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Ecoregional Assessment of Biological Diversity in East Kalimantan

Ecoregional Assessment of Biological Diversity in East Kalimantan

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<strong>East</strong> <strong>Kalimantan</strong> <strong>Ecoregional</strong> <strong>Assessment</strong>: Chapter 2 – Target Ecological Systems<br />

In order to achieve an ecoregional scale-appropriate identification for this system type, we<br />

modified exist<strong>in</strong>g vegetation maps, which lacked coverage for a large part to the west <strong>of</strong> <strong>East</strong><br />

<strong>Kalimantan</strong> (and also which did not differentiate between Lower Montane Ra<strong>in</strong>forests and<br />

Upper Montane Ra<strong>in</strong>forest/Cloud Forest), by draw<strong>in</strong>g the upper boundary <strong>of</strong> Lowland<br />

Ra<strong>in</strong>forests at the altitude <strong>of</strong> 1000 m a.s.l. There is, though, the awareness that some parts <strong>of</strong><br />

the map designated as Lowland Ra<strong>in</strong>forest may <strong>in</strong> fact be Montane Ra<strong>in</strong>forest or Cloud<br />

Forest. Such anomalies can be sorted out <strong>in</strong> the more detailed analysis <strong>of</strong> Conservation Area<br />

Plann<strong>in</strong>g once sites are identified for conservation action. This potential for discrepancy<br />

should be kept <strong>in</strong> m<strong>in</strong>d at the results report<strong>in</strong>g section.<br />

Lowland Ra<strong>in</strong>forests <strong>in</strong> Borneo are dom<strong>in</strong>ated by dipterocarp trees (named because <strong>of</strong> their<br />

w<strong>in</strong>ged seeds) and are frequently referred to as Lowland Dipterocarp Forests. Dipterocarps<br />

are extremely tall trees and frequently reach heights <strong>of</strong> 45m. In the richest forests, 10% <strong>of</strong> all<br />

trees and 80% <strong>of</strong> all emergents are dipterocarps, usually belong<strong>in</strong>g to the genera<br />

Dipterocarpus, Dryobalanops and Shorea. The other dipterocarp emergents are commonly<br />

the legumes Dialium, Koompassia and S<strong>in</strong>dora (Ashton 1982).<br />

One <strong>of</strong> the characteristics <strong>of</strong> these forests is that they are considered to be stratified <strong>in</strong>to three<br />

or more vertical layers. However, this stratification is frequently not readily apparent, as<br />

different growth stages <strong>of</strong> plants tend to confuse the simple classical view <strong>of</strong> stratification.<br />

These forests also have flowers and fruits that are commonly borne on their trunks – referred<br />

to as cauliflory- the extent <strong>of</strong> which dim<strong>in</strong>ish with their altitud<strong>in</strong>al occurrence.<br />

Lowland Ra<strong>in</strong>forests also have dense tree crown epiphytes, <strong>in</strong>clud<strong>in</strong>g many ferns and orchids<br />

<strong>in</strong> the upper strata as well as abundant large woody climbers and bole climbers. Many <strong>of</strong> the<br />

enormous trees <strong>in</strong> this forest have very shallow root systems designed to utilize the soil<br />

nutrients limited to the surface soil result<strong>in</strong>g from the decomposition <strong>of</strong> leaf litter. Because<br />

many <strong>of</strong> these trees generally lack stabiliz<strong>in</strong>g taproots, they have evolved huge buttress roots<br />

to support their tall straight trunks. The ground layers are frequently sparse because the<br />

cont<strong>in</strong>uous upper canopy layer allows little light through.<br />

Evidence suggests that if naturally disturbed, through formation <strong>of</strong> tree-fall gaps, these<br />

Lowland Ra<strong>in</strong>forests take 120-140 years to regenerate to their orig<strong>in</strong>al mature structure,<br />

although floristically they may conta<strong>in</strong> the dom<strong>in</strong>ant trees after 60 - 70 years (Riswan et al.<br />

1985). MacK<strong>in</strong>non et al. (1996) estimate that it may take 200 years for a logged-over<br />

lowland forest to completely regenerate to its pre-logg<strong>in</strong>g structure and composition.<br />

The predom<strong>in</strong>ant ecological formation type, or matrix formation, <strong>in</strong> the <strong>East</strong> <strong>Kalimantan</strong><br />

assessment area is Lowland Ra<strong>in</strong>forest. It blankets most <strong>of</strong> the lowlands up to an altitude <strong>of</strong><br />

1,000m. It is far from a homogeneous formation: its topography ranges from<br />

alluvial/colluvial pla<strong>in</strong>s, both flat, gently slop<strong>in</strong>g, and gently undulat<strong>in</strong>g, to hilly areas,<br />

usually lower than 300 m a.s.l., and foothills. The formation occurs on volcanic, sedimentary<br />

and sedimentary substrates, and has a variety <strong>of</strong> soil types. This soil variety is represented <strong>in</strong><br />

the land system types that the ECA team used at a later stage <strong>in</strong> this process to subdivide the<br />

Lowland Ra<strong>in</strong>forest ecological system <strong>in</strong>to discrete occurrences.<br />

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