322 Couquiaudenvironment does not allow the creation <strong>of</strong> a sem<strong>in</strong>aturalhabitat.Artificial facilities are more costly to buildand ma<strong>in</strong>ta<strong>in</strong> than natural enclosures due to thetype <strong>of</strong> construction and the operation <strong>of</strong> the lifesupport system. Yet, they allow for greater control<strong>of</strong> the environment and, possibly, better protectionaga<strong>in</strong>st hazards than natural facilities.Recent artificial environments <strong>in</strong>corporatenaturalistic features and elements. The design <strong>of</strong>coves and islands, the <strong>in</strong>corporation <strong>of</strong> naturalboulders or artificial reefs, topography that simulatesa rocky shore or beach, and wave generatorsthat recreate an ocean swell are all elements thatcan br<strong>in</strong>g a certa<strong>in</strong> degree <strong>of</strong> enrichment to theanimal environment accord<strong>in</strong>g to surveyed facilities.These features <strong>of</strong>ten come with greater space,and multiple and free-shape pools that allow forFigure 4.7. Educational programme <strong>in</strong> an artificial poolFigure 4.8. Feed<strong>in</strong>g and pett<strong>in</strong>g <strong>in</strong> an artificial pool(Photograph from M. L<strong>in</strong>et-Frion)escape from other animals, rest, and diversity<strong>of</strong> swimm<strong>in</strong>g pattern. Although these elements<strong>in</strong>crease the cost <strong>of</strong> construction and ma<strong>in</strong>tenance,they may prove beneficial <strong>in</strong> the long termby reduc<strong>in</strong>g behavioural problems related to poorenvironmental design.Figure 4.5. Show <strong>in</strong> an artificial poolFigure 4.6. Show <strong>in</strong> an artificial poolFunctions <strong>of</strong> PoolsDue to the various needs associated with publicdisplay, research, tra<strong>in</strong><strong>in</strong>g, medical care, breed<strong>in</strong>g,and, most importantly, the animals’ social needs,s<strong>in</strong>gle-pool facilities have evolved <strong>in</strong>to multi-poolcomplexes. Each pool has a ma<strong>in</strong> function, but theyeach can be used for other purposes as well. Names<strong>of</strong> pools may vary among facilities. Accord<strong>in</strong>g toUSA proposed regulations, the denom<strong>in</strong>ation <strong>of</strong>areas <strong>in</strong> SWTD facilities varies from those <strong>of</strong> regulardisplay facilities (Anonymous, 1979-1984,1995; 1998a; 1998b; 1999). The term “pool” isused <strong>in</strong> the follow<strong>in</strong>g section regardless <strong>of</strong> theenvironment, but it is applicable to enclosures andlagoons as well.In practice, pool functions are by no meansexclusively def<strong>in</strong>ed. Many facilities use poolsaccord<strong>in</strong>g to circumstances. In multi-pool facilities,if there is no maternity pool, a pregnant or
4. Types and Functions <strong>of</strong> Pools and Enclosures 323nurs<strong>in</strong>g female is <strong>of</strong>ten put <strong>in</strong>to a hold<strong>in</strong>g poolnot designed for this purpose. If temperamental<strong>in</strong>compatibility occurs between animals, theysometimes need to be separated. When severalspecies are kept together, but <strong>in</strong>ter-species breed<strong>in</strong>gis not favoured, males and females also needto be separated. Therefore, a facility may veryquickly face space limitations. The plann<strong>in</strong>g <strong>of</strong>a new facility too <strong>of</strong>ten responds to immediateneeds without foresee<strong>in</strong>g further developments<strong>in</strong> the size <strong>of</strong> the colony or its structure, or themany events <strong>in</strong> the cetaceans’ lives. It is thereforeimportant from the plann<strong>in</strong>g stage on to anticipatethose needs by provid<strong>in</strong>g extra space and extrapools, and by design<strong>in</strong>g habitats that ensure thatthe animals’ behavioural requirements are met.Ma<strong>in</strong> PoolThe ma<strong>in</strong> pool, also called the show pool <strong>in</strong> displayfacilities, is generally the largest pool, whichis used for display purposes and <strong>of</strong>ten is accessibleto public view<strong>in</strong>g. Animals usually haveaccess to this pool at all times. In the daytime,they may be kept temporarily <strong>in</strong> another poolbefore shows or public sessions. All pools whereanimals are housed permanently, and a fortior<strong>in</strong>ot given access to other pools, should followregulation requirements for the ma<strong>in</strong> or permanent,pool. Bassos & Wells (1996), as well as thesurveyed facilities, recommend, however, that allanimals have access to at least two pools to caterfor escape and varied behaviours.The dimensions and shape <strong>of</strong> the ma<strong>in</strong> pool willbe designed to allow the animals to display all species-specificbehaviours, <strong>in</strong>clud<strong>in</strong>g fast swimm<strong>in</strong>g,breach<strong>in</strong>g, and unison swimm<strong>in</strong>g. It is importantto provide ample space to allow for the development<strong>of</strong> social hierarchy and to provide adequatedistance between <strong>in</strong>dividuals <strong>in</strong> the case <strong>of</strong> multispeciesexhibits.Cetacean species are known to have very differentdiv<strong>in</strong>g and forag<strong>in</strong>g behaviours. It is thereforeencouraged that enough space be provided forspecies known to be pelagic, and thus deep divers,and for those that exhibit gregarious behaviour.Hold<strong>in</strong>g PoolThe pool used for temporary hous<strong>in</strong>g or for tra<strong>in</strong><strong>in</strong>gpurposes is called the hold<strong>in</strong>g pool or pen <strong>in</strong>natural environments. It is also known as the secondarypool or tra<strong>in</strong><strong>in</strong>g pool. Hold<strong>in</strong>g pools are asimportant <strong>in</strong> controlled environments as the ma<strong>in</strong>pool. They are used for tra<strong>in</strong><strong>in</strong>g, temporary hous<strong>in</strong>g,and separation. This pool is usually smallerthan the ma<strong>in</strong> pool.Hold<strong>in</strong>g pools can be used to house animalspermanently only if their dimensions are similarto those <strong>of</strong> the ma<strong>in</strong> pool. Subdivid<strong>in</strong>g withpartitions to allow a larger number <strong>of</strong> animalsor different species to be ma<strong>in</strong>ta<strong>in</strong>ed separatelyshould be avoided because the dimension requirementswill no longer apply. If some <strong>in</strong>dividualsneed to be separated permanently from the rest <strong>of</strong>the group because <strong>of</strong> behavioural problems, poolswith space requirements similar to the ma<strong>in</strong> poolwill be provided.Many surveyed facilities recommend that am<strong>in</strong>imum <strong>of</strong> two hold<strong>in</strong>g pools be connected tothe ma<strong>in</strong> pool to allow for multiple functions.They also advise that a maternity pool be <strong>in</strong>corporatedif breed<strong>in</strong>g is encouraged. If this pool isnot <strong>in</strong>corporated <strong>in</strong> the design, one <strong>of</strong> the hold<strong>in</strong>gpools will need to be used for this purpose,thereby reduc<strong>in</strong>g the overall space available toother animals.Medical PoolThe medical or quarant<strong>in</strong>e pool, also called the husbandryor isolation pool, is a small pool designedfor handl<strong>in</strong>g and isolat<strong>in</strong>g animals requir<strong>in</strong>g medicaltreatment (Geraci, 1986). It is preferably connectedto the other pools, with the possibility forisolation with watertight gates. In artificial facilities,a medical pool preferably will have a separatewater system that can be isolated from the ma<strong>in</strong>one to prevent contam<strong>in</strong>ation <strong>of</strong> other pools andso that it can be dra<strong>in</strong>ed and refilled very quickly.It will ideally provide options for both salt- andfreshwater. Dehydrated animals may be successfullyrehydrated by be<strong>in</strong>g kept <strong>in</strong> freshwater forlimited time (Sweeney & Samansky, 1995). Foranimals be<strong>in</strong>g treated for contagious diseases thatrequire isolation, or for stranded animals, it ispreferable to use a pool physically separated fromthe pools hous<strong>in</strong>g resident animals. In naturalenvironments where water control is not possible,a similar option can be considered.The medical pool can be located between thema<strong>in</strong> pool and one <strong>of</strong> the hold<strong>in</strong>g pools or betweentwo hold<strong>in</strong>g pools to facilitate the transfer <strong>of</strong> ananimal. (For the design, equipment, and special features<strong>of</strong> medical pools, please see Chapter 5.) If themedical pool also is used to treat p<strong>in</strong>nipeds, appropriatedra<strong>in</strong>age and fenc<strong>in</strong>g will be provided.Maternity PoolThe maternity or nursery pool is an alternativetype <strong>of</strong> hold<strong>in</strong>g pool dedicated to pregnant andparturient females, and to nurs<strong>in</strong>g females withtheir calves.To promote successful breed<strong>in</strong>g and rear<strong>in</strong>g <strong>of</strong>healthy calves, a facility can have a pool specificallydesigned for this function. Many problemshave occurred <strong>in</strong> the past due to unsuitable pooldesign (Amund<strong>in</strong>, 1986; survey). Pools with arestricted surface area or too shallow depth may
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8. Husbandry 373Figure 8.3. Milk sa
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8. Husbandry 375reintroducing a new
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8. Husbandry 377Rescue and Rehabili
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8. Husbandry 379Appendix II include
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8. Husbandry 381& R. J. Harrison (E
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Appendix 383Dolphinarium YaltaDolph
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Appendix 385Aomori Prefectural Asam
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Subscription Rates forAquatic Mamma