NY); Irian Jaya, Sorong, D.R.Pleyte 744 (A, K, L, P); Papua New Guinea: Buso, Lae Sub-Distr., H.Streiman NGF45157 (K); Papua New Guinea, Paiawa Valley, Morobe Sub-Distr., R.J.Johns NGF47316 (K, L). 178
CHRYSOBALANACEAE (Parestemon) Figure 35. Parastemon urophyllus. A, habit; B, flower; C, 1 /2 flower; D, petal; E, infructescence (A–E, P.Chai S17282); F, fruit, × 3 (W.Meijer SAN38933). Scale bars: A, E = 2 cm; B–D = 1 mm; F = 5 mm. Drawn by Margaret Tebbs. 179
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