116 H. PETER HALLBERG .A.ND REINHOLD IV .A.RSSONtrue heather heath (Hylocomieto-:-Ca1lunetum accordingto DAMMAN 1957, see also MALMER'spaper inthe present book), apart from Oalluna, also otherdwarf shrubs dominate, e.g., Arctostaphylos uva-ursi(on rocks and boulders), Empetrum nigrum, andV accinium vitis-idaea. On strongly exposed cliffsin the bare skerry region, Empetrum often growsnearer to the sea than Oalluna (GILLNER 1964b,p. 8, cf. DAMMAN 1957, p. 371). Exposure to strongwind causes Calluna to be replaced by Deschampsiaflexuosa. Other plants frequenting the heatherheath are some other grasses (see below), herbs likePotentilla erecta, mosses, and lichens.The heaths are nowadays seldom grazed, and thetree and bush vegetation on these heaths is graduallyincreasing, particularly Sorbus aucuparia havingbecome much more widespread. The heather heathcorresponds in the woody vegetation to Vaccinietummyrtilli (IVARSSON 1962, p. 98) under a tree andbush layer of varying composition.On dry ground the heather heath is rich in grasses,mosses, and lichens. Among these species,Agrostis canina ssp. montana, Anthoxanthum odoratum,Deschampsia flexuosa, Hylocomium splendens,and Oladonia silvatica sens. lat. may be mentioned.On thin soil on rock the heather heath is replacedby grass heath, rock crevice vegetation, or lichencommunities (see above).Carex nigra, C. panicea, Oornus suecica, Ericatetralix, Eriophorum angustifolium, E. vaginatum,and Molinia coerulea belong to heath (moor) communitieson wet ground. They are, as a rule, countedin Sweden among the so-called wet heaths, whichconsist of a whole series of plant communities,transitional between heather heath and peat-formingvegetation, meadows, and pastures. In wetheaths, which resemble the fens closely, Nartheciumossifragum grows together with Drosera rotundifolia,Erica tetralix, and M olinia coerulea, etc. in a matof different Sphagnum species. However, N artheciumis more common inland where the precipitationis higher and the peaty areas niore extensive(cf. H. FRIES 1945, p. 42, Fig. 6).Lakes and fensReal lakes are few (cf. LYSEN 1960) but smallbodies of water and wet fen areas are common. Inthe littoral zone, particularly in the bare skerryregion, rock pools are numerous, whose brackishwater and high pH values reduce the vegetation toa green soup of Enteromorpha species. Inland ofthe littoral, the vegetation of the ponds has agreater variety of species. Limosella aquatica andTillaea aquatica are characteristic of a type ofshallow, periodically desiccated pools. The deeperponds near tlie sea shore are the habitats of Sparganiumangustifolium, Typha latifolia, and severalScirpus species, e.g., S. maritimus. Not infrequently,Scorpidium scorpioides may appear in these localities,sometimes indicative of the presence of shelldeposits. Typical Lobelia lakes are very rare in thebare skerry and deciduous woodland regions (cf.H. FRIES 1945, map 178).Owing to the peat cutting, which was formerlyvery frequent, a number of minor bodies of water,originally peat-hags, are to be seen in the small peatareas formed in the rocky depressions. These peatdeposits were often dug out right down to the rock.Typical plants of the peat-hags are Equisetumfluviatile, Glyceria fluitans ad M enyanthes trifoliata,sometimes also . Iris pseudacorus. In thedeeper trenches, colonies of Hippuris vulgaris,Nymphaea alba and Sparganium angustifolium maybe seen, among others. Remains of bog vegetationmay be observed here and there in the peat-hags(IVARSSON 1962, p. 12). The hummocks are cappedwith, for instance, Calluna, Empetrum nigrum,Erica tetralix, Sphagnum spp. and, occasionally,Rubus chamaemorus, which is rare in the coastalregion. The margins of ponds and peat-hags aremostly overgrown, forming transitional stages betweenthe communities of the open waters andthose of the moist meadow, wet heath, and firmerpeat-forming vegetation.In wet and poorly nutrified fens, Carex rostrata,Equisetum fluviatile and Drepanocladus fluitans,grow among others, while Carex vesicaria, Eriophorumlatifolium, Phragmites communis, Oalliergongiganteum, and Scorpidium scorpioides occur in thewell nutrified fens. On slightly drier ground, Oarexnigra, Lythrum salicaria and Potentilla palustris arecharacteristic, while Oarex flava and 0. hostianainhabit firm fens richer in nutrients..Acta Phytogeog1·. Suec. 50
Vegetation of coastal Bohusliin 117Non-calcareous grasslandPASTURES.-With the exception of the vegetationon shell deposits (see below), that of meadowsand pastures has been little studied. Some indicationsof its composition are to be found e.g. inworks by GILLNER (1960, pp. 60, 80, 87, and 1964b)and IvARSSON (1962, pp. 20, 30-31). Fairly commonis a type of pasture-land which is closely related tothe heather heath and occurs partly in greenpatches in that heath. These pasture communitiesinclude such species as Agrostis tenuis, Anthoxanthumodoratum, Luzula campestris, Nardus stricta,Sieglingia decumbens, Lathyrus montanus, Polygalavulgaris, Potentilla erecta, Solidago virgaurea, Succisapratensis, Veronica officinalis, Hylocomium splendens.,,Pleurozium Schreberi, Rhytidiadelphus squarrosus,and others. In the interior coastal regionArnica montana is often a member of these communities,which are referred by PREISING (1949,1950) to Nardo-Callunetea (Nardetalia).The pastures in the clay valley district are partlywhite with Trifolium · repens during the heightof summer, and in late summer yellow withLeontodon autumnalis. Cynosurus cristatus growsscattered, but is noticeably less common towardsthe north. Other typical species are, in the bottomlayer, Climacium dendroides and Rhytidiadelphussquarrosus, in the field layer Achillea millefolium,Agrostis tenuis, Festuca rubra, Poa pratensis,Taraxacum (V ulgaria), and . Trifolium pratense.Certain variants also contain Carex leporina, Deschampsiacaespitosa, H olcus la natus, and J uncuseffusus. These Cynosurion communities (TUXEN1955, p. 170) pass into the Alchemilla glaucescens Anthyllis vulneraria association on dry ground withshell deposits (see below), on wet soil into moistmeadow communities (Molinietalia, see TuxEN1955, p. 171), and on soil deficient in nutrients intoNardetalia (see above). In the woody vegetationthey are represented, in the first place, by Agrostidetumtenuis and Filipenduletum ulmariae (seelater sections of this paper).An outstanding feature in this district is theoccurrence of communities dominated chiefly byFilipendula ulmaria. Communities of this kindhave in recent years gained a greater distributionalong ditches and roads. Apart from Filipendulaulmaria, Cirsium palustre, Holcus lanatus, Juncusconglomeratus, J. effusus, and Lychnis flos-cuculiare common in this vegetation. They border onforest of Alnus glutinosa with Filipendula ulmariain the field layer (see end of paper).GRASSLAND ON SAND.-In contact with theElymo-Ammophiletum of the sandy shores (seeabove), or with cer ,tain salt marsh communities,e.g. Juncetum Gerardi (see GILLNER's contribution),a Festuca ovina - Galium verum v. litorale associationis to be seen on sand deficient in lime, and aTortulo-Phleetum arenarii on calcareous sand.Owing to pasturing and trampling these communitieshave become rather widely distributed, particularlyin northern Bohuslan.·The Festuca ovina - Galium verum v. litoraleassociation corresponds to a community describedunder the same name in Central Europe (BRAUNBLANQUET & DE LEEUW 1936, p. 367, RAABE 1950,p. 25, and others). This community has been referredto briefly by DEGELIUS (1939, p. 59) and FRODIN (1921). Some of its typical species are Armeriamaritima, Rumex acetosella, Sedum acre, and Trifoliumarvense. Fruticulose lichens are abundant.In the initial phases of this community, Carexarenaria plays an important part (cf. DEGELIUS),sometimes also Spergula vernalis, Viola canina, andV. tricolor. A variant of this community is connectedwith the Oalluna heath by species like Agrostiscanina, A ntennaria dioeca, Call una vulgaris, Violacanina, and Dicranum scoparium. Another variant,on a wet or slightly clayey substratum, is relatedto Cynosurion (see above).Vegetation on shell depositsIt has been emphasized earlier that the soils ofBohuslan are, as a rule, acid. However, also richlybasic soils are present, owing to varying contentsof shells. These shells are either scattered in depositsof different textures, or collected in plentiful shelldeposits, so-called shell banks (HESSLAND 1943,SANDEGREN 1931). Most of the shell deposits arelocalized to the coastal region; where, owing to thepaucity in woods, they have a more apparent effecton the composition of the vegetation than the shelldeposits in the wooded interior of the province.Acta Phytogeog.1·. Suec. 50
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ACTA PHYTOGEOGRAPHICA SUECICAEDIDIT
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Printed with support fromH. M. King
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TABLE OF CONTENTSREGIONAL SURVEYFea
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REGIONAL SURVEYFeatures of Land and
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Features of land and climate 3Fig.
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Features of land and climate 5bould
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8 HUGO SJORSFig. 9. The length of d
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10 HUGO SJORSYearly precipitation i
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12 HUGO SJORSrainfall, and in winte
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Fig. 2. On the lee side of an expos
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16 MATS WJERNmetres below the surfa
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18 MATS WlERNmon in shallow water,
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20 MATS W.LERNthe West Coast is cau
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22 MATS WlERNalways sterile). Ceram
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24 MATS WlERNparticular species of
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26 MATS WlERNRich in species. Scatt
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The Vegetation of Swedish LakesBy G
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30 GUNNAR LOHAMMARby compact underw
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32 GUNNAR LOHAMMARmake up only a sm
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34 GUNN AR LOHAMMARoccur patches of
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36 GUNNAR LOHAMMARFig . 7. Exposed
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38 GUNN AR LOHAMMARFig. ll. South s
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40 GUNN AR LOHAMMARFig. 15. Vastann
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42 GUNN AR LOHAMMARFig. 19. Lake Ta
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44:GUNNAR LOHAMMARLappmark it goes
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46 GUNNAR LOHAMMARice cover. From l
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Forest RegionsBy HUGO SJO RSA count
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50HUGO SJORSFig. 2. Biotic zonation
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52 HUGO SJORSFig. 4. Beech forest w
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54 HUGO SJORSFig. 6. Coastal oakwoo
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56HUGO SJORS·the growth of deciduo
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58HUGO SJORSFig. 8 . . vernal hazel
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60 HUGO SJORSthrough basal shoots o
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62 HUGO SJORSdebted to their collea
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The Mountain Regions of IJapplandBy
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ASPECTS OF THE NORTHThe BorderlandB
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The Borderland169'Fig. 2. The same
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The marked closeness of the iso-lin
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The Borderland 173level, much of th
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The Borderland 175Fig. 4. The Nitte
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Glimpses of the Bothnian coast 177d
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Glimpses of the Bothnian coast . 17
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Regional ecology of mire sites and
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Regional ecology of mire sites and
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Regional ecology of mire sites and
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Regional ecology of mire sites and
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Algfloarna, a mixed mire complex in
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Algjloarna, a mixed mire complex in
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Calcareous fens in J iimtland193CAL
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Calcareous fens in Jamtland 195Fig.
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Calcareous fens in J iimtland 197FE
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The major rivers of Northern Sweden
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The major rivers of Northern Sweden
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The major rivers of Northern Sweden
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The North-East CornerBy ERIK JULIN
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Terrestrial vegetation and flora 20
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Notes on aquatic vegetation 209SPON
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Notes on aquatic vegetation 211Fig.
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Notes on aquatic vegetation 213Fig.
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Lappland East of the MountainsBy JI
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South-facing hills and mountains 21
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Vegetation and flora of alpine outl
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Regional aspects on the flora 221Fi
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Regional aspects on the flora 223Fi
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Regional a.spects on the flora 225O
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Cultural influence on the flora 227
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Notes on the vegetation of lakes in
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Notes on the vegetation of lakes in
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Lost and Living Lakes In the Upper
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Lost and living lakes in the upper
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P:....,.>::."tl;:3"..,.cc-'!lJ'.J-,
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Lost and living lakes in the upper
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Subalpine tall herb vegetation, sit
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Subalpine tall herb vegetation, sit
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Subalpine tall herb vegetation, sit
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Subalpine tall herb vegetation, sit
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Mountain MiresBy AKE PERSSON1 ntrod
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Mountain mires 251Fig. I. Part of a
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Mountain mires 253Fig. 3. Carex jun
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Mountain mires 255Spring vegetation
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Plant Cover of the Alpine RegionsBy
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low-alpine belt most of the other s
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Ohionophobous plant communities 261
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Ohionophilous plant communities . 2
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Ohionophilous plant communities 265
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The high-alpine region 267as a pred
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PAST AND PRES ENTThe Late-Quaternar
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The Late-Quaternary vegetation of S
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The Late-Quaternary vegetation of S
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The Late-Quaternary vegetation of S
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The Late-Quaternary vegetation of S
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The Late-Quaternary vegetation of S
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Reindeer Grazing ProblemsBy ELIEL S
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Reindeer grazing problems 283Fig. 2
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Botanical Indications of Air Pollut
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Botanical indications of air pollut
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Recent changes in flora and vegetat
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Recent changes in flora and vegetat
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Recent changes in flora and vegetat
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BibliographyBy AK E SJODINThe year
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Bibliography 297Torfmooren. Diss. -
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Bibliography 2991934, Jungfrun. - S
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Bibliography 301(Das Gebiet des Ben
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Bibliography 303Humiditat des Klima
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Bibliography 305- 1960, Bidrag till
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Bibliogmphy 307land. ("Uber die Gef
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Bibliography 3091958, Dynamik och k
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Bibliography 311- 1926, Stockholms
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Bibliography 313- Verh. Int. Verein
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SVENSI
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Pris 72 kronor Printed in Sweden 19