Vol. 1 part 2 - Species Plantarum Programme

CHRYSOBALANACEAE (Parinari)
different gender, and have accepted that original publication of names under Parinarium
should be maintained under Parinari, with appropriate correction of the gender of their
epithets. Index Kewensis listed all specific names up to Supplement 7 (1929) under
Parinarium, but from Supplement 8 (1933) onwards under Parinari, irrespective of which
was adopted by the authors of the species names. A few authors, however, notably
J.E.Dandy, have argued that Parinari and Parinarium are different names and not mere
variants and that a new combination is required to transfer an epithet from one to the other.
This might have been supported by the International Code up to 1994 in its list of conserved
generic names, where, for example, Pupalia and Pupal, Canavalia and Canavali, and
Cajanus and Cajan respectively, were listed as different names, though the examples of
Naravelia and Naravel, and Vochysia and Vochy were listed as mere orthographic variants
respectively. However, from 1994 onwards the list of conserved names has been amended so
that Pupalia, Canavalia and Cajanus are listed as orthographic variants, as are Naravelia and
Vochysia. The case of Parinari and Parinarium thus seems to be clearly resolved in favour
of treating them also as orthographic variants, not requiring new combinations when epithets
are transferred from one to the other. In this account, when names were originally published
under Parinarium, this is indicated after the date of publication.
1 Stomatal crypts absent from leaf lower surface; leaf lower surface
glabrous or with a persistent lanate pubescence and then with large
persistent stipules 7–40 mm long
[sect. Anareolata]
2 Leaf lower surface glabrous; stipules small and caducous
3 Leaves elliptic to oblong or obovate-elliptic, 9.5–20.5 × 4.5–8.5 cm;
primary veins 11–16 pairs; panicles large and silvery pubescent
1. P. argenteosericea
3: Leaves ovate, 5–9 × 2–4.5 cm; primary veins 7–11 pairs; panicles
small and subsericeous brown pubescent
2. P. canarioides
2: Leaf lower surface densely lanate pubescent, but when removed no
stomatal crypts present; stipules large and persistent, 7–40 mm long,
3–5 mm broad at base
4 Leaves oblong-lanceolate, 5–18 cm long on flowering branches,
thickly coriaceous, with base cuneate
3. P. elmeri
4: Leaves elliptic to oblong, 11–28 cm long, chartaceous, with base rounded 4. P. parva
1: Stomatal crypts present on leaf lower surface; leaf lower surface lanate or
at least pubescent in crypts; stipules usually small (larger in species 5–8)
[sect. Parinari]
5 Stipules 10–40 mm long, persistent, semi-amplexicaul
6 Leaf base cuneate or subcuneate; young branches with dense stiff
ferrugineous hairs
5. P. rodolphii
6: Leaf base usually cordate or subcordate, sometimes rounded; young
branches usually grey- to brown-tomentellous to -puberulous, without
stiff ferrugineous hairs (but ferrugineous-tomentose in P. alvimii)
7 Leaf lower surface with a dense golden-ferrugineous pubescence
obscuring stomatal cavities; leaf base rounded or only slightly
subcordate; with marginal glands at end of primary veins
6. P. hypochrysea
7: Leaf lower surface grey to brown tomentose; leaf base cordate or
subcordate; without marginal glands
8 Leaf apex bluntly acute; primary veins 17–25 pairs 7. P. alvimii
8: Leaf apex acuminate; primary veins 14–16 pairs 8. P. campestris
5: Stipules caducous, usually small (1–8 mm), if larger not clasping stem
9 Plants rhizomatous suffructices, usually less than 30 cm tall, rarely up
to 2 m
10 Leaf lower surface with white-lanate pubescence obscuring the
stomatal cavities; leaves oblong to elliptic, cuneate at base; primary
veins 11–16 pairs
9. P. obtusifolia
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