Edge effects on ground beetles 7evolve. The physiognomy of the edges in the pres<strong>en</strong>t study area,which were g<strong>en</strong>erally quite abrupt because of frequ<strong>en</strong>t managem<strong>en</strong>toperations, could exp<strong>la</strong>in the abs<strong>en</strong>ce of ground beetlespecies associated with edge habitats. As a consequ<strong>en</strong>ce of thisabs<strong>en</strong>ce of edge-associated species and the pres<strong>en</strong>ce of speciesfrom both adjac<strong>en</strong>t habitats in the woodlot–op<strong>en</strong> habitat edges,we found that the species assemb<strong>la</strong>ge of ground beetles in theedge zone constituted a transition betwe<strong>en</strong> the assemb<strong>la</strong>ges ofwoodlots and op<strong>en</strong> habitats.DEI on the species assemb<strong>la</strong>ge of ground beetlesin woodlotsQuantifying the DEI on ground beetles in for<strong>est</strong>s and woodlotshas led to contrasting results being reported in the literature. Forground beetles, values explicitly reported in previous studies arein the range 5–22 m <strong>des</strong>pite the fact that they rely on differ<strong>en</strong>ttrapping <strong>des</strong>igns and concern very diverse ecological andbiogeographical contexts such as an agro-for<strong>est</strong>ed <strong>la</strong>ndscape inEng<strong>la</strong>nd (5 m; Bedford & Usher, 1994), boreal for<strong>est</strong>–clearcutboundaries (
8 A. Roume et al.and the movem<strong>en</strong>t of op<strong>en</strong> habitat species into woodlot edg<strong>est</strong>o protect themselves from disturbance or to overwinter.Acknowledgem<strong>en</strong>tsWe sincerely thank colleagues of UMR DYNAFOR andreviewers for their useful comm<strong>en</strong>ts on the manuscript andD. Goodfellow for improving the English. We also gratefullyacknowledge the farm<strong>la</strong>nd and woodlot owners who allowedus to install traps on their private <strong>la</strong>nd. This study was fundedby the joint Midi-Pyr<strong>en</strong>ees-Aquitaine Regions’ program ‘For<strong>est</strong>Biodiversity Dynamics under Global Change’ and the ANR-Biodiversity ‘BiodivAgriM’ program. 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