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A flora of Manila - Rainforestation

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272<br />

A FLORA OF MANILA<br />

70. SIMARUBACEAE (Quassia Family)<br />

Trees or shrubs, sometimes climbing, with usually bitter bark, and alternate,<br />

pinnate, leaves. Inflorescence axillary or terminal, racemose,<br />

cymose, or paniculate. Flowers regular. Calyx 3- to 5-lobed. Petals 3<br />

to 5, hypogynous. Disk annular or elongated, rarely none. Stamens as<br />

many as, or twice as many as, the petals, free. Ovary superior, free,<br />

1- to 6-celled, entire or deeply lobed; ovules 1 or few in each cell; styles<br />

free or more or less united. Fruit drupaceous or capsular; seeds usually<br />

solitary.<br />

Genera 32, species about 130, tropical and subtropical regions in both<br />

hemispheres, 8 genera and about 12 species in the Philippines.<br />

Somewhat climbing, armed with short spines; petioles not winged; flowers<br />

small, white 1. Harrisonia<br />

Erect, unarmed; petioles and leaf-rachis winged; flowers large, red.<br />

1. HARRISONIA R. Brown<br />

2. Quassia<br />

Erect or somewhat climbing, spiny, glabrous or pubescent shrubs.<br />

Leaves odd-pinnate. Flowers perfect, racemose or cymose. Calyx small,<br />

4- or 5-fid. Petals 4 to 5, longer than the calyx. Stamens twice as many<br />

as the petals, the filaments with small scales at the base; ovary globose,<br />

4- or 5-lobed, 4- or 5-celled; ovules solitary. Fruit berry-like, fleshy, glo-<br />

bose. (In honor <strong>of</strong> J. Harrison, an English horticulturist.)<br />

Species 3 or 4, tropical Africa and Asia through Malaya to Australia,<br />

2 in the Philippines.<br />

1. H. perforata (Blanco) Merr. (H. paucijuga Benn.). Asimao, Laiya,<br />

Mamiquil (Tag.).<br />

Somewhat climbing or nearly erect, 2 to 4 m high, the branches armed<br />

with short sharp spines. Leaves 5 to 10 cm long, slightly pubescent or<br />

nearly glabrous; leaflets oblong-ovate, 1.5 to 3.5 cm long, entire or crenate,<br />

acute or obtuse. Racemes 3 to 5 cm long, terminal. Flowers white; petals<br />

about 6 mm long; filaments villous at the base. Fruit globose, fleshy,<br />

glabrous, 1 to 1.5 cm in diameter. (Fl. Filip. pi 23 Fagara piperita.)<br />

In dry thickets near Fort McKinley, fl. Apr.-June; <strong>of</strong> local occurrence<br />

in the Philippines. India to China and Malaya.<br />

2. QUASSIA Linnaeus<br />

A glabrous erect shrub, intensely bitter. Leaves alternate, odd-pinnate,<br />

the petiole and rachis winged. Flowers large, perfect, in terminal simple<br />

racemes or panicles. Calyx small, 5-partite. Petals 5, elongated, erect,<br />

not spreading. Stamens 10, inserted at the base <strong>of</strong> the large columnar<br />

torus. Ovary 5-lobed. Drupes 5 or fewer, spreading. (Named after<br />

Quassi, or Coissi, a negro slave in Surinam who used the plant for<br />

medicine.)<br />

A monotypic genus <strong>of</strong> tropical America.<br />

1. Q. AMARA L. Corales (Sp.-Fil.) ; Quassia.<br />

A glabrous shrub 2 to 3.5 m high. Leaves alternate, about 20 cm long,<br />

the petiole and rachis broadly winged; leaflets 5, sessile, elliptic-oblong.

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