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94 ON THE SEXUAL OKGANS OF THE CYCADACE^.
tissue is slightly different, but there is no definite membrane admitting
' of separation. I cannotcl early comprehend what Gottsche means by the
yellow membrane which, according to him, covers the endosperm (Bot.
Zcit. 1. c. p. 398), unless he has m view the remains of the nucleus.
It is not unusual to see germination commence iu the undetached
seed itself; the radicle pierces the remains of the summit of the nucleus
and appears externally (Plate XCII. fig. 11). We must interpret in
this way the figure of the embryo of Maa-ozomia which I published in
1845 (Ann. des Sc. Nat. 1. c). In this instance the embryonic con-
dition had been passed and the first leaves developed.
It must be remarked that in all Cycads the plumule is only composed
of a few scales. On this and other points I may refer to my
communications in the Ann. des Sc. Nat. 1. c, and in ' Linnaea ' xix.
p. 5. The parts which produce the pollen,* and which compose
the male cone, are in every point the morphological equivalents of the
carpophylls. It would be perfectly correct to call them anthers, but
from their analogy to the female organs, and to avoid the risk of con-
founding them with their loculi (which many authors continue very
improperly to call anthers), it will be perhaps preferable to adopt the
terra androphylh. They are always smaller and simpler in shape than
the carpophylls, !)ut their structure presents no essential difference.
Tlie polygonal cells of the epidermis are much thickened, and the sto-
mata are deeply situated among them.
The principal point which arises here relates to the development of
the numerous polliniferous loculi (' lotjettes ') which are situated on
the under surface of the androphylls, and which were formerly looked
upon as distinct anthers.f Mohl has observed with reason that the
way in which I had explained in my monograph the evolution and
morjihological meaning of these loculi was not consistent (' A'^ermischte
Schriften,' p. 57).
They are placed on the lower face of the androphyll (which is also
their situation in the peltate forms) and on either side of the median
line, and they spring in groups of two to four, or rarely of five from a
* The numerous figures wliic-li exist of these ovgans may be consulted.
t Tills was the opinion of Kichai'd (Diet. Class, d'ilist. Nat. tome v. p.
21G). Other views as to their nature are discussed by R. Brown in the Appendix
to Captain Kiug's Voyage (Miscellaneous Bot. Works, vol. i. p.
459, et seq). Lindley (Veget. Kiugd. p. 223), adopting Richard's view, terms
tiie androphylls " antheriferous cone scales."— W. T. D.