The ecology of eelgrass meadows in the Pacific Northwest: A ...
The ecology of eelgrass meadows in the Pacific Northwest: A ...
The ecology of eelgrass meadows in the Pacific Northwest: A ...
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<strong>the</strong> area dur<strong>in</strong>g a period <strong>of</strong> ris<strong>in</strong>g sea<br />
level.<br />
North <strong>of</strong> <strong>the</strong> Salmon River all rock<br />
formations are Tertiary. Eocene<br />
formations are widespread and <strong>in</strong>clude<br />
volcanic (basalts with tuffs and breccias)<br />
and sedimentary rocks (siltstones and<br />
sandstones) (proctor et al. 1980b). In <strong>the</strong><br />
01 igocene 1 irnited sedimentary formations<br />
developed (siltstones, shales,<br />
sandstones). In <strong>the</strong> Miocene extensive<br />
basalt flows occurred <strong>in</strong> nor<strong>the</strong>rn Oregon<br />
and southwestern Wash<strong>in</strong>gton. Dur<strong>in</strong>g <strong>the</strong><br />
Pleistocene, extensive erosion occurred<br />
dur<strong>in</strong>g coastal uplift<strong>in</strong>g.<br />
It appears that <strong>the</strong> late Pliocene and<br />
Pleistocene were characterized by<br />
alternat<strong>in</strong>g coastal submergence and<br />
uplift, with result<strong>in</strong>g deposition and<br />
erosion, respectively. <strong>The</strong> Puget Sound<br />
Bas<strong>in</strong>-Willamette Trough was formed <strong>in</strong> <strong>the</strong><br />
late pliocene (ca. 3 million years ago)<br />
when a general north-south uplift, form<strong>in</strong>g<br />
<strong>the</strong> Olympic and Cascade Mounta<strong>in</strong>s, was<br />
accompanied by a downwarp between <strong>the</strong> two<br />
ranges (Yosh<strong>in</strong>aka and Ellifrit 1974).<br />
Present coastal con£ igurations and<br />
sediments <strong>in</strong> estuaries to <strong>the</strong> north are<br />
<strong>the</strong> result <strong>of</strong> <strong>the</strong> melt<strong>in</strong>g <strong>of</strong> lobes <strong>of</strong> <strong>the</strong><br />
Vashon glacier <strong>of</strong> <strong>the</strong> Wiscons<strong>in</strong> period<br />
(17,000 to 9,01i70 years ago) (Wiedemann et<br />
al. 1974; Yosh<strong>in</strong>aka and Ellifrit 1974;<br />
Jefferson 1975). As <strong>the</strong> glaciers melted,<br />
erosion occurred with bay deposits form<strong>in</strong>g<br />
beds, fill<strong>in</strong>g <strong>in</strong> all river mouths<br />
(Jefferson 19751.<br />
Coastal submergence<br />
occurred aga<strong>in</strong>, creat<strong>in</strong>g <strong>the</strong> general<br />
features <strong>of</strong> <strong>the</strong> present coastl<strong>in</strong>e. Even<br />
though <strong>the</strong> glaciers did not extend south<br />
<strong>of</strong> <strong>the</strong> olympic Mounta<strong>in</strong>s (~umler 1969),<br />
<strong>the</strong> glacial melt<strong>in</strong>g did result <strong>in</strong> a<br />
general rise <strong>in</strong> sea level 120 to 150 m<br />
(400-500 ft) above its recent level<br />
(Jefferson 1975).<br />
1.2 CLIMATE OF REGION<br />
<strong>The</strong> climate <strong>of</strong> <strong>the</strong> <strong>Pacific</strong> <strong>Northwest</strong><br />
coastal region is mar<strong>in</strong>e (Proctor et al.<br />
1980b). ~t is characterized as a midlatitude,<br />
west-~oast-m~ he type, with wet<br />
w<strong>in</strong>ters and dry summers. <strong>The</strong> region is<br />
located <strong>in</strong> <strong>the</strong> Center <strong>of</strong> <strong>the</strong> prevail<strong>in</strong>g<br />
westerlies, with local w<strong>in</strong>ds rang<strong>in</strong>g from<br />
northwest to southwest througlmut most <strong>of</strong><br />
<strong>the</strong> year. <strong>The</strong> w<strong>in</strong>ds exchange heat with<br />
<strong>the</strong> ocean and become nearly saturated with<br />
moisture from evaporation. <strong>The</strong> air rises<br />
as it reaches land, cools, and gives up<br />
considerable moisture. Thus <strong>the</strong> climate<br />
is moderated by <strong>the</strong> <strong>Pacific</strong> ocean and<br />
characterized by high ra<strong>in</strong>fall and a<br />
moderate temperature range. <strong>The</strong>re is<br />
little north-to-south variation <strong>in</strong><br />
temperature and only a slight <strong>in</strong>crease <strong>in</strong><br />
precipitation to <strong>the</strong> north <strong>in</strong> <strong>the</strong> region.<br />
Two semipermanent pressure patterns, <strong>the</strong><br />
Aleutian Low and <strong>the</strong> North <strong>Pacific</strong> high,<br />
control <strong>the</strong> climate. <strong>The</strong> Aleutian Low,<br />
consist<strong>in</strong>g <strong>of</strong> a series <strong>of</strong> low pressure<br />
centers, lies over southwest Alaska and<br />
<strong>the</strong> Aleutian Islands. W<strong>in</strong>ds blow<br />
counterclockwise around <strong>the</strong> low, br<strong>in</strong>g<strong>in</strong>g<br />
a series <strong>of</strong> wet onshore frontal storms<br />
which are <strong>of</strong>ten characterized by strong<br />
w<strong>in</strong>ds. This pressure system dom<strong>in</strong>ates <strong>the</strong><br />
w<strong>in</strong>ter wea<strong>the</strong>r from Octcber to March. <strong>The</strong><br />
low shifts southward dur<strong>in</strong>g w<strong>in</strong>ter,<br />
result<strong>in</strong>g <strong>in</strong> an <strong>in</strong>creased frequency <strong>of</strong><br />
frontal storms reach<strong>in</strong>g <strong>the</strong> coast. Eighty<br />
percent <strong>of</strong> <strong>the</strong> annual precipitation falls<br />
dur<strong>in</strong>g this period able 1).<br />
<strong>The</strong> North <strong>Pacific</strong> High dom<strong>in</strong>ates <strong>the</strong><br />
summer wea<strong>the</strong>r <strong>of</strong> <strong>the</strong> region. As summer<br />
approaches, <strong>the</strong> high expands over <strong>the</strong><br />
eastern <strong>Pacific</strong> north <strong>of</strong> 2g0 N almost to<br />
Alaska and westward to 16g0 E. Storms are<br />
veered northward from <strong>the</strong> region by <strong>the</strong><br />
high. Only 5% <strong>of</strong> <strong>the</strong> annual precipitation<br />
falls <strong>in</strong> July and August. W<strong>in</strong>ds flow<br />
clockwise and br<strong>in</strong>g predom<strong>in</strong>antly fair<br />
wea<strong>the</strong>r. Dur<strong>in</strong>g this time coastal w<strong>in</strong>ds<br />
are from <strong>the</strong> northwest and north.<br />
Seasonal shifts <strong>in</strong> w<strong>in</strong>d direction<br />
<strong>in</strong>fluence nearshore ocean currents,<br />
result<strong>in</strong>g <strong>in</strong> upwell<strong>in</strong>g which br<strong>in</strong>gs cooler<br />
water to <strong>the</strong> surface near slmre <strong>in</strong> summer.<br />
'This produces fog, reduced <strong>in</strong>solation, and<br />
lclwer air temperatures.<br />
Precipitation <strong>in</strong> w<strong>in</strong>ter varies from a high<br />
<strong>of</strong> 32 cm (13 <strong>in</strong>ches) at Tatoosh Island <strong>in</strong><br />
northwestern Wash<strong>in</strong>gton to 27 cm (11<br />
<strong>in</strong>ches) at North Bend <strong>in</strong> southwestern<br />
Oregon to 18 cm (7 <strong>in</strong>ches) at Eureka <strong>in</strong><br />
nor<strong>the</strong>rn California, all <strong>in</strong> December<br />
(Proctor et al. 1980b). In summer,<br />
precipitation varies from a low <strong>of</strong> 5 cm (2<br />
<strong>in</strong>ches) <strong>in</strong> August at Tatoosh Island, to 2<br />
cm (0.8 <strong>in</strong>ch) <strong>in</strong> July at North Bend, to 0<br />
at Eureka <strong>in</strong> July.