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- 14 ­<br />

into eight categories: mangrove swamps and associated wetlands; deciduous<br />

forests; evergreen forests; high humid tropical forests; coniferous forests;<br />

low lying subtropical and tropical forests; savannahs; and coral reefs (Fig.<br />

4-1). Active volcanic areas could also be considered a specific ecosystem<br />

type, and many integrated ecotones exist depending on soil type,<br />

precipitation, fire history, elevation and human activity. The length of the<br />

rainy season has a major influence on the types and distributions of<br />

ecosystems present throughout the isthmus. The complexity of CA ecosystems is<br />

enormous and perhaps best exemplified by Costa Rican natural history (Janzen,<br />

1983). The Tropical Science Center in San Jose, Costa Rica, for example,<br />

recognized for its work carrying out national and international ecological<br />

classification, lists 22 Roldridge life zones in Costa Rica (Fig. 4-1) from a<br />

"reduced version" map which includes only forest types.<br />

4.2.2 Biological Diversity<br />

The biotic history and paleogeography of the CA dispersal<br />

route for flora and fauna between North and South America has formed one of<br />

the most diverse regions on Earth. Even an abbreviated description of each of<br />

the major life zone of the isthmus would be outside the scope of requirements<br />

of this Environmental Assessment. In general, however, and in spite of its<br />

relatively small size, the natural environment of Central America can be<br />

characterized as an admixture of steep mountains, tropical rain forests,<br />

deserts, inland wetlands, highly variable coastal ecosystems on both the<br />

Atlantic Ocean and Caribbean Sea, significant mangrove ecosystems, and the<br />

second largest coral reef in the world. The region is also characterized by<br />

frequently recurring natural disasters, including earthquakes, volcanism,<br />

hurricanes and flooding, which result in a unique array of seral stages.<br />

While this complicates the classification of regional biotypes, it also--with<br />

the overlay of ancient and new agricultural activities--results in some of the<br />

very richest and biologically diverse ecosystems for its area found anywhere<br />

in the world.<br />

Two ecosystems are of special Interest<br />

projects<br />

with respect<br />

in Central<br />

to development<br />

America: (1) mangrove swamps and associated<br />

(2) forests.<br />

wetlands<br />

About<br />

and<br />

8980 square kilometers of mangrove<br />

wetlands<br />

swamps<br />

were present<br />

and associated<br />

in Central America in 1983 (Leonard<br />

are<br />

1986).<br />

not necessarily<br />

Mangroves<br />

diverse in terms of vegetation, but they:<br />

very<br />

(1)<br />

high<br />

tend<br />

primary<br />

to have<br />

and secondary productivity; (2) are<br />

cycles<br />

critical<br />

of a range<br />

to the<br />

of<br />

life<br />

species, serving as spawning grounds<br />

estuarine<br />

for many<br />

organisms;<br />

marine and<br />

(3) are important habitats for<br />

robins,<br />

migrating<br />

gulls,<br />

species<br />

terns,<br />

such<br />

swallows,<br />

as<br />

warblers, flycatchers, and<br />

as sea<br />

such<br />

turtles:<br />

marine species<br />

(4) are important ecosystems for nutrient<br />

highly complex<br />

cycling;<br />

in<br />

(5)<br />

terms<br />

are<br />

of their structure and function;<br />

buffers<br />

and<br />

against<br />

(6) are<br />

erosional<br />

important<br />

forces (e.g., hurricanes) along<br />

Mangrove<br />

some coasts<br />

systems<br />

in<br />

are<br />

CA.<br />

currently under pressure in much of the<br />

and<br />

region--directly<br />

indirectly--from agricultural use, fish farming,<br />

development,<br />

wood cutting,<br />

and<br />

"urban"<br />

a range of Industrial operations. While<br />

are<br />

many<br />

said<br />

mangrove<br />

to be substantially<br />

sites<br />

depleted, however, significant locations are<br />

believed to remain virtually untouched.

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