Vol. 1 part 2 - Species Plantarum Programme

CHRYSOBALANACEAE (Exellodendron)
3: Lamina to 7 cm long, with acumen less than 8 mm long
4 Leaves coriaceous, with a distinct but shorter acumen 2–3 mm long;
primary veins prominent on lower surface
4: Leaves chartaceous, long-apiculate (c. 1 cm long); primary veins
inconspicuous
4. E. gardneri
5. E. gracile
1. Exellodendron coriaceum (Benth.) Prance, Fl. Neotrop. Monogr. 9: 197 (1972)
Parinari coriacea Benth., J. Bot. (Hooker) 2: 213 (1840), as Parinarium coriaceum; Ferolia coriacea (Benth.)
Kuntze, Revis. Gen. Pl. 1: 216 (1891). T: Guyana, R. & R.Schomburgk 65; holo: K; iso: BM, BR, CGE, E,
GH, L, NY, OXF, P, W.
Illustration: G.T.Prance & F.White, Philos. Trans. B 320: 101 (1988).
Map: G.T.Prance, Fl. Neotrop. Monogr. 9S: 202 (1989).
Tree to 10 m tall. Leaves with stipules to 2 mm long, caducous; petioles 5–6 mm long,
canaliculate above; lamina oblong-elliptic, 9–12 × 3–7.5 cm, cuneate to subcuneate at base,
with acumen 1–1.5 cm long, glabrous above, densely arachnoid beneath; midrib prominent
on both surfaces; primary veins 13–19 pairs, not prominent. Inflorescences dense flowered
axillary or terminal panicles, much shorter than the leaves, with silver-grey pubescence.
Bracts and bracteoles 1–3 mm long, caducous. Receptacle subcampanulate, swollen slightly
to 1 side, silver-grey-tomentose on both interior and exterior. Calyx lobes sharply pointed.
Petals 5, white, caducous. Stamens 7, with staminodes represented solely by swellings in the
staminal circle or occasionally with 1 or 2 short filaments. Ovary and style densely villous.
Fruit round to oval, basally narrowed, 1.5–2.5 cm long, 1.2–2 cm broad; epicarp smooth and
glabrous; mesocarp thin and fleshy; endocarp thin, hard, with a smooth surface, sparsely
lined on interior with short straight silvery hairs. Fig. 37A–F.
Guyana, adjacent Venezuela and C, E and N Amazonian Brazil. 82: GUY, VEN. 84: BZN
(AM, PA, RR). Savannas, savanna margins and riversides. Map 238.
82. GUYANA: 2.5 km NW of Karanambo ranch, U. Takutu-U., Essequibo region, B.Hoffman 1242 (K, US);
Rupununi R., Monkey Pond Landing, SW of Mt. Makarapau, P.J.M.Maas et al. 7601 (K, U). VENEZUELA:
Amazonas, Caño San Miguel, Río Guainía, Ll.Williams 14914 (NY); Bolívar, Cerro Cotorra, between La
Paragua and San Pedro de las Bocas, J.A.Steyermark 86843 (VEN). 84. BRAZIL NORTH: Amazonas,
Manaus, Igarapé Cachoeira Grande, Tarumã, A.Ducke RB 35185 (K, US, RB); Amazonas, Baía de Buiaçú,
S Bank Rio Negro, G.T.Prance 24216 (INPA, NY); Roraima, Caracaraí, R.L.Fróes 22991 (IAN, NY); Pará:
Alter do Chão, A.L.K.M.Albernaz 149 (K, NY).
2. Exellodendron cordatum (Hook.f.) Prance, Fl. Neotrop. Monogr. 9: 197 (1972)
Parinari cordata Hook.f., in C.F.P.Martius, Fl. Bras. 14(2): 50 (1867), as Parinarium cordatum; Ferolia
cordata (Hook.f.) Kuntze, Revis. Gen. Pl.: 216 (1891). T: Brazil Northeast: Piauí, G.Gardner 2560; holo:
K; iso: BM, CGE, US, W.
Illustration: G.T.Prance, Fl. Neotrop. Monogr. 9: 198 (1972).
Map: G.T.Prance, Fl. Neotrop. Monogr. 9S: 201 (1989).
Shrub or small tree. Leaves with stipules to 1.5 mm long, caducous; petioles 4–8 mm long,
canaliculate above, pubescent, with 2 glands at lamina base; lamina ovate, 3.5–9 × 2.25–3.5 cm,
usually distinctly cordate, rarely rounded at base, rounded at apex, or with a small acumen to
3 cm long, glabrous above, glabrous or with densely lanate easily rubbed off pubescence on
lower surface; midrib prominent on both surfaces; primary veins 9–12 pairs, not prominent.
Inflorescences terminal and axillary panicles. Bracts and bracteoles 1–2 mm long, caducous.
Receptacle subcampanulate, swollen slightly at 1 side, silvery grey-tomentose on exterior,
villous to base within. Calyx lobes acute. Petals 5, white. Stamens 7, with 7 filamentous
staminodes opposite them. Ovary and base of style villous. Fruit (immature) oblong; epicarp
smooth, glabrous; mesocarp thin; endocarp thin, hard, with a smooth surface, densely brownpubescent
within.
C and NE Brazil. 84: BZC (GO), BZE (BA, MA, PI), BZN (TO). Cerrado. Map 239.
84. BRAZIL NORTH: Tocantins, Rio do Sono, A.C.Sarmento 604 (RB). BRAZIL NORTHEAST: Piauí,
Município de Uruçuí, R.P.Orlandi 599 (RB); Palmeiras, A.B.Sousa 83 (RB); Maranhão, Fazenda Parnaíba,
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